European Journal of Taxonomy 75: 1-123

http://dx.doi.org/10.5852/ejt.2014.75

www. europeanj ournaloftaxonomy. eu

2014 • David G. Notton

ISSN 2118-9773

This work is licensed under a Creative Commons Attribution 3.0 License.

Monograph

um:lsid:zoobank.org:pub:E02C736E-661D-407F-9322-24BFB95E233E

A catalogue of the types of Diapriinae (Hymenoptera, Diapriidae)

at the Natural History Museum, London

David G. NOTTON

Department of Entomology, the Natural History Museum, Cromwell Road,

London, SW7 5BD, United Kingdom

[email protected]

um:lsid:zoobank.org:author:2E452EF2-703C-47C0-8432-6AB9C05AC06A

Abstract. The types of nominal species of Diapriinae in the collection of the Natural History Museum,

London, are catalogued. Lectotypes are designated for the following taxa: Diapria peraffinis Ashmead,

1896; D. smith ii Ashmead, 1896; Galesus bipunctatus Ashmead, 1894; G. (G.) foersteri var. nigricornis

Kieffer, 1911; G. sexpunctatus Ashmead, 1893; G. walkeri Kieffer, 1907; Idiotypa nigriceps Kieffer, 1909;

I. nigriceps Kieffer, 1911; I. pallida Ashmead, 1893; I. pallida Ashmead in Riley, Ashmead & Howard,

1894; Paramesius angustipennis Kieffer, 1911; P. cameroni Kieffer, 1911; Phaenopria cameroni Kieffer,

1911; P. halterata Kieffer, 1911; P. magniclavata Ashmead, 1896; Tropidopsis clavata Ashmead, 1893;

T. clavata Ashmead in Riley, Ashmead & Howard, 1894. New combinations are proposed: Aneuropria

bifurcata comb. nov. for Mantara bifurcata Dodd, 1920; Basalys quadridens comb. nov. for Microgalesus

quadridens Kieffer, 1912; Coptera cratocerus comb. nov. for Galesus cratocerus Cameron, 1912; Coptera

sexpunctata comb. nov. for Galesus sexpunctatus Ashmead, 1893; Doliopria magniclavata comb. nov. for

Phaenopria magniclavata Ashmead, 1896; Spilomicrus aterrimus comb. nov. for HoplopriaaterrimaDodd,

1920; Spilomicrus campbellanus comb. nov. for Antarctopria campbellana Yoshimoto, 1964; Spilomicrus

coelopae comb. nov. for Antarctopria coelopae Early, 1978; Spilomicrus diomedeae comb. nov. for

Antarctopria diomedeae Early, 1978; Spilomicrus helosciomyzae comb. nov. for Malvina helosciomyzae

Early & Horning, 1978; Spilomicrus insulae comb. nov. for Malvina insulae Early, 1980; Spilomicrus

latigaster comb. nov. for Antarctopria latigaster Braes in Tillyard, 1920; Spilomicrus punctatus comb. nov.

for Malvina punctata Cameron, 1889; Spilomicrus rekohua comb. nov. for Antarctopria rekohua Early,

1978; Trichopria bouceki comb. nov. for Oxypria bouceki Masner, 1959; Trichopria nigriceps comb. nov.

for Tropidopria nigriceps Ashmead in Riley, Ashmead & Howard, 1894; Trichopria nigriceps comb. nov.

for Xyalopria nigriceps Kieffer, 1907; Trichopria spinosleeps comb. nov. for Acidopria spinosiceps Dodd,

1920; Trichopria walkeri comb. nov. for Diapria walkeri Dalla Torre, 1890. New replacement names are

proposed: Coptera mosselensis nom. nov. for G. nigricornis Nixon, 1930 preocc.; Coptera pijiguaorum

nom. nov. for G. sexpunctata Montilla & Garcia, 2008 preocc.; Spilomicrus kozlovi nom. nov. for S.

punctatus Kozlov, 1978 preocc.; Trichopria fluminis nom. nov. for T. nigriceps (Kieffer, 1907) preocc.;

T. thermarum nom. nov. for T. nigriceps (Kieffer, 1913) preocc. New specific synonyms are proposed:

Basalys cursitans (Kieffer, 1911) = B. pedisequa (Kieffer, 1911) syn. nov. (the former removed from

synonymy with B. parvus Thomson, 1858); B. iphicla Nixon, 1980 = B. macroptera (Kieffer, 1911) syn.

nov.; Coptera bipunctata (Ashmead in Riley, Ashmead & Howard, 1894) = G. sexpunctata (Ashmead,

1893) syn. nov.; Idiotypa nigriceps Kieffer, 1911 =/. nigriceps Kieffer, 1909 syn. nov.;/. pallida Ashmead

in Riley, Ashmead & Howard, 1894 = I. pallida Ashmead, 1893 syn. nov.; Psilus nigricornis (Kieffer,

European Journal of Taxonomy 75: 1-123 (2014)

1911) = 72 fuscipennis (Curtis, 1831) syn. nov.; P. walkeri (Kieffer, 1907) = P. fuscipennis (Curtis, 1831)

syn. nov.; T. boucehi (Masner, 1959) = T. conotoma (Kieffer, 1911) syn. nov.; Trichopriahalterata (Kieffer,

1911) = 77 halter ata (Kieffer, 1909) syn. nov. New generic synonyms are proposed: Antarctopria Braes in

Tillyard, 1920 = Spilomicrus Westwood, 1832 syn. nov.; Malvina Cameron, 1889 = Spilomicrus Westwood,

1832 syn. nov.; Mantara Dodd, 1920 = Aneuropria Kieffer, 1905 syn. nov.; Microgalesus Kieffer, 1912

= Basalys Westwood, 1833 syn. nov.; Xyalopria Kieffer, 1907 = Trichopria Ashmead, 1893 syn. nov.

(Xyalopria is removed from synonymy with Megaplastopria Ashmead, 1903). A brief account of some

aspects of the history of these types is given.

Keywords. Hymenoptera, Diapriidae, Diapriinae, types, The Natural History Museum (London).

Notton D.G. 2014. A catalogue of the types of Diapriinae (Hymenoptera, Diapriidae) at the Natural History

Museum, London. European Journal of Taxonomy 75: 1-123. http://dx.doi.org/10.5852/eit.2014.75

Introduction

Diapriinae Haliday, 1833 is a cosmopolitan subfamily of diapriid wasps including about 1000 described

species (Johnson 1992). Most are pupal or puparial endoparasitoids of Diptera or more rarely Coleoptera

or Formicidae. Diapriinae are often a major component of the microhymenopteran fauna attacking

Diptera in a range of habitats, but despite this they remain poorly known. The location and recognition

of type specimens and their placement in currently recognised taxa is a necessary preliminary step to

revisionary studies which, in order that they have lasting value, must be based on knowledge of species

described already. This problem is particularly severe for neglected families such as the Diapriidae

where there is a large and fragmented literature, with the older descriptions not always mentioning type

depositories. There are a number of publications which refer to types in the Natural History Museum,

London (BMNH), most usefully Masner (1965), which mentions types of 135 nominal species of

Diapriinae, and Johnson’s (1992) World catalogue, which cites publications mentioning types. A number

of types have been added to the BMNH collection since 1965 thanks to the generosity of Lubomir

Masner and other colleagues, and some types have been discovered which were previously overlooked.

The current study aims to catalogue all types of Diapriinae in London in the context of the history of

those types, modern assessments of type status and current ideas on classification of Diapriinae, in

order to make them accessible to future workers as a basis for future revisions of World Diapriinae. The

types of more than 250 nominal species are treated, a significant proportion of the World fauna when

compared to the circa 1200 valid species listed in the last World catalogue (Johnson 1992). The concept

of Diapriinae followed here is that of Masner & Garcia (2002) and Notton (2004), including all the

genera traditionally placed in Psilini and some genera for which tribal placement is problematic. A brief

history of some of the people associated with these types is given here.

William Harris Ashmead (1855-1908)

William Ashmead, was Assistant Curator, Division of Insects at the United States National Museum,

(Smithsonian Institution, Washington D.C.) and one of the foremost Am erican workers of his t im e

in the systematics of Hymenoptera (Anon. 1908; 1909; Bethune 1908, 1909; Crawford 1909; Horn

1909; Howard 1908; Howard, Crawford & Banks 1909; Semenov-Tian-Shanskii 1909; Smith 1910).

Types of Ashmead’s species of Diapriinae found in the BMNH are from material collected by Herbert

Huntingdon Smith (1852-1919) in the West Indies as part of a project coordinated by the West India

Exploration Co mm ittee The BMNH Entomology Registers include donations of numerous insects of

all kinds from the Committee and its agents; the record for lot 1899-331 shows the Committee presented

2205 Hymenoptera from St. Vincent and Grenada, including 610 species and 347 types. New taxa of

Hymenoptera were described in a series of papers, mainly by Ashmead (Ashmead 1893, 1896, 1900;

Howard 1897; Riley, Ashmead & Howard 1894). It is likely that the intention of the Committee was to

NOTTON D.G., Diapriinae types at The Natural History Museum, London

deposit the majority of the specimens at the BMNH since the Committee was a British endeavour set

up by the Royal Society and the British Association, and St Vincent and Grenada were longstanding

British colonies at the time. Published reports on the Parasitica do not mention type repositories, except

for Ashmead (1893), which generally has series split between London and Washington. On the basis

of what has been found in the BMNH collection, unique specimens and most important individuals are

generally in the BMNH, and any remaining specimens appear to have been split between London and

Washington in recognition of Ashmead’s effort in working up the material, although only a few were

recognised by Masner & Muesebeck (1968).

Ashmead did not designate holotypes, so type series are syntypic. Where there are series, Ashmead’s

type labels cannot be taken as evidence that the specimens are holotypes, as he did not consistently label

types or add determination labels. Consequently, all available undetermined H.H. Smith material has

been assessed for type status, and it has become apparent that much previously unlabelled material in

the BMNH is syntypic. Unfortunately, much H.H. Smith material was mounted on short bendy pins and

a number of specimens have been damaged in the past as a result. Many of these were repinned during

the current project so they could be examined safely. Lectotype designations have been based on BMNH

specimens by previous authors, and it seems pragmatic to continue this, insofar as BMNH specimens

are taxonomically suitable.

Peter Cameron (1847-1912)

Peter Cameron was an energetic amateur student of Hymenoptera, describing many species despite

meagre means and poor health, and donating much of his type material to the BMNH ( Anon. 1913;

GMW 1913; Morice 1913; Morley 1913). Cameron is an important figure not just because of the species

of Diapriinae he described himself, but also because he loaned material to Kieffer (q.v. below) to

describe, mainly from Scotland and England. In his own descriptions, Cameron did not always designate

holotypes or give a clear indication of the numbers of specimens on which he based his descriptions

(Notton, Buffington & Van Noort 2009; Quinlan 1974); in these cases the specimen(s) have been treated

as syntypes (ICZN 1999: Reco mm endation 73F) even when there is only one in the BMNH, since

Cameron is known to have dispersed syntype series.

Abbe Jean-Jacques Kieffer (1856-1925)

Jean-Jacques Kieffer was a Catholic priest and teacher at a secondary school in Bitche in Lorraine,

Northern France. In his spare time he published hundreds of taxonomic papers on parasitic Hymenoptera

and gall midges (Cecidomyiidae) (Nomine 1925, 1926; Kelner-Pillault 1958; Gagne 1994; Notton

2004; Vlug 1995). As Kieffer described many of the species covered in the catalogue below, it is worth

mentioning the history of his types. Kieffer based his descriptions on specimens in his own collection, but

also on material borrowed from other collectors and museums. Much of Kieffer’s personal collection, at

least the Diapriidae, has survived at the Museum national d’Histoire naturelle, Paris, France, despite the

doubt cast on the survival of Kieffer material belonging to some other taxa (Gagne 1994 and references

therein; Vlug 1995). In brief, P.F.G. Benoit, Head of the Invertebrates Section at the Musee du Congo

Beige at Tervuren, found Kieffer’s collection at the College de Bitche where Kieffer taught. Thanks to the

Recteur, R.P.P.J. Schmitt, the collection was transferred to the Entomology Faboratory of the Museum

in 1957 (Kelner-Pillault 1958). Otherwise, types which Kieffer described from material received from

collectors or museums were usually returned to them and so their current location depends on the fate of

their collection of origin. Thus, many types can be traced where Kieffer states the collector in original

descriptions. For example, some of those based on du Buysson and de Gaulle material are now in Paris,

whereas those based on Cameron specimens are in Fondon (Notton 1995, 2004 and this study). Kieffer

also exchanged specimens of myrmecophile Hymenoptera with E. Wasmann; hence, some syntype

series of Wasmann and Kieffer myrmecophile diapriids are split between Paris and Maastricht (Dessart

1975; Notton 2004).

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Many of Kieffer’s diapriid types in BMNH were described from Scottish and English specimens from the

collection of Peter Cameron, q.v. above. Cameron posted the specimens to Kieffer, who numbered and

described or identified them and sent them back. Rather than adding determination labels, Kieffer wrote

lists of specimen numbers and identifications on postcards and sent them back to Cameron separately.

The specimens and postcards were given to the BMNH by Cameron, with the addition of a list written

by him of the species represented, and the specimens were registered as lots 1910-55 and 1910-302.

The specimens can be recognised today by Kieffer’s number labels and one of the two register numbers,

and they may also have labels in Cameron’s handwriting or identification labels added later by BMNH

curators based on Kieffer’s lists. The postcards sent by Kieffer, together with secondary lists based

on them prepared by Cameron and BMNH curators, are now held by the BMNH Entomology library

(Entomology Accession Registers and Insect Room Lists: 29, 32, 82, 83, 84). All specimens which could

be recognised from these lists were reassessed for their type status, giving most weight to Kieffer’s

original number labels and postcard lists.

Cameron had very poor handwriting, did not always label his specimens, and never labelled them with

the country which created problems, both for him and for Kieffer, when it came to reporting localities

correctly in publications. The documentary evidence of individually numbered specimens and Kieffer’s

numbered lists make it certain that Kieffer and Cameron were working from the same material, and

yet there are many differences between the localities reported by Cameron (1910), the type localities

given by Kieffer for species described from Cameron’s material and the actual localities written on

the specimens concerned, where it has been possible to decipher them. It appears that Kieffer assumed

that most of Cameron’s specimens came from Scotland when in fact many really came from England.

Consequently, a number of type localities have been revised in the catalogue below where it is evident

that the data on the specimens does not match the published type locality. Even without the problems

of Cameron’s labels, it appears Kieffer had a poor grasp of geography because the postcards he wrote

to Cameron at Whitle, New Mills, near Stockport in England are addressed to Scotland. A further

complication for Kieffer’s names is that Cameron (1910) published many as nominanuda before Kieffer

made them available. Similarly, it appears that Kieffer assumed that Walker material of Galesus walkeri

was from England when it was actually from Norway.

Another major group of Kieffer types is based on material collected by the Percy Sladen Trust expedition

to the Seychelles (Kieffer 1912a; Huggert 1979) and is split between the BMNH and the Zoology

Museum, University of Cambridge, Cambridge, UK. The existence of the Cambridge specimens only

became apparent at a late stage in the preparation of this work, and unfortunately there was not time

to include them in the current work. On the basis of dating evidence presented by Evenhuis (1994),

Kieffer’s report on material from the Sladen expedition in the Transactions of the Linnean Society

(Kieffer 1912a) was published before his Genera Insectorum v. 124 (Kieffer 1912b). This conclusion is

supported by the fact that Kieffer quoted page references from the Transactions in Genera Insectorum.

Lastly, another problem with Kieffer’s names is that he sometimes described the same species twice by

mistake. In these cases, to avoid future nomenclatural problems lectotype designations are used here to

ensure that one specimen is the primary type for both nominal species, thus creating objective synonymy.

Material and methods: notes on the recognition of types and the arrangement and

format of the catalogue

The status of nominal species represented by type material was considered and details of type material

given. Within the catalogue, original combinations are arranged alphabetically by author, then in

order of publication date and page number. Full label data are quoted for primary types, except where

illegible ([illeg.]), slashes are used to indicate the end of a line (/), where a slash occurs in the label

data this has been replaced by a dash (-), a semicolon is used between labels (;) and a full stop between

NOTTON D.G., Diapriinae types at The Natural History Museum, London

mounts (.). Specimen condition is noted where this may help in the recognition of primary types, as well

as comparative notes on original descriptions.

Each specimen was assessed for type status. For difficult cases, the criteria of Notton (2004) have been

followed when recognising syntypes. Earlier authors did not always designate holotypes or give a clear

indication of the numbers of specimens on which they based descriptions; in these cases the specimen(s)

have been treated as syntypes (ICZN 1999: Recommendation 73F) even when there is only one in the

BMNH, since types series have often been dispersed. Inevitably, some specimens were found to bear

labels with incorrect type status. A degree of latitude was necessary when matching specimens with the

descriptions ofKieffer and Ashmead, which are sometimes inaccurate.

The species described by Borgmeier (1939) from type series (i.e. two or more specimens) are all

considered to be based on syntypes, even where these were labelled “paratype” or “cotype”, because he

did not designate a holotype in the original publication, instead referring to all type specimens equally

as “typos” (types). Of course, the mention of the word “type” or equivalent expression on a label is not

necessarily evidence that the specimen is fixed as a holotype (Article 72.4.7). Similarly, the species

described by Nixon (1930) from type series are all considered to be based on syntypes even where these

were labelled by Nixon as “type” or “paratype”, because he did not designate a holotype in the original

publication. In cases where type series included both sexes, Nixon labelled one specimen of each sex

as “type”, so in this particular work he clearly did not use the word type consistently to mean holotype.

The identity of each nominal species is given with species placed in currently recognised genera, using

the generic keys of Nixon (1980), Masner & Garcia (2002) and various other sources cited in Notton

(2004). Antennal segments are referred to as follows: Antennal segment 1 (scape) = al; segment 2

(pedicel) = a2; segment 3 (flagellar segment 1) = a3; and so on. Article numbers cited refer to the

International Code of Zoological Nomenclature (ICZN 1999).

Lastly, a nomenclatural summary is given. Nominal taxa mentioned in the catalogue are listed. More

complete synonymies will be found in Johnson (1992) and other published sources. The tribe Psilini

is available from Fallen (1812) and although it predates Diapriidae Haliday, 1833, Diapriidae retains

priority (Article 35.5) which, in the interest of stability, gives precedence to names in use at a higher

rank before 1999. Muesebeck & Walkley (1956) and Johnson (1992) have been consulted to confirm

the type species of genera involved in new generic synonymy. Diapriid genus names are often based on

the ending -pria, of which the first was Diapria Latreille, 1796. The derivation of this name is from the

Greek SioutpiG), to saw in two, a reference to the incision at the base of the large tergite. This name is

based on a Greek word Latinized with a change of ending to the feminine Latin gender ending - a , and

so takes the feminine gender (ICZN 1999: Art. 30.1.3). Later names ending in -pria are also based on

7ipicov, to saw, by analogy with Diapria and similarly have feminine gender. Basalys Westwood, 1833

is derived from Greek (luoiq (foot) + oAuq (listlessness); an appropriate name for a wasp which will not

sit still. Since aZrx; is masculine this genus has masculine gender (M.A.Alonso-Zarazaga pers. comm.;

ICZN 1999: Article 30.1.2). Westwood was somewhat of a classical linguist and commonly used Greek

words when composing the names of genera, so while the species of this genus usually have a distinct

basal vein, linguistically Basalys is not connected with the Latin word basalis (basal) except possibly that

Westwood may have intended it as a homophonic pun. The Latin epithets pedisequus (variant spelling

pedissequus ) a male foot servant or lackey and pedisequa (or pedissequa) a waiting-lady are nouns, and

so do not decline. All names considered here which were originally described as varieties are deemed

to be available at subspecific rank from their date of publication (Article 45.6.4). All taxonomic acts are

attributable to the author except for the new name Coptera pijiguaorum nom. nov. which is attributable

to Notton, Montilla and Garcia.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Results

Type Catalogue

Tropidopsis clavata Ashmead, 1893: 402, pi. xvii, figs 2, 2a.

