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https://archive.org/detaiis/annaismagazineof1919unse
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY,
INCLUDING
ZOOLOGY, BOTANY, and GEOLOGY.
(being a continuation op the ‘annals’ combined with LOUDON AND
CHARLESWORTII’S ‘ MAGAZINE OP NATURAL HISTORY.’)
CONDUCTED BY
ALBERT C. L. G. GUNTHER, M.A., M.D., Ph.D., F.R.S.,
WILLIAM CARRUTHERS, F.R.S., F.L.S., F.G.S.,
AND
WILLIAM FRANCIS, F.L.S.
VOL. XIX.— SEVENTH SERIES.
LONDON:
PRINTED AND PUBLISHED BY TAYLOR AND FRANCIS.
SOLD BY SIMPKIN, MARSHALL, HAMILTON, KENT, AND CO., LD. ;
BAILLIERE, PARIS: HODGES, FIGGIS, AND CO., DUBLIN :
AND ASHER, BERLIN.
1907.
r
“Omnes res creatso sunt divinaj sapientiaj et potentise testes, divitiaj felicitatis
hurnanae: — ex harum usu honitas Creatoris; ex pulchritudino sapientia Domini ;
ex oeconomia in conservatione, proportione, renovatione, potentia majestatis
elucet. Eanim itaque indagatio ab hominibus sibi relictis semper acstimata ;
h vero eruditis et sapientibus semper oxculta ; male doctis et barbaris semper
inimica fuit.” — Linnaeus.
“ Quel que soit le principe de la vie animale, il ne faut qu’ouvrir les yeux pour
voir qu’elle est le chef-d’eeuvre de la Toute-puissance, et le but auquel se rappor-
tent toutes ses operations.” — Buuckner, ThSorie du 8ysteme Animal, Leyden,
1767.
The sylvan powers
Obey our summons ; from their deepest dells
The Dryads come, and throw their garlands wild
And odorous branches at our feet; the Nymphs
That press with nimble step the mountain-thyme
And purple heath-flower come not empty-handed.
But scatter round ten thousand forms minute
Of velvet moss or lichen, torn from rock
Or rifted oak or cavern deep : the Naiads too
Quit their loved native stream, from whose smooth face
They crop the lily, and each sedge and rush
That drinks the rippling tide : the frozen poles.
Where peril waits the bold adventurer’s tread.
The burning sands of Borneo and Cayenne,
All, all to us unlock their secret stores
And pay their cheerful tribute.
J. Taylor, Norwich, 1818,
aces ubrary
CONTENTS OF VOL. XIX.
[SEVENTH SERIES,]
NUMBER CIX.
Page
I. Descriptions of new Pyralidce of the Subfamilies Hydrocampince
and Scopariancs. By Sir George E. Hampson, Bart., B.A.,
’F.Z.S., &c
1
II. On some new Species of Blattidce in the Oxford and Paris
Museums. By R. Shelford, M.A., E.L.S 25
III. New Eastern, Australian, and African Heterocera. By
Colonel C. Swinhoe, M.A., F.L.S., &c 49
IV. Descriptions of Six new Species of Coleoptera from New
Zealand. By Major T. Broun, F.E.S 66
V. Descriptions of Three new Fishes from Yunnan, collected by
Mr. J. Graham. By C. Tate Regan, B.A 63
VI. Diagnoses of new Central- American Freshwater Fishes of
the Families CyprmodontidcB and Mugilidcs. By C. Tate Regan,
B.A 64
VII. A Contribution towards a Knowledge of the Entozoa of
British Marine Fishes.— Part I. By William Nicoll, M.A.,
B.Sc., Gatty Marine Laboratory, St. Andrews. (Plates I.-IV.) . . 66
VIII. Descriptions of Fifteen Terrestrial Mollusca from South
Africa. By James Cosmo Melvill, M.A., F.L.S., and John
Henry PoNSONBY, F.Z.S. (Plate VI.) 94
IX. On Three Mollusk-infesting Trematodes. By Marie V.
IjEBOUR, B.Sc., Armstrong College, Newcastle-upon-Tyne. (Plates
Vlt. & VHI.) 102
0
103468
IV
CONTENTS.
Page
X. Preliniinary Diagnoses of Six new Mysidoi from tlie West
Coast of Ireland. Dy W. M. Tattersall, H.Sc., Department of
Agriculture and Technical Instruction, Fisheries Branch, Dublin . . 106
XI. On further new Mammals obtained by the Ruwenzori Expe-
dition. By Oldfield Thomas 118
XII. Note on Ochotona {Conothoa) aurita, Blanf., from Ladak.
By J. Lewis Boniiote, M.A 124
XIII. On Two new Parasitic Coleoptera (Fam. Staphylinidce)
from South America. By Gilbert J. Arrow, F.E.S 125
New Books : — Catalogue of the Indian Decapod Crustacea in the
Collection of the Indian Museum. Part III. Macrura.
Fasciculus 1. The Prawns of the Peneus Group. By A.
Alcock, M.B., LL.D., C.I.E., F.R.S. — A Treatise on Zoology.
Edited by E. Ray Lankester, M.A., LL.D., F.R.S. Part V.
Mollusca. By Paul Pelseneer, D.Sc 127
Note of Correction : Parorchis, n. nom., for Zeugorchis, Nicoll, 1906,
by Win. Nicoll, M.A., I3.Sc., Gatty Marine Laboratory, St.
Andrews 128
4
NUMBER CX.
XIV. On the Bats of the Family Meg ader mat idee. By Knud
Andersen and R. C. Wroughton 129
XV. Note on the Crab Hymenosoma depressum, Jacquinot and
Lucas. By Charles Chilton, M.A., D.Sc., F.L.S., Professor of
Biology, Canterbury College, New Zealand. (Plate V.) 146
XVI. Descriptions of new Species of Trochomorpha, Cochlostyla,
Ampliidromus, Bulimulus, Brymeeus, Placostylus, Stenogyra, Lepto-
poma, Cyelophornis, Cyclotus, and Alycceus. By Hugh C. Fulton.
(Plates IX. & X.) 149
XVII. Notes on Papuina and Pupina. By Hugh C. Fulton . . 158
XVIII. Description of a new Cyprinoid Fish, Acheilognathus
signifer, from Korea, with a Synopsis of all the known Rhodeince.
By L. S. Berg (St. Petersburg) 159
XIX. Description of a new Cyprinoid Fish, Paraleucogohio nota-
canthus, from N. China. By L. S. Berg (St. Petersburg) 163
XX. Description of a new Species of Eulota from Formosa. By
G. K. Gude, F.Z.S 164
XXI. On some undescribed Phytophagous and Parasitic Hynieno-
ptera from the Oriental Zoological Region. By P. Cameron .... 166
CONTENTS.
V
Page
XXII, On tho Black-and-tan Pattern of Domestic Dogs {Cams
familiaris). By li. I. Pococx, P.L.S., F.Z.S., Superintendent of
the Zoological Society’s Gardens 192
XXIII. Seminiila'. a Note by Arthur Vaughan, B.A., D.Sc.,
F.G.S ■ 194
Neio Book: — Illustrations of British Blood-sucking Flies, with
Notes by Ernest Edward Austen 199
Proceedings of the Geological Society 200
NUMBER CXI.
XXIV. New Species of Eastern Heterocera. By Colonel C.
SwiNHOE, M.A., F.L.S., &c 201
XXV. Notes on Scorpions, with Descriptions of Two new Species.
By A. S. Hirst 208
XXVI. Descriptions of Three new Lizards and a new Frog, dis-
covered by Dr. W. J. Ansorge in Angola. By G. A. Boulenger,
F.R.S 212
XXVII. Diagnoses of new Species of Corhula and Bithinella from
Lower Bengal. By H. B. Preston, F.Z.S 215
XXVIII. Descriptions of Nine new Species of Land-Shells from
New Caledonia. By II. B. Preston, F.Z.S 217
XXIX. Descriptions of new Genera and Species of Sijntomid<B,
Arctiadce, A qaristidce. and Noctuidce. By Sir George F. Hampson,
Bart., F.Z.S. 221
XXX. Descriptions of Six new Freshwater Fishes from Mexico
and Central America. By C. Tate Regan, B.A 258
XXXI. Descriptions of Two newCharacinid Fishes from Argentina.
By C. Tate Regan, B.A 261
XXXII. On some new Species of Chrysochloris. By R. Broom,
M.D., D Sc., C.M.Z.S 262
XXXIII. New Species of Sphegidce from Australia. By R. E.
Turner 268
NUMBER CXII.
XXXIV. Rhynchotal Notes. — XLI. By W. L. Distant 277
XXXV. On a Freshwater Decapod Crustacean collected by
W. J. Burchell at Para in 1829. By W. T. Calman, D.Sc.,
British Museum (Natural History) 295
VI
CONTENTS.
Pag®
XXX VI. Descriptions of new Species of Ileterocera belonging
to the Families SyntomidcB, Hypsid(X, CyllopodidcBf Dioptidce, and
Erateininm. By Herbert Druce, F.L.S. &c 299
XXXVII. On new Species of Histeridce and Notices of others.
By G. Lewis, F.L.S 311
XXXVlll. On a new Species of Karschia from Tibet. By A. S.
Hirst 322
XXXIX. Descriptions of Three new Snahes discovered by Mr. G.
L. Bates in South Cameroon. By G. A. Boui.enger, F.ll.S 324
XL. The Synonymy and Generic Position of certain Species of
Miiscidce {sens, lat.) in the Collection of the British Museum,
described by the late Francis Walker. By Ernest E. Austen . . 326
XLI. Some new Species and Genera of Lamellicorn Coleoptera
from the Indian Empire. By Gilbert J. Arrow 347
Neio Book : — Catalogue of the Lepidoptera Phalaenae in the British
Museum. Vol. VI. (Catalogue of Noctuidse, Vol. HI.) 360
NUMBER CXIH.
XLII. Descriptions and Records of Bees. — XIII. By T. D. A.
Cockerell, University of Colorado 361
XLHI. New Mammals from Lake Chad and the Congo, mostly
from the Collections made during the Alexander-Gosling Expe-
Charles Chilton, M.A., D.Sc., F.L.S., Professor of Biology,
Canterbury College, New Zealand. (Plate XI.) 388
XLV. On Barbus aureus, Cope, from Natal. By G. A.
Boulenger, F.R.S 390
XLVI. Description of a new Cyprinid Fish of the Genus Laheo
from the Transvaal. By G. A. Boulenger, F.R.S 392
XLVII. Descriptions of Two new African Lizards of the Genus
Latastia, By G. A. Boulenger, F.R.S ib.
XLVIII. Description of a new Frog of the Genus Telmatohius
from Brazil. By G. A. Boulenger, E.R.S 394
XLIX. Rhyuchotal Notes. — XLII. By W. L. Distant 395
L. Some new Species and Genera of Lamellicorn Coleoptera from
the Indian Empire. By Gilbert J. Arrow. — Part II 416
LI. On a remarkable Mountain Viscacha from Southern Pata-
nonia, with Diagnoses of other Members of the Group. By Oldfield
Thomas 439
CONTENTS.
Vll
I’ago
IjII, On tlic Occurrence of the Edible Dormouse (Genus Glis) in
Sardinia. By Oldfield Thomas 444
LIII. A new Genus and Species of Phlebotomic MuscidtE from
Aden. By Ernest E. Austen 446
New Boohs: — Fishes of Japan, an Account principally on Economic
Species. By Keinosuke Otaki, Tsunenobu Fujita, and
Tadashi Higurashi. — The British Woodlice, being a Mono-
graph of the Terrestrial Isopod Crustacea occurring in the British
Islands. By Wilfred Mark Webb and Charles Sillem.
448, 450
Proceedings of the Geological Society 450, 451
Modern Helminthological Nomenclature, by Dr. von Linstow .... 451
NUMBER CXIV.
LIV. On the Classification of the Decapod Crustaceans. By
L. A. Borradaile, M.A., Lecturer in Natural Sciences at Selwyn
College, Cambridge 457
LV. Descriptions of new Lizards in the British Museum. By
G. A. Boulenger, F.R.S 486
LVI. On the Variations of Stereolepis gigas, a great Sea-Perch from
California and Japan. By G. A. Boulenger, F.R.S 489
LVn. On an African Barbel hitherto confounded with Barhus
trimaculatus, Peters. By G. A. Boulenger, F.R.S 492
LVni. Spirochceta {Trypanosoma) Balhianii (Certes), its Move-
ments, Structure, and Affinities ; and on the Occurrence of Spiro-
chceta anodontoe (Keysselitz) in the British Mussel, Anodonta cygnea.
By H. B. Fantham, B.Sc., Derby Research Scholar, University
College, London ; and St. Mary’s Hospital Medical School 493
LIX. On Pterocyon, Rousettus. and Myonyctei'is. By Knud
Andersen 601
LX. Notes on the Quagga and Burchell’s Zebra in the Paris
Museum. By R. I. Pocock, Superintendent of the Zoological
Society’s Gardens London 616
LXI. New Mammals from Lake Chad and the Congo, mostly
from the Collections made during the Alexander-Gosling Expe-
dition. By Oldfield Thomas, F.R.S., and R. C. Wroughton . . 620
LXII. On Two Spiders of the Genus Selenocosmia. By A. S.
Hirst 622
Vlll
CONTENTS.
Page
LXIII, Brachiopod Nomenclature : the Genotype of Terehratula.
Jiy S. S. Buckman, F.G.S. (Plate XII.) 525
LXIV. Descriptions and Records of Bees. — XIV. By T. D. A.
OocKEiiELL, University of Colorado 531
New Books : — The Crawfishes of the State of Pennsylvania. By
Dr. A. E. Ortmann. — A Natural History of the British
Butterflies, their World-wide Variation and Geographical
Distribution. A Textbook for Students and Collectors. By
J. W. Tutt, F.E.S. Vol. 1 540, 542
Index 543
PLATES IN VOL. XIX.
Entozoa of British marine fishes.
Ilymenosoma depressum.
Mollusca from South Africa.
Mollusk-infesting Trematodes.
New species of shells.
Phreatogammarus propinqiuis, sp. u.
Terehratula terehratula (Limio).
THE ANNALS
AND
MAGAZINE OF NATGEAL HISTOEY.
[SEVENTH SERIES.]
“ per litora spargite museum,
Naiades, et circixm vitreos coi^idite fontes :
Pollice virgineo teneros hie calrpite flores :
Floribus et pictum, divse, replete canistrura.
At VOS, o Nymp^iae Orataridei, itebub IxLdasi,
Ite, recurvato variata corallia truneo
Vellite museosis ^ rnpibug, dt m>lii eonehas
Ferte, Deaj pelagi, et pingui eonehyiia sueeo.”
N.Parthenii Giannettasi,T^Q\. 1.
No. 109. JANUARY 1907.
I. — Descriptions of new Pyralidse of the Subfamilies Hydro-
campinse and Scoparianse. By Sir George F. Hampson,
Bart., B.A., F.Z.S., &c.
[Concluded from vol. xviii. p. 472.] '
Genus Pseudlithosia, nov.
Palpi upturned, short, not reaching vertex of head,
smoothly scaled, the third joint moderate ; maxillary palpi
minute, filiform ; frons rounded ; antennse simple in both
sexes ; legs long and slender, the spurs short ; retinaculum
of male a corneous bar. Fore wing very long and narrow ;
vein 3 from angle of cell ; 4, 5 from angle ; 6 from well
below upper angle ; 7 from angle ; 8, 9, 10 stalked from
long before angle. Hind wing with veins 3 and 5 separate,
4 absent ; 6 from upper angle ; 7 from well before upper
angle and anastomosing with 8.
(1.) Pseudtithosia Schausi, sp. n.
. Head and thorax black-brown ; abdomen fulvous-
brown. Fore wing black-brown. Hind wing pale brown.
Ann, (Sc Mag. N. Hist. Ser. 7. Vol. xix. 1
2
Sir G. F. IlaiYipson on new Pyralidse
? . Fore wing with diffused white scales forming traces of
antcmedial, medial, and subterminal bands.
Hah. Mexico, Jalisco, Guadalajara [Schaus), 1 $ type.
Exp., ^ 32, ? 40 mm. Type in Coll. Schaus.
(4.) Daulia argyrophoralis, sp. n.
Mid tibia of male without groove and tuft of hair ; fore
wing with vein 10 from cell or stalked with 8, 9.
Head and thorax ochreous tinged with brown ; fore legs
tinged with fuscous ; abdomen white slightly tinged with
ochreous. Fore wing ochreous yellow irrorated with a few
black scales, more thickly on disk ; a silvery fascia below
base of cell, then obliquely bent downwards to middle of
inner margin ; some silvery scales in middle and end of cell
and beyond discocellulars, on which there is a slight fulvous
lunule ; a silvery subtcrmihal slightly curved band defined
on each side by fine black lines; a narrow silvery terminal
band defined on inner side by a slight black line ; cilia
yellow at base, silvery white at tips. Hind wing uniform
ochreous white.
Hab. Argentina, Florenzia, Gran Chaco [S. R. Wagner),
2 c? ^ 1 $ type. Exp. 20 mm.
(4 a.) Luma flavimarginalis, sp. n.
Head and thorax orange-yellow ; palpi with the second
joint black above ; abdomen fuscous, the extremity and
ventral surface yellow. Fore wing fuscous brown with a
purplish gloss ; the base orange-yellow ; an orange-yellow
terminal baud expanding widely to costa, the outer edge of
brown area being strongly curved. Hind wing fuscous
brown with a slight purplish gloss ; an orange-yellow terminal
band expanding slightly to costa.
Hab. Ceylon, Maskeliya {Green, de Mowbray), 1 1 $
type. Exp., S' 28, $ 36 mm.
(6 a.) Luma holoxantha, sp. n.
$ . Head, thorax, and abdomen orange-yellow. Fore
wiDg uniform glossy yellow. Hind wing pale glossy yellow.
Hab. Mashon ALAND, Salisbury [Marshall), 1 ? type.
Exp. 30 mm.
(2.) Margarochroma fuscalis, sp. n.
Antennae annulate.
S . Black-brown ; pectus, legs, and ventral surface of
3
of the Subfamily Ilydrocam pinse.
alxlomeii whitish ; wings mixed with greyish. Fore wing
with a pale siibterminal line, oblique with an inward curve
from costa to vein 5, then minutely dentate and indistinct
to tornus. Hind wing with indistinct minutely dentate
subterminal line.
Hab. Celebes, Bonthain, Mdrulaman, 2300' {Everett).
Exp. 14 mm. Type in Coll. Rothschild.
(6 a.) Draccenura chrysochroa, sp. n.
S . Head, thorax, and abdomen orange-yellow ; palpi
white at base, black at tips ; lower part of frons blackish ;
pectus and ventral surface of abdomen white ; mid tibise
black at base. Fore wing orange-yellow, rather deepei
orange towards termen, the costa tinged with fuscous ; a
slight, dark, somewhat sinuous and oblique antemedial line ;
a black point in middle of cell and slender discoidal bar ; a
brown postmedial line, slightly bent outwards to costa, then
erect ; cilia black at base, grey at tips. Hind wing orange-
yellow, deeper orange towards termen; a slight obliquely
curved brown line from below middle of costa to above
tornus ; cilia black at base, white at tips.
Hab. S.W. New Guinea, Kapaur {Doherty) y 2 S type.
E.xp. 22 mm.
(6 b.) Bradina costalis, sp. n.
$ . White ; palpi brown at tips ; neck brown ; shoulders
with a brown stripe ; abdomen dorsally tinged with brown.
Fore wing with dark brown moderately broad costal fascia ;
both wings with diffused brown terminal band, narrowing to
a point at submedian fold of hind wing ; the cilia white.
Hab. Solomons, Guadalcanar. Exp. 26 mm. Type in
Coll. Rothschild.
(17 a.) Bradina glaucinalisy sp. n.
Head, thorax, and abdomen fuscous brown, the last with
white segmental rings ; pectus, legs, and ventral surface of
abdomen whitish. Fore wing pale glossy gf’ey-brown, with
traces of dark discoidal point and oblique postmedial line ; a
fine dark terminal line ; cilia broAvnish white, with dark line
at base. Hind wing uniform pale glossy grey-brown, with
fine dark terminal line ; cilia brown at base, whitish at tips.
Hab. New Guinea, Kapaur {Doherty), 1 c?, 1 ? type;
Solomons, Florida 1. {Meek), 1 ? . Exp. 22-30 mm.
) 1*
4
Sir G. F. Ilanipson on new Pyralldse
(18«.) Bradina dentalis, sp. n.
Pale silky brownish ochreous ; palpi and frons fuscous ;
anal tuft vvliite. Fore wing with the costal area suffused
with purplish fuscous ; an antemedial black line bent inwards
to costa ; a point in middle of cell and discoidal lunule ; a
minutely dentate postmcdial linc^ bent outwards below costa^
then oblique. Hind wing with oblique discoidal striga ; a
minutely dentate curved postmedial line.
Hab. Sum BA [Doherty) ; Selaru [Micholitz), 1 ^ type.
Exp. 22-24 mm.
(26 a.) Bradina pwniUalis, sp. n.
Ilead^ thorax, and abdomen grey tinged with brown and
irrorated with black ; palpi and legs banded with black ;
abdomen with dorsal black band before anal tuft. Fore
wing grey tinged with brown and irrorated with black ; the
costa blackish to beyond middle ; diffused black subbasal and
antemedial lines, the latter expanding into a patch below
cell ; a rather x -shaped black discoidal spot ; postmedial
line black, diffused, rather maculate, excurved between veins
5 and 3, then retracted to below end of cell ; a terminal
series of blackish striae. Hind wing grey suffused with
fuscous brown.
Hab. SuMBAWA [Doherty), 1 ^ type ; Pura [Doherty),
1 $ . Exp. 10 mm.
(27 a.) Bradina punctilinealis , sp. n.
(^ . Very pale yellow; abdomen dorsally tinged with
fuscous at middle. Fore wing with black point in middle
of cell and slight oblique discoidal lunule ; a curved post-
medial series of minute black streaks on the veins. Hind
wing with oblique postmedial series of minute dark streaks
on veins 6-2.
Hab. Fiji, Navua I. [de la Garde), 1 type. Exp.
22 mm.
\ ^(28 a.) Bradina neuralis, sp. n.
^ . Hel(id and thorax whitish tinged with brown ; palpi
black at tips ; antennae ringed with brown; fore tibiae and
tarsi banded with black ; abdomen white tinged with brown,
the terminal half dorsally banded with fuscous. Fore wing
white, the costal and apical areas suffused with cupreous
brown, the veins streaked with brown ; a dark spot on costa
of the Subfamily Ilydrocampina). 5
near base ; an elongate spot in middle of cell, n itb diffused
brown band from it to inner margin ; a dark discoidal spot ;
a dark postmedial line excurved between veins 5 and 2.
Hind wing white ; the veins stieaked with brown except at
base; a discoidal point; a postmedial line excurved between
veins 5 and 2 ; apical area suffused with brown, narrowing
to a point at vein 2.
Ilab. Samoa, Pago I. [de la Garde), 1 S type. Exp.
22 mm.
(30 a.) Bradina hemiphcealis , sp. n.
S . Head and thorax fuscous black ; pectus and abdomen
whitish suffused with fuscous brown. Fore wing deep
fuscous brown with slight dark irroration ; faint traces of
a darker discoidal spot and postmedial line excurved at
middle ; cilia rather paler fuscous brown. Hind wing white
slightly tinged with fuscous brown ; traces of a slightly
diffused postmedial line, excurved at median nervules, then
obsolete; a rather diffused fuscous terminal line from apex
to vein 2 ; cilia white, with faint brown line near base
towards apex ; the underside more strongly tinged with
fuscous brown, the costal area fuscous, the terminal area
suffused with fuscous.
Hah. Br. E. Africa, Kikuyu, Nairobi Plains [Crawskay) ,
1 S type. Exp. 30 mm.
(34.) Bradina purpurascens, sp. n.
$ . Deep purplish fuscous ; palpi below, pectus, and
ventral surface of abdomen white. Fore wing with oblique
antemedial black line, slightly angled on median nervure ;
a point in cell and discoidal lunule ; the postmedial line
slightly excurved below costa, oblique to vein 5, bent out-,
wards between veins 5 and 2, retracted to below end of cell,
and slightly excurved above inner margin, defined by white,
on outer side between costa and vein 5 ; the apical area
suffused with black. Hind wing with the postmedial line
bent outwards between veins 5 and 2, then retracted to
below end of cell and excurved above inner margin; cilia
white at tips. Underside purplish grey.
Hah. Jamaica. Exp. 20 mm. Type in Coll. Rothschild.
(4.) Coptobasis lopkocera, sp. n.
Antennae of male without tuft and excision at base, the
shaft w'ith large tuft of hair at three fourths from base ; fore
G
Sir G. F. Ilampson on new Pyrallcise
femora and mid tibiae with tufts of hair ; bind tibiae flat-
tened, greatly eurved and shortened, the tibiae, spurs, and
tarsi with immensely developed tufts of hair.
$ . Uniform very dark brown ; palpi at base below, pectus,
legs, and ventral suii'ace of abdomen whitish ; fore tibiae
with fuscous band.
Hab. S. Celebes {Doherty), 1 S type. Eorp. 36 mm.
(5.) Ccelorhjncidia jmrpurea, sp. n.
Hind tibiae of male with tufts of hair on inner side and a
thickly scaled process on outer side at extremity.
Black suffused with brilliant purple; palpi and legs at
base and underside of abdomen white. Fore wing with
indistinct curved antemedial black line ; both wings with
discoidal spot ; the postmedial line exeurved from costa to
vein 2, then retracted to lower angle of cell. Hind wing
with the cilia white. Underside with the discoidal spots and
postmedial line defined by whitish.
Hab. Amboina {Dohey'ty), 1 ^ typ®- Exp. 24 mm.
(6.) Coelorhyncidia nitidalis, sp. n.
Head, thorax, and abdomen glossy grey slightly tinged
with brown ; palpi blackish, white below ; pectus, legs, and
ventral surface of abdomen white; fore tibia at extremity
and terminal joints of tarsus black. Fore wing uniform
glossy grey slightly tinged with browm. Hind wing rather
paler glossy grey with a faint brown tinge; the underside
pure white.
Hab, New Guinea, Kapaur {Doherty), 1 $ , Milne Bay
{Meek)y 1 1 $ type. Exp., ^ 26, ? 30 mm.
(7.) Diathrausta cymialis, sp. n.
^ . Head and thorax cupreous brown ; palpi white below ;
pectus and legs whitish ; abdomen cupreous brown with
slight white segmental rings, the ventral surface white. Fore
wing cupreous brown irrorated with fuscous ; some white
scales below the cell near base ; antemedial line whitish
defined on each side by black scales, exeurved ; the medial
area somewhat whiter except costal area ; an obscure dark
spot at upper angle of cell; postmedial line white defined on
each side by black, oblique to vein 7, inw^ardly oblique to
vein 2, retracted to upper angle of cell, incurved and out-
wardly oblique to vein 1 and erect to inner margin ; a slight
white mark on costa towards apex ; termen blackish, with
some irregular white marks, the mark above vein 6 some-
7
of the Subfamily IIy(lrocampina3.
what dentate ; cilia cupreous at base, with black medial line
and whitish tips. Hind wing cupreous l)ro wn suffused in
parts with fuscous ; an indistinct blackish discoidal bar
defined on each side by whitish ; a whitish line from lower
angle oC cell to inner margin defined on each side by black;
a curved white postmedial line defined on each side by blaek ;
a white subterminal line defined on each side by black, ex-
curved from costa to discal fold, where it is interrupted, then
oblique, and ending on termen at vein 1 ; cilia cupreous at
base, with black medial line and whitish tips.
Hab. Brazil, Castro Parana (Jones), 1 S fype. Exp.
16 mm.
(2.) Deuteropliysa micralis, sp. n.
Antenme of male annulate ; fore wing with the apex not
produced and without fovea above vein 7.
Purplish grey; palpi fuscous, wdiite at base; legs
whitish ; wings glossy, whth very fine striae. Fore wing with
the first line almost medial, fine, brown, somewhat oblique
from eosta to submedian fold, where it is slightly bent
inwards; postmedial line brown, eurved ; cilia dark brown.
Hind wing with slight discoidal striga ; a faint line from
beyond lower angle of cell to inner margin ; an obliquely
eurved line from costa beyond middle to termen at sub-
median fold ; cilia reddish at base, followed by a black line
and white tips ; the underside whitish.
Hab. Jamaica, Runaway Bay [Walsingham), 1 type.
Exp. 12 mm.
Genus Lasiogyia, nov.
Palpi porrect, straight, the second joint about twice length
of head and fringed with hair above, the third about length
of head ; frons with pointed conical prominence ; antennae
minutely ciliated ; fore femora and tibiae of male fringed
with rough hair above, the first tarsal joint very long and
the joints fringed with very long hair on both sides; abdo-
men long, with slight lateral expansion towards extremity.
Fore wing with vein 3 from close to angle of cell ; 4, 5 from
angle ; 7 straight and well separated from 8, 9, 10. Hind
wing with the cell short ; vein 3 from close to angle ; 4, 5
somewhat approximated for a short distance ; 6, 7 from upper
angle, 7 anastomosing with 8.
(1.) Lasiogyia xanthozonata, sp. n.
S . Head and thorax orange-fulvous, with patches of pale
yellow on head, tegulse, patagia, and metathorax ; fore legs
8
Sir G. F. Ilampson on new Pyralulse
■with the tufts brown ; abdomen orange, 'v\dth pale yellow
dorsal band on first segment and segmental white lines on
the others. Fore wing fuscous, bceoming fulvous at margins ;
a yellow spot at base; a broad yellow medial band with
waved edges, contracting somewhat below the cell. Hind
wing fuscous, the inner area yellowish; a medial yellow
band, with two small teeth on its outer edge beyond lower
angle of cell ; cilia fulvous.
Hub. N. Guinea, Fergusson I. [Meek). Exp. 22 mm.
Type in Coll, llothschild.
(3 o.) Stenia desertalis, sp. n.
$ . Ilead and thorax ochreous tinged with brown ; palpi
rufous, wdiite below ; antennae ringed ~ brown and white ;
abdomen white. Fore wing ochreous white irrorated with
brown and faintly tinged with rufous towards costa ; a dark
diffused antemedial line, oblique from costa to below cell and
obsolescent towards costa; a slight dark discoidal lunnle; post-
medial line rather diffused, obliquely excurved from costa to
below angle of cell, then erect ; a slight dark terminal line ;
cilia white, with faint dark lines at middle and tips. Hind
wing white, wdth traces of postmedial line towards costa and
slight terminal line ; the underside with slight dark spot at
upper angle of cell and diffused postmedial line from costa
to vein 5, then slight and excurved to vein 2.
Hub. Cape Colony, Zuurberg [Bair stow), 1 ? type.
Exp. 24j mm.
[7 a.) Stenia costalis, sp. n.
JjlepharomastLv colubralis, Druce, Biol. Centr.-Am,, Het, ii. p. 2G8
(part.), iiec Guen.
Wliite ; palpi and vertex of head fuscous ; patagia with
fuscous spots ; abdomen dorsally suffused wdth fuscous
except towards base, leaving white segmental rings. Fore
wing with the costal area fuscous brown; a dark antemedial
line obsolescent towards inner margin ; a point in middle of
cell and discoidal lunule ; the postmedial line bent outw^ards
between veins 5 and 2, then retracted to below end of cell
and sinuous to inner margin ; a terminal series of dark points.
Hind wing with fuscous discoidal spot ; the postmedial line
bent outwards between veins 5 and 4, then retracted to
below end of cell and ending at tornus, obsolescent except
towards costa; a terminal series of points.
Hab. Guatemala, Vera Paz 2 S type; Costa
Bica, Irazu [Rogers), 2 S > Goduian-Salvin Coil. ; Colombia,
R. Dagua [Rosenberg) . Exp. 26 mm.
of the Subfamily Ilydrocamplnce.
(7 b.) Stenia semifascalis , sp. n.
"White ; liead, front of thorax, and abdomen dorsally
fuscons except at base. Fore wing with the costal half fuscous,
expanding on terminal area to tornus ; a dark antemedial
line, obsolete towards inner margin ; a point in cell and
discoidal lunule ; a line from vein 2 below end of cell to
inner margin; a nearly straight line from costa beyond
middle to tornus ; a fine pale line at base of cilia. Hind
wing with discoidal point ; a fine postmedial line bent out-
wards between veins 5 and 2, then retracted to below end of
cell and obsolescent ; the apex tinged with fuscous.
Hab. Ecuador, Cachabe [Rosenberg) , 1 1 ? ; Paramba
[Rosenberg) , 1 S type. Exp. 22-26 mm.
(7 c.) Stenia interruptalis , sp. n.
$ . Head, thorax, and abdomen white tinged with brown ;
palpi black above, white below. Fore wung white, the costal
area suffused wdth fuscous brown, at base extending to
median nervure ; the apical area broadly fuscous brown down
to vein 3, the white extending to termen between vein 3 and
submedian fold, the tornal area fuscous broAvn to near post-
medial line ; an indistinct erect brown antemedial line ; a
small brown annulus in middle of cell; a brown discoidal
lunule with white centre ; postmedial line slightly waved,
erect from costa to vein 5, bent outwards between veins 5
and 2, then retracted to lower angle of cell and erect to inner
margin ; a terminal series of blackish points ; cilia wEite,
tinged with fuscous brown at tips. Hind wing white ; a
slight discoidal dark lunule with white centre ; a slight dark
postmedial line, erect from costa to vein 5, excurved and
obsolescent between veins 5 and 2, then retracted to lower
angle of cell and oblique to tornus ; a large apical fuscous-
brown patch ; a slight dark mark on termen at submedian
fold.
Hab. Ecuador, Chimbo, 1 $ type. Exp. 20 mm.
(7 d.) Stenia fuscilunalis, sp. n.
Blepharomastiv colubralis, Druce, Biol. Centr.-Am., Het. ii. p. 268
(part.) nec Guen.
S . Head, thorax, and abdomen white suffused with
browm ; palpi black above, white below ; pectus, legs, and
ventral surface of abdomen white. Fore wing white ; the
costal area suffused with fuscous browu ; the apical area
Sir G. F. Hampson on new Pyralidse
suffused with fuscous brown to vein 4 ; a slight erect ante-
medial line; a dark point in middle of cell and small dark
discoidal lunule ; postmedial line minutely waved^ ereet from
costa to vein o, bent outwards between veins 5 and 2,
retracted and obsolescent to lower angle of cell, then erect
to inner margin ; a terminal series of dark points ; cilia
brownish white, with a dark line near base. Hind wing
white; a slight dark discoidal lunule, with an oblique line
from it to tornus ; postmedial line erect from costa to vein 5,
then bent outwards and oblique to termen at submedian
fold ; the apex slightly suffused with brown, a dark line on
termen, and slight brown line through cilia towards apex.
$ . Fore wing with the terminal points more distinct ;
hind wing with the apex not suffused with brown and with
punctiform terminal line from apex to vein 2.
Hab. Guatemala, Vera Paz {Champion'), 1 $ ; Costa
Rica, Irazu [Rogers), 1 S Cache [Rogers), 1
R. Sucio [Rogers), 1 Panama, Chiriqui [Champion),
2 $ . Godman-Salvin Coll. Exp., 24, $ 20 mm.
(7 e.) Stenia irroratalis, sp. n.
J . Head, thorax, and abdomen whitish suffused with
brown; palpi blackish, white below; genital tufts ochreous
white. Fore wing whitish tinged and irrorated with yellow-
browm, the costal area and terminal area to vein 3 suffused
with brown ; traces of a dark antemedial line ; a slight dark
discoidal lunule ; a faint, dai’k, postmedial, minutely waved
line, erect from costa to vein 5, bent outwards to vein 2,
then retracted and almost obsolete to lower angle of cell and
erect to inner margin ; a terminal series of slight dark points ;
cilia ochreous white, with a slight brown line through them.
Hind wing tinged with ochreous ; a faint dark discoidal
lunule ; traces of a postmedial line erect from costa to vein 5,
bent outwards to vein 2, retracted to lower angle of cell and
oblique to tornus ; some slight dark points on termen
towards apex.
Hab. Guatemala, Cerro Zunil [Champion), 3 type,
Godman-Salvin Coll. ; Brazil, Rio Janeiro, 1 c^. Exp.
26 mm.
(8 a.) Stenia biannulalis, sp. n.
$ . Head, thorax, and fore w ing white slightly tinged with
pale red-brown ; palpi black above ; fore tibiae and tarsi
tinged with fuscous. Fore wing white ; the costal edge pale
reddish brown ; a diffused subbasal reddish-brown line from
of the Suhfaniilij Ily Jrocampinse.
costa to submedian fold ; aiitemedial line reddish brown,
rather diflused, obliquely curved ; an annulus in middle of
cell defined by rather diflused reddish brown and a similar
elliptical discoidal annulus ; postmedial line reddish brown,
diffused, with small black spot at costa, incurved from costa
to vein 2 near termen, then retracted to lower angle of cell
and rather outwardly oblique to inner margin ; a diffused
red-brown terminal band except at apex and tornus ; a dark
striga on termen from apex, followed by a series of slight
points ; cilia white, brownish at tips. Hind wing white ; a
diffused pale reddish-brown aiitemedial line; a diffused pale
reddish-browm postmedial line excurved between veins 5 and
2 and ending on termen above tornus ; a diffused pale
reddish-brown band just before termen from apex to vein 2 ;
cilia white faintly tinged with red-browm except tow^ards
tornus ; the underside with rather quadrate discoidal
annulus.
Hab. Brazil, Amazons, Pebas, 1 type. Exp. 20 mm.
(8 5.) Stenia aphenice^ sp. n.
Lederia lilienice, Druce, Biol. Centr.-Am., Het. ii. p. 249 (nec Cram,).
. Head and thorax purplish fuscous ; antennse yellowish
white with some dark points on shaft ; abdomen purplish
fuscous, the medial segments and ventral surface ringed with
white ; anal tuft yellowish ; wings white. Pore wing with
purplish-fuscous costal fascia ; an inwardly oblique straight
antemedial band ; both wings with outwardly oblique medial
and postmedial bands terminating before and above tornus ;
a terminal band expanding towards apex.
Hab. Mexico, Vera Cruz (^Schaus), 1 ^ type. Exp.
24 mm.
(8 c.) Stenia mallaleuca, sp. n.
<^. Pure white. Pore wing with some dark points on
costa ; the lines pale yellow-brown, the antemedial curved,
the postmedial excurved between veins 5 and 2, then
retracted to below end of cell ; traces of a discoidal spot.
Hind wing with indistinct discoidal spot ; the postmedial
line excurved between veins 5 and 2, then retracted to below
end of cell.
Hab. Brazil, Castro Paraha, 1 ^ type ; Sao Paulo.
Exp. 22-26 mm.
1
Z Sir G. F. llampson on new Pjrali'Ise
(10«.) Stenia 2>haofipilalis^ sp. ti.
Head and tliorax brownish grey; pectns white in
front; fore legs except tarsi fuscous ; abdomen grey. Fore
wing brownish grey irroratcd with a few black scales ; a very
ol)lique punctiform black antemcdial line ; a slight blackish
s[)ot in middle of cell and another at upper angle; post-
medial line black, rather punctiform, excurved from costa to
vein 3, then strongly incurved, and with blackish patch
from it to tornus below vein 2; a terminal series of slight
black points. Hind wing semihyaline whitish tinged with
brown, the termen slightly browner except towards tornus.
Hab. Bii. E. Africa, Teita [F. J. Jackson), 2 ^ type.
Eap. 14 mm.
(3 «.) Piletocera albipictalis , sp. n.
Antenme of male with a tuft of hair at one third, followed
by a ridge of hair, then a tuft at two thirds, followed by a
sinus ; fore wing with the basal half of costa lobed ; hind
wing with the termen excised below vein 3 and produced to
a rounded lobe at vein 1, the tornus truncate ; fore femora
with tuft of hair at extremity.
. Head black-brown, the vertex pure white; antennse
whitish, with the tufts at one third and two thirds blackish ;
thorax white, the outer edge of tegulse and patagia and the
tips of patagia dark brown, the dorsum of thorax tinged with
brown ; pectus and legs black-brown, the hind tarsi white ;
abdomen black-brown, the first three segments with wdiite
dorsal bands, the tw^o terminal segments with white dorsal
spots, the ventral surface white, the genital tufts ochreous.
Fore wing black-brown, the cell and basal half of inner area
pure white, confluent with a triangular patch between veins
5 and 2 extending to postmedial line ; a black point in middle
of cell and quadrate spot in end of cell, with its lower edge
indented by wdiite ; a browm striga on vein 1 « at inner
margin; postmedial line defined by wdiite on outer side,
somewdiat excurved below costa, slightly bent outw^ards
betw een veins 5 and 2, then retracted to near base of vein 2,
oblique and sinuous to middle of inner margin ; a white
point on termen above tornus ; cilia white at tips towards
tornus. Hind wdng blaek-browm ; some ‘whitish at base of
inner area ; a pale, oblique, minutely waved medial line,
obsolescent on costal half, whitish on inner half; an irregu-
larly quadrate white patch on terminal area between veins
5 and 2, leaving a dark terminal line ; cilia white from
of the Suhfamily Ilyclrocainpina?.
\3
vein 3 to the lobe ; tlie underside with maculate wliite medial
band from costa to vein 2.
Hab. Solomons^ Bougainville I. {Meek)^ 2 S type- Exp.
26 mm.
(3 b.) Piletocera slyyialis^ sp. n.
Antennae of male with a tuft of hair at one thirds followed
by a ridge of hair, then another tuft, followed by a sinus ;
fore femora with tuft of hair at extremity; fore wdug with
the basal half of costa lobed.
. Head, thorax, and abdomen black-brown ; antennae
with the tuft at two thirds of shaft white, the shaft beyond
it white above; tarsi with slight pale rings ; abdomen with
the terminal segment ochreous below, the genital tufts
ochreous white ; wings black-brown. Fore wing with very
indistinct pale postmedial line, slightly excurved below costa,
bent outwards between veins 5 and 2, then retracted towards
lower angle of cell and again slightly excurved. Hind Aving
with very indistinct pale postmedial line, slightly excurved
beloAV costa, bent outwards between veins 5 and 2, then
retracted to near lower angle of cell and oblique to tornus.
Hab. Solomons, Choiseul I. {Meek), 1 J type; Guadal-
canar I. {Meek), 1 ; Kulambangra I. {Meek), 1 . Exp.
24 mm.
(4 a.) Piletocera dentico stalls, sp. n.
Fore wing with tufts of hair on inner area below pro-
jecting over hind wing.
$ . Head whitish ; palpi with the extremity of second
joint and the third joint brown ; antennae brown to beyond
the tufts ; thorax cupreous brown ; pectus and legs white,
fore legs with the tuft of scales at extremity of femora
black. Fore wing cupreous brown ; a small whitish spot in
middle of cell, with others below it above and below vein 1 ;
a whitish discoidal lunule ; postmedial line with whitish bars
at costa and inner margin, excurved and obsolescent between
those points. Hind wing cupreous brown ; a dark post-
medial line defined by whitish on outer side, nearly straight
from costa to termen at vein 2, then retracted to lower angle
of cell and oblique to tornus ; the underside with the basal
area whitish, an indistinct discoidal annulus.
Hab. Solomons, Gizo I. {Meek), 1 ^ type. Exp. 20 mm.
Sir G. F. Ilampson on new Pyrali Jai
(4 b.) Piletocera infernalis, sp. n.
Fore win" with tufts of hair on inner area below projeeting
over hind wing.
(^ . Uniform black-brown.
Hah. Solomons^ Kulambangra I. [Meek), 2 type. Exp.
22 ram.
Subsp. 1. — Palpi at sides_, except third joint, and outer side
of fore legs whitish.
Hah. Solomons, Guadalcanar I. [Meek), 1 Exp.
18 ram.
(14 a.) Piletocera microdontalis, sp. n.
$ . Head, thorax, and abdomen cupreous brown ; palpi
in front, pectus, legs, and ventral surface of abdomen whitish.
Fore wing cupreous brown ; traces of a dark antemedial line ;
a white bar across cell before the dark discoidal lunule with
whitish centre ; subtermirial line white, somewhat puncti-
form, arising from costa towards apex nearly straight from
costa to vein 2 near termen, except that it is toothed outwards
at vein 5, retracted at vein 2 and almost obsolete except for
a small whitish spot in submedian interspace ; cilia whitish
just below vein 2. Hind wing cupreous brown ; postmedial
line pale, arising from costa towards apex, sinuous to termen
at vein 2 and toothed outwards below vein 5, below vein 2
retracted to lower angle of cell and almost obsolete, then
again whitish and oblique to tornus ; cilia with a fine white
line at base ; the underside with faint dark spot in middle of
cell and discoidal annulus.
Hah. Woodlark I. [Meek), 1 $ type. Exp. 18 mm.
(15 a.) Piletocera hadesialis, sp. n.
^ . Black ; pectus and ventral surface of abdomen yellowish
white. Fore wing with traces of a dark antemedial line ; an
ochreous point in middle of cell and discoidal somewhat
8-shaped spot. Hind wing with traces of dark discoidal
spot and postmedial line retracted at vein 3 to angle of cell.
? . Fore wing with the discoidal spot larger and with
traces of a dark postmedial line on its outer edge, retracted
below it.
Hah. Penang [Curtis). Exp. 16 mm. Type in Coll.
Rothschild.
of the Siihfamily ITy(liocamj)lria3.
V
(32«.) Piletocera phaeocraspedalis, sp. ii.
$ . Head and thorax orange-yellow tinj^ed with brown ;
abdomen yellow, dorsally sulfiised with brown exeept at
base, the ventral surface whitish. Fore wing orange-yellow,
the terminal area broadly fuscous ; the costa brown ; two
small brown spots in cell near base; a round spot in end of
cell with a short streak below it ; a yellow discoidal lunule
defined by brown ; postmedial line dark brown, somewhat
dentate, expanding slightly to costa, oblique to vein 7, angled
inwards in discal fold, then excurved to the dark terminal
area, at vein 2 retracted towards lower angle of cell, then
slightly excurved again ; a yellowish mark on termen at
vein 2; cilia dark brown, whitish above tornus. Hind wing
orange-yellow, the terminal area broadly fuscous; a dark
discoidal spot with oblique line from it to above tornus ; a
dark postmedial spot below costa with traces of the post-
medial line from it excurved between veins 5 and 2 ; some
yellow on termen at vein 2; cilia fuscous, whitish at sub-
median interspace ; the underside with discoidal annulus
defined by brown.
Hab. Solomons, Choiseul I. [Meek), 1 $ type. Exp.
18 mm.
(34 a.) Piletocera rotundalis, sp. n.
Very dark brown; palpi at base, pectus, legs, and
ventral surface of abdomen whitish. Fore wing short and
broad, the apex rounded ; a dark-edged annulus at middle
of cell and discoidal lunule with whitish spot between them ;
traces of an antemedial line ; a minutely dentate postmedial
line defined by pale brown on outer side, excurved between
veins 5 and 2, then retracted to lower angle of cell. Hind
wing with traces of ante- and postmedial lines. Underside
of fore wing with fascia of black scales in and beyond upper
angle of cell and whitish patch beyond the cell.
Hab. Bali [Doherty), 2 $ type. Exp. 16 mm.
Subsp. 1. — Fore wing more uniform dark brown, the
markings obscured; the white spot reduced to a point;
underside without the fascia of black scales, a small white
spot beyond cell.
Hab. Talaut [Doherty), 1 Woodlark I. [Meek),^^-,
St. Aignan [Meek), 1
Sir G. F. Ilampson oyi upao PyralidaB
(37 «.) Piletocera micralis, sp. n.
Antennse of male laminate ; hind wing with the termen
excurved at middle and excised below apex and towards
tornus.
S . Pale yellow ; fore tibise with black band at extremity ;
abdomen with black dorsal line before anal tuft. Fore wing
with black points at base on costa and below cell ; an
ill-defined fuscous subbasal line ; aiitemedial line excurved
from median ncrvure to inner margin ; a small dark spot in
middle of cell and discoidal lunule defined by fuscous^ with
small blackish spot above it on costa ; two black points on
postmedial part of costa, the postraedial line arising from the
outer point, excurved below vein 5 and ending at tornus,
towards which it is somewdiat diffused ; a rather strong black
terminal line ; cilia whitish with blackish tips. Hind wing
with slight dark discoidal spot, with sinuous line from it to
inner margin ; postmedial line sinuous, excurved below
vein 5 and ending at tornus ; a rather strong blackish
terminal line from apex to vein 2 ; cilia whitish, fuscous at
tips on apical half.
Hab. PoRNEo, Kuching {Shelford), 1 J' type. Exp.
8 mm.
(37 h.) Piletocera cumulalis, sp. n.
Antennse of male laminate and without tufts ; fore wing
normal.
$ . Head, thorax, and abdomen pale yellow, slightly irro-
rated with fuscous ; palpi with black bars at extremity of
first and second segments ; abdomen tinged with orange at
extremity and with subdorsal black spots on terminal
segment ; pectus, legs, and ventral surface of abdomen
whitish, the fore tibise with fuscous band at extremity. Fore
wing pale yellow sparsely irrorated with black and with
slight blackish suffusion on terminal area below apex ; a
black subbasal line, slightly angled inwards in cell, then
oblique and somewhat maculate ; antemedial line oblique
from costa to just below median nervure, then erect ; a small,
rather indistinct, dark annulus in middle of cell ; a discoidal
lunule with yellow centre ; postmedial line incurved from
costa to vein 4y then erect to vein 2, then retracted to lower
angle of cell and oblique to above inner margin ; a terminal
series of small black spots from costa to vein 4 and a spot
above tornus; cilia fuscous at tips. Hind wing pale yellow
with fuscous suffusion between lower angle of cell and tornus
of the Subfamily llydrocamplnio.
atul beyond the postmcdiiil line below eosta ; a small Ijlaek
diseoidal spot and line t‘roin lower atiirle of eell to inner
margin near tornus ; postrnedial line bent outwards at
vein 5 and again at vein 2, ending at tornus ; a black ter-
minal line from apex to vein 3 and towards tornus ; eilia
blaekish at tips.
Hub. Borneo, Kueliing {Shelford), 1 type. Erp.
IG mm.
(38 fl.) Piletocera ranalis, sp. n.
(d' . Head and thorax fulvous yellow slightly tinged with
brown ; abdomen pale fulvous yellow, the anal tuft blackish.
Fore wing fulvous yellow slightly tinged with brown, the
basal costal area rather darker ; a curved, slightly waved
dark antemedial line; a black diseoidal spot; postrnedial
line black, slightly angled outwards below costa and inwards
in discal fold, then excurved, at vein 2 retracted towards
lower angle of cell and again slightly angled outwards at
vein 1 ; a series of small black spots just before termen; a
fine terminal fuscous line ; cilia whitish. Hind wing fulvous
yellow slightly tinged with brown; a small oblique blackish
diseoidal spot ; postrnedial line blackish, excurved below
costa, angled inwards in discal fold, then excurved, at vein 2
retracted to lower angle of cell, then oblique to inner margin ;
a series of small black spots just before termen ; a tine
fuscous terminal line ; cilia whitish.
$ much more suffused with fuscous ; the antemedial
line of fore wing on inner side and the postrnedial line of
both wings on outer side defined by ochreous.
Hab. Brazil, Lr. Amazons, Breves {Austen), 3 3^, 3 $
type ; Parana de Buyassu, 1 1 ? . Exp., ^ 16, ? 20 mm.
(38 b.) Piletocera albicilialis, sp. n.
$ . Head, thorax, and abdomen browm mixed with yellow ;
pal[)i brown, white below; pectus, legs, and ventral surface
of abdomen whitish ; fore tibiae with fuscous band at extre-
mity. Fore wing yellowish almost wholly suffused with
brown, leaving the medial part of costa yellow ; an indistinct
dark antemedial line, oblique from costa to median nervure,
then erect; an indistinct dark diseoidal annulus; two
blackish semicircular marks on costa just beyond middle;
postrnedial line indistinct, dark slightly defined by yellowish
on outer side, minutely dentate, erect from costa to vein 2,
then retracted to lower angle of cell and slightly excurved at
Ann. Sf Mag. N. Hist. Ser. 7. Vol. xix. 2
•fs
Sir G. F. Hampson on new Pyralidse
vein 1 ; a terminal series of small black spots ; cilia yellow
at base followed by a black line, the tips white slightly inter-
sected with fuscous. Hind wing yellowish almost wholly
suffused with brown ; a faint dark postmedial line slightly
defined by whitish on outer side, nearly straight from costa
to vein 2, then retracted to lower angle of cell and oblique
to inner margin; a fine black terminal line ; cilia yellow at
base followed by a black line, the tips pure white.
Hab. Brazil, Lr. Amazons, Parana de Buyassu {Austen)^
1 ? type. Exp. 20 mm.
(38 c.) Piletocera rufulalis, sp. n.
Fore wing of male with no fovea in cell.
^ . Pale rufous ; fore tibiae with black band at extremity ;
tarsi whitish with fuscous bands ; abdomen with black dorsal
line before anal tuft ; wings slightly irrorated with fuscous.
Fore wing with subbasal blackish points on costa and in ceil ;
antemedial line blackish, oblique from costa to submedian
fold ; a blackish spot in middle of cell and discoidal lunule ;
three small black annuli on postmedial part of costa, the
postmedial line arising from the outermost, excurved between
veins 5 and 2 along which it is retracted, then again excurved ;
a terminal series of small black spots ; cilia with series of
slight fuscous spots at tips. Hind wing with small blackish
discoidal spot ; postmedial line bent outwards between veins
5 and 2, then retracted to below end of cell and oblique to
tornus ; a terminal series of small blackish spots ; cilia with
series of slight fuscous spots at tips.
Hab. Sierra Leone [Clements), 2 ^ type. Exp. 20 mm.
(38 d.) Piletocera fidvalis, sp. n.
$ . Head, thorax, and abdomen fulvous yellow ; palpi
fuscous above, white below ; pectus, legs, and ventral surface
of abdomen whitish, the fore tibiae with fuscous band at
extremity. Fore wing fulvous yellow ; the costal edge
fuscous ; antemedial line indistinct, oblique, slightly angled
just below median nervure ; a small rather ill-defined ocellus
in middle of cell ; a discoidal lunule with yellow centre ;
postmedial line expanding into a small black spot at costa,
incurved to vein 5, then excurved to termen at vein 2, then
almost obsolete and retracted to just below lower angle of
cell and again distinct and slightly excurved ; a maculate
terminal line from below apex to vein 2 ; cilia with a slight
fuscous line at middle. Hind wing fulvous yellow ; a small
of the Subfamily llydi’ocamplnai.
fuscous discoidal spot ; postmedial line fuscous, cxcurvcd
from vein 5 to termcu at vein 2, then almost obsolete and
retracted to just below lower angle of cell and again distinct
and oblique to inner margin ; a maculate blackish terminal
line ; cilia with a slight fuscous line at middle.
Hab. Bii. Guiana, Potaro 11. [Kaye), 1 ? type. Exp.
18 mm.
Genus Neogenesis, nov.
Palpi porrect, triangularly scaled, downcurved at extremity
and extending about twice the length of head ; maxillary
jialpi with pointetl tuft at extremity ; frons flat and oblique ;
antennse annulate; tibiie with the spurs long and nearly
even ; abdomen very long and slender. Fore wing of male
with the costa strongly arched and fringed with short hair,
the apex rounded and truncate; a fringe of very long hair
on inner area below ; vein 3 from well before angle of cell ;
4, 5 stalked; 7 straight and well separated from 8, with
w'hich 9, 10, 11 are coincident; female with vein 9 absent,
10, 11 stalked with 8. Hind wing with vein 3 from before
angle of cell ; 4, 5 stalked ; 6, 7 stalked, 7 anastomosing
with 8.
A development from Clupeosoma.
(1.) Neogenesis flaviplagialis, sp. n.
Pale rufous ; palpi dark below ; throat pure white ; fore
tibiae banded with fuscous and white ; abdomen ringed with
white; wings semihyaline. Fore wing with indistinct den-
tate subterminal line defined by pale yellow on inner side,
most strongly towards inner margin, towards which it is
bent inwards, the area beyond it pink ; termen yellow with
series of black points. Hind wing suffused with pink except
costal area ; a dentate postmedial line with large yellow
patch on its inner side between veins 6 and 2, the termen
and cilia yellow with series of black points.
Hab. Br. N. Guinea, Moroka [An'hony], 1 1 ? type,
Milne Bay [Meek), I . Exp,, c? 2f, ? 22 mm.
(4 a.) Clupeosoma laniferalis, sp. n.
^ . Fore wing with a fold the whole length of submedian
interspace containing a fringe of long hair on upperside ;
a large patch on upperside beyond the cell clothed witli
rough hair directed towards the centre.
2^
Sir G. F. Ilampson on new Pyralidae
Head, thorax, and abdomen rufous ; throat, greater part
o£ tibiae, the tarsi, and segmental rings on abdomen wliite.
Fore wing rufous, the costal area towards apex and termen
pink; tlie cilia yellow. Hind wing pale rufous; a pink
patch in and below end of cell followed by a large yellow
lunulatc patch, the area beyond it black down to vein 2 ;
cilia yellow.
Hah. Louisiades, St. Aignan [Meek). Exp, 14 mm.
Type in Coll. Rothschild.
ScOFAHIAN^.
(3.) Microglossa JIavidaHs, sp. n.
. Head, thorax, and abdomen yellowish suffused with
fuscous; third joint of palpi and basal joint of antennse
blackish ; legs whitish, the fore tibiae and the tarsi banded
with blackish ; abdomen with the anal tuft rufous. Fore
wing yellowish irrorated with fuscous; a black subbasal line
emitting short streaks below cell and on inner margin; two
blaek antemedial spots on costa, a slight mark in cell and a
band from cell to inner margin expanding into a patch below
median nervure, a black point beyond it in cell ; a rather
quadrate black discoid al spot ; two small black postmedial
spots on costa, then a yellowish line defined on each side by
blackish, excurved from below costa to vein 4, then incurved
to below angle of cell and slightly waved to inner margin ;
terminal area rather more suffused with black, with diffused
yellowish marks below apex and at middle; cilia yellowish
intersected with fuscous at base, whitish at tips. Hind wing
whitish slightly tinged with brown.
Hah. W. China, Chang-Yang [Pratt) ^ 1 ^ type. Exp.
14 mm.
(3.) Eclipsiodes cuprealis, sp. n.
. Head and thorax dark cupreous brown ; palpi whitish
below at base; tarsi with slight whitish rings; abdomen
cupreous brown with a greyish tinge, the anal tuft with some
whitish hairs below. Fore wing dark cupreous brown with
slight dark irroration ; a faint diffused dark discoidal spot ;
traces of a dark postmedial liiie excurved from costa to
vein 4, then oblique ; a faint dark terminal line. Hind
wing cupreous brown with a greyish tinge, the cilia whitish
at tips.
Hah. Victoria, Gisborne [Lyell), 1 c^type. Exp. 24 mm.
of the Suhfamihj Scoparianai.
21
(4.) Eclipsiodes striatalis, sp. n.
Ilcafl and thorax pale grey irrorated witli fuscous ; abdo-
men l)rownisli, the basal segment dorsally orange, lore
'vving grey-white strongly irrorated with fuseous ; the veins
streaked vvitli blaek ; a prominent black streak just below
submedian fold with a whitish streak below it ; a similar
streak in cell from before middle to extremity with a more
prominent white streak below it ; a short streak beyond the
cell connected by a diffused oblicpie streak wdth the apex ;
a series of dentate marks on termen. Hind wing tinged
with browm, especially towards apex ; cilia w bite with a brown
line through them.
Hab. W. Australia^ Sherlock R. {Clements), 2 J type.
Ea:>j?. 24 mm.
(84 a.) Scoparia rufitinctalis , sp. n.
Head and thorax yellowdsh white ; sides of palpi and
shoulders pale rufous ; fore legs tinged with rufous ; pectus,
mid and hind legs, and abdomen white. Fo e wdng yellowish
white rather sparsely irrorated with rufous ; a curved ante-
medial rufous band diflPused on outer side and with slight
dark streaks on it in and below cell ; a diflused rufous patch
from middle of costa to lower angle of cell with black dis-
coidal point on it ; an oblique rufous postmedial line, slightly
excurved at middle and with darker point at costa, a broad
band of rufous suffusion from just beyond it, constricted at
middle ; the termen suffused with rufous, expanding some-
what at discal fold and with obscure series of dark points on
it; cilia yellowish white with a brown line near base. Hind
wing white, faintly tinged with ochreous towards termen.
Hab. U.S.A,, Washington Terr., 4 ^ , Calitornia, Shasta
Co., Pit R. ( Walsingham), 7 cf 5 ^ ? type. Exp. 20 mm.
(88 a.) Scoparia poliophcealis , sp. n.
Head and thorax dark reddish brown mixed with some
white scales; abdomen pale reddish brown; pectus, legs,
and ventral surface of abdomen white, the fore t]bi^e and the
tarsi banded wdth brown. Fore wing whitish thickly irrorated
and suffused with reddish brown, the suffusion forming dark
bands beyond the ante- and postmedial lines ; an indistinct
rather diffused whitish subbasal line; antemedial line diffused,
white, oblique from costa to vein 1, where ic is angled, then
angled inwards above inner margin ; a slight dark spot
22
Sir G. F. Ilampson on new PyralIJa3
l)cyoTi(l it in cell ; a reniform cliscoidal stifjma incompletely
(lefined by diffused dark brown, and with some greyish
siiffnsion beyond it before the postmedial line, which is white
defined on inner side by a rather punctiform brown line,
slightly angled inwards above vein 6, then excurved to vein 4,
and obli(pie to itiner margin ; a diffused whitish snbterminal
line somewhat angled inwards at diseal and submedian folds;
a series of small dark brown spots just before termen ;
cilia whitish tinged with brown and with a series of brown
strim near base. Hind wing greyish brown, the cilia white
•with a brown line near base ; the underside greyer with traces
of a curved postmedial line.
Hah. SyiiiA, Lebanon [Pratt)^ 1 $ type. Exp. 2G-
30 mm.
(90 c^.) Scoparia microdontalis, sp. n.
Head and thorax mixed with fuseous and brown ; palpi dark
brown at sides, the base and the maxillary palpi at tips white ;
pectus and legs white, the fore tibiae and the tarsi banded
with fuscous ; abdomen grey tinged with brown, the ventral
surfaee white. Fore wing white tinged with brown and
irrorated wdth black ; antemedial line indistinctly double filled
in with white, oblique to’wards costa, angled outwards on
median nervure and inwards in submedian fold, with short
black streaks beyond it in cell and submedian fold; a
slight dark patch on middle of eosta ; a rather X -shaped
black discoidal spot ; postmedial line double filled in wdth
white and with some dark suffusion beyond it, minutely
dentate, excurved from costa to vein 5, then oblique ; a dark
patch on middle of termen; a terminal series of small, some-
what dentate black spots w ith white strise betw'een them ;
cilia chequered pale brown and white with a dark line near
base. Hind wing white tinged with brown.
Hob. Japan, Hakodate [Andrews), 1 c^, 1 $, Kiushiu,
1 $ , Yokohama [Jonas), 1 ^ type. Exp. 16-22 mm.
(105 a.) Scoparia isochroalis, sp. n.
$ . Head, thorax, and abdomen pale reddish brown, the
ventral surface paler ; fore tibiae and tarsi banded fuscous
and white. Fore wing wdjitish almost wholly suffused with
red-brown and irrorated with fuscous ; slight subbasal dark
spots in cell and above inner margin ; antemedial line defined
by whitish on inner side, oblique towards costa, slightly
angled inwards in submedian fold, then excurved, a short
23
of the Subfamily Scopariana?.
black streak beyond it in submedian fold ; some whitish in
end of cell followed by two black discoidal bars with a black
point beyond them; postmedial line indistinct, detined by
whitish on outer side, oblique towards costa, cxcurved to
vein 4, then inwardly oblique and minutely dentate, some
darker brown suffusion beyond it on costal area; a series of
short black streaks in interspaces of terminal area, the streak
above vein 4 extendinj^ to postmedial line ; cilia whitish
with a dark line near base. Hind wing whitish suffused
with pale red-brown ; cilia ochreous white with a fine dark
line near base.
Hab. Japan, Hakodate {Andrews), 1 $ type. Exp. 18 mm.
(107 a.) Scoparia met aleiic alls , sp. n.
S . Head and thorax grey tinged with brown and fuscous,
the palpi at base and tips of maxillary palpi white ; pectus
and legs whitish, the tarsi ringed with fuscous ; abdomen
ochreous white. Fore wing wdiite slightly tinged with
ochreous and irrorated with dark brown ; short dark streaks
from base in cell and above inner margin; a diffused dark
curved antemedial line wuth dark spot beyond it in cell and
suffusion from median nervure to inner margin ; a large
diffused dark brown rounded discoidal patch with some dark
suffusion above it on costa ; postmedial line whitish defined
on inner side by a brown patch on costa, then by short streaks,
and wdth browui suffusion beyond it on costal area and from
vein 3 to inner margin, excurved from below costa to vein 4,
then incurved ; the termen dark with a diffused patch at
middle ; cilia white with a dark line near base. Hind wing
pure w hite.
Hah. W. China, Pu-tsu-fang, 1 $ type. Exp. 18 mm.
(109 a.) Scoparia luteiisalis, sp. n.
$ . Head and thorax oehreous yellow mixed with rufous ;
abdomen ochreous white. Fore wing ochreous yellow ; a
diffused reddish-brown streak below base of cell with some
blackish scales on it and a slight streak above base of inner
margin ; a diffused brown antemedial band, oblique from
costa to submedian fold with a blackish streak from it in cell
and two shorter streaks below the cell ; a diffused brown
postmedial band with some blackish scales on it, oblique from
costa to submedian fold, where it is angled inw^ards, and
expanding into a discoidal spot; a diffused reddish-brown
subterminal band, bent outwards to apex and with slight
24
Sir G. F. Ilampson on new Pjrallda?.
(larl< streaks on it in the interspaees ; the termen tinged with
reddish browni with a series of black points. Hind wing
semiliyalinc wdiite faintly tinged with brown ; a slight brown
terminal line and slight line through the cilia.
Jlah. Azores [J. J. Walker), 1 ? type. E.xp. 22 mm.
(Ill a.) Scoparia melanographa, sp. n.
Scoparia slenota, Warr. Nov. Zool. xii. p. 447, nec Wlstn.
^ . Head and thorax black mixed wdth some grey-while;
palpi white below at base; pectus whitish; legs whitish
banded wntli fuscous ; abdomen black with white segmental
lines, the ventral surface white with fuscous bands. Fore
wing narrow, wdiite irrorated with black ; the base suffused
with black, followed by a band formed of short diffused black
streaks ; a diffused black antemedial line, rather angled
inwards below cell, connected wdth an obscure diffused
annulus from just below' costa to median nervure and with
a patch of blackish suffusion beyond it on inner area ; a black
streak on middle of costa connected by a bar with the rather
X -shaped black discoidal spot, with a slight line from it to
the black jiatch on tornal area; postmedial line black,
strongest at costa, angled inwards at discal fold, then out-
wards at vein 4 and oblique to the patch on tornal area ; a
black patch from costa before apex to the triangular patch
on middle of termen ; a black terminal line ; cilia grey with
a black line near base, blind wing narrow', whitish tinged
with fuscous especially towards termen ; a black terminal
line ; cilia w hitish wdth a tine dark line near base.
Hah. Azores, S. Jorges, 1200' [0. -Grant), 1 $ , Terceira,
2200' [O.-Grant), 1 S fyie. Exp. 14 mm.
(124 «.) Scoparia albifusalis, sp. n.
$ . Head, thorax, and abdomen fuscous ; palpi white at
base; pectus and ventral surface of abdomen white; legs wdiite
and fuscous. Fore wing fuscous sparsely irrorated with
wdiite ; antemedial line white, oblique from costa to sub-
median fold ; an ill-defined dark discoidal spot ; postmedial
line white, obtusely angled at vein 6 ; large wdiite patches
on termen at apex, discal and submedian folds connected by
the white terminal line. Hind wing greyish fuscous w ith a
fine pale line at base of cilia.
Hub. Ceylon, Bogowantalawa, Maskeliya [Pole), 3 ? type.
E.xp. 16 mm.
On some new Species of Blattlclaa.
25
II. — On some new Species o/Blalti(la3 m the Oxford and
Paris Museums. By II. Siielford, M.A., F.L.S.
Subfam. Ectobinje.
Genus Anaplectoidea, Shelf.
Anaplectoidea Dohertyi, sji. n.
$ . Clear testaceous, nitid. Head rufo-testaceous, antennae
testaceous. Pronotum posteriorly truncated, exposing the
scutellum, testaceous, lateral margins hyaline. T( gmina with
fifteen costal veins; anterior ulnar with six branches, some of
which are oblique, posterior ulnar simple ; anal vein strongly
curved, well marked ; four axillary veins ; the part of the
right tegmen overlapped by the left reticulated. Wings
hyaline, suffused with a pale flavid tint, with ten costal veins
incrassated at the apex ; medio-discal field crossed by eleven
transverse veinlets ; ulnar vein curved upwards, with five
branches, the medio-ulnar field only one third the breadth
of the medio-discal field, first anal vein quadriramose ; apical
area small, barely one fifth of total wing-length, its basal
margin obtusely angled, its apex slightly emarginate, divided
almost equally by a longitudinal vein. Supra-anal lamina
produced, its apical margin straight ; subgenital lamina
ample, semiorbicular ; cerci elongate, nine-jointed.
Length of body 7 mm. ; length of tegmina 6 mm.
Sangir [IV. Doherty) ; one example (Oxford Mus.).
Genus Hololampra, Sauss.
Hololampra minuta, sp. n.
$ . Head rufo-castaneous, antennae fuscous with exception
of first two basal joints, which are testaceous. Pronotum
covering vertex of head, trapezoidal, the angles rounded, sides
deflexed, posterior margin nearly straight ; all the margins
testaceous, most broadly at posterior angles ; disk rufo-casta-
neous, with a posterior transverse castaneous vitta which is
notc hed anteriorly. Tegmina ovate, smooth, shining, hyaline-
testaceous, extending as far as middle of fourth abdominal
tergite, the veins testaceous; eight costal veins, the last three
ramose, discoidal sectors oblique and few in number, anal
vein not impressed. Wings scale-like. Abdomen testaceous
above, banded with black, beneath black margined with
26
Mr. 11. Shelford on some
testaceous, last segment and subgenital lamina rufo-casta-
ucous ; supra-anal lamina short, trigonal, with a median
impressed line; subgcnital lamina ample, semiorbicular,
projecting beyond the supra anal lamina; cerci moderate,
black. Legs rufo-testaeeous ; femora very sparsely armed,
not more than two spines on the anterior and posterior
margins of each.
Total length 6 mm. ; length of tegmiiia 4 mm.
Andrahoniana, South Madagascar [Ch. Alluaud, 1901) ;
twm examples (Paris Museum).
One of the smallest species of the genus.
Subfam. Phyllobromin^e.
Genus Isciinoptera, Burm.
Ischnoptera Ridleyi, sp. n.
S. Head ferruginous, a V-shaped darker mark between
the antennal sockets ; antennae ferruginous, longer than the
body. Pronotum rounded, posteriorly truncate, sides slightly
deflexed, not covering vertex of head ; disk rufous, with two
broad vittae, black in colour, not reaching the posterior
margin, their outer margins sinuate, their inner margins
straight. Tegmina ferruginous ; radial vein bifurcated at
the middle ; sixteen costal veins ; discoidal area with eleven
longitudinal sectors ; seven axillary veins. Wings hyaline,
veins fuscous, the marginal field suffused wdth flavid ; radial
vein bifurcated at the middle, mediastinal vein wfith five
branches; fourteen costal veins ; median vein sinuate ; ulnar
vein with fourteen branches, six of which go to the apex of
the w'ing. Abdomen infuscated above, ferruginous below^ ;
the seventh tergite notched in the middle of its posterior
margin, the eighth very narrow ; the eighth sternite reduced
to a pair of lateral lappets ; the supra-anal lamina quadrate,
its posterior margin notched and produced on each side of
the notch into tw o curved and slender processes ; the sub-
genital lamina subquadrate, its posterior margin much
thickened, forming two asymmetrical swellings wLich are
grooved and furnished with numerous minute denticles ;
styles absent. Front femora with the anterior border beneath
armed throughout its length with long stout spines.
Length of body 22 mm. ; length of tegmina 20 mm.
Singapore [H. N. Ridley, March to May, 1906) ; one
example (Oxford Museum).
This species, like so many of its Oriental congeners, presents
unusual modifications of those external parts which are
27
neto Species of Blattida3.
related to reproduction. The form of tlie subgenital lamina
in I. lUdleyi recalls that of Hemithyrsocera hisirio, Burrn.
Ischnoptera per'pulchra, sp. n.
? . Head testaceous, antennae (mutilated) testaceous at
base. Pronotum trapezoidal, posterior margin slightly pro-
duced ; disk flavo-testaceous, encircled by a broad pcnannular
ring of black, open on the anterior margin, lateral and poste-
rior margins testaceous. Tegmina castaneous, margin testa-
ceous ; eighteen costal veins ; radial vein bifurcated beyond
the middle, the lower branch sending ramifications to the
apex of the wing ; discoidal field with nine longitudinal
sectors, the most internal of which are angled. Wings
hyaline, veins fuscous, marginal field bordered with testa-
ceous ; mediastinal vein with six branches, ten to twelve
costal veins, radial vein bifurcated beyond the middle ; ulnar
vein with seven branches, three of which run towards the
dividing vein. Abdomen flavo-testaceous above, except at
the apex, which is fuscous, testaceous below ; supra-aual
lamina produced triangular, subgenital lamina ample. Legs
testaceous ; front femora armed on the anterior margin
beneath with strong spines throughout its length, the most
basal the longest; formula of apical spines j-; front
femora without a genicular spine, unless the most anterior
apical spine is to be regarded as such.
Length of body 12 mm. ; length of tegmina 14 mm.
Macassar, Celebes {W. Doherty, 1896) ; one example
(Oxford Museum).
This species is rather a puzzling one ; the angulation of
some of the discoidal sectors of the tegmina suggests the
genera Fseudomops and Pseudothyrsocera, but this is a
character that also crops up in Phyllodromia, and, taken by
itself, is not of the greatest importance ; it is on account of
the branching of the vena ulnaris alarum, so characteristic of
Ischnoptera, that 1 have referred this species to that generic
position.
%
Ischnoptera cavernicola, sp. n.
S. Head castaneous, mouth-parts testaceous; antennae
testaceous, one and a half times longer than the body.
Pronotum rufo-castaneous, with lateral and posterior margins
narrowly castaneous. Tegmina clear testaceous, radial vein
bifurcated, twelve costal veins, six discoidal sectors. Wings
hyaline, mediastinal vein with three branches, seven costal
28
Mr. H. Slielford on some
vcins^ radial vein not bifurcated, ulnar vein sending three
brandies to tlie dividing vein and three to the apex of the
"vving. Abdomen rufo-tcstaccous, supra-anal lamina sliglitly
jiroduccd, rounded ; subgenital lamina produced, narrow,
with two stout styles; cerci elongate. Legs rufo-testa-
ceous ; front femora with anterior margin beneath armed
throughout its length with a series of short spines, the
most distal the shortest; apical spines {, j- ; genicular
spines 1, 1, 1.
Length of body 10 mm. ; length of tegmina 9 5 mm.
In cave at Bidi, Sarawak, Borneo {R. Shdford) ; one
example (Oxford Museum). [No. 22.]
The species, which can he readily recognized by the pale
pronotum with dark margins, was found in some numbers in
a large and quite dark cave in the limestone formation at
Bidi, Sarawak ; it is somewhat remarkable that this cock-
roach, a Stenopelmatid (? Dolichopoda) , and a species of crab
[Potamon hidiense, Lanch.), which were all found in con-
siderable numbers in the darkest part of the cave, show no
reduction in size of the eyes ; one can only suppose that
the caves have been peopled within quite recent times by
these Arthropoda. The antennae of the Stenopelmatid are
of great length, but this is not the case in Ischnoptera
cavernicola.
Genus Ellipsidium, Sauss.
Ellipsidiwn castaneum, sp. n.
$ . Head dark rufous, with a black spot between the
antennal sockets ; antennae with the basal half strongly
incrassated, black, the two basal joints rufous, apical half
with a testaceous band occupying ten lower joints, the
remaining joints fuscous. Disk of the pronotum rufo-
testaceous, with a symmetrical black design, anterior and
posterior margins pale testaceous, lateral margins hyaline.
Tegmina castaneous, with the densely reticulated veins pale
testaceous, the mediastinal field hyaline. Wings infuscated,
veins flavid. Abdomen black beneath, sternites with white
margins ; subgenital lamina with the apex slightly cleft ;
cerci black, with castaneous legs. Legs castaneous, the
coxae black, with white borders, the tarsi black.
Total length 14 mm. ; length of tegmina 12'5 mm. ;
pronotum 4x6 mm.
Humboldt Bay, New Guinea {W. Doherty, 1896) ; one
example (Oxford Museum).
29
new Species of Blattidae.
Tin’s well-marked speeies somewhat extends the range of
the genus, hitherto known only 1‘rom Australia.
Genus Piroblatta, iiov.
Differs from Chrastohlatta, Sauss. & Zehnt., by the less
prominent vertex, by the shape of the pronotum, by the
greater breadth of the tegrnina, and by the presence of a
prominent triangular apical area in the wings.
Head almost covered by the pronotum ; pronotum trape-
zoidal, anterior margin truncate, sides deflexed, posterior
margin slightly arcuate ; scutellum exposed ; tegmina longer
than the body, discoidal sectors oblique. Wings with a
large apical triangle, projecting beyond the anterior part
of the wing ; ulnar vein bifurcate and sending also two to
three branches to the dividing vein. Front femora unarmed
beneath, mid and hind femora very sparsely armed on both
borders beneath ; genicular spines present on all the femora.
Supra-anal lamina in the male somewhat quadrately pro-
duced, in the female triangularly produced.
On a re-examination of the species described by me as
Theganopteryx Bouvieri (Trans. Ent. Soc. 1906, p. 236) I
have come to the conclusion that a new genus must be
established for this and for the species described below. In
general appearance both species are very like Theganopteryx,
but the branching of the ulnar vein of the wings is sutFicient
to place them close to the genus Chrastoblatta, from which,
however, they may be distinguished by the points enumerated
above. Both species are testaceous in colour and quite
unlike the conspicuous Chrastoblatta dhnidiata, Sauss. &
Zehnt., and C. tricolor, Sauss. & Zehnt. The females are
shorter and a little broader than the males.
Piroblatta Alluaudi, sp. n.
. Head rufo-eastaneous ; maxillary palpi and antennae
testaceous, the latter longer than the body ; pronotum rufo-
testaceous, the lateral margins hyaline. Tegmina testaceous
hyaline ; fifteen to sixteen costal veins, eight oblique discoidal
sectors. Wings hyaline; veins fuscous, marginal field
flavid ; fourteen to fifteen costal veins, their apices very
slightly incrassated ; ulnar vein bifurcated and sending three
branches to the dividing vein; first axillary vein quadri-
ramose, triangular apical field large. Legs and cerci rufo-
testaceous. Abdomen piceous ; supra-anal lamina trigonal,
subgenital lamina semiorbicular ; the left style stout and
curved, the right style minute.
m
Mr. K. fSlielibrd on some
$ . Similar to the male, but shorter ; supra-anal lamina
triangular ; sul)genital lamina ample.
S . Length of body 7 mm. ; length of tegmina 8 mm.
2 . Length of body 7 mm. ; length of tegmina 7*5 mm.
Diego Suarez, Madagascar (Alluaud, April 1896) ; eight
examples (Paiis Museum).
Piroblatta Boiivieri, Shelf.
The(/anopteryx Bouvieri, Shelford, Trans. Ent. Soc. London, 1906,
p. 266.
The female has the pronotum less strongly marked with
testaceous than the male; the supra-anal lamina is trigonal,
the subgenital lamina ample and semiorbicular ; the tegmina
measure 8 mm., as against 10‘8 mm. in the male, the body
8 mm., as against 9 5 mm. in the male.
Genus Phyllodromia, Serv.
Phyllodromia pictiirata, sp. n.
A . Head testaceous, with castaneous markings, forming a
symmetrical design. Pronotum transversely elliptical, lateral
margins hyaline ; disk of the pronotum pale testaceous, with
castaneous markings composed of irregular spots and two
central longitudinal lines ; a few minute castaneous points
in the hyaline margins. Tegmina hyaline testaceous, Avitli
castaneous spots disposed along the veins, denser at two
points in the marginal field, forming two indistinct maculae;
an oblique castaneous fascia on the right tegmen extending
from the middle of the anal field to the apical third of the
radial vein ; mediastinal vein with two branches, radial vein
not bifurcated, twelve costal veins ; anterior ulnar vein
sending several branches to the sutural margin, posterior
ulnar vein simple ; anal vein somewhat sinuate, its apex
suddenly bent inwards. Wings hyaline; nine costal veins,
the six basal clavately incrassated, the apical three ramose;
ulnar vein with five branches. Abdomen infuscated above,
testaceous, marbled with fuscous below ; supra-anal lamina
short, transverse ; subgenital lamina ample, somewhat irre-
gular, the left style larger than the right and curved (cerci
mutilated) . Legs testaceous ; tibise banded wntli castaneous ;
tarsi fuscous except the basal two thirds of the first joint;
front femora not armed beneath ; mid and hind femora
sparsely spined ; apical spines T, -}-, } ; no genicular spine on
front femora.
31
new Species of BlattlJa3.
Length of body 9 imn. ; leiigtli of tcgmiiia 10 mm.; pro-
notum 2*2 x 4' 5 mm.
Singapore, Botanic Gardens {II. N. Ridley) ; one example
(Oxford Museum).
The s[)ecies in the eharactcrs presented by the femora and
supra-anal lamina resembles the Ectobinje, but the wing-
structure is typically Phyllodromine.
Pliijllodromia albovarieyata, sp. n.
$ . Head dark eastaneous, with two diverging testaceous
lines running from the vertex to the sides of the clypeus;
clypeus rufo-testaceous ; palpi fusco-testaceous ; antennae
testaceous at base, the rest fuscous. Pronotum trapezoidal,
dark castaneous, with a narrow central line testaceous, lateral
margins hyaline. Tegmiiia dark castaneous, paler towards
the apex, mediastinal field and base of marginal field hyaline ;
a transverse white fascia extending from the marginal field
to the apex of the anal field, not meeting its fellow of the
opposite side ; twelve costal veins, the apical four branched,
discoidal sectors oblique. Wings infuscated, costal margin
very narrowly flavid ; eleven costal veins, their apices iticras-
sated ; ulnar vein 5-ramose, the branches joined by trans-
verse venulse; a prominent apical triangle. Abdomen
fuscous, with a fulvous patch on the disk below ; supra-anal
lamina short, transverse ; subgenital lamina large, produced,
its apex cleft ; cerci long, testaceous. Legs testaceous, front
femora armed with several spines on the anterior margin
beneath, the more distal being the shortest of the series ;
mid femora strongly spined; hind femora sparsely spined,
apical spines j-, \ ; no genicular spine on front femora.
Length of body 6 2 mm. ; length of tegmina 7 mm.
Fernando Po (L. Conradt, 1901) ; one example (Paris
Museum) .
Phyllodromia nimbata, sp. n.
Testaceous. Head with a rufous band between the
eyes and sometimes with a narrower band between the an-
tennal sockets. Pronotum trapezoidal ; lateral margins
hyaline ; disk testaceous, with rufous markings symmetri-
cally disposed. Tegmina clear testaceous ; ten costal veins,
five longitudinal discoidal sectors. Wings hyaline; medi-
astinal vein with two branches ; eight to nine costal veins,
their apices clavately incrassated ; ulnar vein with four
branches. Abdomen infuscated above, testaceous below, with
32
Mr. R. Slielford on somo
fuscous margins ; supra-anal lamina trigonal, slightly emar-
ginatc ; suhgcnital lamina triangularly produced, with two
styles; ccrci elongate, testaceous. Legs testaeeous ; front
femoi'a armed on anterior margin beneath with a series ol:
s])ines, the most distal short and serried ; apieal spines j-, j-, } ;
all the femora with gcnieular spines.
?. Head entirely rufo-tcstaccous ; supra-anal lamina
transverse ; suhgenital lamina ample, semiorbicular, posterior
margin slightly but widely cmarginate.
cf.
Length of body 11 mm. 8*1 mm.
Length of tegmina 10 mm. 9*8 mm.
Knelling, Sarawak, Borneo; five examples (Oxford
IMuseum). [No. 29.]
Idle species is undoubtedly closely allied to Phyllodromm
lityrifera^ StM, the type of which is now before me, but
differs in the following points: — It is smaller; the head is
not marked with three castaneous bands ; the coloration of
the tegmina is difierent ; the wings are clear hyaline and
their veins pale testaceous; the costal veins are more nume-
rous; the supra-anal lamina is slightly more produced and
its apex is emarginate ; the subgenital lamina is narrower.
It is quite evident that de Saussure correctly identified
P. liturijera, StM (Mel. Orthopt. ii. p. 56, 1869), and his
detailed description of the species is perfectly accurate in
every point.
Phyllodromia nebulosa, sp. n.
^ . Head testaceous ; four longitudinal lines on the vertex,
three indistinct transverse bands on the face, castaneous.
Pronotum trapezoidal, lateral margins hyaline, disk marbled
with castaneous and testaceous. Tegmina clear testaceous,
with numerous irregular castaneous markings occurring
between the veins ; ten to eleven costal veins, five longitu-
dinal discoidal sectors. Wings hyaline ; veins testaceous,
mediastinal vein with two branches ; eight costal veins, their
apices clavately incrassated ; ulnar vein with four branches.
Abdomen infuscated ; supra-anal lamina triangular; sub-
genital lamina ample, semiorbicular, v\ith two styles, the
margin of the lamina emarginate at their points of insertion ;
cerci long, testaceous, base and apex fuscous. Legs testa-
ceous, the tibiae banded with fuscous ; armature of femora
as in the preceding species.
$ . All the castaneous markings on the head more distinct ;
33
neio Species of l^latti(U\3.
snpra-anal lamina transverse, slightly emarginate ; subgenital
lamina very large, its posterior margin slightly and asym-
metrically emarginate.
d. $.
Total length 10 mm. 11 mm.
Length of tegmina 9 mm. 8 mm.
Kuching, Sarawak, Borneo ; three examples (Oxford
Museum). [No. 30.]
The small size of the species and the marbling of the
tegmina with darker markings serve to distinguish it from
any of the described Oriental forms ; its nearest ally appears
to be P. ignohiliSy Wlk., from Sula Islands.
Phyllodromia Hewittiy sp. n.
c^. Fulvo-castaneous. Antennae equal to total length of
body, fuscous except at base. Pronotum trapezoidal, smooth,
shining, sides deflexed, not covering vertex of head, with
obscure darker markings, posterior margin slightly produced.
Tegmina with radial vein bifurcated, twenty-one costal veins,
discoidal area with nine longitudinal sectors. Wings with
marginal area somewhat coriaceous ; mediastinal vein with
five branches, radial vein bifurcated from near base; twelve
costals ; ulnar vein with three branches ; a prominent
triangular apical area. Front femora with eleven long spines
on anterior margin beneath, the more distal closely set
together ; formula of apical spines -{ ; no genicular
spine on front femora. Supra-anal lamina triangular ; sub-
genital lamina produced, highly irregular in appearance ; no
styles.
Total length 21 mm. ; length of body 18 mm. ; length of
tegmina 18 ram.
Kuching, Sarawak. [No. 27.]
This species, which I have pleasure in naming after
Mr. J. Hewitt, Curator of the Sarawak Museum, has all the
appearance of an Ischnopteray but the wing- venation is that
of a typical Phyllodromia ; it is, perhaps, most nearly allied
to P. ferrugineay Br.
Phyllodromia (?) japonicay sp. n.
$ . Rufo-castaneous, nitid, broad, short. Head with a
darker mark between the eyes ; antennae longer than the
body, fuscous except at base. Pronotum trapezoidal, sides
deflexed, not quite covering vertex of head, posterior margin
Ann. <&; Mag. N. Hist. Ser. 7. Vol. xix. 3
34
j\Ir. K. Sliclfoi’d on some
very angled ; a sliort black line on each side in
front. Tegmina not longer than abdomen ; radial vein bi-
furcate; eleven costals ; seven axillaries ; discoidal sectors
oblique. Wdiigs suffused Avith rufo-testaceous ; mediastinal
vein with three branches, radial vein bifurcated ; eight
costals; ulnar vein with three rami reaching apex of wing
a id two irregular and anastomosing rami which are directed
towards the apex of wing but do not reach it ; no apical
triangle. Abdomen dark castaneous above and beneath,
margined with paler ; supra-anal lamina triangular ; cerci
moderate ; subgcnital lamina ample. Legs rufo-testaceous,
all the femora strongly armed ; no genicular spine on front
femora ; formula of apical spines j, { .
Total length 15’2 mm. ; length of tegmina 12 mm.; pro-
notum 5*5 x 7'9 mm.
Kiou-Kiou, Oshima, Japan ; three examples (Paris
Museum).
This is a very puzzling species, and I include it in the
genus Phyllodromia with considerable doubt ; in general
appearance it approaches the Madagascar species of Allacta,
but it certainly does not belong to that genus. The wing-
venation is suggestive of the genus Ischnoptera, but does not
strictly conform to that type nor to the wing-venation of
Phyllodromia.
Genus Pseudophyllodkomia, Br.
Pseudophyllodromia elegans^ Shelf.
$ . Head flavo-testaceous ; a rufous band on the vertex
and between the eyes ; antennae fuscous except three basal
joints, which are testaceous ; second and third joints of
maxillary palpi black. Pronotum transversely elliptical,
anteriorly truncate, posteriorly very slightly angulate ;
lateral margins broadly, posterior margin narrowly hyaline
testaceous ; disk dark castaneous, with four testaceous
markings ; an anterior median line, stopping short before
the middle of the disk ; two dots on each side of the middle
line in the centre of the disk, a median posterior dot, in
addition a rufescent marking on each side of the disk in
front. Tegmina fusco-castaneous, marginal area and area
between radial and anal vein testaceous hyaline ; twelve to
thirteen costal veins, five discoidal sectors, five axillary veins.
Abdomen fusco-castaneous above; supra-anal lamina short,
trigonal ; abdomen beneath and legs flavo-testaceous ; sub-
genital lamina ample, tipped with fuscous, its posterior
margin cleft in the middle ; cerci moderate, testaceous.
r\ero Species o f Blatticl.'c. 35
Total length 11*5 mm. ; length of body 9 mm. ; length of
tegmina 9' 2 mm.
Maroni, Freneh Guiana (F. Gemj, 1903) ; one example
(Paris Museum).
F. histrio, Sauss., appears to be the nearest ally of this
speeies, which is ivcll marked by the dark tegmina with one
liyalinc band.
Genus Pseudectoiua^ Sauss.
Pse.udectohia AUuaudi, sp, n.
$ . Rufo-testaceous. Antennae and mouth-parts testa-
ceous. Pronotum coA^ering vertex of head^ trapezoidal ;
anterior margin truncate, posterior margin obtusely angled,
lateral margins pellucid, with an opaque testaceous sub-
marginal band bordered inw^ardly by a rufous suffusion.
Tegmina convex, nitid, venation of anal and discoidal fields
obsolete; anal vein deeply impressed, arcuate, reaching
sutural margin at a point on one half of its length ; fifteen
costals. Wings small, hyaline ; radial vein bifurcated near
its apex ; twelve costal veins ; ulnar vein triramose ; apical
triangle well defined, projecting beyond the anterior margin.
Abdomen broad ; supra-anal lamina triangular ; subgenital
lamina semiorbicular, projecting slightly beyond the supra-
anal lamina. Front femora with eleven spines along the
anterior margin beneath ; hind femora with four pairs of
spines ; genicular spines and a pair of apical spines on each
femur.
Total length 10 mm. ; length of tegmina 7 mm.; pro-
notum 3x5 mm.
Diego Suarez, Madagascar {Alluaud, April 1896) ; one
example (Paris Museum).
De Saussure created this genus or subgenus for the recep-
tion of the species Luneli, Sauss., liturifera, Stal, insularis,
Sauss., regarding the apical triangle and branched ulnar
vein of the wings of prime importance. As I have shown
(Trans. Ent. Soc. London, p. 231, 1906), the presence of an
apical triangle in the wings is a feature that appears in so
many subfamilies of Blattidae, that, taken by itself, it is of
small value for purposes of generic distinction. I do not
believe that liturifera, Stal, and insularis, Sauss., are con-
generic, and 1 have seen the types of both species ; Luneli,
Sauss., is congeneric with liturifera, Stal, and I do not see
how either species can be separated from the genus Phyllo-
dromia ; insularis^ Sauss.^ is a broad convex insect, very
]\Ir. R. Slielford on some
r,G
(lifTcrent from the other two species, and may well be selected
as the type of tlic genus, which can be distinguished by the
following cliaractcrs : —
Broad, convex insects ; the tegraina not projecting much
beyond the tip of the abdomen, their venation sometimes
obsolete in the anal and discoidal fields. Wings with an
apical triangle, the ulnar vein ramose. Femora generally
strongly armed. Supra-anal lamina variable, but usually
produced.
Type of the genus P. insulans, Sauss.
The other species of the genus are P. hipunctata, Wlk.,
P. adimonialis, Wlk. ( = Lupparia adimonialis)^ possibly
P. latipennis, Br. ( = Phyllodromia latipennis), and the new
species described above. Of the species previously included
in the genus, P. pallidula, Boh, and P. voeltzkowiana, Sauss.
& Zehnt., have the supra-anal lamina produced, the abdomen
is missing in P. Luneli, Sauss., and in P. intermedia, Sauss. &
Zehnt. ; it is probable that the shape of this tergite is as
variable as in the’ genus Phyllodromia, and too much reliance
should not be placed on its transverse form in P. subpectinata,
Sauss. & Zehnt., and P. antiguensis, Sauss. & Zehnt. In all
these species the armature of the femora is most variable.
They cannot be placed in the genus Theganoptery x on account
of the ramose character of the vena ulnaris alarum {cf. Trans.
Ent. Soc. p. 232, 1906), but the presence of a triangular apical
area in the wings does not forbid their entry in the genus
Phyllodromia, seeing that this character does occur in several
well-marked and well-recognized species of that genus.
P. punctulata , Sauss. & Zehnt., must be referred to the genus
Theganoptery X,
Subfam. Nyctiborin^.
Nyctibora bicolor, sp. n.
? . Head entirely black, with a scanty erect pubescence
on the front ; ocelli minute, testaceous ; antennae incrassated,
black, apical joints rufescent, densely pubescent. Pronotnm
transversely elliptical, anterior border not nearly covering
vertex of head ; posterior border more arcuate than anterior,
covering the scutellum; disk with tw^o converging impressions,
flavo-testaceous, with a shield-shaped black mark on the disk,
posterior margin with a fine black line. Tegmina coriaceous,
serio-punctate and reticulate between the raised veins, not
pubescent ; the basal three fifths flavo-testaceous, the apical
two fifths dark castaneous; radial vein black throughout its
new i,Sjn<?c/e.9 oy’ Blattidio. 37
length, sutural margins black, anal vein deeply impressed,
mediastinal vein sending several l)i’anelies to the margin,
costals numerous. Wings dark castaneous, ulnar vein with
nine branches. Abdomen entirely black, except apical halt*
of subgenital lamina, which is flavo-testaceous ; supra-anal
lamina triangular, produced ; subgenital lamina ample, its
lateral margins shortly produced ; cerci black (mutilated).
Legs black ; front femora unarmed, mid femora with four
spines on posterior margin, none on anterior margin ; hind
femora with two spines on posterior margin, none on anterior
margin ; genicular spines on second and third femora; formula
of apical spines f , -}-, 1.
Total length 26’8 mm.; length of body 17 mra. ; length
of tegmina 23'8 mm. ; pronotum 6 mm. X 8’5 mm.
Yarimaguas, Peru [Salle, 1886) ; one example (Paris
Museum).
This species, that described below, and N. crassicornis,
Burm., should probably be included in a new genus, charac-
terized by the incrassated antenuje, shape of the pronotum,
and sparse armature of the femora.
Nyctibora nigrocincta, sp. n.
^ . Head entirely black ; antennae black, except apical
joint, which is castaneous, incrassated, pubescent. Pronotum
as in the preceding species, but with the lateral and posterior
margins slightly reflected and with a very scanty recumbent
pubescence, yellow, with a shield-shaped black mark on the
disk. Tegmina long, coriaceous at base and reticulate, not
pubescent ; four branches to the mediastinal vein, costals
numerous ; colour yellow, radial vein at base, a broad streak
on sutural margin of anal field, a broad band from near the
costal margin to the sutural margin in the apical third, black;
apex castaneous. Wings fusco-castaneous ; a broad pre-apical
yellow band ; costals irregular, six branches to the ulnar vein.
Abdomen black above and below; supra-anal lamina tri-
angular ; subgenital lamina produced, very convex, with two
styles ; cerci black. Legs black ; front femora with no spines
beneath ; mid femora with three to four spines on posterior
margin, none on anterior margin ; hind femora with five
spines on posterior margin, none on anterior margin ; geni-
cular spines on the mid and hind femora ; formula of apical
spines f
$ . Similar to S ) but rather larger, the black band ex-
tending right across the tegmina, castaneous apex of tegmina
more extended, subgenital lamina as in N. bicolor, mihi.
?>H
IMr. Iv. Slid ford on some
S. Total Icngtli 24 rum.; Icngjtli of body 17‘5 mm.;
Icii^’tli of tej^mina 20 mm. ; pronotum 4’2 mm. X 6 mm.
? . Total lenjijtli 27 mm. ; length of body 20*2 mm. ;
length of tegmina 21 mm. ; pronotum 5 mm. x 8 mm.
Colombia ; four examples (Oxford Museum).
Subfam. Etilamfrinm.
Genus Notolampra, Sauss.
Notolmnpra antillarum, sp. n.
cf . Castaneous. Head and antennae testaeeous, the
former with a few seattered castaneous punetures. Pro-
notum not punctate^ castaneous, the lateral borders broadly,
the anterior margin narrowly, testaceous, semiopaque, with
a few scattered fuscous or castaneous dots. Tegmina
castaneous, the lateral borders testaceous, semiopaque with
scattered castaneous dots, these borders are continuous with
the pronotal testaceous borders and are broadest at the base,
narrowing to the apex ; radial vein marked at its base by a
dark line, no other veins visible ; surface of tegmina serio-
punctate. Wings rufo-testaceous ; intercalated apical tri-
angle small. Abdomen rufo-testaceous ; supra-anal lamina
triangular, slightly notched ; subgenital lamina trigonal,
somewhat asymmetrical with two styles; cerci short. Legs
testaceous spotted with castaneous ; front femora with two
spines only at base of anterior margin beneath, rest of
anterior margin occupied by piliform setae, one spine on
posterior margin ; mid and hind femora with two spines on
anterior margin, four on posterior margin; formula of apical
spines o ’ genicular spine on front femora.
Total length 19 mm. ; length of tegmina 14 mm. ; pro-
notum 6 mm. X 7*5 mm.
Trinite, Martinique [A. Bourgouin, 1901) ; one example
(Paris Museum).
The species is most closely allied to Notolampra punctata,
Sauss., from Brazil, but differs in the testaceous borders of
the tegmina and in its proportions.
Genus Apsidopis, Sauss.
Apsidopis Waltacei, sp. n.
$ . Pale testaceous. Frons concave and transversely
striated ; a chevron-shaped depression at base of clypeus and
two marks between the eyes castaneous ; antemue fuscous
new Species of Blaf lid^e. oO
except basal joint ; eyes 1 J mm. a[)art. Pronotum cuoiillate,
of the form characteiistic of the gcnus^ punctate and with
nnmcrons minute fuscous or castancous rnaculte. Tej^rnina
semicoriaceous, densely serio-punctate between the veins in
the basal two thirds, in the apical third the punctures merge
into quadrangular interspaces between numerous reticulated
interstitial veins, a few minute fuscous dots are scattered
over the tegmina ; mediastinal vein with ten branches; seven
ramose costal veins ; nine anal veins, the first ramose. Wings
angulated at the apex as in the genus Derocar'dia, Sauss. ;
marginal area testaceous with fuscous spotting at apex, the
apex semicoriaceous; mediastinal vein multiran)ose ; costal
veins irregular, the interspaces filled by reticulated interstitial
veins ; ulnar vein with nineteen branches. Supra-anal lamina
ample, prominent, bilobed ; subgenital lamina semiorbicular ;
cerci slender and short. Front femora with five spines on
middle of anterior margin beneath, piliforrn* setse extending
from them to apex, two spines on posterior margin; mid
and hind femora with three to four spines on each lower
margin; formula of apical spines {, J; minute genicular
spines on mid and hind femora, none on front femora ;
posterior metatarsus shorter than remaining joints, its
pulvillus produced proximally.
Length of body 30 mm. ; length of tegmina 35 mm. ; pro-
notum 11 mm. X 12*5 mm.
Sarawak (Wallace; Wilson Saunders collection, Oxford
Museum) .
The species is close to A. oocyptera, Wlk., also from Borneo,
which exhibits the same characters of punctuation of the
tegmina; but A. oxyptera is smaller, more rufous in colour,
the proportions of the pronotum are different and the pro-
notum is less closely punctate, but more densely covered
with castaneous dots. Both species can be distinguished
from A. acutipenniSj Sauss., by their larger size.
Subfam. Blattin^.
Genus Blatta, L.
Blatta Rothschildi, sp. n,
S . Rufo-castaneous. Head with four darker markings
between the eyes; ocelli and clypeus testaceous; antennae
much longer than the body, first two joints and apical third
rufo-castaneous, remainder fuscous. Pronotum trapezoidal ;
anterior and posterior borders truncate, flavo-testaceous, a
40
^Ir. K. Sliclford on some
castaiicous marking like an inverted W on the disk, a central
testaceous line. Tcgrnina abbreviated, not extending much
beyond the second abdominal tergite, surface reticulate, anal
vein reaching internal posterior angle of tegmina. Wings
rudimentary. The first and second abdominal tergites flavo-
testaeeous, third to fifth flavo-testaceous with castaneous
lateral and posterior borders; sixth rufo-castaneous, enlarged;
seventh flavo-testaceous with central castaneous macula,
narrow, posterior margin sinuate and slightly emarginate in
the middle; supra-anal lamina quadrate, broadly emarginate,
rufo-castaneous with a testaceous macula at the postero-
lateral angles. Cerci black, apices rufo-castaneous. Abdo-
men beneath rufo-castaneous, lateral margins castaneous and
a castaneous stigma on second to fourth sternites ; sub-
genital lamina notched on each side, the long slender styles
springing from the notches. Legs rufo-castaneous. Meta-
tarsus scarcely equal to remaining joints, spined beneath,
its pulvillus minute.
$ . Head black ; ocelli, clypeus, gense, vertex flavo-
testaceous ; antennse rufo-castaneous. Pronotum as in ^ ,
but the discal black marking much enlarged, so that it
occupies all the disk, leaving only a narrow sinuate flavo-
testaceous margin, no central testaceous line. Tegmina
squamiform, not extending beyond metanotum, black, with
a yellow line at base ; mesonotum, metanotum, and first five
abdominal tergites black, with a broad central transverse
flavo-testaceous band; sixth tergite enlarged, concavely
depressed, black with flavo-testaceous lateral and posterior
margins ; seventh tergite somewhat triangularly produced,
black, apex slightly emarginate, flavo-testaceous ; supra-anal
lamina produced, narrower than in ^ , broadly emarginate.
Abdomen beneath and legs black, disk of abdomen rufo-
castaneous ; COX86 margined outwardly with flavo-testaceous,
tibial spines and tarsi castaneous.
S . Total length 21 mm. ; length of tegmina 10 mm. ;
pronotum 5*8 mm. x 7*2 mm.
$ . Total length 26 mm. ; length of tegmina 4 mm. ;
pronotum 8 mm. x 10 mm.
South of Lake Rudolph, Brit. E. Africa {Maurice de
Rothschild, 1905) ; five examples (Paris Museum).
The nearest ally of the species appears to be J3. nianca,
Gerst., from W. Africa.
neAO Species of Blattlilac.
41
Subfam. Oxyhaloin.e.
Genus Oxyhaloa, Br.
Oxylialoa variabilis, sp. n.
? . Rufo-castaneous. Vertex of head rufo-castaneous, a
clear testaceous band between the antennae, genae testaceous ;
frons, clypeus, labrum, palpi, and antennae shining black.
Pronotum with two oblique impressions anteriorly, with a
few minute punctures from which spring short slender hairs.
Tegmina with the veins fuscous, sparsely pubescent, very
variable in length, in some examples reaching tip of abdomen,
in others lanceolate and extending no further than the third
tergite ; thirteen costals ; discoidal field reticulate. Wings
as variable in length as the tegmina, flavid at base, the rest
infuscated; veins fuscous, ulnar vein with eight to nine rami,
the basal ones transverse. Abdomen broad, black above,
the margins of the segments narrowly rufous, beneath rufo-
castaneous ; supra-anal lamina short wdth rounded posterior
angles, not emarginate ; subgenital plate projecting beyond
the supra-anal lamina, fuscous, ample, its margin sinuated ;
cerci short, fuscous, tipped with rufous. Legs black, apices
of coxas and femora rufous, tibial spines rufous.
Total length from 16 mm. to 13*5 mm. ; length of body
from 16 mm. to i2'5 mm. ; length of tegmina from 11 mm.
to 8 mm. ; breadth of pronotum from 6*2 mm. to 5 mm. ;
length of pronotum from 4*5 to 4 mm.
Interior of Djibouti (Hermann) ; one example (Paris
Museum).
This is the smallest species of the genus, and is remarkable
on account of the variation in size of the wings and tegmina ;
apparently this variation bears no relation to the variation
in size of the individual, for one of the smallest specimens
has long tegmina and one of the largest has these organs
much reduced.
Genus Paraplecta, nom. nov.
(= Cii'phisy StSl.)
The name Cirphis, created by Stal in 1876 (ffifv. Vet.-
Akad. Forh. xxxiii. p. 74) for a cockroach (C. pallipes) from
Damara Land, is preoccupied, having been applied by VValker
in 1865 to a genus of Noctuid moths.
42
Mr. R. Slielford on some
Paraplecta adhiopica, sp. n.
. Castancous, smooth, riitid ; vertex of head not covered
by pronotnm ; ocelli, apex of clypeus, mouth-parts, and
antennae testaceous ; minutely punctured. Pronotum trape-
zoidal, with rounded posterior angles, minutely punctured;
])Ostcrior margin truncate, exposing the scutellum Tegmina
scmicoriaccous, barely reaching apex of abdomen ; eleven or
twelve costals ; discoidal field reticulate, anal vein impressed ;
eight axillaries. Wings with a large apical reflected area,
two fifths of total wing-length, its basal margin obtusely
angled; costals highly irregular and obsolescent ; median vein
consisting of two parallel branches, with one or two trans-
verse venulse connecting them; ulnar vein with seven branches.
Abdomen castaneous above, supra-anal lamina produced ;
abdomen rufo-castaneous below, subgenital lamina asym-
metrical with one style (the left) ; cerci short, acuminate,
4-jointed. Legs testaceous ; femora spineless, tarsal claws
without arolia.
$ . Similar to cT , hut larger, tegmina and wings (when
folded) not extending beyond the sixth abdominal tergite ;
supra-anal lamina produced quadrately; subgenital lamina
ample, produced, narrowed posteriorly.
^ . Total length 9 mm. ; length of tegmina 8 mm. ; pro-
notum 3 mm. X 3*2 mm.
$ . Total length 11 mm. ; length of tegmina 7 mm.; pro-
notum 3*5 mm. x 3*8 mm.
Fernando Po (L. Conradt, 1901) ; six examples (Paris
Museum).
The species can readily be distinguished from P. pallipes,
Stal, by the wing-structure : in StaFs species there is a con-
spicuous triangular apical area which in P. (Ethiopica has
become extended to form an apical reflected area ; the
venation is very similar in both species, but in pallipes the
costals are better marked and the rami of the ulnar vein are
more numerous, the double median vein is common to both
species.
Genus Choristima, Tepper.
Choristhna, Tepper, Trans. Roy. Soc. S. Austral, xix. p. 165 (1895).
AphlehideAi, Brancsik, Jahresh. Ver. Trencsin. Com. xix. Si xx. p. 56
(1897).
Kirby in his ^ Synonymic Catalogue of Orthoptera ^ (1904),
following Brancsik, places Aphlebidea in the Ectobinse ; but
as the femora are unarmed beneath and a triangular apical
field is present in the wings, the genus falls naturally into tlic
43
new Species 0/ Blatlidaj.
subfamily Oxylialoiii8c( = Plcctopterinse). Aphlebidea is un-
doubtedly the same as Tepper’s genus Choristima^ described
two years previously, and A. Bnmneri, Brancs., if not
identical with Choristinia palerucoides, Wlk., is most closely
allied. Blatta- apicifera, Wlk. (Cat. 151att. B. M. p. 110,
1868), is the male of C. galerucoides, Wlk. : the type, which
is in the British Museum, is in extremely poor condition, the
abdomen and antennae being missing; it is smaller than
the female, the tegmina and wings are relatively longer and
would, 1 imagine, extend beyond the tip of the abdomen.
Tepper^s diagnosis of the genus is extremely brief, but
Brancsik^s description of Aphlebidea is detailed enough to
render it readily recognizable. The species included in the
genus may be distinguished as follows : —
a. Subgenital lamina in 5 somewhat
cucullate.
h. Rufous C. B runner i, Brancs.
hh. Testaceous C. Kershatoi, Tepp.
aa. Subgenital lamina in $ not cucullate.
h. Piceous above C. hydrophoroides, Wlk.
bh. Rufous C. galerucoides, Wlk.
(syn. C. hftyemis, Tepp. ;
C. apicifera, Wlk.).
Chorisoneura pectinata, Sauss. (Mel. Orthopter. iv. p. 131,
1872), may be referred to a new genus, on account of the
fusion of the radial and ulnar veins of the tegmina— a con-
dition which obtains also in Ectobia, Westw., but not in the
genera Chorisonew'a, Br., or Choristima, Tepp., to which it is
most nearly allied.
Genus Ectoneura, nov.
Allied to Choristima, Tepper, but the radial and uliiar
veins of tegmina fused and emitting oblique veins to both
margins. Tegmina and wings somewhat reduced in the
female, but not to so great an extent as in Choristima ;
triangular apical area large and conspicuous. Supra-anal
lamina transverse or slightly produced; subgenital lamina
of the male narrow, triangular, of the female large and
subquadrate. Femora spineless beneath, except for apical
spines, the formula of which is -J, j, } .
Type, jE. pectinata, Sauss.
Ectoneura figurata, sp. n.
. Head fusco-castaneous with a pale testaceous band
44
jMr. R. Slielford on some
l)etwcen the eyes ; antennae testaceous. Prothorax trans-
versely elliptical, margins hyaline; disk fusco-castaneous with
the centre testaceous, on which ai*e some fuscous markings.
Tegmina hyaline with pale fuscous spots along the veins;
twelve costal veins, nine discoidal rami. Wings hyaline,
veins fusccscent ; nine costal veins ; medio-discai area twice
as broad as medio-ulnar, crossed by several irregular trans-
verse bars ; nlnar vein simple ; first axillary vein biramose,
upper branch irregularly bifurcate. Abdomen fuscous, with
testaceous markings; cerci long, fuscous, with a broad
testaceous band. Legs testaceous.
Total length 8 mm. ; length of tegmina 6"5 mm.
Five males without locality (Oxford Museum).
It is possible that this is the Blatta marcida of Erichson,
a species placed by Brunner with some doubt in the genus
Ectobia, Westw.
Genus Chorisoneura, Br.
Cliorisoneura Brunneri, sp. n.
$ . Head fuscous, vertex rufo-castaneous ; antennse fuscous
at base, the remainder testaceous. Prothorax transversely
elliptic, disk fuscous, margins hyaline. Tegmina castaneous
with hyaline costal margin and with fuscous humeral vitta,
veins white ; sixteen costal veins, very irregular; median vein
distinct, longitudinal, discoidal vein with four irregular
branches ; the whole surface of the tegmina is much re-
ticulated, and the veins are rather obscured thereby ; the
part of the right tegmen covered by the left is infuscated.
Wings fuscous, the edge of the marginal field yellowish ;
eleven costal veins, their ends swollen, joined by transverse
bars ; medio-discai field crossed by several transverse bars ;
ulnar vein arcuate, simple; apical triangle unevenly divided,
its base acutely angled, its apex subtruncate. Abdomen
fulvous, legs testaceous. Cerci long, testaceous.
Total length 8'5 mm. ; length of tegmina 7‘5 mm.
Rio Grande do Sul. Two examples labelled in Brunner
V. WattenwyPs band writing Chorisoneura, sp. n.'’^ (Oxford
Museum) .
The species appears to be distinct from anything described ;
perhaps it shows most affinity with C. anomala, Sauss. &
Zehnt.
Chorisoneura morosa, sp. n.
. Head rufo-fuscous, with a pale narrow transverse
line between the eyes ; antennsc (mutilated) testaceous ;
45
new Species o/Blattida3.
pronotum with the disk fuscous, marj^ins hyaline. Tegmina
pale fulvous, surface not reticulated, veins prominent and
white, a fuscous humeral vitta ; twelve costal veins, the last
two biramose; the humeral vein also giving off four rami to
the sutural margin, the discoidal vein giving off only three;
ulnar vein triramose. Wings infuscated, edge of marginal
field yellow ; fourteen costal veins, medio-discal field crossed
by numerous transverse bars ; ulnar vein forked at apex,
axillary vein triramose; apical area unequally divided,
acutely angled at base, its apex subtruncate. Body and legs
testaceous beneath.
Total length 9 mm. ; length of tegmina 7 mm.
Cachabi, Ecuador (W. F. H. Rosenberg coll., Dec. 1896) ;
one example (Oxford Museum).
The species is evidently allied to C. traiislucida, Sauss.,
from Mexico.
Subfam. Panchloein^.
Genus Zetobora, Burm.
Zetobora lata, sp. n.
^ . Head black ; labrum golden ; antennse at base black,
nitid, the rest fuscous, villose. Pronotum with anterior
margin strongly rounded in the middle, less so laterally, the
margin slightly reflected, the posterior margin only slightly
curved, the postero-lateral angles slightly notched ; disk with
a humeral carina on each side, the ‘Miood'’-’ with a few
tubercles, nitid, the sides with numerous tubercles of various
sizes, a few granules on the front ; castaneous except for a
testaceous hyaline patch, semicircular in shape, in front ;
scutellum with central carina and a few punctures. Tegmina
broad, scarcely exceeding the abdomen in length, testaceo-
hyaline, anal field and humeral vein castaneous ; marginal
field very broad, its outer border slightly thickened and
margined ; mediastinal area with large shallow punctures ;
anal field reticulate-punctate ; discoidal field reticulate ;
apex broadly rounded. Supra-anal lamina quadrate, notched ;
subgenital lamina produced, rather asymmetrical, with one
style (the right). Abdomen castaneous ; the dorsal tergites
with their posterior angles strongly produced backwards
and projecting considerably beyond the sternites, very much
as in the genus Capucina, Sauss.
46
Mr. 11. Sbelford on some
Total Icnj^th 22 mm. ; length of tegmina 25 mm. ; breadth
of tegmeri 13 mm. ; pronoturn 10 mm. x 16*2 mm.
The species is characterized by its great breadth in pro-
portion to its length ; the notched postero-lateral angles of
the pronoturn are also distinctive.
Subfam. Corybinm.
Genus Euthyrrapha.
Euthyrrapha bigeminata, sp. n.
$ . Very similar to E. pacifica^ Coq., but the pronoturn
entirely fuscous, a round pale testaceous spot on eaeh tegmen
beyond the middle ; the abdomen beneath is orange with the
apex fuscous. The tegmina at base are slightly rugose,
minutely punctate, and furnished with an erect pubescence ;
the apex of the tegmina appears velvety. Legs fuscous,
coxal joints and tibial spines castaneous. Subgenital lamina
strongly carinate.
Total length 7’5 mm. ; length of body 5 mm. ; length of
tegmina 6 mm.
Ivory Coast, W. Africa (G. ThoirSj 1901) ; one example
(Paris Museum).
Subfam. Ferisprmrinjj:.
Genus Paranauphceta, Br.
Paranaupheeta Brunneri, sp. n.
$ . Closely allied to P. rufipes, Haan, but smaller. Head
with three maculae on the vertex ; the ocelli, genae, clypeus,
and basal joints of the palpi testaceous; apical joints of
antennse not testaceous. Pronoturn with the testaceous
margins much narrower than in P. rufipes. Tegmina and
wings as in P. rufipes. Abdomen above less strongly marked
with testaceous ; subgenital lamina ample, produced, emar-
ginate. Coxae testaceous, castaneous at base ; femora
testaceous at base, remainder castaneous ; tibiae and tarsi
r uf o-castaneous .
Total length 20 mm.; length of body 18 mm. ; length of
tegmina 17T mm.
Kuehing, Sarawak [Shelford, 1900) ; one example (Oxford
Museum).
47
nen) Sj.ecics of l>lattida3.
Genus Eustegasta_, Gerst.
Eusiegasta agrilidina, sp. n.
^ . Black, nitid, with dark green metallic reflections.
Head entirely of this colour except the labrum, which is
testaceous ; antennje with five basal joints, black, nitid,
remainder fuscous. Prouotum of typical form, broadly
bordered laterally with orange. Tegmina with an orange
spot at base of discoidal area, another in apical third of
marginal area. Abdomen above fuscous, broadly margined
with orange, beneath orange; supra-anal lamina short,
trigonal, fuscous ; subgenital lamina small, asymmetrical,
without styles, black ; cerci pale testaceous. Front legs
orange ; mid and hind coxae black, outwardly bordered with
white, femora and tibiae orange, all the tarsi fuscous; front
femora with two spines on anterior margin beneath ; mid
femora with one spine on anterior margin, none on posterior
margin ; hind femora with one spine on anterior margin,
three on posterior margin ; formula of apical spines -J, 1 ;
no genicular spine on front femora.
Total length 1.2-5 mm.; length of body 10 mm.; length
of tegmina 10*1 mm.
N'^Kogo, Freueh Congo {H. Bonnet, 1903) ; one example
(Paris Museum).
The nearest ally of the species is E. metallica, Sauss.
Eustegasta variegata, sp. n.
(f . Head testaceous ; occiput, a cordate patch on the
frons, the genae, two spots on the clypeus, and the palpi cas-
taneous or black ; antennae black, the six basal joints nitid.
Prouotum testaceous, the disk rufo-castaneous, with two
black longitudinal vittae of irregular shape. Tegmina rufo-
castaneous ; mediastinal area and two thirds of marginal area
clear testaceous, the stripe at its termination expanding into
a spot that extends on to the outer part of discoidal field ;
a humeral stripe ; the anal vein and a longitudinal stripe in
the anal field black, faint indications of a testaceous spot at
base of discoidal field. Wings flavo-hyaline ; median vein
bifurcate ; ulnar vein with ten rami, four of which reach the
apex of the wing. Abdomen orange above and beneath ;
supra-anal lamina quadrate, posterior angles acute, not
emarginate ; subgenital lamina asymmetrical, with one style,
the right ; cerci orange. Coxse black, outwardly bordered
with testaceous; front femora castaneous, the other femora
48 ^Ir. H. Slielford on some new Species o/Blattida3.
and all tlie tibia3 davo-testaceous; tibial spines rufo-castaneous^
a[)ical joints of tarsi fuscous ; femoral spines as in preceding
species, except that the mid femora have no spines beneath.
Total length 13'8 mm. ; length of body 10 mm. ; length
of tegmina 10*8 mm.
Congo [Dyhowski, 189G) ; two examples (Paris Museum).
Nearest to E. Lueci, Dom.
Genus Ellipsica, Sauss. & Zehnt.
Ellipsica rugosa^ sp. n.
$ . Blacky narrowly elliptical^ convex. Head punctate;
clypeus_, mouth-parts, and antennse flavo-testaceous. Sides
of the thoracic tergites strongly deflexed, their posterior
angles acute and produced backwards, their surface beset
with tubercles, between which are numerous punctures ; on
the meso- and metanotum the tubercles are in a double row,
on the pronotum they are less regularly placed. Abdominal
tergites transversely divided by a sulcus into two unequal
portions ; the anterior narrow portion is im punctate and
smooth, the posterior portion is marked by a double row of
punctures followed by a double row of tubercles ; the
transverse sulci are laterally very deep, especially in the
posterior segments, and one row of punctures lies in these
sulci; on the seventh tergite the tubercles are larger and
less regularly arranged than in the preceding segments.
Supra-anal lamina quadrate, margined, tuberculate. Abdo-
minal sternites transversely divided by sulci like the tergites,
punctate but not tuberculate; subgenital lamina ample,
punctate. Legs black, tarsi flavo-testaceous, arolia large.
Total length 15 mm. ; pronotum 5 mm. x 7 mm.
West coast of Madagascar {Lantz, 1882) ; two examples
(Paris Museum).
The species is quite unlike any other of the genus in its
tuberculate characters.
Genus Pseudoglomeris, Br.
Pseudoglomeris magnifica, sp. n.
$ . Brilliant metallic green or blue-green. Head punc-
tate ; eyes 1 mm. apart ; base of antennse and palpi
flavid, apical half of antennse fuscous, middle section
castaneous. Pronotum semiorbicular, posterior angles back-
wardly produced ; anterior border margined and slightly
Colonel C. Swinhoe on new Ihterocera.
49
reflected, densely punctate ; anteriorly the punctures are
irregular, producing a I'eticulate appearance; a few irregular
smooth spaces on all the thoracic tergites ; thorax bcneatli
black, shining. Abdomen with large shallow punctures
above; lateral margins of fifth and sixth segments slightly
sinuate, four or five punctures in the sulci of third to sixth
segments ; supra-anal lamina quadrate, slightly concave,
angles rounded ; abdomen beneath mrieous, densely punctate,
lateral smooth stigmata on each segment ; subgenital lamina
ample, sinuate, striato-punctate. Cerci rufous. Coxce, apices
of femora, tibiae, and tarsi rufo-testaceous ; femora castaneous.
Total length 23 mm. ; pronotum 7‘8 mm. x 14 mm.
Tuyen-Quan, Central Tonkin (A, Weiss, 1901) ; a long
series of specimens in all stages of growth (Paris Museum).
This is one of the most brilliantly-coloured cockroaches
known to science ; its colour is reminiscent of the gorgeous
metallic-green Cetoniid beetles of the Eastern Tropics.
111. — New Eastern, Australian, and African TIeterocera.
By Colonel C. Swinhoe, M.A., F.L.S., &c.
Family Bombycida.
Andraca apodecta^ nov.
3" ? . Of a uniform ochreous-fawn colour, the male slightly
suffused with brownish and darker than the females ; shafts
of the antennm dark brown above, white at the sides, pectina-
tions of the male brown ; some whitish suffusion along the
outer portions of the costa in the male, not present in the
females : fore wings with three transverse indistinct grey
lines or thin bands — antemedial, medial, and discal, the two
former sinuous, the latter deeply angled outwardly below the
costa : hind wings with two medial lines and with the ground-
colour of the wings paling towards the costa : the hairs on the
abdominal margin dark brown, this margin curved inwards
but without any excision. Underside paler, lines as above.
Expanse of wings, ^ inches.
Patiang, W. Sumatra ; one male, two females.
The fore wings are not so produced as in the type species
of the genus, bipunctata. Walker, from India, and it is
without the excision of the abdominal margin of the hind
wing as in that species, but it has every other character of
the genus.
Ann. & Mag. N. Hist. Ser. 7. Vol. xix. 4
50
Colonel C. Swinhoe on new Easiern^
Family Aganiidae.
Asota pTiilippinensiSj nov.
(3'. Palpi black, first and second joints with white stripes;
frons white, with a black central spot ; head and body deep
chrome-yellow ; a black spot at the base of each antenna; a
black and white collar, a black thin band behind it; a black
spot on each shoulder, another on the thorax near the base of
the abdomen, a black stripe on the thorax at the base of the
fore wings, black dorsal spots on the first five segments of
the abdomen, and a double row on each segment below : fore
wings dark purplish grey, as in A. paphos^ with a deep
chrome-yellow subbasal patch, which does not touch the
costa, which is there black intersected by white, making four
square marks ; the immediate base is white, with a black
spot, followed by another on the inner edge of the chrome
patch ; an elongated spot on the lower side of the patch, an
angled spot on the middle of the outer side and a round spot
above it ; veins whitish, most prominently on the outer
portions of the wing : hind wdngs black, whitish above the
cell ; some whitish longitudinal short streaks in the disk ; a
long streak in the abdominal marginal space and an outwardly
recurved w hitish discal line or thin band, which ends near the
extremity of the long streak. On the underside the wings
are slaty grey for two thirds, slaty black with whitish veins
on the outer third ; a large patch or spot at the end of each
cell ; a small spot near the base on the fore wings and one
in the middle of the cell on the hind wings; the veins are
marked with slaty black on both wings ; the discal recurved
band is clear on the hind wings, and indications of a some-
what similar band on the fore wings; cilia white above and
below.
Expanse of wings 2/^ inches.
Cagayancillo, Philippines, Jan. 6, 1903 (type in B. M.).
A handsome insect, belonging to the papkos group.
Digama costimacula^ nov.
. Antennae simple ; palpi white, with black spots at the
tips of each joint ; frons, head, thorax, and fore wings dark
slaty grey, frons and head with white and black spots at the
sides ; abdomen pale dull pink, with dorsal grey dots : fore
wings with three prominent subcostal white spots very close
to the costal line, the spots more or less square, the middle
one the largest, the first smooth, one fifth from the base, the
last at one fourth from the apex : hind wings yellow, w’ith
51
Australian^ and African Fleterocera.
blackish-brown outer marginal border, terminating abruptly
one third from the anal angle. Underside : body and wings
yellow ; fore wings with a black cell-spot ; the outer third of
the wing dark slaty grey, the middle portion of its innor
margin extending- in an angle into the wing.
Expanse of wings inch.
Gold Coast (IK. H, Johnston) (type in B. M.).
Belongs to the Hearseyana group.
Digama lithosioides^ nov.
^ . Palpi white, with black bands at the ends of the last
two joints ; antennae simple ; frons and head white, two
black spots in front of the antsnnse, one between them, two on
the collar; thorax and fore wings dull ochreous grey, veins
grey and rather prominent ; hind wings and abdomen dark
ochreous ; the wings are rather longer than usual, and the
example looks like a Lithosid ; there are no markings above
or below, except a small black dot at the end of cell of the
fore wings below.
Expanse of wings 2 inches.
One male, Hills of Terta, Kilimanjaro, E. Africa (type
in B. M.).
Digama meridionalis^ nov.
^ . Antennse ciliated ; palpi grey, with blackish bands at
the ends of the last two joints ; head, thorax, and fore wing^
dark grey, much as in the common Indian form Hearseyana^
Moore; two black spots in front of the antennae, one imme-
diately behind, two on the collar, and two on the thorax in
front : fore wings with black spots on the costa, three inside
the cell, one at the base, and two beyond it below' the cell;
discal and medial transverse, brown, erect bands below the
cell and an outw^ardly curved, very highly dentated, discal,
brown band and similar but smaller dentations on the outer
margin ; abdomen and hind wings yellow, the latter without
markings ; the abdomen is more ochreous and darker in colour
than the hind wings and has black dorsal spots : the wings
are long ; fore wdng narrow, with the hinder margin curved
outw’ards before the middle. The underside is uniform pale
yellow ; wings without markings, except a black dot at the
end of the cell of the fore wings ; abdomen with lateral black
spots.
Expanse of wings inch.
Johannesburg [A, T. Cooke) ; one male.
4^
52
Colonel C. Svvinlioe on new Eastern^
Digama africana^ nov.
$ . Antennai of male bipectinate ; palpi grey, with black
bands at the tips of the last two joints ; frons and head white,
thorax greyish white ; two black dots on the collar, four in a
transverse row on the fore part of the thorax ; abdomen
ochreous, with dorsal black segmental dots : fore wings dark
ochreous grey, with a purplish tinge, one fifth at base white,
divided by a grey transverse line, containing a black spot in
the cell and one near hinder angle ; a white medial band,
consisting of a large white square patch containing a black
spot on the costa and a much smaller patch on the hinder
angle ; another white patch on the costa at two thirds, with a
tliin white band running down from it to the hinder angle,
angled outwards in its middle, with a curious incomplete
white circular mark near the inner side of the angle.
The above is a description of a female from Tonga ; the
otlier female and the male from Samburu have all the white
bands more or less traceable, but very obscure from greyish
suffusion ; the hind wdngs are dark ochreous yellow, with a
narrow band at the apex, not continued on the costa or outer
margin. On the underside all three examples are identical,
being entirely yellow, the fore wings having some traces of
the upperside markings and the costal spot rather prominent.
Expanse of wings, ^ inch.
Samburu, British East Africa {Betton) ; one male. Tonga,
British East Africa ; two females. Types in B. M.
Digama elongata^ nov.
^ . Antennae with the shafts white, pectinations black ;
palpi black, white beneath ; frons, head, body, and wings
white; frons with a large black spot, a small one on the top
of the head, three on the thorax down the middle, and three
on each side ; abdomen with a dorsal row : fore wings with
pale blackish broad bands, first basal, for one fifth with a
white mark at the base, its outer margin with an outward
dentation above the middle and extends in a streak along
the hinder margin, joined on the costa to the second band,
which has irregular margins on each side and runs obliquely
from the middle of the costa to the liinder margin one third
from the angle ; the third band is discal and runs down only
a short distance from the costa, and narrows down to two little
spots; the fourth band is composed of elongated spots on the
Australian^ and African ITeterocera. 53
outer margiti : liiiul wings dull white, with a rather broad,
grey, marginal band.
Expanse of wings ineh.
Eb Urm, British East Africa (Betto?i) (type in B. M.).
The fore wing*s longer than usual in the genus.
Family Acontiidae.
Eublemma acarodes^ nov.
. Palpi, frons, head, and fore part of the thorax pure
white, rest of the thorax, abdomen, and wings pale greyish,
veins on both wings darker grey : fore wings with the costal
line dark grey ; a very broad, dark pink, oblique band, occu-
pying more than half the outer portion of the wing, its inner
margin dark grey and well defined, its outer margin sufi’used
w ith the dark grey colour of the outer margin of the wing :
hind wings without markings. Underside : body, legs, and
wdngs white : fore wings suffused with grey except for a
broad subcostal ochreous streak from the base for more than
half the length of the wing.
Expanse of wings yV inch.
Bihd, W. Africa ; one example.
Somewhat resembling E. derogata^ Walker, from India.
Tar ache amydray nov.
. Palpi white, with grey bands on the last two joints;
head, body, and fore wings dark iron-grey ; frons white, with
two grey spots ; abdomen with white segmental lines : fore
wdngs with the orbicular and reniform w^hitish, the former
containing one, the latter two dark grey spots ; the entire
wing picked out with many whitish marks, transverse,
sinuous, ante- and postmedial, and submarginal lines, and
some whitish dots on the costa : hind wings uniform greyish
white ; cilia of both wings grey, with a whitish interline.
Underside : fore wings uniform pale grey, with some blackish
and ochreous spots on the outer half of the costa ; hind wings
pale, with a discal curved grey line ; body and legs whitish,
the latter with blackish bands.
Expanse of wings y^y inch.
Bihe, W. Africa ; one example.
Tarache apateliay nov.
? . White, palpi wdth last joints black ; the frons projec-
tion with two black spots touching each other; thorax with
54
Colonel C. Svv inhoe on new Eastern,
black spots ; abdomen suffused with grey, with black seg-
mental bands : fore wings almost entirely covered with dis-
located blackish bands, subbasal, ante- and postmedial, and
marginal, the last the broadest, only extending two thirds
upwards, with a black spot on the costa : hind wings white,
with grey marginal borders and a grey dot at the end of the
cell. Underside paler, slightly ochreous tinged, markings
showing through : hind wings with an additional large grey
spot on the costa beyond the middle : body white; legs wiiite,
with grey bands.
Expanse of wings 1 inch.
Bihd, W. Africa ; one example.
There are several unnamed examples from British East
Africa in the B. M. Trifid Draw’er 195 which are identical
w'ith my specimen.
Family Pyraustidae.
Isocentris char oj) alls, nov.
d* Above clear ochreous-yellow ; palpi dark brown at the
base : fore wings with a black subcostal spot near the base ;
transverse lines browm, antemedial nearly erect, medial
descending to the hinder margin from a lunular line at the
end of the cell, and postmedial from costa to vein 2; a sub-
marginal line, follow’ed by a silvery line, and a marginal
chocolate thin band, all close together ; cilia wliite, thickly
interlined with chocolate-brown: hind wdngs with a discal
'line, curving deeply inwards at vein 4, the marginal marks
as in the fore wings. Underside : pectus and body white ;
legs yellow above, white beneath ; tarsi with blackish-brown
bands ; wings pale yellowish ; fore wings with some of the
interior lines visible ; hind wings without markings.
Expanse of wings inch.
Queensland j one example.
Blepharomastix liedycliroalis, nov.
Bright ochreous yellow, transverse lines brown: fore
wings with a short line on the lower half very near the base;
an antemedial, outwardly curved, even line ; a lunular line
at the end of the cell ; a somewhat irregular and crenuhited
discal line, which bends outwards below the costa, then curves
deeply inwards to the lower end of the cell, then again bends
outwards near the hinder margin, to which it then runs down
straight ; a row of brown points near outer margin : hind
55
Australian^ and African Heterocera,
wings whitish in the cell and costal space, with a discal,
irregular, recurved line; cilia of both wings yellow. Under-
side paler than above ; fore tarsi with narrow blackish-brown
bands.
Expanse of wings inch.
Port Blair, Andaman Islands; one example.
Calamochrous homochroalisj nov.
$ . Pale greyish ochreous ; upper side of palpi, frons, and
top of head chocolate-brown ; lines brown, tliin and crenu-
lated : fore wings with an antemedial line, not reaching the
costa ; a lunular line at the end of the cell ; a discal line
fairly erect from costa to vein 2, where it bends inwards and
then downwards to the hinder margin : hind wings with a
discal recurved line. Underside with the face and body
white; legs and wings pale whitish yellow, without markings.
Expanse of wings inch.
Port Blair, Andaman Islands ; one example.
Nacoleia dairalis,
Botys dairalis, Walker, xviii. 698 (1859).
Nacoleia dairalis, Hmpsn. P. Z. S. 1898, p. 697.
Merotoma dairalis, Swinhoe, Cat. Het. Mas. Oxon. ii. p. 483 (1900).
Saraw’ak, Borneo.
Type (female), Sarawak, in B. M., and four others (all
females) from the same locality. I have received a pair
lately from Sarawak the male of which has bipectinate an-
tennse and has not the characters ascribed to the genus
Merotoma, Meyrick ; it will come into a fresh section of
Hampson’s genus Nacoleia,
Merotoma Merjriclci, nom. nov.
Merotoma dairalis, Meyrick (nec Walker), Trans. Ent. Soc. 1894,
p. 460.
Pulo Laut, Celebes.
A pair received from Mr. Meyrick from Pulo Laut are in
the B. M. with dairalis, Walker.
Pyrausta plinthinalis, nov.
^ . Pale primrose-yellow ; palpi above,, frons, head, and
thorax dark yellowish brown; palpi below w'hite; abdomen
5G
Major T. Broun on new
w'hlte, witli a brown dorsal stripe : fore wings with the costa
broadly suffused with ochreous brown, the basal portion of
the suffusion very dark ; a prominent, almost square, blackish-
brown spot fills the end of the cell ; lines thin, pale, ante-
and postmedial, the latter with an outward curve in the
middle; an outward marginal liim, much thickened and dark
brown towards the apex; cilia dark greyish ochreous : hind
wings nearly white ; a brown spot at the end of the cell ; a
discal line with an outward bend in its middle, and an outer
marginal line as in the fore wings ; cilia white. Underside
white; fore wings with a brown dot in the middle of the
cell, the spot at the end and some of the other markings
showing through the wing; hind wings with only a small
brown mark at the apex ; body and legs white without
markings.
Expanse of wings inch.
Padang, W. Sumatra ; one example.
IV. — Descriptions of Six new Species of Coleoptera from
New Zealand. By Major T. Broun, F.E.S.
Group Pogonidae.
Oopteriis pygmeatuSy sp. n.
Subovate, slightly convex, nitid, piceo-fuscous, sometimes
more rufescent ; legs and antennae fusco-testaceous or
reddish.
Head narrow^er than thorax, interocular furrows well
marked. Eyes large, moderately prominent. Thorax
cordiform, about one-third broader than long, distinctly
rounded and marginated laterally, much narrowed but not
sinuated behind the middle, posterior angles rectangular;
its surface with fine indistinct transverse rugae, discoidal
groove entire and rather feebly impressed, basal foveae
moderately large. Elytra ovate-oblong, large, marginal
channels well developed, posterior carinae distinct ; they are
more or less finely punctate-striate, sometimes irregularly
or indistinctly, but, excepting the sutural striae, they are
without sculpture near the apices; the third interstices have
three small punctures and fine setae.
Underside shining, pitchy or reddish ; fourth ventral seg-
ment wdth four setigerous punctures at the extremity, the
57
Coleoptera from New Zealand.
other segments with one on each side of the middle in the
female, in the male there are only two at the apex of the
fourth segment.
Tibioi straight, the intermediate pair and the tarsi distinctly
setose. The other characters agree with those defined hy
me on page G09, vol. xi. 1903, of the Ann. & Mag. Nat. Hist.
When compared with O. parvulus, this new species may
be at onee distinguished by the more ample and oblong, but
evidently less convex, hind body, which, moreover, does not
possess the rather coarse seulpture so apparent in the former
species. In two or three examples I notice an additional
seta at the thoracie hind angles.
Length If ; breadth | lines.
Found under stones near a stream on the Garrick Range,
near Obelisk Peak, at an elevation of 4100 feet, by Mr. J.
II . Lewis.
Group Oxytelinae.
Phloeon(jeus cordicollis, sp. n.
Elongate, subdepressed, slightly shining, rufo-testaceous,
elytra paler, finely pubescent.
Head shining, with an indistinct transverse impression
behind the eyes, behind this it is so minutely sculptured as
to appear smooth ; in front, near the eyes, it is punctate, and
seems there a little asperate and hairy ; the antennal tubercles
are prominent. Eyes rather flat, distinctly facetted, quite
lateral. Mandibles curvate, inwardly bidentate. Antennce
finely pubescent, basal articulation elongate and gradually
incrassate; the terminal three joints nearly twdee as broad as
the intermediate ones. Thorax about as long as it is broad,
truncate and widest in front, gradually curvedly narrowed
towards the rounded base, and without perceptible hind
angles ; its punctuation is moderate and rather shallow, it is
almost smooth along the middle, and the greyish pubescence
is disposed transversely. Elytra pallid, short, broader than
long, subtruncate behind, posterior angles oblique or rounded;
their surface is not definitely sculptured, but bears evident
greyish hairs. Hind-body elongate, parallel, marginated, its
five exposed segments are nearly equal and distinctly pubes-
cent, the sculpture is ill-defined ; the terminal segment is
short and retracted and is trituberculate at the extremity.
Legs of moderate length, rather slender ; tibise minutely
spinose externally, the anterior outwardly curved, with small
terminal spurs. Tarsi slender, the basal two joints, taken
58
Major T. Broun on new
togetlier^ are shorter than the apical one. Paljji with sub-
ulate terminal joints.
This may be readily identilied by its simple cordiform
thorax. M. FauveFs P. punctiventris has a shorter bisulcate
thorax and longer wing-cases.
Length IJ; breadth lines.
Collected by Mr. J. H. Lewis at Ocean Beach near
Dunedin.
Group Byrrhidae.
Pedilophorus Lewisi, sp. n.
Ovate, moderately convex, aeneo-niger ; tibiae pitchy red ;
tarsi and antennae rufescent, the basal joint of these latter,
however, piceous. Body apparently nude, but really bearing
some minute greyish setae.
Head closely and distinctly punctured, narrowed anteriorly,
the forehead subtruncate or obtusely rounded, but not
sharply defined in front, the short vertical frontal portion
nearly smooth. Labrum punctate-rugose, more or less
pilose, lateral grooves deep. Eyes large, evidently facetted,
most prominent in front. Thorax twice as broad as it is
long, narrowed anteriorly, lateral margins well-defined, front
angles rectangular and slightly projecting ; its surface closely
aJid distinctly punctured, more coarsely towards the sides.
Scutellum almost cordiform, distantly punctate. Elytra
large, moderately narrowed posteriorly, not so closely
sculptured as the thorax, much more distantly and finely,
almost obsoletely, punctured tow^ards the apices. The
female with six or seven more or less obvious striae on each
elytron, the sutural most distinct. The male with two or
three short ill-defined striae, and with series of coarse
punctures on each elytron.
Tihice finely pilose, with short terminal spines, the anterior
pair grooved along the lateral portion of the frontal face for
the reception of the tarsi, the external edges of all minutely
setose. Tarsi elongate, pilose underneath ; fourth joint
small, third prolonged below.
$ . Antennce elongate, finely pubescent, very gradually
dilated from the fifth joint onwards; basal joint stout, sub-
cylindric, and finely sculptured ; second much shorter and
narrower than the first, tapering apically ; third nearly as
long as the first, but slender; fourth shorter than the con-
tiguous ones ; sixth and seventh longer than broad ; joints
59
Coleoptera from New Zealand.
eight to ten subquaclrate ; eleventh elongate-oval ; the basal
three artieulations are less pubescent than the following
ones. My female specimen, owing to malformation, dilfers
considerably in antennal structure.
Underside, including the femora, shining pitchy-black,
moderately closely j)uncturcd, the metasternum most coarsely.
Flanks of prosternnm smooth, the intercoxal process broad,
but not prolonged and fitting into the deep angular meso-
sternal cavity. The posterior femora are accommodated in
rather large deep cavities. The abdominal sutures are nearly
straight, the fourth only being somewhat incurved.
Length 3^--3f ; breadth 2-2^ lines.
Named in honour of Mr. J. H. Lewis, who found this
fine species under stones near a stream on the Old Man
Range, Otago, at an elevation of 4100 feet. It may be
easily recognized by its rather large size and more or less
obvious elytral striae.
Group Melolonthidas.
Scythrodes squalidus.
This new genus and species of our Southern Alps was
founded (Man. N.Z. Coleopt. p. 955) on a single female
discovered by Mr. G. M. Thomson, about tw^elve years ago,
at an elevation of 6000 feet on Mount Tyndall. Quite
recently, Mr. J. H. Lewis kindly sent me some males which
he had found on Obelisk Peak, Garrick Range, at an elevation
of 4100 feet. These males exhibit several discrepancies of
sculpture and antennal structure when carefully compared
with the typical female, so it was considered best to draw up
the subjoined description so that others may not make the
mistake of treating the sexual forms as distinct species.
Body broadly oviform, only moderately convex, nigrescent,
nearly nude.
^ . Atilennee rather short, without pubescence. The basal
joint equals the following four conjointly in length, but the
portion visible from above barely exceeds the second in
length, it is gradually thickened towards the oblique apex,
which bears three or four elongate setae ; second nearly twice
as long as broad, somewhat pyriform, but unsymmetrical,
the inner side being straight whilst the outer is swollen ;
third subtriangular, obviously smaller than the contiguous
ones ; fourth triangularly cordiform, about twice the breadth
of the preceding one ; fifth remarkably short, as wide as the
GO
Major T. Broun on new
extremity of tlie fourth. Club moderately short, rather
closely })uneturcd and bearing minute brassy setae, tri-
articulate, its third lobe either slightly concave or grooved
along the outside so as, in some lights, to indicate a fourth
joint, which, however, docs not exist.
Head coarsely and irregularly punctured, the punctures
twice as numerous as in the typical female; it is sparingly
fringed with somewhat rufescent outstanding setae, and in a
small fovea close to the front of each eye there is a cluster
of about six setae. Thorax twice as broad as it is long ; the
front angles attain the middle of the eyes, the posterior are
obtusely rectangular and rest on the shoulders; the sides
are finely margined and so little curved as to appear nearly
straight except where they are narrowed towards the front ;
its surface is distinctly but irregularly punctured, between
the middle and each side the punctuation, on some spots, is
not so close, but there are no very evident smooth spaces,
the punctures usually contain greyish scale-like or sappy
matter. Elytra almost as broad as they are long, moderately
transversely convex, shoulders oblique, sides regularly
slightly rounded, apex rounded ; each with six rather broad
but not deep striae, beyond these, laterally and apically, the
seulpture is rugose and punctate, the striae, however, are not
very definitely punetured, but the whole surface being more
or less rugose some parts of the striae appear crenate ; the
interstices are broad and punctate ; the lateral margins are
not distinctly reflexed, but the channels are broad, with
coarse, transverse, and rather shallow sculpture.
Underside sparingly pilose, glossy blaek, the ventral seg-
ments with a transverse series of setigerous punctures on
each.
Tibi(B sparsely setose or hairy, the anterior a little expanded,
oblique at apex, tridentate externally, the inner calcar attains
the base of the second tarsal joint.
^ . Length 8 ; breadth 5^ lines.
Group Cryptorhynchidse.
Acalles fuscatus f sp. n.
Ovate- oblong, transversely convex, opaque, rufo-fuscous,
densely covered with depressed pale broAvn scales and sappy
matter and numerous greyish-yellow erect setae; antennae
and tarsi ferrugineous.
Rostrum as long as thorax, red, squamose at base, very
gradually and moderately narrowed towards the middle,
CoJeoptera from Neio Zealand, GL
finely punctured and irregularly longitudinally strigose.
Head immersed up to the rather flat eyes. Thorax one-third
broader than long, much contracted in front, base very
feebly sinuated, sides medially rounded, but narrowed behind ;
its punctuation close and moderately coarse, but concealed
by the covering. Scutellam invisible. Elytra oblong,
rounded and nearly vertical posteriorly, coarsely striate-
punctate.
Underside rufescent, moderately coarsely and closely
punctured, with greyish-yellow squamae and a few setae.
Pectoral canal profound, limited behind by the semi-
circularly raised margins of the mesosternum. Metasternum
very short. Abdomen large ; basal segment, in the middle,
as long as the following three conjointly, second as long as
the next two, third and fourth short, with deep straight
sutures, fifth about the length of the preceding two, with a
median fovea-like impression.
Legs long and stout, their clothing like that of the body.
Tarsi stout, third joint bilobed, but only moderately expanded.
Antenn(R elongate, inserted just before the middle of the
rostrum, their pubescence yellowish ; scape clavate and
slightly flexuous, reaching backwards to the eye ; funiculus
rather longer than the scape, second joint nearly as long as
the first, but not so stout, joints 3-7 of nearly equal
size, the seventh, however, rather broader than the others ;
club elongate, oval, apparently triarticulate.
In superficial appearance this insect is somewhat similar
to the much larger Syrnpedius vexatus (no. 890).
Length, rost. excl., 1^-1 breadth | lines.
Two examples from Mr. J. H. Lewis.
Acalles multisetosus, sp. n.
Convex^ subovate, fuscous, covered with brownish-grey
scales ; coarse, greyish, squamiform setae form a sort of
fringe round the thorax, on the hind-body there are many
outstanding, but more slender and sometimes infuscate setae,
whilst the legs and underside bear conspicuous coarse grey
ones.
Rostrum stout, slightly rufescent, squamose, finely punc-
tured. Thorax of nearly equal length and breadth, con-
stricted in front, rounded at the middle ; usually with two
grey, squamose, longitudinal streaks near the base, its surface
IS moderately coarsely punctured. Elytra oviform, obviously
broader than the thorax j they are striate-punctate, the
sculpture, however, is not easily seen.
62
On new Goleoptera from New Zealand.
Legs stout ; tibiae slightly reddish ; tarsi almost testaceous.
The pectoral canal extends almost to the middle of the
intermediate coxae. Metasternum very short, but the basal
ventral segment is large, the terminal one is red.
This minute species is most nearly allied to Pascoe^s
A. perpusillus (no. 870).
I-iength, rost. excl., | ; breadth f line.
This species also is from the collection of Mr. Lewis.
Group Cossonidae.
Agastegnus rufescenSj sp. n.
Elongate^ subdepressed, subcylindric, finely pubescent,
moderately glossy, infuscate-red ; rostrum and thorax quite
red, head darker.
Rostrum shorter than thorax, nearly cylindric, but con-
tracted at the base and slightly narrowed between the middle
and the apex, finely and distantly punctured in front, more
closely behind. Head immersed up to the eyes, much
curvedly narrowed anteriorly, so punctured as to appear
rather dull. Eyes flat, lateral. Thorax somewhat oviform,
but constricted near the apex ; its disk subdepressed and
medially furrowed, distinctly yet finely and not closely
punctured ; it bears distinct decumbent golden hairs near
the sides. Scutellum small, but distinct. Elytra elongate,
somewhat gradually narrowed towards the base, broadly, but
not very obviously, transversely depressed between the middle
and the base, sparingly clothed with yellow and greyish
hairs ; punctate-striate, intermediate striae shallow or obso-
lete, the two suturals deep at the base ; interstices apparently
impunctate, the third carinate towards the apices.
Legs long, rather stout, finely and sparingly pubescent;
tibiae flexuous, with well-developed hooks. Tarsi 4-jointed,
penultimate moderately expanded and sub-bilobed, the apical
joint nearly as long as the other three conjointly, their soles
with scattered outstanding slender hairs.
Antennee reddish, scape flexuous and a little thickened
towards the extremity, quite as long as the funiculus, which
is 5-jointed, with the basal joint obconical, second nearly
twice as long as broad and more slender than the contiguous
ones, third and fourth apparently transversel}^ quadrate,
though slightly narrowed near the base, fifth transverse,
broader than the preceding ones ; the club densely but finely
pubescent, oblong-oval, large, indistinctly articulated, about
the length of joints 2—5 of the funiculus taken togefher.
63
On Three new Fishes from Yunnan,
A. distinctus, also from Otago, may be distinguished by
its thicker, almost parallel-sided rostrum, stouter and shorter
antennae, evidently shorter second joint of the funiculus, by
its deeper thoracic channel, and the double elytral trans-
verse impressions, one being behind the middle femora and
the other in line with the posterior pair.
Length, rest, inch, 1^; breadth quite | lines.
Described from a single mounted specimen sent by
Mr. J. H. Lewis, who found it at Otago.
V. — Descriptons of Three new Fishes from Yunnan^
collected by Mr. J. Graham. By 0. Tate ReGAN, B.A,
Schizothorax taliensis.
Depth of body 5 to 6 in the lengtli, length of head 4| to 5.