Valid name

Basalys clavatus (Ashmead, 1893).

Summary of types

Lectotype §, BMNH number 9.916. Lectotype here designated.

Primary type data

St. Vincent/ W.I./ H.H. Smith/ 230; Tropidopsis/ clavatus/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

This species was described from an unspecified number of females and males. Although Ashmead

states that the types are in the Smithsonian Institution, Washington D.C., only one male syntype is there

(Masner & Muesebeck 1968) and a female syntype is in the BMNH. Another specimen in the BMNH

was labelled by Ashmead as “ var. ” and is much darker than the description and so is not considered part

of the type series. Since the male syntype is a Trichopria Ashmead, 1893 and the female syntype is a

Basalys , the female is designated here as lectotype in order to stabilise the current generic placement

established by Masner and Muesebeck. The lectotype is mounted on a card point, with the right antenna

lost and left antenna lost beyond a8.

Idiotypa pallida Ashmead, 1893: 403, 463, pi. 17, fig. 3.

Valid name

Idiotypa pallida Ashmead, 1893.

Summary of types

Lectotype $, BMNH number 9.1004. Lectotype here designated. Paralectotype f.

Primary type data

St. Vincent/W.I./H.H. Smith/ 158; Idiotypa/ pallida/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

This species is named at the end of the generic description of Idiotypa with a reference to the figure of

the female; the figure serves as an indication to make this species name available (Article 12.2.7). The

explanation of the plates (Ashmead 1893: 463) shows that Ashmead saw both sexes and it is reasonable

to suppose that the syntype series is the same as for Idiotypa pallida Ashmead in Riley, Ashmead &

Howard, 1894. The same specimen is designated as the lectotype for both/, pallida Ashmead, 1893 and

NOTTON D.G., Diapriinae types at The Natural History Museum, London

I. pallida Ashmead in Riley, Ashmead & Howard, 1894 q.v. to ensure objective synonymy. The lectotype

is mounted on a card point, is missing part of the right hind leg, and has the wings stuck together.

Galesus sexpunctatus Ashmead, 1893: 408, 463, pi. 17, fig. 6.

Valid name

Coptera sexpunctata (Ashmead, 1893) comb. nov.

Summary of types

Lectotype $, BMNH number 9.1003, here designated. Paralectotype S-

Primary type data

St. Vincent/ W.I./ H.H. Smith; Galesus/ bipunctatus/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

This species is named at the end of the generic description of Galesus Haliday in Curtis 1829 with a

reference to the figure of the female and male; the figure serves as an indication to make this species

name available (Article 12.2.7). Although Ashmead (1893) says this species was from St. Vincent, in his

study of St. Vincent diapriids the following year, the only species of Galesus mentioned is G. bipunctatus

Ashmead in Riley, Ashmead & Howard, 1894. It is reasonable to assume that this is one and the same

species redescribed under another name because: a) both species were described from St. Vincent from

type series including both sexes; b) both species are referred to as having six punctures; G. sexpunctatus

in the specific epithet and G. bipunctatus in its original description which includes the phrase “vertex

with six small punctures”; c) an examination of Ashmead’s labels shows how the change of names

could have happened as 6-punctatus and bipunctatus look quite similar in Ashmead’s handwriting.

Minor differences between the figures of G. sexpunctatus in Ashmead (1893) and the description of G.

bipunctatus Ashmead in Riley, Ashmead & Howard (1894) may be discounted because the figures in

Ashmead (1893) are often inaccurate in details. It is reasonable therefore to regard the syntype series of

G. sexpunctatus as the same as that of G. bipunctatus. The original description of G. bipunctatus was

based on two females and two males, of which a pair was found in the BMNH. The same specimen is

designated lectotype for both G. sexpunctatus Ashmead, 1893 and G. bipunctatus Ashmead in Riley,

Ashmead & Howard, 1894 q.v. to ensure objective synonymy. The lectotype is glued to a micropin

and is entire. G. sexpunctatus belongs in the genus Coptera. Coptera sexpunctata Montilla & Garcia,

2008 is now a secondary junior homonym and the new replacement name Coptera pijiguaorum Notton,

Montilla & Garcia nom. nov. is proposed. Pijiguaorum is the genitive plural of a Latinized form of the

name of the Pijiguaos people of Venezuala.

Idiotypa pallida Ashmead in Riley, Ashmead & Howard, 1894: 243.

Valid name

Idiotypa pallida Ashmead, 1893 syn. nov.

Summary of types

Lectotype §, BMNH number 9.1004, here designated. Paralectotype S-

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Primary type data

St. Vincent/ W.I./ H.H. Smith/ 158; Idiotypa/ pallida/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

Although already described under the same name in 1893, Ashmead redescribed this species as new in

1894, from two females and one male, of which a pair was found. It is reasonable to assume that the

syntype series is the same for both I. pallida Ashmead, 1893 and I. pallida Ashmead in Riley, Ashmead

& Howard, 1894, and the same specimen is designated lectotype for both names to ensure objective

synonymy. The lectotype is mounted on a card point, and is missing part of the right hind leg, and has

the wings stuck together.

Hemilexis latipennis Ashmead in Riley, Ashmead & Howard, 1894: 244.

Valid name

Entomacis latipennis (Ashmead in Riley, Ashmead & Howard, 1894).

Summary of types

Holotype S, BMNH number 9.687, by monotypy.

Primary type data

1500 feet; St. Vincent/W.I./H.H. Smith; Type/H.T.; Hemilexis/latipennis/ 3 Type Ashm./ unique; type

lost and/ and absolutely no fragments in/ the drawer/N.D.M.F. 23-iii-1977.

Type locality

West Indies, St Vincent.

Remarks

Only the mount and labels of the holotype remain. It has been missing since before 1965 (Masner 1965).

On the basis of Ashmead’s description there is no doubt that this species belongs to the genus Entomacis.

Hemilexodes filiformis Ashmead in Riley, Ashmead & Howard, 1894: 244.

Valid name

Entomacis filiformis (Ashmead, 1984).

Summary of types

Holotype S, BMNH number 9.917, by monotypy.

Primary type data

Leeward side/ St. Vincent, W.I./H.H. Smith/ 201; W.Indies/ 99-331; Entomacis m/ filiformis Ash.

Type locality

West Indies, St Vincent.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

The holotype is mounted on a card point with the left antenna missing beyond scape.

Tropidopsis clavata Ashmead in Riley, Ashmead & Howard, 1894: 245.

Valid name

Basalys clavatus (Ashmead, 1893).

Summary of types

Lectotype $, BMNH number 9.916, here designated.

Primary type data

St. Vincent/ W.I./ H.H. Smith/ 230; Tropidopsis/ clavatus/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

This species was redescribed in error from the same type series as Tropidopsis clavata Ashmead, 1893

q.v. The lectotype of T. clavata Ashmead, 1893 q.v. is also designated as lectotype of T. clavata Ashmead

in Riley, Ashmead & Howard, 1894 to create objective synonymy. Hence, the condition of the lectotype

and generic placement of T. clavata Ashmead, 1984 are the same as for T. clavata Ashmead, 1893.

Paramesius thoracicus Ashmead in Riley, Ashmead & Howard, 1894: 245.

Valid name

Paramesius thoracicus Ashmead in Riley, Ashmead & Howard, 1894.

Summary of types

Syntypes 3 $$, 2 SS-

Primary type data

Leeward side/ St. Vincent, W.I./ H.H. Smith/ 201; Paramesius/ thoracicus/ $ Ashm.; W.Indies/ 99-

331 (syntype $). St. Vincent/W.I./H.H. Smith/ 198; W.Indies/ 99-331 (syntype §). Leeward side/ St.

Vincent, W.I./H.H. Smith/201; W.Indies/ 99-331 (syntype $). 1500 feet; St. Vincent,/W.I./H.H. Smith;

Paramesius/ thoracicus/ S Ashm.; W.Indies/ 99-331 (syntype (?). 1000 feet; St. Vincent,/ W.I./ H.H.

Smith; W.Indies/ 99-331 (syntype S).

Type locality

West Indies, St Vincent.

Remarks

This species was described from four females and four males, of which three females and two males

were found. Another female specimen labelled by Ashmead as this species is from Grenada, so is not

syntypic. The three female syntypes are mounted on card points, the one with Ashmead’s determination

label is entire, the one numbered 198 is missing its head, the third is missing its head and right fore leg.

The two male syntypes are glued to the points of micro-pins, both are entire.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Spilomicrus aneurus Ashmead in Riley, Ashmead & Howard, 1894: 246.

Valid name

Spilomicrus aneurus Ashmead in Riley, Ashmead & Howard, 1894.

Summary of types

Syntypes $, 3

Primary type data

St. Vincent/ W.I./ H.H. Smith; Spilomicrus/ aneurus/ $ Ashm.; W.Indies/ 99-331 (syntype §). St.

Vincent/ W.I./ H.H. Smith/ 237; Spilomicrus/ aneurus/ $ Ashm.; W.Indies/ 99-331 (syntype $). Sea

level; St. Vincent/ W.I./ H.H. Smith; W.Indies/ 99-331 (syntype $). 1000 feet; St. Vincent/ W.I./ H.H.

Smith; W.Indies/ 99-331 (syntype $).

Type locality

West Indies, St Vincent.

Remarks

This species was described from a female and five males, of which one female and three males were

found. The female syntype is glued to a micropin and is entire; the male with Ashmead’s determination

label is mounted on a card point and is entire; the male labelled “sea level” is glued to a micropin and is

missing the ends of both antennae; the male labelled “1000 feet” is micropinned and is entire.

Spilomicrus vulgaris Ashmead in Riley, Ashmead & Howard, 1894: 246, 247.

Valid name

Spilomicrus vulgar is Ashmead in Riley, Ashmead & Howard, 1894.

Summary of types

Syntypes 11 $$, 16 SS-

Primary type data

Mountain forest. 3000ft/ in rotting leaves, March; St. Vincent/ W.I./ H.H. Smith; Spilomicrus/ vulgaris/

9 Ashm.; W.Indies/99-331 (syntype $). 2000 feet; St. Vincent/W.I./H.H. Smith; Spilomicrus/ vulgaris/

9 Ashm.; W.Indies/ 99-331 (syntype §). 2000 ft; St. Vincent/ W.I./ H.H. Smith; W.Indies/ 99-331

(syntypes 4§§). St. Vincent/ W.I./ H.H. Smith; W.Indies/ 99-331 (syntypes 2$$, 5fS). St. Vincent/

W.I./ H.H. Smith/ 158; W.Indies/ 99-331 (syntype $). St. Vincent/ W.I./ H.H. Smith/ 227; W.Indies/

99-331 (syntype $). St. Vincent/ W.I./ H.H. Smith/ 244; W.Indies/ 99-331 (syntype §). Leeward side/

St. Vincent,/ W.I./ H.H. Smith/ 45; Spilomicrus/ vulgaris/ S Ashm.; W.Indies/ 99-331 (syntype S).

1500 feet; St. Vincent/ W.I./ H.H. Smith; W.Indies/ 99-331 (syntypes 6fS). 500 feet; St. Vincent/W.I./

H.H. Smith; W.Indies/ 99-331 (syntype f). Sea level; St. Vincent/ W.I./ H.H. Smith; W.Indies/ 99-

331 (syntype c?)- Leeward side/ St. Vincent, W.I./ H.H. Smith/ 240; W.Indies/ 99-331 (syntype f). St.

Vincent, W.I./H.H. Smith/ 241; W.Indies/ 99-331 (syntype f).

Type locality

West Indies, St Vincent.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

This species was described from 14 females and 22 males, most of which were found. One specimen

labelled by Ashmead as this species is excluded from the type series because it is from Grenada. Most

of the syntypes are entire.

Galesus bipunctatus Ashmead in Riley, Ashmead & Howard, 1894: 248.

Valid name

Coptera sexpunctata (Ashmead, 1893) syn. nov.

Summary of types

Lectotype §, BMNH number 9.1003, here designated. Paralectotype S-

Primary type data

St. Vincent/W.I./H.H. Smith; Galesus/bipunctatus/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

The original description was based on two females and two males, of which a pair was found. As

explained above it is reasonable to suppose that the syntype series of G. bipunctatus and G. sexpunctatus

are exactly the same specimens. The same specimen is designated lectotype for both G. sexpunctatus

Ashmead, 1893 q.v. and G. bipunctatus Ashmead in Riley, Ashmead & Howard, 1894 to ensure objective

synonymy. The lectotype is glued to a micropin and is entire. This species is a Coptera , and Coptera

sexpunctata is a new combination.

Loxotropa thoracica Ashmead in Riley, Ashmead & Howard, 1894: 249.

Valid name

Basalys thoracicus (Ashmead in Riley, Ashmead & Howard, 1894).

Summary of types

Holotype §, BMNH number 9.740, by monotypy.

Primary type data

Leeward side/ St Vincent, W.I./H.H. Smith/ 242; Type/H.T.; Loxotropa/ thoracica/ $ type Ashm./unique.

Type locality

West Indies, St Vincent.

Remarks

The holotype is mounted on a card point, the head and tips of the fore wings are lost.

Tropidopria triangularis Ashmead in Riley, Ashmead & Howard, 1894: 249.

Valid name

Acanthopria triangularis (Ashmead in Riley, Ashmead & Howard, 1894).

European Journal of Taxonomy 75: 1-123 (2014)

Summary of types

Lectotype S, BMNH number 9.937, selected by Masner (1965).

Primary type data

St. Vincent/ W.I./ H.H. Smith; Tropidopria/ triangularis/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

The lectotype is glued to a micropin, and is a little faded, with the left antenna missing beyond the scape.

This species belongs to Acanthopria Ashmead, 1896 where it was placed by Masner (1965).

Tropidopria nigriceps Ashmead in Riley, Ashmead & Howard, 1894: 249, 250.

Valid name

Trichopria nigriceps (Ashmead in Riley, Ashmead & Howard, 1894) comb. nov.

Summary of types

Syntypes 3 SS-

Primary type data

1500 feet; St. Vincent/ W.I./ H.H. Smith; Tropidopria/ nigriceps/ $ Ashm.; W.Indies/ 99-331 (syntype

S). 1000 feet; St Vincent/ W.I./ H.H. Smith; W.Indies/ 99-331 (syntype $). St Vincent/ W.I./ H.H.

Smith/ 158; W.Indies/ 99-331 (syntype $).

Type locality

West Indies, St Vincent.

Remarks

This species was described from a female and four males. The three male syntypes agree well with the

description except that the scutellum is weakly to obscurely carinate; however, there is the superficial

appearance of a strong carina because the cuticle is transparent so that a dark gap between the internal

scutellar muscle blocks is visible in the place where a carina might be, so probably Ashmead misinterpreted

this character and this should not exclude these males from the type series. One additional female

standing over this name is not considered syntypic as it disagrees with the description in several points,

most notably the antennal club which is sub-four-segmented and has the apical four segments darkened.

The syntype labelled “1500 feet” is glued on a micropin and the left and right fore legs and left mid

leg are missing. The syntype labelled “1000 feet” is glued on a micropin and both antennae are missing

beyond a4. The syntype numbered “158” is mounted on a card point, with the right antenna missing

beyond a3 and the right hind leg missing. The new combination Trichopria nigriceps (Ashmead in Riley,

Ashmead & Howard, 1894) is a senior secondary homonym of Trichopria nigriceps (Kieffer, 1913), so

Kieffer’s species is given the new replacement name Trichopria thermarum nom. nov. Thermarum is

the genitive plural of therma , Latin for a thermal bath, or hot spring, and refers to the type locality of

Los Banos.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Tropidopria pallida Ashmead in Riley, Ashmead & Howard, 1894: 249, 250.

Valid name

Trichopriapallida (Ashmead in Riley, Ashmead & Howard, 1894).

Summary of types

Syntypes 10 $$, 3 S<3-

Primary type data

St. Vincent/ W.I./ H.H. Smith; W.Indies/ 99-331 (syntypes 5$$, (?). St. Vincent/ W.I./ H.H. Smith/

158; Tropidopria/ pallida/ $ Ashm.; W.Indies/ 99-331 (syntype $). St. Vincent/W.I./H.H. Smith/ 158;

W.Indies/ 99-331 (syntypes 2§§). St. Vincent/ W.I./ H.H. Smith/ 201; W.Indies/ 99-331 (syntype $).

St. Vincent/ W.I./ H.H. Smith/ 6; W.Indies/ 99-331 (syntype $). May; St. Vincent/ W.I./ H.H. Smith;

Tropidopria/ pallida/ S Ashm.; W.Indies/ 99-331 (syntype S). St. Vincent/ W.I./ H.H. Smith/ 73;

W.Indies/ 99-331 (syntype $).

Type locality

West Indies, St Vincent.

Remarks

This species was described from twelve females and six males, of which ten females and three males

were found. The three females labelled “158” are mounted on card points, two are entire and one is

missing the antennal tips; female syntype labelled “201” is mounted on a card point, and is missing the

antennal tips; five remaining female syntypes are glued to micropins, four are entire and one is missing

its head; male syntype labelled “May” is glued to a micropin and is entire; male syntype labelled “73”

is mounted on a card point and is entire; third male is glued to a micropin and is missing its front legs.

Diapria mellea Ashmead in Riley, Ashmead & Howard, 1894: 251.

Valid name

Trichopria mellea (Ashmead in Riley, Ashmead & Howard, 1894).

Summary of types

Syntypes 2 $$, $.

Primary type data

St. Vincent/W.I./H.H. Smith/ 158; Diapria/mellea/ $ Ashm.; W.Indies/99-331 (syntype §). St. Vincent/

W.I./ H.H. Smith/ 158; W.Indies/ 99-331 (syntype §). Windward side/ St. Vincent, W.I./ H.H. Smith;

Diapria/ mellea/ $ Ashm.; W.Indies/ 99-331 (syntype $).

Type locality

West Indies, St Vincent.

Remarks

This species was described from three females and two males, of which two females and one male were

found. The female syntype with Ashmead’s determination label is mounted on a card point and is entire.

The other female syntype is mounted on a card point and is missing most of the antennae. The male

European Journal of Taxonomy 75: 1-123 (2014)

syntype was glued to a micropin, and has evidently fallen off and been reglued on the same micropin at

some point, so that the tip of the left antenna and tips of some of the legs are glued separately.

Tricliopria insularis Ashmead in Riley, Ashmead & Howard, 1894: 251, 252.

Valid name

Trichopria insularis Ashmead in Riley, Ashmead & Howard, 1894.

Summary of types

Syntype §.

Primary type data

St. Vincent/W.I./H.H. Smith; Trichopria/ insularis/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

This species was described from one specimen of each sex. One female syntype was found; it is glued

to the end of a micropin, and is entire.

Trichopriapleuralis Ashmead in Riley, Ashmead & Howard, 1894: 251, 252.

Valid name

Trichopria pleuralis Ashmead in Riley, Ashmead & Howard, 1894.

Summary of types

Syntype $.

Primary type data

Leeward side/ St. Vincent, W.I./ H.H. Smith/ 239; Trichopria/ pleuralis/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

This species was described from one specimen of each sex, of which the male was found. The male

syntype is mounted on a card point and has much of the right flagellum missing; it has a fine but distinct

basal vein and triangular marginal vein and belongs to the genus Basalys. It appears that the syntype

series includes two species because Ashmead says the female syntype has a four-segmented antennal

club, so it is unlikely to be a Basalys and more likely a Trichopria. It will therefore be necessary at some

point to designate a lectotype to determine the application of the name Trichopria pleuralis ; however,

no designation is made here since designating the male would create homonymy elsewhere and change

the established generic combination, so future revisers are recommended to locate and consider the

female syntype if possible before resolving this problem. The established generic combination is

retained for now.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Trichoprici atriceps Ashmead in Riley, Ashmead & Howard, 1894: 251, 253.

Valid name

Trichopria atriceps Ashmead in Riley, Ashmead & Howard, 1894.

Summary of types

Syntypes $, S-

Primary type data

Leeward side/ St. Vincent, W.I./H.H. Smith/ 72; Trichopria/ atriceps/ 5 Ashm.; W.Indies/ 99-331 (syntype

5 ). St. Vincent/W.I./H.H. Smith/ 158; Trichopria/ atriceps/ $ Ashm.; W.Indies/ 99-331 (syntype S).

Type locality

West Indies, St Vincent.

Remarks

This species was described from two females and one male. A pair of syntypes was found; the female

syntype is mounted on a card and is entire; the male is mounted on a card point and is entire.

Phaenopria subclavata Ashmead in Riley, Ashmead & Howard, 1894: 253, 254.

Valid name

Trichopria subclavata (Ashmead in Riley, Ashmead & Howard, 1894).

Summary of types

Syntypes ?, S-

Primary type data

St. Vincent/ W.I./ H.H. Smith/ 238; Phaenopria/ subclavata/ 5 Ashm.; W.Indies/ 99-331 (syntype §).

Leeward side/ St. Vincent, W.I./ H.H. Smith/ 239; Phaenopria/ subclavata/ S Ashm.; W.Indies/ 99-331

(syntype $)■

Type locality

West Indies, St Vincent.

Remarks

This species was described from two females and two males, of which a pair was found. Thirteen other

females standing over this name, collected by H.H. Smith, are not syntypic as they are from Grenada.

Both syntypes are mounted on card points: the female is missing the left flagellum and right fore leg;

the male is entire.

Phaenopria simillima Ashmead in Riley, Ashmead & Howard, 1894: 253, 254.

Valid name

Trichopria simillima (Ashmead in Riley, Ashmead & Howard, 1894).

Summary of types

Syntype §.

European Journal of Taxonomy 75: 1-123 (2014)

Primary type data

St. V., West/ Indies, Smith; Phaenopria/ simillima/ $ Ashm.; W.Indies/ 99-331.

Type locality

West Indies, St Vincent.

Remarks

This species was described from one female and one male. One female syntype was found; this is glued

to the end of a micropin and is entire.

Loxotropa pleuralis Ashmead, 1896: 803.

Valid name

Basalys pleuralis (Ashmead, 1896).

Summary of types

Holotype §, BMNH number 9.742, by monotypy.

Primary type data

Balthazar/Windward side/ Grenada, W.I./H.H. Smith; Type/H.T.; Loxotropa/ pleuralis/ $ type Ashm.

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from a single female. One specimen was found which has quite a small

scutellar pit; however, it agrees well with the description in all other respects and is certainly the

holotype. The holotype is mounted on a card and is entire.

Loxotropa grenadensis Ashmead, 1896: 803.

Valid name

Basalys grenadensis (Ashmead, 1896).

Summary of types

Holotype §, BMNH number 9.741, by monotypy.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith/ 20; Type/ H.T.; Loxotropa/ grenadensis/ $ Type

As hm

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from one female. The holotype is mounted on a card point and is entire.

The type is a Basalys with 12-segmented antenna, an abrupt three-segmented club, and fore wing with

a basal vein.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Acanthopria crassicornis Ashmead, 1896: 804.

Valid name

Acanthopria crassicornis Ashmead, 1896.

Summary of types

Syntype $, BMNH number 9.691. Syntypes 2 33-

Primary type data

Type/ H.T.; St. John’s Riv./ Leeward side/ Grenada, W.I./ H.H. Smith/ 19; Acanthopria/ crassicornis/

9 type Ashm. (syntype 9 ). Mount Gay Est./ Leeward side/ Grenada, W.I./ H.H. Smith/ 26; Type;

Acanthopria/ crassicornis/ 3 type Ashm.; W.Indies/ 99-331 (syntype 3)- Mount Gay Est./Leeward side/

Grenada, W.I./H.H. Smith/ 25; W.Indies/ 99-331 (syntype 3)-

Type locality

West Indies, Grenada, Balthazar and Mount Gay Estate.

Remarks

This species was described from one female and four males, of which one female and two males were

found. The female differs from the description in that it does not have two faint grooves posteriorly on

the mesonotum, but the mesoscutal setae are stuck down in a way which suggests this, so this most likely

an error of observation on Ashmead’s part and this specimen can still be considered syntypic. Also this

specimen is labelled St John’s River but still is considered syntypic since this river is near the Mount

Gay Estate. All three syntypes are mounted on card points, the female is entire, the male numbered “26”

has the right hind leg missing, the male numbered “25” is entire.

Diapria smithii Ashmead, 1896: 804, 805.

Valid name

Trichopria smithii (Ashmead, 1896).

Summary of types

Lectotype 9, BMNH number 9.710, here designated. Paralectotype 3-

Primary type data

Grenada, W.I./H.H. Smith/ 51; Type/H.T.; Diapria/ smithii/ 9 Type Ashm.

Type locality

West Indies, Grenada.

Remarks

This species was described from a female and a male specimen, both of which were found. The female

is a Trichopria and the male is an Acanthopria. The female is designated as lectotype to maintain the

current combination (Kieffer 1916; Masner 1965). The original description included the multiple original

spellings smithii and smithi ; smithii is to be considered the correct original spelling following the first

reviser action of Dalla Torre (1898). Consequently the spelling smithi is unavailable.

European Journal of Taxonomy 75: 1-123 (2014)

Diapria grenadensis Ashmead, 1896: 804, 805.

Valid name

Trichopria grenadensis (Ashmead, 1896).

Summary of types

Lectotype $, BMNH number 9.727, designated by Masner (1965). Paralectotypes 5 ?$,(?.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith/ 20; Lecto/ type/ C.M.; W.Indies/ 99-331; Diapria/

grenadensis/ $ Type Ashm.; selected as/ lectotype by/ L. Masner xi.61.

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from nine females and two males, of which six female and one male syntypes

were found. Masner (1965) designated a lectotype under the unnecessary replacement name Trichopria

(Planopria) grenadicola Kieffer, 1916. The lectotype is mounted on a card point and is entire; it is a

Trichopria.

Diapria peraffinis Ashmead, 1896: 805, 806.

Valid name

Trichopria peraffinis (Ashmead, 1896).

Summary of types

Lectotype S, BMNH number 9.709a, here designated. Paralectotype §, BMNH number 9.709b.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith/ 40; Type; Diapria/ peraffinis/ $ Type Ashm.;

W.Indies/ 99-331.

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from a female and two males, of which one female and one male were

found. The female syntype is a Doliopria Kieffer, 1910 and the male syntype is a Trichopria. The female

syntype has only eleven antennal segments, not twelve as Ashmead described it, however it is very small

and Ashmead most probably miscounted, otherwise it agrees with the description and bears Ashmead’s

type label, so is considered to be syntypic here. The male is designated as lectotype in order to maintain

the current generic combination. The lectotype is mounted on a card point and is missing the apex of the

right hind tarsus.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Diapria melanopleura Ashmead, 1896: 805, 806.

Valid name

Trichopria melanopleura (Ashmead, 1896) stat. rev.

Summary of types

Holotype S, BMNH number 9.726, by monotypy.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith; Type/ H.T.; Diapria/ melanopleura/ $ Type

Ashm.

Type locality

West Indies, Grenada, Balthazar.

Remarks

The holotype is mounted on a card and has the tip of right hind tarsus missing. This species was included

in Xyalopria Kieffer, 1907 by Masner (1965), then Megaplastopria Ashmead, 1903 by Arias-Penna

(2003); however, this species does not belong to Megaplastopria. While it has a slender propodeal spine,

it lacks key characters of Xyalopria such as the occipital spine and carina, grooved scape and flared

corners of the metasomal tergites, and is returned here to Trichopria. It belongs to the group of species

in which the males have whorled hairs on the antennal flagellum.

Diapria unicolor Ashmead, 1896: 805, 806.

Valid name

Trichopria unicolor (Ashmead, 1896).

Summary of types

Holotype §, BMNH number 9.708, by monotypy.

Primary type data

Balthazar/Windward side/ Grenada, W.I./H.H. Smith/ 38; Type/H.T.; Diapria/ unicolor/ § Type Ashm.

Type locality

West Indies, Grenada, Balthazar.

Remarks

The holotype is mounted on a card point and is entire.

Ceratopria grenadensis Ashmead, 1896: 807.

Valid name

Basalys grenadae (Kieffer, 1912).

Summary of types

Syntype §, BMNH number 9.745.

European Journal of Taxonomy 75: 1-123 (2014)

Primary type data

Mount Gay Est./ Leeward side/ Grenada, W.I./ H.H. Smith/ 25; Type/ H.T.; Ceratopria/ grenadensis/ $

Type Ashm.; W.Indies/ 99-331.

Type locality

West Indies, Grenada, Balthazar and Mount Gay Estate.

Remarks

Described from two females of which one syntype was found. The syntype is mounted on a card point

and is entire. It is a Basalys with the antenna 12-segmented, an abrupt three-segmented club, and fore

wing with basal vein.

Ceratopria flavipes Ashmead, 1896: 807.

Valid name

Basalys flavidipes (Kieffer, 1912).

Summary of types

Holotype §, BMNH number 9.744, by monotypy.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith/ 40; Type/ H.T.; Ceratopria/ flavipes/ $ Type

As hm

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from a single female. The holotype is mounted on a card point and is entire.

Trichopria grenadensis Ashmead, 1896: 807, 808.

Valid name

Trichopria ashmeadi Kieffer, 1912.

Summary of types

Syntype $, BMNH number 9.712. Syntype f.

Primary type data

GrandEtang/Windward side/ 1900ft/Grenada, W.I./H.H. Smith/13; Type/H.T.; Trichopria/grenadensis/

5 Type Ashm. (syntype §). Balthazar/Windward side/ Grenada, W.I./H.H. Smith/ 37; Type; Trichopria/

grenadensis/ $ Type Ashm.; W.Indies/ 99-331 (syntype $).

Type locality

West Indies, Grenada, Balthazar and Grand Etang.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

This species was described from one female and two males, of which one female and one male syntype

was found. Both syntypes are mounted on card points, are entire, and both belong to Trichopria.

Trichopria bifoveata Ashmead, 1896: 808.

Valid name

Trichopria bifoveata Ashmead, 1896.

Summary of types

Syntype §, BMNH number 9.711.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith/ 20; Type/ H.T.; Trichopria/ bifoveata/ $ Type

Ashm.

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from two females, of which one was found. The syntype is mounted on a

card point, is entire, and has the wings matted and dirty.

Trichopria affinis Ashmead, 1896: 808.

Valid name

Trichopria neotropica Masner, 1965.

Summary of types

Holotype $, BMNH number 9.707, by monotypy.

Primary type data

Balthazar/Windward side/ Grenada, W.I./H.H. Smith/ 20; Type/H.T.; Trichopria/ affinis/ f Type Ashm.

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from a single specimen. It is mounted on a card point and is entire.

Phaenopria nigricornis Ashmead, 1896: 810.

Valid name

Trichopria obscura Masner, 1965.

Summary of types

Holotype BMNH number 9.722, by monotypy.

European Journal of Taxonomy 75: 1-123 (2014)

Primary type data

St. John’s Riv./ Leeward side/ Grenada, W.I./ H.H. Smith/ 16; Type/ H.T.; Phaenopria/ nigricomis/ cf

Type Ashm.

Type locality

West Indies, Grenada, St John’s River.

Remarks

This species was described from one male. The specimen agrees well with the description in all respects,

except that a4 is angulate; probably Ashmead missed this character. It is mounted on a card point and is

entire.

Phaenopria angulifera Ashmead, 1896: 810 lines 13, 29.

Valid name

Trichopria angulifera (Ashmead, 1896).

Summary of types

Holotype S, BMNH number 9.718, by monotypy.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith; Type/ H.T.; Phaenopria/ angulifera/ m Type

As hm

Type locality

West Indies, Grenada, Balthazar.

Remarks

Described from one male specimen; it is mounted on a card, is entire, and the wings are matted.

Phaenopria grenadensis Ashmead, 1896: 809 (as P. grenadensis ),

810 line 44 (as P. angulifera [sic]).

Valid name

Trichopria confusa Masner, 1965.

Summary of types

Syntype §, BMNH number 9.713. Syntype §.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith; Type/ H.T.; Phaenopria/ grenadensis/ $ Type

Ashm. (syntype §). Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith/ 20.; W.Indies/ 99-331

(syntype $).

Type locality

West Indies, Grenada, Balthazar and Mount Gay Estate.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

This species was described from six females, of which two syntypes were found. The specimen

numbered 9.713 is mounted on a card and is missing part of the right fore leg; the other is mounted on

a card point and is missing the tip of the right hind leg. Both belong to Trichopria. In his treatment of

this species Ashmead used the specific name grenadensis in the key (p. 809) and angulifera heading

the description (p. 810 line 44 - species number 3) as is evident from the correspondence between the

characters used in the key and the description, hence P. angulifera Ashmead, 1896: 810 line 44 and P.

grenadensis are multiple original spellings. Ashmead (1900) is the first reviser under Art. 24.2.4, and

he chose P. grenadensis as the correct original spelling. P. angulifera Ashmead, 1896: 810 line 44 is

thus an incorrect original spelling, has no separate availability and cannot enter into homonymy with P.

angulifera Ashmead, 1896: 810 lines 12, 29 (species number 2) q.v. above.

Phaenopria nigriclavata Ashmead, 1896: 809, 811.

Valid name

Trichopria nigriclavata (Ashmead, 1896).

Summary of types

Holotype §, BMNH number 9.721, by monotypy.

Primary type data

Chantilly Est./ Windward side/ Grenada, W.I./ H.H. Smith/ 14; Type/ H.T.; Phaenopria/ nigriclavata/ y

Type Ashm.

Type locality

West Indies, Grenada, Chantilly Estate.

Remarks

This species was described from one female. The holotype is mounted on a card point and is entire.

Phaenopria balthazari Ashmead, 1896: 809, 811.

Valid name

Trichopria balthazari (Ashmead, 1896).

Summary of types

Syntype §, BMNH number 9.719.

Primary type data

Balthazar/ Windward side/ Grenada, W.I./ H.H. Smith/ 20; Type/ H.T.; Phaenopria/ balthazari/ $ Type

As hm

Type locality

West Indies, Grenada, Balthazar.

Remarks

This species was described from two females, of which one was found. The syntype is mounted on a

card point and is entire.

European Journal of Taxonomy 75: 1-123 (2014)

Phaenoprici magniclavata Ashmead, 1896: 809, 812.

Valid name

Doliopria magniclavata (Ashmead, 1896) comb. nov.

Summary of types

Lectotype §, BMNH number 9.720, here designated. Paralectotype §.

Primary type data

Grand Etang/ Windward side 1900ft./ Grenada, W.I./ H.H. Smith/ 13; Type/ H.T.; Phaenopria/

magniclavata/ $ Type Ashm.

Type locality

West Indies, Grenada, Grand Etang.

Remarks

This species was described from three females, of which two were found. The two syntypes differ

from the description in having an eleven-segmented antenna and belong to Doliopria\ however, this

discrepancy is accounted for by the fact that the antennae are covered with glue which obscures the

number of segments and there is no reason not to consider them syntypes. A lectotype is designated in

order to stabilise the new combination. The lectotype is mounted on a card point and is entire.

Zacranium oahuense Ashmead, 1901: 295.

Valid name

Zacranium oahuense Ashmead, 190E

Summary of types

Holotype §, BMNH number 9.680, by monotypy.

Primary type data

29; Type/H.T.; WaianaeMts.,/ Oahu, ft/Perkins. 4.1892; Sandwich Is./ 1912-215; Zacranium/ oahuense/

5 Type As hm

Type locality

Hawaii, Oahu, Waianae Mountains.

Remarks

This species was described from one female. The holotype agrees well with the description and has

Ashmead’s type label. Although Ashmead says it was collected in June, the label says April, an error

on Ashmead’s part. The holotype is mounted on a card and is entire. Zacranium Ashmead, 1901 has a

slender fore tibial spine and is most probably allied to the Diapria genus group.

Platymischoides molokaiensis Ashmead, 1901: 296.

Valid name

Platymischoides molokaiensis Ashmead, 1901.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Syntype §, BMNH number 9.679. Syntype §.

Primary type data

163; Type/ H.T.; Molokai Mts.,/ 74000 ft+/ Perkins ixl893.; Sandwich Is./ 1912-215; Platymischoides/

mololcaiensis/ $ Type Ashm. (syntype $). Hawaii: Molokai/ Mountains/ 74000 ft+: R.C.L./ Perkins:

ix. 1893; Sandwich Isles/ BMNH(E)/ 1912-215; $ Platymischoides/ molokaiensis Ashm./ Type det.

Ashmead (syntype §).

Type locality

Hawaii, Molokai.

Remarks

Described from three females, of which two were found, agreeing with the description except they

were collected in September, not October. This is probably an error on Ashmead’s part as this part of

the label is not as clear as it could be. The two syntypes were originally mounted together on the same

card, but one has been remounted on a card point; the specimen still on the original card has a red ink

circle around it and has the left apical flagellomere separated but present; the other specimen is entire.

Platymischoides Ashmead, 1901 and Platymischodes Ashmead, 1901 are multiple original spellings.

In choosing Platymischoides , Ashmead (1903) appears to be the first reviser and Platymischodes is

consequently unavailable (Article 24.2.4).

Phaenopria hawaiiensis Ashmead, 1901: 296.

Valid name

Trichopria hawaiiensis (Ashmead, 1901).

Summary of types

Syntype §, BMNH number 9.723. Syntype <$■

Primary type data

Type/ H.T.; Molokai Mts.,/ 3000ft./ Perkins 19 & 20ixl893.; Sandwich Is./ 1912-215; Phaenopria/

hawaiiensis/ $ Type (syntype $). Type/ Molokai Mts.,/ 8000ft./ Perkins 19 & 29ixl893; Sandwich Is./

1912-215; Phaenopria/ hawaiiensis/ $ Type Ashm. (syntype $).

Type locality

Hawaii, Molokai.

Remarks

This species was described from one female and one male. Although Ashmead says July, both specimens

were collected in September. This is probably an error on Ashmead’s part since the dates on the labels

are almost illegible. Similarly, although Ashmead says 3500 ft, one specimen is labelled 3000 ft (or

possibly 5000 ft) and the other 8000 ft; again the altitudes are written on the labels in small, barely

legible handwriting, and in one case the number is transfixed by the pin, so probably this was an error in

transcription on Ashmead’s part. The syntypes are both carded: the female is entire; the male is missing

its left antenna beyond a5 and its right mid and hind legs beyond the coxae.

European Journal of Taxonomy 75: 1-123 (2014)

Mimopria comes Borgmeier, 1939: 532, figs 1,13.

Valid name

Mimopria comes Borgmeier, 1939.

Summary of types

Syntype $.

Primary type data

Campinas, Goiaz/ Schwarzmaier/ 24.T.30/E. crassicorne; Cotypus; Mimopria/ comes Borgm./Paratype/

det. Borgmeier; Brit. Mus./ 1950-553.

Type locality

Brazil, Goias, Campinas and Inhumas; Sao Paulo, Barrettos; Pernambuco, Tapera.

Remarks

This species was described from “numerous” females. Other syntypes were apparently in Borgmeier’s

collection. One female was found agreeing with Borgmeier’s description, and although this specimen is

labelled as a “paratype” and “cotype”, since neither a unique holotype or lectotype has yet been fixed,

the type series is still syntypic (Article 73.2). The syntype is mounted on a card point and has the head

missing.

Mimopria barbata Borgmeier, 1939: 534, figs 14,19.

Valid name

Mimopria barbata Borgmeier, 1939.

Summary of types

Syntype $.

Primary type data

Eciton/ crassicorne; Campinas/ Goias 9.iii.36/ Schwarzmaier; Cotypus; Mimopria/ barbata Borg./

Paratype/ det. Borgmeier; Brit. Mus./ 1950-553.

Type locality

Brazil, Goias, Campinas.