Diameter of eye in the length of head, lengtli of snout
3§, interorbital width 3. Mouth terminal, with the jaws
equal anteriorly ; maxillary extending to below anterior edge
of eye ; fold of the lower lip not continuous ; barbels short.
85 to 100 scales in the lateral line, 24 to 29 between dorsal
fin and lateral line, 12 to 15 between lateral line and root of
ventral. Scales irregular ; thorax and a median abdominal
strip naked. Dorsal IV 7, its origin nearly equidistant from
tip of snout and base of caudal ; the compound serrated ray
strong, its length f to J the length of head ; free edge of the
fm concave. Anal III 5, when laid back extending to the
base of caudal. Pectoral | the length of head, extending -J
or a little more than ^ of the distance from its base to the
origin of ventrals. Ventrals 9- or 10-rayed, originating a
little behind the origin of dorsal, extending ^ to § of the
distance from their base to the origin of anal. Caudal
forked. Caudal peduncle twice as long as deep. Silvery ;
back bluish.
Four specimens, 130 to 210 mm. in total length, from the
Tali Fu Lake, 300 miles W. of Yunnan Fu.
Discognathus yunnanensis.
Depth of body 5 in the length, length of head 4|. Eye
slightly nearer to edge of operculum than to tip of snout, its
diameter ^ the length of head and ^ the width of the slightly
convex interorbital region. Barbels 2 on each side, short,
64
Mr. C. T. Regan on
subequal. 40 scales in a longitudinal series, 5 between
dorsal fin and lateral line, 3 between lateral line and root of
ventral fin. Dorsal III 8 ; origin nearly equidistant from
tip of snout and base of caudal ; longest ray nearly as long as
the head ; free edge of the fin slightly coticave. Anal III 5.
Caudal deeply forked. Pectoral as long as the head, not
reaching the ventrals, which originate below the middle of
the dorsal and extend to the origin of anal. No well-defined
spots or markings.
A single specimen, 53 mm. in total length, from the lake
at Yunnan Fu.
Silurus Grahami,
Depth of body 6 in the length, length of head 5. Breadth
of head If in its length, diameter of eye 8, interorbital width
3^, length of snout 3f. Lower jaw projecting; vomerine
teeth in separate patches ; 4 barbels, the maxillary ones
reaching the ends of the pectorals, the mental ones f as long.
Dorsal 4, its distance from tip of snout slightly more than J
its distance from the caudal. Anal 73, continuous with the
caudal, which is subtruncate. Pectoral with I 13 rays ;
spine with the outer edge weakly denticulated and the inner
edge entire, its length | that of the fin, which is f as long as
the head and does not reach the ventral. Ventrals 12-rayed,
extending to the third or fourth ray of anal. Greyish brown.
A single specimen, 260 mm. in total length, from the
Cliien Kiung Lake, 30 miles S.E. of Yunnan Fu.
This species is very close to S. mento^ Yunnan
Fu Lake, which has shorter barbels, the inner edge of the
pectoral spine serrated, and the ventrals 10-rayed.
VI. — Diagnoses of new Central- American Freshwater Fishes
of the Families Cyprinodontidae and Mugilidse. By C.
Tate Regan, B.A.
1 . Rivulus flahellicauda.
D. 9. A. 12. Sc. 42. Diameter of eye 4 in the length
of head. End of anal below middle of dorsal. Brownish,
edges of scales darker ; vertical fins with some small darker
spots ; a caudal ocellus.
Jriab. Costa Rica, Juan Venas [Underwoodf
Total length 70 mm.
new Central- American Fishes.
G5
2. RivuJus Godmanni.
D. 8. A. 11. Sc. 35. Diameter of eye 3 in the length
of head. End of anal below middle of dorsal. Olivaceous, a
darker spot on each scale ; operculum blackish ; vertical fins
dusky, the caudal with a narrow pale edge and below with
a blackish intramarginal stripe; caudal ocellus sometimes
present.
llah. Guatemala (^Godman).
Total length 40 mm.
3. Pcecilia salvatoris.
Poecilia thermalis (non Steind.), Giinth.. Cat. Fish. vi. p. 341 (1866).
D. 10-11. A. 8-9. Sc. 27-30. Closely allied to
P. splienops, C. & V., but with the body not so deep, the
interorbital space broader, and the free edge of the dorsal fin
straight instead of eonvex. Olivaceous ; males with more or
less distinct cross-bars ; dorsal with 2 series of vertically
expanded blackish spots, sometimes absent in females ;
caudal, in the males, with oblong blackish spots.
Ilah. San Salvador, in warm springs [Dow],
Total length 55 mm.
4. Xiphophorus strigatus.
Xiphophorus Helleri (non Heck.), Meek, Zool. Pub. Columbian Mus. v.
1904, p. 157.
D. 12-14. A. 9-10. Sc. 28-30. A blackish lateral stripe
from eye to base of caudal ; no additional stripe in the males.
Hah. Southern Mexico, Vera Cruz and Oaxaea.
The true X. Helleri is the species named X. jalapce by
Meek, males of which have two lateral stripes.
5. Xiphophorus brevis.
D. 13-15. A. 9-10. Sc. 27. Depth of body 2|- to 2| in
the length. 3 or 4 indistinct dark longitudinal stripes at the
edges of the series of scales on the sides ol the body.
Hah. British Tdonduras, Stann Creek [Robertson).
Total length 75 mm.
6. Agonostomus macracanthus.
D. IV, I 8. A. II 10. Sc. 41-43. Upper lip very thick,
as in A. nasutus. Maxillary extending to below middle of
eye. Dorsal spines strong, the first J-| the length of head or
Ann. & Mag, N. Hist. Ser. 7. Vol. xix. 5
G6
]\Ir. W. Nlcoll on the
the distance between tlie origins of tlie dorsal fins and as
long as the longest rays of the second dorsal. Pectoral | the
length of head, extending to or a little beyond the vertical
I'rorn the origin of spinous dorsal.
Hah, Guatemala, Uio Guacalate {Salvin).
Total length 210 mm.
7. Agonostomus Salvini.
D. IV, I 8. A. II 10. Sc. 38-40. Closely allied to
A. nasutusj but upper lip not so thick, eye smaller, lower jaw
a little longer, pectoral fin shorter. Maxillary extending to
below anterior ^ or of eye. Dorsal spines moderate, the
first I (adult) or J (young) the length of head or of the
distance between the origins of the dorsals. Pectoral f the
length of head or less, not extending to below the spinous
dorsal.
JIah. Guatemala, Rio Nacasil [Salvin).
Total length 270 mm.
VH. — A Cont?'ibutio7i towards a Knowledge of the Entozoa of
British Marine Fishes. — Part I. By William Nicoll,
M.A., B.Sc., Gatty Marine Laboratory, St. Andrews.
[Plates I.-IV.]
The following aecount of an as yet little-known province of
British marine zoology can hardly be regarded as more than
a mere preliminary. The original intention was to have
treated the subject in a systematic manner, dealing wdth the
Entozoa of each of the natural groups of fishes — Gadidae,
Pleuronectidae, &c. — separately ; but as this w^as found im-
possible for the time being, the most satisfactory remaining
course seemed to be to study the easily accessible fishes as
they came to hand. It will be seen that these fall under
two classes : (1) the commoner littoral fishes, gunnel, stickle-
back, &c. ; (2) the commoner food-fishes, haddock, dab, &c.
A large number of each species, except in a few^ instances,
having been examined, the results obtained may be regarded
as fairly accurate and the parasites from each host as com-
paratively typical. Special attention has been devoted to
the Trematode forms ; cestodes occurred but rarely, except
in the form of scolices. Nematodes and Acanthocephala
were frequent, but, for the most part, assignable to common
species. Several forms remain unnamed, mostly young
Ascarids, which are difficult to diagnose.
G7
Entozoa of British ]\ [urine Fishes.
No Elasinobranch fishes are noted in tliis paper^ and of
the Teleostei, three groups — Pharyrigognathi^ Lophobranehii,
and Pleetognathi — are unrepresented. Ofthe remaining three
groups upwards of 60 speeies are reeorded from St. Andrews
Bay, but only 50 of these are at all eommon. About half
of these are dealt with here. A few yielded no parasites, a
notable case being Blennius pholis, which will be referred to
later; amongst the others an individual not harbouring its
quota of parasites was exceptional.
A striking feature of the herein-noted results is the large
number of instances in which a parasite is recorded from a
new host, although the new species are few. This may be
due to a particular host not having been examined before
or to its not having harboured Entozoa in the localities where
it had previously been examined. A third possibility sug-
gesting itself is that two species of fish may have been
confused or regarded as identical, and the parasites of one
attributed to the other. A case in point is that of Coitus
scorpiuSy Bloch, and C. bubalis, Euphr., between which I
have sometimes found difficulty in deciding. I have, how-
ever, carefully compared each specimen examined with Day^s
descriptions of the two species, and in so far as he is to be
depended upon the results may be regarded as correct. The
large number of forms new to Coitus bubaiis may be explained
in this way; moreover, almost every one occurred in an
example which was unmistakably Coitus bubaiis, Euphr.
This leads us to a consideration of the work of previous
observers. Two most important attempts have been made
towards a systematic and exhaustive knowledge of the
Entozoa of the North Sea fishes — the one by Olsson* in
1867, the other by P. J. van Benedenf in 1870. The work
of the former was done in the waters off the west coast of
Norway, which can hardly be regarded, strictly speaking, as
the North Sea; but since the majority of the species which
he examined are common to our shores, we can include his
results under those for fish taken within the North Sea.
Yan Beneden deals with fish from the Belgian coast and
from further out at sea. Both examined a very large number
of fishes, especially the commoner varieties. Amongst the
littoral fishes neither Olsson nor Van Beneden makes mention
of Gobius Ruthensparri and Liparis Montagui. Olsson, in
addition, omits Zoarces viviparus ; Van Beneden, Coitus
O
* “ Entozoa lios Skandinaviska liafsfiskar,” Lunds Univ. Arsskrift,
vol. iv. 1867, no. viii.
t “ Les Poissons des Cotes de Belgique,” M(5m. Acad. Belg. xxxviii.
1871.
5*
C8
]\[r. W. Nicoll on the
huhalis. From tlicir observations no parasite appears to
infest the fish haunting the pools and crevices along the
shore in the same way as Podocotyle atomon^ llud., has been
found to do at St. Andrews. The presence of this Trematode
is quite a feature ; it occurs in eight out of the nine species
examined. Another common form is Echinorhynchus acus,
llud._, which was found in four species. Amongst the fishes
from deeper waters the widely-spread Hemiurus forms and
Derogmes various, Muller, are recorded very frequently l)y
both the above-mentioned observers. As might have been
expected, the same is true in the present instance.
Fish, such as the herring and haddoek, which roam far and
wide in their search after food tend to exhibit the same para-
sitic fauna in whatever quarter they are taken, but the littoral
fish, with their more circumscribed lives, are dependent on
local faunae for food, and in consequence their parasites vary
according to the locality.
A case like that of Podocotyle atomon, Fud., would almost
lead one to enunciate the hypothesis that the Entozoa of a
particular fish depend more on its environment than on the
species to which it belongs ; that is to say, that no matter
what the species of fish the parasites found in it ought to be
the same as those found in other species from the same
neighbourhood. This raises the very important question of
idiosyncrasies in feeding, for it is well known that different
species, although living quite close together, have often
entirely different modes of feeding. Van Beneden took some
pains to note the food of the various fish which he examined ;
but into this matter I do not propose at present to enter. One
case, however, that of the shanny, Blennius phoUs, cannot be
overlooked. It occurs here commonly, and I had an oppor-
tunity of examining a large number, but in no instance did
a single parasite present itself. Such a fact is not remark-
able ill itself, but, in view of the frequency with which other
fish in the vicinity were infected, it is striking enough.
Van Beneden found the food of the shanny to consist of
crustaceans, Balanus for the most part. In addition to
crustaceans I have met with excessive numbers of small
gastropods, especially young Littoi'ina. Thus the food is
apparently not so very diflerent from that of other species.
The presence of the large number of shells in the intestine
might be offered as a reason for the absence of parasites,
but shells and other hard debris are found in fish in which
parasites abound. Another explanation might be sought in
some constitutional peculiarity of the shanny which renders
it an unsuitable host. Similar instances — e. g. Agonus
CD
Kulczoa of British J\farine I'ulies.
(Asp'ulophorus) catnphracim, Cienolabrus rupcstris, and
Jh'osmius brosme — occur in Olsson’s observations^ altliougb lie
makes no comment upon them.
Neither Olsson nor Van Beneden made any a^^^mpt
towards a solution of the life-history of the forms with '''diicli
they deal. The subject is difficult and requires sj)ccial and
long continued study. Leviriscn* made an extensive seareli
amongst the marine invertebrata of Greenland for inter-
mediate forms, and was suecessful in three or four eases,
llis results have not been confirmed. Latteily some
important work has been done in this direction by Miss
Lebour in Northumberland t- She has investigated many
of the commoner Mollusea and has reeorded from them quite
a number of sporocysts and eercaria3. The difficulty of
tracing these to their final host still remains.
In the Irish Sea at the Liverpool Laboratory considerable
additions have been made to the British Entozoa fauna by
Johnstone Very few Trematodes are mentioned, Cestodes
receiving most attention. His note on Distomum valdem-
flalum, Stoss., is interesting and will be referred to later.
'\ he occurrence of Hemiunis appendiciclaius in very small
(^-1 in.) plaice and dabs is curious, for this Trematode has
never been recorded from these fish in their adult state.
The mention of a form {Distomum gulosum, Linton) first
described from America is also noteworthy. Linton^s work
on the American fishes § reveals many forms which are the
same as or almost identical with species already known from
corresponding European fish. He himself is unable in
several eases to establish their identity with absolute
certainty. His work is valuable on account of the eare with
which he has studied and measured the various important
organs, but unfortunately little exact information is to be
derivecl from his plates.
Some experiments which I h.ad an opportunity of conducting
with sticklebacks may conveniently be described here. The
fact that the sticklebacks from brackish ditches were infeeted
with Fodocotyle atomon, Rud., to as great an extent as those
from the marine roek-pools suggested ascertaining the effect
^ “ Bidrag til Kimdskab om Groiilands Trematodfauna,” Oversigt Kgl.
Bansk. Selskab. 1881, pp. 52-84, pis. ii., iii.
t Notes on Northuinhrian Trematodes,” Northumberland Fisheries
Report for 1905.
X ‘‘ Internal Parasites and Diseased Conditions of Fishes,” Lancashire
Sea-Fisheries Report for 1904, p. 98 ; 1905, p. 151.
§ Proc. U.S. Nat. Mus. xx. pp. 423-456, pis. xxvii.-xxxiv. and pp. 507-
548, pis. xl.-liv.
70
Mr. \V. Nicoll on the
of pure fresh 'water. Some examples^ both from pools and
from (litdies^ were confined in a tank of fresh water. For
two months they were supplied with })ond-weeds, snails,
grubs, &c., which, liowever, they did not accept very readily.
At the end of* tliat time sorric were examined and found to
contain the parasites as frequently as before. No new
])arasites had made their a})peatance. For another two
months the sticklebacks received no food at all, and on
examination thereafter no diminution in the number of
])arasitcs was observable, although by this time the fish were
in a very poor condition. Tliis is at variance with the
observations of Zschokke and others, who found that in fish
migrating from the sea to rivers the parasites acquired during
their sojourn in the sea were gradually killed off by the fresli
water of the river. The above experiment appears to indicate
that the disappearance of the parasites in certain cases is not
W'holly attributable to the effect of the fresh w^ater, but that
some other factor must enter into account. The length of
time (four months) during which the fish were confined to
the fresh water was ample for any effect due to the water
to have taken place. The only noteworthy circumstance
observed was the largely increased number of ova extruded
from the parasite in the intestine of the host.
The following is a list of the hosts examined, with their
respective parasites ; the habitat is also noted. An asterisk
prefixed to the habitat denotes that the parasite is recorded
for the first time from this situation; an asterisk prefixed
to the name of the parasite denotes that it is here recorded
for the first time from this host. The fish are named
according to Day^s ‘British Fishes.'’
Acanthoptebygii.
Gasterosteus aculeatus, Linn. 3-spined
Podocotyle atomon, Mud.
( = Psilostomum redactum, mihi.
^Ascaris sp. (jiiven.).
Coitus scorpius, Bloch. Bullliead.
Podocotyle atomon, Mud.
(=l)istomimi simplex, Mud.)
Derogenes varicus, Muller.
Bistomum sp.
Echinorhynclms acus, Mud.
Bothiiocephalus punctatus, Mud.
Coitus buhalis, Euphr. Father-lasher.
Podocotyle atomon, Mud,
^Ilemiums appendiculatus. Mud.
■^Derogenes varicus, Muller.
*Prosorhynchus squamatus, Odhner
Stickleback.
Stomach, intestine.
Body- cavity.
Intestine.
Stomach and ^intestine.
Gills in capsules.
Intestine.
Intestine.
Intestine.
Stomach.
Stomach.
Intestine and pyloric
a2)pendages.
71
Kntozoa of British Marine Fishes.
■’i^Ascans sp.
■^Ascai’is capsularia, Rtul.
^Ascarop.sis nioiTlmrc, v. Ben.
*EcliiiU)rliynclui8 acus, Rud.
liotlirioceplialus punctatus, Rucl,
^Scolox polyiiiorplius, Rud.
^Diatom uiii sp. (juveii.).
liody-cavity.
I’eritoiieimi.
Intestine.
Intestine.
Intestine.
Intestine and rectum.
capsules.
DouLle- spotted (iub3^
Stomach and intestine.
Gills in capsules.
Ilody-cavity.
Intestine.
Gohius Ruthens^jarri, Euphr.
^Podocotyle atomon, Rud.
^Distomum sp. (juven.).
^Ascaris sp. (jiiven.).
■^Scolex polymorph us, Rud.
Ci/clopterus lumpus, Linn. Lump-sucker.
Scolex polymorph us, Rud.
Liparis Montagui, l)onov. Montague’s Sucker.
^Podocotyle atomon, Rud. Intestine,
^Prosorhynchus squamatus, Odhner. Intestine
^Stomach and intestine.
^Echinorhynchus acus, Rud.
*'Ascaris sp. (juven.).
Centronotus gunnellus, Linn. Gunnel.
^Podocotyle atomon, Rud.
^Hemiurus appendiculatus, Rud.
^Ascaris sp. (juven.).
and pyloric
appendages.
Intestine.
Body-cavity.
Intestine and rectum.
Intestine.
Peritoneum.
Zoarces viviparus, Linn. Viviparous Blenny.
^Podocotyle atomon, Rud.
^Echiuorhynchus acus, Rud.
An.vcanthini.
Gadus ceglefinus, Linn. Haddock.
Lepodora rachisea, Cobhold.
*'llemiurus communis, Odhner.
Ascaris communis. Dies.
Ascaris clavata, Rud.
*A scaris sp.
^Agamonema commune, Dies.
^Ileterakis foveolata, Rud.
^Ascaropsis morrhuoe, v. Ben.
Echinorhynchus acus, Rud.
Bothriocephalus rugosus, Rud.
^'Scolex polymorphus, Rud.
^Scolex sp.
Gadus merlangus, Linn. Whiting.
Derogenes varicus, Muller.
Ascaris clavata, Rud.
Filaria echinata, v. Linstoio.
Motella mustela, Linn Five-bearded
^Podocotyle atomon, Rud.
Ascaris capsularia, Rud.
Ammodgtes tohianus, Linn. Sand-Eel
^Ilemiurus communis, Odhner.
Intestine.
Intestine.
Intestine.
Stomach.
*Body-cavit3^
^Stomach.
Intestine.
Liver and pyloric ap-
pendages in capsules.
Intestine.
Mouth, oesophagus,
stomach, intestine.
Intestine.
Intestine and pyloric
appendages.
Intestine.
Gall-bladder.
^Intestine.
Intestine.
Intestine.
Bockling.
Stomach and intestine.
Peritoneum.
(Esophagus, stomach,
intestine.
72
Mr. W. Nicoll on the
Bracliypliallua crenatus, Rud.
J^ecitliaster gibbosus, Rud.
•=*^Ascaria sp. (juven,).
^ Ascai-is sp.
'^JCcliinorlijncbiia aciia, Rud.
(?) Scolex amniodytis Tobiani, v.Ben.
Jlippof/lossus vuU/aris, Flein. Halibut.
*tStephanocbasniiis baccatus, sp. n.
^Ilemiurus apijeiidiculatus, Rud.
l^erogeues various, Muller.
*Derogeiies cacozelus, sp. n.
Ascaiis capsularia, Rud.
Ascaris collaris, Rud.
<*^Ascaris sp.
^Ascaropsis morrliuse, v. Ben.
*Filaria echiuata, Linstoiv.
Ileteraliis foveolata, Rud.
Scolex polymorphus, Rud.
Rhombus maximus, Linn. Turbot.
*Zoogonoides viviparus, Olsson.
^Derogenes various, Muller.
Lotliriocepbalus punctatus, Rud.
Scolex polymorplius, Rud.
Rhombus Icevis^ lioiidelet. Brill.
Berogenes various, Midler.
^Ascaris collaris, Rud.
Bleuronectes limanda, Linn. Bab.
Z'jogonoides viviparus, Olsson.
Steringopliorus furciger, Olsson.
^•Berogenes various, Muller.
^Berogenes cacozelus, sp. n.
Ascaris capsularia, Rud.
* Ascaris sp.
Scolex polymorphus.
'Bleuronectes platessa^ Linn. Plaice.
Zoogouoides viviparus, Olsson.
Heteralds foveolata, Rud.
Scolex polymorphus, R^id.
Bleuronectes microcephcdus, Flem. Lemon-
Bistomum sp.
^Ascaris sp.
^Ascaropsis (?) sp. (juven.).
Stomach and intestine.
Intestine.
Body-cavity.
Intestine.
Intestine.
Intestine.
Ilectum.
Stomach.
Stomach.
Intestine and rectum.
lAritoneum.
Intestine.
Bectum.
Stomach.
Bectum.
Stomach.
Intestine and ^rectum.
Bectum.
Mouth, oesophagus, and
stomach.
Intestine.
Intestine.
^Stomach.
Intestine.
Intestine and rectum.
Stomach and intestine.
Mouth, oesophagus, and
stomach.
Intestine.
Peritoneum.
Intestine and pyloric
appendages.
Intestine.
Intestine and rectum.
Intestine.
Intestine.
dab.
Intestine.
Intestine.
Intestine.
PllYSOSTOMI.
Clupea harengus^ Linn. Herring.
^Hemiurus Liihei, Odhner.
Agamonema capsularia, Dies.
Anguilla vulgaris, Tint. Eel.
Hemiurus appendiculatus, Rt(d.
Lecithochirium rufoviride, Rud.
^Scolex polymorphus, Rud.
Stomach and cascum.
Peritoneum.
Stomach.
Stomach.
Intestine.
Kntozoa of British Marine Fishes.
73
T R E M A T O D A.
Podocotyle atomon, llud. (PL I. figs. 2.)
Bistoma simplex, Uiid. ?, Olsson, Lovinsen, Gronlanda Trematodfauna,
]). 8, pi. iii. lig. 1.
Allocrendmm atomon (Hud.), Odlmer, Zool. Jalirb. Syst. xiv. p. 500,
pi. 33. figs, 0, 10.
Podocotyle atomon (Rud.), Odlmer, Fauna Arctica, iv. (2) p. 320,
pi. ii. figs. 9, 10.
Psilostomum redactum, sp. n., Nicoll, Ann. & Mag. Nat. Hist. (7)
xvii. p. 525, pi. xiii. figs. 9, 10.
This is a species which Odlmer regards as wrongly
identified by Olsson* * * §, although the latter had some doubt
on the matter himself. Olsson assigned his specimens to
Distoma simplex, Rud., and was followed by Levinsen and
Linton. Odlmer, by elucidating the structure of Distomum
atomon, Rud., shows that they ought really to have been
assigned to this species. He also includes Distomum re-
flexum, Creplin, under this species, although he excludes the
forms which Olsson t and Zschokke J identified as such.
As Psilostomum redactum I described what I considered to
be a distinct species, but I must now regard it as identical
with Podocotyle atomon, Rud.
Of RudolphPs Distoma simplex { = Fasciola aglefine,
Muller) no specimens remain, so that Odlmer regards the
species as unidentifiable. He omits notice of the fact that
Van Beneden § found in Gadus eeglefinus what he apparently
regarded as Distomum reglefine, Muller. Van Beneden is
slightly confusing, for wdiile he marks the form sp. nov.^’
in his notes, without any attempt at description, the accom-
panying figure is marked D, aglefiue, Muller, and we may
suppose that this was his real intention. Von Linstow ||
notes both D. simplex, Rud., and D. aglefine, van Beneden,
under Gadus <jeglefinus, so that he was either misled by van
Beneden or he regarded the two species as distinct. Stossich^
falls into the same error. Van Beneden^s figure is meagre,
but, so far as it goes, exhibits a certain resemblance to the
form we are here dealing with. The large elliptical ventral
sucker, the position of the genital aperture, the male genital
* Entozoa iaktt bos Skandiiiaviska bafsfiskar,” in Lunds Univ.
Arsskrift, iv. p. 34, pi. iv. figs. 81, 82.
O
f Lunds Univ. Arsskrift, iv. (8) p. 52.
j Verliandl. Gesell. Basel, Till. 8, lift. 3, p. 789, pi. xi. fig. 1.
§ “Poissons des Cotes de Belgique,” p. 50, pi. iv. fig. 14, in M^m.
Acad. Roy. Belg. xxxviii.
II Conipend. d. Helminthol. p. 236.
^ List. d. Pesci, p. 01.
71
Mr. W. Nicoll on the
apparatus^ tlic vitelline glands^ testes, and ova, all point to
its being a member of the Allocreadium. The ova are
excessively large and the ovary is absent, and on this account
it is impossible to assign this form to any of the already
known members of the genus, so that the difficulty of
])roving the identity of Dist. simplex, Rud., still remains.
Olsson found this parasite occurring in Sebastes norveyievs
(one or two fairly often), Gadus melanostomus (frecpiently
numerous), niger (once, in great wxxmhQv^), Anguilla
vulgaris (a single specimen). Levinscri found it fairly often
in Coitus scoipius and Coitus gobio {^Fhobetor ventralis) fi’om
Greenland. Rudolpbi’s specimens were from the stomach of
Pleuronec'es flesus. Odhner adds that he has met with it
in a very considerable number of other Scandinavian marine
fishes from the west coast of Sweden,^^ although I have seen
no list of such forms. I have already recorded it from
G aster osteus aculeatus (var. trachurus) , and to this I have
to add Coitus bubalis, Cottas scorpius, Gobius Ruthensparri,
Centronotus gunnellus, Zoarces viviparus, Motella mustela, and
Liparis Montagui. About 70 per cent, of the total number
of fish of these species examined were infected, usually with
three or more adult parasites and often a large number of
young. Thus it may be understood that this form is ex-
ceedingly frequent.
Olsson determines the length of his specimens at 3-9 mm. ;
Levinsen found somewhat smaller examples in Coitus scorpius
(3-5 mm.) ; Odhner gives the average length as about
2 mm., and is inclined to regard Olssoids larger specimens
as a variety. He also differentiates a medium-sized variety
corresponding with Levinseids specimens and also with
Jdist. reflexum, Crepl. My examples only in rare cases
exceed 3 mm., the majority being l'5-2‘5 mm., so that as
far as length is concerned they correspond v/ith Odhner’s
smallest variety, i. e. with the Rudolphi type. Fully de-
veloped adults were found as small as 1 mm. ; one small
example from the stickleback measured 1*01 mm., and
contained seven ova measuring ’081 x *047 mm., i. e. of
fully normal size. The largest immature individual observed
was *90 mm. in length ; it contained no ova, but the penis
Avas well developed, and the testes were as large as *28 x
’ll mm.
The general shape oE the body is elongate-ovoid, depressed,
somewhat attenuate anteriorly, more rounded posteriorly.
Like the other species of the genus, it is extremely mobile,
the ant-acetabular region being capable of great extension
and contraction, the post-acetabular part less so, but the
Kntozoa of British Marine Fishes. 75
cuticle of tlie bitter is often thrown into irregular wrinkles.
The eonstrietion at the level of the testes, noted by Olsson,
is not of invariable oeeurrenee, and the outline of })reserved
(pressed) speeimens is eomparatively even. Tiie eolour
viewed by transmitted light is greenish yellow, darker
posteriorly from the presenee of the yolk- glands and golden
yellow centrally from the ova.
The eutiele is of no great thiekness, devoid of spines, and
striated longitudinally and transversely. The suekers are
fairly well developed : the oral sueker is subterminal and
globular, with a eircular aperture ; its diameter is usually
about of the length of the body, but it is proportionately
larger in the younger examples than in the older, the
observed limits being *12 mm. in the smaller (I’O mm.)
and -29 mm. in the largest (3T5 mm.). The ventral sucker
is more variable ; it is always elliptical in outline, with a
corresponding aperture, the long axis being transverse and
usually about half as long again as the diameter of the oral
sucker, i. e. \ of the total body-length, but here again the
proportion decreases with increase in size. The measured
dimensions were ’22-42 mm. These figures agree very
closely with those of Odhner, viz. *12-*25 mm. for the oral
and ’25-’44 mm. for the ventral sucker. Linton^s American
specimens yield pretty much the same figures.
Another feature to which some importance is attached is
the distance between the suckers (i. e. length of neck). My
observations coincide with those of Levinsen, viz. of the
length of the body and ^ in young individuals. Odhner
also determines the limits at 1-J.
The alimentary system conforms to the genus type, except
that a distinct prepharynx is present, first noted by Odhner.
It appears as a dilated tube about half as long as the pharynx
and considerably wider than the oesophagus. The pharynx
is almost globular, with a diameter of about TO mm., but its
breadth usually exceeds its length slightly. The oesophagus
may be twice as long as the pharynx, but on contraction it
is bent in the form of an S, and appears short then. The
bending takes place either laterally or doiso-ventrally.
The excretory vesicle is a long simple closed sac extending
as far forward as the level of the ovary, and opening pos-
teriorly by a terminal pore. To it run down two narrow
convoluted tubules, one on either side.
The genital aperture lies almost midway between the
two suckers and also midway between the median line and
the extreme left edge of the body. It is thus to the left of
the oesophagus and between the pharyirx and intestinal
Mr. W. NIcoll on the
7lj
l)if’nrcation. Tlie cirrus-pouch is long and narrow^ extending
some distance behind the ventral sucker. It contains at its
])osterior end a large bipartite vesicula scminalis, from the
anterior end of which issues the ductus ejaculatorins. The
latter bends back almost immediately to lie alongside the
vesicula seminalis^ but bending again it passes forward to
the penis. This organ is long and slender^ and when well
extended is curved. It has a squarely-cut end, but instead
of being inflated at its termination, as Olsson represents it,
it is somewhat tapering. The configuration of tiie internal
genitalia is precisely as Odhner* represents it.
In one case I was fortunate enough to witness the fer-
tilization and formation of the ova. The worm was under
pressure, and the unfertilized ova passed along the oviduct
I’airly rapidly. As each arrived opposite the receptaculuni
seminis, it was surrounded by sperms, and shortly afterwards
two or three yolk-plugs were congregated round it. The
mass was forced on towards the uterus, receiving as it went
a shell-coating and gradually acquiring the characteristic
elliptical shape. The whole process did not last longer than
ten minutes.
The vitelline glands consist of numerous follicles, not
by any means so regular or so large as Odhner represents
them. He is correct in saying that they do not extend in
front of the ventral sucker, although occasionally a follicle
or two is to be found in the neck.
The uterus is short and rarely contains more than 20-40
eggs ; Odhner says 20-30, and yet in his figure f he repre-
sents upwards of 80 eggs. The ova are light yellow to
yellowish brown in eolour, and measure, aceordiiig to my
observations, *075-’084 mm. in length by 040--054 mm. in
breadth. These rather wide limits include measurements of
the ova from various hosts. The details in four cases are as
follows : —
mm. mm.
From Gasterosteus aculeatus .... •07o--0Sl X •040-’044
,, Coitus hibalis •07o-’081 x •044-‘0o4
„ Ceiitronotus gnnnellus .... ‘075 X ‘OoO-'OS^
„ Zoarces vivipams •03l-'084 X •046-050
Odhner^s figures are '060- 084 mm. x •0I0-'045 mm.
The minimum length-limit seems rather small. I have
certainly never observed ova (mature, at least) approaching
that small size. My measurements were usualH determined
* Log. cit. pp. 510-511, pi. 33. fig. 10.
t Log. Git. pi. 33. lig. 9.
77
Entozoa of British J^lnrine Fishes.
from ova as near tl.e terminal portion of the uterus as
possible. Linton^s figures for the ova of his Arneriean
specimens are *084 x *04 mm., so that they also corrcs[)ond
with other observations.
This parasite, is usually eonfiiied to the intestine of its
host, though occasionally one or two specimens are to be
found in the stomach and in one host (Gadus m,elcmostomus) ,
according to Olsson, it occurs in the pyloric caeca.
While studying this form, I examined sticklebacks from
three diflerent regions, viz. (1) rock-pools, (2) brackish
ditches communicacing with the sea, (3) streams near their
entrance into the sea. In the first two instances 1 obtained
Gastei'osteus aculeatiis, var. trachurus ; in the third, var.
gymnurus. The river sticklebacks were much smaller than
those from the sea, and in no case did they harbour Podo-
cotyle atomon. The specimens from the pools and ditches
w'ere identical and were each equally subject to infection.
Levinsen asserts that the intermediate host of Distomum
simplex is Themisto lihellula. This crustacean is not recorded
from the St. Andrews district, so that it cannot be the
intermediate host here.
Lepodora rachicea, Cobbold*. (PI. I. figs. 3, 4.)
Distomum rachion, Cobbold, Trans. Linn. Soc. xxii. p. 158, pi. xxxi.
figs, y, 10; Stossich, List. d. Pesci, p. 43 ; Linton, Proc. U.S. Nat.
Mus. XX. p. 538, pi. liii. figs. 3-7.
Lepodora rachiesa, Odliner, Fauna Arctica, iv. (2) p. 332, pi. ii.
dgs. 12-15.
To reconcile Cobbold^s figure of Distomum rachion with
the one which 1 herewith submit seems at first sight difficult.
The disposition of the genital glands and uterus appears to
effectually separate them. The resemblance in other respects,
however, is striking enough, and as it is possible to interpret
correctly the structures misrepresented by Cobbold, I have
little doubt that his specimens and mine are identical, and
this is endorsed by the facts that the organization of the
parasite is definitely characteiistic and that Gadiis ceglefinus
figuies as the best in both instances.
Cobbold^s description is embodied in a few lines, but this
brevity is remedied by a boldly drawn figure. In inter-
preting this figure w^e must suppose either that he drew from
the living animal, when the tliickness of the body would
prevent him seeing clearly the organs in the posterior part,
or that his preserved specimens were not sufficiently cleared.
Odhner’s amended descriptions of this species and others did not
come to hand until the present paper was in the press.
78
m. AV. Nicoll on the
The anterior region is eorrect exeopt tliat tlie genital aper-
ture is rather far forward and the eutieular spines have
assumed enormous proportions. The ventral sueker has a
somewhat powerful appearance^ and the vesieula semitialis
extends as far baek as the anterior testis. • Tlie testes are
correctly enough placed^ but the ovary is posterior to them
instead of in front. The uterus displays tlie greatest
divergence; it is represented as two tubes^ winding down^
one on each side of the body, to the extreme posterior end.
It is to be presumed that Cobbold here confused the uterus
with the vitelline glands, the rounded follicles of which
might be mistaken for ova. The uterine walls must have
been supplied from imagination.
Cobbold fancies a resemblance between this species and
Distomum scabmm, Zed., and D. appendiculatum, Rud.
Wherein this lies I fail to perceive, even in Cobbold^s figure,
for the organization of the alimentary system with its long
pre-pharynx, the presence of the large spines, and the dispo-
sition of the entire genital system are widely different from
the coi’responding structures in the above-mentioned species.
Stossiclds description of this form is merely a brief trans-
lation of Cobbold^s. Along with von Linstow* he falls
into the error of attributing it to the cod [Gadas morrhua)
instead of Gadus ceglejinus, although Cobbold is perfectly
clear on this point.
A not uncommon parasite of the haddock, it occurred in
rather more than 50 per cent, of individuals examined,
always in the intestine and never in large numbers. It is
of moderate size : length l-88-I‘47 mm., maximum breadth
•61-‘90 mm. ; average size 2'65 x '59 mm. ; body of elon-
gated oval outline, rounded at both ends, somewhat attenuate
anteriorly. It is of compact build, sluggish in movement,
and does not long survive removal from its host. The
cuticle is beset with stout regularly arranged spines cover-
ing the whole of the ant-acetabular region and gradually
disappearing behind the ventral sucker. A few spines are
to be found laterally within a short distance of tne tip of
the tail. The length of the spines is about '012 mm., but
they are shorter in front and longer posteriorly. They have
comparatively broad bases and are arranged so that the
spines of each row alternate with those of adjacent rows.
The oral sucker is almost terminal, of fair size (diameter
•24-'38 mm.), but not very muscular. The ventral sucker is
remarkably small and feeble ; it is situated almost centrally
* Oompend. d. Ilelminthol. 1878, p. 238.
Entozoa of J3rillnh Marine Fishes. 70
and has a diameter of '11— '23 mm. Both arc circular in
outline with circular apertures.
The alimentary system is well developed. Tlic mouth
opens into a long pre-pharynx (about *3 mm. long in an
average specimen), which is followed by a pharynx of large
size ('21 X'lG mm.) ; the oesophagus is very short and the
diverticula into which it divides are of great width and
extend to the end of the body.
The excretory vesicle is a simple sac of no great extent, in
the posterior end of the body and opens by a terminal pore.
The testes are median, one behind the other, in the pos-
terior third of the body. They are globular or ovoid in
shape and of considerable size ('25 mm, diameter). The
edge of the posterior testis is at a distance of about '6 mm.
from the end of the body. The ovary lies directly in
front of the testes and is much smaller than them. The
receptaculum seminis is an elongated vesicle lying between
the ovary and the anterior testis. The vitelline glands are
extensive and well-defined ; situated laterally from the level
of the ventral sucker to the posterior extremity of the body.
They consist of numerous, compact, irregular follicles.
The ova are few in number (about 30) and are confined to
the space bounded by the ovary, the ventral sucker, and the
two intestinal diverticula. Light yellow in colour, ovoid
in shape, they measure *059-'068 mm. in length and
•033-*040 mm. in breadth.
The genital aperture is in front of the ventral sucker, but
to the left of the middle line; it is within the intestinal
fork. The sphincter muscle surrounding it is sometimes
very prominent.
Idle penis-sac (PI. I. fig. 4) is of large size and divi ded
into two portions by a constriction. The anterior part is
the penis-sac proper, containing the retracted penis and the
prostate, neither of which is very large. This part is
regularly ovoid and lies in front of and dorsal to the ventral
sucker. Joined to this by a narrow neck is a large sac con-
taining a long rather narrow vesicula seminalis. This sac is
capable of considerable extension and contraction, and in
the latter state the vesicula seminalis becomes bent up, as is
shown in the figure. There is a short non-prostatic part of
the ductus ejaculatorius issuing from the anterior end of the
seminal vesicle.
Linton describes a form from Gadiis callarias, which he
identifies as Distomum rachion, Cobbold, or a species very
near it. His description is short, but the measurements lie
notes for the various organs show a remarkable agreement
80
Mr. W. Nicoll on the
■with tliose obtained from my specimens. It deals mainly
witli the external appearancc_, and as his figure is poor little
exact knowledge of tlie more important internal organs is to
be derived from it. The suckers and alimentary canal are
distinctly reproduced, as are also the testes. The penis-sac
cirrus-pouch occupies its proper position, but the
genital a})crture is in the middle line almost directly over
the intestinal bifurcation. Two round bodies are figured in
front of the testes, but their nature is not noted. From
analogy the posterior of the two bodies would represent the
ovary, but it is much larger than I am accustomed to see it,
and, in fact, is as large as either of the testes. In the same
way the anterior body would be the vesicula seminalis, but
it is much further behind the ventral sucker than in my
specimens. The yolk-glands are not well indicated. A most
important feature of difference lies in the arrangement of
the spines, which, if Linton’s figure is to be depended on,
would distinctly differentiate his specimen. He represents
them as scales (from the anterior region), in close array,
overlapping but not arranged alternately. On no part have
I observed such an arrangement; the spines are certainly
scale-like on the neck, but they are well spaced and always
alternate with those of the next row. Thus, in the absence
of more exact information, it is impossible to determine
whether Linton’s one specimen is Lepodora rachiesa, Cobbold,
or not, but it is certainly very near it.
Subfamily Eciiinostomin^, Looss.
Genus Stephanochasmus, Looss.
1899. Steplianostomum, Lss. Zool. Jalirh. Syst. xii. p. 576.
1900. Stephanochasmus, Lss. Zool. Anz. xxiii. p. 603.
Stephanochasmus baccatus, sp. n. (PI, II. figs. 5-7.)
Of this species I have been able to obtain only one speci-
men. It occurred in the rectum of a halibut [Hippoglossus
vulgaris) , and at first sight appeared to correspond so closely
with my recollection of Stephanochasmus cesticillus, Molin,
that I regarded it as such and placed it aside. A note
by Looss * on some examples of this latter form, drawing
attention to an error or variation in the number and
arrangement of the circumoral spines, induced me to re-
examine my specimen. Several features of difference at
once presented themselves; moreover, comparison with the
^ Zool. Jaliuh. Sjst. xii. p. 696.
81
Kntozoa of Jirif/sh Marine FlsI/es,
other species of the genus did not admit its inclusion witli
any of them. It thus falls to be described as a new species.
The following description as well as the figure are from
the preserved specimen, so that the measurements may
admit of modification : —
The body is depressed, elongate-ovate, somewhat attenuate
in front, rounded behind, with a small but distinct terminal
prominence. Length 3’34 mm. ; maximum breadth (at
ventral sucker) ’75 ram. Ant-acetabular region (neck) com-
prises of length of body. Anteriorly the cuticle is beset
with numerous irregularly arranged spines, becoming fewer
behind the ventral sucker and absent from the greater part
of the post-acetabular region. Closely apposed to the margin
of the oral sucker are two rows of large spines. The spines
in the first row are shorter than those in the second, the
lengths being about ’031 mm. and *037 mm. respectively,
but there is some variation. The number and disposition of
these spines are characteristic of the species, and serve to
distinguish it from other species of the genus. They occur
in two regular uninterrupted rows ; there are 28 spines in
each row, making a total of 56, and the spines of one row
alternate with those of the other. In no other species are
the spines so numerous, the nearest approximation being
48, as recorded by Looss in Stephanochasmus caducus, Lss.
Steph. pristis, Deslongch., according to Looss, has 36 spines.
A point which Looss lays stress on is that in his examples
of Steph. cesticillus the second row of spines numbers one
less than the first. This is due to the absence of a spine of
the second row in the mid- ventral line. No such arrange-
ment occurs in Steph. haccatus\ each row is complete, so
that there is no gap in the mid-ventral line. In Steph. ces-
ticillus, moreover, the spines of the first row are longer
than those of the second. Between the two species another
feature of difference presents itself in the disposition of the
other cuticular spines. In Looss^s figure these do not start
immediately behind the cephalic spines, so that a small
triangular bare area is left. This does not occur in Steph.
baccatus, for irregular scattered spines are seen on the neck
immediately behind the cephalic spines.
The suckers are comparatively small and not very mus-
cular. The oral sucker is terminal and cup-shaped ; its
diameter is ’23 mm., while the aperture is T9 mm. At the
lateral margins of the rim, projecting into the aperture, a
little nodule is apparent. Whether this is a natural con-
dition or the result of preservation 1 am unable to say. The
ventral sucker is at a distance of *87 mm. from the anterior
Ann. & Mag. N. Hist. Ser. 7. FeZ. xix. 6
82
Mr. W. Nicoll on the
end. Tt is approximafely globular^ and lias a diameter of
•J3.S mm. with a eircnlar aperture.
The aliment, ry system is fairly typical. The prepharynx
is shorter than in Stej/h. cesticiUus^ being only *17 mm. long.
The pharynx is large and almost horseshoe-shaped^ with the
convexity directed backwards; it measures *21 x '16 mm.
The oesophagus is extremely short. The diverticula extend
to the posterior extremity, and are wide and irregularly
dilat((l. The bifurcation occurs just a little in front of the
ventral sucker.
The testes are situated in the ])osterior third of the body,
one behind the other in the middle line. In shape they are
ovoid, and have a greatest diameter (longitudinal) of ’38 mm.
The ovary lies directly in front, almost median or a little to
tlie right, and is less than the testes (*21 mm ). The vitelline
glands are lateral, extending from the posterior ei d of ti c
body to a short distance behind the ventral sucker ; they
consist of small compact follicles.
There are very few ova (17), but they are of large size,
measuring '087-*091 mm. x •0ff-'053 mm. The shape is
unusual, pointed at one end and truncated at the other
The uterus lies entirely between the ovary and the ventral
sucker, and is contined laterally by the intestinal diverticula.
There is a long club-shaped penis-sac extending some
distance behind the ventral sucker, though not so elongated
as in bteph. cesticillus. The posterior portion is occupied by
a large ovoid vesicula seminalis. There is a considerable
pnrs prostatica. The genital aperture is median, directly in
front of the ventral sucker.
It is evident that this species difftrs very considerably
from already knowm forms. In the first place the body is
much less elongated, and the suckers are proportionately
further apart and larger. The cephalic spines, besides being
more numerous than in any other species, are differently
disposed, those in the anterior row^ being shorter than those
Three ova were observed lying within the ventral sucker. This
recalls a condition already noted (Ann. & Mag. Nat. Hist. (7) xvii. p. 620)
in a parasite inhabiting the cloaca of L(mis argentatus. The suppesition
which I then hazarded seems to be strengthened by this case, liulh are
forms living in the terminal portion of the gut of their host, and the
possibility suggests itself that in both the ova may be retained for some
time within the ventral sucker in order to pi event their being excreted
in too immature a condition. Many forms having a similar habitat
display adaptations in the ova, having apparently the same end in view.
The precocious development of eye-spots in several species and the
ciliated embryos in Diatomum viviparum, Olsson (from tl e end-gut of
rieuronectid fishes), are illustrative cases.
83
Entozoa of British Marine. Fishes.
in tlie postci’ior. In Sleph. minulus, Looss*, they are of
equal size in the two rows, hut in the other species the
anterior row contains the larger spines. In addition the
ova are much larger than those of any other species except
Steph. cesiicillus,
Zoo(/onoldes vivlparus, Olsson.
(PI.* 11. fig. 8 ; PI. III. fig. 9.)
Distomum viviparum. Olsson, Lunds Univ. Arsskrift, 1867, iv. no. 8,
p. 28, pi. iv. figs. 73-75.
ZoogoTuis viviparus, Looss, Centralbl. Bakt. Iste Abtlieil. xxix. p. 440.
Zoogonoides viviparus, Odlmer, Centralbl. Bakt. Iste Abtlieil. xxxi.
p'. 62, fig. 2.
This species, first discovered by Olsson in Pleuronectes
microcephalus and incorrectly described by him, has been
fully described by Odlmer. Looss assigned it to the genus
Zoogonus, Lss., along with Z. miruSj Lss., but Odhner with
reason regarded it as the type of a new genus. Olsson was
only able to discover two specimens, occurring, as he believed,
in the stomach of the host. Odhner, however, correctly
points out that the true habitat of the species is the terminal
portion of the intestine. He found it in Pleuronectes flesus,
PL limanda, PL platessa, PL microcephalus , Hippoglossoides
platessoides, and Callionymus lyra. I have found it here in
PL limanda, PL platessa, and Rhombus maximus, and
always in the lower reaches of the intestine, particularly the
rectum. It always occurs in large numbers, usually accom-
panied in Pleuronectes limanda by tiering ophor us furciger,
Ol'Son.
Odhner^s description of the species is almost exhaustive.
The limits of size which he gives, however, are rather
narrow. I have found mature examples as small as *8 mm.
and, in Rhombus maidmus, as large as 1*6 mm. ; in Pleuro-
nectes limanda they never exceed 1*2 mm. The maximum
breadth is •31-*42 mm. The diameter of the oral sucker
lies within Odhner’s limits, viz. ‘Id-HG mm. - but the ventral
sucker is never twice as large, as Odhner has it, and its
aperture is nearly circular, or if elliptical the eccentricity is
small. The cuticular spines are very minute, arranged in a
regular diamond pattern, and cover the whole body except
a small part at the posterior end. The intestinal diverticula
do not extend beyond the posterior border of the ventral
sucker.
The testes are two ovoid bodies, situated about the level
• Centralbl. fiir Bakter. Iste Abtheil. xxix. p. 604, figs. 5, 5 a.
6*
Si
Mr. W. NIcoll on the
of the ])oslerior border of the ventral sueker and symmetri-
eally plaeed, one on citlier side of tins. In the living speei-
men it is diffieult to fix the exaet position of the testes and
ovary, for they move baekwards and forwards with the move-
ments of the animal. Sometimes one testis is a little further
forward than the other, and sometimes the ovary is on a level
with one or both. In preserved spceimens tlie position is
])retty mueh as shown in the figure (PI. II. fig. 8, T).
^riie ovary is to the rear of the ventral sucker, almost median
or somewhat to the right. It is smaller and more globular
than the testes. I find the size of the miracidium-containing
capsule to be •088-'094 x •0I2-*04I mm., which is larger
than Odlmer has it.
Subfam. IIemiurinje (Looss, ex p.), Liilie.
Looss * included in this subfamily the appendieulate
Distomes and their congeners. Ltihet, however, saw fit to
restrict the name to a certain group of these forms repre-
sented by two genera, Hemim'us, Rud. (ex p.), and Lecitho-
cladium, n. g. Odhner later J removed Hemiurus crenatus,
Rud., from the former genus and made it the type of a
distinct genus, Br achy phallus. Of the members of this sub-
family we have here to deal with Hemiurus appendiculatus,
Rud., and Brachyphallus crenatus, Rud.
Genus Hemiurus, Rud. (ex p.), Liihe.
Within this genus Liihe includes only H. appendiculatus
(Rud., 1802), H. Stossichii (Montic., 1891), H. crenalus,
Rud., 1802 {=Distomum ocreatum., Olsson, 1867), H. Icevis,
Linton, and H. grandipurus, Moiin. These all agree in
having the yolk-glands compact, rounded or only slightly
lobed, situated close behind the ovary and almost touching
each other. The features in which they present differences
are, amongst others, the relative sizes of the suckers, the pro-
portionate length of the appendix, the position of the genital
aperture, the length of the pars prostatica, and the position of
the vesicula seminalis.
I have had before me during my investigation specimens
of appendieulate Distomes belonging to the genus Hemiurus
from four different hosts: — Clupea harengus, Hippoglossus
* Zool. .Tahrb. Syst. xii. p. 640.
t Zool. Anzeig. xxiv. p. 694.
t Fauna Arctica, iv. (2) pp. 3o2, 653.
85
Entozoa of British Marine Fisher,
vuUjaris, Gadus (pghfinus, and Ammodtjtes toblanus. From
the first tlirce Distomuni appendiciilaliim, Rud., has already
been recorded, but, as Ltdie points out, identification has
in many cases been totally erroneous. Each of the four
varieties which I have examined agree well enough on
superficial inspection with the accepted idea of Distomum
appendiculatum, Rud., but on closer investigation they are
found to differ each from the other to a greater or less
degree and all from Liihe’s amended definition * of Hemiurus
appendiculatus, Rud.; they approach more nearly to H. Stos-
sichii (Montic.), Liihe t. The chief features of difference
between the above-mentioned varieties are the situation of
the testes and the length of the pars prostatica and con-
sequent position of the seminal vesicle. In these respects
the examples from the haddock, sand-eel, and halibut agree
closely with each other and differ markedly from the herring
specimens. We must therefore regard the former as distinct
from the latter. To fully reconcile either with already
existing descriptions and in particular with the definitions
of Liihe is impossible. Eor the present, however, I shall
include the specimens from Clupea harengus under Hemiurus
Liihei, Odlmer, while those from Ammodytes tobianu^ are
noticed under Hemiurus communis^ Odhner.
Hemiurus Liihei, Odhner \^ = H. Siossichii (Montic.), Liihe],
From the stomach and emeum of Clupea harengus.
The already recorded host of this form is Clupea pilchardus,
and the fact that this fish is a member of the same family as
the herring strengthens the probability that the forms from
both are identical. Yv^ithout attempting a full description
of the species I shall endeavour to make clear wherein my
specimens differ from Liihe'’s definition.
Its occurrence in the herring was frequent and in great
numbers, and it w’’as the only intestinal parasite to be met
with. The body is very ranch elongated. The length,
including the extended appendix, is 2'70-4T4 mm. Of this
the appendix comprises about l~\ {i. e. the appendix = of
the rest of the body). The breadth was measured at two
points, (1) just behind the ventral suckers, (2) immediately
in front of the appendix ; it was found remarkably constant,
viz. *28-’33 mm. x •35-*40 mm., representing a somewhat
more attenuate condition than Liihe admits in H. Stossic/iii.
The length of the neck (taken as the distance between the
* Zool. Anzeig. xxiv, p. 396.
t Ibid. p. 39S.
8(5
Mr. W. Nicoll orj (he
centres of llie two suckers) has also a fairly constant pra-
])ortion. It was inrn., representing a pro[)ortion of
of the body (without appendix). The oral sucker has a*
diameter of T1-T3 mm. and the ventral •18-24 mm. The
relative proportion may therefore be approximately 1 : 2 or
2 : 3, not invariably 2 : 3 as Liihe has it. Tlie pharynx is
contiguous with the oral sucker and has a length of 'Ob mm.
with a breadth of •06-*07 mm. The oesophagus is very
short or absent. The intestinal diverticula may extend into
the appendix, but usually do not. The cuticle of the body
is ringed geringelt that of the appendix is deeply
striated.
The genital aperture lies a short distance behind the
ventral lip of the oral sucker ; the genital sinus is long and
extends almost as far back as the ventral sucker. The pars
prostatica is also of great length ; the prostate cells are very
numerous and form a dense mass behind the ventral sucker.
In consequence of the length of the prostate, the vesicula
seminalis is at a great distance behind the ventral sucker.
The distance as measured is from *6 mm. to *9 mm., i. e,
rather more tlian ^ of the body-length. The seminal vesicle
is double and usually lies towards the left side. The testes are
just behind it and one is almost directly behind the other,
the obliquity being very slight. They are globular or some-
what ovoid and of fair size (diam. •14-'16 mm.). The ovary
and vitelline glands lie almost midway between the testes
and the beginning of the appendix. The ovary is always
elliptical, the long axis being transverse, and measures
•13-T8 X Tl-’12 mm. It is thus about the same size as the
testes. The yolk-glands lie close behind and ventral to the
ovary, and are each about the same size as the ovary ; they
are slightly lobed, one usually having three, the other four
lobes. This coincides, curiously enough, with Liihe’s
observations in the cases where the yolk-glands are tubular.
The glands are at a distance of *4-'6 mm. from the appendix.
The uterus is very voluminous and may extend a short dis-
tance into the appendix, but usually does not. Tlie ova are
numerous and measure •022-’028 mm. X •011- 012 mm.
Htmiurus coimuunis , Odhner.
From the oesophagus, stomach, and intestine of Ammodytes
iobianus and from the stomach of Gadus ceglefinus.
This species shows very great resemblance both to
H. appendiculaius and to H. Liihei. It is ditferentiated from
the former by the situation of the testes, wdiich are close
87
Entozoa of Urltish Marine Fishes.
beliiiul tlie ventral sucker and markedly obli(|ue ; also by
the relative sizes of the suckers. Ih’om H. Lu/iei it differs
ill the length of the a-sophagiis, in the shortness of the pars
prostatica, and in the forward position of the vcsiciila senii-
nalis and testes. It differs from both in the length of the
appendix.
Mature examples were found in the sand-eel, varying in
length (including appendix) from r5-2 5 mm. Of this the
appendix often comprised as much as | (i. e. | of the rest of
the body) ; usually somewhat smaller, but never less than
half as long as the rest of the body. The examples from the
liaddock and the halibut agree in this respect. The body is
not so attenuated as in the form from Ciupea harengas^ the
breadth being ’37-'88 mm. just behind the ventral sucker
and '42-‘57 in front of the appendix. The neck (distance
between centres of suckers) is also much longer than in the
above form, being ^ of the body-length. The oral sucker
has a diameter of T1-T8 mm. and the ventral a diameter of
T6-’26 mm., so that the relative size is from ^ to more
usually the latter. The pharynx is contiguous with the oral
sucker and measures ’06 x '07. There is a distinct cesophagus,
about as long as or longer than the pharynx. The intestinal
diverticula may extend a considerable distance into the
appendix, but often stop short of it. The cuticle of the
body is ringed, but the rings become faint towards the
appendix. The striations on. the latter are not very distinct.
It IS usually divided into two portions of almost equal length,
the posterior of which becomes invaginated within the other
when the appendix is retracted.
The genital aperture occupies the same position as in the
preceding species; the genital sinus is not quite so long.
The pars prostatica is shorter and the seminal vesicle, which
is double, is immediately behind the ventral sucker. The testes
are globular or ovoid and are not far behind the seminal
vesicle ; somewhat obliquely situated, the left is almost
entirely in front of the right. They have a diameter of
*10-' 16 mm. The ovary is ovoid, the long axis being trans-
verse, and measures Tl-'16 mm. X '09 mm. The yolk-glands
are behind the ovary and contiguous with it; they are usually
slightly lobed as in the preceding species. The uterus is
voluminous and may extend into the appendix. The ova
are numerous and measure •022-'031 mm. x '009-013 mm.
The specimens from Gadus ceglefinus present slight
variations from the above and tend more to H. appendix
culatus, but they certainly cannot be included under that
species according to Llihe’s definition.
88
.Mr. W. Nicoll on the
An interesting oceurrence was witnessed while a living
speeirnen from the sand-eel was- being examined. The left
yolk-gland was observed to gradually, but rapidly disappear,
so that not a traee was left. The preserved speeirnen
exhibits only one yolk-gland. This may afford a possible
explanation of Monticelli^s observation of ordy one gland
in Hemiurus S^tossichii, a case which Liihe has difficulty in
reconciling with his own observations
In an example of Hemiuriifi appendiculatus from the halibut
(PI. III. fig. II) a rare condition was met with, namely,
pigment-spots in the ova. The presence of eye-spots in the
ova of Hemiurinse is not usual, and on that account I
regarded the case as suspicious. On close examination some
of the ova displayed two spots, one at each pole ; others had
several spots congregated at one end, but in the majority
the appearance simulated the usual occurrence of eye-
spots. Indications of pigmentation, however, were found
throughout the body and large patches were discovered in
the substance of both suckers. Another smaller example
showed no spots in the ova, but the suckers contained several
black patches. The explanation of this seems difficult ; the
case is certainly unique in my experience, and I can only
attribute it at present to some diseased condition.
By' achy phallus ci'enaiiis (Hud.), Liihe.
Distoma crenntum, Rud. Entoz. Hist. ii. p. 404, pi. v. fig. 1.
O
Distoma ocreatum, Olsson, Lunds Univ. Arsskrift, iv. (8) p. 48, pi. v.
figs. 90-98.
Hemiurus crenatus, Liihe, Zool. Anzeig. xxiv. p. 399 ; Lander, Bull.
Mus. Harvard, xlv. no. 1.
Brachyphallus Odhner, Fauna Arctica, iv. (2) p. 352.
Amongst the numerous examples of Hemiurus communis
from the sand-eel several specimens of this species were
found. It is at once differentiated by the almost equal size
of the suckers. The length of my examples, which were
pretty nearly all of one size, was 2T 2-2*38 mm., with an
appendix of 1*08-1*18 mm., i. e. the appendix equals about
half the length of the rest of the body. This corresponds
very well with the observations of Lander and Odhner. Liihe
makes the length I *25-1 '35 mm. and the appendix J as long
as the trunk. The breadth behind the ventral sucker was
*52-‘64 mm., and near the appendix it was '7\-‘77 mm.
These figures are much in excess of any found by other
observers.
* Zool. Aiizeiger, xxiv. p. 399, note 13.
89
Entozoa of British Marine Fishes.
The diameter of tlie oral suekor was •21-’26 mm., that of
the ventral sucker *28-‘8() mm., the pro[)ortioii therefore
l)ein^ () : 7, which is identic al with that found by Lander and
Odhner. The genital aperture lies midway between the
suckers: Lander says nearer the oral sucker, Odhner nearer
the ventral. The appendix always apj)ears in two parts and
the intestinal diverticula reach to nearly its extreme end.
It is deeply striated.
The })harynx is contiguous with the oral sucker. Between
the observations of the above observers there is some diver-
gence, Odhner^s dimensions being much less than those of
Lander. I have found it to vary, the breadth as often as
not exceeding the length. Length '09-T2 mm., diameter
'09-Tl mm. These figures agree more with Lander’s. In
my specimen the left testis is further forward than the right,
which disagrees with both Odhner and Lander’s figures. The
ovary is globular or ovoid and smaller than the testes. The
yolk-glands are usually compact, but sometimes they are
more or less scattered, a part being occasionally found at
some distance from the main mass and connected witli it by
a narrower portion. In one case single follicles were ob-
served scattered throughout the body near the appendix.
The ova measure •02I-’023 mm.X •015-’0I6 mm. ; they are
thus somewhat broader than those measured by Odhner.”
Subfamily Lecithochieiin^, Liihe.
Genus Lecithaster, Liihe.
Lecithaster gibbosus, Bud. [ = L. mollissimus^ Levinsen] .
From a sand-eel [Ammodytes tobianus) I obtained what I
believe to be a specimen of this parasite. It occurred only
once, although more than fifty sand-eels were examined, so
that it is extremely rare. It was, unfortunately, damaged
during inspection, but not before I had observed the following
particulars : —
A large vesicula seminalis lay at the level of the ventral
sucker and dorsal to it. A ductus ejaculatorius, with pro-
static cells, stretched forward to open a little way behind the
oral sucker. In the posterior part of the body the yolk-
glands were arranged in seven or eight lobes radiating from
a common centre. There was a small caudal appendix. In
the position of the testes were situated two circular bodies,
displaying internally a system of concentric rings. For this
appearance I am unable to account. The ovary could not be
90
Mr. W. Nicoll on the
distinguished, and tliere were no ova (so tliat the specimen
was probably immature). Together with the size of the
body and the condition of the suckers these characters seem
sufficient to estal)lish identity with Distomum mollissimum as
described by Levinsen
Genus Derogenes, Liilie.
Verogenes various, Muller.
This has proved one of the most widely distributed Trema-
todes in this locality, and has occurred in the following
hosts : — Gadus merlangus (intestine), HippogJossus vulgaris
(stomach). Rhombus maximus (oral cavity, oesophagus, and
stomach b Rhombus Icevis (stomach), Pleuronectes lunanda
(oral cavity, oesophagus, and -stomach), and Coitus scorpius
(intestine). It is here recorded for the first time from
Rhombus maximus and Pleuronectes limanda, and in these it
has occurred most frec^uently.
Its characters are so well known that it calls for no descrip-
tion here. The limits of size are wide, Stossich giving them
as T5-7 mm. I have found adult specimens less than 1 mm.
long, and the largest example obtained was 3*4 mm. long
from the stomach of the turbot. The most common size is
1*5-2 0 mm. The oral sucker in individuals of that size has
a diameter of *19-*24 mm., and the ventral sucker *33-*40 mm.
The ova vary greatly in specimens from different hosts. The
limits in length are *050-*062 mm., and in breadth *025-
*034 mm. Those in front of the ventral sucker are always
larger than those behind, and it is in the latter that the
greatest variations are to be observed. Thus in an example
from the halibut the ova in the anterior part of the body
measured *059 X *032 mm., in the posterior part *057 X *0i7
mm. The approximate size of the ova just about to be
extruded may be taken as *056-*060 x *031- 034 mm.
Derogenes cacozelus, sp. n. (PI. III. fig. 10.)
Found in the intestine and rectum of Hippoglossus vulgaris
and the intestine of Pleuronectes limanda. It is not at all
common, only a few specimens having been obtained. It
bears a close resemblance to Derogenes various, Muller, but
the large vesicula seminalis and the small ova differentiate it
at once from that species.
The body is elongated, cylindrical, broadest in the middle.
♦ Orersigt Kgl. Uansk. Selskab. 1881, pp. o9-61, pi. ii. fig. 4.
91
Entozoa of British Marine Fishes,
la|)erlii«* towards cadi end. Tjcii^tli •87-1*51 mm. The
section is approximately circular. Tlic cuticle is unarmed,
l)iit very faint tiansvcrsc wrinkU's apjicar on the surface,
^riic oral sucker is subtcrminal, globular, with a diameter of
•16 mm. (in a specimen of length 1*5 inrn.). The vcntial
sucker is somewhat prominent, globular, and lies in front of
tlie middle of the body (*57 mm. ITom anterior end) ; its
diameter is *23 mm. Both have circular apertures.
There is a large muscular pharynx, almost globular, with
a diameter of *08 mm. The oesophagus is extremely short
and the intestinal bifurcation takes place immediately behind
the pharynx. The diverticula are wide, irregularly dilated,
and extend to the posterior extremity of the body. The
excretory system resembles that of Derogenes varicus.
The testes are obliquely placed behind the ventral sucker;
they are two ovoid bodies, with a maximum diameter of
*12 mm. The ovary is more nearly globular : diameter
*10 mm. It is situated almost midway between t’ e ventral
sucker and the tip of the tail. Behind it lies a pair of large
vitelline glands having the same structure and disposition as
in Derogenes varicus The uterus is much convoluted and
fills a large portion of the body. The eggs are very numerous
and rather small, ovoid in shape. Size 021-*023 mm. X
•014- 016 mm.
There is a large ovoid vesicula seminalis lying on the same
level as the ventral sucker and dorsal to it, measuring
•23x‘17 mm. From it issues a somewhat narrow ductus
ejaculatorius, leading forward to the penis-sac. The duct is
surrounded for two thirds of its length by numerous
prostate-gland cells lying free in the body-substance. The
penis-sac resembles that in Derogenes varicus. It is some-
what pear-shaped or almost globular. The aperture is
situated in the raid- ventral line at a distance of ‘21 mm.
from the edge of the oral sucker.
Distomum sp. (PI. III. fig. 12 ; PI. IV. fig. 13.)
From the muscles &c. of Coitus bubalis.
While collecting one day I observed a Coitus lying in a
small rock-pool. I approached it cautiously, but it seemed
unaware of my presence until I seized it, and even then its
struggles to escape were feeble. In a tank it took no notice
of food or objects placed near it, but lay torpid. It appeared
to be blind. On dissection the cause was revealed. The
whole body, skin, muscles, bones, and the layers of the eye
were impregnated w'itli small masses of black pigment,
92
Mr. W. Nicoll on the
accompanied by cysts contairiinj^ Trematode cercariie. The
only ])arts not affected were the brain and the abdominal
or^atis. The pigment-spots appeared to follow the course of
tlie })lood-vcssels, as is evident from fig. 13^, and they are
probably spread throughout the body by means of the blood.
The nature of tlie pigment T did not ascertain. The Trema-
tode is a small tailless larva^ having the body entirely
covered vvitli minute spines, two small suckers, and with
intestinal diverticula extending to the tip of tlie tail. I am
at present unable to assign it to any known species.
Idiis occurrence may admit of explanation in the same
way as Johnstone* accounts for a similar infection of
Plearoriectes limanda, although the parasites in the two cases
are not identical.
Distomum sp.
From gills of Cottas scorpius and Gobius Rathensparri in
capsules.
This occurred rarely, but in infected specimens the
numbers were large. The wall of the cyst was thin, so that
the eni3losed larva could be easilv^ seen. It possessed few
distinctive features, so that identification was, for the time
being, impossible.
N EMATODA.
Ascaropsis morrhu(je, van Beneden. (PI. IV. figs. 14-16.)
Ascaropsis morrhuce, van Beneden, Mem. Acad. Belg. 1871, xxxviii.
p. 56, pi. hi. fig. 11.
Van Beneden appears to have instituted this genus and
species at one and the same time, bat he gives no definition
of the genus or description of the species except a few words
in a footnote and drawings of the head, tail, and ovum.
From these, however, I am able to establish the identity of
the specimens which I assign to this species.
Van Beneden found it in the intestine and pyloric caeca of
the cod [Gadus morrhua). I have to record it from Gadus
(sgleJinuSj Hippoglossus vulgaris, and Coitus bubalis. In each
of the two latter only one specimen occurred, but in the
haddock it was extremely numerous and was met with
throughout the whole anterior part of the alimentary canal.
The body is elongated, narrow, and cylindrical, of almost
uniform girth ; attenuated anteriorly and posteriorly, with a
* Keport Lancashire Sea-Fisheries Laboratory for 190-1, p. 101.
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Kntozoa of British Marine Fishes.
ventral excavation at the tail. Length 6-8 niin., breadth
*8-’9 nun. Cuticle annnlatcd by furrows, which are con-
tinnons for more than one annulus. Van l^cneden represents
each ring as complete, i)robably owing to bis not having
studied the character of the annulation. Towards the ante-
rior end the furrows become faint and disappear. Projecting
forward from the head are two small spines, omitted by
van Beneden ; these are largest in the example from Coitus
bubalis, and almost invisible in that from the halibut.
The month is terminal and appears to consist of two
equal-sized but little-differentiated lips. There is a long
oesophagus and a long simple intestine pursuing an uneven
course towards the anus, wliieh opens in the excavation at
the tail. In adult specimens the ovary and uterus occupy
almost the whole of the remainder of the body. The ovary
arises at the posterior extremity of the body and passes
forward, twisting lound the intestine in its course. The
middle third of the body is usually completely filled with ova.
These have the characteristic shape, with the two flagella at
one pole, noted by van Beneden. Size •039-’040 ram. x
•021 -‘022 mm. These measurements are almost uniform in
the specimens from each of the three hosts.
A. form somewhat resembling this was taken from the
intestine of Pleuronectes microcephalus. It displayed the
cliaracteristic annulation, but on the head, instead of two
spines, appeared two prominent circular marks. No ova or
genitalia were present, so that the specimen was immature
and identification was impossible.
EXPLANATION OF THE PLATES.
The following letters apply to all the ligures : —
BSN. Ventral sucker.
CB. Penis-sac.
DB. Ductus ejaculatorius.
l)Sf. Vitelline glands.
B.v. Excretory vesicle.
J. Intestinal diverticula.
KSt. Ovary.
LC. Laurer’s canal.
MSA'. Oral sucker.
Oe. QEsophagus.
Ov. Ova.
P. Penis.
PG. Genital aperture.
Ph Pharynx.
PPh. Prepharynx.
PPr. Pars prostatica.
Pr. Prostate glands.
1\, 1\. Testes.
BS. Receptaculum seminis.
SD. Shell-gland.
Ut. Uterus.
1"'g. Vagina.
/ S. Vesicula seminalis.
Plate I.
Fig. 1. Podocolyle ntomon, Rud. Anterior part, to show prepharvnx •
dorsal view. * ’
01
^.lessrs. J. C. lUelvIIl and II. Ponsonby on
Fig. 2. Vodocotyle atomo7i, Kud. Sliell-gland complex ; ventral view.
J)G, yolk-duct; YR, yolk-receptacle; KG, oviduct.
Fig. 3. J.epodora rachicea, Cokbold. Ventral aspect.
Fig. 4. Ditto. Male genital apparatus ; peuis retracted : dorsal view,
Plate II.
Fig. /). Sti‘phanoehns77ius haccatus, sp. u. Ventral aspect.
Fig. 6. Ditto. Head, showing spines ; ventral aspect.
Fig. 7. Ditto. Outline of ovum.
Fig. 8. Zoogonoides viviparus, Olsson. Ventral aspect. Em, embryo.
Plate III.
Fig. 9. Zoogonoides viviparus, Olsson. Penis-sac ; penis retracted,
showing the spines ; ventral view.
Fig. 10. Derogenes cacozelus, sp, n. Right lateral aspect.
Fig. 11. Hemiurus appendiculatus, Rud., from Hippoglossus vulgaris.
Loop of uterus, showing ova with pigment-spots.
Fig. 12. Head of Coitus hubalis, Euphr. Eye removed, to show pigment-
spots in the eye-socket.
Plate IV.
Fig. 13. Head of Coitus buFdis, Euphr. Lower jaw divided and turned
to the sides ; gills bent hack, to show the pigment-spots in
the roof of the mouth, following the course of the blood-
vessels.
Fig. 14. Ascaropsis morrliuce, van Ben,, from Gadus ceglefinus. a, cepha-
lic spines ; b, termination of annulating furrow ; c, end of
oesophagus ; d, ova.
Fig. 15. Ascaropsis morrliuce, van Ben., from Coitus buhalis. Highly
enlarged view of head, showing the two spines.
Fig.\Q>. Ditto. Tail; ventral aspect. intestine ; anus.
The drawings, with the exceptions of figs. 2, 4, 8, 9, were made from
preserved specimens.
VIII. — Descriptions of Fifteen Terrestrial MoUusca from South
Africa. By James Cosmo Melvill, M.A., F.L.S., and
John Henry Ponsonby, F.Z.S.
[Plate VI.]
Three year.s having now elapsed since our last communica-
tion we now venture to otfer an eigditeenih contribution
upon the subject, maiidy consisting of the descriptions of seven
Ennecie several Helicoids, mostly collected by Miss Hickey,
Messrs. J. McBean, J. Farquhar, and H. C. Burnnp, to
whom our best acknowledgments are due, while we would
* Ann. k Mag. Nat. Hist. ser. 7, vol. xii. (December 1903), pp. 595 sipj.
05
Mollusca fro)u South A frica.
take this opportunity of especially thanking the last-named
for his continued valuable critical assistance.
Ennea Alicioij sp. n. (PI. VF. tig. 1.)
E. testa rimulata, obtuso-cylindrica, laevissima, pellucida, tenui,
nitida, lubrica ; anfractibus 9, quorum apicales 3 subconici, caeteris
ventricosulis, apud suturas improssis, ultimo paullum producto, ad
basin arete striate ; apertura semicirculari, peristomate incras-
sato, nitido, plicis dentibusve quatuor munito : plica parietali
acinaciformi, acuta, intrante, dente labiali bi- vel tritido, magiio,
plica basali incurva, columellari omnirio interna, bimamillata,
magna, aperturam fere claudente.
Long. 9-5, lat. 4*5 mm.
Ilah. Makowe, Zululand (Burnup) .
A large and handsome species, slightly recalling E. for-
mosa., M. & P., iti general appearance; but it is a much
smoother shell, with subconical apex, whorls broader and
tun-shaped, mouth-processes differing in the absence of a
colurnellar tooth, in the tendency of the labial tooth to become
trifid, the internal colurnellar plait being almost identical in
both species.
To l\lrs. Alice Burnup, companion of her husband on so
many successful expeditions in South Africa, this conspicuous
Ennea is dedicated.
Ennea johanneshurgensis, sp. n. (PI. VI. fig. 2.)
E. testa parva, subrimata, dolioliformi, solidula, pallide straminea,
apice obtusissimo ; anfractibus 7, paullum ventricosis, apud suturas
impressis, undique longitudinaliter obliquistriatis ; apertura fere
rotunda ; peristomate ^bo, nitido, incrassato : plica parietali
acuta, intrante, fere recta, dente labiali acuto, basali obtuso, plica
coiuuiellari conspicue et omnino intrante, rotundata, mammaeformi.
Long. 6, lat. 2 mm.
Hah. Johannesburg, Transvaal [McBean, Johnson).
More than a dozen examples of this species have been
inspected, besides one or two that were received a few years
since from Mr. Johnson, shoi tly after his first arrival in !South
Africa. One of these slightly exceeds 6 mm. in length. It
belongs to the alliance of S. eximia^ Gollieri^ Wottoni^
M. & P., or infans, Craven, from which it chiefly differs in
the presence of an obtuse basal tooth.
Ennea phragma ^ , sp. n. (PL VI. fig. 3.)
E. testa cylindrica, dolioliformi, apud basin pone labrum conspicue
scrobiculata, recta, parva, pallide albo-straminea, apicem versus
* ([ipayna, a palisade or fence.
Messrs. J. 0. Melvill and J. IT. Ponsonby on
Of)
planata ; anfractibiis, inclusis apicalibus, 7, ad suturas impressis,
undi(pie arete lonj^itudinaliter liratis, liris obliquis; apertura
ovata; ])eristornate incrassato, sinuoso, albo, nitido, dentibus
j)licisve quatiior muiiito : plica parietali conspicua, magnopere
iiitraiite, dente labiali crasso, acute, basali parvo, plica colu-
luellari bifida, superficialiter dentiformi, multum intraute, subtus
mammaeforrai.
Alt. 3-5, diam. 1’5 mm.
Hah. Waku District, Catlicart, near the Klipplatz River
{Miss Ilickei/').
Smaller, narrower, and more cylindrical than E. drakens-
hergensis^ M. & P., and in general form more resembling
E. caryatis^ M. & P., but differing from that species entirely'
as regards the disposition of the oral plaits and teeth. It
may be compared also with E. montana, M. & P., from
Mountain Drive, Grrahamstown, the aperture of which is
trigonous, with finer and closer spirals on the whorls.
Ennca stauroiua*, sp. n. (PI. VI. fig. 4.)
E. testa subrimata, ovato-oblonga, laevigata, nitida ; anfractibus,
apicali incluso, 8, omnibus pellucidis, perlaevibus, vitreis ; aper-
tura rotunda ; peristomate incrassato, nitente, albido, plicis
dentibusve complicatis fortiter munito : plica parietali crassa,
labiali magnopere intrante, 3- vel 4-partita, devia, dente basali
minore, squarroso, plica columellari omnino interna, tridentata,
magna, aperturam semiclaudente.
Alt. 6, diam. 3 mm.
Hah. Melmoth, Zululand, at 3000 feet, with no water
near the spot'’^ [AHss Ilich-y).
Several examples of a pellucid, very smooth and shining
Ennea^ eight-whorled, oval or oval-oblong in form, with very
complicated peristomatal processes, the most peculiar being
the thrice or four times divided labial tooth or plait, proceeding
deeply within, and irregularly serrulate. The moderately-
sized basal tooth is squarely oblong, the columellar process
wdiolly internal, half closing the orifice below.
We take as the ty])e the ovate form. One specimen is
decidedly oblong, in all other respects similar.
Ennea vallaris^^ sp. n. (PI. VI. fig. 5.)
E. testa breviter cylindrica, crassa, parva, solidula ; anfractibus 8,
apice ipso obtusissimo, cseteris arctissime obliquicostulatis ;
apertura subrotunda ; peristomate albo, nitido, crassiusculo,
* cTTavpwixa, a palisade, from the oral processes,
t Val/ans, belonging to a fortification or rampart.
97
Mollnsca from South Africa.
plicis dentibusve 5 praedito : plica parietali acinaciformi, intrantc,
plicis duabu8 labialibus, cxtus dentiforrnibus, longe intrantibua,
denfce basali aimplicc, aciito, parvo, plica columellari raammaoformi,
profunda, aperturam fere claudente.
Alt. 4, diam. 1-5 mm.
Hah. Meirnotb, Ziiluland (ilfm Tficlcey).
Four examples, all precisely similar, of a small peculiarly
incrassate species, something like E. crassidem^ Pfr., and
other members of that alliance in miniature. The peiisto-
matal processes are very marked, almost between them closing
the roundish aperture.
Erinea xysila sp. n. (PI. VI. figs. 6, 6 «.)
E. testa recta, cylindrica, variabili, parum rimata, nunc majore,
solida, albo-cinerea, longitudinaliter indistincte et oblique crassi-
costata, nunc (var. hyalina) laevi, pellucente, nitida ; anfractibus
7-9, quorum apicales obtusissimi, subplanati, cmteris rectis ;
apertura ovata vel ovato-rotunda ; peristomate nitido, albo, in-
crassato, dentibus plicisve quinque instructo : plica parictali acuta,
longe intrante, dentibus duobus labialibus, quorum superior
bifidus, dente basali conspicuo, acuto, plica columellari omnino
interna, magna, maraillata.
Long. 5, lat. 2 mm. (sp. min., var. hyalina).
„ 8, „ 3 „ (sp. maj.).
Hah. Johannesburg, Transvaal {Johnson^ McBean) ; Pre-
toria (^McBean) .
A variable species, the smaller, very smooth, shining,
transparent form (var. hyalina) offering considerable contrast
to the large, more coarsely longitudinally striate shell which
attains the maximum development as given above, and was
collected by Mr. Johnson.
The mouth-processes in both are virtually identical. With
the larger form E. vanstaadensis^ M. & P., is comparable, as
also are E. instahilis and ampullacea., Stur. [ = ohovataj Pir. ?),
but our species differs in the more cylindrical contour and
bolder disposition of peristomatal plaits. Another marked
characteristic in E. xysila is the bifurcation of the upper
labial tooth.
Our best thanks are due to Mr. Henry C. Burnup for
having devoted much time to the elucidation of this very
interesting but variable species.
Ennea zelota'\, sp. n. (PI. VI. fig. 7.)
E. testa ovato-oblonga, politissima, obtecte rimata, apud apicem
* ^vaiKos, smoothish.
t (rjXcoTos, enviable.
Ann. (k Mag. N. Hist. Ser. 7. VoL xix.
7
98
Messrs. J. C. Melvill and J. H. Ponsonby on
obtusa, albo-hyalina, vel pallide straminea ; anfractibus 8, parum
ventricosis, loevissirais, striis longitudinalibus fere evanidis, non-
nisi apud basin (vel, in uno speciraine, obscurissirae juxta
suturas) ; apertura ovata ; peristomate albo, nibido, sex plicis
dentibusve instriicto : plica parietali conspicua, dente ad sinistram
minute addito, dentibiis labialibus duobus, inferiore forti, bifido,
superiore minute, dente basali bene definite, celiimellari magna,
baud prefunde intrante.
Leng. 5, lat. 2 mm.
Hah. Port Shepstone {Bm^nup).
A remarkable A'/iweri, without a near ally in the disposition of
its very elaborate peristomatal processes. The sliell is smooth,
shining, oblong-ovate, remarkably obtuse, with only occa-
sional faint signs of striation ; there exists a small parietal
tooth near the strong deep-seated plait; the labial teeth are
two in number, the lower being very well defined and bifid ;
the basal and columellar teeth are likewise conspicuous.
Naialina arguta sp. n. (PL VI. fig. 8.)
N. testa profunde umbilicata, Igete rufo-brunnea, depresso-orbiculari,
undiqiie arete longitudinaliter striata, apud basin nitidiore ; an-
fractibus ad 5, ventricosulis, ultimo magno ; apertura rotundo-
lunari ; peristomate, prsecipue apud regionem columellarem,
paullum reflexo.
Alt. 14, diam. 21 mm.
Hah. East London (Biirniip).
Allied to A. Trimeni^ M. & P., that species being more
globose, of olivaceous rather than rufous-brown hue, not so
regularly striate throughout, the base being smoother and far
more shining, the umbilicus narrow, and aperture slightly
oblique. N. arguta^ in brief, is a brighter-coloured and
handsomer form in every way.
Natalina instgnts, sp. n. (PI. VI. fig. 9.)
A. testa magna, depresso-globosa, profunde sed anguste umbilicata,
sordide olivacea ; anfractibus ad 5, convexiusculis, longitudinaliter
rudi-striatis, striis irregularibus, subtus apud basin nitidam fere
evanidis ; apertura rotundo-lunari ; peristomate apud marginem
paullum incrassato, ad umbilicum triangulatim reflexo.
Alt. 23, diam. 30 mm. (sp. maj.).
Huh. Teafontein, Grahamstown.
May be compared with N. euynactaj M. & P., a more
♦ Arffutus^ clear, distinct.
99
Mollusca from South Africa,
globular species, and lihytida Kraussii, Pfr.* We have only
as yet seen two examples of this interesting mollusk, the first
being in the British Museum (Natural History). This is
taken as the type.
Ildicarion astlienes t, sp. n. (PI. VI. fig. 10.)
H. testa ampla, suborbiculari, laete olivacea, vel rufescenti-cornea,
teimi ; spira dcpresso-conica, tenui ; anf'ractibus 5, ultimo rapide
accrescente, magno, superficie undique sericea, paullum nitida
(prmcipiie ad basin in statu juvenili) ; apertura lata, ovali ;
peristomate tenuissimo.
Alt. 13, diam. 21 mm.
Hah, Cradock (/. Farquhar),
A brightly coloured shell, o£ a yellowish or rufous olive,
somewhat depressed, thin, effuse ; surface entirely silky,
shining only in young examples basally. In size approxi-
mating//./wsaWor, M. & P., which is only four-whorled,
with slightly iridescent epidermis, and of a peculiar warm
sepia coloration.
Trachycystis paula^ sp. n. (PI. VI. fig. 11.)
T. testa minuta, umbilicata, depresso-globosa, cornea, tenui ; an-
fractibus 4, ventricosis, undique longitudinaliter sparsim lirato-
plicatis, liris irregularibus, rarius arctis, hie illic rugulosis,
obliquis ; apertura rotunda ; peristomate tenui, apud umbilicum
triangulatim reflexo.
Alt. 1, diam. 1*75 mm.
Hah, Johannesburg [J, McBean).
Very small, with narrow but deep umbilicus, depressedly
globose, four-whorled, the plaited lirse being irregular,
obliquely disposed, sometimes wrinkled, and rarely close-set.
A few specimens. It seems comparable with T, somerset-
ensis^ M. & P.J, both of these species being only provisionally
located in the genus Trachycyslis.
Trachycystis rotula, sp. n. (PI. VI. figs. 12, 12 a.)
T. testa minuta, depressa, profundissime umbilicata, cornea, delicata,
tenui, interne circa basin concentrice sulcata ; anfractibus 5-6,
* The opportunity may be taken to mention that further comparison
of Dorcasia inhluzana, M. & P. (Ann. & Mag. Nat. Hist. 1894, vol. xiv.
p. 91, pi. i. fig. 4; 1895, vol. xv. pi. xii. figs. 6, 0«), with Rhytida
Kraussii, Pfr., convinces us they must, if not varieties of each other, be
placed in the same genus,
t d<rdhr)s, from its fragility.
300 Messrs. J. C. Melvill and J. H. Ponsonby on
angustis, ventricosuHs, apud siituras multura impressis, undiqiie
sub lente arctissime lirafcis, liris tenuibus ; apertura lunari, peri-
stomate tenui.
Alt. 75, diam. 1’5 mm.
II ah. Fern Kloof, Grabamstown [J. Farquhar).
Another exceedingly minute species, with deep perspective
umbilicus. With the aid of a lens the whole surface, which
is horny and subpellucid, is seen to be very closely lirato-
striate. There is a single broadish spiral groove present,
running through the centre of the body- whorl, but not quite
extending to the orifice, this being an unusual character, and
one which we have not before met with in any other South-
African Helicoid. A few examples only.
Trachycystis spissicosta sp. n. (PI. VI. fig. 13.)
T. testa parva, depresso-globosa, umbilicata, tenui, subpellucida,
cornea ; anfractibus 4, ad suturas multum impressis, apice ipso
Isevi, aliter undique arctissime longitudinaliter tenuiliratis ; aper-
tura rotundo-lunari, peristomate tenuissimo.
Alt. 4, diam. 6*5 mm.
Hah. Near the racecourse, Grahamstown {J. Farquhar) .
A pretty species, of the same alliance as T. hisculpta,
Bens., hathycosle and epetrima^ M. & P., &c., very strongly
costulate, thin, horny, globosely depressed in form, and
narrowly but deeply umbilicate.
? Fliasts sollers'l, sp. n. (PL VI. fig. 14.)
Ph. testa anguste umbilicata, tenui, delicata, pallide straminea,
globoso-conica ; anfractibus 5, apud suturas impressis, undique,
praecipue superne, irregulariter et oblique plicato-liratulis, super-
ficie Omni, praecipue ad basin laevigatam, spiraliter tenuissime
Bub lente sericeo-striata, ultimo ad peripheriam angulato ; aper-
tura late lunari ; peristomate tenui, marginem apud columellarem,
supra umbilicum, triangulatim reflexo.
Alt. 4*5, diam. 7 mm.
Hah. Melmoth, Zululand [Miss Hickey).
A very few examples only have yet been received of a
little Helicoid with a certain resemblance to Phasis capensis^
Pfr., but entirely differing in texture, as seen Ly the above
description. The spiral, silky, close striation is only visible
with a pov/erful lens. We are not quite sure of its right
generic location, but provisionally place it in Phasis.
* Spissus, costa, with close-set ribs,
t Sollers, adroit, expert.
AruuAM(/^.Nal.JIi.si.S.7. Voi.IlX. PI VI.
UNWLHSiTir
Mollusca from South Africa. 101
Euonyma platyacme^ sp. n. (PI. VI. fig. 15.)
E. testa elongata, imperforata, albida, epidermide olivacea contacta,
hand iiitcnte, tenui ; anfractibiis 12, quorum apicales duo obtusi,
claviilati, parvi,'tres su])eriu liis proximi recti, angusti, caeteris
leiiiter accrcscentibus, uiidi(jue irregulariter obli(]uo rudi-plicatis,
ultimo et peiiultimo tore rectis ; apertura ovata ; peristomate
tenuissimo.
Alt. 38, diam. 11 mm. (sp. maj.).
Hah, Kei Road Bush {Miss Ilickei/^ May 1906).
Semi-fossilized specimens, with but scant remains of
epidermis, liave as yet come to hand, and though in the
opinion of tlie discoverer it may be an “extinct species” (Miss
Hickey, in litt.), we consider the evidence of the fragmentary
epidermis militates against this theory. The gradually
increasing whorls, club-shaped apex, and absence of any
umbilical perforation characterize this species, which differs
considerably from any form hitherto known from the South-
African region. We place it in the genus Euonyma^ recently
monographed * by Pilsbiy, who has included in it not only
the original sinistral type {E. loeocochlis ^ M. ^ P.), but like-
wise ^Hulimus^^ turriformis, Krauss, and its allies. Com-
pared with its nearest ally, E. Lanceolata (Pfr.), the propor-
tionate. size of the last whorl is striking. This last species
is, so far as is yet known, confined to Natal.
EXPLANATION OF PLATE VI .
Fig. 1. Ennea Alicice.
Fig, 2, johanneshurgensis.
Fig. 3. phragma.
Fig. 4. stauroma.
Fig. 5. vullaris.
Fig. 6. xysila.
Fig. Qu. , var. Jiyalina.
Fig. 7. zelota.
Fig. 8. Natalina arguta.
Fig. 9. insignis.
Fig. 10. Helicarion asthenes.
Fig. 11. Trachycystis paula.
Fig. 12. rotula.
Fig. 13. spissicosta.
Fig. 14. ? Fhasis sollers.
Fig. 15. Euonyma platyacme^
* Man. Conch., Pulmonata, vol. p. 38,
102
M iss M. Y. Lebour on
JX. — On I'hree Molhislc-infesting Trematodes. By Marie V.
Lebour^ B.Sc., Armstrong College, Newcastle-upon-
Tyne.
[Plates VII. & VIII.]
Muddy and sandy flats which are left uncovered by the tide
for several hours of the day and are the haunts of numerous
sea-birds are specially good ground for larval Trematodes and
the mollusks and other invertebrates which harbour these
worms. Fenham Flats, the land between Holy Island and
the mainland of Northumberland, is one of these good localities,
and the shore of Loch Byan, at Stranraer, in Galloway, is
another. Numerous Trematodes are to be found in both
places, three of which seem to be specially worthy of attention
and are here described.
The greater part of the flats near Holy Island is uncovered
for several hours each day, but the Mussel Scaup, which lies
to the south -w^est, remains almost completely covered except
at spring-tides. Here the ground is more gravelly and many
mollusks abound. Cockles are fairly numerous, but perhaps
the commonest shell is the little white whelk Purpura
lapiUus^ which preys continually on the mussels and is found
in clusters all over the scaup. It is, however, a very small
form.
The shores of Loch Eyan at Stranraer are chiefly made
up of muddy sand, with large stones strewn about, and the
tide leaves a long stretch of ground uncovered for several
hours each day. The usual common mollusks are found,
Purpura lopillus being particularly large and fine.
Tlie Trematodes to which I should like to draw attention
are all species of Distomum. The first infests the cockle,
Cardium edule, and w^as noticed some time ago in one cockle
out of many examined from Budle Bay, Northumberland,
and was imperfectly described and figured by me North-
umberland Fisheries Eeport^ for 1905, p. 100, pi. ii. fig. 3).
Mr. James Johnstone, of the Liverpool Fisheries Labora-
tory, sent me another specimen of the cockle infested with
this parasite from Yeoman Wharf, Morecambe Bay, and kindly
allows me to use his drawing of the living sporocysts
(PI. YII. A). In October 1906 I again found this Trematode
in a cockle from the Mussel Scaup, Holy Island, and am now
able to describe it more in detail. The visceral mass of the
cockle, which is whitish and unhealthy-looking, is almost
completely riddled with sporocysts, especially wdiere the
103
Mollusk-vifesting Trematodes.
gonad usually is, but tliat organ is quite obliterated. The
liver is more or less infected, in the Holy Island specimen
hardly at all. Each sporocyst is about O'G mm. long and
O’ 15 mm. broad. It is colourless, transparent, and some are
very contractile, assuming all kinds of shapes, while others
are inert and move little. Inside the sporoeysts are small
granules and large opaque round masses from two to twelve
in number, which, on closer examination, are seen to be
encysted cercariae. More or less developed tailed cercari^
are also sometimes to be seen in the S[)orocysts along with
the encysted forms. They are very contractile and are con-
stantly shifting their position inside the sporocyst, the tail
moving incessantly. The cercaria sheds its tail before
encysting, and in several sporoeysts tails were to be seen
moving quickly round the encysted worms when only these
latter were present in the sporoeysts. The free tails were
excessively active. The cyst is about 0’14 mnn across and
is very thin. The enclosed cercaria is seen to be covered
with short spines and has two suckers, both usually visible
when the worm is curled up in the cyst. When pressed out
from the cyst it is about 0*19 mm. long (see PI. VII. B), the
spines covering it are conspicuous and form circular rows
round the anterior sucker and all over the body. The ante-
rior sucker is large (O’OI mm. across), and leads by a short
oesophagus to a thick-lipped pharynx {which, however, often
appears to be continuous with the sucker owing to the con-
tracting of the animal). The pharynx leads to the intestine,
which almost immediately bifurcates into two lobes, reaching
nearly to the posterior end. The small ventral sucker is
slightly posterior to the centre of the body. There is a large
clear excretory sac opening posteriorly, but excretory canals
could not be distinctly made out The tailed cercaria (see
PI. VII. C) is much the same, but the spines had not appeared
in those I saw and the intestine did not branch so soon,
probably because the worm was stretching itself more than
the encysted form. Two ducts each side of the anterior
sucker can be indistinctly seen, probably coming from the
glands which serve for the secretion of the material for the
cyst, as they disappear in the later stages, after the worm
has encysted.. The intestine is very indistinct, owing to the
opacity of the animal, and can only be completely seen by
staining.
The less-developed cercarise in the sporoeysts are small,
granular, oval masses, then a trace of a tail appears, after
that the clear excretory sac is seen, then the anterior sucker,
and later the posterior sucker. The cercaria may be half the
104
Miss M. V. Lebour on
size of the encysted form and possess both suckers, pharynx,
excretory sac, and tail.
It is remarkable that the cercaria sliould possess a tail
W’hen it encysts in tlie sporocyst, as tlie organ must be quite
nseless. The encysting within the sporocyst is not usual,
but Filippi states tliat the Tretnatode wliich he describes as
Cercaria echinatoides encysts in tlie same Paludina in which
it was developed from redia3 without quitting that mollusk,
and it has a tail wliich it throws off before encysting (Ann.
des Sci. Nat. 4® ser., Zool. 1854, p. 255).
The above-mentioned Trematocle from the cockle is not
common — one infested specinnn occurring from Budle Bay
in March 1905 out of two hundred examined, one found
by Mr. James Jolinstone from MorecambeBay in April 1906,
and one out of one hundred examined from Holy Island in
October 1906. In the two first-mentioned specimens found
in March and April no tailed cercaria^ w^ere seen. They were
found only in the cockle from Holy Island . in October.
Possibly examination of many cockles in the summer months
may show us younger stages of the worm.
The second Trematode to be described is from the common
w helk. Purpura lapillus, from Holy Island and also on the
shores of Loch Pyan to the west of Stranraer. Two hundred
and eight specimens from the Mussel Scaup, Holy Island,
w'ere examined in October 1906, and the liver in two of these
was found to be infested by a Trematode. At Stranraer at
the beginning of September 1906 it occurred more abun-
dantly, five specimens out of thirteen containing it. The
liver in healthy specimens of Purpura lapillus is a bright
yellow, but wdien this parasite is present it is a yellowish
white and is full of long and not very active rediae, which are
transparent and colourless, wfith the exception of the sac-like
intestine, which is coloured yellow from granules of food-
material (PI. VIII. B). Each redia is about 1’8 mm. long,
with a conspicuous pharyiix and intestine, and is full of
rather opaque cercarise in various stages of development.
The cercaria (see PI. VHI. A) is tailed, and when full-grown
is about 0*45 mm. in length without its tail, wdiich is about
two thirds the length of tlie body. The head-region is more
transparent than the rest of the body, and is separated from
it by a slight constriction ; the body is coarsely granular and
opaque and is covered (but only in fully developed speeimens)
with blunt spines. Tlie tail is blunt, not so opaque as the
body, and, as usual, comes off at the slightest touch. A large
anterior sucker leads into a short oesophagus, which is often
obscured by contraction, and this leads by a thick-lipped and
105
MoUusk -infest in g Tre rn a todes .
conspicuous pharynx to a narrow intestine bifurcating about
a third of the way down the i)ody, each lobe reaching nearly
to the posterior end. Two clear excretory canals beginning
in the head-region run down the sides of the body, and join
just belbre they reach the clear excretory sac opening
posteriorly. Tiie ventral sucker, which is larger than the
aTiterior, is placed slightly behind the centre of the body,
d’he cercaria is very contractile, and when moving uses its
body more than its tail, shortening and elongating itself
continuall3^
It is an interesting fact that these specially large and fine-
looking Purpura lapillus from Stranraer are much more
frequently infested with this Trematode than the poor under-
sized specimens from Holy Island. One would imagine that
the reason for this is the greater prevalence at Stranraer of
the host in which the adult worm lives, in all probability a
sea-bird.
The third Trematode is not unlike the second, but is some-
what slimmer in build and has a distinct double row of spines
round the head. It occurred in one specimen of the common
limpet, Patella vulgata^ out of sixteen examined from the
shores of Loch Hyan, east of Stranraer, in September 1906,
completely obliterating the gonad and riddling the liver.
The worm occurred as long narrow redige about 1*8 mm. long,
transparent and colourless (see PI. YIII. Dj . A pharynx can
be distinctly seen, but I could make out no enteron in any of
the specimens. Tailed cercarise fill the redige and move about
inside them, contracting and elongating in much the same
way as those from Purpura lapillus. The blunt tail is less
than half the length of the animal (see PI. VIII. C). The
cercaria is about 0*50 mm. long without the tail ; its head is
transparent, surrounded by a collar of two rows of fine spines,
and is eonstricted off slightly from the rest of the body, which
is coarsely granular and opaque, and in fully developed
specimens covered with short blunt spines. A large anterior
sucker leads to a narrow oesophagus, which leads by a small
pharynx to a narrow intestine running down for about two
fifths of the length of the body before it branches into two
lobes nearly reaching to the posterior end. Two very
granular excretory canals begin in the head-region, run down
the sides of the body, and pass into a clear excretory sac
opening posteriorly. A large ventral sucker occurs behind
the centre of the body. This worm seems to be an Echino^
stomum^ and the last species was so like this that one would
expect it to belong to the same subgenus; but no spines were
seen on the head.
106
Mr. W. M. Tattersall on new
I think these tliree larval Trematodes are new to our
fauna ; no other stages in their life-history have yet been
foimd.
EXPLANATION OF THE PLATES.
Plate VII.
Trematocle from Carclium edule.
A. Sporocysts enclosino- cercarise.
IL Cercaria pressed out of cyst.
C. Tailed cercaria.
Plate VIII.
A & B. Trematode from Fw'imra lapillus.
A. Cercaria.
B. Bedia enclosing cercarioe.
C & D. Trematode from Pattella vuUjata,
C. Cercaria.
D. Redia enclosing cercariae.
X. — PreUminary Diagnoses of Six new Mysidse from the
West Coast of Ireland. By W. M. Tattersall, B.Sc.,
Department of Agriculture and Technical Instruction,
Fisheries Branch, Dublin.
The six new forms, of which preliminary descriptions are
now offered, were captured off the southern part of the west
coast of Ireland in depths ranging from 465 to 800 fathoms,
by the S.S. ^ Helga,'’ the fishery cruiser of the Department
of Agriculture for Ireland. These depths have been but
rarely reached by the bottom-fishing apparatus of the ^ Helga,^
and the fact that thus early in their exploration six new
species of Mysidse have been brought to light suggests the
existence of a fauna rich in undescribed forms.
All six species belong to the subfamily Leptomysince * of
the Alysidse. Two are types of new and interesting genera,
while the other four belong to two recently defined deep-
water genera characterized by the imperfectly developed eyes
possibly modified for tactile functions.
Genus Metamblyops, nov.
Characters generally as in Amblyops, G. O. Sars, except : —
Carapace produced in front into a well-developed, rather
long and acute rostrum.
I cannot agree with Norman’s recent proposal to raise his sub-
families to family rank (c/i Norman and Scott, ‘ Crustacea of Devon and
Cornwall,’ London, 1906y
‘-IBHAfty
UNIVEfisiVonu
Ah„. .V Ma.</. A,il. \ 7. r»/. A7,V. /'/. 17/,
Ana. <S)‘ M(((f. Aa/. Ilisf. A. 7- \ ol. XL\. /*l. 17//.
0^
JUNIYEKSITY oi ILUNOib
107
^lyslclai from the West Coast of Ireland.
Eyes well developed, normal in appearance and structure,
pigment light reddish brown.
Telson entire, lauciforni in shape, its margin armed with
more or fewer spines, median setae absent.
Inner uropod with a few spines on its inner margin in the
region of the otocyst.
Female with only two pairs of incubatory lamellae.
Type species, MetamhJyops oculata.
The genus Chalcophthalmiis, lllig, 1906, would appear to
be rather closely allied to the present one, but judging from
lllig’s figures Metamblyops is a more compact and robust
form, the carapace covers all the thoracic segments, the eye
is much larger and its papilla quite minute, and the antennal
scale comparatively much longer. The chief distinction lies
in the structure of the first thoracic limbs, which in Chalco-
phthaimus are described as being devoid of endopods, while
in M eiamhlyops both endopod and exopod are fully and
normally developed.
Metamblyops oculata ^ sp. n.
Carapace covering all the thoracic segments ; produced in
front into a slightly upturned, acute rostrum reaching as far
as, or a little beyond, the eyes, and partially covering the
eye-stalks ; terminal angle about 60°, its apex produced into
a short acute point; evenly rounded at the antero-lateral
corners and slightly emarginate behind.
Pleon longer than carapace ; the first segment one and a
half times as long as the second, which is subequal to the
third and fourth ; fifth segment slightly longer than the
fourth ; sixth segment twice as long as the fifth.
Eyes large, well developed and normal in structure ; ex-
tending to the distal margin of the first joint of the anten-
nular peduncle ; pigmented portion equal in width to the
last pleon-segment, a minute papilla on the inner distal part
of the peduncle where it joins the cornea ; visual elements
well developed ; pigment light reddish brown.
Antennular peduncle about twice as long as the eye; third
joint a little shorter, but considerably stouter than the first
and more robust in the male than in the female ; second
joint small ; male appendage well developed and densely
hirsute, but otherwise of normal appearance.
Antennal peduncle short, not extending beyond the distal
end of the second joint of the antennular peduncle and com-
posed of three subequal quadrangular joints.
Antennal scale about one-third as long again as the anten-^
108
Mr. W. M. Tattersall on neio
imlar peduncle and twice as long as the antennal ; about
three and a half times as long as broad ; outer margin entire
and terminating in a strong spine, beyond which tlie apex of
tlie scale is not produced ; spine on the outer distal margin
ot‘ the basal joint quite short
Mouth-parts not exhibiting any striking points of difference
from those of Amhhjops ahbreviatci, except that the second
joint of the mandibular l)alp is considerably broader.
First thoracic limb with the endopod almost exactly as in
Amblyups abbreviata.
Second thoracic limb with the endopod of the same form as
in A. abbreviata, but comparatively much longer ; twice as
long as that of the first thoracic limb and longer than its
own exopod.
Remaining thoracic limbs rather long and slender, with
the tarsus longer than the merus and composed of three
joints, the third joint longer than the second ; dactylus well
developed.
Eocopods of all thoracic limbs having the basal joint
lamelliform with a small spine at the outer distal corner ;
flagelliform part composed of nine to ten joints.
Incubatory lamellce of the female, two pairs.
Pleopods in the male agreeing essentially with those of the
males of the genus Amblyops.
Telson not quite so long as the last segment of the pleon
and twice as long as broad at its base, where the margins are
slightly expanded ; entire and lanciform in shape, tapering
distally to a narrowly rounded apex j the distal two thirds of
its margins armed with from twenty-eight to thirty-two
spines increasing in length towards the apex ; terminal spine
about one sixteenth of the length of the telson ; median
setae absent from the apex.
Uropods slender : inner, about one and a half times as long
as the telson, with six spines on its internal margin in the
region of the otocyst ; outer, about twice the length of the
telson.
Length of the largest female 16 mm., of the largest male
15 mm. Female with about twenty young in the mar-
su})ium.
Locality. Fourteen females and thirteen males from
S.R. 352, 92 miles S.W. by W. of Bull Rock, Co. Kerry,
lat. 50° N., long. 11° 40' W., 800 fath., August 1906,
Petersen trawl at 750-800 fath.
The external appearance of this species with its large well-
developed eyes and long acute rostrum at first suggests a
species of Boreomysis-, such as B. arctica, but the details of
109
Mjsidse from the West Coast of Ireland.
tlie various appcndaf^es, the number of incubatory lamellae
in the female, and the form of the telson and uropods clearly
indicate its position in the Leptomysime, among the numerous
genera of which Amblyops seems to be its nearest relative.
The characters of the rostrum, eye, antennal scale, and telson
combined abundantly distinguish it from all other genera in
the subfamily.
Genus Dactylerythrops, Holt & Tattersall, 1905.
Non Dactylerythrops, Illig, 1906.
This genus when first described was compared with
Meterijthrops, S. I. Smith. The discovery of two further
species and of the closely allied genus Dactylamblyops,
H. & T., indicates that it is perhaps more nearly related to
the genus Amblyops, G. 0. Sars, and it may thus be more
accurately redefined in the light of this new material as
follows : — Characters generally as in the genus Amblyops,
G. O. Sars, except : —
Eyes small ; not exhibiting any definite eye- stalk, but
joined at their bases by a membranous integument ; visual
elements imperfectly developed, not reaching to the surface
of the eye, but rather deeply seated in its tissues ; outer
distal corner produced into a rather long digitiform flexible
process.
Telson entire, rather small, subtriangular or lanceolate in
shape, lateral margins armed distally with more or fewer
spines, median apical pair of setae present or absent.
Incubatory lamellce in the female, two pairs.
Type species, Dactylerythrops dactylops^ H. & T.
Dactylerythrops arcuata, Illig, should more properly be
referred to the genus Dactylamblyops.
Dactylerythrops bidigitata, sp. n.
Carapace covering all the thoracic segments ; cervical
sulcus well marked ; produced in front into a broadly
rounded obtuse rostrum which reaches to about the centre of
the eyes; antero-lateral corners rounded ; emarginate on its
posterior border.
Pleon longer than the carapace ; the first segment one and
a half times as long as the second, which is subequal to the
third and fourth; fifth segment slightly longer than the
fourth, sixth segment twice as long as the fifth.
E?jes small with their basal parts covered by the rostrum ;
as far as can be seen, joined to each other at the base by a
no
Mr. W. M. Tattersall on new
membranous integument such as is described for D. dactijlops ;
the outer distal corner produced into a rather long digitate
and flexible process ; a shorter and firmer process on the
inner and upper face of the eye, which a raised ridge connects
to the main parts of the eye, so that the whole organ is tri-
angular in cross-section and not flat ; visual elements much
more numerous than in D. dactylops, confined to a triangular
area on the outer part of the eye at the base of the outer
process; pigment confined to the visual elements, pale
purplish pink in freshly preserved specimens.
Antennular peduncle rather short, with the third joint a
little longer and very much stouter than the first and more
swollen in the male than in the female ; second joint small ;
male appendage well developed, but only slightly hirsute, as
the specimen is still immature.
Antennal peduncle shorter than the antennular and having
the second joint slightly larger than the subequal first and
third joints.
Antennal scale almost twice the length of the antennular
peduncle ; about four to four and a half times as long as
broad ; outer margin entire and ending in a strong spine,
beyond which the apex of the scale is not produced ; no
spine on the basal joint.
Mouth-parts fundamentally as in the genus Amblyops.
First and secojid thoracic likewise agreeing with those
of the genus Amblyops, except that the second limb is com-
paratively longer.
Remaining thoracic limbs broken away.
Exopods of all the thoracic limbs with the outer distal
corner of the basal joint slightly acuminate ; flagelliform
part well developed and composed of about seventeen joints.
Incubatory lamellce of the female, two pairs.
Pleopods in the only male, which is immature, have the
inner branch of the first pair already more developed than in
D. dactylops and armed with many more setae ; otherwise
they conform to the Amblyops type.
Telson comparatively short ; about two thirds (slightly
less) of the length of the last segment of the pleon ; sub-
triangular in shape ; one and a half times as long as broad
at its base; margins tapering evenly to a bluntly rounded
apex ; the distal third of its margins armed on each side
with ten rather stout spines increasing slightly in length
towards the apex ; a pair of median plumose setae situated
at the apex between the terminal spines of the margins.
Uropods moderately slender : inner, one and two-thirds of
Ill
Mysldfe/rom the West Coast of Ireland.
tlie length of the telson, apparently without spines on its
inner margin ; outer, twiee the length of the telson.
Length of a mature female 16 mm., of an immature
female 14 mm., and of an immature male 15 mm.
Locality. Two females and one male from S.R. 352, 92
miles S.W. by W. of Bull Rock, Co. Kerry, lat. 50° 22' N.,
long. 11° 40' W., 800 fath., August 1906, Petersen trawl at
750-800 fath.
This species while obviously congeneric with D. dactylops
is abundantly distinguished from it by the better developed
rostrum, the two processes of the eye, the longer antennal
scale, and the greater number of spines arming the margins
of the telson.
Dactylerythrops gracilura, sp. n.
Carapace covering all the thoracic segments ; broadly and
evenly rounded in front, without any trace of a rostral
projection ; antero-lateral corners rounded ; emarginate on
its posterior border.
Pleon a little longer than the carapace ; first segment a
little shorter than the second, which is subequal to tlie third
and fourth ; fifth segment slightly longer than the fourth ;
sixth segment one and a half times as long as the fifth.
Eyes very small, almost entirely covered by the carapace ;
very thin and membranous ; united at their bases by a
membranous integument ; outer distal corners produced into
a very acute fairly long flexible process ; a shorter and less
acute process on the inner corner ; visual elements very
imperfectly developed, confined to a small deeply-seated mass
at the base of the outer process ; no pigment observed in
preserved specimens.
Antennular peduncle rather short ; about half as long as
the antennal scale ; basal joint flattened and broader than
the other two ; second joint small ; third joint the longest,
moderately stoutly built, produced ventrally betw^een the
bases of the two flagella into a short process armed with
about six strong setae ; the whole appendage in lateral view
appears curiously contorted.
Antennal peduncle slightly shorter than the antennular
peduncle and likewise slightly contorted in lateral view ;
rather stout ; second joint the largest.
Antennal scale almost twice as long as the antennular
peduncle ; from two and a half to three times as long as
broad ; outer margin entire, terminating in a strong spine,
beyond which the apex of the scale is produced for a length
112
Mr. W. M. Tattersall on new
equal to about one-eighth of the total length of the scale ;
no spine on the basal joint.
Mouth-parts and first and second thoracic limbs not differing
in any important point from those of the last species.
Remaining thoracic limbs with the tarsus equal to the merus,
three-jointed, the first joint the longest ; nail well developed,
but shorter than the last joint of the tarsus.
Exopods of all the thoracic limbs with the outer distal
corner of the basal joint slightly acuminate ; flagelliform
part of ten joints.
Incubatory lamellcE of the female, two pairs.
Telson diverging somewhat from the type met with in
D. dactylops and D. bidigitata, and shaped almost exactly as
in the genus Meterythrops ) equal in length to the last seg-
ment of the pleon and twice as long as broad at its base
where the margins are somewhat inflated ; margins rapidly
converging to a very narrow truncate apex armed with two
spines set close together and equal in length to one twelfth
of the length of the telson; median setge absent from the
apex ; distal half of the lateral margins armed with about
nineteen short spines.
Uropods slender : inner, about one and a half times as
long as the telson, without spines on its inner ventral margin ;
outer, nearly twice as long as the telson.
Length of a mature female 15 mm.
Locality. Four females from S.F. 352, 92 miles S.W. by
W. of Bull Rock, Co. Kerry, lat. 50° 22' N., long. 11° 40'
W., 800 fath., August 1906, Petersen trawl at 750-800 fath.
This form diverges somewhat from the other two species
of the genus in the shape of the telson, which in its narrowly
lanceolate form and want of apical setm approaches that of
members of the next genus. The eyes, however, conform to
the general type of Dactylerythrops , and to that genus the
species is provisionally referred. The eyes are rather remark-
able and enable the species to be readily distinguished ;
they are almost entirely covered by the carapace, only the
two digitate processes projecting beyond the latter.
Genus Dactylamblyops, Holt & Tattersall, 1906.
Syn. Dactylerythrops, Ulig, 1906.
This genus is undeniably very closely allied to the pre-
ceding one, but may be dislinguished by the following
characters : —
Eye small, with distinct and definite eye-stalks; more or
less pyriform in shape ; visual elements, though imperfectly
113
J\rysl Jix} //’om the West Coast of Ireland.
formed, arc better developed and more numerous than in
Dactylenjthrops, reaching to tlie surface of the eye and
probably directly functional as organs of sight; outer distal
corner rounded and not produced into a digitiform process ;
a short process ahvays present on the inner and upper surface.
Type species, llactijlamhhjops Hodysoni, H. & T.
The type ami the two new species described below appear
to form a natural group chiefly distinguished from the
genus Dactijlerythrops by the above points, and in the present
state of our knowledge of the group this generic division
may well be allowed to stand.
Dactylerytkrops arcuata, Illig, should be referred to this
genus, and is, in fact, synonymous with the type species,
D. Hodgsoni.
Dactylamhlyops thaumatopSy sp. n.
Carapace covering all the thoracic segments except the
la'it ; produced in front into a short, broadly rounded,
obtuse rostrum, which extends to the distal end of the first
joint of the antennular peduncle and partially covers the
eye-stalks ; evenly rounded at the antero-lateral corners and
emarginate behind ; cervical sulcus well marked.
Pleon longer than the carapace ; the first segment a little
longer than the second, which is subequal to the third,
fourth, and fifth; sixth segment twice as long as the fifth.
Eyes small, extending forwards to the distal end of the
second joint of the antennular peduncle ; pyriform in shape,
with distinct eye-stalks ; each eye with a short digitiform
process on the inner and upper face ; a broad membranous
ledge projecting at right angles to the surface of the cornea
starts at the outer lateral part of the eye-stalk and runs
equatorially round the outer part of the eye, terminating
just ventral to the digitiform process and dividing the cornea
into a dorsal and ventral portion ; the ledge is broadest
about the centre of the cornea and narrows off at either end ;
visual elements imperfectly developed, numerous, reaching to
the surface of the eye; pigment pale purplish pink.
Antennular peduncle about twice as long as the eye and
three quarters of the length of the antennal scale; third
joint slightly longer than the first; second joint small.
Antennal peduncle about half as long as the scale ; the
three joints roughly subequal in length.
Antennal scale about one third as long again as the
antennular peduncle and twice as long as the antennal ;
about four times as long as broad ; outer margin entire and
A7in. cb Mag. N. Hist. Ser. 7. VoL xix. 8
114
]\rr. W. M. Tattersall on new
terminating in a spine^ beyond which the a])cx of the scale is
not produced ; spine on the outer distal corner of the basal
joint obsolete.
Mouth-parts and first and second thoracic limbs not differing
in any striking way from those of the type species.
Remaining thoracic limbs missing.
Exopods of all the thoracie limbs with the outer distal
corner of the basal joint slightly acuminate and the fiagelli-
form part composed of ten joints.
Incubatory lamellce of the female, two pairs.
Telson not quite so long as the last segment of the pleon
and once and two thirds as long and broad at its base, where
the margins are somewhat expanded; entire and laneiform
in shape, tapering distally to a narrowly rounded apex ;
distal two thirds of its margins armed with about twenty-four
short spines, increasing in length towards the apex ; median
apical setae absent.
Uropods moderately slender : inner, about one and a half
times as long as the telson, otoeyst rather large, with appa-
rently no spines on its inner margin ; outer, broken in both
specimens.
Length of both specimens (immature females) 11 mm.
Locality. Two immature females from S.li. 352, 92 miles
S.W. of Bull Rock, Co. Kerry, lat. 50° 22' N., long. 11°40'W.,
800 fath., August 1906, Petersen trawl at 750-800 fath.
This species is readily distinguished by the remarkable
structure of the eye, which is one of the most wonderful
among the many varied forms met with in Schizopoda. The
exact function of the external membranous Pdge is not quite
clear. In other characters the species is rather closely allied
to the type.
Dactylamblyops goniops, sp. n.
Carapace covering all the thoracic segments except the
last, which is fully exposed ; evenly rounded in front and
not produced into a rostral projection ; antero-lateral corners
rounded; emarginate behind; cervical sulcus well marked.
Pleon longer than the carapace, first segment slightly
longer than the second, which is subequal to the third,
fourth, and fifth ; sixth segment rather long, two and a half
times as long as the fifth.
Eyes a little larger than in the last species, set close
together and rather subquadrangular than pyriform in dorsal
outline; the digitiform process on the inner and upper face
more slender and a little longer than iu either the type or
115
MjsIJaj from the West Coast of Ireland.
the last species ; visual elements imperfectly developed,
numerous; pigment pale purplish pink.
Antennular peduncle in the female extending sliglitly more
than halfway up the scale, third joint rat’ier shorter and
not wider than the first; in the male relatively a little
longer, with the third joint slightly longer and more robust
than in the female ; male process well developed and hirsute.
Antennal peduncle only slightly shorter than the an-
tennular; third joint the longest and most robust.
Antennal scale extending for rather less than half its
length beyond the antennular peduncle; about three times
as long as broad ; outer margin slightly sinuate and ter-
minating in a very strong spine which projects for its entire
length beyond the apex of the scale ; outer distal corner of
the basal joint prolonged into a long acute spine.
Telson about three quarters of the length of the last
segment of the pleon and slightly less than twice as long as
broad at its base; entire and lanciform in shape, tapering
distally to a narrowly rounded apex; the distal two thirds of
its margin armed with about eighteen long slender spines,
increasing in length towards the apex ; the terminal spines
about one seventh of the total length of the telson ; a single
very slender median spine at the apex between the terminal
spines of the margins ; median setce absent.
Uropods slender : inner, only a little longer than the telson
plus the terminal spines and with four long spines on its
ventral inner margin in the region of the otocyst ; outer,
about one and a half times as long as the telson.
Length of an adult and mature male and female, 10 mm.
The third to the eighth thoracic limbs are broken off in
both specimens. The first and second thoracic limbs and
the mouth-parts, as well as the male pleopods, are in sub-
stantial agreement with the same parts in both the type and
the foregoing species, except that the last joint of the
mandibnlar palp is sliorter and somewhat more robust.
Locality, An adult male and female from S.R. 359,
56 miles W. by N. of Tearaght, Co. Kerry, 465-492 fath.,
August 1906, tow-net on trawl.
Tliis species is at once distinguished from its congeners by
the characters of the eye, antennal scale, telson, and inner
uropods. The sixth pleon-segment is also relatively longer
than in either of the other two species. The antennal scale
resembles rather closely that figured for Paramblyops ros~
trata, H. & T., 1905.
8*
11(5
Mr. W. M, Tattersall on new
Genus Batiiymysis^ nov.
Carapace evenly rounded in front, without any traee of
rostral i)rojection.
Eyes set close together, apparently without definite eye-
stalks, somewhat flattened and sub'quadrangular in shape ;
visual elements imperfeetly developed and unpigmented in
preserved specimens.
Antennal scale shortly lanceolate in shape, setose all round.
Mouth-organs and first and second thoracic limbs as
deseribed by Sars for the genus Leptomysis^ except that the
terminal joint of the palp of the seeond maxilla is expanded
at its apex and armed with numerous short stout spines, the
whole appendage being generally as figured by Sars for
Schistomysis spiritus.
Tarsus of the remaining thoraeie limbs four-jointed ; nail
long and slender.
Telson fairly long ; very deeply cleft, the cleft serrated ;
lateral margins armed throughout their length with spines.
Inner uropod with a row of spines all along its inner
margin.
Pleopods in the male exactly as for the genus Leptomysis.
Type species, Bathymysis Helgce.
In the general structure of the appendages of the thorax
(with the exception of the second maxillse), and especially of
the pleopods of the male, this genus agrees almost exactly
with Leptomysis, G. O. Sars. The chief points of difference
are to be found in the second maxillie, telson, and eyes.
The first two of these structures are interesting as exhibiting
a form met with in many of the genera of the subfamily
Mysince, while the eyes appear to have undergone speeializa-
tion and reduction along lines very similar to Amhlyops^ the
eyes of which they strongly recall, though rather smaller in
size. Bathymysis also bears considerable resemblance to the
genus Pseudomysis, G. O. Sars, but the greatly different
form of the telson at once distinguishes it.
Bathymysis Helgce, sj). n.
Carapace coveiing all the thoracic segments; evenly
rounded in front and at the antero-lateral corners ; without
trace of rostral projection.
Pleon longer than the carapace ; the first segment equal in
length to the fifth and slightly longer than the subequal
117
Myskla'/row2 the West Coast of Ireland.
second, third, and fourth segments ; sixth one and a half
times as long as the liftli.
Eyes strongly recalling those of Amblyops, rather small,
somewhat flattened and SLd)quadrangular in shape, not
reaching to the distal end of the first joint of the antennular
peduncle ; apparently without definite peduncles ; set very
close together; visual elements imperfeetly developed and
without pigment in preserved specimens.
Antennular peduncle a little shorter than the telson, mode-
rately stout, third joint equal in length to the basal two
combined ; male appendage well developed and densely
hirsute.
Antennal peduncle a little shorter than the antennular,
slender, the second joint one and a half times as long as the
third.
Antennal scale equal in length to the telson and a little
longer than the antennular peduncle ; about four and a half
times as long as broad ; shortly lanceolate or oval in shape ;
setose all round ; spine on basal joint almost obsolete.
Mouth-parts and thoracic limbs as described above in the
generic definition. The exopods have the outer corner of the
basal joint rounded and the flagelliform part composed of
twelve joints.
Pleopods of the male as described for Leptomysis. The
fourth pair have the outer ramus longer than the inner;
the last three joints are devoid of setm ; the antepenulti-
mate joint carries a single long and powerful plumose or
barbed spine; a similar but much shorter spine is found on
the penultimate joint, while the terminal joint has two of
these long barbed spines.
Telson a little longer than the last segment of the pleon
and twice as long as broad at its base ; narrowing slightly
towards the apex, where its breadth is equal to one third of
the total length ; cleft very deep and fairly wide, extending
for one third of the total length ; the apical lobe on each
side of the cleft bluntly rounded at its tip ; cleft serrated,
with about thirty spines on each side ; lateral margins armed
throughout their entire length with about forty spines,
which increase slightly in length towards the apex, but there
is no single outstanding long spine at the apex of each lobe
as seen in species of Mysis and Schistomysis.
Uropods broken in the specimen ; inner one with a row of
spines all along its inner ventral margin, thirty-four being
counted on that part of the uropod wdiich remained and
which extended a little beyond the apex of the telson.
1J8
Mr. 0. Thomas on new
Length of the only specimen (a mature male) 15 mm.
J.ocaiity. S.ll. 36T N., lon^. ] 29' W., to
lat. 51° 25' N., long. 11° 3G' W., 620-G95 fath., August
1906, fine iict on trawl.
Two species new to the British and Irish list may also
here he noticed, Hansenomysis Fyllce (Hansen) and Erythrops
microphthalma, G. O. Sars, having been taken in 400-800
fathoms off the coast of Kerry, in August 190G.
List of Authorities quoted.
Holt & Tattklsall, (1905). — “ Scliizopoda from the North-east
Atlantic Slope. ’ Fisheries, Iieland, Sc. Invest, iv. 1902-3.
Holt A Tatteesall, (1906). — “ Iheliiuinary Notice of the Schizopoda
collected by IJ.M.S. ‘ Discovery’ in the Antarctic Region.” Ann.
& Mag, Nat. Hist. ser. 7, vol. xvii.
Illig, (1906).— “ Bericht u. die neiien Schizopoden-Gattung iind
Arten der Deutschen Tiefsee Expedition 1898-1899.” Zoologischen
Anzeiger, Bd. xxx. No. 7.
Saes, (1870-9). — “ Carcin. Bidrag t. Norg. Fauna.” Monog. o. Mysider.
AI. — On further new Mammals obtained by the LUiwenzori
Expedition. By Oldfield Thomas.
Sylviscrex Granti^ sp. n.
A medium-sized species, with tail about equal to the head
and body.
Size much less than in the other Buwenzori specks,
S. lunaris.^ about the same as in the Nyasan S. sorella^.
Fur very long ; hairs of back over 7 mm. in length. General
colour dark slaty grey above, little paler below, but as the
only specimen is in spirit, the tones cannot be described with
accuracy. Hands and feet pale brown, the digits rather
lighter. Tail almost as long as the head and body, finely
haired, brown above, rather paler below.
Skull short, broad, and rounded, not unlike that of S. sorella,
but with a broader flatter brain-case and an even shorter
muzzle. Teeth practically as in S. sorella, the second and
third upper unicuspids rather more nearly subequal.
Dimensions of the type (a spirit-specimen) : —
Head and body 55 mm.; tail 54; hind foot 13T.
Skull : condyio-basal length 17 ; breadth across palate 5*3 ;
* P. Z. S. 1897, p. 930.
1L9
Mammals from Ruwenzori.
breadth across bralii-case 8'0 ; front of to back of wd 0*4 ;
length of lower tooth-row 7.
Uah. lluweiizori East. Alt. 10,000'.
Tt/pe. Adult male in spirit. B.M. no. 6. 12. 4. 85. Col-
lected Olli April, 190G, by E. Dent.
This species is of the intermediate size found in S. sorella,
beiiif^ markedly smaller than S. Innaris and morio^ ami
equally larger than the pigmy S. Johnstoni. b^’om S. sorella
it is at once distinguished by Its far shorter tail.
I have had much pleasure In naming this distinct mountain-
sbrew in honour of my colleague Mi-. W. 11. Ogilvie-Grant,
to whose exertions as originator and manager science Is
indebted for the remarkable results obtained by the lluwenzori
Expedition of Mr. Woosnam and his companions. No less
than twenty new species and subspecies have now been
described from their collections, and the specimens registered
in the Museum collection amount to over three hundred.
Mangos^ gracilis proteus^ subsp. n.
A very variable form, usually strongly affected by
melanism.
In the single wholly non-melanistic specimen the colour
throughout, of body, limbs, and tail (apart from the black
terminal pencil), is grizzled tawny ochraceous, darkened on
the posterior back, the upper surface of the hands and feet
terminally rich rufous. 4h*om this there is a complete series
of intergradations to one in which the body is blackish bistre,
the tail even darker, practically black throughout, and the
feet deep glossy black. The type is an intermediate speci-
men, its general colour mummy- brown, but its feet and tail
wholly blackish.
Dimensions of the type (measured in the flesh) : —
Head and body 806 mm. ; tail 260 ; hind foot 59 ; ear 25.
Skull: condylo-basal length 61.
Ilah. iiuwenzori East. Alt. 7000^ (ranging from 5000').
Type. Adult female. B.M. no. 6. 12. 4. 35. Original
number 115. Collected 13th March, 1906, by R. E. Dent.
This remarkably variable mungoose forms a parallel to the
Alpine squirrels, which are commonly affected by melanism
in a similar way. Possibly something of the same sort occurs
in Abyssinia, where the dark mutgigella” has been con-
sideied to be the same species as the true gracilis. Elsewhere
^ Mungos, E. Geoff. & G. Ciiv. Mag. Encycl. ii. pp. 184-187 (1795).
Herpestes, 111. Prodr. Syst., Mamin, p. 135 (1811).
Cf. Palmer, Index Gen. Mamm. 1904.
120 Mr. 0. Thomas on new
in Africa forms of the gracilis group are very constant in
colour.
No members of this group^ liglit or dark, liavc been
described from the Lake region of Central Africa.
Crossarchus fasciatus macrurus, subsp. n.
Size larger than in irne fasciatus and tail markedly longer.
Colour quite as in Zululand specimens of fasciatus^ the
general tone similarly deep and rich, and the shoulders equally
shaggy haired and suffused with fulvous (not clear grey).
Hands and feet nearly wholly deep glossy black.
Dimensions of the type (measured in the flesh) : —
Head and body 378 mm. ; tail 250 ; hind foot 70 ; ear 2G.
Skull : condylo-basal length 71 ; basal length G7 ; zygo*
inatic breadth 38*5 ; median length of nasals 12*5 ; palatal
length 38 ; greatest diameter of 6.
llah. Kuwenzori South-east. Alt. 3400'.
Type. Male, adult but not old. B.M. no. 6. 12. 4. 20.
Original number 133. Collected 30th April, 1906, by It. E.
Dent. A second specimen (no. 671), collected the same day
by E. B. Woosnam.
Two examples of fasciatus^ collected in Zululand by
C. H. B. Grant, measure — head and body 333 and 335 mrn.,
tails 207 and 209 mm. respectively. No. 671 of macrurus
measures — head and body 380, tail 270 mm. There is
evidently, therefore, so material a difference in size and in
length of tail between the Iluvvenzori form and the South-
African that subspecific distinction is necessary, in spite of
the close resemblance in other characters.
Sciurus rufohracJiiatus semlikiiy subsp. n.
Most closely allied to 8. r. nyansce, Neum.*, with which it
agrees in the extent of the rufous on the limbs, but distin-
guished by the speckling of the back being very much finer
and by its greyer general colour. In nyansce there is a strong
suffusion of buffy or fulvous in the dorsal colour, the rings of
the longer hairs and the ends of the wool-hairs being markedly
tinged with buffy ; in semlilciij on the other hand, the former
are almost white and the latter are dull greyish buffy. As a
result the general tone of semlikii is a nearly pure dark grey,
between grey no. 6 and “smoke-grey"’^ of liidgway, the
middle line of the back only with slight yellowish suffusion.
Dimensions of the type (measured in the flesh) : —
Head and body 225 mm. ; tail 251 ; hind foot 49; ear 16.
* SB. Ges. mit. Fr. Berl. 1002, p. 56.
121
Mammals from Ihiwenzori.
llah. Beni, Scmliki Biver. Alt. 3000'.
Tt/pe. Adult male. B.M. uo. G. 12. 4. 64. Original
number 153. Colleclecl 22nd July, 1906, by E. E. Dent.
Jdiis handsome squirrel is no doubt nearly allied to
8. r. iiyanso?^ but in the very hue speckling of the body and
its greyer colour shows a relationship to the form found in
the Gaboon, to which the name of 8. r. Auhryi, M .-Edw.'*, is
applicable. The latter, however, has not the richly rufous
feet characteristic of both the Central- African subspecies.
It may be noted that the Euwenzori squirrel described by
Schwann as a subspecies of 8, rvfohrachiatus t proves, on the
arrival of a good series, to be a quite distinct species, charac-
terized not only by its greenish olivaceous colours, the absence
of red on the limbs, and its white ventral line, but by its
possession of two upper premolars, 8. rufohrachiatus having
only one. It should therefore stand as L. rincenzorii.
Tth\MNOMYS, gen. nov. [Mnrince).
Type X TJi. venustus, sp. n.
In pointing out recently the advisability of generically
distinguishing Micromys from § on account of its posses-
sion of the or postero-iriternal cusp on its first and
second upper molars, I mentioned that two African species,
hitherto assigned to Mus — M. arhorarius and rutilans —
possessed the same character ; and I now take this occasion
to separate them and their allies also from the parent genus,
which greatly needs reducing. From Micromys^ which is
entirely Palcearctic, they may be distinguished by being-
free- or bush-, instead of terrestrial miee, and by having
therefore the usual pencilled tail found in such forms.
The genus Thamnomys divides into two sections — the one
consisting of Peterses Mus rutilans and the new Euwenzori
species Th. venustus^ which I propose to take as type of
the genus; and (2) the less strongly marked group containing
Mus dolichurus^ Smuts, il/. arhorarius ^ Peters, and their
allies.
* Sciurus Auhryi, M.-Edw. Rev. Zool. xix. 228 (1867). Stated by
Jentink and Trouessart to be based on a young specimen of the S. rufo-
brachiatus group. Collected by xYubry Le Conte in the Gaboon.
t Ann. & Mag. Nat. Hist. (7) xiii. p. 71 (1904).
t Th. venustus is selected as the type in order to avoid any possible
complication which might arise should our examples of 3Ius rutilans,
Peters, prove to be wrongly determined. This course is the more advis-
able as Peters says of rutilans, “ Backzahne von gewohnlichen Propor-
tionen,” as though there were nothing special about the dentition.
§ Ann. & Mag. Nat. Hist. (7) xv. p. 492 (1905). See also op. cit. (7)
xvii. p. 85 (1906).
122
Mr. O. Thomas on new
While tlie first of tliese groups unquestionably deserves
distinction from Mus, the second is more or less intermediate
in its tooth-characters; but since it shows a distinct tendency
to the true Tharnnomys type of teeth, and has absolutely the
same cxterrial characteristics, 1 think it advisable to refer it
to the new genus rather than to leave it in Mas,
In the typical venustus-rutilans group all the supplementary
cusps of the teeth are highly develoj)ed, and the x cusp in
particular is a distinct separate cusp almost equalling in size
the other internal cusps in front of it, and there are con-
sequently nine well-marked cusps along the inner side of the
upper tooth-row.
On the other hand, in the dolicharus group the x cusp is
but little developed, and in fact is usually represented by a
mere ridge running backwards from the hinder edge of the
second internal cusp of and nt^ and joining the inner side
of the median cusp of the hinder lamina of each tooth.
In the lower jaw the postero-external supplementary cusp
of mi is decidedly larger than in Mas, nearly as large in area
and at least half as high as the main posterior cusp against
which it lies.
Externally, as already noted, the mice of the genus
Tharnnomys are characterized by their terminally pencilled
tails, in the same way as the analogous forms Rh’pidomys
and (Ecomys in S. America. The feet are similarly modified
for climbing, with broad rounded sole-pads, and the fifth
hind toe readies nearly to the end of the second phalanx of
the fourth, and is therefore nearly as long as the second.
Tharnnomys venustus, sp. n.
Allied to Th. rutilans, Peters, but larger and with much
longer fur.
h'ur of back about 15 mm. in length. General colour
above dark rufous (in spirit), a brighter line along the sides ;
belly white, with a faint huffy suffusion, the basal halves of
the hairs slaty. Ears dull greyish. Upper surface of feet
buff, becoming whiter on the toes. Tail long, pencilled ter-
minally, uniformly dark brown, as in Th. rutilans. Mamnue
0—2 = 4.
Skull decidedly larger than in Th. rutilans ; muzzle long,
parallel-sided ; interorbital region narrow, the edges not so
widely expanded as in rutilans ; palatine foramina long, not
expanded mesially, reaching back to the level of the front of
ni^ ; bullae rather larger than in rutilans. Molars large,
heavy, the series markedly larger than in rutilans.
Dimensions of the type (measured on the spirit-specimen) : —
Head and body 125 mm.; tail 181 ; hind foot 25 ; ear 18.
123
Mammals from Buwenzori.
Skull : gToatcst kuigtli 34*5 ; basilar length 28 ; greatest
breadth 17'2; nasals 12‘5 ; interorbital bia adth 4'9 ; breadth
of bi.ain-ease 14*3; ))ahitilar length 15; diastema 9‘4 ;
palatal foramina 8‘2 ; length of upper molar series G*l.
JIah. lluwenzori East. Alt. 7000'.
Type. Adult female in alcohol. lEM. no. 6. 12. 4. lOG.
Oiig'inal number G15. Collected by K. B. Woosnam.
Unfortunately only one specimen of this fine species was
obtained by the expedition, and that was preserved in spirit.
However, its longer skull, larger teeth, longer fur, and the
slaty mixture of its belly-colour will readily distinguish it
from Th. rutilanSj to which alone it is nearly allied.
Thamnomys dryas, sp. n.
A member of the Th, dolichuriis group, but with the
mammae only 0 — 2 = 4.
Quite similar to the more rufous forms of dolichiirus,
e. g. those from Nyasa and East Africa, the general colour
tawny, greyer on the liead, licher on the rump, sides paler, a
vell-rnarked huffy or ochraceous line edging the belly, wliich
is wdiite, sometimes tinged with huffy, the hairs not slaty at
base. Ears with proectote tawny brown or blackish ; meten-
tote rich buffy ; a pale huffy spot behind their posterior base.
Upper surface of hands and feet pale huffy. Tail long,
pencilled, uniformly dark brown. Mammae 0 — 2 = 4.
8kull lightly built. Palatal foramina reaching just to the
level of the front of vd. Bulla3 small. Molars light and
delicate.
Dimensions of the type (measured in the flesh) : —
Head and body 118 min.; tail 185; hind foot 24’5 ;
ear 19.
Skull: greatest length 31*3; basilar length 24*5; zygo-
matic breadth 15*3 ; nasals 11*5 ; interorbital breadth 4*7 ;
breadth of brain-case 13*6 ; palatal ioi amina 7*2 ; length of
u])per molar series 4*3.
IJab. Ruwenzori East. Alt. GOOo-7000'.
Type. Adult female. B.M. no. G. 7. 1. 136. Original
number 207. Collected 30lh December, 1905, by D. Car-
ruthers. Nine specimens, of which six are females, including
one in spiilt.
The presence of only four mammas in this mouse is a very
curious fact, as there are six in all the other members of the
27i. dolichurus group, even those from compaiatively near
localities. Specimens from Shoa (^Zophirof Mt. Elgon
{Jackson), Nyasa {Johnston), Angola {Ansorgef and South
Africa ail show this latter number.
124
On Ocliotona (Conotlioa) aurlta.
XJl. — Note on Ocliotona (Conotlioa) aurlta, Blanf,^
from Ladah, By J. Lewis Boniiote, M.A.
Since writing rny paper on the genus Ocliotona (P. Z. S.
1904, ii. pp. 205 et seqq.) I have through the kindness of
Col. A. E. Ward reeeived some specimens of an Ocliotona
from the Pangong Lake, the typical locality of BlanfoiHPs
O, aurita^ which, from reference to his descri))tion and plate,
1 have no hesitation in referring to that species.
In my paper quoted above 1 considered, in default of
S])ecimcns, that Blanfordbs 0. aurita was probably ideiitieal
with Gunther’s O. macrotis) this, however, is not the case,
though in view of the stress laid by the describer on the
large size of the ears, as also evinced by the name, the
mistake was perhaps natural and pardonable. The following
is a deseription of Col. Ward’s speeimens from the l^angong
Lake, Ladak, and Nubia Valley : —
General appearance very similar to Blanford’s plate.
Colour above buffish grey grizzled with black, each hair
being black at its base with a buffish subterminal portion
and a black tip, which tips are frequently worn otf to a
greater or less extent. The feet are white with a tendency
to yellowish in their median portion. The underparts are
])ure white with an indistinct narrow median line of yellowisli.
Between the ears, across the shoulders, and reaching down
the sides of the neck is a broad band of pure buff, divided in
two in the middle line by a few of the hairs having black
tips. The whole of the head is rather brighter and yellower
than the rest of the body, but there is no trace of the rufous
suffusion of this part which is found in both 0. Iloylei and
0. inacrotis. The skull is typical of the Carzonioe ( = subgen.
Conotlioa) group. But in some specimens the anterior
portion of the large palatine foramen does not immediately
broaden out from its anterior end, but the two sides run
parallel for a short distance ; they, however, never show any
tendency to approach each other and can therefore not be
confounded with the rnfescens ( = subgen. Ocliotona) group.
In general features the skull resembles most closely that
of 0. Curzonice^ from which there is little to distinguish it,
except that the postorbital process of the zygoma is not so
long and narrow ; in size it is about the same. Erom
0, Iloylei the skull may be distinguished, apart from iis
rather smaller size, by the fact that the palatal foramen is
more triangular in general outline, the posterior portion
tending to open out more on either side.
125
On Two new rarasitic Coleoptera,
The skull of 0. macrotis is larger and the foramen resembles
that of 0. Roylei ; it basin addition two oval foramina in the
frontal bones which are not found in 0. aurita.
Dimensions of 0. aurita from Pangong Lake (Coll. A. E.
Ward^ no. 48) —
Head and body 170 mm. ; hind foot 31 ; ear 21.
Skull: greatest leiii^th 40; basal length 34; zygomatic
breadth 18; length of nasals 13; length of molar series 7'5.
There can be no doubt, from a comparison of the skins,
that the present species is closely allied to 0. Gurzonice. It
is apparently merely rather darker and the light buff patches
behind the ears are deeper in colour as a rule than in typical
0. Gurzonice^ though this seems to be a rather variable
feature.
Good series of both 0, Gurzonice and 0. aurita will
probably prove the latter to be a geographical form dis-
tinguished by some minor but constant differences. The
name aurita is unfortunate, as the ears are by no means
conspicuous or large. 0. macrotis^ except for its large ears,
is not unlike a pale greyish 0. Roylei, a resemblance which
is to some extent borne out by the skulls.
The following are the references to 0. aurita : —
Lagomys auritus, Blanford, J. A, S. B. voL xliv. p. Ill (Oct. 1875) ;
id, loc. cit. xlvi. p, 326 (1877) ; id. Yark. Mainin. p. 74, pi. vi. %. 2,
pi. vii. a, dg. 2 (1879).
XIII. — On Two new Parasitic
li nidge) from South America.
F.E.S.
Goleoptera {Fain. Staphy-
By Gilbert J. Arrow,
Five species have hitherto been described of the curious
parasitic genus of which four have been brought
from the mountain-chain of Western South America and the
fifth from the yjlains of La Plata. I am now adding two
more species, one from the north and the other from the
extreme south of the continent, and each represented by a
single specimen in the British Museum collection. The
circumstances in which they were found have not been re-
corded, but it may be assumed that, like the other species,
they are parasites of burrowing rodents.
Amblyopinus angustus^ sp. n.
Pallide testaceiis, vix nitidus, elongatus, scutello, elytris abdo-
mineque flavo-pubescentibus ; capite satfortiter punctato, postice
12() On Two new Parasitic Coleoptera.
rco'iilariter dilatai-o, liaud angulaho, oculis miiiutis, baud pro-
minciitibus ; protboraco rugose punctato, latcribus aniico arcuatis,
ap])roxirnal,is, postieofcre rectis,aiigulis posticis baud late arcuatis,
niargine basali distinctc sinuata ; scutello rnagno, lato, crebre
])UiJct;ito, apice vix angulato ; clytris punctato-rugosis ; abdomine
6u])tilitcT rugose punctato, segrnentis postice longitudine valdo
crcscentibus, ultimo quam latitudincm longiore; anteunis pedibus-
(}iie sat brevil)us.
Long. 8 mm.
Jlah. British Guiatia, Mt. Roralina. Found by Mr. J.
J. Quelcli.
Tills species has a very distinctive aspect, owing to its
elongate form. It is small and of a uniformly pale colour,
and has the liead and thorax closely punctured and not at all
shining, the elytra rugose and closely pubescent, and the
abdomen finely pubescent, but more shining than the rest of
the body. Tliere are two minute black spots at the posterior
])art of tlie liead, symmetrically placed, but perhaps not con-
stant. The first three joints of the antennas are elongate and
the remainder rather short and equal. The head is dilated
behind, but not very strongly, the protliorax is distinctly
narrowed in fiont, and the scutellum is very large, occupying
at the base fully a third of the breadth of the insect from
shoulder to shoulder. The elytra, beneath which there are no
wings, are not elongated in proportion to the rest of the body.
The legs and antenuEe are shorter than in most of the species,
but hardly as short as in A. Gahani^ Fauvel.
Amhlyopinus fuegensisy sp. n.
Rohustus, sat latus, rufo-ferrugiueus, capite prothoraceque nitidis,
glabris, elytris, scutello abdomiueque deuse fulvo-pubescentibus ;
capite crebre sed distiucte puuctato, post oculos leviter dilatato,
his parvis, paulo promiueutibus ; prothorace subtiliter punctulato,
la'eribus arcuatis, augulis posticis obliteratis, margiue basali
leviter siuuata, augulis anticis rotuudatis ; scutello sat parvo,
triaugulare, apice distiucte augnlato ; elytris brevibus, sicut
abdomine hand distiucte punctatis, hujus segmento ultimo sat
lato, apice vix emarginato.
Long. 8 mm.
Hah, Tierra del Fuego, Useless Bay. Found by Capl. R.
Crawshay, in December 1904.
This is like A. Jelskii, Solsky, but rather larger. The
head is relatively larger, the prothorax more circular, with
the sides more curved, the posterior angles less apparent, and
the surface less punctured and nmre shining. The scutellum
127
Bihliogra'pli ical Notices,
is longer and more angnlated and tlie elytra and abdomen
less distinctly sculptured, ddie antcniue are rather slender,
^vith the first joint almost twice as long as the second and
the three last as broad as they are long, the terminal one
having a short iinger-like process at the end.
EIBLIOGllAPIIICAL NOTICES.
Catalogue of the Indian Decapod Crustacea in the Collection of
the Indian Museum. —Vnci 111. Macrura. Fasciculus 1. The
Prawns of the Peneus Group. By A. Alcock, M.B., LL.D.,
C.I.E., E.ll.S. Calcutta : Printed by Order of the Trustees of
the Indian Museum. 1900. Price 7 rupees.
This is the Third, but an independent, Part of a Monograph of the
Decapod Crustacea of that portion of the Oriental llegion which
lies wdthin the political boundaries of British India ; and it deals
only with the prawns of the maniple Peneus. Nevertheless it
should prove of real service to all interested in the study of the
Crustacea, for there are few who can draw upon so wdde a know-
ledge of this group as Dr, Alcock. This part, like those which have
preceded it, is profusely illustrated by means of a large series of
plates, and this should add much to the value of the work.
Dr. Alcock points out that the Penei swarm in the warm waters
of the Indian seas, and form a large part of the food of many fishes.
But, “ beyond this,” he remarks, “ they in themselves constitute a
not inconsiderable part of that plenteous harvest of the sea which
in this country [India] still runs to waste for want of capital and
enterprise. What the prawn-fisheries of India might be wmrth it
is difficult to say ; but a statement published by Kishinouye, in the
‘ Journal of the Fisheries Bureau of Tokyo ’ for the year 1900, that
the dried prawns annually exported from Japan to China are valued
at 200,000 yen (or a little over <£20,000), shows that there must be
possibilities in them.”
A Treatise on Zoology. Edited by E. Bat Laxkester, M.A., LL.D.,
E.ll.S. — Part V, Mollusca. By Paul Pelseneek, D.Sc. London:
Adam A; Charles Black, 1906.
When we say that Dr. Pelseneer’s volume on the Mollusca is in
every way worthy of the earlier volumes of this great Treatise, we
have bestowed high praise ; and in doing so we entertain no fears
but that'this expression of opinion will meet with the approval of
all those who will have occasion to consult its pages.
Written originall)’ in another tongue, it has been translated, and
128
Miscellaneous.
ably translated, by Dr. Gilbert TlDurne, the Linacre Professor of
Comparative Anatomy at Oxford.
Designed for the advanced student, this volume, like its pre-
decessors, is of a highly technical character, and bristles with
terminology as yet unfamiliar even to those for whom it is in-
tended. 8o far from this being a drawback, however, it is, on the
contrary, a valuable feature — making for clearness, exactitude, and
condensation.
Not since the appearance of Ray Lankester’s article “ Mollusca,”
in the ninth edition of the ‘ Eucyclopjedia Britannica,’ has any work
on this group appeared comparable to the present volume ; hence
there can be no question about the welcome that this latest contri-
bution will receive. The ‘Encyclopaedia’ article marked an epoch
in the study of the Mollusca, and for many years, indeed, remained
the only effectual work of reference on the subject : no more
striking testimony as to its solid worth can be found than the fact
that all the more important conclusions therein arrived at find a
place in the volume now before us.
In one or two minor matters we find cause for complaint. The
chief of these concerns the unnecessarily vague statements as to
the number of living species of Mollusca. Thus on p. 35 we read ;
“ Descriptive zoologists have enumerated more than 28,000 species
of living Mollusca, of which more than half are Gastropods ” ; while
on p. 142 we meet with the statement that “ Some thirty thousand
species of Gastropoda have been enumerated, of which twenty
thousand belong to the present epoch.” Comment of this kind may
savour of “ quibbling ” : we do not mean to be hypercritical, but
desire simply to draw attention to a small point which might be
altered in a future edition.
MISCELLANEOUS.
Note of Correction.
Parorchis^ n. nom., for Zeucjorchis, Nicoll, ] 90G. By Wm. Nicoll,
M.A., B.Sc., Gatty Marine Laboratory, St. Atidrews.
The name Zeugorehis., proposed (Ann. & Mag. Nat. Hist. (7) xvii.
p. 519) for a Trematode genus represented by Z. acanthus, mihi,
from the cloaca of Larus argentatus, having previously been assigned
by Stafford (Zool. Anz. xxviii. (1905) p. 691) to a genus parasitic
in Reptilia, it is necessary to alter this. In its place I propose the
name Parorchis.
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. no. FEBRUARY 1907.
XIV. — On the Bats of the Family Megadermatidge.
By Knud Andersen and R. C. Wroughton.
The following notes are based on an examination of the
material in the British Museum of Natural History.
Key to the Genera,
A. Frontal shield of skull wider in front than be-
hind. Cusp 2 of m}^ much reduced in size or
almost entirr'iy disappeared. Antero-internal
basal cusp of upper canines always distinct.
a. t present. Prenasal notch of skull rounded
posteriorly. (Oriental and Malayan.)
a\ Prenasal notch of skull shallower, extending
I 9nly to level of front of m^. Tragus longer,
/more slender. General size smaller I. Megaderma.