Remarks

This species was described from ten females of which one was found agreeing with Borgmeier’s

description, and although this specimen is labelled as a “paratype” and “cotype”, since neither a unique

holotype or lectotype has yet been fixed, all of the type series is still syntypic (Article 73.2). The syntype

is mounted on a card point and entire.

Mimopria splendens Borgmeier, 1939: 534, figs 1,15.

Valid name

Mimopriella splendens (Borgmeier, 1939).

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Syntype §.

Primary type data

Campinas, Goyaz/E. goldii 756/ 15.4.36; Cotypus; Mimopria/ splendens B./Paratype/ det. Borgmeier;

Brit. Mus. 1950-553.

Type locality

Brazil, Goias, Campinas and Trindade.

Remarks

This species was described from seven females of which one was found agreeing with Borgmeier’s

description, and although this specimen is labelled as a “paratype” and “cotype”, since neither a unique

holotype or lectotype has yet been fixed, all of the type series is still syntypic (Article 73.2). The syntype

is micropinned and entire. This species was transferred to Mimopriella Masner & Garcia, 2002 by

Masner & Garcia (2002).

Philolestespronotalis Borgmeier, 1939: 536, figs 5,11,18.

Valid name

Notoxoidespronotalis (Borgmeier, 1939).

Summary of types

Syntype $.

Primary type data

Eciton/ dulcius; Campinas/ Goias 5.xi.33/ Schwarzmaier; Cotypus; Philolestes/ pronotalis Borgm./

Paratype/ det. Borgmeier; Brit. Mus./ 1950-553.

Type locality

Brazil, Goias, Campinas.

Remarks

This species was described from numerous dealate females, one alate female and four males of which

one female was found agreeing with Borgmeier’s description. Although this specimen is labelled as a

“paratype” and “cotype”, since neither a unique holotype or lectotype has yet been fixed, all of the type

series is still syntypic (Article 73.2). The syntype is micropinned, is missing its head, and is mounted

together with an ant. Philolestes Kieffer, 1922 was synonymized with Notoxoides Ash mead, 1903 by

Masner (1977).

Notoxopria pedissequa Borgmeier, 1939: 538, figs 2,12.

Valid name

Notoxoides pedissequa (Borgmeier, 1939).

Summary of types

Syntype $.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Primary type data

Campinas, Goyas/ Schwarzmaier/ 17.xii.28/ E. pseudops; Cotypus; Notoxopria/ pedissequa/ Borgm.

Paratype/ det. Borgmeier; Brit. Mus. 1950-553.

Type locality

Brazil, Goias, Campinas.

Remarks

This species was described from numerous dealate females and one alate female, of which one was found

agreeing with Borgmeier’s description. Although this specimen is labelled as a “paratype” and “cotype”,

since neither a unique holotype or lectotype has yet been fixed, all of the type series is still syntypic

(Article 73.2). The syntype is micro-pinned and entire and it is mounted together with an ant. Notoxopria

Kieffer, 1910 was synonymized with Notoxoides by Masner (1977). The epithet “pedissequa” is a noun

in apposition and is indeclinable.

Asolenopsia schwarzmaieri Borgmeier, 1939: 542, fig. 10.

Valid name

Asolenopsia schwarzmaieri Borgmeier, 1939.

Summary of types

Syntype §.

Primary type data

Campinas/ Goyaz/ E. pseudops 686/ 4.2.36; Cotypus; Campinas/ Goias/ Schwarzmaier; Asolenopsia/

schwarzmaieri/Borgm. Paratype/ det. Borgmeier; Brit. Mus./ 1950-553.

Type locality

Brazil, Goias, Campinas.

Remarks

This species was described from numerous females, of which one was found agreeing with Borgmeier’s

description. Although this specimen is labelled as a “paratype” and “cotype”, since neither a unique

holotype or lectotype has yet been fixed, all of the type series is still syntypic (Article 73.2). The syntype

is mounted on a card and is entire.

Neivapria penicillata Borgmeier, 1939: 544, figs 3, 4, 6, 7.

Valid name

Neivapria penicillata Borgmeier, 1939.

Summary of types

Syntype $.

Primary type data

Campinas, Goyas/ Schwarzmaier/ T22 mimense/ 11.3.36; Cotypus; Neivapria/ penicillata/ Borgm.

Paratype/ det. Borgmeier.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Type locality

Brazil, Goias, Campinas.

Remarks

This species was described from numerous females of which one was found agreeing with Borgmeier’s

description. Although this specimen is labelled as a “paratype” and “cotype”, since neither a unique

holotype or lectotype has yet been fixed, all of the type series is still syntypic (Article 73.2). The syntype

is mounted on a card point and the head is lost.

Loxotropa fuliginosi Box, 1921: 16.

Valid name

Basalys pedisequa (Kieffer, 1911).

Summary of types

Syntype §, BMNH number 9.762.

Primary type data

Loxotropa/ fuliginosi/ Woking/ 30.V.20; Assoc, with/ Lasius fuliginosus; Ex coll./ Donisthorpe/

B.M. 1934-4; Loxotropa/ fuliginosi Box; Holo-/ type.

Type locality

England, Woking.

Remarks

This species was described from an unspecified number of females from Donisthorpe’s collection. The

sole surviving syntype is mounted on a card and is entire. Nixon’s (1980) interpretation of Basalys

pedisequa (Kieffer, 1911) is followed here.

Aulatopria tucumana Brethes, 1927a: 164.

Valid name

Bruchopria tucumana (Brethes, 1927).

Summary of types

Syntypes $, (?•

Primary type data

Reared; Rosenfeldy/Barber, colls.; Est. Exp./A.C. No. 115; Tucuman/1-22-1913; Aulatopria/tucumana

Brethes; Pres. By Imp. Bur. Ent./Brit. Mus./ 1928-347 (syntypes §, S).

Type locality

Argentina, Tucuman.

Remarks

The description for this species was published twice, in the Revista industrial y agricola de Tucuman

(Brethes 1927a) and translated into English in the Bulletin of Entomological Research (Brethes 1927b).

It is most probable that the Revista came first, as this work was published every two months at that

European Journal of Taxonomy 75 : 1-123 ( 2014 )

time; volume 17(7-8) contains weather reports up to 31 January 1927 and was probably published

shortly afterwards, whereas the English translation in the Bulletin has the cover date December 1927.

Brethes mentions that the specimens were reared during studies of the sugar-cane moth borer Diatraea

saccharalis Fabricius, 1794 by Rosenfeld and Barber and published in 1914 also in Revista industrial

y agricola de Tucuman. Rosenfeld and Barber’s paper could not be traced during this study, but is

mentioned here since it may throw further light on the composition of the type series. Two syntypes were

found which had been reared by Rosenfeld and Barber in 1913, labelled by Brethes, and received at the

Imperial Bureau of Entomology in London, shortly after the date of the description. Both syntypes are

mounted on card points and are somewhat dirty: the female is missing the left fore wing and has the right

fore wing chipped; the male is entire. This species is now placed in Bruchopria Kieffer, 1921 following

Masner & Garcia (2002).

Paramesius fasciatipennis Cameron, 1888: 437, pi. 18, fig. 18.

Valid name

Spilomicrus fasciatipennis (Cameron, 1888).

Summary of types

Syntype §, BMNH number 9.639. Syntypes §, <$.

Primary type data

Bugaba,/ Panama/ Champion; Type/ H.T.; B.C.A. Hymen. I./ Paramesius/ fasciatipennis/ Cam.;

Paramesius/ fasciatipennis/ Cam. type (syntype §). Fasciati-/ pennis; V.de Chiriqui/ 25-4000ft/

Champion; PCameron Coll./ 1914-110 (syntype $). Maculi-/ fasciati -/pennis; V.de Chiriqui/25-4000ft/

Champion; PCameron Coll./ 1914-110 (syntype 3).

Type locality

Panama, Bugaba, Volcan de Chiriqui.

Remarks

This species was described from at least one specimen of each sex. Two female and one male syntypes

were found. Cameron appears to have later changed his mind about the identification of the male,

crossing out the name fasciatipennis and replacing it with maculipennis ; however, this specimen is

clearly the male referred to as it agrees so well with the description, and not with that of maculipennis.

Another female labelled “fasciatipennis” differs significantly from the description in having shallow

notauli, a shorter petiole, different antennal proportions, the altitude is wrong, and it is not considered

syntypic. All three syntypes are mounted on cards and are entire.

Paramesius maculipennis Cameron, 1888: 438, pi. 18, figs 11-12.

Valid name

Spilomicrus maculipennis (Cameron, 1888).

Summary of types

Syntype §, BMNH number 9.640b.

Primary type data

Fasciati/ maculi-/ pennis; Bugaba/ Panama/ Champion; PCameron Coll./ 1914-110 (syntype $).

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Type locality

Panama, Bugaba.

Remarks

This species was described from at least one specimen of each sex. One female was found which agreed

exactly with the description and had been labelled by Cameron as “maculipennis”. The male and female,

mounted together, from Vera Paz formerly regarded as types (Masner 1965) are not considered syntypic

because they are not from the type locality. A fourth specimen labelled “maculipennis” did not agree

with the description and is considered here to be a syntype of P. fasciatipennis q.v. which Cameron later

incorrectly reidentified. The syntype is mounted on a card and is entire.

Paramesius canaliculatus Cameron, 1888: 439, pi. 18, fig. 24.

Valid name

Spilomicrus canaliculatus (Cameron, 1888).

Summary of types

Syntype §, BMNH number 9.638.

Primary type data

Cordova; Mexico/ Salle coll.; Type/H.T.; B.C.A. Hymen. 1/Paramesius/ canaliculatus/ Cam. Type.

Type locality

Mexico.

Remarks

This species was described from an unspecified number of females, of which one syntype was found.

The syntype is mounted on a card and is entire.

Paramesius chiriquensis Cameron, 1888: 439, pi. 18, fig. 15.

Valid name

Pentapria chiriquensis (Cameron, 1888).

Summary of types

Syntype §, BMNH number 9.641.

Primary type data

V.de Chiriqui/ 8000ft/ Champion; B.C.A. Hymen. I./ Paramesius/ chiriquensis/ Cam.; Type/ H.T.;

Paramesius/ chiriquensis/ P.439 Cam.

Type locality

Panama, Volcan de Chiriqui.

Remarks

This species was described from an unspecified number of females of which one was found. The syntype

agrees with the description except that Cameron miscounted the number of keels on the petiole - there

are seven. The syntype is mounted on a card and is entire.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Spilomicrus tinctipennis Cameron, 1888: 440, pi. 18, fig. 17.

Valid name

Idiotypa tinctipennis (Cameron, 1888).

Summary of types

Holotype §, BMNH number 9.747, by monotypy.

Primary type data

Bugaba/ Panama/ Champion; B.C.A. Hymen. I./ Spilomicrus/ tinctipennis/ Cam.; Type/ H.T.;

Spilomicrus/ tinctipennis/ Cam./B.C.A. ii 440.

Type locality

Panama, Bugaba.

Remarks

The description of this species was based on an unspecified number of females. One specimen was

found which agreed with the description, except that it did not have a blunt, projecting, basal tooth on

the hind femur; however, a piece of dirt was found on the base of the right hind femur which gave the

appearance of a tooth (the base of the left femur was obscured). It is therefore considered that Cameron’s

description is incorrect on this point, and should not prevent the specimen from being a type. It is most

unlikely there would be another specimen with a piece of dirt on the base of the hind femur giving the

appearance of a tooth, so it is considered that this was the only specimen Cameron saw and is therefore

a holotype by monotypy. The holotype is mounted on a card and is dirty with the left wings crumpled.

The piece of dirt which gave the appearance of a tooth on the femur has been removed, but the specimen

is now clearly labelled so it can still be recognised.

Malvina punctata Cameron, 1889: 13.

Valid name

Spilomicruspunctatus (Cameron, 1889) comb. nov.

Summary of types

Syntype $, BMNH number 9.681. Syntypes 3 $ §.

Primary type data

Type; Greymouth; Cameron Coll./ 98-124; Malvina/ punctata/ Cam. Type/ New Zealand (syntype §).

N.Zeal. (syntype §). Cameron./ 99-30.; N.Zeal. (syntypes 2§5).

Type locality

New Zealand, Greymouth.

Remarks

The description was made from an unspecified number of females, certainly more than one since

Cameron says “coxae usually black”, and since the specimen labelled by Cameron as type has the petiole

obscured by its wings he must have seen another to describe the petiole. Besides the specimen labelled

by Cameron as a type, three other specimens were found in the collection, two with register labels

saying they were from Cameron’s collection, and the third with a similar mounting style and locality

NOTTON D.G., Diapriinae types at The Natural History Museum, London

apparently written in Cameron’s hand. All four are considered syntypic. In a later paper Cameron (1898)

noted there were four specimens in this collection. The syntype with register number 98-124 is mounted

on a card and is missing its left hind leg; the syntype without a register number is mounted on a card

point and is missing the right antenna beyond a2; of the two syntypes with register number 99-30, one

is mounted on a card point and is missing parts of all the tarsi and the other is mounted on a card and is

missing the right hind tarsus. Malvina punctata appears to be a part of a small group of related species

with pronounced punctation and sometimes striation on the head and mesosoma; these appear to be little

more than a derived species group of Spilomicrus Westwood, 1832. Cameron (1889) noted that this

species has a metanotal spine, which is a rare character in Diapriinae; however, it is not the metanotum,

but the anterior propodeum which is spinose, a feature found in some other Spilomicrus. This species is

transferred to Spilomicrus here. Since M. punctata is the type species of Malvina Cameron, 1889, this

genus becomes a synonym of Spilomicrus syn. nov. The pronotal groove, which is supposed to be a

feature of Malvina according to Early (1980), is virtually absent in the females of the type species and is

found in some other species of Spilomicrus which had not been included in Malvina. The other species

currently placed in Malvina can be accounted for as follows: M. quadriceps (Smith, 1878) is returned

to Spilomicrus stat. rev., q.v. below; M. helosciomyzae Early & Horning, 1978 and M. insulae Early,

1980 are both transferred to Spilomicrus (new combinations). Spilomicrus punctatus (Cameron, 1889)

comb. nov. is a senior secondary homonym of Spilomicrus punctatus Kozlov, 1978; the latter is given

the replacement name Spilomicrus kozlovi nom. nov.

Loxotropa tricarinata Cameron, 1912: 69.

Valid name

Trichopria tricarinata (Cameron, 1912).

Summary of types

Holotype §, BMNH number 9.751, by monotypy.

Primary type data

Type; Z13; Kuching/ J.H./ 100; P.Cameron Coll./ 1914-110; Loxotropa/ 3-carinata/ Cam. Type/Borneo.

Type locality

Malaysia, Borneo, Sarawak, Kuching.

Remarks

The description of this species was made from at least one female. The type specimen agrees broadly

with the description; there are a number of errors in the description, in particular Cameron’s erroneous

observation of a non-existent basal cell in the fore wing, which may be explained by the fact that the

venter, metapleura, propodeum, petiole and wings are all covered with glue and the wings are folded

together, obscuring part of the body. Since the errors of observation relate to the particular condition of

this specimen it is considered that Cameron based his description only on this specimen; hence it is a

holotype by monotypy. The holotype is mounted on a card and is entire.

Galesus cratocerus Cameron, 1912: 69.

Valid name

Coptera cratocerus (Cameron, 1912) comb. nov.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Summary of types

Syntype §, BMNH number 9.672.

Primary type data

Type; Kuching/ J.H.; P.Cameron Coll./ 1914-110; Galesus/ cratocerus/ Cam. Borneo.

Type locality

Malaysia, Borneo, Sarawak, Kuching.

Remarks

This species was described from an unspecified number of females. One syntype was found, although

there may have been others as Cameron made detailed observations on the length of the hairs on the

antennae, propodeum and petiole, which in the surviving syntype are all covered with glue. The syntype

is mounted on a card, much matted with glue, right antenna missing beyond a5, and left antenna obscured

by glue but probably missing beyond the scape.

Hoplopria curvispina Cameron, 1913: 136.

Valid name

Spilomicrus curvispina (Cameron, 1913).

Summary of types

Syntype BMNH number 9.918.

Primary type data

Type; 359; P.Cameron Coll./ 1914-110; Hoplopria/ curvispina/ Cam. Type/ Br. Guiana.

Type locality

Guyana (as British Guiana).

Remarks

This species was described from an unspecified number of males. One male was found. Reference to

the “male” on p. 135 could equally mean the male sex, so is not clear evidence that there was only one

specimen, and the type is treated here as a sole surviving syntype. The syntype is mounted on a card,

with the head reglued upside-down, the left hind wing has become detached and is now stuck to the

scapes, and the left antenna beyond a6 and parts of the left fore and mid tarsi are missing. The name

curvispina does not change its ending in the new combination since it can be considered based on the

Latin noun spina , meaning a thorn (Article 31.2.2).

Hoplopriapicicornis Cameron, 1913: 136.

Valid name

Spilomicrus picicornis (Cameron, 1913).

Summary of types

Syntypes 2 $5, BMNH number 9.647.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Type; 358; P.Cameron Coll./ 1914-110; Hoplopria/ picicomis/ Cam. Type/ Br. Guiana (syntypes 2 $ §).

Type locality

Guyana (as British Guiana).

Remarks

The description is based on an unspecified number of females; although Cameron refers to “the female”

on p. 135, this could refer to the female sex and is not clear evidence for a unique type. Two female

syntypes were found, mounted on the same card, the left specimen is entire, and the right specimen is

missing its head and the tip of the right hind tarsus.

Hoplopria aterrima Dodd, 1920: 373.

Valid name

Spilomicrus aterrimus (Dodd, 1920) comb. nov.

Summary of types

Holotype $, BMNH number 9.635, by original designation. Paratype <$■ Lectotype designation by

Masner (1965) invalid.

Primary type data

Lecto/ type/ L.M.; Kuching/ J.H.; P.Cameron Coll./ 1914-110; Brachyaulax/ striaticollis/ Cam. Type/

Borneo; Hoplopria/ aterrima Dodd/ §; selected as lectotype by/ L. Masner 25.xi.01.

Type locality

Malaysia, Borneo, Sarawak, Kuching.

Remarks

This species was described from one specimen of each sex, of which Dodd said one was type and the

other cotype (effectively holotype and paratype). There is, however, no clear evidence to say which of

the two specimens is the type either from Dodd’s description or labels. Masner (1965) tried to resolve

the situation by designating a lectotype; however, this action is invalid since only syntypes can be

designated as lectotypes and Dodd had already designated a holotype. It is prudent to recognise the

female specimen as Dodd’s holotype because a) Dodd described the female first and in more detail and

so it is most likely he considered the female specimen the type, and b) it is in the interests of stability to

follow Masner in recognising the female as the name bearing type. The holotype is mounted on a card

and is entire. Following the synonymy of Hoplopria Ashmead, 1893 with Spilomicrus presented by

Masner & Garcia (2002), the position of this species in Spilomicrus is confirmed.

Hoplopria obsoleta Dodd, 1920: 375.

Valid name

Spilomicrus obsoletus (Dodd, 1920).

Summary of types

Holotype BMNH number 9.636, by monotypy.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Primary type data

Omilteme/ Guerrero/ 8000ft./ Aug. H.H. Smith; Godman-Salvin/ Coll. 1904.-1.; Type/H.T.; Hoplopria/

obsoleta/ Dodd $.

Type locality

Mexico, Guerrero, Omilteme.

Remarks

This species was described from one female. The holotype is mounted on a card with the right fore and

hind wings missing.

Hoplopria affinis Dodd, 1920: 375.

Valid name

Spilomicrus affinis (Dodd, 1920).

Summary of types

Holotype §, BMNH number 9.637, by original designation. Paratype §.

Primary type data

Teapa/Tabasco/March. H.H.S.; Godman-Salvin/ Coll. 1904.-1.; Type/H.T.; Hoplopria/ affinis/Dodd §.

Type locality

Mexico, Tabasco, Teapa.

Remarks

This species was described from two females, which Dodd designated type and cotype, of which both

were found. The holotype is mounted on a card and has part of the left hind tarsus missing.

Hemilexis paucisetis Dodd, 1920: 377.

Valid name

Spilomicrus paucisetis (Dodd, 1920).

Summary of types

Holotype §, BMNH number 9.724, by original designation.

Primary type data

Mt. Wellington,/ S.Tasmania./ 12-21 Mch. 1913.; 1,300 to 2,300 ft./R.E.Turner./ 1913-212.; Hemilexis/

paucisetis/ Dodd $.

Type locality

Australia, Tasmania, Mount Wellington.

Remarks

The holotype is mounted on a card point, with the head, left fore wing and part of right hind tarsus

missing.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Bakeria rugosci Dodd, 1920: 377.

Valid name

Doddius rugosus (Dodd, 1920).

Summary of types

Holotype §, BMNH number 9.676, by original designation.

Primary type data

Omilteme/ Guerrero/ 8000 ft./ Aug., H.H. Smith.; Godman-Salvin/ Coll. 1904.-1.; Type; Bakeria/

rugosa/Dodd, §.

Type locality

Mexico, Guerrero, Omilteme.

Remarks

The holotype is mounted on a card, all wings missing except left hind wing, right antenna beyond scape

mounted separately, and left fore leg and both mid legs damaged. Masner & Garcia (2002) described

the genus Doddius with Bakeria rugosa as the type species. Arias-Penna (2003) placed this species

simultaneously in Doddius and Pentapria Kieffer, 1905; the latter is an error.

Paramesius longior Dodd, 1920: 378.

Valid name

Paramesius longior Dodd, 1920.

Summary of types

Holotype §, BMNH number 9.663, by original designation. Paratype S-

Primary type data

Type; Hewitt/ Kuching; J2; PCameron Coll./ 1914-110; Brachyaulax/ rufipes Cam. §/ Type Borneo;

Paramesius §/ longior Dodd.

Type locality

Malaysia, Borneo, Sarawak, Kuching.

Remarks

This species was described from a specimen of each sex which had been labelled with the manuscript

name “ Brachyaulax rufipes ” by Cameron. One female and four males were found labelled “ B . rufipes ”

in Cameron’s hand. Of these two of the males were mounted together; these could not have been seen by

Dodd as he only mentioned one male. A third male is labelled as a type but is not, as the antennae are the

wrong colour. The female and remaining male are the types. The female is labelled “type” and the male

“cotype”, following Dodd’s published, designation so the female is the holotype by original designation.

The holotype is mounted on a card, and has its wings torn and partly missing.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Mantara bifurcata Dodd, 1920: 380.

Valid name

Aneuropria bifurcata (Dodd, 1920) comb. nov.

Summary of types

Holotype §, BMNH number 9.677, by original designation. Paratypes §, <f.

Primary type data

1081.; Type; Madeira./ Wollaston./ 55.7; Mantara/ bifurcata/ Dodd §; Holotype §/ Mantara/ bifurcata

Dodd/ L.Huggert-81.

Type locality

Madeira.

Remarks

This species was described from two females and a male, referred to in the original description as

a “type” and “cotypes”. Three specimens matching the description were found, one of the females

has Dodd’s identification label, and this is considered to be the holotype by original designation. The

holotype is mounted on a card, with the left antenna mounted separately. Mantara bifurcata is actually a

species of Aneuropria Kieffer, 1905. Since Mantara bifurcata is the type species of Mantara, this means

Mantara Dodd, 1920 is a junior synonym of Aneuropria Kieffer, 1905 syn. nov. Aneuropria bifurcata

comb. nov. is clearly a different species (in head shape, antennal proportions and the lack of notauli)

from the widespread European A. foersteri Kieffer, 1910, which is also found in Madeira.

Acidopria spinosiceps Dodd, 1920: 380.

Valid name

Trichopria spinosiceps (Dodd, 1920) comb. nov.

Summary of types

Holotype §, BMNH number 9.692, by original designation. Paratype §.

Primary type data

Type; Kuching/ J.H.; P.Cameron Coll./ 1914-110; Brachyaulax/ erythrocerus/ Cam. Type Borneo;

Acidopria/ spinosiceps/ Dodd $ type.

Type locality

Malaysia, Borneo, Sarawak, Kuching.

Remarks

This species was described from two females referred to as “type” and “cotype”. Two specimens were

found corresponding to the description, one of which had Dodd’s type label and is regarded as the

holotype by original designation. The holotype is mounted on a card and has parts of five legs missing.

This species was placed by Dodd in Acidopria Kieffer, 1913 (now a junior synonym of Basalys) because

of the spines on the head; however, it is a species of Trichopria. It differs from those species of Basalys

with spinose heads in having the spines behind the ocellar triangle rather than in front of it. A non-type

NOTTON D.G., Diapriinae types at The Natural History Museum, London

male was found which agrees closely with the females and it shows that this species belongs to the group

of Trichopria with whorled hairs on the male antenna.

Galesus muscidorum Dodd, 1920: 381.

Valid name

Coptera muscidorum (Dodd, 1920).

Summary of types

Holotype §, BMNH number 9.654, by original designation.

Primary type data

Type/ H.T.; Reared/ from pupa found/ in palpalis/ breeding ground/ similar holes can/ often be found in/

palpalis pupae/ G.D.H.Carpenter/Uganda/Aug 10.1910.; Galesus/ muscidorum/Dodd $.

Type locality

Uganda.

Remarks

The holotype is micropinned, with the left antenna missing beyond a8 and mounted with a fly puparium,

presumably the host. The generic placement established by Masner (1965) is confirmed here.

Aneurhynchus indicus Dodd, 1920: 382.

Valid name

Aneurhynchus indicus Dodd, 1920.

Summary of types

Holotype §, BMNH number 9.634, by original designation.

Primary type data

Type; Kangra Valley/4,500 ft./Apr. 1899/Dudgeon.; Punjab./ G.C.Dudgeon./ 1903-37.; Aneurhynchus/

indicus Dodd $.

Type locality

India, Punjab Province, Kangra Valley.

Remarks

This species was described from a single female. The holotype is mounted on a card and is entire.

Antarctopria coelopae Early, 1978: 209, 210, figs 1-9.

Valid name

Spilomicrus coelopae (Early, 1978) comb. nov.

Summary of types

Paratypes 12 17 SS-

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Remarks

An additional male from the Otago peninsula labelled as a paratype is not a type; specimens from

this locality are listed under “other material” and so are not types. All current Antarctopria species,

A. coelopae,A. campbellanaYoshimoto, 1964, A. diomedeae Early, 1978, A. latigaster Brues in Tillyard,

1920 and A. rekohua Early, 1978 appear to belong to a derived species group of Spilomicrus showing

pronounced wing reduction and the associated suite of characters that go with it. Extra setae on the

gastral tergites are found on wing-reduced representatives of a number of other diapriid genera and

is not considered here a character of generic value. Consequently, these five Antarctopria species are

transferred to Spilomicrus (new combinations), and since Antarctopria latigaster is the type species of

Antarctopria , this genus is now a synonym of Spilomicrus syn. nov.

Antarctopria diomedeae Early, 1978: 210, 223, figs 28-32.

Valid name

Spilomicrus diomedeae (Early, 1978) comb. nov.

Summary of types

Paratypes 2 S3-

Remarks

For notes on the new generic combination, see A. coelopae above.

Spilomicrus pilgrimi Early, 1978: 210, 224, figs 33-38.

Valid name

Spilomicrus pilgrimi Early, 1978.

Summary of types

Paratype §.

Hemilexomyia spinosa Early, 1980: 154,160, figs 12-20, 23-24.

Valid name

Hemilexomyia spinosa Early, 1980.

Summary of types

Paratypes 3 $$,(?.

Entomacis subaptera Early, 1980: 154,163, figs 21-22, 25.

Valid name

Entomacis subaptera Early, 1980.

Summary of types

Paratypes §, f.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Spilomicrus barnesi Early & Horning, 1978: 231, figs 1-6,12.

Valid name

Spilomicrus barnesi Early & Horning, 1978.

Summary of types

Paratypes 6 5 5, 4 SS-

Rostropria simplex Early & Naumann, 1990: 529, 530, figs 1-18.

Valid name

Rostropria simplex Early & Naumann, 1990.

Summary of types

Paratypes 5, $.

Rostropria spiniventris Early & Naumann, 1990: 529, figs 19-38.

Valid name

Rostropria spiniventris Early & Naumann, 1990.

Summary of types

Paratypes §, S-

Trichopria tachinidarum Ferriere, 1933b: 104, fig. lla-b.

Valid name

Trichopria tachinidarum Ferriere, 1933.

Summary of types

Syntype §, BMNH number 9.714. Syntypes 3 $ §, 2 SS-

Primary type data

Type; Java/ Buitenzorg/ xi. 1930/ R.W.Paine/ Par. TJ’; Hyperparasite/ from puparium/ of Tachinid

‘B7 of Tirathaba; Pres, by/ Imp. Inst. Ent./ B.M. 1933-375; Trichopria/ tachinidarum §/ Ch. Ferriere

Type, (syntype $). Java/Buitenzorg/ xi. 1930/R.W.Paine/Par. TJ’; Buitenzorg: xi-30. R.W.P./Diapriid,

Trichopria sp.?; Hyperparasite from puparium of/ Tachinid ‘B’ of Tirathaba.; Pres, by/ Imp. Inst.

Ent./ B.M. 1933-375; Trichopria/ tachinidarum §/ Ch. Ferriere Cotype (syntype §)■ Java/ Buitenzorg/

xi. 1930/ R.W.Paine/ Par. ‘U’; Hyperparasite from puparium of/ Tachinid B' of Tirathaba.; Pres, by/

Imp. Inst. Ent./B.M. 1933-375; Trichopria/ tachinidarum §/ Ch. Ferriere Cotype (syntypes 2 $ §). Java/

Buitenzorg/ xi.1930/ R.W.Paine/ Par. ‘IT; Hyperparasite from puparium of/ Tachinid ‘B’ of Tirathaba.;

Pres, by/ Imp. Inst. Ent./ B.M. 1933-375; Trichopria/ tachinidarum $! Ch. Ferriere Cotype (syntypes

2 SS).

Type locality

Indonesia, West Java, Kota Bogor (as Java, Buitenzorg).

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Remarks

Since Ferriere’s paper was published in two parts, the name of this species was introduced first as a

nomen nudum (Ferriere 1933a: 86) and made available shortly after (Ferriere 1933b: 104). This species

was described from six females and three males, of which four females and two males were found.

Although Ferriere labelled one specimen “type” and five “cotype”, there is no type designation and

all the types are syntypic. Nor does the mention of a type and paratype by Huggert (1982) count as an

original holotype designation since it is not in the original description. All the syntypes are mounted on

card points: the first female is entire; second female entire; third female missing head and gaster; fourth

female missing left hind wing and left fore wing and antenna mounted on a microslide attached to the

same pin; first male missing left antenna beyond a5; second male missing right antenna, gaster and hind

legs, the antenna and dissected gaster are mounted on a separate slide.

Trichopria capensis var. robustior Ferriere, 1935: 329, 338.

Valid name

Trichopria capensis robustior Ferriere, 1935.

Summary of types

Syntypes 17 55,8 55 -

Primary type data

Zululand/ Empangen/ 16.V.1923/ R.H.Harris; Sent by/ Div. of Ent./ Pretoria; primary parasite/ of

Glossina pallidipes/ puparia; Ser. No./ 1186; Trichopria/ capensis Kieffer/ var. robustior, Silv./F.Silvestri

det. (syntype $). Zululand:/Empangen: 16.V.1923/ primary parasite of/ Glossina pallidipes/ puparium:

R.H.Harris; Ser. No. Div. of Entom./ Pretoria/ South Africa/ 1186; Pres, by/ Imp. Bur. Ent./ Brit. Mus./

1929-43 (syntype §). Zululand:/ Empangen: 16.v. 1923/ primary parasite of/ Glossina pallidipes/

puparium: R.H.Harris; Sth. Africa/ Pretoria/ Div. of Entom.; Pres, by/ Imp. Bur. Ent./ Brit. Mus./ 1929-

43; Trichopria/capensisKff7 v. robustior S./ Silvestri det. (syntype §). Zululand:/Empangen: 16.v. 1923/

primary parasite of/ Glossina pallidipes/ puparium: R.H.Harris; Sent by Division of/ Entomology,

Pretoria,/ S.Africa, ser. no. 1186; Pres, by/Imp. Bur. Ent./Brit. Mus./ 1929-43 (syntypes 5 $$, 5 55 )-

Zululand:/Empangen: 16.V.1923/ primary parasite of/ Glossina pallidipes/ puparium: R.H.Harris; Sent

by Division of/ Entomology, Pretoria,/ S.Africa, ser. no. 1186:/ Pres, by/ Imp. Bur. Ent.:/ Brit. Mus./

1929-43 (syntypes 9 $$, 3 55 )-

Type locality

South Africa, KwaZulu-Natal Province, Empangeni (as Natal, Zululand, Empangen).

Remarks

This species was described from seventeen females and eight males. All specimens were located,

together with a fly puparium, presumably the host. No holotype was designated so all are syntypes. All

the syntypes are mounted on card points, and most are entire. This variety was attributed to Silvestri

by Ferriere, but Silvestri never published the name and does not appear to have been responsible for

Ferriere’s description, so it is attributable to Ferriere (1935).

Abothropria lloydi Ferriere, 1935: 329, 338, fig. 3a-c.

Valid name

Lepidopria lloydi (Ferriere, 1935).

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Syntype 5, BMNH number 9.678. Syntypes 10 $ §, 9 33 -

Primary type data

Type; Tanganyika/ Bugambwa/ 7.vi.l933/ J.E.M.Lloyd; Ex Glossina/ palpalis pup.; Pres by/ Imp. Inst.

Ent./ B.M. 1935-231; Abothropria/ lloydi §/ Ch. Ferriere Type (syntype $). Tanganyika/ Bugambwa/

7.vi. 1933/ J.E.M.Lloyd; Ex Glossina/palpalis pup.; Pres by/Imp. Inst. Ent./B.M. 1935-231; Abothropria/

lloydi §/ Ch. Ferriere Cotype (syntypes 7 §§). Tanganyika/ Bugambwa/ 7.vi. 1933/ J.E.M.Lloyd; Ex

Glossina/ palpalis pup.; Pres by/ Imp. Inst. Ent./ B.M. 1935-231; Abothropria/ lloydi c?/ Ch. Ferriere

Cotype (syntypes 3 33 )- Tanganyika T./ Bugambwa, 7.vi.l933/ J.E.M.Lloyd/ Ex Glossina/ palpalis,

pupa; Diapriidae:/ Abothropria/ lloydi cotype/ Ch. Ferriere $ (syntype §). Tanganyika T./ Bugambwa,

7.vi.l933/ J.E.M.Lloyd/ Ex Glossina/ palpalis, pupa; Diapriidae:/ Abothropria/ lloydi cotype/ Ch.

Ferriere 3 (syntype 3 )- Tanganyika T./ Bugambwa, 7.vi.l933/ J.E.M.Lloyd/ Ex Glossina/ palpalis,

pupa; Diapriidae:/ Abothropria/ lloydi cotypes/ Ch. Ferriere 5c? 2$ (syntypes 2 5 33 )-

Type locality

Tanzania, Mara Region, Buganbwe Island (as Tanganyika, Bugambwa).

Remarks

This species is said to have been described from 11 $ $ and 11 33 , of which the following were found:

8 55, 3 33 pinned; 3 $$, 6 33 on slides; and two Glossina Wiedemann, 1830 puparia, presumably

the hosts. These are all syntypes as no holotype was designated. Four other specimens of this species

could not be assessed as they were on loan. The pinned syntypes, 8 $ $ and 3 33 , are mounted on card

points: the first female is entire; the second female has the left antenna missing and the head dented;

the third female is entire; the fourth female has the right antenna mounted separately and is not fully

developed - it appears to have been dissected from the host; the fifth female is entire; the sixth female is

entire and has the head flattened; the seventh female has no wings; the eighth female has the left wings

crumpled; the first male is not fully developed and has a teratological right antenna; the second male is

entire; the third male has the right antenna missing beyond the scape and the left wing crumpled. The

other 3 $ $ and 6 33 syntypes are slide mounted and mostly entire. This species was transferred from

Abothropria Kieffer, 1913 to Lepidopria Kieffer, 1911 by Notton (2004), but it may well deserve a

genus of its own.

Spilomicrus kiefferi Fouts, 1925: 150.

Valid name

Spilomicrus atriclavus Ashmead, 1893.

Summary of types

Paratype $.

Trichopria ( Trichopria ) tabanivora Fouts in Cameron, 1926: 39.

Valid name

Trichopria tabanivora Fouts in Cameron, 1926.

Summary of types

Paratypes §, 3 -

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Loxotropa nigrescens Fouts, 1927: 165,169, fig. 1.

Valid name

Basalys nigrescens (Fouts, 1927).

Summary of types

Paratypes 2 $ $ •

Remarks

Two paratypes have come to the BMNH via the Whittaker collection.

Paramesius laetus Fouts, 1927: 170.

Valid name

Paramesius laetus Fouts, 1927.

Summary of types

Paratype $.

Remarks

The paratype has come to the BMNH via the Whittaker collection.

Labolips innupta Haliday, 1857: 173, pi. 10, figs 1-20.

Valid name

Labolips innupta Haliday, 1857.

Summary of types

Syntype §, BMNH number 9.905.

Primary type data

59-4; Labolips/ extincta §; Britain/ purchased from/ T.V.Wollaston/ named by Haliday/ BM 1859.4;

syntype §/Labolips/ innupta Haliday/ det. D.Notton, 2003.

Type locality

Ireland, County Down, between Belfast and Holywood, Knocknagoney, Clifden House.

Remarks

Haliday evidently collected a long series of this species, much of which he dispersed before making a

formal description in an effort to discover whether it was previously known or described (Haliday 1857).

Hence, as well as the twelve syntypes in Dublin, syntypic material of Labolips innupta is also to be found

in the collections of the Natural History Museum, London (one female, via T.V. Wollaston) and in the

Dale Collection at the Hope Department of Entomology, Oxford University Museum of Natural History

(two females). The London and Oxford syntypes are all labelled “ Labolips extincta ”, a manuscript name

Haliday used for this species before settling on the published name. There may also be syntypes in the

Forster collection at the Naturhistorisches Museum Wien, Wien, Austria, as the description of Labolips

Forster, 1856 was made by Forster from two females sent to him by Haliday (Forster 1856: 125; Notton

NOTTON D.G., Diapriinae types at The Natural History Museum, London

& O’Connor 2004). The London syntype is mounted on a card and is set with the wings spread in a

similar fashion to the other syntypes.

Trichopria stomoxydis Huggert, 1977: 205, figs 1-7.

Valid name

Trichopria stomoxydis Huggert, 1977.

Summary of types

Holotype §, BMNH number 9.801, by original designation. Paratypes 20 $$, 8 33 -

Primary type data

Holo-/ type; Uganda/ C.I.B.C./25.vii.1975/ C.I.E.A8124; ex pupa of/ Stomoxys nigra/ from cut/ elephant

grass; $ Trichopria sp./ nr. inconspicua?/ det. Z.Boucek, 1975; Holotypus/ Trichopria §/ stomoxydis/

n. sp./L.Huggert-76.

Type locality

Uganda.

Remarks

The holotype is mounted on a card point, the left hind wing is missing and the left fore wing and left

antenna are mounted on a microslide on the same pin.

Trichopria pseudosaxatilis Huggert, 1977: 214, figs 32-37.

Valid name

Trichopria pseudosaxatilis Huggert, 1977.

Summary of types

Holotype 3, BMNH number 9.802, by original designation.

Primary type data

5; Silhouette, ‘08/ Seychelles Exp.; Diapria/ saxatilis/ J.J.Kieffer/ Paratype; Holotypus/ Trichopria 31

pseudosaxatilis/ n. sp./ L.Huggert-76.

Type locality

Seychelles, Silhouette, high country near Pot-a-eau.