* jTh4 three cusps forming the tips of the W of a typical molar in
inseejiti’^ ^rous bats are termed, in antero-posterior direction, respectively
1, 2,, 3 ; the two cusps forming the base of the W, 4 and 5 ; the lingual
“ heiel” ' of the upper molars, when single, 6, when double, G and 7 ; see
Herlu ? Winge, “ Om Pattedyrenes Tandskifte, isser med Hensyn til
Teen delrnes Former,"’ in Vidensk. Meddel. Naturhist. Foren. Kbhvn. 1882,
pp. 15 -69, pi. hi. (We prefer Winge’s designations to those proposed
by Os^born, which, in our opinion, are based on an erroneous idea of the
succejjsion of the cusps, and give a mistaken interpretation of the cusps
of the' lower as compared with those of the upper molars.)
t T'he small anterior upper premolar.
Ann,, & Mag. N. Hist. Ser. 7. Vol, xix. 9
130 Messrs. K. Andersen and E. C. Wroughton on
b'. Prenasal notch of shull deeper, extending to
level of front of m^. Tragus shorter, broader.
General size larger II. Eucheira.
b. absent. Prenasal notch of skull pointed
posteriorly. Tragus very short and broad.
General size very large. (Australian.) III. Macroderma.
B. Frontal shield of skull not wider in front than be-
hind. Cusp 2 of nd not or scarcely reduced in
size, p^ absent. (Ethiopian.)
a. Posterior pair of angles of frontal shield pro-
duced to form postorbital processes ; upper
surface of shield comparatively flat, m'- quite
normal. Antero-internal basal cusp of upper
canines present. Nose-leaf very large. Tra-
gus long, narrow. Pollex small IV. Lavia.
b. Posterior pair of angles of frontal shield not
produced to form postorbital processes ; upper
surface of shield strongly concave. Cusp 3 of
moved backward. Antero-internal basal
cusp of upper canines absent. Nose-leaf
small. Tragus short, broad. Pollex larger. . V. Cardioderma.
I. Megaderma, Geoff.
1810. Megaderma, Geoffroy, Ann. Mus. d’Hist. Nat. xv. p. 187.
Diagnosis. — See Key above, p. 129.
Skull. — Frontal shield wider in front than behind, its
anterior pair of angles situated at the bases of the maxillary
processes of the zygomatic arches ; the ridges joining them
with the posterior pair of angles forming quite normal supra-
orbital ridges ; looked at from above, the anterior pair of
angles are obscured by anteorbital swellings, which, separated
by a distinct depression, occupy the whole anterior part of the
frontal shield. Prenasal notcli shallower, measured from the
cingulum of canine equal to about ^ of total length of skull.
Teeth. — A small present. Cusp 3 of is moved back-
ward, while cusp 2 is moved inward and reduced in height.
In the typical molar of an insectivorous bat the anterior and
posterior triangles are practically of the same size; ih Mega-
derma the posterior triangle (formed by cusps 2, 3*, 5) is
distinctly larger than the anterior (formed by cusps 1^, 2^, 4),
though not to 'the same extent as in Eucheira. Antero-
internal basal cusp of upper canines quite distinct, but no't so
strongly developed as in Eucheira. , ^
Nose-leaf. — Posterior leaf in shape a broad ellipse, :|rather
longer than the horseshoe, with a median longitudina I |fold
which, seen from the front, appears as a longitudinal ridg^^> ; at
its base this ridge expands into a broad, heart-shaped, imedian
leaf, which overlaps and conceals the lateral margins c-‘f the
horseshoe.
i:u
the Bats of the Family Megadermatidae,
Tragus. — Both lobes propoitionally long and narrow, as
compared with those of Eucheira.
Range. — From the Indian Peninsula and Ceylon eastward
as far as the Philippines, Celebes, and Ternate.
Species. — Three, viz. : M, spasmaj M, carimatoe, M. na~
tunce.
Nomenclature. — The name Megaderma was proposed by
Geoffrey in 1810 {1. s. c.). The author describes the four
species included by him in Megaderma in tlie following
order: — M. lyra^ M. frons., M. tr {folium ^ M. spasma. But
M. spasma is mentioned in the description of the genus in
advance of all other species je me suis cm fond^a considerer
le V. spasma et ses congenbres comme formant un groupe
isole/^ p. 190), and in a short summary (p. 197) which
professes to give a view of the species in their proper order
(‘‘dans Fordre de leurs rapports ^^) Geoffroy heads the list
with M. trifolium^ i. e. the western race of M. spasma* In
accordance with the generally accepted view we therefore
regard M. spasma as the type of the genus.
Key to the Forms*
A. Size smaller : skull 25-27 mm.
a. Ears shorter: from crown 27'0-30’0 mm. . . 1. M. spasma.
a' . Lower leg averaging shorter : 28-29-5 mm. la. M. s. spasma.
b'. Lower leg averaging longer : 29'o-33'5 mm. 15. M. s, trifolium.
b. Ears longer : from crown 31'4-35 mm 2. M. carimatce.
B. Size larger : skull 28'5 mm 3. M. natunce.
1. Megaderma spasma^ L.
Diagnosis. — Size in every respect smaller than in M. na~
tunas ; ears shorter than in M. carimatce.
Details. — Distinguishable at a glance from M. natunce by
the conspicuously smaller size, especially of the skull and
teeth. Total length of skull to front of canines 25-27 mm.,
in M. natunce 28'5 mm. ; length of upper tooth-row [c-m^)
9‘5-10*5 mm., in M. natunce 11*4 mm. ; forearm 53-62 mm.,
in M* natunce 63 mm.* From M. carimatce it differs only
by its somewhat shorter ears.
Range. — The same as that of the genus.
Races. — Two, viz. : M. spasma spasma^ M. spasma trifolium,
* We give in the letterpress a few principal dimensions only; a
detailed comparative table of measurements will be found at the end of
the paper (p. 144).
V
132 Messrs. K. Andersen and R. C. Wroughton on
1 a. Megaderma spasma spasma, L.
1734. Glis volans Ternatunus, Seba, Thesaur. p. 90, pi. Ivi. fig. 1.
1758. Vespertilio spasma, Linnaeus, Syst. Nat. ed. x. p. 32.
1810. Meyaderma spasma, L. ; Geoffroy, Ann. Mus. d’llist. Nat. xv.
p. 195.
1843. Meyaderma philijppinensis, Waterhouse, P. Z. S. p. 09.
Diagnosis. — Lower leg averaging shorter; 28-29*5 mm.
Specimens examined. — 6 (2 skins) and 6 skulls, from
Qelebes (3), Philippines (3).
Range. — Ternate,^^ Celebes, Philippines.
Nomenclature. — Linn^’s Vespertilio spasma was based on
Seba^s ^^Glis volans Ternatanus.” Having had no speci-
mens from Ternate for examination, we use the name spasma
for the race here under consideration, the habitat of which is
the nearest to Ternate. Waterhouse described AI. pfiilippin^
ensis from specimens collected by Cuming, some of which are
now in the British Museum Collection. They are in every
respect indistinguishable from Celebes individuals.
1 h, Alegaderma spasma trifolium^ GeofF.
1810. Meyaderma trifolium, Geoffroy, Ann. Mus. d’Hist. Nat. xv.
p. 193.
1851. Meyaderma , Horsfield, Cat. Mamm. Mus. E. I. Co. p. 32.
1863. Meyaderma Horsfieldi, Blyth, Cat. Mamm. Mus. As. Soc. Beng.
p. 23.
Diagnosis. — Lower leg averaging longer : 29*5-33*5 mm.
Details. — AI. s. trifolium can only be distinguished from
AI. s. spasma by its rather longer lower leg. The skull, teeth,
and external characters are otherwise the same in the two
races.
Specimens examined. — 29 (16 skins) and 29 skulls, from
Ceylon (3), W. India (4), Siam and Cochin China (8),
Penang (2), Singapore (3), Sumatra (2), Java (6), Borneo (1).
Geoffroy’s M. trifolium. — Type locality : Java. Geoffroy
compared his Javan bat with Seba’s plate of Glis volans
Ternatanus (Linnets V. spasma), and found it to differ in the
shapes of the posterior leaf and the tragus; he therefore
described it as a new species, Al. trifolium. . AVe have
examined specimens from Java and can find no validity in
these alleged characters. The name trifolium is the earliest
available for the form here under consideration.
Blythes AI. Horsfieldi. — In 1851 (/. s. c.) Horsfield described
a Aiegaderma “ from Continental India, contributed by
E. Blyth, Esq., on behalf of the Asiatic Society of BengaL";
no name was proposed by Horsfield ; it was characterized by
133
the Bats of the Family Megadermatidas.
tlie absence of transverse lines ” from the membrane along
the sides of the abdomen, and the presence of regularly
parallel ridges at the base of the ear/'’ The British Museum
possesses a specimen (skin, no. 60. 5. 4. 13), received from
the India House Collection, which is iji all probability the
actual individual referred to by Horsfield ; the peculiarities
emphasized by him are partly due to bad preparation of the
specimen and partly are quite individual. In 1863 {1. s. c.)
Blyth proposed the name Horsjieldi for the species described
but left unnamed by Horsfield. 'When the late Dr. J.
Anderson (Cat. Mamm. Ind. Mus. p. 21, 1881) registered as
“ types ” of M. Horsjieldi two specimens obtained by
Mr. Theobald in Tenasserim he was undoubtedly in error.
Blytl/s M. Horsjieldi was, as already pointed out, based on
Horsfield'’s Megaderma ” from Continental India.
2. Megaderma carimatoBj Miller.
1906. Megaderma carimatce, Gerrit S. Miller, Jun., Proc. U.S. Nat.
Mus. xxxi. no. 1481, p, 65.
The species is known to us only from the published account.
Type locality: Karimata Island. According to Miller, it
differs from M. spasma only in the smaller size and larger
ears. The former character must be dropped ; Miller gives
as length of the forearm (five males and four females) 53*6-
58'6 mm., as against 57-61 mm. in Al, spasma (seven females,
Malay Peninsula, i. e, M. s. trijolium of the present paper) :
but, first. Miller has compared his il/. carimatce with females
only of Af, spasma, and females of this species average larger
than males ; second, we find in twenty-nine individuals of
Al. s. trijolium the forearm varying between 53 and 62 mm.
There remains the alleged greater length of the ears in
AI. carimatce : measured from the crown “ 31*4-35 mm.’^
(Aliller) against 27-29 mm.'’"’ {Alillei') in AL spasma ; we
find that similar measurements for M, s, trijolium range from
27-30 mm. (Malay specimens 28-29 mm.).
3. Alegaderma natunce^ sp. n.
Diagnosis. — In every respect larger than AI, spasma.
Details. — The differences between AI. natunce and AI. spasma
have been pointed out above under the latter species (p. 131).
Type. — ^ ad. (in ale.). Bungaran Island, N. Natunas.
Collected by A. Everett, Esq. British Museum no.
94. 9. 28. 30.
Range. — As yet known from the type specimen only.
134 Messrs. K. Andersen and R. C. Wrougliton on
II. Eucheira, Hodgs.
1847. Eucheira, Hodgson, J. A. S. B, xvi. p. 891, footnote (September
1847).
1872. Lyroderma, Peters, MR. Akad. Berl. p. 195 (18th March, 1872).
Diagnosis. — See Kev/^ above, p. 130.
Skull. — Frontal sliield as in Megaderma. Prenasal notch
deeper, measured from the cingulum of canine about ^ of
total length of skull.
Teeth. — A small present. Cusp 3 of moved con-
siderably backward ; the posterior triangle (formed by cusps
2, 3, 5) at least double the size of the anterior (formed by
cusps 2, 4). Cusp 2 moved inward and tending to dis-
appear, being represented only by a small tubercle, much
below the level of the other cusps. Antero-internal basal
cusp of upper canine very strongly developed, more so than
in Megaderma.
JSose-leof. — Posterior leaf a rectangle, with slightly convex
sides, twice as long as the median leaf; median longitudinal
ridge (fold) as in Megaderma, but its junction with median
leaf forming an obtuse angle on each side, strongly con-
trasting with the sharply acute angles formed in Megaderma.
Range. — Indian Peninsula; S. China; there seems as yet
to be no record of this genus from Burma.
Species. — Two, viz. : E. lyra, E. sinensis.
Hodgson^s Eucheira. — Type species, E. schistacea, Hodgs.,
= Megaderma lyra, Geoff. The generic name Eucheira was
proposed by Hodgson because the phalangeal system [of
E. schistacea] is apparently irreconcilable with Cuviei-’s
general and Geoffroy^s particular definitions^^ of the common
structure of the wing in Chiroptera; but at the same time the
description and figure of the wing of schistacea, as given by
Hodgson himself, are those of a quite normal Megadermine
wing, and liis schistacea is undoubtedly nothing but the well-
known “ Megadermad^ lyra. Although, therefore, the whole
basis on which Hodgson founded the genus Eucheira is a
mistake, still the name, as being the earliest in date, will
have to stand according to the current nomenclatural rules.
The name Eucheira has apparently been overlooked by suc-
ceeding writers and is not mentioned in Palmei-’s ^ index
Geneium Mammalium.^
Feters^ Lyroderma. — Type species. Megaderma lyra, Geoff.
The characters given by Peters for the “ subgenus Lyro-
derma are the shape of the nose-leaf, the flatness of the
Irontal shield, and the absence of postorbital processes. The
name Lyroderma is antedated by Hodgson^s Eucheira.
the Bats of the Famity Megaderm atidoo.
135
Key to the Forms.
A. Skull smaller: 27’8-29 mm. ; preriasal notch pos-
teriorly wider, flattened 1. E. lyra.
a. On the average larger : forearm 65-69 mm \a, E. 1. lyra.
h. On the average smaller : forearm 63-68 mm. . . \h. E. 1. caurina,
B. Skull larger : 29'3-32 mm. ; prenasal notch
narrower, rounded posteriorly 2. E. sinensis.
1. Eucheira lyra, GeofF.
Chief characters. — Skull and teeth markedly smaller than
in E. sinensis. Total length of skull 27*8-29 mm., in
E. sinensis 29*3-32 ram. ; upper tooth-row 10*8-11*5 mm.,
in E. sinensis 11*5-12*1 mm. Prenasal notch proportionally
rather shorter, wider behind, posterior margin flattened.
Range. — Indian Peninsula.
Races. — Two : E. lyra lyra and E. lyra caurina,
1 a. Eucheira lyra lyra, GeofF.
1810. Megaderma lyra, Geoffroy, xAnn. Mus. d’Hist. Nat. xv. p. 190.
1839. Vespertilio (^Megaderma) carnatica, Elliot, Madr. Journ. vol. x.
p. 96.
1814. Megaderma spectrum, Wagner, in Hiigel’s Kaschmir, iv. p. 569.
1847. E. sckistacea, Hodgson, J. xA. S. B. xvi. p. 889.
Characters. — The present race can only be distinguished
from E. 1. caurina by average characters : skull 29-29*5 mm.,
upper tooth-row 11*1-11*5, forearm 65-69, against 27*8-28*3,
10*8-11, 63-64 respectively in E. 1. caurina.
Specimens examined. — 18 (16 skins) and 20 skulls, viz. :
^‘Madras^^ (11); Secunderabad (1), Bengal (6).
Range. — Indian Peninsula, east of 75° E.
Geojfrofs Megaderma lyra. — Type locality : one of the
Dutch factories in India, probably East Coast of Madras.
Geoftroy separated lyra as a distinct species on account of the
shape and size of the nose-leaf (“Feuille rectangulaire,
la follicule de moitie plus petite, 1. s. c.).
Elliotts Megaderma carnatica. — Type locality : Southern
Maratha Country. Elliot relied on the presence of only
three teeth in the upper row behind the canine to justify the
separation of carnatica, but he seems himself to have been in
doubt as to the validity of the species, inasmuch as he adlei
“ M. lyra ? ” The British Museum has several specimens
collected by Elliot, all of them unquestionably lyra.
Hodgson’s Eucheira schistacea. — Type locality: Siligori,
N.E. Bengal. Hodgson left Nepal for good in 1844, and
the context (/. s. c.) shows that he had never seen any species
136 Messrs. K. Andersen and R. C. Wrougliton on
of the family Megadermatidae until he obtained the examples
on which he based schistacea ; these specimens, when later on
acquired by the British Museum, were wrongly labelled
Nepal (see Scully, J. A. S. B. Ivi. pt. ii. no. 3, p. 234,
1887). Hoclgson^s specimens, as well as his published figure
of the bat, show that schistacea is nothing but E. lyra lyra.
Wagner^ s Megaderma spectrum. — Type locality : Kashmir.
Apparently based on a single specimen (Baron Hiigebs
collections). According to Wagner, M. spectrum has only
four teeth behind the upper canine (/. e. one premolar only) :
but bearing in mind that is present in all known Oriental
representatives of the family Megadermatidas (this small tooth
is wanting only in the Ethiopian genera Lavia and Cardio-
derma and the Australian Macroderma) ; farther, that
Wagner^s description of M. spectrum is based throughout on
a comparison with M. frons {Lavia frons\ not with E. lyra^
with which last he seems to have been unacquainted ; again,
that p^j when present in bats of this family, is extremely
small, hidden on the internal side of the tooth-row, and
therefore very easily overlooked when not searched for ; and,
finally, that Wagner^s figure of M. spectrum differs in no
appreciable respect from an ordinary E. lyra — there can be
no reasonable doubt that W agner^s statement as to the
number of teeth was wrong ; if so, the whole basis of the
supposed new species breaks down.
1 h, Eucheira lyra caurina^ subsp. n.
Characters. — See above, under E. 1. lyra (p. 135).
Specimens examined. — 8 skins and skulls, all from the
West Coast of India.
Type. — ? ad. skin. Surat District, W. India. Collected
by il. C. Wrougliton, Esq. B.M. no. 98. 4. 2. 2.
Range. — India, west of 77° E.
2. Eucheira sinensis ^ sp. n.
Characters, — Size somewhat larger than E. lyra ; see
comparative measurements under that species above (p. 135).
Prenasal notch proportionally rather longer, narrowed behind,
posterior margin rounded.
Specimens examined, — 2 skins and skulls, viz. Swatow (1),
Amoy (1).
Type, — Adult (unsexed), skin and skull. Amoy, S. China.
Tomes Collection. B.M. no. 7. 1. 1. 339.
Range. — S. China.
the Bats of the Family Megaclermatida3.
137
III. Macroderma, Miller.
1900. Macroderma, Gerrit S. Miller, Jun., Proc. Biol. Soc. Wash. xix.
p. 84 (4th June, 1906).
Diagnosis. — See “ Key/^ p. 130.
Skull. — Frontal shield wider in front tlian behind ; its
anterior angles raised into the same plane as the posterior (in
the two preceding genera the anterior angles are depressed to
the bases of the maxillary processes of the zygomatic arches) ;
anteorbital swellings obliterated (distinct in Megaderma and
Eucheira) ; as a consequence of these modifications the whole
area of the pentagonal frontal shield flat and all its angles
sharply and strongly defined. Prenasal notch still longer
than in Eucheira (measured from cingulum of canines about
i of total length of skull) ; posteriorly pointed, not rounded
or flattened as in Megaderma and Eucheira.
Teeth. — yS absent. Cusp 3 of moved backward as in
Eucheira j cusp 2 moved inward to a line between cusps 1
and 5, almost obliterated ; cusp 4 much reduced, markedly
below the level of the remaining cusps. Antero-internal
basal cusp of upper canines strongly developed.
Eose-leaf. — Posterior leaf ovoid as in M.egadermaj double
the length of the median leaf as in Eucheira ; median leaf
shaped as in Eucheira. Front margin of horseshoe plicate,
suggesting a rudimentary form of the much more complicated
structure in Lavia.
Tragus. — Both lobes short and broad, even more so than
in Eucheira.
Range. — Central Queensland.
Species. — One, viz. M. gigas.
Eomenclature. — The genus Macroderma was established by
Miller (Z. s. c.) iox Megaderma gigas and characterized by the
absence of the shape of the frontal shield, and the much
greater development of the cartilaginous premaxillaries.
Macroderma gigas, Dobs.
1880. Megadei'ma gigas, Dobson, P. Z. S. p. 461, pi. xlvi.
Diagnosis. — Forearm 103-104 mm.
Specim.ens examined. — 2 (1 skin) and 2 skulls, from Central
Queensland.
Range. — As yet known only from Central Queensland.
138 Messrs. K. Andersen and R. C. Wrougliton on
IV. La VIA, Gray.
1838. Lavut^ Gray, Mag. Zool. Bot. ii. p. 490.
.DiognosU. — See “ Key,^^ p. 130.
Skull. — Frontal shield parallel-sided, but appearing wider
behind, owing to development of the posterior pair of angles
into long ])rocesses; otherwise as in Prenasal
notch about the same proportional length as in Eucheira^
showing a tendency to become pointed behind in some indi-
viduals (compare Macrodermd).
Teeth. — j/ absent. All five cusps of m} equally developed
and placed in a regular W. Antero-internal basal cusp of
upper canine about as in Megaderma.
Nose-leaf. — Posterior leaf in shape a long triangle, with
sliglitly convex sides and truncated apex, three times as long
as the median leaf ; median longitudinal fold gradually
widening anteriorly, the junction with the median leaf
forming no appreciable angle on each side ; median leaf
small, with a lobe on each side anteriorly which reaches to
the margin of the horseshoe and covers the nostrils ; horse-
shoe with a notch on each side in front, between which the
front margin of the horseshoe is produced into a free lobe ;
the sides of this lobe folded together downward, and then the
whole turned backw^ard to lie on the face of the median leaf.
Tragus. — External lobe very long.
Range. — W. Coast of Africa, from Cape Verd to the
mouth of the Niger ; Upper Nile Valley; Uganda; British
and German East Africa.
Species. — One, viz. Lavia frons.
Nomenclature. — Type species of the genus, Megaderma
frons, Geoff. Gray proposed the generic name Lavia on
account of the shape of the nose-leaf and frontal shield and
the absence of ju'h
1. Lavia frons, Geoff.
Characters and Range. — Those of the genus.
Races. — Two, viz. Lavia frons frons and Lavia frons
affinis.
1 a. Lavia frons frons, Geoff.
1759. Za Feuille, Daubenton, M^m. Acad. Sci. Paris, p. 388.
1800. ? Vespertilio mecjalotis, Bechstein, in Pennant’s Uebersiebt der
vierfiiss. Thiere, p. 022.
1810. Megaderma frons, Geoffrey, Ann. Mus. d’llist. Nat. xv. p. 192.
1905. lAivia rex, Gerrit S. Miller, Jim., Proc. Biol. Soc. Wash, xviii.
p. 227 (9th Becember, 1905).
139
the Bats of the Family Megaderm atlda3.
Diagnosis, — Average size larger.
Details. — Can only be discriminated from L. f. affinis by
average characters : forearm 5G-62 mm., skull 21:‘5-26, upper
tooth-row 9-10, as against 52-58 rnm., 23'5-21, 8*7-9 re-
spectively in L.f. affinis.
Specimens examined. — 33 (21 skins) and 23 skulls, viz.
Gambia (0), Kumasi (1), N. Nigeria (2), Kordofan (4),
liuweiizori (4), Uganda (0), British East Africa (10).
Range. — Tlie same as that of the genus (above, p. 138),
with the exception of Bahr-el-Ghazal.
Bechsieiids V. megalotis. — Under the name V. megalotis
Bechstein describes a bat taken by Levaillant in Great
Namaqualand, which, from the absence of the tail and the
presence of nose-leaf and tragus, was clearly a member of
the family Megadermatid^e. But no species has in recent
times been recorded from Africa south of 15° S., and the
measurements quoted by Bechstein are so strange (rendered
from German inches into millimetres: nose-leaf 35 mm.;
body 78 mm.; ear 70 (!) mm.; expanse 210 mm.) that it
appears safer to leave Levaillanf’s bat unidentified.
Geoffroy’s Megaderma frons. — Type locality : Senegal.
Geoffrey’s descidption is based, not on actual specimens, but
on Daubenton'’s description of La Feuille"’'’ (/. s. c.), which
is undoubtedly the species here under consideration, being a
'bat with an ovate nose-leaf, ^^posee verticalement, qui
ressemble a une feuille,^^ huit lignes de longueur sur six de
largeur,^^ with the ears prbs de deux fois aussi grandes que
la membrane’^ [f. e. the nose-leaf], and united par la moitid
de la longueur de leur bord interne,^^ with a long, narrow,
pointed tragus, with no tail, with the fur “ d^une belle couleur
cendiee, avec quelque teinte de jaun^tre,"’^ with § incisors,
^ cheek-teeth, and inhabiting Senegal.
Milled s Lavia rex. — Type locality ; Taveta, German East
Africa. Miller relies for the discrimination of L. rex on its
greater external dimensions, longer mandible, and heavier teeth:
“ forearm 60 mm.,^'’ “ mandible 17*8,^^ maxillary tooth-
row 9*2,^^ instead of ^*56 mm.,'’^ ‘^15*2'’^ mm., and “8*2 mm."’'
respectively in L. frons. These measurements, as well as all
the others given by Miller, place it beyond doubt that his
L. rex is L. frons frons and that the reason for his describing
it as new was that he compared it, not with the true L. frons
f'ons, but with the smaller race described in this paper as
L. frons affinis.
140 Messrs. K. Andersen and R. C. W rougliton on
1 h. Lavia frons affiais, subsp. n.
Diagnosis. — Average size smaller.
Details. — Comparative measurements are given above
under L. frons frons (p. 139).
Type. — S at]* skin. Kaka, White Nile. Collected by
R. M. Hawker, Esq. B.M. no. 1. 8. 8. 3.
Specimens examined. — 8 (7 skins) and 6 skulls, viz. White
Nile (5), Lake No (1), Lado (1), Wadelai (1).
The British Museum possesses a somewhat damaged skull
without skin from Cape Coast Castle (Gold Coast) which
seems to belong to this race, and Miller^s specimen of
L. frons referred to above under L. f. frons (p. 139)
appears also to belong to the present race ; so it is possible
that the range of affinis extends westward to the coast.
V. Cardioderma, Peters.
1873. Cardioderma, Peters, MB. Akad. Berlin, p. 488 (23rd June,
1873).
Diagnosis. — See Key,^^ p. 130.
Skull, — Frontal shield parallel-sided; the centre longitu-
dinally depressed, so as to form a ‘‘ trough ; all the angles
sharply defined, but not produced into processes as in Lavia,
Prenasal notch about as long as in Megaderma total length
of skull), flatly rounded posteriorly.
Teeth, — p"^ absent. Cusp 3 of about as in Megaderma,
Antero-internal basal cusp of upper canine absent.
Nose-leaf. — As in Megaderma^ but a distinct trace of
antero-lateral lobes of the median leaf, as in Lavia ; in
Megaderma, Eucheira^ and Macroderma these lobes are merely
indicated by faint depressions in the margin of the median
leaf.
Tragus. — Both lobes veiy short ; the inner flatly rounded
at top and excavate at the base in front, so that it seems
directed inward rather than upward.
liange. — East Africa : Zanzibar, Mombasa, Somali.
Species. — One, C, cor.
Nomenclature. — Type species of the genus, Megaderma cor.
In 1872 Peters described Megaderma cor, and in the following
year proposed for this species the subgeneric name Gardio-
derma without any further characterization.
141
the Bats of the Family Megadermatidoe.
Cardioderma cor^ Ptrs.
1872. Megaderma coVy Peters, MB. Akad. Berl. (18th March, 1872)
p. 194.
Specimens examined. — 10 (3 skins) and 8 skulls, viz.
Zanzibar (2), Mombasa (2), Somali (6).
Peters^ Megaderma cor, — Type locality : Abyssinia. Peters
based his diagnosis of Megaderma cor on the shape and size
of the nose-leaf and tragus.
Wing-structure.
The four families Nycteridse, Megadermatidae, Hippo-
sideridae, and Khinolophid^ are rather closely inter-related ;
they have probably had a common origin. When trying to
form an idea of the stage of development at which the wing-
structure of the Megadermatidae has arrived, it is therefore
fair to compare them with such species of the other families
as have, in this respect, remained on a low level, f. i. Hippo-
siderus diadema.
In H. diadema the third metacarpal is the longest, the
fifth the shortest, the indices of the third, fourth, and fifth
metacarpals being, respectively, 716, 696, and 640; that this
is a primitive condition needs hardly any comment (a similar
mutual length of the metacarpals is found in many primitive
fruit-bats). In the Megadermatidse the third metacarpal is
the shortest, the fifth the longest, the indices being 727, 784,
and 845 ; the third metacarpal, it will be noticed, has retained
practically the same length (727) in proportion to the forearm
as in II. diadema (716), whereas the fourth and, still more,
the fifth have been very considerably lengthened.
In H. diadema the first phalanx of the third digit (index :
329) is somewhat less than half the length of the third
metacarpal (716) ; in the Megadermatidae it is decidedly
lengthened (index: 404), being always more than one half of
the metacarpal (727). The first phalanx of the fourth digit
has retained the same length in proportion to the forearm as
in H. diadema (242 against 237); the first phalanx of the
fifth digit is slightly lengthened (280 against 247).
In H. diadema the second phalanx of the third digit (327)
is only equal in length to the first phalanx (329) ; in the
Megaderniatida3 it is enormously lengthened (693), being
almost \ of the first phalanx (404). In H. diadema., t\\Q
distal phalanges of the fourth and fifth digits are considerably
shorter than the proximal phalanges; in the Megadermatidae
the second phalanx of the fourth digit is at least equal to
J42 Messrs. K. Andersen and R, C. AVrougliton on
[Macroderma^ Lavia), but often much longer than [Mega-
dermoy EucheWa, Cardioderma)^ the first phafanx ; the
second phalanx of the fifth digit is in all genera of Mega-
dermatidge approximately equal to, or rather longer than, the
U3
the Bats of the Family Megadermatldae.
first phalanx, except in Macroderma^ in which it has remained
rather short.
Tlie total result of these modifications is best realized by a
comparison of tlie total index of tlie third, fourth, and fifth
dio;its: in Megaderm atidae 1824, 1303, and 1404, as against
1372, 1106, and 1081 in H. diadema and allied species.
The greatest increase (452) falls on the third digit, the next
(323) on the fifth, the smallest (197) on the fourth; i. e. the
area of the wing in the family Megadermatidae is enormously
increased in size, the wing being at the same time much
more pointed (lengthening of third digit in proportion to
fourth) and much broader (lengthening of fifth digit in
proportion to fourth).
General Remarks.
The five genera of Megadermatidae are referable to two
fundamental types. In the one, represented by Megaderma^
Eucheira^ and Macroderma^ the frontal shield of the skull is
but moderately developed, the median external cusp (cusp 2)
of the upper m’- is more or less on the point of disappearance,
and, with the exception of the somewhat aberrant Macro-
derma, they have preserved the small anterior upper premolar
(p^). In the other group, represented by Lavia and Gardio-
derma, the frontal shield is largely developed, giving the
skull a quite peculiar aspect, the median external cusp of the
upper is of normal (or almost normal) size and has
completely disappeared. The former group is Oriental,
Malayan, and Australian, the latter Ethiopian.
Megaderma and Eucheira, from the Malay Archipelago and
S. Asia, are very closely related, differing in no other
important respects than the degree oi modification of the cusps
of the upper molars, the size of the prenasal notch, and the
size and shape of the tragus. In having cusp 2 oi nd still
more reduced in size, cusp 3 still more posterior in position,
and the prenasal notch still deeper, the strictly continental
Eucheira is clearly on a higher level of development than the
Malayan and continental Megaderma.
The Australian Macroderma is undoubtedly an offshoot of
the Megaderma-Eucheira branch ; broadly speaking, it
accords with these latter genera in the general shape of the
frontal shield, the strong reduction of cusp 2 of and the
posterior position of cusp 3 of nd, but it has completely lost
and the prenasal notch is unusually deep and different in
shape.
The two Ethiopian genera Lavia and Cardioderma, forming
the second section of the family, are closely related inter se.
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145
On the Bats of the Family Megadcrmatldoc.
Lavia is the more primitive : the frontal shield is not so
profoundly modified as in Cardioderma^ is quite normal
in structure. In Cardioderma the frontal shield attains its
highest degree of modification, the posterior of the external
cusps (cusp 3) of- is moved somewhat backward, and the
antero-internal basal cusp of the upper canines, present in all
other genera, has quite disappeared.
Summary. — Selecting the most primitive of the cranial
and dental characters preserved in the five living genera of
Megadermatidge, we are able to draw up a rough sketch of
the skull and teeth of the unknown prototype of the family.
It was a bat with the frontal shield not very considerably
different from that of Megaderma and Eucheira, with the five
primary cusps of ml practically of equal size, as in the typical
molar of an insectivorous bat, and with a small From
this type of bat originated, on the one side the Malayan and
Oriental Megaderma and Eucheira : frontal shield not largely
modified, preserved, but cusp 2 of m} more or less reduced,
cusp 3 more Or less moved backward; and the Australian
Macroderma', essentially as Megaderma and Eucheira^ but^^
lost ; on the other side the Ethiopian Lavia and Qardioderma :
molar cusps almost normal, but frontal shield profoundly
modified, and lost.
The subjoined diagram gives a view of probable inter-
relations and phylogeny of the genera : —
Macroderma.
Cardioderma.
Ann. & Mag. N. Hist. Ser. 7. Vol. xix.
10
146
Prof. G. Chilton on
XV. — Note on the Crah Ilymenosoma depressum, Jacquinot
and Lucaff. By Charles Chilton, M.A., D.Sc., F.L.S.,
Piofessor of Biology, Canterbury College, New Zealand.
[Plate y.]
The little crab Ilymenosoma Jacquinot and Lucas,
was described many years ago*, but does not appear to have
been recognized since, and some confusion and doubt has
therefore arisen with regard to it. As I happen to have seen
some specimens which, in my opinion, undoubtedly belong to
the species intended by Jacquinot and Lucas, it is only just
to their memory that I should endeavour to establish the
validity of the species in question ; at the same time I can
add one or two facts which are, perhaps, not without interest.
White, in 1846, established Ilalicarcinus as a subgenus of
Ilymenosoma, and the second of the two species placed under
it by him was Ilymenosoma depressum, Jacquinot, a species
which was transferred to Ilymenicus by Dana in 1852.
According to Miers, however, the specimens referred to
Jacquinot^s species by White were distinct from it, and he, in
1876, named them Elamena Whitei'\. Miers had not seen
any specimens really referable to Ilymenosoma depressum,
Jacquinot and Lucas, and in the ^ Catalogue of the N. Z.
Crustacea,^ p. 51, gives this species under the name
‘^Ilymenicus depressusl,” quoting Jacquinot and Lucases
description, and adding: —
“ The carapace is represented as nearly circular, the front
very narrow, not longer than the eyes, and of equal width
throughout. There is apparently a small tooth outside the
eyes. Third joint of the external maxillipeds much the
largest. I have not seen specimens of this species.'’^
H ere the story of this species appears to end so far as
printed records are concerned. The new facts that I am
able to add are as follows : —
In April 1888, in the collection of Mr. E. Helms, of
Greymouth, which was submitted to me for examination, I
found a dried crab which, by its yellow colour, flat depressed
carapace, and long legs evidently belonged to the species
described and figured by Jacquinot and Lucas. In 1893,
in the collection of the late Mr. S. H. Drew, of Wanganui, I
found two specimens of the same kind which were labelled as
* Voj. Pole Sad, Zool. iii. Crustaces, p. G2, pi. v. figs. 34-39.
t See Stebbiag, P. Z. 8. 1900, p. 522.
llymenosoma depressiim, Jacquinot and Lucas. 147
liaving been dredged on tlie Grxiyinouth bar in 20 fatliomg ;
very probably they had been eollected by Mr. Ilolmg at the
same time as the speeimen in his own eollection.
I have notes on these speeimens, made at the time, stating
tliat they appear 'to agree well with the original description
and figures by Jacquinot and Lucas, with which I had com-
pared them, and that Miers^s remarks as to the figure are not
altogether correct, but that the front is really ‘‘ dentiform,"’^
{. e. tooth-like, and narrowing to its extremity, instead of
being of equal width throughout,’’^ and that there could
scarcely be said to be a small tooth outside the eyes
either in the figure or in the specimens. In my MS. notes I
also mentioned the fact that the terminal joints of the last
four pairs of legs were fringed with hairs and looked as if
they were used as swimming-organs, .like the legs of the
water-boatmen (Notonectidae).
T saw no further specimens of this species till December
1905, when Mr. Marriner brought me a dried specimen
found at Sumner, Canterbury, precisely like those already
referred to.
This specimen is a male, while the original description and
figure were drawn up from a female specimen, so that I can
add to it the few points in which the male differs from the
female.
The well-marked fringe of long hairs on the distal joints of
the last four pairs of legs is worthy of notice, for tliis, com-
bined with the flat depressed carapace and the very long legs,
shows, I think, that the crab is capable of swimming.
Jacquinotks specimen is said to have been taken under stones
at low tide on the shores of the Auckland Islands ; the two
in Mr. Drew^s collection were dredged on Greymouth bar at
20 fathoms ; I have no definite information as to the circum-
stances under which the other two specimens which I have
seen were taken, but there is nothing in these records incon-
sistent wdth the powder of swimming. Most of the ordinary
swimming-crabs have the last pair of legs specially flattened
for the purpose, but probably many other crabs possess some
power of progression by swimming even without special
modification of the legs ; this is certainly the case with the
young of Petrolisthes elongatuSj Milne- Edwards, just hatched
out from the last Zoeea-form, as I know from personal observa-
tion that it can swim with considerable agility.
The whole of the Hymenosomidm of New Zealand require
careful revision, and until that is done the generic position of
this species must remain an open question ; in the meantime,
however, I give a new description of the species, which, with
10^
148 On Ilymcnosoma depressum.
the figures, will, I hope, fa'cilitate its identification in the
future.
IJymenosoma depressum^ Jacquinot and Lucas.
(PI. V. figs. 1-4.)
ITymenosoma depresmm, Jacquinot and Lucas, Voy. Pole Sud, Zool.
iii. Crustaces, p, 62, pi. v. tigs. 34-39.
Male. — Carapace nearly circular, very slightly longer than
broad ; flat, depressed, lateral margins without teeth ; the
slightly raised margin of carapace continued over the base of
the rostrum. Rostrum short, not longer than the eyes,
narrowing to the subacute extremity ; orbits not interrupting
the margin of the carapace, the lateral portion of the orbit
separated by a slight cleft from the upper portion, so that the
lateral portion in dorsal view gives almost the appearance of
a small tooth outside the eyes *.
Anterior legs (chelse) rather short, only moderately en-
larged j meros and carpus with inner surface supplied with a
few short scattered hairs, outer surface smooth ; propod with
inner surface nearly flat, with short scattered hairs which
extend along the lower border of the fixed finger, outer surface
strongly convex, smooth ; fingers slender, their inner surfaces
with short setae, fixed finger with a small tooth near the base,
movable finger with a larger tooth placed a little more
distally, the rest of the opposing margins with small irregular
teeth, that of the movable finger also bearing small tufts of
setae.
Tiie succeeding four pairs of legs of approximately equal
length, greatly elongated, fully two and a half times as long
as the carapace, slender, without spines; the meros con-
siderably longer than any of the other joints except the
propod ; the carpus about half as long as the propod ; dactyl
slender and acute, slightly longer than the propod ; in all
the legs the proximal joints bear a few scattered setae, while
a continuous row of closely-packed set^ fringes the hinder
margin of the three or four distal joints, the setae being
longer than the joint is wide.
Abdonien of male short, triangular, only slightly longer
than its breadth at the base, and reaching only to the poste-
rior margin of the sternum corresponding to the fourth pair
of legs.
Length of carapace (without rostrum) 12*5 mm., breadth
* These points are not well shown in fig. 1 ; an attempt has been made
to show them more accurately in hg. 2.
^nrr ^ Hia L J.7 VoL AJX J^L \
HVMEN050MA DEPRES5UM , Jacq & Luc
LAF and C.C del.
On new Species of Shells.
Ud
„ .t «»i. '«« >*"•
without teeth.
Auckland Islands.^
explanation of plate V.
Ilymenosoma depressum, Jacq. & Luc.
XYL-Descnptwnsofne,^8p^ DlymsZ'fcSylus
Alycseus. By Hugh C. Fulton.
[Plates IX. & X.]
Trochomorpha modesta^ sp. n. (Pk o )
Shell rather widely
colour; spire co.uc, whorls
stri* crossed on * “£;[td ne“ last sharply
6, convex, f ^ !he margin ; aperture very oblique ;
but is smaller,
r!‘ j5 JuTby tlm Xght depression just above the suture and
'^"iadllf stilted from T. castra, Bens., by its wider
umbilicus and uniform coloration.
• [There are specimens of thi^species “ more slender
S " c"
150
Mr. II. C. Fulton on
Trocliomor'plia crassicarinaia^ sp. n. (PI. IX. fig. 2.)
Shell moderately solid, widely umbillcated, apex yellowish,
remainder o£ shell of a uniform dark brown colour ; whorls 6^,
slightly convex above, the last decidedly so below, first two
smooth, others with rather conspicuous and irregular oblique
strite, last whorl compressed and rounded at the keel ; aper-
ture very oblique, dark within ; peristome simple, sliglitly
thickened at basal and columellar portion.
Maj. diam. 20, alt. 8 mm.
Hah. Nias Island, N.W. Sumatra.
Chiefly characterized by its dark coloration and thickened
keel.
Trochomorjpha niasensiSj sp. n. (PI. IX. fig. 3.)
Shell rather thin, subtransparent, moderately umbilicated,
light yellowish brown, suture of lower whorls and keel of last
edged with dark brown ; whorls 6, very slightly convex
above, first one and a half smooth, others with oblique striae
or growth-lines; aperture whitish within, oblique; peristome
thin, somewhat thickened at columellar portion.
Maj. diam. 21, alt. 8 mm.
Hah. Nias Island, Sumatra.
This species differs from T. crassicarinata by its thinner
substance, lighter coloration, narrower umbilicus, and its
more rapidly increasing whorls, more especially noticeable on
a comparison of the last whorl.
Cochlostyla (Anixa) propitia^ sp. n.
(Pl.TX. figs. 4-6.)
Shell imperforate, solid, subglobosely depressed, upper
])art light reddish, the lower dark, covered with a lighter
yellowish-brown epidermis, which consists of oblique narrow
streaks crossed on the last whorl by more or less distinct,
narrow, interrupted bands of arrow-like markings; this
coloration is continued to a point just below periphery of last
whorl, the remainder of the underside being of a blackish-
brown colour; whorls nearly 5, moderately convex, suture of
lower whorls narrowly impressed, last whorl subangular and
descending slightly at its termination ; aperture suboval,
bluish white within ; peristome thickened and expanded,
slightly reflexed, blackish brown, except the upper expanded
columellar portion, which is whitish ; columella oblique,
thickened ; margins of peristome connected by a slightly
raised transparent callus.
151
new Species of Shells.
I\Taj. (liani. 40, alt. 30 mm.
Var. A (fig. 5). — Sharply keeled at the periphery. Maj.
diam. 40, alt. 26 mm.
Var. B (fig. 6). — Globose. Maj. diam. 30, alt. 28 mm.
JJab. Cebu Island, Philippines.
(= Moreleti, MlIdfF., non Pf., Berichb d. Senek. natur.
Gesellseh. 1890, p. 238.)
Tiiis shell has been distributed by MollendorfF and others
as Cochlostyla (^Anixa) Moreleti, Pf., but that s{)ecies is
(judging from the figure and description and specimens in
tiie British Museum) but a specimen of C. (Anixa) Mont-
fortiana, Pf., that has lost its epidermis.
C. propitia is closely allied to G. carhonaria, the chief
difference being that tlie former is a thicker shell with a
much lighter-coloured epidermis than tlie latter. G. propitia
is also much larger generally and broader in proportion to
height, and its peristome is more expanded ; but these cha-
racters are not constant in the large series under examination.
The great variation in the form of G. propitia is shown by
the three shells selected for description.
Ampliidromus cognatus, sp. n. (PI. IX. fig. 7.)
Shell sinistral, moderately solid, minutely perforate, smooth, ~
shining, whitish ground, covered on lower whorls by a bright
yellow periostracum ; whorls nearly 6, slightly convex, first
two dark reddish brown, remainder with dark bluish-grey,
narrow, spiral bands, one at tlie sulure and one above tlie
middle, tliree on the last whorl, one about 2 mm. wide at the
periphery, a similar one encircling the umbilicus, and a
narrow one about 2 mm. below the suture, lines of growth
rather conspicuous ; aperture subovate, white, with the outer
band showing clearly through ; peristome white, moderately
expanded.
Maj. diam. 17, alt. 31 mm.
Hah. ?
In the position of the colour-bands and their showing
clearly through the aperture this species greatly resembles
A, hemicyclus, Bochebrune, but the latter is much narrower
— so narrow tliat one almost doubts the correctness of the
dimensions given, viz. : long. 30, lat. 10 mm.
Amphidromus niasensis, sp. n, (PI. IX. fig. 9.)
Shell sinistral, moderately thin, finely obliquely striated,
nucleus of a semitransparent waxy colour, rest of shell witii
light reddish ground ornamented by rather broad oblique
152
lUr. II. C. Fulton on
stripes which are intersected in the middle by a narrow spiral
band of lighter colour ; the under part of last whorl has a
rather broad dark brown band situated just below the peri-
phery and a narrower one lower down, between which is a
yellow one, umbilical area reddish ; whorls 6, convex, rather
slowly increasing ; aperture with outer markings showing
through ; peristome slightly expanded, flesh-colour ; columella
vertical, flesh-coloured, rounded and slightly expanded at
point of insertion.
Maj. diam. 17, alt. 30 mm.
Tlab, Nias Island, Sumatra.
Although totally distinct in coloration, the details of this
species agree very closely with A. Sowerbyi\ but the whorls
of niasensis are slightly more coiivex and increase a little
slower in size than those of A. Sowerhyi.
It is also similar to pcecilochroa, Fult., in form and
markings, but is thinner and its wdiorls are more convex.
Amphidromus Sowerhyi^ sp. n. (PI. IX. fig. 10.)
Shell sinistral, thin, umbilicus almost closed, nucleus dirty
white with a brown spot at the apex ; lower whorls yellow,
with six narrow dark brown spiral bands on the middle
whorls, the last whorl having two broader bands in front, one
situated just below the periphery and the other a little lower,
umbilical area yellow', finely obliquely striated ; whorls 6|-,
moderately convex, rather slowly increasing ; aperture with
the outer bands showing through ; peristome very narrowly
expanded, w’axy colour ; columella vertical, narrowly expanded
at point of insertion.
Maj. diam. 17, alt. 31 mm.
Hah. Nias Island, Sumatra.
This new form bears a great resemblance in coloration and
markings to some of the varieties of Helix nemoralis, Linn.
The number of bands varies, one specimen before me having
only a single narrow peripheral band.
Named in honour of my esteemed and genial colleague,
G. B. Sowerby, Esq., F.L.S.
Amphidromus Webhi^ sp. n. (PI. IX. fig. 8.)
Shell sinistral, moderately solid, earlier whorls yellow,
fading to cream on lower wdiorls, with a broad lightish
chestnut band about 14 mm. wide encircling the last whorl
and continued as a narrow ever-diminishing band at suture of
penultimate whorl ; whorls 6|, moderately convex, with
inconspicuous oblique growth-lines, last ascending slightly at
153
neic Species of Shells.
its termination ; aperture subovate, white within ; peristome
rather broadly expanded and slightly reflected ; columella
vertical, expanded above.
Maj. diam. 31, alt. 51 mm.
Hab. Nias Island, Sumatra.
A handsome and distinct new form, which can be readily
separated from A. enganoensiSj Fult., by its much less convex
whorls and less broadly dilated columella.
Named in honour of Walter F. Webb, Esq., of Rochester,
New York.
Bidimulus [Protoqh/ptus) dejectus, sp. n.
(Pl.X. fig. 1.)
Shell very narrowly urabilicated, acuminately elongate,
rather thin, covered with yellowish-brown epidermis, wdiite
beneath, apical structure consisting of oblique, microscopic,
wrinkled or granular striae, lower part with inconspicuous
oblique striae or lines of growth crossed by close-set spiral
rows of minute hairs ; whorls 8, slightly convex, regularly
increasing, the last not deflected ; suture rather deep, simple;
aperture suboval, wfliite within ; peristome very slightly
expanded, broader at point of insertion of columellar portion,
margins joined by a thin transparent callus.
Maj. diam. 10, alt. 29 mm.
Hab. Santa Catharina [fide Linn^a Institute label).
The nearest species to this known to me is crepundia^ Orb.,
but that is readily separated by its less cylindrical form.
Drymceus volsusj sp. n. (PI. X. fig. 2.)
Shell elongately fusiform, minutely rimate, rather thin,
almost smooth to the eye, but under the lens the usual
Brymceus sculpture on nuclear whorls, while the lower-
whorls have oblique blunt costae or lines of growth crossed by
microscopic close-set spiral striae ; whorls 65, slightly convex,
first three and a half dirty white, lower with cream ground
ornamented by irregular, oblique, somewhat zigzag, dark
brown stripes which are broken by narrow white lines and
dots ; aperture oblong-oval, dark brown markings within ;
peristome rather broadly expanded, especially at the basal
portion, pale yellow ; columella narrow and cord-like,
entering spirally.
Maj. diam. 12J, alt. 30|- mm.
Hab. Ecuador.
I know of no other species wdth which to make a helpful
comparison.
154
Mr. II. C. Fulton uw
]\Ir. S. I. Da Costa, who has made a special study of this
^enus, is unable to identify it with any species known to
him.
riacostylus [Euplacostylm) cyUndricus. sp. n.
(PI. X. fig. 3.)
Shell elongate, almost imperforate, solid; spire reddish,
lower part covered by a greenish-brown cuticle; whorls 6J,
slightly convex, rather rapidly increasing, last two witli
longitudinal growth-lines, crossed on middle whorls by obscure
spiral lines, giving the appearance under the lens of a some-
what reticulated suif'ace, last whorl indistinctly malleated in
parts ; aperture ear-shaped, dirty white within ; peristome
thickened, expanded inwardly, wliite, outer edge yellowish,
margins connected by a thin callus, columella with a wdiite,
thick, spirally entering fold.
Maj. diam. 23, ait. 71 mm.
Peristome : maj. diam. 27, alt. 33 mm.
Ilah, Isabel Island, Solomons [Meek),
Separated from Seemani, Dohrn, by its narrow cylindrical
form, more rapidly increasing whorls, and much shorter
peristome.
From koroensis^ Garrett, to which it has some resemblance
in form, it can be easily distinguished by its larger size and
dark-coloured epidermis. Of the three specimens before me
none appear to have the punctures found on the spiral whorls
of the two above-mentioned species ; but that character may
have been worn off.
Stenogyra [Euonyma) Beckeri, sp. n.
(PL X. fig. 7.)
Shell elongately fusiform, moderately thin, polished, sub-
transparent, nucleus whitish, lower whorls of a pale olive-
greenish colour ; apex obtuse, rounded ; whorls 12, slightly
convex, first three smooth, slowly and regularly increasing,
lower whorls with indistinct lines of growth, slightly crenu-
lated at the suture ; aperture oblong-oval ; columella slightly
curved, white, slightly expanded ; peristome continuous with
columella, thin.
Maj. diam. 10, length 44 mm.
Length of aperture 9, width 5 mm.
11 ah. Pondoland [Dr. H. Becker').
Differs from S. Purcellij Melv. & Pons.* (the type of
^ Arm. & Mag. Nat. Hist. 1901, vol. viii. p. 317, pi. ii. fig-. 0.
155
new Species of Shells.
wliicli is a young shell), by its broader form and more
rapidly increasing whorls. In Beckeri the first two whorls
are smooth, whereas in PurcelU they are distinctly crenulated
at the suture.
Tliis comparison was made with the type of PurcelU^ no\y
in the British Museum, and a young specimen of S. Deckeri.
In the description of S. PurcelU the species is characterized
as having no sculpture^ but that is not correct ; the figure also
gives one the idea of a broader form than that of the actual
type.
Leptopoma niasense^ sp. n. (PL X. fig. 6.)
Shell globosely conic, narrowly umbilicated, thin, sub-
transparent, very light brownish ground with darker-coloured
raised spiral striae, about 6 on middle whorls ; between these
are close-set microscopic spirals, a patch of darker brown at
umbilical area ; whorls 5, moderately convex, last subcarinate
in front; aperture circular, rather dark w'ithin ; peristome
expanded, whitish both front and back, margins approxi-
mating and joined by a thin callus ; operculum corneous,
thin, 8 whorls.
Maj. diam. 14, alt. mm.
Ilab. Xias Island, N.W. Sumatra.
Similar to L. pellucidumj Grat., but can be readily sepa-
rated by its slightly broader peristome and dark-coloured
umbilical area.
The species varies somewhat in coloration, some specimens
having patches of a smoke-colour and others being banded.
Cyclophorus iTheohaldius) Dautzenhergi. sp. n.
(PI. X. fig. 8.)
Shell depressed, spire plane, moderately solid, widely
umbilicated, dark brown, with rather distant oblique streaks
of lighter colour ; wdiorls 4|, convex, closely obliquely
striated, upper part of last whorl with five or six more or
less distinct spiral striae, which stop short on the penultimate
whorl, third of last whorl slightly and gradually descending ;
aperture circular, bluish white within ; peristome with
yellowish-white edge, surrounded by a dark-coloured flange,
giving the peristome a duplex appearance ; operculum
corneous, fairly solid, about 12 whorls.
Ma,j. diam. 25, alt. 10 mm.
Hah. Nias Island, N.W. Sumatra.
Bearing a likeness to the Ceylon species G. Layardij Ad.,
15G Mr. II. C. Fulton on
but easily distinguished by its flatter form and smaller
aperture.
Some speeirnens are almost a uniform brovvm, whilst others
are strikingly marked with yellowish-white, narrow, oblique
streaks.
Named after Mons. Ph. Dautzenberg, one of our most
esteemed conehologists.
CyclopJwrus {Cyclohelix) Kihleri^ sp. n.
(Pi. X. fig. 4.)
Shell turbinate, very narrowly umbilieated, very solid,
nueleus consisting of whorls, first one and a half smooth,
the next with rather distant curved stride, remainder of shell
with fine oblique striae, crossed by close-set microscopic
spirals which are waved on the underside of body-whorl, rich
dark brown colour, with numerous irregularly shaped spots
and streaks of lighter colour ; on the underside of shell the
markings take a more regular spiral pattern ; whorls 5^,
convex, the last a little flattened below ; aperture orange-
coloured within, very oblique, subirregularly oval; peristome
orange, well thickened, margins approaching and connected
by a transparent callus, columellar portion sloping towards
the right, with a blunt tooth-like projection about the middle;
operculum corneous, thin, about 8 whorls.
Maj. diam. 31^, alt. 27 ram.
Hab, Nias Island, N.W. Sumatra.
This shell is of a similar form to C. turho^ Chem., from
the Nicobar Islands, but cannot possibly be confused with
that or any other species of the genus known to me.
Cydotus niasensis, sp. n. (PI. X. fig. 9.)
Shell depressed, spire almost plane, apex dark coloured,
somewhat roughened and slightly exserted, rather thin,
covered with a somewhat thick closely striated epidermis ;
whorls 4^, moderately convex, last shortly and slightly
descending, suture deep ; aperture circular, bluish white
within; peristome with narrow outer flange; operculum
calcareous, whorls 9.
Maj. diam. 15, alt. 6 mm.
Hah. Nias Island, Sumatra.
The nearest to this known to me is C. discoideus^ but
C. niasensis is smaller, the peristome does not descend so far,
and the epidermis appears to be thicker.
/
7. Vnl.m. PLJX.
oI.'Ga?een del- et litK.
rl^:
LIBRARY
OF THE
UNIVERSITY of ILLINOIS
(J. Green del et lith.
LIBR/iRY
OF THt
UNlVEKSlTy ul ILLINOIS
new Species of Shells.
157
Alycceus [Chamah/cceus) Smithij sp. n.
(PI. X. fig. 6.)
Shell depressed-turbinate, dirty white, rather widely
umbilicated ; whorls 4, first smooth, remainder with oblique
striae which are rather widely spaced on first half of last
whorl, then suddenly become very close, widening out again
immediately before the strangulated portion, whicfi is situated
about one third of a whorl from the peristome, last whorl
sliortly descending; aperture circular; peristome rather
thick, duplex ; sutural tube ratlier long ; operculum thin,
whitish, concave, with 9 or 10 wdiorls.
j\Iaj. diam. 4, alt. 2 mm.
Hah. Mengtzu, Yunnan [fide Carl Bock).
Very like A. 'plicilabris ^ Molldff., but has no plicae on the
peristome, the striae immediately beliind the strangulation are
similar, but the striae on the earlier part of the whorl are
closer than in Mollendorff’s species.
From A. rathousianus^ Heude, it differs by being smaller,
has a more strongly produced duplex peristome, and its
sculpture on the last wdiorl is finer.
Named in honour of Edgar A. Smith, Esq., I.S.O.
EXPLANATION OF THE PLATES.
Plate IX.
Fig. 1. Trochomorpha modesta.
Fig. 2. crassicarinata.
Fig. 3. niasensis.
Fig. 4. Cochlostyla propitiu.
Fig. 5. , var. A.
Fig. 6. , var. B.
Fig. 7. Amphidromus cognatus.
Fig. 8. Wehhi.
Fig. 9. niasensis.
Fig. 10, Sowerhxji.
Plate X.
Fig. 1. Bulimulus dejectus.
Fig. 2. Drgmcexis volsus.
Fig. 3. Flacostglus cglindricus.
Fig. 4. Cgclophorus Kibleri.
Fig. 5. Algcceus Smithi.
Fig. 6. Leptopoma niasense.
Fig. 7. Stenoggra Beckeri.
Fig. 8. Cgclophorus Dautzenbergi.
Fig. 9. Cgclotus niasensis.
16S
Mr. II. C. Fulton on Papuina and Pupina.
XVII. — Notes on Papuina and Pupina.
By Hugh C. Fulton.
On the Identity of Papuina (Dendrotrochus) pumila, Falt.^
loith Dendrotrochus mentuni, Tledley.
Since tlie publication of P. pumila in the ^Journal of
Malacology/ 1905, vol. xii. p. 22, pi. 6. figs. 5 & 6, Mr. Chas.
Hedley has written me that it is probably his Dendrotrochus
mentuiUy described in the ‘ Records Australian Museum/
1899, p. 153, fig.
Unfortunately I had omitted to look up the subgenus
Dendrotrochus in the Record, only having looked for Papuina,
To make certain of the matter, I recently sent a co-type of
P . pumila to Mr. Hedley, who now informs me that it is
identical with his D. mentum ; my species therefore becomes
a synonym.
On Papuina Sellersi, Gox, and P. migratoria, Pfr,
Dr. Cox described P. Sellersi in the P. Z. S. 1876, p. 646,
pi. 52. fig. 9, and appears to have fixed no particular speci-
men as the type, but to have based the species on a number
of specimens ; the measurements of one specimen are given —
viz. : diameter, greatest 0’78, least 0*56, height 0*60 of an
inch/’ — but they do not agree with the figure given.
After examination of a number of specimens bearing a
label in Dr. Cox’s handwriting Typical P. Sellersi^ I can-
not separate it from Pfeiffer’s P. migratoria^ a species most
variable in form and coloration.
The two forms described as migratoria, Pfr., and Sellersi,
Cox, are linked closely together by specimens before me, which
show them to be one species having great variation in form
and coloration : some specimens are sharply carinated, whilst
others have the last whorl rounded; a few are quite imperforate,
whilst others are slightly perforate.
Coloration varies from specimens all white, except for a
narrow dark brown sutural band, to specimens of yellowish
ground with broad spiral bands.
All the varieties have inconspicuous oblique strise, crossed
on the lower whorls by oblique, forward descending wrinkles,
which vary in strength and are almost absent in the var.
leucophcea, Cox.
On Pupina Thomsoni, Forbes, and P. bidentata,
C. E. Beddoine.
After comparison of typical specimens of the above-
mentioned species, I am forced to the conclusion that P. hi
150
On a new Cij’prinovl Fish.
dentala is simply Thomsoni with the callus-folds situated at
the anterior area of the back of the shell more strongly
developed.
In a series of co-types from the collection of the late
Mr. C. E. Beddome there are some specimens entirely without
these foldsj and a selected series seems to demonstrate that
these folds are only fully developed in the last stage of
growth.
It is significant that in the description of P. hidentata there
was no comparison made with the obviously closely allied
P. Thomsoni.
XVIII. — Description of a new Cyprinoid Fishj Acheilo-
gnathus signifer, Korea, with a Synopsis of all the
known E,hodeina3. By L. S. Berg (St. Petersburg).
Acheilognathus signifer.
Cdosely allied to A. cyanostigma^ Jordan & Fowler, 1903,
from Japan (Lake Biwa), differing chiefly in coloration and
longer barbels.
No dark shoulder-spot above gill-opening, no longitudinal
band on body, neither on tail. Dorsal deep hrown^ with a
broad marginal whitish hand. Anal with several darker
longitudinal cross-bars. Ventral blackish.
D. II 8. A. II 8. L. 1. 36 j.
Teeth 5 — 5, not serrated. No spines in dorsal and anal.
Barbels present, rather long, somewhat more than half the
length of the eye, reaching to the vertical from the anterior third
of eye. Mouth small, subinferior. Maxillary reaching to the
nostrils. Depth of body 2| in its length (without caudal),
length of head 4 ; diameter of eye J length of head, slightly
less than interorbital space, equal to snout, in postorbital
space. Caudal peduncle 3j^ in body, twice as long as deep,
its depth 2| in that of body. Origin of dorsal somewhat
behind the root of ventrals, midway between end of snout and
root of caudal ; its end opposite to the third branched ray of
anal. Base of dorsal in body, height 6. Upper margin
of dorsal straight. Origin of anal below the posterior
dorsal rays; its base in body, its height 6^. Pectonds
not reaching ventrals, 5f in body ; ventrals 6, reaching origin
of anal. Lateral line slightly decurved ; 6 scales between
IGO
Mr. L. S. Berg on a
lateral line and middle of belly. Anus midway between
origin obthe ventrals and anal.
A single adult male (with two semicircular excrescences
on snout), measuring about 57 mm. (47 without caudal),
taken by the late O. Ilerz in Pungtung, Korea, in 1888.
Type specimen N. 10265 in the Zool. Mus. of the Acad, of
►Sciences at St. Petersburg.
The following is a synopsis of all the species known to me
of the group Rhodeinse * : —
I. Rhodeus, Agassiz, 1835.
Lateral line incomplete. No spine in dorsal and anal. Pharyngeal
teeth 5 — 5, not serrated. More than 7 branched rays in anal.
rt. No barbels; D. Ill 9-10, A. ITT 8-10,
1. 1. 34-40.
b. Third suborbital not more than half the
length of the eye. (Central Europe,
N.W. and S. Russia, Macedonia,
Anatolia, Caucasus, basin of Amur,
Manchuria f-)
Jih. sericeus (Pallas), 1776 { = Rh. amarus, Bloch, 1782).
bh. Third suborbital more than half
the length of the eye. (Southern
China.) Rh. sinensis, Gunther, 1868.
aa. Barbels present ; I). Ill 8, A. Ill 9,
1.1. 32; teeth 5 — 5 (not serrated?).
(Japan, Kiu-siu.) Rh. (?) oryzce, .Iordan & Seale, 1906 f.
II. Pararhodeus, gen. nov.
Lateral line incomplete. No spine in dorsal and anal. Pharyngeal
teeth 5 — 4, slij^htly serrated. D. II 7, A. Ill 6, 1. 1. 48-49. — A single
species : P. syriacus (Lortet), 1883. (Syria, Damascus.)
III. Achbilognathus, Bleeker, 1860.
Lateral line complete. Pharyngeal teeth 5 — 5, not serrated. Barbels
present or absent. Spine in dorsal and anal absent or present.
A. Barbels present. No spine in dorsal and
anal.
a. Shoulder with a more or less distinct
* Messrs. Jordan and Fowler (Prcc. U.S. N. Mus. xxvi. 1903, pp. 812,
822) refer Capoeta elonyata, Schl., and C. gracilis, Schl., to the group
Rhodeinae (“teeth one-rowed”). Although Schlegel does not indicate
the dentition of these species, Bleeker (Atlas ichth. iii. 1863, p. 117 ;
Ned. Tijdsch. Dierk. ii. 1865, p. 138), who has had specimens from Japan,
states that they are three-rowed, 1.3. 5 — 5 .3.1 (“ dentes raptatorii ”)
as in Barhince, which induced Dr. Gunther (Cat. Fish. vii. 1868, p. 136)
to refer the Bleekerian genus Gnathopogon to Barbus.
t A very remarkable instance of an interrupted distribution : not yet
known from Siberia.
\ Scarcely belonging to this genus.
new Cijpruioid Fish.
lOL
ilark shoulder-spot
openiiii:
above gill-
h. Barbels short, about half the length
of the eye ; shoulder-spot and lateral
band very distinct. 1). Ill 10,
A. In 10. (Japan.) A. limbatus (Schlegel), 1846.
hb. Barbels more than half tlie length
of the eye ; shoulder-spot and lateral
band indistinct or absent. D. Ill 9,
A. Ill 9. (Japan.) A. lanceolatus (Schlegel), 1846.
au. Shoulder without dark spot above
gill-opening.
c. Barbels short, not more than half the
length of the eye.
d. A longitudinal dark band ; dorsal
with several darker longitudinal
cross-bars ; anal dark, with a
broad marginal whitish band.
Maxillary barbel very short.
B. Ill 8, A. Ill 8, 1. 1. 39.
(Japan.) A. cyanostigma, Jordan & Fowler, 1903.
dd. No longitudinal dark band ;
dorsal dark, with a broad mar-
ginal white band ; anal with
several darker longitudinal cross-
bars. D. II 8, A. II 8, 1. 1. 36.
Maxillary barbel about half the
eye. (Korea.) A. signifer, Berg, 1906.
cc. Barbels long, as long as eye. A
longitudinal dark band along the
middle of tail ; lower half of dorsal
with a series of blackish dots ;
I). II 9, A. II 11-12, 1. 1. 34. (For-
mosa.) A, himantegus, Giinther, 1808.
AA. Barbels present. Spine in dorsal and
anal present.
e. I). II 11, A. II 9, 1. 1. 36; depth 2^ in
length of body, snout shorter thai
eye. (Shanghai.) . A. barhatulusj Gunther, 1873.
ce. D. Ill 11-13, A. Ill 11, 1. 1. 35-36 ;
depth 2J—2i in length of body ; snout
as long as eye ; a dark blue spot
above gill-opening. (Korea, Seoul.)
A. coreanuSf Steindachner, 1892.
AAA. Barbels absent *.
f. Dorsal and anal without spine.
D. II 14-15, A. II 14-15, 1. 1. 35-38.
(Japan, L. Biwa.) A. lonyipinnis, Regan, 1905.
ff. Dorsal and anal with spines.
D. Ill 12-13, A. Ill 10, 1. 1. 35-36.
(Basin of Amur, L. Khanka or [Dyb.l, 1872.
Hanka.). . A. chankaensis (Dybowski) { = Devario chankaensiSf
* Acheilognathus mesemh'inum, Jordan & Evermann (Proc. U.S. N.
M. XXV. 1902, p. 323, fig. 6), from Formosa, apparently does not belong
to the Rhodeinae.
Ann. (k Mag. N. Hist. Ser. 7. Vol. xix.
11
162
Mr. L. S. Borg on a
IV. Paracheilognathus, Bleeker, 1863.
Lateral line complete. Pharyngeal teeth 5 — 5, deeply serrated. No
spine in dorsal and anal.
a. Barbels (veiy minute) present ; mouth
inferior ; D. II 14, A. Ill 10, 1. 1. 37-38.
Head 4g in length of body, depth 2|,
eye 4 in head. (Japan.) P. rhombeus (Schlegel), 1840.
aa. No barbels; month oblique.
b, 13. II 10, A. II 12, 1. 1. 35. Head 4 in
length of body, depth 3, eye 3 in
head. (China.) [berbis, Gthr.).
P. imberbis (Giinther), 1868 { — Achilognathus im-
bb. D. Ill 13-14, A. Ill 10-11, 1. 1. 35.
Head 4 in length of body, depth 2^,
eye 2f in head. ( Yang-tse-kiang,
Tien-tsin.). . . . P. Bleekeri, n. n. ( = P. imberhis, Bleeker, 1871,
[non Achilognathus imberbis, Giinther^).
V. PsEUDOPERiLAMPus, Bleeker, 1863.
Lateral line incomplete. Pharyngeal teeth 5 — 5, deeply serrated.
No spine in dorsal and anal. No barbels.
a. H. HI 10-11, A. Ill 10-11, 1. 1. 55-65.
(Japan.) P. typus, Bleeker, 1863.
aa. D. Ill 11-12, A. HI 10-11, 1. 1. 33-34.
(Shanghai, Y'ang-tse-kiang.) P. 06W/a^ws,Kner, 1865-67.
VI. Acanthorhodeus, Bleeker, 1871.
Lateral line complete. Teeth deeply serrated. Spine in dorsal and
anal present. Barbels present or absent.
a. Barbels present.
D. Ill 17-18, A. Ill 12-1.3, 1. 1. 35 ; head 5 in
length of body, depth 2^. (Yang-tse-
kiang.) A. macropterus, Bleeker, 1871.
H. Ill 17-18, A. Ill 13-14, 1. 1. 36-40 ; head
4|-4^ in length of body, depth 2-2|.
(Yang-tse-kians: ; Shanghai.) A. Guichenoti, Bleeker, 1871.
D. Ill 17, A. Ill 13-14, 1. 1. 34 head 4| in
length of body, depth 2^. (China ;
Swatow.) A, dicmis, Butter, 1898.
D. Ill 15, A. HI 11, 1. 1. 36; head 4 in length
of body, depth 2. (Upper Tonkin.) A. tonkinensis, Vaillant, 1892.
aa. Barbels absent.
D. HI 14-15, A. HI 12-1.3, 1. 1. 30. (Yang-
tse-kiang.) A. hypselonotus, Bleeker, 1871.
D. II 12-13, A. II 10-11, 1. 1. 35. (Shanghai.).
A. atranalis, Gunther, 1873.
D. II 16-17, A. II 13-14, 1. 1. 35-36 ; second
suborbital twice as long as deep ; dorsal
* Cf. Gunther, Zool. Becord, 1871, p. 107.
1G3
new Cyprinoid Fish.
and anal twice as long as high. (Shang-
hai.) A. tcBniatialis, Gunther, 1873.
D. III. 16-18, A. Ill 12-14, 1. 1. 36-39 ; second
suborbital about as long as deep ; dorsal
and anal less than twice as long as high.
(Lower Amur, Ussuri.) A. Asmussi (Dybowski), 1872
[_{=Devario Asmussi^ Dyb.).
XIX. — Description of a new Cyprinoid Fish, Paraleacogobio
notacanthus, N. China. Bj L. S. Berg (St. Peters-
burg).
Paraleucogobio, gen. nov.
Allied to Leucogohio, Gunther, from which it differs in
having the dorsal provided with a spine and inserted behind
'centrals.
Body compressed, deep ; scales large, about 35 in lateral
line; lateral line in the middle of body, anteriorly slightly
decurved, posteriorly straight. Dorsal short, with 7 branched
rays and with a strong, but flexible at the top, spine, in-
serted a little behind origin of ventrals, nearer to end of
snout than base of caudal. Anal short, with 6 branched
rays, wnthout spine, its origin beyond the end of dorsal,
nearer to base of ventrals than to caudal. Anus a little before
origin of anal. Breast scaled. Belly not keeled. Mouth
small, anterior : jaws of equal length ; lower without tubercle,
its edge somewhat trenchant, semilunar. Hind margin of
maxillary reaching the vertical of the middle between
nostrils. Tip of snout about on the level of the lower margin
of eye. Lips thin, the lower interrupted at the symphysis.
A minute barbel at the angle of mouth, equal to | of the
diameter of eye, not reaching the vertical of front margin of
eye. Preorbital reaching the eye. Suborbitals long, narrow.
Gill-membranes attached to isthmus below the hind margin
of eye. Peritoneum silvery, with some black points. Pha-
ryngeal teeth two-rowed, 2 . 4 — 5 . 3, compressed, subuncinate;
upper teeth of the longer series conical.
Faraleucogohio notacanthus^ sp. n.
D. II 7. A. Ill 6. Lin. lat. 34 37.
Snout blunt ; interorbital space convex. Back behind
nape flattened. Pectorals not reaching ventrals, ventrals not
11*
1G4
lyir. G. K. Glide on a new
reacliing anus. Upper margin of dorsal and anal straight.
First unarticulated ray of dorsal about | of second (the
longest). Scales with radial striae. 5 longitudinal series of
scales between lateral line and middle of belly. Depth of
body 3| in length (without caudal) ; length of head 4|-.
Depth of head in its lengthy width IJ ; eye 3*8 in length
of head, 1‘4 in interorbital space, 1*2 in length of snout, 1*8
in postorbital part of head. Caudal peduncle 4*8 in body,
1’6 as long as deep, its depth 2*3 in that of body. Length
of dorsal 7 in body, height 5*2; length of anal 10‘1, height
7'0 ; length of pectoral 5*4, of ventral 5*8. Pectoral in
distance from its base to origin of ventral ; ventral \\ in
distance from its base to origin of anal. Antedorsal space
2 in body, postdorsal 2|.
Total length of a single specimen (mature female) about
95 mm. (without caudal 81 mm.).
In general shape resembles Leucogohio tcematus^ Gunther,
in colour Leuc. Herzensteini^ Gunther. Many longitudinal
dark bands along the sides of body ; a rather broad blue
band above the lateral line. Each scale of the lateral line
with two dark spots (as in Alburnus hipunctatus) . Dorsal
wdth two series of dark spots.
Taken in a rivulet, Je-hol, at Cheng-tu-fu (or Je-hol),
tributary of Lu-ang-ho (N.E. of Pekin), by Colonel D.
Putiata in May 1901. Type specimen N. 9873 in the ZooL
Mus. of the Acad, of Sciences at St. Petersburg.
XX. — Description of a new Species of Eulota from Formosa,
By G. K. Gude, F.Z.S.
Eulota [Euhadra') Warhurgij sp. n.
(Figs. 1 & 2.)
Shell narrowly umbilicated, conoid, shining, finely irregu-
larly striated, with obsolete shallow spiral furrows here and
there; chalky white, with one distinct, narrow, yellowish-
brown band at the periphery and a fainter one above and
below, the upper one very indistinct. Spire depressed- conoid,
suture deep, apex prominent. Whorls nearly 6, increasing
slowly and regularly, convex above and at the periphery,
slightly flattened below ; last whorl scarcely descending in
front. Aperture little oblique, subovate ; peristome thick-
ened and expanded, but slightly reflexed ; margins distant,
Species of Eulota from Formosa. 165
columellar margin strongly thickened and reflexed, slightly
receding over tlie narrow deep umbilicus.
Diam. maj. 27, min. 23 ; alt. 18 mm.
Ilah. Dunes at Long-Krau, South Formosa (^Warburg
leg.^ Feb. 1888).
Type in my collection.
Fig. 1.
b
c
Professor G. Pfeffer, of the Naturhistorisches Museum,
Hamburg, kindly placed at my disposal three specimens of an
undescribed Eulota. All tlie shells appear to be sand-blown,
being denuded of cuticle. The second specimen has the bands
much fainter and is smaller, measuring: — diam. maj. 24,
min. 20 ; alt. 15*5 mm. The third specimen (fig. 2) is still
Fig. 2.
smaller, but has a more elevated spire than the type ; it
measures : — diam. maj. 21, min. 18’5 ; alt. 15*5 mm.
The new species is related to Eulota irrediviva^ Pils. & Hir.,
from Okino-erabu-shima, N.E. Loo-Choo, which it resembles
in the banding ; but tliat species is much larger, measuring
36 mm., although possessing only 5 whorls; in E. Warburgi
the whorls are more convex, the suture is deeper, and the
peristome is less reflexed.
Another near ally is E. ohinoerahuensisj Pils. & Hir. ; but
the new species differs from that shell in being more de?
pressed, smoother, and with weaker striae, in not being spirally
striated, and in having a wider umbilicus.
166
Mr. P. Cameron on Ilymenoptera
XX T. — On some undescribed Phytophagous and Parasitic
Ilymenoptera from the Oriental Zoological Region, By
P. Cameron.
For tlie species described in this paper 1 am indebted to
Col. C. T. Bingham.
TenthredinidaB.
Tentliredo lepchOf sp. n.
Luteous; the antennal scape in the middle above_, the
flagellum, the sixth to eighth and the sides of the ninth
abdominal segment broadly laterally, and the hind legs, except
the coxae and trochanters, black. Wings yellowish hyaline,
the apex from the end of the stigma fuscous, tinged with
violaceous; the apex of the hinder with a narrower luscous
cloud. ? .
Lengtii 16 mm.
Sikkim.
Smooth, shining; the face, clypeus, labrum, mandibles,
lower outer orbits, and base of legs pale yellow. The vertex
in the centre with a short black pile, the face, clypeus,
labrum, and base of mandibles sparsely covered with long
pale hair ; the thorax above thickly covered w’ith short
fuscous pubescence. Apex of clypeus broadly, roundly
incised ; the labrum punctured at the root of the hairs. Apex
of mandibles black. Upper half of the front with a wide
shallow depiession, W’hich is sparsely haired ; the lower with
a deeper, smooth, bare depression, the two parts being clearly
sepal ated ; they are bordered by a broad keel. Centre of
vertex (including the ocelli) bordered by a narrow^, but dis-
tinct, furrows H ead lai'ge, wider than the thorax, the temples
wide, rounded ; occiput sharply margined. The third and
fourth joints of the antenna? are almost of equal length ; the
antennae are densely covered with short stiff black pubescence,
^cutellum prominent, obliquely sloped at the base and
apex.
This species comes close to T. xanthoptera^ Cam.; the two
may be separated thus : —
Hind femora, base of hind tibiae, and apex of abdomen
liileous; the third joint of antennae distinctly longer
than the fointh ; the upper part of the frontal area
roundly curved outwardly, the sides distinct, reaching
to the outer ocelli, the apex of the lower part distinctly
obliquely narrowed xanihoytera.
167
from the Oriental Zoological Region,
Hind legs, except the coxee and trochanters, and the apical
segments of the abdomen black; the third joint of
the antennjB not distinctly longer than the fourth ;
the upper part of the frontal area not curved out-
wardly, the sides not distinctly keeled, not reaching
to the hind ocelli, the lower part not distinctly
obliquely narrowed lepcha.
Siobla ru/O’halfeataj sp. n.
Black, the head and mesonotam tinged with blue ; the
labruni, a line round the pronotum, narrowest in the middle,
the anterior tibiae in front and the outer side of the posterior,
white ; the second to sixth abdominal segments rufous.
Wings hyaline to the first transverse cubital nervure, smoky
violaceous beyond ; the nervures and stigma black. Head,
npper part of thorax, and legs densely covered with
white pubescence, the antennae thickly with stiff black
pubescence. $ .
Length 8 mm.
Yunzalin Valley, Upper Tenasserim ; November ((7. T.
Bingham) .
Front and vertex closely, finely punctured, shining; above
each antenna is a wide, deep furrow, roundly dilated above
and reaching halfway to the ocelli ; in the middle below is
a deep, slightly longer than wide, fovea; there is a wide,
deep furrow between the ocelli in the middle. Scutellum
minutely punctured. Cenchri large, white. The black on
the apex of the abdomen has a violaceous tint ; below it
extends on to the sides of the sixth segment. Middle lobe of
mesonotum furrowed, the furrow wide at the base, gradually
narrowed towards the apex. The middle of the vertex is not
bordered by furrows, the part behind the ocelli forming one
piece. Temples narrow ; occiput not quite transverse, being
slightly curved inwardly. The clypeus is more strongly
punctured than the rest of the head and becomes gradually
narrowed to a point. The third joint of the antennae is dis-
tinctly longer than the fourth ; it is also thinner, becoming
slightly dilated towards the apex; the apical four joints are
thicker than the others, they become gradually narrowed
towards the apex.
Selandria hasilineaia^ sp. n.
Black, slightly tinged with blue ; the clypeus, a narrow
line on the edge of the pronotum, a broader one on the apex
of the fiist abdominal segment, the apex of the middle coxae
1G8
Mr. P. Cameron on Ilymenoptera
narrowly, of the hinder broadly, tlie apieal joint of the middle
trochanters slightly, the greater part of the posterior, the
anterior tibia3 in front and the basal half of the four posterior
al'ove, white. Fore wings hyaline to near the stigma, the
rest fuscous ; the stigma, costa, and nervures black ; the hind
wings hyaline.
Length 9 mm.
Sikkim.
Antennae stout, densely covered with black, stiff pubes-
cence ; tapering towards the apex, thickened towards the
middle. Head slightly broader tlian the thorax; the temples
broad, rounded behind ; the front and vertex forming one
piece ; the ocellar region not bounded by furrows; there is a
deep, small, clearly defined, semicircular fovea in the middle
of the front below ; on each side, above it, is a smaller, less
distinct one, the three forming a triangle. Apex of clypeus
broadly rounded ; the pubescence on the front long, dense,
pale. Upper part of thorax almost bare, the sides and breast
bearing white pubescence. The second and following abdo-
minal segments, at the apex, are narrowly, obscurely lined
with white. The first transverse cubital nervure is faint in
the middle.
Allied to S. fuscinervis, Cam., and S. cceruleicepSy Cam.;
it may be known from both by the white clypeus and by the
Avhite band bordering the basal segment of the abdomen. It
is a broader species than either.
Chalcididae.
Oncochalcis marginiscutis, sp. n.
Black, densely covered with silvery pubescence, the apex
of all the femora, the four anterior tibiae, except the front
pair (which are marked with black broadly in the centre
behind) and the middle pair (which are similarly marked in
front and behind), the hinder broadly at the base and apex
behind (about one third), the larsi and the tegulae, bright
lemon-yellow ; wings hyaline, the nervures black. ? .
Length 6 mm.
Haundiaw Valley, Middle Tenasserim ; August [C, T.
Dingham).
►Sides of front longitudinally reticulated ; the vertex
covered with round, interlacing punctures. Face irregularly
rugosely punctured, with an aciculated space in the middle ;
the space wide in the middle, obliquely narrowed above and
below. Clypeus punctured somewhat strongly in the middle.
1G9
from the Oriental Zoological Region,
Labriim large, aciculated, depressed in the middle, almost
semicircular, the sides margined. Pro- and mesonotum and
scntellum closely, somewhat strongly punctured; tlie base of
))ronotum sligjitly projecting in the middle, the sides broadly
curved. Apex of scntellum broadly rounded, with a raised
margin, behind which is a crenulated furrow. Metanotum
coarsely reticulated, the areola longer than wide, the apical
half abru})tly widened, its apex almost transverse. In the
centre of the propleurse is a semicircular closely striated space,
proje cting into the punctured upper part, the sides irregularly
rnaiked with stout keels; at the bottom are two irregular
depressions. On the lower part of the smooth, flat base of
the metapleurse are three irregular, but distinct fovege, one
below^ the other, the upper longer than wide, ovoid, the lower
two wdder than long; the part below them coarsely, rugosely
punctured, more or less reticulated; the middle stoutly
striated, the striae distinctly separated and becoming weaker
below ; the apex projects and bears weak indications of striae.
Metapleurae closely, rugosely reticulated. Basal three seg-
ments of abdomen bare, shining; the fourth and fifth are
})unctured and covered with white pubescence at the apex ;
the sixth is large, strongly punctured and covered with white
pubescence. There are twelve teeth on the hind femora, the
apical stronger than the basal ; the apex of the hind tibiae
has a distinct projection, w'hich is curved and becomes
gradually narrowed.
Evaniidae.
Evania Binghamij sp. n.
Black, the antennal scape and the basal three joints of the
flagellum, mandibles, the fore legs, the middle except the
tarsi and the liind trochanters and femora, red; the head and
thorax closely covered with white hair, the thorax also more
densely wdth white pubescence ; the legs thickly with longer
hair, which is white on the coxae and trochanters, fuscous
on the rest. Wings hyaline, clouded in the discoidal cellule;
the costa, stigma, and nervures black, the apical nervures
pale. $ .
Length 7 mm.
Haundraw Valley, Middle Tenasserim ; August ((7. T.
Bingham) .
Face strongly striated, the striae clearly separated, con-
verging towards the centre below and with a stouter central
longitudinal keel. Sides of front strongly striated, the wider
170
Mr. P. Cameron on Tlymenoptera
central part smootli, with a stout keel down tlie middle; the
malar space strongly, obliquely striated and with some
})uncturcs. Eyes prominent, very slightly converging above,
separated from the mandibles by their own length. Temples
short, oblique ; the occiput margined, transverse. Base of
thorax transverse, the sides of pronotum oblique. Pro- and
mesonoluin coarsely, rugosely punctured ; the apex of the
latter with widely separated keels enclosing depressions.
Scutellum stoutly reticulated-punctured. Median segment
with large, round, deep reticulations; the apical slope de-
pressed in tlie middle at the apex, thickly covered with white
pubescence. Propleurse with longitudinal keels, with a large
ibvea between them at the apex ; the meso- and metapleurm
closely reticulated, the upper basal half of the former smooth,
bare, and shining, and having the top bordered by an area
with four stout keels, of which the anterior pair are closer to
each other and converge below. Metasternal process stout,
the basal part long, broad, the apical forks roundly diverging.
Abdomen shining, the petiole irregularly striated at the base,
the middle witli deep punctures. Hind coxa?, except at the
enlarged base, stoutly punctured ; the trochanters and femora
with scattered punctures ; the tibia? more strongly, closely,
rugosely punctured, their calcaria rufo-fulvous, the long spur
not reaching to the middle, more than one third of the length
of the metatarsus; the tarsal joints with rufous spines at the
apex; the metatarsus sparsely spinose. The transverse
median nervure is interstitial. The antennse are shorter and
thicker than usual, especially beyond the middle of the
flagellum. Tlie ocelli are almost in a line, the anterior hardly
projecting beyond the posterior ; these are separated from
eacli other by a distinctly greater distance than they are
from the eyes.
The legs and antennse are shorter and stouter than usual.
Braconida).
Iphiaulax domdamiensis^ sp. n.
Luteous, the antennse and ocellar region black. Wings
yellowish hyaline to the parastigrna, the yellow forming a
triangular cloud in the lower basal part of the first discoidal
cellule ; the first cubital cellule for the greater part hyaline,
slightly tinged with yellow ; the basal two thirds of the hind
wings yellowish hyaline. Basal lour segments of abdomen
as long as the head and thorax united, hardly wider than the
thorax, smooth, except the second segment, wdiich is strongly
171
from the Oriental Zoological Begion.
striated to the lateral depressions, and the third, which is
tinely closely striated to the beginning of the lateral furrows,
the strijB extending to near the middle. There is a
distinct keel down the middle of tlie second segment, with a
small, smooth, triangular plate at the base. Suturiform
articulation closely crenulated, the apical lateral branch
smooth ; there is a smooth curved furrow at the base of the
fourth segment in the middle ; there are no lateral apical
furrows. The hind tarsi are black from near the middle of
the basal joint. Temples broad, rounded, slightly narrowed ;
the occiput transverse. The ocellar black spot is rounded
behind ; in front it is gradually narrowed and reaches
close to the antennae ; there is a distinct furrow down the
front. Ocelli browm, the anterior lighter coloured than the
posterior. $ .
Length 13, terebra 9 mm.
Domdami Valley, Middle Tenasserim ; September ((7. T.
Binghani).
The apex of metanotum and the second and third abdo-
minal segments are suffused wdth fuscous. Face smooth,
with sparse black longish hairs. Tips of mandibles black.
Second cubital cellule about two and a quarter times longer
than wdde ; the recurrent nervure is received shortly behind
the cubital.
Iphiaulax liaundrawensis^ sp. n.
Luteous; occiput, vertex, upper half of front, a line
leading from this black part to each antenna, antennje, three
broad lines on the mesonotuin (the central shorter and broader
than the lateral, which extend from near the base to the
scutellum), the third and following segments of the abdomen
and laige spots on the ventral, the apical two thirds of the
hind tibite, their calcaria, and the tarsi, black. Wings
yellow'ish hyaline to near the base of the stigma, the yellow
extending beyond the lower part of the transverse basal and
the transverse median nervure and the basal half of the
hind wings ; the rest fuscous, except the usual hyaline spots
below the stigma ; the costa, except at the apex, and the
stigma to shortly beyond the middle, pale luteous. $ .
Length 11, terebra 3 mm.
Haundraw Valley, Middle Tenasserim; August ((7. T,
Bingham).
Abdomen as long as the head and thorax united and
broader than them ; longish oval ; the first segment stoutly
keeled in the middle, the sides with a few irregular stout
172
Mr. P. Cameron on Jlymenopiera
striae ; the second segment clearly wider than long, it^
cejitre stoutly longitudinally striated, without a smooth
basal area and keel ; the suturiform articulation irregularly
striated, the sides smooth and triangularly dilated ; there is a
distinct crenulated furrow on the base of the fourth and fifth
segments and a smooth one on their apices ; the pubescence
on the basal two segments is pale fulvous, on the others
black, on the apical dense ; the apices of the sixth and seventh
have a narrow pale border. Wings ample; the recurrent
nervure is interstitial ; the second cubital cellule is three
times longer than wide.
Ipliiaulax stramineuSj sp. n.
Pale yellow, the lobes of the mesonotum broadly pale
brown ; the antennte, centre of front broadly, the vertex
broadly in front, entirely behind to near the middle of the
outer orbits, and the occiput, black ; the face and orbits pale
yellow. Wings yellowish hyaline to the lower part of the
transverse basal nervure ; beyond fuscous violaceous, with
the usual hyaline spots below the base of stigma; the
fuscous cloud in the hind wings is broadest behind ; the
stigma is yellow to the commencement of the radius. The
middle of the first abdominal segment, the second entirely,
and the basal two thirds of the third are closely longitudinally
striated ; there is no area on the base of the second segment,
it being there striated like the rest of the segment. ? .
Length 13, terebra 4 mm.
Haundraw Valley, Middle Tenasserim ; August.
Abdomen elongate-oval, the basal three segments as long
as the thorax ; suturiform articulation wide, deep, crenu-
lated ; there is a narrower but distinct crenulated furrow on
the base of the fourth ; there are no apical furrows. Meta-
iiotum thickly covered with longish wliite pubescence. The
second cubital cellule fully three times longer than wide.
Tills species is related to I. spilocephaius, Cam., but is
much paler in colour, the abdomen longer and narrower,
and the second cubital cellule much longer.
Iphiaulax hurmaensisy sp. n.
Eufo-luteous, the flagellum of antenme black. The wings
yellowish hyaline, the apex with a narrow pale fuscous
border ; the base of first cubital cellule with a small black
square cloud, reaching to the base of the stigma ; the costa,
stigma, and nervures bright luteous. The raised middle of
173
from the Oriental Zoological Region*
first abdominal segment closely, strongly, longitudinally
striated, with a distinct keel down the centre ; the basal two
thirds of the second closely, somewhat strongly, irregularly,
more or less obliquely striated, and without a basal defined
area. ^ .
Length 11, terebra 9 mm.
vShwegyin, Lower Burma; November (C. T. Bingham).
This species is remarkable for the asymmetry of the furrows
on the second abdominal segment; they are not placed oppo-
site each other, that on the left side, looking from the head,
being near the middle and straight, oblique ; the other is
near the apex and is roundly curved, the part beyond it is
closely striated, that on the opposite side to it at the apex is
smooth ; beyond it is a narrow, deep, closely striated furrow,
from which the following segments become gradually
narrowed. Parapsidal furrows indicated on basal slope only ;
the apex of mesonotum flat. Temples roundly narrowed ; the
occiput slightly roundly incised. Face sparsely punctured
laterally, each puncture with a longish black hair. Pubes-
cence moderately dense, fulvous. Wings longer than the
body, the second cubital cellule as long as the third, the
transverse median nervure not quite interstitial, the re-
current nervure received near the apex of the first cubital
cellule.
Iphiaulax lineaticarinatus^ sp. n.
Luteous, the flagellum of antennas black. The wings
yellowish hyaline, the costa, stigma, and nervures luteous;
the parastigma and apex of stigma black ; a black cloud at
the base of the stigma, oblique, slightly narrowed in front,
extending to the recurrent nervure, which it covers ; a broad
lighter-coloured band on the apex, commencing near the base
of the radial and second cubital cellules, broader and deeper
tinted behind ; there is a similar apical cloud in the hind
wings. Smooth, shining; a distinct narrow keel of equal
width down the middle of the first and second segments and
a less distinct one down the third ; there is a broad depression
down the sides of the second segment, curved, narrowed and
oblique on the inner side of the base ; there is a distinctly
defined smooth furrow on the basal half of the third ; the
suturiform articulation wide, crenulated. Temples wide,
obliquely narrowed ; the occiput rounded inwardly, the sides
broadly rounded. Face covered with long black hair.
Clypeus clearly separated, forming a distinct semicircle.
First abdominal segment twice longer than it is wide' at the
174
Mr. P. Cameron on Ihjmenoptera
apex, the second longer than wide, the others wider than
long ; the transverse median nervure not quite interstitial. $ .
Length 15 mm.
Sikkim.
This species is closely related to /. hhotanensis^ Cam.; that
may be known, inter alia, by the absence of a middle keel
on the third segment, by the much weaker keel on the first,
and by the keel on the second being triangularly dilated at
the base.
Also to 1. pauperatiiSj Cam. ; tliat has the keel on the
second abdominal segment dilated at the base, and it does
not reach to the apex of tlie segment ; the space on either
side of it is obliquely striated, not perfectly smooth as in the
present species.
Ipliiaulax sikkimensisj sp. n.
Luteous, the abdomen darker coloured, the antennae black.
The wings yellowish hyaline to near the top of the transverse
basal nervure, dark fuscous beyond, the base of the discoidal
cellules being yellowish ; the basal half of the stigma yellow ;
the area on base of second abdominal segment narrow, longish,
gradually narrowed, running into the keel, stoutly longitu-
dinally striated. Abdomen longish oval, broader than the
thorax; the basal three segments stoutly longitudinally
striated, the striae strongest on the first. Temples broad,
roundly narrowed, tbe occiput transverse. ? .
Length 12, terebra 4 mm.
Sikkim.
Face sparsely punctured, covered with long bright fulvous
hair. Front and vertex smooth, sparsely haired. Basal
half of mandibles luteous, the apical black. The depression
on the base of the second segment is closely obliquely
striated; the striae on the middle of the segment are strong
and intertwine. The basal two abscissae united are as long
as the third ; the second cubital cellule is fully three times
longer than wide along the cubitus.
Comes near to 7. spilopteriiSj Cam.
Iphiaulax Hooker sp. n.
Luteous, the head paler, more yellowish in tint ; a large
broad mark on the front, extending backwards to shortly
behind the ocelli, where it is narrowed and rounded ; the
third and following segments of the abdomen, tbe bind tibiae
from near the base, the hind tarsi, the hind spurs, and the
antennae black. Wings yellowish hyaline to tbe top of the
175
from the Oriental Zoological Region.
transverse basal nervure, blackish fuscous beyond ; the stigma
black. Abdomen smooth ; the apex of first segment finely
striated in the middle, the second much more strongly and
irregularly striated ; the keel large, the basal half roundly
narrowed, smooth, the apex gradually narrowed to a point,
followed by a keel nearly as long as the dilated basal part,
and which does not quite reach to the apex of the segment;
it is bordered in the middle by five or six stout oblique
strise, the parts surrounding the base and apex being
smooth. Suturiform articulation deep, closely stoutly crenu-
lated ; there is a curved lateral branch, which is not quite
so strongly crenulated ; the sides of the segment irregularly
roughened. $ .
Length of body and ovipositor 13 mm.
Sikkim.
Head large, cubital ; the temples longer than the eyes,
slightly rounded, not narrowed ; the occiput transverse.
Face distinctly sparsely punctured ; the clypeus is separated
from it by a distinct curved furrow, laterally by a depression
in which is a round distinct fovea. Mandibles black, except
at the base. Palpi testaceous, covered with white hair.
Antennal scape slightly more than twice longer than wide,
slightly dilated at the apex below. Recurrent nervure inter-
stitial. First abdominal segment clearly longer than the
second, about one fourth longer than it is wide at the apex.
Malar space about one fourth shorter than the length of the
eyes.
Iphiaulax Camphelli^ sp. n.
Black ; the head, the prothorax, the mesonotum laterally
at the base, a line bordering the middle lobe, the centre
broadly from behind the middle to the apex, and a large
oblique mark below the fore wings testaceous ; a mark
bordering the sides of the scutellum and a broad line on the
lower part of the mesopleuise rufous ; the first abdominal
segment, except for a mark, longer than broad, in the centre
at the apex and the ventral surface, pale testaceous. Legs
black ; the anterior apical joint of middle trochanters, the
apical half of middle femora, a streak in their middle above,
and the base of middle tibiae testaceous. Wings almost
hyaline; the stigma black, testaceous in front and at the base
behind, the costa and nervures black ; the recurrent nervure
received shortly before the first transverse cubital. $ .
Length 8, terebra 3 mm.
Sikkim.
17G
Mr. P. Cameron on Hymenoptera
Abdomen longisli ovate, in tlie middle clearly wider than the
thorax, which is equal to it in length ; its first segment a little
longer than it is wide at the apex ; the central part gradually
roundly narrowed from the apex to the base, longer than
wide ; the apical half with a central keel, widest at the base;
from it stout strise radiate towards the apex. The central
part of the second segment is irregularly longitudinally
reticulated-striated, the sides finely rugose, intermixed with
strife and punctures ; the suturiform articulation deep, closely
crenulated, without an apical lateral branch ; tlie other
segments opaque, alutaceous. The sixth and apical segments
are whitish testaceous. Antennal scape fully three times
longer than wide. The base of the cubitus is straight and
runs parallel with the costa before curving down towards the
first transverse cubital nervure.
Iphiaidaxl tenasserirmnsis, sp. n.
Luteous, the flagellum of the antennae black. The wings
yellowish hyaline, the nervures and stigma luteous ; the apex
of the fore wing from the radius with a narrow smoky border,
the hind wings with the apical third smoky ; there is a
square blackish cloud between the transverse basal nervure
and the base of the stigma, the costa at its apex being also
black. The first abdominal segment, except the basal slope,
the second entirely, and the third to near the middle, closely,
finely, longitudinally striated. ? .
Length 13, terebra 7 mm.
Domdami Valley, Middle Tenasserim ; October {Bingham).
Smooth, except on the basal abdominal segments, densely
covered with fulvous pubescence, except on the face, where
it is long, sparse, and blackish. Abdomen nearly twice the
length of the thorax, not dilated in the middle, as wide as the
thorax ; the suturiform articulation wide, deep, striated ; the
other furrows narrow, smooth, indistinct. The third abscissa
of radius is nearly as long as the basal two united. Temples
broad, roundly narrowed ; the occiput transverse. Meso-
notum flat behind, the furrows distinct in front. There is no .
depression on the sides of the second segment; there is a
short, broad, curved furrow on the sides of the third.
Ichneumonidae.
Tryphoninje.
Ciiltrarius purpureotinctas^ sp. n.
Black, the apical three segments of the abdomen purple ;
the edges of the facial shield (the top more narrowly), the
177
from the Oriental Zoological Region.
upper eye-orbits below, and tlie underside of the antennal
scape yellow ; the flagellum of antennae brownish below; the
underside of the four anterior tibiae and tarsi, the hind
femora, a mark on the sides of the first abdominal segment,
a broader continuous band on the apices of the second and
third, and one on the lateral fourth of the fourth dark red ;
the genital armature large, rufo-testaceous. Wings hyaline,
very iridescent, the radial cellule and the apical half of the
third cellule smoky. .
Length 14 mm.
Takvar, Darjiling, 4000 feet; April {G. T. Bingham),
Head somewhat strongly but not very closely punctured ;
the lower part of the face, outside the shield, and the clypeus
fully more strongly and more closely punctured ; the occiput
and temples smooth, densely covered with white pubescence.
Pro- and mesothorax strongly closely punctured, the pleurae
more strongly than the top; thescutellum still more strongly
punctured, its basal depression wide, deep, with five stout
keels ; the end of the apical slope striated. Postscutellum
densely covered with long white hair. Areola 6-angled,
clearly longer than it is wide at the base, the apical half
narrowed, the apex half the width of the base : it has a
raised smooth line in the centre ; the rest irregularly wrinkled ;
there is a large lateral area, wider than long, its apex broadly
rounded, the centre with some large punctures, the apex
with a few striae, the apical slope closely rugosely punc-
tured. Metapleurae more strongly punctured than the rest,
the punctures larger and more widely separated than on the
inesopleurae. Abdomen closely strongly punctured ; the first
segment raised in the middle, the raised part bordered by
keels, which are higher and rounded at the base and depressed
in the middle ; the second and fifth segments are indistinctly
keeled in the middle, the third and fourth have a more
distinctly defined central keel which projects on the apex of
the third; the apical two segments are densely covered with
short black hair. Mesopleural furrow wide, deep. Areolet
large, the nervures not uniting in front ; the recurrent nervure
received in the middle. Scutellum uniformly punctured;
the lateral teeth large, triangular. Fore calcaria testaceous,
the others white.
Cultrarius areolatiiSj sp. n.
Length 10 mm. .
Sikkim.
This species resembles closely C. purpnreotmetus in colora-
Ann. ^ Mag, N. Hist. Ser. 7. Vol. xix. , 12
178 Mr. P. Cameron on ITymenoptera
tion ; tlie differences between them may be expressed
thus : —
Areola clearly longer than wide, G-angled, the apical
half obliquely narrowed, the apex transverse, the
lateral area strongly punctured ; scutellum with-
out a deep furrow in the middle ; the keels on the
centre of the first abdominal st'gment not uniting
at the apex, straight ; arc olet not appendiculated ;
the hind femora rufous piirpureotincfus.
Areola as long as wide, not angled, its apex broadly
rounded, the lateral area smooth ; scutellum with
a deep furrow in the middle ; the keels on first
abdominal segment roundly curved and uniting .
at the apex ; the areolet shortly appendiculated ;
the hind femora black areolatus.
The coloration is the same otherwise, except that the
purple tint on the abdomen commences on the second segment,
that the scutellar keels are yellow, and that the lines on the
second to fifth abdominal segments are narrower, of more
uniform thickness, and more yellowish in colour. The
puncturation is pretty much the same ; on the second seg-
ment of the abdomen it is stronger and more irregular, that
segment, too, being shorter compared with the width ; the
keels on the second to fourth segments are narrow, but
distinct. The calcaria are all whitish. The abdomen is
shorter, too, compared with the thorax.
Opuionin^.
Eyiiscospilus xanthocephalus^ sp. n.
Luteous ; the head pale yellow. The wings hyaline, the
nervures black, the stigma testaceous : there are two horny
points, the basal large, broad and tiansverse at the base; the
base above narrow^, transverse ; from there it becomes
gradually roundly narrowed to a fine point at the apex ; the
second point is close to the middle of the basal abscissa of
the radius ; it is longish, narrow, roundly curved towards the
radius. Scutellum distinctly keeled laterally from the base
to the apex ; the basal half smooth, the apical finely, closely,
longitudinally striated. Base of metanotuin smooth ; the
depression with some stout stride ; the rest is stoutly striated,
the striae clearly separated ; in the centre of the base is a
short straight stria ; the striae at the sides of this are longi-
tudinal and straight at the base, then curving out obliquely
to the sides; the striae on the apical slope are roundly curved
from side to side. The centre of propleurae finely obliquely
striated ; the lower half of the mesopleurae longitudinally
179
from the Oriental Zoological Region.
striated, tlie strlje weaker above and all clearly separated.
Base of radius broadly curved downwards; the discocubital
nervure broadly roundly curved, but not much, at the apex,
its apex widely distant from the commencement of the apical
abscissa of the radius. $.
May be known from E. reticulatus^ Cam., and E. striatus,
Cam., from the Khasias, by the wings having two horny
points in addition to the other differences.
Length 27 mm.
Haundraw Valley, Middle Tenasserim ; August (0. T.
Bingham) .
Cryftinj!.
Melcha ornatipennis^ sp. n.
Black ; the scutellum, median segment entirely, the apex
of mesopleurte, base of first abdominal segment to shortly
beyond the middle, all the femora, and the four hinder coxae
red ; the four anterior tibiae and the fore tarsi of a paler red,
the middle tarsi fuscous, the anterior coxae and trochanters
black ; the hind tibiae and the basal and apical joints of the
tarsi black, the middle joints white ; the basal half of pro-
notum, apex of postpetiole broadly, the apex of the second
segment slightly more narrowly and more irregularly, the
apex of the sixth and the apical entirely, white. Wings
hyaline, a cloud in the fore wings extending from the base
of the stigma to the areolet and the recurrent nervure ; the
stigma and nervures black. ^ .
Length 9 mm.
Salween Valley, Middle Tenasserim ; July (C. T.
Bingham).
Thorax and head thickly covered with white pubescence.
Base of metanotum smooth ; there are two short keels in the
middle at the transverse keel ; the rest of the metanotum
rather strongly reticulated, almost smooth in the middle at
the base ; the metapleurge strongly but more closely reticulated.
Propleurge at the apex from shortly above the middle covered
with close, stout, curved, longitudinal strise. Areolet not
quite square, being slightly longer along the radius than
along the transverse cubital nervures ; it is of equal widtli.
Allied to M. maculipennis , Cam., from Borneo ; that
species has the femora black and the scutellum white.
Silsila spilonota, sp. n.
Black ; the face, clypeus, a line on the inner orbits, one on
the lower third of the outer, the malar space, mandibles
12*
180
Mr. P. Cameron on IJymenoptera
(except the teeth), palpi, the dilated base of the pronotum,
tegulse, an elongated conical mark at the apex of the midlle
lobe of the mesonotum, the broad end at the base and trans-
verse, scutellums, scutellar keels, a large curved mark behind
the liind wings, the apical slope of the metanotum (the mark
continued backwards in the middle, this line becoming
gradually narrowed), the apex in the centre with a black
line, the lower edge of the propleurse, the line narrowed in
front, tubercles, a large mark on the lower part of the meso-
pleurae, the top broadly incised, the apex above smaller than
the base, a curved mark at the apical half of the sternal
furrow, a large elongated conical mark on the metapleura3
and broad bands on the apices of the abdominal segments,
yellow. Four front legs pale yellow, infuscated above; hind
coxse black, broadly yellow in the middle above, the tro-
chanters black ; the femora fulvous, their base narrowly and
the apex from near the middle black ; the tibise yellow,
tinged with fulvous, the base narrowly and the apical fourth
black; the tarsi white, the apex of last joint black. Wings
hyaline, the stigma and nervures black. Antennae broadly
ringed with white. .
Length 13-14 mm.
Haundraw Valley, Middle Tenasserim [C. T. Bingham').
Face strongly but not closely punctured; the clypeus with
a few punctures above. Front and vertex smooth, a short
keel and a few striae on the front below. Mesonotum
smooth, the inner half of the outer lobe irregularly reticulated
at the apex. The space at the sides of the scutellums irregu-
larly stoutly striated. Basal depression of metanotum
stoutly crenulated ; the base at the sides stoutly irregularly
punctured ; the rest stoutly transversely reticulated. Pro-
and mesopleurae stoutly longitudinally striated, the strias
roundly curved, the lower part of the latter strongly punc-
tured. Metapleuraj strongly punctured above, the punctures
distinctly separated, the puncturation below coarser, the
])unctures running into reticulations. Pro- and mesonotum
sparsely haired, the metanotum much more densely pilose
and with the hair longer.
May be known from the two described Himalayan species
of ISilsila (fulvipes^ Cam., and hilineata, Cam.) by the large
conical mark on the mesonotum.
Buodias rugifrons, sp. n.
Black ; the face, clypeus, basal half of mandibles, palpi, a
181
from the Oriental Zoological Region,
outer, a line on the base of propleurae extending broadly on
to the pronotuni, a line on the sides of pronotum, a small
conical spot on the apex of the middle lobe of pronotum,
scutellum, base of metanotum (the lines narrowed on the
inner, dilated on the outer side), a square mark in the centre
behind the keel (the middle part narrowed at the base, the
line becoming gradually dilated to a triangle), the apical slope,
a spot below the hind wings, and the apical three fourths of
the metapleuras, the apex of the second abdominal segment,
the apical and the ventral segments, pale yellow. Four
front legs pale yellow, the femora slightly tinged with
fulvous ; the hind legs rufo-fulvous ; the trochanters, apical
half of femora, and the tibise from near the middle black ;
the tarsi white, narrowly black at the base. Antennae
broadly ringed with white. Wings hyaline, the nervures
and stigma black. ^ .
Length 10 mm.
Sikkim.
Face closely and strongly punctured, the clypeus smooth.
Front closely rugosely reticulated, keeled down the middle,
the lower part with a smooth transverse line, clearly separating
the rough part ; vertex closely punctured. Mesonotum closely
distinctlj?- punctured, except at the sides ; the scutellum
smooth. Base of metanotum with scattered punctures ;
inside the spiracles is a keel reaching from the base to the
apex; there is a keel on the apical half of the white central
mark ; the rest is stoutly longitudinally reticulated, mixed
with some punctures; the lateral teeth are small. Areolet of
almost equal width, longer along the radius than along the
transverse cubital nervures, the recurrent nervure received
near the middle.
This species is smaller than the others ; the areolet is
larger and the metanotal spines smaller than usual.
Cry plus Binghami, sp. n.
Black ; a narrow line on the inner orbits, a similar one on
the lower two thirds of the outer, and the tubercles yellowish
testaceous ; the legs red ; the coxae, trochanters, and the
underside of the four anterior femora to shortly beyond the
middle, and a short line on the apex of posterior femora,
black ; the apical joints of the four anterior tarsi infuscateci,
joints 2-4 of the posterior yellow. Wings hyaline, the
nervures and stigma black. $ .
Length 13, terebra 4 mm.
Darjeeling, 5000 feet ((7. T. Bingham),
182
Mr. P. Cameron on llymenoptera
Postscutellum smootli, the sides aeiculated-striated. Pro-
pleurae rugose above, the rest stoutly obliquely striated^ tlie
striae intermixing more or less; the meso- and meta-
pleuise closely coarsely reticulated-punctured. Tibiae sparsely,
the tarsi closely spinose. Palpi and mandibles black, the
former covered with white pubescence. Head, thorax, and
legs densely covered with white pubescence. The front is
deeply depressed from the eyes and is strongly, closely,
transversely striated.
This species is not unlike C. lucuJentus, Cam. ; the two
may be separated thus : —
Apical keel on metanotum interrupted in the middle ;
four anterior femora not lined with black ; the
three middle joints of hind tarsi not yellow .... luculentus, Cam.
Apical keel on metanotum complete ; the four ante-
rior femora broadly lined with black ; the three
middle joints of hind tarsi yellow Binghami.
C. Binghami is a more slenderly built species than lucu-
Jentus ] the front is more deeply depressed and is more
distinctly transversely striated ; the scutellum is more promi-
nent and has the sculpture rougher and more irregular.
The male of Binghami has the inner orbits more broadly
yellow than in the female ; it differs further in the face
having in the middle of the top above a yellow mark, longer
than wide and narrowly projecting at the sides above ; the
clypeus and mandibles are for the greater part yellow, as are
also the greater part of the anterior coxae and trochanters ;
the middle coxae and trochanters are yellow on the outer side,
C, luculentus has been taken by Col. Bingham at Darjiling,
. 7000 feet, in March.
IcRNE UMONINJE.
ICHNEUMONINI.
Lagarista, gen. nov.
Scutellum not much raised, quadrate, stoutly keeled
laterally to the top of the apical slope. Metanotum regularly
areolated ; the areola about two and a half times longer than
wide, rounded and slightly narrowed at the base, rounded
inwardly at the apex ; the lateral basal areas confluent, the
petiolar area absent. Antennae distinctly dilated towards the
apex. Apex of clypeus broad, transverse. Apical tooth of
mandibles long, narrowed gradually towards the apex ; the
Bubapical tooth short, turned inwardly. Base of petiole
broader at the base than it is thick dorso-ventrally — broader
183
from the Oriental Zoological Regioa.
tlian in Ichneumon^ but not so broad as in Plati/lahls. There
aie seven abdominal segments ; the last broad at the apex,
two thirds of the length of the penultimate ; the ventral fold
extends to the apex ot the fourth segment. Arcolet 5-angled ;
the discocubjtai nervure unbroken ; the transverse median
nervure interstitial. Tarsi closely spinose. The ovipositor is
long, distinctly projecting, as long as the apical two segments
united.
The head is as wide as the thorax ; it is large, but not
behind, the temples being short, the occiput rounded in-
wardly, the cheeks sharply margined. Labrum hidden. The
antenna3 are longer than the body, slender, except at the
dilated apical part ; the basal joints of the flagellum elongate,
the first slightly longer than the second. Postpetiole striated.
The systematic position of this genus is not clear. The
form of the metanotum is as in the Iclmeumonini, and it
cannot therefore be referred to the Joppini. For the present
I refer it to the former tribe. It is clearly related to the
Ceylonese Aluinaj Cam. ; that genus may be known by the
areola being much wider compared with the length, by the
much wider temples, and by the discocubital nervure being
broken by a stump. The form of the metanotal areola
separates the Ceylonese Deniya from it; that genus may be
further known by the almost obsolete temples, transverse
occiput, and by the apex of the clypeus being rounded.
Lagarista mucaUscatis^ sp. n.
Black ; the head except the centre of the vertex (covering
the ocelli), the front, and the occiput except the outer edges,
a line on the top and bottom of prothorax, two narrow longish
lines in the middle of the mesonotum, scutellar keels, scutel-
lum except for a conspicuous black conical mark in the centre,
postscutellum, an irregular mark in the centre of the lateral
basal arese of the metanotum, the apical lateral areas, the
yellow extending slightly below them, the greater part of the
lower half of the inesopleurEe, a large spot behind the hind
wings, the metapleurae from behind the middle, and the apices
of the abdominal segments (the lines on the basal three
broad and dilated laterally — that on the fifth narrow and
interrupted in the middle, that on the sixth in the centre
only, and the last entirely), pale yellow. Legs fulvous; the
four anterior coxae and trochanters pale yellow, the hinder
femora more reddish in tint ; the hind coxae yellow, fulvous on
the outer side and blackish at the apex above; the apex of
the hind femora, of the hind tibiae more broadly, and of the
184
Mr. P. Cameron on Ihjmc)\oj)lera
basal half of the metatarsus black ; the hind tarsi white.
Wings hyaline, the stigma testaceous, the nervures blackish.
Flagellum of antennae broadly ringed with white. ? .
]jength 11 mm.
Sikkim.
Face distinctly but not closely punctured, the clypeus with
a few scattered punctures above; front below the ocelli finely
closely striated. Mesonotum closely, regularly, somewhat
strongly punctured ; the scutellum with a few large punctures
in the centre. Posterior median area furrowed round the
edges except at the apex ; it is smooth, as are also the parts
at its base; the rest of the segment closely strongly punc-
tured except the posterior median area, which is strongly
irregularly striated. Meso- and metapleurae closely strongly
punctured ; the propleurae smooth, striated at the apex.
Abdomen closely punctured, the centre of postscutellum finely
striated.
J 0 P P I N I.
LissOPHADNUS, gen. nov.
Scutellum rather flat, distinctly keeled laterally to the
commencement of the apical slope. Areola represented by a
small, smooth, flat, curved tubercle, widely distant from the
base of the metanotum. Posterior median area with the apical
half narrower than the basal ; the apical lateral area com-
mences at the end of the basal part, it is narrow, sharply
pointed at the top, becoming gradually widened towards the
apex ; the basal lateral arese are large, square, confluent,
there being no petiolar area; the apical large, wide at the
base, gradually narrowed towards the apex ; the spiracular
area distinct. Clypeus not separated, its apex transverse.
Labrum projecting. Occiput deeply incised, margined.
Areolet large, 5 -angled ; transverse median nervure received
distinctly beyond the transverse basal. Petiole long, narrow,
the postpetiole gradually widened. Mandibles unequally
dentate.
Body uniformly rufo-testaceous; the wings hyaline. 'Disco-
cubital nervure broken by a stump, as is also the recurrent
nervure. The second and following segments of the abdomen
closely punctured, the first smooth and shining.
The form of the metanotum and its areola is as in Dimcetha^
which genus may be known from the present by the scutel-
lum not being flat and not keeled, by the areolet being
narrowed in front, 4-angled, by the discocubital and recurrent
nervures being unbroken, by the wings being clouded at the
apex, and by the postpetiole being strongly punctured.
185
from the Oriental Zoological Region.
Lissophadnus testaceusj sp. n.
Testaceous, tending to rufous-ferruginous ; the antennae
beyond the sixteenth joint black, the middle joints tinged
with yellow. • Wings clear hyaline, highly iridescent, slightly
violaceous towards the apex. $ .
Length 22 mm.
Runjit Valley, 1500 feet, Sikkim; April [G. T. Bing-
ham) .
Base of metanotum at the sides sparsely but strongly
punctured ; the lateral middle area rugosely punctured at
the base, transversely reticulated-punctured at the apex ; the
posterior median area irregularly transversely striated ; more
or less reticulated at the apex ; the lateral apical arese are
coarsely reticulated ; the segment is thickly covered with
pale pubescence. Pieurse closely punctured ; the lower apical
part of propleurae striated. Clypeus somewhat strongly but
not closely punctured in the middle ; the sides and face
smooth, almost bare. Labrum fringed with long golden
hair. Front depressed; the inner orbits margined. Tips of
mandibles black. Tarsi closely spinose.
Acanthojopp)a flavo-orhitalisj sp. n.
Luteous; the sides of the face, clypeus broadly, the orbits
broadly, and the pleurae tinged with yellow : the middle of
antenna? tinged with yellow ; tlie eighteenth and following
joints black. Wings hyaline, the basal half tinged with
fulvous ; the stigma and costa rufo-testaceous, the nervures
black. $ .
Length 14 mm.
Domdami Valley; October ((7. T. Bingham) ,
Face strongly closely punctured, the upper part of the
clypeus as strongly but not so closely punctured. Front
and vertex smooth, the upper part of the front raised,
surrounded at the sides and below by a smooth, bare, shining
depression. Mesonotum closely finely punctured, depressed
on either side laterally at the base ; the scutellum more
strongly punctured and thickly covered with long pale fuscous
pubescence; the apical incision broad, rounded, not deep.
Base of metanotum irregularly punctured ; the areola round
the sides and base reticulated, the apex in the middle closely,
strongly, longitudinally striated; the second lateral arese
closely reticulated ; the apical slope strongly transversely
striated. Propleurse on the upper half perpendicularly
striated, below irregularly, more strongly reticulated-striated.
18G
Mr. P. Cameron on ITi/menoptera
]\Icso])leura3 closely, strongly, irregularly punctured, more or
less striated in the middle and at the apex belov\\ Meta-
pleurae coarsely rugosely punctured, more or less coarsely
striated below. Petiole smooth ; the second segment is
more strongly punctured than the others. The transverse
cubital nervures meet in front ; the recurrent nervure is
received shortly beyond the middle.
The areola is large, longer than wide, rounded at the base,
transverse at the apex, the sides bulging out where the trans-
verse keel joins them ; the pubescence on the apical slope
long, dense, fuscous; the spines broad, rounded; its centre
raised.
Comes near to A. tinctipennis, Cam.
Xantliojoppa latihalteata^ sp. n.
Pale yellow ; a line covering the ocelli and extended down
to near the antennm, three broad lines on the rnesonotum (a
transverse one uniting them at the apex), the scutellar de-
pression, a line round the base of the metanotum, the greater
part o£ the areola, the posterior median area, a broad band
across the pronotum, a line on the top of the mesopleurge, one
down the base of the metapleurse, one behind the hind coxae,
a mark (longer than wide) commencing near the middle and
extending on to the postpetiole, a band on the second segment
extending from the base to shortly beyond the middle, one on
the basal half of the third with the apex transverse, and semi-
circular ones on the basal half of the fourth, fifth, and sixth,
the underside of the hind coxae except at tlie base, the mark
continued above obliquely to the upper inner edge, the hind
femora except below, the apical two thirds of the hind tibiae,
and the hind tarsi, black. Wings hyaline, iridescent, the
stigma testaceous, the nervures black. ,
Length 12 mm.
Domdanu Valley, Middle Tenasserim ; October ((7. T,
Bingham),
Areola retieulated ; the upper third of the posterior median
area longitudinally, the rest transversely, reticulated, the
rest of the segment closely rugosely punctured, the apical
slope thickly covered with fuscous pubescence. Pro- and
mesothorax distinctly punctured, closely on the rnesonotum ;
the scutellum more strongly punctured and tliickly covered
with longish fuscous pubescence. Scape and pedicle of an-
tennae rufo-testaceous, the rest black, the basal joints testaceous
below. Petiole smooth ; the second and third segments are
closely distinctly punctured ; the gastrocoeli small, transverse,
187
from the Oriental Zoological Region,
pmootli, testaceous ; behind them are two long keels, with a
shorter keel on the outer and inner side. Areolet 4-angled,
distinctly narrowed in front ; the recurrent nervure is received
shortly beyond the middle ; the discocubital nervure unbroken,
the transverse median received beyond the transverse basal.
Allied to X. 2>-lineata^ Cam., which may be known by the
absence of black on the hind legs, by the black line on occiput,
and by the narrower abdominal black bands.
Amhlyjoppa Binghami, sp. n.
Black ; the basal three segments of abdomen red ; the
face (except for an anchor-shaped black mark down its
centre), the clypeus (except for a small conical mark on its
apex in the middle), the upper inner orbits (the line not
reaching to the ocelli), a line on the lower two thirds of the
outer (the line becoming gradually wider below, extending
obliquely to the outer side of the mandibles), a broad line on
the pronotum, tegulse, tubercles, a squarish mark on the lower
side of the mesopleurae extending on to the sternum, two
lines in the centre of mesonotum, scutellar keels, scutellum
from shortly behind the middle, postscutellum, a mark
almost filling the apical lateral arege of metanotum, and the
broad apical part of the sides of postpetiole, yellow. Legs
yellow, densely covered with white pubescence; the hind
femora red ; the outer apical half of the fore femora, the
middle femora (except the apical half below), the hind coxa3
(except for a large mark above), the base and apex of hind
femora narrowly, the apex of the hind tibiae more broadly,
and the hind tarsi, black. Wings hyaline to the base of
stigma, smoky violaceous beyond, the nervures and stigma
black. Antennal scape yellow below, the flagellum witli a
narrow yellowish band. ? .
Length 20 mm.
Sikkim, Runjit Valley, 1000 feet ; April T. Bingham).
Head and thorax closely strongly punctured and thickly
covered with white pubescence. The areola small, roundly
narrowed to a point behind ; the point dilated, the bordering
keels smooth, shining, the apical narrower, less distinct,
broadly rounded inwardly ; the inner sides furrowed, the
centre shining, irregularl}" wrinkled. The basal arege acicu-
lated at the base, the rest irregularly punctured ; the other
area3 strongly punctured, the posterior median stoutly trans-
versely striated. Scutellum slightly roundly raised, the sides
keeled to the middle. Pleurae closely punctured, the lower
half of propleurse irregularly striated. Petiole keeled laterally.
188
Mr. P. Cameron on Ilymenoptera
and with a less distinct keel down the middle ; the post-
])etiule closely strongly striated in the middle, the sides
almost smooth ; the second to sixth segments closely punc-
tured, the puncturation becoming gradually weaker ; the
second and third striated down the middle. Gastrocoeli deep,
smooth.
This species is not unlike A. rufocincta, Cam., in colora-
tion ; it differs in the scutellum being longer compared with
the width, more raised at the base, and with the lateral keels
longer and more distinct, and the areola is smaller and more
narrowed — more sharp-pointed — at the base.
Amhlyjoppa iridipennis, sp. n.
Dark rufo-testaceous ; the apical two thirds of the meso-
pleura3 above, the apical half below, the metapleurae, rneta-
notum, the basal segment of abdomen, and the second to
shortly beyond the middle, the mesosternum (except for a
triangular space on either side at the base), the hind coxae,
and the base of the trochanters above, black. The basal
sixteen joints of antennae yellow, tinged with red, the rest
black. Wings uniformly fuscous violaceous, the nervures
and stigma black. $ .
Length 18 mm.
Sikkim.
Head and thorax somewhat strongly closely punctured ;
the eye-orbits are tinged with yellow, the yellow tinge
broader on the lower outer orbits. The apex of mesonotum
lighter-coloured in the middle. Base of metanotum strongly
punctured (except for a smooth space bordering the areola) ;
the latter is gradually narrowed from the apex to the base,
which is rounded, narrow; the apex is rounded inwardly,
the keel narrow, irregular; the outer edge of the inner side
is irregularly narrowly depressed ; in the centre is an irregular
depression, and there are two smaller ones at the apex; the
apical slope is irregularly strongly reticulated, the posterior
median area more strongly than the lateral. Abdomen
closely punctured, strongly at the base, weaker towards the
apex ; the second segment depressed at the base, between the
gastrocoeli, closely striated ; the latter has four stout clearly
separated keels on the inner half and one near the outer
side.
Amhlyjoppa ? maculicolliSj sp. n.
Black ; the face, clypeus, mandibles, and the orbits nar-
rowly, yellow; the front and vertex yellow, suffused with
189
from tlte Oriental Zoological Region,
rufous ; the occiput and temples rufous, the mesonotum an I
scutellum of a deeper rufous colour, tlie scutellum darker
coloured ; a yellowisli-rufous line, dilated behind, on the
pronotum ; the anterior legs reddish testaceous; the middle
tibise in front testaceous, the middle femora fuscous in front.
Wings fuscous violaceous, the nervures and stigma black ;
the areolet 4-angled, the nervures meeting in front. Areola
moderately large, horseshoe-shaped ; the apex rounded
inwardly. ^ .
Length 15 mm.
Middle Tenasserim, Salween Valley ; July {C. T, Bing^
ham) .
Basal half of areola smooth, bordered laterally by a deep
furrow; the apical half coarsely rugosely punctured. Head
closely punctured, the apex of clypeus with only a few"
scattered punctures. Front furrowed down the centre, the
furrow narrowed tow^ards the apex. Thorax closely distinctly
punctured, the scutellum more strongly than the mesonotum
and thickly covered with blackish-fuscous hair. The rneta-
notum at the base strongly punctured, the punctures distinctly
separated ; on the rest the puncturation is closer, more
rugose, and more or less running into reticulations, on the
posterior median area transversely reticulated. The meso-
notum is densely covered with short fulvous pubescence ; on
the median segment the pubescence is longer and black, on
it also the black is suffused with brown. Abdomen strongly
closely punctured, the puncturation becoming weaker towards
the apex; gastrocoeli strongly regularly striated, the space
between them being also striated. The malar space is
black ; a black spot connects the ocelli to the eyes.
This species differs from the typical form in the transverse
cubital nervures being united in front. This may not,
however, be a constant difference.
Amhlyjoppa rujicepsy sp. n.
Black ; the head reddish, the face and clypeus more
yellowish in tint ; a squarish black mark joins the ocelli to
the eyes ; the fore legs testaceous, darker above ; the antennal
scape reddish testaceous, the flagellum brownish below at the
base. Wings uniformly fuscous violaceous, the nervures and
stigma black. J .
Length 14 mm.
Middle Tenasserim, Salween Valley; July {C. T. Bing,
ham).
Areola longer than wide, transverse at the base, becoming
190
]\rr. r. Cameron on Ilymenoj^tera
gradually obliquely widened to beyond the middle ; the sides
at the apex not quite straight ; the apex not clearly margined,
broadly rounded inwardly; the segment is closely rugosely
j)unc(ured, more closely rugosely on the apical slope ; there
is a longish narrow triangular area bordering the apical half
of the posterior median ; it is coarsely transversely striated.
The pro- and mesothorax are closely strongly punctured, the
inetapleurae more strongly than the rest ; below the middle
is a stout curved keel. The scutellum is more strongly but
not so closely punctured as the mesonotum and is thickly
covered with black pubescence. Abdomen closely punc-
tured, the ])ostpetiole more coarsely than the rest, the apical
segments more weakly. Gastrocoeli deep, bearing stout
curved strige. Tegul^e testaceous. Tubercles dark rufous,
d’he black on the upper part of the body is slightly tinged
with brown, especially on the metanotum ; the middle coxa3
are fuscous. There is a short but distinct stump on the
discocubital nervure.
Apart from the difference in coloration and in the form of
the areola, this species differs from A. macuUceps in having a
stout curved keel on the lower part of the metapleurge.
Amhlyjoppa maculiceps^ sp. n.
Black ; the face, clypeus, labium, and base of mandibles
yellow ; the front, vertex, outer orbits, and occiput yellow,
largely suffused with rufous ; a line on the pronotum, dilated
at the apex, red, tinged with yellow ; the mesonotum (except
at the sides and base) narrowly rufous; the fore legs (except
the coxte, trochanters, and base of femora) black ; the middle
tibiae tinged with testaceous at the base and in front. Wings
uniformly dark fuscous violaceous, the nervures and stigma
black. Areola semicircular. Antennal scape yellow, the
flagellum brownish below. A •
Length 17 mm.
Middle Tenasserim, Salween Valley; July ((7. T. Bing-
ham) .
Face and clypeus strongly, closely, regularly punctured ;
the labrurn as closely but not so strongly punctured ; the
vertex distinctly but not so closely or so strongly punctured
as the face ; a smooth space at the sides of the ocelli, which
are joined to the eyes by a broad black spot ; there is a deep
clearly defined furrow in the centre of the front, bordered by
a raised distinctly punctured part ; it extends to the middle ;
the part below is very smooth and shining, bare ; the sides
are smooth excc})t near the eyes, where they are punctured.
191
from the Oriental Zoological Region.
Pro- and mesothorax distinctly regularly punctured; the
Kcutellum is more strongly punctured and is thickly covered
with black pubescence. The metanotum is much more
strongly and closely, almost rugosely punctured ; the areola
wider than long, broadly rounded at the base, semicircular ;
the apex irregular and slightly turned inwardly ; inside it is
irregularly rugosely punctured, depressed on either side at the
base, and with a smooth transverse space at the base ; the
segment is tliickly covered wdth black pubescence. The
basal four segments of the abdomen are closely punctured,
the puncturation becoming gradually weaker; the base of
the ])etiole is smooth in the middle; the postpetiole closely
rugosely punctured.
The antennae are as long as the abdomen, serrate, distinctly
tapering towards the apex.
Amhlgjopiia rnficaada^ sp. n.
Black ; the head, prothorax, mesonotum, the fourth and
following segments of the abdomen, the anterior legs, the
apex of the middle femora broadly, of the hinder more
narrowly, and the four posterior tibiae and tarsi, rufo-testa-
ceous ; the nineteen to twenty basal joints of antennas reddish
yellow, the rest black. Wings uniformly dark fuscous
violaceous, the nervures and stigma black. ^ .
Length 14-19 mm.
Salween Valley, Middle Tenasserim; July {C. T. Bing-
ham).
Face and clypeus closely, somewhat strongly and uniformly
punctured ; the front and vertex are not so closely, regularly,
nor so strongly punctured. Pro- and mesothorax closely
distinctly punctured ; the mesonotum densely covered with
fulvous pubescence ; the scutellum and metanotum (except
at the base) densely covered with longer black pubescence.
The metanotum is more strongly rugosely punctured ; the
posterior median area is more or less transversely striated, the
striae running into reticulations ; the outer apical areae
coarsely reticulated, the apex of the spiracular area is more
coarsely punctured and more or less striated-reticulated than
the rest. The lower part of the propleurae is irregularly,
stoutly, obliquely striated. Abdomen closely punctured,
strongly at the base, becoming weaker towards the apex ;
the apical segments are densely covered with fulvous pubes-
cence. Gastrocoeli deep, large, the base and inner side with
curved striae, the rest more strongly longitudinally striated.
The areola is broader at the apex than it is long, the base
192
Mr. E. I. Pocock on the
broadly rounded, half the width of the apex, wliicli is sligditly
rounded inwardly; the bordering keel is shining, flat,
sniootli, broad at the base ; the apex is broadly depressed,
flnely rugosely punctured, the rest irregularly punctured and
aciculated.
XXII. — On the Blaclc-and~tan Pattern of Domestic Dogs
(Canis fainiliaris). By R. I. PocoCK, F.L.S., F.Z.S.,
Superintendent of the Zoological Society's Gardens.
In ^ The Variation of Animals and Plants under Domestication,^
pp. 33-35 (ed. 1905), the black-and-tan pattern of domestic
dogs is discussed at some length. Darwin was led to investi-
gate the question somewhat fully in the hope that he might
thereby discover a clue to the origin of our breeds amongst
wild species; but failing to find tlie tan-coloured spots over
the eyes either depicted in any drawings of wolves, jackals,
and other species of Canis^ or visible on any skins in the
collection of the British Museum, he came to the conclusion
that the coexistence of these spots with tan-coloured paws is
probably a case of correlated variation.
Apart from suggesting that a now extinct species involved
in the pedigree of domestic dogs may have possessed these
spots, he offered no other explanation of the highly remark-
able^^ fact of the occurrence of these spots in extremely
different breeds, living in various parts of the world."’'’
The phenomenon, however, appears to me to be susceptible
of a quite simple explanation.
In a typical black-and-tan dog, whatever the breed, the
tan is distributed as follows : — on the sides of the muzzle and
lips, the lower half of the cheeks, and the throat; a spot
over the inner corner of the eye, very frequently on the inside
of the ear, and as a large patch on each side of the chest above
the base of the fore legs ; on the paws of the fore legs and on
the hind legs below the hock ; to a somewhat variable degree
on the inner sides of the legs, but extending over the front of
the hind leg up to the body ; on the circumanal area and on
tlie underside of the tail, at least in its proximal portion. The
rest of the animal is black. If a dog thus coloured be com-
|)ared with many of the common wild species of Canidae, it
will be seen that the tan occurs over areas which in the wild
species are paler than the rest of the body, owing to the
fading or absence of the black annuli which prevail in the
193
Blade- and-tan Pattern of Domestic Dogs.
liair elsewhere, and tlmt the black corresponds to the darker
portions of the body, where the hair is richly pigmented, in
the wild animals. This statement only needs qualification
with respect to the tan spots over the eyes, the homologues of
which are by no means always visible in wild dogs, or, at all
events, are not sufficiently evident to carry absolute conviction
as to their presence. This is the case with examples of the
following species now living in the Zoological Gardens : —
Canis latranSj anthus^ lupasteVy aureus^ and mesomelas^ \ Guon
duhhunensis ?c(\di alpinus. On the other hand they are detect-
able, though minute, in some examples of Vulpes vulpes^ and
visible, though not conspicuous, in two Dingos which I have
reason to think have a mongrel strain of domestic dog. In
the case, however, of the wolves now or lately living in the
Gardens, namely in a Siberian example of C. lupuSj in three
specimens of C, lupus occidentalism and in one of C, pallipesm
there is no possibility of overlooking them. Although not
emphasized by a setting of jet-black hair, they nevertheless
show up as pale spots relatively as large as the corresponding
tan spots in dogs. Their conspicuousness in these species
suggests, though it does not prove, a preponderance of the wolf
strain over that of jackals in our breeds of domestic dogs.
Black-and-tan dogs may be termed melanescent, or,
preferably, nigrescent sports. Were they completely melan-
istic or perfect ^‘melanos^^ fkey would be black all over, as
many dogs are. It is evident that the tan stands in the same
relation to the pale areas as the black does to the more
heavily pigmented areas of the wild species; and it is a
highly interesting fact that the nigrescent sport throws back
to the type of pattern characteristic of a parent form. Tan
is merely one of the shades of that class of colour which is
commonly called erythristic ; and, assuming the truth of
the above-given explanation of the occurrence of tan in
dogs, it appears that albinism, erythrism, and melanism are
three consecutive stages in colour-variation, erythrism being
the incipient stage either of albinism or melanism, according
as the organism is albescent or nigrescent — that is to say,
assuming or tending to assume the albino or melano livery.
I believe this law of colour-change will prove to be
capable of wide application in the Mammalia and probably
outside the limits of that class. For example, there are in
the Zoological Gardens at the present time some black-and-tan
mouHon [Ovis musimon) . In these animals the white portions
of the typical wild sheep are tan and the rest of the coat
* This is not strictly true of all examples of this species.
Ann. dc Mag. N. Hist. Ser. 7. Vol. xix. 13
11)4
Dr. A. Vaughan on Seminula.
lieavily suffused with black, exactly as in the case of black-
and-tan dogs. They are intermediate in colour between a
ty])ical mouflon and a completely melanistic sport of that
species; and, be it noted, they commonly produce perfectly
black lambs. They furnish an instance of the tan being
the halfway stage between white and black in organisms
assuming a black pelage.
Conversely, as examples of the tan or red being the inter-
mediate stage between the normal and the albino sport, may
be cited yellow or ‘‘ginger” varieties of domestic cats, which
frequently at all events, and perhaps always, have the pads
of the feet pink instead of black ; and also red-haired blue-
eyed types of some Jews, whose colour Prof. Haddon speaks
of as a kind of minor albinism. In support of this 1 may add
that in the ‘ Sketch ^ for Nov. 14th, 1906, there was a photo-
graph of a Kaffir reported to have had a white skin, pale blue
eyes, and short, woolly, yellow hair. It is well known that
some wholly white, or partially white cats like Siamese have
blue eyes. Hence the blueness of the iris appears to be a
sign of albescence both in the human and the feline species.
XXIII. — Seminula : a Note hi/
Aethur Vaughan, B.A., D.Sc., F.G.S.
In the Ann. & Mag. Nat. Hist. ser. 7, vol. xviii., Nov. 1906,
pp. 321-327, ]\lr. S. S. Buckman, F.G.S. questions the
accepted use of certain Carboniferous generic names and
suggests somewhat drastic alterations.
He has, in the case of Seminula^ drawn attention to as
pretty a tangle as palaeontologists have ever made, and I
have to acknowledge his very courteous aid in my attempt to
unravel it.
I shall content myself with the statement of the results
which I have obtained from a careful examination of Sj'^eci-
mens and figures at the British Museum, and, in this work, I
am under a pleasant sense of obligation to Dr. F. A. Bather
and Mr. C. D. Slierborn. The question of priority in names
must be left to experts in nomenclature.
Seminula.
The genotype is stated by Mr. Buckman to be Terehratula
fentuedra^ Philh, and in this opinion he has the support of
Hall & Claike and Schuchert. It is only necessary, there-
fore, to discover to what genus this species belongs.
195
Dr. A. Vciuglian on Semlnula.
The figure of the species given in the ‘ Geology of York-
shire/ pt. 2, pi. xii. fig. 3, is a good and truthful representa-
tion of a specimen in the Gilbertson Collection (Brit. Mus.)
which has been type-labelled without any expression of
doubt. The figure is very slightly enlarged, as is usual with
all Phillips’s drawings, but there cannot be any reasonable
doubt that this specimen is the holotype of the species, and
no such doubt has been previously suggested.
Description of the Holotype of Terehratula pentaecha, Phill.
(the Genotype of Seminula) .
The type specimen has lost the greater part of its test,
and consequently the characters of the external ornament are
not so obvious as could be desired.
Form. — The size and outline of the shell agree closely with
those of the holotype of Spirifer ambignus^ Sow., with which
I have carefully compared Phillips’s specimen ; the two shells
have the same pentagonal outline, the same position of
maximum width, and the beak is ‘‘ produced in the same
manner in each. Whereas, however, Spir, amhiguufi is
strongly convex. Ter. pentaedra is rather strikingly flattened
(a character reminiscent of Ter. hastata^ Sow.).
Hinge-line and Fold. — Tiie valve-intersection is continu-
ously curved in the beak-region ; it lies completely in one
plane, except for the shallow lobe which marks the extremity
of the mesial fold.
The mesial fold is marked out on the cast by two strong
radial ridges, separated by a broad shallow depression, and
the whole fold is raised but little above the level of the
flanks.
In Spir. amhiguus, Sow., the mesial fold is formed on the
same pattern, but is prominently raised above the flanks of
the shell.
Pedicle- valve. — The test being partly removed from the
beak-region, the cast of the pedicle-cavity is satisfactorily
exposed ; it exhibits a narrow flattened area, bounded on
either side by a precipitous drop, indicating the presence of
strong dental plates. These characters are possessed in
common by all Athyrids.
There is no evidence as to the size of the aperture.
Structure of Test. — The test, where preserved, is minutely
fibrous and impunctate, but it is doubtful whether the outer-
most layer of the test is anywhere present.
Surface-ornament. — (a) Concentric. Two or three con-
centric ridges on the cast indicate marked growth- halts.
19G
Dr. A. Vaughan on Semlnula.
(Z>) Radial. There is no clear indication, either on the
cast or on the remnant of the test, of the impressed glabri-
striation which characterizes the Athyrids provided with
fringed or ribbed flounces [Cleiothyris and Actinoconchus) .
[On the same tablet with the type specimen of Ter, pent-
aedra in the Gilbertson Collection are nine other specimens.
Eight of these have well-preserved double- valves, and can be
assigned with certainty to an Athyrid provided with fringed
or ribbed flounces, by reason of the radial impressed ornament
and the low beak. It was probably on the evidence of these
specimens that Phillips accredited a minute perforation to his
species Ter, peyitaedra. These specimens differ completely
from the type specimen in form, beak, ornament, and nature
of fold.
The remaining specimen on the tablet (a cast) approaches
more closely to the type specimen both in form and beak,
but differs in exhibiting conspicuous glabristriation. Where
broken it exhibits part of a spire.]
Discrimination of the Genotype of Seniinula from other
Genera.
The holotype of Terehratula pentaedra, Phillips, differs
from the following genera in the characters subjoined.
From Camarophoria [genotype: Terehratula Schlotheimi ^
von Buch] in the fact that the dental plates do not unite,
absence of mesial septum, absence of angular plaits, low mesial
fold, &c.
From Martinia [genotype : Spirifer glaher (Martin)] in
presence of dental plates, absence of area, excavated mesial
fold, &c.
Yyou\ Dielasma [genotype: Terehratula elongatus (Schlo-
theim)] in impunctate test and the nature of fold.
From the Athyrids with fringed or ribbed flounces \_Gleio-
thyris (King), genotype : Atrypa pectinifera, J. de 0. Sow.]
in the outline, the nature of the fold, the absence of glabri-
striation, and the produced beak.
From Actinoconchus [genotype : Actinoconchus paradoxus^
M‘Coy (which is united with Spirifer planosulcatus^ Pliilh,
by Davidson)] in the deflection of the valve-intersection and
the presence of a mesial fold, the absence of a mesial septum
in the pedicle-valve, and the absence of glabristriation.
On the other hand, the type specimen of Seminula agrees
in all its generic characters with Spirifer amhiguus^ Sow,,
from which it only differs specifically in the lower fold and
more flattened shell [as was pointed out by Davidson (Pal.
197
Dr. A. Vaughan on Seininiila.
Soc., Garb. Bracli. p. 78) in explanation of the fact that he
queried his own assertion that Ter. pentacdray Phill. was a
synonym of S^^irifer ambiguuSj Sow.].
Ter'ehratula serninula^ Phillips, Geology of Yorkshire, pt. 2,
pi. xii. figs. 21, 22, 23.
Since M‘Ooy himself, in his ‘Palaeozoic Fossils^ (1855),
subsequently stated that this species was the type of Semi-
nulttj I also examined the specimens which are assigned to
this species in the Gilbertson Collection.
From the catalogue of the collection it appears that there
were originally fifteen specimens ; there are now only ten,
and one of these is a small Spiriferid. The remaining nine
all obviously belong to the same species and are closely
related to Camarophoria glohulina^ at least generically.
(Davidson t very justly draws attention to the presence of a
strong mesial septum in the pedicle-valve, visible through the
test.)
It cannot be reasonably doubted that the specimen actually
figured by Phillips (fig. 21, loc. cit.') had the same general
characters, although none of the actual nine specimens can
be definitely considered to agree with his figure.
Hence, in all probability, and as Davidson stated, the type
of Terehratula seminula^ Phill., is a Camarophoria (so long
only as that genus continues to include Terehratula glohulina^
Phillips).
COMPOSITA, Brown.
As Mr. Buckman has pointed out, Brown states very
definitely that “ the genus [i. e. Gompositai] is founded
upon the Spirifer arnhiguus of Sowerby.^^ Unfortunately it
is equally clear that Brown had an erroneous conception of
Sowerby’s species, for his figures of Gomposita amhigua
(‘ Fossil Conchology^ (1845), pi. liv.* figs. 4 & 5) represent
Spirifer [Martinia] glaber (Martin), and not Sowerby’s
species, Spirifer ambiguus.
Brownes figure exhibits the following differences from
the holotype ]: of Spirifer ambiguus. Sow. : —
(1) Large size : the figure measures l''*2 x 1", very un-
usual dimensions for a Seminula.
t Loc. cit. p. 116.
j This holotype, which is represented in the two uppermost figures on
pi. ccclxxvi. Min. Conch., is a small double-valved specimen, preserved
in the Sowerby Collection in the British Museum. The other two figures
of the plate depict a very imperfect pedicle- valve and a mere fragment of
a brachial valve, neither of which can be accepted as adequate definition
of a species.
198
Dr. A. Vaugliaa on Seiniiiula.
(2) Marked traiisverseiiess : Spirifer ambigaas is typically
an elongated shell.
(d) The sides of the shell have a continuoasly uniform
curvature : in Spinfer amhigans the curvature is
strong at the point of maximum width, but the outline
becomes rapidly straight above and below that
poii.t.
(4) The fold is uniformly convex in transverse section : in
S. amhigans the section is flattened and usually
excavated on the top of the fold.
(h) The fold is strongly semiconical and extends to the
beak : a character very common in the Spiriferids,
but almost unknown in the Athyrids.
(6) A short straight hinge-line, extending the width of
the beak of the brachial valve (as indicated by the
approximation of the valve-intersection to a horizontal
direction on either side).
(7) The absence of any concentric growth-halts and com-
plete smoothness : Seminula usually exhibits strong
concentric growth- halts.
The depiction of a perforated beak is quite negligible, for
the perforation, as drawn, is much too large to be the unbroken
aperture of S. ambiguuSj and, unfortunately, specimens of
Martinia glabra with the beak perforated in this way are
sadly common.
As a matter of mere personal experience, many specimens
of Martinia glabra may be collected which agree with Brownes
figure of Composita ambigua, but I have never seen a specimen
of S. ambiguus which has any right to be identified with it.
Hence it seems clear that, though undoubtedly Brown
thought that his conception of Composita included Spirifer
ambiguus, Sow., he did so in error.
Here is another problem for the expert in nomenclature!
Luckily, Martinia has precedence over Composita, and conse-
quently escapes the danger of rejection in favour of Brownes
genus.
I should like to tender my thanks to Dr. Ivor Thomas, of
H.M. Geol. Survey, for assistance in examining the Gilbertson
specimens, and also for pointing out to me, several months ago,
that Seminula was sickening for a serious illness. As
Dr. Thomas will [>robably explain his own views, 1 need say
no more beyond expressing the hope that my old and trusted
friend Seminula will “ pull through.’''
Blhliogrrtphlcal Notice.
199
BIBLIOGRAPHICAL NOTICE.
Illustrations of British Bloodsucking Flies, luith Notes by Ernest
Edward Austen. London : Printed by Order of the Trustees of
the British Museum, 1906.
The student of British Diptera has many difficulties to contend
with, and not the least of these is the absence of any good descrip-
tive handbook, especially one that is illustrated. The Lepidoptera,
Coleoptera, Aculeate Hymenoptera, Hemiptera, the Dragonflies and
the Sawflies have all been monographed and well illustrated by
British authors, but with the exception of Walker’s work, published
in the “ fifties” and now scarce and out of print (containing a few
plain lithographic plates), and the few plates in Curtis’s ‘ British
Entomology,’ there is nothing to help the collector of two-winged
flies. A book like the one now under consideration will therefore
be eagerly welcomed by the small but rapidly growing band of
British dipterists. Although the scope of the work, as indicated by
its title, does not admit of a purely scientific arrangement, yet the
volume gives us what is practically a pictorial monograph of the
Culicidae, Tabanidae, and Hippoboscidae, while a few members of the
Chironomidae, Simuliidae, and Muscidae are necessarily included. Of
the thirty-four plates it is impossible to speak too highly. Executed
by the three-colour process, on paper which is reputed to be perma-
nent, they are superb specimens of this comparatively new art, and
far surpass anything which has been attempted before in illustration
of this order of insects, either in this country or abroad, save, of
course, the splendid Monograph on the Tsetse Flies by the same
author and issued under the same auspices. No one attempting to
name examples in the three families more fully represented need
have any difficulty with the present work before him, and yet in
the past even the large and handsome Tabanid® were far from easy
to determine. Take, for example, the two species of Cleg, Ilcemato-
■pota pluvialis and H. crassicornis, both of fairly wide distribution.
It is impossible to imagine more beautiful and accurate figures of
these two species than those given on plates xi. and xii., while the
specific distinctions {e. g. the basal joint of the antenna, the black
spots on the frons, the light lines on the thorax, and the mottled
pattern on the wings) are strikingly shown.
The text is short but useful. It might have been more useful if
attention had been paid to the published records of the flies in question,
instead of limiting the notes on distribution to a list of specimens
actually in the British Museum collection. Thus, for example,
Atylotus fulvus has been recorded from Scotland, but there is no
mention of the fact on the page devoted to this species. But it is
easy to be too critical, and in spite of this slight sin of omission (which
can easily be rectified in a new edition) the volume is a great boon to
the collector of the Diptera of the British Islands. And not only docs
it appeal to the entomologist in this country, but also to those going
200
G eo I o(j tea I Soc lety.
abroad who, being interested in blood-sucking insects and their
connection with tropical diseases, will find in this handy and
beautiful book as perfect illustrations as they could wish for of
typical examples of all the families of Diptera possessed of such
pernicious habits. P. H. G.
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
November 21 st, 1906. — Sir Archibald Geikie, D.C.L., Sc.D., Sec.R.S.,
President, in the Chair.
The following communication was read : —
‘ On the Skull and greater portion of the Skeleton of Gonio-
pJiolis crassidens from the Wealden Shales of Atherfield (Isle of
Wiglit).’ By Reginald Walter Hooley, F.G.S.