Remarks

This species was described from a single male. This holotype is also a paralectotype of Diapria saxatilis

Kieffer, 1912 q.v. It is possible to be more precise about the type locality than Huggert (1977) based on

information given about this specimen by Kieffer (1912a). The holotype is mounted on a card; the left

wings and left antenna are mounted on a microslide on the same pin.

Trichopria nirva Huggert, 1977: 215, figs 38-43.

Valid name

Trichopria nirva Huggert, 1977.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Summary of types

Holotype §, BMNH number 9.803, by original designation.

Primary type data

145; Mahe, ’08-9./ Seychelles Exp.; Diapria/ saxatilis/ J.J.Kieffer/ Paratype; Holotypus/ Trichopria §/

nirva n. sp./ L.Huggert-76.

Type locality

Seychelles, Mahe.

Remarks

This species was described from a single female. The holotype is also a paralectotype of Diapria

saxatilis q.v. The holotype is mounted on a card; the right antenna beyond the scape is missing and the

left antenna and wings are mounted on a microslide on the same pin.

Trichopria euvulgaris Huggert, 1979: 19, figs 22-27.

Valid name

Trichopria euvulgaris Huggert, 1979.

Summary of types

Holotype S, BMNH number 9.806, by original designation.

Primary type data

Mahe, ’08-9./ Seychelles Exp./ 95; type; Type/ H.T.; Seychelles Islands./ Percy Sladen Trust/

Expedition./ 1913-170.; Rhopalopria/ vulgaris/ J.J.Kieffer/ Type; Holotypus/ Trichopria $1 euvulgaris/

n. sp./ L.Huggert -77.

Type locality

Seychelles, Mahe.

Remarks

This species was described from a single male. The holotype is also a paralectotype of Rhopalopria

vulgaris Kieffer, 1912 q.v. The holotype is mounted on a paper-covered cork block with the left hind

wing missing and the left antenna and fore wing mounted on a microslide.

Trichopria ensia Huggert, 1979: 23, figs 35-40.

Valid name

Trichopria ensia Huggert, 1979.

Summary of types

Holotype §, BMNH number 9.807, by original designation.

Primary type data

Mahe, ’08-9/ Seychelles Exp/ 45; Seychelles Islands/ Percy Sladen Trust/ Expedition./ 1913-

170; Paratype; Diapria/ seychellensis/ J.J.Kieffer/ Paratype; Holotypus $/ Trichopria/ ensia n. sp./

L.Huggert-77.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Type locality

Seychelles, Mahe.

Remarks

This species was described from a single female. The holotype is also a paralectotype of Diapria

seychellensis Kieffer, 1912 q.v. The holotype is mounted on a paper-covered cork block, the left antenna

is missing beyond scape, and the left wings and right antenna are mounted on a microslide on the same

pin.

Trichopriapercyi Huggert, 1979: 24, figs 41-46.

Valid name

Trichopria percyi Huggert, 1979.

Summary of types

Holotype BMNH number 9.808, by original designation.

Primary type data

Mahe, ’08-9/ Seychelles Exp/ 97; Seychelles Islands./ Percy Sladen Trust/ Expedition./ 1913-

170.; Paratype; Diapria/ seychellensis/ J.J.Kieffer/ Paratype; Holotypus c V Trichopria/ percyi n. sp./

L.Huggert-77.

Type locality

Seychelles, Mahe.

Remarks

Trichopria percyi was described from a single male. The holotype is also a paralectotype of Diapria

seychellensis q.v. The holotype is mounted on a paper-covered cork block, the left mid leg is mounted

separately, and the left antenna and wings are mounted on a microslide on the same pin.

Townesella marjoriae Huggert & Masner, 1983: 67, 70, 71, figs 14,16,18-20.

Valid name

Townesella marjoriae Huggert & Masner, 1983.

Summary of types

Paratype §.

Trichopria myrmicae Huggert & Masner, 1983: 68, 77, figs 39-49.

Valid name

Trichopria myrmicae Huggert & Masner, 1983.

Summary of types

Paratype $.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Neurogalesus carinatus Kieffer, 1907: 298.

Valid name

Neurogalesus carinatus Kieffer, 1907.,

Summary of types

Holotype BMNH number 9.673, by monotypy.

Primary type data

Type; Mackay/ 3.01; Queensland./ R.E.Turner./ 1907-94; Neurogalesus/ carinatus $/ type. Kieff./

named by/ J.J.Kieffer.

Type locality

Australia, Queensland, Mackay.

Remarks

This species was described from an unspecified number of females; however, Kieffer describes the

scutellum as “vom beschadigt”, i.e., damaged in front, a rather unusual thing to say. The specimen

recognised as the type has been pinned at exactly this point, destroying the basal scutellar pit. Had

Kieffer seen any other specimens it is likely they would not have been damaged in precisely this place

and he would have described the distinctive bifoveate scutellar pit, so it is considered that he saw only

this specimen; hence it is a holotype by monotypy. The holotype is micropinned through the mesosoma

and has the head missing. Despite what Kieffer says about its resemblance to a Galesus, this species

belongs to the Spilomicrini.

Xyalopria nigriceps Kieffer, 1907: 300.

Valid name

Trichopria fluminis nom. nov.

Summary of types

Syntype §, BMNH number 9.690.

Primary type data

Type; Rio/ 59-2lx; Xyalopria/ nigriceps/ Kieffer; named by J.J.Kieffer.

Type locality

Brazil, Rio de Janeiro.

Remarks

This species was described from an unspecified number of females, of which one was found. The syntype

is mounted on a card, with part of the right mid tarsus missing and the left mid tarsus mounted separately.

This species was included in Megaplastopria by Arias-Penna (2003); however, it does not belong there.

While it is large, elongate and has a long, slender propodeal spine, it lacks significant characters of

Megaplastopria such as the occipital spine and carina, grooved scape and flared corners of the metasomal

tergites, and is here transferred to Trichopria instead. Since the new combination T. nigriceps (Kieffer,

1907) is a junior secondary homonym of T. nigriceps (Ashmead in Riley, Ashmead & Howard, 1894),

Kieffer’s species is given the new replacement name Trichopria fluminis nom. nov. The word fluminis ,

NOTTON D.G., Diapriinae types at The Natural History Museum, London

genitive of flumen (Latin for river) refers to the type locality Rio de Janeiro. Since X nigriceps Kieffer is

the type species of Xyalopria, Xyalopria is removed from synonymy with Megaplastopria and becomes

a junior synonym of Trichopria syn. nov.

Galesuspunctulatus Kieffer, 1907: 301.

Valid name

Copterapunctulata (Kieffer, 1907).

Summary of types

Syntype BMNH number 9.653.

Primary type data

Type; Pt/ Natal/ 56-63; Galesus/ punctulatus/ Kieffer/ named by J.J.Kieffer; BMNH(E) 1856-63/ South

Africa:/ Port Natal:/ coll. Plant:/ purch. Stevens.

Type locality

South Africa, Durban (as Port Natal).

Remarks

This species was described from an unspecified number of males, of which one was found. The syntype

is micropinned, the left fore wing is mounted separately and the metasoma, left hind wing and part of

the right mid tarsus are missing.

Galesus walked Kieffer, 1907: 301.

Valid name

Psilus fuscipennis (Curtis, 1831) syn. nov.

Summary of types

Lectotype BMNH number 9.919, here designated.

Primary type data

Alten/ Finnmark/ F.Walker/ Feb.37/ 201.

Type locality

Noted as “wohl England”, i.e., probably from England, but actually from Norway, Finnmark, Alta.

Remarks

This species was described from an unspecified number of males, said to be in the British Museum

by Kieffer. One male specimen was found which agreed with the description, except that Kieffer says

the collector was J. Walker, whereas the label says “F. Walker” although the “F.” is written like a “J.”

Kieffer says the specimens were probably from England, suggesting that the label was difficult to read

and that he assumed the specimen(s) was English because it was collected by Walker. The label on the

type is difficult to read but actually says “Alten, Finnmark”, no doubt a specimen collected by Francis

Walker on his 1836-7 tour of Scandinavia (Christy & Walker 1837). The close agreement with the

description, Kieffer’s mistake in reading the “J.” for “F.” because of the peculiar style of writing and the

near illegibility of the label which caused him to guess the locality (erroneously) convince me that this

European Journal of Taxonomy 75 : 1-123 ( 2014 )

is a type. This specimen belongs to Psilus fuscipennis following the interpretation of Nixon (1980) and

the two species are synonymized here. Alectotype is designated to stabilise the synonymy. The lectotype

is mounted on a card and is missing the left fore leg and the tips of three tarsi.

Galesus gracilipes Kieffer, 1907: 302.

Valid name

Psilus cornutus Panzer, 1801.

Summary of types

Syntype BMNH number 9.648.

Primary type data

Type; Galesus/ gracilipes/ Kieffer/ Named by/ J.J.Kieffer; milleri/ [illeg.].

Type locality

England.

Remarks

This species was described from an unspecified number of males. The syntype is mounted on a card and

has the left wings missing.

Loxotropapedestris Kieffer, 1907: 302.

Valid name

Basalys parvus Thomson, 1858.

Summary of types

Holotype §, BMNH number 9.728, by monotypy.

Primary type data

Uxbridge; Type; Loxotropa/ pedestris/ Kieffer/ Named by/ J.J.Kieffer.

Type locality

England, Uxbridge.

Remarks

This species was described from an unspecified number of females, but Kieffer (1911b: 926) says “type

au Musee de Londres”, so evidently there was only one type, thus a holotype by monotypy. The holotype

is mounted on a card and is entire.

Phaenopria halterata Kieffer, 1909: 381.

Valid name

Trichopria halterata (Kieffer, 1909).

Summary of types

Lectotype S, BMNH number 9.715, designated by Notton (1995).

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Galloway; Type; 478; Cameron Coll./ 1910-55.; halterataK.

Type locality

Scotland, Galloway.

Remarks

This species was described from an unspecified number of males. The lectotype agrees well with the

description, although it has a weak (and easily overlooked) scutellar pit. The lectotype is mounted on a

card, left fore leg mounted separately, most tarsi broken.

Phaenopria cameroni Kieffer, 1909: 382.

Valid name

Trichopria cameroni (Kieffer, 1909).

Summary of types

Lectotype $, BMNH number 9.717, designated by Notton (1995).

Primary type data

M; Type; suspecta Nees; 231; Cameron Coll./ 1910-55.; cameroni K.

Type locality

Probably England, Micldeham.

Remarks

This species was described from an unspecified number of females and it is quite likely that the type

series contained at least two specimens from Clyde at Newton and Micldeham (Cameron 1910). A single

syntype was located which had been designated as the lectotype by Notton (1995). Since the lectotype

has an “M” written on the mount, it is probably from Mickleham in England. Although Kieffer gave the

type locality as Scotland, there is no reason to exclude the Mickleham specimen from the type series

because Kieffer is known to have misinterpreted many of the localities on Cameron’s specimens. The

lectotype is mounted on a card and has become twisted on its mount, breaking the antennae, and a 12 of

the right ante nn a is missing.

Trichopria inermis Kieffer, 1909: 386.

Valid name

Trichopria nigra (Nees, 1834).

Summary of types

Lectotype $, BMNH number 9.705, designated by Notton (1995).

Primary type data

Thornhill/ llix.; Type; 238.; Cameron Coll./ 1910-55.; inermis K.; Holotype; $ Lectotype/ Trichopria/

inermis/Kieffer, 1909/ des. D.Notton, 1995.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Type locality

Scotland, Thornhill.

Remarks

This species was described from an unspecified number of males of which one specimen was found. The

lectotype is mounted on a card and is entire.

Idiotypa nigriceps Kieffer, 1909: 386.

Valid name

Idio typa n igriceps Kie ffer, 1909.

Summary of types

Lectotype §, BMNH number 9.746, here designated.

Primary type data

Lochawe; Type; 230.; Cameron Coll./ 1910-55.; nigriceps K.

Type locality

Scotland, Lochawe village.

Remarks

This species was described from an unspecified number of females. A single female syntype was found

which is designated here as lectotype to ensure objective synonymy with Idiotypa nigriceps Kieffer,

1911 q.v. The lectotype is mounted on a card and is entire.

Paramesius inchoatus Kieffer, 1911a: 756.

Valid name

Paramesius rufipes (Fonscolombe, 1832).

Summary of types

Lectotype §, BMNH number 9.813, designated by Nixon (1980). Paralectotype §.

Primary type data

Q.B1.; 469.; Paramesius/ inchoatus, K.; Cameron Coll./ 1910-302.; Determined by/Dr. Kieffer.; Selected

as/ type of/ Paramesius/ inchoatus K/ by G.E.J.N. 1976.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of females. Two specimens were located, of

which one had been designated as a lectotype by Nixon (1980). The lectotype is mounted on a card and

is entire.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Paramesius subinermis Kieffer, 1911a: 757.

Valid name

Paramesius rufipes (Fonscolombe, 1832).

Summary of types

Lectotype S, BMNH number 9.812, designated by Nixon (1980). Paralectotype <$■

Primary type data

Bishop; Bishopton; 446.; Paramesius/ subinermis, K.; Cameron Coll./ 1910-302.; Determined by/ Dr.

Kieffer; selected as/ type of/ Paramesius/ subinermis/ K/ by G.E.J.N. 1976.

Type locality

Scotland, Bishopton.

Remarks

This species was described from an unspecified number of males. Three specimens were found, of

which one had been designated as lectotype by Nixon (1980). The lectotype is mounted on a card and

parts of the right mid and hind tarsi are disarticulated.

Paramesius angustipennis Kieffer, 1911a: 762.

Valid name

Paramesius brachypterus (Thomson, 1858).

Summary of types

Lectotype $, BMNH number 9.929, here designated.

Primary type data

Mugd/ 26-7; Mugdock; 455.; Paramesius/ angustipennis; Cameron Coll./ 1910-302.; determined by/ Dr.

Kieffer.

Type locality

Scotland, Mugdock.

Remarks

This species was described from at least two females, of which one syntype was found. Macek (2002)

synonymized this species with P. brachypterus without seeing the type. The syntype belongs to

P. brachypterus following the interpretation of this species by Nixon (1980) and Macek (2002) and

Macek’s synonymy is confirmed. A lectotype is designated to stabilise the application of the name and

its synonymy. The lectotype is mounted on a card, with the left fore wing missing. The species epithet

augustipennis is an incorrect original spelling, and is to be corrected (Article 32.5.1), since there is clear

evidence in the original publication of an inadvertent error, i.e., in the description Kieffer says “Ailes

mince... ”, i.e., wing narrow which when formed into a Latin name from angustus (narrow or straight) +

penna (a wing) would become angustipennis not augustipennis.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Paramesius nigricornis Kieffer, 1911a: 764.

Valid name

Paramesius rufipes (Fonscolombe, 1832).

Summary of types

Syntype §, BMNH number 9.661.

Primary type data

Galloway; Type; 265.; Cameron Coll./ 1910-55.; nigricornis K.

Type locality

Scotland and Norway, Solum, Overhaiden.

Remarks

This species was described from an unspecified number of females, of which one was found. The syntype

is mounted on a card and has the left hind wing disarticulated.

Paramesius cameroni Kieffer, 1911a: 767.

Valid name

Paramesius brachypterus (Thomson, 1858).

Summary of types

Lectotype 3, BMNH number 9.927, here designated. Paralectotypes 2 S3-

Primary type data

Mugdock/ 9-8; Mugdock; 445.; Paramesius/ cameroni, K.; Cameron Coll./ 1910-302.; Determined by/

Dr Kieffer.

Type locality

Scotland, Mugdock.

Remarks

This species was described from at least three males. Three syntypes were located, of which one is

designated a lectotype. Macek (2002) synonymized this species with P. brachypterus without seeing

the type. The lectotype belongs to P. brachypterus following the interpretation of this species by Nixon

(1980) and Macek (2002), so Macek’s synonymy is confirmed. The lectotype is designated to stabilise

the application of the name and its synonymy. The lectotype is mounted on a card and is entire.

Paramesius dentatus Kieffer, 1911a: 768.

Valid name

Paramesius rufipes (Fonscolombe, 1832).

Summary of types

Syntype 3, BMNH number 9.662.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Mug/ 26-7; Type; 253.; Cameron Coll./ 1910-55.; dentatus K.

Type locality

Scotland.

Remarks

Described from an unspecified number of males. The syntype is mounted on a card and is entire.

Spilomicrus hemipterus var. pedissequus Kieffer, 1911a: 774.

Valid name

Spilomicrus hemipterus Marshall, 1868.

Summary of types

Syntype $, BMNH number 9.782.

Primary type data

(3.; Type/ of var.; Spilomicrus/ hemipterus,/ var.-/ pedissequus/ Kief. §.; ?Aneurhynchus sp./ In nest

of Lasius/ fuliginosus/ Oxshott. p/ (15th-) 12-4-01./ 27; Holotype of/ Spilomicrus/ pedisequus K./

N.D.M.Fergusson, 1974.

Type locality

England.

Remarks

This species was described from an unspecified number of females. One specimen was found which

agreed well with the description except for having the basal pit bifoveate. The syntype is mounted on a

card, missing the tips of both antennae and the left mid leg; four tarsi are broken but parts of two of them

are still present. It appears that pedissequus is the correct original spelling, as there is no evidence in

the text of the original description that it is an inadvertent error (Article 32.5.1). The epithet pedisequus

is therefore an unjustified emendation attributable to Kieffer (1916) and is a junior objective synonym.

Spilomicrus integer var. variicornis Kieffer, 1911a: 777.

Valid name

Spilomicrus integer Thomson, 1858.

Summary of types

Syntype §, BMNH number 9.644.

Primary type data

[illeg.]/ Glen. 20-5; Type; 465.; Cameron Coll./ 1910-55.; integer K. v.; integer var./ variicornis, K.

Type locality

Scotland.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Remarks

This species was described from an unspecified number of females, of which one was found. The syntype

is mounted on a card and is entire.

Spilomicrus crassiclavis Kieffer, 1911a: 788.

Valid name

Spilomicrus crassiclavis Kieffer, 1911.

Summary of types

Lectotype $, BMNH number 9.645a, designated by Notton (1999).

Primary type data

Mugdock/ 11-7; Cameron Coll./ 1910-302.; Mugdock; Determined by/ Dr Kieffer.; Spilomicrus/

crassiclavis, K.; Spilomicrus/ formosus Janss./ G.Nixon det. 1972; § Lectotype/ Spilomicrus/

crassiclavis/Kieffer, 1911: 788/ des. D.Notton, 1999.

Type locality

Scotland, Mugdock.

Remarks

This species was described from an unspecified number of females and males. The lectotype is mounted

on a card, left antenna missing, right fore wing disarticulated. Spilomicrus crassiclavis was previously

confused with Spilomicrus formosus Jansson, 1942 by Nixon (1980), but this is a different species

(Notton 1999).

Spilomicrus annulicornis Kieffer, 1911a: 788.

Valid name

Spilomicrus annulicornis Kieffer, 1911.

Summary of types

Syntype $, BMNH number 9.642. Syntype §.

Primary type data

27-69/Mickleham; Type; 249.; Cameron Coll./1910-55.; annulicornis K. (syntype $). 275.; Spilomicrus/

annulicornis, K.; Cameron Coll./ 1910-302.; Determined by/ Dr. Kieffer (syntype $).

Type locality

Scotland and England, Mickleham.

Remarks

This species was described from an unspecified number of females. Kieffer gave the type locality as

Scotland; however, one of the syntypes is labelled almost illegibly “Mickleham” and the other has no

locality label. Since the specimens agree with the description, there seems no reason to exclude either

syntype from the type series because Kieffer is known to have misinterpreted many of the localities on

Cameron’s specimens. Both syntypes are mounted on cards, the one from Mickleham has lost the tip of

its right hind tarsus, the other has lost the tip of its left hind tarsus.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Spilomicrus minor Kieffer, 1911a: 791.

Valid name

Paramesius rufipes (Fonscolombe, 1832).

Summary of types

Syntype §, BMNH number 9.664. Syntype §.

Primary type data

Dairy; Type; 242.; Cameron Coll./ 1910-55; minor K. (syntype $). 55-69/ Box Hill; 241; Cameron

Coll./ 1910-55 (syntype §).

Type locality

Scotland and England, Box Hill.

Remarks

This species was described from an unspecified number of females; two females were found which

agreed with the description. Although the published type locality is Scotland, there is no reason to

exclude the English specimen from the type series because Kieffer is known to have misinterpreted

many of the localities on Cameron’s specimens. Both syntypes are mounted on cards; the one from

Scotland is entire, the one from Box Hill has the tip of the left ante nn a missing.

Spilomicrus tripartitus Kieffer, 1911a: 792.

Valid name

Spilomicrus stigmaticalis Westwood, 1832.

Summary of types

Syntype $, BMNH number 9.643.

Primary type data

[illeg.]/ viii; Type; 248.; Cameron Coll./ 1910-55.; tripartitus K.

Type locality

Scotland; England; France, Mesnil-le-Roy and Versailles.

Remarks

This species was described from at least four specimens including both sexes. The syntype agrees well

with the description and while the notauli are percurrent, it is worth noting they are deep posteriorly and

weak anteriorly. The syntype is mounted on a card and is entire.

Idiotypa nigriceps Kieffer, 1911a: 814.

Valid name

Idio typa n igriceps Kie ffer, 1909.

Summary of types

Lectotype §, BMNH number 9.746, here designated.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Primary type data

Lochawe; Type; 230.; Cameron Coll./ 1910-55.; nigriceps K.

Type locality

Scotland, Lochawe village.

Remarks

This species was described from an unspecified number of females, apparently including the type series

of Idiotypa nigriceps Kieffer, 1909 with additional material from England. The specimen which is the

lectotype of I. nigriceps Kieffer, 1909 is also designated here as the lectotype of I. nigriceps Kieffer,

1911 to create objective synonymy. The lectotype of/, nigriceps Kieffer, 1911 is mounted on a card and

is entire.

Aneurhynchus obliquus Kieffer, 1911a: 828.

Valid name

Aneurhynchus obliquus Kieffer, 1911.

Summary of types

Syntype BMNH number 9.633.

Primary type data

Dumf; Type; 437.; Cameron Coll./ 1910-55.; obliquus K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. The syntype is mounted on a card and

has the left hind tarsus mounted separately.

Galesus ( Schizogalesus ) mayeti var. hispanicus Kieffer, 1911a: 845.

Valid name

Copter a hispanica (Kieffer, 1911).

Summary of types

Syntype §, BMNH number 9.651.

Primary type data

Giberalter [sic]; Type; Gibraltar; 401.; Cameron Coll./ 1910-55.; mayeti K. v./ hispanicus K.

Type locality

Gibraltar.

Remarks

This species was described from an unspecified number of females. The syntype is mounted on a card

and the left fore tarsus is disarticulated.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Galesus (G.)parvulus Kieffer, 1911a: 853.

Valid name

Psilusparvulus (Kieffer, 1911).

Summary of types

Syntype 3, BMNH number 9.650.

Primary type data

Clober; Type; 407.; Cameron Coll./ 1910-55.; parvulus K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. The syntype is mounted on a card and

is entire.

Galesus (6.) foersteri var. nigricornis Kieffer, 1911a: 856.

Valid name

Psilus fuscipennis (Curtis, 1831) syn. nov.

Summary of types

Lectotype 3, BMNH number 9.922, here designated. Paralectotypes 2 S3-

Primary type data

Glen Lyon; Type; 408.; Cameron Coll./ 1910-55.; atricornis.

Type locality

Scotland, Glen Lyon.

Remarks

The original description was based on an unspecified number of males, of which three were found. All

the syntypes agree well with the description. Kieffer evidently changed the name between labelling the

specimens and writing the description; the connection is clear from the names: atricornis and nigricornis

are closely related, since ater and niger are both Latin words meaning black. The lectotype is mounted

on a card and is entire. This specimen belongs to Psilus fuscipennis following the interpretation of Nixon

(1980) and the two species are synonymized here.

Galesus (6.) bispinosus Kieffer, 1911a: 865.

Valid name

Psilus fuscipennis (Curtis, 1831).

Summary of types

Syntype §, BMNH number 9.920.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Primary type data

Scar/ Scarbr; Type; 406.; Cameron Coll./ 1910-55.; bispinosus K.; Galesus/ bispinosus/Holotype Kieff./

G.Nixon 1977.

Type locality

Probably England, Scarborough.

Remarks

This species was described from an unspecified number of females. Despite Nixon’s label, this specimen

cannot be a holotype as Kieffer did not designate a holotype. The type locality is most probably Scarborough

in England, based on the data reported by Cameron (1910), rather than Scotland. The syntype is mounted

on a card, the left antenna missing beyond a4, wings crumpled and right fore wing broken.

Galesus (6.) cameroni Kieffer, 1911a: 865.

Valid name

Psilus fuscipennis (Curtis, 1831).

Summary of types

Syntype $, BMNH number 9.649. Syntype §.

Primary type data

[illeg.]; Type; 403.; Cameron Coll./ 1910-55.; cameroni K. (syntype §). Mickleham/ 31-76; 404.;

Cameron Coll./ 1910-55 (syntype §).

Type locality

Probably England.

Remarks

This species was described from an unspecified number of females. The syntypes agree well with the

description; although Kieffer says the head has five anterior teeth, this probably means four plus the

median ocellus which protrudes anteriorly; also, the “arete en avant jusque entre les antennes” is weak

but observable from the side. Although the published type locality is Scotland, it appears from the

locality (Caterham) given by Cameron (1910) that Kieffer saw English specimens, so English specimens

have not been excluded from the type series. In any case, Kieffer is known to have misinterpreted many

of the localities on Cameron’s specimens. Both syntypes are mounted on cards; the syntype numbered

403 has the metasoma missing; syntype 404 is entire.

Phaenopria halterata Kieffer, 1911a: 880.

Valid name

Trichopria halterata (Kieffer, 1909).

Summary of types

Lectotype $, BMNH number 9.715, here designated.

Primary type data

Galloway; Type; 478; Cameron Coll./ 1910-55.; halterata K.

Type locality

Scotland, Galloway.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

This species was described from an unspecified number of males, apparently from the same type series

as for P. halterata Kieffer, 1909. The type agrees well with the description, although it has a weak

scutellar pit. A lectotype is designated here; this is the same specimen as the lectotype of P. halter ata

Kieffer, 1909, creating objective synonymy between the two names. The lectotype is mounted on a card,

left fore leg mounted separately, and most tarsi broken.

Phaenopria subimpressa Kieffer, 1911a: 880.

Valid name

Trichopria subimpressa (Kieffer, 1911).

Summary of types

Lectotype $, BMNH number 9.716, designated by Notton (1995).

Primary type data

13-69/ Hounslow; Type; Diapria tritoma; 232.; Cameron Coll./ 1910-55.; subimpressa K.

Type locality

England, Hounslow.

Remarks

This species was described from an unspecified number of females. Although the published type locality

is Scotland, Cameron (1910) says he has no definite locality for this species, and the specimen label

says Hounslow in England. There is no reason to exclude this specimen from the type series because

Kieffer is known to have misinterpreted many of the localities on Cameron’s specimens. The lectotype

is mounted on a card, with its head disarticulated, right antenna with a4-10 missing, mesosoma broken,

propodeum and metasoma disarticulated from the rest of the mesosoma, left fore wing folded and left

hind wing missing.

Phaenopria cameroni Kieffer, 1911a: 881.

Valid name

Trichopria cameroni (Kieffer, 1909).

Summary of types

Lectotype §, BMNH number 9.717, here designated.

Primary type data

M; Type; suspecta Nees; 231; Cameron Coll./ 1910-55.; cameroni K.

Type locality

Probably England, Mickleham.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Remarks

This species was described from an unspecified number of females and apparently from the same

type series as P. cameroni Kieffer, 1909 q.v. A single syntype was located and is designated here as a

lectotype. The lectotype ofE cameroni Kieffer, 1911 is also the lectotype of P. cameroni Kieffer, 1909,

creating objective synonymy. Type locality and condition of lectotype as for P. cameroni Kieffer, 1909.

Tetramopria donisthorpei Kieffer, 1911a: 891.

Valid name

Tetramopria cincticollis Wasmann, 1899.

Summary of types

Lectotype 3, BMNH number 9.892, designated by Notton (1995).

Primary type data

c?/ Tetramopria/ Donisthorpei/ Whitsand/ Bay/ 14.iv. 1909; Type; Ex coll./ Donisthorpe/ B.M. 1934-4;

Assoc, with/ Tetramorium/ caespitum; Tetramopria <31 donisthorpei Kief.; Lectotype Tetramopria/

donisthorpei/Kieffer, 1911/ det. D.G.Notton, 1992; $ Tetramopria/ cincticollis/ det. D.Notton, 1993.

Type locality

England, Cornwall, Whitsand Bay.

Remarks

This species was described from an unspecified number of females and males from Whitsand Bay and

two females and a male from Hohscheid. The lectotype is mounted on a card and is entire. An additional

female labelled as a cotype is not syntypic since it was collected in June; the original date can be seen

on the mount.

Tetramopria donisthorpei var. femoralis Kieffer, 1911a: 892.

Valid name

Tetramopria cincticollis Wasmann, 1899.

Summary of types

Lectotype $, BMNH number 9.893, designated by Notton (1995). Paralectotypes 2$ §.

Primary type data

c?/ Tetramopria/ femoralis/ Whitsand/ Bay/ 15.iv. 1909; Type; Ex coll./ Donisthorpe/ B.M. 1934-4;

Assoc, with/ Tetramorium/ caespitum.; Tetramopria/ donisthorpei var./ femoralis Kief.; Lectotype

(57 Tetramopria donisthorpei var./ femoralis Kieffer, 1911/ det. D.G.Notton, 1992; $ Tetramopria/

cincticollis/ det. D.Notton, 1993.

Type locality

England, Cornwall, Whitsand Bay.

Remarks

This species was described from an unspecified number of females and males. The lectotype is mounted

on a card and the gaster, right hind wing and left mid leg are missing.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Planopriapedestris Kieffer, 1911a: 896.

Valid name

Platymischus dilatatus Westwood, 1832.

Summary of types

Syntype §, BMNH number 9.764.

Primary type data

Planopria/ pedestris/ Luccombe/ Chine/ 1 .viii. 1909; Ex coll./ Donisthorpe/ B.M. 1934-4; Assoc, with/

Lasius niger; Planopria/ pedestris Kief.; Type.

Type locality

England.

Remarks

This species was described from an unspecified number of females. The syntype is mounted on a card

and is entire.

Bcisalys collaris Kieffer, 1911a: 903.

Valid name

Basalys collaris Kieffer, 1911.

Summary of types

Syntype §, BMNH number 9.748.

Primary type data

York; Type; 225.; Cameron Coll./ 1910-55.; collaris K.

Type locality

Probably England, York.

Remarks

This species was described from an unspecified number of females. One syntype was found which

agrees well with the description; however, the collar is not red edged whitish as stated by Kieffer “collier

de feutrage roux borde de blanchatre”, but the nucha and pronotal cuticle underneath it is reddish and

the hairs of the collar are whitish, the overall effect resembling Kieffers description. Although Scotland

is the published type locality, Cameron (1910) noted a specimen from York and so this specimen is not

excluded from the type series. The syntype is mounted on a card and is entire.

Basalys rufiscapus Kieffer, 1911a: 909.

Valid name

Basalys formicarum (Kieffer, 1911).

Summary of types

Syntype S, BMNH number 9.743.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Primary type data

Mull; Type; 243.; Cameron Coll./ 1910-55.; rafiscapus K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. The syntype is mounted on a card, and

is missing its right fore wing and right antenna beyond a9.

Loxotropa cursitans Kieffer, 1911b: 916.

Valid name

Basalys pedisequa (Kieffer, 1911) syn. nov.

Summary of types

Syntype §, BMNH number 9.921.

Primary type data

Lade at/ Cadder/ 2-8; 209.; Cameron Coll./ 1910-55.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of females. The specimen formerly regarded

as the type of L. cursitans by Masner (1965) and Nixon (1980) is actually a syntype of L. cursitans var.

alacris Kieffer q.v. A different specimen was found which is considered here to be a true syntype of the

nominate L. cursitans. This type has much shorter wings (Kieffer says “remplacees par une ecaille”),

and a shorter head, and agrees better with Kieffer’s description. This syntype is mounted on a card

and is entire. Reinterpretation of the true syntype of L. cursitans shows that it is the same as Nixon’s

(1980) interpretation of Basalys pedisequa. Consequently it is removed from synonymy with B. parvus

Thomson, 1858.

Loxotropa cursitans var. alacris Kieffer, 1911b: 916.

Valid name

Basalys parvus Thomson, 1858.

Summary of types

Syntype §, BMNH number 9.729. Syntypes 2$ §.

Primary type data

Eccles [illeg.]; Type; 208.; Cameron Coll./ 1910-55.; cursitans/ Kieff (syntype §). Cladich; 210.;

Cameron Coll./ 1910-55 (syntype §)• 212; Cameron Coll./ 1909-182 (syntype §).

Type locality

Scotland.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

This variety was described from an unspecified number of females. Three syntypes were found, of which

the one numbered as type 9.729 had erroneously been regarded as the type of L. cursitans q.v. by Masner

(1965) and Nixon (1980). All syntypes have the wings reaching the basal quarter of the metasoma, a

gradually expanded triangular marginal vein, and the head not higher than long and thus agree better

with Kieffer’s description of L. cursitans var. alacris. All three syntypes are mounted on cards; syntype

number 212 is missing part of the left antenna and part of the left fore tarsus and the other two syntypes

are entire. The synonymy of this species with Basalys parvus Thomson, 1858, following Nixon’s (1980)

interpretation of B. parvus , is confirmed here.

Loxotropa macroptera Kieffer, 1911b: 929.

Valid name

Basalys macroptera ( Kieffer, 1911).

Summary of types

Syntype §, BMNH number 9.730.

Primary type data

Loch Awe; Type; 227.; Cameron Coll./ 1910-55.; macroptera K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of females. The syntype is mounted on a card

and is entire.

Loxotropa sulcata Kieffer, 1911b: 931.

Valid name

Basalys macroptera (Kieffer, 1911).

Summary of types

Syntype $, BMNH number 9.731. Syntype <$■

Primary type data

Cladich; Type; 456.; Cameron Coll./ 1910-55.; sulcata K. (syntype §). Mugdock; 457.; Loxotropa/

sulcata, K.; Cameron Coll./ 1910-302.; Determined by/Dr Kieffer; Basalys/ iphiclaNix./ 1979/ G.Nixon

det. 1978 (syntype S).

Type locality

Scotland.

Remarks

This species was described from an unspecified number of females and males. Both syntypes are

mounted on cards and are entire.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Loxotropa convexa Kieffer, 1911b: 932.

Valid name

Basalys abruptus Thomson, 1858.

Summary of types

Syntype §, BMNH number 9.732. Syntype §.

Primary type data

Type; 237.; Cameron Coll./ 1910-55.; convexa K. (syntype $). Clober/ 87; 240; Cameron Coll./ 1910-

55 (syntype §).

Type locality

Scotland.

Remarks

This species was described from an unspecified number of females. Two females were found matching

the description. Both syntypes are mounted on cards; the syntype from Clober is entire and the other is

missing the right antenna beyond a8. This species was synonymized with Basalys abruptus by Nixon

(1980); however, only one syntype agrees with his concept of B. abruptus. A lectotype needs to be

designated for Loxotropa convexa , but it is recommended that Thomson’s type of Basalys abruptus is

examined first to ensure that the synonymy established by Nixon (1980) is maintained if at all possible.

Loxotropa longipennis Kieffer, 1911b: 932.

Valid name

Basalys longipennis (Kieffer, 1911).

Summary of types

Syntype §, BMNH number 9.734.

Primary type data

Mull; Type; 211.; Cameron Coll./ 1910-55.; longipennis K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of females. The syntype is mounted on a card

and is entire.

Loxotropa nigricornis var. subterranea Kieffer, 1911b: 939.

Valid name

Trichopria nigricornis (Marshall, 1868).

Summary of types

Lectotype BMNH number 9.785, designated by Notton (1995).

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Loxotropa/ subterranea/ Blackgang Chine/ Aug. 17/ 1908; Type; Ex coll./ Donisthorpe/ B.M.1934-

4.; Assoc, with/ Tasius flavus; Toxotropa/ subterranea Kief.; Tectotype $1 Toxotropa/ nigricomis/

subterranea/ det. D.G.Notton.

Type locality

England, Isle of Wight, Blackgang Chine.

Remarks

This species was described from an unspecified number of males. The lectotype is mounted on a card

and is entire.

Loxotropa bifoveata Kieffer, 1911b: 940.

Valid name

Basalys bifoveatus (Kieffer, 1911).

Summary of types

Syntype BMNH number 9.735.

Primary type data

Cladich; Type; 467.; Cameron Coll./ 1910-55.; bifoveata K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. The syntype is mounted on a card and

is entire; it has both wings twisted.

Loxotropa luctuosa Kieffer, 1911b: 941.

Valid name

Basalys luctuosus (Kieffer, 1911).

Summary of types

Syntype S, BMNH number 9.733.

Primary type data

Dumfries; Type; 470; Cameron coll./ 1910-55.; luctuosa K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. This syntype agrees well with the

description, although the basal scutellar pit is not clearly bifoveate; however, this part of the specimen

European Journal of Taxonomy 75 : 1-123 ( 2014 )

is dirty, so probably Kieffer was mistaken. The syntype is mounted on a card and has three of the tarsi

broken.

Loxotropa atricrus Kieffer, 1911b: 941.

Valid name

Basalys fumipennis Westwood, 1833.

Summary of types

Syntype $, BMNH number 9.749. Syntype $.

Primary type data

[illeg.]; Type; 458.; Cameron coll./ 1910-55; atricrus K. (syntype <$). [illeg.]’ 461.; Loxotropa/ atricrus,

K.; Cameron Coll./ 1910-302.; Determined by/ Dr Kieffer (syntype f).

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. Both syntypes are mounted on cards:

the specimen numbered “9.749” has one tarsus broken; the other is entire. The specific epithet is derived

from ater , Latin for black, and crus , Latin for leg, and is indeclinable.

Loxotropa scotica Kieffer, 1911b: 942.

Valid name

Basalys scoticus (Kieffer, 1911).

Summary of types

Syntype BMNH number 9.750. Syntype $.

Primary type data

[illeg.]; Type; 462.; Cameron Coll./ 1910-55; scotica K. (syntype c?)- Dumfries; 463.; Cameron Coll./

1910-55; Basalys/ scotica K./ G.Nixon det. 1978 (syntype $).

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. Both syntypes are mounted on cards:

the one numbered 9.730 is missing its left fore wing; the other is entire.

Loxotropa ciliata Kieffer, 1911b: 943.

Valid name

Basalys ciliatus (Kieffer, 1911).

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Syntype S, BMNH number 9.737.

Primary type data

Dumfries; Dumfries; Type/ G.N.; 269.; Loxotropa/ ciliata K.; Cameron Coll./ 1910-302; selected as/

type of/ Loxotropa/ ciliata K./ by G.E.J.N. 1934.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males, of which one was found. Masner

(1965) notes that Nixon labelled this specimen as a lectotype, but Nixon did not publish a designation so

it is still a syntype. The syntype is mounted on a card and has one tarsus broken.

Loxotropa unifoveata Kieffer, 1911b: 943.

Valid name

Basalys bifoveatus (Kieffer, 1911).

Summary of types

Syntype S, BMNH number 9.736.

Primary type data

[illeg.]; Type; 527.; Cameron Coll./ 1910-55.; unifoveata K.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of males. The syntype is mounted on a card and

is entire.

Loxotropa formicarum Kieffer, 1911b: 944.

Valid name

Basalys formicarum (Kieffer, 1911).