In the late autumn of 1904, at a place locally called ‘ Tie Pits,’
near Atherfield Point, a huge mass of the cliff, comprising many
thousand tons of the Wealden Shales, subsided, pushing its foot
across the beach until below low-water line. As the sea washed
away the base, the mass continued to sink, and fresh horizons were
denuded. In 1905 a series of heavy ‘ ground-seas ’ cast up blocks
of limestone and ironstone, containing crocodile-bones, which were
discovered on the sand between high- and low-water marks. The
skull came ashore in six pieces. Fragments of bones, and scutes
were constantly picked up ; and the Author is indebted to
Prof. T. McK. Hughes for the block which had been picked up
and sent to the Sedgwick Museum at Cambridge. The specimens
were derived from a horizon 80 to 90 feet below the top of the
Wealden Shales. A history of the British Goniopholidae from the
foundation of the genus by Owen in 1841 is given, and it is noted
that the frame in the Mantell Collection, now in the British Museum,
not only contains the two type-blocks, but a smaller one with the
impression of the orbital region of the skull, a fragment of the
frontal bone, and the impression and fragments of a moiety of
the right ramus. The skull and bones of the new specimen are
next described, and a detailed comparison is instituted between
G. simus and G. crassidens^ with the result that the specimen
is referred to the latter species, differing in several important
particulars from the former. Comparisons are also made with
other species of Goniopholis, with dsannosuclius and OweniasneJms.
In conclusion, the Author notes that, while in certain features the
species comes nearer to the Teleosaurs than G. simus, it is farther
removed than the latter from them in the position of the posterior
nares.
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTM SERIES.]
No. 111. MARCH 1907.
XXIV. — Neio Species of Eastern and African ITeterocera.
By Colonel C. SwiNHOE, M.A., F.L.S., &c.
Family Syntomidae.
Syntomis sympJiona, nov.
$ . Antennae black, with white tips ; palpi black ; frons
ochreous, with the centre black ; head and body black, an
ochreous band behind the head j an ochreous spot on each
shoulder ; a longitudinal ochreous streak on each side of the
thorax ; abdomen with an ochreous band at the base and
another on the fifth segment, and some small ochreous marks
on the other segments : fore wings hyaline ; veins thickly
black, broadly so on the interior vein, which is joined to the
median - vein by a broad black bar, the interspace below
ochreous, the discoidal vein broadly black, the black running
up the interspace between veins 5 and 6 for two thirds its
length; costa broadly black, the outer marginal band nar-
rower, but broad at the tip : hind wings with the costa black,
the outer margin with a black band which runs up in the
middle in a circular form to the centre of the wing, and then
curves back to the anal angle, all the interior portion of the
wing being ochreous.
Expanse of wings inch.
Tenom, British North Borneo; one example.
Ann. May. N. Hist. Ser. 7. VoL xix.
14
202
Colonel C. Swinlioe on new
Family Arctiidae.
UtetJiesia IcalUmaj nov.
S . Palpi white, with black tips ; frons and head white, a
large black spot on lower part of the frons ; collar and shoul-
ders yellow, with black spots ; thorax and abdomen white, two
black spots on the middle of the thorax : fore wings bright
dark crimson ; six transverse bands of black spots encircled
with white, two spots close to the base, the other bands at
nearly equal distances apart, the sixth is marginal, the fourth
throws out a branch from the middle and curves close to the
fifth, the space between them here being white : the hind
wings are white, a short black band covering the discoidal
vein and a broad black band on the outer margin slightly
narrowing hindwards, but fairly broad all round, witli its
inner edge somewhat sinuous ; cilia of both wings ochreous
white, with grey spots. Underside : body and legs white,
with black spots ; tarsi black ; abdomen with lateral black
spots : fore wings with the spots confluent, forming rather
broad and prominent bands ; hind wings with two black
costal spots, and in the outer marginal black band are three
white spots — a large one at the apex, a small one in the
middle, and another near the anal angle.
Expanse of wings inch.
Angola; one example.
Family Agaristidae.
Pseudospiris j ucunda,
Pseudospiris jucunda, Jordan, Nov. Zool. xi. p. 444 (1904).
? . Similar to the male, except that the hind wings are
yellower, there is a large black and rather thick lunular mark
at the end of the cell, and a fairly broad blackish-brown
border on the outer margin, with large ochreous spots on the
margin.
Angola; four males, two females.
Dr. Jordan described this species from eight males from
the same locality ; the sexes differ much as they do in the
type of the genus {P. paidiformis, Butler, P. Z, S. 1895,
p. 267, pi. XV. figs. 8, 9).
Family EupterotidaB.
Hypsoides cleoiis^ nov.
d- Anfennge black; palpi, head, and body above and
203
Eastern and African Heterocera.
below ochreous : wings white, some ochreous hairs at base of
fore wings and abdominal margin of hind wings ; fore wings
with the veins, a band on the costa, and the apical space
blackish, this colour running narrowly down the outer margin
and is diffuse inwardly ; cilia blackish: hind wings without
markings, upper half of cilia blackish : the abdomen has grey
segmental bands ; legs ochreous, with black stripes ; tarsi
black.
Expanse of wings inches.
Madagascar ; one example.
Allied to H. hipars^ Butler, also from Madagascar, but the
outer third of the fore wings of that species is black, with a
sharply defined inner margin to that colour.
Family Lymantriidae.
Euproctis Butleri, nom. nov.
Ai'oa immaculataf Butler, Ann. & Mag. Nat. Hist. (5) x. p. 227 (1882).
Duke of York Island (type in B. M.).
I have it in both sexes from New Guinea and Borneo. I
must rename it, because immaculata is twice preoccupied in
the genus Euproctis^ and this insect is a true Euproctis. I
overlooked the species in my memoir on the Lymantriidse in
Trans. Ent. Soc. 1903.
Aroa niasana, nov.
S . Palpi and frons ochreous ; antennae purplish black ;
body and wings of a uniform dark black, tinged slightly with
pink ; a pale longitudinal streak below the cell on each wing ;
a pale lunular mark at the end of the cell of the fore wings ;
the wings coloured similarly above and below ; the abdomen
below and the legs are ochreous.
Expanse of wings inch.
Nias; two examples.
Dasycliira Dudgeoni, nov.
$ . Antennas, palpi, head, thorax, and fore wings dark
brown ; the fore wings variegated in parts with pinkish grey,
two brown ringed spots of that colour near the base; an
oblique indistinct band of lunular marks from middle of
hinder margin to costa at one third from the apex ; a double
row of similar pale markings near outer margin ; cilia
variegated, with a pale line at the base : abdomen and hind
14^
204
Colonel C. Swinlioe on new
wings pale brownish grey, without any markings. Under-
side wdiitish grey, with indications of a transverse, discal,
darker grey band across both wings.
Expanse of wings, $ IJ, $ inch.
Sikkim [Dudgeon) {(^ type in B. M.).
Khasia Hills, $ (fjpe ?)•
Abdomen without dorsal crests.
Dasychlra cymata^ nov.
d . Antennfe, palpi above, head, thorax, and fore wings
dark olive-brown, the last with subbasal, medial, discal, and
submarginal erect lunular black lines, finely marked with
white in places, the second line containing two rather large
dentations ; a rather large lunular ochreous stigma, ringed
wdth black, at the end of the cell : hind wings pale greyish
brown without markings : abdomen grey, with a thin brown
dorsal stripe. Underside pale whitish grey, tinged with
ochreous; the fore wings with pale brown suffusion.
Expanse of wings 1-]% inch.
Darjiling (Moore Coll.) (type in B. M.).
Family Notodontidae.
Tarsolepis javanOj nov.
(^ . Antenna0 brown; palpi, frons, and head ochreous
chestnut-colour ; fore part of the thorax grey, with a thin
black line in front; rest of thorax brown; patagia pinkish
grey, nearly white ; abdomen ochreous grey, with pale brown
segmental bands, abdominal tuft ochreous : fore wings with
broad costal and outer marginal pale pinkish-grey bands ;
the inner portion of the wing dark chestnut-brown, with pale
streaks and a pale space at the base; two hyaline elongated
spots as in T, Sommeri, Hiibner, but proportionately shorter,
and the upper spot with its outer margin curved inwards ;
the outer marginal band contains two or three indistinct grey
lines ; the outer margin is slightly crenulated and has long
black lunules ; the cilia are ochreous, interlined witli grey :
hind wings white, tinged with pale pinkish, the veins promi-
nent, the outer margin with indistinct grey lunular marks.
On the underside the wings are whitish ; a minute black dot
at the end of the cell of the fore wings and a large black spot
on the hind wings.
Expanse of wings 2 inches.
East Java; one example.
Eastern and African lleterocera.
205
Dudusa nohills.
Eudusa nohihs, Walker, xxxii. 447 (1865).
Crinodes Vethi, Snellen, Vetli’s Mid.- Sum., Lep. p. 40, pi. iv. fig-s. 1, 2
(1880).
The type from N. Cliiiui is in the B. M.
Snellen’s types were from Sumatra. 1 have a female from
Ichang-, a female from Makassar, Celebes, and a male from
the Khasia Hills. There is an example in Mus. Oxon. from
Singapore, and Ilampson records it from the Khasia Hills,
Malacca, and Celebes. They appear to me to be identical.
Dudusa synopldy nov.
$ . Antennse black ; palpi dark brown at the outer sides,
the inner sides, frons, head, and thorax ochreous brown ;
crest in the front of the thorax large and upstanding; abdo-
men pale ochreous brown, the sides with broad blackish-
brown segmental bands, the first band crossing the abdomen
above ; anal tuft ochreous grey, with black feathery tips :
fore wings pale ochreous fawn-colour, the veins somewhat
prominent ; a short subbasal blackish band and a broad
oblique blackish band from the costa a little before tlie
middle, where it is more or less obsolescent, to the outer
margin one fourth from the hinder angle, much as in
D. nohilis ) a narrow greyish band from the outer margin
below the apex, running in a slightly waved form to the
hinder margin near the base, where it broadens and becomes
diffuse ; indications of antemedial, double, transverse grey
lines and two similar lines from the coTa one third from the
apex, where it bends inwards, to the middle of the hinder
margin, these lines somewhat crenulate; submarginal pale
lunular marks, with a short brown lunular line in each and
marginal brown lunular marks : hind wings dark brown, with
duplex marginal brown lunules. Underside greyish ochreous ;
a blackish-brown spot at the end of each cell, and a crenulated
brown discal line across both wings, bent inwards below the
middle on the fore wings, evenly curved outwards on the
hind wings; marginal lunules as above; thorax dark brown,
tarsi black ; abdomen greyish ochreous.
Expanse of wings 4^ inches.
Khasia Hills ; three examples.
The antennae are as deeply bipectinate as in the male of
D. nohills^ of which I have both sexes; the wings are much
broader.
206
(volonel C. Swirihoe on new
Fydna hrunnea^ nov.
^ , Antennae, palpi, bead, body, and fore wings ocbreous
brown : fore wings with some pale shades, some costal
blackish-brown dots, a spot one third from apex, a spot below
the middle of the cell, and an outwardly curved discal row of
black dots : hind wings blackish brown, much darker than
the fore wings, no markings. Underside : body and both
wings pale greyish ochreous ; the fore wings with the interior
portions suffused with blackish brown and some marginal
black dots on the upper half ; hind wings with a brown spot
at the end of the cell and a thin discal band, its upper half
crenulated.
Expanse of wings inches.
Khasia Hills; one example.
There is a worn example of a male from Ceylon in the
B. M. unnamed.
Notodonta nodyna, nov.
^ . Antennae, palpi, and thorax brown ; a broad band
behind the head blackish brown ; abdomen paler than the
thorax, with a broad dorsal dark brown band : fore wings
dark brown, with a pink tinge, with three indistinct longitu-
dinal dark brown stripes from near the base to the apex and
a blackish submarginal stripe from the outer margin above
the middle to the hinder margin one third from the angle,
accompanied by a similar stripe on the margin, which extends
along the margin and stops at one third from the base;
between these stripes the colour of the wing is paler : hind
wings white, semihyaline, the veins prominent; some grey
suffusion towards the costa and abdominal margins ; a grey
thin band near the outer margin, its lower portion dark and
brownish; a marginal brown band and brown interlined
cilia.
Expanse of wdngs inches.
Khasia Hills ; one example.
Very similar in shape to N. gigantea^ Elwes, of which I
have several examples from the same locality, but the outer
margin, especially of the fore wings, is highly crenulate.
Hyperceschra plana, nov.
? . Head, thorax, and fore wings dark olive-brown, the
latter with a black longitudinal basal streak and many in-
distinct thinner streaks on the outer portions of the wings ;
indications of an angulated discal transverse line ; a sub-
207
Eastern and African Ileterocera,
marginal brown thin band, sinuous and dentated in parts :
hind wings pale grey, with a dark greyish-brown broad
marginal band, the entire wing suffused with pinkish ;
marginal line of both wings blackish brown ; cilia pinkish
grey, with brown patches on the fore wings ; with a brownish
middle band and white tips on the hind wings. Underside
pale uniform grey, with an indistinct pale discal band across
the hind wings ; hind tarsi with brown spots.
Expanse of wings inch.
Omei-shan, W. China (type in B. M.).
HypercescJira tusa, nov.
d . Dark grey : fore wings with more than the basal half
much darker than the outer portions, limited by a thick black
line from the costa at two thirds to vein 4, then curved in-
wards to vein 1, then obliquely inwards to the production at
the middle of the hinder margin, where it is very thick ; a
black mark just inside this on the margin, and two black
angles between them, forming the commencement of a trans-
verse line ; an indistinct, grey, transverse, sinuous band in the
disc ; hind wings grey, without markings ; cilia of fore wings
ochreous grey, of hind wings pale grey, both with darker
patches : antennae, body, legs, and wings on the underside
pale grey, uniform in colour, a slightly darker medial band
across both wings.
Expanse of wings 2 inches.
Japan [Schaus) (type in B. M.).
Allied to H, hasalis, Moore.
HypercescJira curvilinea, nov.
$ . Palpi blackish brown, white beneath ; head and body
ochreous grey ; fore wings with nearly the basal half pinkish
ochreous grey, with some longitudinal black streaks in it,
bounded by an outwardly deeply curved, thick, black, sinuous
line, much retracted a little above the hinder margin, then
straight to the margin at the middle ; the remainder of the
wing dark grey, containing a prominent black, outwardly
curved, highly sinuous line not far from the first line ; two
or three indistinct grey lines ; a submarginal row of black
spear-shaped marks and a black, longitudinal, subapical line :
hind wings grey, with a large black patch on the outer
margin near the anal angle ; cilia of both wings grey, with
small darker patches with a white dot in each patch. Under-
side pale greyish brown, the hinder marginal space of the
208 Mr. A. S. Hirst — Notes on Scorpions.
fore wings and all but the costal space of the hind wings
whitish ; costa of both wings suffused with ochreous, with
some black marks on the outer half of the costa of the fore
wings and indications of a discal line ; a curious angular
mark beyond the middle, below the costa of the hind wings ;
body and legs whitish.
Expanse of wings 2 inches.
Durban, August 1900 (type in B. M.).
LopJiopteryx uniformis^ nov.
$ . Antennae with the shafts brown, plumes ochreous ;
head, thorax, and fore wings dark chestnut-brown, with a
pink tinge; antemedial and postmedial indistinct bands of
pale lunular marks, straight and slightly oblique ; cilia con-
colorous, with a pale basal band : hind wings grey, pale on
the basal half; cilia grey, with a whitish basal band : abdo-
men wdiitish, tinged with pink, with thin grey segmental
bands. Underside: fore wings uniform dull purplish brown,
hind wings grey, no markings ; body and legs grey.
Expanse of wings 2y\ inches.
Mashonaland {Dohhie) (type in B. M.).
XXV. — Notes on Scorpions., with Descriptions of Two new
Species. By A. S. Hikst.
Family Buthidae.
Babycurus somalicus, sp. n.
Colour brownish yellow or yellow; vesicle, hand, and
lower surface of body paler ; fingers of hand light brown ;
aculeus black, the basal portion excepted.
Carapace slightly trapezoidal, the frontal edge weakly
concave ; ocular tubercle granular, the groove between the
eyes smooth. Distribution of granules apparently much as
in zamhonelUj the surface being covered wdth numerous
granules, unequal in size and leaving several small spaces
(devoid of granules), which are especially noticeable in the
median part. Median eyes bordered on each side by an
area furnished with minute granules. Median area of fronttd
region with an aggregation of large granules.
Abdomen. Tergites minutely granular in their anterior
parts, posteriorly provided wiHi large shining granules in
Mr. A. S. Hirst — Notes on Scorpions. 20i)
addition to the minute ones. Keels granular, incomplete
anteriorly. Tliird and fourth sternites with sparse granules
laterally ; fifth sternite granular and provided with four
granular keels.
Tail, Keels distinct and granular In all segments. First
segment with ten keels, segments 2-4 with eight, fifth seg-
ment with five. Dorsal surfaces concave in segments 1-4 ;
fifth caudal segment slightly convex dorsally and provided
with a narrow and shallow median groove ; posterior portion
of this last segment narrowed, but exceeding the vesicle in
breadth. Intercarinal spaces minutely granular ; the granules
very sparse on the dorsal surfaces of the first four segments ;
fifth caudal segment more strongly granular. Vesicle hairy,
granules absent ; spine under aculeus laterally compressed
and terminating in a little knob.
Palp. Hand with scattered granules on the inner edge,
outer and inner keels of the dorsal surface minutely but
distinctly granular; finger-keel granular only in that portion
which is situated on the hand itself. Movable finger pro-
vided with eight series of granules exclusive of the short
apical series, six series ending on the outer side in two large
granules, the last pair of granules situated at the base of the
prebasal series. Upper surface of the femur of the palp
minutely and densely granular.
Pectinal teeth 19-20 in number.
Measurements in rnm. Length of cephalothorax 5'5, of
tail (vesicle excl.) 23, of movable fingerfi, of hand-back 3*6 ;
breadth of hand 2*25, of tibia 1'75; total length 47.
Mab. A $ specimen (the type) labelled Berbera and
Durbar, Somaliland, sea- level to 400 ft.,^^ and a second from
the Wagar Mountains behind Berbera, 3000-4000 ft. The
specimens were collected by Mr. G. W. Bury.
Remarks. This species is closely allied to zamhonelli,
Borelli, from which it apparently differs in the granular keels
of the manus, the little knob terminating the spine under
the aculeus, the distinct keels of the last caudal segment, the
minute granules of the upper surface of the femur of the palp,
and in the fact that the movable finger is provided externally
with two large granules situated at the base of the prebasal
series of granules.
Genus ISOMETROIDES, Keys.
There are two species of this genus in the collection of
the British Museum and each is represented by a single
specimen. I believe them to be the two described species
210 Mr. A. S. Hirst — Notes on Scorpions.
I. angusticaudus, Keys., and I. vescuSj Karscli ; I have not
seen the typical specimens of these species, however. The
more important differences are given below,
Isometroides angusticauduSy Keys.
Isometroides angusticaudus, Keyserling, Araoh. Austral, vol. ii. p. 19
(1884-89) ; Kraepelin, Das Tierreich, Scorpiones et Pedipalpi,”
p. 40 (1899).
Fourth caudal segment with eight granular and well-marked
keels; surface granular, the granules larger than in the pre-
ceding segments, with few punctures. Surface of fifth caudal
segment with numerous punctures, rugulose. Pectinal teeth
twenty-three in number. Measurements in mm. : length of
tail (vesicle excl.) 23, of fifth caudal segment 6*5 ; breadth
of fifth caudal segment 3’5 ; total length 41.
Hah. A single example from Port Lincoln, Australia.
Isometroides vescus^ Karsch.
Isometrus vescus, Karsch, SB. Ges. naturf. Berlin, p. 58 (1880).
Isometroides vescus^ Keyserling, Arach. Austral, vol. ii. p. 17 (1884-89) ;
Kraepelin, Das Tierreich, “Scorpiones et Pedipalpi,” p. 40 (1899).
Fourth caudal segment provided with eight indistinct keels;
devoid of granules and with distinct punctures. Fifth caudal
segment smooth and shiny, with numerous punctures.
Pectinal teeth 24-25 in number. Measurements in mm. :
length of tail (vesicle excl.) 2P5, of last caudal segment 6;
total length 39 ; breadth of last caudal segment 2'5.
Ilah. Kalgorlie near Coolgardie, W. Australia.
Tityus Kraepeli'ii^ Pocock,
Tiiyus Kraepelini, Pocock, Ann. & Mag. Nat. Hist. (7) x. p. 379
(1902).
The name Kraepelini being preoccupied *, I propose that
of Pococki for this species.
Hah. Abundant at Merida, also found at Obama, Venezuela.
Family Scorpionidae.
Diplocentrus nitidus, sp. n.
Colour. A very dark brown, maims paler, legs and under
surface of a much paler colour.
* Borelli, Boll. Mus. Torino, xiv. no. 315 (1899).
211
Mr. A. S. Hirst — Notes on Scorpions,
Carapace minutely punctured, more sparsely so on the
raised portions ; surface smooth and polished, devoid of
granules. Median groove present in front of the eyes.
Anterior border with a deep bay.
^5<iome«.'Tergites smooth and polished, minutely punc-
tured ; provided on their hinder edges with a row of granules,
which is incomplete in the median line. Last tergite with
large isolated granules laterally. Sternites smooth and
polished, minutely punctured ; the punctures more distinct
on the last sternite, which is furnished with four smooth
keels.
Tall stout ; superior dorsal keels of the first four segments
formed by a few fairly large granules, the posterior granule
of each series enlarged. Ventral keels much as in antillanus,
smooth, and each bearing a few setiferous depressions.
Accessory lateral keels ill-defined. Vesicle very stout, its
surface minutely punctured and devoid of granules.
Fectines with nine teeth.
Feet with minute punctures, tarsi angular and with but
few punctures; tarsi of third and fourth legs provided with six
spines on the inner edge and with a proximal crescent of four
granules, externally with 6-7 spines.
Manus narrow, lobe reduced ; dorsal surface smooth and
keelless, less densely punctured than is the case in antillanus
and with frequent unpunctured spaces which give it a reti-
culate appearance.
Measurements in mm. Length of carapace 5, of tail (in-
cluding vesicle) 20, of first two segments of tail 5*5, of last
segment of tail 4*75, of movable digit of hand 5 5; total
length 36 ; breadth of last segment of tail 2*6, of tibia of
palp 2*25, of vesicle 3 ; greatest breadth of hand 4 ; length
of vesicle (aculeus excl.) about 4.
Hah, San Eamon, Kio Wanks, about 185 miles above
Cape Gracias, Nicaragua. A single specimen collected by
Mr. G. Palmer in the month of July 1905.
Remarks, Allied to antillanus, Pocock, from which it may
be distinguished by the weakly lobed, keelless hands, the
dorsal surfaces of which are punctured in a reticulate fashion,
and also by the stout tail and vesicle, the coarse and less
numerous granules of the superior dorsal keels of the tail,
and the deep bay of the anterior margin of the cephalo-
thorax.
212
Mr. G. A. Boulenger on new
XXVI. — Descriptions of Three new Liz irds and a new Frog,
disco oered hi) J)r. W. J. Ansorge in Angola, By G. A.
BoulengerJ F.li.S.
Phy Uodactylus An sorgi i.
Head ratlier small^ oviform, much longer than broad ; snout
not longer than the distance between tiie eye and the ear-
opening, which is small and oval. Body very elongate ;
limbs moderate. Digits moderately depressed, with large,
sublrapezoid terminal expansions ; eight lamellae under the
fourth toe. Head and body covered with uniform, smooth,
flattened granules, which are larger on the snout and on the
belly. Rostral twice as broad as deep, without cleft above ;
symphysial small, a little longer than broad ; ten upper and
as many lower labials ; rostral and flrst upper labial entering
the nostril ; no chin-shields. Tail cylindrical, (apering,
covered with uniform, small, quadrangular, smooih scales.
A curved transverse series of 8 or 9 enlarged prseanal scales
(indicating prmanal pores in the male?). Bale greyish brown
above, with a series of large whitish spots along each side of
the back ; a dark streak on each side of the head and neck,
passing through the eye; upper lip and lower parts white,
with small brown spots.
mm.
Total iengtli 75
Head 10
Width of head 6
Body 35
Fore limb 12
Tlind limb 15
Tail 30
Two female specimens from Maconjo, Benguella.
The more slender form and the enlarged pra3anal scales
well distinguish this species from P. porphyreuSy Daud.
Mahuia Icevis.
Head and body much depressed. Snout short, pointed.
Lower eyelid with a very large undivided transparent disk.
Nostril behind the vertical of the suture belwcen the rostral
and the flrst labial ; a postnasal ; anterior loreal in contact
with the first labial ; supranasals narrow and widely separated
by the fronto-nasal, which forms a suture with the rostral and
with the frontal ; frontal as long as fronto-parietals and inter-
parietal together, in contact with the first, second, and third
213
Lizards Jrom Angola.
supraocLilars ; four supraoculars ; six supraciliarles ; parletals
in contact behind the interparietal, followed by a pair of large
nuchals ; five upper labials anterior to the subocular, wdiich
is large and not much narrowed interiorly. Ear-opening
large, suboval, with projecting granules on its anterior
border. Scales perfectly smooth, dorsals largest, 32 round
the middle of the body. The hind limb reaches the elbow of
the aJpressed fore limb. Toes moderately long, slender;
subdigital lamellae with a tubercular keel. Tail depressed,
with a series of transversely enlarged scales above and
beneath. Head red, body and tail bluish grey; a broad
black vertebral band, bifurcating on the nape; a black lateral
band, passing through the eye, confluent with large spots
below it oi\ the neck and body ; ’ limbs bluish grey, with
the scales black-edged.
mm.
Total length 130
Head 13
Width of head 7
Body 39
Fore liinh lo
Hind limb 22
Tail ^ 78
A single specimen, rather damaged, from Macon jo,
Benguella.
Mahuia Ansorgii.
Snout short, obtuse. Lower eyelid wdth a large undivided
transparent disk. Nostril anterior to the vertical of the
suture between the rostral and the first labial ; a postnasal ;
anterior loreal not in contact with the first labial ; supranasals
in contact behind the rostral ; fronto-nasal broader than long,
forming a very narrow suture with the frontal ; frontal as
long as the fronto-parietals and the interparietal together, in
contact with the second and third su})raoculars ; four supra-
oculars, second largest ; five supraciliaries, second largest ;
fronto-parietals distinct, a little smaller than the interparietal;
parietals just meeting behind the interparietal; a pair of
nuchals ; five or six labials anterior to the subocular, which
is much narrowed inferiorly. Ear-opening oval, as large as
the transparent palpebral disk, with four short pointed
lobules anteriorly. Dorsal and nuchal scales strongly
quinquecarinate ; 42 scales round the middle of the body,
dorsals largest. The hind limb reaches the elbow of the
adpressed fore limb. Scales on the soles sharply keeled,
spinose ; subdigital lamellje sharply tricarinate, spinose.
214
On a new Frog from Angola.
Yellowish brown above, with two pairs of broad black lonf^i-
tudinal bands on the back and one on each side from the eye
to the tail ; each pair of dorsal bands separated by a narrow
light line and confluent into a single band on the tail; lips
and border of the ear pink ; lower parts whitish, chin spotted
with brown.
mm.
Total length (tail reproduced) 142
Head 17
Width of head 12
Body 69
Fore limb ’ 27
Hind limb 37
A single, somewhat damaged specimen from Caconda,
Benguella.
Closely allied to M. Bocagii^ Blgr.
Eana hunoderma.
Vomerine teeth in two short oblique series commencing
from the inner front edge of the choanse. Head moderate ;
snout pointed, as long as the orbit ; canthus rostralis obtuse ;
loreal region deeply concave ; interorbital space much nar-
rower than the upper eyelid ; tympanum distinct, two thirds
the diameter of the eye. Fingers moderate, blunt, first not
extending beyond second ; toes moderate, one-third webbed ;
subarticular tubercles small but very prominent ; a small,
oval, inner metatarsal tubercle, no outer tubercle. Tibio-
tarsal articulation reaching a little beyond the tip of the
snout. Back with very prominent, tectiform, large warts,
forming irregular longitudinal series ; no lateral fold. Olive-
grey above, with large roundish black spots on the body
and cross-bars on the limbs ; a fine light vertebral streak ;
a light cross-bar between the eyes and a light spot on the
tympanum ; lips black ; hinder side of thighs reddish brown,
dotted with white ; lower parts wdiite.
From snout to vent 35 mm.
A single female specimen from Caconda, Benguella.
Intermediate between R. mascareniensisj D. & B., and
R. Grayij Smith.
On new Shells from Lower Bengal,
215
XXVII. — Diagnoses of new Species of Corbula and Blthinella
from Lower Bengal, By H. B. Preston, F.Z.S.
The species described in the present paper form part of a
small collection recently sent to me for identification by the
authorities of the Indian Museum, Calcutta. All were
collected in the brackish pools near Port Canning, at the
northern extremity of the district known as the Sandarbans
in the Ganges Delta, and are the property of the Indian
Museum.
Corbula abhreoiata^ sp. n. (Fig. 1.)
Shell nearly equivalve, roundly ovate, inflated, thin,
greyish white, posteriorly obliquely rounded, anteriorly
sharply abbreviated, sculptured with rather coarse concentric
lines of growth ; umboes small, situated somewhat anteriorly.
Alt. 3*5, length 4*25 mm.
Hab, Port Canning, Lower Bengal; in brackish pools.
Type in Indian Museum, Calcutta.
Fig. 1.
Corhula ahbreviata.
Fio-. 2.
Corhula Alcocki.
Fig. 3.
Corhula calcaria.
Fig. 4.
Corhula gracilis.
Fig. 5.
Corhula Pfefferi.
Fig. 6.
Bithinella canningensis.
Corbula Alcocki^ sp. n. (Fig. 2.)
Shell irregularly rhomboidal, rather ventricose, tliin,
whitish, posteriorly rounded, anteriorly produced, angled
21G
On new Shells f rom Lower Bengal.
from the umboes, sculptured with fine concentric lines ot
growth; uinboes small, nearlj central.
Alt. 3*25, length 5 rum.
Jlah. Port Canning, Lower Bengal ; in brackish pools.
Type in Indian Museum, Calcutta.
Corhula calcaria, sp. n. (Fig. 3.)
Shell elongatelj oval, depressed, thin, chalky white, poste-
riorly sharply rounded, anteriorly truncate; umboes small.
Alt. 3, length 5 mm.
Hah. Port Can ning. Lower Bengal ; in brackish pools.
Type in Indian Museum, Calcutta.
Corhida gracilis^ sp. n. (Fig. 4.)
Shell oblong-ovate, thin, whitish, rather ventricose, obso-
letely concentrically ribbed, posteriorly rounded, anteriorly
elongately produced, truncate ; umboes small, situated some-
V hat posteriorly.
Alt. 4, length 6 mm.
Hal). Port Canning, Lower Bengal ; in brackish pools.
Type in Indian Museum, Calcutta.
Corhida Pfefferi^ sp. n. (Fig. 5.)
Shell ovate, somewhat triangular, moderately ventricose,
thin, white, posteriorly obliquely rounded, anteriorly bluntly
beaked, sculptured with rather coarse, smooth, concentric ribs
and very fine transverse stria3 or scratches, these last being
especially noticeable on the right valve; umboes small,
central.
Alt. 3*5, length 5 mm.
Hah. Port Canning, Lower Bengal ; in brackish pools.
Type in Indian Museum, Calcutta.
Bithinella canningensis^ sp. n. (Fig. 6.)
Shell subperforate, pyramidal, dull yellowish-brown horn-
colour ; remaining whorls 3, somewhat convex ; sutures well
impressed; aperture oval; peristome simple, continuous;
operculum horny, paucispiral.
Alt. 1*75, diam. maj. 1 mm.
I ah. Port Canning, Lower Bengal ; in brackish pools.
Type in Indian Museum, Calcutta.
On new Land- Shells from Xew Caledonia.
217
XXVIII. — Descriptions of Nine new Species of Land- Shells
from New Caledonia. By H. B. PiiESTON, F.Z.S.
When recently working tlirough a quantity of New Caledonian
land-shells I found a number of forms which seemed to be
difficult of determination; these I submitted to Mr. G. K.
Gude, who, with his unfailing courtesy, assisted me in identi-
fying a certain number. There were, however, several
species which Mr. Gude suggested might be new, and having
compared these carefully with all available material, I now
venture to describe them as follows : —
Charopa ahena^ sp. n. (Fig. 1.)
Shell depressed, subdiscoidal, thin, bronze-coloured; whorls
3-4, sculptured with moderately fine arcuate ribs; sutures
presenting an almost channelled appearance ; umbilicus
rather wide; aperture lunate ; peristome simple.
Alt. 2, diarn. maj. 4*5 mm.
Aperture : alt. 1*25, diam. 1 mm.
hab. New Caledonia.
4'his form shows some affinity ^^ith C. dispersa, Gass.*,
the chief characters which separate it from that species bemg
its more depressed spire, rather coarser sculpture, and
shallower and narrower umbilicus ; it has also one whorl
less than C. dispersa.
Charopa Margueritce, sp. n. (Fig. 2.)
Shell depressed, rather thin, yellowish-browm, blotched,
streaked, and spotted with chestnut ; whorls 4-5, sculptured
with fine arcuate ribs; spire slightly sunken ; sutures deeply
impressed ; umbilicus wide ; peristome simple ; columella
descending very obliquely ; aperture obliquely lunate ; inte-
rior of shell bearing a callus on the parietal whorl.
Alt. 2’25, diam. maj. 6*25 mm.
Aperture : alt. 2’25, diam. maj. 1 mm.
hah. New Caledonia.
In some respects allied to C. vetula, Gass.f. It is, however,
much larger and more depressed ; it is not so closely coiled,
* Gassies, ‘‘Faune Concliyliologique Terrestre et Fluvio-lacustre de
la Nouvelle Calddonie,” pt. i., Actes Soc. Linn. Bordeaux, xxiv, 1861,
p. 233.
t Gassies, ibid. pp. 233-234.
Ann. & 'Mag. N. Hist. Ser. 7. Vol. xix. 15
218
Mr. H. B. Preston on new
and the chestnut painting readily separates it from that
species.
Fig. 1.
Charopa ahena.
Fig. 2.
Charopa Marguerites.
Fig. 3.
Charopa Gioendolince.
Fig. 4.
Charopa Marioru^,
Fig. 5.
Charopa vicina.
Fig. 6.
Charopa Gassiesiana.
Fig. 7.
Charopa {Tr.) Gudei,
Fig. 8.
Ostodes vitreus. Acmella {8ol.) turbinata.
Charopa OwendoUnce^ sp. n. (Fig. 3.)
Shell orbicular, bearing traces of having been covered with
a rich brown periostracum ; spire concave ; whorls 5-6,
rather closely coiled, sculptured with fine, closely-set, arcuate
ribs, the last whorl descending somewhat abruptly near the
peristome; sutures impressed; umbilicus deep, moderately
wide; peristome simple; aperture lunate.
Alt. 3, diam. maj. 6 mm.
Aperture; alt. 3, diam. 1 mm.
Hah. New Caledonia.
Easily separated from the other Charopce described from
this region by its sunken spire and closely coiled whorls.
Tjand- Shells from New Caledonia.
219
Charopa Marionce, sp. n. (Fig. 4.)
Shell depressed, subcaririiite, somewhat pellucid, pale
yellowish horn-colour, indistinctly blotched and streaked with
light chestnut; whorls 3, sculptured with closely set arcuate
ribs, the last whorl flattened obliquely ; sutures impressed ;
umbilicus very wide : peristome simple; aperture ovate.
Alt. 2, diam. maj. 6'25 mm.
Aperture: alt. 2, diam. 2*25 mm.
Hnh. New Caledonia.
Chiefly remarkable for the excessively wide umbilicus,
which easily distinguishes it from any other species hitherto
described from New Caledonia.
Charopa vicina, sp. n. (Fig. 5.)
Kesembling C. Marionce in general outline; it is, how-
ever, rather smaller, higher in the spire, more carinate, and
of a uniform rich brown colour; the arcuate ribs are much
finer, the sutures not so well impressed, and the umbilicus
is deeper and rather narrower.
Alt. 2*75, diam. maj. 5’75 mm.
Aperture : alt. 2, diam. maj. 1*75 mm.
Hab. New Caledonia.
Charopa Gassiesiana^ sp. n. (Fig. 6.)
Shell narrowly perforate, thin, somewhat transparent,
yellowish horn-colour; whorls 3, regularly sculptured, with
rather distant broad and rounded ribs ; sutures impressed ;
peristome simple ; columella descending obliquely and slightly
reflexed outwards ; aperture roundly lunate, a callosity
appearing inside the shell upon the parietal whorl.
Alt. 2*5, diam. maj. 5 mm.
Aperture : alt. 2, diam. maj. 1*5 mm.
hah. New Caledonia.
Mr. Gude considers this species to be closely allied to
C. melitcej Gass.* ; it is, however, much smaller and more
swollen, and shows no signs of carination ; the umbilicus is
also very much narrower and the sculpture is a great deal
coarser than is the case with C. melitce.
I have much pleasure in dedicating this pretty little species
to the memory of the late Jean Baptiste Gassies, whose
* Gassies, “ Faime Concliyliologique Terrestre et Fluvio-lacustre de
la Nouvelle Caledonie,” pt. ii,, Actes Soc. Lina, Bordeaux, xxviii. 1871,
pp. 24-25.
15^
220 On new Land- Shells from New Caledonia.
Monos:raph of the Terrestrial and Fluviatile Mollusca of
New Caledonia is invaluable to the student of the concho-
logical fauna of that island.
Charopa [Tropidotropis *) Gudei^ sp. n. (Fig. 7.)
Shell discoidal^ sharply carinate, covered with a slightly
lamellose periostracum ; spire flat; whorls 3-4, painted with
purple flame-markings, which appear as blotches near the
sutures ; sutures impressed base of shell very convex ;
umbilicus moderately broad, deep; peristome simple;
aperture securiform.
Alt. 2*75, diam. maj. 7*25 mm.
Aperture : alt. 2, diam. 3 mm.
Hah. New Caledonia.
Readily separable from its ally T. trichocoma, Crosse f, by
its narrower umbilicus and by its less lamellose periostracum ;
moreover the spire is not quite so flat as is the case with
2\ trichocoma.
Ostodes vitreus^ sp. n. (Fig. 8.)
Shell depressedly turbinate, transparent, pale hoin-colour ;
whorls 4-5, the last three sculptured with spiral lirse, obso-
lete only on the base near the aperture ; sutures well im-
pressed ; umbilicus rather wide and deep ; peristome serrated
by the termination of the spiral lirse ; aperture subcircular ;
columella curved, a callosity joining it with the lip above ;
operculum paucispiral, horny, concave.
Alt. 5*25, diam. maj. 7*5 mm.
Aperture: diam. 2’5 mm.
Hah. New Caledonia.
The present species may be compared with Ostodes upolensis^
Moussa, from Upolu ; it is, however, more depressed, the
sculpture is coarser and more regular, and there are no traces
of crenulation ; moreover it is much thinner and more
transparent than is the case with 0. npolensis.
Acmella [Solenomphala) turhinata, sp. n. (Fig. 9.)
Shell perforate, turbinate, smooth, polished, brownish
horn-colour, indistinctly streaked with grey ; whorls 5, the
last three very convex ; sutures deeply impressed ; umbilicus
moderately wide; peristome simple; columella descending
* Ancey, Bull. Soc. Mai. Fr. v. p. 370.
t Journ. de Conchyl. vol. xyi. pp. 158-160.
On new Genera and Species 0/ Syrifomidse. 221
in a curve and reflc^xed outwards over the umbilicus ; aperture
roundly ovate ; operculum concave, horny, paucispiral.
Alt. o'16, diam. rnaj. 3*25 mm.
Aperture: alt. 1‘25, diam. 1 mm.
Ilab. New Caledonia.
Somewhat closely allied to Hydrocena caledonica^ Crosse * ;
it is, however, smaller, has one whorl more, is not so globose,
and is darker in colour ; the sutures are not so deep and the
base is less convex ; moreover it is easily recognizable from
H. caledonica by its much less variegated appearance, the
grey streaks in A. turhinata being only visible in a strong
XXIX. — Descriptions of new Genera and Species of Synto-
midse, Arctiadse, Agaristidse, and Noctuidee. By Sir
Geoege F. Hampson, Bart., F.Z.S.
The following paper forms a fourth supplement to the first
six volumes of the ‘ Catalogue of Lepidoptera Phalaense in the
British Museum,’ the former papers having appeared in the
Ann. & Mag. Nat. Hist. ser. 7, vol. viii. pp. 165-186 (1901),
vol. xi. pp. 337-351 (1903), and vol. xv. pp. 425-453 (1905).
The numbers before the species indicate their position in the
classification adopted in those volumes. The types are in the
British Museum.
Syntomidae.
11 a. Geryx alhipuncta^ sp. n.
S . Head, thorax, and abdomen black-brown ; frons
white; tegulse with lateral white patches; patagia with
white streaks on outer edge ; dorsum of thorax with white
streak, metathorax with whitish patch ; sides of pectus and
coxae with white patches ; hind femora streaked with white ;
abdomen with dorsal series of small whitish spots except at
extremity and complete ventral series of whitish bands.
Fore wing black-brown ; a wedge-shaped hyaline patch in
cell ; a patch in submedian interspace from base to near
termen, its lower edge indented by a small dark tooth at
middle ; a spot above base of vein 2 ; a postmedial streak
above vein 6 and slight streak above 7 ; elongate spots above
veins 4, 3. Hind wing black-brown, with small hyaline
* Journ. de Conch, xvii. 1869, pp. 24 & 25.
222
Sir G. F. llainpson on new
spot in lower extremity of cell and large patelies in sub-
median interspace and above vein 2 extending nearly to
termen.
Jiah. Philippines, Luzon, Benguet Prov., Irisan
(McGregor)y 1 ^ type. Exp. 30 mm.
45 a. Myopsyche xanthosorna^ sp. n.
$ . Head black ; palpi and proboscis orange ; antennae
Avhite at tips ; thorax black,' with some orange scales; the
tegulae and patagia orange ; legs mostly orange ; abdomen
orunge, the last three segments black at sides and below.
Fore wing hyaline, the veins and margins black ; the base
with orange patch ; a black discoidal patch, emitting a
fascia between veins 5, 6 to the terminal band, which ex-
pands widely on apical area and into a bidentate patch below
vein 2. Hind wing hyaline, the veins and margins black,
the base, cell, and costal area to beyond middle orange ; the
terminal band expanding widely on apical area and below
vein 2.
Ilab. Uganda, Entebbe (S’. A. Minchin)^ 1 ? type.
Exp. 30 mm.
100 a. Syntomis philippinensis, sp. n,
A. Head, thorax, and abdomen black-brown; frons and
vertex of head orange; antennse white at tips; tegulse
orange ; patagia orange, with black edges ; metathorax with
orange patch ; pectus and coxae mostly orange ; abdomen
with orange basal patch and bands on six following segments.
Fore wing black-brown ; an orange hyaline wedge-shaped
patch in cell, a wedge-shaped spot below base of cell, an
oblique wedge-shaped patch from below base of vein 2 to
near termen, an elongate streak above vein 6 and slight
streak above 7, and elongate spots above veins 4, 3. Hind
wing black-brown, with orange patch in and below cell
extending to near tornus, and a pair of spots above and
below vein 3.
$ . Vertex of liead black ; abdomen with five bands
beyond tlie basal patch. Fore wing with the patches above
veins 4, 3 and below 2 more elongate.
Hah. Philippines, Luzon, Manila {Fletcher) J', 2 7
type; Mindora (McGregor)., 1 Mindanao, Cagayancillo
[McGregor), 1 J. Exp. 22-26 mm.
223
Genera and Species of Syntomidse.
100 h. Spntoniis ticaonisj sp. n.
Black-brown ; antennae white at tips ; frons and tegulae
orange ; pectus with lateral orange patches ; fore femora
and tibiae streaked with orange; abdomen with basal orange
patch, followed by six bands in male, five in female. Fore
wing with wedge-shaped orange-hyaline patch in end of cell,
a triangular patch below base of cell, an oblique wedge-
shaped patch from b^low base of vein 2 to near termen, an
elongate streak above vein 6, sometimes with traces of a
streak above 7, and elongate spots above veins 4, 3. Hind
wing with orange patch in and below cell to near tornus and
pair of spots above and below vein 3.
Hah. Philippines, Ticao {McGregor) , 3 c? , 1 ? type.
Exp. 24-28 mm.
115 Z>. Sgntomis phceohasis, sp. n.
$ . Head and thorax black shot with metallic green ;
antennse white at tips ; fore coxae, first joint of fore and mid
tarsi, and first three joints of hind tarsi white; abdomen
metallic green, the three medial segments dorsally brilliant
blue with scarlet bands. Fore wing black shot with blue,
the base, basal half of cell, and costal area metallic green ;
a small quadrate hyaline antemedial spot below the cell, a
large spot in end of cell, an oblique narrow spot below base
of vein 2, and small elliptical postmedial spots above veins
6, 4, 3. Hind wing black shot wdth blue; a bar-shaped
hyaline antemedial spot below the cell and a small round
spot beyond lower angle.
Hab. Uganda, Nsadzi I. {E. A. MincMn)^ 1 $ type.
Exp. 32 mm.
132 h. Syntomis Cholmlei^ sp. n.
Black suffused with brilliant metallic blue ; shoulders with
orange spots ; fore femora and tibiae slightly streaked with
orange ; abdomen with dorsal orange patch on basal segment,
lateral orange patches on second and third segments, and
dorsal bands on fourth and fifth segments. Fore wing with
oblong semihyaline orange subbasal patch below cell, an
oblong patch in end of cell, an oblique wedge-shaped patch
below base of vein 2, and postmedial patches above veins
6, 4, 3. Hind wing with almost basal semihyaline orange
patch in and below cell and on inner area, and a rounded
patch beyond the cell above and below vein 4.
Huh. Br. E. Africa, Mola {Cholmley)^ 3 c? > 2 ? type.
Exp. 28-30 mm.
224
8ir G. F. Ilampsoii on new
2C0 a. Syntomis layosensis, sp. n.
Black ; frons yellowish white ; antennse white at tips ;
pectus and coxae with some yellowish-white scales; abdomen
with dorsal yellowish-white patch at base and band on fifth
segment, the ventral surface with the segments fringed with
yellowish white. Fore wing with wedge-shaped hyaline
patch in cell, an elongate patch below the cell from near base
to near termen, and elongate spots beyond the cell above
veins 6, 4, 3. Hind wing with hyaline patch below the cell
and spot above base of vein 2.
JIab. Lagos, Ebute Meta {Boay), 1 , 2 ? type. Exp.
26 mm.
Genus PAEALiETHiA, nov.
Type, P. sub/o7’micina, Beth.-Baker.
Proboscis fully developed ; palpi porrect, not extending
beyond frons ; antennae of male ciliated ; fore tibiae and tarsi
of male thickly fringed with scales on inner side; mid and
hind tibiae with the spurs short. Fore wing with veins 2, 3
from a point long before angle of cell ; 4, 5 from angle ; 6
from long below upper angle ; 7, 8, 9, 10, 11 stalked. Hind
wing with vein 2 from long before angle of cell ; 3, 4
absent; 5 from angle; 6, 7 stalked, 6 curved downwards.
271a. Apisa metarctiodes, sp. ii.
d . Head and thorax reddish fulvous ; palpi, frons, branches
of antennae, and tips of tegulae brown ; pectus and fringes of
hair on femora and tibiae brown ; abdomen fulvous, tinged
with brown towards extremity, a slight whitish dorsal band
at base, and subdorsal white spots on second segment ; the
ventral surface brown, with the first three segments fringed
with whitish. Fore wing reddish fulvous, the medial area
suffused with dark brown except at costa, narrowing below
the cell and embracing all the spots, tlie veins of terminal
area with dark streaks ; a quadrate semiliyaline white spot
in middle of cell and an irregularly elliptical spot below
middle of cell extending to below vein 1 ; a trifid spot beyond
upper angle of cell from above vein 7 to below 6 and a bifid
spot above and below vein 4 ; cilia dark brown. Hind wing
wdiitish, the inner area tinged with flesh-yellow^ and the inner
margin and terminal area suffused with brown ; a small
brown spot below end of cell above base of vein 2 ; cilia
225
Genera and Species of Syntomidge.
yellowish; the underside flesh-yellow, the disk whitish; a
curved brown discoidal band.
Huh. Uganda, Ruwenzori, 6000' [Euwenzori Exp.)^ 1
type. Exp. 64 mm.
287 a. Metarctia pidverea, sp. n.
$ . Head and thorax rufous ; abdomen greyish tinged with
rufous. Fore wing rufous thickly irrorated with black; a
small blackish spot above base of vein 2 ; a diffused blackish
discoidal patch. Hind wing greyish tinged with rufous.
Hah. Uganda, Ruwenzori, 6000' [Euwenzori Exp.)^ 1 ?
type. Exp. 42 mm.
290^. Metarctia, Jlaviciliata^ sp. n.
^ . Head and tegulae scarlet ; palpi, sides of frons, and
antennae fuscous ; thorax fuscous, the patagia with scarlet
edges towards base, the vertex with scarlet stripe; tibise and
tarsi fuscous ; abdomen scarlet, the last two segments and
ventral surface with fuscous bands. Fore wing fuscous
brown, the cilia ochreous yellow. Hind wing pale fuscous,
the base whitish, the inner margin yellow, with some scarlet
hair towards base ; cilia ochreous yellow. Underside of fore
wing yellowish, the costal and apical area to vein 2 suffused
with fuscous brown, the cell clothed with scarlet hair; hind
wing yellowish, with the costal area suffused with fuscous.
$ . Abdomen with narrow fuscous dorsal bands and broad
band just before extremity ; fore wing fuscous black, hind
wing fuscous, the cilia of both wings yellow, the underside
similar.
Hah. Uganda, Beni Semliki {Legge ds Wollaston),^ 1 ,
1 $ type. Exp.j S 4:8, ? 58 mm.
299. P seudapiconoma elegans.
? . Head and thorax blue-grey tinged with brown ; palpi,
sides of Irons, basal joint of antennae above, and back ot head
crimson ; pectus (except in front), sides of coxae, femora
above, and tibiae on inner side crimson ; abdomen blue-grey
tinged with brown, the first two segments with diffused sub-
dorsal orange bands, slightly connected dorsally on second
segment ; the next five segments with subdorsal black bands,
narrowing at middle; the lateral tufts of hair brown; the
ventral surface with crimson subventral patches on first six
segments and slight marks on anal segment. Fore wing pale
22G
Sir G. F. llampsoM on neto
ved-brown, tlie inner area and a faint bar across end of cell
tinged with blue-grey. Hind wing orange. Underside of
fore wing with the basal half of costal area, the cell, and area
below it to termen at vein 2 orange ; a crimson spot in upper
end of cell.
Hah. Uganda, Entebbe {Minchin'). Exp. 62 mm.
637 h. Earota nigricinctaj sp. n.
S . Head and thorax black-brown ; tegulse yellow at
sides; metathorax with yellowish-white bar; pectus with
yellow stripes below wings ; abdomen orange, a dorsal black
patch on first segment and dorsal bands on the other seg-
ments, the extremity black ; a lateral series of black spots;
the ventral surface yellowish white, with black bands. Fore
wing black-brown ; a basal yellow patch wdtli irregular outer
edge, angled inwards at vein 1 ; a wedge-shaped hyaline
patch in end of cell and quadrate patch below the cell ;
a postmedial series of five hyaline spots between veins 7 and 2,
the spots below veins 5, 4 extending to near termen, and the
spot below 3 minute and near termen. Hind wing black-
brown ; a small yellow patch at base with irregular outer
edge ; a hyaline patch beyond the cell between veins 7 and 2,
the spots above veins 6 and 2 small.
Hah. Argentina [0. W. Thomas), 2 type., Exp.
28 mm.
892 a. F seudaclytia Jlavidorsia, sp. n.
Head black, the vertex orange; thorax orange, the
outer edge of patagia, pectus, and legs black ; fore coxa3 with
white patches; abdomen black. Fore wing greyish, the
veins black, the inner area suffused with black ; the inter-
spaces of terminal area streaked with black. Hind wing
hyaline ; the veins black ; the inner area black, with slight
semihyaline streaks in submedian interspace; costal area
greyish ; termen black.
Venezuela, Caura Valley (A'%es), 2 cT type. Exp.
16 mm.
1073 a. Eucereon phazophlehiaj sp. n.
Hind wing of male with the inner area truncate towards
tornus ; a fold on underside containing a fringe of yellow
hair.
(^ . Head and thorax yellow-brown; palpi pink at base
and fuscous at sides ; coxa? pale pink ; abdomen fuscous
227
Genera and Species of Arctiadae.
brown, the ventral surface pale pink except at extremity.
Fore wing yellow-brown, the veins darker. Hind wing
fuscous brown.
Hah. Argentina (0. W, Thomas) ^ 1 type. Exp.
30 mm.
Arctiadae.
Nolinm.
13 a. Celama leucoscopula^ sp. n.
Head, thorax, and abdomen white, slightly tinged with
pale rufous ; anal tuft pure white. Fore wing white, tinged
in parts with pale brown ; antemedial line black, strong,
angled outwards in cell, then oblique, with brownish suffu-
sion before it; medial and postmedial oblique elliptical
patches from costa; postmedial line very ill-de{lned, bent
outwards below costa, tlien oblique ; traces of a sinuous sub-
terminal line. Hind wing white slightly tinged with brown
and with a faint discoidal spot.
Hah. Ceylon, Ambalangoda [J. Pole)^ 1 S type; Pera-
deniya, 2 ?; Matale, 1 ?; Pattalam, 1 $ ; Hambantota,
2 ? . Exp. 12 mm.
61a. JSola ochrographa^ sp. n.
Head and tegular ochreous white mixed with some
brown ; palpi and sides of frons black-brown ; tliorax brown,
mixed with black ; abdomen greyish suffused with black-
brown, the anal tuft tinged with ochreous. Fore wing
greyish suffused and thickly irrorated with dark brown, the
medial area suffused with red-brown below the cell and on
inner side of postmedial line ; antemedial line black, defined
by ochreous white on inner side, acutely angled outwards in
cell, then very oblique and ending at vein 1 ; the tuft of
scales in cell blackish, the tuft at upper angle greyish, with a
slight black line on its inner edge from costa, angled out-
wards below costa ; postmedial line ochreous white, defined
by black on inner side, very oblique and almost straight ;
subterminal line ochreous white, arising from costa just
beyond postmedial line, excurved to vein 6, then straight; a
fine dark terminal line; cilia fuscous, with a fine pale line at
base. Hind wing ochreous white, the costal and terminal
areas suffused with brown except towards tornus ; the under-
side irrorated with dark brown ; a dark discoidal lunule.
llab. Uganda, Entebbe {flinchin)^ 1 $ type. Exp.
22 mm.
228
Sir G. F. IIiiiiipsoii on new
I'l a. Nola hiconica^ sp. n.
Head and thorax grey-white irrorated with fuscous ; tarsi
fuscous ringed witli white ; abdomen grey tinged with fuscous.
Fore wing white irrorated with brown ; a brown fascia on
base of costa ; anternedial line arising from a small triangular
brown patch on costa, blackish, oblique below submedian
fold, and not reaching inner margin ; the tufts of scales at
middle and end of cell brown, the latter placed at extremity
of a small triangular costal patch, and with an oblique waved
line from it to inner margin ; postmedial line more or less
punctiform, angled outwards below costa, incurved to vein 6
and below 4 ; subterminal line strongly angled outwards at
vein 7 and excurved at middle ; some fuscous on termen.
Hind wing whitish tinged with fuscous ; the underside white
irrorated with browm ; a black discoidal spot.
Hah. Panama, Cana Mines (7)//eco^6), 2 Br. Guiana,
St. Jean Maroni {Schaus)^ 1 ; Brazil, Organ Mts.,Tijuca
(Wagrier)^ 1 ? type; Sao Paulo (Jones) y 1 Exp.
12-16 mm.
74 Nola argyrolepisj sp. n.
$ , Head and thorax ochreous white mixed with rufous
scales i; abdomen ochreous white tinged with rufous. Fore
wing ochreous thickly irrorated with rufous and with a few
black and silvery scales; subbasal line rufous, with a black
point at costa, sinuous, from costa to submedian fold ; ante-
medial line slight, irregularly waved, with black point at
costa; medial line rather diffused, angled outwards at discal
and submedian folds, a small black spot beyond it at upper
angle of cell ; postmedial line angled outwards below costa
and inwards at vein 6, excurved to vein 3, then oblique and
irregularly waved, with black point on it above vein 1; sub-
terminal line indistinct, rufous, angled outwards at vein 7,
excurved at middle, and ending at tornus ; a terminal series
of slight black points. Hind wing ochreous white, tinged
wn’th rufous.
llab. Mashonaland [Dohhie)y 1 $ type. Exp. 22 mm.
75 a. Nola poliotisy sp. n.
? . Head and thorax whitish mixed with fuscous, the
head whiter ; abdomen white suffused with grey. Fore wdng
grey-white irrorated with fuscous ; an indistinct dark ante-
medial line, very oblique from discal fold to inner margin ;
tufts of dark scales in upper part of cell towards extremity
229
Genera and Species 0/ Arctladse.
and at upper angle; an indistinct, oblique, dentate, postmedial
line with blackish points on the veins, somewhat excurved at
middle and incurved in submedian interspace ; a faint dentate
subterminal line, angled outwards at vein 7 and excurved at
middle ; a terminal series of minute dark points. Hind wing
white faintly tinged with brown ; the underside with the
costal area slightly irrorated with brown.
Hah. Transvaal [Chohnley\ 3 ? type. Exp. 26 mm.
76 nt. Nola leucalea^ sp. n.
? . Head and thorax white, sometimes tinged with pale
brown ; palpi brownish at sides ; abdomen white tinged with
brown. Fore wing white irrorated with brown and sometimes
tinged with pale brown ; antemedial line fine, dark, excurved
below costa and incurved at median nervure ; medial line
oblique towards costa, then slightly sinuous ; tufts of dark
scales in cell before antemedial line, in middle, and at upper
angle ; postmedial line slightly incurved from costa to vein 4,
then oblique and slightly sinuous ; subterminal line oblique
towards costa and excurved at middle ; a terminal series of
black points. Hind wing white tinged with brown.
Hah. Transvaal, Piet Retief {^Crawshay^ Janse')^ 3 7
type. Exp. 20-22 mm.
155 5. Roeselia pallidiceps, sp. n.
. Head and tegulge ochreous white; palpi and lower
part of frons black-brown ; thorax grey mixed with fuscous ;
abdomen grey. Fore wfing grey thickly irrorated with
fuscous brown, the terminal half slightly paler ; a dark
slightly curved medial line ; a black discoidal bar, the post-
medial line conjoined to its upper and lower extremities and
excurved beyond cell ; an ill- defined line from costa beyond
it joining the subterminal line at vein 4, and with a dark
striga from costa between them ; tlie subtcrminal line ill-
defined, slightly angled outwards at vein 7 and inwards at
vein 2. Hind wing grey thickly irrorated with fuscous ;
cilia with a fine pale line at base.
Hah. Ceylon [Alston)., 1 A type. Exp. 14 mm.
158 a. Zia ectrocta, sp. n.
^ . Head and thorax white ; legs blackish, the tarsi ringed
with white; abdomen white tinged wfith fuscous. Fore
wing pure white ; a small black spot on costa near base ; a
medial triangular black patch from costa to origin of vein 2,
ii30 Sir G. F. Ilampson on new
its outer edge excised in cell ; two small discoidal tufts of
raised scales with a few dark scales round them; postmedial
line black with small tufts of raised metallic scales on it,
oblique and obsolescent from costa to vein 6, slightly incurved
at discal fold and strongly below vein 4, the area beyond it
rufous except at apex ; subterminal line represented by a
dark point on costa, then on the rufous area white, defined on
inner side by black from below apex to vein 3, excurved
below vein 7 and at middle and below vein 2 angled inwards
to near postmedial line; cilia rufous intersected with white.
Hind wing white, the terminal area slightly tinged with
brown from apex to vein 3.
Ab. 1. Fore wing with tlie postmedial line more angled
inwards below vein 4 and with large black patches beyond
it on inner area and at middle, the latter connected with
terraen below apex by an oblique black fascia.
Hah. Ceylon, Haputale (^Mackwood), 1 type, Maskeliya
[de Mowbray) j 1 , Exp. 20-24 mm.
Lithosianjl.
222 a. Tigrioides termineola^ sp. n.
$. Head and thorax fulvous orange; palpi, frons,tibi8e,and
tarsi blue-black ; antennae blackish; pectus mostly fuscous ;
abdomen yellow with lateral fuscous stripes. Fore wing-
fulvous orange ; a black streak in submedian fold from
middle to termen ; a terminal blue-black band, expanding
below apex and narrow towards tornus ; cilia blue-black.
Hind wing pale ochreous, the veins brownish ; a dark ter-
minal line expanding into a narrow purplish-fuscous band
towards apex, cilia purplish fuscous except towards tornus.
Hab. Ashanti, Kumassi {Whiteside)^ 1 $ type. Exp.
28 mm.
254 a. Ilema barbata, sp. n.
Antennae of male serrate with fascicles of long cilia; fore
wing with the cell clothed with long downturned scales
on upperside, narrow, veins 3, 4 curved downwards, the
discocellulars oblique, an areole.
d. Head and thorax orange; antennae tinged witli
fuscous ; abdomen greyish yellow. Fore wing orange-
yellow, the large downturned scales in cell rather deeper
orange ; a small brownish sj)Ot on costa above end of cell.
Hind wing pale yellow.
Genera and Species of Arctladie. 231
Ilah, Philippines, Luzon, Benguet Prov., Irisan
{McGregor) j 1 type. Exp. 20 mm.
• 301a. llema pentaspilaj sp. n.
? . Head, thorax, and abdomen oclireous ; fore tibiae and
tarsi suffused with fuscous. Fore wing ochreous ; a post-
medial somewhat oblique series of five small black spots,
below costa, in end of cell, a spot below veins 3, 4 displaced
outwards, a short streak-like spot in submedian fold and a
spot above inner margin. Hind wing pile ochreous.
Hab. Singapore {Ridley), 1 $ type ; Borneo, Kuching
{Shelford), 2 $ , Exp. 26-30 mm.
361 h. Ilema atrifrons, sp. n.
Head, tegulae, patagia, pectus, and legs fulvous
yellow ; palpi, frons, and fore legs in front fuscous ; dorsum
of thorax and base of abdomen grey-white, the rest of abdomen
yellow. Fore wing yellow with a whitish suffusion. Hind
wing whitish yellow.
Hah. Nicobars [G. Rogers), 1 type. Exp. 26 mm.
Genus Metagylla, nov.
Type M. miroides.
Proboscis fully developed ; palpi upturned, short, the
second joint fringed in front with short rather downturned
hair, the third small with pointed tuft in front; antennae of
male with bristles and cilia; tibiae with the spurs moderate ;
abdomen of male with large dorsal and lateral anal tufts of
hair. Fore wing ratiier narrow ; veins 3, 4, 5 from angle
of cell ; 6 from below upper angle ; 7, 8, 9 stalked, 7 from
before 9 ; 10, 11 from cell. Hind wing with vein 3 shortly
stalked with 4, 5 ; 6, 7 coincident ; 8 from middle of cell ;
male with a patch of androconia and fringe of long hair in
and beyond end of cell on upperside.
392 a. Metagylla miroides, sp. n.
$. White; head and thorax above and fore tibiae and
tarsi in front tinged with fuscous ; palpi with the tuft on
third joint fuscous ; abdomen with the dorsal anal tuft
fulvous, the lateral tuft fuscous. Fore wing with the costal
area and inner area to cell and vein 2 sliglitly tinged with
232
Sir G. F. Hampson oji new
fuscous. Hind wing with tlie patch of androconia deep
rufous with some ochreous suffusion beyond it, the fringe of
hair ochreous.
Hah. Fr. Guiana, St. Jean Maroni (^Schaus)^ 1 ^ type.
E.rp. 28 mm.
‘522 a. Aglossosia latifusca, sp. n.
^ . Head and thorax deep orange ; antennse blackish, the
shaft whitish above ; palpi fuseous above ; legs striped with
fuscous ; abdomen orange, with black dorsal stripe and seg-
mental bands except at base. Fore wing fuscous; a yellow
fascia on costa narrowing to a point before apex; a yellow
fascia on inner margin; cilia yellow. Hind wing pale yellow,
the costal area suffused with fuscous or suffused irregularly
with fuscous to submedian fold, leaving the inner area and
termen yellow. Underside of both wings with slight dark
discoidal spot.
Hah, Uganda, Ketoma, 5000' {Doggett) 2 J' type.
Exp. 40 mm.
523 a. Caripodia albescens, sp. n.
$ . Head orange-yellow ; thorax and abdomen whitish
tinged with yellow; pectus, legs, and ventral surface of
abdomen orange-yellow. Fore wing white, tinged with
yellow on costa and inner margin. Hind wing white, tinged
with yellow. Underside suffused with orange-yellow.
Hah. Nigeria [Capt. Richardson), 1 ? type. Exp.
30 mm.
586 a. Flalone flavinigra, sp. n.
^ . Head, thorax, and abdomen fuscous, the vertex of
head, base of shaft of antennae, and tegulae yellow. Fore
wing orange-yellow with irregular oblique outer edge ; a
postmedial black band angled inwards below cell and with
rather dentate edges; some diffused fuscous before termen.
Hind wing pale fuscous.
Hah. S. India, Palni Hills, 6000' (IF. H. Campbell),
1 type. Exp. 20 mm.
894 a. Neasura taprohana, sp. n.
A . Ochreous yellow ; antennce at tips, fore legs in front,
and extremities of mid and hind tibice fuscous. Fore wing
233
Genera and Species of Arctladye.
witli black point in base of cell; the costa fuscous to the
curved diffused antemedial line; a blackish discoidal point
on some fuscous suffusion ; postmedial line very diffused and
ill-defined, waved, emitting streaks inwards on the veins and
outwards on veins 7, 6, 4. Hind wing with slight fuscous
suffusion below apex.
Hah. Ceylon, Maskeliya {J. Pole)^ 1 type. Exp.
24 mm.
897 h. Trlcholepis xanthopera^ sp. n.
$ . Head and thorax fulvous yellow; abdomen yellowish,
dorsally tinged with brown. Fore wing pale brown, the base,
costa, and terminal area pale yellow. Hind wing pale yellow
tinged with brown except terminal area.
Hah. Singapore {Ridley)^ 1 S type. Exp. 16 mm.
937 a. Asura toxodes^ sp. n.
d . Head and thorax pale ochreous slightly mixed with
fuscous ; antennse and extremities of tibiae fuscous ; abdomen
ochreous white. Fore wing pale ochreous; costal edge
blackish on basal and terminal areas ; a black point in base
of cell ; some fuscous in subraedian fold ; a highly curved
antemedial line ; a medial line angled inwards in cell ; post-
medial line confluent at costa and inner margin with the medial
line, with which it forms a bow-shaped mark, very oblique
from costa to vein 6 and from 4 to inner margin; a very
irregular subterminal line angled outwards at veins 6 and 4 ;
a fine black terminal line. Hind wing pale semihyaline
ochreous.
Hah. Andamans (^G. Rogers)^ 1 d fjp®* Exp. 24 mm.
966 a. Asura phantasma, sp. n.
A- Whitish ochreous; antennae and fore legs in front
fuscous. Fore wing with the base of costa black ; a small
black spot in base of cell ; an indistinct antemedial series of
spots strongly excurved in cell and less so below it, some-
times almost conjoined into a line ; a medial line oblique
from costa to subcostal nervure, then excurved, often almost
obsolete ; a small discoidal spot ; a postmedial series of points,
sometimes almost obsolete, those on veins 6 and 4 nearer
termen ; one or two points on terrnen sometimes present.
Hind wing pale ochreous, the apex sometimes faintly tinged
with fuscous.
Hah. Andamans {G. Rogers), .5 A type. Exp. 18 mm.
Ann. dj Hag. N. Hist. Scr. 7. Fu/. xix. 16
234
Sir G. F. Ilampson on new
1017 a. Miltochrista ocellata, sp. n.
$. Head and thorax orange-yellow ; patagla and prothorax
with black spots ; tibiae banded with black, the last joint of
tarsi black ; abdomen greyish ochreous, the ventral surface
blackish. Fore wing orange-yellow, small black spots at
base of costa and cell ; fuscous spots below costa, cell, and
above vein 1 before the antemedial line, which is interrupted
at submedian fold and angled inwards above inner margin ;
a large annulus at end of cell ; postmedial line strongly bent
outwards below costa, then highly and irregularly dentate,
strongly incurved below vein 4 and conjoined to antemedial
line above and below submedian fold ; a subterminal series
of small spots on the veins. Hind wing yellow.
Hah, Ceylon, Ohiya [de Mowbray) j 1 $ type. Exp.
40 mm.
1037 a. Miltochrista citrona^ sp. n.
? . Head, thorax, and abdomen orange-yellow ; antennae
and fore legs in front fuscous. Fore wdng pale orange-
yellow ; small black spots on base of costa and in cell ; ante-
medial part of costa streaked with black ; a curved series of
four wedge-shaped black spots before the curved, maculate
antemedial black line; traces of a curved orange medial line;
a black discoidal spot; postmedial black line quadrately
angled outwards betw^een veins 6 and 4, with a series of
strong black streaks on the veins from it to termen ; cilia
black. Hind wdng pale orange-yellow, with short black
streaks on veins towards apex ; cilia blackish on apical
half.
Hah, SiNGAPOEE (Ridley) j 1 ? type. Exp. 24 mm.
1103 a. Palpidia melanotrichay sp. n.
Palpi hardly reaching above vertex of head, the second
joint moderately scaled in front ; antennae of male ciliated ;
fore wing with vein 10 stalked with 7, 8, 9 ; hind wing with
veins 6, 7 stalked.
. Head, thorax, and abdomen white mixed with ochreous ;
palpi with fuscous patch on second joint at sides; tarsi
slightly banded with fuscous. Fore wing white, finely
striated with golden brown and irrorated with long black
hair-like scales on basal, medial, and postmedial areas ; ante-
medial line dark brown, from costa to submedian fold,
straight; postmedial line red-brown, oblique from costa to
235
Oenera and Species of Avctiadae.
discal fold, tlien slightly incurved; an ill-defined, waved, white
subterminal line, excurved below costa and at middle, angled
inwards at discal and submedian folds; a fine black terminal
line ; cilia white, the tips tinged with brown. Hind wing
white, the veins, inner and terminal areas tinged with brown;
a fine black terminal line ; the underside with the costal area
irrorated with brown, a black discoidal striga and interrupted
maculate fuscous postmedial and subterminal lines.
Hah. Jamaica, Runaway Bay (Walsingham) ^ 2 S Ijpe*
Exp. 16 mm.
Arctianm,
1677 a. Mcenas ramosa^ sp. n.
? . Head and thorax white ; palpi yellow ; antennae black;
tegulae slightly edged with pale brown ; patagia and vertex
of thorax streaked with pale red-brown ; legs streaked with
brown; abdomen orange, the anal tuft and ventral surface
white, a dorsal series of black spots except at base and
extremity, subdorsal black points on three medial segments
and a sublateral series. Fore wing white ; a red-brown
fascia in base of cell, then along subcostal nervure and to
costa before apex expanding at angle of cell and at costa ;
a fascia along median nervure and to termen below vein 6,
emitting branches on basal half of veins 2, 3, expanding
and partially enclosing a white spot above middle of
vein 4 ; a fascia on vein 1 ; an oblique fascia from apex to
middle of vein 6 ; a spot on termen at extremity of vein 2.
Hind wing white tinged with yellow; the underside with
minute discoidal point.
Hah. German E. Africa, Dar-es-Salaam, 1 ? type.
Exp. 34 mm.
1761 cr. Diacrisia hypogopa^ x\,
$ . Head and thorax pale ochreous, the vertex of thorax
with black streak; palpi black, the first joint orange; frons
black ; femora crimson above, the fore and mid tibiae fuscous,
the hind tibiae streaked with fuscous, the tarsi fuscous; abdomen
orange, with dorsal and lateral series of black spots, the
ventral surface ochreous with sublateral series of black points
except on basal segments. Fore wing ochreous ; a black
point at base ; an antemedial oblique series of small elongate
black spots from costa to median nervure and spots on inner
area above and below vein 1 ; a point at upper angle of cell
with oblique spot from costa above it and a point just beyond
16*
2.3T) Sir G. F. Ilampson on new
lower anj^le; traces of an oblique postmedial line witli black
point below vein 4 just beyond lower angle of cell, points
above and below veins 3 and 2 and spots above and below
vein 1, the spot on inner margin slightly confluent with the
antemedial spot ; an oblique series of black points from costa
near apex above and below veins 8, 7, 6, and points near ter-
inen above and below veins 5 and 4. Hind wing pale yellow ;
a black discoidal spot and a subterminal series of spots above
and below veins 7, 5, 2 and on vein 1 ; cilia white.
Hah. Singapore [Ridley)^ 1 ? type. Exp, 56 mm.
1765 a. Diacrisia lioloxaniha^ sp. n.
$ . Head and thorax brownish orange ; palpi black at
tips ; fore and mid tibiae black, hind tibiae black at base and
extremity, the tarsi black ; abdomen bright orange with
dorsal, lateral, and sublateral series of black points. Fore
wing uniform brownish orange. Hind wing orange-yellow.
Underside of both wings with blackish discoidal points and
terminal points above and below vein 5.
Hah. Nigeria, Old Calabar [Sampson)^ 1 ? type. Exp.
44 mm.
1794 a. Diacrisia melanodisca^ sp. n.
Head and thorax deep rufous; antennae whitish;
patagia with black spots and paler tips ; tibiae and tarsi dark
brown ; abdomen dull orange with dorsal and lateral series of
blackish spots. Fore wing buff tinged with rufous, especially
towards costa ; a black point at base ; an antemedial series
of black points, angled outwards just below cell ; a large
blackish patch in end of cell confluent with a medial maculate
band, which is angled on median nervure ; an indistinct post-
medial line with dark points on the veins, oblique from costa
to vein 6, incurved at discal fold and oblique below vein 4,
some small black spots beyond it towards costa and from
vein 3 to inner margin, met at vein 6 by an oblique fascia
from costa near apex ; cilia with a series of black spots.
Hind wing pale dull yellow with small black discoidal lunule
and subterminal spots at veins 5, 2, 1.
$ . Fore wing with ground-colour bright rufous irrorated
with brown.
Hah. Uganda, Kuwenzori, 6000' [Ruwenzori Exp.)y 1 (^ ,
1 7 type. Exp., S 38, 7 40 mm.
Genera and Species of Arctladce.
237
1814 Diacrisia coccinea^ sp. n.
J . Head white ; palpi scarlet, the third joint blackish;
antennse black; tegula3 and patagia white edged with scarlet,
the latter with black spot; thorax scarlet with pair of dorsal
white streaks ; pectus scarlet ; legs white, the femora scarlet
above, the tibiae and tarsi streaked with brown ; abdomen
scarlet, with dorsal, lateral, and sublateral black spots on
medial segments, the ventral surface white. Fore wing
scarlet, the interspaces of disk thinly scaled, the costa and
veins towards base streaked with w'hite, the inner area with
slight blackish irroration ; small antemedial black spots
below costa and above and below vein 1 ; small postmedial
spots above and below vein 1 ; cilia whitish. Hind wing
semihyaline, the costal and inner areas and termen suffused
with scarlet.
$ . Fore wing with the disk not thinly scaled ; hind wing
wholly scarlet, the cilia whitish.
Hah. Philippines, Luzon, Benguet Prov., Irisan
[McGregor) j 1 (5" ? 1 ? fjpe. Exp.^ d mm.
1824 a. Acantharctia airiramosay sp. n.
$ . Head and thorax ochreous white ; antennse black,
ochreous at base; legs black, the femora above orange;
abdomen pale orange tinged with brown, the base white, the
extremity and ventral surface greyish, lateral series of black
spots and sublateral stripes. 4’ore wing ochreous white ;
a black fascia from middle of subcostal nervure to apex,
emitting a short spur on vein 6 ; a black fascia on median
nervure, emitting liner branches on veins 4, 3, 2 ; a black
streak on vein 1 to termen. Hind wing white.
Hah. Uganda [Doggett)^ 1 ^ type. Exp. 44 mm.
1860 Z>. Estigmene neuriastisy sp. n.
S . Head whitish ; palpi and antennae blackish ; thorax
greyish, the tegulae and patagia edged with orange; tibiae
and tarsi brownish streaked with black ; abdomen orange
with dorsal black segmental bands, the ventral surface
yellowish white with lateral and sublateral series of small
black spots. Wings yellowish white, the veins finely streaked
with blackish ; fore wing with the margins rather yellower.
Hah. Angola, Bihe, 1 type. Exp. 44 mm.
238
Sir G. F. Hampsoii on new
1870 Estigmene flavice;ps j sp. n.
. Head orange-yellow ; palpi and antennae black ;
thorax white ; pectus and legs orange-yellow, the tibiae and
tarsi striped with black; abdomen orange-yellow, the basal
segment and ventral surface white, a dorsal series of black
bands except at base and extremity, and lateral series of
black spots and sublateral series of points except at base and
extremity. Fore wing white, the costal edge orange-yellow.
Hind wing white.
Hab. Sierra Leone [Quinton)^ 2 S type. Exp, 36 mm.
1878 a. Pericallia nephelistis, sp. n.
$ . Head and thorax grey mixed with fuscous brown ;
patagia, meso- and metathorax with paired blackish spots;
pectus and legs yellow, the tibiae and tarsi greyish, the
latter banded with black; abdomen yellow with dorsal series
of black patches, except on two basal segments, and lateral
series of spots. Fore wing brownish grey, mostly clouded
with fuscous blotches and spots, some of them forming a
subbasal band from costa to submedian fold with two small
spots beyond it in submedian interspace, an antemedial
band angled outwards in cell, then oblique to vein 1, a
medial patch in and below cell connected with a patch on
costa, some confluent markings on middle of inner margin;
a diseoidal patch confluent with a patch on costa, and some
very irregular and partly confluent postmedial and sub-
terminal spots; a scries of small spots on termen and cilia.
Hind wing pale yellow, with diseoidal blackish spot and
point at lower angle of cell ; two small spots on basal part of
vein 2 and two irregular spots on vein 1 ; irregular sub-
terminal patches from costa to vein 4, vein 3 to submedian
fold and above tornus.
Hah, Angola, Bihe, 3 $ type. Exp, 52 mm.
1994 a. Turuptiana sanguinea^ sp. n.
Head, thorax, and abdomen black ; femora crimson above ;
abdomen with crimson subdorsal fasciae not reaching base.
Fore wing black ; an irregular curved antemedial crimson
band expanding on costa towards base and extending to
vein 1 ; a medial, ratlier triangular band from costa to
vein 1, indented by black streaks on the veins; a small spot
beyond the cell ; a curved postmedial band from costa to
vein 1 with slight dark streaks on the veins; apical part
239
Genera and Species of Arctiadae.
of costa and cilia pale yellowish tinged with crimson. Hind
wing crimson, a little black at base; a discoidal limule;
a terminal black band narrowing to a point at vein 1.
Ilah, Bolivia, La Baz, 9000', 1 c^, 1 $ type. Exp,
28 mm.
2028a. Antarctia rhodosoma^ sp. n.
d . Head fuscous with tufts of orange hair on basal joint
of antennas ; palpi crimson with the third joint black ; tegulae
black, with dorsal crimson streak; patagia orange; thorax
black; pectus and legs dark brown, the coxae and femora
crimson ; abdomen crimson, with dorsal and lateral series
of small black spots, the ventral surface black. Fore wdng
pale reddish brown, the base yellowish, the subbasal area,
cell, and a discal patch tinged with fuscous. Hind wing
pale semihyaline brown, the cell and a streak in discal fold
sliglitly tinged with fuscous.
Hah. Chili, Maquahue, Temuco {Middleton)^ 1 S tjpe.
Ex^). 38 mm.
2034 a. Antarctia atrifascia^ sp. n.
d . Head and thorax rufous, tinged with brown ; palpi
blackish ; frons and pectus in front with blackish hair
mixed ; tore tibiae blackish fringed with rufous hair ; abdomen
ochreous suffused with brown. Fore wing brownish ochreous
irrorated with fuscous; a diffused black streak below
median nervure to origin of vein 2 ; a slight streak in lower
part of cell from middle to above middle of vein 5 ; a black
discoidal point ; a terminal series of small black spots.
Hind wdng pale ochreous suffused and irrorated with fuscous.
Underside of fore wing suffused with fuscous ; hind wing
with black discoidal point.
Hab. Bk. E. Africa, Aberdare Range {R. Ford)^ 1 S
type. Exp. 36 mm.
2086 a. Utetheisa pulchelloideSj sp. n.
Differs from U. pulchella in the antennae of male being
serrate instead of ciliated and in the hind wing having the
fold and tuft on inner area.
It varies much in the same way as U. pulchella^ but never
seems to lose the black spots of fore wing, which usually has
the ground-colour rather white ; in specimens from the
Rew Hebrides and Solomons the black terminal band on
hind wing is largely developed, and it appears to be conhned
to Oceanic and other islands and to N. Australia.
240
Sir G. F. Hampson on nev)
Hah. Seychelles, Praslin (Fletcher), 2 ^ , 1 $ ; Car-
GADOS CarejoS (Fletcher), 1 ^ COETIVY (Fletcher), 1 ,
1 $ ; Amirantes {Fletcher), Poivre, 1 , 1 $ , D’Anos, 1 ,
St. Josepli, Eagle, 1 c? j Desroches ; Chagos Is., Peros
Banlios (Fletcher), 1 ^ type, Salomon Atoll, Diego Garcia ;
Ceylon, Kandy (Green), 3 , 2 Peradenyia, Ham-
Lantota, Trincomali (Fletcher), 1 , 1 $ ; Cocos-Keeling I.
(Darwin, Wood- Jones), 3 ^ ; CHRISTMAS I. (Andrewes) , 1 ^ j
2 ?; Singapore (Ridley), 1 (^, 1 ?; Formosa (Seehohm,
Dickson, Ilohson), 1 , 5 ?; LOO-CHOO Is. (Fryer), 1
New Guinea (Mathew), 1 , 3 ? ; N. Australia, Baudin I.
(J. J. Walker), 1 , 1 $ , Pt. Darwin (J. J. Walker), 3 S i
2 Queensland, Cooktown (de la Garde), 2 $; SOLO-
MON Is., Alu (Woodford, Mathew), 2 S', Gilbert Is.
(WoodfojA), 2 S', Marshall Is. (Mathew), IS, 2 ?;
Ellice Is. (Woodford, Mathew), 4 S, I ?• Exp.
34-44 mm.
I am indebted to Paymaster T. Bainbrigge Fletclier for
pointing out the distinctions and suggestiug the name of
this species. The larva feeds on Tournefortia argentea.
Specimens from the mainland of Africa and Asia, Ceylon,
Colombo, Nicobars, Java, New Guinea, Port Moresby, New
South Wales, Tasmania, and Fiji are IJ. pulchella.
2087 a. Utetheisa pectinata, sp. n.
S . Differs from U. pulchella only in the male having the
antennae bipectinate with very short branches, and the hind
wing having the fold and tuft on inner area.
Hah. N. Australia, Port Essington, 1 S Exp.
36 mm.
This species belongs to the same section of the genus
as U. antennata, which also has the fold and tuft on inner
area as well as the pectinate antennae.
2107 a. Rhodogastria atrivena, sp. n.
$ , Pure white; palpi tinged with yellow, black above;
fions black at sides and with two black spots above; vertex
of head with black point ; antennae brownish ; tegulae,
shoulders, patagia, and piotliorax with pairs of small black
spots, those on tegulae larger ; legs yellow, the knees with
black spots; abdomen with lateral black points except on
basal segments and minute sublateral black streaks on two
medial segments. Fore wing with fine black-brown streaks
241
Genera and Species of Agarlstidse.
on tlie veins and minute discoidal point. Hind wing with
minute discoidal point.
Hah. Uganda, Entebbe {Minchin)j 1 ? type. Exp.
60 mm.
Agaristidse.
81. Xanthospilopteryx llornimani,
Subsp. Minchini^ nov.
? . Fore wing with the antemedial and medial yellowish-
white patches confluent, leaving a slight black streak on
middle of subcostal nervure with small spots below it in and
below middle of cell ; a yellowish-white streak above inner
margin, except towards base and tornus ; the oblique band
beyond the cell much broader, a fascia between veins 3, 2
from below end of cell to near termen ; the spot above tornus
slight. Hind wing with the terminal band narrower and
with slightly waved inner edge.
Hah. Uganda, Entebbe {Mincliin')^ 1 ? type. Exp.
82 mm.
Genus Acantuerta, nov.
Type, A. {Tuerta) thomensis, Jord.
Proboscis fully developed ; palpi upturned, the second joint
fringed with long hair in front, the third porrect and some-
what dilated at extremity ; frons with truncate conical
prominence wdth raised rim in front and corneous plate below
it, very narrow above between the eyes, which are naked ;
antennae somewhat dilated towards extremity; mid and hind
tibiae spined ; abdomen with dorsal crests on first two seg-
ments. Fore wung wflth veins 3 and 5 from near angle
of cell ; 6 from upper angle ; 9 from 10 anastomosing with 8
to form the areole ; 11 from cell. Hind wdng with veins 3, 4
from angle of cell ; 5 obsolescent from middle of disco-
cellulars ; 6, 7 from upper angle, 8 anastomosing with the
cell near base only.
161 a. Tuerta cyanopasta^ sp. u.
7 . Head and thorax black-browm ; pectus pale orange at
sides; fore tibiae and the tarsi with greyish rings; hind
tibiae with the basal half mostly orange and fringed with
orange hair; abdomen orange, the dorsal crests black, the
extremity blue-black, the ventral surface w'ith the basal half
brownish, the terminal half blue-black and forming an
242
Sir G, F. Hampson on new
elliptical depression. Fore wing deep chocolate-brown
suffused with silvery blue to the subterminal line ; the base
of inner margin black-brown ; a waved antemedial black bar
from costa to median nervure and an almost medial waved
bar from submedian fold to inner margin ; blue stigmata
strongly defined by black in end of cell and on discocellulars,
the former oblique elliptical and conjoined at lower ex-
tremity to the latter, which is lunulate and dilated at lower
extremity ; postmedial line strong, black, excurved below
costa and at middle, angled inwards at discal fold, strongly
incurved below vein '6 to lower edge of reniform, then
waved ; the outer edge of the blue area dentate, with small
blackish spots in the interspaces, angled outwards at
veins 6, 4, 3, then incurved; a terminal series of small
black spots and a fine waved terminal line; cilia bluish
fuscous with a slight pale line at base. Hind wing deep
oiange with black-brown terminal band, its inner edge in-
curved at discal fold, narrowing to a point at tornus; cilia
with a fine pale line at base. Underside of fore wing orange,
except costal area and the broad terminal area, a black spot
in end of cell and broad oblique discoidal bar to the terminal
band at vein 2; hind wing with golden-brown suffusion on
costal half except at base, and before the terminal band
except towards tornus.
Ilab, Be. E. Afeica, Njoro ( Cholmley), 1 ? type. Exp.
46 mm.
NoctuidsB.
A GROUND.
58 a. CJiloridea jiavigera^ sp. n.
$ . Head, thorax, and abdomen yellow, the thorax
suffused with pale pink, the abdomen pale at base with twm
pink crests; palpi suffused with purplish red, tibiae and
ventral surface of abdomen irrorated with red. Fore wing
yellow with a slight greenish tinge; the costa suffused with
red-brown, a subbasal red-brown striga from costa and point
below cell ; an antemedial red-brown striga from costa and
traces of a waved line; claviforrn with its extremity faintly
defined ; orbicular with brown point in centre and a faint
round circumference ; reniform with grey centre defined by
red-brown and traces of brown circumference, confluent with
a brown mark from costa above it; postmedial line very
indistinct, bent outwards below costa, then minutely waved,
incurved below vein 4 ; subterminal line with brown tri-
243
Genera and Species o/Noctuidse.
angular patch on costa, then very indistinct, angled outwards
at vein 7 ; cilia red-brown, with series of darker lunules and
greyish line near tips. Hind wing yellow, with slight
discoidal lunule and diffused sinuous subterminal dark band,
extending to terinen below apex. Underside of both wings
with the costal and terminal areas irrorated with rufous ;
fore wing with a point in cell, dark discoidal spot, curved
postmedial and indistinct subterminal lines ; hind wing with
slight discoidal lunule, curved postmedial line from costa to
vein 2 with minute dark streaks on the veins, and diffused
subterminal band.
Hah. Khodesia, Buluwayo {Marshall)^ 1 ? type. E.vp.
32 mm.
175 a. Timora latimgraj sp. n.
Head and thorax bright rufous; antennae whitish at base,
blackish at tips; a tuft of blackish hair below patagia;
palpi, frons, tibiae, and tarsi black; abdomen greyish fuscous,
the anal tuft pale rufous. Fore wing ochreous tinged with
cupreous red, especially on costal area ; the costal edge pure
white ; a broad black fascia on median nervure, attenuate
towards base and extending to near termen ; an antemedial
black point on vein 1 ; a postmedial series of small black
spots except towards costa, oblique below vein 4 ; some
black on costa towards apex ; a terminal series of small
black spots ; cilia pale crimson. Hind wing ochreous white
tinged wdth brown, especially on terminal area ; a terminal
series of black points ; cilia pale crimson ; the underside
whitish.
Bab, Uganda, Mulema {Doggett\ 17 J', 4 7 type. Exp.
30 mm.
738 a. A gratis eloiopis, sp. n.
7 . Head and tegulae red-brown ; thorax fuscous ; palpi
fuscous ; pectus, legs, and abdomen whitish tinged with
brown. Fore wing red-browm with slight dark irroration ;
subbasal line slight, dark, from costa to submedian fold ;
antemedial line indistinct, double, sinuous, rather punctiform ;
claviform absent; orbicular and reniform small, fuscous with
whitish annuli slightly defined by black, the former elongate,
acute at extremity, and touching the latter ; postmedial line
represented by a double series of black points, bent outwards
below costa, excurved to vein 4, then oblique ; subterminal
line absent ; cilia rufous with a fine whitish line at base.
Hind wing white with dark discoidal point, postmedial series
of minute streaks on the veins and fine terminal line ; the
214
Sir G. F. Ilampson on neio
underside witli the markings more prominent^ the costal area
slightly irrorated with fuscous.
Ilah. Uganda, Kuwenzori, 6000^ [Ruwenzori 1 $
type. Exp. 30 in in.
795 a. MetaJepsis faegensis^ sp. n.
. Head and thorax clothed with white and brown hair ;
palpi black at sides; lower part of frons black and brown, a
y('llow mark between antennae, which are black; tegulae with
black medial line followed by a brown patch ; patagia with
brown line near U))per edge ; dorsum of thorax mostly
brown ; abdomen grey, suffused with brown. Fore wing
grey-white, suffused with rufous in the interspaces, except on
costal area; the veins slightly streaked with black and
prominently defined by grey except on terminal area ; a
short black and rufous streak below base of costa ; a black
streak below base of cell and a stronger streak above inner
margin before middle; antemedial line obsolete; claviform
represented by a slight white streak with short black streak
beyond it ; orbicular and roniform small, white with a black
streak below them, the former elongate elliptical, the latter
a minute lunule; a short black streak beyond upper angle of
cell ; postmedial line obsolete ; a siibterminal series of small
black spots in the interspaces interrupted by the grey streaks
defining veins 7, 6, 4, 3 ; cilia white with a brown line
through them. Hind wing grey, suffused and irrorated with
fuscous ; cilia white with a fuscous line through them ; the
underside paler.
Hah. Tierra del Fuego, Cheena Creek (^Crawshay)^
1 $ type. Exp. 30 mm.
817 a. EpisiUa clavata^ sp. n.
^ . Head, thorax, and abdomen dark reddish brown
mixed with grey ; tarsi with pale rings. Fore wing grey
tinged with red-brown, the medial area red-brown exce[)t
towards costa and inner margin ; subbasal line represented
by a black striga from costa ; a strong sinuous black streak
below base of cell with yellow streak above it to the clavi-
form, which has a yellowish annulus denned by black and
intersects the oblique sinuous antemedial line ; orbicular and
reniform grey with brownish centres and defined by black,
the former oblique elliptical, open above, the latter a narrow
lunule very strongly angled on median nervure to below
orbicular, some blackish in cell before and between them ;
245
Genera and Species of NoctuI(I[ie.
postmediul line strongly bent outwards below costa, tlien
dentate, strongdy incurved below vein 4; subterminal lin.i
very indistinct, greyish, slightly angled outwards at vein 7
and excurved at middle, the veins beyond it with slight dark
streaks ; a terminal series of slight brown lunules ; cilia
with fine brown line near base, blind wing grey suffused
and irrorated with brown, a dark terminal line; the under-
side witli dark discoidal lunule and diffused curved postmedial
line.
IJah. Punjab, Kulu {Dudgeon)^ 1 c? lyp^. E.cp.
30 mm.
856 c. Episilia arenacea^ sp. n.
d . Head and thorax pale brownish ochreous ; tarsi
fuscous with pale rings ; abdomen pale ochreous, dorsally
irrorated with fuscous. Fore wing pale brownish ochreous
slightly irrorated with fuscous ; a double, waved, subbasal
line from costa to submedian fold ; antemedial line in-
distinctly double, oblique, strongly waved, interrupted ;
orbicular and reniform with slight yellowish annuli in-
completely defined by fuscous, the former round, the latter
large ; postmedial line double at costa, then indistinct, bent
outwards below costa, then dentate and produced to a series
of black points on the veins, oblique below vein 4, some pale
points beyond it on costa; subterminal line ochreous white,
slightly defined by fuscous on inner side at costa, then by
slight dentate marks, angled outwards at vein 7 and slightly
excurved at middle ; a terminal series of black points.
Hind wing whitish suffused with pale brown; cilia yellowish
white ; the underside white, the costal area tinged with
ochreous, a small discoidal spot and punctiform postmedial
line.
Ilah, BeloochisTAN, Quetta {Nurse), 1 type. Exp,
46 mm.
867 a, Episilia rhodopea^ sp. n.
. Head and thorax fuscous slightly mixed with grey
and rufous ; abdomen whitish suffused with brown. Fore
wing red-brown with slight dark irroration ; subbasal line
represented by double oblique striae from costa ; antemedial
line with double black strim from costa, then minutely
dentate, above inner margin angled outwards almost to the
postmedial line; claviforni absent; orbicular and reniform
with fuscous centres and undefined whitish annuli, the former
small, narrow, oblique elliptical ; a medial daik shade,
oblique below the cell ; postmedial line represented by a
246
Sir G. F. Hampson on new
double series of black points on the veins ; subterminal line
absent, the terminal area slightly darker ; a terminal series
of black points. Hind wing whitish tinged with brown;
a dark discoidal spot, rather diffused sinuous postmedial line
and terminal series of striae; cilia ochreous white with a
faint dark line through them ; the underside with the
markings more prominent, the costal area tinged with rufous
and iiTorated with fuscous.
Ilab, Uganda, Kuwenzori, 12,600^ [Ruwenzori Exp.'),
1 S type. Exp. 34 mm.
890 c. Episilia rujisigna^ sp. n.
Head and thorax ochreous white tinged with brown; sides
of palpi and frons blackish ; pectus tinged with rufous ;
abdomen fuscous brown above, whitish below. Fore wing
white tinged with ochreous and sparsely irrorated with black ;
the lines absent ; orbicular represented by a black point ;
reniform rufous defined by brown, quadrate, angled inwards
on median nervure ; a diffused oblique red-brown shade from
apex ; a terminal series of black points. Hind wing whitish
suffused with reddish brown ; a dark discoidal spot ; cilia
whitish ; the underside whitish irrorated with brown, a dark
discoidal spot, small subapical spot on vein 7, and terminal
series of small spots.
Hah. S.E. Peru, St. Domingo, 1 \ ?. Exp.
52 mm.
913 a. LycopJiotia ecUpticaj sp. n.
^ . Head and thorax rufous ; tegulse with triangular deep
rutous patch at base; abdomen pale rufous mixed with
greyish. Fore wing rufous with an ochreous tinge; a slight
curved rufous subbasal line ; antemedial line rufous, angled
outwards below costa, incurved in submedian interspace and
bent outwards to inner margin ; claviform represented by a
faint point at its extremity ; orbicular a small almost obsolete
annulus ; reniform narrow, fuscous defined by whitish ;
traces of a medial shade ; postmedial line rufous, slightly
bent outwards below costa, then minutely waved, excurved
to vein 4, then incurved, slightly angled inwards in discal
and submedian folds ; subterminal line almost obsolete, with
slightly darker shade before it at costa ; cilia grey-brown
with a dark browm line through them. Hind wing white,
the veins, costal area, and termen tinged with brown ; a
terminal series of dark striae ; the underside with the costal
247
Genera and Species 0/ N’octuiilaj.
area su ffused with fuscous, a small discoidal lunule and
indistinct sinuous postmedial line, the terminal strise black.
Hah. Tierra del Fuego, Rio McClelland {Crawshaij),
1 type. Exp. 36 mm.
916 a. Lycopliotia air {media ^ sp. n.
S . Head and thorax bright rufous slightly irrorated with
white ; tarsi fuscous with pale rings ; abdomen pale brown,
the anal tuft and ventral surface rufous. Fore wing bright
rufous, the costal area slightly irrorated with white ; an in-
distinct curved subbasal line from costa to submedian fold ;
antemedial line bent inwards to costa, then slightly sinuous
and rather oblique, indistinct towards inner margin ; clavi-
form narrow defined by black ; orbicular and reniform
irrorated with white and defined by black, the cell before and
between them and the area between reniform and postmedial
line suffused with black, the orbicular rounded, open above ;
the postmedial line slightly bent outwards below costa, then
very minutely waved, excurved to vein 4, then incurved,
some pale points beyond it on costa; subterminal line very
indistinct, with somewhat darker shade before it at costa,
slightly angled outwards at vein 7 and excurved at middle.
Hind wing white, the costal area slightly irrorated with
brown ; the underside with the costal area suffused with
rufous.
Hah. Tierra del Fuego, Rio McClelland ( Grawshay) ,
1 S fjpe. Exp. 42 mm.
923 a. Lycopliotia atrifascia^ sp. n.
Head and thorax grey mixed with brown ; pectus with
some ochreous hair ; abdomen ochreous white. Fore wing
grey tinged with brown and irrorated with fuscous especially
on the veins ; subbasal line represented by traces of double
black striae from costa and cell ; antemedial line double,
oblique, waved ; orbicular and reniform small, incompletely
defined by black, the former round, a black fascia in cell
between them and slight streak beyond the reniform ; post-
medial line indistinct, double, bent outwards below costa,
then minutely dentate and produced to black and white points
on the veins, excurved to vein 4, then oblique ; subterminal
line only defined by the area beyond it being tinged with
fuscous, excurved below vein 7 and at middle ; a fine whitish
line at base of cilia. Hind wing pure white ; the underside
with the costal area slightly irrorated wfith brown.
Hah. Argentina, Mendoza [Bain)^ 1 , 3 ? type. Exp.
32-38 mm.
248
Sir G. F. Hampson on new
924 a. Lycopliotia melanoleuca ^ sp. n.
^ . Head and thorax black -brown, tlie scales of head and
thorax slightly tipped with grey ; abdomen fascous, the base
paler, the anal tuft ochreous. Fore wing black-brown with
a slight reddish tinge and leaden gloss ; subbasal line in-
distinct, waved, from costa to submedian fold ; antemedial
line double, the inner line very indistinct, waved, somewhat
obli,que ; claviform minute, very indistinctly defined by black
scales ; orbicular and reniform defined by black, the former
round ; traces of a dark medial shade ; postmedial line defined
by paler colour on outer side, bent outwards below costa, then
very minutely waved, excurved to vein 4, then incurved,
some pale points beyond it on costa; traces of a dark sub-
terminal line, angled outwards at vein 7 and excurved at
middle. Hind wing white, the costal area and veins slightly
tinged with brown ; a fine brown terminal line ; the under-
side with the costal area irrorated with fuscous.
Hab. Tierka del Fuego, Rio McClelland {Crawshay),
1 ^ type. Exp. 42 mm.
951 a. Lycopliotia poliadeSj sp. n.
^ . Head and thorax white with a few fuscous hairs ; tarsi
banded with black ; abdomen white with tufts of long ochreous
hair from the lateral stigmata. Fore wing grey-white
slightly tinged with pale rufous in parts and irrorated with
fuscous, the veins with dark streaks ; a slight black streak
below base of cell ; subbasal line represented by black sfrise
from costa and cell ; antemedial line represented by a black
point on costa ; claviform defined by a few black scales ;
orbicular represented by a short black streak defined by
white, the reniform by an undefined white lunule with
fuscous spot at lower angle of cell ; medial and postmedial
black points on costa, with some slight white points beyond
them ; subterminal line indistinct whitish defined on inner
side by slight dentate rufous marks, angled outwards at
vein 7 and inwards at discal fold ; a terminal series of slight
black lunules. Hind wing pure white.
Hab. BeloOCHISTAN, Quetta [Nurse), 1 ^ fjpe. Exp.
36 mm.
1004 a. Lycopliotia hucoplaga, sp. n.
$ . Head and thorax clothed with rufous and whitish
scales ; tegulae whitish with rufous tips ; metathorax with
rufous- tinged whitish patch ; abdomen fuscous brown. Fore
2i9
Genera and S/jecies of Noctulda?.
wing rufous with slight dark irroration, the costal area whitish
at base and with some grey beyond middle, the terminal area
grey-wliite ; subbasal line absent ; antemedial line repre-
sented by faint dark points on the veins ; claviform and
orbicular absent ; reniform represented by a diffused dark
patch, a quadrate wliite patch in cell before it connected with
costa by an oblique wliite bar ; postmedial line represented
by a white striga from costa, bent outwards below costa, then
by a faint series of dark points on the veins ; subterminal
line indistinct, grey with dark patch before it on costa, the
apex white; a terminal series of slight black stria3 ; cilia
rufous with fine pale line at base. Hind wing pale fuscous,
the basal area whitish ; a slight dark discoidal point and fine
terminal line; cilia whitish with a dark line through them ;
the underside whitish irrorated with brown, the costal and
terminal areas suffused with brown, a slight discoidal lunule
and postmedial series of minute dark streaks on the veins.
Hab. Uganda, Ruwenzori, 6000' {Ruioenzori Exp.)^ 1 ?
type. Exp. 28 mm.
1020 a. Ufeus carnea^ sp. n.
$ . Head and thorax pale flesh-colour mixed with brown ;
abdomen pale grey-brown. Fore wing pale flesh-pink
slightly irrorated with fuscous ; the medial area (except
towards costa) and a patch on costa beyond postmedial line
suffused with fuscous ; subbasal line represented by double
striae from costa and cell ; antemedial line rather indistinct,
waved, incurved to costa and angled inwards on vein 1 ;
claviform minute, defined by blackish ; orbicular and reni-
form pale pinkish defined by fuscous, the former rather
oblique elliptical, the latter with some brownish in centre and
angled inwards on median nervure ; traces of a vv^aved medial
line; postmedial line indistinct, bent outwards below costa,
then minutely waved, incurved below vein 4, some pale
points on costa beyond it ; a subterminal series of small
dentate black marks angled outwards at vein 7, then oblique;
a terminal series of minute dark points ; a fine pale line at
base of cilia. Hind wing ochreous suffused with brown ; the
underside whitish tinged with flesh-colour and irrorated with
brown, a small discoidal spot and indistinct sinuous post-
medial line.
Hah. Kashmir, Naikundah {McArthur)^ 1 ? type. Exp.
44 mm.
Ann. As Mag. N. Hist. Ser. 7. Vol. xix.
17
250
Sir G. F. Hampson on new
Genus Anhausta, nov.
Type, A. expj’imata, Stand.
Proboscis absent; palpi porrect to well beyond frons,
fringed with long hair ; irons smooth ; eyes large^ rounded,
overhung by long cilia; antennae of female ciliated; head
and thorax clothed with hair only and without crests ; fore
tibiae without spines, mid and hind tibiae with a few spines;
abdomen with dorsal crest at base only. Fore wing with
veins 3 and 5 from near angle of cell ; 6 from upper angle ;
9 from 10 anastomosing with 8 to form the areole ; 11 from
cell. Hind wing with veins 3, 4 from angle of cell ; 5 obso-
lescent from middle of discocellulars ; 6, 7 stalked ; 8 anas-
tomosing with the cell near base only.
1021 e. Anytus leucocyma, sp. n.
? . Head and thorax red-brown mixed with fuscous ; frons
with lateral black bars ; tegulae with slight medial black line ;
abdomen grey-brown. Fore wing red- brown irrorated with
grey and black on basal half, the veins streaked with black ;
a sinuous black streak below base of cell ; subbasal line
absent; antemedial line represented by two l3lack striae from
costa, then very indistinct, strongly dentate, oblique ; clavi-
form narrow, defined by black and with blackish streak from
it to postmedial line ; orbicular defined by black, oblique,
wedge-shaped ; reniform indistinctly defined by black and
with slight whitish lunule on outer edge, its lower extremity
produced ; postmedial line double at costa, bent outwards
bilow costa, then dentate and produced to streaks on the
veins, oblique to vein 5, then inwardly oblique and angled
inwards in submedian fold, some white points beyond it on
costa; subterrninal line white, waved, angled outwards at
vein 7 and to termen at veins 4, 3, with black streaks
beyond it in the interspaces ; a fine waved black terminal
line; cilia greyish and fuscous intersected with white. Hind
wing whitish tinged c\ith brown, the veins and terminal area
suffused with brown ; traces of a waved white subterminal
line ; cilia white with a slight dark line through them ;
the underside white irrorated with fuscous ; a discoidal
spot, slight waved postmedial line, and traces of subterminal
line.
H(xb. Kashmir, Nubra [McArthur), 2 ? type. Exp.
50 mm.
Genera and Species of Noctuidse.
251
1025 a. Anytiis negrita, sp. n.
\ $ . Head and thorax black-brown irrorated with grey ;
tarsi ringed with white ; abdomen grey-brown, the dorsal
crests rather darker. Fore wing black-brown slightly suffused
with grey ; an indistinct double waved subbasal line from
costa to submedian fold ; a slight black streak above inner
margin near base; antemedial line indistinct, oblique, waved;
claviform short, deep red-brown defined by black ; orbicular
and reniform with brown centres and grey annuli defined by
black, the former round, the latter constricted at middle and
extending to below cell ; postmedial line indistinct, slightly
bent outwards below costa, then waved, excurved to vein 4,
then oblique, some grey points beyond it on costa ; a sub-
terminal series of slight grey and black dentate marks ; a
terminal series of slight black strife. Hind wing white, the
veins brown, the inner area, and terminal area broadly, fuscous
brown ; cilia pale brown, whitish towards tornus, a slight dark
line near base ; the underside with the costal and terminal
areas fuscous suffused with grey, a black discoidal point.
Hah. Uruguay, Monte Video [de la Garde), 1 ? type.
Exp. 36 mm.
Genus Blepharita, nov.
Type, B. arnica, Treit.
Proboscis fully developed ; palpi oblique, the second joint
fringed with long hair in front ; frons smooth ; eyes large,
rounded, overhung by long cilia; antennse of male bipectinate
with moderate branches to near apex ; head and thorax
clothed chiefly with scales, the pro- and metathorax with
spreading crests ; tibim fringed with hair, the mid and hind
tibise spined; abdomen with dorsal crests on basal segments.
Fore wing with the termeii crenulate ; veins 3 and 5 from
near angle of cell ; 6 from upper angle ; 9 from 10 anas-
tomosing with 8 to form the areole ; 11 from cell. Hind
wing with veins 3, 4 from angle of cell ; 5 obsolescent fro.m
middle of discocellulars ; 6, 7 from upper angle ; 8 anas-
tomosing with the cell near base only.
Genus Blepharoa, nov.
Type, B, {Agrotis) mamestrina, Butl.
Proboscis fully developed ; palpi obliquely upturned, the
second joint fringed with long hair in front; frons smooth; eyes
17*
252
Sir G. F. llampson on new
larp^e, rounded, overhung by long cilia ; antennae of male bi-
])ectinate with moderate branches, the apical third ciliated ;
head and thorax clothed chiefly with scales, the pro- and
metathorax with spreading crests; hind tibiae with one spine
between mid and terminal spurs; abdomen with dorsal series
of crests. Fore wing with veins 3 and 5 from near angle of
cell ; 6 from upper angle; 9 from 10 anastomosing with 8 to
form the areole ; 11 from cell. Hind wing with veins 3, 4
from angle of cell; 5 obsolescent from middle of disco-
cellulars ; 6, 7 from upper angle ; 8 anastomosing with cell
near base only.
Genus Pal^agrotis, nov.
Type, P. inops, Led.
Proboscis fully developed ; palpi obliquely porrect, fringed
with hair in front; frons with slight truncate prominence;
eyes large, rounded ; antennae of male ciliated ; head and
thorax clothed with hair overlying scales, the pro- and
metathorax with slight crests ; mid and hind tibiae with one
or two spines, fore tibiae without spines ; abdomen with
dorsal crest at base only. Fore wing with veins 3 and 5
from near angle of cell ; 6 from upper angle ; 9 from 10
anastomosing with 8 to form the areole ; 11 from cell.
Hind wing with veins 3, 4 from angle of cell ; 5 obsolescent
from middle of discocellulars ; 6, 7 from upper angle; 8
anastomosing with the cell near base only.
JIadeninje,
Genus POLIA. Insert : —
Miselia, Ochs. Schmett. Eur. iv. p. 72 (1816), non
descr. ; Treit. Eur. Schmett. v. (1) p. 386 (1825),
which has precedence
Chera, Hiibn. Verz. p. 211 (1827)
Hama, Steph. 111. Brit. Ent., Ilaiist. iii. p. 4 (1829). . . .
1269 a. Miselia pyro soma ^ sp. n.
d . Head and thorax deep red ; tegulae with blackish marks
at base and two leaden-grey medial lines ; patagia with
leaden-grey line near upper edge ; palpi and the hair on
pectus and femora fiery red; tibiae and tarsi fuscous, the
latter ringed with white; abdomen fiery red, whitish at base
and wdth the crests deep red. Fore wdng deep red suffused
Type.
conspersa.
serratilinea.
aliena.
253
Genera and Species of Xoctuidae.
with black-brown, the veins streaked with fuscous and leaden
grey ; subbasal line represented by a leaden-grey striga from
costa and an oblique pale yellow striga from cell defined by
black on inner side and with minute yellow striga before it ;
a curved yellow mark with black suffusion above it above
inner margin before the antemedial line, which is black defined
by leaden grey on inner side, angled outwards below costa
and inwards on median nervure, then oblique to vein 1, where
it is angled outwards; claviform faintly defined by black and
with wxdge-shaped yellow spot defined by black beyond it ;
orbicular and reniform leaden grey faintly defined by black
and with red lines in centres, the former oblique elliptical,
the latter angled inwards on median nervure to below the
former ; postmedial line black defined by leaden grey on
outer side, slightly bent outwards below costa, then minutely
dentate, oblique to vein 5, then inwardly oblique and angled
inw^ards on vein 1, some wdiitish points beyond it on costa ;
subterminal line represented by minute yellow streaks between
veins 7 and 2, with blackish streaks beyond them and an
oblique yellow striga below vein 2 ; a narrow leaden-grey
terminal band with waved inner edge with some white points
on it ; cilia red at base mixed w'ith grey and black at tips.
Hind wing red-browm tinged with fuscous ; cilia white at
tips; the underside red tinged wdth browm and irrorated with
black, the inner area whitish, a black discoidal spot and
curved postmedial line.
Ilah. S.E. Peru, St. Domingo, 2 S ^yp^- 42-48 mm.
1293 a. Miselia pliimipeSj sp. n.
S . Head, thorax, and abdomen bright rufous with a few
dark scales ; some of the hair on fore tibise fuscous ; abdomen
with sublateral black striae. Fore wing bright rufous slightly
irrorated with black ; subbasal line represented by double
black striai from costa ; antemedial line double, slightly
waved ; oibicular and reniform small, the former oblique
elliptical with whitish annulus defined by black, tlie latter
open above and below, concave towards base, defined at sides
by white between black lines, the white on outer side forming
a small triangular mark ; postmedial line double, bent out-
wards below costa, excurved to vein 4, then oblique, sinuous,
some white points on costa beyond it ; subterminal line pale,
angled outwards at vein 7 and excurved at middle, the area
beyond it darker ; a terminal series of small black luiiules.
Hind wing rufous, the cilia whitish at tips ; the underside
greyish irrorated with rufous, especially on costal area ; a
254
Sir G. F. Hampson on new
small discoidal spot, minutely waved postmedial line, and
some small black lunules on termen from apex to vein 2.
Hah, Costa Kica, 1 type. Exp, 28 mm.
1361a. Miselia gercea^ sp. n.
$ . Head and thorax grey-white mixed with brown ; frons
with lateral black bars ; tegulse with medial black line ;
patagia edged with black ; tarsi black ringed with white ;
abdomen grey-white, the dorsal crest at base with black line
near tip, the extremity tinged with rufous. Fore wing grey-
white irrorated with brown and fuscous, the veins with slight
dark streaks ; a sinuous black streak below base of cell ; sub-
basal line represented by slight black striae from costa and
cell ; traces of a double antemedial line excurved from costa
to submedian fold and angled inwards on vein 1; claviform
narrow, defined by black ; orbicular and reniform with slight
whitish annuli defined by black, the former elongate elliptical ;
a diffused curved medial shade ; postmedial line with double
black points at costa, then represented by slight black streaks
on the veins, bent outwards below costa, excurved to vein 4,
then oblique ; subterminal line represented by a series of
faint dentate brown marks, the area beyond it somewhat
browner except at base and tornus ; a terminal series of black
points. Hind wing white, the veins with dark streaks towards
termen, which is tinged with brown ; the underside wdth the
costal area slightly irrorated with fuscous, a terminal series
of slight black striae.
Hab, Br. E. Africa, Taveta [K. St. A. Bogers), 1 ? type.
Exp. 34 mm.
1930 a. Cirphis clavifera^ sp. n.
$ . Head and thorax white slightly tinged vvith brown and
irrorated with black ; frons with lateral black bars ; patagia
with some black scales near upper edge ; abdomen white,
dorsally tinged with ochreous. Fore wing white, slightly
irrorated wdth black and faintly tinged with brown except
costal area to beyond middle and inner area to middle, the
submedian and discal folds tinged with yellowish ; the median
iiervure with fine white streak ; a sliglit black streak below
base of cell ; antemedial line represented by a black point on
vein 1 ; claviform indicated by slight black marks above and
below ; a rather wedge-shaped white mark in end of cell ;
postmedial line represented by a series of black points on the
veins, bent outw'ards below costa and oblique below vein 4 ;
255
Genera and Species of Noctuidae.
a slight whitish streak below terminal part of vein 7 and
slight subterminal striga above tornus ; a terminal series of
black points. Hind wing pure white.
Hah. Br. E. Africa, Taveta (K. St. A. Rogers), 1 ? type.
Exp. 32 mm.
1972 a. Borolia ustata, sp. n.
S . Head, tegulae, and prothoracic crest ochreous mixed
with dark brown ; the tegulae with three black lines ; thorax
yellow mixed with fiery red ; pectus and legs ochreous mixed
with dark brown ; abdomen ochreous suffused with brown.
Fore wing yellow suffused with fiery red, the costal and
inner areas with slight dark irroration, the veins whitish
defined by slight brown streaks, the interspaces with slight
brown streaks ; the costal edge wdiitish, the costal area
brownish on terminal half and an oblique brownish fascia
from termen below apex ; subbasal line represented by a
series of black points strongly excurv’-ed below the cell ; a
white streak on extremity of median nervure ; orbicular and
reniform yellow with some rufous in centres, ill-defined ;
a curved postmedial series of black points with traces of a
crenulate line between them ; a terminal series of black
points; cilia brown. Hind wing ochreous white, the terminal
half suffused with brown ; a terminal series of black points
from apex to vein 2 ; cilia ochreous white witli a brown line
through them ; the underside with the costal and terminal
areas suffused with rufous and irrorated with black ; a small
discoidal spot, rather diffused postmedial line from costa to
vein 2, and terminal series of black points.
Hah. Transvaal, White R. {Cooke), 1 type. Exp.
28 mm.
1973 a. Borolia pyrostrota, sp. n.
? . Head and thorax ochreous mixed with red ; sides of
palpi and frons brownish ; patagia streaked with brown at
sides ; abdomen reddish ochreous irrorated with fuscous.
Fore wing yellowish tinged with fiery red, the veins white
defined by fine fuscous streaks, the interspaces of terminal
area with fine fuscous streaks, the basal half of costal area
pale yellow ; antemedial line represented by two obliquely
placed black points in submedian interspace ; a black point
at lower angle of cell ; a postmedial series of black points,
slightly bent outwards below costa, angled inwards in discal
fold and oblique below vein 4 ; diffused dark shades along
subcostal and median nervures and thence to apex, from
which an oblique pale fascia extends to vein 4 ; a terminal
250
Sir G. F. Ilampsoii on new
series of black points. Hind wing oclireous, nearly uniformly
suffused with fuscous ; cilia pale yellowish ; the underside
pale yellowish irrorated with fuscous, a black discoidal spot,
postmedial series of short streaks on the veins, and terminal
series of points.
Hah. Uganda, Ruwenzori, 6000' [Ruwenzori Exp.), 1 $
type. Exp. 32 mm.
1977 a. B or olia fascia, sp. n.
$ . Head and thorax pale oclireous tinged with reddish
brown ; antennae blackish ; tarsi fuscous ; abdomen pale
oclireous tinged with fuscous brown. Fore wing pale
brownish oclireous, the cell and area beyond it running
obliquely to termen below apex pale rufous, the medial part
of submedian interspace tinged with rufous, the inner area
slightly irrorated with black except towards base ; a sinuous
black streak below basal half of cell ; the terminal half of
median nervure with prominent white streak; veins 3,4 with
slight white streaks with fine streaks above them ; an oblique
whitish fascia from apex ; a subtermiiial black point on vein 4
and a terminal series of black points ; cilia grey-brown.
Hind wing oclireous whitish, the terminal half suffused with
fuscous brown; cilia white; the underside brownish white
with terminal series of black points.
Hob. Lagos {Boag), 1 $ type. Exp. 40 mm.
1998 a. Borolia metasarca, n. sp.
f . Head and thorax pale pinkish brown ; patagia with a
few black scales near upper edge ; legs slightly irrorated with
fuscous; abdomen oclireous brown. Fore wing pinkish
fleBh-colour, the cell and area beyond it to postmedial line,
the area below it to submedian fold, and a fascia below medial
part of vein 1 olive-brown ; a whitish streak on median
nervure and above vein 3 to termen, some sparse black
iiToration ; a prominent black spot at lower angle of cell ;
postmedial line represented by a series of black points, bent
outwards below costa and oblique below vein 4 ; the veins of
terminal area streaked with olive-brown ; a pale oblique
fascia from apex defined above and below by fuscous shades ;
a terminal series of black points ; cilia pinkish intersected
with brown. Hind wing with the cell and area beyond it to
termen flesh-pink slightly suffused with fuscous, the costal
and inner areas })ale brownish oclireous ; cilia pale yellow ;
the underside with the costal area slightly iiroratcd witli
257
Genera and Species o/Noctuicl?o.
black, a discoidal limiile, curved postmedial series of points
and terminal series. »
Hal). Ashanti, Kumassi ( Whiteside) ^ 2 $ type. Exp.
32 mm.
2003 a. Borolia phceopastaj sp. n.
$ . Head and thorax pale ochreous irrorated with fuscous ;
abdomen ochreous irrorated and suffused with fuscous. Fore
wing ochreous tinged with red and thickly irrorated with
fuscous ; traces of a waved antemedial line ; orbicular repre-
sented by a dark point, the reniform by a dark lunule on
yellowish patches ; traces of a postmedial line with dark
streaks beyond it on the veins, slightly bent outwards below
costa and oblique below vein 4 ; subterminal line represented
by the area beyond it being somewhat darker, angled out-
wards at vein 7 and excurved at middle ; a terminal series
of black points. Hind wing ochreous uniformly suffused with
fuscous ; cilia pale ochreous ; the underside ochreous irrorated
with fuscous, a slight dark discoidal spot and curved post-
medial line.
Hah. Uganda, Huwenzori, 6000' (Ruwenzori Exp.), 1 $
type. Exp. 32 mm.
Ctjcvllianm.
2589 a. Trichoridia ethiopica, sp. n.
. Head and thorax deep red-brown, the head and tegulae
redder ; palpi dark brown fringed with rufous; pectus and
legs pale brown mixed wdth fuscous ; abdomen pale browni
suffused with fuscous browm, the anal tuft rufous. Fore wing
pale red-brown, the basal area and costal area to near apex
suffused with dark brown leaving the costal edge rufous, the
medial area deep red-brown except on costal and inner areas
and defined from subcostal nervure to vein 1 by the slight
pale ante- and postmedial lines, the former outw^ardly oblique
below submedian fold, the latter inwardly oblique below^
vein 5 ; orbicular and reniform whitish tinged with browm,
the former very oblique oblong and botli confluent with a
large elliptical patch on extremity of median nervure extend-
ing to well beyond low^er angle of cell ; a slight brown
terminal line. Hind wing pale brown suffused with fuscous
brown ; cilia pale rufous ; the underside whitish tinged with
rufous and irrorated with fuscous, a fuscous discoidal lunule
and diffused slightly curved postmedial line.
Hah. Bk. E. Africa, Aberdare Range {R. Ford), 1 S type.
Exp. 34 mm.
258
Mr. C. T. Regan on
XXX. — Descriptlom of Six new Freshwater Fishes from
Mexico arid Central America. Bj C. Tate Regan, B.A.
Pimelodus Boucardi,
Depth of body in the length, length of head 4. Breadth
of head 1 J in its length, length of snout 2|, diameter of eye 6,
interorbital width 3. Lower jaw nearly as long as the
upper ; maxillary barbel extending to origin of adipose fin.
Head covered with smooth skin ; occipital process rather
strong, extending the distance from its base to the origin of
dorsal. Dorsal 1 6, the spine slender, the fin rounded, as
high as long. Adipose fin J the length of the fish. Anal 14.
Pectoral spine with serrated inner edge, about | the length of
tlie fin and J the length of head; humeral process long,
nearly reaching the middle of the fin ; ventrals originating
nearly below the last dorsal ray, extending more than f of
the distance from their base to the origin of anal. Caudal
deeply notched, the lobes rounded, the lower the larger.
Least depth of caudal peduncle the length of head.
Blackish.
Hah. Yucatan [Boucard).
A single specimen, 190 mm. in total length.
Fimelodus hrachycephalus.
Depth of body to in the length, length of head 5^
to 5|. Breadth of head Ih in its length, length of snout 2|
to 3, diameter of eye 5 to 6, interorbital width 3 to 3^.
Lower jaw a little shorter than the upper ; maxillary barbel
extending to the basal part of pectoral. Head covered with
smooth skin ; occipital process short. Dorsal I 6, the spine
slender, the fin rounded, as high as or higher than long.
Length of adipose fin 3^ to 3|; in the length of the fish.
Anal 12-13. Pectoral spine with serrated inner edge, about
I the length of the fin and | the length of head ; humeral
proeess short ; ventrals originating behind the dorsal, ex-
tending I to I the distance from their base to the origin of
anal. Caudal moderately notched, the lower lobe rounded
and rather shorter than the upper. Least depth of caudal
peduncle § or more than the length of head. Brownish.
JIab. Guatemala, Rio Nacasil [Salvin).
Six specimens, measuring up to 190 mm. in total length.
new Freshwater Fishes.
259
Plmelodus Roger si.
Depth of body 6 in the length, length of head 5. Head 1^-
as long as broad. Diameter of eye 6 in the length of head,
interorbital width 3, length of snout 3. Head covered with
smooth skin ; fontanel not extending beyond the level of
posterior margin of eye ; occipital process short. Maxillary
barbel extending to middle of pec'^oral fin. Dorsal I 6, the
spine slender, the fin rounded, higher than long. Adipose
fin as long as or a little longer than its distance from the
dorsal, about f the length of the fish. Anal 12-14. Pectoral
spine nearly | the length of the fin and nearly | the length of
head, its inner edge with a series of rather small denticula-
tions. Ventrals originating behind the last dorsal ray, ex-
tending J the distance from their base to the origin of anal.
Caudal with a shallow notch. Least depth of caudal peduncle
I the length of head. Brownish.
Hah. Costa Pica, Irazu [H. Rogers).
Pour specimens, measuring up to 130 mm. in total length.
Allied to P. tSalvinij Gthr., which has a longer adipose
fin (f the length of the fish), a shorter pectoral spine (less
than the length of iiead), a shorter anal fin with 11 rays,
and the caudal more deeply notched.
Gambusla annectens.
Depth of body 3 to 3| in the length, length of head 3J to
3|. Snout as long as or shorter than eye, the diameter of
which is 3 to 3| in the length of head ; interorbital width
equal to the distance from middle or posterior part of eye to
free edge of operculum. 28 to 31 scales in a longitudinal
series. Dorsal 10-12; origin equidistant from tip of snout
and middle or posterior part of caudal ; free edge of the fin
convex. Anal 9-10, pointed, originating a little in advance
of the dorsal. Pectoral about f the length of head. Caudal
rounded or subtruncate. Least depth of caudal peduncle
about f the length of head. Scales with dark edges, forming
series of spots on the lower part of the side ; often a series of
short dark vertical bars along the middle of the side; a series
of dark spots on the lower part of the dorsal fin.
Hah. Costa Pica, Carrullo and Juan Venas [Underwood) ;
Iiazu [Rogers).
Numerous specimens, measuring up to 70 mm. in total
length.
This species is very close to O. episcopi, Steind., from
Panama, which has only 8 or 9 dorsal rays and also ditfers in
having a blackish spot on the anal fin. It is also close to
2G0
On new Freshwater Fishes.
G. terrahensis^ Jiegan, and G. Jonesii^ Giinther and with
tliern forms a series froiri G. episcopi to G. {Pseudoxipho-
jdwrus) bimaculata, Heck., so that the genus Pseudoxipho-
phurus can no longer be maintained.
Gamhusia terrahensis.
Depth of body to 3^ in the length, length of head 3-^ to
3i. Snout shorter than eye, the diameter of which is 3 to 3^
in the length of head ; interorbital width equal to the distance
from posterior margin of pupil to free edge of operculum. 29
to 31 scales in a longitudinal series. Dorsal 12-14; origin
equidistant from tip of snout and base of caudal or a little
nearer the latter. Anal 9-10 ; oi’igin below or a little in
advance of the middle of dorsal. Pectoral nearly J the
length of head. Caudal subtruncate. Least depth of caudal
peduncle nearly § the length of head. Scales of the upper-
part of the body with dark edges ; an interrupted dark lateral
stripe ; dorsal fin with a basal series of vertically expanded
blackish spots and with a second series of smaller spots ; anal
dark at the base and also distally ; posterior part of caudal
dusky.
Hah. Costa Rica, Rio Grande de Terraba, Pacific Slope
(L/. Pittier').
Six specimens, measuring up to 48 mm. in total length.
Sicydium Piitieri.
Depth of body 6 in the length, length of head to 5|.
Diameter of eye to 6 in the length of head, intei-orbital
width 4 to 4J. Snout obtuse ; mouth subterminal, with
horizontal cleft, extending to below the middle of eye; u])per
teeth bicuspid ; teeth of the outer series in the lower jaw
sometimes concealed. Occipital region covered with small
scales; 75 to 80 scales in a longitudinal series; abdomen
with a median naked area or strip. Dorsal VI, 1 10; rays
of first dorsal (in the males) produced into filaments, the
longest, when laid back, nearly reaching the end of the base
of second dorsal ; longest rays of second dorsal a little longer
than the head. Anal I 10 ; origin equidistant from eye and
base of caudal. Pectoral longer than the head, twice as long
as the ventrals. A dark spot on each scale ; dorsal fins witli
dark vermiculations ; anal with a daik edge.
llah. Costa Rica, Rio Grande de Terraba (/i. Pittier).
Two specimens (males), 120 and 135 mm. in total lengih.
This species is nearest to H. Salrini, Grant, from Panama
and Western Ecuador.
* MolUenisia Jonesii, — Psetaloxiphophorus pauciradiaius, 1-Jogan.
On new CJiaracinid Fishes.
2(31
XXXI. — Descriptions of Two new Gharacinid Fishes from
Argentina. By C. Tate Reg AN, B.A.
Dr. L. Reh, of the Hamburg Natural History Museum,
recently sent a few specimens to Mr. Boulenger for deter-
mination, describing them as aquarium fishes from Argentina.
These have been handed over to me and referred to tliree
species, viz. Tetragonopterus cordovensis, Gthr., and two other
Characinids, each the type of a new genus and species, which
are described below.
PoGONOCHARAX, gen. nov.
Closely allied to Pgrrhulina, C. & V., differing in the
toothless mouth, the presence of two barbels on each side,
respectively attached to the distal ends of the praemaxillary
and maxillary, and in the more posterior position of the dorsal
fin, which is opposite the anal.
Pogonocharax Rehi^ sp. n.
Depth of body equal to length of head, in the length of
the fish. Snout shorter than eye, the diameter of which is
3^' in the length of head and less than the interorbital width.
Mouth small, toothless ; lower jaw flat, shovel-shaped (as in
Pyrrhxdina Jilamentosa) ; pra3maxillary barbel about as long
as the eye ; maxillary entirely in front of the eye, bearing a
barbel which is | as long as the fish. Scales rather large.
cycloid, apparently about 25 in a longitudinal series ; no
lateral line. Dorsal 8, rather elevated anteriorly. Anal 8;
origin below that of the dorsal ; fin larger than the dorsal
and with the posterior rays much branched. Pectoral long,
262
Dr. R. Broom on some
pointed, 1^ as long as the head ; outer ray of ventral some-
what produced, reaching the anal. Caudal peduncle twice
as long as deep. Perhaps a dusky lateral band.
A single specimen, 45 mm. in total length.
The barbels at once distinguish this remarkable fish from
any other member of the family.
Phoxinopsis, gen. nov.
Allied to Lehiasina^ C. & V., but with the teeth conical
instead of tricuspid and tlie anal fin longer.
Phoxinopsis ty pirns ^ sp. n.
Depth of body nearly e'qual to the length of head, 4 in the
length of the fish. Snout much shorter than eye, the
diameter of which is 3 in the length of head and a little less
than the interorbital width. Teeth conical, in a single series ;
maxillary toothless, extending to below the anterior edge of
eye. Scales cycloid, 33 in a longitudinal series ; lateral line
developed anteriorly, on 6 or 7 scales only. Dorsal 10;
origin a little nearer to base of caudal than to tip of snout ;
longest ray a little shorter than the head. Anal 16 ; origin
below end of dorsal ; free edge emarginate. Pectoral shorter
than the head, extending a little beyond the base of ventrals,
which do not quite reach the anal. Caudal peduncle a little
longer than deep. A dark linear lateral streak.
A single specimen, 34 mm. in total length.
XXXII; — On some new Species of Chrysochloris.
By R. Broom, M.D., D.Sc., C.M.Z.S.
Dobson, in his monograph on the Insectivora, published in
1883, recognizes only five species of Chrysochloris as in-
habiting South Africa, viz. Chrysochloris aurea [ = asiatica)^
new Species 0/ Clirysocliloris.
C. vilhsaj C. Trevdyani^ C. rutilans ( = hottentota) , and
C. ohtusirostris ; and W. L. Sclater, in his ‘ Mammals of
South Africa,’ recog'nizes only the same five. Recently, as
the result mainly of the researches of Mr. C. II. B. Grrant,
Oldfield Thomas has added a few new species belonging to the
subgenus Amblysomus^ viz. A. clirysillusj A. irisj A. Corricej
and a subspecies A, hottentottus pondolice.
Tliough I have devoted comparatively little attention to
systematic zoology, I have from time to time made pretty
extensive collections of the bones of small mammals, and the
best hunting-grounds I have invariably found to be the
haunts of owls. In the disgorged pellets often found in great
abundance in rock-clefts the small mammal skulls are usually
preserved uninjured, and the owls frequently obtain specimens
which the collector of skins will not readily come across.
Having been recently studying the development and varia-
tions of the teeth of Chrysochloris^ I have gone over witli
care the collection of small mole skulls that I have made from
time to time, and among them I have discovered four new
species. Through the kindness of Mr. Peringuey, Director
of the South African Museum, I have examined all the
specimens in the museum, and have found in the collection
the skins of two forms of which I had already the skulls, and
also another new species.
The examination of these new forms has led me to doubt
the wisdom of making Amblysomus a distinct genus, and I
shall therefore retain, provisionally at least, the generic name
Chrysochloris for all the species.
Chrysochloris Sclaieri, sp. n.
Of this new species two specimens have been in the South
African Museum for some years. Both are preserved in
spirit and come from Beaufort West.
The nose-pad at once distinguishes the species from
C. asiatica, as its outer end is not produced into a sharp
point as in the common species, but rounded. The general
colour of the back is reddish brown, not unlike many speci-
mens of C. hottentota. The lips and cheeks are dull cream-
yellow, and the light patches extend to a little behind the
ocular region, over the temporal area, and meet each other
above the nasal pad. On the upper surface of the head a few
brown hairs are mixed with the yellow, and above the nasal
pad so many as to make the area pale brown. The general
colour of the abdomen is a pale dirty brown.
The skull differs from that of C. asiatica in being much
2G4
Dr. R. Broom on some
narrower and in having no trace of the protuberance into the
temporal well. In general proportions the skull is much
more like that of (7. hottentota^ but much smaller.
The teeth are forty in number and resemble pretty closely
in general structure those of C. hottentota, though smaller.
The second last molar is not unlike the last molar of G. hotten-
tota j owing to the posterior style being rudimentary. The
lower teeth are almost identical in structure with those of
C. hottentota^ the premolars and molars having the well-
developed posterior basal ledge. The second last molar has
the ledge which is absent in G. hottentota^ and the last molar
is small. The whole structure of the skull and teeth shows
this species to be much more nearly related to G, hottentota
than to G. asiatica ; and if this species, because it has forty
teeth, is retained in the genus Ghrysochloris, it seems scarcely
advisable to make a distinct genus for G. hottentota merely
because the minute last molar is missing.
Dimensions of the type (probably slightly shrunk by
spirit) : —
Head and body 96 mm. ; hind foot (s. u.) 11.
Skull: greatest length 23‘2 ; basal length 18*8; greatest
breadth 15‘6 ; greatest height 12 ; interorbital breadth 7*2 ;
front of to back of rn^ 10 ; palate across posterior pre-
molars 7’ 7.
Eab. Beaufort West, Cape Colony. Also probably much
further east.
Type. Specimen no. 3148 in South African Museum.
Young female.
I have named the specimen after Mr. W. L. Sclater, late
Director of the South African Museum.
GhrysochJoris Wtntom] sp. n.
About ten years ago I obtained a couple of specimens of
moles at Port Nolloth. Unfortunately at that time I had
not seen G. asiatica, and assumed that the Port Nolloth
animals belonged to the ordinary species. The skin of one
was sent to the British Museum, but the skull was destroyed
for the sake of the brain. Prom the other specimen a
skeleton was prepared. As I had kept the lower jaw of the
first specimen with a preparation of the tongue, I have for
some time been aware that it belonged to an undescribed
species. Luckily there is in the collection of the South
African Museum a specimen from Port Nolloth which,
though rather badly preserved in spirit, may be taken as the
type.
new Species of Clirysoclilorls. 265
The nose-pad at the sides is intermediate in shape between
that of C. asiatica and C. Sclateri, and forms a blunt angle.
The digging portion of the snout is narrower and longer than
in the other species. The back is pale slaty grey in colour,
with a greenish iridescence, while the whole of the upperside
of the head and the lips are yellowish grey, which becomes
darker above the nose-pad. The abdomen is the same colour
as the back. The fore and hind feet are relatively broader
than in most species. The fore foot has a large pad on the
inner side of the first digit, doubtless to facilitate digging in
the sand. The fourth toe of the front foot is fairly well
developed.
The skull is chiefly remarkable for the great size of the
posterior olfactory region. In this it differs from all other
known species. Whereas in all other species the interorbital
region is flat or concave, here it is convex. There is less of
a crest between the parietal and occipital regions than in
(7. asiatica^ and the projection formed by the head of the
malleus is smaller, though quite distinct.
The teeth are forty in number and resemble those of
C. asiatica^ except in being smaller and in the relatively
smaller size of the first premolar and the last two molars.
Lower molars have no basal ledge.
Dimensions of type
Head and body 90 mm. ; hind foot (s. u.) 10*3.
Skull : greatest length 21*3 ; greatest breadth 16*2 ;
greatest height 11*2; interorbital breadth 9’2; front of to
back of w? 9 ; palate across posterior premolars 7*8.
Hah, Port Nolloth, Cape Colony. Among the sand-dunes.
Type, Specimen no. 1917 in South African Museum.
The species is named after Mr. W. E. de Wintori, who
has done much towards the study of South African mammals,
and to whom I have been indebted at different times for kind
assistance.
Chrysochloris Grantiy sp. n.
Of this species I have four well-preserved skulls from
Garies, Namaqualand; but the skin was unknown till 1
discovered a specimen in the South African Museum without
locality or history, but which doubtless also comes from
Namaqualand.
The nasal pad is rather small, the hard portion very short
and the outer soft portion very narrow at the sides. The fur
is extremely long, the hairs on the back being about 20 mm.
in length. The general colour on the back is greyish
Anru Mag. N. Hist. Ser. 7. Vol. xix. 18
2G6
Dr. D. Broom on some
brown, the tips ot the hairs being light and the inner part of
the fur dark. In spirit the fur shows a violet iridescence.
The whole head is light yellowish; it is unusually broad
and the nose short. The fore feet have a much smaller pad
to the first digit than G. Wintoni, while the fourth digit is
a fairly well developed functional toe, larger than in any other
known species, and with a rather large claw.
The skull is like that of C. asiatica in being relatively
broad, but differs in being much smaller, in having a rela-
tively much shorter snout, and in having scarcely any trace
of a protuberance into the temporal fossa.
The teeth are forty in number, and the most noteworthy
feature is that the first upper premolar is of small size and
single-rooted. This character is constant in all five skulls.
In the lower jaw of the type there are only nine teeth on the
right side.
Dimensions of type : —
Greatest length 82 mm. ; hind foot 9.
Skull : greatest length 19*2 ; basal length 15 ; greatest
breadth 17; greatest height 11*5; interorbital breadth 7;
front of p- to back of 8*5; palatal width across posterior
premolars 7*6.
Hah, Namaqualand.
Type, In South African Museum.
The species is named after Mr. C. H. B. Grant, who has
done so much in the last few years to advance our knowledge
of South African Mammals.
Chrysochloris namoquensis, sp. n.
This new species is represented by four fairly well preserved
skulls, but nothing is known of the skin ; but as the skulls
are easily distinguished from those of any known species, and
as the skull and teeth are much safer guides than the skins, 1
think it well to give the form a name.
The skull is about the same size as in G. Granti^ but is
narrower and longer. It is further differentiated by having a
large protuberance in the posterior wall of the temporal fossa,
as in G. asiatica. The teeth resemble, so far as preserved,
those of G. asiatica^ the first upper premolar being large and
double-rooted. The second upper incisor is relatively larger
than in G. asiatica.
Skull : greatest length 20 mm. ; basal length 16*5 ;
greatest breadth 15*5 ; greatest height 11 ; orbital breadth G’5 ;
front of to back of 8*5 ; palate across posterior pre-
molars 7*8.
2G7
neio Species of Clirysoclilorls.
There are forty teeth.
Uah. Garies, Namaqualand.
Type. The type skull will be deposited in the South
African Museum and a co-type given to the British Museum.
CJirysochloris tenuis^ sp. n.
This species is only represented by two skulls, both slightly
imperfect, from Garies, It differs from all the preceding
species in having, like C. hottentota., only thirty-six teeth,
and thus belongs to the subgenus Amhlysomus. It differs,
however, from all the known species of Amhlysomus in
having a large protuberance encroaching on the temporal
fossa as in G. asiatica.
Skull : greatest length 20*5 mm. ; basal length 16 ; greatest
breadth 14*5 ; greatest height 10*2 ; interorbital width 6*5 ;
front of T to back oT 9*6 ; palate across posterior pre-
molars 7*3.
Hah. Garies, Namaqualand.
Type, The type skull will be deposited in the Soutli
African Museum and a co-type given to the British Museum.
Tl'e following may be taken as a synopsis of the species,
omitting one or two imperfectly known and very doublfiilly
distinct forms : —
Synopsis of the Species.
I. Teeth 40.
A. Bony projection on posterior wall of temporal
fossa ; zygomatic arch small ; ratio of breadth
of skull to length 7o to 85 ; 100.
a. Interorbital region narrow.
a'. Skull, breadth to length ratio over 80 : 100 . C. asiatica.
b' . Skull, breadth to length ratio 75:100 .... C. namaquensis.
h. Interorbital region expanded C. Wintuni.
B. Bony projection on posterior wall of temporal
fossa very small or absent ; zygomatic arch
small.
a. Skull, breadth to length ratio 90:100 C. Granti.
b. Skull, breadth to length ratio 66: 100 C. Sclateri.
C. Bony projection on posterior wall of temporal
fossa very small or absent ; zygomatic arch
greatly expanded.
a. Length of skull 33 ram C. villosa.
b. Length of skull 42 mm C. Trevelyani.
II. Teeth 36.
A. Bony projection on posterior wall of temporal
fossa ; ratio of skull breadth to length 68 : 100. . C. tenuis.
268
Ml*, rj. E. Turner on
B. No bony projection on posterior wall of temporal
fossa.
a. Skull, breadth to lenp-th ratio about GO : 100.
a'. Length of skull 28 mm C. hottentota.
C. Corrice.
b'. Length of skull 25 mm C. iris,
h. Skull, breadth to length ratio about 72 : 100 . . C. ohtusirostris.
. C. chrysillus.
Chrysoclihris Corrice and C. chrysillus^ recently described
by Thomas, appear to be closely allied to C, hottentota and
C. ohtusirostris respectively, and no marked cranial characters
are stated by which they can be readily distinguished. I am
not acquainted with either of these recently described species.
Victoria College, Stellenbosch,
Nov. 30, 1906.
XXXIII. — New Species of Sphegidte from Australia.
By B, E. Turner.
Only two genera allied to Ampulex have hitherto been
recorded as Australian, each containing a single Australian
species. Aphelotoma was described by Westwood for a
Tasmanian species, and I am now able to describe two
.additional species from the mainland. Dolichurus carbonarius^
Sm.., belonging to a small but wide-ranging genus, was
described by Smith from Champion Bay, W.A. I can now
record its occurrence at Mackay, on the Queensland coast. I
also describe three species of a new genus, which is, perhaps,
nearest to Aphelotoma,
Genus Aphelotoma, Westw.
Aphelotoma, Westw, Proc. Ent. Soc. Loudon, 1840, p. 12.
Type, Aphelotoma tasmanica^ Westw.
Aphelotoma aterrima, sp. n.
^ . Head and thorax opaque, delicately reticulate. Clypeus
broadly rounded anteriorly, without a median carina. Plead
broad, the eyes not quite touching the base of the mandibles
and convergent towards the vertex. Pronotiim narrowq
elongate, depressed anteriorly, broadest posteriorly, with a
median longitudinal sulca. Mesonotum short and broad.
2G9
neio Spliegicla; from Amir alia.
tlie surface divided into three almost equal parts by a longi-
tudinal sulca on each side. Median segment broad, sub-
quadrate, only slightly narrowed posteriorly, abruptly truncate,
the angles above the truncation produced on each side into a
minute spine, the dorsal surface of the segment coarsely
rugose, the surface of the truncation irregularly obliquely
striate. Abdomen shining; the three basal segments large,
the second slightly constricted at the base ; sparsely punc-
tured, the sides of the segments (especially of the third) very
finely and closely punctured; the apical segments small and
withdrawn. The tibiae are smooth, not serrate. The inter-
mediate tibiae have two apical spines.
Black ; a spot beneath the scape of the antennae at the
apex pale yellow ; abdomen dark fuscous. Wings hyaline,
slightly tinted with yellow, the radial and second and third
cubital cells clouded with pale fuscous. Tegulae and nervures
testaceous.
The first recurrent nervure is received close to the apex
of the first cubital cell, the second just at the base of the
third cubital cell, almost interstitial with the second trans-
verse cubital nervure. The second cubital cell is very
narrow on the radial nervure; the first transverse cubital
nervure is bent sharply outwards near the base.
Length 5 mm.
$ unknown.
Hah. Mackajq Queensland.
This species is very distinct from A. tasmanica in colour
and general facies, but there does not seem to be any marked
structural difference.
Aphelotoma auriventris, sp. n,
$ . Clypeus broad and short, truncate anteriorly, sparsely
punctured ; head opaque, the front strongly sculptured,
irregularly longitudinally punctured striate, vertex reticulate.
Pronotum subquadrate, about half the width of the meso-
notum, very coarsely rugose, with an obscure indication of a
median sulca; a small tubercle on each side before tlie
posterior angle, behind the tubercles the pronotum is obliquely
depressed to the posterior margin. Mesonotum and scutelluin
very coarsely rugose, the mesonotum reaching to the hind
margin of the tegulse, the scutellum broadly rounded at the
apex. Median segment at the base as broad as long, flat
above, truncate posteriorly, the lateral margins raised, the
dorsal surface strongly longitudinally striate, the striie about
270
Mr. H. E. Turner on
ten in number ; tlje surface of the truncation coarsely rugose.
Abdomen short, the two basal segments shining and finely
punctured, the third segment delicately reticulate and pubes-
cent, the apical segments small and withdrawn.
Black ; the antertnae, mandibles, tibiae, and tarsi fusco-
ferruginous ; abdomen shining golden bronze. Wings
hyaline, iridescent, with faint fuscous clouds near the centre
and in the second cubital cell. The first transverse cubital
nervure is sharply bent outwards near the base and throws
out a shoit branch inwards, so that the first cubital cell is
imperfectly divided.
Length 6 mm.
Hah. Victoria.
Genus Auchenophorus, nov.
$ . Head as broad or broader than the thorax ; the cheeks
well developed ; the eyes reaching to the base of the man-
dibles, convergent towards the vertex. Antennae with the
apical joints somewhat stouter than the basal, shorter than
the head and thorax, not covered at the base by a tubercle.
Mandibles acute at the apex, not bidentate ; the apical outer
angle of the maxilla produced into a prominent lobe, the
maxillary palpi five-jointed, the joints in the typical species
rather short and subequal ; labial palpi four-jointed ; labrum
transverse, straight, strongly ciliate on the anterior margin.
Pronotum narrower than the mesonoturn, depressed anteriorly;
mesonotum short, the posterior margin not reaching beyond
the posterior margin of the tegulse. Median segment rather
long, flat above, truncate posteriorly, marked dorsally with a
straight median carina from the base, continued on the
surface of the truncation to the apex, also with a carina on
each side converging from the base and forming a triangle.
Abdomen pseudosessile or subpetiolate, the apical segments
not very short, none of the segments constricted at the base.
The trochanters long, the intermediate and posterior tibiae
slightly spinose, the intermediate tibiae with only one apical
spine ; the joints of the tarsi slender and elongate. Wings
short, with only one cubital and one discoidal cell, the radial
cell short, subtruncate at the apex ; the cubital cell large,
receiving the recurrent nervure at about two thirds of tlie
distance from the base to the apex. Tlie anal cell of the
hind wing is short ; the apex of the median cell touches the
costa; the radial and cubital nervures are only faintly
indicated beyond the median cell.
new S[)liegida3 from Australia,
271
This curious genus belongs to Kohl’s Group X. of the
Sphegidae, which includes Ampulex and the allied genera,
but in the neuration it approaches much more nearly to some
of the genera in Group XL It should probably come next
to Aphelotoma.
Type, A, coruscans^ sp. n.
In size and colour these insects resemble some of the
Mutillidae.
Auchenophorus coruscans, sp. n.
$ . Clypeus broad, rounded anteriorly, the apical margin
depressed ; head shining, very minutely punctured ; a short,
delicate, longitudinal carina from between the antennae
reaching halfway to the anterior ocellus. Pronotum convex,
depressed anteriorly, narrow, with a slight depression in
the middle of the posterior margin, which is strongly
depressed. The sides of the prothorax are concave and
the posterior angles do not reach the base of the wings.
Mesonotum short and broad ; the whole thorax shining,
minutely punctured. Median segment long, flat above,
abruptly truncate posteriorly, very coarsely rugose ; a
median carina starting from the base and continued to the
apex on the dorsal surface and also on the surface of the
truncation, with a carina on each side converging from the
base and meeting above the truncation, forming an isosceles
triangle; the surface of the truncation is irregularly trans-
versely striated. Abdomen pseudosessile, smooth and
shining, the first segment long, the apical segment pointed.
Bright metallic blue ; the mandibles, labrum, and scape of
the antennae dark ferruginous. Antennae fuscous. Pro-
thorax, mesonotum, tegulae, anterior tibiae above, the three
basal abdominal segments and the base of the fourth bright
ferruginous red. Tibiae and tarsi fuscous. Wings hyaline,
crossed by two fuscous bands, one crossing the base of the
discoidal cell, the other broader, from the radial cell across
the apex of the cubital cell. Xervures black.
Length 10 mm.
cJ unknown.
Hab. Mackay, Queensland (October and November).
Auchenophorus ceneuSy sp. n.
$ . Clypeus broad, rounded anteriorly, with a straight
transverse carina before the anterior margin, which is
depressed. Head and thorax opaque, very finely reticulate ;
272
Mr. R. E. Turner on
tlie liead broadly emarginate posteriorly ; pronotum narrowed
anteriorly, the posterior angles elevated, subtuberculate ;
inesonotum short. A depressed transverse line, smooth and
shining, at the base of the scutellum. Median segment sub-
quadrate, flat above, obliquely truncate posteriorly, rugose
above, the lateral margins raised, forming carinse ; a median
Carina from the base, extending along both the dorsal surface
and the surface of the truncation to the apex, a carina on
each side converging from the base and meeting before the
truncation, forming an equilateral triangle, the space included
longitudinally striated ; the surface of the truncation trans-
versely striated. Abdomen subpetiolate, sharply pointed at
the apex; shining, the segments almost smooth at the base,
finely punctured along the apical margins.
Opaque black ; the median segment metallic blue-green ;
the abdomen shining bronze-green ; the mandibles, the scape
and two apical joints of the antennae, the mesopleurse, the
sides of the median segment, the prosternum, the base of the
first abdominal segment, and the legs ferruginous. The
femora and coxae marked with green-bronze above. Wings
hyaline, crossed by two fuscous bands, one crossing the base
of the discoidal cell, the other broader and close to the apex.
Length 8 mm.
$ unknown.
JIah. Mackay, Queensland (February).
d-he joints of the maxillary palpi are much longer in this
species than in coruscans ; the three apical joints are elongate
as in the genus Aphelotoma, Westw.
Auchenophorus falvicornisj sp, n.
^ . Head opaque, finely reticulate, the eyes converging
towards the vertex, the ocelli placed between the eyes near
their apex. Antennae as long as the head and thorax united,
the scape short, the two apical joints stouter than the others.
The three terminal joints of the maxillary palpi much elon-
gated. Pronotum short, much depressed anteriorly, broadened
posteriorly, the angles prominent but not spined. Meso-
notum finely reticulate, opaque, not reaching the posterior
margin of the tegulse. Median segment short, flat above,
obliquely truncate posteriorly, the margins of the dorsal
surface raised, forming an enclosed space, rounded at the
apex and longitudinally striated within ; the surface of the
truncation obliquely striated at the base, smooth at the apex.
Abdomen subpetiolate, shining, the two basal segments
smooth, the rest very finely and closely punctured.
273
new Spliegidse from Australia.
Black ; the antennae and the first abdominal segment at
tlie base testaceous; the prothorax, the mesopleurse, the sides
of the median segment, and the legs ferruginous ; tlie median
segment aboye obscure bt-on^e^ green, testaceous at the
extreme apex ; the abdomen very dark bronze-green. Wings
hyaline, nervures fuscous ; a broad fuscous band crossing the
discoidal cell, and another, broadly interrupted in the middle,
near the apex.
Length 5 mm.
$ unknown.
Hah. Kuranda, near Cairns, Queensland (January).
Genus Psen, Latr.
Psen lutescenSj sp. n.
$ . Clypeus broader at the apex than at the base, as long
as broad, covered with silver pubescence; the eyes diverging
towards the clypeus and towards the vertex. Head shining,
the front delicately reticulate, with a fine median sulca from
the anterior ocellus; a large tubercle between and below the
ba,se of the antennae ; from the base of the tubercle on each
side springs a carina, which skirts the base of the antennae
below, but does not reach the eye. The vertex is broad,
smooth, and polished ; the ocelli are large. Thorax shining,
finely punctured, the pronotum linear, transverse, very
broad ; the scutellum transverse, with a slightly depressed
row of strong punctures at the base. Median segment con-
vex, oblique, with a transverse depressed space at the base
and a very deep median sulca from the base to the apex ;
finely rugulose. Petiole long ; abdomen shining, very finely
punctured.
Yellow ; the abdomen and posterior legs luteous ; the
head, a square spot on the middle of the mesonotum, the
depressed line at the base of the scutellum, and a transverse
mark on each side of the scutellum black ; the antennae
fuscous ; the scape and mandibles yellow. Wings hyaline,
iridescent. Tlie first recurrent nervure is interstitial with the
first transverse cubital nervure, the second recurrent nervure
is received by the third cubital cell near the base.
Length 9 mm.
S unknown.
Hah. Mackay and Cairns, Queensland (November to
February).
Ann. (So Mag, N. Hist. Ser. 7. Vol. xix.
19
274
Mr. E. E. Turner on
Genus POLEMISTUS, Sauss.
Polemistus exul, sp. n.
$ . Mandibles very broad at the apex, produced into a
tooth at the outer angde, clypeus slightly produced at the
apex, smooth. Head narrow, the eyes just touching the base
o£ the mandibles, divergent towards the vertex. Front very
long, concavely hollowed ; the scape of the antennse long,
resting in the hollow, the flagellum short and stout. The
ocelli placed on the vertex near the posterior margin. A
row of large punctures surrounding the eyes, interrupted
narrowly at the base of the mandibles ; a transverse, emargi-
nate carina between the eyes above the concave front, a
longitudinal carina running from the anterior ocellus to the
transverse carina, and another short one on each side,
separated from the eye by the row of punctures only. Pro-
notum very short, transverse, showing only a row of large
punctures, the anterior margin straight, with prominent
angles. Mesonotum longer than broad, with a longitudinal,
median double carina, the very narrow space enclosed by the
caringe punctured ; on each side are two longitudinal carinae,
each with a row of punctures on the outer side, and another
short arched carina above the tegula. Scutellum broad,
smooth, broadly truncate at the apex, with a punctured de-
pression at the base divided by a very short longitudinal
carina ; and a depressed transverse row of punctures at the
apex. Median segment longer than broad, abruptly truncate
at the apex, very coarsely rugose ; the surface of the trun-
cation concave, divided by a flne median sulca, rugulose.
Abdomen short, smooth, and shining ; the first segment
vertically truncate anteriorly, with a short petiole ; the
apical segment compressed, acute. The legs smooth, not
spinose.
Black ; the mandibles at the apex ferruginous ; the
antennse, legs, and tegulse testaceous ; the vertex of the head,
the thorax, and the median segment dark green-bronze.
Wings hyaline, nervures testaceous.
Length 5 mm.
Hab. Mackay, Queensland (December to April) .
Two species only of this genus have been described, both
by Saussure, the localities being Madagascar and Mexico.
Genus Paracrabro, nov.
? . Head very large and broad, half as wide again as the
thorax ; the mandibles large, very strongly tridentate at the
new Spliegldae //*c>m Australia, 275
apex. Cljpeus small, transverse, truncate anteriorly. The
antennae far apart at the base, inserted low down, close to
the base of the clypeus, the scape long, much more than half
the length of .the flagellum, which is short. Front broad,
concavely hollowed. Eyes entire, reaching the base of the
mandibles, convergent towards the vertex; cheeks and vertex
very broad ; the ocelli placed close together between the
eyes, near their apex, far removed from the posterior margin
of the head. Pronotum very short, almost perpendicularly
depressed, narrower than the mesonotum ; scutellum trans-
verse. Median segment with a broadly subtriangular enclosed
space at the base, rounded and truncate posteriorly. Abdo-
men petiolate, the petiole linear and rather short. Fore wing
with two cubital cells ; the first quadrilateral, the width
about two thirds of the length, receiving the recurrent
nervure some distance before the apex ; the second less than
half the length of the first, almost square. The radial cell is
long and pointed, almost reaching the apex of the wing, not
appendiculate. The medial cell of the hind wing is pro-
duced at the apex, reaching downwards from the costa, the
anal cell is long. Legs short, tibiae feebly spinose, the tarsi
much longer than the tibiae.
The neuration is similar to that of StigmuSj but the general
form is very near Crahro.
Type, P. Froggattij sp. n.
Paracrahro Froggaiti, sp. n.
? . Head very large, quadrilateral, almost smooth, sub-
opaque. Mesonotum sparsely and very minutely punctured,
with a faintly impressed median line, and a faint longitudinal
one on each side, not reaching the posterior margin. Median
segment with a broad subtriangular space at the base enclosed
by carinae, the enclosed space longitudinally rugose striate ;
the segment behind the carinae oblique, rounded and striate,
then abruptly truncate and rugose to tho apex. Abdomen
smooth and shining.
Black ; the mandibles, antennae, tegulae, and legs fulvo-
ferruginous. Wings hyaline, tinged wdth yellow; nervures
testaceous at the base, fuscous at the apex.
Length 7 mm.
Hah. Victoria.
Genus Sericophorus, Sm.
SericopJiorus dipteroides, sp. n.
? . Clypeus broad, finely punctured, depressed transversely
276
On new SpliegldaB from Australia.
before the apex, the apical margin subtruncate. Head
broad, very delicately reticulate, eyes convergent towards
the vertex ; antennai very short, the apical joints thickened.
Pronotum very short, depressed below the mesonotum, which
is broad, opaque, and very delicately reticulate. Median
segment short, oblique, vertically truncate posteriorly, a
transverse depressed row of coarse punctures at the base,
from which a similar longitudinal row runs down the middle
of the segment, the rest of the segment smooth. Abdomen
almost smooth, triangular, the first segment vertically trun-
cate anteriorly, the surface of the truncation concave.
Castaneous; the head black; the antennae and clypeus
castaneous ; the mandibles dark ferruginous, black at the
apex ; the second to the fifth abdominal segments blackish
brown, the apical margin of the segments testaceous. Wings
hyaline, nervures fuscous.
Length 7 mm. ; expanse of wings 13 mm.
Hab. Cairns, Queensland (April) .
Nearest to S. hicolor^ Sm., from the Swan Liver.
SericopJiorus funehris^ sp. n.
$ . Clypeus broad, much elevated in the centre, projecting
at the apex almost at right angles. Head opaque, almost
smooth, the front above the clypeus depressed concavely,
clothed with silver pubescence ; a very short, longitudinal,
median carina above the base of the antennae. Pronotum
short, strongly depressed ; mesonotum large, very delicately
reticulate. Median segment almost smooth, narrowed from
the base, truncate posteriorly, with a median carina from the
base continued on the surface of the truncation to the apex,
a small tubercle on each side of the carina just above the
truncation, the lateral margins of the segment raised at the
base, forming short carinse ; the sides of the segment
obliquely striate. Abdomen shining, very delicately punc-
tured, the apical margin of the segments smooth ; the first
segment is not truncate, and is narrower at the base and
longer than in the allied species.
Black ; the mandibles yellow, an interrupted line on the
pronotum and the tegulse pale yellow ; the tibia3, tarsi, and
the apex of the femora testaceous. Wings hyaline, nervures
testaceous.
Length 6 mm. ; expanse of wings 9 mm.
Hab. Mackay, Queensland (November).
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. 112. APRIL 1907.
XXXIN.—Rhynchotal Notes,--XlA, By W. L. DISTANT.
Fam. FtilgoridaB (continued from vol. xviii. p. 356).
Subfam. CixiiNJs.
Genus Mundopa.
Mundopa, Dist. Faun. B. I., Rhynch. iii. p. 263 (1906).
Type, M. cingalensis^ DIst.
Mundopa pollens,
Cixius pallens, Walk. Journ. Linn. Soc., Zool. i. p. 149 (1857).
Hob. Borneo.
Bajauana, gen. nov.
Head somewhat broad, the vertex widened anteriorly, its
lateral margins strongly ridged, the anterior and posterior
margins less strongly so, base angularly emarginate; face
moderately broad and angularly marginally ampliate beyond
middle, anterior margin truncate, surface flat, lateral margins
strongly ridged, with a central longitudinal ridge which
is continued through clypeus, containing a very indistinct
ocellus a little before clypeus; pronotum exceedingly short ;
mesonotum v^ry long, almost twice as long as vertex and
Ann, (k Mag, N, Hist, Ser. 7. Vol, xix. 20
278
I\Tr. W. L. Distant on iromoptera.
pronotum together, trlcarinate, the interspaces between tlie
carinations forming somewhat raised flat processes, the bases
of wliich are angularly narrowed ; legs moderately short and
robust; tarsi robust, first joint longest ; tegmina widened or
ampliated towards apex, costal margin moderately sinuate
from base to stigma, before the latter it is distinctly notched,
stigma elongate, a series of irregularly placed transverse veins
near apex defining a series of longitudinal apical areas ; claval
vein extending to a little beyond middle of claval margin.
Type, B. rufula^ Walk. (^Brixia).
Bajauana rnfula.
Brlvia rufula, Walk. Journ. Linn. Soc., Zool. x. p. lOG (1837).
JIah. New Guinea.
Bajauana tenehrosa.
Brixia tenehrosa^ Walk. Journ. Linn. Soc., Zool. x. p. 103 (1867).
llah. Ceram.
Bajauana palUcepff.
Brixia palliceps, Walk. Journ. JJnn. Soc., Zool. x. p. 106 (1867).
llah. My sol.
Bajauana marginafa.
Brixia marginata, Walk. Journ. Linn. Soc., Zool. x. p. 107 (1867).
Hah. New Guinea.
Bajauana varia.
Brixia varia, Walk. Journ. Linn. Soc., Zool. x. p. 108 (1867).
JIah. Sula.
Bajauana variegata.
Brixia variegata, Walk. Journ. Linn. Soc., Zool. x. p. 108 (1867).
Hah. Sula.
Bajauana hicolor.
Brixia hicolor, Walk. Journ. Linn. Soc., Zool. x. p. 100 (1867).
Ilab. New Guinea.
Bajauana puncticosta.
Brixia puncticosta. Walk. Journ. Linn. Soc., Zool. x. p. 109 (1867).
Hah. My sol, Morty.
279
^fr. W. L. Distant on Ilonioptera.
Bajauana tr aliens.
Cixius trdhemy Walk. Journ. Linn. Soc., Zool. i. p. 149 (1857).
Hah. Borneo.
Genus Brixia.
Biixia, StSl, Ofv. Vet.-Ak. Fork. 1856, p. 162.
Type, B. natalicola, Stal.
Brixia migratoria, sp. n.
Head, pronotum, and abdomen above brownish ocliraceous ;
lateral angles of pronotum and mesoiiotum ocliraceous ;
abdomen beneath, sternum, and legs stramineous ; abdomen
above more or less cretaceously tomentose ; tegmina very
pale brownish ocliraceous, the veins thickly minutely spotted
with fuscous, three transverse spots on costal area, the central
one connected with a broad transverse discal fascia, a spot
near apex of costal margin, a spot on inner margin beyond
end of clavus, and an oblique streak at base chocolate-
brown ; wings pale iridescent creamy white ; face extending
considerably in front of and above eyes, the margins strongly
ridged, centrally carinate.
Long., excl. tegm., mm.; exp. tegm. 10 mm.
Hah. Queensland {F. P. Dodd, Brit. Mus.).
This appears to be the first described Australian species of
this Oriental, Malayan, and Ethiopian genus. Brixia has
already been traced to New Guinea.
Hamba, gen. nov.
Head with the vertex narrow, longer than broad, with two
longitudinal carinations and the lateral margins also strongly
raised ; face elongately subtriangular, the anterior margin
convex, widened towards clypeus, with the lateral margins
strongly ampliate and moderately recurved, very strongly
centrally longitudinally carinate ; clypeus triangular, cen-
trally carinate ; pronotum small, centrally tricarinate ; meso-
notum tricarinate ; tegmina elongate, narrow, more than
twice longer than broad, costal margin moderately convex,
apical margin rounded, inwardly obliquely widened from apex
of claval area; veins longitudinal, some discal transverse
veins a little beyond middle, and a series of continuous trans-
verse subapical veins defining a series of short apical areas ;
wings a little wider than tegmina ; legs moderately short and
robust, basal joint of tarsi very long.
Type, H. perjplexa, Walk.
20*
280
Mr. W. L. Distant on ITomoptera.
Hamha perplexa.
Civius perplexus, Walk. Journ. Linn. Soc., Zool. i. p. 147 (1857).
Hah, Borneo.
Hamha inclinata.
Cixius inclinatus, Walk. Journ. Linn. Soc., Zool. i. p. 147 (1875).
Hah. Borneo.
Taloka, gen. nov.
Head with the vertex broad, a little longer than broad, mode-
rately ampllated anteriorly, tricarinate, the lateral carinations
curved inwardly anteriorly and from each side of which there
emerges a strong oblique caririation in front of eyes ; face
very large, clypeus very small, the first tricarinate, the lateral
carinations oblique and terminating considerably before
clypeus, a prominent transverse rugosity on each side of
central carination at region of eyes, and the appearance of an
obsolete ocellus on each side of the same before clypeus ;
pronotum considerably shorter than head and centrally tri-
carinate ; mesonotum indistinctly seen owing to insertion of
pin in the unique typical specimen, but apparently strongly
tricarinate ; tegmina moderately broad, with two series of
transverse veins and with a distinct ovate cell beyond claval
area ; legs moderately short and robust.
Type, T. opaca^ Walk.
Taloka opaca.
Brixia opaca, Walk. Journ. Linn. Soc., Zool. x. p. Ill (1867).
Hob. New Guinea.
Genus Oliarus.
Oliarus, StSl, Berl. ent. Zeitschr. vi. p. 306 (1862).
Type, 0. Walkeri^ Stal.
Oliarus maculifrons.
Cixius maculifrons, Walk. List Horn. ii. p. 343 (1851).
Hah. Sierra Leone.
OUnriis vicarius.
Cixius vicarius, Walk. List Horn. ii. p. 343 (1851).
Hah. Florida.
281
Mr. W. L. Distant on Iloinoptera.
Oliarus testaceus.
Cixius testaceus, Walk. List Horn. ii. p. 344 (1851).
Hab. ?
Oliarus simplex.
Cixius simplex, Walk. Journ. Linn. Soc., Zool. i. p. 147 (1857).
Hab. Borneo.
Oliarus reductus,
Cixius reductus, Walk. Journ. Linn. Soc., Zool. x. p. 105 (1867).
Hab. Mjsol.
Oliarus intertectus,
Brixia intertectus, Walk. Journ. Linn. Soc., Zool. x. p. 114 (1867).
Hab. New Guinea.
Oliarus privatus.
Brixia privata, Walk. MS.
Hab. Mysol.
The unique typical specimen is without head, and therefore
description at present is unwarranted.
Oliarus Melichari, nom. nov.
Oliarus frontalis, Melich. Wien. ent. Zeit. xxiv. p. 282 (1905) : nom.
praeocc. Melich. (1904).
Hab. E. Africa.
Oliarus funebris.
Cixius funehris. Walk. Ins. Saund., Horn. p. 41 (1858).
Hab. Natal.
Oliarus modicus.
Cixius modicus. Walk. Journ. Linn. Soc., Zool. i. p. 148 (1857).
Hab. Borneo.
Oliarus latifrons.
Cixius latifrons. Walk. List Horn. ii. p. 844 (1861).
Hab. ?
Oliarus fulvus.
Cixius fulvus. Walk. List Horn., Suppl. p. 77 (1853).
Hah. Santarera.
282
^Ir. W. L. Distant on Homoptera.
Oliarus efferatus,
Cixius ejferatus, Walk. Journ. Linn. Soc., Zool. i. p. 87 (18.56).
IJah. Singapore.
Oliarus subpunctatus,
JBrixia subpimctata, Walk. Joum. Linn. Soc., Zool. x. p. 112 (1867).
Hab. Flores.
Oliarus dingkana^ sp. n.
Head and pronotum black ; vertical ridges to head, mar-
gins of pronotum, and carinae to mesonotum pale testaceous ;
face and cljpeus black, margins and a central carination
traversing both, pale testaceous ; body beneath imperfectly
seen in carded specimens now before me ; legs pale testaceous,
femora more or less testaceous with their apices ochraceous ;
tegmina hyaline, the venation and stigma piceous, interior
margins to about end of clavus piceous ; vertex slightly nar-
rowed anteriorly, the margins strongly ridged ; face with the
base truncate, obliquely ampliated towards clypeus, before
which it is a little convex, the central carination is a little
bifurcate at base, enclosing a small black sulcus ; pronotum
short, angularly narrowed to between base of eyes ; meso-
notum with five carinations, the two outermost strongly bent
beyond middle ; tibiae deeply longitudinally grooved, posterior
tibiae without spines.
In some specimens the outer mesonotal carinations are
black, and the interior margin of the tegmina only piceous at
apical half of clavus.
Long., inch tegm., 7 mm.
Hab. Queensland ; Peak Downs (Colls. DIst. and Biit.
Mus.).
Oliarus lubrOy sp. n.
Allied to the prectding species, but larger ; lateral margins
of face from beyond middle and just extending to base of
clypeus ochraceous; mesonotum much broader, with the
lateral angles pale testaceous ; tegmina broader and the
transverse veins on a})ical area more pronouncedly piceous.
Long., inch tegm., 8 mm.
Hab. Queensland ; Peak Downs (Colls. Dist. and Brit.
Mus.).
I have placed cotypes of this and the preceding species in
the National Collection.
Mr. W. L. Distant on Hoiiioptera.
283
Genus Mnemosyne.
Mnemosyne, StSl, Hem. A.t‘r. iv. p. 1-50 (1866).
Type, xl/. plahicpps^ Fabr.
Mnemosyne planiceps.
Flata planiceps, Fabr. Syst. Rhyng. p. 48 (180-3).
Civius columbue, Walk. List Horn. ii. p. 339 (1851).
Hab. Central America ; Cuba.
Mnemosyne Dohertyi^ sp. n.
Head, pronotum, and mesonotum castaiieous ; abdomen
above piceous; face, sternum, and legs ferruginous brown ;
abdomen beneath dark castaneous, the segmental margins
ochraceous ; tegmina subhyaline, trie- like, sprinkled with
minute ferruginous setigerous spots or granules, veins pale
castaneous, stigma, upper apical margin, a sinuate streak
from end of radial area bifurcating to middle of apical margin
and outer angle of interior margin, claval area and inner
margin ferruginous brown, outer half of stigma piceous ;
wings very pale fuliginous, the veins and extreme base
piceous, apical maigin narrowly ferruginous brown ; vertex
a little narrowing anteriorly, lateral margins ridged and with
a more obscure central carination ; face centrally ridged,
posteriorly angularly emarginate ; clypeus elongate, globose,
finely centrally ridged ; pronotum very finely punctate, with
a central longitudinal ridge; mesonotum with five longitudinal
tarinations, those on each side of the central carination more
discal and obscure ; tegmina elongate, apices angularly
rounded ; anterior legs thickened.
Long., excl. tegm., 10^ mm.; exp. tegm. 24 mm.
Hab. South-east Borneo {Doherty).
Allied to M. phiiippina^ Stal.
Mnemosyne cameranensis, sp. n.
Head, pronotum, and mesonotum castaneous, extreme apex
of mesonotum and abdomen above testaceous, the segmental
margins ochraceous, anal appendage piceous ; face, clypeus,
sternum, and legs brownish testaceous ; abdomen beneath
piceous, the segmental margins pale ochraceous ; tegmina
subhyaline, talc-like, the veins ferruginous brown, a series of
similarly coloured spots round the apical margin, of which
the two lowermost near apex of inner margin are largest,
284
Mr. W. L. Distant on Homoptera.
stigma pale ocliraceous, with its outer half ferruginous brown ;
wings very slightly fuliginous, the veins fuscous, apical
margin ferruginous brown ; vertex slightly narrowed ante-
riorly, the margins ridged, centrally somewhat obscurely
longitudinally carinate; face and clypeus with a continuous
central fine carination, face anteriorly convexly narrowed,
posteriorly strongly angularly emarginate ; pronotum short,
centrally carinate ; mesonotum with five carinations, the one
on each side of the central carination more discal and obscure ;
tegmina with the apex rounded, the stigma moderately
small and narrow; anterior legs not prominently incrassated.
Long., excl. tegm., 1\ mm. ; exp. tegm. 17 mm.
Hob, Cameroons {Escahra, Brit. Mus.).
Genus Bodecia.
Bodecia, Walk. Journ. Linn. Soc., Zool. x. p. 117 (1867).
Type, B. varipesy Walk.
The unique type of this genus is in a most mutilated
condition, and its proper location is a question of doubt and
difficulty. The structure of the tegmina apparently allies it
to Mnemosyne^ near which I have (awaiting better and further
material) placed it.
Genus Cixius.
Cixius, Latr. Hist. Nat. Ins. xii. p. 310 (1830-4).
Type, C. nervosus, Linn.
Cixius per sicus^ nom. nov.
Cixius longipennis, Melich. Annuaire Mus. St. P^tersb. vii. p. 86 (1902) :
nom. preeocc. Walk. (1851).
Cixius merula^ sp. n.
Vertex of head and pronotum ochraceous, disk of vertex
more or less piceous ; face black, basal margin and lateral
and central ridges ochraceous; mesonotum entirely jet-black ;
tegmina subhyaline with a very pale ochraceous tint, thickly
and minutely speckled or spotted with fuscous ; body beneath
and legs ochraceous, lateral areas of sternum, clypeus, and
basal .and lateral margins of abdomen black; vertex some-
what broad, narrowed anteriorly, margins very strongly
ridged and with a more or less obsolete central ridge ;
285
Mr. W. L. Distant on Homoptera.
face narrowed anteriorlj, moderately roundly ampliated to-
wards clypeiis, central and lateral ridges very pronounced ;
mesonotum with the disk a little depressed, tricarinate ; the
central carination weakest ; sternal segmental margins
ridged.
Long., inch tegm., to 4^ mm.
Hah. Queensland ; Peak Downs (Colls. Dist. and Brit.
Mus.).
Somewhat allied in general appearance to the Palsearctic
C. simplex, Herr.-Sch.
Genus Benna.
Benna, Walk. Journ. Linn. Soc., Zool. i. p. 90 (1856).
Type, B. capitulata, Walk.
Benna guttata.
Brixia guttata^ Walk. Journ. Linn. Soc., Zool. x, p. 110 (1867).
ditiasa, Walk. MS.
Hah, New Guinea, Sula.
Species in too mutilated a condition for Generic Determination,
and some prohahly not even belonging to this Subfamily.
nanula.
Brixia nanula^ Walk. Journ. Linn. Soc., Zool. x. p. 112 (1867).
Hah. New Guinea.
concinnula.
Brixia concinnula, Walk. Journ. Linn. Soc., Zool. x. p. 110 (1867).
Hah, Sula.
ruhiginosa, MS. ?
despectus.
Cixius despectus, Walk. Journ. Linn. Soc., Zool. i. p. 148 (1857).
Hah. Borneo.
— vilis.
Cixius vilis, Walk. Journ. Linn. Soc., Zool. i. p. 148 (1857).
Hah. Borneo.
286
j\Ir. W. L. Distant on Moino{>tera.
e quits.
Ci.rfus equKS, ^^'alk. Journ. Liun. Soc., Zool. i. p. 147 (1857).
Ilah. Borneo.
Jiffinis.
Civius diffinis, Walk. Journ. Linn. Soc., Zool. i. p. 146 (1857).
linh. Borneo.
congrua.
Brixia congrua^ Walk. Journ. Liun. Soc., Zool. x. p. 110 (1867).
Ilah. Mysol.
terminalis,
Brixia terminalis, Walk. Journ. Linn. Soc., Zool. x. p. Ill (1867).
Ilab. New Guinea.
muriitus.
Cixius munitus, Walk. Journ. Linn. Soc., Zool. i. p. 149 (1857).
Uab. Borneo.
• suhlucida.
Brixia sublucida, Walk. Journ. Linn. See., Zool. x. p. 1C7 (1867).
Hab. New Guinea.
insuetus,
Cixius insuetus, Walk. Journ. Linn. Soc,, Zool. i. p. 150 (1867).
Uab. Borneo.
deductus.
Cixius deductus, Walk. Journ. Linn. Soc., Zool. i. p. 149 (1867)
JJab. Borneo.
atratula.
Brixia aU alula, ^^■alk. Journ. l.inn. Soc., Zool. x. p. 107 (1867).
Uab. IMoity,
287
]\Ir. AV. L. Distant on ITomopfera.
Subfam. Tropiducrinje.
Genus Ficakasa.
Ftcarasa, "Walk. Journ. Linn. Soc., Zool. i, p. 162 (1857)
Type, F, pallida j W alk.
Ftcarasa straminea^ sp. n.
Body and legs dark stramineous ; tegmina pale strami-
neous ; wings pale hyaline ; vertex more than twice as broad
as long, marginally and centrally carinate, anteriorly convex,
laterally straight, posteriorly strongly angularly emarginate ;
f)ronotum about as long as vertex, centrally and laterally
carinate, strongly subconically produced between eyes, pos-
terior margin angularly emarginate ; mesonotum longer
than vertex and pionotum together, ti icarinate, the lateral
carinations obliquely incurved anteriorly and joining the
central carination a little befoie base of pronotum ; face
broad, but longer than broad, the lateral margins curved
outwardly at about two thirds from base and then obliquely
narrowed to clypeus, anterior and lateral margins strongly
caiinate, the disk ti icarinate, an oblique carination on each
side of the central one, commencing a little before clypeus
and terminating near anterior angles; tegmina with the
costal area strongly transversely veined, apical area divided
by two transverse series of transverse veins, many of the
outer series bifurcate.
Long., excl. tegm., 9 mm. ; exp. tegm. 27 mm.
Hah. Malay Aichipelago ; Ilalmaheira {W, Doherty).
Ftcarasa australasice^ sp. n.
Pale ochraceous, in fresh specimens probably pale virescent ;
vertex broader than long, anterior and lateral margins
ridged, centrally longitudinally carinate ; face broadly
elongate, a little ampliate towards clypeus, anterior and
lateral margins ridged, centrally longitudinally carinate ;
clypeus centrally ridged, not much more than half the length
of face ; pronotum angularly produced between the eyes,
tricarinate ; mesonotum tricarinate ; tegmina with the costal
area obliquely transversely veined, two series of transverse
veins beyond middle enclosing a space much narrower than
that between it and the apical margin ; posterior tibiae with
three spines.
Long., excl. tegm., mm. ; exp. tegm. 21^ to 23 mm.
llab. Queensland; Kuranda [F. P. Dodd^ Brit. Mus.).
288
Mr. W. L. Distant on llomoptera.
Magia, gen. nov.
Vertex wide, a little longer than broad, flat, lateral mar-
gins straight, anterior and lateral margins ridged, centrally
longitudinally carinate ; face with the lateral margins a little
concave, prominently angularly ampliate on each side of
posterior margin, tricarinate, the lateral carinations curved
and anteriorly joining the central carination, clypeus centrally
ridged ; pronotum a little shorter than vertex, not produced
beyond base of head, tricarinate ; mesonotum about as long
as pronotum and vertex together, tricarinate, the lateral
carinations posteriorly outwardly directed; abdomen equal in
length to space between anterior margin of vertex and
posterior angle of mesonotum ; posterior tibiae with two
spines ; tegmina somewhat long and narrow, anterior claval
area with three transverse veins, apical area with numerous
transverse veins, some of which form a continuous apical
submarginal series ; wings narrow, two series of transverse
veins beyond middle.
Type, M. subocellata, Dist.
Magia suhocellataj sp. n.
Body and legs pale ochraceous ; two spots at anterior
margin of face, two small spots at base of anterior coxae, a
spot on each lateral margin of mesosternum, lateral and
posterior margins, lateral carina3, two spots between them,
and two very small spots near each lateral angle of pronotum,
lateral margins, carinae, two spots near each side of anterior
margin, and two very small spots near base of mesonotum,
posterior margins and angles of abdominal segments above,
and the tarsal claws, black; carinations to face testaceous ;
tegmina subhyaline, very slightly pale fuliginous, the
venation fuscous, stigma ochraceous, a lower subapical black
spot with a white pupil at its upper margin ; wings sub-
hyaline, the venation fuscous.
Long., excl. tegm., 8 mm.; exp. tegm. 21 mm.
Hab. Queensland, Kuranda {F. P, Dodd^ Brit. Mus.).
Genus Paricana.
Paricana, Walk. Journ. Linn. Soc., Zool. i. p. 158 (1857).
Type, P. dilatipennisj Walk.
Paricana curvifera.
eurvifera^ Walk. MS.
Vertex, mesonotum, abdomen above, body beneath and
289
Mr. W. L. Distant on Homoptera.
legs ochraceous ; pronotum, base and apical area of abdomen
above, and a central fascia to face and scutellum testaceous
red ; posterior angle to mesoiiotum black ; legs ochraceous,
the tarsi and posterior femora piceous ; tegmina hyaline, the
veins ochraceous, margins, the stigma and a curved fascia
connecting it with inner margin ferruginous brown ; vertex
very narrow ; face smooth and shining, the central reddish
area a little raised and grooved on each side ; pronotum and
mesonotum tricarinate, the latter with the lateral carinations
curved and anteriorly joining the central carination ; posterior
tibiae with two spines.
Long., inch tegm., 8 mm.
Hah. Aru {Wallace^ Brit. Mus.).
This species was standing under the genus Gixius in the
British Museum and labelled curviferay Walk., but 1 cannot
trace any published description.
Subfam. Achilin^.
Genus Faventia.
Faventia, Stal, Hem. Afr. iv. p. 181 (1866).
Type, F, pustulataj Walk.
Faventia guttifer.
Cixius guttifer^ Walk. Journ. Linn. Soc., Zool. i. p. 146 (1857).
Hah. Borneo.
Mahuna, gen. nov.
Vertex of head about as long as broad, a little pointed in
front, lateral margins oblique and carinate, centrally more
obscurely carinate, basal margin broadly subconcave ; face
much narrower anteriorly than before clypeus, strongly
centrally ridged, lateral margins moderately ridged, roundly
ampliated before clypeus, which is only about half its length
and centrally and laterally carinate ; pronotum narrow and
centrally carinate ; mesonotum elongate, about twice as long
as head and vertex together, tricarinate; abdomen broad
and flattened; legs simple; tegmina somewhat long and
narrow, apical margin rounded, claval margin subangularly
dilated near base and then narrowed to claval apex, thence
obliquely widened to apex, claval vein apically curved down-
ward and terminating at about one third before apex, a few
transverse veins near apex of costal margin ; wings con-
siderably broader than tegmina.
Type, M. co7ispersa, Dist.
290
Mr. AV. L. Distant on Ilomoptera.
]\[ahuna conspersa, sp. n.
Dody ocliraceons or greenish ocliraceous; legs paler,
atiterior and intermediate tibiae darkly annulated ; tegmina
creamy white, opaque, sprinkled with fuscous, more pro-
minently spotted on costal and apical margins, two small
piceous spots on claval margin, one near middle, the other
at apex; wings very pale fuliginous.
Long., excl. tegm., 5^ mm. ; exp. tegm. 15 mni.
Hah. Queensland [F. I\ Dodd^ Brit. Mus.).
Tudea, gen. nov.
Head with the vertex about as broad as long, the lateral
margins veiy strongly ridged, centrally obscurely carinate,
the base subconcavely emarginate, face concave, anterior
margin truncate, lateral margins convexly rounded, centrally
obscurely carinate, about twice as long as clypeus, which is
obscurely centrally carinate and has its lateral margin
ridged ; rostrum reaching the intermediate coxae ; pronotum
a little longer than vertex, anteriorly produced between eyes,
tricarinate, the lateral carinations almost meeting in front of
the central carination ; mesonotum almost twdce as long as
pronotum and vertex together, obscurely tricarinate; abdo-
men above centrally ridged and obliquely deflected on each
lateral area ; legs simple, posterior tibiae with a short sub-
central spine, posterior tarsi wdth the basal joint a little
longer than the other two joints together ; tegmina moderately
broad and short, apex truncately rounded, interior margin of
clavus angularly dilated and then obliquely narrowed to
claval apex, thence obliquely widened to apical margin, radial
area very broad, a few transverse veins near apex of costal
margin, veins on apical area longitudinal, almost entirely
wdthout transverse veins, claval vein terminating at about
one third before apex ; wings broader than tegmina, more
densely longitudinally veined at apical angle.
Type, T. picturataj Dist.
Tudea piciurata.^ sp. n.
Head and pronotum ocliraceous ; eyes, central base of pro-
notum, and the mesonotum chocolate-brown ; abdomen above
and beneath virescent, the base greyish white ; face, sternum,
and legs greyish white, anterior area of face and lateral
margins of prosternum pale ocliraceous ; tegmina stramineous,
upper half sprinkled with dark spots, a large spot on dilated
291
Mr. W. L. Distant on Ilomoptera.
basal claval area, and a subapical marginal fascia broadly
bifurcating on interior margin, pale purplish brown, this
fascia is broken near upper apical angle ; wings creamy
while, hyaline.
Long., excl. t'egm., mm.; exp. tegm. 19-20 mrn.
Ilab. Queensland (F. P. Dodd^ JLut. Mus.).
Booneta, gen. nov.
Booneta, ^IS., Brit, Mas.
Head much shorter than pronotum, vertex shorter than
broad, the margins ridged, centrally carinate, the anterior
angles appearing prominent from above ; face long, con-
vex ly deflected, rather more than twice as long as clypeus,
nearly equally broad throughout, the lateral margins straight,
both face and clypeus strongly centrally and marginally
carinate; rostrum not passing the posterior coxag, the apical
joint elongate ; posterior tibiae unarmed ; pronotum about as
long as vertex, centrally tricarinate, the lateral carinations
curved and joining the central one at anterior margin ; rneso-
notum about as long as pronotum and vertex together, tri-
carinate, the lateral carinations sinuate; tegmina short and
broad, apical margin obliquely truncate, inner margin
obliquely arnpliate from end of clavus, costal area broad,
veins longitudinal, a transverse subapical line formed by
transverse veins, and three transverse veins on disk; wings
a little narrower than tegmina.
Type, B. ferruginea^ Walk.
Booneta ferruginea.
Cixius ferrtigineus, Walk. Journ. Linn. Soc., Zool. x. p. 104 (1867).
Oixius caliginosus, Walk, loc, cit. p. 105.
Ilah. New Guinea, Mysol.
Booneta lurida.
Cixius Iw’idus, Walk. Journ. Linn. Soc., Zool. x. p. 104 (1867).
Hah. Mysol.
OuwEA, gen. nov.
Head much narrower than pronotum, vertex short, very
much broader than long, its anterior lateral angles prominent ;
face about twice as long as clypeus, somewhat flat, with the
lateral margins strongly laminately reflexed, narrowing
towards clypeus, its posterior margin sinuated for the recep-
tion of the clypeus, centrally longitudinally carinate ; clypeus
292
J\Ir. W. L. Distant on Homoptera.
strongly centrally keeled, its lateral margins lammately
carinately reflexed ; rostrum with the apical joint elongate;
pronotum a little shorter than vertex, centrally carinate ;
rnesonotum about twice as long as pronotum and vertex
together, its disk moderately flatly raised, distinctly deflected
before pronotum, obscurely tricarinate ; abdomen broad, con-
siderably longer than space between apex of head and posterior
angle of rnesonotum ; legs simple, posterior tibise with one
spine, basal joint of posterior tarsi elongate ; tegmina with
the apical margin obliquely rounded, obliquely broadened
from apex of clavus, apical third of costal area obliquely
transversely veined, lower discoidal area very strongly
sinuated, a subapical sinuate transverse line consisting of
transverse veins, defining a series of longitudinal apical areas;
wings broader than tegmina.
Type, 0, Doddi^ Dist.
Ouwea Doddiy sp. n.
Body brownish ochraceous, paler beneath than above ;
rnesonotum with three longitudinal ochraceous lines above
the obscure carinations, its lateral and apical margins ochra-
ceous ; first and second abdominal segments above more or
less ochraceous; face and sternum mottled with ochraceous and
castaneous ; legs very pale ochraceous, femora obsoletely
annulated with brownish ; tegmina bronzy ochraceous,
beyond middle becoming darker between the veins, three
large spots in costal area, a large sublunulate apical spot,
some spots near posterior angle of inner margin, two spots
in claval area, and a discal spot, greyish white ; wings
fuliginous brown, the basal area very much paler and
subhyaline.
Long., excl. tegm., c? 7, ? 8 mm. ; exp. tegm., ^ 20-
21, ? 25 mm.
Hah, Queensland ; Kuranda [F. P. Dodd, Brit. Mus.).
Genus Rhotala.
Rhotala, Walk. Journ Linn. Soc., Zool. i. p. 152 (1857).
Errada, Walk. loc. cit. x. p. 116 (1867).
Type, R. delineata. Walk.
Head moderately conically produced in front of eyes,
centrally and laterally ridged, between the ridges the surface
foveate, face a little longer (not twice as long) than clypeus,
subconical, beyond middle rather abruptly convexly widened
to base of clypeus, both moderately longitudinally carinate,
Mr. W. L. Distant on Ilomoptera.
293
lateral margins of clypeus distinctly carinate; antennae
robust, longly produced, inserted a little beneath eyes, apical
joint longest and stoutest ; pronotum somewhat long and
tricarinate, the carinated area convexly produced between
the eyes, the posterior margin angularly sinuate, lateral
margins oblique ; mesonotum about as long as pronotum,
tricarinate, the lateral carinations inwardly obliquely bent
behind middle and joining the central carination at base;
femora slightly dilated, beneath strongly centrally carinate ;
posterior tibiae with five spines, the first near base, the fifth
and largest at about one fourth from apex ; tegmina narrow,
the costal margin slightly undulate and convex, apical
margin rounded, claval area inwardly angularly widened and
subobliquely narrowed to apex, beyond claval apex again
slightly widened, costal area obliquely transversely veined
from about middle, radial area obliquely transversely veined,
apical area longitudinally veined and containing a few obscure
transverse veins, at the base of apical area and from beneath
radial area an irregular dividing-line formed of transverse
veins, claval vein not reaching apex of claval area ; wings
about twice as broad as tegmina, and containing an irregular
series of subapical transverse veins.
In his description of this genus Walker makes no mention
of the five spines to the posterior tibiae, and in his figure of
the typical species only four are delineated.
Fowler (Biol. Centr.-Amer., Bhynch. Horn. ii. p. 137)
places the genus as of uncertain position, and writes it most
probably must be placed with the Cixiinae. He. however,
also states that the genus in some respects is closely allied to
Helicoptera. In this latter view I agree, and consider its
position is with the Achilinae.
Ehotala delineata.
Wiotala delineata, Walk. Journ. Linn. Soc., Zool. i. p. 152, t. vii. fig. 6
(1857).
Vertex of head and pronotum ferruginous ; mesonotum and
face piceous, the latter with the lateral margins flecked with
greyish ; body beneath piceous ; abdomen above pale testa-
ceous, basal segment and anal appendage castaneous brown ;
legs pale castaneous brown with ochraceous annulations;
tegmina pale greyish brown, the veins so far as commence-
ment of apical area spotted with castaneous, four somewhat
large castaneous spots in costal area and two on disk, the
dividing-veins at base of apical area suffused with dark
castaneous, apical area paler and more unicolorous, with
Ann, dc Mag, N, Hist, Ser, 7. Vol, xix. 21
294
Mr. W. L. Distant on Homoptera.
some greyish suffusions ; wings fuliginous, with the veins
fuscous ; face very finely and thickly granulose, somewhat
foveate, centrally longitudinally carinate, margin broadly
ridged ; carinations to vertex, pro- and mesonota strongly
pronounced.
Long., excl. tegm., 10 mm. ; exp. tegm. 24 mm.
Hah. Borneo ( Wallace, Brit. Mus.).
RTiotala philippinensisj sp. n.
Vertex of head and pronotum ochraceous, the carinations
a little paler in liue, lateral areas of the pronotum piceous;
mesonotum ochraceous, the disk between the central carina-
tions piceous, and an anterior marginal piceous spot on each
side of the outer carinations, posterior angle ochraceous ; face
and clypeus piceous, anterior angle or base of first ochraceous ;
body beneath more or less piceous ; legs ochraceous ; tegmina
ochraceous, the veins finely spotted with piceous, some larger
and somewhat obscure, irregularly shaped piceous spots on disk
and costal area, two distinct cretaceous white spots on disk
beyond middle, the outermost just touching the inner margin
of apical area, which is paler in hue and unspotted ; wings
pale fuliginous, a little darker at apex ; vertex of head
foveate above and at sides, the lateral margins distinctly
raised above the level of the pronotum, the latter with its
anterior margin conically produced to the latitude of the
anterior margins of the eyes ; posterior tibiae with five spines,
the two nearer base smaller and placed closer together ;
apical margin of tegmina obliquely rounded.
Long, (abdomen mutilated) ; exp. tegm. 21 mm.
Hah, Philippines {Whitehead, Brit. Mus.).
Talaloa, gen. nov.
Head narrower than the pronotum, vertex obliquely
narrowing to apex, its lateral margins strongly ridged, and
appearing as angularly prominent at apex ; face long and
narrow, but not longer than clypeus, elongately subtriangular,
broadening to clypeus, centrally and laterally ridged ; clypeus
long, slender, narrowing to apex, centrally and laterally
ridged ; rostrum long, its apical joint elongate ; antennse
robust, longly prominent, second joint nearly twice the length
of first ; pronotum shorter than vertex, centrally carinate ;
mesonotum longer than pronotum and vertex together, tri-
carinate; legs simple, posterior tibiae unarmed, basal joint
of posterior tarsi elongate; tegmina and wings subhyaline,
the first broadly rounded at apex, inner margin moderately
295
On a Freshwater Crustacean from Para,
obliquely ampliated from end of clavus, which is angularly
posteriorly dilated near base, a curved line formed by trans-
verse veins crossing tegmen at about one fourth from apex
enclosing a series of narrow longitudinal apical areas ; this is
preceded by three transverse veins enclosing four discoidal
areas ; wings a little broader than tegmina, three oblique
transverse veins on disk, apical veins mostly forked.
Allied to Rhotala^ Walk., by the robust and longly pro-
duced antennse; the tegmina give the appearance of a
Tropiduchid, but the lateral carinate margins to the clypeus
induce its being regarded as an Achilid.
Type, T. pallescens^ Dist.
Talaloa pallescens, sp. n.
Body above and antennae castaneous brown ; body beneath
and legs pale ochraceous ; abdomen beneath pale brownish,
with the posterior segmental margin and a central discal line
pale ochraceous ; posterior tarsi annulated with pale brownish ;
tegmina and wings pale subhyaline, the venation fuscous ;
tegmina slightly fuliginous, the stigma fuscous, traversed at
base by a creamy-white line which is widened posteriorly.
Long., excl. tegm., 4J-6 mm. ; exp. tegm. 14-16 mm.
Hab, Queensland; Kuranda [F,P. Dodd^ Brit. Mus,).
Genus ?
Eurial globulifera^ Walk, Ins. Saund., Horn. p. 108 (1858).
HaF ?
This species does not belong to the Tropiduchid genus
Euria,— Tropiduchus^ Stal, but is an Achilid. As it is an
unlocalized species, 1 refrain from describing the genus.
XXXV, — On a Freshwater Decapod Crustacean collected
by W, J. Burchell at Pard in 1829. By W. T. Calman,
l3.Sc., British Museum (Natural History).
Amonc^ a few dried Crustacea belonging to the Burchell
collection submitted to me for determination by Prof. Poulton
is a small shrimp-like animal, little more than half an inch
in length, labelled in Burchell‘’s handwriting From the well,
4.9.29.'’^ Prof. Poulton informs me that, at the date indicated
21*
296
Dr. W. T. Caiman on a
(Sept. 4, 1829), Burcliell was living in Par4*. The fragile
and shrivelled specimen had been laid aside as indeterminable
wlien my attention was, by chance, directed to Mr. E. J.
Miers^s description and figures of a form which he assigned
to a new genus of Pala3monida3 under the name Earyrhynchus
Wrzesmowskii j", and of which tv/o specimens were obtained
from a well in Cayenne by Prof. Wrzesniowsky, of Warsaw.
One of Miers’s types exists, in fragments, in the British
Museum collection, and a comparison with Burchelhs specimen
shows that the two are certainly congeneric and possibly even
specifically identical. Since, however, there are definite,
though not very important, differences between the two, I
propose to refer to Burchelhs specimen under the new specific
name of E, Burchelli.
The figures given by Miers are somewhat unsatisfactory
and his description is incomplete, especially in the absence of
any account of the mouth-parts. The genus has therefore
remained in obscurity, and, so far as I have been able to
discover, the only further reference to it is by Prof. Kingsley,
who, in his revision of the Palsemonidse j, quotes Miers’s
description and leaves the systematic position of the genus
doubtful. In view of this uncertainty, it seems worth while
to record the results of an examination of the two specimens.
Burchell’s specimen, in spite of its great age, was found, on
being carefully relaxed, to be in fairly good condition.
The following diagnosis includes the characters which
seem to be of generic importance.
Family Palsemonidae.
Genus Eukyehynchus, Miers.
Bostrum very short, flattened, without teeth above or below.
Carapace with an antennal spine, but no supraorbital, hepatic,
or branchiostegal spines. Outer flagellum of antennules com-
pletely divided. Mandible without a palp. Third maxilli-
peds slender. Second peraeopods much stronger than the
first.
Type species, E. Wrzesniowshii^ Miers.
The absence of the mandibular palp brings this genus
within a group which lies near the boundary-line between
the Palsemonidae and Pontoniidae and renders this line, as
Schenkel has remarked, somewhat indefinite. The fact that
* Cf. Poulton, Ann. & Mag. Nat. Hist. (7) xiii. pp. 45-56, pi. iii.
t Proc. Zool. Soc. 1877, p. 662, pi. Ixvii. tigs. 2-2 6.
f Proc. Acad. Nat. Sci. Philadelphia, 1879, p. 424.
297
Freshwater Crustacean from Pard.
the two branches of the outer flagellum of the antennules are
divided to the base may, however, be taken as indicating for
the genus a place within the Palsemonida3. The other genera
usually incliuled- in this family which have no palps on the
mandibles are Pahemonetes, Heller, Borra-
daile, and Ancijlocaris^ Schenkel. From these the present
genus is distinguished by the form of the rostrum and by the
absence of spines except the antennal on the antero-lateral
region of the carapace.
Euryrhynclius WrzesniowslcUy I\Iiers. (Fig. 1.)
Euryrhynchus Wrzesniowskiiy Miers, Proc. Zool. Soc. 1877, p. 662,
pi. Ixvii. tigs. 2-2 h.
Merus of second perasopods with two inconspicuous rounded
lobes at the distal end below ; carpus without a spine on the
inner side ; fingers hardly longer than the palm. Telsori
with the tip rather narrowly rounded, projecting nearly as
far as the long lateral spines.
Fig. 1.
Euryrhynchus Wrzesniowshiij Miers. Distal end of telson.
I can find no trace of the small spine between the eyes
and the rostrum described by Miers, and the telson, as
figured, is much too narrow.
Locality. Cayenne, in a w^ell.^^ Co-type in British
Museum.
Euryrhynchus Burchelli^ sp. n. (Figs. 2-8.)
Menis of second peiaeopods with two acute spiniforra teeth
at the distal end of its lower surface ; carpus with a sharp
tooth on the inner side near the distal end ; fingers distinctly
longer than the palm. Telson with the tip broadly rounded,
not projecting nearly as far as the long lateral spines.
Lccalitij. Paid, 4th September, 1829 ; Burcheli Coll. Type
in Hope Museum, Oxford.
* Mr. Borradaile has pointed out to me that this name was first used
by Stimpson in 1871 (Atm. Lyc. Nat. Hist. New York, x. p. 128), but
apparently only as a nomen nudum. Whether Borradaile’s later use of
the namehs illegal may be left to the decision of those interested in such
questions.
298
On a Freshwater Crustacean from Pard.
Fig. 2.
'Euryrliijnchus Burclielli.
2, — Anterior part of carapace, from the side.
Fig. 3. — Anterior part of cara]>ace, with eyes, antennules, and auteunre,
from above (set£e omitted).
Fig. 4. — Mandible.
Fig. 5. — Distal end of merus and carpus of second perajopod from outer
side.
Fig. 6. — Distal part of second permopod (left side) from below.
Fig. 7. — Dactylus of third permopod.
p’ig. 8, — Telson and uropod. Marginal set;e of uropod omitted.
On new Species of Ileterocera,
299
Comparative Measurements,
E. Wrzesniowskii.
E. Burchelli.
mm.
chela
palm
The fact tliat both the forms referred to this genus were
found in wells suggests that they belong to the subterranean
fauna. The eyes, however, although small, are well-pig-
mented and facetted. Certain blind forms referred to the
genus F alcemo7ietes have been described from subterranean
liabitats in Texas and in Cuba. They are still imperfectly
known and it does not seem certain that they are correctly
assigned to that genus. They are certainly specifically, and
probably generically, distinct from the forms here dealt
with.
XXXVI. — Descriptions of new Species of Ileterocera hehnqiag
to the Families Syntomidj^e, Hypsida?, Cyllopodhlae, Diop-
tidse, and Erateininse. By Herbert Druce, F.L.S. &c.
Family Syntomidge.
Agijrta varuna^ sp. n.
Male. — Head, antennae, and thorax black, the shaft of the
antennae white ; the collar and palpi bright crimson ; tegulae
black, edged with white ; abdomen metallic blue above,
white on the underside ; a bluish-white dorsal stripe extends
from the thorax to the anus. Primaries black, with a semi-
hyaline white band from the base as far as the end of the
cell; a white spot near the apex: secondaries pale yellow,
the apex and outer margin broadly black shot with brilliant
blue; the fringes of both wings black. Underside primaries
very similar to the upperside, but shot with brilliant blue :
secondaries with the costal margin banded with white and an
300 Mr. II. Druce on
oval-sliapecl wlilte spot close to the apex, the black margia
glossed with blue.
Expanse inch.
Hah. Peru, Pichbs Road, 3000 feet {Watlcins^Mus. Druce).
Allied to Agyrta flavitincta^ Ilampson, from Bolivia.
Family Hypsidae.
CalUmorplia solai^ sp. n.
Head, antennae, palpi, collar, tegulse, and thorax black;
abdomen above orange-yellow banded with black ; the under-
side, anus, and legs black. Primaries yellowish cream-
colour, the costal margin, apex, and outer margin black, the
inner edge of the black outer margin deeply dentated about
tlie middle ; a black spot above the anal angle and a black
band near the apex ; a spot at the end of the cell and the
base of the wing black : secondaries dark orange, broadly
bordered with black ; a black spot at the end of the cell ; the
secondaries are almost identical with those of CaUimorpha
TheJwalHj Druce. Underside almost identical with the
upperside, but the primaries dark orange-yellow, the same as
the secondaries.
Expanse 2| inches.
Hah. British East Africa {Mus. Druce).
Specimens are also in the National Collection.
Eucyane dilutana, sp. n.
Head, antennae, palpi, collar, tegulce, thorax, abdomen, and
legs all black ; the underside of the head and front of thorax
yellow. Primaries black, the veins slightly greyish ; a
yellow band crosses the wing beyond the cell from the costal
margin to the anal angle, where it becomes quite narrow;
the fringe black ; secondaries black ; a submarginal yellow
band extends from close to the apex to the anal angle, be-
coming rather reddish ; tlie outer margin black. The under-
side very similar to the upperside, but the bands more shaded
with red.
Expanse 2 inches.
Hah. Amazons [Mus. Brit.).
Allied to Eucyane kedar^ Druce.
Pericopis anadema^ sp. n.
Male. — Head, antennm, palpi, and legs black ; two white
spots on the collar ; tegula3 yellow, with a black line
301
new Species oj Hekrocera,
down the middle; thorax blackish brown, with two yellowish-
white spots at the base ; abdomen above black, striped with
yellow on each side, the anal segments black, the anal tuft
yellow; the underside of ihe abdomen pale yellow. Pri-
maries semihyalinc, shaded with brown; the apex, outer
margin, and inner margin brownish black ; two brownish-
black lines cross the wing from the costal margin, the first
from the end of the cell to the outer margin, the second
about the middle of the wing; a marginal row of minute
white dots extends from the apex to the anal angle : second-
aries orange-yellow in some specimens, the veins blackish;
the outer margin black, with a marginal row of very minute
white dots. Underside very similar to the upperside ; the
fringe blackish brown. — Female. Primaries pale brownish
yellow, crossed from the costal margin beyond the middle by
a wide broken yellow* band : secondaries orange-yellow, the
outer margins black, the white dots very indistinct.
Expanse, S $ 3 inches.
Hah. Colombia, Minca, 2000 feet [H. H. Smitli^j 10 S)
1 $ (l/w.9. Druce).
One specimen in the National Collection from Panama.
Pericopis Forhesi^ sp. n,
Male. — Head, antennas, and palpi black; collar black,
spotted with yellow ; tegulae black, yellow at the base ; thorax
and abdomen black, the abdomen banded with yellow on each
side; the underside yellow, with a central black line; the
anus reddish yellow; the legs black; secondaries brownish
hyaline, the veins, apex, outer and inner margins black ; a
wide black band crosses the wing beyond the cell from the
costal margin to the outer margin above the anal angle ; a
second black band crosses the wing about the middle : second-
aries yellowish hyaline, the veins, apex, and outer margin
broadly black ; a narrow black band at the end of the cell.
Underside very similar to the upperside, but with a small
red spot at the base of each wing.
Expanse 3 inches.
Hah. South Brazil [Mus. Druce).
A very worn and broken specimen is in the National
Collection.
Pericopis mosera, sp. n.
Male. — Head, antennm, palpi, collar, tegulae, thorax, and
legs black; collar spotted with yellow; abdomen black,
banded on the sides with bluish grey ; the underside pale
302
Mr. H. Druce on
yellow; tlie anal tuft dark yellow. Primaries greyish
liyaline, the costal margin from the base to end of the cell and
a streak on the inner margin from the base almost to the anal
angle reddish brown ; the apex, outer and inner margins
black excepting at the apex; a black band crosses the wing
at the end of the cell from the costal to the outer margin ; the
band is semihyaline black in the middle ; a short, slightly
curved black band crosses the wing about the middle of the
cell ; a marginal row of minute white dots extends from the
apex to the anal angle : secondaries hyaline, shaded with
black above the anal angle ; the costal margin, apex, and
outer margin broadly black ; a black band at the end of the
cell ; a marginal row of white spots extends from the apex to
the anal angle. Underside similar to the upperside; the
primaries with a reddish-brown band at the apex and a
reddish-brown spot about the middle of the outer margin ;
the costal margin of the secondaries reddish brown.
Expanse inches.
Hah. Peru, Pio Colorado, 2500 feet {Watidns & Tom'-
linsorij Mus. Brit.),
Family Cyllopodidse.
Devara hicolorata^ sp. n.
Male. — Head, antennse, collar, tegulse, thorax, legs,and
the abdomen black ; the underside of the abdomen greyish
white. Primaries black, with a large round white spot about
the middle of the wing, some white scales near the base, the
veins black : secondaries black ; the fringes of both wings
black. Underside : primaries, the white spot larger, ex-
tending to the base of the wing ; the apex dark grey, covered
by the black veins : secondaries dark grey, the veins all
black, the outer margin slightly white.
Expanse inch.
Hah. Peru, San Kemon, 3000 feet [Watidns^ Mus, Druce) ;
Polivia, Yungas la Paz [M^is. Brit.).
The specimen from Bolivia is slightly paler in colour on
the underside of the secondaries, but in all other respects it is
identical with the Peruvian specimens. It is allied to
Devara huhona, Druce, from Ecuador.
Devara xanihion^ sp. n.
Male. — Head, antennse, collar, tegulse, thorax, and abdo-
men black, the sides of the abdomen yellow ; legs black.
Primaries black ; a reddish-brown streak extends from the
^03
new Species of Ileterocera,
base almost to the middle of the wing ; a rather large yellowish-
white square-shaped spot close to the costal margin nearest
the apex ; the fringe black : secondaries black, the central
part of the wing reddish brown irrorated with black scales ;
the fringe black. Underside of primaries very similar to the
upperside ; the white spot larger and reaching the costal
margin ; several small yellowish-white streaks at the apex
and a white dot on the outer margin nearest the apex ;
secondaries pale yellowish white, the veins all black, broadly
bordered wdth blackish brown from the apex to the anal
angle, some of the spaces between the veins yellowish white.
Expanse inch.
Hab. S.E. Brazil, Bio Janeiro [Mas. Brit.).
Allied to B. hifenestrata^ Herr.-ScliafF., and D. pallida^
Druce.
Devara trebonia^ sp. n.
Head, antennas, collar, tegulae, thorax, abdomen, and legs
greyish black. Primaries black, the cell semihyaline white ;
a large round yellow spot beyond the cell nearest the apex ;
the fringe black : secondaries white, broadly bordered with
black from the apex to the anal angle. Underside very
similar to the upperside.
Expanse inch.
Hab. Peru, Rio Colorado, 2500 feet ; La Mercede, 3000 feet
[Wathins^ Mas, Bruce) \ San Remon, 3000 feet [Watkins,
Mas. Brit.),
Devara chiliorij sp. n.
Male. — Head, antennse, collar, tegulas, thorax, and upper-
side of the abdomen brownish black ; the underside of the
abdomen and legs greyish white. Primaiies brownish black;
three greyish-white streaks near the base and a white spot
close to the apex ; the veins yellowish brown ; the fringe
black : secondaries wdiite, broadly bordered with black from
the apex to the anal angle; the inner margin dusky white.
The underside very similar to the upperside, but blacker.
Expanse 1-j^- inch.
Hab, Peru, Rio Colorado, 2500 feet (fMus. Bruce).
A specimen of this species is in the National Collection
from the same locality.
Devara don, sp. n.
Male. — Head, antennae, collar, tegulae, thorax, abdomen,
and legs black ; the underside of the thorax and abdomen
304
Mr. H. Driice on
greyish white. Primaries black, the basal third white j a
white spot close to the apex : secondaries white, broadly
bordered with black I’rom the apex to the anal angle ; the
inner margin black ; fringes of both wings black. Under-
side : ])rimaiies very similar to the ujiperside ; two white
spots close to the apex ; the apex greyish : secondaries
greyish white, the veins black ; a black mark at the apex
and one close to the anal angle, the outer margin irroratecl
with black scales.
Expanse 1 inch.
Hah. Bolivia, Yungas la Paz (dfws. Brit.) j 2 ^ and 1 $
[Mus. Bruce).
Devara picroideSj sp. n.
Male. — Head, antennae, collar, tegulae, thorax, and abdo-
men above black ; the underside of the thorax and abdomen
greyish white; legs greyish. Primaries black, the basal
half of the wing white, the base irrorated with black scales :
secondaries white, edged with black, broadest at the apex ;
the fringe black. Underside: primaries very similar to the
upperside, excepting that the costal margin and apex are
greyish white : secondaries glossy white, the veins black.
Expanse inch.
Hah. Bolivia, Yungas la Paz [Mas. Brit.).
Devara cressida^ sp. n.
Male. — Head, antennae, and thorax black ; collar and
tegulm grey ; abdomen black, banded with grey, the under-
side white ; legs black. Primaries black; an elongated spot
at the base whitish hyaline, and a small white spot on the
costal margin near the apex : secondaries creamy white,
broadly bordered witli black from the apex to tlie inner
margin. Underside : primaries very similar to the upperside,
but with the apex greyish, crossed by the black veins and a
small white spot about the middle of the outer margin :
sccondaiies creamy white, the veins all black, the aj)ex and
outer margin clouded greyish black ; a small white spot near
the apex.
Expanse Inch.
Uab. Peru, liio Colorado [WatJeins^ Mas. Brace).
Devara prhtea^ sp. n.
Female. — Head, antenna?, thorax, and upperside of the
abdomen black, tlie underside white ; collar orange ; tegulre
305
new Species of Heterocera.
black, edged with white. Primaries black; a curved hyaline
white band extends from the base to the end of the cell ; the
inner margin from the base to about the middle greyish ; a
round white spot near the apex : secondaries white, broadly
bordered with black from the apex to the anal angle. Under-
side very similar to the upperside, but not so black ; the outer
margin of the secondaries lias an indistinct greyish-white line
on the black border which extends from the apex to the anal
angle ; the fringes of both wings black.
Expanse inch.
Hab. Peru, La Mecede, 2000-3000 feet {Watkins^ Mas,
Dr ace) .
JSelo cosyra, sp. n.
Male. — Head, antennae, collar, tegulae, thorax, abdomen,
and legs brownish black. Primaries brownish black, the
costal margin from the base almost to the apex orange-red;
a wide orange-red band crosses the centre of the wing from
the costal margin to the anal angle; the fringe dark brown :
secondaries brownish black. Underside: primaries very
similar to the upperside ; secondaries pale brown, the veins all
black.
Expanse inch.
Hub.- Peru, San Remon, 2000-3000 feet ( Wathins')^ 10 S
{Mus. Druce), 1 S [Mas, BriL).
Allied to Nelo coccienatay Walk.
Nelo creteSy sp. n.
Male. — Head, antennae, collar, tegulae, thorax, abdomen,
and legs black, the underside of the abdomen reddish. Pri-
maries black, with a white spot shaded with blue beyond the
cell nearest the outer margin : secondaries black; the fringes
of both wings black. Underside : primaries very similar to
the upperside, the white spot more distinct, the apex and half
of the outer margin red : secondaries red, the veins black.
Expanse IJ inch.
Hab. Peru, Rio Colorado, 2500 feet; La Mercede, 2000 feet
[Watkins j Mas. Druce)^ IS S : ^ S [Mus. Brit.).
Nelo dolopiay sp. n.
Male. — Head, antennae, collar, tegulse, thorax, abdomen,
and legs dark brown. Primaries dark brown, crossed about
the middle by a wide red band, which is straight on the inner
side and much curved on the outer side : secondaries dark
306
Mr. II. Druce on
brown. Underside very similar to the upperside, but much
paler brown, the veins black.
Expanse inch.
Ilah. Bolivia, La Paz {Garlepp, Mas, Druce)] Peru,
Oroya, 1000 feet {Mas. Brit.),
Nelo cunaxa^ sp. n.
Male, — Head, antennae, collar, tegulae, thorax, abdomen,
and legs brownish black. Primaries brownish black, with a
large orange-red spot extending from the costal margin
nearest the apex to the anal angle ; secondaries dark brown ;
the fringe of both wings dark brown. Underside : primaries
similar to the upperside, but paler in colour, the orange-red
spot edged with white on the costal margin : secondaries pale
brown, the veins black ; a white streak on the costal margin
just above the apex.
Expanse 1;^ inch.
Ilah. Peru, Bio Colorado, La Mercede, 2000-3000 feet,
5 S (J^Ius, Druce),
Nelo cypharUj sp. n.
Male. — Head, antennae, collar, thorax, abdomen, and legs
black. Primaries deep blue-black, with a large, almost round,
orange-red spot beyond the cell : secondaries deep blue-black.
Underside very similar to the upperside, excepting that the
costal margin, apex, and outer margin are pale brown :
secondaries brown, greyish round the outer margin, the veins
all black.
Expanse If inch.
Ilah, North Peru, Huancabamba, 6000-10,000 feet ;
Pozuzo, 5000-10,000 feet [Native Collector) ; Bio Colorado,
2500 feet ( Watkins)^^ 16 S [Mas, Druce),
A specimen of this species is in the National Collection
from Peru.
Nelo donuca^ sp. n.
Male, — Head, antennse, collar, tegulae, thorax, abdomen,
and legs black. Primaries and secondaries black, primaries
slightly glossed with dull blue near the apex. Underside of
both wings black.
Expanse IJ inch.
Hab, Peru, Pozuzo, 5000-10,000 feet (Native Collector^
Mas. Druce),
Allied to Nelo philodamea^ Druce, but very distinct.
new Species of lleterocera.
307
Sangala cydrara, sp. n.
Male. — Head, collar, teo'ulie, thorax, and abdomen black,
the point of the .tegulm red. Primaries dark brown, with a
large central red patch that does not reach either the costal
or inner margin : secondaries dark brown. Underside: pri-
maries pale brown, the veins black ; the red spot smaller than
above : secondaries pale brown, a red spot at the base, the
veins black.
Expanse IJ- inch.
Hab. Bolivia, Yungas la Paz [Mas, Brit.').
Sangala anasa^ sp. n.
Male. — Head, antennae, collar, tegalse, thorax, abdomen,
and legs black. Primaries bright glossy dark blue : second-
aries dark brownish black, in some lights slightly glossed
with blue. Underside ; both wings pale brown, the veins
black.
Expanse If inch.
Hah. Peru, La Mercede, 2000-3000 feet [WathinSj Mas.
Druce) ,
Allied to Sangala antiphates, Druce, but very distinct on
the underside.
Sangala marpesia^ sp. n.
Male. — Head, antennae, collar, tegulas, thorax, abdomen,
and legs black, the sides of the abdomen streaked with red
and yellow. Primaries dark brown, with a large central
reddish-orange spot crossing the wing from the costal margin
nearly to the inner margin ; the fringe dark brown : second-
aries dark brown. Underside : primaries pale brown, the
veins black, the costal margin from the base to the end of the
cell red ; a red spot at the end of the cell : secondaries pale
brown, the veins black ; a small red spot at the base and two
large white spots close to the apex.
Expanse 1 j inch.
Hob. Bolivia, La Paz [Qarlepp^ Mas. Druce),
Sangala cynara^ sp. n.
Male. — Head, antennse, collar, thorax, upperside of abdo-
men, and legs all black, underside of abdomen reddish brown.
Primaries dark brown, with a large red spot at the end of
the cell ; the wing is glossed with bright blue from the base
almost to the outer margin ; the red spot varies greatly in
308
Mr. H. Druce on
size in the eight specimens before me : secondaries dark
brown, in some specimens the wing is slightly glossed with
blue on the outer margin. Underside : primaries pale brown,
the veins black ; an orange-red, rather wide streak at the end
of the cell : secondaries pale brown, greyish brown at the
apex, the veins black.
Expanse IJ ineh.
11 ah, Peru, Upper Rio Toro {Mas. Dr ace).
In some specimens the red spot on the underside of the
primaries is entirely wanting. This species is allied to
Sangala antipJiateSy Druce.
Taraxineura quadripuncta, sp. n.
Male. — Head, antennae, tegulae, thorax, abdomen, and legs
black; collar orange-yellow ; anal tuft yellow. Primaries
black, a large square white spot at the end of cell ; fringe
black : secondaries black, a small white round spot at the end
of the cell. Underside similar to the upperside.
Expanse inch.
Hah, British Guiana, Roraima [WMtehj) ] Paramaribo
(U. W. Ellacoinhe, Mas. Druce)^ 2 (^ , 1 $; Potaro River
((7. B. Roberts j Mas. Brit,)^ 1 J.
Family Dioptidae.
Lauron halizoaj sp. n.
Female. — Head, antennae, and thorax black ; tegulae orange-
red ; abdomen brownish black, with a greyish-white line on
each side ; legs brownish black. Primaries reddish orange,
semihyaline near the base, the veins mostly black ; a band of
five white spots crosses the wing near the apex from the costal
to the outer margin ; the fourth and fifth spots are quite
small: secondaries semihyaline brownish black; the under-
side of both wings brownish black; the costal margin of the
prirnaiics reddish orange.
Expanse 2 inches.
Hah. Jamaica {Mus. Brit.).
Dioiitis quiriteSj sp. n.
Male. — Head, antennae, collar, tegulae, thorax, abdomen,
and legs black ; the front of the head and underside of the
thorax and abdomen white. Primaries hyaline, the veins
black, the apex and outer margin broadly black ; a semi-
hyaline white band partly crosses the wing near the apex,
new Species of FTeterocera. 301)
but does not reach the outer margin ; a round semihyaline
white spot at the anal angle : secondaries hyaline, the veins
black ; the costal margin, apex, and outer margin black ; a
submarginal pale yellow line edged with black on the inner
side extends from the aj)ex to the anal angle. The under-
side very similar to the upperside.
Expanse IJ inch.
Hah, Colombia, Bogota [Mus. Brit.).
From the Crowley Collection. Allied to Dioptis Trailii^
Butler, and Dioptis cheledonis^ Druce.
Dioptis albifasciata.^ sp. n.
Male. — Head, antennae, collar, tegulai, thorax, abdomen,
and legs black; underside o£ the abdomen white. Primaries
hyaline, the veins black; a black line crosses the wing
beyond the cell from the costal margin to the anal angle,
beyond which a wide white Ijand crosses from the costal to
the outer margin ; the apex and outer margin broadly black :
secondaries hyaline, the veins, costal and outer margins
black. The underside very similar to the upperside.
Expanse inch.
Ilab, Peru, La Mercede, 2000-3000 feet [Watkins.^ Mas.'
Druce).
Loclia hermeSj sp. n.
Female. — Head, antennae, collar, tegulae, thorax, abdomen,
and legs black. Primaries hyaline, the veins all black; the
costal margin, apex, outer and inner margins black; a black
band crosses the wing at the end of the cell from the costal
to the outer margin ; a small orange-red spot at the base of
the wing : secondaries hyaline, the veins, costal and outer
margins black. The underside the same as the upperside.
Expanse inch.
llah. Colombia [Mus. Brit.).
Allied to Loclia hyalina^ Walk.
Phanoptis taxila^ sp. n.
Female. — Head, antennae, collar, thorax, abdomen, and
legs black ; tegulae black, with a white spot at the base.
Primaries hyaline, the costal margin, apex, outer and inner
margins, and a wide band crossing the wing beyond the cell
all black glossed with brilliant blue : secondaries liyaline, the
costal margin, apex, and outer margin and veins black glossed
Ann. (F Mag. N. Hist. Scr. 7. Vol. xix. 22
310
On new Species of TIeterocera.
wltli brilliant blue. Underside similar to the upperside, but
not so much glossed with blue.
Expanse 1| inch.
llah. Colombia, Bogota {Mus Brit.).
From the Crowley Collection.
FJianoptis Ipdia^ sp. n.
Female. — Head, antennae, and tegulse black; collar bright
red; thorax and abdomen bright blue, abdomen with a white
central line, underside of the abdomen white. Primaries
hyaline, the veins, costal margin, apex, outer and inner
margins, and a band crossing the wing beyond the cell all
black glossed with brilliant blue: secondaries hyaline, the
veins and margins black glossed with brilliant blue ; the
fringe of the secondaries white. Underside very similar to
the upperside, but without the blue gloss.
Expanse 2 inches.
llab. Peru, La Mercede, 2000-3000 feet [Watkins, M us.
Druce) .
Allied to Bhanoptis cyanomelas, Felder.
Family Erateininae.
Eratina amazonia, sp. n.
Female. — Head, thorax, and abdomen black; collar white;
abdomen banded with white above, the underside white; legs
black, while on the underside. Primaries pale greyish brown
from the base to about the middle ; the apical half black,
crossed from the costal margin almost to the anal angle by a
yellowish-white band : secondaries greyish brown, the outer
margin broadly black; a submarginal row of yellowish-white
spots extends from near the apex to the anal angle; a red
spot at the anal angle ; the fringe alternately black and
white. Underside : primaries similar to the upperside, but
redder and with a submarginal white band from the costal
margin near the apex to the anal angle ; secondaries white,
streaked with yellow and red bands, the outer margin reddish
brown.
Expanse inch.
Uab. Amazons, Banos [Mas. Brit.).
Eratina Garlep>2ji^ sp. n.
Male. — Head, antenna}, collar, tegulcC, thorax, and upper-
side of the abdomen black, underside of the abdomen white.
Primaries black, with white streaks from the base, the inner
811
Oyi new Species of Plistericlic.
margin white ; a wide semihyaline white band crossing the
wing about the middle, but not reaching either margin ; a
second small band near the apex ; the fringe black ; the
semihjuxline bands are glossed with dark blue : secondaries
black, the central part of the wing bluish hyaline white ; the
fringe alternately white and black. Underside of both wings
very similar to the upperside, but the black parts reddish in
tint.
Expanse l-J inch.
Ilab, Bolivia, Yungas la Paz i^Gavlepp)^ 3 $ {flm,
Dnice).
A specimen in the National Collection from the same
localitj\
Erathia albonulataj sp. n.
Female. — Head, antennae, tegulse, thorax, and abdomen
above black ; collar and underside of the abdomen pale
yellow, abdomen banded with white on the upperside. Pri-
maries black, slightly greyish at the base; a large oval-
shaped white spot about the middle of the wing : secondaries
black. Underside : primaries, the base streaked with yellow
and white, the wdiite spot as above, but extending to the costal
margin ; a submarginal line of bright yellow spots from near
the apex almost to the anal angle: secondaries pale yellowish
w’hite, the veins yellow, and a submarginal yellow line; the
apex reddish.
Expanse inch.
Uab. Bolivia, Yungas la Paz {^Garlepp^ Mas. J)ruce) ; one
specimen {Mas. Brit.).
XXXVIT. — On new Species of Histeridie J Notices of
others. By Gr. Lewis, F.L.S.
This is the thirtieth paper of this series.
In the last few years certain Histeridae have been found iti
India which show in a marked degree a connexion between
the fauna of our eastern possessions and that of Japan.
Mr. E. P. Stebbing has found Niponius canalicollis Lew.,
on the mountain ot Siiinghun, in the northern part of Zhob
in Baluchistan, at an elevation of 7500 feet; it w^as found
feeding on a species of Polygraphus which infests the Hima-
layan pine, Finns Gerardiana. The locality is of interest,
as it is probably the western geographical limit of Niponius,
312
Mr. G. Lewis on
as Japan is of its eastern boundary. The best time for
searching for the speeies is June and early in July; they are
diurnal, and occur in the higher regions of large forests. I
found the species in Japan chiefly on deciduous trees, elm,
oak, and celtis, and they seemed to feed on various sub-
cortical and wood-boring Coleoptera. It is likely that tliere
are unrecognized species in collections, as one from Borneo
remained in the Pascoe collection for many years until
detected by Mr. Arrow.
Species of two other peculiar genera have been recently
discovered in the Nilgiri Hills by Mr. H. L. Andrewes which
have a like distribution from Japan in the east to India in
the west, viz. Try'peticus and Pachylomahis, and in botli
instances the species are very similar to each other. Histei'
punctulatuSj Wiedem., one of the most curious species in the
family, occurs in Bengal, and I have found it in China and
Japan ; the species should now be placed in Zahromorphus,
as it has longitudinal antennal fossas.
It is by the peculiar genera and species of a family that the
connecting-links in a fauna are principally shown, not by the
ordinary genera, such as TIister and Saprinus, which are
world-spread.
List of Species.
ITololepta sci&sonia, Mars, (maiira,
Leiv.).
Trypan£ens thoracicus, F.
torpedo, Lew.
Caiupylorhahdus Poop-ei^ Har.
Teinotarsus latipes, Lew.
Ilister oyassas.
Silantjevi, Shir.
Carciiiops arqnatus.
Paro mains, Erich son.
Isolomaliis, pen. nov.
elongatus.
Tsolomalus truncatus.
Microloinalus, gen. nov.
Discoscelis argentine.
Abneus orientalis.
Sapriinis caesopygus, Mars.
Euspilotus, gen. nov.
zonalis.
flavopictns, Letu.
■ decoratns, Er.
bisignatns, Er.
Gnatlioncns brevisternns.
Ilololepta maura.^ Lew., 1885, = //. scissoma, Mars., I860.
In 1885 1 knew' //. scissoma by description only. //. maura,
Lew., is a variety in which the first dorsal stria is interrupted
in the middle; in more than fifty examples I have since seen
the first dorsal stria is complete, as shown in Marseul’s figure
(pi. xi. fig. 10, Monograph, 18G0).
Trypanaais thoracicus, F., and T. torpedo., Lew.
The females of these species are very similar to each other,
313
new Species of Ilisterida3.
except that the latter is more robust and tlie lateral thoracic
stria is conspicuously stronger ; but the male of torpedo has a
rostrum which has a protuberance on either side, and in this
respect agrees with the males of T, carincerostris, noxiusj
nasicornis, and volvulus^ Mars. Two figures are given to
show the differences of the outlines of the rostrum.
Fig. 1. Fig. 2.
Trypanceus thoracicus, F. Trypanmis torpedo, Lew.
Campyloriiabdus, Schmidt, 1889.
Teinotarsus Poggei, Har., is undoubtedly congeneric with
CampyJorhahdus singularis, Sch. There is a very great
resemblance in the sculpture and form of the sterna and in
the peculiar dorsal strife of both species.
Teinotarsus latipeSy Lew., 1904.
I give a figure of this species, drawn from the type example
in the British Museum.
Fig. 3.
Ulster nyasscBj sp. n.
Breviter ovalis, convexus, iiiger, nitidus ; fronte biimpressa, stria
recta ; pronoto lateribus impiinctato, marginato et bistriato ;
elytris sfriis 1-3 et subliumerali integris, 4 basi evanescenti, 5
Mr. G. Lewis on
?>U
a])icali, suiurali utrinque abbreviata ; propygidio pygidioque
densissime puiictatis ; prosterno aiigust-ato baud striato ; meso-
steriio truncate, margiiiato ; tibiis anticis 3-dentatis.
L. mill.
This species is extremely similar to II. Marshalli, Lew.,
from Mashonalancl, but it is a little larger, rather less convex,
and somewhat more oval. The thorax is smooth laterally,
not conspicuously punctate, and the pygidiaare more densely
punctured. In Mar.skalli there is a short basal outer humeral
stria which is wanting in 7iyassce, but in both species the
niesosternum is truncate and has a fine marginal stria, and
the anterior tibia? are tridentate, with the apical tooth very
robust.
Ulster ceneus.^ angoniensis, africanus^ and sulcipijgus^ Lew.,
with afevj Payk., may be associated with the above, as they
all have a truncate niesosternum and a complete subhurneral
stria ; the thoracic stria? and the form of the anterior tibiae
are also similar.
Ilab. Nyassaland, Central Africa.
I give the following description for convenient reference: —
Ulster Silantjevij Shirjajev. E,ev. Russed’Ent. iii.
p. 16 (1903).
Corps ovalaire, assez convexe, luisant, antennes noires.
Stria frontale a peine sinuee. Pronotum a bords lateraux
renhes, avec une strie laterale unique, non parallble an bord
mais s’en rapprochant davantage en avant qu’en arribre;
dessus ponctue contre la strie laterale (interieurement) .
Elytres avec une strie subhumerale entibre et fortement
arquee, a sa base, vers I’interieur ; epaules avec une striole
oblique, bien distincte, reliant la partie basilaire de la strie
subhumerale au bord del’elytre; stries dorsales 1—3 entibres,
la 4® excedant legbrement le milieu des elytres, la 5® n’atteig-
nant jias leur tiers, la b® egale a la 4®. Epipleures ponctues
assez lortement. Propygidium ponctue un peu rnoins dense-
ment et un peu ])lus grossierement que le pygidiurn. Tibias
rougeatres, les anterieurs munis de cinq dents dont la derniere
est simple et assez aigue.
“ Long. mill.
“ Cette espbee est notamment voisine de II. marginatus^ Er.,
mais la strie laterale du pronotum n’atteint pas son bord
})Oslerieur, la base des elytres n^offre aucun vestige d’une 5
slide dorsale et les tibias antbricurs n’ont que 5 dents h Icur
315
nem Species of Ilistericla3.
bord externe. Elle se rapproclic egalement de II, carbon-
ariuSj HofFni., mais le proiiotum est ponctue cotitre la strie
laterale, interieureinent, etc.
“ Uii specimen ^ de cette interessante espbce dans le
district de Starobielsk, gouv. de Kharkov."”
Carcinops arquatus, sp. n.
Ovalis, convexiis, iiiger, nitidus ; fronte stria Integra ; pronoto
piinctato, punctis ssepe conflaentibus ; elytris striis subhumerali,
1-5 dorsalibus integris, 5 pone scutellum, conspicue incurvatis,
suturali parte tuberculiformis ; mesosterno antice leviter sinuato,
stria marginali late interrupta; tibiis anticis 3-dentatis.
L. 1| mill.
Oval, convex, blacky and shining ; the head, frontal stria
complete, punctulation rather sparse and interspaces with
microscopic points ; the thorax, marginal stria complete and
finely crenellate behind the head, punctured somewhat like
the head, but the points are more distinct and many are
confluent, along the basal edge there is a row of punctures ;
the elytra, strite, inner humeral fine and complete, 1-3 are
complete, 4 crenellate on the apical half and bent inwards at
tlie base, 5 apical half crenellate, rather widening out before
the middle and bending to the suture at the base. The fifth
striae appear to meet at the suture and togetiier form a com-
plete arch. The sutural stria is composed of three small
dorsal tubercles, with one point anterior to them, and in their
line are some apical points; the propygidium is transversely
punctured in the middle ; the pygidiuin is irregularly and
sparingly punctate ; the prosternum, lobe punctate and
marginate, keel smooth, stria joined anteriorly and sinuous at
the sides ; the mesosternum is feebly sinuous and the marginal
stria widely interrupted, closely behind the anterior edge
there is an arched stria which continues in a parallel line
along the sides to the base of the metasternum, and there is
no visible suture between the sterna ; the anterior tibiu3 are
feebly 3-dentate.
This species resembles G, striatisternus^ Lew., in that the
meso- and metasterna are connate and in the remarkable
lateral stria which is common to both.
Hab. Nilgiri Hills [H. L. Andrewes).
“ Found in a nest of a harvesting-ant, Pheidologiton
diversus, in a decaying Ficus on the Barwood Estate, altitude
3500 feet.”
kSIx examples ; Paussidse and other Coleoptera have been
found with the same ant.
31G
]\Ir. G. Lewis on
Paromalus, Eilclison, Jalirb. p. 107 (1834),
Type^ comylanatus^ Panz.
After founding the two new genera, as below, tlie species
now remaining in this genus are: — acistrigus^ biai-calus, clavis^
des'pectuSj evanescens^ Forestieri^ honoratus^ keicoluy khongiuSy
miliariSy musicus, oceanitisj RoherU\ aculptipygus^ ttibodwy
uwbiUcatiiSj Victorice^ vittula. Mars.; cequalis^ Say ; AUuaudiy
derasuSy lenticula, Modiglianiiy Schultheissi, Sch.; amelias ^
hrevipeSy commeatusy Fe<ey fugisanus, goliath, indicas, locellas,
loiigicornisy mendteaSy montivaejusy niponensis, oblisus, per-
siihilisy submetallicus, tardipeSy vermicidatuSy viaticuSy Lew. ;
ceglanicus, Motsch. ; complanatus, Panz. ; digitatuSy Woll. ;
exlgnuSyVhXw. ; LudoviciySauciuSy terroi-reginoi, \iW.\javanuSy
liedt.
The prosternum of P. complanatus and all the above are
marginate (see fig. 4, p. 318).
In my Catalogue of 1305 not more thati 100 species of
l\iromalus were given, but the number of species yet
uudescribed in collections is considerabhu
ISOLOMALUS, gen. nov.
Type, verminosusy Lew.
Body more or less convex ; outline varying considerably,
being sometimes elongate-oblong or sometimes shortly oval;
dorsum with a sutural stria, abbreviated anteriorly ; the pro-
sternum, anterior lobe somewhat short, keel narrowed before
the coxoe, without stria3, and semicircular at the basal edge
(see fig. 5) ; the mesosternum is wide posteriorly.
Tlie other species are : — addendusy irregularisy Sch.; bi-
ductus, bilineatusy causticusy concentricusycordgpggus, didgmus,
harioluSy hispaniolce, ir>Jimus, inunctuSy Luderiiy malusy ocuh-
qyygiiSy productusy rugigeniuSy trifoliumy Mars. ; bistriatusy
seminuhimy Erich. ; com plexus y mancuSy JissuSy jejuna Sy
notabilisy orbusy pupiltnSy rogalisy selectus, similisy sincerusy
sobrmasy verminosus. Lew.; and dijjicHisy Horn.
There are four species in which the sutural stria is obsolete
or wanting, but the general form of tlie body is the same,
-\VL. cunvexusy Mars. ; divaricatiiSy injiatuSy and sulcatuSy Lew.
The figure of the last species ( Biol. Ceiit.-Ainer., Col. vol. ii.
pt. 1, tab. vi. fig. 7) shows that it has a trace of a sutural
stria.
y. Ludertiy Mars., has made a settlement along the borders ’
of the Mediterranean, having doubtless been introduced with
Opuntia largely grown there.
neic Species of IIistci ida3.
317
Isolornalus elongafus, sp. n.
Elong-atus,parallelus, 8ubdeprcssiis,niger, niridiis, undique punctahis ;
froiite plana, punctulata ; mesosterno antice hand margiriato ;
pygidio postice margine elevato, antice vermiculato ; tibiis anticis
5-dentatis.
L. 2|-2| mill.
Elongate, parallel at the sideSj somewhat depressed, black
and shining ; the head clearly punctured, stria complete ;
the thorax, marginal stria obsolete behind the neck, lateral
punctures somewhat oblong and well marked, punctures on
the disc resemble those of the liead, but they are evanescent
behind the neck ; the elytra are more distinctly punctured
than the thorax, but on the same plan, largest latterly, finer
on the dorsum, with the sutural margin smooth, sutural stria
fine and reaching just beyond the middle; the propygidium
is flat and very distinctly punctate ; the pygidium S has a
raised rim with a crescent sulcus within it, the disc is
irregularly not finely vermiculate ; the prosternum, lobe
slightly punctured and with the keel microscopically strigose ;
the mesosternum is deeply margined at the sides only, with
two fine detached striee at the suture formed like inverted V’s ;
the pygidium ? is densely punctulate ; the anterior tibiai
5- dentate.
llah. Para and Santarem, Amazon River {II. IT. Smith).
Isolornalus truncatus^ sp. n.
Parum clongatus, parallelus, depressus, niger, riitidus; stria froiitali
Integra ; elytris apicibus anguste imj)iiuctatis ; propygidio disco
conspicue jmuctato ; prosterrio baud striato ; mesosterno stria
sLiturali interrupt a et biarcuata.
L. 21 mill.
Rather long, parallel, depressed, black and shining ; the
head clearly and rather closely punctured, frontal stria com-
plete ; the thorax punctured, finely strigose laterally, marginal
stria complete, disc finely punctulate; the elytra, sutural
stria abbreviated before the middle, sutural margin smooth,
surface punctate and feebly strigose longitudinally, apical
margin smooth ; the propygidium, disc conspicuously punc-
tate, outer margin smooth ; the pygidium, S apex obtusely
triangular and punctured, base deeply, irregularly, and
unevenly sculptured, $ unknown ; the prosternum, anterior
lobe rather coarsely punctured, keel more finely pointed and
without striai ; the mesosternum immarginate anteriorly,
laterally. there is a well-marked stria, the sutural or transverse
318
Mr. G. Lewis on
stria consists of two sliort, detached, bent striie; the meta-
sternum is laterally widely and microscopically strigose, with
a shallow rugose sculpture ; the anterior tibise are 4-5-
denticulate.
ddie form of the meso-metasternal transverse stria connects
this species with 7. elongatas.
Ilab. Serra de Comrnunaty, Pernambuco (^Gounelle).
MiCROLOMALUS, gen. nov.
This genus is established to receive species hitherto
included in Paromalus. The body is somewhat cylindrical
and elongate, but not much depressed as in the type of the
genus Paromalus [complanatuSj Panz.), and the form of the
}jrosternum is on a different plan. The prosternum is with-
out stripe and the keel is narrowed anteriorly and not much
flattened out, the metasternum throughout its length is
relatively more narrow, see tig. 6.
The species are jiavicornis^ 'parallelepipeduSy Herbst ;
omineus^ Lew.; j^/aw,E-eit.; soul pti pectus^ Mars.; and
simplicisternus, Sch. M. sculpti pectus^ from Java, I only
know by description ; the other species are from the north-
temperate zone of the Old World.
Fig’. 4. Fig. 5. Fig. 6.
AAA.
Fig. 4. — Faromalus complcmatus, Panz.
Fig. 5. — Isolomalus verniinosufi, Lew.
Fig. 6. — Microlomalus Jiavicornis, Herbst.
Discoscelis argentince, sp. n.
Late ovata, convexa, nigra, nitida ; fronto cum clypeo marginata ;
prouoto stria marginal! antice recta, disco oblique striato ; elytris
striis subhumerali et dorsalibus 1-2 integris ; pygidio apice
longitudinal! sulcato ; tibiis anticis emargiuatis.
L. 3| mill.
Broadly oval, convex above, black and shining ; the head
slightly impressed anteriorly, faintly and irregularly strigose,
lateral stria elevated and continued round the clypeus, which
has a widely sinuous edge; the thorax, anterior angles
obtuse and slightly explanate, marginal stria fine and con-
tinued behind the head, where it is markedly straight, at the
319
new Species (?/Histerida3.
base, in a line with the second dorsal stria, there is a fine
oblique stria bent inwards at the basal edge and in length
about one third of that of the thorax ; the elytra, the inner
humeral stria is complete and carinate in tlie basal half,
1 dorsal fine and complete and incurved at the apex, 2 com-
plete and similarly fine and turns inwards at the base, 3 is
very short basal and hamate, between the third stria and the
suture along the base is a very fine widely arched stria, which
apparently represents the fourth and sutural striae, there is no
fifth stria, but there is a very fine stria round the apical
margin which continues partly along the suture ; the surfaces
of the pygidia are scantily and microscopically punctulate,
the propygidium is margined anteriorly with a fine stria,
and the pygidium at its apex has a series of longitudinal
grooves, about 24 in all, these constitute a well-marked
character and may be sexual ; the prosternum, the stride are
didymous and widen out behind the coxa3 and in front the
two parts of each stria join ; the mesosternum has a marginal
stria along its edge and beliind it there is a transverse stria
which is sinuous in the middle ; the tibiae are disciform, the
anterior pair are minutely serrate on the outer edge and
markedly emarginate on the inner edge at the tarsal end.
D. arechavaletce^ Mars., is similar but larger, and has a third
dorsal stria and the 4-6 very short and apical. The oblique
thoracic strim resemble those of Renimus meticulosus^ Lew.,
and if more species of Discoscelis and Renimus are at any
time detected the genera may require revision.
llah. Tucuman, Argentina.
AhrcEus orientalis^ sp. n.
Suborbicularis, convexus, piceus, opaciis, ruguloso-punctatus, supra
setulis erectis fiavescentibus ; prosterno basi coiispicue pimctato;
mesosterno stria marginali late interrupta ; mesosteruo meta-
sternoque subocellato-punctatis ; tibiis obscure rufo-brunneis.
L. 1| mill.
Circular in outline, convex, rugosely punctured and setu-
lose above ; the head slightly impressed anteriorly ; the pro-
sternum straight at the base, narrowly marginate anteriorly,
with an extremely fine stria behind the marginal stria, across
the middle there is a feeble ridge and behind the ridge the
punctures are very distinct and arranged in two rows, anterior
to the ridge the punctures are irregular and inconspicuous,
basal margin narrowly smooth not striate ; the mesosternum,
marginal stria widely interrupted and the punctures, which
are similar to those of the metasternum, are in two irregular
820
Mr. G. Lewis on
rows; the nietasterniim, suture straight and rather coarsely
crenellate, the crenellations numbering about twelve, the
punctures are deep, circular, very evenly placed, somewhat
ocellate and closely but not densely set ; the legs are obscurely
reddish brown.
This is the fifth species known from India ; it is of ster-
coraceous habits.
Ilah. Nilgiri Hills (//. L, Andrewes).
Soprinus ccesopygus^ Mars.
I have received examples of this species from Tucuman,
Argentina; there is therefore no longer any doubt as to its
being a native of S. America. S. pygidialis^ Lew., is a
similar species, and in both the curious form of the pygidium
is a masculine character only.
EuSPILOTUS, gen. nov.
Body oval, shining, metallic ; head without a definite
frontal stria ; thorax transverse; elytra maculate, sutural stria
with an arcuate basal extension ; prosternum, keel narrow,
bistriate, with a strong and straight carina on either side, as
shown in fig. 7 ; mesosternum widely sinuous and the
margin interrupted ; anterior tibise multidenticulate.
The species to be included are Saprinus hisignatus^ de-
coratus^ lepidus, Er.; crenatipes, Sober; Mars. ;
dichrousj jiavopictusj Icesits, and zonalis (type). Lew.
The elytral markings of Saprinus interru2)ius, Pajk., a
Central Asian species, are remarkably similar to decoratus, and
1 give a figure of the prosternum to show that the species
are structurally very different.
Euspilotiis zonalis, Lew. Saprinus viterniptus, ILyk.
Euspilotus zonalis^ sp. n.
Ovutus, jeneus, nitidus ; fronte dense punctata; pronoto ciliato,
lateribus late rugoso-punctatis ; clytris macula flava lata, striis
1-2 brevissiniis, suturali Integra basi arcuata, cooteris iiullis ;
mesosterno stria marginali interrupta ; tibiis anticis mulL-
denticulatis.
L. 3|-I mill.
321
new Species of Ilistci'icla3.
Oval, brassy and sinning ; the head densely punctate,
stria obsolete ; the thorax broadly rngose-piinctate hiterally,
and narrowly at the base, smooth behind the head anfl on the
disc, lateral edge with flavons hair; the elytra, humeral
angle smooth, striae 1-2 basal and very short, interstices
strigose, sutural stria complete with an arcuate basal extension,
the other striae are wanting, the flavous band has a lob^-
shaped outline on the sutural disc and extends to the outer
margin, anteriorly it is obtusely pointed on the inner side of
the second stria (see fig. 9) ; the pygidium is somewhat closely
but not densely punctate ; the prosternum bistriate ; the
niesosternum evenly not closely punctured, with the marginal
stria interrupted in the middle ; the anterior tibiae have
12-13 small teeth.
This species differs from E. flavop ictus, Lew., in the thorac'c
band of punctures not continuing behind the head ; in the
form of the flavous band it is most similar to hisignatus, Er.
Hab, Chubut, Patagonia. Many examples.
As the elytral markings are fairly constant in these species
1 give outlines of the patterns oi E.zonaUs (tig. jlavopictus
(fig. 10), Lew., decoratus(f\g. 11) and hisignatus (fig. 12), Er.,
as an aid to their identification.
Fig. 9. Fig. 10, Fig. 11. Fig. 12.
GnafJionciis hrevisternus, sp. n.
Ovalis, convexiusculus, nitidns ; fronte parum convexa, parco
punctulata ; elytris striis 1-4 dorsalibus dimidiatis, 4 })asi
crenata ad suturam continuata ; prosterno striis brevibiis antice
obtuse coeimtibus.
L. 2| mill.
Oval, rather convex, black and shining; the head, forehead
slightly convex, almost smooth, punctures being fine and
sparse; the thorax clearly punctured laterally, punctures
niuch finer on the disc, stria interrupted anteriorly, scutellar
fovea well-marked; the elytra, striae 1-4 dimidiate, iriner
stri^ gradually shorter, 4 continues along the base to the
suture as a crenellate stria, sutural wanting; the pygidia
finely and evenly not densely punctulate ; the prosteinum,
322
]\Ir. A. S. Hirst on a
striae are very distinctly sliorter and wider npart than those
of G . na7inetensis, Mars., and rotandatus^ Kiigel, and meet
more obtusely in front.
Except for the characters given above this species is similar
to 7'otundatus.
I Jab. Yunnan.
XXXVIII. — On a neio Species r>/ Karschia Tibet.
I3y A. S. Hirst.
Karschia tibetana^ sp. n.
S . — Colour. Cephalic and thoracic segments pale yellow,
the cephalic plate being tinged with brown in the anterior
and lateral parts ; ocular tubercle dark. Abdomen greyish
white ; the tergal plates black ; ventral surface of this part of
the body marked laterally with a longitudinal series of dark
spots. Dorsal surface of mandible ornamented with three
dark longitudinal stripes ; tibia, metatarsus, and tarsus of
])alp, together with tl.e corresponding segments of the fourth
leg, infuscate dorsally, the remaining segments of these
appendages and the other legs being light yellow in colour or
but little darkened.
Mandible. Edge of upper finger of mandible with thirteen
teeth; the first, tliird, fourth, sixth, seventh, ninth, tenth, and
eleventh from the anterior end being of rather large size, the
others small. Lower finger bearing five teeth, of which the
posterior three are minute.
The two main bristles of the inner side of the mandible are
long, slender, and provided throughout the greater part of
their length with a ventral row of minute hairs, which are
absent from the basal part (fig. 2).
Flagellum long and spiral, retained in position by a hook
or thorn placed above it and by a long whip-like seta below ;
a small spine present on the lower surface towards the
terminal portion, which is long and very slender ; otherwise
the flagellum is smooth (fig. 2).
Palp. Tarsus and metatarsus of palp furnished below with
long and slender thorns, which are six in number in the case
of the tarsus. Metatarsus on the inner side with many
minute tufted hairs.
Abdomen. Third abdominal segment with many (over
twenty on each side) setae arranged in several irregular rows
towards the hinder edge of the ventral surface. Posterior
323
new Species of from Tibet,
margin of fourth abdominal segment with a single row of
nineteen long and slender setae.
Ocular tubercle beset anteriorly with numerous bristles,
several of which are of fairly great length, the chief of these
being a pair of diverging and upwardly directed bristles,
situated near the summit of the tubercle.
Fig. 1.
Fig. 1 . — Left mandible of $ from the outside.
Fig. 2. — Eight mandible of (S from the inside.
Measurements in mm. Length of mandible (finger incl.)
5*25, of palp 24-5, of first leg 18*5, of second leg 15, of third
leg 20, of fourth leg 30, of cephalic plate 2*5 ; total length
(mandible incl.) about 19; breadth of mandible (at base) 2,
of cephalic plate 4.
? . — Mandible. Teeth of upper finger of mandible fifteen
in number, the fourth and eighth from the anterior end beino-
of large size; first, third, seventh, and tenth fairly large”;
second, sixth, and ninth small, and the fifth granular; the
324
Mr. G. A. Boulanger on
row of teetli divides after the tenth into two branches, an
outer of two and an inner of tliree denticles. Lower finger
of mandible provided with two teeth of large size, the first of
which is preceded b)^ three or four small denticles, the two
large teeth being separated from one another by three teeth,
of which the median one is minute ; posterior enlarged tooth
followed by three small teeth (fig. 1).
Abdomen. Ventral surface of fourth abdominal segment
furnished posteriorly with a fringe of nineteen long set^e.
Palp. Tarsus and metatarsus of palp provided with a few
long hairs ; thorns absent.
Size. The female is of much larger size than the male,
which has appendages of greater length.
Measurements in mm. Length of mandible (finger inch) 7,
of palp 21’5, of first leg 16’7e5, of second leg 13‘75, of third
leg 18, of fourth leg 24*5, of cephalic plate 3*5 ; total
length 24 ; breadth of mandible (at base) 3, of cephalic
plate 6.
Ilah. Fourteen males and five females of this species were
collected by Captain H. J. Walton, I.M.S., at Gyantse,
Tibet, in the year 1904,
Remarks. Allied to K. nasuta^ from which it apparently
differs in the disposition and size of the teeth of the man-
dibular fingers, in the slenderness of the terminal portion of
the flagellum, and in the absence of the tooth which is
situated antero-dorsally towards the inner side of the upper
finger of the mandible in K. nasuta. The female differs
from that of persica., which is the only other species of this
genus in which this sex is known, in that the ventral surface
of the fourth abdominal segment is provided with nineteen
long setre on the hinder edge. It is necessary to state that I
have not been able to examine specimens of the two species
mentioned above.
XXXIX. — Descriptions of Three new Snakes discovered hi/
Mr. G. L. Bates in South Cameroon. By G. A.
Boulenger, F.R.S.
Jbisijpeltis macrops.
Eye larger than in I). sc(d)ra. Rostral nearly twice as
broad as deep, just visible from above ; internasals as long as
ora little shorter than the ])r0efrontals ; frontal a little longer
325
new Snakes from South Cameroon.
than broad, longer than its distance from the end of the
snout, as long as or a little shorter than the parietals ; one
prse- and two postoculars ; two superposed temporals, followed
by keeled scales-; seven upper labials, third and fourth
entering the eye ; a pair of large chin-shields, followed by a
smaller pair or by the first ventral shield. Scales in 20 to
23 rows, the smaller oblique lateral scales with serrated keel
reduced to two or three series. Ventrals 237-239 ; anal
entire ; subcaudals 79. Olive above, with very indistinct
yellowish cross-bars on the baek ; upper surface of head with
black vermiculations ; belly plumbeous grey or olive-grey,
more or less speckled with blackish.
Total length 900 mm.
Two specimens, ^ (Sc. 20; V. 239; C. 79) and ? (Sc. 23 ;
V. 237 ; C. ?), from Efulen.
The larger size of the eye, combined with the low number
of scales and the coloration, induces me to regard this snake
as specifically distinct from any of the numerous forms of
Dasy'peltis hitherto described. In the larger ( $ ) specimen
(900 mm.) the head measures 25 mm. and the eye 4 ; in the
smaller (d') specimen (765 mm.) the head measures 18 mm.
and the eye 4. In adult specimens of D. scabra the eye
measures only 3 mm.
Dipsadohoa isolepis.
Rostral a little broader than deep, just visible from above ;
internasals a little broader than long, half as long as the
prsefrontals ; frontal a little longer than broad, as long as its
distance from the end of the snout, shorter than the parietals ;
loreal deeper than long ; one praeocular, forming a suture
with the frontal ; two postoculars ; temporals 1 + 2 ; eight
upper labials, third, fourth, and fifth entering the eye ; six
lower labials in contact with the anterior chin-shields, which
are longer than the posterior. Scales in 19 rows, not oblique,
vertebrals scarcely enlarged. Ventrals 199 ; anal entire ;
subcaudals 56. Blackish grey above and under the tail ;
upper lip and lower surface of head and body yellowish
white.
Total length 435 mm. ; tail 70.
A single female specimen from Efulen.
This species connects the genera Dipsadohoa and Leptodira.
Aparallactus Baiesii,
Diameter of eye greater than its distance from the oral
margin. Rostral a little broader than deep, the portion
Ann. &; Mag. N. Hist. Ser. 7. Vol. xix. 23
326
Mr. E. E. Austen on the Synonymy and
visible from above nearly half as long as its distance from the
frontal; internasals much shorter than the prsefrontals ;
frontal once and a half as long as broad, longer than its
distance from the end of the snout, shorter than the parietals ;
nasal divided, in contact with the piseocular ; two post-
oculars, lower very small, upper in contact with the fifth and
sixth upper labials ; a single temporal; seven upper labials,
third and fourth entering the eye, sixth largest and forming
a long suture with the parietal ; first lower labial in contact
with its fellow behind the syrnphysial ; two pairs of chin-
shields, posterior longer. Scales in 15 rows. Ventrals 145 ;
anal entire ; subcaudals 48. Snout and upper parts of body
blackish, upper lip and back of head yellowish, lower parts
yellowish white.
Total length 235 mm.; tail 40.
A single female specimen from the forest 5 miles inland of
Kribi,
XL. — Synonymy and Generic Position of certain Species
of Muscidse (sens, lat.) in the Collection of the British
Museum^ described by the late Francis Walker. By
Ernest E. Austen.
Although nothing in the shape of general interest can be
claimed for the following notes, it is hoped that they may
at any rate prove useful to compilers of catalogues who
wish to avoid a long list of unidentified Walkerian species,
chiefly at the end of the genus Tachina. Even with the
types before him the present writer has often found it a
matter of great difficulty to assign Walker’s species of
Muscidse to their proper genera, owing to tlie fact that the
species are almost invariably based upon single specimens,
usually in very poor condition. It would have taken far too
long to correct the manifold inaccuracies and imperfections
of the original descriptions, and in the majority of cases no
attempt has been made to do so. As proving that Walker
described the specimen, and not the species, the characters of
which he was generally incapable of grasping, it may be
mentioned that he is responsible for no fewer than eleven
synonyms of the well-known Eiitachina rustica, Mg., the
description in every case being based upon a single
specimen.
The present paper contains but a first instalment of notes
upon Walker’s species of Muscidae, since it is hoped to deal
with other species in a similar manner at a later date.
327
Generic Position of certain Musciilaj.
Phasia argentifrons (List Dipt. Ins. in Coll. Brit. Mus.
part iv. (1849) p. 691. — Interior of South Africa). — This
species may be assigned provisionally to the genus Hijalomyia,
Kob.-Desv., though possibly it should be transferred to yet
another genus : the posterior transverse vein is nearer to the
bend of the fourth vein than to the small transverse vein.
Ocgptera picta {ibid. p. 695. — Sierra Leone). — Apparently
belongs to ParalophosiUj Br. & v. Berg.
Trichopoda lateralis {ibid. p. 697. — Brazil). — Belongs to
Homogenia, v. d. Wulp.
Trixa apicalis {ibid. p. 699. — Locality unknown) =
Microphthalma disju7icta, AVied.
Tachina melanax {ibid. p. 700. — Venezuela) is a Pejeaniay
Rob.-Desv.
Tachina alterna and T. aurifera {ibid. pp. 701-702. —
Venezuela) belong to Saundersia, Schin.
Tachina diversa {ibid. p. 703. — Venezuela) = Saundersia
ornata, Macq. {Micropalpus ornatus, Macq. Dipt. Exot. ii. 3,
p. 47).
Tachina ceciuabilis and T. constans* {ibid. pp. 704-705. —
Venezuela), as also T. transiens {ibid. p. 706. — Quito,
Ecuador), T. alligans {ibid. p. 713. — Venezuela), T. leuco-
melana’\ {ibid. p. 714. — Locality unknown), and T. con-
traria J {ibid. p. 716. — Mexico) belong to Saundersia, Schin.
Tachina notata, T. varia, T. lativitta, and T. vittata
(Insecta Saundersiana. — Diptera, pp. 267, 268, 269, 273. —
Colombia), and T. iransversa [ibid. p. 274. — Brazil) belong
to Saundersia.
Tachina signata (List Dipt. Ins. in Coll. Brit. Mus. part iv.
(1849) p. 709. — Locality unknown §) is a Hystricia, Macq.
Tachina zelica {ibid. p. 711. — New Zealand). — As stated
by Hutton (Trans. N. Zealand Institute, vol. xxxiii. (1901)
p. 51), this is a synonym of Hystricia {Musca) lupina,
* Saundersia nigropilosa, v. d. Wulp (Biol. Centr, -Americana, Diptera,
ii p. 23. — Mexico ; Costa Rica), is a synonym of this species : it may
also be noted that S. bipartita, v. d. Wulp {ibid. p. 25. — Mexico) =
S. bicolor, Will.
t Saundersia unicolor, v. d. Wulp {pp, cit. p. 23. — Mexico), is a
synonym of this species. The palpi” of Walker’s description are
really pollinia of a flower !
J Saundersia ruf tibia, v. d. Wulp {op. cit. p. 24. — Mexico), is a
synonym of this species.
§ The species is from New Zealand, and Hystricia pachyprocta,
Nowicki (Beitr. zur Kenntn, Diptf. Neu-Seelands (1875), p. 25), is a
synonym.
23*
328
Mr. E. E. Austen on the Synonymij and
Svederus (K. Vet.-Akad. Nya Handl. viii. (1787) p. 289) :
according to Svederus, the type of the species is in Mus.
Dom. Banks/-’ but it is not now to be found in the Banksian
Collection.
Tachina patula (JMd. p. 712. — Locality unknown) is a
Hystricia. The face is hairy.
Tachina anthemon [ibid. p. 733. — Brazil), T. amisias {ibid.
p. 734. — Locality unknown), T. epileuca [ibid. p. 716. —
Jamaica), and T. caliginosa (Ins. Saund. — Dipt. p. 268. —
Brazil) belong to the genus Hystricia. In H. epileuca the
face is hairy : H. amisias is possibly a synonym of H. an-
themon, but in the type of the former species, which is the
only specimen in the Museum, the macrochsetse near the
ront margin of the second abdominal segment are differently
disposed.
Echinomyia furiosa (Trans. Ent. Soc. Lond. iv. (1858)
p. 194. — Rio Grande, Brazil) is a Hystricia : the face is
hairy.
Tachina busalis^ (List Dipt. Ins. &c. pait iv. p. 713. —
Jamaica) is a Jurinia, Rob.-Desv.
Tachina decisa [ibid. p. 715. — Hudson’s Bay, Nova Scotia)
is a Jurinia', J. punctata, v. d. Wulp (Biol. Centr.-
Americana, Diptera, ii. p. 468. — Mexico) is a synonym of
this species.
Tachina iterans [ibid. p. 727. — Nova Scotia) = Peleteria
tessellata, Eahr. The statement that there are no bristles
on the side of the face is precisely the opposite of the fact ;
the same remark applies to the same statement in the case
of Tachina punctif era [ibid. p. 728. — Massachusetts), which,
as recorded by Coquillett (Revision of the Tachinidse of
America North of Mexico. U.S. Dept, of Agriculture.
Division of Entomology. Technical Series, no. 7 (1897),
p. 141), is likewise a synonym of P. tessellata, Eabr.
Tachina sacontala [ibid. p. 728. — Nepaul) apparently =
Tachina fera, Linn.
Tachina orbilius [ibid. p. 736. — England) =
flavipes, Mg., ? .
Tachina. mesula [ibid. p. 737. — England) = Demoticus
plebejus, Fin.
* Jurinia adusta, v. d. Wulp [op. cit. p, 28. — Mexico), is very closely
allied to this species; but in the former the antennae, especially the third
joints, are more reddish beneath and not wholly dark, and the dorsum
of the thorax is more yellowish pollinose in front.
329
Generic Position of certain MusciJse.
Tachina crisia [ibid, p. 738. — England) = Ernestia (Eri-
gone) radicum, F.
Tachina tyche [ibid. p. 738. — England) and T. amphiro
[ibid. p. 749. — England) = Blepharidea vulgaris, Fin.
Tachina megaleas [ibid. p. 739. — England), T.pitho [ibid.
p. 740. — England), T, admete {ibid. p. 743. — England),
T. pamesos [ibid. p. 744. — England), T. medoacus [ibid.
p. 746. — England), T. telestho, T. cerceis [ibid. p. 747. —
England), T. philonis and T. nymphidius {ibid. p. 751. —
England) rustica, Mg.
Tachina separata (Insecta Britannica. — Diptera, ii. p. 67
(1853). — England) devia, Fin.
Tachina lucifera (Insecta Saundersiana. — Diptera, pt. iv.
(1852) p. 282. — Locality ?) perhaps belongs to the Section
Blepharipoda, Br. & v. Berg., but the writer is at present
unable to determine its proper genus with certainty.
Tachina inornata and T. nervosa (Trans. Linn. Soc. Lond.
xvii. (1836) p. 349. — Cape Gregory and Port Famine,
Strait of Magellan) belong to the Section Pscudodexia,
Br. & von Berg., near the genus Macquartia, Rob.-Desv.
The eyes are bare, or but faintly pubescent, and the face
and arista are also bare. The face has a distinct keel and the
jowls {Backen) descend posteriorly. The types of both species
are males. The two species are very similar in appearance
and extremely closely allied ; in T. inornata, however, the
palpi and tibiae are orange and the last joint of the tarsi is
ochraceous, while on the underside of tlie abdomen there is
a long and very prominent tuft of stiff black hair on either
side of the hypopygium. In T. nervosa the palpi are black,
the tibiae blackish at the extremities, and the anal hair-tufts
are wanting.
Tachina piceiventris [ibid. p. 350. — S. America ; precise
locality uncertain) is a Masicera.
Tachina trifasciata [ibid. p. 350. — S. America ; precise
locality uncertain) belongs to the “ Section’^ Masicera, Br. &
von Berg. A new genus will probably have to be foundea
for this species, differing from Ceromasia, Rond., in the
narrowness of the body, the depth of the jowls being only
one-sixth of that of the eyes, the long and slender arista,
and the third vein being setigerous nearly as far as the
small transverse vein.
Tachina albifrons [ibid. p. 351. — S. America; precise
locality uncertain) is a Sisyropa, Br. & von Berg.
330
Mr. E. E. Austen on the Synonymy and
Tachina chrysocephala [ibid. p. 351. — S. America; precise
locality uncertain) is a Vhoroceray Rob.-Desv.
Tachina basalis [ibid. p. 351. — Port Famine, Strait of
Magellan) belongs to the Section Phorocera, Br. & v. Berg.,
and ap])arently to a new genus near ClKstoycedia, Br. & v.
Berg. Eyes bare ; depth of jowls, which descend notice-
ably behind, equal to half that of the eyes ; facial ridges
ciliated to above the middle ; face with a row of bristles
(a continuation of the frontal series) which descends nearly
to level of lower margin of eye; abdominal macrochsetse
discal and marginal ; bend of fourth vein somewhat rounded.
Tachina rnaura [ibid. p. 352. — Port Famine, Strait of
Magellan) is a synonym of the foregoing species.
Tachina atrivcntris (Ins. Saundersiana. — Diptera, pt. iv.
(1852) p. 290. — India) is assigned by Walker himself at the
commencement of his description to the genus Nemorcea.
It is not, however, congeneric with N. pellucida, Mg., and
apparently a new genus, possibly referable to the Section
P ar am acr onychia, Br. & von Berg., will have to be founded
for its reception. — Eyes hairy ; arista bare ; antennae long ;
jowls not descending; abdomen thickly clothed with erect
hair ; first, second, and third segments with marginal but no
discal macrochsetae ; costal spine wanting ; no appendix or
Faltenzinke to bend of fourth vein ; claws in male very
long.
Tachina alia [ibid. p. 293. — India). — The type of this
species proved to be an unrecognizable fragment, and has
consequently been destroyed. The name should be deleted
from van der Wulp^s ‘ Catalogue of the Described Diptera
from South Asia.^
Furigaster languida (Trans. Ent. Soc. Lond. iv. (1858)
p. 198. — India) is a Blepharipoda.
Tachina tricincta (Ins. Saundersiana. — Diptera, pt. iv.
(1852) p. 301. — India) belongs to the genus Frontina —
sens. lat. Br. & von Berg.
Masicera incivica (Trans. Ent. Soc. Lond. n. ser. v. (1861)
p. 305. — Asia. — ? India) is an Aporomyia, Rond. The
facial ridges are ciliated to the level of the descending
orbital setae — i. e. to above the middle. The length is in-
correctly given by Walker as 4, instead of 2 lines.
Tachina adusta (Ins. Saundersiana. — Diptera, pt. iv.
(1852) p. 292. — India) is a Frontina, Mg.
Echinornyia stolida (Trans. Ent. Soc. Lond. vol. iv. (1858)
Generic Position of certain Muscidse. 331
p. 195. — New South Wales) is the male of Tachina ohtasaj
Walk. (Ins. Saund. — Diptera, pt. iv. 1852, p. 274. — New
South Wales). .Under the latter name Walker professed
to describe both sexes, but the female alone is now to be
found in the Museum collection. It is quite possible that the
type of E. stolida is really the specimen previously described
by Walker as the male of Tachina obtiisa. This species
may be assigned provisionally and with doubt to the 'genus
Microtropesa, Macq. There is no noticeable keel on the
face, and in the female the third joint of the antennae is
somewhat square and distinctly shorter than the second (the
antennae are wanting in the male), so that the species might
be regarded as belonging to the genus Fabricia. The head,
however, is rather wide, and in an obviously closely allied
blit undescribed species from Tasmania in the Museum
collection the third antennal joint is as long as or slightly
longer than the second. In both species there is a row of
from six to eight small admedian macrochaetae on the margin
of the second abdominal segment, which are absent in
Microtropesa sinuata^ Don., the typical species of the genus.
In general appearance, however, T. obtusa and the species
from Tasmania resemble Microtropesa more than Fabricia.
Tachina despicienda (Trans. Ent. Soc. Lond. n. ser. v.
(1861) p. 306. — New South Wales) apparently belongs to
the Section'’^ Pyrrhosia, Br. & v. Berg., bat the condition
of the typical specimen is such that it is impossible to
determine the genus. The first posterior cell is closed at
the margin of the wing.
Tachina hebes (Ins. Saund. — Dipt. pt. iv. (1852) p. 289. —
Tasmania) is not a female as stated by Walker, but the
male of Tachina densa {ibid. p. 288. — New South Wales).
The species apparently belongs to a new genus near Exorista^
Mg. The antennae are inserted above the level of the middle
of the eye; the face is broad and the frontal bristles descend
to the level of the arista, while a number of small bristles
descend still lower ; the facial ridges are ciliated on the
lower third; depth of jowls about one-third of that of the
eye ; abdominal macrochaetae only marginal.
Eurigaster tasmanice (Trans. Ent. Soc. Lond. n. ser. iv.
(1858) p. 197. — Tasmania) belongs to a new genus, near
Frontinaj Mg., Section Phorocera, Br. & von Berg. : —
Eyes hairy ; ocellar bristles wanting; face hairy; oral margin
very prominent; abdominal macrochaetae marginal ; claws in
male short, as in Frontma. The type is a male, not a female
as stated by Walker; the tip of the abdomen (last segment)
is dull, not shining as stated in Walker^s description.
332
Mr. E. E. Austen on the Synonymy and
Tachina australis (Ins. Saund. — Dipt. pt. iv. (1852)
p. 279. — New South Blepharipoda (Tachina)
zebina, Walk.
Tachina leocrates (List Dipt. Ins. Brit. Mus. pt. iv. p. 475
(1849). — South Africa) is perhaps to be assigned to the
genus Brachelia, Rob.-Desv. (Ess. sur les Myod. p. 61) —
the type of which is Tachina Westermanni, Wied. (Auss. zw.
Ins. ii. p. 291 )_, from the Cape of Good Hope. On the
genus Brachelia, cf. Brauer & von Bergenstamm, Denkschr.
math.-naturw. Cl. k. Akad. Wiss., Bd. lx. 1893, p. 184.
Tachina cassotis (ibid. p. 761. — Sierra Leone). — Head of
type wanting ; the species apparently belongs to the genus
Rhynchomyia, Rob.-Desv.
Tachina verritus (ibid. p. 774. — South Africa) belongs to
the Section Phorocera, Br. & von Berg., and may, pro-
visionally at any rate, be assigned to the genus Doria
(sens'd Rondani) . The statement in Walker’s Latin diagnosis
as to the colour of the palpi, antennae, and legs is misleading;
in tlie typical specimen (a female) the palpi, so far as it is
possible to see them, appear to be brown with yellowish tips ;
in a male in the Museum collection from Malvern, Natal,
March 1897 (G. A. K. Marshall), and a second male from
Leopoldville, Congo Free State, Dec. 13, 1903, In bush’"’
(Drs. Dutton, Todd, ^ Chy'isty), the palpi are entirely yellow ;
in both sexes the antennae and legs are black, the front
femora greyish beneath. The arista is long and slender,
with its second joint greatly elongated in the male; the
third vein is bristly nearly to the small transverse vein, and
the bend of the fourth vein has a Faltenzinke.”
Tachina subaurata (Ins. Saund. — Dipt. pt. iv. (1852)
p. 298. — Cape of Good Hope) belongs to the genus Ceromasia,
Rond.
Tachina ocychus (List Dipt. Ins. Brit. Mus. pt. iv. p. 770
(1849). — Jamaica) is a Frontina, Mg.
Tachina basifulva {ibid. p. 725. — Jamaica) is correctly
referred by Aldrich (Cat. N. Amer. Diptera, p. 485 (1905))
to the genus Archytas, Jaenn.
Tachina infiy^ma (ibid. p. 719. — Chile) is an Archytas,
Jaenn.
Tachina pilosa (Ins. Saund. — Dipt. pt. iv. (1852) p. 266
(^‘ Musca pilosa‘1 Drury ’’). — S. America) = Archytas hystrix,
Eabr.
Tachina latij'rons (ibid. p. 284. — S. America), correctly
333
Generic Position of certain Muscidse.
referred by Walker himself {loc. cit.) to the genus Blephari-
peza, is, as staled hy Aldrich {op. cit. p. 472), a synonym of
B. leucophrySy Wied.
Tachina coniermina {ibid. p. 285. — S. America) is a
Belvosia, and perliaps = {Tachina) atrata, Walk.
Tachina divisa {ibid. p. 270. — =■ Archytas analis,
Fahr.
Tachina apicalis {ibid. p. 245. — California : — not Colum-
bia,^'’ as stated by Walker) = robnsta, Wied.
Trixal sejuncta (Trans. Ent. Soc. Lond. iv. (1858) p. 200. —
Cape of Good Hope) belongs to the genus Dexiosoma^ Rond.
Microphthalma capensis, Schin. (Reise Novara. — Diptera,
p. 322), is a synonym of this species.
Echinomyia albiceps (Trans. Ent. Soc. Lond. n. ser. v.
(1860) p. 295. — Brazil) is an Archytas^ near A. (Tachina)
basifulva, Walk.
Echinomyia ludens {ibid. — Brazil) is devoid of palpi. The
species would be a Saundersia but for the fact of the presence
of a strong bristle on the face on each side, just above the
level of the bottom of the eye. The same character is
exhibited by Saundersia truncaticornis , v. d. Wulp, from
Panama, and a new genus in the near vicinity of Saundersia
will probably be needed for these two species. In Walker’s
species the front tarsi are not expanded in the female,
although they are in S. truncaticornis.
Tachina similis (Ins. Saund. — Dipt. pt. iv. (1852) p. 266. —
New South Wales) apparently belongs to the genus Chcet-
ophthalmus, Br. & von Berg. (Denkschr. math.-naturw. Cl. k.
Akad. Wiss., Bd. Iviii. (1891) p. 383). — Section^’ Micro-
palpus, Br. & von Berg. So far as can be seen from the type,
which is in very poor condition, the palpi are entirely
wanting.
Tachina basalis {ibid. p. 281. — Locality unknown) is an
Aporia, Macq., near A. {Macquartia) venusta, v. d. Wulp.
Tachina vulgata {ibid. p. 300. — S. America). The head
of the type is wanting, and since this species, like the
majority of those described by Walker, is based upon a
single specimen, it is impossible to be certain as to the
genus. From Walker’s description of the head, however,
tfie species would appear to belong to the genus Phorocera,
Rob.-Desv.
Masicera longiuscida (Trans. Ent. Soc. Lond. iv. (1858)
p. 198.— S. America). — Section ■” Phorocera, Br. & von
llcrg. : genus uncertain.
331: Mr. E. E. x\usten on the Synonijmi/ and
Tachina chrysotelus (Tns. Saund. — Dipt. iv. (1852) p. 296.
— Brazil) = Tachina tincta, Walk. [ihid. p. 287). The species
is a Phorocera with the abdominal macrochaetae confined to
the margins of the segments. The Museum collection
contains a female of this species from Santa Catharina,
Brazil (Crowley Bequest).
Tachina compacta (ibid. p. 294. — Brazil) is a Blepharipoda^
Br. & von Berg. : Masicera alacris, Walk. (Trans. Ent. Soc.
Loud, new ser. vol. v. (1861) p. 301) is a synonym of this
species.
Tachina scita (Ins. Saund. — Dipt. iv. (1852) j). 302. —
Brazil). — Tlie type is a female, not a male as stated by
Walker. The face is slightly hairy; the lower anterior
angle of the third joint of the antenna is somewhat
prominent ; the facial ridges are ciliated on rather less than
the lower half ; the abdomen has only marginal macrochaetse,
and the second and third segments are serrate below. — Genus
uncertain : Section Pseiidodexia, Br. & von Berg., near
Decodes, Br. & von Berg., and Gymnostylia, Br. & von Berg.
(?zec Macq.) ; Aldrich (Cat. N. Amer. Dipt.) the species
is probably a Masicera.
Tachina sordida (ibid. p. 297. — S. America). — Genus
uncertain ; Section Pseudodexia, Br. & von Berg. ? —
The type is a male, not a female as stated by Walker; the
palpi are orange, not “ black. The vibrissfu are well above
the oral margin, and the jowls descend considerably behind;
the abdominal macrochsetse are confined to the margins of
the segments.
Tachina umbrifera (ibid. p. 294. — Brazil). — Genus im-
certain, possibly new ; near Ptilodeyeeria, Br. & von Berg.
(^^ Section Pseudodexia, V>\\ & von Berg ). Eyes bare;
jowls somewhat descending behind ; abdominal macrochaetae
only marginal,
Tachina cincta (ibid. p. 303. — Brazil). — The type (a male,
not a female as stated by Walker) is in poor condition, since
the abdomen has been damaged. The species belongs to the
Section Pseudodexia, Br. & von Berg. ; genus uncertain,
perhaps new; near Degeeria, Mg., but the facial ridges are
bare, and the arista is pubescent.
Tachina squamata (ibid. p. 279. — Colombia) apparently
belongs to the genus Lepidodexia, Br. & von Berg. (Deukschr.
math.-naturw. Cl. k. Akad. Wiss., Bd. Iviii. (1891) p. 379) ;
however, in the type, at any rate, the bristles on the third
vein do not extend quite so far as halfway between the
335
Generic Position of certain Museidse.
base and the small transverse vein. Walker^s alulae/^ in
the description of this and other species^ are in reality the
sqnamse.
Tachina tincta (ibid. p. 287. — Brazil) is a Phorocera. —
Vide supra, under T. chrysotelus.
Tachina atratula {ibid. p. 305. — Brazil) is a Phorocera.
Tachina ruficornis (ibid. p. 304. — S. America) is a Chryso-
tachina, Br. & von Berg. The Museum collection contains
two specimens of this species from Atoyac and Teapa,
Mexico (presented by Messrs. Godman ^ Salvin), which
were assigned by van der M^ulp (Biol. Centr. Amer., Dipt.^
Suppl. p. 480) to Gymnochceta alcedo, Lw.
Sarcophaga parva {ibid. p. 321. — Para^ Brazil) is not a
Sarcophaga. The species should perhaps be assigned to the
Section Paramacr onychia, Br. & von Berg.^ but the type
and solitary representative is in such poor condition that it
is impossible to be certain as to the genus. The face is
narrow and somewhat receding, otherwise the species might
perhaps be regarded as allied to the genus Sphixapata,
Rond.
Tachina candens (List Dipt. Ins. Brit. Mus. iv. (1849J
p. 720. — Nova Scotia) is an Archytas. M^alker^s name is
given by Aldrich (Cat. N. Amer. Diptera, p. 486) as a
synonym of Archytas {Jurinia) lateralis, Macq., but at any
rate the type is not conspecific with specimens in the
Museum collection determined as belonging to the latter
species by Prof. Tyler Townsend ; the pile on the face in
WalkeCs type is golden yellow instead of black, the
second, third, and fourth joints of the front tarsi are much
more expanded, and the ])ollinose covering of the thorax and
scutellum is denser and deeper yellow. It may be noted,
however, that according to Macquart the pile on the face of
A. lateralis is white Face d’un jaune pale, a duvet
blanc^^), so that Townsend’s identification is probably
incorrect.
Tachina speculifera {ibid. p. 731. — N. America), of which
the type is a female, is a Micropalpus, Macq. {Linncemyia,
Rob.-Desv.), near M. pictus. Mg. In neither of these
species are the palpi reduced to mere stumps.
Tachina degenera (ibid. p. 732. — Hudson’s Bay) is an
Ernestia, Rob.-Desv. Contrary to Walker’s statement, the
eyes are hairy.
Tachina melobosis (ibid. p. 743. — Florida) is a Phorocera ,
33G
Mr. E. E. Austen on the Synonymy and
Rob.-Desv. : syn. Tachina addita, Walk. (Ins. Saund. — Dipt,
iv. (1852) p. 290) j as correctly stated by Coquillett & Aldrich.
Tachina dydas {ibid. p. 748. — Hudson^s = Eutachina
rustica, Mg.
Tachina masurins (ibid. p. 753. — N. America) is an
Accmyia, Rob.-Desv._, apparently distinct from A. dentata,
Coq., and A. tibialis, Coq.
Tachina clesides {ibid. p. 757. — N. America) is a Phori-
chmta, Rond. P. tricincta, Rond. (Dipt. Ital. Prodr. iv.
(1861) p. 103), described from Italy, is apparently a synonym
of this species.
Tachina hybreas (ibid. p. 785. — Hudson’s Bay). — The head
of the type is missing : discal as well as marginal raacro-
chsetse are present on the abdomen, and the species may
perhaps be referred provisionally to Ceromasia, — sensiX
Br. & von Berg.
Tachma epicydes (ibid. p. 786. — Hudson’s Bay). — Genus
Exorista : Walker's name is given by Aldrich (op. cit.
p. 455) as a synonym of Exorista (Tachina) affinis, Fin.,
but the shape of the third joint of the antenna in Walker's
type does not agree with Fallen’s description.
Jurinia innovata (Trans. Ent. Soc. Lond. new ser. vol. v.
(1861) p. 296. — Mexico) is doubtfully referred by Aldrich
(op. cit. p. 486) to Archytds, but does not belong to that
genus, in spite of the fact that the proboscis is slender and
polished, with small labella. The sp cies must apparently
be assigned to the Seetion ” Erigonc, Br. & von Berg., in
which a new genus will probably have to be founded for it
near Ernestia, Rob.-Desv. (Erigonc, olim'). The head is
proportionately somewhat small; eyes bare; face hairy;
second joint of antennse somewhat elongate ; palpi small
and slender, orange, not black as stated by Walker; third
vein bristly for rather more than half the distance from the
base to the small transverse vein.
Tachina atra (Ins. Saund. — Dipt iv. (1852) p. 273. —
Georgia) is correctly given by Aldrich (op. cit. p. 485) as a
synonym of Archytas aterrima, Rob.-Desv.
Masicera expergita (Trans. Ent. Soc Lond. new ser. vol. v.
(1861) p. 304. — Mexico) is a Degeeria, Mg. (Medina of
Aldrich’s Catalogue), of whieh Degeeria longipes, v. d. Wulp,
from comparison of the types, is a synonym. Apud
V. d. Wulp (Biol. Centr.-Amer.j Dipt., Suppl. p. 485)
7). longipes is a synonym of D. (Oglisa) nigrifacies, Big.
337
Generic Position of certain Muscidte.
(Ann. Soc. Eut. Fr. 1888, p. 268). — The facial ridges in
Walker’s type are not bare, as stated by Walker, but are
somewhat sparsely ciliated up to the middle.
Masicera gentica (ibid. p. 302. — Mexico) apparently
belongs to Ceromasia^ — sensu Br. & von Berg. The type is
now a mere fragment.
Tachina ancilla (Ins, Saund. — Dipt. iv. (1852) p. 299. —
United States) is correctly assigned by Coquillett (Rev.
Tachinidae, p. 106) to the genus Frontina, Mg.
Eurigaster saginata (Trans. Ent. Soc. Lond. n. ser. v.
(1861) p. 298. — Mexico) does not belong to Exorista, to
which it is referred by Aldrich (op. cit. p. 459). The head
of the type is missing^ and the genus must therefore remain
uncertain, but the species must apparently be assigned to
the Section ” Plagia^ Br. & von Berg. — Costal spine
wanting; third vein bristly as far as the small transverse
vein ; posterior transverse vein much more oblique than the
apical portion of the fourth vein, which is very abruptly
bent up ; bend of fourth vein with a very small appendix ;
abdominal macrochsetae marginal.
Tachina ampelus (List Dipt. Ins. Brit.^ Mus. iv. (1849)
p. 732. — Nova Scotia) is an Ernestia (Panzeria of Coquillett
& Aldrich), but is not synonymous with radicum, Fabr., as
erroneously stated by the American writers. — It would
appear from the fact that specimens of another species in
the Museum collection, from Vernon, British Columbia,
April to May, 1902 (^Miss Ricardo), were determined by
Coquillett as Panzeria radicum, Fabr.,” but have in reality
nothing to do with that species, that the interpretation of
Musca radicum, Fabr., by American authors is incorrect.
Miss Ricardo’s species is one of two species of Ernestia
obtained by her at the same locality, and both remarkable
for the smallness of the eyes. The species wrongly identified
by Coquillett has the abdomen unicolorous shining bronze-
black, while in the other the second, third, and fourth
segments have a pollinose band in front.
Tachina aids [ibid. p. 710. — Locality unknown) — - Hystricia
(Tachina) signata, W^alk. (op. cit. p. 709), from New Zealand.
Tachina metallifera (ibid. p. 717. — Locality unknown) is
an Archytas.
Tachina damippus (ibid, p. 719. — Locality according to
the List ” unknown, but from a label on the specimen—
Archytas analis, Fabr.
338
Mr. K. E. Austen on the Synonymy and
Tachina carbonifera [ibid. p. 721. — Locality unknown) is
an Archytas, and apparently = ^. [Tachina) metallifera, Walk.
Tachina fimbriata [ibid. p. 724. — Locality unknown) is a
true Tachina^ Mg. {sensu Meigen and Br. & von Berg.).
The species is very likely from India, since it is closely
allied to Tachina nitida, Walk.
Tachina pagasus {ibid. p. 750. — Locality unknown)
= Eutachina rustical Mg.
Tachina thyamis [ibid. p. 756. — Locality unknown). — The
specimen in the Museum collection which figured as the
type of this species did not agree with Walker’s description;
the name must therefore be cancelled as unrecognizable.
Tachina thyamis [ibid. p. 771. — Locality unknown)
= Pelatachina tibialis, Fin.
T'achina enarette [ibid. p. 758. — Locality unknown). — The
type is a mere fragment and the species consequently
unrecognizable.
Tachina nysas [ibid. p. 758. — Locality unknown) is a
Rhynchomyia, Rob.-Desv.
Tachina petalus [ibid. p. 759. — Locality unknown) is also
a Rhynchomyia.
Tachina segonax [ibid. p. 762. — Locality unknown). —
Genus uncertain : near Xysta, Mg.
Tachina sosicles [ibid. p. 772. — Locality unknown) is
apparently a Frontina. 31ie type is in very poor condition.
WalkeFs statement “sides of the face without bristles is
precisely the opposite of the fact.
Tachina otichestus [ibid. p. 773. — Locality unknown)
belongs to the genus Bothria, Bond, [sensil Br. & von Berg.).
Tachina nepia [ibid. p. 774. — Locality unknown) =
Baumhaueria goniceformis, Mg.
Tachina ipsea [ibid. p. 776. — Locality unknown). — Genus
uncertain ; resembles Metopia in venation, except that the
])Osterior transverse vein is much more oblique ; front not
prominent; sides of face but little receding, fringed nearly
to level of base of third joint of antennae with stout sparsely-
set bristles ; jowls narrow. This species has been placed
near Metopia in the Museum collection.
Tachina opiter [ibid. p. 776. — France) belongs to the
genus Sisyropa, Br. & von Berg., and perhaps=8. ['Tachina)
excisa, Fin.
'Tachina calliphon [;ibid. p. 777.— Locality according to
339
Generic Position of certain MusciJaj.
the List unknown; the type, however, bears a small
label with the word Picton on it, and so is presumably from
either New S.outh Wales or Canada). — Apparently an
Exorista-, the type, which is accompanied by its puparium,
is in very poor eondition.
Tachina scotinus [ibid. p. 742. — Loeality unknown) is
apparently a Pseudo pachystylum,, Wlk. (Wien. ent. Z. x.
(1891) p. 208). — The arista, however, is not geniculate,
although its second joint is elongate ; the face, except next
the eyes, is very bristly to the level of the bottom of the
eyes, but its sides are not fringed with bristles as stated
by Walker.
Tachina broteas (ibid. p. 763. — England) is a Thryptocera,
probably T. pilipennis, Fin.
Tachina rhmo (ibid, p, 778. — Locality unknown) is a
Masicera.
Tachina cymelus [ibid. p. 790. ^Locality unknown) is a
Blepharipoda, Br. & von Berg.
Phorocera expellens (Journ. Proc. Linn. Soc. v. (1860)
p. 155. — Amboyna) is an Exorista.
Nemoraa tenebrosa (ibid. iv. (1860) p. 123. — Macassar,
Celebes) belongs to the Seetioii Blepharipoda, Br. &
von Berg., and is apparently allied to the genus Cteno-
phoroCera, Br. & von Berg. (Denkschr. math.-naturw. Cl.
k. Akad. Wiss., Bd. Iviii. (1891) p. 312), so far as can be
judged from the deseription of the latter. In the type
(a male, not a female as stated by Walker), and in another
male from Macassar [Wallace), the tirst and second abdo-
minal segments are totally devoid of macrochsetse in the
middle line ; a female from the same locality, however, has
a pair of marginal macrochsetse on the second segment. The
hind tibice, not the hind femora as stated by Walker, are
strongly ciliated. The second joint of the antennae is
somewhat elongated, and the ciliation of the facial ridges,
consisting of fine bristles, extends to the level of the base of
the third joint.
Nemoraa postulans [ibid. v. (1861) p. 240. — Dorey, New
Guinea) is an Exorista. The abdomen is narrow, and the
wings are narrow and elongate.
Masicera vicaria [ibid. i. (1857) p. 20. — Singapore) is a
Sisyropa, Br. & von Berg.
Masicera guttata [ibid. iii. (1859) p. 99.— Aru Is.). — Genus
uncertain, Section Phorocera, Br. & von Berg. Contrary
340
]\[r. r]. E. Austen on the Synonymy and
to Walker’s statement, the faeial ridges are eiliated to rather
above the middle ; eyes bare ; arista minutely pubescent to
just beyond the middle ; abdominal macrochaetae discal and
marginal ; first posterior cell opening close to tip of wing. —
An examination of the type of M. guttata shows that the
“ row of white dots along each side on the fore borders of
the segments ” of the abdomen are really transverse silvery
bands, which are interrupted and become more or less obsolete
in the median dorsal region : the depth of the band on the
fourth segment is twice that of the bands on the second and
third segments.
Masicera horrens {ibid. iv. (1860) p. 124. — Macassar,
Celebes) may be referred, at least provisionally, to the
genus Trichohjga, Rond. The second joint of the antennae
IS not noticeably elongated, but the bend of the fourth vein
has a long Faltenzinkey This species presents a strong
superficial resemblance to Masicera morio, Dol., but may be
distinguished at once by its hairy eyes.
Masicera sarcophagata (ibid. vii. (1864) p. 235. — Ceram)
may provisionally be referred to Eutachina, Br. & von Berg.,
but a new genus near Eutachina will perhaps have to be
founded for its reception, since the second joint of the arista
is not elongated, while the third joint is long and tapering
and not noticeably incrassated towards the base ; the oral
margin, too, is not at all prominent.
Masicera dotata (ibid. iv. (1860) p. 123. — Macassar,
Celebes) belongs to the Section ” Pseudodexia and Sub-
Section ” Thelaira, Br, & von Berg., where a new genus allied
to Thelaira will doubtless have to be founded for its recep-
tion.— Walker’s statements are misleading in some respects.
The eyes are hairy, the arista pubescent ; facial ridges ciliated
with stout bristles on the lower third ; one or two stout
bristles on the jowds beneath the eyes ; orbital setae stout in
the $ (S as yet unknown) ; abdominal macrochsetae discal
and marginal ; third vein bristly almost to small transverse
vein, other veins bare ; bend of fourth vein abrupt, deeply
incurved, with an appendix.
Masicera immersa (ibid. p. 124. — Macassar, Celebes) is a
Blepharipoda, Br. & von Berg., and —B. (Tachina) ophirica,
Walk.
Masicera mysolana (ibid. vii. (1864) p. 213. — Mysol,
E. Indian Archipelago) is the ? of Exorista (Nemorcsa)
postulans, Walk.
341
Generic Position of certain Mu^ciJa’.
Masicera manifesta {ibid. v. (1870) p. 154. — Amboyna) is
an Exorista.
Phorocera convertens {ibid. v. (1861) p. 240. — Dorey, New
Guinea) is apparently the ? of Masicera notabilis^ Walk.
{ibid. iii. p. 97), described from the Aru Is. This species
may be left provisionally in the genus Masicera, although
from its general facies, and especially its elongate shape, it
can scarcely be congeneric with Masicera sylvatica, Fin., the
type of the genus.
Masicera prominens {ibid. v. (1860) p. 155. — Amboyna)
may be allowed to remain, at least for the present, in the
genus Masicera.
Masicera (?) tentata {ibid. iii. (1859) p. 98. — Aru Is.)
apparently belongs to a new genus of the Section*’^ Pseudo-
dexia, Sub-Section Thelaira, Br. & von Berg., allied to
Thelaira, Rob.-Desv. The arista is pubescent on its basal
half; the third vein haa a few bristles at the base, the other
veins are bare.
Masicera solemnis {ibid. p. 98. — Aru Is.) is a Blepharipoda,
Br. & von Berg., closely allied to and perhaps identical with
B. {Tachina) ophirica, Walk.
Masicera simplex {ibid. p. 99. — Aru Is.) may be allowed
to remain for the present in the genus Masicera.
Masicera ? ficta {ibid. v. (1861) p. 286. — Batchian,
Molucca Is.) : head of type now missing ; true genus
uncertain.
Eurygaster conglomerata {ibid. iv. (1860) p. 126. — Ma-
cassar, Celebes) is a Sisyropa, Br. & von Berg.
Eurygaster prominens {ibid. p. 127. — Macassar, Celebes)
is a Sisyropa, Br. & von Berg. The palpi are yellow, not
black as stated by Walker.
Eurygaster interdicta {ibid. vii. (1864) p. 213. — Mysol). —
The type is in poor condition. Genus uncertain ; perhaps
new, and belonging to the Section ” Erigone, Br. & von
Berg. The oral margin descends considerably below the
yibrissse ; above the few small bristles next to the vibrissa3
the facial ridges are ciliated with fine hairs nearly to the
level of the arista ; eyes hairy, not bare as stated by
Walker; palpi short and slender; antennse short ; abdominal
macrochsetoe discal and marginal ; no appendix to bend of
fourth vein.
Eurygaster apta {ibid. iv. (1860) p. 126. — Macassar,
Ann. tk Mag. N. Hist. Ser. 7. Vol. xix. 24
342
j\rr. E. E. Austen on the Synonymy and
Celebes) is a Blepharipoda, Br. & von Berg. : the palpi are
yellow, not blaek as stated by Walker.
Eurygaster fingens (ibid. viii. (1865) p. 132. — Salwatty I.
New Guinea) is a Phorocera. The eyes are hairy, not bare
as stated by Walker ; the first portion of the third longi-
tudinal vein, to a point midway between the base and the
small transverse vein, is bristly.
Eurygaster contract a {ibid. iv. (1860) p. 128. — Maeassar,
Celebes). — Genus uncertain; near Blepharipoda^ Br. 8c von
Berg., but hind tibise not uniformly ciliated ; eyes large,
jowls very narrow, linear ; tips of palpi yellow ; four
post-sutural dorso-central bristles; abdominal macrochaetse
marginal.
Eurygaster progressa {ibid. — Macassar, Celebes). — Genus
uncertain, probably new ; near Walk., and Leskia^
Rob.-Desv. {^‘ Section ” Pyrrhosia, Br. 8c von Berg.). — Eyes
pubescent, not bare as stated by Walker; arista minutely
pubescent ; oral margin only slightly prominent ; jowls
bearing a pair of bristles on their central portion and
descending somewhat posteriorly ; number of post-sutural
dorso-central bristles uncertain, owing to condition of
type; abdominal raacrochsetse marginal, in pairs ; a pair of
admedian bristles and a lateral pair on each side on each
segment.
Sarcophaga ingens (List Dipt. Ins. Brit. Mus. iv. (1849)
p. 816. — Locality unknown). — A series of specimens in the
Museum collection from Trinidad, B.W.I. {J. H. Hart :
taken in a flower of Aristolochia gigas) and a male from
Union I., Grenadines, B.W.I. {H. H. Smith), apparently
belong to this species.
Sarcophaga edax {ibid. p. 832. — France ? •’■’) = Myio-
stoma {Estheria) cristatum, Mg.
Sarcophaga inoa {ibid. p. 832. — Galapagos Is.) is not a
Sarcophaga, but belongs to the Section ” Paramacr onychia,
Br. & von Berg., — genus uncertain, perhaps new, near
Arrenopus, Br. & von Berg. Frontal bristles small ; face
sparsely clothed with minute hairs ; epistoma prominent ;
eyes bare ; two post-sutural dorso-central bristles ; ab-
dominal macrochfietse confined to a row on the hind margins
of the fourth and fifth segments.
Sarcophaga! punctipennis (Trans. Ent. Soc. Lond. iv.
(1858) p. 208. — Colombia) is not a Sarcophaga, but appa-
rently belongs to a new genus allied to Phorichceta, Rond.
(‘" SectioiE^ Thryptocera, Br. 8c von Berg.). The type is very
Generic Position of certain Mu-scidie. 313
mouldy, so that it is not easy to make out its characters ;
the face, however, on each side has a row of stout bristles
descending from the front. The apical portion of the fourth
vein and the posterior transverse vein are abruptly bent in
and out in an unusual manner ; the basal portion of the
third vein is bristly almost as far as the small transverse
vein ; abdominal macrochaetse discal and marginal.
Sarcophaga chrysotelus (Ins. Saund. — Dipt. iv. (1832)
p. 329. — S. America) is an Exorista.
Dexia aurinia (List Dipt. Ins. Brit. Mus. iv. (1819)
p. 847. — Locality unknown) = Dexia vacua^ Fin., $ .
Dexia posio (ibid. p. 844. — Cape of Good Hope) is a
Microphthalma, Macq., with the first posterior cell closed
well before the margin of the wing, and long-stalked : owing
to this peculiarity in the venation it may eventually be
advisable to found a new genus for this species.
Dexia australis (Ins. Saund. — Dipt. pt. iv. (1852) p. 314.
— Australia) is a Thelaira.
Dexia notata (ibid. p. 309. — New South Wales) is a
Myiostoma, Bob.-Desv.
Dexia randa (List Dipt. Tns. Brit. Mus. iv. (1849) p. 852.
— Brazil) belongs to the genus Mesembrinella, Giglio-Tos.
Dexia ohsciira (Ins. Saund. — Diptera, pt. iv. (1852) p. 307.
— Brazil). — Genus uncertain, probably new (‘^^ Section"’'’ Dexia,
Br. & von Berg.). — Arista feathered with long hairs; septum
between antennse and grooves for latter well-marked ; pro-
boscis slender, polished, and somewhat elongate ; abdominal
macrochsetae marginal on second segment, discal and marginal
on third ; terminal portion of fourth vein bent up sharply,
then incurved ; bend with a small appendix. — In spite of the
greater development of the facial septum and of the an-
tennary grooves, this species is probably congeneric with
Rhamphinina picta. Bigot (Ann. Soc. Ent. Fr. ser. 6, t. viii.
(1888) p. 265), from Cuba, the type of which, through the
generosity of Mr. G. H. Verrall, is now in the British
Museum. According to Brauer (SB. k. Akad. Wiss.,
math.-naturw. Cl., Bd. cvi. (1897) p. 359. 105) Rh. picta,
Big., with Dexia potens, Wied., belongs to Stomatodexia
according to the head, and to Leptoda according to the bend
of the fourth vein. The proboscis, however, is very different
from that of stomatodexia.
Dexia muscaria (fbid. p. 308. — Brazil) is congeneric with
the foregoing, though discal and median marginal macro-
chsetse are absent from the second abdominal segment, and
discal macrochsetse from the third segment.
24*
34 i l\Ir. E. E. Auston on the Synonymy and
Dexia angmta {ibid. p. 314. — Brazil) is a Stomatodexia,
Br. & von Berg., near, or perhaps identical with, S. (Dexia)
diadem, a, Wied.
Dexia plana (ibid. p. 315. — Brazil) is a Stomatodexia, near
S. diadema, Wied.
Dexia svffusa [ibid. p. 317.- — Locality unknown) appa-
rently belongs to a new genus o£ the Section Pseudo-
dexia, Sub-Section Thelaira, Br. & von Berg., near
Xanthodexia, v. d. Wulp. This species presents a deceptive
resemblance to Xanthodexia sericea, Wied., from which,
however, it can at once be distinguished by the frontal
stripe being wide and by the presence of marginal macro-
chmtseon all the abdominal segments. Orbital setae in female
stout and conspicuous ; face and jowls narrow ; antennae
rather below level of middle of eyes ; bend of fourth vein
rounded.
Dexia? albicans (Trans. Ent. Soc. Bond. iv. (1858)
p. 204. — R. Amazons, Brazil) belongs to the Section
Sarcophaga, Br. & von Berg., and apparently to a new
genus. — Very narrow and somewhat resembling a Scato-
phaga in shape and general appearance; head in profile
square, with rather prominent front ; eyes semicircular in
outline; face bare; arista feathered on rather more than
basal half; abdomen, femora, and front and middle tibise
clothed with short w^oolly hair ; hind tibiae in male shortly
ciliated on inner side ; first joint of front tarsi somewhat exca-
vated on underside ; abdomen totally devoid of macrochaetae
except on hind margin of fourth segment ; third vein bristly
for three-fourths of the distance from the base to the small
transverse vein ; remaining veins bare.
Dexia insolita (Ins. Saund. — Dipt. iv. (1852) p. 318. —
Brazil) may, provisionally at any rate, be referred to Hystri-
chodexia, von Roder : there is a row of stout macrochaetae on
the hind margins of the second and third abdominal segments,
but on the disc of the latter segment macrochaetae appear to
be wanting.
Tachina similis (ibid, p, 269. — S. America) is a Peleteriay
near P. robusta, Wied,
Dexia harpasa (List Dipt. Ins. Brit. Mus. iv. (1849)
p. 840. — N. America) = Plilodexia tibialis, Rob.-Desv.
Dexia cerata (ibid. p. 847. — N. America) is a Rhyncho-
dexia, v. d. Wulp (Bigot, joro joaWc).
Dexia pristis (ibid. p. 841. — Massachusetts) belongs to
the genus Aporia^ Macq. The type is in very poor
condition.
345
Generic Position of certain Muscida?.
Dexia cremides [ibid. p. 842. — N. America) must be can-
celled as unrecognizable. The specimen that the writer
found in the Museum collection doing duty as the type does
not agree with the description, and is a female of Dinera
grisescens, Fin., a species apparently not included in AldriclFs
‘ Catalogue of North American Diptera,^ where (p. 500) Dexia
cremides, Walk., is placed under Myiocera.
Dexia ogoa (ibid. p. 841. — Nova Scotia) must also be
cancelled as unrecognizable, since the type is not now to be
found in the Museum collection, and Walker states that
when he described it the head was missing.
Dexia dirphia [ibid. p. 836. — Locality unknown) is a
Myiocera, Rob.-Desv.
Dexia prexaspes (ibid. p. 837. — Georgia, U.S.A.) is a
Ptilodexia, Br. & von Berg., to which genus Estheria abdo-
minalis, Rob.-Desv., from Nova Scotia (assigned by Aldrich,
Cat. N. Amer. Dipt, p; 501, to Dexia), also belongs.
Dexia abzoe [ibid. p. 846. — Georgia, U.S.A.). — Type not
now to be found in the Museum collection ; name should
consequently be cancelled.
Tachina corythus (ibid. p. 797. — Georgia, U.S.A.). —
So far as it is possible to determine from a comparison of
Walker’s type and the descriptions, Coquillett (Rev. Tachi-
nidae, p. 73) is apparently correct in quoting T. corythus,
Walk., as a synonym of Xanthomelana (Pkasia) atripennis.
Say.
Tachina cdops [ibid. p. 796. — Georgia, U.S.A.) . — Cor-
rectly referred by Coquillett [op. cit. p. 73) to the genus
Beskia, Br. & von Berg.
Dexia hypsa [ibid. p. 866. — Locality unknown) is a
Fycnosoma, Br. & von Berg.
Musca liris [ibid. p. 882. — Locality unknown) is the
female of Rutilia minor, Macq. (Australia and Tasmania),
which apparently should be referred to a new genus near
Rutilia.
Dexia albifrons (Ins. Saund. — Dipt. iv. (1852) p. 317. —
United States) is a Rhynchodexia, v. d. Wulp (Bigot, pro
parte) .
Dexia canescens [ibid. p. 310. — United States) is a Ptilo-
dexia, near and perhaps only a dark form of P. tibialis,
Rob.-Desv.
Dexia pedestris (ibid. p. 313. — United States) has nothing
to do with Dexia. It is synonymous with Tachina menapis.
Walk. (List Dipt. Ins. Brit. Mus. iv. (1849) p. 769), the
34 r> On the Synonymy &c. of certain MusclJgp.
type of wliicli was stated to be from Upsala, Sweden. The
present writer is unable to assign this species to its proper
genus, but it perhaps belongs to the Section Masicera,
13r. & von Berg. The front is prominent and the face
hairy ; depth of jowls rather greater than half the height of
the eye ; first posterior cell opens at or close to tip of wing.
Gymnosiylia invita (Journ. Proc. Linn. Soc. v. (1861)
p. 243. — Dorey, New Guinea) = Dexia alulifera, Walk.
(ibid. p. 157 ; described from Amboyna). — Genus uncertain,
probably new Section Pseudodexia^ Sub-Section
Thelaira, Br. & von Berg.) .
Gymnosiylia luteicornis {ibid. vi. (1862) p. 10. — Gilolo,
E. Indian Archipelago) belongs to an apparently new genus
near Thelaira, Rob.-Desv. — Eyes large, occupying whole
depth of head, so that jowls are reduced to a mere line ;
antennae below level of middle of eyes ; abdominal macro-
chaetae discal and marginal ; first and third veins bristly.
Trichoprosopa^ marginalis {ibid. v. (1860) p. 157. —
Amboyna). — A new genus will have to be founded for this
species close to Ocyptera, Latr., from which it is distin-
guished by the dorso-central bristles being greatly reduced,
as well as by the depth of the head, narrowness of the face,
and length of the third joint of the antennae.
2'richoprosopal divisa {ibid. vii. (1864) p. 213. — Mysol)
appears to be a Plesiocyptera^ Br. & von Berg., although
differing from P. {Ocyptera) bicolor, Wied. (the type of the
genus), in the face being narrow and the proboscis not
slender. In the typical specimen, at any rate, the second
and third abdominal segments are devoid of admedian
marginal macrochaetae.
Tacliina titan (List Dipt. Ins. Brit. Mus.iv. (1849) p. 735.
— Sylhet) is a Nemorcea, Bond., near, but apparently distinct
from, N. iropidobothra, Br. & von Berg. Van der Wulp
(Cat. Descr. Dipt. S. Asia, p. 126) assigns this species to the
genus Micropalpus, doubtless misled by Walker having
headed the page on which the description appears Group II.
Linncemyia, Desv. Essai Myod. 52.^^
Musca aluta (ibid. p. 911. — ‘^Lapland?, France?'”). —
The type is a female of Myiospila meditabunda, Fabr.
Tachina zebina (ibid. p. 772. — North Bengal) is a Ble-
pharipoda, Br. & von Berg. (i.e. Sturmia, Bob. -Desv., apud
Coquillett& Aldrich), and perhaps a synonym of B. (Tachina)
cilipes, Macq. (Dipt. Exot. ii. 3 (1843), p. 62, tab. 6. fig. 6).
The following Walkerian species are synonyms of B. zebina-. —
'Tachina fusiformis (List Dipt. Ins. in Coll. Brit. Mus. iv.
3’47
On new Coleoptera from the Indian Empire.
(1849) p. 1161) ; Eurygaster mutans (Jourii. Proc. Linn.
Soc. V. (1861) p. 240) ; Tachina australis (Ins. Sannd. —
Dipt. iv. (1852) p. 279) ; and Nemorcea amplificans (Journ.
Proc. Linn. Soc. iv. (1860) p. 122). The extensive series of
specimens in the Museum collection shows that B. zehina is
among the commonest of the larger Tachininse in India and
Ceylon^ and also occurs in Burma^ Malacca^ Java, the Sunda
Is., Celebes, Dorey (New Guinea), Queensland, and New
South Wales.
Tachina grandis (Ins. Saund. — Dipt. iv. (1852) p. 278,
pi. vii. fig. 1. — India) belongs to the genus Nemorcea, Rob.-
Desv., and may be the female of N. tropidohothra, Br. &
V. Berg., as stated by van der Wulp (Tijdschr. v. Ent. xxxvi.
p. 161, & Cat. Descr. Dipt. S. Asia, p. 129). But, since
Walker^s type does not altogether agree with the description
by Brauer and von Bergenstamm, it seems advisable, for the
present at any rate, to regard the two species as distinct.
XLl. — Some new Species and Genera of Larnellicorn Goleo-
ptera from the Indian Empire, By Gilbert J. Arrow.
This paper contains descriptions of some of the most important
Indian Lamellicornia which are yet unnamed in the British
Museum collection, together with observations upon allied
forms which have accrued in the course of its preparation.
Part I.
CetoniidaB.
Heterorrhina dispar, sp. n.
8at elongata, crebre punctata, processu sternali hand longo, sab
acuto.
(5 . Viridi-olivacea, abdomine pedibusque riifis, nitida; capite inermi,
excavate, clypei margine antica valde retlexa et arcuata ; pedibus
sat gracilibus, tibiis anticis vix dentatis.
5 . Obscure castanea, opaca ; clypeo valde excavato, antice nasuto,
Venice cornu decumbente truncato armabo; pedibus crassioribus,
tibiis anticis latis, bidentatis.
Long. 20-22 mm.
Hah. N. India, Darjeeling.
The form is elongate, not much depressed, rather strongly
and uniformly punctured above. Head rugosely punctured,
with the clypeus rather broader than long and the front
margin prominent in the middle. Prothorax coarsely and
closely punctured, with the interstices extremely finely punc-
tulated. iScutelluin punctured except along the middle line.
348
Mr. G. J. Arrow on neic
Elytra rugosely punctured, some of the punctures forming
double rows. Pygidium transversely rugose. Sternal process
short, but rather sharp. Metasternum smooth in the middle
and coarsely punctured at the sides. Abdomen finely
punctured.
. Olive-green, shining, with the abdomen and legs
reddish. Head unarmed behind. Clypeus somewhat exca-
vated, with the front margin curved, reflexed, and slightly
produced in the middle. Sides of the prothorax strongly
angulated in the middle and nearly straight in front and
behind.
? . Purplish black, with abdomen and legs castaneous.
E orm more elongate, with the upper surface more opaque.
Clypeus strongly excavated, with the front margin ratlier
strongly produced upwards in the middle and the vertex armed
with a horizontal process freely produced and truncated in
front. Prothorax more convex and less contracted in front
than in the male. Legs stouter, with the front tibiae strongly
bidentate and all the tarsi shorter and thicker than in the
other sex.
In the peculiar differences of form and colouring between
the two sexes this species shows relationship only to
H. mutahilis, Hope, from which it is quite easily distin-
guished. It is larger and has less distinctly costate elytra,
besides which the sternal process, although short, is much
less so than in that species, and the clypeus is longer and
quite differently shaped in both sexes.
A male and female have formed part of the British Museum
collection for fifty years, but have not hitherto been distin-
guished from the allied species.
Anomalocera subopaca, sp. n.
Olivaceo- vel purpureo-viridis, antennis tarsisque nigris, elongata,
paulo depressa ; capite granulate, cl}'peo quadrato, antice paulo
dilatato, marginibiis fere rectis ; prothorace crebre punctato,
medio subtilissime, lateraliter grosse et rugose, lateribus vix
arcuatis, medio leviter angulatis ; scutello elongate, acuto ; elytris
parum profunde rugose punctatis, obsolete striatis, postice mar-
ginibus grosse rugulosis ; pygidio dense rugoso, breviter setoso ;
processu sternali sat longo, parum curvato, metasterni lateribus
dense punctatis, piliferis ; abdomine fere Hvi.
(5 . Angustior, prothorace antice magis contracto, abdomine sul-
cato, clava antennali longissima ; tibiis auticis muticis, posticis
dense et longe fulvo-ciliatis.
Long. 22 mm.
llah. N. India, Manipur,
Green, with pinkish reflections ; elongate, ))arallsl-sided.
349
Coleoptera from the Indian Empire*
ratlier flat above, and not bigldy glazed. Clypeus granu-
lated, about as long as it is broad, slightly widening towards
the front, with the anterior and lateral margins nearly straight.
Prothorax very finely punctured upon the disk and very
coarsely and rugosely at the sides, rather shorter than in
A. glaherrima and Mearesi^ and with the sides rather more
distinctly angulated in the middle and the base strongly
trisinuate. Elytra finely and shallowly but ratlier closely
strigosely punctured, some of the punctures forming rows
anteriorly, with the apical and posterior lateral parts coarsely
strigose, but scarcely setose, and the apical angles slightly
produced. Pygidium densely rugose and clothed with short
and not closely-set setae. Sternal process moderately long,
depressed, blunt, and not much curved. Metasternum
smooth and deeply grooved at the middle and densely punc-
tured and pubescent at the sides. Abdomen almost smooth.
The male is more elongate, with the prothorax more
narrowed in front, the abdomen deeply channelled, the an-
tennal club very long, and the hind tibiae thickly fringed.
Half a dozen specimens were collected by Mr. Doherty.
The species is closely related to A. glaherrima and Mearesi^
but is rather more elongate and depressed, and the upper
surface is much less glassy, being rather closely sculptured
all over. The male has the prothorax less narrowed in front
than in those two forms.
The' genus Anomalocera was formed for A. Mearesi^ Hope,
alone, but that species is only peculiar in the rather greater-
length of the antennal club of the male than that of its
allies, and a natural group is formed by associating with it
the species which have been placed in Ileterorrhina and
Rhomborrhina^ characterized by a tapering sternal process,
the clypeus simple in both sexes, and the hind tibias straight.
The genus Heterorrhina will then be confined to the forms in
which the clypeus is armed in one (the female) or both sexes,
and Rhomborrhina to those in which the sternal process is
broadly transverse and the clypeus of the characteristic
spatulate shape.
The species I include in Anomalocera as thus defined are
A. Mearesi^ Hope, glaherrima^ Westw., mbopaca^ sp. n.,
microcephalaj Westw., Mellii, G. & P., heros^ G. & P.,
resplendens^ Schonh., rujitibiis^ Bates, unicolor , Motsch.,
Fortunei^ Saund., and olivacea^ Jans. There still remains
one isolated species, which, while it has the clypeus simple
in both sexes and the sternal process long and slender, cannot
be associated with this group. The antennal club is equally
short in both sexes, the clypeus is nearly square in shape,
the elytra are very strongly and regularly punctate-striate.
350
Mr. G. J. Arrow on new
and in particular the hind tibiae of the male are strongly bent
and furnished with a thick brush of hairs. This is Cetonia
loeta, F., with which Heterorrhina sylhetica^ Thoms., is
synonymous. It was described by Thomson from North
India and by Fabricius from Java, but it has a continuous
range through Burma and the Malay Peninsula, and the
differences mentioned by Thomson have no reality when a
series of specimens is compared. For this form I propose
the new genus Euchloropus.
The chief differential characters of the Heterorrhina group
of genera may be tabulated as follows : —
Clypeus armed in one or both sexes Heterorrhina.
Clypeiis unarmed.
Hind tibiae of strongly curved Euchloropus.
Hind tibiae of S straight.
Sternal process elongate Anomalocera.
Sternal process transverse Rhomborrhina.
Macronota gracilis^ sp. n.
Nigra, supra opaca, elytris obscure rufis, macula postscutellare ad
humeros producta, fascia media transversa apicibusque nigris,
scutelli apice, suturae medio lineolisque duabus transversis post-
humeralibus albis, sterni abdominisque lateribus plus minusve
albis ; sat parva, angustata, capite (tuberculo laevi postico excepto),
prothorace pygidioque omnino granulosis, clypeo modice emar-
ginato ; prothorace fere circular!, postice leviter lobato et depresso,
angulis vix perspicuis ; singulo elytro fortiter unicostato, lateribus
post humeros sat leviter sinuatis ; pedibus gracilibus, tibiis
anticis tridentatis, posterioribus omnibus sine dentibus.
Long. 15 mm.
Hah. Assam, Naga Hills.
Black, with the elytra dark red except for a black patch
behind the scutellum produced to the shoulders, a transverse
median fascia, and the apical margins, and decorated with
white markings consisting of a spot behind the scutellum,
another at the middle of theelytral suture, and two transverse
marginal spots on each elytron. The sides of the sternum
and the margins of the basal segments of the abdomen are
also marked with white.
The form is very elongate, tapering behind, and the legs
are slender. The head, pronotum, and pygidium are coarsely
granulated. The head is flat, with a smooth tubercle on the
vertex and moderately notched in front. The prothorax is
almost circular in shape, with all the angles almost obsolete,
and moderately depressed behind. The ely^tra have a silky
sheen, and each has a strong costa and is feebly sinuated
351
Coleoptera from the Indian Empire.
beliind tlie shoulder. The front tibiae have three slight teeth
and the four posterior tibiae are without teeth or spines at the
middle. The antennal club is of moderate length.
Glycosia hiplagiata^ sp. n.
Nigra, nitida, elytris opacis, plaga suturali nitida excepta, singulo
post medium flavo-maculato ; elongata, depressa, ad humeros
lata, deinde augustata, capite parvo, crebre punctato, antice sat
emarginato ; prothorace convexo, grosse punctato, sat late
raarginato, lateribus medio angulatis ; scutello sat parvo, impunc-
tato ; elytris striato-punctatis, lateribus post humeros valde
sinuatis, apicibus sinuatis, angulis suturalibus acutis ; pygidio
parvo, fere laevi ; corpore subtus nitido, lateribus grosse punc-
tatis : S xu’othorace basi latiore, tibiis anticis minus distincte
tridentatis.
Long. 20 mm.
Hah. Andaman Is.
Shining black, with the elytra opaque and sooty except at
the inner margins, and with a lemon-yellow patch beyond
the middle of each, broad at the outer margin and pointed at
its inner extremity. The form is depressed, broad at the
shoulders, with the head small and the elytra straight at the
sides and strongly narrowing towards the extremity. The
head is strongly punctured, with the clypeus long, narrowing
towards the front, where it is rather deeply notched, and not
at all reflexed at the margin. The prothorax is convex,
coarsely punctured, with the sides strongly margined, and
angulated in the middle. The scutellum is rather small,
pointed, and impunctate. The elytra are coarsely striate-
punctate, with the margins strongly sinuated behind the
shoulders and minutely excised at the extremities, and the
apical angles acute. The pygidium is feebly punctured and
the metasternum and abdomen coarsely, except at the middle.
The sternal process is compressed, truncated in front, and
directed obliquely downwards. The front tibim are 3-dentate
in the female, but the uppermost tooth is almost obsolete in
the male. In the latter the prothorax is broader at the base
and the apical angles of the elytra are strongly spinose.
The type specimen is a female found by EoepstorfF in the
Andamans. There is a male in the Calcutta Museum labelled
Rangoon,” probably by mistake.
The only other known species of Glycosia are G. tricolor,
Oliv., and G. palliata, Mohn.
Protoitia hidentipes, sp. n.
Elongata, parallela, depressa, nigra vel purpureo-nigra, supra velu-
tina, subtus nitida, maculis capitis 2, prothoracis 6-8, utriusque
352
Mr. G. J. Arrow on new
elytri 9, ])ygidii 2, flavis ; caj)ifcc crebre punctato, anlice profunde
excise ; prothorace laxe punclato, transverse, lateribus inedie
angulatis, pestice fere parallelis ; elytris punctato-striatis, apico
paule spinesis ; pygidie crebre punctate, breviter flavo-setese ;
tibiis anticis acute bidentatis.
Leng. 18 mm.
Hab, Nicobar Is.
Sooty black or piceous black, with the head, legs, and
underside shining, decorated with yellow spots distributed as
follows : — a pair upon the vertex of the head, a pair at the
middle and three at each lateral margin of the pronotum, the
two posterior ones sometimes uniting ; three placed in an
oblique line upon the anterior half of each elytron, two ad-
joining the suture posteriorly, and four adjoining the lateral
margin, and a large patch at each side of the pygidium.
There are also patches upon the mesosternal epimera and the
sides of the sternum and abdomen. The head is thickly
punctured, with the clypeus long and deeply notched in front.
The prothorax is very transverse, distinctly but not densely
punctured all over, with the sides strongly angulated in the
middle, and nearly parallel from there to the base, which is
strongly emarginate before the scutellum. The scutellura is
rather narrow. The elytra are parallel-sided, punctate-striate,
with the sutural angles rather spinose. The mesosternal
process is moderately prominent, nearly cireular, and not
much dilated at the end. The metasternum is rugose at the
sides and the abdomen sparsely punctured. The front tibise
are bidentate in both sexes. The pygidium is pubescent in
two female specimens in the British Museum collection, but
in a male in the Calcutta Museum labelled (no doubt
wrongly) “ Rangoon the setae are scarcely visible. The
yellow markings in that specimen are also of a deeper colour.
Clinteria truncataj sp. n.
Nigra vel obscure cuprea, opaca, capite, pedibus corporeque subtiis
nitidis, prothoracis lateribus anguste llavo-marginatis, elytrorum
maculis 2 vel 4 minutis post medium transverse positis, apicalique
minuta, pygidii maculis duabus sat magnis ; depressa, sat
parallela, postice parum attenuata ; capite elongate, dense sat
regulariter punctato, antice valdo excise ; prothorace subtiliter
punctato, postice fortiter sat acute lobato ; elytris grosse seriato-
punctatis, costis duabus distiuctis ; processu sternali valido,
conico, acuminato, abdomine medio glabro.
Long. 15-18 mm.
IJah. S. India, Nilgiri Hills.
Black or very dark coppery, velvety and opaque above.
353
Coleoptera from the Indian Empire,
with the head, legs, and underside sliining, and with white or
yellow markings, consisting of a narrow line on each side of
the prothorax, frequently interrupted or absent, a spot upon
the mesostern'al epimeron, two small spots placed close
together behind the middle of each elytron and frequently
coalescing, a minute external apical spot, a large patch on
each side of the pygidium, and a row of spots on each side of
the sternum and abdomen.
The head is closely punctured, rather long, and deeply
notched in front. The prothorax is finely punctured,
attenuated in front, and strongly and rather sharply lobed
behind. The elytra are rather parallel-sided and little
narrowed towards the extremity ; they are coarsely punctured
in rows, with two well-marked costae upon each. The sternal
process is conical, rather long, and acuminate.
In the female there are three sharp equidistant teeth to the
front tibia. In the male the uppermost tooth is distant from
the other two and much shorter.
The pale markings are liable to reduction, and in one
specimen in the British Museum have disappeared entirely
I have seen a considerable number of specimens of this
form, which is generally confused with Clinieria guttifera,
Burm. [=C. valida^ Lansb.). It differs by its straight and
more parallel sides, the median spots of the elytra placed
farther back and not obliquely, and the more sharply pointed
sternal process.
I at hrst regarded this as possibly the C. modesta^ Blanch.,
but the examination of a co-type kindly sent from the Paris
Museum by M. Lesne shows tlie latter to be a variety of
G. Klugi, Hope (= C. Jtavopicta^ Bl.), with the pale markings
very much reduced.
Anthracophora hufo, sp. n.
Ovata, sat depressa, piceo-nigra, velutina, supra et subtus fulvo-
variegata, prothoracis, metasterni abdominisque medio costisque
duabus elytrorum basi glabris, immaculatis ; capite grosse punc-
tate, prothoracis lateribus grossissime et crebre, medio la^vius sed
fortiter punctatis, marginibus valde sinuatis, basi angulatim
emarginata ; scutcllo baud longo, punctate, variegate ; elytris
irregulariter striatis et punctatis, costa distincta basali, lateribus
vix sinuatis ; pygidio rugoso, variegato ; corpore subtus grosse
punctate, lateribus rugose punctate, opaco et variegato, processu
sternali breviter acuminate ; pedibus brevibus, variegatis, tibiis
anticis minute bidentatis.
Long. 16 mm.
Hah. Sylhet.
Deep red-brown, irregularly speckled above and below with
354
Mr. G. J. Arrow on new
yellow markings, and opaque and velvety except at the
middle of the prothorax, metasternum, and abdomen, and a
strong costa on the anterior half of each elytron.
The form is ovate and depressed. The clypeus is broad,
entire, and strongly punctured. The prothorax is strongly
but not closely punctured in the middle, very coarsely and
rugosely at the sides, with the lateral margins distinctly
angulated at the middle and strongly sinuated behind. The
scutellum is rather short, punctured, opaque, and variegated.
The elytra are irregularly punctured and striated, and each
has a smooth curved costa on the basal half. The pygidiurn
is rugose and the metasternum and abdomen are strongly
punctured and shining in the middle, and opaque and closely
sculptured with crescent-shaped impressions at the sides. The
legs are very short, opaque, and decorated like the body, and
there are two very short teeth to the front tibia.
A single specimen from the Bowring Collection has been
many years in the Museum.
Rutelidae.
Fruhstorferia hirmanica^ sp. n.
Ilufo-castanea, corpore subtus pygidioque sat longe rufo-hirsuta.
Corpus breve, robustum, grosse punctatum, modice nitidum, capite
rugoso-punctato, antice angustato, apice curvato, carina oculari
minuta, baud acuta ; prothorace transverse, grosse punctate,
medio indistiricte siilcato, lateribus bisinuatis, angulis anticis fere
rectis, posticis obtusis, margine basali leviter trisinuato ; scutello
lato, obtuse angulato, punctate ; elytris crebre irregulariter
punctatis, lineis indistinctis nonnullis ; pedum 2 anteriorum
tibiis tridentatis, 4 posteriorum unguibus externis profunde
fissis.
(S . Corpore breviore, magis parallelo, pygidio incurvato ; sat
nitido, mandibulis omnino exsertis, ad capitis longitudinem
sequalibus, valde recurvatis, acutis, tarsis anticis incrassatis
ungueque externo multo majore.
Long, (sine mandibulis) 16 mm.
5 . Multo longiore, ovali, pygidio producto, crebre punctato, corpore
supra grossius punctato, punctisannuliformibus, mandibulis parvis,
obtusis, elytrorum lateribus post medium callosis.
I-ong. 20 mm.
Hah. Burma, Euby Mines (^Doherty').
Chestnut-red, strongly punctured, with the pygidiurn and
lower surface clothed with tawny hair. The legs are stout,
with the front tibise tridentate and the outer claws of the two
posterior pairs strongly bitid. There is a short tubercular
prosternal process tufted at the end.
355
Coleoptera from the Indian Empire.
^ . Short, robust, and parallel-sided, with the head coarsely
punctured, the clypeus small, narrowed and rounded at the
apex, the mandibles produced (about as long as the head),
strongly recurved, and acute at the tips. The prothorax is
transverse, rather strongly punctured, with the sides parallel
behind and the base feebly trisinuated. The scutellum is
very short and finely punctured. The elytra are strongly
and irregularly punctured, some of the punctures forming
lines. The pygidium is turned inwards and sparingly punc-
tured. The front tarsi are thickened and the outer claw
much larger than the inner one.
$ . Longer and more oval, with the mandibles not pro-
duced or acute, the head more rugose and the prothorax,
elytra, and pygidium more coarsely and closely punctured.
The elytra have a small but well-marked fold beyond the
middle of the outer edge, and the pygidium is prominent.
A single pair of this species was received with the Fry
Collection. They are about equal in size to small speci-
mens of F. ^-maculata, Kr., but are more robust and convex
than that species. It is more strongly punctured above and
more hairy beneath, and differs also by the prominent
pygidium and well-marked lateral thickening of the elytra in
the female and the form of the mandibles in the male. It is
likely that the latter attain a greater development than in the
type, but they are peculiar by their twisted appearance and
abruptly acuminate tips. The maxillary palpi are much
stouter than in the other species.
Dr. Ohaus has sent me for comparison a female from
^Tonkin in his collection. It is like the Burmese female, but
more elongate and rather more finely sculptured upon the
elytra, and may prove to represent another species.
Desmonyx, gen. nov.
Labium elongatum, acuminatum, sine ligula distincta, palpis prope
apicem positis, triarticulatis, articulo ultimo grandi, fusiformi.
Maxillse reductae, absque lobis aut dentibus, paljjis prope apicem
positis, 4-articulatis, articulo ultimo grandi fusiformi.
Mandibulae porrectae, falciformes.
Labrum porrectum, angustum, integrum, dense ciliatum.
Clypeus antice latus, trilobatus, ad antennarum basin valde con-
strictus.
Oculi parum prominentes.
Antennae ( 6 ) longae, 10-articulatae, articulo primo valde clavato,
2-6 subaequalibus, 7 sat brevi, 8-10 longissime flabellatis, ad
praecedentes omnes longitudine aequalibus.
Prothorax transversus, lateribus basique arcuatis.
Scutellum latum, fere semicirculare.
356
Mr. G. J. Arrow on new
Sternum muticum.
Pedum coxm anticm prominentes, tibiae 4-dentatae, tibiae 4 posteriores
extus spinosae, posticae apice minute serratae : pedum anticorum
( (^ ) uugue externo lato, valde dentato, articulo penultimo subtus
lobato : pedum 4 posteriorum unguibus aequalibus, gracilibus,
intcgris.
Desmonyx humeralis^ sp. n.
Late ovatus, brunnens, antennarum clava, vertice, prothoracis medio,
scutello elytrorumque parte antica obscurioribus, singulo elytro
antice bimaculato, maculis flavis, approximatis, una prope scutel-
lum alteraque minore posteriore prope marginem lateralem.
Corpus supra grosse et rugose punctatum ; clypeo subtiliter rugoso,
lato, antice trilobate, medio acute, postice valde constricto ;
fronte prothoraceque grosse irregulariter punctatis, laxe hirsutis,
hujus medio longitudinaliter impresso, basi leviter arcuata,
lateribus arcuatis, antice paulo approximatis ; scutello sat minute
punctato ; elytris grosse punctato-striatis et rugosis, singulo apice
arcuato ; pygidio nitido, minute et laxe punctate ; corpore subtus
longe hirsute.
Long. 9-10 mm.
Hah. Burma, Buby Mines {Doherty).
Bark mahogany colour, with the back of the head, the
middle of tlie prothorax, the scutellum, and the anterior part
of the elytra, as well as the club of the antennae, black, and
with two yellow spots near the anterior border of each elytron —
one near the scutellum and a rather smaller one a little behind
and outside the first. The body is robust and convex,
coarsely and rugosely punctured above, wdth the vertex of
the head and the prothorax hairy. The clypeus is finely
rugose, broad and trilobed in front, strongly constricted at
the base of the antenna?. The prothorax is channelled down
the middle, broadly rounded at the base, with the front angles
acute and the hind angles rounded. The scutellum is nearly
semicircular and finely punctured. The elytra are coarsely
])unctate-striate with the intervals rugose. The pygidiurn is
minutely punctured and shining.
The female is unknown. The antennal club is very long
in the male, the last joint of the front tarsus is enlarged, the
inner claw thickened and very widely cleft, and the fourth
joint produced beneath the claw-joint. The other claws are
simple, slender, and equal.
This is an isolated and very remarkable genus, which must
be placed amongst the primitive and polymorphic Kutelidae
of the Parastasia group, although it has considerable affinities
with the Dynastida?. The slender and symmetrical claws of
the four posterior feet are quite foreign to the llutelida? and
357
Coleoptera from the Indian Empire.
the clypeus and the organs of the mouth are very peculiar.
Another aberrant genus to which Desmonyx seems to show
some relationship is OryctomorphuSy a Chilian genus for- which
Lacordaire formed a special group of Dynastidae (the Orycto-
morphides), associating with it an African genus, Homoeo^
morphuSj and an Australian one, Corynophyllus. This
curious assemblage is quite unnatural and a better resting-
place has to be found for Oryctomorphus. Its claws are
movable and slightly unequal on all the feet, and those of
the front feet of the male are exactly as in Desmonyx, the
labrum has a thickened front margin which is visible
externally, and in other respects, conspicuously in its bright
colouring and sexual differences, it agrees less with any
other genus of Dynastidae than with the Parastasia group of
Rutelidae. The latter embraces a variety of forms already
recognized as connecting the Rutelidae and Dynastidae, but
the latter family, if these aberrant members are excluded
from it, becomes fairly homogeneous. H. W. Bates in
forming the genus Metapachylus has pointed out its relation-
ship to Oryctomorphus and also to Parastasia and Polymoeclius,
but without definitely assigning it to either family.
Another genus, Mesystoechus of Waterhouse, placed by
its founder among the Anoplognathini, seems to belong also
to the present group. Its labrum is quite unlike that
characteristic of the former group.
Melolonthidae.
Dejeania lineata, sp. n.
Fusco-nigra, supra squamosa, subtus nitidior, pectore pedibusque
sparse, abdomine dense, griseo-setosis ; clypeo semicirculari, undo,
rugoso, fronte subtiliter rugosa, setosa ; prothorace globose, fusco-
squamoso, marginibus lateraliter atque postice griseis; scutello
fusco ; elytris fusco-squamosis, lineis 4 griseis, externa abbre-
viata ; pygidio dense flavido-squamoso ; prothoracis margine
postico ante scutelli angulos minutissime inciso ; scutello lato ;
elytris postice ad suturam baud angulatis.
(S . Tarsis intermediis sat brevibus, pedibus postiois validis, tro-
chanteribus longis, paulo spinosis.
Long. 7 mm.
Hah. Burma, Shan States.
This species is nearly related to D. Hoplia ”) marginatusy
Non fried, which its author has referred to a wrong group.
It is larger than that insect and in addition to the dark
margins of the elytra has three dark stripes of equal width
to the pale interspaces. The white scales upon the pronotum
are more restricted behind. I have seen only males of both
Ann. d) Mag. N. Hist. Ser. 7. Vol. xix. 25
358
Mr. G. J. Arrow on new
species, which sex, in each, has the hind trochanters very
long and spinose. In D, lineata^ but not in the allied species,
the middle tarsi are shortened and thickened also in the male.
Geotrupidae.
Ochodceus deceptor^ sp. n.
Rufo-testaceus, hreviter ovatus, corpore supra omnino sat dense
granulate, subtus laeviore, flavo-hirsuto ; clypeo parvo, semi-
circulari, maudibulis extus regulariter arcuatis ; prothorace
latissimo, margine lateral! recto, angulis anticis rectis, posticis
baud late arcuatis ; scutello elongate, apice acute ; elytris dis-
tincte striatis, interstitiis dense setosis.
Long. 5’5-6*5 mm.
(S . Tibiae anticae angulo interne producto, femore postico ante apicem
dentate.
2 . Capitis vertice tuberculis duobus geminatis armato.
Hah. S. India, Bombay, Madura, Kanara.
This species has a close resemblance to the European
0. chrysomelinus, F., but the sculpture of tlie whole upper
surface is rather less dense, the side margins of the prothorax
straighter, the posterior angles less broadly rounded off, and
the scutellum rather narrower and more acute at the apex.
The secondary sexual distinctions of the male are the same,
but the female is distinguished by a slight crest upon the
forehead consisting of two closely approximate tubercles.
These are absent in 0. chrysomelinus, which has instead a
single tubercle near the tip of the clypeus.
Ochodceus pallidusj sp. n.
Pallide testaceus, ovatus, capite prothoraceque minute et dense
rugosis, setiferis, elytris subnitidis, minute punctulatis et setiferis ;
clypeo brevissimo, lato ; mandibulis extus sinuatis, apice paulo
produetis, intus leviter dentatis ; prothorace lato, margine laterali
recto, angulis anticis fere acutis, posticis baud late arcuatis ;
scutello mediocre, baud acute angulato; tibiae anticae dente
apicali longo. Laud acuto, secundo sat acuto, tertio minutissimo.
Long. 6-5 mm.
Hah. N. India.
Two specimens, both females, were collected by Capt. Boys
and have been in the Museum since 1848. The species is
more elongate than the preceding one and the elytra are
paler and more shining, the sculpture being much less dense.
In addition the mandibles are more produced, the clypeus
shorter and broader and the scutellum less elongate. The
hind tibiae and tarsi are longer and slenderer and the second
tooth of the front tibiae more acute.
359
Coleoptera from the Indian Empire,
Westwood lias mentioned an Indian Ochodceus to which
the MS. name of 0. indianus was given by Reiche, but
wliioh the former did not consider distinct from 0. chryso-
melinus. This is probably one or other of the two species
here described.
Hybosoridae.
Phceochroops indicus^ sp. n.
Pyriformis, fuscus, undique longe rufo-hirtus, capite prothoracequo
densissime punctatis, illo angustato, margine regulariter arcuato ;
prothorace parvo, transverso, lateribus elev.atis, denticulatis,
regulariter arcuatis, angulis anticis productis, posticis fere rectis ;
scutello parvo, angiisto ; elytris valde convexis, postice fortiter
ampliatis, dense variolose punctatis, costis tribus modice distinctis;
corpora subtus subtiliter rugoso, metasterni medio pedibusque
politis ; tibiis anticis extus denticulatis, dentibus tribus validis
rectis, posticis longe ciliatis, singulo extus medio minute carinato.
Long. 11'5 mm.
Hah, S. India, Nilgiri Hills, Anamalai Hills.
Three specimens have been found by Mr. H. L. Andrewes.
They are dark olive-brown, clothed all over with long tawny
hairs. The head and prothorax are exceedingly densely,
and the elytra closely, covered with circular pits. The lower
surface is finely rugose, with the middle of the metasternum
alone Simooth and shining. The labrum is very prominent,
quadrate, and coarsely punctured and the clypeus parabolic.
The pronotum is very narrow in front and rather broader
behind, with the lateral margins raised, denticulate, and gently
and uniformly curved, and the posterior angles right angles.
The elytra are regularly ampliated from the shoulders and
have each three feebly raised cost^.
The genus Phceochroops ^ with two others, Phceochridius
and Pantolasius, have been placed by their authors, Candeze
and Lansberge, amongst the Trogidae, and excluded from the
Hybosoridae by the presence of five ventral segments only.
These forms, however, have certainly a much closer relation-
ship to Hyhosorus^ Chcetodus^ &c., than to Trox and appear to
me to exhibit no difference of any consequence in the abdomen.
A sixth segment is always visible laterally, although partially
hidden beneath the hind coxae, and Lacordaire has observed
that this is sometimes the case in Trox itself. The form of
the antennae and labrum, however, is very difierent to that
occurring in Trox^ and if it is considered desirable to retain
both families I think the Trogidae should be restricted by
removing the genera Liparochrus and Anaides to the
Hybosoridae, with which they have most in common.
3G0
Bihliographical Notice*
BIBLIOGRAPHICAL NOTICE.
Catalogue of the Lepidoptera 'Phalaenoe in the British Museum.
Vol. VI. {Catalogue of Noctuidce, Vol. III.) London : Printed
by Order of the Trustees. 8vo. Pp. xiv, 532. Pis. xcvi.-cvii. ;
text-figs. 172.
The present volume of Sir George Hampson’s great work is devoted
to the subfamily Cucullianse, and includes 692 species distributed
among 111 genera, many of which are described as new. While
the Noctuidse, as a rule, are smaller and less showy than the moths
classed under Sphinges and Bombyces, they are more interesting to
European entomologists, as a larger proportion inhabit temperate
climates. Under the Cucullianae, as here defined, we find Cucullia,
Cleophana., Calophasia, Aporophyla., Cloantha, Xylina ( = Calocampa,
auct.), AgriopiSy Dasypolia, Cosmuiy and other well-known Euro-
pean genera. A series of additional species (uncoloured) are re-
presented, with neuration &c., in the text-figures. Although many
of these moths are dull- coloured, yet the larvae of some of them
(e. g. those of Cucullia and Xylina) are extremely beautiful, those of
Cucullia being gregarious and feeding in clusters on Verhascum &c.
(the larva of Cucullia verhasci is figured on p. 2). They are a great
contrast to the dull brown subterranean larvae of the Agrotinae,
called “ Cut-worms ” by the Americans. The moths of the genus
Cucullia are popularly called “ Sharks ” in England. They some-
what resemble small Sphingidae in their long, narrow, pointed
wings, and fly over flowers at dusk in the same manner. Our
species are all light brown, grey, or whitish, but several of the
Continental and Siberian species are of a most beautiful green, and
others brilliant silvery white, none of which, however, are figured
in the work before us.
It is less than two years since the publication of vol. v., and
vol. vi. is the third volume of Noctuae, vols. i.-iii. having been
devoted to Arctiidae &c. Those who are best acquainted with what
catalogue and descriptive work means will know best how to
appreciate the energy and the industry of the Author, and least
inclined to find fault on account of any error or omission which
they may be able to discover. For our own part, we may say that
the previous high standard of the work seems to be fully maintained
in the present volume both as regards the descriptions and tables,
and we are glad to see that the larvae, when known, are also briefly
noticed.
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. 113. MAY 1907.
XLII. — Descriptions and Records of Bees. — XIII.
By T. D. A. Cockerell, University of Colorado.
Osmia viridiorj sp. n.
d .—Length fully 12 mm. ; anterior wing a little over 8 ;
width of abdomen 4.
Head very dark blue-green ; mesothorax and scutellum
yellow-green ; the rest of thorax black with a greenish
lustre; legs black, not at all metallic; abdomen shining
blue-green, dark but very decidedly green. Head and thorax
densely punctured, clothed above with copious long hair, that
of face white, but of sides of vertex and cheeks strongly
intermixed with black ; head normal, mandibles strongly
bidentate ; anterior edge of clypeus straight, very smooth and
shining ; clypeus otherwise dull and densely rugoso-punctate ;
antennae black, third joint with a fine reddish tomentum,
only noticed in certain lights ; flagellum not at all moniliform ;
thorax above with long creamy-white hair, not at all mixed
with black ; pleura with long hair, anteriorly dull white,
posteriorly black. Legs with black hair, but shining reddish
on inner side of anterior tarsi, long and dull white on anterior
femora behind, and some dull white at apex of middle femora
behind ; spurs and tarsi normal ; tegulae shining black.
Wings hyaline, the apical margin broadly pale brownish, and
a daik streak in marginal cell; first r. n. joining second s.rn.
Ann. d' J/ap. X. /list. Ser. 7. Vol. xix. 26
362 Mr. T. D. A. Cockerell — Descriptions and
twice as far from its base as second from its apex. Abdomen
parallel-sided, first segment with white hair (some black at
extreme sides) ; second with thin white hair and shorter
black hairs intermixed, especially at sides, but inconspicuous ;
remaining segments with black hair, but third with a little
silvery white on margin laterally ; sixth segment with a
broad shallow notch, seventh bidentate ; second and third
ventral segments shallowly emarginate, with a very little
shining reddish hair in the emargination.
Very close to 0. marginipennis^ Cresson, but distinguished
by the strongly green mesothorax and scutellum, the paler
hair of thorax above, the edge of clypeus not subsinuate,^^
and the apical margin of wings not so dark. It also much
resembles 0. cyaneonitens^ CklL, but differs thus: —
Smaller ; hair of pleura all white ; middle tibia with con-
spicuous white hair cyaneonitens.
Larger : hair of pleura partly black ; middle tibia with
black hair viridior.
Hah, Boulder, Colorado, May 20, 1906 [S. A, Rohwer),
Osmia olivacea, sp. n.
d. — Length about or nearly 10 mm.; anterior wing 7^ ;
width of abdomen 3j.
Olive-green, the sides of the thorax dark bhie-green ; hair
of head and thorax long and abundant, white on clypeus,
cheeks below, and lower part of pleura, but otherwise tawny,
with very long black hairs intermixed on thorax above,
vertex, cheeks, and a few at sides of face; head normal,
lower edge of clypeus straight, the bidentate mandibles with
their teeth strongly divergent ; hair of upper part of face
strongly tawny; antennae long, black; flagellum slightly
moniliform or crenulate ; vertex and mesothorax rough with
exceedingly dense punctures ; a little shining space bordering
anterior ocellus in front. Legs black, the hind coxse and
femora dark green, and a slight green tint on middle ones ;
anterior and middle femora with long pale hair behind, hind
femora with dusky hair ; hair of tibiae (except anterior ones
behind) dark ; hair on inner side of hind tarsi golden ; spurs
normal; tegulse green. Wings dusky, hyaline before the
broad apical area; first r. n. joining second s.m. about IJ
times as far from its base as second from its apex. Abdomen
shining olive-green, the hind margins of segments con-
colorous; first two segments with pale hair, the others with
black hair, quite long, and a little pale intermixed; sixth
363
Records of Dees.
with a good deal of light hair ; sixth segment entire or with
the faintest suggestion of a notch ; seventh bidentate ; first
ventral segment subemarginate ; third emarginate, with a
tuft of orange hair in the notch.
In Robertson^s table of genera segregated from Osmia this
runs to 3, and runs out because of the structure of the sixth
abdominal segment. Because of its coloration it could be
mistaken for 0. inurhana^ Cress., but it is easily known from
that by the coarse black hair on the abdomen &c. It is also
exceedingly like 0. Latreillei, Spinola (which I have from
Tangier), but is known from that by the same characters
which separate it from inurhana *.
Hah. Boulder, Colorado, at flowers of Pulsatilla hirsutis-
Sima, April 20, 1906 [W. P. Cockerell).
Osmia pulsatillcej sp. n.
d . — Length about mm. ; anterior wdng about 5 ; width
of abdomen
Green, the head and thorax above rather yellowish green,
the abdomen blue-green (Prussian green). Head normal,
eyes prominent, mandibles with two widely divergent teeth ;
face covered with dull w’hite hair, with some long black hairs
at sides and many long black hairs on front and vertex ; hair
of cheeks white, with black hairs immediately adjacent to
the eye. Antennae rather long, black ; flagellum normal ;
hair of thorax long and dull white, dorsally w'ith black
hairs intermixed ; mesothorax dull, with exceedingly dense
punctures ; tegulae largely green. Wings dusky ; first r. n.
joining second s.m. about 1§ as far from base as second from
apex. Legs dark green, with dull white hair, that on inner
side of hind basitarsus dark fuscous ; spurs normal. Abdo-
men with white hair on first three segments, long only on
first ; the other segments have the hair mixed black and
silvery, but short and inconspicuous ; sixth segment entire ;
seventh bidentate ; ventral segments normal, second apically
with fine white hair.
This is an Osmia s. str. in the sense of Robertson. It is
very like 0. proooima^ Cresson, but is readily distinguished
by the mixture of dark hairs on the thorax above. The
abdomen is not so globular as in proximo^ but is distinctly
parallel-sided.
Hah. Boulder, Colorado, prox. 5350 ft., at flowers of
Pulsatilla hirsutissima, April 20, 1906 (W. P. Cockerell).
* An Algerian male of 0. Latreillei, also before me, has the hair of
the thorax much redder.
26^
3G4
Mr. T. D. A. Cockerell — Descriptions and
Osmia aprilina^ sp. n.
. — Lengtli about 7 mm.; anterior wing about 5; width
of abdomen 2^.
Si ui liar to 0. jmlsatillm, but hair of thorax above pale
reddish, without black hairs interspersed. Head larger, much
wider above. Abdomen broader, more globose, and yellowish
green ; sixth segment with a conspicuous rounded notch.
The antennse are entirely dark ; hair of face pale, white
below, yellowish above, without any black ; vertex with an
admixture of black hairs ; cheeks with long black hairs in
front, but the hair otherwise white. Wings dusky. Legs
with green tints. Abdomen with pale hair, but a good deal
of black from middle of third segment on ; seventh segment
bi dentate.
Known from 0. proxima by the dark flagellum and dark
Iiairs on cheeks and vertex. Similar characters and the
structure of the sixth abdominal segment separate it from
0. pumila. It is an Osmia s. str. in the sense of Robertson.
Ilah. Boulder, Colorado, at flowers of Pulsatilla hirsutis-
stma, April 20, 1906 (IK. P.& T. D. A. Cockerell) ; Boulder,
May 1906 {S, A. Rohwer). Four specimens.
Osmia phacelice^ sp. n.
$ . — Length 8 mm. ; width of abdomen 3.
Blue-green, with yellowish-green tints about the head and
thorax. Legs black, the hind femora faintly greenish ; scopa
black, some white hairs at extreme sides. Hair of head and
thorax mainly white, but many coarse black hairs on face and
vertex and fewer on scutellum and mesothorax ; seen from
above, the hair of the clypeus appears short and black and
that on each side of face longer and nearly all silvery white ;
head and thorax very densely punctured ; head ordinary,
clypeus normal, mandibles 3-dentate, antennse short and
black ; hair of legs largely black, that on inner side of middle
and hind tarsi nigro-fuscous ; tegular largely green. Wings
dusky, first r. n. joining second s.m. at least three times as
far from base as second from apex. Abdomen shining blue-
green, hind margins of segments concolorous ; hair of second
segment wdiite but very short and scanty, with longer dark
hairs toward sides ; following segments with the usual dark
hairs, but the apical one, except at apex, with fine silvery
toraentum.
Distinguished from 0. faceta^ Cress., by the smaller size,
greei]cr abdomen, and colour of hair on cly[)eus. It looks
just like 0. atriventris^ but is easily separated by tlie lar^-e
amount of black hair on head. 0. alholateralis, Ckll., is
much larger and has not the fine appressed white hairs seen
on abdominal segments 4 and 5 \w i)haceUm. It is an Osmia
s. str. in the sense of Kobertson.
Hah. Ward, Colorado, July 1905, at flowers of Phacelia
( W. P. (jf* T. I). A. Ckll.) ; also one from Boulder, Colorado,
June 17, 1905 {W. P, Cockerell).
Osmia Hendersoni, sp. n.
$ . — Length 10 mm. ; width of abdomen 4^.
Robust ; head and thorax densely punctured, black, with
strong blue and green tints at sides of face, and the pleura,
scutellum, and metathorax greenish ; abdomen very short and
broad, very shiny, with sparse punctures, the segments olive-
green, with the hind margins broadly purple, their extreme
edge more or less reddish. Head rather large, with broad
cheeks, but not otherwise remarkable ; clypeus normal, the
anterior edge gently concave ; mandibles with three large
but short teeth ; antennse black ; hair of face entirely black ;
the black hair goes as far back as the ocelli, and behind this
the hair of the top of the head is all dull white, except at tlie
extreme sides ; cheeks nude ; hair of thorax above entirely
yellowish white, of pleura black ; tegulas rufo-piceous.
Wings hyaline with yellow stains; first r. n. joining second
s.m. about twice as far from base as second from apex. Legs
black, with black hair ; anterior tarsi with long pale hair ;
first abdominal segment with long white hair, the others with
rather long black hair ; sixth apically with a little reddish
appressed hair ; scopa black.
A beautiful species, separated from 0. nigrifrons by the
shining abdomen, the second segment with black hair, &c.
The 3-dentate mandibles and long hair of pleura readily
separate it from 0. hrevihirta.
Hah. Arapahoe Peak, Colorado, Sept. 1, 1906 (aS^. A.
Rohwer) .
Named after Judge J. Henderson, who was present when
it was caught, in recognition of his work on the Arapahoe
Glacier and other contributions to the natural history of
Colorado. The locality is high alpine.
Osmia hrevihirta^ sp. n.
$ . — Length about 11 mm. ; width of abdomen barely
over 4.
Dark blue, with green tints ; the abdomen brilliant, more
3G6 Mr. T. D. A. Cockerell — Descriptions and
shining than in 0. nigrifrons, the hind margins concolorous.
Head large ; clypeus normal ; mandibles 4-dentate, the
inner two teeth very short ; near the interval between the
first and second teeth is a little patch of shining orange hair ;
antennae black; hair of face black and dull white, the black
coarse and mainly on the clypeus and above, the white prin-
cipally at sides ; black hairs about ocelli, but hair on top of
head dull white ; cheeks with black hair, but anteriorly bare,
with a very little short white hair which glistens in certain
lights ; thorax above with yellowish-white hair, not mixed
with black ; hair of pleura short and scanty, dark, a little
light above posteriorly. Legs black, with black hair ;
tegulse black, with a blue spot. Wings broadly dusky on
apical margin; first r. n. joining second s.m. about 1| as far
from base as second from apex. Abdomen with light hair on
first segment and middle of second ; otherwise the hair is
black except that the apex of the last segment is covered
with chocolate-brown tomentum ; scopa black.
This may prove to be the female of 0, cyaneonitens, CklL,
which (cT) was also found at Boulder by my wife, June 4,
9, and 10, 1905.
Hah. Boulder, Colorado, June 10, 1905 {W. P, Cockerell).
Osmia Tiypochrysea Bohwerif subsp. n.
$ . — Length about 10 mm.
Differs from true hypocJirysea in being somewhat larger
and more robust, the patch of orange hair on the mandibles
little developed, the hair on inner side of hind tarsi dark
ferruginous (black in type). The clypeus is quadridentate
as in the type.
Hah. Boulder, Colorado, May 1906 {S. A. RoJiwer).
The locality is more than 2500 ft. lower than that of the
typical form,
Osmia hypoleuca^ sp. n. {pentstemonis^ subsp. ?).
$ . — Length about mm.; anterior wing 5 J ; width of
abdomen about 2^.
Head and thorax deep blue ; abdomen brilliant purple-
blue, the hind margins of the segments broadly olive-green ;
head and thorax very densely punctured, their hair long and
coarse, but not dense, black on head except on occiput and a
little on hindmost part of cheeks, dull white mixed with
black on thorax above, light on tubercles, black on upper,
but white on lower part of pleura. Head oblong, clypeus
367
Records of Bees,
normal, mandibles 3-dentate, antennse black ; tegulas piceous,
blue in front. Wings dusky ; first r. n. joining second s.m.
about or hardly 1^ times as far from base as second from
apex. Legs black, with black or sooty hair ; hind coxae and
femora strongly bluish, hind basitarsus flat and rather broad.
Abdomen with inconspicuous black hair on segments 3 to 5 ;
scopa black.
Perhaps only a variety of 0. pentstemoniSj Ckll., but that
has the hair of pleura all light, and the last dorsal abdominal
segment with fine whitish tomentum.
Hah. Boulder, Colorado, June 9, 1905, at flower no. 10
(W. P. Cockerell).
0. pentstemonis occurs at higher altitudes ; in Boulder
County we have taken it at Ward, about 9000 ft., at flowers
of Pentstemon, July 1905. In my original account of
0. pentstemonis I wrote : — It may prove to be the female of
0. Wheelerid^ It is, perhaps, significant that at Boulder,
June 4, 1905, my wife took a male Osmia which I liave
referred to 0. Whtehri^ CklL, variety.
Osmia pikeiy sp. n.
$ . — Length about 8 mm. ; width of abdomen 3.
A short broad species, with abdomen almost exactly circular
in outline. Dark greenish blue, the abdomen shining ; hair
of head and thorax long and coarse, black on clypeus, cheeks,
and pleura, dull white with black intermixed on front, vertex,
and thorax above j some white hair also on sides of face;
head and thorax densely punctured ; head rather large ;
mandibles with four prominent sharp teeth, the apical one
long ; clypeus normal, the two orange brushes below its
anterior edge well developed; antennae black, flagellum very
faintly reddish beneath ; scutellum and hind part of meso-
thorax yellowish green, contrasting with the blue-black
metathorax ; tegula3 black, greenish in front. W ings dusky ;
first r. n. joining second s.m. only a little further from base
than second from apex. Legs black, with black hair, fine
pale tomentum on underside of hind femora and tibia? ; hair
on underside of hind tarsi reddish except at sides. Abdomen
very sparsely punctured; first segment with pale hair,
second with pale and black, the others with black, the last
(except apically) with some appressed pale hairs; scopa
black.
Distinguished especially by its comparatively small size,
broad form, and 4-dentate mandibles. The colour of the
abdomen is the same as in 0. propinqua.
3G8 Mr. T. D. A. Cockerell — Descriptions and
Ilah, Halfway House, Pikers Peak, Colorado, at flowers of
Salix, May 30, 1904 {Cockerell).
The locality is 8400 ft. above sea-level.
Osmia senior j sp. n.
? . — Length about 12 mm. ; width of abdomen 3|.
Abdomen approximately parallel-sided, broadest about the
apex of the third segment. Head dark blue below, green
above; mesothorax and scutellum green, pleura dark blue.
Abdomen blue-green, the margins of segments concolorous,
but segments 2 and 3 have an indistinct pinkish-purple band
across the middle, while 4 and 5 show coppery-red tints.
Head and thorax densely punctured, head rather large ;
antennse black ; clypeus normal ; mandibles with four very
distinct teeth, the apical one not very long ; hair of clypeus
black, of sides of face dull white, of front mixed white and
black, of vertex black, of occiput pale, of cheeks pale behind
and black in front ; hair of thorax above scanty, dull white
with black intermixed, of pleura black, but of tubercles
abundant and light, conspicuously contrasting ; tegulse rufo-
piceous, shaded with green. Wings dusky, first r. n. joining
second s.m. about 1§ as far from base as second from apex.
Legs black, with black hair, the hind femora and coxae very
faintly metallic ; middle femur with a few pale hairs apically
beneath. Abdomen with light hair on first segment and
short black hair on the others, each with a scarcely noticeable
subapical band of shining pale hairs, mostly failing in the
middle ; sixth with fine pale tomentum ; scopa black.
0. gaillardice^ CklL, is allied, but larger, and without
appressed light hair on cheeks. The white hair at sides of
face separates 0. senior from 0. wardiana. By the colour
and shape of the abdomen 0. senior is like densa, but that
has light hair on pleura.
Hah. Boulder, Colorado, June 12, 1905 {W. P. Cockerell).
Osmia wardiana., sp. n.
$ . — Length a little over 12 mm; width of abdomen 4.
liobust ; abdomen oblong. Head and thorax black, the
sides of face and supraclypeal area green, the scutellum and
hind part of mesothorax greenish ; middle of mesothorax
purplish black, with an area in which the strong punctures
are separated, showing the shining ground; head large, the
cheeks very full and rounded ; antennae black ; clypeus
somewhat produced, with the apex shallowly subeniargiiiate ;
3G9
Records of Bees.
mandibles 4-dentate, the apical tooth much the largest, the
others subequal ; face with coarse black hair, cheeks with
quite long black hair ; vertex witii hair mixed black
and dull white ; thorax above with dull white hair, with
long black hairs intermixed; pleura and tubercles with
black hair ; tegulse with a large ferruginous spot. Wings
hyaline, with reddish stains; first r. n. joining second s.m.
about or scarcely times as far from base as second from
apex. Legs black, with black hair, that on inner side of
anterior tarsi reddish. Abdomen quite dark, with tints of
purple and green, the hind margins of the segments purple ;
hair on first segment long and pale, except a tuft of black
on each side toward base ; hair on remaining segments short
and black ; scopa black.
E-esembles O. rtigrifrons^ Cresson, but that has 3-dentate
mandibles, entirely black tegulae, &c.
Hah. Ward, Colorado, 9200 ft., at flowers of Senecio. July
{W.P.^ T. D. A. CM.).
Osmia Titusij Ckll.
Las Cruces, New Mexico, April 10, at flowers of Dithyrea
Wislizeni ; two females collected by Prof. C. H. T.
Townsend.
New to New Mexico.
Andrena Hitei^ sp. n.
$ .— Length rather more than 11 mm.; anterior wing
about 8^ ; width of abdomen 3J.
Black, the thorax and first four abdominal segments
covered dorsally with bright fox-red hair, exactly as in the
European A./ulva, of which it looks like a small example.
Cheeks and face below antennae with black hair, front and
vertex with dull reddish hair, paler and not so bright as that
of thorax ; antennae dark, scape with black hair ; facial foveae
very broad, not divergent from eye, dark, but with a pale
tint in certain lights ; clypeus shining, strongly but not very
densely punctured, with a broad smooth median band ;
process of labrum large, truncate, with sloping sides; third
antennal joint conspicuously longer than the next two united ;
mesothorax dull and granular, minutely punctured ; area of
metathorax triangular, dull and granular, small, defined only
by an impressed line ; hair of lower part of pleura black, of
metathorax red, but at sides are long, curled, black hairs ;
tegulge dark. Wings dusky, stigma ferruginous, nervures
370 Messrs. 0. Thomas and R. C. Wrougliton on
fuscous ; second s.m. about as broad as high, receiving first
r. n. almost at its end. Legs black, with black hair; spurs
clear ferruginous ; apical fimbria and hair of underside of
abdomen black j abdomen very feebly punctured, second
segment without any visibly depressed area.
Hob. Boulder, Colorado, May 21, 1906 {Glenn Hite).
A most unexpected find, like nothing I have seen from
America, but closely allied to the European A. falva.
Boulder, Colorado, U.S.A.,
Feb. 3, 1907.
XLTII. — New Mammals from Lake Chad and the Congo,
mostly from the Collections made during the Alexander-
Oosling Expedition. By Oldfield Thomas and B. C.
Weoughton.
It is hoped that a general account of the zoological collections
made during the recent Alexander-Gosling expedition from
Nigeria by way of Lake Chad to Shari, Oubangui, and
Welle to the Upper Nile will be published later, but in the
meanwhile such new mammals as we have been able to
distinguish are here described.
Both Capt. Gosling, whose death we have to deplore, and
Mr. Boyd Alexander took very great interest in the collec-
tion of mammals, and considering the difficulties inseparable
from such an expedition the number and condition of the
specimens obtained afford striking testimony to the pains
they took in this direction. As a result they have discovered
a remarkable number of new and interesting forms, one of
these, Colomys Goslingi, representing a new genus.
We have also included descriptions of a few other mammals
from the same area, their characteristics having become
evident during the work on the Alexander-Gosling specimens.
The whole of the collections obtained by the expedition
have been presented to the National Museum by Mr. Boyd
Alexander and the executors of Capt. Gosling.
Rhynchocyon Claudi, sp. n.
A white-tailed Rhynchocyon, with the usual pattern
brightly marked ; about the size of Stuhlmanni, Mats.
General ground-colour above a rather bright red-brown
(darker and less bright in the female), below a bright orange-
buff. A patch behind the ears, and the checks, a lighter
neio Mammals from Lake Chad and the Congo. 37 L
shade ; two dark (almost black) streaks from the shoulders
to the tail, broken at regular intervals, from the middle of
the back backwards, by whitish-buff spots; outside these on
each side two other dark stripes, not extending quite so far
forward as the median pair, broken by five and four pale
spots (the general effect is that of a rather regular chess-
board pattern on the lower back, rump, and haunches).
Individual hairs of the back, outside the pattern-area, grey at
their bases, then a bright buff (almost orange) with black
tips, the black tips varying very much in length, from two
thirds the whole length of the hair to 1 or 2 mm. only at the
distal end. Fore feet dark-coloured, clothed with short
hairs which are black at base and tip, with a median bright
buff ring ; hind feet still more sparsely clothed with even
shorter hairs of the same pattern. Tail at base black for a
short distance (30-35 mm.), then white and clothed with
sparse, very short, white hairs.
The measurements of the type specimen are as follows : —
Head and body 314 mm. ; tail 222; hind foot 80; ear 30.
Skull : greatest length 67 (circ.) ; length of nasals 19 ;
length of frontal suture 27*5 ; greatest breadth 35 ; inter-
orbital breadth 22 ; length of palate 35 ; length to
inclusive 20.
Hah. Beritio, Welle River.
Type. Old male. Original number 58. Collected Feb-
ruary 14th, 1906, by Mr. Boyd Alexander. (Four specimens
and a young skull examined.)
The present species is at once separable from all others
except Stuhlmanni by its white tail, and from that it differs
by its paler brighter colouring. In the type of Claudi, though
a very old animal, the chessboard pattern on the back is
brightly and clearly marked.
We have named this handsome species and the duiker
described further on in honour of Major Claud Alexander,
the leader of the Alexander-Gosling Expedition up to the
time of his death.
Erinaceus spicalus^ sp. n.
A medium-sized hedgehog, belonging to the group with
four toes, with the colouring and short slender spines of
alhiventris.
Spines short (15 mm.), slender and absent on the summit
of the crown along a band about 5 mm. broad.
Ear of medium size, smaller than in Adansoni from
Senegal, but larger than in the Soudanese alhiventris. Hind
372 ]\Iessrs. 0. Thomas and 1\. C. Wrougliton on
foot larger than in either of these species. The claws of the
toes (especially of the two inner) exceptionally long and
stout, even more so than in Adansoni ; that of the innermost
toe measures 7x2^ mm.
Skull in size as in Adansoni, but that of the latter much
stouter and broader : thus, in Adansoni the width of the
brain-case at the roots of the zygomata is 19*5 mm., against
18*3 in the present species; similarly the breadth at the
postorbital constriction and that across the zygomata are 11
and 30 mm., as compared with 10’3 and 26. In Adansoni
the sagittal crest is strongly defined and carried forward to
the middle of the frontals, while in spiculus it is much less
distinct and cannot be traced beyond the front edge of the
parietals. In Adansoni the nasals (15 mm.) are longer than
in spiculus (12‘5 mm.), but nevertheless the muzzle is
shorter ; the distance from the front of p^ to the front of i^
in the two species is 10 and 11 mm. respectively.
The following are measurements of the type (those of the
body taken in the flesh) : —
Head and body 220 mm. ; tail 10; hind foot 30 ; ear 20.
Skull : greatest length 44 ; palatal length 26 ; brain-case,
breadth 18'3 ; zygomatic breadth 26; length p^, and
10.
Hah, Maifoni, near Lake Chad.
Type. Old male (skin and skull). Original number 18.
Collected by Mr. Boyd Alexander on 17tli January, 1905.
(Three specimens examined.)
The nearest neighbours of spiculus are alhivent'ris, Wagn.,
from the Soudan and Adansoni, Bochebrune, from Senegal.
It resembles albiventris externally by its short fine spines,
but is distinguishable from it by its longer hind foot and
very much stronger toe-claws. In skull-characters it approxi-
mates to Adansoni and is easily distinguishable from albi-
ventris by ils markedly larger molars, broader para- and
mesopteiygoid fossas, and especially by the absence of the
frontal depression so marked in albiventris ; the points in
which it differs from Adansoni have already been noticed.
Poiana Pichardsoni ochracea, subsp. n.
A paler and more ochraceous form of P. Ricliardsoni.
General characters, so far as can be judged from a native
skin, as in P. lliclmrdsoni, but the spots are smaller and more
scattered, and the ground-colour is nearer “ clay-colour,’*’ but
verging slightly towards tawny. Under surface, instead of
being nearly white, of a bright buffy ochraceous colour.
new Mammals from Lake Chad and the Congo. 373
Limbs approximately clay-colour. Ground-colour of tail like
body, the dark rings rather narrower than in Richardsoni]
no trace of the fine intermediate dark rings generally present
between the broad ones.
Size apparently rather less than in true Richardsoni.
Hah. Near Yambuya, Aruwimi Kiver, Congo (about 25°
E., 1° N.).
Type. Native skin without skull. B.M. no. 7.1. 2. 4.
Obtained by E,. B. Woosnam.
This beautiful eastern representative of the West-African
Poiana Richardsoni may be readily recognized by the marked
difference in its general body-colour.
Crossarchus Alexandria sp. n.
Allied to C. ohscurusy but larger and with much longer
skull.
General appearance very much as in G. obscurus, the long
loose coat and grizzled colour as in that animal, tliough the
general tone is not so dark. Fur, when the animal is in full
pelage, very long, the hairs from 50 to 60 mm. in length; an
underfur of short woolly hairs present in some specimens.
General colour a coarse mixture of black and pale clay-colour,
the dorsal hairs pale clay-colour for their basal half (an
inconspicuous dark ring present about the middle of their
lower half), then broadly black, with a terminal or subterminal
band of dull whitish. Wool-hairs pale brown basally, lighter
terminally. Under surface similar, but the browner basal
part of the pelage more obvious. Crown and middle line of
face blackish brown, sometimes sharply contrasted with the
grey or buffy cheeks and sides of muzzle. Ears pale brown.
Lips and chin pale buffy. Upper surface of forearms, hands,
and feet black. Claws very large, the anterior longer than
the posterior. Tail long-liaired at base, evenly tapering,
mixed black and greyish like the body.
In some specimens, taken probably at a different season,
the long loose hairs are few or absent, and the whole body is
clothed in a short coat of greyish underfur.
Skull conspicuously different from that of C. obscurus by
its very much greater length, the breadth being about the
same. Teeth much longer throughout.
Dimensions of the type (measured on the skin) : —
Head and body 450 mm. ; tail (c.) 290 ; hind foot (s. u.) 77 ;
longest front claw (above) 14*5 ; longest hind claw 9.
Skull: condylo-basal length 81; basilar length 74*8;
greatest breadth 36*3 ; length of nasals in middle line 15*3 ;
374 Messrs. 0. Thomas and R. C. Wroughton on
interorbital breadth 15 ; mastoid breadth 31 ; palatal
length 44*5 ; breadth of palate outside 23*2 ; greatest
diameter of 7*1, of rn? 5*7, of 5*1.
Hah, Northern Congo. Type from Banzyville, Ubanghi.
Type obtained from a trader by Mr. Boyd Alexander. Five
specimens examined.
This remarkable species differs widely from any previously
known, its nearest ally, G. ohscurusy being only about two
thirds its size.
It is a curiously variable animal, both in colour and in the
condition of its pelage, though this latter may be a question
of season. A young example obtained by Emin Pasha in
Niam-Niam is absolutely without the long hairs characteristic
of two of the Alexander-Gosling specimens, while a third
is in an intermediate condition.
An example of Crossarchus DyhowsMiy Pousargues, was
also obtained by the Expedition.
Crossarchus Talhoti, sp. n.
A striped Crossarchus of medium size and very pale
coloration.
Hair of back short (15-20 mm.) and rather harsh.
The usual colour-pattern of the group; ground-colour a
dingy white, grizzled with black on the neck and shoulders,
and becoming pinkish buff'’"’ when alternating with black
in the stripes of the back.
Approximate dimensions : —
Head and body 450 mm.; tail 300.
Hah, Bornu, North Nigeria.
Type, Adult. B.M. no. 5,5.13.2. Collected by Mr. P.
A. Talbot and presented to the Natural History Museum.
A second specimen (young) which had been kept in
captivity by Capt. Gosling shows that the pale coloration is
quite normal and not an individual characteristic.
The species most resembling Talhoti is somalicusy Thos. ;
but in that species the dark stripes of the colour-pattern are
rather dark brown than black, while the paler are a red-
brown approaching hazel,” so that the pattern is obscured,
while in Talhoti the contrast between the black and pale bars
is strongly marked, though the general colour is even paler
than in somalicus. The short hair of Talhoti serves further
to distinguish it from the long-haired (40 mm.) somalicus^
and, lastly, it has a black tip to the tail which is wanting in
somalicus.
new Mammah from Lake Chad and the Congo. 375
Lycaon 'pictns aharicus., sp. n.
A medium-sized, bright-coloured Lycaon with very short
hair.
Size intermediate between typical pictus from Nyasa and
pictus lupinus from British East Africa.
Hair short (15 mm.), but not so short as in L. p. somalicus.
The hair of neck behind ears, above and below, lengthened
(*20-25 mm.), suggesting a ruff; that of distal § of tail long
(60 mm.).
General colour above ochraceous buff, somewhat irregularly
mottled with black and in the type specimen with some white
blotches on the back ; below mottled white, buff, and dark
brown, the hair of the belly, however, so short and sparse
that the mottled skin shows through, looking almost naked.
Muzzle black as far as the eyes, behind the eyes as far as
the ears buffy similar to the ground-colour of the back,
but very much less orange ; a black median line on crown,
extending along neck and indistinctly along back ; tail for
first third rich buffy, remainder black except for a con-
spicuous white patch almost immediately following the buffy
basal portion and for the white tip. Chin dark brown, almost
black. Anterior throat, corresponding with neck behind the
ears above, mottled black and white. All hairs the same
colour from base to tip.
Skull resembling that of typical pictus^ especially in the
narrowness between the orbits, but the brain-case distinctly
fuller than in that form ; pterygoid fossa narrow, greatest
width 22 mm. against 25 mm. in p. lupinus and 27 mm. in
typical pictus ] bullae smaller than in typical pictus^ about
as in lupinus. Teeth small, much smaller than in pictus^
smaller even than in lupinus.
The following are measurements : —
Head and body 1030 mm. ; tail 360 ; hind foot 212 ;
ear 110.
Skull; greatest length 207 ; basal length 178; length of
nasals diagonally 66 ; interorbital breadth 38 ; breadth of
brain-case 68 ; palate length 95 ; length of upper carnassial
19*5 ; greatest breadth of same 9 ; length of bullas 28.
Hah, Maui, Shari River.
Type, Adult female. Original number 36. Collected
lOlh May, 1905, by Capt. G. B. Gosling.
The colour-pattern is more nearly bilaterally symmetrical
than in any other specimen in the Museum, but probably this is
an indiviuual character. Its smaller size serves to distinguish
sharicus from typical pictus^ as its greater size does from
376 Messrs. 0. Thomas and R. C. Wrougliton on
hipinm, while its short fur differentiates it from all other
forms exeept pictm somalicus^ Tlios. This last form has still
shorter hair than sharicus and is very markedly smaller.
Mellivora concisa^ sp. n.
A medium-sized Mellivora with the usual white mantle
wanting on the rump and tail.
Size smaller than in typical ratel.
Hair of back 30 mm. long.
Colour-pattern as in most of the members of the genus,
i. e. black with a mantle of white extending from the forehead
to halfway down the tail, except that the white begins to die
out in the median line from about the middle of the back
posteriorly, disappearing completely on the rump and base of
the tail.
Skull not differing materially from that of ratel, except in
its markedly smaller size.
Measurements of the type (those of the body taken in the
flesh) : —
Head and body 580 mm. ; tail 153 ; hind foot 94; ear 19.
Skull : greatest length 135 ; basal length 125 ; breadth of
brain-case 60 ; interorbital breadth 33 ; palatal length 63 ;
length of upper carnassial 12’5 ; greatest breadth of
same 11*3.
Hah. Yo, Lake Chad.
Type. Adult male. Original number 62. Collected 7th
December, 1904, by Capt. G. B. Gosling.
The complete absence of white on the rump and base of the
tail suffices to distinguish M. concha at a glance from any other
form. Even should this modification of the usual colour-
pattern prove hereafter to be merely individual, which,
however, is most improbable, its marked inferiority in size
justifies its separation from typical ratel.
Funisciurus Alexandri, sp. n.
A small Fimisciurus of the same pattern as Emini, Stuhl.,
but much smaller.
Fur soft but short (8 mm. on the back).
General colour above a greenish yellow, approaching
'^olive-buff” of Ridgway ; from the shoulders to tlie base of
the tail a broad (5 mm.) median band, reddish orange in
colour, flanked on each side by a black band half its width,
and outside these again by a narrow pale yellowish band on
each side. Colour below the same as the ground-colour of
the back, but tinged with bright can:iry-ycllow, ospeeially on
new Mammals from Lake Chad and the Congo, 377
the throat and chest. The ground-colour of the back extends
over the whole head and face, except the ears, which are
conspicuously white. The hairs of the tail (about 15 mm.)
ringed pale and dark and pale tipped, except those at the
extreme tip, which are longer and broadly black tipped.
Skull, though smaller in all ways, shaped much as in
Emini ; interorbital area, however, proportionally much
broader and distinctly flatter, and postorbital processes very
much further back than in Emini,
The following are measurements of the type specimen, the
body-dimensions being those recorded by the collector : —
Head and body 110 mm. ; tail 115; hind foot 26; ear 13.
Skull : greatest length 30 ; basilar length 22 ; brain-
case 14’5 ; interorbital breadth 9 ; length of nasals 8*5 ;
upper tooth-row 5*3; true molars 3-7; bullae 7. (The
measurements of the second specimen scarcely differ from
these.)
Hab. Gudima, R. Iri, Upper Welle.
Type, An adult male. Original no. 122. Collected on
29th August, 1906, by Mr. Boyd Alexander, (Two speci-
mens examined.)
Its small size and conspicuous white ears suffice to distin-
guish Alexandri at a glance from any other species.
Funisciurus AntonicB, sp. n.
A very small striped species, like F, Alexandria but the
ears not white and the striping different.
Size slightly greater than in F, Alexandria with which it
forms a special section of the group characterized by small
size and the short blunt-nosed skull. General grouna-colour
pale greenish yellow, paler and greyer than in F, Bbhmi and
Alexandri, Striping of back exactly as in F, Bbhmi Emini,
i, e, with four distinct black stripes, but the outer narrow
pair only running halfway up the body from the loins.
Median pale stripe about 5 mm. broad, pale yellowish ; outer
light stripes 2|-3 mm. broad, yellowish white, whiter than
the median. Head of the general ground-colour, a dark
mark running through the eye, slight in front of it, distinct
behind it, succeeded below by a yellow line from whiskers to
ear ; upper eyelid also yellow. Ears of the general pale
greenish-yellow colour, not conspicuously white as in
F, Alexandri, Under surface broadly washed from chin to
anus with bright yellowish buff. Limbs yellowish grey
externally, buffy on their inner aspects. Tail slender, grizzled
black and pale yellow, as in the allied species.
Skull larger than that of F, Alexandri, but of the same
Ann, & Mag, N, Hist, Ser. 7. Vol, xix. 27
378 Messrs. 0. Thomas and Ji. C. Wrougliton on
general shape, much shorter than in F. Bohmi. Upper
ineisors very pale yellow in front.
])imensions ot the type (measured in the flesh) ; —
Head and body 104 rnm.; tail 126; hind foot 25; ear 12.
Skull: greatest length 31*5; basilar length 24*2; greatest
breadth 19*4; interorbital breadth 9*6; palatilar length 12*2 ;
length of upper tooth series (exclusive of the minute 5*1.
Hah. Ponthierville, above Stanley Falls, Upper Congo.
Alt. 2000'.
7ype. Adult female. Original number 351. Collected 21st
February, 1907, by Douglas Carruthers. Two specimens.
This beautiful little squirrel is no doubt most nearly allied
to F. Alexandri of the Welle, but in colour it almost exactly
mimics the much larger F. Bohmi Emini which occurs
with it.
Tatera lacustris, sp. n.
A Tatera of the Emini group, paler coloured than that
species and with smaller bullae.
Size about the same as Emini', fur long and fine, length
on the back 15 mm., against 10 mm. in Emini', general
colour above a rusty buff, pure on the flanks, much grizzled
with black on the back, below pure white. Basal three
fourths of dorsal hairs a silvery grey, tips buff, but in a
certain percentage of the hairs of the back the terminal
fourth is black. Cheeks, a patch above the eye and one
behind the ears, and the whole of the under surface and inner
sides of the limbs pure white. Tail above coloured like the
back, below a silvery buff ; hairs of terminal third length-
ened to form a tuft, which is markedly darker than the rest
of the body.
Skull almost as in Eminij but the nasals a shade longer
and narrower, giving the skull a lighter appearance, and the
interparietal more horizontal, making the skull slightly
longer. The bullae appreciably smaller than in Emini.
The following are measurements of some specimens : —
new Mammals from Lake Chad and the Congo, 379
Hah, Lake Chad.
Type, An adult male. Original number 14. Collected
by Capt. G. B. .Gosling on the 9th February, 1905. (Nine
specimens examined.)
Wroughton, in his key to the genus Tatera (Ann. & Mag.
Nat. Hist. xvii. 1906, p. 477), placed Emini by itself at the
end of the N.- African forms as the only species with a band
of hair across the sole of the foot. Since then Thomas has
described Harringtoni from Abyssinia (Ann. & Mag. Nat.
Hist, xviii., Oct. 1906, p. 303), which has the same character,
and the present is a third species to be now added to the
group. They may be fitted into Wroughton’s key as
follows : —
Sect. II. — B. h. Band of hair across sole of foot.
a^. Size larger (head and body 140 mm., hind foot
30). Colour darker ; lengthened hairs of tail
extending over more than half its length.
Length of skull 36 mm. ; upper molar series
5-2 ; hullse 10 4
A*. Size larger (head and body 120 mm., hind
foot 30). Colour paler ; lengthened hairs of
tail extending over distal third of tail only.
Length of skull 36 mm. ; upper molar series 5’6 ;
bullae 9
«*. Size smaller (head and body 100 mm., hind
foot 28). Colour darker; lengthened hairs
of tail extending over more than half its
length. Length of skull 30’7 mm. ; upper
molar series 4 ’3 ; bullae 9
COLOMYS^, gen. nov. (Alurince),
Like Malacomys and Deomys in external form, with simi-
larly elongated feet, but the skull wholly different from that
of either. Teeth strictly murine.
General form light and slender. Ears large. Forearms
thin; thumb with a somewhat elongated nail. Hind limbs
also very slender, the feet much lengthened, especially in the
metatarsal region ; metatarsi apparently somewhat loosely
bound together, as though they might splay out from each
other in walking on swampy soil. Tail of medium length,
very thinly haired, not pencilled.
Skull of medium proportions, not in the least elongate
as it is in Malacomys, Supraorbital and parietal regions
scarcely ridged. Anterior plate of zygoma-root not projected
forward, its front edge barely anterior to that of the
bridge above. Palatal foramina large and open, but on the
• KcoXop, a limb. The Greek term for stilts, KooXo^aOpov, is based on
this word.
Emini.
lacuatris.
Harringtoni.
27*
'^S0 Messrs. O. Thomas and C. Wrougliton on
posterior third of the septum between them there are a pair of
horizontal bony expansions projecting laterally and partly
closing the foramina below. Palation just a little behind the
back of level with the front edge of the parapterygoid
fossae ; these fossae are broad and well defined anteriorly, but
posteriorly the ectopterygoids soon become obsolete. Bullae
of medium proportions.
Incisors narrow, simple, flat or faintly concave in front, but
they cannot be said to be grooved. Molars essentially as in
Alus (comparison made with Al. rattus)^ but the pattern
rather more zigzagged, the median and lateral cusps higher
and the valleys between them deeper. No trace of postero-
internal secondary cusps, but on there is a small antero-
external as well as the usual large antero-internal cusp.
Cusps of lower molars very upright, not or but little slanted
forwards ; minute extra cusps present externally between the
second and third lamiiise of and the two laminae of also
mesially at the hinder edge of both and m2 ; m3 distinctly
bilaminate, the second lamina rather more than half the
breadth of the first.
Type. Colomys Goslingi^ sp. n.
This striking genus has clearly no real relationship to the
other long-footed forms, Malacomys and Deomys^ found in the
same region. Deomys is a member of the Dendrorayinae and
Malacomys has an elongated skull, dififerent in almost every
detail from that of Colomys.
We have therefore here a very interesting case of geogra-
phical isomorphism, three unrelated genera all showing the
same elongated metatarsi, and we trust that some observer
on the spot may be able to discover what special form of
surroundings has produced their common characteristic.
Colomys Gosliiigi^ sp. n.
About half the bulk of Mus rattus. Fur short, rather
crisp and velvety in texture, but not spiny ; hairs of back
7-8 mm. in length. General colour above between wood-
brown and cinnamon, a posterior dorsal area more blackened.
Under surface pure sharply defined white, the hairs white to
their bases ; line of demarcation very high on cheeks and
sides, fully halfway up the body, the white area taking in
the whole of the fore limbs, which are white throughout, but
the hind limbs have a narrow darker line running down
behind them to the ankles. Ears fairly large, practically
naked, grey ; a prominent white spot on the side of the head
below the notch. Upper surface of hands white, of feet
flesh-colour. Tail finely ^scaled, 12 scales to the centimetre.
new Mammals from Lake Chad and the Congo. 381
practically naked, the few hairs at the end about a milli-
metre in length, uniformly grey above and below.
Skull and teeth as described above.
Dimensions of the type (measured on the skin) : —
Head and body (c.) 140 mm.; tail 156; hind foot 38*5 ;
ear (c.) 17.
Skull : greatest length 32*5 ; basilar length 26 ; zygo-
matic breadth 15’8 ; nasals 12*2 x 4 ; interorbital breadth 4*8 ;
brain-case, breadth 14 ; interparietal 5 x 10*5 ; anterior
zygomatic plate 2*4 ; palatilar length 15 ; diastema 9*1 ;
palatal foramina 7*2 x 3’2 ; length of upper molar series 5*2.
Hab. Gambi, Welle R.^
Tyjpe. Adult male. Original number G. B. G. 92. Col-
lected 31st January, 1906, by Capt. G. B. Gosling. One
specimen only.
Monbuttu name Monbongolo.^’ — G. B. G.
We have named this handsome species, the type of a most
distinct new genus, in honour of its captor, Capt. Gosling,
whose untimely death has deprived science of one who took
an intense interest in the collection of Mammalia, and who we
had hoped would have taken up their study at home with the
same zeal that he devoted to their capture abroad.
Thamnomys kuru, sp. n.
A Thamnomys of the venustus-rutilans group, but markedly
smaller than either of those species.
Fur of back rather long (10 mm.) and silky. General
colour above dark brown, grizzled with yellowish buff, below
white, more or less stained with pinkish buff.
Whole upper surface of body a dark brown grizzled wdth
orange-buff (the hairs dark slate-colour, a large proportion,
almost all at the root of the tail, tipped with orange-buff, the
remainder with very dark brown, almost black), paling rather
suddenly at the sides into the white of the belly (the hairs of
which are white to their bases), a certain amount of orange-
or pinkish-buff suffusion modifying the white colour of the
under surface of the body, especially on the chest and on
inner side of limbs. Tail dark, regularly but sparsely clothed
on proximal half with very short brown hairs, thence distally
length of hairs increasing until at tip they measure 5-6 mm.
in length ; rings of tail 20-21 to the inch.
The skull much smaller than in rutilans ; comparing with
a specimen of that species of like age from the Cameroons
the skull is one fifth shorter and proportionally narrower and
the upper molar series is 5*5 mm. only, instead of 6’2 mm.
The first and second molars show the characteristic third
3S2 Messrs. 0. Thomas and R. C. VVh’ougliton on
postero-internal cusp quite as plainly as either venustus or
rutilans.
Dimensions of the type (taken on the skin) : —
Head and body 135 mm. ; tail 160 ; hind foot 24 ; ear 15.
Skull : greatest length 29 ; basilar length 23 ; greatest
breadth 15 ; nasals 10 ; interorbital breadth 5’6 ; brain-case
breadth 12*7 ; palatilar length 12*5 ; diastema 8 ; palatal
foramina 6’5 ; upper molar series 5*5.
Mr. Alexander has recorded the body-dimensions of
another specimen (skull missing), taken the same day and
obviously older, as follows : — Head and body 145 mm. ;
tail 200 ; hind foot 35 (? 25) ; ear 15. These measure-
ments agree fairly closely, but for the shorter ear, with those
of rutilans and venustus.
Hah. Angu, Welle River.
Type. Young adult female. Original number 87. Col-
lected by Capt. G. B. Gosling on the 30th January, 1906.
This species, though undoubtedly closely allied to venustus
and rutilansj and, except for the smaller ears and some small
differences of fur, texture, and colour, not easy to distinguish
externally, is readily separable by the smaller skull and
teeth.
Lophuromys major^ sp. n.
A large Lophuromys about the size of Ansorgei^ de Wint.,
but with the colouring of aquilus. True.
General colour above a very dark brown, minutely speckled
with cinnamon, merging without any sharply defined line
into ‘‘ vinaceous cinnamon ” below.
Hair of the back &c. rather short (8 mm.), shorter than in
Ansorgeiy rather harsher than in aquilus and much more so
than in Ansorgei ; basal ^ or f of each hair cinnamon, the
remainder black, with a subtermiiial cinnamon ring ; on the
belly the hairs of one colour from base to tip. Crown of head
and face coloured like the back ; ears and tail looking almost
naked to the unaided eye, but covered with minute black
hairs through which the skin-colour shows. Chin, throat,
and inner side of limbs coloured like the belly, but in a
rather darker shade.
Skull markedly larger than in any other species of Lophu-
romys except Ansorgeiy with which it closely agrees in size
and shape. Teeth as in Ansorgei.
Dimensions (those of the body measured in the flesh) : —
Head and body 145 mm. ; tail 72; hind foot 25; ear 15.
Skull: palatilar length 15 ; brain-case breadth 13 ; inter-
new ^lammah from Lake Chad and the Congo, 383
orbital breadth 7 ; length of nasals 14*7 ; upper molar tooth-
row 5‘5.
LJab, Bwanda, R. Ubanghi.
Type. Old male. Original no. 44. Collected bj Mr. Bojd
Alexander, 25th December, 1905.
The species at present known in this genus fall into three
well-marked colour-groups, viz. : — (1) general colour without
any speckling, either dark brown, as in sikapusi from the
West Coast and Ansorgei from Lake Victoria, or olive-grey,
as in Woosnami ; (2) general colour a dark brown, finely
speckled with dark butf or cinnamon, as in aquilus from
Kilimanjaro and the present form ; and (3) general colour a
dark brown, coarsely speckled with pale buff or yellow, as
in Jlavopunctatus from the extreme north-east of Africa. By
its colouring the present species falls in the aquilus group,
but is markedly larger than any form in that group, and
though it closely agrees in all dimensions with Ansorgei ^
it is sharply differentiated from all the sikapusi group by
its colouring.
Lophuromys laticeps, sp. n.
Near L. aquilus-, the brain-case broader and lower and
the palatal foramina shorter.
External characters very much as in L. aquilus, though
the speckling may be a tritie stronger. General colour of the
same warm vandyke-brown above and dull russet below. In
a young specimen the posterior back is unspeckled, as in
L. sikapusi, and the belly is more strongly tawny.
Skull shorter and broader than in L. aquilus, the brain-case
peculiarly broad, rounded and low, the height from bulla to
crown a millimetre or more less than in that species ; ridges
practically undeveloped behind postorbital processes ; ante-
rior root of zygoma projected forward as in aquilus, not
narrow and slanting as in L, Woosnami ; palatal foramina
unusually short, widely open.
Dimensions of the type (measured in flesh) : —
Head and body 108 mm. ; tail (broken, in an immature
specimen 54 mm.) ; hind foot 20 ; ear 18.
Ski^ll : greatest length 29 3; basilar length 24; zygo-
matic breadth 15*2; nasals, length 11*5; interorbital
breadth 6’3; breadth of brain-case 13‘7 ; height of brain-
case including bullae 10'6; palatilar length 12; palatal
foramina 6’2 ; length of upper molar series 5.
Jdab. Lake Kivu. Alt. 4900'.
384 Messrs. O. Thomas and R. C. Wroughton on
Type. Adult female in British Museum. Original num-
ber *318. Collected 29th November, 1906, by Douglas
Carruthers. A second (immature) specimen from the
Mfumbiro volcanoes.
This species is no doubt closely allied to tlie common
L, aquilus of British East Africa, but may be distinguished
by its differently shaped brain-case and shorter palatal
foramina.
Thryonomys Harrisoni, sp. n.
A Thryonomys of the gregorianus-Sclateri group, more
closely allied to the former ; the remarkable postorbital pro-
cesses so characteristic of the latter entirely absent.
Size, judging from the skulls, rather larger than in
gregorianus.
As compared with the skull of gregorianus that of Harrisoni
is markedly longer and narrower ; the frontal depressions less
marked ; postorbital processes even less developed ; the
lacrymal bone smaller and the anterior edge of malar broader,
so that the distance between these two bones is very markedly
smaller, scarcely more than one third the same distance in
gregorianus) the nasals longer and the portion of the pre-
maxillary flanking them narrower. The following measure-
ments taken on the type skulls of the two species demonstrate
these differences very clearly : —
Harrisoni. gregorianus.
Greatest length 91 82
Greatest breadth 52 54
Nasals, length 30 26'5
Frontals, length 25*5 23
Interorbital breadth 28’5 30
Greatest width of rostrum posteriorly, I ^4
Posterior breadth of nasals . 13 13
Distance on the orbit between malar I c.fr 9
and lacrymal \
Upper molar series 16 16
Hah. Loka, 60 miles S.W. of Fort Berkeley, Lado
District.
Type. Skull and imperfect skin. B.M. no. 4.9.28.2.
Collected and presented by Col. J. J. Harrison.
Lepus chadensis^ sp. n.
A very pale-coloured long-eared hare of rather large size.
Size rather larger than in cethiopicus^ Hawker ij &c.
neio Mammals from Lake Chad and the Congo. 385
Fur of back short (15 mm.), but soft and silky.
General colour above a drab-grey, with only a very faint
buffy suffusion on the back ; below pure white. Eye-ring
white ; exterior margin of ears white, inner margin fringed
with pale buff ; fringe at tip of ears black outside, pale buff
inside. Chin white ; neck, above to half the length of the
ears, below to the level of the fore legs, pinkish buff.” The
hairs of the back pale grey from their bases for half their
length, remaining half about equally black and pale buff.
Measurements of the type (taken in the flesh) : —
Head and body 434 mm. ; tail 115 ; hind foot 121 ; ear 97.
Hah. Kadde, Lake Chad.
Type. Adult male. Original number 19. Collected by
Mr. Boyd Alexander on 29th December, 1904.
The present species very closely resembles L. Hawkerij
Thos., from the Soudan, which, however, is smaller, darker,
and more sandy-coloured.
Cephalophus rufilatus ruhidtor^ subsp. n.
A Cephalophus related to rufilatus^ Gr., from Gambia, of
about the same size, but much redder in colour.
Fur rather long and harsh. General colour a dark red-
brown, near ‘‘burnt-sienna,^^ with a narrow median dorsal
patch much darker, with a slight bluish tinge. Below a paler
shade of the dorsal colour.
Colour-pattern as in rufilatus^ the dark dorsal area hardly
so sharply defined, the “ ochraceous buff ” ground-colour of
that species replaced by “ burnt-sienna,” and the “ smoke-
grey ” of the nape, dorsal patch, and feet by slate-grey.”
Chin and inner side of thighs white in both forms.
Skull entirely as in rufilatus, except that the bullae are
larger.
Dimensions of type : —
Head and body 800 mm. ; tail 115 ; hind foot 65; ear 65.
Skull : greatest length 155 ; basilar length 135 ; greatest
breadth 68 ; length of nasals 50 ; length of rostrum in front
of orbit 74 ; length of frontals 65; bullae 22’5.
Hah. Basin of the Upper Welle.
Type. Young adult male. Original number 65. Collected
by Mr. Boyd Alexander on 30th January, 1906.
A second specimen, obtained at the same time and place,
though a female and younger, is exactly like the type. Con-
sidering the great distance separating the habitat of this form
from that of rufilatus on the Gambia, it is perfectly clear that
3SG Messrs. O. Thomas and R. C. Wroughton on
it should be distinguished from that animal ; but as the
Museum contains a specimen (unfortunately quite young)
from Nigeria which seems to be intermediate between the
two, we prefer for the present to consider it merely as a
subspecies.
Cephalophus Claudi^ sp. n.
A Cephalophus allied to and about the size of nigrifrons^
Gray, from the Gaboon, but much darker and richer in
colour.
Fur as in mgrifrons. General colour above near “ burnt-
sienna ” ; only slightly paler on the haunches and belly.
Colour-pattern of the head and face as in nigrifronSj but
the actual colours darker; a strong suffusion of black on the
shoulders : tail for basal third of its length same colour as
back ; second third very sparsely covered wdth long whitish
hairs ; terminal third forming a w^ell-furnished black tuft, the
extreme tip with tendency to show white. Throat, belly,
and inner side of limbs only slightly paler than back ; the
belly W'ith a median dark stripe from the chest to the navel.
The fore legs blue ” (as in nigrifrons and many others),
but the body-colour extending down to the metatarsus, much
lower than in nigrifrons.
Skull : as compared with that of nigrifrons markedly
longer in front of the orbit, narrower between the orbits,
flatter on the forehead ; teeth markedly broader ; bullae
larger.
Dimensions of the type (taken in the skin) : —
Head and body 920 mm. ; tail 150; hind foot 210 ; ear 85.
Skull : greatest length 185 ; basilar length 170 ; greatest
breadth 77 ; length of rostrum in front of orbit 102 ; length
of nasals 75; length of frontals 70; interorbital breadth 38 ;
bull^ 27.
Bob. Bambili, Welle Basin.
Type, Adult female. Original number 118. Collected
by Capt. G. B. Gosling on 15th April, 1906.
Cephalophus rubidus, Thos., from Ruwenzori is, equally
with nigrifrons^ closely related to the present form. The
presence of a white chin-patch and the complete absence of
the dark mantle, belly-stripe, and colouring above the hock
suffice to distinguish it from Claudi. The skull-characters
indicated above as separating the present form from nigrifrons
differentiate it still more markedly from rubidus, as is shown
by the following comparative measurements : —
V
neio Mammals from Lake Chad and the Congo, 387
Oarehia GosUngi, sp. n.
An Ourehia of the size of hastata^ Peters, from Nyasa,
larger than kenyoe^ Meinertzh., of British East Africa, and
with longer horns than montana, Cretzschm., from the Soudan,
with a striking black patch on the forehead between the horns,
extending on to the base of the ears.
Colour-pattern and colouring as in other members of the
genus, but there is a distinct darkening of the back (even
blackish in the type specimen), and a well-marked black
patch on the forehead between the horns, extending on to the
bases of the ears, curiously recalling the somewhat similar
marking in the very distinct 0. oribi of S. Africa.
Skull only equalled in size by that of hastata ; distinguished
by a marked and characteristic convexity of the rostrum,
commencing in front of the frontal depression, and extending
to the unusually depressed tips of the nasals. Horns longer
than in montana^ but shorter and slighter than in kenyce.
Dimensions of the type : —
Head and body 940 mm.; tail 94; hind foot 300;
ear 107.
Skull: greatest length 182; basilar length 160; greatest
width 73; length of rostrum in front of orbit 98; length of
nasals 65 ; length of frontals 56 ; interorbital breadth 44 ;
bullae 20.
Hah. Niangara, Upper Welle Basin.
Type. Adult female. Original number 125. Collected
by Capt. G. B. Gosling on 9th June, 1906. (Examined two
complete specimens and one head-skin and skull.)
The frontal black patch of this species is a very distinctive
character, and is equally distinct in all three of the specimens
examined. In the Natural History Museum collection there
is a specimen of montana (?) from the White Nile which has
a distinct dark brown patch on the vertex ; this, however
does not extend on to the bases of the ears; in no other form
is a black patch like that of the present species to be found
except in the South African 0. oribi, a species geographi-
cally barred from any close affinity. The larger skull
differentiates 0. Goslingi from either montana or kenyce^
and from hastata it is separated as well by its geographical
position as by the peculiar profile of its skull and its black
frontal patch.
Prof. C. Cliilton on a
:]88
XLIV. — A new Freshwater Gammarid from New Zealand.
By Charles Chilton, M.A., D.Sc., F.L.S., Professor of
Biology, Canterbury College, New Zealand.
[Plate XL]
IbiROUGH tlie kindness of Mr. J. Crosby Smith, of Invercargill,
I am able to describe a new Gammarid from the surface
freshwaters of New Zealand. He obtained it from a small
pool near the top of Mt. Anglem in Stewart Island at a
height of about 2800 feet above sea-level. Unfortunately
only one specimen was obtained ; however, this is quite
sufficient for me to identify the animal satisfactorily, and
as it is undoubtedly different from any Gammarid hitherto
described from the freshwaters of New Zealand, I venture to
describe it as a new species. It is evidently pretty closely
allied to the subterranean species Phreatogammarus frag ills ^
and I am referring it to that genus, though it differs from
the definition of the genus as given by Mr. Stebbing in
having the first gnathopod distinctly smaller than the second
and of distinctly different form, for in his generic diagnosis
Mr. Stebbing says “ First and second gnathopods equal ”
His diagnosis was, however, drawn up from the single type
species, and where a genus is based on minute points of
difference as exhibited by a single species it is easy to restrict
the characters of a genus too narrowly. I give a brief
specific diagnosis and a fuller description of some points in
the single specimen I have examined.
Phreatogammarus propinquus, sp. n.
*(P1. XI. figs. 1-6.)
In general appearance, antennae, perseopoda, and uropoda
closely resembling P. fragilis, but differing in the gnathopoda.
First gnathopod smaller than second and of different form,
having carpus longer than propod, the latter widening distal ly
and with palm transverse. Second gnathopod with carpus
short, subtriangular ; pro[)od twice as long as carpus, ovoid ;
palm very oblique.
Colour nearly white.”
Length of body 5 mm.
Ilah. Small pool near top of Mt. Anglem, 2800 ft. above
sea, Stewart Island, New Zealand.
* ‘ Das Tierreich,’ Amphipodn, I. Gammaridea, p. 453.
new Freshwater Gammarid.
389
Bemarks, — I have given above the points that seem to
distinguish tliis species from P. fragilis. The following
fuller account is based on the single specimen before me : —
Body rather slender ; pleon-segments 4 to 6 with one or
two fine hair-like setae on dorsal surface, the fourth with a
stout spine on lower margin. Head without rostrum. First
antenna rather more than half as long as the body, first
joint with a tuft of small tactile setae on the upper margin
near the base, a few hair-like setae at the distal end, and a
small spine-like one on lower side of distal end ; second joint
about two thirds the length of first, with some slender setae
at extremity ; third joint about half as long as second ;
flagellum of about twenty joints, about twice as long as
peduncle ; accessory flagellum of four joints. Second an-
tenna about two thirds as long as first; flagellum shorter than
peduncle.
I have not examined the mouth-parts in detail, but they
appear to be closely similar to those of P . fragilis.
First gnathopod smaller than the second ; carpus longer
than propod, suboblong, posterior margin densely fringed
with setse, a few on the anterior margin ; propod widening
towards distal end, where its width is about equal to its
length ; palm transverse, well defined, 'with a few small spine-
like setae and some long hairs ; dactyl rather stout. Second
gnathopod with carpus short, triangular, produced on posterior
side into a rounded lobe bearing several fine setae ; propod
about twice as long as carpus, ovoid, narrowing distally, palm
very oblique, occupying two thirds length of posterior
margin, supplied with a double row of spine-like setae and a
few^ fine hairs ; dactyl rather stout, closely serrate on inner
margin. The third to fifth peraeopoda moderately long, but
not greatly increasing in length posteriorly, the last reaching
as far as the end of pleon ; in each the second joint (basos)
is moderately broad, about two thirds as broad as long.
First uropod with peduncle longer than the rami, its
upper margin with three small spines and a large one at
extremity ; rami equal, each with a few spine-like setse ;
second uropod similar, but with peduncle only as long as
rami ; third uropod extending much beyond the others,
peduncle only about one half as long as rami, which are equal,
not narrowing distally, and each with two groups of three
spines and a terminal tuft of setse. Lobes of telson with the
posterior margin rounded and bearing two or three fine
hairs.
When alive the animal was, Mr. Crosby Smith says,
390
Mr. G. A. Boulenger on
nearly white ” in colour ; whether it is blind or not I
cannot say for certain, but I can find no undoubted indications
of eyes in the specimen before me.
EXPLANATION OF PLATE XL
Phreatogammarus propinquus^ sp. n.
P'ig. 1. First antenna, X 60.
Fig. 2, Second antenna, X 60.
Fig. 3. First gnathopod, X 105.
Fig. 4. Second gnathopod, X 105.
Fig. 5. Fourth peraeopod, X 60.
Fig. 6. End of pleon with uropoda, X 60.
XLV. — On Barbus aureus, Cope, from Natal.
By G. A. Boulenger, F.R.S.
A FISH obtained by Dr. Alden Grout at Umvoti, near the
boundary between Natal and Zululaiid, was described by the
late Prof. E. D. Cope in 1869 (Tr. Amer. Philos. Soc. (2)
xiii. p. 406) under the name of Laheobarhus aureus. The
original description was so meagre as to make it impossible
to assign the species its position in the system. Having
recently had to describe several new Barbels from the eastern
parts of South Africa, I felt extremely anxious to know
something more of this Laheobarhus aureus^ the types of
which are preserved in the Museum of the Academy of
Natural Sciences, Philadelphia. In answer to an application
made through my friend Dr. A. Erwin Brown, I have been
favoured by the Curator of the Museum with the loan of one
of the types, from which I have drawn up the following
definition. The other specimen, which I have not seen, has
been compared by Mr. W. H. Fowler, who states that he can
detect no difference of any importance between the two.
The fish is not referable to the group named Laheobarhus
by Euppell. It has the thin lips and the trenchant lower jaw
which characterize the genus Capoeta as defined by Gunther ;
but its affinities are with L. Bowkeri^ Blgr., from Natal,
which has the edge of the lower jaw rounded and the lower
lip continuous across the chin. It affords a further instance
of the unsatisfactory arrangement of the species of this genus
according to the structure of the mouth and lips, to which I
have alluded on previous occasions when describing species
from Morocco, Abyssinia, East Africa, and the Transvaal,
Jfat Jfist 5.7 VoLXlXPlXI
Qias Chilton, del
Phreatogammarus propinquus sp.nov
library
Of THE
UNIVERSITY of ILLINOIS
Barbus aureus, Cope^from Natal, 391
which show remarkable agreement in all characters except
the mouth and lips.
Barbus aureus may be thus defined : —
Depth of body equal to length of head, 3§ times in total
length. Snout rounded, feebly projecting beyond the mouth,
\ length of head ; diameter of eye4J^ times in length of head,
interorbital width 3 times; mouth moderate, evenly curved,
its width 3J times in length of head ; lower jaw with a sharp
edge; lips thin, not extending across the chin; two pairs of
equal barbels, measuring diameter of eye. Dorsal III 8,
third ray not at all enlarged, articulated, smooth; the fin,
which is equally distant from the occiput and from the root
of the caudal fin, has the upper border concave, and its
longest ray measures f length of head. Anal III 5, longest
ray f length of head. Pectoral | length of head. Ventral
below^ origin of dorsal. Caudal deeply forked. Caudal
peduncle once and ^ as long as deep. Scales 37 ^1, 2| be-
tween lateral line and root of ventral, 16 round caudal
peduncle.
Total length 195 mm.
As stated above, B, aureus resembles B. Bowheri^ differing
in the structure of the mouth. In this respect it agrees with
B. rhodesianusj Blgr., another close ally, in which, however,
the barbels are only about half the diameter of the eye and
the scales are larger (30-32 ||, 2J-3, 12). B. marequensis ^
A. Smith, to which Cope regarded B. aureus as most nearly
allied, has the lower lip continuous, longer barbels, the last
simple ray of the dorsal strong and bony, and larger scales
(33 I 3, 12).
392
j\Ir. G. A. Boulenger on
XLVI. — Description of a new Gyprinid Fish of the Genus
Labeo from the Transvaal. By G. A. Boulenger, F.R.S.
Laheo Ruddi.
Body strongly compressed, its depth 3| to 4 times in total
length ; length of head 4 J to 4J times in total length.
Snout rounded, J length of head ; eye perfectly lateral, 4J
times in length of head ; interorbital width half length of
head ; mouth rather small, its width 3^ times in length of
head ; lips without transverse plicje, with a fringe of conical
papillae; rostral lobe not fringed; no barbels. Dorsal III 9-10,
upper edge concave, last simple ray as long as or a little
shorter than head ; its distance from caudal equals its
distance from anterior border of eye. Anal III 5. Pectoral
a little shorter than head. Ventral below middle of dorsal.
Caudal peduncle once and J to once and ^ as long as deep.
Scales 40-41 p, 5 between lateral line and ventral, 18-20
round caudal peduncle. Dark olive-brown above, whitish
beneath.
Total length 210 mm.
Four specimens from the Klein Letaba, tributary of the
Olifant River (Limpopo System), obtained along with
L. Rosce^ Stdr., and L. Darlingi, Blgr., by Mr. Claud Grant
(Rudd Expedition to S. Africa) in August 1905.
XLVII. — Descriptions of Tiuo new African Lizards of the
Genus Latastia. By G. A. Boulenger, F.R.S.
Latastia Johnstoni,
Head small, rather elongate ; snout obtusely pointed. Two
superposed postnasals ; frontal narrowed posteriorly, grooved
anteriorly ; two large supraoculais, with a few small shields
in front and behind and a series of granules between them
and the supraciliaries ; interparietal narrow, much longer
than broad, in contact with a small occipital ; a band-like
supratemporal ; a curved tympanic ; temporal scales granular,
smooth ; no auricular denticulation ; subocular bordering the
lip, between the fifth and sixth or sixth and seventh upper
labials. Gular scales smooth ; collar toothed, with 7 or 8
neia African Lizards.
393
sliieKls. Dorsal scales small, rliomboidal, feebly imbricate,
sharply keeled, 50 to 52 across the middle of the body.
Ventral plates in 6 longitudinal and 24 or 25 transverse
series; the })lates of the two median series nar:owest. Two
large piaeanals, one before the other, or three, one in front and
two behind. The hind limb reaches the shoulder or the neck.
15 or 16 femoral pores on each side. Upper cauJal scales
strongly keeled, basal subcaudals smooth. Four black
streaks along the back, and a fifth on the nape or on the
nape and greater part of the back ; sides of neck and body
with numerous irregular vertical black bars ; lower parts
uniform white.
Total length
Head
Width of head
From end of snout to fore limb
Fore limb
Hind limb
Tail ....
mm.
220
14
8
20
60
17
34
160
Two male specimens from the Nyika and Masuka Plateaus,
British Central Africa (alt. 6000-7000 feet), presented by
{Sir Harry Johnston, G.C.M.G., K.O.B., in 1897.
Laiastia Baril.
Head small, elongate ; snout acutely pointed. Two super-
posed postnasals ; frontal narrowed posteriorly, grooved
anteriorly ; two large supraoculars, with a few small shields
in front and behind, and a series of granules between them
and the siipraciliaries ; interparietal much longer than broad,
separated from the small occipital by a small shield ; four
supraternporals, first longest; a small tympanic; temporal
scales minute, granular, smooth ; no auricular denticulation ;
subocular bordering the lip, between the fifth and sixth or
sixth and seventh upper labials. Gular scales smooth ;
collar toothed, with 9 or 11 shields. Dorsal scales small,
hexagonal, juxtaposed, keeled, 42 to 45 across the middle of
the bod3^ Ventral plates in 6 or 8 longitudinal and 23 to 25
transverse series; the plates of the two median series a little
narrower than the adjacent ones. A large praeanal, with a
smaller one in front of it. The hind limb reaches bet.veen
the collar and the ear. 12 to 14 femoral pores on each side.
Upper caudal scales strongly keeled, basal subcaudals smooth.
Bluish grey in front, pale reddish brown behind ; three black
Ann. d; Mag. V. Hist. Ser. 7. Vol. xix. 28
On a new Frog from Brazil.
lines along tlie nape and two blaek streaks on each side of
the head and neck ; the median nnehal line and the two
lateral streaks conlinned on the bodv, fading to reddish
brown behind ; lower parts uniform white,
rnm.
Total leng’tli lo7
TTead...;
AVbdth of head
From end of snout to fore limb
Fore limb ,
TIind limb
Tail
Two male specimens from near Berbera, Somaliland (alt.
up to 400 feet), collected by ]\lr. G. W. Bury.
This species comes very near to L. Boscw, Bedr., which
differs in Itaving the dorsal scales smooth (except near the
tai!) and the dark lines in even number.
19
o2
10
. /)
XLYIir. — Descripfion of a new Frog of the Genus
Telmatobius/rom Brazil. By G. A. BOULENGER, F.R.S.
Ttlmatohius asper.
Vcmeiine teeth in two rounded groups behind the level of
the choarse. Head a little bia ader than long ; snout rounded,
longer than the ej’e ; no canthus rostralis ; nostril nearer the
end of the snout than the eye ; interorbital space a little
broader than the upper eyelid ; no tympanum. Fingers
moderate, witli slightly swollen tips, first not extending quite
as far as second ; toes with swollen tips, nearly entirely
webbed; subarticular tubercles well developed, flat; an oval
inner and a rounded outer metatarsal tubercle. The tibio-
tarsal articulation reaches the eye. ^kin of upper parts
closely studded with small warts, each bearing a pearl- like
horny tubercle; lower parts smooth. Blackish brown above,
with or without large yellowish blotches on the back and a
cross-bar between the upper eyelids; limbs with yrllowish
cross-bars; lower parts brown. Male with an internal vocal
sac.
From snout to vent 50 mm.
Four i^pecimens from Theresopolis, Santa Catharina,
collected by Mr. J. MichaGis.
]\Ir. W. L. Distant on Ilomoptera.
395
XLIX. — Rhynchotal Notes. — XLII. By W. L. Distant.
Fam. Fulgoridse (continued from p. 295).
Subfam. Derbinjs.
During the preparation of these pages my attention was
drawn to a ‘Report of Work of the Experiment Station of
the Hawaiian Sugar Planters’ Association/ Bull. No. 1,
pt. 9 (Honolulu, 1906). This part is entitled “ Leaf- Hoppers
and their Natural Enemies,” by G. W. Kirkaldy, but in
fact consists very largely, if not principally, of descriptions
of Australian species of Fulgorida3, with some figures and
many indications of new genera. It is singular that, as most
of the species described in this fugitive report are from
Queensland, there should be so little identity with the
species of Derbinm from the same locality described in this
paper. I have, however, carefully studied Kirkaldy’s indica-
tions of his new genera : those he has figured cause little
difficulty ; but those without illustration are quite different,
as in most instances he has scarcely referred to the tegmina
and wings, so important in this subfamily, and thus his
publication is left in a rudimentary condition. The figures
here given will, however, prevent further confusion, and if
any synonymy is found to exist, which I doubt, it may induce
Mr. Kirkaldy in future to amplify his generic indications.
Genus Dekbe.
Derhe, Fabr. Sjst. Rliyng. p. 80 (1803).
Type, D. hcjeniorrhoidalis, Fabr.
Dtrhe longitadi nails., sp. n.
Body pale brownish ochraceous, above with a broad central
longitudinal piceous fascia occupying the whole of vertex of
head and continued to apex of abdomen ; face and clypeus
fuscous brown; body beneath and legs pale ochraceous;
abdomen above with the lateral margins and apex fuscous
brown ; tegmina and wings pale luteous, the veins fuscous
brown, on basal half of tegmen there are also some inter-
mediate longitudinal fuscous-brown streaks, of which the
most prominent are in the subcostal, radial, and claval areas ;
face concave, with a central incised line, the lateral margins
strongly ridged ; clypeus centrally and laterally carinate ;
mesonoturn tricarinate ; pronotum centrally carinate : scutel-
lum strongly and broadly centrally sulcate.
28*
39G ]\lr. W. L. Distant on Ilomoptera.
Long., excl. tegm., min. ; exp. tegm. 26 to 2S mm.
Hah. Bolivia (J. Sleinhach, Brit. Mus.) ; Ecuador;
(yficliabd (Rosp.yiherg , Brit. Mus.).
In tlie smaller and varietal specimen from Ecuador tlie
subapical area to tlie tegmina and the apex of the wings
are also fuscous brown.
Genus Mysidia.
Mysidia, Westw. Trans. Linn. Soc. xix. p. 5 (1842).
Type, M. ixdlida, Fabr.
Mijsidia nehidosa.
Derhe nehulosa, Germ, in Then, Ent. Arch. ii. 2, p. >56 (18-30).
Mysidia nehulosa, Walk. List Horn., Suppl. p. 97 (1858).
Walker {supra), evidently with a rio-ht determination,
described this species as new. Fowler (Biol. Centr.-Am.,
Bhynch. Horn, ii, p. 73) has followed Walker.
Mysidia Steinhachij sp. n.
Body and legs pale stramineous; tegmina and wings
creamy white, subhyaline, the first with a fuscous spot
extending from costa at about one third from base across
radial area, between this spot and apex is another very pale
brownish transverse costal spot not reaching middle of
tegmen, some of the discal transverse veins also of the same
colour ; wings with a pale fuscous costal spot at about one
third from base which crosses radial area, the discal transverse
vein pale brownish ; face narrow, broadened towards clypeus,
lateral margins strongly ridged ; clypeus not or very
obsoletel}^ carinate.
Long., excl. tegm , ,3 mm. ; exp. tegm. 17 mm.
flah. Bolivia (J. Sleinhach, Brit. Mus.).
From the description apparently allied io 1\[. y^nneta, Fabr.
]\hjsidia jamaicensis, sp. n.
Body and legs very pale luteous; tegmina and wings pale
creamy, semiopaque, the venation darker, two black s[)ots
above clavus, the smallest near its base, the largest near i(s
a])ex, two black subapical spots, the uppermost smallest, two
black dots in basal third of costal area, remaining costal area
a little darker with pale spots; wings with a transverse
series of three small irregular spots at about one third from
base.
Mr. W. L. Distant on Ilomoptera. 31)7
Ijong’., excl. tegin., 3 mm.; exp. tegra. 13 mm.
JIah. Jamaica; Moneague (Cruise of ‘Valhalla/ M. J.
JSicoll, Brit. M'us.).
Mysidia glauca, sp. n.
Head and pronotum very pale brownish ochraceous ;
sternum with sanguineous markings ; body beneath and legs
very pale brownish, somewhat greyishly tomentose ; abdomen
above pale bluish; tegmina and wings pale bluish, in some
lights with a bronzy tint ; mesonotum distinctly tricarinate ;
tace and clypeus centrally sanguineous, the lateral margins
ridged.
Long., excl. tegm.^ 3 mm. ; exp. tegm. 15 mm.
Hah. Lower Amazons; Parana de Buyassu [E.E. Austen^
Brit. Mus.).
Genus Phenice.
Fhenice, Westw. Trans. Linn. Soc. xix. p. 10 (1842),
Type, P. fritillanSj Boh.
Phenice australis^ sp. n.
Head and pronotum dark chocolate-brown, the carinations
greyish white ; metaiiotum and scutellutn greyish white ;
abdomen above black, with a central longitudinal greyish-
white ridge ; head and sternum beneath dark chocolate-brown ;
abdomen beneath black ; legs stramineous ; tegmina black,
a large claval spot extending upward, but not reaching radial
vein, a large irregular spot beyond middle extending from
radial vein to inner margin, a series of spots to costal area,
becoming duplicated near apex, and apical marginal spots
greyish white, in the large pale spot beyond middle, but not
in the claval spot, the veins are black ; wings piceous brown,
posterior margin broadly greyish white divided by piceous
veins.
Long., excl. tegm., 3 mm.; exp. tegm. 17 mm.
Hah. Queensland [E. P. Dodd.^ Brit. Mus.).
Allied to the Indian species P, moesta^ Westw.
Akfaka, gen. nov.
Head (including eyes) very much narrower than pronotum ;
vertex narrow, triangular, widened posteriorly, distinctly
pioduced in front of eyes, centrally longitudinally deflected ;
lace laterally compressed, convex, centrally very narrow,
linear, a little widened anteriorly, centrally longitudinally
398
Mr. W. L. Distant on Ilomoptcra.
narrowly silicate; clypens about as lon^ as face, tricarinate ;
antemia3 inserted beneath eyes, second joint moderately long
and incrassate ; ])ronotum short and strongly laterally emar-
ginate posteriorly, strongly centrally carinate ; meson otum
long, tricarinate, anteiiorly somewhat strongly narrowed ;
legs moderately long and slender, posterior tarsi with the
basal joint long and moderately thickened ; tegmina broadened
before middle, a little convexly narrowed to apex, which is
truncate, costal area with five transverse veins, upjier apical
area with four large cells, radial area witli a transverse vein
)iear middle, and a broken series of discal transverse veins in
longitudinal sequence ; wings narrow, less than half the
length of tegmina, a transverse vein before middle.
Allied to Pamendanga, Dist., but differing by the narrow
face, longer clypeus, different shape of the mesonotum, &c.
Type, A. decisa^ Dist.
At f aka decisa, sp. n.
decisa, "Walker, M8.
Body and legs stramineous ; vertex of head, pronotum,
and carinations and posterior margin to mesonotum, greyish
white; on each side of pronotum, behind eyes, a testaceous
spot ; eyes brownish testaceous ; apices of the tarsi black ;
tegmina and wings pale hyaline, with the venation prominent
and black.
Long., excl. tegm., 4 mm.; exp. tegm. 16 mm.
Hah, New Guinea {Wallace^ Brit. Mus.).
A specimen of this species in the British Museum is labelled
d(ci'sa in Walkei-’s handwriting, but I cannot trace any
publication of a description.
SiKAiANA, gen. nov.
Head (including eyes) very much narrower than pronotum ;
vertex narrow, triangular, widened posteriorly, its apex a
little prominent; face extremely narrow and linear, appearing
as a line only between the eyes ; clypeus about half the length
of face, centrally and laterally carinate; antenna3 inserted
beneath eyes, very robust, second joint longer than third ;
])ronoturn short, very broadly transverse, very strongly
centrally carinate; mesonotum tricarinate; legs moderately
long and slender; tegmina broadened towards middle, a little
convexly narrowed to apex which is rounded, costal membrane
with some indistinct transverse ^eins, radial area short, dis-
coidal areas four, aj)ical areas six, a series of transverse veins
Mr. W. L. Distant on IIomop!era. 39D
roiintl apical margin ; wings mucli less tlian half the lengtli
of tegmina, discoidal areas long and narrow, a transverse
vein near middle of anal area ; apical areas four.
Allied to Pamendangn ^ DIst.
'^I’yi^e, S. hyalinata^ Dist.
]ly tlie shape of the pronotum this genus seems to be allied
to Basil eocephal us, Kirk., but the antennae are not short
as described in that genus, nor are the lateral areas of the
pronotum “ foliaceous, recurved, practically enclosing the
antennie.’'’ The only reference to the tegmina and wings
given by Kiikaldy is that the first are elongate, narrow,”
and the second are not mentione 1. It is therefore impossible
to locate the po^^itioti of Basileoceplialus in the Derbinse.
Sikaiana liyalinata, sp. n.
Body and legs very pale tawny yellow, slightly greyishly
tomentose ; eyes black; tegmina and wings pale hyaline, tlie
venation darker or more ochraceous; tegmina with some pale
sanguineous spots on costal area, becoming more numerous
Fig. 1.
near apex, Inner and apical margins a little clouded with pale
fuscous; wings with the apical veins and the transverse vein
on anal area very slightly and palely fuscous.
Long., excl. tegm., 2 mm. ; exp. tegm. llj mm.
llah. Queensland {k\ P, Dodd, Brit. Mus.j.
Sikaiana maculosa, sp. n.
Head with the margins of vertex greyish white, the eyes
dark castaneous ; pronotum veiy } ale tawny; rnesonotum
pale tawny brown, the carinations gieyish; abdomen abjve
400
Mr. W. L. Distant on ITomoptera.
and beneatli ocliraceons, with some castaneous suffusions,
especially on the lateral areas ; sternum and legs stramitieous ;
tegmina and wings pale hyaline with some opaline reflections ;
tegmina with the base narrowly pale ochraceous, terminating
in a subbasal fuscous spot on costal area, a somewhat large
fuscous spot before apex of costal area, another near apex, a
series of smaller fuscous spots on apical margin, and a still
smaller spot on upper vein of radial area ; wings with some
subapical marginal spots; antennae robust, ochraceous, third
joint infuscate.
Long., excl. tegm.y 2 mm. ; exp. 10 mm.
Hah. Queensland [F. P. Dodd, Brit. Mus.).
Genus ZORATDA.
Thracin, Westw, Trans. Linn. Soc. xix. p. 10 (1842), nom. prqeocc.
Zoraida, Kiik. Entomologist, 1900, p. 242, n. nom.
Type, Z. sinuosa, Westw.
Zoraida nivifera.
Thracia nimfera, Walk. Journ. Linn. Soc., Zool. x. p. 137 (1867).
Thracia abrujda^ Walk. loc. cit.
Hah. Batchian, Gilolo.
Zoraida costalis.
Thracia costalis, Walk. Jonrn. Linn. Soc., Zool. x. p. 136 (1867).
Thracia dorsalis, Walk. loc. cit. p. 137.
Hah. Batchian, New Guinea.
Zoraida scutellaris.
Thracia scutellaris. Walk. Journ. Linn. Soc., Zool. x. p. 135 (1867).
Thracia anticalis, Walk. loc. cit. p. 136.
Thracia sea notata, Walk. loc. cit. p. 139.
Hah. Morty, Mysol, Aru.
Zoraida Walkeri, n. nom.
Thracia nervosa, AValk. Journ. Linn. Soc., Zool. x. p. 135 (1867), nom.
piaeocc.
Hah. Mysol, New Guinea.
Zoraida horneensis, sp. n.
Head, pronotum, mesonotum, body beneath, and legs
stramineous ; abdomen above brownish ochraceous, it and
401
Mr. W. L. Distant on llomoptera.
the scutellnm more or less eretaeeously sericeous ; teg-
miiia hyaline, basal third suffused witli ochraceous, the costal
area creamy white containing a few brown spots, the sub-
costal area beyond middle is mostly black, with the veins
defining it sanguineous, and beneath the lower vein the black
is continued in some prominent angulations, inner and apical
margin with small piceous spots at the* apices of the veins,
the apex has also a subapical transverse line of small spots,
most of the transverse veins fuscous ; mesonotum prominently
tricarinate, the central carination continued through pro-
noturn ; clypeus centrally and laterally carinate.
Long., excl. tegm., 5 to 5^ mm. ; exp. tegm. 28 mm.
Hah, Borneo ; Sandakan {W. B. Fryer ^ Brit. Mus.),
Zoraida erythractiSj sp. n.
Body and legs dark ochraceous ; vertex of head, lateral
margins of pro- and mesonota, abdominal anal segment,
longitudinal streaks to femora and sternum, and abdomen
beneath sanguineous ; tegmina hyaline, crossed by three
broad pale fuscous fasciae, the first about one third from base,
the second just beyond middle, and the third at apex; in
these dark fasciae the veins are red, remaining venation
yellowish; costal area yellowish, reddish from base to first
dark fascia and from second dark fascia to apex ; wings
hyaline, with the venation darker; second joint of antennm
reddish, longer than head and pronotum together; nieso-
notura distinctly tricarinate, the central carination continued
through the pronotum.
Long., excl. tegm., 5 mrn.; exp. tegm. 25 mm.
Hub. Borneo; Kuching.
Zoraida cycnoptera^ sp. n.
Pronotum, mesonotum, scutellum, body beneatli, and lec^s
stramineous; abdomen above, posterior disk of pronotum
vertex of head, face, and clypeus testaceous ; tegmina hyaline,
tlie venation fuscous, costal area for about one third from
base fuscous brown, with creamy-white spots, remainino’ area
ochraceous, with a prominent black spot before apex, costal
and subcostal veins testaceous red, upper apical area piceous
apical inargiii and extremities of apical veins testaceous red
in S in $ the tvvo central extremities stramineous ; wino-s
hyaline, the venation fuscous; mesonotum tricarinate, the
central carination continued through pronotum ; clypeus
centrally and laterally carinate.
402
Mr. W. L. Distant on llomoptera.
Long., excl. tegm., ^ 4^^ ? G mm. ; exp. tegm., ^ 24,
? oO mm.
JIah. Queensland {F. P. Dodd^ Brit. Mus.).
Allied to Z. Essingtoni, Westw.
Zoraida eupoecila, sp. n.
Head, antenna^ pronotum, mesonotam, scntellum, and
abdomen tawny brown ; sternum and legs stramineous ;
carinations to pronotum and m.esonotum distinctly paler and
pale ocliraceous in hue ; eyes black ; tegmina very pale
fuliginous, subliyaline, infuscated on basal area, wliere there
is a distinct white spot beneath radial area, veins fuscous,
costal area greyish white, with an elongate fuscous spot near
middle and another nearer apical area; beneath the costal
area the colour is narrowly fuscous, broken with a large
creamy- white spot just before the second dark spot in costal
area, costal margin at apex testaceous, and the apical margin
creamy white, before the latter a transverse series of four
small fuscous spots placed on the veins, a large fuscous spot
at end of clavus; wings very pale fuliginous, with the veins
tu'cous ; second joint of antennjc robust, as long as head and
thorax together.
Long., excl. tegm., 5 mm.; exp. tf^gm. 22 mm.
Hah. Queensland; Kuranda [F. P. Fodd, Brit. Mas.).
Zoraida consanguineay sp. n.
Allied to Z. eiipoecilay and differing principally by the
tegmina, hich are very much more broadly and regularly
fuscous beneath the costal area, the latter creamy white, witli
three elongate fuscous spots on basal third, subapical trans-
verse spots obsolete, veins todiscoidal areas more transversely
and less longitudinally oblique.
T.ong., excl. tegm., 4 mm. ; exp. tegm. 18-23 mm. '
llah. Queensland (A. P. Dodd, Brit. Mus.).
Zoraida cydistay sp. n.
Head (including eyes) ferruginous brown ; thorax above
stramineous, mesonotum a little transversely piceous near
base; scutellum cretaceously white; abdomen above testa-
ceous, basal half of disk cretaceously white; body beneath
and legs stramineous; tegmina hyaline, venation fuscous;
costa, costal and subcostal veins purplish red, beyond middle
tlie spaces between these veins are ])iceous ; beneath and
attached to the lower red veins some })iceoiis spots, of vvdiich
403
^Ir. W. L. Distant on TTomopteva.
tlie most prominent are one about middle and another nearer
apex, apical marg’in between the veins piceous ; wings hyaline,
the venation fuscous; antennae with the second joint stra-
mineous, its apex ocliraceous, as long as head ami thorax
together ; mesonotum distinctly tricarinate, the carinations
very slightly paler.
I.orig., excl. tegm., 3 mm.; exp. tegm. 19-21 mm.
Hub. Queensland {F. P. Dodd^ Brit. iMus.).
Zoraidi picta^ sp. n.
Above and beneath pale ocliraceous ; eyes black ; pro-
notum stramineous ; abdomen above with angulate fasciate
sanguineous markings and with a curved black discal line on
each side a little beyond middle ; tegmina hyaline, the veins
fuscous, costal area pale testaceous, an oblique black spot in
subcostal area at about two thirds from base and a black
spot at its apex, the veins emanating from the subcostal vein
black at their bases ; mesonotum faintly tricarinate ; second
joint of antennse minutely black-speckled, with its apex
dai’ker ocliraceous, as long as head and thorax together.
Long., excl. tegm., 4^ mni. ; exp. tegm. 21: mni.
Huh. Queensland [F. P. Dodd^ Brit. Mus.).
Genus DroNA.
Drona, Dist. Faun. B. I., Rhyncli. hi. p. 305 (1906).
Type, Jd. carnosuy Weslw.
In describing this genus, founded on the type of Westwood’s
carnosa contained in the British Museum, and a somewhat
antique specimen, I could not see any carinations to the
mesonotum, nor could the artist who drew the figure. ISince
then I have been able to examine specimens in better con-
dition, and the mesonotum niUFt be described as tricarinate,
even if in some examples faintly so. The male anal armature
is a very prominent character.
Drona lanius.
Derhe lanius, Stal, Ofv'. Vet -Ak. Forli. 1855, p. 94.
Thracia lanius, Still, Hem. Atf. iv. p. 195 (I860).
llah. Caffraria.
Drona opicalis.
Thracia apicalis, Hagl. Ofv. Vet.-Ak. Forh. 1899, p. C4.
404
]\lr. W. L. Distant on llomoptera.
Drona Graliaml^ sp. n.
Docly and legs testaceous red ; tegmina subliyaline, sliglitly
fuliginous, veins a little darker, costal margin and subcostal
vein piceous ; costal area and stigma tlavescent ; wings
flavescent, posterior margin broa lly piceous, a somewhat
large spot near middle of anterior margin black ; male with
three long anal a])pendages, the uppermost defieeted from
near base arid directed downwards, its apex acute, the two
lowermost directed sti’ongly outwards and then as strongly
directed inwardly and downward; mesonotum tricarinate.
Jiong., excl. tegm., 3 mm.; exp. tegm. IG mm.
IL.h. Ashanti; Obuasi (IV, M. Graham ^ Brit. Mus.).
Drona hiclavata.
Derhe (The7}{ce?) bichivaia, Westw. Ann. & Ma^. Nat. Ilkt. (2) vii.
p. 209 (1851).
Tlah. Congo.
A remnant of the type of this species is in the British
Museum collection, which, with the description given by
^V estwood, leaves no reasonable doubt that it should be
included in the genus Drona.
Camma, gen. nov.
Vertex of head projecting a little in front of eyes, a little
narrowed anteriorly, the margins strongly ridged, the anterior
margin notched ; face elongate, widest at base, margins
strongly ridged, downwardly recurved, shorter than clypeiis;
antennse inserted beneath eyes, second joint moderately long
and globose ; pronotum narrow, its posterior margin strongly
concavely sinuate, centrally longitudinally carinate ; meso-
notum tricarinate, the lateral carinations sinuate, the central
carination straight ; legs simple ; tegmina about three times
as long as broad, somewhat narrow at base, the costal area
strongly convexly ampliated to about one third from base,
radial area veiy long, crossed near middle by a transverse
vein, a cential longitudinal series of transverse veins defining
a series of discoidal areas, apical areas short, four in number ;
wings little more than one tliird the length of tegmina,
nanow, apically widened.
d'ype, C. ctUatata^ Westw.
My. W. L. Distant on llomoptera. 405
Gamma dllatata.
Devhe {Phenice?) dilatata, "Westw. Ann. & Mag'. Xat. Hist. (2) vii.
p. 209 (1851)'.
llah. Sierra Leone (type in Brit. Mas.).
Fig'. 2.
Camma dilaiata, Westw.
Genus VlVAHA.
Vivaha, Dist. Faun. B. I., Rhyncli. iii. p. 307 (1900).
Type, V. facialisj Dist.
Vivaha saniosa, sp. n.
Body and legs very pale stramineous ; vertex (excluding
margins), upper lateral sides of vertex and front of face, and
a central longitudinal fascia to mesonotum and scutellum
sanguineous ; tegraina pale semihyaline, the venation pale
stramineous, inner margin to a little beyond end of clavus
and a spot above apex of clavus sanguineous, beyond the red
inner margin the apical margin is ochiaceous; wings pale
hyaline ; head jjroduced in front of eyes into a broad flattened
laminate process.
Long., excl. tegm., 4 mm. ; exp. tegm. 16 mm.
IJab. Queensland (A". P. Doddj Brit. Mus.).
Vivaha delicata.
Interamma delicata, Walk. Joiirn. Liim. Soc., Zool. x. p. 118, pi. iii.
tig'. 0 (1807).
Hub. New Guinea.
The InUramma angunta, Walk., may also belong to the
genus Vivaha^ but the unique typical specimen in the
British Museum is without a head.
408
]\rr. W. L. Distant on llomoptera.
Arunta, gen. nov.
Vertex of head longly produced in front of eyes, broad,
laterally sinuate, apically widened and centrally notched on
anterior margin ; antenna? inserted at some distance beneath
eyes, second joint long and broad ; head laterally broad and
sj)a(ulate, very much compressed, face very long, attenuated
centrally, marginally strongly ridged ; cly[)eus short, globose ;
pronotum moderately triangularly anteriorly produced, its
])Osterior margin equally angularly sinuate ; mesonotum
tricarinate, apical joint of rostrum very short ; legs slender;
tegmina elongate, gradually broadening to apex, which is trun-
cate, discoidal areas four, lowermost longest, second shortest
and triangular ; wings a little shorter than tegmina, but
considerably broader, two subapical transverse veins, third
apical area short, triangular.
Type, A. rubrovenoba, Dist.
Allied to Phantasmatocera^ Kirk., from which it differs by
the different shape and structure of the head, the longer
antenna?, apically truncate tegmina, with short and less
numerous apical areas. In his short generic indication
Kirkaldy does not refer to the wings.
Arunta ruhrovenosa^ sp. n.
Head creamy white, margins of face as far as eyes black ;
Fig. 3.
face before eyes pale ochraccous ; clypeus and inner margnn
of eyes black ; eyes and second joint of antennie luscous
407
]\Ii’. W. L. Distant on Ilomoptera.
brown; pronotum creamy white; mesonotiun oeliraceou^,
with a black fascia near each anterior lateral margin ; tegmina
creamy white, subhyaline, interior area from base to lower
apical area pale fuscous, the discoidal longitudinal veins san-
guineous, tlie transverse veins fuscous; wings pale creamy
white, subhyaline.
Long., excl. tegm., 2^ mm. ; exp. tegm. 8 mm.
llah. Queensland [F, F. Dodd, Brit. Mas.).
Kuranda, gen. nov.
Vertex of head somewhat longly produced in front of eyes,
narrowed anteriorly, lateral margins very strongly ridged ;
head laterally much compressed, face long and slender, slightly
widened towards clypeus, 'which is very much shorter than
face; antennte inserted at some distance beneath eyes, the
second jh)int very long and robust ; pronotum very short ;
mesonotum subelongate, faintly carinate; tegmina more than
three times longer than broad, apically rounde.l, apical areas
short and continued round apex of costal margin, costal area
moderately broad ; wings broader and shorter than teginina.
Type, K. notata. List.
liuranda notatOj sp. n.
Body and legs pale ochraceous, eyes and antennae black ;
tegmina very pale creamy ochraceous, with four piceous spots,
I'ig. 4.
Kuranda notata, Dist.
the smallest near base, two discal, and one near apical margin ;
wings pale hyaline.
408
]\Ir. W. L. Distant on lIomo{)tera.
Var. — Tc'grnina with an irregular ])iceous fascia extending
from the basal spot to near apex of inner margin.
Jjong., exel. tegm., 8-^ mm. ; exp. tegm. 10 mm.
JIah. Queensland ; Kuranda {F. P, Doddj Brit. Mus.).
Makula, gen. nov.
Vertex of head broad, modei-ately produced in front of
eyes, triangularly foveate on disk, lateral areas of head broad,
compressed, face narrow, curved downward, not continuous
with the clypeus, which is very robust ; pronotum very
narrow ; mesonoturn elongate, tricarinate, tlie central carina-
tion straight and almost percurrent, the lateral carinations
short and curved inwardly; legs simple; tegmina about
three times as long as broad, costal margin sinuate, apex
obtusely rounded, two series of transverse veins, the first a
little beyond middle, and non-continuoiis, defining five
discoidal areas, the second series more continuous, subapical,
and enclosing the apical areas ; wings shorter and broader
than the tegmina.
Type, M. ornata^ Dist.
Makula ornata^ sp. n.
Head, pronotum, scutellum, body beneath, and legs stra-
mineous ; abdomen above fuscous, the lateral areas and apex
Fig. 5.
Maliula ornata, Dist.
stramineous ; tegmina greyish white, subhyaline, a large
oblique spot before middle, commencing near inner margin
409
W. L. Distant on Ilomoptera.
and terminatlno* at lower raargni of radial area, a small
streak on costal margin beyond middle, and a broken apical
spot which is rnore faintly continued to inner margin, fuscous
or piceous, on the area of these apical fuscous suffusions the
veins are sanguineous; wings pale hyaline.
Long., excl. tegm., 2 mm. ; exp. tegm. 10 mm.
llab. Queensland (Z^. P. Dodd^ Brit. Mus.).
Malaila parviceps.
Briria parvice.ps ^ Walk. Journ. Linn. Soc., Zool. x. p. 11.3 (1867).
Hab. My sol.
Makiila pictipennis.
7?rm« Walk. Journ, Linn. Soc., Zool. x. p. 113 (1807).
Ilah. Morty.
Makula testacea.
Brixia testacea, Walk. Journ, Linn. Soc., Zool. x. p. 115 (1867).
Ilah. My sol.
Genus BhoTANA.
Bhotana, Walk. Journ. Linn. Soc., Zool. i. p. 160 (1857).
d’ype, li. latipennisj Walk.
Walker gave a wrong reference to his figure of the type.
It should be pi. viii. fig. 1, and the type of Paricancij Walk.,
should be pi. viii. fig. 2.
Rhotana ramentosa, sp. n.
Body very pale testaceous, more or less greyishly tornen-
tose ; eyes black; legs stramineous; tegmina subhyaline,
talc-like, more or less iridescent, the veins ochraceous, the
series of transverse veins separating the apical areas, and
most of the apical veins, broadly ochraceous, costal area pale,
with two dark costal marginal spots, one before and one at
apex, a small dark spot on costal margin above the series of
transverse subapical veins, and another beneath these veins
on inner margin, a similar small spot on apical margin, a
transverse creamy-white spot in costal area beyond middle ;
wings pale hyaline ; tegmina with the transverse subapical
series of veins posteriorly curved and terminating near apex
of clavus ; apical areas short.
Long., excl. tegm., 3 mm.; exp. tegm. 12 mm.
Hah. Queensland (F. P. Dodd^ Brit. Mus.).
Ann. (k i\[ag. N. Hist. Ser. 7. Vol. xix.
29
410
j\Ir. W. L. Distant on liomoptera.
Rhotana transversa^ sp. n.
Body above ochraceous, beneath with legs paler ; scntel-
lurn pale sanguineous ; tegmina and wings subhyaline, talc-
like^ the veins more or less ochraceous ; tegmina crossed by
two narrow very pale fuscous transverse fascise, the first at
about one third from base and not extending to the costal
area, the second subapical, between is a third very short
fascia, commencing on costal margin and only extending
about one third across tegmen, a spot on costal area above
the first fascia and a series of small spots on apical margin.
Allied to R. ramentosay Dist., but to be easily separated
from that species by the direction of the subapical transverse
fascia, which follows the direction of the subapical transverse
veins and is not curved inwardly to apex of clavus as in
R. ramentosay but terminates near posterior angle of the inner
margin.
Long., excl. tegm., 2 mm.; exp. tegm. 9 mm.
Hub. Queensland (A. P. Doddy Brit. Mus.).
Rhotana opalinOy sp. n.
Body and legs pale ochraceous; disk of mesonotum,
scutellum, and body beneath sanguineous; eyes black;
tegmina subhyaline, talc- like, with opaline redections, the
venation ochraceous, a subapical undulating fascia, preceded
by two short fascia?, extending from costa to near middle of
tegmen, the outer half of inner margin^ and a series of small
apical marginal spots very pale fuscous, a central edge-like
spot to costal margin, edge of apical margin, and the apices
of the upper apical veins white ; wings pale hyaline ; tegmina
with the costal margin somewhat strongly sinuate beyond
middle, the transverse series of subapical veins not poste-
riorly curved inwards, but nearly straightly transverse.
Long., excl. tegm., 2 mm.; exp. tegm. 9 mm.
Hab. Queensland (A. P. JJoddy Brit. Mus.).
Rhotana septemmaculatay sp. n.
Body and legs pale ochraceous, beneath a little paler than
above ; tegmina and wings pale hyaline, talc-like, the vena-
tion ochraceous; tegmina with a broad, outwardly ajigulate,
pale ochraceous lascia at about one third from base, and two
narrow, transverse, subapical, pale fuscous fasciae, a cluster
of about seven prominent black spots, which are somewhat
arranged in two longitudinal series on inner apical area.
Long., excl. tegm., 2^ mm.; exp. tegm. 9 mm.
Hab. Queensland (F. P. Roddy Brit. Mus.).
]\Ir. \V. L. Distant on llonioptera .
411
Rhotana quadrimacalata, sp. n.
Body and legs ocliraceous ; tegmina pale sliming oclira-
ceous, with opaline reflections, the subapical transverse veins
and the longitudinal veins to apical areas testaceous, a central
apical marginal series of four black spots, each placed near
the apex of a longitudinal vein ; wings pale hyaline, with
opaline lustre; the subapical series of transverse veins
continued inwardly to apex of clavus.
Long., excl. tegin., mm. ; exp. tegm. 11 mm.
Hah, Queensland [b\ P. Dodd^ Brit. Mus.).
Kirkaldy (Rep. Exp. IStat. Haw. Plant. Assoc, pi. ix.
p. 4o5) has described two species of Rhotana from Queens-
land. Of the first, R, chrysonoe^ the British Museum
possesses a single carded specimen, which appears to agree
with the description and has now been doubtfully labelled as
that species in the collection. The description of the second
species is inadequate for identitication without seeing a
typical specimen. ^
Genus Nisia.
Nisia, Melich. Horn. Faim. Ce}don, p. 53 (1903) ; Dist. Faun. B. I.,
Khyncli. iii. pp. & 309 (1900j ; Kirk. liep. Exp. IStat. flaw.
Plant. Assoc, pt. ix. p. 427 (1900).
Kirkaldy writes: ‘^This genus seems to be but slightly
differentiated from Kermeaia, Melich.” On the contrary,
the shape of the tegmina is quite divergent and the number
of apical areas different, eight or nine in Kermesia, five in
JSisia,
Genus PlIACONEURA.
Pliaconeura, Kirk. Rep. Exp. Stat. llaw. Idant. Assoc, pt. ix. p. 427
(1900).
Type, P. Froggatti, Kirk.
It IS necessary to point out that Kirkaldy, in the short
description of his typical species, refers to pi. xxix. figs. 5—6,
whereas it should be, as later on correctly stated m the
“Description of Plates,” j)l. xxix. figs. 3-4.
Fenuahala, gen. nov.
Vertex of head projecting a little in front of eyes, gradually
narrowing to apex, its margins very strongly ridged ; lateral
areas of iiead compressed; face long and narrow, medially
attenuated; clypeus almost as long as face ; antennal inserted
nearer base of clypeus than to eyes, ihe second joint long,
412
Mr. W. L. Distant on Homoptera.
broad, and spalulate ; apical joint of rostrum minute ; pro-
noturn veiy short, triangularly sinuate ; mesonoturn with its
anterior margin strongly angulate, discally tricarinate ; legs
sim[)le; tegmina about three times as long as broad, apically
ampliated, apical margin subtruncate, a subapical line formed
by transverse veins separating a number of short apical areas,
which commence at apex of costal margin and are con-
tinued to near end of clavus, wdiere they are larger, a trans-
verse vein beyond apex of radial area; wings extending to
posterior angle of inner margin of tegmina, a prominent
transverse vein beyond apex of radial area.
Type, F. infiiscata^ Dist.
This genus appears to have some affinity wdth Heronao'^
Kiik., but it is impossible to form a definite opinion. In the
indications given of that genus Kirkaldy writes: Allied
to Futara, Westwmod, but the venation is different.” lie,
however, abstains from any description of the venation, nor
does he mention the tegmina either as regards shape or
breacfth, or in relation to the wings, wdiich are also not
alluded to.
FennahaJa iufuscata^ sp. r.
Body above pale dull umber-browm, abdomen with darker
and paler suffusions; body beneath a little paler than above;
Tig. (t.
Fem(ah(da infvscdta, Dist.
legs stramineous, apices of femora and annulations to tlbim
fuscous brown ; tegmina pale hyaline, talc-like, the veins
413
Mr. ^\^ L. Distant on Ilonioptera.
fuscous brown, the same colour more or less suffuses the
apical areas and irregularly appears on disk and more palely
so in clavus, costal area vvdth transverse fuscous spots; near
centre of subapical margin is a prominent piceous spot ;
wings pale liyaline.
Long., excl. tegm,, 3 mm. ; exp. tegm. Id mm.
llah. Queensland; Kuranda [F.P. Dodd^ Brit. Mus.).
Fenaahala rubrinervis^ sp. n.
Body above fuscous, more or less greyishly tomentose,
lateral margins of the vertex of head greyish white ; antennag,
body beneath, and legs very pale ochraceous ; eyes dark
castaneous ; tegmina greyish subhyaline, with dark fuscous
markings, the principal greyish areas being costal and claval,
three small discal spots, and three large irregular spots on
a})ical area, three small fuscous spots in costal area beyond
middle, veins to apical marginal areas sanguineous; wings
very pale fuligitious with a slight opaline lustre, veins dark
fuscous.
Long., excl. tegm., mm.; exp. tegm. 14 mm.
Hab. Queensland ; Cairns District [F. F. JJodd^ Brit,
Mus.).
Fenuahala juno^ sp. n.
Body above piceous ; body beneath and legs stramineous,
lateral margins of sternum and lateral marginand apical area
of abdomen piceous ; tegmina piceous, the veins to apical
marginal areas sanguineous, on under surface the costal
margin beyond middle also sanguineous; wings very pale
fuliginous, the veins darker.
Long., excl. tegm., 2 mm. ; exp. tegm. 12 mm.
Hah. Queensland; Kuranda [F. P. Dodd, Brit. Mus.).
A small species, to be recognized by its very distinct
coloration.
FenuaJtala pallescensj sp. n.
Body above pale tawny, more or less greyishly tomentose ;
vertex of liead cretaceous white ; eyes piceous ; body beneath
and legs stramineous, apex of abdomen slightly testaceous ;
tegmina and wings subhyaline, with a slight opaline lustre,
the veins very pale dull ochraceous ; tegmina with the inner
claval margin pale dull ochraceous.
Long., excl. tegm., 2^ mm. ; exp. tegm. 12^ mm.
llab. Queensland [F. P. Dodd^ Brit. Mus.).
414
^Ir. AV. L. Distant on Ilomoptcra.
Urabunna, gen. nov.
Vertex of head moderately ])rojecting beyond eyes, nar-
rowing to apex, which is angularly sinuate, margins strongly
ridged ; face narrow, lateral margins stiongly ridged, a little
widened posteriorly, longer than clypeus ; rostrum with the
apical joint minute ; antennre short, globose, inserted imme-
diately beneath eyes ; pronotum short, concavely sinuate ;
mesonotum tricarinate; legs simple; tegmina elongate, more
than three times longer than broad, costal margin prominently
arched a little beyond base, thence somewhat narrowed to
apex, which is moderately rounded, two transverse veins
near middle, one beyond clavus, another limiting penultimate
apical area, and two or three subapical, which help to detine
a series of short apical areas continued round apex of costal
margin ; wings ample, much broader than tegmina, about
reaching posterior angle of tegminal inner margin, the apical
veins very pronounced.
Type, U. lineata^ Dist.
Urahunna lineata, sp. n.
Body and legs dull ochraceous; abdomen somewhat
greyishly tomentose ; tegmina pale greyish subhyaline, much
suffused wdth fuscous brown, costal area pale, traversed by a
broken, longitudinal, piceous, linear fascia Avhich almost
Fig. 7.
reaches apex ; wings hyaline, the apical veins })roniinent
and infuscate.
Long., excl. tegm., 3 mm. ; exp. tegm. 13-14 mm.
Hah. Queensland [F. F. Dodd, Brit. Mus.).
]\Ir. W. L. Distant on Ilornoptera.
415
Appendix.
I have furtlier examined Mr. Kirkaldjhs paper describing
Australian Fulgoridie under the title of “Leaf-Hoppers and
their Natural Enemies/’ published at Honolulu, in comparison
with the descriptions of some Queensland Fulgoridse which
have appeared in these pages and elsewhere. The following
notes are necessary.
8 u b fa m . E ur ybraciiybinm.
Dardus ohscnrm.
Dardus ohscurm, Dist. Trans. Ent. Soc, Lond. 1892, p. 283.
Dardus immaculatm, Kirk. Rep. Exp. Stat. Haw. Riant. Assoc, pt. ix.
p. 446 (1906).
Hob. Queensland.
Subfam. Dictyoruarinje,
Eictyopliara hifasciata.
Dictyophora hifasciata, Hist. Trans. Ent. Soc. Lond. 1892, p. 279.
Thanatodictya (^Lucinda) lucinda, Kirk. Rep. Exp. Stat. flaw. Plant.
Assoc, pt. ix. p. 392 (1906).
Ilab. Queensland.
DictyopJiara insignis,
Dictyophora insignis, Hist. Trans. Ent. Soc. Lond. 1892, p. 279.
Thanatodictya {Niculda) anadyomene, Kirk. Rep. Exp. Stat. Haw.
Plant. Assoc, pt. ix. p. 393 (1906).
S u b f a m . CixiiNAE .
In the genus Oliarus Kirkaldy has described nine Aus-
tralian species, and I have described two from Queensland,
which, even by the aid of his tabular synopsis, I cannot
decide as synonymic or otherwise. One thing, however, is
certain, we have both used luhra as a specific name ; I
therefore alter mine.
Oliarus incerta^ n. noin.
Oliarus lubra, Dist. Ann. & Mag. Nat. Hist. (7) xix. p. 282 (1907).
41G
^Ir. G. J. Arrow on new
Subfam. Ttlovidvctiinm.
Genus Ficarasa.
Ficarasa, Walk. Journ. Linn. Soc., Zool. i. p, 1G2 (18o7).
Feltodictya, Kirk. Rep. Exp. Stat. Haw, Plant. Assoc, pt. ix. p. 414
(1906).
The species I described as F. australasice (Ann. & Mag.
Kat. Hist., antOy p. 287) is apparently closel}" allied to the
one described by Kirkaldy as P. hurandcE^ but nothing defi-
nitely can be decided from Kirkaldy’s description, which is
simply — Grass-green. Eyes and genital segments partly
brownish. Veins lightly and shortly piligerous — and is
applicable to almost any species in the genus.
Fa nua viiiemis.
Vanua viliemis, Kirk, Rep. Exp, Stat. Ilaw. Plant. Assoc, pt. ix. p. 416,
pi. xxvii. figs. 6-7 (1906).
It may be useful to point out that Kirkaldy, in his
description, has referred this species to pi. xxvii. figs. 7-9;
it should be figs. 6-7.
Subfam. Ackilinm.
Genus Aneipo.
Aneipo, Kirk, Rep. Exp. Stat. Haw. Plant, Assoc, pt. ix. p. 425 (1906).
2'udea, Hist. Ann. Mag. Nat. Hist. (7) xix. p. 290 (1907).
Aneipo diva.
Arieipo diva, Kirk. Rep. Exp. Stat. Haw. Plant. Assoc, nt. ix. p. 425
(1906).
Tudea picturata, Hist. Ann. & Mag. Nat. Hist. (7) xix. p. 290 (1907).
Hah. Queensland.
L. — Some new Species and Genera of Lamellicorn Coleoptera
from the Indian Empire. 13y Gilbert J. Arrow.
Part II.
Copridae.
Cassolus humeralis, sp. n.
Rufo-piceus, capite prothoraceque rufo-cupreis elytrorumque Imraeris
flavo-maculatis : breviter ovatus, convexus ; capite crebro punc-
tato, antice 4-dentato ; prothorace paulo minus crebre puiictato,
417
Coleoptera from the Indian Kinpire.
elytrorum laiitudiiio, latcribus fortiter curvatis, antice vix angus-
tatis, angulis posticis vix perspicuis ; elytris profunde punctato-
striatis, inters titiis locvibus; pygidio fortiter punctate; tibiis
anticis extiis minute denticulatis, dentibusque tribus validis
acutis armatis, tibiis 4 posterioribus valde curvatis, tarsorum
posticorum articulo primo quam secundum distinct© longiore.
(S . Tibia antica subtus unispinosa, tibiis posticis gracilioribus
femoribusque posticis subtus dilatatis.
Long. 4 5 mm.
JIah. Assam, Sudiya, Patkai Mts.
Piceous black, with a sligdit metallic tinge, and liaving* the
liead and prothorax coppery and the elytra decorated upon
each shoulder with a small round j^ellovv spot.
The form is short and compact. The head is closely
punctured and without visible sutures or elevations, but the
front maroiii is armed with four teeth, of which the middle
pair are the longest and most acute. The prothorax is
moderately closely punctured, but very shining ; it is as
broad as the elyTra and not much narrow^er in front, with the
sides strongly and uniformly curved and the hind angles
obsolete. Tlie elytra are strongly punctate-striate and the
pygidium strongly and rather closely punctured. The front
tibise are strongly tridentate and very finely denticulate
between and above the teeth. The middle and hind tibife
are slender and curved, and the tarsi have the first joint
nearly twice as long as the second and the remainder nearly
equal.
The male has a perpendicular tooth beneath the front
tibia, the hind tibia is longer, more slender and more strongly
curved than that of the female, and the hind femur is strongly
dilated, the flange forming an obtuse angle near the knee.
Several specimens were collected by Doherty.
Although two species of Cassolus have already been de-
scribed by Sharp and Lansberge {G. nudus, Sharp, Siam,
and C. sumatranus, Lansb., Sumatra), the very interesting
sexual characters have not yet been noticed. In the appendix
to this paper I have described another species in which other
sexual differences are found. I have not yet seen either of
the previously described forms, but the present insect appears
to be like C. nudus, Sharp, but sufficiently distinguished by
the yellow shoulder-spots and the puncturation of the ])ygi-
dium, which is closer than that of the prothorax.
Panelus assamensis, sp. n.
Castaneus, laevis, nitidus, breviter ovatus, postice latior : P. parvulo
simillimus, sed prothoracis parte posteriore laevigata, linea elevata
418
Mr. Gr. J. Arrow 0)l new
arciiata distincte demarcata ; elytris paiilo distinctiiis striabis efc
])unctatis ; metasterrio medio antice valde prodiicto, acute angu-
lato ; pedibus gracilibus, tibiis omnibus curvatis, tarsis 4 poste-
rioribus longis, robustis, anticis brevibus.
liOng. 2'5 mm.
Ilah. Assam, Sudiya, Patkai Mts.
Three specimens were collected by Mr. Doherty, in which
no sexual difFerence is apparent.
This little insect, one of the smallest Lamellicorn beetles
yet known, has a very great resemblance to the Japanese
Panelm parvulas^ Waterh., figured by Mr. George Lewis in
the Ann. & Mag. Nat. Hist. 1895, (6) xvi. p. 375, tigs. 1 & 2,
the only species hitherto described, although I have seen
several. It is a little smaller and relatively shorter, being
a trifle less produced behind. It is similarly sculptured,
but the elytra are a little more strongly striated and punc-
tured, and the prothorax has a smootli unpunctured strip at
the base, which is separated by a distinct dividing-line from
the remaining strongly punctured surface. This smooth
area exists in P. parvalus^ but is not sharply defined. The
most pronounce 1 difFerence between the two forms, however,
is found in the lower surface. The metasternum, instead of
meeting the mesosternum in a broad lobe, is produced into
a very acute angle, apparently almost dividing the meso-
sternum into two. The sides of this process form quite
straight lines extending backwards to the posterior ends of
the middle coxse. These lines are finely raised, and under-
neath them can be traced another strongly pigmented line,
which probably represents the true meso-rnetasternal suture.
This line of dark pigment does not follow the direction of
the carinate lines in the posterior part, but branches off to
the middle of the intermediate coxae.
Genus Ontiiopiiagus.
[a] Male with two imcoimected cephalic horns.
0 nthophacjm gladiator^ sp. n.
iEneo-niger, opacas, depressus, latus ; capito dense punctate, breviter
setose, margine integro, antennis tlavis ; protiiorace producto,
angulis anticis hand acutis, posticis distiuctis, lateribus basicpie
medio angulatis, vix a’^cuatis ; elytris planis, subtiliter striatis,
interstitiis minute punctulatis ; pygidio leviter punctato, meta-
sterni lateribus fusco-liirsutis.
<S . Capite posticc cornul)us duobus gracilissimis, basi coiinexis,
quarn corporc toto paruiu brcvioribus, postice subtus serratis,
4U)
CoJeoptera from the Indian Kmpivp.
armato ; prothorace fere impnnctato, antice profundo excavato,
dorso ])rocessu longo, compresso, antice oblique directo, armato.
$ incognita. •
l.ong. 17-19 mm.
Hah. Tiulia.
Obscurely bronzy, broad and depressed, with the upper
surface opaque. The head is punctured and setose, witli the
front margin entire and nearly straiglit in front. The pro-
thorax is rather broader than long, with the margins recti-
linear, tlie hind angles not rounded off, and the base and sides
angulated in tlie middle. The elytra are flat, faintly striated
and very finely punctured. The pygidiuin is lightly punc-
tured, tiie sides of the metasterninn clothed with fine hair,
and the abdomen almost smooth.
^ . The sutural carina of the head is strongly angulated
and the head is provided behind with a pair of backwardly-
directed horns almost as long as the body when well-developed ;
they are united at the base, gently curved, slender and
serrated beneath at the hinder part. The prothorax is
excavated beneath, and from the posterior margin of the
cavity proceeds a long, compressed, straight horn, directed
obliquely forwards and blunt at its extremity.
J have not seen the female.
This species is closely related to 0. Mouhoti^ liar., but is
broader and more flattened and has the margins of the pro-
thorax less curvilinear. The cephalic horns differ in being
serrated beneath, and the thoracic horn is placed farther
back, and in our specimen is more slender. The thoracic
cavity extends farther baek, but rather less from side to side.
Two specimens, without precise locality, were received with
the Bowring collection.
Onthopliapus rubripennis, sp. n.
Niger, subtus nitidus, supra opacus, capite prothoraceque obscure
cupreis, elytris rufo-castaneis ; capite rugose punctate, semi-
circulari, integro ; prothorace fortiter et dense punctato, mar-
ginato, angulis anticis paulo productis, hand acutis, posticis
obsoletis ; elytris stria tis, striis subtiliter et lineare punctatis,
interstitiis planis, fere Icevibus ; pygidio grosse et crebre punctate.
S . Capite postice cornubus duobus longis curvatis, valde genicu-
latis et divergentibus, armato ; prothorace antice longitudiualiter
excavato.
2 . Capite leviter bicarinato, prothorace medio leviter sulcato.
Long. 11-13 mm.
Hah. Sikkim, Kurseong,
Black, with the head and prothorax slightly bronzy, the
4 20 Mr. G. J. Arrow on neio
elytra dull brick-red, and the antennas red with a pale yellow
club.
The form is broad and convex, with the head semicircular,
rug’osely punctured, and scarcely emarginate in front. The
])rothorax is coarsely and densely punctured, completely
margined, with the base slightly angulated in the middle,
the anterior angles rather produced but not sharp, and the
posterior angles almost obsolete. The elytra are shallowly
striated, the striae containing very elongate })unctures having
each a raised margin, and the interstices are flat, opaque, and
almost smooth. The pygidium is flat, opaque, and coarsely
and densely punctured, and the metasternum is rather closely
but not strongly punctured and not prominent in front.
TTie front tibiae are moderately long and armed with four
strong but not sharp teeth.
. The head bears behind a pair of long, curved, and
widely divergent horns, like those of the European 0. tauras,
but strongly elbowed internally near the point of origin.
The prbthorax has a slight longitudinal excavation in front.
$ . There are two slight transverse carings upon the head
and the prothorax is only feebly channelled along the middle.
There are specimens in the British Museum and M. Rene
Oberthiir’s collection.
Var. suhcribratus.
The punctures of the prothorax are exceedingly dense and
confluent, so that the metallic lustre is scarcely traceable,
and the anterior excavation in the male is limited behind by
two rather sharp tubercles, the corresponding elevation in
the Sikkim form being quite blunt.
Hah. Bootan, Khasia Hills, N. Chin Hills.
Onthophagus hufo^ sp. n.
ISTger, iioununquam obscure seneus, parum nitidus, ovatus ; capitc
baud producto, cl}'peo sat crebre punctato, margine leviter
elevato, antice paulo inciso ; protliorace parum convexo, lateribus
valde arcuatis, aiigulis posticis baud distinctis, basi baud dis-
tincte angulato aut marginato; elytris distincte striatis, intcr-
stitiis parcissime granulatis ; pygidio sat convexo, parce punc-
tato ; corpore subtus lu)vi, mctasterno ubique sat grosse punctato,
abdomine inipunctato.
(S . Capitis Carina suturali oblitcrata, vertice cornubus duobus,
postice directis, fere rectis, basi dilatatis, baud coimexis, armato,
spatio incluso parcius punctato, antice cariua vix porspicua
limitato ; protborace antice perparum declivo.
5 . Clypeo rugoso, carina antica recta, parum elevata, alteraqiu'
421
CoJeoptera from the Indian Kmpire.
medio broviter acuminata, vertice leviter excavato, piinctato,
postice bituberculato ; prothorace baud declivo.
J.ong. 6-8 mm.
Jlah. N.W, India, Baiinn, Gwalior.
Black, sometimes with a faint bluish or greenish tinge,
and not very shining above. It is a small species, elongate-
ovate in shape. The head is not produced, but is slightly
emarginate in front. The prothorax is evenly and not very
finely punctured, with the sides strongly curved, the posterior
angles rounded off and the base not angulated or margined.
The elytra are moderately deeply striated, with sparsely
scattered granules upon the interstices. The pygidium is
thinly punctured, the metasternum evenly but not very
strongly, and the abdomen im punctate.
d . The clypeus is moderately punctured, and the vertex
armed with two straight horns of moderate length, directed
slightly backward and dilated at their bases but not connected
together. There is a very slight ridge just in front of the
horns, and the enclosed space is concave and thinly punctured.
The prothorax is feebly declivous and smooth in front.
$ . The clypeus is finely rugose, with a very fine anterior
Carina and a well-marked posterior carina acuminate in the
middle, and the horns are represented by a pair of short
tubercles. The intervening space is concave and punctured.
This species is allied to 0. tragus^ F., and 0. darna^ F.,
but is smaller and more elongate than either.
Onthopliagus rectecornutus, Lansb., is an interesting species
belonging to the same group which needs further investi-
gation as regards both its variation and geographical distri-
bution. It is stated by Lansberge (Leyden Museum Notes,
1883, p. 50) to inhabit Java, Surnbawa, and Ceylon, and to
differ from 0. Juridus (I)ej. MS.) only by the curved and
diverging horns of the latter. The two forms thus distin-
guished appear to me to be specifically inseparable, although,
in addition to the development of the horns, the luridiis form
has the prothorax more globose and much more contracted
in front and the head more narrow. These are all characters
peculiar to the male, and in other respects there is so much
constancy and individuality that I feel obliged to regard this
as a single species with two forms of male. Lansberge
remarked that the female was unknown to him, but probably
looked for a hornless form and did not notice the real sexual
differences, which, contrary to the general rule, are not found
in the armature. Male and female of the rectecornutus form
are alike armed with a pair of erect parallel horns and a
422
j\Ir. G. J. Arrow on new
transverse carina upon the cl y peal suture ; they may be
distinguished by the front tibiic, which, in the male, are
elongate, slender, and strongly curved, with the teeth feebler
and farther apart. The specimens of the hiridm form are
all males. The three forms have been taken too-ether in
Ceylon and Travancore, wliere they a[>pear to be common.
L have only seen a single female from Java, and there is a
single male in our collection from Darjeeling, while Dejean
gives China as the habitat of the var. laridas^ so that the
species has evidently a very wide range.
0 nthoi>lia(jus hrevicollisj sp. n.
Niger, nitidus, anteimis tarsisque rufis ; prothorace valde convexo,
piinctato, quam corpore ad Immeros miilto latiore, margine
postico arCLiato, marginato, hand angulato ; elytris striatis, striis
puiictatis, iuterstitiis coovexis, politis, vix punctulatis ; pygidio
parce punctate.
C5* . Capite polite, lateribus aiitice fere rectc contractis, margine
antice panic emarginate, carina sutnrali parnm arcnata cornu-
bnsqne duobus posticis graeilibns, parallelis, erectis ; prothorace
antice verticali, la)vigato, postice leviter punctate, carina dorsali
transversa.
2 . Clypeo seinicirculari, vix emarginato, pnnctato-rugoso, carina
sutnrali distincta, valde curvata ; capite postice loeviore, bituber-
culato ; prothorace antice medio subverticali, carina dorsali
transversa.
Long. 9-5-13 mm.
JIab. S. India, Nilgiri Hills, Belgaum.
Black and shining, with the anteinue and tarsi reddish.
The prothorax is finely but not closely punctured all over,
except upon the anterior declivity, and the elytra are sulcate,
with the interstices convex and scarcely punctured. The
pygidium is sparingly punctured and the median part of the
inetasternuni has only a very few punctures near the sides.
The spur of the front tibia is bent at a right angle and acute.
d. The head is octagonal, slightly emarginate in front
and irnpunctate except just in front of the eyes. There is a
distinct carina upon the clypeal suture, and two slender,
unconnected horns arise almost vertically from the back of
the head, with a slight forward curvature. The prothorax
is very short and broad, vertical and smooth in front, very
convex and distinctly punctured behind, with the anterior
edge of the dorsal ])art carinated in front and the posterior
edge curved, finely margined and not angulated. The Iront
tibial spur is broader than in the female.
^ . 'The clypeiis is curvilinear and very feebly emarginate
423
Coleoptera from the Jiulian JCmpire.
in front_, and Its surface is ratlier finely rugose. The carina
is stronger, and the horns are represented by a pair of conical
tubercles. Tlie prothorax is more elongate and only slightly
ret use in front.
[h) Male with two connected cephalic horns.
Onthophagus productus, sp. n.
Xiger, convexus ; capite parvo, subtilitcr rugoso-punctato, antico
rotundato, vix emarginato , prothorace postice ante basin fere
carinatim elevato, lateribus valde angulatis, margine postico angu-
lato, angiilisque posticis distinctis ; elytris striatis, interstitiis
convexis, crebre et rugose punctatis, minutissirae setosis ; pygidio
sat grosse punctate ; metasterni medio uitido, parce sat grosse
punctate.
. Capite postice cornubus duobus curvatis, baud longis armato;
prothorace nilido, minute baud crebre punctato, lateribus antice
vix punctatis, antice valde angustato, angulis anticis acutis,
fossulatis, dorso antice medio elevato ; tibiis anticis elongatis, sat
gracilibus.
5 . Capite carina suturali aliaque frontali medio leviter acuminata
armato ; prothorace rugose punctato et setose, carina transversa
antica parum elevata.
Long. 10-12 mm.
Ilah. N. India, Sikkim.
Black, clothed with short dark setae, except at the middle
of the metasternum and the prothorax of the male, which is
shining. The head is small, finely and rugosely punctured
and scarcely emarginate in front. The prothorax is strongly
angulated at the middle of the sides and distinctly at the
middle of the base, and the hind angles are not rounded
away. The posterior part of the disk is elevated into a blunt-
edged carina just before the margin. The elytra are striated,
with the interstices convex, finely and rugosely punctured
and setose. The pygidium is rather coarsely punctured, and
the middle of the metasternum shining and sparingly
punctured.
. The head is produced behind into a pair of slender
but not very long horns, united by a sharp ridge at the base
and forming v/ith it rather more than half of a perfect circle.
The prothorax is shining, finely and not closely punctured
and not setose, except in front, where it is slightly rugose.
It is strongly contracted in front, elevated in the median
part, and much depressed at the sides, with a round pit in
each front angle. Tiie front tibiae are elongate and slender.
$ . The head bears two transverse carinas, of which the
frontal one is minutely acuminate in the middle. The pro-
424
Mr. G. J. A rrow on new
thorax is not greatly narrowed in front and is strongly and
thicldy punctured and minutely setose, with an interrupted
transverse carina in front.
0. pi'oducius is very closely related to 0. rngulosus, liar.,
from N. China. It is less densely pujictured above, especially
upon the prothorax of the male, which is also still more
narrowed in front, and the front tibiae in the same sex are
more elongate.
There are examples in the British Museum and in M. Reue
Oberthiir^s collection.
OntJiophagus [Caccohius) gallinas, sp. n.
Niger, capito prothoraceque obscure cupreus, subnitidus, crebre et
rugose punctatus, ubique breviter sotosus ; clypeo ernarginato,
grosse punctato ; prothorace dense punctato, marginato, angulis
anticis baud acutis, posticis obsoletis ; elytris striatis, inter-
stitiis granuiatis ; pygidio grosse punctato, metasterno sat
pinictato ; tibiis anticis fortiter tridentatis.
(S . Capitc nitido, leviter ernarginato, carina anteriore arcuata
posterioreque recta valde elevata, surnma tritida,-dentibus duobus
exteruis longioribus, recurvatis ; prothorace dorso elevato, antice
fere acumiriato, parte antica excavata, polita ; metasterno postice
impresso.
2 . Capite rugose punctato, paulo producto, antice acute bidentato,
carina anteriore arcuata posterioreque fere recta ; prothorace
antice medio minute prominente.
Long. 4-5 mm.
Hah. S. India, Nilgiri Hills, Belgaum.
Deep bronzy black, with the head and prothorax generally
more distinctly metallic, clothed above and below witli
greyish sette. The prothorax is densely punctured, the
elytra striated and rugosely granulated, and the pygidium
coarsely punctured. The metasternurn is strongly but rather
sparsely punctured and- not prominent in front. The anterior
tibia3 bear three strong teeth, above which they are finely
denticulated.
d . The head is shining, thinly punetured, slightly notched
in front, with a curved sutural carina and a very strongly
elevated frontal one terminating in a triangular median tooth
and a pair of short recurved lateral horns. The prothorax is
convex behind, slightly excavated and smooth in front, the
upper margin of the excavated part forming a slight tooth in
the middle.
9 . The head is rugosely punctured and sharply bidentate
in front, with two slight carina;.
This species belongs to the closely connected grou[) of
425
Coleoplera from the Indian Empire.
small Indian 0 ntliophagi of wliicli 0. valcanuSj Fab., is the
type. It is most nearly related to Gaccohius tortus, Sharp, but
is more closely sculptured, especially upon the prothorax, and
the elytra are without the yellow apical spot distinguishing
that species and 0. vulcanus. It is peculiar also in having
the middle of the prothorax angularly instead of broadly
prominent in the male. It has been found in numbers by
Mr. H. L. Andrewes in the Ouchterlony Valley, Nilgiris
(3000 ft.), in April and July.
Another species of the same group, although not hitherto
associated with it, is 0. farculusy Fab., in which the elytra
are marked with yellow in front and behind, and the margin
of the thoracic excavation is not sharply raised. This is
found in the same localities and also in Ceylon.
(c) Head of male produced into a single horn behind.
Onihophagus ephippioderus^ sp. n.
Niger vel obscure nigro-aeneus, robustus, convexiis, sat nitidiis,
autennis flavis ; elytris punctato-striatis, interstitiis vix convexis,
parce et minute punctulatis, lateraliter fortius ; pygidio leviter
pimctulato.
S . Capite grosse punctato, antice et postice producto, antice
fortiter uasuto, postice acute cornuto, cornu gracili, baud longo,
basi late angulate laminate ; prothorace sat punctato, antice
retuso, laevigato, dorso antice utrinque oblique tuberculato.
$ . Capite antice ruguloso, postice punctato, margine antico integro,
leviter producto, carina suturali valde curvata laminaque postica
erecta paulo bifida ; prothorace crebre punctato, carina antica
obsoleta.
Long. 12-14 mm.
Hah. Mysore, Bangalore, Nilgiri Hills.
Black or slightly seneous, with pale antennae. The shape
is broad and convex, and the body smooth and beneath
almost impunctate. The elytra are shallowly punctate-striate
and the interstices very lightly punctured. The prothorax
is strongly punctured and margined, but not angulated at
the sides or base, the posterior angles being rounded away.
The pygidium is finely but not closely punctured.
d . The head is produced into a blunt point in front, the
sutural carina is well curved but not strongly marked,
and the posterior part of the head is produced backwards,
forming a reflexed angular plate behind each eye and a
slender curved horn in the middle. The prothorax is refuse
and smooth in front, with the dorsal ridge produced obliquely
on each side. The spur of the fiont tibia is short and
blunt.
Ann. cb Mag. N. TUst. Scr. 7. Vol. xix.
30
426
Mr. G. J. Arrow on new
$ . The head is strongly rugulose in front of the sutural
Carina, which is more pronounced tlian in the male, and
punctured behind it. The clypeal margin is subangulate
and the posterior margin of the head bears an elevated trans-
verse lamina which is ernarginate at its upper edge. The
protliorax is closely and coarsely punctured. The spur of
the front tibia is slender and acute.
This species resembles 0. semculus, F., the male of which
is without cephalic armature, but the processes upon the
prothorax attain a greater development in that insect, and
the clypeus is less developed in front as well as behind.
OnthopJiagus manipurensis, sp. n.
Niger, iiitidus, corpore lateraliter brnnneo-hirto, antennis pallide
testaceis ; clypeo ruguloso ; prothorace baud fortiter punctato,
postice laeviore, medio obsolete sulcato, antice excavate, lateribus
basique marginatis, basi medio angulata, angulis posticis obso-
letis ; elytris striatis, interstitiis planis, subtiliter pimctulatis.
(S . Capite antice fere acute angulato, medio sat fortiter bicuber-
culato, postice retrorsum producto, cornu recurvato, acuminate,
basi lato quadrate ; prothorace antice valde retuso, supra leviter
emarginato.
2 . Capite antice arcuato, parum producto, medio fortiter recte
carinate, postice breviter producto, processu erecto truncate;
prothorace antice paulo excavate, supra bituberculato.
Long. 15-19 mm.
JIab. N. India, Manipur.
Black and shining, with testaceous antennse and reddish-
brown hair at the sides of the sternum and abdomen. The
form is short and compact and the sculpture fine. The pro-
thorax is smooth and very sparingly punctured above ; it is
distinctly margined at the sides and base, the base is distinctly
angulated in the middle and the bind angles are rounded
away. The elytra are distinctly striated and the interstices
slightly convex and feebly })unctured. The pygidium is
moderately punctured. The front tibim are broadly quadri-
dentate and the first joint of the middle and hind tarsus is very
large and flattened. The tibial spurs are blunt and that of
the front tibia is strongly bent.
J . The clypeus is rather elongate and pointed, with the
sides straight. The clypeal suture is marked by two strong
tubercles, which divide the width of the head into three
nearly equal parts. The posterior part of the head is smooth
and produced backwards into a lamina, which narrows into a
slender recurved horn. The prothorax is excavated and
rugosely punctured in front and scarcely punctured behind.
Coleoptera from the Indian Empire, 427
and the upper edge of the excavated part is slightly emarginate
in the middle.
^ . The clypeus is rounded, scarcely produced, with a
short, straight, and strongly raised sutural carina, and a small,
vertical, and truncated posterior horn. There is a rudimentary
excavation at the front of the prothorax and a pair of slight
tubercles at its posterior edge.
This species is very closely related to 0, rubricoUis^ Hope,
but differs from it by its uniform black colour, the feebler
sculpture of the elytra, and the presence of the two clypeal
tubercles in the male. The basal joint of the antenna has a
serrate carina, as in 0. ruhricolliSj diaholicuSj and other allied
species.
Onthophagus cupreiceps^ sp. n.
Obscure cupreus, prothorace, elytris pygidioque fuscis, dense opacis,
parce et minutissime setiferis ; capite nitenti, Isevi, antice leviter
punctate, margine antice laevissime excise ; pretheracis lateribus
leviter sinuatis, angulis anticis paule preminentibus, hand acutis,
pesticis tote obsoletis ; elytris sat planis, hand distincte striatis ;
corpore subtus pedibusque seneo-micantibus, sat grosse punctatis.
J . Capite postice retrersum producto, cornu recurvato, acuminate,
basi laminate, quadrate ; prothorace antice leviter retuso,
laevigato.
2 . Capite antice magis punctate, sutura clypeali valde carinata,
semicirculari, vertice carina angusta, supra emarginata armato.
Long. 8-5-10 mm.
Hab. Sikkim, Khamba Jong, Tungu (13,000-16,000 ft.).
Head, legs, and underside shining coppery, with prothorax,
elytra, and pygidiurn very opaque, pitchy black, and very
minutely setose. The head is impunctate except in front,
and the front margin is very feebly refiexed and slightly
excised in the middle. The prothorax has a barely perceptible
angulation at the middle of the base, the hind angles are
completely rounded off and the sides feebly sinuated, the
front angles being prominent but not sharp. The anterior
part is shining and slightly retuse in both sexes. The elytra
are faintly striated.
^ . There is a short, sharp, curved horn arising from the
posterior edge of the head and forming a quadrate lamina
at the base. Only the front margin of the clypeus is
punctured.
$ . The horn of the male is represented by a strongly
raised truncate carina. There is a curved sutural carina, in
front of which the clypeus is rugosely punctured.
This species is very close to 0, concolor^ Sharp, but differs
30*
428
Mr. G. J. Arrow on neio
by its metallic head and underside, and the very opaque
sooty prothorax and elytra.
Onthopliagus tihetanus, sp. n.
Niger, capite, pedibus corporeque subtus micantibus ; prothorace,
elytris pygidioque sat opacis, minute setiferis ; capite antice
rugose, postice parce punctate, margine antico minute excise ;
prothorace angustissime marginato, basi medio laevissime angu-
lato, lateribus simplice arcuatis, angulis omnibus rotundatis ;
elytris leviter striatis, interstitiis vix convexis ; corpore subtus
pedibusque sat grosse punctatis.
(S . Capite postice retrorsum producto, cornu recurvato, acuminate,
basi laminate, quadrate ; prothorace antice leviter retuso,
lo3vigato.
2 . Sutura clypeali valde carinata, semicircular!, vertice carina
angusta, supra leviter emarginata armato.
Long. 7*0-10 mm.
Hob. Brahmapoutra Valley, Chaksam (12,000 ft.),
Gyangtse (13,000 ft.).
0. tihetanus is exceedingly close to 0. concolor and
0. cupreiceps^ but is easily distinguished from both by the
uniform curvature of the sides of its prothorax, which com-
pletely rounds off the front angles. The upper surface is
more opaque than that of 0, concolor ^ but less so than that
of 0. cupreiceps.
Onthopliagus vividus^ sp. n.
Yiridis vel violaceus, capite prothoraceque splendide rufo-cupreis
vel viridibus ; capite ruguloso, clypeo paulo producto, integro ;
prothorace granulate, postice medio Iseviore ; elytris planis,
subopacis, subtiliter striatis et punctatis; pygidio grosse punctato;
metasterni medio polito, antice paulo prominente.
J . Capite crebre punctato, postice cornu brevi, simplice armato ;
prothorace antice paulo et anguste excavate.
$ . Clypeo rugose, carina arcuata distincta, capite postice breviter
tuberculato ; prothorace antice nonnihil excavate.
Long. 11-15 mm.
Hah. Bangalore, Trivandrum, Nilgiri Hills.
Deep blue or green, with the abdomen, tibim and tarsi,
and the front of the clypeus nearly black, the antennm yellow,
and the head and prothorax fiery red, bronzy green, or bright
metallic green.
The form is short and compact. The clypeus is slightly
produced and pointed. The prothorax is coarsely granu-
lated, except at the middle of its posterior part, where it is
nearly smooth ; it is finely margined all round and bluntly
429
Coleoptera from the Indian Empire.
angulated in tlie middle of the base, with a slight anterior
excavation in both sexes^ which is limited behind by a pair
of opaque prominences. The anterior angles are rather
acute. The elytra are finely striated and their interstices
flat and closely punctured. The pygidium is coarsely and
evenly punctured. The front tibia2 are stout and the terminal
spur strongly bent.
d . The short cephalic horn is directed backwards with a
slight curvature, and is moderately broad at the base and
acuminate at the end. The sutural carina is not well-marked.
$ . The sutural carina of the head is distinct and the horn
is very short and blunt, generally having the appearance of
two coalescing tubercles.
This is allied to 0. igneus^ Vigors, but is generally larger
and more convex and the sculpture is stronger. In the male
the head is less produced both in front and behind. The
sexes of the new species differ little, and the less developed
males can only be distinguished from the females by a close
examination.
Onihophagus hronzeus, sp. n.
0. vivido affiais, sed obscure cupreus, elytris hand metallicis,
opacioribus, autennis pallide testaceis ; capite ruguloso, clypeo
vix producto ; prothorace ubique dense granuloso ; elytris planis,
subtiliter striatis, punctatis et setiferis.
d . Capite carina suturali curvata vix elevata raunito, postice
breviter cornuto, cornu postice inclinato, basi vix dilatato ;
prothorace antice paulo excavato et tuberculis duobus minutis
armato.
$ . Capite carina suturali curvata distincta antice raunito, cornu
postico perparura elevato ; prothorace tuberculis duobus vix
distinctis antice arraato.
Long. 12 ram.
Ilah. Nilgiri Hills [Hampson)^ Kanara [T. R. D. Bell).
This is a species very nearly related to the preceding one,
but deep bronze-coloured, with the elytra brown and quite
opaque. The prothorax is evenly granulated all over and
more finely than in 0. vividas, and the elytra are clothed
with a fine but distinct pubescence.
The horn of the male is more abrupt and scarcely widened
at the base, and the clypeus is scarcely produced in front.
0. pollicatasj Har., is another closely allied species, but
it is more elongate and the elytra are granulated and not
j)unctured.
I may note here that Harold’s supposition that 0. turhafnSj
430
Mr. G. J. Arrow on new
Walker, was described from a female of 0. sjnnifer^ F., is
not correct. The types of both are in the British Museum
collection and tlie two species are quite distinct, altho-ugh
nearly related, belonging to the group in which the male has
a single very long and slender horn curving backwards.
0. spinifer^ F., is steel-blue, with rugose elytra, and 0. tur-
hatus, Walker^ is coppery, with the elytra finely punctured.
(d) Head of the male without horn.
Onthophagus coeruleicollis^ sp. n.
Caeruleus, subtus nigro-cseriileus, pedibus piceis vel rufo-piceis ;
antennis elytrisque flavis, horum sutura fasciisque duabus trans-
versis interruptis nigris ; capite piano, dense rugoso, antice
product©, baud acuto, margine postico arenato, medio fere
angulato ; prothorace valde gibboso, ubique dense granulato,
postice lobato, fere angulato, lateribus valde arcuatis, angulis
anticis rotundatis, posticis obsoletis ; elytris subtiliter striatis,
iuterstitiis planis, rugose puuctatis et minutissimo setiferis ;
pygidio grosse punctato ; corpore subtus ubique punctate, parce
griseo-hirto.
Long. 5-5-8-S mm,
lldh. S. India, Dharwar, Belgaum.
Metallic blue, darker beneath, with the legs piceous or
piceous-red, the antennae testaceous and the elytra yellow,
with the suture and two transverse rows of spots black, the
spots of the posterior row sometimes coalescing into an
irregular band.
This is a small species, moderately elongate and very
convex. The head is small, flat, finely rugose, and without
horn or carina, and the clypeus is bluntly produced in the
middle. The posterior margin of the head forms a sharp
edgOj but is not produced. The prothorax is very globose,
finely granulated, and without armature or excavation, with
the sides strongly rounded, the posterior angles obliterated,
and the base lobed but scarcely angulated, the lobe indicated
by' a slight impression on each side. The elytra are finely
striated and the interstices rugosely' punctured and very
minutely setose. The pygidiurn and underside are coarsely
])unctured ■ and clothed with short greyish hairs. The front
tibiae are broad and strongly but very bluntly quadiidentate.
The sexes are apparently alike. There is no trace of horn
or carina, but a very minute tubercle sometimes traceablo
near the middle of the head probal)ly indicates the male.
This species and the following one belong to a group of
which several have been described fi’om Tropical Africa, and
of which 0. maculatuSy F., is the type. The elytra are less
rugose than is usual in this group.
Coleoptera from the Indian Empire.
431
Onthophagus regalis^ sp. n.
Laete rufo-cupreus, subtiliter flavo-setosus, corpore subtus obscure
ciipreo, elytris ochraceis minute nigro-maculatis, macula uiia
humerali, tiibus medianis quatuorque posticalibus, pygidio nigro ;
capite piano, undique transverse rugoso, antice paulo obtuse
producto, margine postico medio minute dentato, sutura clypeali
subtiliter carinata ; prothorace globoso, undique dense granuloso,
lateribus simplice arcuatis, angulis posticis obsoletis, margine
basali lobato, fere angulato ; elytris subtiliter rugosis, striatis,
interstitiis planis ; pygidio corporeque subtus dense punctatis,
bujus lateribus flavo-hirsutis ; tibiis anticis dentibus tribus
robustis munitis.
Long. 8-5 mm.
Hah. Ceylon.
Coppery-red, with the legs, pygidium, and underside
black or obscurely metallic, the antennae pale yellow, and the
elytra orange-red with about eight minute black spots on each,
one of them near the shoulder and the rest forming two
transverse lines.
The size and form are almost those of the last species.
The head is flat and finely transversely rugose, slightly
produced and recurved at the apex, with the clypeal suture
feebly carinated and an impression near the middle of the
posterior margin, which is slightly toothed. The prothorax
is very globose and finely granulated, with a slight impression
in front and one on each side of the posterior lobe, the hind
margin of which is very obtusely angular. The sides are
strongly curved and the posterior angles rounded off. The
elytra are finely striated and minutely rugosely punctured,
with an almost imperceptible yellow pubescence. Tlic
pygidium and underside are closely and deeply punctured
and the sides of the body are clothed with yellow hairs.
Specimens have been found by Mr. E. E. Green and
Col. J. W. Yerbury. I have found no external sexual
characters.
Onthophagus myrmecophilus^ sp. n.
Breviter ovatiis, depressus, politissimus, obscure cupreus ; elytris
nigris vel piceo-nigris, macula humerali plus minusve distiiicta
rufa ; corpore subtus seueo-nigro, nudo, pedibus ferrugineis ;
capite omnino Isevi, inermi, margine crebre punctato, paulo
reflexo ; prothorace subtiliter baud dense punctato ; elytris sub-
tiliter striatis, striis punctatis, interstitiis planis, Imvissimis ;
pygidio convexo, polito, minutissime punctato ; metasterni medio
polito, lateribus crebre punctatis ; pedibus brevibus, tibiis anticis
fortiter 4-dentatis, et supra minutissime serratis.
432
Mr. G. J. Arrow on neio
d . Capite ant semicirculari ant paulo ant valde prodncto, apice
minnte cmarginato ; prothorace antice valde retnso, dorso medio
late prominente, antice parnm impresso, fossa laterali sat pro-
funda, extns carinata, Carina antice pins minnsve acnte producta.
2 . Capite semicirculari ; prothorace aeqnaliter convexo, inermi.
Long. 4*5-6 mm.
JJdb, Nilgiri Hills (Barwood Estate).
Very smooth and shining, slightly coppery, with the
elytra black and marked with a more or less distinct reddish
patch on each shoulder. The form is broadly oval, rather
flat, and the legs are short. The head is very smooth,
without horns, caringe, or visible sutures, and the margin of
the clypeus is finely and closely punctured and reflexed.
The prothorax is finely punctured, with the sides regularly
curved, the posterior angles very obtuse and the base gently
curved. The scutellum is invisible. The elytra are very
finely punctate-striate, with the interstices flat and impunctate.
The pygidium is convex and ver}^ finely punctured, and the
underside smooth with the sides of the metasternum rather
closely punctured. The front tibiae are armed with four
strong teeth and very finely serrated above these. In the
four posterior tarsi the first joint is equal in length to the three
succeeding joints, but is not very much flattened.
$ . The head is produced in front, rather broad at the
apex, and very feebly bilobed. The prothorax has a broad
dorsal elevation shallowly impressed in front, and on each
side a longitudinal carina more or less acutely produced in
front, with a deep lateral cavity on each side between the
latter and the dorsal hump.
? . The head is semicircular and the prothorax quite
simple.
There is another male form in which the head is like that
of the female, although the thoracic armature may be well-
developed.
This little beetle is entirely peculiar both in habits and
appearance. It was found in some numbers by Mr. H. L.
Andrewes in a nest of the harvesting ant, PMdoJogiton
diversus, in a decaying Ficus trunk. The specimens were
taken frcm the ddbris accumulated near the entrance to the
nest, and Mr. Andrewes believes the beetle to breed in the
nest, which is probably the case. That its manner of life is
quite abnormal is sufficiently evidenced by its appearance,
which is entirely unlike that of the typical Onthophagus. The
round deq)ressed Ibrm, very shining surface, and short legs
produce, in the female at least, a strong suggestion of a
Ilisterid, but stiucturally it does not differ in any important
particular from the present genus.
Coleoptera fi'om the Indian Empire.
433
Appendix.
I liave' brought together here a few observations and
descriptions relating to certain other Oriental Lamellicornia
of tlie genera dealt with in the foregoing paper.
Cetoniidae.
Owing to the first part of this paper being very hurriedly
published and the proofs hastily corrected while absent from
town, it is necessary to rectify one or two oversights and
omissions occurring there.
I find that in re-characterIzIng the genera of the Heteror-
rhina group I have used the name Rhomborrhina in an
inadmissible sense. The type of that genus is not, as I at
first assumed, one of Hope’s species, but is stated by Hope
to be R. heroSj G. & P., one of the forms which I associated
under the name of Anomalocera. Rhomborrhina must there-
fore replace the latter name, and the group of species for
which 1 proposed to retain Hope’s genus must receive a
fresh one. These species are R. distincta^ Hope, with its
varieties flammea, Gestro, carianaj Gestro, ultramarineay
Nonfr., &c., opicalisj Westw., hyacinihina^ Hope, nigra,
Saund., japonica, Hope, and opalina, Hope. These, called
Rhomborrhina in my table (p. 350, supra), I now propose to
call
Torynorrhina, gen. nov.
Type, Rhomborrhina distincta, Hope.
Its distinctive characters are : — Head and margin of clypeus
entirely without prominences in both sexes, the clypeus
widening towards the front and its anterior margin broadly
rounded ; sternal process short, broad, widened in front of
the middle C3X£e and broadly rounded in front; front tibiae
simple and unarmed in the male, bidentate in the female ;
hind legs simple and straight in both sexes; pygidium
pubescent.
I will also formulate here in rather more detail the
characters of the other new genus established in the table
just mentioned : —
Euchloropus, gen. nov.
Type, Cetonia Iceta, Fab.
Head and margin of clypeus entirely without prominences
in both sexes, the clypeus parallel-sided, wdtli the front
margin straight ; sternal process slender, recurved, and
434
Mr. G. J. Arrow on new
pointed; front tibise simple and unarmed in the male, biden-
tate in the female ; hind tibise of the male strongly curved,
with a thick fringe at the inner edge of the posterior part;
those of the female simple.
I omitted to include in my description on p. 350 any reference
to the sexes of the new species Macronota gracilis. This is a
form allied to M. antennata^ Wallace, and was described
from a specimen of each sex. These differ very little, but
the male has the prothorax rather narrower than that of the
female, and the basal lobe more pronounced, and the front
tibise are rather slenderer and their teeth more equal.
The sexes in this genus often differ considerably both in
form and colouring, and have in several cases been described
as different species. I am not aware that it has yet been
recorded that iMacronota sculpticollis^ Thoms., is the female
cf M, 4i-vittata, Schaum, although the insect is common in
Ceylon and the fact has long been recognized. Still more
dissimilar are the sexes of M. Oherthuri^ Lansb., of which
M. liumilisj Lansb., described at the same time and from the
same place, is the female. I have seen the two forms
together in various collections, the male being silky black
with white markings, and the female dull ashy brown. It is
not common, but I associate the two forms without any
hesitation because the closely-allied M. crucicollis^ Lansb.
{—favosparsa^ Wat.), of which I have seen a large number,
has an exactly similar female. In a work in preparation I
hope to give the sexual distinctions of all the Indian species
of this genus.
Another genus in which marked diflferences between the
sexes occur, but have not hitherto been noticed, is Glycyphana^
in which also several so-called species have to be eliminated
in consequence. In his description of G. hinotata^ G. & P.,
Burmeister noted that his examples were females, and all
that I have seen of that form are of the same sex. It occurs,
however, in conjunction with another form generally regarded
(perhaps wrongly) as G. torqaata^ F., and of this our
speeimens are all males. The two differ only in their
coloration, the males having a row of four spots (instead of
only two) across the middle of the elytra, while the lateral
patches upon the pygidium are generally yellow instead of
blood-red.
A still greater difference occurs between the two sexes of
G. regalis^ Volk, of which the female ((r. celehensis, Wall.)
has the pygidium entirely black, while in the male it is
entirely orange, and the elytra have two large lateral patches
435
Coleoptera from the Indian Empire.
instead of a row of four. In the male also the scutellum is
red and each elytron has a longitudinal red stripe. In
G. Forsteni^NoW.^ which is a variety of the male of the same
species, these red markings are absent, as in the female, but
the yellow colouring is that of the male. G. hella^ Wall.,
again, wdiich has a red pygidium and red markings upon the
elytra, is the male of G. A-guttataj Volk, which is without
these.
G. andamanensis^ Jans., is another species with a very-
well-marked colour-dimorphism. A black form was described
by Thomson as G. andamana, and Kraatz recorded that this
was a variety of Janson’s species; but it has escaped obser-
vation that this is the female form, the male being invariably
green, at least in a very larg^e series which I have examined.
The two specimens from which Mr. Janson described
G. suhcinctay another Andaman species, prove to be males,
and the type of G. himaculata^ Kraatz, which I have also
been allowed to examine, is also a male of the same insect.
I have seen two females, which jpppear to me identical in all
structural features, but have an additional spot on the anterior
part of each elytron, a row of four across the middle, and a
patch on each side of the pygidium. I am inclined to regard
this as the other sex of G. suhcincta^ Jans. It appears to
correspond with the description of Getonia torquataj Fab.,
but as tlie locality of Fabricius’s insect (now in the Copen-
hagen Museum) is unknown, only a careiul comparison can
decide the point.
Glycyphana lateraHsf Wall., is only a colour- variety of
G> perviridisy AVaik, of which both sexes in normal specimens
are green. The single type specimen of the variety is a
female, in which sex the pygidium is marked by a broad
furrow.
I may perhaps note here that the Australian SchizorrhinF''
pidchra, Mach, is a species of Glycyphana.
In describing Ileterorrhina horneensis and mitrata from
female speclriiens, Mr. A. K. AA^allace suggested that the male
might prove to be similar to H. dives^ Westw. Both sexes
of the first species are now in the British Museum collection
and, contrary to expectation, there is no difference in the form
of the head, but the male of II. mitrata is still unknown. So
exactly does the female correspond with the figure of the
unique II. dives in Gory and Percheron’s Monograph, that I
regard it as extremely probable that they are tlie same.
Schaum recorded in .1849 that he had seen a female of
II. dives (apparently II. mitrata) in the Linnean Society’s
collection, but this cannot now be found.
43G
Mr. G. J. Arrow on new
Melolonthidse.
Tlie name Dejeania lias been used in tliree different
orders of Insects, the genus having the priority belonging to
the Diptera. The Coleopterous genus should be called
.DichelomorpJia, that name liaving been given by Buvmeister
to a Chinese species [D ichdomorpha ochraceo) placed by him
among the Ilopliinee, but allied to the insect I have described
in the present paper. The type of Blanchard’s genus
{Dejeania alsiosa) is an insect of very different appearance,
but these extreme forms are linked together by the new
species {D, lineata^ Arrow) and others described and un-
described, and all may best remain in one genus until the
time comes for a comprehensive study of them.
Hybosoridae.
PhcEodiroops gigas^ sp. n.
P. hidico proxime affinis, major, minus grosse punctatus ; prothoracis
lateribus baud denticulatis, vix arcuatis, medio leviter angulatis,
angulis posticis magis acuminatis, elytrorum costis paulo laevi-
oribus, magis elevatis ; abdomine magis rugoso ; tibiis 4 posteri-
oribiis hand transverse carinatis.
Long. 15 mm.
Ilah. Borneo, Pengaron ; Perak.
Specimens were found by Mr. Doherty in both the above
localities. They closely resemble the new Indian species,
but are larger, the puncturing of the upper surface is a litt'e
less dense, the prothorax is less uniformly curved at the sides,
which are not denticulated, and the hind angles are more
evident. The elytral costae are a little more elevated and
the hind tibiae are not interrupted at the middle of the outer
edge by a transverse carina.
Phceodiroops acuticollis^ sp. n.
Statnra pracedentium sed opacior, rugosius punctatus, magis regu-
lariter setosus ; prothorace cmnino crebre punctato-rugoso,
lateribus distincte eleYatis, fere red is, angulis posticis paulo pro-
ductis ; eljtris densissimc punctatis, singnlo fortiter tricostato,
costis lacvibus, sat parce lufo-setosis ; tibiis 4 posterioribus baud
transYcrse carinatis, posticis gracilibus.
Long. 14 mm.
JIah. Borneo, Kina Balu.
A single specimen found by Mr. Doherty is in the Biitish
]\Iuseum collection. It is more densely sculptured than either
of the preceding species, the hind angles of the prothorax are
437
Coleoptera from the Indian Empire.
sliglitly produced, the costae of the elytra are sliarp, and the
long reddish hairs are situated at regular intervals upon
them.
Copridae.
Cassolas peninsularisj sp. n.
Latus, convexus, nigro-cupreus, pedibus castaneis, capite crebre
pimctato, qiiadridentato ; prothorace subtilifcer punctato, lateribus
valde arcuatis, angulis posticis fere obsoletis ; elytris imrnacu-
latis, striatis, striis subtiliter puuctatis ; pygidio fortiter puric-
tato; tibiis anticis serratis, dentibus tribiis validis, tarsoruin
posteriorum articulis fere aequalibus.
S . Tibiis anticis subtus unispinosis, dente apicali lato, truncate ;
tibiis posticis gracilioribus, sinuatis, intus denticulatis.
Long. 4-5 mm.
Hah. Perak, Penang.
This is like the Indian species, but rather broader in form
and without the humeral spot. It is also less strongly
punctured, and in the structure of the legs peculiar to the
males there are remarkably pronounced differences between
the two.
Onthophagus huffalo^ sp. n.
Niger, robustus, elytris corporeque subtus breviter brunneo-setosis ;
capite sat lato, subtiliter rugoso-punctato, margine reflexo, haucl
excise ; prothorace fortiter punctato, marginato, postice pro-
fun de longitudinaliter sulcato, lateribus sinuatis, angulis posticis
valde obtusis ; elytris striatis, interstitiis convexis, dense punc-
tatis ; pygidio fere rugose punctato ; metasterni medio nitido,
parce sat grosse punctato.
(S . Capite carinato, postice cornubus duobus curvatis, baud longis
armato, cornubus basi carina medio leviter dentata connexis ;
prothoracis dorso elevato, antice recte truncate, lateribus valde
depressis, fere impuuctatis, nitidis, marginibus valde sinuatis,
antice contractis, baud productis ; tibiis anticis paulo elongatis.
$ . Capite Carina suturali aliaque frontali medio dentata armato ;
prothorace carina transversa distincta munito.
Long. 11-13 mm.
Hah. Java, Borneo, Labuan.
This is very closely related to 0. rugalosus, Har., and
0. productuSj Arrow, but the posterior part of the prothorax
is less shar[,tly carinated and more deeply sulcated longi-
tudinally in the middle. In the male the prothorax is less
narrowed and produced in front, and the anterior angles are
not hollowed out. The head in the same sex is less narrow,
it has a distinct carina upon the clypeal suture, and the
carina connecting the horns (which are of the same form) is
438 On new Coleoptera from the Indian Empire.
dentate in tlie middle. The elongation of the front tibise of
the male is much less marked.
Rufo-cupreiis, elytris nigris ; capite rugoso, absque carinis trans-
versis, antice producto ; prothorace paruin convexo, basi medio
leviter angulato ; elytris sat opacis, subtiliter striatis, interstitiis
minute punctulatis ; pygidio crebre punctato ; metasterno ubique
punctato, antice paulo acuminato ; tibiis anticis minute serratis,
dentibus 4 validis.
S . Corpore supra et subtus breviter fulvo-setoso ; clypeo dilatato,
antice excavato, Imvigato, profunde emarginato, medio longe et
anguste producto, processu late bifido, recurvato, intra oculos
tuberculo parvo, compresso ; prothorace confertim granuloso,
medio antice anguste longitudinaliter carinato, postice medio
obtuse angulato ; an tennis perinsignibus, articulis 4 -6 brevibus,
latis, 7 cupuliformi, 8 biramoso, 9 multiramoso.
5 . Corpore fere nudo, supra et subtus subtiliter punctato ; clypeo
leviter rugoso, antice paulo producto, integro, inermi ; antennis
simplicibus ; prothorace postice medio dentato.
Long. 10-12 mm.
Hah. E. Borneo, Pontianak.
Reddish bronze, with the elytra black, and not very convex
or shining. The male has the surface finely setose above
and below, and the head broad and rugose, except in front,
where it is smooth and hollowed, with a strong emargination
Onthophagus egregius, sp. n.
Head and antennae (seen from above and below) : enlarged.
in front, from the middle of which springs a slender process
curving upwards and bifurcated broadly at the end. There
is a slight longitudinal carina between the eyes. The pro-
thorax is granulated, longitudinally carinated in the middle,
with a slight angulation at the middle of the hind margin.
The elytra are finely striated, with the interstices closely and
Onthophagus egregius^ sp. n.
On a remarkable Mountain Viscacha, 439
finely punctured, and the pygidium and underside are strongly
punctured.
The female is more finely punctured and almost devoid of
setae, with the head simple and slightly pointed in front.
Tlie prothorax is minutely and not closely punctured, without
a median carina, and is rather more sharply angulated at
the middle of the base. The elytra and underside are also
more finely punctured than in the male.
The species is remarkable for the extraordinary structure
of the antennae characteristic of the male, which is quite
unlike anything hitherto known. The last three joints of
the footstalk are broad and close-fitting, and the seventh joint
forms a hollow half-hemisphere, covering the two remaining
joints as seen from beneath. The eighth joint sends off a
slender branch on each side, and each branch is toothed
underneath ; and the last joint has a slender footstalk, which
gives off two similar toothed branches on each side, dimin-
ishing in length, and is broad at the end, partly shutting
in at the extremity of the antenna the whole complicated
apparatus.
The antennae of the female are quite normal.
A series of specimens, chiefly males, have been obtained
by M. Rene Obertliilr’s collectors.
LT. — On a remarkable Mountain Viscacha from Southern
Patagonia, with Diagnoses vf other Members of the Group.
By Oldfield Thomas.
The British Museum owes to its generous and indefatigable
contributor, Mr. J. A. Wolffsohn, C.M.Z.S., three examples of
a fine Viscaccia^ from a locality far south of any place where
members of this group have been p)eviously recorded, ddie
animal is quite distinct from any hitherto known, and I
* In his paper on the nomenclature of this group, Dr. F. Lahille
(An. Soc. Cient. Argent. Ixii. p. 39, 1906) seems to have come to quite
correct conclusions, including the allocation of Viscaccia to the Mountain
Viscachas, formerly known as Lagidium. But with the fate that has
as yet always befallen writers on this most difficult and complicated
subject, he has made a mistake in crediting the name Viscnccia to Molina,
for although the latter did say in 1810 that the animal ought to have a
special geueric name, he did not give it one, only using Viscaccia in such
a way that it cannot be distinguished from the vernacular term.
However, fortunately, Oken in 1816 used the term Viscaccia in a techni-
cally valid manner, with Legus chilensis ’’ as its type, choosable as
such whether by elimination or by the first-species rule.
440
Mr. 0. Thomas on a
propose to name it in lionour of its donor, to whom tlie
JMuseum is indebted for series of all the mammals of Central
Chili.
Viscaccia Wolffsohni, sp. n.
A large species with long fur strongly suffused with orange ;
tail very bushy.
Size large, form stout and robust, the size appearing even
larger than the truth owing to the very long rich fur, of which
the wool-hairs are over 35 mm. in length and the long hairs
upwards of 50 mm. General colour above of head and body
approaching clay-colour (Ridgway), though brighter and
clearer; the wool-hairs, which give the predominating colour,
brownish slaty for four- fifths their length, their ends creamy
buff darkening to clay-colour. Below, on the lower cheeks,
throat, chest, and belly, the terminal colour becomes richer
and redder, attaining almost to ‘Gawny.” A distinct white
spot on each axilla and on each side of the inguinal region.
Dorsal dark line little marked. Head like back. Ears
comparatively short, thickly and closely haired, their backs
black, their inner surfaces with whitish hairs, and there is a
marked line of creamy-tipped hairs running across their bases
above. Arms with tawny-tipped hairs, the tawny or yellowish
colour extending to the tips of the toes. Hind limbs duller,
more browmish clay-colour; the feet very large and heavy.
Tail far finer than in any other form, immensely bushy, the
hairs of its dorsal crest attaining over 150 mm. ; in colour
the upper, crested side is mixed black and buff or ochraceoiis
buff ; underside black, finely grizzled with glossy ochraceous
buff.
Bkull comparatively large and heavy ; nasals expanded in
front; palatal foramina rather short; bullre not so much
swollen as usual.
Dimensions of the type (measured in the flesh) : —
Head and body 470 mm.; tail 305 ; hind foot 107 ; ear 65.
Skull: greatest length 91 ; basilar length 75*5 ; zygomatic
breadth 50; breadth of brain-case on parietal bones 34*2;
length of upper tooth series (crowns) 22.
Hah. Sierra de los Baguales y de las Vizcachas, lat.
50° 50' S., long. 72° 20' W., on the boundary between Chili
and Argentina.
Type. Adult female. B.M.no. 7. 4. 5. G. Original number
277. Collected 1st February, 1907, by Mr. John A. Wolffsohn.
Three specimens.
In the interesting account which Mr. W olffsolm has
sent me of the capture of these specimens, he states that
remarhahle Mountain Viscacha.
441
Mr. Ferrier_, who owns a farm in that district, at the foot of
Mount Payne, says that the Sierra de los Baguales is known
to be the most southern part in which Vizcachas occur, and
between that range and much farther north there are none
at all.”
The species is readily distinguishable from all other
members of the genus by its large size, rich colour, long fur,
immensely bushy tail, and short black ears.
With characteristic modesty Mr. Wolffsohn has suggested
that the species should be named after Mr. McClelland, the
President of his Company, to whom he owed the pleasure of
his trip to Patagonia, but in view of the immense amount
of help w^e have received from Mr. Wolffsohn, I have
ventured to disregard his request, and to name this fine
animal after him.
AVhile working out Viscaccia Wolffsohni I have found the
different races of this genus, of which the British Museum
possesses a large number of specimens, to be both numerous
and yet locally so constant as to deserve recognition by name.
The fine series obtained by the late Mr. Perry O. Simons
during his exploration of the Andes is especially rich in
noteworthy forms.
Asa preliminary I propose to give short diagnoses of such
as I believe to be new, hoping to return to the subject later,
when still further material is available.
Of the old names : —
Viscaccia. viscaccia, Molina, is the large deep grey animal
from the Chilian Andes, with a short ill-defined dorsal line,
yellow belly, the tail with long grey crest and with its
underside, although blackish, not sharply defined black.
V. Cuvieri, Benn., and, more doubtfully, V.pallipes, Benn.,
appear both to be referable to a strongly yellowish form
found in Northern Chili, localized specimens from Iquique
and Tarapacd being in the Museum.
F. peruana, Meyen, I cannot certainly identify, as of none
of the specimens before me can it be said that auf dem
Bauche, besonders zwischen den hintern Extremitaten, sind
die Haare zierlich weiss gefarbt,” all having the belly of a
more or less yellow colour, which is generally deepest in the
inguinal region. Some form agreeing with Meyen^s descrip-
tion will no doubt yet be found.
In the absence of exact knowledge of the local relationship
of the different forms to each other, I provisionally use
binomial names for all.
Ann. & Mag. N. Hist. Ser. 7. Vol. xix.
31
442
Mr. 0. Thomas on
Viscaccia inca, sp. n.
Pale grey (between grey no. 8 and smoke-grey), without
or with very faint trace of dorsal stripe. Wool-hairs about
30 mm. Belly well-defined pale yellow, the axillary white
spots distinct. Feet creamy white. Tail not sharply bi color,
the underside only slightly darkened proximally.
Hind foot (on skin) 85 mm. ; skull, greatest length 81.
Hab. Junin, Peru. Type from Incapirca, Zezioro.
Type. Adult female. B.M. no. 94. 8. 6. 20. Collected by
J. Kalinowski, 8th May, 1890. Three specimens.
Viscaccia arequipoe, sp. n.
Posterior body grey like V. inca.^ but the fore-back and
shoulders more buffy. Dorsal line present, though incon-
spicuous. Wool-hairs of back about 30 mm. Undersurface
pale yellow. Feet white. Tail distinctly and sharply bicolor,
the under surface and end black.
Hind foot (measured fresh) 92 mm. ; skull, greatest
length 79.
Hah. Sumbay, near Arequipa. Alt. 4000 m.
Type. Subadult male. B.M. no. 0. 10. 1. 93. Collected
4th July, 1900, by P. O. Simons. Four specimens.
Viscaccia suhrosea^ sp. n.
The grey distinctly suffused with pinkish buff, the re-
sulting general colour approaching broccoli-brown.^’ Dorsal
line scarcely perceptible. Under surface inclined to pink,
nearest to ‘^salmon-buff.” Feet pinkish buff. Tail not very
sharply bicolor, although more so than in V, inca.
Hind foot (fresh) 85 mm. ; skull, greatest length 77.
Hah. Galera, W. of Oroya, Dept. Lima. Alt. 4800 m.
Type. Adult female. B.M. no. 0. 7. 7. 53. Collected
24th February, 1900, by P. O. Simons. Four specimens.
Viscaccia saturata^ sp. n.
Colour much darker than in the previous species, an olive-
grey nearly as dark as Ridgway’s “ olive,” sometimes
marbled with black along the dorsal area ; no distinct dorsal
line. Wool-hairs about 26-27 mm. in length. Under
surface cream-buff. Feet white. Tail distinctly bicolor, the
under surface glossy black. Skull broader than usual, the
nasals much swollen.
Hind foot (fresh) 95 mm.; length of skull 80.
443
new Species of Viscaccla.
Ilah. Limbane, Inambari, Dept. Puno. Alt. 3500 m.
Type, Adult male. B.M. no. 1. 1. 1 . 52. Collected 26th
July, 1900, byP. 0. Simons. Three specimens.
Viscaccia punensis^ sp. n.
Size small. Colour more brownish grey, the area across
tbe shoulders specially brown. Dorsal line fairly distinct.
Wool-hairs about 24 mm. Under surface dull creamy
whitish, the brown basal part of the hairs less hidden than
usual. Axillary white spots well marked. Feet white.
Tail sharply bicolor, the light part of the crest nearly white;
the under surface and end black.
Hind foot (on skin) 90 mm. ; length of skull 76*5.
Hah. Puno, Lake Titicaca. Alt. 3800 m.
Type. Adult female. B.M. no. 97. 10. 3. 51. Collected
1st April, 1896, by J. Kalinowski. Six specimens.
Viscaccia cuscus, sp. n.
Size large. Colour deep grey, about grey no. 5 ; dorsal
line very distinct, black, about 300 mm. in length. Wool-
hairs about 23 mm. Under surface ^‘bufF” or “pinkish
buff^’ ; axillary white patches unusually large. Feet cream-
colour. Tail grizzled grey, not markedly bicolor, the prox-
imal part of the under surface above blackish.
Hind foot (fresh) 102 mm. ; length of skull 91.
Hab. Paratani, Bolivia (about 66° W., 17° 5' S ). Alt.
2600 m.
Type. Adult female. B.M. no. 2.1.1.104. Collected
9th April, 1901, by P. O. Simons. One specimen.
Viscaccia lutea, sp. n.
Colour between cream-buff and clay-colour ; dorsal line
fairly distinct, about 150 mm. in length ; wool-hairs about
29-30 mm. Under surface cream-buff ; the axillary spots
well marked. Feet creamy. Tail bicolor, the upper crest
mixed with blackish.
Hind foot (on skin) 95 mm. ; length of skull 85.
Hah. Esperanza, Sahama, Bolivia. Alt. 4000 m.
Type. Adult male. B.M. no. 98.3.16.22. Collected
12th May, 1897, by Gustav Garlepp. Four specimens.
Viscaccia perlutea^ sp. n.
Like V. lutea^ but the colour throughout a richer buffy.
Face clearer grey. Dorsal line very strong, nearly 200 mm.
31*
444 Occurrence of the Edible Dormouse in Sardinia.
in length ; wool-hairs about 28 mm. Under surface rich
buff instead of cream-buff. Feet cream-buff. Tail bicolor,
not sharply defined, the light hairs of the crest yellowish,
instead of the usual greyish white.
Hind foot (fresh) 100 mm. ; length of skull 82.
Hah. Pampa Aulliaga, Bolivia (67° W., 19° 30' S.). Alt.
3800 m.
Type. Adult male. B.M. no. 2. 2. 2. 89 bis. Collected
22nd October, 1901, by P. 0. Simons. Two specimens.
Viscaccia tucumana^ sp. n.
Colour clear grey, with scarcely a trace of drab ; dorsal
stripe short, fairly distinct, rather over 100 mm. in length.
Hairs of back about 24 mm. in length. Under surface
buff and cream-buff; the axillary white spots distinct. Feet
creamy. Tail not markedly bicolor ; the crest intermixed
black and greyish white, the underside black proximally,
greyer distally.
Hind foot (on skin) 91 mm. ; length of skull 85.
Hah. Cumbre de Mala-Mala, Sierra de Tucuman. Alt.
3000 m.
Type. Adult female. B.M. no. 5. 10. 29. 6. Collected
10th April, 1904, by L. Hinelli. Six specimens.
LTI. — -On the Occurrence of the Edible Dormouse {Genus
Glis) in Sardinia. By Oldfield Thomas.
Up to the present time no trustworthy record has existed of
the occurrence of the edible dormouse either in Sardinia or
Corsica, such references to ‘‘ Ghiri ” as have been published
possibly, indeed probably, referring to Eliomys.
From the palaeontological list Glis is also absent*; but
Dr. Forsyth Major has been good enough to inform me that
he knows of an undoubted Glis from the Pleistocene of
Corsica, a fact which has an important bearing on the question
as to the really indigenous position of the animal I am about
to describe.
The British Museum owes to Sr. Giuseppe Meloni, of
Lanusei, Eastern Sardinia, a number of examples of a
dormouse discovered by him in the neighbourhood of that
place, and a careful comparison with a series of Glis italicus
from various parts of Italy convinces me that the Sardinian
form is distinct enough to require a name. I therefore
propose to term it
Willi the, exception of a reference by Studiati in T.a Marmora’s
^ Geology of Sardinia,’ and this may, again, be very possibly an Ulwfnys.
On a new Genus and Sj.)ecies of Muscida3.
445
Glis Melonii, sp» n.
Like G. italicusy but greyer in colour and with a different
tail.
General characters and size as in the Italian species, the
upper premolar small and comparatively simple as in that
animal. Colour, instead of being drab or drab-grey as is the
case in italicus, clear grey, without drab suffusion, approxi-
mating to grey no. 6 of Ridgway. Tail very thick and
bushy at the base, tapering terminally to the comparatively
narrow tip ; quite evenly broad throughout in G. italicus.
In colour, while at least the terminal half, and sometimes
more, of the tail in italicus is brown or blackish, in Melonii
only the tip is darkened, the grizzled grey extending for from
two thirds to three fourths of the length of the tail ; the tip
itself is also darker, often really black ; under surface with
the usual white line along the proximal two thirds.
Skull and dentition as in G. italicus.
External dimensions apparently about as in G. italicus,
but the skins seem to be stretched, and no useful purpose
would be served by measuring them.
Skull of type : greatest length 4^ mm. ; basilar length 34;
greatest breadth 35 ; nasals 13* *5; interorbital breadth 5*5 ;
length of upper tooth series 7*6.
Hah. (of type). Marcurighe, Urzulei, Ogliastra, Sardinia..
Other specimens from Monte Nieddu.
Type. Adult male in British Museum. Collected by
Sr. Giuseppe Meloni. Twelve specimens examined.
Signor Meloni informs me that this dormouse is only found
in the Forests of Urzulei and Orgosolo, while it appears to
be quite absent from the other forests of the island.
LIII. — A new Genus and Species of Phlebotomic Miiscidge.
from Aden. By Ernest E. Austen.
Stomoxybinjs.
Stygeromyia *, gen. nov.
In some respects intermediate between Stomoxys, Geoff.,
Bcematobia'\ j Rob.-Desv.,and Lyperosia, Rond, (sensu Bezzi).
* (TTvyepos, hateful ; fxv7a, a fly.
t The present writer cannot agree with Speiser (Zeitschr. f. wiss.
Insektenbiol. Bd. i. (1905) p. 461), whom he regrets to see has recently
been followed by Bezzi (“ Mosche Ematofaghe,” Rendiconti del R. 1st.
Bomb, di sc. e lett., serie ii. vol. xl. 1907, p. 17 [sep. imp.] ; and ‘ Katalog
UG
Mr. E. E. Austen on a neio
Li general appearance and form of body similar to Stomoxys^
but in shape of proboseis and palpi resembling Ilcematohiaj
though with arista feathered only on upper side, as in Sto-
moxys and Lyperosia. — Head somewhat flattened from front
to rear, with basioccipital region only slightly swollen ; pro-
boscis short, stout, and shining, of uniform thickness through-
out, not tapering to the tip, chitinous, but terminated by a
pair of small fleshy labella ; palpi equal to proboscis in
length, large, clavate towards the tips, curving upwards, and
with stout bristles on the outer side at the distal extremity.
Bristles of thorax : — Humeral^ 3. Post-humeral^ 1. Noto-
pleural^ 2. Prcesutural, 1. Supra-alar^ 1. Intra-alar^ 1.
Post-alar j 2. Dorso-central, 6 (1 in front of and 5 behind
the suture). Inner dorso-central ^ 1. Scutellar^ 4 (1 prse-
basal, 1 basal, 1 discal, 1 apical) Mesopleuraly 9 or 10,
wider apart than in Stomoxys or Ilcematohia. Stern opleural^ 1
(posterior, as in Stomoxys, instead of 1 : 1, as in Ilcematohia) .
Wings with first posterior cell narrowly open at the tip,
the width of the opening being precisely that seen in the same
cell in the wing oi Musca corvina, Fabr., and less than half
der Palaarktischen Dipteren,’ Bd. iii.), in transferring to this genus, from
its time-honoured position among the Tachiuinse, Meigen’s name Siphona.
So far from Meigen’s diagnosis, published in 1803 (Illiger’s ‘ Magazin fiir
Inselitenkunde,’ Bd. ii. p. 281), applying just as well to Hmnatohia, Ilob.-
W&y., to Bucentes, Latr. { = Siphona, Mg., Syst. Beschr. iv. (1824),
p. 154, et aiict.), as erroneousl}^ asserted by Speiser, the statements con-
cerning the bare arista and “ gebrochen ” proboscis show that it does
nothing of the kind. On the contrary, it must be evident to any unbiassed
investigator that, when writing his diagnosis of 1803, Meigen had in view
identically the same genus as that subsequently characterized by him in
greater detail in the volume of the ‘ Systematische Beschreibuug’ pub-
lished in 1824, and illustrated in tab. xxxvii. of that work, tigs. 18-25.
Strangely enough, in view of the course that he has seen tit to adopt, this
contention is actually advanced by Bezzi (toe. cit. pp. 17-18, sep. imp.) !
In dealing his perfectly gratuitous blow at the stability of Muscid nomen-
clature, Speiser relies chietly on the fact that Meigen appended to his
1803 diagnosis of Biphona, as “ type ” or ‘^example,” the name “ Stomoxys
irriUms, Fabr.,” which is now admitted to be a synonym of Ilcematohia
{Stomoxys) stimulans, Mg. This argument, however, goes for naught in
view of the statements in the diagnosis itself, to which attention has
already been drawn ; and, as Bezzi remarks {loc. cit. p. 18, sep. imp.),
there appears to have been some mistake as regards the species given as
the type of the genus Siphona. It may well be that ‘‘ irritans,’’’ Fabr.,
was simply a layisus calami for “ minuta'd Fabr., since in Syst. Beschr. iv.
p. 155, Stomoxys minuta, Fabr., is given by Meigen himself as a synonym
of Sipho7ia (Mnsca) yeniciilata, Beg., the species which there follows
immediately after the detailed description of the genus Siphona.
* For the nomenclature of the bristles of the scutellum, cf. Girschnor,
“ Ueber die Scutellarbeborstung der Musciden,” Wiener entoinologische
Zeitung, xx. Jahrg. (1901), pp. 71-72, Taf. i. tigs. 4-7.
447
Genus and Species o/Muscidoe.
of that exhibited by the first posterior cell in the case of
Stomoxys calcitrans^ L. ; apical portion of fourth vein beyond
the bend perfectly straight^ not bent inwards at the extremity,
as in Stomoxys and Hcematohia.
Typical species, Stygerornyia maculosa^ sp. n.
Stygeromyia maculosa j sp. n.
S . Length mm. ; length of wing 6^ mm. ; width of
head mm. ; width of front at narrowest part ^ mm., slightly
wider at vertex j length of proboscis from base of chitinous
• portion to tip of labella 1^ mm.; width of thorax at trans-
verse suture 2^ mm. ; width of abdomen at hind margin of
second segment 2| mm.
Entirely smoke-grey striped and spotted with hromn^
bristles and hair entirely black palpi and tibice orange-buff
(anterior surface of hind tibiae suffused with grey) ; wings
hyaline^ basal portion of third vein with some 7 or 8 bristles,
wide apart and exceedingly minute.
Head. — Face and front yellowish silvery, posterior surface
grey ; ground-colour of face ochraceous ; frontal stripe
walnut-brown, narrovy, slightly narrower than orbits in middle,
expanding above and below, lower extremity enclosing usual
silvery spot above base of antenna; antennce^ first and second
joints and base of third joint on inner side below orange-
rufous, remainder of third joint and arista dark brown ; hairs
on upper side of arista long, 11 or 12 in number ; palpi with
short black bristles on outer side on distal two-thirds, three
or four bristles at extreme tip of each palpus longer than
remainder, and very conspicuous when insect is examined under
a lens ; middle portion of lower margin of each palpus beset
with a series of fine and rather long black hairs ; proboscis.
dark chestnut-brown. Thorax. — Brown markings on dorsum
as follows : — a pair of narrow admedian stripes extending
from anterior margin across transverse suture and terminating
at one-fourth the distance between suture and prsescutellar
furrow ; a somewhat lighter-coloured median stripe., which,
commencing a short distance in front of the inner dor so-central
bristles, disappears close to hind margin, and reappears again
as a faintly marked fleck on base of scutellum ; two broader
spots on each side near transverse suture, one in front of
suture, immediately behind post-humeral bristle, the other
behind suture, between supra-alar bristle and the dorso-central
row, and nearer to the latter; the last-mentioned spot is
^ For names of colours see Ridgway, ‘ A Nomenclature of Colors for
Naturalists’ (Boston : Little, Brown, and Company, 1886).
448
Bibllograjyhical Notices,
somevvliat elongate, and all the thoracic markings, except the
short posterior median stripe and the fleck on the base of the
scutellum, are somewhat shining : tlie general arrangement of
the thoracic markings, except for the presence of a fleck on
tlie base of the scutellum, is similar to that seen in tlie case
of the $ of Iloimatohia stimulans, Mg. Abdomen rounded,
the upper surface strongly arched ; first segment unspotted ;
second^ third, and fourth segments with an elongate median
spot of mummy-brown^ and a lateral spot on each side) the
median spots on the second and third segments are guttate,
and that on the second segment is in contact with the front
margin, but does not quite reach the hind margin ; the
median spot on the third segment is not in contact with either
front or hind margins; the same spot on the fourth segment
takes the form of a narrow median stripe extending from the
front margin to a point one-third of the length of the segment
from the hind margin ; the lateral spots on the second segment
are the largest of all and transverse. Legs. — Femora greyish,
except tips, which are orange- buff ; front tarsi cinnamon-
rufous, middle and hind tarsi darker, last joint of all tarsi
ferruginous.
One specimen. Little Aden, Arabia. 8. ii. 1895. {Lieut.-
Colonel Yerhury.) Type in British Museum (Natural
History).
The British Museum is indebted to the generosity of
Colonel Yerbury, whose remarkable energy as a collector of
Hiptera is well known, for the enrichment of its collection
with the type of the new genus and species described above.
Although the donor has unfortunately been unable to find in
his diaries any reference to the habits of the insect, there can
be no doubt, from its evident affinities and from the shape
of its proboscis, that Stygeromyia maculosa is a blood-sucker ;
and it is to be hoped that we may ere long receive, in addition
to a series of specimens of both sexes, full details as to the
bionomics of this striking addition to the Stomoxydinae.
BIBLIOGRAPHICAL NOTICES.
Fishes of Japan, an Account principally on Economic Species. By
Keinosukb Otaki, Tsunenobu Etjjita, and Tadashi Higuhasui.
Text roy. 8vo ; plates imper. 4to. Tokyo, 1903-7. (Publishers :
Shokwabo, Tokyo.)
This work has been in progress for several years. Excellent in its
plan, object, and execution, it is characteristic of the thoroughness
with which the Japanese Naturalists have mastered western
449
Blhliograpliical Notices,
scientific methods, as well as of their national aptitude for rendering
their work generally useful by combining practical instruction with
purely technical detail.
The text of each number consists of two separate portions : the
first, which is written in English, contains a technical description ot*
each species with an account of its distribution in Japan, its habits,
propagation, economic value, and, in a condensed form, of the
implements and modes of capture. The second portion is a
Japanese reproduction of the former, but in it the authors enter
more fully into such details as possess a special local interest ; it is
illustrated by numerous text-figures of the implements and
apparatus employed by the Japanese fishermen, many of which are
very ingenuous, while a few are admittedly foreign importations.
Some spirited sketches of fishing-operations cannot fail to delight
the fisherman’s heart, and a study of them may be useful or give
new ideas to the expert of other lands.
The figures of the fishes have been taken from fresh specimens
and are most faithful representations ; although every attention is
paid to an accurate delineation of structural details, ichthyologists
will readily recognize in them an artistic family-likeness to the
illustrations in Siebold’s ‘ Fauna Japonica.’
The authors do not hamper themselves with a systematic
sequence of the species admitted in their work. Of the four parts
that have appeared up to the present, the first (1903) treats of
Lateolahrax japonicus^ Latilus sinensis, Thynnus affinis, and Seriola
quinquei'adiata. The importance of the last two may be gathered
from the fact that in 1899 the catch of the Tunny amounted to
79,124,002 pounds, valued at about .£394,000, and that of the
Seriola in 1901 to 44,731,405 pounds, valued at i209,000.
Of the food-fishes treated of in the second part (1904) — Scombrops
chilodipteroides, Scomber colias, Trachurus japonicus, Caranx
muroadsi, Paralichthys olivaceus — the most valuable is the Spanish
Mackerel, identical with the European form, its capture being
valued on an average at <£120,000 a year.
In the third part (1906) Spams Schlegelii, Thynnus Sclilegelii,
Clupea melanosticta, Chaioessus punctatus, and two freshwater fishes,
Cyprinus carpio and Carassius auratus, are included. As to the
Carp, it is interesting to notice that this fish is regarded as “ one
of the table dainties,” and that its culture has been practised in
Japan as far back as the first century. Goldfish are generally
distributed over the islands, even the smallest streams being
stocked with this species ; its culture is known to have been well-
developed in the ninth century.
Part 4 (1907) contains descriptions and figures of nine species :
Pagrus major, Oncorhynclius keta, Plecoglossus altivelis, Salmo
mason, Hypomesus olidus, Salanx microdon, Anguilla japonica,
Murcenesox cinereus, and Conger anago. The first of these is of
great commercial value, estimated at £326,000 per annum. The
small Plecoglossus is highly esteemed for the table, and aflfords
principally sport with Cormorants ; some places in the Miuo
450
Geological Societg.
Province have been famous for this mode of fishing since the reign
of the Emperor Jimmu, the founder of the nation more than 2500
years ago. A. G.
The British Woodlice, being a Monograph of the Terrestrial Isopod
Crustacea occurring in the British Islands. By Wilfred Mirk
Wedb and Charles Silleh. With 25 Plates and 59 Figures in
the Text. [Reprinted from the ‘ Essex Naturalist/ vol. xiv.
1905 -6.] 8vo. Duckworth & Co., 1906. Pp. x, 54. Qs. net.
Hitherto the British terrestrial Arthropoda other than the most
attractive insects have received comparatively little attention from
the general public, and we are glad to welcome a well-executed
monograph of one of these neglected groups on which hitherto there
has been no popular or easily accessible work. Moreover, as it is
a small group, it has been possible to deal with the subject in greater
detail than if a great number of species required to be noticed in a
limited space. In the present work seventeen species are described
and figured from Essex, including Ligia oceanica^ a sea-shore species,
which was very properly included as being closely allied to the
terrestrial species and too important and interesting to be omitted.
Besides these, eight species are described which have occurred in
other parts of the British Islands but have not yet been recorded
from Essex, making twenty-five British species in all. Now that
attention has been called to the group, others will doubtless soon be
added to the list.
The book commences Avith a well-written account of the position,
geological history, and structure of the group, the structure and
anatomy being well illustrated. This is followed by remarks
on habits, use in medicine, names, cpllecting and preservation,
classification, and tables of genera. Then follows the detailed
description of genera and species, and a good Bibliography closes a
volume which deserves the attention of all who are interested in
the zoology of the British Islands. W. F. K.
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
Januaiy 9th, 1907. — Sir Archibald Geikie, D.C.L., Sc.D., Sec.R.S.,
President, in the Chair.
The following communications were read : —
1. ‘ On the Cretaceous Formation of Bahia (Brazil) and on the
Yertebrate Fossils contained therein.’ By Joseph Mawson, F.G.S.,
and Dr. Arthur Smith Woodward, F.R.S., F.L.S., F.G.S.
This paper relates to a series of estuarine and freshwater deposits
originally described to the Geological Society by the late Samuel
Allport, in 1859. The results of thirty years’ collecting of fossils
Miscellaneous.
4j1.
are summarized, and the distribution of the formation, so far as
determined, is marked on a map. The strata are disturbed by
numerous dislocations and discordant dips, and no regular succes-
sion of zones or horizons can be discovered. All the more
important vertebrate fossils collected are now in the British Museum
(Natural History). From these a few remains of new species are
selected for special description. A mandibular symphysis of a
very large crocodile, with a long garial-like snout, belongs to one
of the Goniopholidae. Some Dinosaurian vertebrae seem to belong
to the Iguanodont group. A large fish-skull represents a new
genus allied to Macropoma^ and indicates a species five or six times
as large as any Coelacanth previously discovered. The discussion of
a complete list of the fossil Vertebrata proves that the formation is
of Cretaceous age, and suggests that it may be Lower Cretaceous,
as supposed by Hartt.
2. ‘ On a New Dinosaurian Reptile from the Trias of Lossiemouth,
Elgin.’ By Arthur Smith Woodward, LL.D., E.R.S., F.L.S., F.G.S.
Mr. W^illiam Taylor, of Elgin, has recently discovered two
skeletons of a small new reptile in the Triassic sandstone of
Lossiemouth. Two imperfect skeletons of the same species are
also shown on a slab of the same sandstone in the British Museum
(Natural History). The head and trunk measure only 4 inches in
length, but there is a very long and slender tail. The head is
relatively large, and resembles that of Ornithosuchus in many
respects ; but the fossils do not exhibit any teeth. There are about
twenty-one presacral vertebrae, of which nine are cervical. There
are distinct traces of a plastron of delicate abdominal ribs. The
limb-bones exhibit a large internal cavity. The fore-limbs are very
small, with a humerus as long as the radius and ulna. The hind-
limbs are relatively large, and the ilium is extended autero-
posteriorly for the length of four vertebrae. The femur is almost as
long as the tibia and fibula ; while the metatarsus is especially
remarkable, being half as long as the tibia and consisting of four
metatarsals of nearly-equal length firmly fused together. The toes
are long and slender, with sharply-pointed claws. The Author
concludes that this must have been a running or leaping reptile,
and that it represents a new genus of Dinosauria related to the
American Triassic Hallopus.
MISCELLANEOUS.
Modern Helminthological Nomenclature.
By Dr. vox Linstow.
Within the last few years such far-reaching changes have been
effected in helminthological nomenclature that it appears well worth
while to subject them to a critical examination.
In the first place the principle has been established that the name
452
Miscellaneous.
that shall be valid is the oldest, and not the one that has hitherto
been in use and is well known ; the consequence is that the species
are quoted under names some of which were never current, and
have to be explained by the addition of those previously used, since
their meaning is unknown ; the oldest name has the right of
priority.
On this point Looss remarks in his latest work, ‘ On New and
Previously Described Trematodes from Marine Turtles ’ (‘ Uber neue
und bekannte Trematoden aus Seeschildkroten,’ Jena, 1902) ; — “A
name has no right of priority unless accompanied by statements
by means of which the species in question can be re-identihed. The
bare possibility that by an older name a certain species is meant to
be understood is not sufficient to justify its re-introduction ; the
right of priority can be claimed for an old name only when it is
recognizably defined, otherwise it is invalid, since the question
turns upon the meaning of the name. With an exchange of names
there must also be connected an exchange of concepts ; the old
name must, in order to supersede tlie later one, be justified by its
meaning ; the introduction of old names to which no one is able to
attach a certain definition could only give rise to differences of
opinion.”
That this is to be demanded of generic just as much as of specific
names is self-evident ; generic names that convey no meaning are
valueless.
It is sad that there should be any necessity to lay stress upon
these self-evident principles ; we shall see, however, that it was
bound to happen, for modern systematists deal largely in words
that are wanting in meaning, and we are involuntarily reminded
of Goethe’s
Denn eben wo Begrifte fehlen,
Da stellt ein Wort zur rechten Zeit sich ein ;
Mit Worten lasst sich trefflich streiten,
Mit Worten ein System bereiten ;
An Worte lasst sich trefflich glauben,
Von einem Wort lasst sich kein Jota raiiben” *
This is the answer that the foolish scholar receives to his
objection ; —
“ Doch ein Begriff muss bei dem Worte sein ” f.
The word Fasciola is substituted for the well-known and
customary I)istomum\ almost all authors loyally write Fasciolidae
instead of Distomidae, according to the law of priority laid down by
the Zoological Congress.
^ For just where ideas are wanting
Comes in a timely word ;
With words can we raise a rare dispute,
With words a system institute ;
Belief on words may we well bestow,
No single jot will a word forego.
t Yet with the word an idea must be.
Miscellaneous,
453
The “ genus ” Fasciola was founded by Linnaeus (‘ Fauna Suecica/
ed. ii. (Holmiae, 1746), p. 505, no. 2075) for Fasciola hepatica ovata,
by which name- no fewer than three species are meant : — Distomum
liepaticum, Abildg. ; Dendrocoelum lacteum, Oerst. ; and Schisto-
cephalus solidus, Rud. It follows therefore that a Trematode, a
Turbellarian, and a Cestode are equally entitled to be termed
Fasciola, if under these circumstances Fasciola can stand as a
generic name. There can, however, surely be no doubt that a
generic name which at the same time denotes a Trematode, a
Turbellarian, and a Cestode is nonsense from a scientific point of
view. No idea is conveyed by the word, but to this modern
systematists apparently attach no importance.
Let us see how the name Fasciola was subsequently applied ; —
Fasciola intestinalis, Linnaeus, = digramma, Creplin ; Fasciola
alata, Rudolphi, = Hemistomum alatum. Diesing ; Fasciola excavata,
Die&in^, — Hemistomum excavatum^ Diesing ; Fasciola strigis, Gmelin,
= Holostomum variahile, Nitzsch ; Fasciola suhclavata, Schrank, =
Diplodiscus suhclavatus, Diesing ; Fasciola has been used to designate
fifty species of Distomum.
Fasciola hepatica, Muller, = A^nphistomum conicum. Rudolphi ;
Fasciola verrucosa, Schrank, = Monostomum verrucosum, Zeder ;
Fasciola uncinata, Q(me\m, = Polystomum integerrimum, Rudolphi ;
Fasciola harhata, JAmidd\x.s,, = Rhynchohothrium paleaceum, Rudolphi ;
Fasciola truttae, Rcederer, = ZV’^cieuop^oms nodulosus, Rudolphi ;
Fasciola marmorosa, Muller, = marmorata, Hempr. &Ehrg. ;
Fasciola glauca, MAWcr, = Monocelis glauca. Diesing ; Fasciola lactea,
Mxl\QY, — Planaria lactea, Muller.
This is the result of the foundation of the genus Fasciola by
Linnaeus ; the definition of Fasciola is a fiattened worm, which
tapers at both ends.
Looss discusses the question whether in applying the law of
priority we should go back to Linnaeus, ‘ Systema Naturae,’ ed. x.
1758, or to Rudolphi, ‘ Synopsis,’ 1819, and decides in favour of
the latter year, since Rudolphi was to a certain extent the founder
of scientific helminthology, and the only means of interpreting the
obscure old descriptions correctly is the study of the types. He
holds, however, that, as a general rule, the introduction of old and
unknown names not hitherto in use is a retrograde step, and that
disinterred antiquities should not again be employed.
As a matter of fact, when we, only in accordance with the law of
priority, write Vesicaria truttoe instead of Taenia longicollis, Halysis
latus instead of Bothriocephalus latus, Lumhricus teres instead of
Asca7'is lumhricoides, Gordias medinensis instead of Dracunculus
medlnensis, Filaria locustae instead of Gordius aquaticus, Cucullanus
ranee instead of Strongylus am'icularis, and Taenia hceruca instead of
Echmorhynchus angustatus, satisfaction has been rendered to the
right of priority. Not all the species here mentioned are the first
and typical ones for which the old genus was founded ; but it
was only a question of showing what a confusion of ideas is occa-
sioned by the old names, and with the growing modern inclination
Miscellaneous,
4r>t
to form where possible a speeial genus for each species the prospect
of being obliged to return to the old generic names indicated
becomes constantly greater.
It is an arbitrary proceeding to lay down 1758 and 1819 as
limits of time from which the law of priority shall commence to
apply ; what really matters is the signification of the old names.
If investigators of the importance of a Leuckart and others did
not resuscitate these old names, they must have had their reasons
for the course that they adopted ; they, too, must surely have con-
sidered such a procedure as a step in the wrong direction.
In strange contrast to the endeavours to replace later names by
old ones are the equally frequent attempts to substitute quite recent
names for the latter.
The genus Tetrabothrium, lludolphi, with the typical species
cylindraceum, Eud., and macrocephalum^ End., has been broken up
and replaced by Prosthecocotyle, Monticelli and Fuhrmann, and
Bothriotcenia, Lonnberg. As a reason for this proceeding it is
asserted that Diesing has employed the designation Tetrahothriiim,
Eudolphi, in a sense different from that in which it is used by
Eudolphi, but this, however, in no way concerns the latter and his
genus.
The old and well-characterized genus Amphistomiini, Eudolphi,
has been broken up by Fischoeder, who substitutes his genus
Parampliistomum. The reason given is that Eudolphi described an
Ampliistomum macrocephalum, which, however, according to the
laws of priority must be called Strigea. The oldest name for this
species is not Strigea, but Planaria teres, Goeze(l782); it was
afterwards termed Festucaria strigis, Schrank (1788), and subse-
quently (1793) Fasciola strigis, Gmelin ; then for the first time
Strigea, Abildgaard (1793), later Amphistoma macroceplialum,
Eudolphi (1801), and finally Holostommn variahile, Nitzsch (1819).
The latter is the name of the species to-day, and consequently it
is not an Ampliistomum ; the typical species of this genus are
conicum, Eud., and suhtriquetrum, Eud., hut the name Parampthi-
stomum, which has not the slightest justification, has been adopted
by modern systematists.
Trichina, Owen, is now called Trichinella, Eailliet, since Meigen
applied the name Trichina to a Dipteron in 1830. Medicine,
veterinary science, and agriculture will not employ the terms
“ Trichinellae,” “examination for Trichinellac,” and “ Trichinellosis.”
For more than thirty years the whole of the educated world has
known the meaning of Trichinae, but what Trichinellae are it does
not know and will, moreover, not learn ; there is no risk of con-
fusion ; when we read that someone is suffering from Trichinae or
that they have been found in a pig we do not think of flies ; 1
consider it wrong to bring about a change of name that will never
be adopted.
The genus Monostomum, Zeder, was abolished by the bestowal
ii])on its species of numerous other generic names; afterwards.
^fisceUaneous.
455
when Monostomidse were spoken of, it was remarked that the
genus Monostomum had disappeared ; it had slipped through our
fingers ; the only species remaining was Monostommn prismaticum^
Zeder, which, however, according to Monticelli is a Disiomum ;
according to Looss the species is absolutely indeterminable. Thus
Zeder’s genus Monostomum has ceased to exist, and a similar fate is
in store for many another old genus if the course adopted is
followed further.
While entirely sharing the above-quoted views of Looss with
reference to the value of the old names, I cannot approve of the
action of this author and others in founding wherever possible a
new genus for each new species, nor can I assent to his definition
of the idea conveyed by the latter term. Looss declares that, if two
different species exhibit anatomical differences, they at least belong
to different genera ; two species of the same genus must be in
perfect agreement as regards their anatomical structure, and may
only differ in the relative size and position of their organs and in
actual bodily size. If we examine the figures of the six Distomes
given by Looss on p. 860 of his work already referred to, which are
stated to represent six genera, the conviction takes hold of us that
specific differences are here mistaken for generic : in fig. 5 the
viteilarium lies beneath and outside the limb of the intestine, in
fig. 6 only outside it ; otherwise the two figures agree in even the
smallest detail ; and these are said to belong to two genera. If
trivialities of this sort are regarded as generic differences, we shall
soon have as many genera as species. Looss asserts that the old
generic designation Distomum no longer signifies anything whatever.
I consider Distomum to be an excellently characterized genus,
which, like Tcenia, only possesses the inconvenience of being too
bulky ; we shall therefore do well to effect a division into subgenera,
and to write, for example, Distomum {Apohlema) appendiculatum
and Tcenia {Davainea) frontina. The definition of a genus adopted
by Looss is, however, inapplicable to other classes, e. g. birds, fishes,
and insects. I regard the genus as the aggregate of species united
into closer association by means of common characters ; thus we
have genera such as Felis^ Anas, Cyprinus, Dana, Vipera, Vanessa,
and in botany Quercus and Ranunculus ; they are not based upon
anatomical differences, however ; Looss’s definition ignores common
characters and only takes into consideration the differences ; for the
concept of a genus, as I apprehend it, it is not the differences but
the common features that are decisive.
In ornithology we have got beyond this period of nomenclature ;
time was when the gulls found on German shores bore the generic
designations Rodostethia, Xema, Hydrocolceus, Gavia, Melagavia,
Cephus, Laroides, Rissa, Chimonea, Pagophila, Cetosparactes, Leucus,
Glaucus, Clupeilarus, and Dominicanus — fifteen generic names for
twelve species ; they are now in the majority of cases all called
Larus once more, the three-toed gull at the most is assigned to
Rissa.
45G
Miscellaneous,
Owing to the confusion occasioned by modern nomenclature, it
has already become impossible to make an alphabetical catalogue of
Helminthes ; for where, for instance, are we to place Distomum
maculosum, which has been assigned to five different genera ? The
s})ecies is called Dlstommn maculosum, Rudolphi, Fasciola maculosa,
lludolphi, Bicrocoellum maculosum, Olsson, Brachyleimus maculosus,
Stossich, and Playiorclius maculosus, Braun ; among these genera
the reader may make his choice.
The method of writing the names has also been influenced by the
modern passion for innovation. The rule has been laid down that
specific names are to be written with a small initial letter. Ascaris
linnoei means Linne’s Ascaris ; the word is a genitive, and whoever
writes the name Linnoeus in the nominative and linnoei in the
genitive perpetrates an orthographical error ; are we, for example,
also to write Tcenia van henedeni instead of van Benedeni ? Three
mistakes are comprised in the specific name of Ascaris gadi-hrandti,
for the nominatives are Gadus and Brandtus, and a hyphen is
unknown in Latin. There is no advantage in this new fashion,
since everyone knows that in zoological names the first is the
generic and the second the specific name. The disadvantages are
manifold ; in the first place the modern way of writing is wrong ;
secondly, we are led to consider words with small initial letters as
adjectives, which is not the case ; and lastly, when a name has
been bestowed by an author, no one has the right to alter it
according to his own particular fancy.
On enquiring as to the origin of these interfering changes, we are
told that it is a question of the principle of stability in nomen-
clature ; ‘‘ our first consideration in nomenclature should be
stability,” says Stiles. In the setting-up and observance of the
laws of stability of nomenclature, however, the names, which after
all were the end in view, have been entirely forgotten, for the
stability of the names has been utterly destroyed. We have now
introduced names which are scientifically impossible : in the place
of the old well-known names are found new and unknown ones ;
instead of new and legitimate ones we have old senseless names
that are mere words. The attempt is made to cancel the old idea
of a genus, and to place almost every new species in a genus of its
own ; the names are wrongly written, and this is called stability.
Science also has its fashions, and we just go along with them ;
we do not want to be old-fashioned, but desire to stand on the
scientific summit ; we swim with the tide.
Science, however, is free, and no one, not even a zoological
congress, has the right to give it precepts which injure it. That the
course which has been adopted by the prevailing helminthological
nomenclature is a serious disadvantage to science I have no doubt
whatever. — Zoologischer Anzeiger, Bd. xxvi. no. G92 (January 26,
1903), pp. 223-229 : from a separate impression supplied by the
Author.
THE ANNALS
AND
MAGAZINE or NATURAL HISTORY.
[SEVENTH SERIES.]
No. 114. JUNE 1907.
LIV. — On the Classification of the Decapod Crustaceans,
By L. A. Borradaile, M.A., Lecturer in Natural Sciences
at Selwyn College, Cambridge.
In the following pages proposals as to the classification of the
Decapod Crustaceans which I have made in a number of
papers during the last few years are stated in a connected
form and with certain additions, so as to form a complete
conspectus of the higher divisions of the group. I hope that
this systematic summary may prove of practical use and
that some remarks introductory to those portions of it which
have not yet been published will be of service as a contribu-
tion to the discussion of vexed questions of phytogeny and
classification.
The necessity for keeping the article within reasonable
limits has compelled me to choose between the ordinary
method of stating the diagnostic characters of the divisions of
the classification under headings and that known as a key."*’
I have adopted the latter as being better suited to bring out
the resemblances and contrasts on which a phylogenetic
arrangement is based, and because it is of more immediate
use to anyone unfamiliar with the group. Questions with
which I have dealt elsewhere are not discussed in detail here,
Ann. ck Mag, N. Hist. Ser. 7. Vol. xix. 32
458
Mr. L. A. Borradaile on the
but a fuller treatment of them may be found in the origina
papers*, especially those in the ‘ Fauna of the Maldives.’
I.
The true position of the order Decapoda in the Crustacean
system is very well shown by Dr. Caiman’s elaboration of
Hansen’s classification, published in this Journal in 1904 f-
To this classification I would give my adhesion, only pointing
out that, if the Crustacea be given rank as a subphylum of
the Arthropoda, equivalent to the Arachnoidea, Tracheata,
and Prototracheata, the Malacostraca become a class and the
Eumalacostraca a subclass.
For our present purposes the most Important of the points
made by Dr. Caiman is the close relationship between the
Euphausiacea and the Decapoda. We shall assume that the
two orders have a common origin and regard the subdivisions
of the Decapoda as primitive in proportion as they approach
the Euphausiacea, though it will at the same time be necessary
to bear in mind that the Decapods with four rows of gills,
representing, as we shall see, both epipodite and proepipodite,
cannot be descended from the modern Euphausiaceans, which
have only one row.
* A Kevision of the Pontoniidoe,” Ann. & Mag. Nat. Hist. (7) ii.
pp. 376-391 (1898). [History, affinities, and limits of the (sub-) family,
pp. 376-379.]
“ On the Stomatopoda and Macrura brought by Dr. Willey from the
South Seas,” Willey’s Zool. Eesults, iv. pp. 395-428, pis. xxxvi.-xxxix.
(1900). [Palmnonopsis, a Palaemonid with affinities to Pontoniidae, p. 410 ;
doubtful validity of Latreutidae, p. 414.]
“ On some Crustaceans from the South Pacific. — Part IV. The Crabs,”
Proc. Zool. Soc. 1900, pp. 508-596, pis. xl.-xlii. [Primary subdivisions
of the Crabs, p. 571 ; subfamilies of Atelecyclidae, Cancridae, and Por-
tunidae, pp. 675-577.]
“ Marine Crustaceans ” in Gardiner’s ‘ Fauna and Geography of the
Maidive and Laccadive Archipelagoes.’ — Part III. The Xanthidae and
some other Crabs, vol. i. pp. 237-271, text-figs. 41-60. [Characters and
Classification of Xanthidae, pp. 237-238.] — Part IV. Some Eemarks on
the Classification of the Crabs, vol. i. pp. 424-429, text-fig. 110. — Part VI.
The Sand-Crabs (Oxystomata), vol. i. pp. 434-439, text-figs. 115-117,
pi. xxii. [Cliaracters and Classification of Oxystomata, p. 434.] — Part IX.
The Sponge-Crabs (Dromiacea), vol. ii. pp. 574-578, pi. xxxviii. [Cha-
racters and classification of Dromiacea, pp. 574-670.] — Part X. The
Spider-Crabs (Oxyrhyncha), vol. ii. pj). 681-690, text-figs. 122-126,
pi. xlvii. [Characters and classification of Oxyrhyncha, pp. 681, 682.] —
Part XI. On the Classification and Genealogy of the Eeptant Decapods,
vol. ii. pp. 090-698, text-figs. 125, 120, pi. xlviii. — Part XIII. The
Ilippidea, Thalassinidea, and Scyllaridea, vol. ii. pp. 750-754, pi. Iviii.
[Characters and classification of the groups.]
“ On the Classification of the Thalassinidea,” Ann. & Mag. Nat. Hist.
(7) xii. pp. 534-651 (1903).
Classification of the Decapod Crustaceans, 459
II.
1. The earliest of the surviving classifications of the
Decapoda is that established by Latreille in 1806 in which
the order is subdivided into Macrura or tailed forms and
Brachyura or Crabs. Roughly speaking, this division
depends on the condition of the abdomen, which in the
Macrura is carried at length and in the Brachyura is folded
under the thorax. In framing a definition, however, it is not
possible to rely on the above criterion^ for in the Porcellanidag,
the Hippidea, and the Lithodidas, which are undoubtedly
nearly related to tailed forms, the abdomen is carried as in
the Crabs. The absence from the Brachyura of the limbs of
the sixth abdominal segment is a better character of separa-
tion, but even this breaks down in the case of the Lithodidae,
which were, indeed, placed by Latreille with the Crabs.
Another criterion which is all but absolute is given by the
fusion of the carapace at the side to the epistome. This is
found in the Crabs, but only in the Scyllaridea and Eryonidea
among the Macrura. No single difference, however, can be
found which will absolutely and sharply define the Brachyura
from the Macrura.
2. The next important step in the working out of the
system was the establishment by H. Milne-Edwards in 1834
of a third suborder, the Anomura, intermediate between the
two of Latreille. In the new group were placed certain of
the higher Macrura (Paguridse, Hippidse, Porcellanidge) and
lower Brachyura (Dromiidse, Homolidse including LithodeSj
Raninida?), the abdomen in all these forms being more or less
modified from the primitive macrurous condition, but keeping
the sixth pair of limbs, except in the last two families.
Milne-Edwards’s Anomura has had a chequered history in
the hands of various authorities, having been alternately
added to or reduced, retained or parcelled out again between
the Brachyura and Macrura. In a recent paper f I have
tried to show that the macrurous members of the original
suborder, with the addition of the Galatheinea and Thalas-
sinidea, form a natural group, and must be retained as such
in the classification.
3. The last important proposal for the modification of the
classification of the Decapoda was made by Boas in 1880 J.
On the basis of an examination of the anatomy of a number
* Gen. Crust. Insect, i. Fabricius’s two classes Kleistagnatha and
Exochnata of “ Insects,” and Lamarck’s Cancri brachyuri and Cancri
macrouri, had much the same extension.
f Gardiner’s ‘ Fauna of the Maldives,’ vol. ii. p. 690.
t Kongl. Danske Vidensk. Selsk. Skrifter, (6) i. p. 23.
32«*
4G0
Mr. L. A. Borradaile on the
of typical genera Boas came to the conclusion that the
existing arrangement was unnatural in that it contrasted the
Brachyura and Anomura — single branches of the Decapoda
— with a heterogeneous assemblage (the Macrura) consisting
of the whole of the rest of the tree, some branches of which
are more nearly related to the Brachyura and Anomura than
they are to the rest of the Macrura. He accordingly proposed
to divide the order into two suborders — the Reptantia,
containing the Crabs, Anomurous forms, Thalassinidea,
Homaridea (Nephropsidea), Scyllaridea, and Eryonidea ; and
the Natantia, containing the Penaeidea (including Steno-
pida?) and the Caridea The names of these suborders
indicate the main difference by which, on the whole, they are
separated ; a more accurate diagnosis will be found below.
4. There can be little question of the correctness of Boas’s
view that the members of his E-eptantia form a natural
group. The characters that they hold in common are too
numerous and too specialized to admit of doubt on this point.
But it by no means follows that the same is true of the
Natantia. Unless it can be shown that the former group
arose from the Decapod stem before the separation of the
forms which constitute the latter, Boas’s classification will be
open to the objection that he raised against Latreille’s — that
is to say, it will be based, not on the divergence of two groups,
but on the elevation of a branch to the same rank as the
parent-stem. This, as it happens, is precisely the impression
conveyed by the tree figured by Boas on p. 27 of his paper.
The following considerations, however, serve to show that
Boas’s tree is wrong and his classification true.
A search for the most primitive group of the Decapoda
leads, beyond all doubt, to the PenEeidea. This is seen, (1) in
their primitive life-history, recalling that of the Euphausiacea ;
(2) in the peculiar copulatory armature of the male, which
suggests the same relationship ; (3) in the small number of
special features, unshared by other Decapoda, which the
group possesses, and the number of characters that they have
in common with one or other of the remaining groups — •
thus, with the lower Reptantia they share the shape of the
first three pairs of legs, which are fairly alike except some-
times in size, and all chelate t, whereas those of the Caridea
often differ much and their third pair is never chelate, the
structure of the maxillipeds, which lack special modifications
found in the Caridea, and the absence of the CEiridean bend
in the abdomen, while they share with the Caridea all those
Eukijphotes of Boas,
t Exc('pt in the aberrant Sergcstida).
461
Classification of the Decapod Crustaceans.
characters, enumerated below, which separate the Natantia
from the Keptantia ; (4) in their early appearance in the
earth’s liistory (probably in the Trias), thaugli it is true that
the remains of Reptantia are found fully as early ; (5) possibly
in the structure of their gills, if, as Boas thinks, the phyllo-
branchise of the Caridea and the trichobranchiae of the lower
Reptantia be both derived from the dendrobranchiae of the
Penaeidea. In any case it is impossible to regard the phyllo-
branch condition as the original one, but whether dendro-
branchiae or trichobranchiae are ta be regarded as the starting-
point of the gills of the Decapoda is much more doubtful.
It would be possible to support either theory by cases among
the Euphausiacea which might be regarded as substantiating it.
The evidence for the primitive nature of the Penaeidea is
therefore strong, but it must not be supposed that the modern
Penaeids were the stock from which the rest of the order arose.
Their loss of the appendix interna of the pleopods which is
found in Euphausiacea and in many Reptantia and Caridea,
is clear evidence that they do not stand in the direct line of
descent of the latter two groups. Moreover, the original
Decapoda must have borne the podobranch on the fourth leg
found in some of the lower Reptantia and the epipodite on
the last leg, of which Coutl6re has found a vestige in many
Caridea. Both these structures have been lost by the
Penaeidea. The most that can be said is that, of modern
Decapoda, the Penaeidea more nearly approach the primitive
condition than any others.
From the original Decapod stock, whose nearest descendants
we have found in the modern Penaeids, the Reptantia and
Caridea must have arisen separately, for it is impossible to
suppose that either of these specialized groups arose from the
other. They have no characters in common which they do
not also share with the Penaeidea, and each, as we have seen,
has characters which it shares with the latter group and not
with the other. There remains, then, the question, which
of the two was the first to leave the early Penaeid stem, and
that this was the Reptantia is shown pretty clearly by the
following facts : —
(1) The Caridea and Penaeidea have undoubtedly more in
common with one another than either of them has with the
Reptantia. This extends to characters which are at least not
obviously primitive, such as the styloeerite ” of the first
antenna.
(2) The gill-series in the lower Reptantia are fuller
than in either Penaeids or Carids, so that it seems likely
^ Except on the second pair of the male.
462
Mr. L. A. Borradalle on the
that the stock from which the latter two groups have
sprung lost a portion of their heritage in this respect after
the differentiation of the former. For, not only have some of
the lower Reptants kept tlie podobranchs on the legs of the
fourtli pair which all the Penseidea* and Caridea have lost,
but on several segments in the Potamobiidse we find the full
possible branchial equipment. Coutibre (‘ Comptes Rendus,’
1905, p. 64) has elaborated an extremely ingenious theory of
the homologies of the several kinds of epipodial structures of
the Decapoda with one anotlier and with those of the lower
Crustacea. Shortly put, this theory is as follows : — The
primitive number of epipodial outgrowths of the thoracic
limb of the Crustacea is two — a distal, the epipodite, belonging
to the coxopodite, and a proximal, the proepipodite^ belonging
to the true basal joint of the limb, which in the Decapoda is
taken into the body during development. Both these struc-
tures are found in Branch'pus in Anaspides. In Schizo-
poda and Decapoda both proepipodite and epipodite divide
into two parts. The epipodite forms in the Lophogastridse
(a) the oostegite and {^) a setiferous tubercle which I shall
call the setohranch. In the Caridea the epipodite forms, when
present, [a) the epipodite ” {rnastigohrancK) and (/3) on the
legs a setobranch of the same form as in the Lophogastridm,
and on maxillipeds 2 and 3 a podobranch and an arthrobranch
respectively ; in the Penseidea it forms (a) the “ epipodite ”
and (y8) the (anterior) arthrobranch, wanting in Caridea and
supposed to be there represented by the setobranch. The
proepipodite forms in the Lophogastridai a divided gill. In
the Decapoda it forms (a) the pleurobranch and (/>) the
(posterior) arthrobranch. In the development of Penwus
this subdivision can actually be seen to take place. The
Euphausiacea have lost their proepipodite.
Now, valuable and suggestive as this theory is, it is to
some extent invalidated by the fact that, in the case of
section ^ of the epipodite, structures which it regards as
alternative developments of the same rudiment can be found
coexisting. For it supposes that one arthrobranch (presumably
the anterior) and the podobranch and the setobranch are
equivalent and alternative structures. But in the Potamo-
biidse all these are present together on several segments of
the body. In Dromia Bohn has discovered what is un-
doubtedly a setobranch on the third maxilliped, where,
though the podobranch is wanting, both arthrobranchs are
present. On the first leg the setobranch is found on the
* It is only in certain of the primitive deep-sea Penaeids that the first
two or three pairs of legs bear podobranchs.
Classification of the Decapod Crustaceans, 463
base of the mastigobrancli and appears as an outgrowth from
it, suggesting strongly that the similar process on the masti-
gobranch of the third maxilliped of many crabs has the same
origin and that the two branches of the forked epipodite ”
of some Penasidse represent the setobranch and mastigo-
branch respectively. Of course there are also cases in the
lower Penaeidse and elsewhere where the podobranch and
both arthrobranchs are found together. I w^ould suggest,
therefore, that in the primitive Decapoda the epipodite divided
not into two but into four structures — (a) the mastigobranch,
(yS) the setobranch, (7) the podobranch, (3) the anterior
arthrobranch— just as in the Lophogastridse the proepipodite
has sometimes as many as four branches. At the same time
it must be remembered that the connexion of the anterior
arthrobranch with the mastigobranch is not a proved fact, as
is that of the posterior arthrobranch with the pleurobranch.
It seems quite possible that the ancestors of the Decapoda
bore not two but three rows of epipodial outgrowths on their
limbs, and that the anterior arthrobranchs represent the
middle of these three rows. Besides the epipodite ” and
proepipodite,” Branchipus bears on the outer side of its
thoracic limbs a third outgrowth of somewhat different form.
This has been doubtfully claimed as the exopodite, but may
quite possibly represent the mastigobranch.
(3) Whereas the Reptantia (Eryonidea &c.) appear in the
Trias, the geological record shows no trace of Caridea till late
Jurassic times. This group, in fact, is a late and somewhat
specialized offshoot from the Penajid stem. The lower
Reptantia have, perhaps, evolved further than the lower
Caridea, but they are still in some respects more primitive
and they took origin much earlier. Boas’s arrangement is
therefore justified. The Natantia are as natural a group
as the Reptantia, and into these two suborders the order must
be divided.
III.
In considering the subdivision of the Natantia it will be
evident from what has been said that the Penaeidea and the
Caridea must stand as two tribes of the suborder. To these,,
however, must be added a third whose position needs some
examination. The little family IStenopida3 was placed by
Boas with the Penaaidea, which it resembles in its tliree chelate
legs and in other respects ; but other authorities have very
* It is quite possible that the trichobratichiate nature of the gills of
the lower Keptantia is another primitive feature lost by the Penaeidea
and Caridea. >
464
Mr. L. A. Borradaile on the
rightly removed it to an independent division, the Stenopidea.
Tiie position of this group is extremely doubtful. It has
clearly no relationship to the Caridea, for it differs from them
and agrees with the Penaeidea and lower Reptantia in all
respeets in which the Caridea are peculiar, but its penseid and
reptant affinities are more evenly balanced. On the one hand,
like most of the Penaeidea it has lost all the podobranchs
behind the second maxilliped and the appendices internee, and
lias legs of the natant form j on the other hand, like the lower
Reptantia, it is trichobranchiate, has a curved mandibular
palp and short endopodite to the first maxilliped, and lacks
the copulatory apparatus of the male penajids and the spine
(stylocerite) on the stalk of the antennule which is so
characteristic of the Penseidea and Caridea.
There would be much to be said for placing this group by
itself as a suborder, but, on the whole, its affinities with the
Natantia seem strong enough to justify its being included
with them.
Since the termination -’idea is used below for groups of a
lower rank, the names of the tribes of the Natantia have, in
the key which follows, been made to end in -ides.
IV.
Within the Reptantia, the Brachyura and the Anomura
stand out as natural groups. With these I have already
dealt elsewhere *. There remain for consideration the
Nephropsidea, Scyllaridea, and Eryonidea. The latter two
of these divisions are closely related. They differ widely
from the Nephropsidea in the fusion of the carapace to the
epistome, the reduction of the rostrum f and of the inner lobes
of the second maxillse and first maxillipeds, the retention of
appendices internee on some of the limbs at least, and the
lack of sharp sutures on the tail-fin, and are very ancient,
whereas Nephropsidea, at least of the modern type, do not
appear till somewhat later. I propose therefore to class the
Scyllaridea and Eryonidea as a single tribe of the Reptantia,
giving to this tribe the name Palinura, which has the same
ending as those of the other tribes of the suborder, and recalls
the fact that the Palinuridse are among its members and the
position in which the abdomen is carried. For the sake of
uniformity, the Nephropsidea may take the name AstacurA,
which will indicate that the tail-fin in all the members of the
group is like that of Astacus, one of its most common
representatives. Thus the old Macrura are completely
dispersed.
* Gardiner’s ‘ Fauna of the Maldives,’ vol. ii. p. 690.
t Except in Palinurellus.
Classification of the Decapod Crustaceans.
465
V.
The following tree illustrates diagram m at ically the relation
ship between the groups which have been discussed : —
Brachyura.
I have grouped the families of the Carides into super-
families,” based on, but not quite the same as, the extremely
suggestive alliances ” proposed by Major Alcock for the
Indian deep-sea families. The shape of the mandible should
not be followed too implicitly as an indication of affinity in
this group. It shows a tendency to division into molar ”
and “ cutting ” halves throughout the order. In the higher
Carides this division is greatly accentuated, but in some cases
a secondary simplicity is reached by the loss of one of the
halves, and this has happened independently in Latreutes
and the Crangonoida (cutting-edge) and, I think, Pasiplueidse
466 Mr L. A. Borradaile on the
(molar process). The palp comes and goes from genus to
genus.
The following tree is an attempt to represent diagram-
matically the course of the evolution of the Oarides : —
Pdlaemonoida.
In discussing the classification of the crabs, I have else-
where * suggested, among other clianges, the abolition of the
distinction between the groups Cyclometopa and Catometopa.
The families gathered under the latter name have probably
genetic affinity, at least in some cases, but they pass into the
* Gardiner’s ‘Fauna of the Maldives/ vol. i. p. 425,
Classification of the Decapod Crustaceans, 467
Cyclometopa by such easy transition and, even in typical
genera, differ from them so little that their separation is a
needless and misleading complication of the system. It
would, in fact, be logically necessary, if a group Cyclometopa
were to be retained, to balance it by dividing the other
brachyrhynchous crabs into equivalent sections somewhat as
follows : — (1) Corystidge, (2) Portunidse, (3) Potamonidae,
(4) Atelecyclidae and Cancridae, (5) Xanthidae and Gono-
placidae, (6) Pinnotherida?, (7) ? Ptenoplacidae and Palicidae:,
(8) Hapalocarcinidae.
VIII.
A Table of the Classification of the Crustacea Decapoda.
Suborder NAT ANTI A.
Tribe Pen^ides.
Families: Penseidae (subfamilies: Cerataspinae, Aristaeinas, Sicyo-
niuae, Penaeinae), Sergestidae (subfauiilies : Sergestinae,
Ampbioniuae, Leuciferinae).
Tribe C a n i d e s.
Superfamily Pasiph^oida.
Families : Bresiliidae, Pasiphaeidae.
Superfamily Hoplophoroida.
Families : Hoplopboridae, Nematocarcinidae, Atyidae.
Superfamily Stylodactyloida.
Family Stylodactylidae.
Superfamily Psalidopodoida.
Family Psalidopodidae.
Superfamily Pandaloida.
Family Pandalidae (subfamilies : Thalassocarinae, Pandaliuae).
Superfamily Palcemonoida.
Families : Alpbeidae, Hippolytidae, Phyncliocynetidae, Pabc-
monidae (subfamilies : Hymenocerinae, Pontoniiuae, Palae-
moninae).
Superfamily Crangonoida.
Families : Gnathophyllidae, [Autonomaeidae ?], Processidae,
Glyphocrangonidae, Crangonidae.
Tribe Stenopides.
Family Stenopidae.
Suborder EEPTANTIA.
Tribe Palinuea.
Superfamily Bryonidea.
Family Eryonidae.
Superfamily Scyllaridea.
Families : Scyllaridae, Palinuridae.
Tribe Astac ura.
Families : Nephropsidae, Parastacidae, Potamobiida;.
4G8
Mr. L. A. Borradaile on the
Tribe Anomura.
Superfamily Galatheidea.
Families ; J^gleidae, Chirostylidae, Galatbeidae (subfamilies :
Galatheinae, Munidapsinae), Porcellanidae.
Superfamily Thalassinidea.
Families : Axiidse, Laomediidse, Callianassidae (subfamilies :
Callianassinae, Upogebiinse), Tbalassinidae.
Superfamily Tacjuridea.
Families : Pylochelidae, Paguridae (subfamilies ; Pagurinae,
Eupagurinae),GcBnob,itidae, Lithodidae (subfamilies : Ilapalo-
gastrinse, Lithodinae).
Superfamily Hi2)pidea.
Families: Albuneidae, Hippidag.
Tribe Brachyura.
Subtribe Dromiacea.
Superfamily Dromiidea.
Families : Homolodromiidae, DromiidcBj Dynomeriidae.
Superfamily Homolidea.
Families : Homolidae, Latreillidae.,
Subtribe Brachygnatha.
Superfamily Brachyrhpncha ( Cancridea).
Families: Corystidae, Atelecyclidae (subfamilies : Tliiinae, Acan-
tbocycliiiae, Atelecycliuae), PTricbiidae, Cancridae (sub-
families; Cancrinae, Pirimelinae), Portunidae (subfamilies:
Carcinidinae, Portumniuae, Gatoptrinae, Carupinae, Portuu-
inae, Capli5U‘inae, Thalamitinae, Podoplithalminae), Potamon-
idae (subfamilies : Potamoninae, Deckeniinae, Potamocarcin-
inae, Trichodactjdinae), Xanthidae (subfamilies : Xantliinae,
Carpilinae, Etisinae, Meiiippinae, Trapeziinae, Eripliiinae,
Oziiuae), Carcinoplacidae (subfamilies : Carcinoplacinae,
Gonoplacinae, Prionoplacinae, Rliizopinae, Hexapodinae),
Pinnotlieridae, Grapsidae (subfamilies: Grapsinae, Varuninae,
Sesarminae, Plagusiinae), Gecarcinidae, Ocypodidae (sub-
families : Ocypodinae, Macroplitlialmiiiae, Mictyrinae), Pali-
cidae, Ptenoplacidae, Hapalocarcinidae.
Superfamily OxyrJiyncha {Maiidea).
Families : Parthenopidas (subfamilies : Partbenopinae, Eiimedo-
ninae), Maiidae (subfamilies : Inacbinae, Acaiitbouycbiuae,
Pirinae, Maiinae), Hymenosomidae.,
Subtribe Oxystomata.
Families : Calappidae (subfamilies : Calappinae, Oritbyinae,
Matutinae), Leucosiidae (subfamilies : Leiicosiinae, Iliime),
llauinidae, Dorippidae (subfamilies : Uorippinae, Tymoliuae).
IX.
A Conspectus of the Classification of the Crustacea Decapoda.
Key to the Suborders.
I. Rostrum seldom reduced or absent, if well
developed almost invariably compressed.
Body almost always compressed. First
abdominal segment not much smaller than
4G9
Classification of the Decapod Crustaceans.
the rest. First antennae irenerally bear a
stylocerite. Second antennal scale generally
large. Legs slender (except sometimes a
stout chelate limb or pair which may be any
one of the first three pairs), with basipodite
and ischiopodite never fused, only one fixed
point in the carpo-propodal articulation,
sometimes exopodites, and podobranchs
hardly ever present on the first three pairs
and never on the last two. Male genital
opening almost always arthrodial. Abdo-
minal limbs 1-5 always present in full num-
ber, well developed, and used for swimming. NATANTIA.
II. Rostrum often reduced or absent, depressed
if present. Body not compressed, generally
depressed. First abdominal segment dis-
tinctly smaller than the rest. N o st5docerite.
Second antennal scale never large, generally
small or absent. Legs strong, the first
usually, the others never, stouter than their
fellows, basipodite and ischiopodite almost
always fused in the first pair, generally also
in the others ; two fixed points in the carpo-
propodal articulation, exopodites never
present, podobranchs fairly often present on
some of the first four paii’S. Male genital
opening coxal or sternal. Abdominal limbs
1-0 often reduced or absent, not used for
swimming REPTANTIA. ‘
Key to the Tribes of the Nat ant hi.
I. Third legs chelate, except in genera in which
the legs are much reduced. Third maxilli-
peds 7-jointed. Second maxillipeds with
normal end-joints. First maxillipeds with-
out the caridean lobe on the base of the
exopodites. Pleura of first abdominal seg-
ment not overlapped by those of second.
Abdomen without sharp bend. Not phyllo-
branchiate (except Amphioninfe).
1. One or both legs of third pair longer and
much stouter than those of first two pairs.
Trichobranchiate. Endopodites of first
maxillipeds short. Mandibular palps
curved. First antennae without stylo-
cerites. First abdominal limbs of male
not as in Penaeides STENOPIDES.
2. Legs of third pair not stouter than those of
first two pairs. Dendrobranchiate (except
Leuciferinae and Amphioninae : see below).
Endopodites of first maxillipeds long.
Mandibular palps straight. First antennae
generally with stylocerites. First abdo-
minal limbs of male bear a sexual
apparatus PENAUDES.
470 Mr. L. A. Borradaile on the
II. Third legs not chelate. Third maxillipeds
4-0-jointed. End-joint in second maxilli-
peds nearly always lies as a strip along end
of joint before it. First maxillipeds have a
lobe on the base of the exopodites. Pleura
of second abdominal segment overlap those
of first. Abdomen has generally a sharp
bend. Phyllobranchiate CARIDES.
Key to the Families of the Penseides.
I. Last two pairs of legs well developed. Gills
many Penaeidae.
II. Last one or two pairs of legs reduced or lost.
Gills few (up to 8) or wanting Sergestidae.
Key to the Subfamilies of the Penaeid«?.
I. Carapace covers le;.fs. Exopodites well deve-
loped. [Podobranchs on some legs.] Cerataspime ,
II. Carapace of normal size. Exopodites re-
duced or lost.
1. Well-developed podobranchs on some legs.
[Exopodites on maxillipeds and sometimes
on some legs. Arthrobranchs in double
series. First antennae without leaf-like
appendage on first joint.] Aristcsince.
2. No podobranchs on legs (vestige on first legs
of Halijjorus).
i. No exopodite behind first maxillipeds.
Arthrobranchs in single series. No
leaf-like appendage on first joint in first
antennae Sicyonince.
ii. Exopodites on all maxillipeds and usually
some legs. Arthrobranchs in double
series. A leaf-like appendage on inner
side of first j oint in first antennae .... Penceince.
Key to the Subfamilies of the Sergestidae.
I. All the thoracic limbs biramous. Gills
present and resemble phyllobrancliiae Amphionincp.
II. Last seven thoracic limbs uniramous. Gills,
if present, are dendrobranchiae.
1. Head not greatly elongated. Gills present. Seryestince.
2. Head greatly elongated. No gills Leucif evince.
Key to the Supevfamilies of the Carides.
I. Second maxillipeds normal. [Exopodites
on some or all legs. Mastigobranchs on
none. First two pairs stouter than the rest,
with normal chelae and undivided wrist-
joints. Mandibles without or with distinct
but small molar process, with or without
palps.] Pasiph^.oida.
471
Classification of the Decapod Crustaceans,
II. Second maxilli^eds with the sixth and seventh
joints articulating separately on fifth. [No
exopodites on legs. Mastigobranchs on first
to fourth pairs. First two pairs of good size,
chelate, with very long fingers and undivided
wrist-joints. Mandibles imperfectly cleft,
with palp.]
III. Second maxillipeds with short seventh joint,
usually applied as a strip to the end of the
sixth.
1. Mandibles imperfectly cleft. Exopodites
usually present on all or some legs. First
two pairs of legs substantially similar, of
moderate size, chelate, with undivided
wrist-joint. [Mastigobranchs present on
some legs (except Limnocaridina).'] ....
2. Mandibles either deeply cleft or simple,
apparently owing to the loss of the cutting-
edge. No exopodites on legs (except in
a very few cases on the first pair). First
two pairs of legs more or less unlike.
i. At least the basipodites of the second
maxillae well developed. Mandibles
rarely simple {Latreutes &c. ). First legs
not subchelate.
(1) First two pairs of legs slender. First
pair simple or minutely chelate.
Second chelate, with wrist divided
into two or more joints. [Mastigo-
branchs generally present on legs.] . .
(2) First legs with both fingers movable,
second with last joint replaced by a
tuft of bristles and undivided wrist-
joint. [No mastigobranchs on legs.]
(3) First two pairs of legs not both slender
(one often very large), chelate. Wrist
of second pair often subdivided.
[Mastigobranchs present or not.] . .
ii. Inner lobes of second maxillae reduced.
Mandibles simple. First legs often sub-
chelate. [Second wrist divided or not.
No mastigobranchs on legs.]
Stylodactyloida.
Hoplophoroiba.
Pandaloida.
PSALIDOPODOIDA.
PALmiONOIDA.
Crangonoida.
Key to the Families of the PasiphsGoida.
I. Rostrum small or wanting. No molar process
on the mandibles. Inner lobes of second
maxillae and first maxillipeds reduced. Exo-
pcdites on all legs Pasiphaeidae.
II. Rostrum well developed. Mandibles with a
distinct molar process. Inner lobes of second
maxillae and first maxillipeds not reduced.
Exopodites on first two pairs of legs only . . Bresiliidae.
Key to the Families of the Hoploplioroida,
I. Both fingers of chelae S]^on-like and ending
in tufts of bristles. Exopodites may be
472 ^Ir. L. A. Borradaile on the
■wanting on some or all legs. Freshwater
forms Atyidae.
II. Chelee not as in Atyidae. Exopodites on all
legs. Deep-sea forms.
1. Last three pairs of legs abnormally long.
A lash on the exopodite of the first
maxillipeds Nematocarcinidae.
2. Last three pairs of legs not abnormally
long. No lash on the exopodite of the
first maxillipeds Hoplophoridae.
Key to the Subfamilies of the Pandalidse.
I. Second wrists undivided Thalassocarinfc .
II. Second wrists subdivided Pandalince.
Key to the Families of the Palaemonoida.
I. Second wrists subdivided.
1. First legs much stronger than rest. Eyes
usually covered by carapace. [Mastigo-
branchs of legs and mandibular palps
present.] Alpheidae.
2. First legs not much stronger than rest.
Eyes not covered by carapace. [Mastigo-
branchs of legs and mandibular palps
present or absent,] Hippolytidae.
II. Second wrists undivided.
1. nostrum movable. Mastigobranchs on
legs. [Mandibular palp present.] ...... Rh5mchocinetid3e.
2. Rostrum not movable. No mastigobranchs
on legs Palaemonidae.
Key to the Subfamilies of the Palaemonidae.
I. First antennae with two flagella (one usually
cleft for some distance from the tip). Third
maxillipeds have third joint flat and often
broad.
1. Mandibles with palps. Propodites of
second legs, third maxillipeds, and one
branch in first antennae broad and flat . . Ilymenocerincc.
2. Mandibles without palps. Limbs not
broadened as in Hymenocerinae Pontoniince.
II. First antennae with three flagella (owing to
cleavage of one almost or quite to the base).
Third maxillipeds pediform. [Mandibular
palps usually present.] Paloimonince.
Key to the Families of the Crangonoida.
I. One or both legs of first pair chelate.
Rostrum short, compressed.
[1. Second legs simple Autonomaeidae.] *
numcca is that Risso, who described it, thought it related to Processa.
473
Classification of the Decapod Crustaceans.
2. Second legs chelate.
i. Both legs of the first pair chelate. Second
■wrists undivided. Third joint in the
third maxillipeds very broad. Rostrum
toothed
ii. One leg of the first pair simple. Second
wrists subdivided. Third maxillipeds
pediform. Rostrum not toothed ....
II. Both legs of the first pair subchelate. Ros-
trum long or short, not compressed.
1. Second wrists subdivided. Inner lobes of
first maxillipeds not reduced. Rostrum
2. Second wrist undivided. luner lobes of
first maxillipeds reduced. Rostrum short.
Gnath ophy llidas .
Processidse.
Glyphocrangonidae.
Crangonidae.
Key to the Tribes of the lleptantia.
I. Third legs like first, either chelate or simple
and subcylindrical. Abdomen macrurous
(straight, symmetrical, well armoured, with
good pleura and strong broad tail-fin, lobes
on the first segment clipping the carapace).
Gnathobases of second maxillae narrow.
Basipoditic lobes of first maxillipeds usually
deep. Exopodites of maxillipeds with lash
directed forwards. Gills numerous. [Last ,
thoracic segment with legs not differing
greatly from the rest and sternum rarely free, j
1. Carapace fused at the sides to the epistome.
Rostrum small or wanting (except Palinu-
rellus). Inner lobes of second maxillae
and first maxillipeds reduced. An ap-
pendix interna on some of the abdominal
limbs, at least in the female, but the exo-
podites of the last pair without sharp
suture. Body often depressed FALINURA,
2. Carapace free from the epistome. Rostrum
of good size. Inner lobes of second max-
illae and first maxillipeds not reduced.
No appendix interna y but the exopodites
of the last abdominal limb divided by a
suture. Body subcylindrical ASTACUIiA.
n. Third legs unlike first *, never chelate. Ab-
domen rarely macrurous. Gnathobase of
second maxillae typically broad. Basipoditic
lobes of first maxillipeds broad but shallow,
their inner edge usually in a line with that of
the coxopodite. Exopodites of maxillipeds
■^'ith lash, when present, nearly always bent
inwards. Gills usually few.
1. Carapace not fused with epistome. Last
thoracic sternum free, its legs differing
* Gebicula nearly forms an exception to this.
Ann. May. N. Hist. Ser. 7. Vol. xix, 33
474
Mr. L. A. Borraclaile on the
always clearly in size and position and
nearly always in size and shape from the
third pair. Abdomen anomurous (reduced
in some of its features, but showing- clear
traces of some function other than that of
reproduction, and almost always carrying-
biramous limbs on the sixth segment) or,
rarely, macrurous. A movable antennal
scale often present. Third maxillipeds
usually narrow ANOMURA.
2. Carapace fused with epistome at sides and
nearly always also in middle. Last tho-
racic sternum fused with rest, its legs
often like the others. Abdomen brachy-
urous (small, straight, symmetrical, bent
under the thorax, showing no traces of
other function than reproduction, and
without biramous limbs on the sixth seg-
ment). Never a movable antennal scale.
Third maxillipeds broad BRACHYURA,
Key to the Superfamilies of the Palinura.
I. Carapace gripped by the first abdominal
segment alone. First joint of second an-
tennae not fused withi epistome ; a scale
present on this limb. All the legs, except
sometimes the last pair, chelate ; the first
larger than the rest. Unbranched limbs on
the first abdominal segment. Tail-fin not
softer behind than before, without sutures.
Telson pointed Eryonidea.
II. Carapace gripped between a lobe on the first
abdominal segment and a knob on the side
of the last thoracic. First joint of second
antennae fused with epistome ; no scale on
this limb. None of the legs much longer
than the rest, ca*, except sometimes the first
pair, chelate. No limbs on first abdominal
segment. Tail-fin divided by indistinct
sutures into a soft hinder half and a harder
front half. Telson roughly square behind. . Scyllartdea.
Key to the Families of the Scyllaridea.
I. Cephalothorax subcylindrical. Eyes not en-
closed in separate orbits formed by the edge
of the carapace. Second antennae with
flagella Palinuridae.
IT. Cephalothorax depressed. Eyes enclosed in
separate orbits formed by the edge of the
carapace. Second antennae with flat scales
in place of the flagella Scyllaridae.
475
Classification of the Decapod Crustaceans.
Key to the Families of the Astacura.
I. Podobranchs not united with the mastigo-
branchs. Last thoracic segment fixed.
[Sexual appendages in male. Four pleuro-
branchs.] Nephropsidae.
II. Podobranchs united with the mastigobranchs.
Last thoracic segment free.
1. Gills have a lamina, but no hooks at the
end of the filaments. Sexual appendages
in male. One pleurobranch or none .... Potamobiidae.
2. Gills have no lamina, but hooks at the end
of the filaments. No sexual appendages
in male. Generally four pleurobranchs . . Parastacidae.
Key to the Superfamilies of the
I. Second to fourth legs with last joint curved
and flattened. First pair styliform or sub-
chelate. [Tail-fin not adapted for swim-
ming. Abdomen bent under thorax. Eos-
trum small or wanting. Third maxillipeds
have no mastigobranchs.]
II. Second to fourth legs with last joint not
curved and flattened. First pair not styli-
form, rarely subchelate.
1. Sixth abdominal limbs adapted for swim-
ming (except in Thalassina, where they
are styliform). Pleura usually well deve-
loped. Abdomen symmetrical.
i. Body depressed. Pleurobranchs to last
legs. Often a transverse suture on
telson. Abdomen more or less bent . .
ii. Body compressed. No pleurobranch to
last leg. N 0 transverse suture on telson.
Abdomen straight
2. Sixth abdominal lim bs, when present, with
branches neither broad nor styliform, but
adapted for holding the body into hollow
objects. Pleura very rare. Abdomen
nearly always asymmetrical, and either
soft and twisted or bent under the thorax.
Anotimra.
Hippidea.
Galathbidea.
Thalassinidea.
Paguridea.
Key to the Families of the Hippidea.
I. First legs subchelate. Carapace flattened,
without wings to cover the legs. Third
maxillipeds narrow, with exopodites Albuneidae.
II. First legs simple. Carapace subcylindrical,
with wings which cover the legs. Third
maxillipeds broad, without exopodites .... Hippidaa.
Key to the Families of the Galatlieidea.
I. Trichobranchiate. Eight arthrobranchs. No
limbs on second abdominal segment of male.
[Abdomen not folded against thorax. Second
antennae with 5-jointed stalk, but no scale.] jEgleidae.
33*
47G Mr. L. A. Borradalle on the
II. Phyllobranchiate. Tenarthrobrancbs. Limbs
on second abdominal segment of male.
1. Arthrobranchs stand on side of thorax.
Second antennae have 5-jointed stalk and
usually a spiniform scale. [Abdomen not
folded against the thorax. Third maxilli-
peds veithout mastigobranch.] Chirostylidae.
2. Arthrobranchs normally placed. Second
antennae have 4-jointed stalk and no scale
(or vestiges only).
i. Abdomen not folded against thorax. Third
maxillipeds with mastigobranchs Galatheidse.
ii. Abdomen folded against thorax. Body
crab-like. Third maxillipeds without
mastigobranchs Porcellanidae.
Key to the Sahfamilies of the Galatlieidas.
I. Eyes well developed. Exopodites of third
maxillipeds with 1-jointed “ flagella ” .... Galatheirus.
II. Eyes reduced. Exopodites of third maxilli-
peds without flagella Mu7iidopsina.
Key to the Families of the Thalassiiiidea.
I. No tinea thalassinica. Both movable and
flxed antennal thorns present, though some-
times minute (? absent in Scptoleptus). Ab-
dominal pleura large. [Last endopodite
without suture. Second legs chelate.] .... Axiidae.
II. JLinea thalassinica present (except Callia-
nidea). Fixed antennal thorn wanting ;
scale reduced to a flattened vestige or wanting.
Abdominal pleura usually small.
1. Sutures on endopodite and exopodite of
sixth abdominal limbs. Abdominal pleura
of a good size ; • • • : Laomediidae.
2. No sutures on sixth abdominal limbs. Ab-
dominal pleura small.
i. Second leg chelate or simple. No podo-
branchs on legs. Abdominal limbs 3-6
broad. A vestige of antennal scale
remains Callianassidae
ii. Second leg subchelate. Podobranchs on
legs 1-3. Abdominal limbs all narrow.
No vestige of antennal scale Thalassinidae.
Key to the Subfamilies of the Callianassidge.
I. Rostrum large. Legs of first pair equal. No
appendix interna on abdominal limbs 3-5 . . Upogehimce,
II. Rostrum small. Legs of first pair unequal.
An appendix interna on abdominal limbs 3 -5. Cidlianassince.
Classification of the Decapod Crustaceans.
477
Key to the Families of the Paguridea.
I. Abdomen straight or twisted. Carapace firm
and more or less compressed in the fore part,
soft in the hinder part, at least at the sides.
Fourth legs unlike third. Rostrum almost
or quite wanting. Sixth abdominal limb
present.
1. Abdomen macrurous and symmetrical, with
all the limbs present. Trichobranchiate. Pylochelidae.
2. Abdomen more or less unsym metrical, some
of the limbs lost. Generally phyllo-
branchiate.
i. Antennal scale well developed (thorn-
like). First antennae with stalk of
moderate length and flagella ending in
a filament. Marine forms Paguridae.
ii. Antennal scale reduced. First antennae
with very long stalk and flagella ending
bluntly. Land forms Coenobitidae.
II. Abdomen bent under thorax. Body crab-
like. Carapace firm all over. Fourth legs
like third. Rostrum spiniform. Sixth ab-
dominal appendages lost Lithodidae.
Key to the Subfamilies of the Paguridse.
I. Third maxillipeds approximated at base.
Chelipeds equal or subequal, or the left much
the larger PagurincB.
II. Third maxillipeds wide apart at base. Right
cheliped usually, left never, much the larger. Eupagurin^.
Key to the Subfamilies of the Lithodidae.
I. Third to fifth abdominal segments imperfectly
calcified. Rostrum short and broad IIapalogastrin<£.
II. Third to fifth abdominal segments well calci-
fied. Rostrum generally narrow and pointed. Lithodince.
Key to the Subtribes of the Bracliyura.
I. Mouth-field (endostome) prolonged forwards
to form a gutter. [Last pair of legs normal
or abnormal. Female openings generally
sternal. First abdominal limbs of female
wanting. Gills few.]
II. Mouth-field roughly square.
A. Last pair of legs abnormal, dorsal. Female
openings coxal. First abdominal limbs of
female present. Gills usually many ....
B. Last pair of legs normal, rarely reduced,
not dorsal, except in Palicus and Pteno-
plax. Female openings sternal. First
abdominal limbs of female wanting. Gills
few
OXYSTOMATA.
DROMIACEA.
BRACHYGNATHA.
478
Mr. L. A. Borradaile on the
Key to the Families of the Oxystomata.
I. Body of the shape usual iu crabs. Abdomen
hidden under thorax. Antennae small.
Legs normal in position.
A. Afferent openings to gill-chambers lie in
front of first legs (chelipeds). Gills 9 on
each side. Male openings coxal
B. Afferent openings to gill-chambers lie on
either side of the mouth at the base of the
third maxillipeds. Gills less than 9 a side.
Male openings sternal
II. Body more or less abnormal in shape. Ab-
domen not hidden under thorax. Antennae
large. Last one or two pairs of legs in a
more dorsal position than the rest.
A. Carapace short. Last two pairs of legs
subprehensile, with hook-like end-joints. .
B. Carapace long. Legs usually have the last
two joints very broad
Calappidae.
Leucosiidae.
Dorippidae.
Haninidae.
Key to the Subfamilies of the Calappidse.
I. Last three joints in third maxillipeds not
hidden by the meropodite. Orbits not sepa-
rated from the antennular sockets.
A. Meropodites of third maxillipeds not elon-
gate nor acute. Exopodites of same limbs
with flagella. Legs not adapted for
swimming Caluppince.
B. Meropodites of third maxillipeds elongate
and acute. Exopodites of same limbs
without flagella. Legs adapted for
swimming . . .^ Orithyince.
II. Last three joints in third maxillipeds hidden
by the meropodite. Orbits more or less
separated from the antennular sockets.
[Exopodites of third maxillipeds with fla-
gella. Meropodite in same limbs elongate
and acute. Legs may be adapted for swim-
ming or not.] Matutince.
Key to the Subfamilies of the Leucosiidae.
I. Meropodites of third maxillipeds more than
half the length of the ischiopodites. Fingers
stout, gradually narrowing from base to tip,
usually shorter than the palm Leucoswuc.
II. Meropodites of third maxillipeds never more
than half the length of the ischiopodites.
Fingers slender, of even width from the base
to near the tip, usually longer than palm . . lliinee.
Key to the Subfamilies of the Dorippidae.
I. Third maxillipeds leave a good part of the
mouth uncovered. Inward openings to the
gills near the base of the chelipeds Dorippina-.
Classification of the Decapod Crustaceans. 479
II. Third maxillipeds almost completely cover
the mouth. Inward openings to the gills
may or may not be near the base of the
chelipeds TymolincB.
Key to the Superfamilies of the Dromiacea.
I. Sternum of female with longitudinal grooves.
Vestiges of sixth abdominal limbs usually
present. Gills 14-20 on each side. Eyes
usually completely sheltered by orbits when
retracted. No linece homolicce Dromiidea.
II. Sternum of female without longitudinal
grooves. No vestiges of sixth abdominal
limbs. Gills 8-14 on each side. Eyes in-
completely or not at all sheltered by orbits
when withdrawn against the body. Linece
homolicce usually present Homolidka.
^ Key to the Families of the Dromiidea.
I. No vestige of sixth abdominal limbs. Cara-
pace longer than broad, with ill-marked
side-edge. [First three legs with mastigo-
branchs, fourth and fifth small, subdorsal,
and prehensile.]
II. Vestiges of sixth abdominal limbs present
(except in Hypoconcha, where also no
mastigobranchs). Carapace usually not
longer than broad, with well-marked side-
edge.
A. Mastigobranchs on first legs (chelipeds)
only or on none. Fourth and fifth legs
small, subdorsal, and usually prehensile . .
B. Mastigobranchs on all the first three pairs
of legs. Fifth legs only small and sub-
dorsal
Homolodromiidae.
Dromiidae.
Dynomenidse.
Key to the Families of the Homolidea.
I. Gills 13 or 14 on each side. Mastigobranchs
on first one or three pairs of legs. First j oint
of eye-stalks not much longer than second . Homolidae.
II. Gills 8 on each side. Mastigobranchs not
found on any legs. First joint of eye-stalks
much longer than second Latreillidae.
Key to the Superfamilies of the Brachjgnatha.
I. Fore part of body narrow, usually forming a
distinct rostrum. Body more or less trian- [(Maude a).
gular. Orbits generally incomplete Oxybhyncha
II. F 'ore part of body broad. Rostrum usually
reduced or wanting. Body oval, round, or [(Cancridea).
square. Orbits nearly always well enclosed. Brachyrhyncha
480
Mr. L. A. Borradaile on the
Key to the Families of the Oxyrhynclia.
I. Carapace thin and flat. First legs (chelipeds)
not long or specially mobile or with fingers
bent at an angle with the hand. Male
opening sternal. [No orbits. Second joint
of antennal stalk slender, fused with epistome
but not with front. No hooked hairs.]. . . . Hymenosomidae.
II. Carapace not thin and flat (except Ocinopus).
First legs either mobile or powerful, with
bent fingers. Male opening coxal.
A. Chelipeds specially mobile, rarely much
greater than the other legs, or with fingers
bent at an angle on the hand. Second
joint of antenna well developed, generally
fused with epistome and often with front.
Orbits generally more or less incomplete.
Hooked hairs almost always present .... Maiidae.
B. Chelipeds not specially mobile, usually
much longer and heavier than the other
legs, and with fingers bent on the hand
at an angle towards the side on which the
fixed finger is set. Second joint of an-
tennae small, short, and not fused with
epistome or front. Orbits well made.
Hooked hairs almost always wanting . . Par then opidae.
Key to the Subfamilies of the Maiidae.
I. Second joint of antennae very slender through-
out its length. [No orbits. Eye-stalks
generally long.] Inachince.
II. Second joint of antennae not very slender.
A. No true orbits (eye-stalks hidden under a
supraocular spine or sunken in the sides
of a great rostrum). Second joint of
antenna truncate-triangular. Eye-stalks
very short Acanihonychir
B. True orbits, containing both supra- and
postocular elements sheltering the eyes,
are more or less completely formed, except
in a few genera where the eye-stalks are
long and slender. Second antenna- joint
broad, usually not truncate-triangular.
Eye-stalks long or short.
1. A large, cupped, usually blunt postocular
process present. Eye-stalks short.
Cornea of eyes not completely hidden
when they are folded back Pisince.
2. Postocular process, if present, usually
sharp and not cupped, but if not so,
then cornea hidden (as also in most
other cases). Eye-stalks usually long . Maiince.
Key to the Sulfamilies of the Partlienopidse.
I. Carapace usually triangular, sometimes sub-
oval or subpentagonal. Rostrum simple.
481
Classification of the Decapod Crustaceans,
Chelipeds much bigger than the other legs.
Branchial regions of the body deeply sepa-
rated from 'cardiac Partlienopinm.
IJarapace usually sharply pentagonal. Ros-
trum cleft into two. Chelipeds of moderate
size. Branchial regions of the body not
deeply separated from cardiac Eurnedonince.
Key to the Families of the Brachyrhyncha.
I. Orbits formed, but more or less incomplete.
Second antennal flagella, when present,
long and hairy. Rostrum present. Body
elongate-oval. Fore edge of the mouth
indistinct
II. Orbits complete (though fissures may remain),
except in the Mictyrinae, where the eyes are
almost or quite unprotected. Body rarely
elongate-oval. Rostrum often wanting.
Second antennal flag<3lla usually short, not
hairy.
A. Carpopodites of third maxillipeds articu-
late at or near antero-internal angle of
the meropodites. Body usually round or
transversely oval. Male openings nearly
always coxal. In many species the right
chela is always larger than the left.
1. Legs more or less distinctly adapted for
swimming. Usually a small lobe on the
inner angle of the endopodite in the first
maxillipeds. [First antennae fold slanting
or transverse.]
2. Legs not adapted for swimming, or, if so
modified, then the vas deferens opens
sternally or runs in a sternal groove
(certain Macrophthalmus and Lihystes).
Inner lobe on the endopodite in the first
maxillipeds wanting.
a. Freshwater crabs with the branchial
region much developed and swollen.
[Body often squarish, but male open-
ing coxal.]
b. Marine crabs, with the branchial region
not greatly swollen,
i. First antennae fold lengthwise.
(a) Carapace subcircular. Second
antennal flagella either long and
hairy or wanting
(b) Carapace broadly oval or hexa-
gonal. Second antennal flagella
present, short, not hairy
Corystidae.
Portunidae.
Potamonidae.
Atelecyclidae
Cancridae.
* Trichia, de Haan, is somewhere in the neighbourhood of this
family.
482
Mr. L. A. Borradaile on the
ii. First antennaB fold slanting or
transversely.
(a) Body usually transversely oval.
Male openings rarely sternal.
Not sharply separated from the
following family
(b) Body usually square or squarish.
Male ducts open on the sternum,
or, if coxal, pass along a groove
in the sternum. Not sharply
separated from the foregoiug
family
B. Carpopodites of third maxillipeds do not
articulate at or near the inner angle of the
meropodites. Body usually square or
squarish. Male openings sternal, except
in Ptenoplax, where the duct passes along
a sternal groove to the coxopodite. In no
species is the right chela always larger
than the left.
1. Small symbiotic crabs, with very small
eyes and orbits. Body usually more or
less rounded
2. Free-living crabs, with eyes not specially
reduced and usually a square body.
a. Last pair of legs dorsally placed and
weaker than the others. Interan-
tennular septum very thin. [No
distinct epistome. Exopodites of
third maxillipeds not hidden.]
i. Front narrow. Female opening in
normal position. Third maxilli-
peds subpediform, not covering the
mouth
ii. Front moderately broad. Female
openings on the sternal segment
corresponding to first pair of walk-
ing-legs. Third maxillipeds cover
the mouth ventrally and have very
small meropodites
b. Last pair of legs not dorsally placed
nor markedly weaker than the rest.
Interantennular septum not very
thin, except in Macrophthalminae.
i. A gap of greater or less size is left
between the third maxillipeds.
Front broad or moderately so.
(a) Sides of the body either straight
or very slightly arched. Shape
square. Rarely true land-crabs.
{b) Sides of the body arched. Shape
transversely oval. Land-crabs .
ii. Third maxillipeds almost or quite
close the mouth. Front mode-
rately or very narrow
C. Meropodite in third maxillipeds small,
bearing terminally a carpopodite of
Xanthidae.
Gonoplacidae.
Pinnotheridae.
Ptenoplacidae.
Palicidae.
Grapsidae.
Gecarcinidae.
Ocypodidae.
Classification of the Decapod Crustaceans. 483
nearly its own width. Ischiopodite very
broad. [Body somewhat oblong. First
antenn® not retractile into sockets.
Parasitic on corals.] Hapalocarcinidae.
Key to the Subfamilies of the Portunidse.
I. Eye-stalk and orbits normal.
A. Basal joint of second antennae narrow.
[Flagella of second antennae not shut out
from orbits.]
i. First antennae sloping. Front with a
median tooth. Generally at least one
pair of walking-legs as long as chelipeds.
1. Last pair of legs not distinctly natatorial. Carcinidin<B.
2. Last pair of legs distinctly natatorial . Portumnincs.
ii. First antennae transverse. Front with a
median notch. Chelipeds longer than
walking-legs.
1. Last joint of fifth legs lanceolate .... CatoptrirnB.
2. Last joint of fifth legs rounded Carupince.
B. Basal joint of second antennae broad.
[Chelipeds longer than walking-legs.]
i. Flagella of second antennae not shut out
from orbits by processes of the basal
joints Portunince.
ii. Flagella of second antennae shutout from
the orbit by processes of the basal joints.
1. Last joint of fifth legs sickle-shaped . . Caphyrince.
2. Last joint of fifth legs fiattened ThalamithuB.
II. Eye-stalks enormously long, orbits extend
across the whole fore edge of the carapace.
[Chelipeds longer than legs. Antennae free ;
basal joint short ; fiagella not shut out from
orbits.] Podophthalmince.
Key to the Subfamilies of the Potamonidse.
I. Outward channels from gill-chamber covered
by first maxillipeds, reaching to front.
[Third maxillipeds with meropodites not
longer than broad, subtriangular, bearing
carpopodites at apex, with good exopodites.]
II. Outward channels from gill-chamber not as
in Beckeniinae.
1. Endostome ridges project on fore edge of
mouth. Exopodites of third maxillipeds
more or less reduced. [Meropodites of
same subtriangular, not longer than broad,
with carpopodites at apex.]
2. Endostome ridges do not proj ect on fore edge
of mouth. Exopodites of third maxilli-
peds not reduced.
i. Meropoditesof third maxillipeds not longer
than broad, subquadrate, with carpo-
podites at inner angles
Deckeniince.
Potamocarcinince.
Potamonince.
484
Mr. L. A. Borradaile on the
ii. Meropodites of third maxillipeds longer
than broad, bearing carpopodites on fore
edge, which slopes inwards Trichodactylince.
Key to the Suhfamilies of the Atelecyclidae.
I. Antennal flagella absent. [Mouth covered
bj third maxillipeds. Front uncleft.] .... Acanihocyclince.
II. Antennal flagella present.
A. Regions not defined. Third maxillipeds
cover the mouth. Front entire or lobed . Thiince.
R. Regions more or less clearly marked out.
Third maxillipeds do not cover the mouth.
Front toothed Atelecyclincri.
Key to the Suhfamilies of the Cancridse.
I. Carapace broadly oval. Epistome not sunken. Cancrince.
II. Carapace hexagonal. Epistome sunken .... FirimelincB.
Key to the Subfamilies of the Xanthidse.
I. Endostome ridges wanting. Shape of body
transversely oval or round.
1. Flagella of second antennae not shut out
of orbital gaps.
i. Second joint of second antennal stalk
cylindrical, reaching front but not
entering orbital gap iKanthince.
ii. Second joint of second antennal stalk as
in Xanthinae, but not entering orbital
gap CarpilincB.
2. Flagella of second antennae shut out of
orbital gaps by part of second joints of
stalks Etisince.
II. Endostome ridges present. Shape of body
often square or squarish.
1. Front less than | and fronto-orbital edge
(front and orbits together) not more than
f the greatest breadth of the carapace.
Front usually makes an arch with antero-
lateral edge. Flagella of second antennae
usually not shut out of orbital gaps.
i. Second joint of second antennal stalk
cylindrical and may or may not reach
the front, with which it is not broadly
in contact. Endostome ridges vary in
shape and size Menippince .
ii. Second joint of second antennal stalk
somewhat irregular in shape and broadly
in contact with front. Endostome
ridges strong and project on fore edge
of mouth Oziina;.
2. Front at least \ and fronto-orbital edge
more than f the greatest breadth of the
carapace. Front makes an angle with the
antero- lateral edge. Flagella of second
antennae always shut out of orbital gaps.
4S5
Classification of the Decapod Crustaceans.
i. Endostome ridges strong and project on
fore edge of mouth Ei'iphiiru^.
ii. Endostome ridges moderate and make no
projection on fore edge of mouth .... TrapeziiruB.
Key to the Subfamilies of the Gonoplacidse.
I. Last pair of legs present.
1. Male openings sternal. Eye-stalks almost
invariably fixed. Eyes often reduced.
Front usually narrow. [Male abdomen
does not nearly cover space between last
two legs.]
2. Male openings coxal. Eye-stalks generally
movable. Eyes normal. Front broad,
i. Front occupies whole breadth of carapace.
(1) Male abdomen does not cover space
between last pair of legs Prionoplacince .
(2) Male abdomen covers space between
last pair of legs Gonoplacinas.
ii. Front does not occupy whole breadth of
carapace. [Male abdomen covers space
between last pair of legs.] Carcinoplacince.
Last pair of legs wanting. [Male openings
sternal. Male abdomen does not cover space
between last pair of legs. Eyes small.
Front narrow.] llexapodinct.
Key to the Subfamilies of the Grapsidse.
I. First antennae placed in clefts of the front
and visible from above. [No oblique line
on the third maxillipeds and no wide gap
between them. Male abdomen covers the
space between the last pair of legs.] Plagusiince.
II. First antennae not visible from above.
1. Third maxillipeds have an oblique hairy
line along the ischiopodite and meropo-
dite, and leave a wide gap between them. Sesarmince.
2. Third maxillipeds have no oblique line.
i. Front strongly deflexed. A wide gap
between the third maxillipeds. Male
abdomen covers the space between the
last pair of legs OcypodmcB.
ii. Front not strongly deflexed. The gap
between the third maxillipeds, if present,
is rarely wide. Male abdomen rarely
covers space between last pair of legs . . Varunirux.
Key to the Subfamilies of the Ocypodidas.
I. First antennae transverse, separated by a nar-
row septum. Front of moderate breadth.
Body shallow, usuallv quadrilateral and
broader than long. [jN"o opening between
the bases of the legs.] Macrophthalmince.
48G Mr. G. A. Boulenger on
II. First antennfB oblique or vertical, separated
by a broad septum. Front narrow. Body
deep.
1. Body subquadrilateral. Openings fringed
with hairs between the bases of the third
and fourth pairs of legs Ocypodince.
2. Body more or less globose. No openings
between the legs Myctirinai.
LV. — Descriptions of new Lizards in the British Museum.
By G. A. Boulenger, F.R.S.
Gecho palmatus.
Head large, oviform, much depressed; snout a little longer
than the distance between the eye and the ear-opening, once
and one third the diameter of the orbit ; forehead concave ;
ear- opening oval, oblique, its greatest diameter about one
third that of the orbit. Body and limbs moderately elon-
gate ; digits strongly dilated, half-webbed. Snout and
supraocular region covered with very small granules, the
granules minute on the rest of the head ; rostral and first
labial entering the nostril ; 11 upper and 10 lower labials ;
symphysial triangular, followed by a pair of chin-shields
which are three times as long as broad. Body and limbs
covered with minute granules ; back with scattered, enlarged,
round, flat tubercles, which are about as large as the feebly
imbricate ventral scales. Greyish above, mottled with
brown, and with brown markings, viz. a dark streak from
the eye to the ear and three wavy cross-bars on the body ;
lower parts white, dotted with brown.
mm.
Total length 150
Head 22
Width of head 17
Body 58
Fore limb 27
Hind limb 36
Tail (reproduced) 70
A single female specimen from the Man Son Mountains,
Tonkin, altitude 3000-4000 feet, collected by Mr. Fruhstorfer.
Prionodactylus Ochendeni.
Snout short ; nostril between two nasals ; fronto-nasal
single; preefrontals forming a median suture; interparietal
narrower than the parietals ; two rows of occipitals, tliree in
•new Lizards in the British Museum,
487
the first row ; three supraoculars, first largest ; a single
loreal and a freno-orbital ; a series of small infraorbitals ;
temple with large shields above; 7 upper and 4 to 6 lower
labials ; chin-shields, one anterior and 3 or 4 pairs, the last
separated by a pair of large gulars; a double longitudinal
series of transversely enlarged gulars in front of the collar,
which is composed of 7 or 9 plates. Dorsal scales elongate,
hexagonal, strongly keeled, imbricate, not mucronate; lateral
scales small, keeled ; 36 to 38 scales round the body (in-
cluding ventrals), 35 to 38 from occiput to base of tail.
Ventral plates quadrangular, in 8 longitudinal and 18 to 20
transverse series, outer smallest and keeled, median largest.
3 or 4 large anal plates in a transverse series. Male with
8 to 10 femoral pores on each side. Pale brown above, dark
grey on the sides; a more or less distinct dark brown verte-
bral line ; a dark brown lateral band, light-edged above,
fading into the grey of the sides below, from the eye to the
tail, usually bearing a series of small black ocelli with white
centres ; a white, dark-edged streak from below the eye to
the side of the neck, lost before reaching the collar; belly
white, with grey dots or round black spots.
d. $.
nun. jnm.
Total length ? 189
Head 16 14
Width of head 10 8
From end of snout to fore limb .... 23 22
„ vent 60 61
Fore limb 19 18
Hind limb 26 26
Body 14 14
Tail • ? 128
Several specimens from Carabaya, E. Peru, altitude
6000-7000 feet, collected by the late Mr. G. Ockenden.
Amphishoina Slateri.
^ Snout rounded, prominent. Rostral small, triangular, not
visible from above ; nasals forming a suture ; a pair of prse-
frontals, the suture between them nearly twice as long as
that between the nasals and a little longer than that between
the frontals, which are much smaller ; a pair of occipitals ;
eye distinguishable under the ocular, which is situated
between the praefrontal and the second and third upper labials
and followed by two superposed postoculars; four upper
labials, fourth very small, second largest ; symphysial fused
with a large chin-shield ; four lower labials, second largest.
488
Oil new Lizards in the British Museum.
210 annuli on the body, 12 on tlie tail; an annulus in the
middle of the body contains 22 segments, 10 dorsal and 12
ventral, the dorsals all longer than broad, the median ventrals
a little broader than long. Lateral line very distinct. 6 anal
segments. 4 prseanal pores. Uniform dark purplish brown.
Length to vent 140 mm. ; tail 12; diameter of body 4.
A single specimen from Peru, obtained in the Rio San
Gaban Valley, Prov. Carabaya, altitude 2000-3000 feet, by
Mr. Thomas Slater, and presented by him, through Prof. G. S.
Boulger, to the British Museum.
The small number of segments in an annulus on the body
well distinguishes this species from A. vermicularis and its
South-American allies.
Nucras Emini,
Body moderately elongate ; head moderate, slightly de-
pressed, its length four and one third times in the distance
from snout to vent. Two superposed postnasals ; no granules
between the supraoculars and the supraciliaries ; interparietal
large, not twice as long as broad, in contact with a small
occipital ; subocular between the fourth and fifth upper
labials; temple covered with large granules; a large tym-
panic shield ; two supratemporals. Collar slightly toothed,
formed of 7 plates. Dorsal scales granular, smooth, 42
across middle of body ; ventral plates in 8 longitudinal and
28 transverse series. 2 large praeanal plates, one before the
other. Hind limb reaching the axil ; foot longer than the
head. 12 femoral pores on each side. Caudal scales strongly
keeled. Pale reddish brown above, with black dots ; a black
lateral band from the eye to the root of the tail, bearing a
series of white spots and light-edged above and beneath ;
lower parts white.
mm.
Total length
Head
Width of head
From end of snout to fore limb
Fore limb . . .
Hind limb .
Foot
Tail (injured)
120
11
7
16
46
15
24
14
74
A single small specimen from the south shore of Lake
Victoria, from Emin Pasha’s collection received in 1890.
Lygosoma meleagris.
Section Siaphos. Body much elongate; limbs small, with
four very short digits ] the distance between the end of the
489
On the Variations of Stereolepis gigas.
snout and the fore limb is contained twice and a half in the
distance between axilla and groin. Snout very short,
obtuse. Lower eyelid scaly. Nostril pierced between two
nasals ; no supranasal ; fronto-nasal broader than long,
broadly in contact with the rostral and wuth the frontal ;
prsefrontals minute ; frontal not much larger than fronto-
parietals, in contact with the first and second supraoculars ;
four supraoculars ; five supraciliaries ; frontoparietals dis-
tinct, larger than the interparietal ; parietals forming a suture
bel)ind the interparietal ; a pair of nuchals ; fifth upper
labial below the centre of the eye. Ear-opening minute.
22 smooth scales round the middle of the body. Median
praeanals scarcely enlarged. The length of the hind limb
equals the distance between the anterior border of the eye and
the fore limb ; third and fourth toes equal. Tail long and
thick. Upper surface of head and back blackish brown,
with small round white spots ; sides of body, belly, hind
limbs, and base of tail uniform orange ; a black streak on the
temple and along the side of the neck ; throat black ; greater
part of tail brown above and white beneath, spotted with
black.
mm.
Total length 166
Head 10
Width of head 7
Body 51
Fore limb 10
Hind limb 15
Tail (reproduced) 105
A single specimen from Mount Ruwenzori, altitude
7000 feet. Presented by the Subscribers to the Ruwenzori
Expedition Fund.
LVI. — On the Variations of Stereolepis gigas, a great Sea-
Perch from California arid Javan. By G. A. Boulenger
F.R.S.
When preparing a revision of the Sea-Perches for the
‘ British Museum Catalogue of Fishes,’ vol. i., published in
1895, a comparison of the descriptions and figures given of
Stereolepis gigas, Ayres, from California, and Megaperca
ischinaga, Hiigendorf, from Japan, failed to bring out any
tangible difference between the two, which I accordingly
proposed to unite under the former name. A comparison of
Ann. & Mag. N. Hist. Ser. 7. Vol. xix. 34
490
Mr. G. A. Boulenger on the
an adult Californian specimen with a photograph of Hilgen-
dorPs types in the Berlin Museum, which I made soon after
(P. Z. S. 1897, p. 917, pi. lii.), confirmed me in this opinion.
Dr. D. S. Jordan, although accepting the generic identifi-
cation of the two fishes, could not be convinced of their
specific identity ; and in a paper which he published last year
(P. U.S. Nat. Mus. XXX. p. 841, fig.)^ conjunction with
Mr. J. O. Snyder, he emphatically declares the Japanese fish
to be “ well separated from Stereolepis gigas^ Ayres, of the
coast of California, by the larger scales, and especially by
the form of the spinous dorsal fin, the spines in Stereolepis
gigas being very much lower. The nominal genus Megaperca^
however, differs but slightly from Stereolepis^ the only
tangible character resting in the marked elevation of the
dorsal spines, the first dorsal being low in Stereolepis, The
scales are a shade thicker and rougher, but the difference is
491
Variations of Stereolepis gigas.
not one of importance.” This latest description, accompanied
by a figure, is taken from a specimen 14 inches long, although
the species is known to grow to five times tliat length. Had
Dr. Jordan not overlooked my description and figure in 1897,
as he courteously informs me he inadvertently did, he could
not have stated that the spinous dorsal fin is appreciably more
elevated in the adult Megaperca than in the adult Stereolepis
(young specimens of the latter are, I believe, still unknown).
It seems hardly credible that so experienced an ichthyologist
as Dr. Jordan should overlook the enormous changes in the
comparative depth of the spinous dorsal which take place
with age in all Bass-like fishes, and in order to emphasize this
point I here give outline-figures (a) of a small (1 foot long)
and {h) a large (4 feet long) Nile Perch {Lates niloticus).
I have specially selected the Nile Perch as an example,
because, having been able to study a large number of
specimens, I have no fear of having confounded two species.
In the young Megaperca the longest spines measure about
half the depth of the body, in the adult (photograph of the
type) exactly one fourth. I therefore cannot accept the
differences in the dorsal spines as being due to anything more
than the usual changes which take place with age, and until
the young of the Californian fish is known we may safely
assume that its first dorsal is much more elevated than in the
adult. In a letter addressed to me a short time ago Dr. Jordan
adds that the Japanese fish has “ much larger scales.” I have,
I think, disposed of the supposed distinction in the dorsal fin ;
I will now give some facts against the second distinctive
character, which, so far as I know, is the only one that would
stand after a comparison of the figures given by me in 1897.
In 1895 I gave the scale-formula, compiled from different
sources (Japanese and American specimens), as 80-100 1|.
In the Californian specimen examined by me in 1897 I
counted 115 In their young Japanese specimen Jordan
and Snyder counted 87 In two specimens from Japan
(Sagami Bay), now preserved in the British Museum, and
measuring 19 and 15 inches respectively, I find 90 in the
first, 105 ^ in the second, and my counting has been verified
by my colleague Mr. Began. These numbers seem to me
to dispose entirely of the alleged difference in the size of the
scales as a specific character.
34*
492 Mr. G. A. Boulenger on an African Barbel,
LVIT. — On an African Barbel hitherto confounded with
Barbus trimaculatus, Peters. By G. A. BoulengER,
F.K.S.
Following the lead of Gunther Steindachner t, and Max
Weber J, I have hitherto referred to Barbus trimaculatus a
small barbel common in British Central Africa, Rhodesia,
Portuguese East Africa, Angola, the Transvaal, and Zulu-
land. Having recently had an opportunity of examining
great numbers of these fishes, I have come to the conclusion
that they cannot possibly be identified with the species so
carefully described and figured by Peters §. As suggested
by the latter author, B. trimaculatus^ established on a single
example from the Kovugo River, Mozambique, is very closely
related to, if not identical with, B. trispilus^ Bleeker, of
which I have recently given a description 11. The species
for which I now propose the name B. decipiens differs from
B. trimaculatus and B. trispilus in its strong ossified last
simple dorsal ray, shorter barbels, and more numerous scales
in the lateral line. It would never have been confounded
with B. trimaculatus but for its deceptive markings, which,
moreover, are by no means constant. I append a description
based on the following series of specimens : —
2, R. Ruo, Brit. Cent. Africa. — Sir H. H. Johnston.
1, between Kondowe and Karonga, Brit. Cent. Africa. —
Sir H. II. Johnston.
6, near Salisbury, Rhodesia. — Guy A. K. Marshall.
13, Mazoe R., Rhodesia. — J. ff. Darling.
6, Beira, Portug. E. Africa. — C. Grant.
2, Groot Olifant R., Transvaal. — Capt. G. E. Bruce.
5, Klein „ ^ ,, ,, ,,
2, near Komati Poort, ,, „
2, Potchefstroom, Transvaal.- — Stenning,
28, E. of Pietersburg, Transvaal. — C. Grant.
4, Umfulosi, Zululand. — C. Grant.
1, Elcheleselwane, Zululand. — Dr. E. Warren.
9, Mossamedes, Angola. — Dr. W. J. Ansorge.
Barbus decipiens.
Depth of body 3 to 4 times in total length, length of head
3| to 44 times. Snout rounded, as long as or a little longer
Proc. Zool. Soc. 1893, p. 619.
t Sitzb. Ak. Wien, ciii. i. 1894, p. 452.
X Zool. Jahrb., Syst. x. 1897, p. 151.
§ Reise n. Mossamb. iv. p. 55, pi. xi. fig. 4 (1868).
|j Ann. & Mag. Nat. Hist. (7) xviii. 1906, p. 33.
On SpirocliaBta (Trypanosoma) Balblanli_, (kc. 493
tlian the eye in the adult ; diameter of eye 3 to times in
length of head, interorbital width 2\ to times ; mouth
slightly inferior, with feebly developed lips, interrupted on
the chin ; barbels two on each side, anterior about as long as
eye, posterior 1 to 1-^ diameters of eye, the distance between
them about J diameter of eye. Dorsal III 7-8, last simple
ray strong, bony, not serrated, nearly straight, or feebly
curved, to once length of head ; free edge of the fin feebly
emarginate ; its distance from the occiput less than its
distance from the caudal fin. Anal III 5, longest ray about
f length of head. Pectoral § to | length of head, not
reaching ventral ; latter below anterior rays of dorsal.
Caudal peduncle 1| to 2 times as long as deep. Scales
32-36 3-4 between lateral line and ventral, 14-16 round
caudal peduncle. Brown above, silvery on the sides and
below ; three more or less distinct round blackish spots often
present on each side, the first and second just above the
lateral line, one in front and one behind the vertical of the
base of the dorsal, the third at the base of the caudal fin and
traversed by the lateral line ; these spots may be absent or
reduced to the one at the base of the caudal fin ; fins whitish,
without spots.
This fish grows to a length of 125 mm., but is usually
smaller, about 100 mm. long.
LVIII. — Spirochaeta (Trypanosoma) Balbianii {Certes), its
Movements, Structure, and Affinities ; and on the Occurrence
of Spirochaeta anodontae {Keysselitz) in the British
Mussel, Anodonta cygnea. By H. B. Fantham, B.Sc.,
Derby Research Scholar, University College, London;
and St. Mary’s Hospital Medical School.
(Preliminary Account.)
Introduction,
There are few more interesting organisms at present under
investigation than those microscopic yet most active forms
known as Spirochaetes, which lie near the border-line of
plants and animals. They are, indeed, veritable members of
Haeckel’s kingdom Protista, and it is still a disputed point
whether they are really Protozoa or Bacteria.
494
Mr. II. B. Fantham on Spirochgeta
Historical,
Spirochceta [Trypanosoma) Balhianii first recorded by
Certes from French oysters in 1882 [i], though Mobius,
writing in 1883 [7], stated that he observed the parasite in
1869 in oysters from Schleswig-Holstein. It was found in
the crystalline style and intestine of the host. Lustrac [6],
in 1896, gave particulars of longitudinal division. In 1901
the famous protistologists Laveran and Mesnil [5] briefly
described the main features of the organism, and stated that
it was really a Bacterium allied to the Spirilla and Spiro-
chsetes. In 1905-06 Perrin [8, 9] described the life-history
of the organism, but still retained it in the genus Trypano-
soma, although it lacks a flagellum, Certes having placed it
in this genus years before because it possessed an undulating
membrane ; but at that period the genus Trypanosoma was
ill-defined. Short notes have since appeared by Swellen-
grebel [10], Vl^s [ii], and Fantham [3].
Spirochceta anodontce was recorded from the crystalline
style of Anodonta mutabilis (which is not a British species)
by Keysselitz in 1906. It has not before been noted in
A, cygnea, so far as I am aware.
Material and Methods,
Spirochceta Balhianii has been studied by me in oysters at
Roscoff last summer, and in this connexion I would especially
thank Professor Delage and M. Fred Vl^s. The further
studies on this organism were continued in London, on more
living material, though infected oysters were only procured
with much difficulty.
As regards Spirochceta anodontce from the crystalline style
of Anodonta cygnea, after repeated attempts, during which
only a few parasites were found, I obtained an infected strain
in Anodons from Godaiming, very kindly supplied to me by
Mr. O. H. Latter, to whom my best thanks are tendered.
I have spent much time examining both these Spirochsetes
in the living condition, in their natural medium as far as
possible. I am convinced that too much stress cannot be
laid on the necessity for this careful examination of living
material ; it is not sufficient to rely on stained preparations
alone.
As regards fixed and stained material, best results were
obtained from thin smears of gut-contents or solutions of the
crystalline style (in a little sea-water in the case of Ostrea,
in fresh water in the case of Anodon), the preparations being
(Trypanosoma) Balbianii and Sr anodontse. 495
fixed wet with osmic vapour. Other fixatives used were
Flemming’s solution, corrosive sublimate and alcohol, and in
the case of some dried smears pure methyl and ethyl alcohols.
The preparations were usually mounted in cedar-wood oil
or balsam.
The most useful stains were gentian violet (Ohlmacher’s
formula), iron-alum hseraatoxylin, thionin. Billet’s modifica-
tion of the Griemsa stain, and Delafield’s hsematoxylin, while
dilute methylene-blue was best for intra vitam staining. Too
much reliance must not be placed on the various modifications
of the Komanowsky stain, for the structure of the membrane
is often only poorly revealed thereby.
Movements of these Spirochcetes.
Previous accounts of these phenomena are most meagre,
and yet descriptions of such movements would be of the utmost
importance. At this juncture it is necessary to state that
a typical Spirochsete possesses an undulating membrane.
The type species of the genus, 8. plicatiUs^ Ehrenberg,
recorded in 1833 from muddy water, was shown by Schaudinn
in 1905 to possess such a membrane.
The movements of each of the two Spirochsetes in question
are most complex and difficult to resolve and interpret. A
Spirochsete moves very rapidly, especially 8. anodontce —
indeed, so rapidly that it is almost impossible to analyze its
motion when travelling at full speed. Its path may be either
in a straight line or more or less in a circle. In the case of
slowly moving specimens it is seen that the organism moves
forward while turning on its long axis. The motion, then,
appears to be resolvable into at least two components — (i.) a
vibratory motion of flexion of the body, mainly for progres-
sion, and (ii.) a spiral or corkscrew movement of the body
as a whole, due to the winding of the membrane. The
corkscrew motion is especially well seen in the case of
8. anodontce, which has pointed ends.
Waves can be seen travelling down the thread-like body in
a direction opposite to that in which the organism is pro-
gressing. Many waves or sinuosities, some eight or ten, can
be seen along the body of rapidly moving forms, while only
some two to four may occur in more slowly moving ones.
The outline of the sinuosities is sometimes a little irregular —
that is, the contour is somewhat broken by much smaller
waves.
The movements occur in jerks. The organism may
suddenly come to a dead stop or just as suddenly proceed
496
Mr. II. B. Fantliam on Spirocliaeta
more slowly. It is a matter of indifference which end of the
body is directed forwards, for the parasite is capable of
suddenly reversing its direction of movement and returning on
its own path, apparently even in an almost exact straight line
or circle. I do not consider this retracing of its path to be
due to unfavourable environment, as suggested by Novy in
the case of S. Ohermeieri^ for I have observed it constantly
taking place inside the crystalline style. The organism can
then travel with or against the current indifferently. A very
great deal of energy seems to be used in the motion of
tlie atiimal. The body of the organism can be distinguished
during motion in the case of S. BaTbianii'^\\}a some difficulty,
but sometimes S. anodontce moves too rapidly for its outline
to be clearly discernible.
Sometimes the organism appears suddenly from a deeper
level of the liquid under examination, and swims, or, rather,
spirally bores its way more or less vertically upwards. In
this condition it twists itself into various peculiar shapes, and
so resembles a Catherine wheel as described by Perrin.
Parasites are sometimes noticed anchored by one end to a
separated or shed epithelial cell (from the gut of the host)
lying in the gut-contents. The free end of the parasite then
executes violent lasliing movements or intermittent flickers.
The free end also in such specimens may coil itself over and
over. S. Balhianii^ with its rounded ends, often has some
difficulty, apparently, in boring its way through debris or
obstacles in its path. It often tries to get through these
instead of changing its direction of motion. However, I
have seen it penetrate free epithelial cells and occasionally
appear to come out of such cells.
/S'. anodontcB^ with its pointed ends, is capable of rather
more rapid motion and more easily bores its way through the
debris of cells floating in the gut-contents of the host.
Spirochsetes may at times be seen vibrating in two halves
about their central points as nodes ; they then resemble, to
some extent, two tuning-forks joined by their single ends
and in vibration.
Slowly moving specimens of S. anodontce may curl up one
end, usually the hinder end, judging by the direction of
motion.
These Spirochsetes seem to move more quickly than
Trypanosomes, and with an added corkscrew motion. Also
the body of a Spirillum seems more rigid than that of a
Spirochsete in motion, and of course flagella are present in
the case of true Spirilla.
Various scientific workers — both zoologists and bacteriolo-
(Trypanosoma) Balbianii and S. anodontse. 497
gists — to whom I have shown these Spirochsetes alive, have
compared their motion to that of an eel, or the embryo of
Filaria, or to that of Nereis^ but with the spiral movement
in addition.
General Structure.
The general shape of each of these Spirochaetes is that of
a long narrow thread. S. Balbianii varies from 50 yu to
150 yu in length, and is 2 yu- to 3 yu broad. Its breadth is
almost uniform and its ends are rounded. S. anodontce is
about 35 yLt to 40 yu long and about 0‘7 yu broad. Keysselitz [4]
did not give the dimensions of his parasite from A. mutahilis.
The ends of S. anodontce are pointed and prolonged into a
filament-like process, which some observers might be disposed
to call a flagellum, though it seems to me to be stiff.
The body is bounded by a well-marked periplast which
encloses a homogeneous cytoplasm. There is no marked
differentiation of this cytoplasm into ectoplasm and endo-
plasm, though I tliink the periplast represents an ectoplasm.
The periplast is not composed of cellulose (as tested by iodine
and sulpliuric acid), but might be composed of fungus-
cellulose ” so far as negative evidence goes. Perrin, judging
from macerated specimens, considered the periplast to be
fibrillar in structure. This is very difficult to corroborate,
but is probably correct, for these fibrillae might equally belong,
in macerated specimens, to the membrane, to be described
in the next section.
In the centre of the thread-like organism is a core of
chromatin, the nucleus, stretching nearly from end to end,
which consists of a filament on which are arranged transverse
bands or rodlets of deeply staining chromatin at more or less
regular intervals. These rodlets may be perhaps termed
“ chromosomes,” as by Perrin ; many are irregularly dumbbell-
shaped, while some are thinner than others. Perrin considered
the connecting spiral or zigzag-shaped filament to be a
karyosome. He figures various nuclear changes in connexion
with gamete formation, which are, however, open to varied
interpretations. In preparation for longitudinal division the
dumbbell-shaped rodlets divide, and leave two rows of
chromosomes,” one on each edge, along the periphery of
the parasite. I have not seen definite evidence of reducing
division.
The nucleus was not observed by the earlier writers on
these parasites. It is of a difi’use character, and there is no
definite blepharoplast, facts first noted by Laveran and Mesnil,
the significance of which is of the highest importance
408 Mr. II. B. Fantham on Spirocliseta
There is a cap or nodule attached to each end of the
periplast in S, Balbianii which stains pinkish with Giemsa’s
solution, and which may be a basal granule.”
The chromatin granules of S. anodontce are difficult to see,
the whole organism being much smaller, but the nucleus
seems to conform to the general arrangement of that structure
in S. Balbianii,
The Undulating Membrane,
This structure does not extend quite to the ends of the
organism. There is some difference of opinion as to its
nature, but its presence is a characteristic feature of the
genus Spirochceta as now defined.
It is, I believe, a spirally wound membrane or lateral
outgrowth of the ectoplasmic periplast, and is composed of
longitudinally arranged fibrillse. These fibrillse are contractile
and may be termed myoneme fibrillae,” though the term
myoneme ” is perhaps not a happy one. There appear to
be some eight or nine principal fibrils, and many less evident
ones parallel therewith, all of them apparently longitudinally
arranged. The border of the membrane is thickened and
stains well with chromatin stains. I agree with Perrin that
the membrane possesses a chromatic border.
In slowly moving parasites the edge of the membrane, and
even the membrane itself, is easily seen (Zeiss DD and ocular 2)
in life. Such forms are curved into about two or three
sinuosities only, and the membrane can be seen in the troughs
of these, loosely arranged and not contracted close to the
body.
It has been suggested by Laveran and Mesnil that this
structure is not really a membrane, but a sheath, only
loosely attached, if at all, to the body of the organism. This
matter is most difficult to settle, and is certainly not to be
passed over lightly by mere dogmatic assertion either one
way or the other. However, one can see in preparations the
spiral arrangement of the membrane in many cases, and its
actual crossing over the body from one side to the other, both
above and belo\^^ And, furthermore, the organism possesses
a distinct spiral motion, evidently guided by the spiral
arrangement of the membrane. Again, if the body moved
more or less freely inside a sheath one might expect to see
some signs of differential motion between the organism and
its sheath. I have never seen such during my investigations.
It seems to me, then, that this structure is a membrane,
spirally wound round the body, and is an outgrowth of the
periplast. It is composed of elastic fibrils and is contractile.
499
(4’rypanosoma) Balbianii and S. anodontae.
and is probably the locomotor agent of the organism. Its
own vibrations are veiy slight, and by its alternate contrac-
tion and expansion it appears to control the movements of
the organism. The structure and function of this organella”
are most difficult to interpret. The myonemes, I think, set
up transverse movements in the surface of the body, manifested
as waves passing down the body in a direction opposite to
that in which the organism progresses.
It has been stated that the nuclear core is contractile
(Swellengrebel [lo]). I am rather inclined to the view that
it is only flexible.
It has been suggested [i i] that there is a ciliate stage in
the life-history of S. Balbianii — in other words, that the
membrane is built up from the agglutination of cilia
flagella ” of English bacteriologists), or even that the
membrane may be decomposed into these flagella. I have
myself preparations of this character made both at Roscoff
and in London. It seems to me that these apparent flagella
or cilia are really elastic fibrils, staining pink with Giemsa’s
or Leishman’s stain, “ myoneme fibrils ” in fact, split off
from the membrane during its rupture, which sometimes
occurs during the violent contortions and death-struggles of
the organism, especially in a damp atmosphere like that of
Roscoff. It may be that the membrane, as suggested in a
private communication, is really a “ ciliated membrane.” I
have never seen these flagella during life.
The fibrillar nature of the membrane is none too well
shown by the various modifications of the Romano w sky
stain. It is best revealed by gentian-violet or iron-
hsematoxylin.
In life, during active movements, the membrane is closely
contracted round the body, and is not easily seen except as a
halo round the organism.
The membrane of 8. anodontce is most difficult to discern,
but it can be seen in favourably placed specimens both in
life and when stained. It appears to conform to the structure
described for 8, Balbianii.
Division.
Longitudinal division has been described before in the case
of these Spirochsetes. I have seen such during life and in
stained preparations.
However, I believe transverse division also occurs, for in
the case of 8. Balbianii both long and short forms are seen.
In some stained preparations there were somewhat long forms
500 On Spirocliseta (Trypanosoma) Balbianii,
with tlie membrane discontinuous in the centre, where a
vacuole-like space occurred ; the edges of this space were
sharp, not torn, while the periplast appeared just continuous
over the gap. Laveran and Mesnil [5] have described trans-
verse fission. I think this mode of division naturally occurs,
and such cases are not to be explained away [8, 9] as merely
two forms which, having divided longitudinally, are now
separated by 180°, just before complete separation. Further,
in living specimens one sometimes sees forms vibrating about
a node, possibly not central, but I have never seen in life
division about such a point.
Affinities.
Spirochsetes are Protists : doubtless all will agree to this.
Next arises the vexed question. Are they Protozoa or
Bacteria ?
Their diffuse nuclei and transverse fission rather suggest
Bacteria ; while, on the other hand, the presence of an undu-
lating membrane, longitudinal fission, and even definite
^^chromosomes” suggest Protozoa. Perhaps, too, their size
is great for Bacteria, though too much importance must not
be attached to questions of size probably.
I am not at all sure about the presence of male and female
forms [8, 9] or of encystment [8, 9]. I think these appear-
ances can be otherwise interpreted, the male and female forms
being possibly the extremes from a more or less continuous
series.
On the whole, I somewhat incline to the Protozoal nature
of these organisms, but prefer to leave further discussion of
this and other points for my longer illustrated memoir, which
will, I hope, appear in the Quart. Journ. Microsc. Sci.
POSTSCKIPT. — To this preliminary account it might be of
interest to add that Spirochceta Balhianii was found moving
about freely in a basin of sea-water in which specimens of
infected English oysters were kept, the shells of these oysters
being still partly covered by mud from their native beds.
I was able to demonstrate this, with living specimens, to
some of my friends.
I have also examined specimens of the Acarid, Atax Bonzi,
Clap., found in the mantle-cavity of Anodonta, but have not
yet succeeded in finding any Spirochsetes in the Acarids,
although Spirochceta Duttoni occurs in ticks.
On Pterocyon, Rousettus, and Myonycteris. 501
References to Literature.
[i] Certes, A. 1882. ^‘Notesur les Parasites et les Commensaux
de riluitre,” Bull. Soc. Zool. de France, vii. pp. 347-353, 4 figs.
[z] . 1891. “ Sur le Trypanosoma Balhianiii' op. cit. xvi.
pp. 95, 130.
[3] Fantham, H. B. 1906-07. Spirochreta BalbianiiL Abstr. of
Proc. Zool. Soc. no. 37. Spirochmta anodontcej''’ op. cit. no. 45.
[4] Keysselitz, G. ‘1906. Spirocliceta anodontce, n. sp.,” Arb.
kaiserl. Gesundheitsamte, xxiii. pp. 566-567, 6 figs.
[5] Laveran, a., et Mesnil, F. ‘‘ Sur la Nature bacterienne du
pr^tendu Trypanosome des Huitres (T. Balbianii, Certes),” C. R.
Soc. Biol. liii. pp. 883-885.
[6] Lustrac, a. de. 1896. ^^Trypanosoma Balbianii, Certes,” Actes
Soc. Linn. Bordeaux, 5® s4r. t. x. pp. 265-275, 2 pis.
[7] Mobius, K. 1883. ^^Trypanosorna Balbianii, CQvtQ&,\m'Kr
stiel Schleswig holsteinischer Austern,” Zool. Anzeiger, vi. p. 148.
[8] Perrin, W. S. 1905. “ A Preliminary Communication on the
Life-history of Trypanosoma Balbianii,'' Proc. Roy. Soc. 76 B,
pp. 308-375, 4 figs.
[9] . 1906. Researches upon the Life-history of
Balbianii," Arch, f . Protistenkunde, vii. pp. ] 31-156, 2 pis.
[10] SwELLENGREBEL, N. H. 1907. “ Sur la cytologie comparee des
Spirochetes et des Spirilles,” C. R. Soc. Biol. Ixii. pp. 214-215.
[11] Vees, F. 1906. Sur la Structure et les Affinit^s de Trypano-
soma Balbianii," C. R. Soc. Biol. Ixi. pp. 408-410.
LIX. — On Pterocyon, Rousettus, and Myonycteris.
By Knud Andersen.
The present paper is a brief summary of notes written down
during a revision of the fruit- bats of the genera Pterocyon,
Rousettus, and Myonycteris in the collection of the British
Museum.
Synopsis of the Genera and Species.
Common characters : tongue normal ; molars
well developed ; occipital region of skull not
elongated and tubular ; incisors | — ; postcanine
teeth f ; second digit clawed ; a short tail.
a. Cranial rostrum longer : front of orbit verti-
cally above posterior half or middle of ;
basicranial axis markedly deflected.
a! . Premaxillaries separated in front; tym-
panies forming a bony auditory meatus ;
in cross section twice the size of an
upper incisor ; w, equal in length to
and together. General size large:
forearm 114-132 mm. Ethiopian .... Bterocxjou.
502
Mr. K. Andersen on
d^. Kostrum relatively longer: front of
orbit to tip of nasals equalling or
exceeding maxillary tooth-row ; fur
longer, more woolly, not closely ap-
pressed ; colour darker. Forearm
127-131 mm. Madagascar
Rostrum relatively shorter; front of
orbit to tip of nasals less than maxillary
tooth-row ; fur very short, closely ap-
pressed ; colour lighter.
a^. Skull larger : total length 54-5-62‘5
mm. ; tooth-rows longer : c-m^ 21-
23'8; molars narrower. Forearm
117'5-132. African continent. . . .
Skull smaller: total length 51 ‘5-55
mm.; tooth-rows shorter: c-m^
19-2-20-8 ; molars broader. Fore-
arm 114-127, S.W. Arabia ....
b' . Premaxillaries in contact or co-ossified in
front ; no bony auditory meatus ;
reduced : subequal in size to an upper
incisor ; wq shorter than and
together. General size moderate : fore-
arm 69’5-99 mm. Ethiopian, Oriental,
A ustro-Malayan
c^. Wings from back of 1st toe,
c®. Pollex (with claw) 30-37-5 mm, ;
2nd phalanx of 3rd digit 50-5-61-5.
Ears not attenuated at tip ; lower
leg 40-45-5 mm.
Skullsmaller : total length 40-5-
43-8 mm. ; rostrum slenderer ;
palate ridges normally 3-1-4
-j-1*. Forearm 89-99. S.
Africa
Skull larger: total length 43-6-
46-7 mm. ; rostrum heavier ;
palate ridges 4 -f- 4-1-1. Fore-
arm 88-99, Angola to Pales-
tine
5^. Ears attenuated at tip ; lower
leg 37-39-5 mm . ; forearm 87-96.
Arabia to Sind
iF, Pollex (with claw) 24-30 mm. ; 2nd
phalanx of 3rd digit 36-47-2.
c^. Molars not unusually narrow ; fur
short ; notopatRgium naked,
cb Wg elliptical in outline ; width
of ears (flattened) 14-5-15-8mm.
not deciduous ; fur on nape
and shoulders not unusually
scarce. Forearm 80-5-87-5
mm. India, Himalayas, to
S. China
Pt. Dupreanus.
Pi. heivus.
Pt. sahcms.
Pousettus.
P. Lecichi.
P. cepyptiacus.
P. arahicns.
P. LeschenauUi.
* 3 anterior, undivided ; 4 middle, interrupted in the median line ;
1 posterior, at palation border.
503
rterocyon, Rouscttus, and Mjonjcteris.
deciduous ; nape and shoul-
ders seminaked. Forearm
79-85*5 mm. Ceylon ....
subcircular in outline ; width
of ears (flattened) 10-13 mm.
c®. not deciduous. Forearm
77-87*2mm. Indo-Malayan.
d^. deciduous. Forearm 69*5-
75 mm. Austro-Malayan . .
#. Molars unusually narrow ; bony
palate narrow ; fur longer ;
notopatagium and tibiae well
haired. Forearm 72*5-75 mm.
Celebes
d\ Wings from back of 2nd toe ; fur
long and dense.
e^. Frontal region of skull between
postorbital processes concave ;
molars of normal breadth ; lower
leg short : 29-31 mm. Forearm
79-83*5. Ethiopian
/®. Frontal region of skull between
postorbital processes flat ; molars
excessively narrow ; lower leg 39-
40 mm. Forearm 88*5-90. Ethio-
pian
h. Cranial rostrum shortened .* front of orbit
vertically above back of p‘^ ; basicranial
axis nearly parallel to alveolar border.
Ethiopian
c'. Small : forearm 60-67 mm. Equatorial
Africa
seminudus.
R. amplexicaudatus* .
R, brachijotiS’
R. celehensis.
R, angolensis.
R. lanosus.
Myonycteris.
M. collaris.
I. Ptekocyon, Pet.
1861. Pterocyon, Peters, MB. Akad. Berlin, p. 423 Pt. helvus.
1881. Leiponyx, Jentink, Notes Leyden Mus. iii. p. 60 [nec
Liponyx^ Vieillot, 1816, a genus of birds] Pt. helvus.
Basicranial axis considerably deflected : alveolar line pro-
jected backward passing through bases of post-tympanic and
paroccipital processes. Rostrum long : front of orbit vertically
above posterior half or middle of wp. Tympanic produced
externally into a short tubular bony auditory meatus (a
peculiarity unique among bats). Premaxillaries separated
in front. Cutting-edges of lower incisors simple (not bifld).
in cross section twice the size of an upper incisor, mi
equal in length to Wg and combined. Palate ridges
4-f3-f 3. Size large : forearm 114-132 mm.
Range. — Madagascar ; African continent, from Sennaar
and Senegambia in the north to Nyasaland and Naraaqua-
land in the south ; S. Arabia.
On R. minor, see p. 509.
504
Mr. K. Andersen on
1. Pterocyon Dupreanus^ Sclil. & Poll.
1866. Pteropm Pupreanus^ Schlegel & Pollen; P. Z. S. p. 419 (N.W.
Madagascar).
Range. — Madagasear.
Cotypes in the Leiden Museum.
2. Pterocyon helvus, Kerr.
1771. Lesser Ternate Bat, Pennant, Syn. Quadr. p. 362. no. 274 b,
pi. xxxi. fig. 1.
1774. Vespertilio Vampyrus (nec L.), var. C, Schreber, Saugth. i.
p. 154.
1777. Pteropus Vampyrus, var. y, Erxleben, Syst. Regn. An., Mamm.
p. 133.
1781. Lesser Ternate Bat, Pennant, Hist. Quadr. ii. p. 552, pi. lii.
% .
1788. Vespertilio Vampyrus, var. y, Gmelin, Linn. Syst. Nat. ed. 13, i.
p. 45.
1792. Pteropus Vampyrus, var. y, Donndorff, Zool. Beytr. i. p. 62.
1792. Vespertilio Vampyrus helvus, Kerr, Anim. Edngd. i. pt. i. pp. xvii,
91, no. 108.
1810. Pteropus stramineus, E. Geofifroy, Ann. Mus. d’Hist. nat. xv.
p. 95 (Timor, err ore).
1861. Pterocyon paleaceus, Peters, MB. Akad. Berlin, p. 423 (Africa).
1861. Pteropus mollipilosus, H. Allen, Proc. Ac. Nat. Sci. Philad.
p. 159 (Gaboon).
1865. Pteropus palmarum, Heuglin, Leopoldina, Heft v. nos. 3-4, p. 34
(Middle and Upper Nile).
1866. Xantharpyia leucomelas, Fitzinger, SB. Akad. Wien, liv. Abth. i.
H. 10, p. 544 (Sennaar).
1881. Leiponyx Biittikoferi, Jentink, Notes Leyden Mus. iii. p. 59
(Liberia).
Range. — Africa, from Somaliland, Sennaar, and Sene-
gambia in the north, to Nyasaland, Mashonaland, and Nama-
qualand in the south.
Type not in existence.
Kerr’s Vespertilio Vampyrus helvus. — The present species
was well known to the early post-Linnean systematists, who
put it down as a variety of Vespertilio (or Pteropus) vam-
pyrus, L. The earliest recognizable figure and description
appear to be those given by Pennant, in 1771 (/. s. c.\ under
the name “ Lesser Ternate Bat,’^ so called because Pennant
considered it a lesser variety of Seba’s “ Cams volans
Ternatanus orientalis.^'' Kerr’s V. Vampyrus helvus
on Pennant’s description and figure of this bat. The type,
originally in Museum Leverianum, has probably been lost.
Ko habitat given by Pennant nor by Kerr. Senegal may
be fixed as the type locality of Pt. helvus.
Jentink’s Leiponyx Biittikoferi. — Type locality : St. Paul’s
505
Pterocyon, Rousettus, a>id Myonycteris.
River, Millsbury, Liberia; type in the Leyden Museum.
Chief characters, according to Jentink : postcanine teeth f;
second digit without claw. But the rest of the descrip-
tion and all the measurements are sufficient evidence that
L, Biittikoferi was based on a Pt. helms. It is important to
note that the skull was not extracted from the specimen (I
conclude from the fact that it is not recorded in Jentink^s
Catalogue of osteological specimens in the Leyden Museum) ;
the small posterior upper molar [rr^) can therefore easily have
been overlooked ; in aged individuals with much worn teeth
it is not rarely lost. The lack of a claw to the index-finger
must be fortuitous or an individual abnormality.
3. Pterocyon sabceuSj sp. n.
Differs from Pt, helvus in the following particulars : —
Skull smaller ; total length (one male ad., six females ad.)
51*5-55 mm., against 54*5-62*2 in Pt. helvus (nineteen males
ad., twelve females ad.) ; maxillary tooth-row 19*2-
20*8, against 21-23*5 ; cranial rostrum slenderer ; posterior
premolar and molars, above and below, markedly broader
than in the larger-skulled Pt, helvus. The external dimen-
sions average slightly smaller.
Range. — 8. Arabia (Lahej, Aden).
Type. — ? ad., skin and skull. Lahej, Aden ; Aug. 19tli,
189k Collected by Mr. Dodson. 13ritish Museum, no.
99. 11. 6. 3.
II. Rousettus, Gray.
1821. Rousettus, Gray, London Medical Reposi-
tory, XV. p. 299 (Apr. 1, 1821 )
1829. Cercopteropus, Burnett, Quart. Journ. Sci.
Lit. Art, xxvii. p. 269
1843. Eleutherura, Gray, List Mamm. B. M.
p. xix. Nomen nudum.
1843. Xantharpyia, Gray, List Mamm. B. M.
pp. xix, 37
1844. Eleutherura^ Grav, Voyage ‘ Sulphur,’ i.
p. 29 ‘
1852. Cynonycteris, Peters, Reise Mossamb.,
Zool. i., Saugeth. p. 25
1870. Senonycteris, Gray, Cat. Monk. &c. p. 115.
Type.
R. cegyptiacus.
R. cegyptiacus.
R. amplexicaudatus.
R. Leachi.
R. Leachi.
R. seminudus.
Basicranial axis considerably deflected: alveolar line pro-
jected backward passing through upper part of occipital
condyle (minimum of deflection) or through base of zygoma
(maximum). Front of orbit vertically above posterior half
or middle of wb Tympanic not produced into a bony
Ann, <Sc Mag, N. Hist, Ser. 7. Vol, xix. 35
506
Mr. K, Andersen on
auditory meatus. Premaxillaries in contact or co-ossified in
front. Cutting-edges of lower incisors (when unworn) bifid ;
the emargination of the cutting-edge continued as a shallow
vertical groove for a short distance down the front face of
the crown, subequal in size to an upper incisor.
shorter than and combined. Palate ridges 4 (or 3) + 3
(or 4) +1 (or 2). Size moderate: forearm 69*5-99 mm.
Range, — The African continent, exclusive of the Mediter-
ranean countries W. of Egypt ; S. Asia, from Palestine
and Cyprus to S. China ; the Indo- and Austro-M^layan
Archipelago, as far east as the Solomon Islands.
1. Rousettus LeacM^ A. Sm.
]823. Pteropus eollaris (nec 111.), Lichtenstein, Verz. Doubl. Mus.
Berlin, p. 3, no. 47 (Terra Caffrorum).
1827. Pteropus amplexicaudatus (nec Geoff.), Temminck, Mon. Mamm.
i. pp. 260-261 (Cape).
1829. Pteropus Leachi, A. Smith, Zool. Joiirn. iv. p. 433 (Cape).
. 1832. Pteropus hottentottus, Temminck, in Smut’s Enum. Mamm.
Capens. p. 3 (Cape).
1843. Cynopterus Irrevicaudatus (nec Is. Geoff.), Gray, List Mamm.
B. M. p. 39.
Distinguished from all other species of the genus by the
combination of the following characters : — Frontal region of
skull between postorbital processes flattened ; premaxillaries
in contact, but rarely co-ossified ; total length of skull 40*5-
43*8 mm. ; palate ridges normally 4 + 3 + 1. Wings from
back of first toe, or interspace between first and second toe ;
pollex (with claw) 31-35*5 mm. ; second phalanx of third
digit 50*5-60 mm. ; second phalanx of fifth digit nearly
always shorter than first phalanx ; ears not attenuated at
tip ; fur short. Forearm 89-99 mm.
Range, — S. Africa : Cape Colony, Natal, Lower Zambesi
(Inham bane).
Cotype in the British Museum.
Illiger’s Pteropus eollaris. — Type locality : die ostlichen
[afrikanischen] Inseln ” ; no type. Illiger’s Pteropus eollaris
(Abh. Akad. Berlin, 1804-11, pp. 78, 84; published 1815)
is Brisson’s Pteropus collo ruhroy Boussette a col rouge ”
(1762), Schreber’s Vespertilio VampyruSy var. B (1774),
Pennant’s “ Rougette ” (1781), Kerr’s Vespertilio Vampyrus
suhniger (1792; earliest available name of the species),
E. GeofFroy’s Pteropus ruhrieollis (1810). In 1823 Lichten-
stein (I, s, c.) misapplied the name Pt, eollaris to the
S. African fruit-bat here under consideration ; but the error,
hidden as it was in the little-known ‘ Verz. Doubl. Mus.
507
Pterocyon, Rousettus, and Myonycteris.
Berlin/ passed for many years unnoticed, the species being
constantly referred to as Pteropus Leachi or Pt. hottentottus.
In 1852 Peters Reise nach Mossambique ’) confirmed
Lichtenstein’s wrong identification of Pt, collariSj and from
about that year the names Leachi and hottentottus gradually
went out of fashion, being replaced by collaris ; from about
1870 Leachi and hottentottus only appear in the lists of
synonyms of collaris,
2. Rousettus cegyptiacus^ E. Geoff.
1810. Pteropm cegyptiacus, E. Geoifroy, Ann. Mus. d’Hist. nat. xv.
p. 96 (Lower Egypt).
1825. Pteropus Geoffroyi, Temminck, Mon. Mamm. i. p. 197, pi. xv.
figs. 14, 15 (skull) (Egypt, “ Senegal ”).
1870. Eleutherura unic(dor, Gray, Oat. Monk. &c. p. 117 (Gaboon).
Similar to R, Leachiy but with larger skull, broader
rostrum, broader frontal region, and heavier teeth; palate
ridges normally 4 + 4 + 1. Forearm 88-99 mm.
Range, — From Loanda and Gaboon to Egypt, Erythrea,
Syria, Palestine, and Cyprus.
Cotypes in the Paris Museum.
3. Rousettus arahicus^ And. & de Wint.
1902. Rousettus arabicus, Anderson & de Winton, Zool. Egypt., Mamm.
pp. 86, 88, 89-90 (Aden).
Similar to R, Leachi^ but with shorter and lower rostrum,
narrower ear-tips, shorter tibia and foot. Forearm 87-96 mm.
Range, — From Arabia (Aden, Muscat) to Sind (Karachi).
Type in the British Museum.
4. Rousettus Leschenaulti^ Desm.
1820. Pteropus LescTienaultij Desmarest, Encycl. M^th., Mamm. i.
p. 110. no. 142 (Pondicbery).
1832. Pteropus sp., Hodgson, J. A. S. B. i. p. 340 (Nepal).
1835. Pteropus pyrivorus, Hodgson, J. A. S. B. iv. p. 700 (Nopal).
1843. Cynopterus affinis, Gray, List Mamm. B. M. p. 39 (Himalaya).
1870. Eleutherura marginata, Gray, Cat. Monk. &c. p. 118 (‘‘Nepal,”
i. e. Nasirabad).
1870. Eleutherura fuliginosa^ Gray, Cat. Monk. &c. p. 118 (Lao Mts.,
Siam).
1870. Eleutherura fusca, Gray, Cat. Monk. &c. p. 119 (“India ? ”).
1873. Cynonycteris mfuscata, Peters, MB. Akad. Berlin, p. 487 (Cal-
cutta).
Allied to R, arahicus^ but smaller, with the muzzle shorter
and slenderer, the tip of the ears not attenuated, the pollex
markedly shorter, wings shorter, especially the first and second
35*
508 Mr. K. Andersen on
phalanx of the third digit, and the foot smaller. Forearm
80-5-87-5 mm.
Bange. — Himalayas (Nepal), extending southward over
the Indian Peninsula (Pondichery), eastward through Bengal,
Burma, Siam (Lao Mts.) to S. China (Amoy).
Type in the Paris Museum.
Peters’s Cynonycteris infuscata. — Type locality : ange-
blich aus Calcutta ” (a dealer’s specimen) ; type in the Berlin
Museum (no. 361). “ Sehr ahnlich der (7. Leschenaultn, in
alien Verhaltnissen kleiner, dunkelbraun von Farbe, mit
schwarzen Krallenund dem ersten falschen Backzahn grosser”;
detailed measurements given ; forearm 68, third metacarpal 42,
tibia 29 mm. The type, I am informed by Prof. Matschie,
is a young, not full-grown individual (“ die Epiphysen an
den Fingergelenken sind noch nicht mit den Phalangen
verwachsen,” Matschie, in Utt.) ; hence its small size.
5. Bousettus seminuduSj Gray.
1870. Xantharpyia seminuda, Gray, Cat. Monk. &c. p. 115 (Ceylon).
Similar to B, Leschenaulti^ but deciduous, nape and
shoulders seminaked, general colour of fur lighter. Forearm
79-85*5 mm.
Bange. — Ceylon.
Type in the British Museum.
Gray’s Xantharpyia seminuda. — Type locality : Ceylon.
‘‘ Pteropus seminudus^ Kelaart,” is a nomen nudum ; in the
paper usually referred to by authors, viz. Blyth’s account in
J. A. S. B. xxi. p. 345 (1852), on a collection of mammals
sent by Kelaart to the Asiatic Society of Bengal, it appears
only as a synonym, without comment, of Ph. Leschenaulti ;
the same is the case in Kelaart’s ^ Prodromus Faunae Zey-
lanicae ’ (1852). The name remained a nomen nudum, until
in 1870 (/. s. c.) Gray published a brief diagnosis of
“ Xantharpyia seminuda ” ; the British Museum specimen
on which Gray based this diagnosis is, therefore, the type of
the species.
6. Bousettus ample xicaudatuSy E. Geoff.
1810. Pteropus amplexicaudatus, E. Geoffrey, Aim. Mus. d’Hist. nat.
XV. p. 96, pi. iv. (whole fig.) (Timor).
1870. Eleuiherura infvmata, Gray, Cat. Monk, &c. p. 118 (Flores).
1870. Eleuiherura philijypinensis, Gray, Cat. Monk. &c. p. 119 (Manila).
1898. Cynonycteris Bocagei, Seabra, J. Sci. Math. Lisboa, (2) v.
pp. 160-161, 169, pi. i. fig. 11 (palate ridges) (Timor).
Similar to R. Leschenauliij but subcircular in outline,
ears narrower. Forearm 77-87*5 mm.
Pterocyon, Roiisettus, and Myonycteris. 509
Range. — Cambodja, Philippines, Borneo, Sumatra, Engano,
Flores, Savu, Alor, Timor.
Type in the Paris Museum.
7. Rousettus minor ^ Dobs.
1873. Cynonycteris minor, Dobson, J. A. S. B. xlii. pt. ii. p. 203, pi. xiv.
fig. 9 (ear) (Java).
I have not, as yet, seen the type of Cynonycteris minor.
In none of the characters given by Dobson is there anything
to prove that R. minor is different from the true R. amplexi-
caudatus (Dobson, it must be remembered, lumped the
continental R. Leschenaulti and the Indo-Malayan, insular
R. ample xicaudatus into one species, Cynonycteris amplexi-
caudata,^' and when describing R. minor probably com-
pared it with R. Leschenaulti, not with the true R, amplexu
caudatus), — save perhaps in one: the length of forearm is
stated to be only 71 mm., whereas I have never seen a fully
adult R. amplexicaudatus with the forearm less than 77 mm.
I prefer to leave open the question as to the validity of
R. minor, until I have had an opportunity of examining the
type.
Range. — Java.
Type in the Calcutta Museum.
8. Rousettus hrachyotis. Dobs.
1877. Cynonycteris hrachyotis, Dobson, P. Z. S. p. 116 (Duke of York
Isl.).
Similar to R. amplexicaudatus, but smaller, with shorter
and narrower ears j deciduous; tooth-rows shorter. Fore-
arm 70-75 mm.
Range. — Amboina, New Guinea, New Ireland, Solomon
Isl.‘
Type in the British Museum.
9. Rousettus celehensis, sp. n.
Diagnosis. — Bony palate and all molariform teeth, above
and below, unusually narrow. Fur longer and richer than in
any of the foregoing species ; notopatagium partly (or wholly)
haired; general size small. Forearm 72*5-75 mm.
Skull. — General size as in R. amplexicaudatus-, rostrum
very low and slender ; bony palate unusually narrow ; width
externally across rn^-m^ 9*7 mm. (two adults), against 10*2-
11*8 in amplexicaudatus (ten adults).
Teeth. — Upper canine and p^ (middle premolar) widely
510
Mr. K. Andersen on
separated ; in the centre of the interspace between these
two teeth, not deciduous ; last premolar and molars above
and below very narrow : nii at least twice as long as broad ;
small, less than half the size of rn} ; p2 in cross section
three or four times the size of a lower incisor ; m2 about half
the length of subcircular in outline.
Palate ridges. — 4 + 3 + 1.
External structure. — Ears essentially as in E. amplexl-
caudaius : narrow, not attenuated below the tip, the tip itself
broadly rounded off ; antitragal lobe small, rounded. General
size of the animal as in R. hrachyotis (thus smaller than
R, amplexicaudatus)^ but digits proportionally considerably
longer than in any other eastern species of the genus
(index of pollex 392, of third digit 1646, against 335-41
(pollex) and 1529-41 (third digit) in all other eastern
species). Tail long, probably about 20 mm. (only dried
skins examined).
Fur. — Longer, richer, and more velvet than in R. amplexi-
caudatus and allied eastern species ; notopatagium clothed
with dense fur ; hairing on forearms, tibiae, interfemoral, and
underside of plagiopatagiura longer and richer ; face more
densely haired.
Colour, — Brighter than in R, amplexicaudatus. Back
light ProuBs browm, rump more inclining to mars-brown
tinged with russet ; sides of back and tibiae next to membranes
almost vandyke-brown ; crown and occiput dark brown,
approaching bister; nape of neck broccoli-brown; a tuft of
glandular mummy-brown hairs on either side of the neck in
both sexes ; entire underside of body dark greyish drab. —
Immature individuals are similar in colour to adults, but
without the mummy-brown neck-tuft.
Range. — Celebes.
Type. — ? ad., skin and skull. Mt. Masarang, Celebes,
3500' ; Oct. 1895. Collected by Dr. Chas. Hose. Brit. Mus.
no. 97. 1. 2. 8. Three specimens examined.
Remarks. — On hasty inspection this species, owing to its
small size, can easily be (and has in fact repeatedly been)
confused with R. hrachyotis. The larger skull, very narrow
palate, narrow molars, not deciduous p^, much longer pollex
(28-30 mm., against 24-26 in hrachyotis)^ longer wings
(chiefly owing to the longer metacarpals), much longer fur,
haired notopatagium, and much more densely haired tibiae
readily distinguish it from R. hrachyotis,
10. Rousettus angolensisy Bocage.
1898. Cynonycteris angolensis, Bocage, Jorii. Sci. Math. Lisboa, (2) v.
pp. 133, 138, fig. (palate ridges) (Pungo Andonga, Cahata, Quibula).
Pterocyon, Rousettus, and Myonycteris. 511
Frontal region of skull between postorbital processes dis-
tinctly concave ; premaxillaries co-ossified in front ; molars
as broad as (or broader than) in R, Leachi\ p2 in cross
section only equalling or slightly exceeding a lower incisor.
Wings from back of second toe ; second phalanx of fifth digit
nearly always longer than first phalanx ; antitragal lobe well
developed ; lower leg very short (29-31 mm.) ; fur long ;
notopatagium haired. Size smaller than in B, Leachi ; fore-
arm 77-83*5 mm.
Range. — Angola, north-westward to Cameroon and Togo,
eastward through the Congo Basin to Ruwenzori and German
East Africa.
Cotypes in the Lisbon and British Museums.
11. Rousettus lanosuSj Thos.
1906. Rousettus lanosus, Thomas, Ann. & Mag. N. H. (7) xviii. p. 137
(Ruwenzori East).
Molars excessively narrow; p2 in cross section twice or
three times the bulk of a lower incisor. Wings from back of
second toe; second phalanx of fifth digit longer than first
phalanx ; antitragal lobe indistinct ; lower leg not shorter
than usual (39-40 mm.) ; fur long and coarse ; notopatagium
haired. Larger than R, angolensis : forearm 88*5-90 mm.
Range, Shoa ; Ruwenzori East, 5000-13, 000k
Type in the British Museum.
III. Myonycteris, Matschie.
1899. Myonycteris, Matschie, Megachiroptcra, pp. 61, 63.
Type.
M. coUaris.
Basicranial axis only slightly deflected : alveolar line pro-
jected backward passing through middle of external auditory
meatus. Tympanic not produced into a bony auditory
meatus. Rostrum shortened, owing to enlargement of orbital
cavity : front of orbit vertically above back of p\ Pre-
maxillaries in contact in front. Cutting-edges of lower incisors
(when unworn) bifid, p^ in cross section subequal to an upper
incisor, rn? and rudimentary, m2 much reduced in size
(from one fourth to somewhat less than half the bulk of m-^,.
Palate ridges : 4-f3 + ?. Size small : forearm 60-67 mm.
Range. Ethiopian.
Matschie^s Myonycteris. — Myonycteris in its original sense
(a subgenus of Xantharpyia ; Matschie, I, s, c.) included two
species, M, torquata (i. e. collaris ; the type) and M. angoU
ensis. But angolensis is a true Rousettus, whereas collaris,
as being in skull and teeth more closely related to Cynopterus
512
Mr. K. Andersen on
than to Eousettus, but clearlj different from both, must be
kept in a separate genus. Matschie^s definition of Mi/o~
nycteris was, however, based not on the species selected by
him as type of the subgenus, viz. collaris^ but on angolensis ;
the diagnosis of the genus as given above is therefore entirely
different from that published by Matschie.
1. My 0 nycteris collaris^ Gray.
1870. Cynopterus collaris, Gray, Cat. Monk. <&c. p. 123 W. Africa ”).
1878. Cynonycteris torquata, Dobson, Cat. Cbir. B. M. p. 76, pi. v.
fig. 1 (Angola).
1889. Cynonycteris hrachycephala, Bocage, Jorn. Sci. Math. Lisboa,
(2) i. p. 197 (San Thome).
Forearm 60-67 mm. Wings from back of first phalanx
of second toe.
Range. — From tlie Congo Basin southward to Angola,
north-westward to San Thome, Liberia, and Sierra Leone.
Type in the British Museum.
Gray’s Cynopterus collaris. — Type locality ; W. Africa^^;
the British Museum register for 1843 proves the specimen to
have been obtained near Congo.^’ Gray’s statement (/. s. c.)
that the specimen is young is incorrect; his quotation of
“ Gray, List Mamm. B. M. (1843),^’ where the specimen is
stated to have been registered under the name Xantharpyia
collaris^ does not refer to the printed text of that book, but to
a hand-written addition by Gray in the British Museum copy
of the book. Prior to 1870 collaris ■” had not been used as
a specific name in the genus Cynopterus ; it is therefore valid,
and antedates Cynonycteris torquata^ Dobson.
Bocage’s Cynonycteris hrachycephala. — Type locality :
S. Thorn^, Gulf of Guinea; type in the Lisbon Museum.
From the description la premibre premolaire et la derni^re
molaire extrernement petites aux deux macho ires ” ; forearm
62 mm.) and the figure of the skull and teeth in palate view
clearly a M. collaris.
General Remarks,
The Genera. — Rousettus is allied to Fterocyon ; the two
genera probably represent diverging branches from one
common stem. They accord in most of their important
cranial, dental, and external characters; in both the basi-
cranial axis is deflected to practically the same degree. In
having the premaxillaries in contact or co-ossified (not sepa-
rated), the tympanic not produced into a bony auditory
meatus, not lengthened, and less reduced in size.
Pterocyon, Rousettus, and Myonycteris,
513
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Mr. K. Andersen on
Rouseltus is more primitive than Pterocyon ; but it is on a
higher level in the rather shorter rostrum, and the more
reduced p^. The range of the genus Rousettus over the whole
of the Ethiopian and Oriental regions, the close affinity
of R, arahicus to the S. African R. Leachij the absence
of any representative of the genus from the whole of the
Mediterranean subregion except Egypt, are evidence that
its origin dates back to a time when, owing to different
physiographic conditions, Africa and S. Asia were much
more intimately connected than now. Pterocyon is a more
specialized Ethiopian offshoot of the common prototype.
In its essential cranial and dental characters Myonycteris
is intermediate between Rousettus and Cynopterus^ though
nearer to Cynopterus. In Rousettus the basicranial axis is
very distinctly deflected ; in Myonycteris^ as in Cynopterus^
it is nearly parallel to the alveolar border. In Rousettus
the cranial rostrum has remained comparatively long, the
anterior edge of the orbital cavity being vertically above
the posterior half or middle of ni} ; in Myonycteris the
rostrum is considerably shortened, chiefly owing to the
fact that the anterior edge of the orbital cavity is pressed
forward to a point vertically above the back of p^ ; in
Cynopterus the rostrum is still shorter and stouter, the
anteorbital rim pressed still a little farther forward, to a
point above the middle of From a glance at the dental
formula it would seem that Myonycteris (molars §) is closely
in accordance with Rousettus (f), and essentially different
from Cynopterus (J), but in reality Myonycteris is also in its
teeth nearer to Cynopterus ] in Rousettus m? is reduced in
size, in Myonycteris quite rudimentary, in Cynopterus lost ;
in Rousettus m<^ is normal, small ; in Myonycteris m2 is
much reduced in size, rudimentary ; in Cynopterus m2
much reduced in size, m3 lost. In short, the cranial and
dental peculiarities (non-deflection of brain-case, shortening
of rostrum, reduction of posterior molars) which distinguish
Myonycteris from Rousettus have been preserved, or carried
still farther, in Cynopterus ; if the skull of Myonycteris were
known from a fossil state only, this bat would undoubtedly
have been declared a “ connecting-link ” between Rousettus
and Cynopterus. Also externally, in the form of the tip of the
muzzle (vertical furrow between nostrils deep and narrow,
inner margins of nostrils abruptly projecting), Myonycteris
closely approximates Cynopterus,
The Species. — The three species of Pterocyon are closely
interrelated. Pt. Dupreanus^ from Madagascar, with its
relatively longer rostrum and less modifled fur-structure, is
Pterocyon, Kousettus, and Myonycteris. 515
apparently the least modified species. Pt, sabceusj from
S.W. Arabia, is a small-skulled and broad-toothed repre-
sentative of the African Pt. helvus.
The eleven known species of Rousettus are referable to five
types : — (1) R, Leachi, cegyptiacuSy and arahicus : rather
heavily built species, with strong rostrum and teeth, the
second phalanx of third digit lengthened, the pollex com-
paratively long ; distributed over Africa generally, Cyprus,
Palestine, Syria, and Arabia, as far east as Sind (Karachi) ;
R, cegyptiacus is a larger-skulled modification of the R. Leachi
type ; R. arahicus is more closely related to the S. African
R. Leachi than to R. cegyptiacus. — (2) R. Leschenaulti^ semi-
nuduSj amplexicaudatus, minor and hrachyotis : very closely
related to the species of the former group, but rather more
delicately built, with slenderer rostrum, feebler teeth, the
second phalanx of the third digit not lengthened, the pollex
comparatively shorter ; the members of this group are, pro-
bably, on the whole slightly less specialized than those of
the former ; R. Leschenaulti (continental S. Asia) and semi-
nudus (Ceylon) come near to the S. African R. Leachi in the
width of the interspace between c and the size and shape
of w?3, the width of the ears, and the length of the tail ; in
the Indo-Malayan R. amplexicaudatus there is a tendency to
a reduction of the diastema c-p^, m^, is smaller and more
circular in outline, the ears narrower, the tail averaging
longer, the general dimensions smaller ; most of these
characters find a climax in the Austro-Malayan R. hrachyotis :
diastema c-p^ still more reduced, p^ deciduous, ears still
smaller, size smaller. — (3) R. celehensis : peculiarly narrow
palate, narrow molars, longer and richer fur, small size, pro-
portionally long wings ; probably a modification of the
R. amplexicaudaius-hr achy Otis type. — (4) R, angolensis (Togo,
Cameroons, and Angola, to Ruwenzori), a peculiar species:
skull and teeth differing in some details, fur long and
dense, coloration richer than usual. — (5) R. lanosus (Shoa,
Puwenzori), the most aberrant species of the genus : molars
excessively narrow, fur very long and dense.
From a more general point of view the first three of these
groups (the members of which are certainly more closely
related to each other than to those of groups 4 and 5) may
be united into one section, giving a long chain of intimately
connected forms from W. Africa to the Solomon Islands ;
R. angolensis and still more R. lanosus are aberrant repre-
sentatives of this widely distributed type of bat.
516
Mr. R. I. Pocock on the
LX. — Notes on the Quagga and BurcheWs Zehra in the
Paris Museum, By R. I. PocoCK, Superintendent of the
Zoological Society’s Gardens, London.
In the ^ Bulletin du Museum d’Histoire naturelle,’ pp. 449-
452 (1906), Dr. Trouessart has given an account, illustrated
by two admirable photogravures, of a quagga and a Burchell’s
zebra preserved in the Paris Museum. The quagga especially
proves to be a specimen of considerable systematic import-
ance; and since Dr. Trouessart omits to mention one or
two points of interest connected with it, and makes some
statements which are contrary to fact, no apology is needed
for supplementing his communication with the following
comments.
In a paper* on the Cape Colony quaggas, with which
Dr. Trouessart does not appear to be acquainted, I pointed
out that the two forms named respectively by Mr. Lydekker f
E. quagga Greyi and E, q. Lorenzi resemble each other and
differ from E. q. quagga and E, q. Danielli in having the stripes
brown and the interspaces creamy yellow ; and, further, that
they may be distinguished from each other by certain cha-
racters, of which the width of the stripes on the neck is one.
In Lorenzi the neck-stripes are exceedingly wide, the inter-
spaces forming distinct but very narrow pale lines, whereas
in Greyi the interspaces are relatively broad and the stripes
correspondingly narrow.
So far as the width of the neck-stripes is concerned the
Paris specimen is more like the type of Lorenzi than is any
other recorded specimen. But the stripes are even wider J
and the intervening areas narrower than in the Vienna
* Ann. & Mag. Nat. Hist. (7) xiv. pp. 313-328 (1904).
t ‘ Knowledge,’ xxv. p. 221 (1902). Dr. Trouessart probably had no
opportunity of consulting this paper.
% In his description of the Paris specimen Dr. Trouessart writes : —
‘‘ Les bandes fonc6s du cou sont doubles par le bas, mais confluentes a
leur partie superieure, de telle sorte que la bande intercalaire blanche est
tres Itroite.” The photograph does not bear out this statement, for on
the left side of the body the stripes, so far as can be seen, are in two
cases confluent below and divided above and in two cases divided below
and confluent above, the rest being entire. Such confluence is not
unusual in the quaggine races of Equus, with which I include the
Burch elline zebras. In any case, the alleged confluence does not affect
the total number of neck-stripes, which is approximately the same in
the Paris and Vienna specimens, as well as in the type of Greyi and
various kinds of zebras of the Burchelli type. It is, moreover, the in-
crease in the width of the stripes, not their confluence, that causes the
narrowness of the intervening pale area.
517
Quagga and BurchelVs Zebra.
example. In the latter, as in the Paris example, the pale
intervening areas are distinct upon the withers ; but behind
the withers in the Paris example they die out, the flanks
being only indistinctly striped and the hind-quarters practi-
cally unstriped. In Lorenzi^ on the other hand, the inter-
vening spaces persist in such a manner as to leave no doubt
that both in pattern and posterior extension the stripes were,
to all intents and purposes, like those o£ typical Burchell’s
zebra {E. quagga Burchelli)^ the so-called “ saddle selle ”
of Trouessart), characteristic of that animal and the more
northern forms related to it,' being quite evident Herein
lies the chief difference between the.. Paris and -Vienna
specimens. To the type of Greyi^ on the contrary, the Paris
specimen shows a close resemblance in the obliteration of the
stripes on the body and hind-quarters.
Further evidence of likeness between the three specimens
above discussed, and especially between the Vienna and Paris
animals, is supplied by Dr. Trouessart’s statement that the
latter has the appearance of a chestnut horse banded with
white, the stripes being brown and the intervening areas
whitish. In the typical quagga and E, q. Danielli^ on the
other hand, the stripes were black and the intervening areas
chestnut.
It will be evident from what has been said that the Paris
specimen is to a great extent intermediate in its characters
between the types of Lorenzi and Greyi. This fact may be
used as an argument in favour of the view held in 1904 by
Mr. Lydekkert, that all the genuine quaggas belonged to a
single species very variable in the degree of development of
its stripes, but not resolvable into geographical races or sub-
species ; and also in support of tlie opinion, maintained by
myself, that there were several local forms of this animal, the
assumption of the probable existence of intermediates justi-
fying the view that only a subspecitic value should be
assigned to the differences between them. Whatever con-
clusion be formed with regard to this matter, the chief interest
of Dr. Trouessart’s paper upon the Paris specimen lies in the
fact that it has proved the former existence of a quagga
* As I have already pointed out, the pattern of the stripes on the body
and hind-quarters of the type of Lorenzi affords convincing evidence of
the nearness of the affinity between this quagga and typical Burchelli.
Dr. Lorenz also was forcibly struck by the similarity between the two
animals in this respect. The resemblance between them makes it im-
possible to draw up a logical definition of “ quaggas ” as distinct from
“ Burchell’s zebras.’
t P. Z. S. 1904, i. pp. 426-431.
518
Mr. E. I. Pocock on the
intermediate in coloration between two specimens that have
been made the types of distinct subspecies, namely Greyi and
LorenzL
In his account of the Burcheirs zebra in the Paris Museum,
Dr. Trouessart incidentally attempts to prove that the forms
named BurchelU and Chapmanni are specifically distinct from
each other. Under Chapmanni he includes the northern
form described as Bohmi (^=Granti) and presumably also
Selousiy Wahlhergi^ and antiquorum. It is of no great
moment whether these forms be regarded as species or sub-
species ; but since a practically complete gradation in the
disappearance of the stripes from the fetlocks upwards to the
root of the tail can be traced from Bohmi and Selousi through
Chapmannij Wahlhergi^ and antiquorum to the various types
of BurchelU (sensu stricto)^ it is illogical to draw a line
between BurchelU and Wahlhergiy classifying the latter with
Bohmi and Selousi and letting the former stand alone.
The distinctions upon which Dr. Trouessart lays stress are
the alleged absence of stripes upon the legs in BurchelU and
the presence of only narrow, faint, and incomplete stripes
upon the hind-quarters, beneath the last complete stripe that
runs from the root of the tail to the groin (“aine”). Contrasted
with, this are the strong complete stripes on the hind-quarters
in Chapmanni and their extension at least as far as the hocks.
It is quite true that typical Chapmanni may be distinguished
from iy^ic2i\ BurchelU by these and other characters ; but the
variation in the development and downward extension of the
stripes over the quarters in individual specimens of BurchelU is
very great. I have before me the photograph of a specimen
that formerly lived in the London Zoological Gardens. In
this there are only about two very faint and narrow stripes
below the one that passes to the root of the tail. The example
in the Bristol Museum* is also very imperfectly striped below
that line. This is also the case in an example that was living
in the Amsterdam Zoological Gardens a year or two ago.
Nevertheless the statement that there are no stripes on
the legs in Burchelks zebra is not true. Stripes are quite
commonly retained both on the knees and hocks ; and by
publishing the photograph of the BurchelU preserved in the
Paris Museum, Dr. Trouessart has supplied additional and
conclusive evidence of the occasional extension of transverse
stripes — narrow and more or less broken up certainly — all
over the hind-quarters down to the level of the junction of the
* For fig. and description, see Pocock, P. Z. S. 1903, ii. p. 196, and
Ann. & Mag. Nat. Hist. (6) xx. p. 41 (1897).
519
Quagga and BurcJielVs Zebra.
femur and the tibia. In the striping of this region, indeed,
this specimen forcibly recalls the example of Wahlbergi in
the Tring Museum, in which the stripes on the lower portion
of the quarters are broken up into an irregular reticulated
pattern To maintain that the Tring specimen of WahU
hergi and the Paris specimen of BurchelU represent distinct
species, and to hold at the same time that the former belongs
to the same species as the types of Selousi and Grantij
obscures the plainest facts of affinity as testified by likeness ;
and the adoption of Dr. TrouessarPs view regarding the
zebras in question, and the nomenclature it involves, renders
abortive one of the primary purposes of systematic naming —
that is to say, the expression of relationships, of which, in
this case at least, resemblances are the sole criteria.
Touching the affinity between so-called zebras ” and
quaggas,” Dr. Trouessart remarks that Burchell’s zebra
approaches the quagga in pattern more than it approaches
its allies, the other zebras. The truth of this statement can
scarcely be admitted, for the likeness between BurchelU and
Wahlbergi is, on the whole, greater than the likeness between
BurchelU and the most zebra ’’-like of all the quaggas,”
namely Lorenzi. Nevertheless it is gratifying to welcome an
adherent of the view that the differences between Burchell’s
zebra"” and quaggas ” are practically equivalent to the
differences between BurchelPs zebra and other zebras of the
same type, such as Chapman’s. Dr. Trouessart, however,
does not admit specific identity between quaggas and
Burchell’s zebras, for the alleged reason that the ground-
colour of the latter is white or clear grey without mixture of
red or yellow. It is difficult to find justification for this
argument, since Dr. Trouessart himself describes the ground-
colour of the quagga in the Paris Museum as white. More-
over, the tint of the ground-colour is not a specific character
in this group of Equidae ; and it is not true that it is always
white in BurchelU^ as the literature on the subject conclusively
proves. For instance, although Gray described the type
specimen as white betAveen the stripes, I pointed out ten
years ago that in the specimen in the Bristol Museum the
ground-colour on the body and hind-quarters is dark ruddy
greyish brown ” ; and in a stuffed specimen in the British
Museum the interspaces are heavily washed with yellowish
brown f. The same variability in tint is shown in Chapman’s
* See Ann. & Mag. Nat. Hist. (6) xx. p. 45 (1897).
t In this specimen the stripes on the hind-quarters extend right down
to the level of the femoro-tibial joint and are more complete and distinct
than in the Paris example.
520 Messrs. O. Thomas and R. C. roughton on
zebras. In the case of two specimens living last year in the
Zoological Gardens in London, the interspaces of one were
white, of the other ochre-yellow. Lastly, as has already
been stated in this and other papers, the ground-colour in
quaggas proper is either chestnut or creamy white.
Hence the reasons advanced by Dr. Trouessart for separating
Burchell’s zebra specifically from quaggas have no foundation
in fact.
LXI. — 'New Mammals from Lake Chad and the Congo^
mostly from the Collections made daring the Alexander-
Gosling Expedition. By Oldfield Thomas, F.R.S., and
R. C. Wroughton.
The following descriptions of two new forms of dassie were,
by oversight, omitted from our paper in last month’s issue of
this Magazine (p. 370).
Procavia Lopesi, sp. n.
A large dassie of the hypsodont group with a buff dorsal
spot.
Size about as in Mackinderi ; fur short (20 mm.) and
harsh as compared with that of Mackinderi (40 mm.) or even
of Jachsoni (30 mm.) ; general colour above near “ raw
umber” of Ridgway, resulting from a mixture of black and
buff ; under surface dark buffy, the hairs greyish at base.
Under-fur of back slate-grey at base, dirty white terminally ;
hairs of outer fur either wholly black, or black with pale buff
tips.
Face grizzled black and white or buffy, the dark patch on
the vertex really black, otherwise the usual colour-pattern of
the genus ; the dorsal spot comparatively broad, its hairs
bright buff from base to tip.
Skull large as in Mackinderi^ much larger than in Jacksoni^
width of frontals much less than in the former, nasals narrow
as in Jacksonij quite different fiom the broad nasals of
Mackinderi.
Dimensions of the type (those of the body taken in the
flesh) : —
Head and body 560 mm.; hind foot 70 ; ear 35.
Skull (Stage VIII.) ; greatest length 100 ; basilar length
90; greatest breadth 57; greatest breadth of frontals 39;
anterior breadth of frontals 22 ; anterior breadth of nasals
new Mammals f rom Lake Chad and the Congo, 521
9*5 ; length of nasal suture 26 ; length of upper molar series
43 ; greatest breadth of m' 8.
Hab. Kodja Ili.ll, Gaima Range ; River Kibali, Monbuttu.
Type, Adult female. Original no. 105. Collected by
Mr. Boyd Alexander, 18th July, 1906. (Two specimens
examined.)
The present species is easily distinguishable from either of
its neighbours Mackinderi and Jacksoni by its very short
fur and black vertex. In skull-characters it resembles
Jacksoni in its long narrow nasals, but differs in its greater
size and stouter teeth ; from Mackinderi, with which it
agrees closely in size, it is at once distinguishable by its
shorter fur, narrow nasals, and slightly larger teeth.
We have much pleasure in dedicating this species to Jose
Lopes, who followed Mr. Alexander so pluckily throughout
his long and arduous journey.
Procavia sharica^ sp. n.
A hypsodont dassie of medium size with a linear orange
dorsal spot.
Size approximately as in the Nigerian Goslingi ; fur short
(15 mm.) and harsh as compared with that of Goslingi ;
general colour above near hair-brown,” resulting from a
mixture of black and ^‘wood-brown”; hairs of undersurface
buffy to their bases; under-fur of back wood-brown with dark
brown bases ; the hairs of outer fur black with pale buffy tips.
Face grizzled black and white; crown dark (near “seal-
brown”), the dark area extending on to the nape; con-
spicuous patches of “ pinkish buff” behind the ears. Dorsal
spot long (50 mm.), narrow (6-7 mm.), its hairs “ ochraceous
buff” from base to tip.
Skull narrower for its size than in Goslingi, larger in all
ways than that of the Soudanese rujiceps at the same stage ;
nasals about the same length as in Goslingi and rujiceps, but
narrowing sharply anteriorly, very markedly more so than in
either of the other species.
Dimensions of type (those of body taken in the flesh) : —
Head and body 456 mm. ; hind foot 70 ; ear 28.
Skull (Stage VI.) : greatest length 85 ; basilar length 73 ;
greatest breadth 47 ; greatest breadth of frontals 35 ;
antep>’’ breadth of frontals 20 ; anterior breadth of nasals 8 ;
length of nasal suture 20 ; length of upper molar series
to 32 ; greatest breadth of 6*5.
Hah, Kajibu, Shari River.
Type, Young male. Original no. 45. Collected by
Ann, do Mag. N, Hist, Ser. 7. Vol, xix. 36
522 Mr. A. S. Hirst on Spiders
Capt. G. B. Gosling on 14th July, 1905. (Four specimens
examined.)
The specimens collected by Capt. Gosling are very uniform
in their characters. The two nearest neighbours of sharica are
Goslingi and rujiceps. The duller colouring and harsh fur
of sharica serve to distinguish it easily from Goslingi, while
its colour separates it at once from the pale, white-bellied
rujiceps. In skull-characters the sudden narrowing anteriorly
of the nasals of sharica differentiates it from both of the
others.
LXIl. — On Two Spiders of the Genus Selenocosmia.
By A. S. Hirst.
Selenocosmia Stalkeri^ sp. n.
$. — Colour Cephalothorax and legs a light brown;
sternum and lower surface of coxae of legs, together with the
dorsal surface of the patellae of the posterior legs, darker in
colour.
Ocular tubercle a little more than twice as long as broad ;
the lateral part low, the central part higher.
Eyes, Front row of eyes procurved ; anterior median eyes
a little larger than the anterior laterals and separated from
them by a diameter (of a median eye), the space between
the median eyes being a little more than a diameter. Poste-
rior median eyes small and placed a little in front of the
posterior laterals, from which they are separated by a short
interval.
Cephalothorax. Length of cephalothorax much greater
than the breadth and exceeding the length of the tibia and
patella of the first and fourth leg. Fovea less strongly pro-
curved than is the case in S. Stirlingi and of rather small
extent.
Sternum elongate in shape ; posterior sigilla situated in
the anterior two thirds of it and distant a little less than a
third of the width from the lateral margins.
Legs. First pair of legs measuring a little less than three
times the length of the cephalothorax. Tibia and patella of
the first and fourth pairs equal in length. Tibia of fourth
pair much shorter than the metatarsus.
Stridulating-hacilla of maxillipalp forming a narrow and
elongate patch, which is convex below and runs along the
523
of the Genus Selenocosmia.
greater part of the length of the anterior surface of the coxa,
the bacilla being arranged in comparatively few rows (fig. 1).
Measurements in mm. Length of cephalothorax 20 ‘5, of
sternum 9, of mandibles 11, of first leg 58, of second leg 51,
of third leg 47*5, of fourth leg 60*5, of stridulating-area of
maxilla 5*5 ; breadth of cephalothorax (at the middle) 16*5 ;
breadth of cephalothorax (anterior edge) 11*75 ; breadth of
sternum 7, of mandible 5 ; greatest breadth of stridulating-
area 1*5.
Fig. 1.
Anterior surface of the coxa of the maxilla of Selenocosmia Stalkeri.
Hah. A single female specimen was collected by Mr. W.
Stalker at Alexandria, Northern Territory of South Australia,
during the month of December 1905. The specimen was
presented to the British Museum by Sir William Ingram
and the lion. John Forest.
Collector’s note : — “ Hole driven in very hard ground ;
about 1 foot in depth and slightly chambered.”
Remarks, This new form differs from S. Stirlingi in that
the posterior sigilla are further removed from the posterior
margin of the sternum and that the fovea of the cephalo-
thorax is less strongly procurved. It also differs from that
species in the shortness of the hair which clothes the legs and
in the relative size and position of the eyes. The structure of
the stridulating-area also affords a character of some import-
ance. S. Stalkeri differs from S. vulpina chiefly in the
characters presented by the eyes and in the bacilla of the
stridulating-area being stouter, and from S. crassipes,
8. strenuus, and S. validus apparently in the relative length
of the legs.
Selenocosmia himalayana^ Pocock.
Colour. Cephalothorax greyish brown; body and legs
black ; dorsal surface of coxae and trochanters of the legs,
together with their patellae, white or greyish white and tinged
with yellow ; abdomen dorsally greyish brown or dark in
colour.
3G-^
524
On Sjnders of the Genus Selenocosmia.
c? . — Cephalothorax of less length than the patella and
tibia of the fourth leg.
Legs. Patella and tibia of first leg a little longer than the
corresponding segments of the fourth, the difference being
due to the slightly greater length of the patella of the first
leg. Metatarsus of fourth leg greatly exceeding the tibia in
length.
Palpal organ (fig. 2) provided with an obtuse projection
or lobe, which is situated on the outer side at the junction
of the elongate portion of the organ with the bulb.
Fig. 2.
External view of palpal organ of Selenocosmia himalaycma.
Measurements in mm. Length of carapace 20, of tibia and
patella of first leg 24, of tibia and patella of fourth leg 22,
of metatarsus of fourth leg 18 ; greatest breadth of cephalo-
thorax 16.
$ . — Cephalothorax equal in length to the patella and
tibia of the fourth leg.
Legs. Patella and tibia of the first leg a little shorter than
the patella and tibia of the fourth. Metatarsus of fourth leg
exceeding the tibia in length.
Measurements in mm. Length of cephalothorax 18, of tibia
and patella of first leg 17, of tibia and patella of fourth
leg 18 ; total length (mandibles inch) 39; greatest breadth
of cephalothorax 25.
Hab. A single adult male from Kasauli, Simla, 6600 feet
(July 1905), and a single adult female from Dalhousie,
N.W. Himalayas, 6000 feet. The specimens were collected
by Col. Barrow.
Remarks. The acquisition of fully grown specimens of
both sexes by the British Museum has enabled me to supple-
ment the description * of this handsome species, which was
hitherto known from a single small-sized specimen.
^ Pocock, Journ. Bomb. N. H. Soc. xii. p. 746 (1899), and Faun. Brit.
Ind., Arachn. p. 200 (1900).
On Brachiopod Nomenclature,
o25
LX III. — Brachiopod Nomenclature : the Genotype of
Terebratula. By S. S. Buckman, F.G.S.
[Plate XII.]
To which author must the generic term Terehratala be
ascribed, what species is the type of the genus, and to what
formation does it belong ? These are the questions which it
is proposed to answer in the following communication. What
opinions are held at present may be shown by citing a few
noted authorities.
Davidson * says Terehratala^ Llhwyd, 1696, type T, vitrea
Linn. sp. ” ; but he states further that Llhwyd’s species
Terebratula minor suhruhra is Terebratula maxillata. In the
Appendix issued in 1856, p. 16, he speaks of Terebratula
maxillata, Sow., as being the fossil type of Terebratula.
Dali, in his Index t, writes “ Terebratula 0. F. Muller
1776 ... . Muller cannot be said to have settled the type.
T. vitrea^ Lam., and T. perovalis^ Sow., are generally
accepted as the types of the genus.”
H. Douvill^, * Genres de Brachiopodes/ says J : — Tere-
bratula, Klein, 1753. Espece type: Terebratula terebratulay
Linne sp. Ce genre a ete fonde par Klein pour les Concha
anomia de Fabio Colonna ; Klein reproduit une des figures
de cet auteur repr^sentant une T4rebratule fossile du groupe
des Biplicatcej trbs-voisine de la T. ampulla du Pliocbiie
d’ltalie. C’est cette figure, k laquelle Linne a appliqu^ le
nom specijtque de Terebratula^ que nous prendrons pour type
du genre.”
He reproduces Colonna’s figure and names it Terebratula
terebratulay Linne sp.”
Hall and Clarke, in their ^ Introduction to Brachiopoda,’
state §: — ^^Terebratula, Klein, 1753 It is inferred
that the species [Linne’s Anomia terebratula^ is a fossil from
the Mesozoic or Tertiary formations, though its geological
horizon is not more precisely known.” They reproduce
Colonna-Klein^s figure and give it the name “ Terebratula
simplex, Klein ” ||.
Schuchert, in his ^ Brachiopoda ’ If, says concisely : —
* ‘ Classification of the Brachiopoda,’ Monograph, vol. i. part 1. n 62
(Pal. Soc.) (1854). ^
t Bull. U.S. Nat. Mus. viii. 1877, p. 70.
X Bull. Soc. G4ol. France, (3) vii. p. 264 (1880).
§ Thirteenth Ann. Keport State Geologist for 1893, vol. ii p 875
(1894).
II See note below, p, 530.
^ Text-book Pal., Zittel, transl. Eastman, vol. i. p. 329 (1900).
526
Mr. S. {S. Buckman on
Terehratulaj Klein, 1753. Genus not well known, Meso-
zoic or Tertiary ” ; and he appends a figure labelled Tere-
hratula Phillipsiy Morris.”
These extracts show current opinion ; but all except Dali
ascribe the name to pre-Linnean authorities, which is contrary
to present custom. If a pre-Linnean authority were to be
taken, it would not be Klein, but Llhwyd, who was the true
originator of the term *. However, modern nomenclature
begins with Linnd t? and as he did not use the term Tere-
hratula in a generic sense, it is necessary to take the first
post-Linnean authority. The following is the result of an
investigation into the practice of early post-Linnean users of
the term : —
1776. 0. F. M,uller, Zool. Dan. Prod. p. 249.
This is the first post-Linnean use of the term Terehratula\,
He mentions three species, the first of which is Terehratula
cranium §, of which he says : — Hsec Terehratula auctorum,
an vero Linnaii? Valvulas enim nec bi- nec triplicatas
invenio.” It is evident that Muller employs Terehratula
as an accepted term in general use, and is not defining it or
giving a type, though he indicates that what Linnaeus called
Anomia terehratula is a basis of reference.
1776. Johann Samuel Schroter, Journ. des Steinreichs
und d. Konch. vol. iii. pp. 372 et scqq.
He speaks of species as belonging to the Terehratulas
acunosas (p. 374), but he does not give binomial names.
1777. Johann Samuel Schroter, Abh. fiber versch.
Gegenstande der Naturgeschichte, 20tli part, p. 358.
Separates Terehratula from Anomia and defines it Das
mag also eine Terebratul seyn eine zweischalige umgleich-
shalige Muschel die eine durchbohrten Schnabel hat.”
1785. Thunbero, Disputationes, vi. p. 99.
' He gives a definition of Terehratula^ and mentions T.pecti-
nata first and Terehratula terehratula sixth. For him
Terehratula includes all Brachiopods except Crania.
^ 1696. Lith. Brit. Iconogr.
t 1758. Linn^, Systema Nat. ed. x.
X My authority for this statement is Mr. C. 1). Sherboru’s excellent
‘ Index Animalium.’
§ That he places T. cranium first is a mere chance, and of no value as
indicating the type.
527
B rac h topod Nomenclature.
1788. A. J. Retzius, Diss. Nov. Test. Gen. p. 13.
Gives a definition of Terehratula. Places therein first
Tereh. caputserpentis^ and fourth Tereh. plieata^ to which
lie gives as a reference Anomia terehratulaj Linnd, though
evidently a mistaken definition. On p. 13 he speaks of
Terehratulas Linneanas^ and how they differ from Anomia.
1792. Bruguiere, ‘^Sur deux Nouv. Esp. de Terebratules
fossiles,” Journ. d’Histoire Nat. i. p. 419.
Bruguibre describes, with figures, two species — Terehratula
cor and Terehr. pileus — which now belong to the genus
Antinomia^ Catullo
1793. A. Modeer, Vetens. Acad. Nya Handl. p. 180.
He defines Terehratula, but gives no type.
1798. Cuvier, C., Tableau Elementaire, p. 434.
The first species which he cites under Terehratula is the
recent species T. vitrea, but he gives as a synonym of this
Anomia terehratula, Lin. He does not fix any type.
1798. J. F. Bolten, Mus. boltenianum, p. 192.
Catalogues certain recent Terebratuloids under the genus
Anomia, Die Terebratel,” and removes Anomia ephippium
into a genus Fenestella.
1799. Lamarck, Prodr. Nouv. Classif. Coq.,” Mdm. Soc.
d’Hist. Nat. Paris, p. 89, Terehratula.
He gives a description and places Anomia terehratula,
Lin.,” as type.
There is no necessity to pursue this part of the subject
further. Muller, it may be said, has given us an indication
that he would regard a plicate Terehratula like the Anomia
terehratula, Linne, as the typical form, and he is doubtful if
T. cranium is a (or the) Terehratula, because it is not plicate.
Then Thunberg definitely includes Terehratula terehratula
among his species of the genus, and, according to the accepted
zoological rule, when no selection has been made a species
with the trivial name the same as the generic becomes type
of the genus. Fortunately the person who first made definite
* S. Buckman, Brachiopod Homoeomorphy,” Quart. Journ. Geol. Soc.
Ixii. p. 433 (1906).
528
Mr. S. S. Buckman on
selection — Lamarck — complied with this rule. He chose
Anomia terehratula^ Lin., to be the type ; thus he may be
said to have confirmed Muller and Thunberg. So we may
write
Genus Terebratula, Muller, 1776.
Genotype Anomia terebratula, Linne.
Syn. Terebratula, Klein, pre-Linnean.
Non Terebratula, Llhwyd, pre-Linnean, y^hich—Epithyris, Phillips* * * §.
It is now necessary to identify Terehratula terebratula
(Linnd). That author describes and cites figures for his
species, thus: — ‘‘A. testa obovata l^vi convexa; valvula
altera triplicata, altera biplicata. — Column, purp. 22 f. 1.
List. angl. 240 t. 8. f. 46. Klein ostr. t. 11. f. 74.”
Of these figures the first and third refer to the same shell,
for Colonna used the same woodcut in various places, and
Klein copied it. But Lister’s figure represents a very
different species : it is not plicate, and therefore it does not
agree with Linnd’s description f* As the Colonna- Klein
figure does, that must be taken as the holotype, which, in
fact, has been the usual practice.
In 1819 Valenciennes I gave the name Terebratula Kleini
to a species, citing Klein’s figures, and quoting as a synonym
Anomia terebratula, Linne. In 1850 Davidson § said that
the name Tereb. Kleini covered a well-known Bajocian fossil
of Normandy (it is from the Maliere), and he figured, in
support of his opinion, a specimen so labelled in the Lamarck
collection.
In 1856 S. Hanley identified \A\me' ^ Anomia terebratula 2i^
Terebr. perovalis, Sowerby ||.
In 1864 Deslongchamps followed Davidson’s line, making
T, Kleini, Davidson (? Lamarck), a variety of Terebratula
perovalis If.
Though Lamarck may have thought that the Bajocian
fossil was the same as Klein had figured, and so may
have ticketed it T. Kleini in his collection, yet it is not
the holotype of T. Kleini \ the specimen figured by Klein
* Ami. & Mag. Nat. Hist, ser. 7, vol. xviii. 1906, p. 322.
t Lister’s smooth non-plicate Terebratulid is from near Grantham,
probably therefore from Middle Lias, and it has much the aspect of one
of the T. punetata series.
I In Lamarck’s Anim. sans Vert. p. 252.
§ Ann. & Mag, Nat. Hist. (2) v. 1850, pi. xiii. Hg. 33.
II Ipsa Linneei Conchylia.
Tal. fran^*., Terr. jur. Brach. p. 197.
529
Bmchiopod Nomenclature.
from Colonna must be that. And I find it impossible to think
that the Bajocian fossil and Colonna^s represent the same
species. Both are biplicate, but there the similarity ends.
This Bajocian fossil is peculiar to Normandy ; but Colonna
figured an Italian shell — he tells that it was found “in
Civitate Andriae.” Now there are in the Tertiary (Pliocene)
beds of Italy various species which are much more like
Colonna’s figure; and there is in the British Museum, Nat.
Hist., no. 83458, a specimen from the Tertiary of Monte
Mario, near Rome, which might almost be the original
drawn by Colonna, so like is it to his figure. This specimen
is represented in the delineations given in Plate XII.
Colonna’s figure differs from the Terehratula Kleini of the
Bajocian of Normandy in having the plications more nearly
equal, running further up the valves, and in being much
more plicate for its smaller size. In the Bajocian shell the
frontal fold is much elevated, somewhat after the T. sella
fashion, and the dorsal sulcus is inconspicuously developed.
Colonna’s shell is distinguished from Terehr, perovalis,
Sow., by having much more pronounced plications, extending
much further up the valves. Again, it is distinguished
from Terehr. Phillipsi^ Morris, by lacking the very pronounced
angular plications, by lacking the pronounced posterior
acumination, and by having an incurved beak : in T. Phillipsi
the beak is not incurved, it projects straight posteriorly.
There is one Mesozoic species to which Colonna’s figure
has rather more resemblance than to those mentioned above.
This is T. Stephanie I)av. ; but its folds are not so strong,
do not extend so far back, and the shell is less elongate in
shape.
The conclusion arrived at is that Terehratula terehratula —
that is, Colonna’s shell — is not a Mesozoic species ; but it is
a Tertiary fossil, closely allied to species which have been
called Terehr. hisinuata, Lamarck, T. sinuosa, Brocchi,
T. ampulla^ Brocchi, and even to T. grandis, Blumenbach.
These species may be said to belong to Terehratula, sensu
stricto ; but it may be doubted if any Mesozoic species would
belong to the genus in this very limited sense.
The conclusions now arrived at confirm the results of
H. Douvilld’s investigations — that Anomia terehratula, Linne,
is the genotype of Terehratula, that the species is a Pliocene
shell near to T. ampulla ; but they differ in the small detail
that the name Terehratula must be ascribed to Muller, 1776,
and not to Klein, 1753.
My application of the name differs from Douville’s. He
would keep Terehratula for the biplicate species (p. 265),
530
On Brachiopod Nomenclature.
but I would restrict Terehratula to the species in actual
genetic connexion with T. terehratula — 1 T. hisinuata. The
forerunners of such species would be non-plicate, perhaps
uniplicate, before becoming biplicate. Biplicate Terebratulids
are polygenetic descendants from many non-plicate forms :
sometime they must be separated from Terehratula into their
special genetic series.
For a series of Cretaceous biplicate Terebratulids, which
will no doubt prove to be separable enough from the Tertiary
series, the name Musculus^ Quenstedt, seems to be available.
Genus Musculus, Quenstedt, 1868.
Type Terehratula acuta., Quenstedt.
Kef. Petref. Deutschl. ii. pp. 4, 27, 384, pi. xlviii. fig. 70.
For a Jurassic biplicate series, the T. maxillata group,
very separable from other Terebratulids, the name Epithyris,
Phillips, is available *.
My best thanks are due to Mr. B. B. Woodward, F.L.S.,
and to Mr. C. Davies Sherborn, F.G.S., for invaluable
bibliographic assistance in connexion with the investigations
for this communication ; also to Mr. W. D. Lang, F.G.S.,
for kind answers to various enquiries.
Appended are a few of the synonyms by which Terehratula
terehratula has been known ; these names have been attached
to the figures of Golonna or Klein, or else to the description
by Linne.
Terehratula terehratula (Linn^).
1616. Concha rarior anomia., &c., Colonna.
1763. Terehratula rugosa, Klein f*
1758. Anomia terehratula, Liim6.
1785. Terehratula terehratula ; Tbimberg.
1787. Lampas terehratula ; Meuschen, Mus. Geversianiim, p. 438.
1788. Terehratula plicata, Retzius.
* See above, p. 528.
t Klein’s statement is “ Terehratula .... § 427 species 1. simplex] &
rugosa, Fab. Columnae de Purp. Cap. xii. § 3, pag. 32 ... . Icon exstat
in Tab. Nostra xi. No. 74.” Terehratula rugosa seems therefore to desig-
nate the biplicate shell figured by Colonna, § 3, p. 32, and reproduced by
Klein ; Terehratula simplex may have been intended to designate the
non-plicate Terehratula figured by Colonna in p. 33, § 5.
. IhihA^r . Uf\sl .S. 7. Vo! . AIX . VI . XU.
J. Greer, del. et lith.
TEREBRATULA TEREBRATULA.iW. ^
r:v:.:.- im^?y ■ ■ ■
CF THE ' ;.;
UNIVERSITY of ILLINOIS. .
.f •••
i
iC
'r - ’■'.v/.
■ ■/
iTii '
/ ,
*v
Descriptions and Records of Bees.
531
1797. Lampas columbina^ Humphreys, Mus. Caloneanum, p. 45.
1798. Terebratula vitrea ; Cuvier.
1799. Terebratula terebratula', Lamarck.
1819. Terebratula 'Kleinii, Lamarck.
1856. Terebratula perovalis ; Hanley, Dali.
1880. Terebratula terebratida ; Douvill^.
1894. Terebratula simplex ; Hall & Clarke.
EXPLANATION OF PLATE XII.
Terebratula terebratula (Linn4).
a, drawn as seen ; c, restored outlines.
LXIV. — Descriptions and Records of Bees, — XIV.
By T. D. A. Cockerell, University of Colorado.
Crocisa Wellmani^ sp. n.
$ . — Length 14 mm. or slightly more.
Black with bluish-white hair-markings. Anterior wings
very dark fuliginous, with a pair of suffused hyaline spots
just beyond the third submarginal cell. Face densely white-
haired ; clypeus very densely finely punctured ; antennge
dark ; pleura with the upper half covered with bluish-white
hair, the lower half apparently nude, but with scanty black
hair, and with very strong well-separated punctures ;
pattern of thorax above simulating a face, with a median
nose-like band, a pair of spots for eyes, and a transverse mark
on each side anteriorly like a moustache, all strongly defined,
the appearance being further aided by a stripe on each side,
curving inwards posteriorly, which outlines the head and
suggests hair ; margin of scutellum of the ^-type, with a
conspicuous round spot of hair on each extreme side, but no
median spot; a white fringe from underneath the middle;
anterior tibiae white-haired on outer side, middle and hind
tibiae with somewhat more than the basal half white-haired
on outer side ; basitarsi with a good deal of white hair ; first
abdominal segment with a pair of large U-like lateral marks,
placed laterally and not joined basally ; the broad bands on
the other segments also broadly interrupted, that on the
second with a pointed projection anteriorly near the side of
the segment; apical plate narrow; last ventral segment
produced ; fifth ventral not keeled.
In Friese’s table (Verb, zool.-bot. Gesell. Wien, 1905,
p. 174) this runs to C. arcuata^ Vachal, but that species is
532 Mr. T. D. A. Cockerell — Descriptions and
somewhat larger, and has the basal band of the abdomen
entire. The scutellum brownish-haired (which is not at all
the case in ours), with its spots decidedly blue.
Hah. Portuguese West Africa. Long. E. 15° 05', lat. S.
12° 44', alt. 1360 metres above sea-level, at flowers of
^olanthus^ Dec. 1906 (middle of rainy season) ; collected by
Dr. F. Creighton Wellman. At the same flowers Dr. Wellman
also took a small (hardly over 8 mm. long) ? of G, meripes,
Vachal.
Anthophora quadrifasciata (Villers), $ .
Same locality as Crocisa Wellmani. Dr. Wellman notes :
“ Stands motionless in the air like a bot-fly. Taken near a
flowering Convolvulus^ but not actually seen in the flowers.
Its hum suggests a bottle fly.”
Anthophora convolvuli, sp. n.
? . — Length about 12 mm.
With mainly fulvous hair ; superficially looking exactly
like A. vestita. Smith, but differing as follows: — labrum
yellow (as in A. capensis, Friese), with a dark spot at each
basal corner ; mandibles yellow with the apex dark ;
clypeus with narrow anterior margin and a narrow median
band yellow ; a small supraclypeal mark ; hair of thorax
yellower ; hair on middle and hind basitarsi, except a tuft
posteriorly, black. The tegulee are clear red, and the hind
margins of the fulvous-haired abdominal segments, as in
vestita^ appear pallid.
Collected by Dr. Wellman in the same locality as Crocisa
Wellmanij in Dec. 1906, at flowers of Convolvulus.
Apis nigritarum, Lepel.
Dr. Wellman sent numerous workers from the same locality
as the Crocisa j &c., taken at various flowers, chiefly
Leguminosae.
Agapostemon coloradensisj Crawford.
The male, not hitherto described, was found by Mr. S. A.
Kohwer on the campus of the University of Colorado, at
Boulder, Oct. 3, 1906. It has the head and mesothorax
above blue, exactly as in A. calif or nicus^ but it is a larger
insect, and the yellow band on the clypeus sends upwards a
little projection in the middle. Abdomen with five yellow
bands, the first with a pair of black spots in the middle ;
533
Records of Dees.
last ventral segment not keeled ; trochanters black, the hind
ones with a yellow spot ; hind tibiae yellow, with a small
black mark in front and a larger one behind.
Augochlora viridula^ Smith.
This insect has two forms, one golden-green, the other
blue-green. Mr. N. Banks has taken females of both at
Falls Church, Virginia : the golden-green at flowers of
Ceanothus in June ; the blue-green on the 2nd of August,
flower not stated. Mr. C. Eobertson has sent me the blue-
green from Southern Illinois. A. viridula was founded on a
male of the golden-green form ; A. hicidula^ Smith, on
females of both. I have some suspicion, though I cannot
produce any proof, that the two forms are distinct, but very
closely allied species ; if this is the case, it may be permissible
to retain the name lucidula for the blue-green one.
Augochlora Banksiella^ sp. n. [pura, subsp. ?).
$ . — Length 1-1\ mm.
Brilliant green, the abdomen golden green, the head and
thorax also with a golden lustre — no blue or purple ;
mandibles with a green basal spot; antennas dark, flagellum
obscure brown beneath ; orbital margins strongly converging
below ; tegulas fusco-ferruginous with a green spot. Wings
greyish ; stigma and nervures testaceous ; first r. n. joining
second t.-c., second s.m. almost as broad as high, marginal cell
appendiculate ; area of metathorax with fine but strong striae.
Legs piceous, anterior and middle femora green on under side ;
hind spur simple, curved ; dorsal surface of abdomen nude, not
hairy, or rather with exceedingly scanty short pubescence ;
no vibrissae ; hind margins of segments exceedingly narrowly
black.
Very closely related in all respects to A, pur a, Say
[Rohertsonij Cklh), but readily distinguished by the strong
golden tints and the usually larger area of raetathorax. It is
perhaps a subspecies of A, pura. The male has the tarsi
ferruginous.
Hab, Glencarlyn, Virginia, May 4-July 20 ; one, June 21,
from flowers of Ceanothus •, Great Falls, Virginia, April 20 ;
Odenton, Maryland, July 18. All collected by Mr. Nathan
Banks.
The genuine A. pura I have from Ithaca, New York
[Banks') and Illinois {Robertson). In identifying the latter
I follow Kobertson, but Say’s original description could be
applied to either form, perhaps agreeing a little better with
534 Mr. T. D, A. Cockerell — Descriptions and
Banksiella. Apparently Say did not separate the two or
three allied species of the pura type, and as he doubtless
included the insect which Kobertson calls pura, this appli-
cation of the name may be allowed to stand.
Augochlora aztecula, sp. n.
$ . — Belongs to group Oxystoglossi, with the hind spur not
pectinate. Length about 7 mm. ; anterior wing about 5.
Golden green, middle of second abdominal segment stained
with coppery red ; mandibles ferruginous, darker at base
and apex ; tubercle of lab rum simple, truncate ; face broad,
orbits very strongly emarginate, and strongly converging
below ; clypeus with strong rather close punctures, and its
anterior edge broadly black; flagellum dull ferruginous
beneath ; mesothorax and scutellum brilliantly coloured,
appearing granular from extremely minute and close punc-
tures ; hind part of scutellum with some fuscous hairs ; area
of metathorax with fine longitudinal ridges, except the
middle subapical part, which has finer transverse strim ;
tegulse ferruginous with pallid margins. Wings dusky,
somewhat reddish ; stigma and nervnres testaceous ; first
r. n. meeting second t.-c. Legs black, with the knees, the
tarsi, and the anterior tibiie (except a cloud behind) ferrugi-
nous, the anterior femora green beneath. Abdomen with the
hind margins of the first two segments more or less piceous,
though very narrowly ; no marginal vibrissae, but the lower
edge of the dark base of the second and third segments,
which slides under the segment before, is very finely and
regularly ciliate ; venter piceous, with a subapical ferruginous
stain. The ventral surface of the abdomen, as well as the
hind legs, carries much pale yellow pollen.
In my table in ^ Canadian Entomologist,’ 1897, p. 4, this
runs nearest to A. labrosa. Say, and A. aurifera, Ckll., but
it is readily separable from both by the colour of the legs.
In Schrottky’s table of Brazilian species it runs near
A, urania, Sm., but that is smaller, with a blue-green
abdomen.
Bob. Tlacotalpam, Vera Cruz, Mexico, April 21 (C. H. T.
Townsend), At the same place, on the same day, Professor
Townsend took Megachile Candida^ Smith, ? .
Augochlora cordioefioris, sp. n.
^ . — Belongs to Oxystoglossi, but with stature of a Ghlora-
lictus ; length about 6 mm.
Head and thorax brilliant green ; abdomen with the first
535
Records of Bees.
segment brassy green, the hind margin broadly black ;
second and third segments similar, but not so green, and the
margin redder ; fourth segment hardly green at all ; apex
black, with short dark hairs ; bases, which slide under the
segments before, strongly reddish ; venter rufo-piceous ;
mandibles ferruginous with dark tips j labrum ferruginous,
with a small tubercle ; clypeus strongly punctured, its anterior
edge broadly dark ; eyes strongly emarginate ; flagellum
ferruginous beneath, except at base ; mesothorax and scutellum
very finely and closely punctured, the colour of the scutellum
especially brilliant ; area of raetathorax plicatulate basally ;
tegulas dark reddish with hyaline margins. Wings scarcely
dusky; stigma and nervures testaceous; first r. n. meeting
second t.-c. Legs bright ferruginous, the anterior and middle
femora infuscated, the anterior ones even slightly metallic ;
coxae dark, the anterior ones metallic ; abdomen with a fine
pruinose pubescence.
Easily known by the small size, red legs, and comparatively
dark abdomen.
Somewhat related, apparently, to A. tisiphone^ Gribodo.
Hah. San Rafael, Vera Cruz, Mexico, on flowers of plant
no. 31 [Gordia, probably G. ferruginea), middle of July
((7. H. T. Townsend). On the same flowers, at the same time
and place. Professor Townsend took Megachile chrysophilaj
CklL, ?.
Hesperapis larrece^ sp. n.
Hesperapis larreee, Viereck, MS., 1902 (no description).
d . — Length about 7 ram.
Black, with rather abundant white hair ; hair of face and
cheeks long and dense ; mandibles reddened at tips ; clypeus
shining and very finely punctured ; vertex shining, almost
entirely impunctate ; antennae dark, flagellum faintly reddish,
scape quite long ; mesothorax shining, but very hairy ; area
of raetathorax nude, smooth, and very shiny ; tegulse testa-
ceous. Wings milky hyaline; stigma amber-colour bordered
with dark brown ; nervures brownish ferruginous. Legs
black, hairy. Abdomen narrow, hairy, roughened with fine
punctures, the bases of the segments depressed, the apical
margins faintly reddish ; apex with a narrow ferruginous
pygidial plate.
The general aspect is suggestive of H. olivice (Ckll.) , but
that has the wings dusky, the abdomen more distinctly
banded, and the flagellum bright ferruginous beneath. The
venation also differs, the basal nervure being more strongly
536 Mr. T. D. A. Cockerell — Descriptions and
bent, and more distant from the transverso-medial, in larrece
than in olivice.
Hah. Mesilla Park, New Mexico, at flowers of Larrea^
May 16 {Cockerell) ; also taken by Mr. Viereck at Alamo-
gordo, New Mexico, at Larrea.
Andrena pyrrhacitay sp. n.
$ . — Length about 11^ mm.
Kobust, black, with very abundant erect hair on head,
thorax, and abdomen. Head broad, facial quadrangle very
much broader than long ; facial fovege black, broad but short :
hair of clypeus, sides of face, and vertex black ; of front and
between antennae, and occiput, yellowish white or pale
ochreous ; of cheeks black, with more or less pale above and
below: antennse black, third joint much longer than 4 + 5,
flagellum faintly brownish beneath ; process of labrum
entire; clypeus dull, with dense minute punctures and a
rather faint median ridge ; thorax dull and granular, with
abundant, long, erect, pale ochreous hair; lowest part of
pleura with black hair ; area of metathorax without rugae or
raised margin. Legs black, with mainly black hair, but it is
long and pale on anterior femora behind, and pale on inner
side of hind tibiae ; tegulae piceous. Wings hyaline, nearly
clear, the apex a little dusky ; stigma and nervures dark
rufo-fuscous ; second s.m. receiving first r. n. beyond the
middle ; abdomen with only minute feeble punctures, not
banded, but covered with erect hair of a rather light ferrugi-
nous colour, whitish on first and fifth segments, black or
sooty at extreme apex ; second segment depressed about one
third ; basal part of venter with coarse black hair.
At first sight this suggests A. Hiteij Ckll., but it has much
less brilliantly coloured hair, a broader, more oval abdomen,
and the sculpture of the clypeus is entirely different. It is a
very distinct and beautiful species.
Hab. Salina, Boulder County, Colorado, 6550 ft. alt., at
flowers of Salioj, April 14, 1907 {W. P. and T. D. A.
Cockerell). Another specimen, with the hair of the abdomen
less brightly coloured, was taken by Mr. G. Hite at Boulder,
Colo., March 25, 1907.
Andrena mimetica Falli, subsp. n.
$ . — Differs from A. mimetica, Ckll. (which occurs in
New Mexico), thus: smaller, length about or just over 11 mm.;
third antennal joint shorter than 4 + 5; second and third
abdominal segments with white hair at base, conspicuous
when the abdomen is seen from the side.
537
Records of Bees.
Hah. Southern California, Mt. Wilson {Davidson?). Also
on Mt. Wilson, Dr. Davidson took that remarkable species
A. cleodora, Viereck, hitherto only known from Oregon
[cf. Oanad. Entom. 1904, p. 161).
The new subspecies is named after Mr. H. C. Fall, the
Coleopterist of Pasadena, California.,
Nomada Edncej sp. n.
. — Length 9 mm. or a little more; anterior wing about
or not quite 8 mm.
Black with lemon-yellow markings, the legs red. Head
and thorax very hairy, the hair light fulvous ; vertex and
mesothorax dull and densely punctured ; eyes in life greenish
grey, delicately suffused with red, especially above ; maxillary
palpi long; head broad; mandibles simple; clypeus not
hairy, but supraclypeal region very hairy; clypeus, broad
lateral corners of face, extending narrowly up the orbital
margins nearly to level of antennae, a mark beneath eyes
behind, labrum and basal two thirds of mandibles, all yellow.
Antennae stout, scape not especially swollen, mainly yellow
in front ; flagellum bright ferruginous, its first four joints
with a large black mark above, that on first narrower than
on the others; last joint normal; second antennal joint
sunken into apex of scape ; third joint much shorter than
fourth ; prothorax above, tubercles, patch on lower part of
pleura, and a pair of large coalescent spots on the gibbosities
of the scutellum, all yellow ; minute red axillar spots; post-
scutellum and metathorax all black ; tegul^e red with a large
yellow spot. Wings strongly dusky at apex ; stigma bright
amber-colour, nervures more fuscous ; b. n. passing a little
basad of t.-m.; second s.m. as large as third, not especially
narrower above, and receiving the r. n. at its middle! Legs
red, hind femora mainly black behind ; anterior coxse un-
armed. Abdomen dullish, with exceedingly fine punctures;
basal half of first segment black, after which comes a yellow
band on a red field, interrupted in the middle ; segments 2 to
6 with very broad entire yellow bands, those on 2 and 3 a
little notched behind laterally ; hind margins of segments
broadly red, those of 1 and 2 infuscated ; apical plate
notched ; venter red, banded with yellow.
A species of the Xanthidium group, which runs in my
table of Rocky Mountain Nomada (Bull. 94, Colorado Exp.
Station) to 47, and runs out because of the comparatively
large size and notched apical plate. It has a strong super-
ficial resemblance to N. ornithica.^ CklL, but that is much less
hairy, and differs in the colour of the legs &c. Among the
Ann. dc Mag. N. Hist. Ser. 7. Vol. xix. 37
538 Mr. T. D. A. Cockerell — Descriptions and
Old World species, it shows a general resemblance to
JS . succincta, Panz.
Hah. Boulder, Colorado, on the campus of the University
of Colorado, at flowers of Taraxacum taraxacum^ in company
with N. cuneata, Rob., Osmia^ &c., April 10, 1907.
Collected by Miss Edna Baker.
Osmia gaudiosa^ sp. n.
$ . — -Length 6 mm. or slightly over.
Brilliant golden green, including legs ; the vertex, front,
and thorax above with a strong suffusion of coppery-red or
almost crimson ; abdomen with strong reddish-golden tints ;
hair of head and thorax long and white ; antennae normal,
slender, not moniliform, black with a slight brown tint ;
tegulae bright green, shining golden ; wings clear. Abdomen
strongly punctured, its hair white ; first ventral segment
feebly subemarginate ; second ventral peacock-green ; sixth
dorsal with a very feeble median depression, hardly emar-
gination ; seventh segment bidentate.
Hah. Boulder, Colorado, on the campus of the University
of Colorado, at flowers of Taraxacum taraxacum^ April 10,
1907 [Edna Baker).
A most lovely little species, very distinct from all others
in North America.
Osmia universitatis^ sp. n.
^ . — Length about 9 mm. j anterior wing about 7.
Head and thorax olive-green, some brassy colour on middle
of front, metathorax bluish green ; abdomen dark green,
approaching bluish green, the hind margins of the segments
narrowly purplish black ; legs black, with no metallic tints.
Head and thorax very densely punctured as usual, their hair
long and abundant, dullish, with a yellowish tinge, but over
clypeus forming a dense shining white beard ; no black or
dark hairs intermixed anywhere ; antennae long and slender,
black, the flagellum obscure ferruginous beneath, not monili-
form ; tegulae black. Wings hyaline, very clear; b. n.
falling just short of t.-m. ; hair of legs mostly pale, but
black or dark fuscous on basitarsi (redder within), and the
hind tibiae have long black hairs on outer side, while the
hind femora show short black hair, easily overlooked because
of the longer pale hair. Hair of abdomen partly black on
fourth and following segments; sixth segment entire;
seventh bidentate, the })oints quite wide apart ; first ventral
entire, third deeply emarginate. Eyes slate-colour. Second
and third joints of middle tarsi broadened and thickened.
Records of Bees, 539
This is an ordinary looking species, having a general
resemblance to the following : —
(1) 0. faceta, Cress., from which it differs by the green
abdomen, without black hair on sides of second
segment ; segment 6 entire, &c.
(2) 0. inurhana, Cress., from which it differs by the clear
wings, different tint of abdomen, different colour of
hair of thorax above, &c.
(3) 0. iridis^ C. & T., from which it differs by the
absence of black hairs on head, the smaller, greener
abdomen, &c.
(4) 0. chloropsj C. & T., from which it differs by the non-
moniliform antennse, &c.
Among the Old World species it resembles 0, Lafreillei.
Spin.
Hah. Boulder, Colorado, on the campus of the University
of Colorado, at flowers of Taraxacum taraxacum, in company
with 0. gaudiosa^ April 10, 1907 {Edna Baker).
Bombomelecta arizonica, CklL
This species was described from a single female. Professor
R. H. Forbes has found a male in a cell of Anthophora Forbesij
Ckll., at Tucson, Arizona, April 13, 1907.
This is the first record showing the host-relationships of
Bombomelecta. The male B. arizonica is very similar to that
of B. Alfredi, but smaller (length hardly 12 mm.), with a very
distinct black band between the wings, consisting of a densely
punctured area free from the white pubescence, but with a
moderate amount of black hair. The middle tibiae are
densely covered with snow-white hair on the outer side ;
the posterior tibiae and tarsi have much wdiite hair on the
outer side. The fourth and fifth ventral abdominal segments
are covered with white hair.
Exomalopsis solidaginis, Ckll.
Mesilla, New Mexico, June 24, ? {Cockerell)] Albuquer-
que, New Mexico, June 30, (^ , with only two submarginal
cells, the same on both sides {Cockerell).
Perdita dasyliru^ sp. n.
Exceedingly close to P. lucioe, Ckll., but with the meso-
thorax, scutellums, and upper surface of metathorax entirely
dark metallic green, the green being also of a bluish tint, not
yellowish as in lucice. It is also close to P. Martini^ Ckll.,
but the male has a large yellow patch on the pleura, and
largely darkened nervures, and the yellow of the face does
510
Bibliographical Notices.
not reach to tlie anterior ocellus, nor are any black dots left
at the sides of the eyes.
$ . — Length about 4^ mm.
Bright but light yellow ; mandibles tipped with rufous ;
head yellow except a broad area from the vertex, a little in
front of anterior ocellus, to the occiput, which is dark green
and granular; cheeks yellow, the dark area not extending
downwards; antennae yellow, a dark dot on second joint
above ; thorax dark above, but yellow at sides and beneath,
except a dark mark just below wings; prothorax above with
a broad yellow margin ; nervures and margin of stigma
fuscous, third discoidal cell absent ; marginal cell broadly
but obliquely truncate ; legs yellow, hind tibiae with a faint
dark shade above. Abdomen yellow with apex a little
reddened ; first segment black with two more or less
triangular yellow marks, and sides yellow; four straight
black bands, not reaching lateral margins, the first three
broad ; venter yellow, rufescent apically.
J . — Length 3|-4 mm.
In general similar to the female, but dark colour extending
a short distance down cheeks, leaving a narrow yellow line
next to the eye; flagellum, except last joint, fuscous above;
sides of thorax dark, but pleura with a very large yellow
patch anteriorly ; hind tibiae and tarsi behind, and a spot at
apex of their femora, dark brown ; apical part of abdomen
orange or brownish yellow, without bands ; there are four
well-formed bands.
Hah. Alamogordo, New Mexico, at flowers of Dasylirion
Wheeleri, Watson, June 6 to 9, very many specimens {H. L.
Vierech) .
University of Colorado,
Boulder, Colorado, U.S.A.,
April 20, 1907.
BIBLIOGBAPHICAL NOTICES.
The Crawfishes of the State of Pennsylvania. By Dr. A. E. Ortmanx.
Memoirs of the Carnegie Museum^ Pittsburg, Pa., vol. ii. no. 10,
pp. 343-523, 7 plates : December, 1906.
Since Huxley, in a well-known paper, first placed the classification
of the Crayfishes on a morphological basis and showed the interest
attaching to their geographical distribution, much attention has
been given to this group of Crustacea. In particular, the very
numerous species occurring in North America have been the subject
of important memoirs by Faxon and others. Dr. A. E. Ortmann,
who is well-known as an authority on the liigher Crustacea, has
produced, in the memoir under review, a study of the Crayfishes of
541
Bibliographical Notices,
Ponnsylvania which deserves attention, not only from Carcinologists
but from all who are interested in the wider problems of zoology.
In the first part of the memoir the author discusses the systematic
characters of the forms found within the limits of the State. Seven
species and one variety are recognized, and their variations are set
forth in great detail. Dr. Ortmann’s conclusions as to the limits of
the species are not to be lightly disputed, were it only on account
of the vast amount of material — between two and three thousand
specimens — at his disposal. Further, his familiarity with the living
animals — most of the material was collected by himself — gives him
a great advantage over the mere museum species-maker.
The next section deals at length with the oecology and distribution
of the various species. Especially interesting are the details given
of the habits of the “ chimney-builders — the burrowing species
which are so called from the chimney-like piles of mud thrown up
at the mouths of their burrows. These species are found often at
considerable distances from open water, burrowing down to reach
the ground-water, and their chimneys are sometimes so numerous
as to “ hamper farming operations by interfering with the harvesting
machines, clogging and ruining them ” ; elsewhere they were
observed “ coming up even between the railroad ties of the Mor-
gantown and Kingwood Railroad.” The distribution of the species
within the State is illustrated by coloured maps, and it is shown that
many of the facts cannot be correlated with the existing physio-
graphical conditions. An explanation is sought in past changes in
the configuration of the country and especially of the river-systems.
How far the author’s speculations as to the pre- and post-glacial
migrations of the various species are justified is a question which
can only be answered by local research, but they are certainly
suggestive and stimulating.
The life-histories form the subject of the following section, and
remarkable differences are shown to exist between the species in
this respect. Two main types of life-history are distinguished, a
“ warm water ” type characteristic of the species found in the larger
rivers, and a “ cool water ” type shown by the species found in
mountain-streams. In the former a definite seasonal cycle is
observed, mating taking place in the autumn and spawning in the
spring, while the cool water species breed at all seasons of the year.
Some details are also given of the remarkable alternation of breeding
and non-breeding phases which, as Faxon discovered, gives rise to
the so-called “ dimorphism” of the males in the American Crayfishes.
A similar phenomenon has recently been described by Mr. G. Smith
in certain crabs and may possibly be found to occur in other
Crustacea.
After a short section dealing with the economic aspects of the
Crayfishes, the memoir concludes with a discussion of some current
theories of evolution in the light of the facts set forth. Many of
the points touched on are of a highly controversial nature, and the
arguments cannot be summarized here. It may be said, however,
that Dr. Ortmann sees no evidence of “ mutation ” in the origin of
species (and here, we believe, many systematic zoologists will agree
542
Bibliographical Notices,
with him), but concludes that “Isolation or Habitudinal Segregation,
as the factor forming species, is clearly seen in every case
discussed.”
It is, perhaps, to be regretted that Dr. Ortmann has seen fit to use
the term “ Crawfish ” as the “ proper American name ” of the
animals he deals with. It is surely pushing the rule of priority to
an absurdity 'to apply it in such a case. Apart from this trivial
matter, however, the memoir is one upon which the author and the
Museum with which he is connected are to be congratulated.
W. T. C.
A Natural History of the British Butterflies, their World-wide
Variation and Geographical Distribution. A Textboolc for
Students and Collectors. By J. W. Ttjtt, F.E.S. Vol. I.
London, 1905-1906. 8vo. Pp. iv, 479 j pis. xx.
We have here another of Mr. Tutt’s enormously detailed and
elaborate volumes on British Lepidoptera, which, he tells us, was
issued in parts and should form vol. viii. of the whole series. The
introductory chapters (Part I. Chapters i.-xiv.) are devoted to
general observations on butterflies ; egg-laying, eggs, and larvae ; and
probably the most interesting will be found to be those on the
association of Ants with Butterfly Larvae, and on the Carnivorous
Habits of Butterfly Larvae, wherein the Author brings together a
large amount of scattered information which it is most useful to
have epitomized. Part II. contains a detailed account of the ten
British species of Urbicolides and Buralides (Skippers and Coppers),
and the work closes with an index of eighteen closely printed pages
in double columns. The twenty plates represent eggs, larval hairs,
perfect insects, &c., and one plate represents an apparatus for
photographing the eggs of butterflies. The chapters on Obtaining
Eggs of Butterflies and on Collecting Butterfly Larvae will be found
very useful and interesting to those lepidopterists who care to
undertake such work. It will probably take two hundred similar
volumes to complete the history of the British Lepidoptera on the
grand scale projected by Mr. Tutt, yet all praise is due to him for
his undertaking the initiative in such a gigantic task, and for
having made an appreciable, if comparatively small, diminution in
the number of volumes required for its completion, though this
would require several lifetimes, unless a numerous band of ento-
mologists were to devote themselves to separate portions of the task
simultaneously.
Of the butterflies described in the present volume, Chrysophanvs
dispar is probably the most interesting. Mr. Tutt devotes forty-
eight pages to this species ; but a very large volume might be
written on the subject, and the notice appears to us to be somewhat
less complete than that of some of the other species which ho
discusses.
We need hardly say that Mr. Tutt’s voiumes on British Lepido-
I>tera form an indispensable mine of wealth to all future lepidopterists
who attempt to take up the study of butterflies seriously.
W. F. K.
543
INDEX TO VOL. XIX.
Abr.®us, new species of, 319.
Acalles, new species of, 60.
Acantharctia, new species of, 237.
Acanthojoppa, new species of, 185.
Acantuerta, characters of the new
genus, 241.
Acheilognathus, new species of,
159.
Acmella, new species of, 220.
Agastegnus, new species of, 62.
Aglossosia, new species of, 232.
Agonostomus, new species of, 65.
Agrotis, new species of, 243.
Agyrta, new species of, 299.
Alycaeus, new species of, 157.
Amhlyjoppa, new species of, 187.
Amhlyopinus, new species of, 125.
Amphidromus, new species of, 151.
Amphishseua, new species of, 487.
Anaplectoidea, new species of, 25.
Andersen, K., on the bats of the
family Megadermatidae, 129 ; on
Pterocyon, Rousettus, and Myo-
nycteris, 501.
Andraca, new species of, 49.
Andrena, new species of, 369, 536.
Auhausta, characters of the new
genus, 250.
Anodonta cygnea, on the occurrence
of Spirochaeta anodontae in, 493.
Anomalocera, new species of, 348.
Antarctia, new species of, 239.
Anthophora, new species of, 532.
Anthracophora, new species of,
353.
Anytus, new species of, 250.
Aparallactus, new species of, 325.
Aphelotoma, new species of, 268.
Apisa, new species of, 224.
Apsidopis, new species of, 38.
Arachnida, new, 208, 322, 522.
Arfaka, characters of the new genus,
397.
Area, new species of, 203.
Arrow, G. J., on new parasitic coleo-
ptera from S. America, 125 ; on
new Indian lamellicorn coleoptera,
347, 416.
Arunta, characters of the new genus,
406.
Ascaropsis morrhuae, note on, 92.
Asota, new species of, 50.
Astacura, definition of the tribal
name, 464.
As Lira, new species of, 233.
Auchenophorus, characters of the
new genus, 270.
Augochlora, new species of, 533.
Austen, E. E., on the synonymy and
generic position of certain Mus-
cidae, 326,* on a new genus and
species of Miiscidae, 445.
Babycurus, new species of, 208.
Bajauana, characters of the new
genus, 277.
Barbus, new species of, 493.
aureus, note on, 390.
Bathymysis, characters of the new
genus, 116.
Batrachians, new, 214, 394.
Berg, L. S., on new Cyprinoid fishes
from Korea and N. China, 169
163. '
Bithinella, new species of, 216.
Black-and-tan pattern of domestic
dogs, on the, 192.
Blatta, new species of, 39.
Blattidae, new, 25.
Blepharita, characters of the new
genus, 251.
544
INDEX.
Blepharoa, characters of the new
g’enus, 2ol.
Blepharomastix, new species of, 54.
Bonhote, J. L., on Ochotona aurita,
124.
Books, new : — Alcock’s Catalogue of
the Indian Decapod Crustacea,
127 5 Lankester’s Treatise on Zoo-
logy, pt. V., 127 ; Austen’s Illus-
trations of British Blood-sucking
Flies, 199 ; Catalogue of the Lepi-
doptera Phalaense in the British
Museum, 300; Otaki, Fujita, and
Higurashi’s Fishes of Japan, 448 ;
Webb and Sillem’s British Wood-
lice, 450 ; Ortmann’s Crawfishes of
the State of Pennsylvania, 540;
Tutt’s Natural History of the
British Butterflies, 542,
Booneta, characters of the new
genus, 291.
Borolia, new species of, 255,
Borradaile, L. A., on the classification
of the decapod crustaceans, 457.
Boulenger, G. A., on three new
lizards and a new frog, 212 ; on
three new snakes from S. Cameroon,
324 ; on Barbus aureus. Cope, from
Natal, 390; on a new species of
I.<abeo from the Transvaal, 392 ;
on two new African lizards, 392 ;
on a new species of Telmatobius,
394 ; on new lizards in the British
Museum, 486 ; on the variations
of Stereolepis gigas, 489 ; on an
African barbel hitherto confounded
with Barbus tiimaculatus, 492.
Brachiopod nomenclature, notes on,
525.
Brachyphallus crenatus, notes on, 88.
Bradina, new species of, 3.
Brixia, new species of, 279.
Broom, Dr, R,, on new species of
Chrysochloris, 262.
Broun, Major T., on new coleoptera
from New Zealand, 56.
Biickman, S. S., on the genotype of
Terebratula, 525.
Bulimulus, new species of, 1 53.
Baodias, new species of, 180.
Burchell’s zebra, note on, 516.
Calamochrous, new species of, 55.
Callimorpha, new species of, 300.
Caiman, Dr. W. T., on a freshwater
crustacean from Para, 295.
Cameron, P., on new phytophagous
and parasitic hymenoptera, lOG.
Camilla, characters of the new
genus, 404.
Canis familiaris, on the black-and-
tan pattern of, 192.
Carcinops, new species of, 315.
Cardioderma, remarks on the genus,
140.
Caripodia, new species of, 232.
Cassolus, new species of, 416, 437.
Celama, new species of, 227.
Cephalophus, new species of, 385.
Ceryx, new species of, 221.
Charopa, new species of, 217.
Chilton, Prof, C., on Hymenosoma
depressum, 146 ; on a new fresh-
water gammarid from New Zea-
land, 388.
Chloridea, new species of, 242.
Chorisoneura, new species of, 44.
Choristima, synopsis of the species
of, 43.
Chrysochloris, new species of, 2G2.
Cirphis, new species of, 254.
Cixius, new species of, 284.
CHnteria, new species of, 352.
Clupeosoma, new species of, 19.
Cochlostyla, new species of, 150.
Cockerell, T. D. A., descriptions and
records of bees, 361, 531.
Coelorhyncidia, new species of, 6,
Coleoptera, new, 56, 125, 311, 347,
416.
Colomys, characters of the new
genus, 379.
Composita, note on the genus, 197.
Coptobasis, new species of, 5.
Corbula, new species of, 215.
Crocisa, new species of, 531.
Crossarchus, new species of, 120,
373.
Crustacea, new, 106, 297, 388 ; on
the classification of the decapod,
457.
Cryptus, new species of, 181.
Cultrarius, new species of, 176.
Cyclophorus, new species of, 155.
Cyclotus, new species of, 156.
Dactylamblyops, new species of,
113.
Dactvlervthrops, new species of,
109.
Dasychira, new species of, 203.
Dasypeltis, new species of, 324.
Daulia, new species of, 2.
Decapod Crustacea, on the classifi-
cation of the, 457.
Dejeania, new species of, 357.
INDEX.
545
Derbe, new species of, 395.
Derogenes, new species of, 90.
Desmonyx, characters of the new
genus, 355.
Deuterophysa, new species of, 7.
Devara, new species of, 302.
Diacrisia, new species of, 285.
Diathrausta, new species of, 6.
Digama, new species of, 50.
Dioptis, new species of, 308.
Diplocentrus, new species of, 210.
Dipsadoboa, new species of, 325.
Diptera, notes on, 326 ; new, 445.
Discognathus, new species of, 63.
Discoscelis, new species of, 318.
Distant, VV. L., rhynchotal notes,
277, 395.
Distomuni, notes on species of, 91,
102.
Dogs, on the black-and-tan pattern
of domestic, 192.
Dracsenura, new species of, 3.
Drona, new species of, 404.
Druce, H., on new species of hete-
rocera, 299.
Drynigeus, new species of, 153.
Dudusa, new species of, 204.
Eclipsiodes, new species of, 20.
Ectoneura, characters of the new
genus, 43.
Ellipsica, new species of, 48,
Ellipsidium, new species of, 28.
Eniscospilus, new species of, 178.
Ennea, new species of, 95.
Entozoa of British marine fishes, on
the, 66.
Episilia, new species of, 244.
Equus quagga, note on, 516.
Eratina, new species of, 310.
Erinaceus, new species of, 371.
Estigmene, new species of, 237.
Eublemina, new species of, 53.
Eucereon, new species of, 226.
Eucheira, new species of, 136.
Euchloropus, characters of the new
genus, 433.
Eucyane, new species of, 300.
Eulota, new species of, 164.
Euonyma, new species of, 101.
Euproctis Butleri, definition of the
new specific name, 203.
Eurota, new species of, 226.
Euryrhynchus, new species of, 297.
Euspilotus, characters of the new
genus, 320.
Eustegasta, new species of, 47.
Euthyrrapha, new species of, 46.
Ann, (k Mag, N, Hist, Ser.
Evania, new species of, 169.
Fantham, H. B., on Spirochteta Bah
bianii and on the occurrence of
Spirochseta anodontae in the
British mussel, 493.
Fenuahala, characters of the new
genus, 411.
Ficarasa, new species of, 287.
Fishes, new, 63, 64, 159, 163, 258,
261, 392, 493 ; on the entozoa of
British marine, 66.
Fulton, H. C., on new species of
Trochomorpha &c., 149 ; on Pa-
puina and Pupina, 158.
Funisciurus, new species of, 376.
Fruhstorferia, new species of, 354.
Gambusia, new species of, 259.
Gecko, new species of, 486.
Geological Society, proceedings of
the, 200, 450.
Glis, new species of, 445.
Glycosia, new species of, 351.
Gnathoncus, new species of, 321.
Goniopholis crassidens, on the skull
and skeleton of, 200.
Gude, G. K., on a new species of
Eulota, 164.
Halone, new species of, 232.
Hamba, characters of the new genus,
279.
Hampson, Sir G. F., on new Pyra-
lidae, 1 ; on new Syntomidae, Arc-
tiadae, Agaristidae, and N octuidae,
221. _
Helicarion, new species of, 99.
Helminthological nomenclature, on
modern, 451.
Hemiurus, remarks on the genus,
84.
Hesperapis, new species of, 535.
Heterorrhina, new species of, 347.
Hirst, A. S., on two new scorpions,
208 ; on a new species of Karschia
from Tibet, 322; on two new
species of Selenocosmia, 522.
Hister, new species of, 313.
Histeridae, new, 311.
Hololampra, new species of, 25.
Homoptera, new, 277, 395.
Hooley, R. W., on the skull and
skeleton of Goniopholis crassidens,
200.
Hymenoptera, new, 166, 268, 361,
531.
Hymenosoma depressum, note on,
146.
Hyperaeschra, new species of, 206.
. Vol, xix. 38
546
INDEX.
]IypsoideS; new species of, 202.
llema, new species of, 230.
Iphiaiilax, new species of, 170.
Ischnoptera, new species of, 26.
Isoceutris, new species of, 54.
Isolomalus, characters of the new
genus, 316.
Isometroides, note on the genus, 209.
Karschia, new species of, 322.
Kuranda, characters of the new
genus, 407.
Labeo, new species of, 392.
J^agarista, characters of the new
genus, 182.
Lasiogyia, characters of the new
genus, 7.
Latastia, new species of, 392.
Lauron, new species of, 308.
Lavia, note on the genus, l38.
Lebour, Miss M, V., on three mollusk-
infesting trematodes, 102.
Lecithaster gibbosus, note on, 89.
Lepidoptera, new, 1, 49, 20], 221,
299.
Lepodora rachiaea, notes on, 77.
lioptopoma, new species of, 155.
Lepus, new species of, 384.
Jjewis, G., on new Ilisteridae, 311.
Linstow, Dr. yon, on modern lielmin-
thological nomenclature, 451.
Lissophadnus, characters of the new
genus, 184.
I^ocha, new species of, 309.
Lophopteryx, new species of, 208.
Lophuromys, new species of, 382.
Ijuma, new species of, 2.
Jiycaon, new subspecies of, 375.
Lycophotia, new species of, 246.
Lygosoma, new species of, 488.
Mabuia, new species of, 212.
Macroderma, remarks on the genus,
137.
Macronota, new species of, 350.
Maenas, new species of, 235.
Magia, characters of the new genus,
288.
Mahuna, characters of the new
genus, 289.
Makula, characters of the new
genus, 408.
Mammals, new, 118, 133, 262, 370,
440, 445, 505, 520.
Margarochroma, new species of, 2.
Mawson, J., on the cretaceous forma-
tion of Bahia, 450.
Megaderma, new species of, 133.
lyiegadermatidae, remarks on the
family, 129,
Melcha, new species of, 179.
Mellivora, new species of, 376.
Melvill, J. C., on mollusca from S.
Africa, 94.
Metagylla, characters of the new
genus, 231.
Metalepsis, new species of, 244.
Metamblyops, characters of the new
genus, 106.
Metarctia, new species of, 225.
Microglossa, new species of, 20.
Microlomalus, characters of the new
genus, 318.
Miltochrista, new species of, 234.
Miselia, new species of, 252.
Mnemosyne, new species of, 283.
Mollusca, new, 94, 149, 164, 215, 217 ;
on three trematodes infesting, 102.
Mungos, new subspecies of, 119.
Muscidse, on the synonymy and
generic position of certain, 326 ;
on a new genus and species of,
445.
Myonycteris, note on the genus,
511.
Myopsyche, new species of, 222.
Mysidae, new Irish, 106.
Mysidia, new species of, 396.
Natalina, new species of, 98.
Neasura, new species of, 232.
Nelo, new species of, 305.
Neogenesis, characters of the new
genus, 19.
Nicoll, W., on the entozoa of British
marine fishes, 66 ; note of correc-
tion, 128.
Nola, new species of, 227.
Nomada, new species of, 537.
Notodonta, new species of, 206.
Notolampra, new species of, 38.
N ucras, new species of, 488.
Nyctibora, new species of, 36.
Ochodaeus, new species of, 358.
Ochotona aurita, note on, 124.
Oliarus, new species of, 282.
Onochalcis, new species of, 168.
Onthophagus, new species of, 418,
437.
Oopterus, new species of, 56,
Orthoptera, new, 25.
Osmia, new species of, 361, 538.
Ostodes, new species of, 220.
Ourebia, new species of, 387.
Ouwea, characters of the new genus,
291.
Oxyhaloa, new species of, 41.
Paheagrotis, characters of the new
genus, 252,
INDEX.
547
Palinura, definition of the tribal
name, 464.
Palpidia, new species of, 234.
Panelus, new species of, 417.
Papuina, note on the genus, 158.
l^aracrabro, characters of the new
genus, 274.
Paralaethia, characters of the new
genus, 224.
Paraleucogobio, new species of, 163.
Paranauphoeta, new species of, 46.
Paraplecta, definition of the new
generic name, 41.
Paricana, new species of, 288.
Parorchis, definition of the new
generic name, 128.
Pedilophorus, new species of, 68.
Perdita, new species of, 539.
Pericallia, new species of, 238.
Pericopis, new species of, 300.
Phseochroops, new species of, 359, 436.
Phanoptis, new species of, 309.
Phasis, new species of, 100.
Phenice, new species of, 397.
Phloeonseus, new species of, 57.
Phoxinopsis, characters of the new
genus, 262.
Phreatogammarus, new species of,
388.
Phyllodactylus, new species of, 212.
Phyllodromia, new species of, 30.
Piletocera, new species of, 12.
Pimelodus, new species of, 258.
Piroblatta, characters of the new
genus, 29.
Placostylus, new species of, 154.
Pocock, K. I., on the black-and-tan
pattern of domestic dogs, 192 ; On
the quagga and Burchell’s zebra,
516.
Podocotyle atomon, notes on, 73.
Pogonocharax, characters of the new
genus, 261.
Poiana, new subspecies of, 372.
Polemistus, new species of, 274.
Ponsonby, J. II., on Mollusca from
S. Africa, 94.
Preston, II. B., on new species of
Corbula and Bithinella, 215 ; on
new land-shells from New Cale-
donia, 217.
Prionodactylus, new species of, 486.
Procayia, new species of, 520.
Protsetia, new species of, 351.
Psen, new species of, 273.
Pseudaclytia, new species of, 226.
Pseudapiconoma, new species of,
225.
Pseudectobia, new species of, 35.
Pseudoglomeris, new species of, 48.
Pseudolithosia, characters of the new
genus, 1.
Pterocyon, new species of, 505.
Pupina, note on species of, 158.
Pydna, new species of, 206.
Pyralidae, new, 1.
Pyrausta, new species of, 55.
Quagga, notes on the, 516.
Rana, new species of, 214.
Regan, C. Tate, on new fishes from
Yunnan, 63; on new Central-*
American freshwater fishes, 64;
on new freshwater fishes from
Mexico, 258; on new Characinid
fishes from Argentina, 261.
Reptiles, new, 212, 324, 392, 486.
Rhodeinse, synopsis of the, 160.
Rhodogastria, new species of, 240.
Rhotala, new species of, 294.
Rhotana, new species of, 409.
Rhynchocyon, new species of, 370.
Rhynchotal notes, 277, 395.
Rivulus, new species of, 64.
Roeselia, new species of, 229.
Rousettus, new species of, 509; re-
marks on species of, 512.
Sangala, new species of, 307.
Schizothorax, new species of, 63.
Sciiirus, new species of, 120.
Scoparia, new species of, 21.
Scorpions, new, 208, 322.
Scythrodes, new species of, 59.
Selandria, new species of, 167.
Selenocosmia, new species of, 522.
Seminula, on, 194.
Sericophorus, neW species of, 275.
Shelford, R., on new species of
Blattidae, 25.
Sicydium, new species of, 260.
Sikaiana, characters of the new
genus, 398.
Silsila, new species of, 179.
Siltirus, new species of, 64.
Siobla, new species of, 167.
Sphegidae, new Australian, 268.
Spirifer ambiguus, note on, 197.
Spirochasta anodontae, on the occur-
rence of, in the British mussel, '
493.
Balbianii, note on, 493.
Stenia, new species of, 8.
Stenogyra, new species of, 154.
Stephanochasmus, new species of,
80.
Stereolepis gigas, on the variations
of, 489.
548
INDEX.
Styo-eromyia, characters of the new
genus, 445.
Swinhoe, Col. C.,on Eastern, Austra-
lian, and African heterocera, 49,
201.
Sylvisorex, new species of, 118.
Syntomis, new species of, 201, 222.
Tachina, notes on species of, 327.
Talaloa, characters of the new
genus, 294.
Taloka, characters of the new
genus, 280.
Tarache, new species of, 53.
Taraxineura, new species of, 308.
Tarsolepis, new species of, 204.
Tatera, new species of, 378.
Tattersall, W. M., on new Irish
Mysidae, 106.
Telmatobius, new species of, 394.
Tenthredo, new species of, 166.
Terebratula, on the genotype of, 525.
pentaedra, note on, 195.
Thamnomys, characters of the new
genus, 121 ; new species of, 381.
Thomas, O., on new mammals from
Euwenzori, 118 ; on new mammals
from Lake Chad and the Congo,
370, 520 ; on a viscacha from
S. Patagonia, 439 ; on the occur-
rence of the edible dormouse in
Sardinia, 444.
Thryonomys, new species of, 384.
Tigrioides, new species of, 230.
Timora, new species of, 243.
Tityus Pococki, definition of the new
specific name, 210.
Torynorrhina, characters of the new
genus, 433,
Trachycystis, new species of, 99.
Trematodes, on three mollusk-in^
■ festing, 102.
Tricholepis, new species of, 233.
Trichoridia, new species of, 257.
Trochomorpha, new species of, 149.
Tudea, characters of the new genus,
290.
Tuerta, new species of, 241.
Turner, R. E., on new Australian
Sphegidae, 268.
Turuptiana, new species of, 238.
Ufeus, new species of, 249.
Urabunna, characters of the new
genus, 414.
Utethesia, new species of, 202, 230.
Vaughan, Dr. A., on Seminula,
194.
Viscaccia, new species of, 440.
Vivaha, new species of, 405.
Woodward, Dr. A. Smith, 'on the
cretaceous formation of Bahia,
450 ; on a new dinosaurian reptile
from Lossiemouth, 451.
Wroughton, R. C., on the bats of
the family Megadermatidae, 129 ;
on new mammals from Lake Chad
and the Congo, 370, 520.
Xanthojoppa, new species of, 186.
Xanthospilopteryx Hornimani, new
subspecies of, 241.
Xiphophorus, new species of, 65.
Zetobora, new species of, 45.
Zia, new species of, 229.
Zoogonoides viviparus, notes on,
83.
Zoraida, new species of, 400.
END OF THE NINETEENTH VOLUME.
PRINTED BY TAYLOR AND FRANCIS,
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191907
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