Summary of types

Not seen.

Primary type data

According to Masner (1965): “Loxotropa formicarum Darenth Wood, 24.ix. 1909; BMNH type-

label”.

Type locality

England, London.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Remarks

This species was described from an unspecified number of males. Although one male type was listed for

the BMNH by Masner (1965), no material was found; nor was this specimen noted by Nixon (1980).

Diapria ( Tropidopria ) inaequalis Kieffer, 1911b: 956.

Valid name

Trichopria aequata (Thomson, 1858).

Summary of types

Lectotype $, BMNH number 9.693, designated by Notton (1995).

Primary type data

Type; 477.; Cameron Coll./ 1910-55.; inaequalis K.; Lectotype §/ Diapria (Tropidopria)/ inaequalis

Kieffer/ des. Notton, 1995.

Type locality

Scotland.

Remarks

Described from an unspecified number of females. The lectotype agrees well with the description apart

from “article ll e ...plus de deux fois aussi gros que le 12 e ...”, which does not make sense and is most

probably a slip of the pen. The lectotype is mounted on a card and the right antenna is missing beyond a6.

Diapria ( Tropidopria ) tetratoma Kieffer, 1911b: 958.

Valid name

Trichopria hyalinipennis (Thomson, 1858).

Summary of types

Lectotype $, BMNH number 9.694, designated by Notton (1995).

Primary type data

Gloster; Type; 228.; Cameron Coll./ 1910-55.; tetratoma K.; Trichopria/ hyalinipennis Thoms./ det.

L.Huggert-89; Holo-/ type; Lectotype $/ Diapria Tropidopria/ tetratoma Kieffer/ des. Notton, 1995.

Type locality

England, Gloucester.

Remarks

This species was described from an unspecified number of females. Although the published type locality

is Scotland, Cameron (1910) records it from Gloucester, so this specimen is not excluded from the type

series. The lectotype is mounted on a card and is entire.

Diapria ( Tropidopria ) conotoma Kieffer, 1911b: 966.

Valid name

Trichopria conotoma (Kieffer, 1911).

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Lectotype $, BMNH number 9.725, designated by Notton (1995).

Primary type data

Alsasua/ Spain 30-6; 475.; Spain; Cameron Coll./ 1910-55.; conotomaK.; Type; Holo-/ type; Lectotype

5/Diapria Tropidopria/ conotoma Kieffer, 1911: 966/ des. Notton, 1995.

Type locality

Spain, Alsasua.

Remarks

This species was described from an unspecified number of females. The lectotype is mounted on a card

and is missing its left fore wing. Although the type locality was recorded as Scotland, Cameron (1910)

records it from Spain, so this specimen is not excluded from the type series. A Spanish locality is also

more likely because this is a largely southern European species which only just extends into south-east

England. British material of both sexes of T. conotoma has been examined (specimens in BMNH); the

male is identical with the description of Oxypria bouceki Masner, 1959 syn. nov.

Diapria (Tropidopria) variipes Kieffer, 1911b: 967.

Valid name

Trichopria aequata (Thomson, 1858).

Summary of types

Lectotype $, BMNH number 9.696, designated by Notton (1995).

Primary type data

Type; 218.; Cameron Coll./ 1910-55; variipes K.; Lectotype $/ Diapria Tropidopria/ variipes Kieffer/

des. Notton, 1995.

Type locality

Scotland.

Remarks

This species was described from an unspecified number of females. The lectotype is mounted on a card

and has both antennae and two legs broken.

Diapria ( Tropidopria ) melanopa Kieffer, 1911b: 968.

Valid name

Trichopria subimpressa (Kieffer, 1911).

Summary of types

Lectotype $, BMNH number 9.697, designated by Notton (1995). Paralectotypes §, S-

Primary type data

Mugd./ 26-7; Type; 471.; Cameron Coll./ 1910-55.; melanopa K.; Lectotype $/ Diapria Tropidopria/

melanopa Kieffer/ des. Notton, 1995.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Type locality

Scotland, Mugdock.

Remarks

This species was described from an unspecified number of females and males. The lectotype is mounted

on a card and is missing its right mid leg. According to the publication dates on page 1015 of Kieffer

(1911b), fascicules 107-8 (including Trichopria subimpress a) predate fascicules 109-110 (including T.

melanopa , which is here confirmed to be a junior synonym).

Diapria ( Tropidopria) nocticolor Kieffer, 1911b: 977.

Valid name

Trichopria oogaster (Thomson, 1858).

Summary of types

Lectotype $, BMNH number 9.698, designated by Notton (1995).

Primary type data

York; Type; 266.; Cameron Coll./1910-55.; nocticolor K.; Holo-/ type; Lectotype Diapria Tropidopria/

nocticolor Kieffer/ des. Notton, 1995.

Type locality

England, York.

Remarks

This species was described from an unspecified number of males. The lectotype is mounted on a card,

the right antenna is broken and the right fore wing and both hind wings are missing.

Diapria ( Tropidopria ) clavatipes Kieffer, 1911b: 978.

Valid name

Trichopria subimpressa (Kieffer, 1911).

Summary of types

Lectotype BMNH number 9.695, designated by Notton (1995).

Primary type data

Bishop.; Type; 267.; Cameron Coll./ 1910-55.; clavatipes K.; Holo-/ type; Lectotype ft Diapria

Tropidopria/ clavatipes Kieffer/ des. Notton, 1995.

Type locality

Scotland, Bishopton.

Remarks

This species was described from an unspecified number of males. By comparison with other specimens,

the type locality “Bishop.” almost certainly means Bishopton in Dumfries and Galloway. The lectotype

is mounted on a card and has four tarsi broken and the right hind wing lost.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Trichopria fimbriata Kieffer, 1911b: 995.

Valid name

Trichopria nigra (Nees, 1834).

Summary of types

Lectotype S, BMNH number 9.703, designated by Notton (1995).

Primary type data

Craigton/ fungus 22-9; Type; 270.; Cameron Coll./ 1910-55.; fimbriata K.; Holo-/ type; Lectotype c V

Trichopria/ fimbriata Kieffer/ des. Notton, 1995.

Type locality

Scotland, Craigton.

Remarks

This species was described from an unspecified number of males. The type locality is most probably

Craigton in Dumfries and Galloway, but might instead by Craigton in Glasgow. The lectotype is mounted

on a card, has both antennae and two tarsi broken, and both fore wings are disarticulated.

Trichopria ciliaris Kieffer, 1911b: 1000.

Valid name

Trichopria basalis (Thomson, 1858).

Summary of types

Lectotype $, BMNH number 9.704, designated by Notton (1995). Paralectotypes 2 $ §.

Primary type data

Clober 1-6; Type; 229.; Diapria/ ciliaris; Cameron Coll./ 1910-55.; ciliaris K.; Lectotype §/ Trichopria/

ciliaris Kieffer/ des. Notton, 1995.

Type locality

Scotland, Clober.

Remarks

This species was described from at least two females, since a range of lengths is given. The lectotype is

mounted on a card and is entire.

Trichopria formicaria Kieffer, 1911c: 385.

Valid name

Trichopria sociabilis Masner, 1965.

Summary of types

Lectotype §, BMNH number 9.768, designated by Notton (1995).

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Primary type data

Trichopria/ sociabilis/ Box/ Hill/ 20.v. 1910; Type; Ex coll. Donisthorpe/ B.M. 1934-4; Assoc, with/

Formica fiisca; Trichopria/ formicaria Kief.; Holo-/ type; $ Lectotype/ Trichopria/ formicaria/ Kieffer,

1911: 385/ des. Notton, 1995/=Trichopria/ sociabilis Masner, 1965.

Type locality

England, Box Hill.

Remarks

This species was described from an unspecified number of females. The lectotype is mounted on a card

and is entire.

Schizopria fallax Kieffer, 1912a: 68, pi. 3, fig. 11.

Valid name

Entomacis fallax (Kieffer, 1912).

Summary of types

Syntype §, BMNH number 9.683. Syntypes 2 $ §.

Primary type data

103; Silhouette, ’08-9,/ Seychelles Exp.; Schizopria/ fallax/ J.J.Kieffer/ type; figured specimen; Type;

Type/ H.T.; Seychelle Islands./ Percy Sladen Trust/ Expedition./ 1913-170 (syntype $). 97; Mahe, ’08-

9,/ Seychelles Exp.; Schizopria/ fallax/ J.J.Kieffer/ paratype; Seychelle Islands./ Percy Sladen Trust/

Expedition./ 1913-170 (syntype §). 84; Mahe, ’08-9,/ Seychelles Exp.; Schizopria/ fallax/ J.J.Kieffer/

paratype; Seychelle Islands./ Percy Sladen Trust/ Expedition./ 1913-170 (syntype §).

Type locality

Seychelles, Silhouette, near Mont Pot-a-eau; Silhouette (as Mahe), Mare aux Cochons dist.; Mahe,

Cascade Estate.

Remarks

Three syntypes were found which agree with the description except for having 13-segmented antenna;

although Kieffer says the antenna is 12-segmented, this may be explained by al2-13 being closely

joined, giving the appearance of 12 segments. The holotype label and paratype labels are erroneous, as

Kieffer did not designate a holotype; consequently, they are all syntypes. The specimen numbered “103”

is mounted on a card and is entire; “97” is mounted on a point and has the head and one antenna mounted

separately; “84” is mounted on a point and is entire. All three syntypes belong to Entomacis.

Schizopria flaviclava Kieffer, 1912a: 68, 69.

Valid name

Entomacis flaviclava (Kieffer, 1912).

Summary of types

Holotype BMNH number 9.684, by monotypy.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

High jungle/ 126; Silhouette, ’08-9./ Seychelles Exp.; Schizopria/ flaviclava/ J.J.Kieffer/ type.; Type;

Type/ H.T.; Seychelle Islands./ Percy Sladen Trust/ Expedition./ 1913-170.

Type locality

Seychelles, Silhouette, Mare aux Cochons.

Remarks

The holotype is mounted on a card and is entire. The antenna is sub-12 segmented, the club apparently

formed of two closely joined segments.

Entomacis longicornis Kieffer, 1912a: 69.

Valid name

Entomacis longicornis Kieffer, 1912.

Summary of types

Syntype $, BMNH number 9.685.

Primary type data

97; Mahe, ’08-9./ Seychelles Exp.; Entomacis/ longicornis/ J.J.Kieffer/type; Type; Type/H.T.; Seychelle

Islands./Percy Sladen Trust/Expedition./ 1913-170.

Type locality

Seychelles, Mahe, Cascade Estate.

Remarks

One syntype was found standing over this name. The holotype label is erroneous, as Kieffer did not

designate a holotype. The syntype is mounted on a card point and is entire.

Entomacis curticornis Kieffer, 1912a: 69, 70.

Valid name

Entomacis curticornis Kieffer, 1912.

Summary of types

Syntype BMNH number 9.686. Syntype §.

Primary type data

97; Mahe, ’08-9./ Seychelles Exp.; Entomacis/ curticornis/ J.J.Kieffer/type.; Type; Type/H.T.; Seychelle

Islands./Percy Sladen Trust/Expedition./ 1913-170. (syntype $). 146; Mahe, ’08-9./ Seychelles Exp.;

Entomacis/ curticornis/ J.J.Kieffer/ paratype.; Seychelle Islands./ Percy Sladen Trust/ Expedition./

1913-170 (syntype §).

Type locality

Seychelles, Mahe, Cascade Estate; Mahe, Anse aux Pins and Anse Royal.

European Journal of Taxonomy 75: 1-123 (2014)

Remarks

Two syntypes were found. The holotype and paratype labels are erroneous, as Kieffer did not designate

a holotype. The male syntype is mounted on a card point and is entire; the female syntype is mounted

on a card and is entire.

Loxotropa exsul Kieffer, 1912a: 70.

Valid name

Basalys exsul (Kieffer, 1912).

Summary of types

Syntype 5 BMNH number 9.738. Syntype 3.

Primary type data

22; Silhouette, ‘08/ Seychelles Exp.; Toxotropa/ exsul/ J.J.Kieffer/ type; Type; Type/ H.T.; Seychelle

Islands./ Percy Sladen Trust/ Expedition./ 1913-170 (syntype $). 20; 20/ Silhouette, ‘08/ Seychelles

Exp.; Loxotropa/ exsul/ J.J.Kieffer/paratype; Seychelle Islands./ Percy Sladen Trust/ Expedition./ 1913-

170 (syntype S).

Type locality

Seychelles, Silhouette, Mare aux Cochons; Mahe, Slopes of Morne Seychellois.

Remarks

Two syntypes were found. The holotype and paratype labels are erroneous, as Kieffer did not designate

a holotype. The male syntype is mounted on a card point and is entire; the female syntype is mounted on

a card point and the left flagellum, gaster and tip of the right hind wing are missing.

Loxotropa semirufa Kieffer, 1912a: 70.

Valid name

Basalys semirufus (Kieffer, 1912).

Summary of types

Holotype BMNH number 9.739, by monotypy.

Primary type data

19/ Silhouette, ‘08/ Seychelles Exp.; Loxotropa/ semirufa/ J.J.Kieffer/type; Type; Type/H.T.; Seychelle

Islands./Percy Sladen Trust/Expedition./ 1913-170.

Type locality

Seychelles, Silhouette, Mare aux Cochons.

Remarks

The holotype agrees well with the description, although it is somewhat faded. It is mounted on a card

and is missing its right wings.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Rhopalopria vulgaris Kieffer, 1912a: 71, pi. 3, fig. 12.

Valid name

Trichopria vulgaris (Kieffer, 1912).

Summary of types

Lectotype $, BMNH number 9.689, designated by Huggert (1979). Paralectotypes §, 3 <$< 3 -

Primary type data

99/ Mahe, ’08-9./ Seychelles Exp.; Paratype; Rhopalopria/ vulgaris/ J.J.Kieffer/ paratype; Seychelle

Islands./ Percy Sladen Trust/ Expedition./ 1913-170; Lectotypus/ Rhopalopria/ vulgaris/ Kieff / design.

5/ L.Huggert-77; Trichopria $/ vulgaris Kieff./ det. L.Huggert-77.

Type locality

Seychelles, Mahe.

Remarks

Five syntypes were found. The holotype and paratype labels are erroneous as Kieffer did not designate a

holotype. The syntype series was analysed by Huggert (1979) who designated a lectotype. The lectotype

is mounted on a cork block and has the left antenna and left fore wing mounted on a microslide and

the left hind wing missing. Huggert considered one paralectotype male of R. vulgaris to be a different

species and designated it the holotype of Trichopria euvulgaris Huggert q.v.

Diapria seychellensis Kieffer, 1912a: 71, 72, fig. 13.

Valid name

Trichopria seychellensis (Kieffer, 1912).

Summary of types

Lectotype $, BMNH number 9.699, designated by Huggert (1979). Paralectotypes 2 $$,(?.

Primary type data

22/ Silhouette, ‘08/ Seychelles Exp./ $1 Diapria/ seychellensis; type; Type/H.T.; Diapria/ seychellensis/

J.J.Kieffer/ Type; figured specimen; Seychelle Islands./ Percy Sladen Trust/ Expedition./ 1913-170;

Lectotypus/ Diapria $! seychellensis/ Kieff./ design./ L.Huggert-77; Trichopria (A.) $! seychellensis

(Kieff.)/ det. L.Huggert-77.

Type locality

Seychelles, Silhouette, Mare aux Cochons.

Remarks

This species was described from three females and five males. Two females and two males were located

by Huggert (1979), of which he designated one as lectotype of D. seychellensis. The lectotype is mounted

on a papered cork block; the left antenna and left wings are mounted on a microslide on the same pin.

Huggert considered two of the paralectotypes were not conspecific with T. seychellensis and designated

one the holotype of Trichopria ensia q.v. and another the holotype of Trichopria percyi q.v.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Diapria scotti Kieffer, 1912a: 71, 72.

Valid name

Trichopria scotti (Kieffer, 1912).

Summary of types

Lectotype S, BMNH number 9.700, designated by Huggert (1979). Paralectotype §.

Primary type data

67; Diapria/ scotti/ J.J.Kieffer/ type; Mahe, ‘08-9./ Seychelles Exp.; Type; Type/H.T.; Seychelle Islands./

Percy Sladen Trust/Expedition./ 1913-170; Lectotypus/Diapria c V scotti Kieff./ design./L.Huggert-76.

Type locality

Seychelles, Mahe, nr. Morne Blanc.

Remarks

The holotype label on the lectotype is erroneous as Kieffer did not designate a holotype. The lectotype

is mounted on a card and has the right hind wing missing and the left antenna and left wings on a

microslide on the same pin.

Diapria mahensis Kieffer, 1912a: 71, 73.

Valid name

Trichopria mahensis (Kieffer, 1912).

Summary of types

Holotype §, BMNH number 9.701, by monotypy.

Primary type data

Diapria/ mahensis/ J.J.Kieffer/ type; Mahe, ’08-9./ Seychelles Exp.; Type; Type/ H.T.; Holotypus §/

Diapria/ mahensis Kieffer/ L.Huggert-76; Trichopria §/ mahensis Kieff./ det. L.Huggert-76.

Type locality

Seychelles, Mahe.

Remarks

The holotype is mounted on a card with the left fore wing and left antenna on a microslide mounted on

the same pin.

Diapria saxatilis Kieffer, 1912a: 71, 73.

Valid name

Trichopria saxatilis (Kieffer, 1912).

Summary of types

Lectotype BMNH number 9.702, designated by Huggert (1977). Paralectotypes §, $.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

48; Diapria/ saxatilis/ J.J.Kieffer/ Type; Praslin, ’08./ Seychelles Exp.; Type; Lectotypus/ Diapria $!

saxatilis Kieff./ design./ L.Huggert-76; Trichopria c V saxatilis Kieff./ det. Huggert-76.

Type locality

Seychelles, Praslin, Cotes d’Or Estate, Vallee de Mai.

Remarks

This species was described from two males and three females. Huggert (1977) found two males and one

female syntype, designating one male as the lectotype of D. saxatilis. The lectotype is mounted on a

card; the left wings and left antenna are mounted on a microslide on the same pin. Huggert considered

that two of the paralectotypes were not conspecific with T. saxatilis and designated one the holotype of

Trichopria pseadosaxatilis q.v. and the other the holotype of T. nirva q.v.

Calogalesus parvulus Kieffer, 1912a: 73.

Valid name

Calogalesus parvulus Kieffer, 1912.

Summary of types

Holotype §, BMNH number 9.675, by monotypy.

Primary type data

Type/ H.T.; Mahe ’08-9/ Seychelles Exp.; Calogalesus/ parvulus/ J.J.Kieffer/ type; Type.

Type locality

Seychelles, Mahe, Cascade Estate.

Remarks

Kieffer based his description on one specimen, hence a holotype by monotypy. The holotype is mounted

on a card point; the left fore wing, right hind wing and some legs are lost.

Microgalesus quadridens Kieffer, 1912a: 74.

Valid name

Basalys quadridens (Kieffer, 1912) comb. nov.

Summary of types

Holotype §, BMNH number 9.674, by monotypy.

Primary type data

Type/ H.T.; Mahe, ’08-9/ Seychelles Exp.; Microgalesus/ quadridens/ J.J.Kieffer/ type.

Type locality

Seychelles, Mahe, Cascade Estate.

European Journal of Taxonomy 75 : 1-123 ( 2014 )

Remarks

Kieffer based his description on one specimen, hence a holotype by monotypy. The holotype is

mounted on a card point; the head, right hind wing and parts of the legs are lost; the right fore wing is

disarticulated. The triangular stigmalis, three-segmented antennal club, notauli represented by a weak

trace, and form of the propodeum are typical of Basalys. The absence of the basal vein is not surprising

since the specimen is so small. The bealc-like mandibles, anteriorly indented head with teeth, and the

transverse petiole are known in other representatives of this genus. Microgalesus quadridens is here

transferred to Basalys. Since M. quadridens is the type species of Microgalesus , Microgalesus is now a

junior synonym of Basalys syn. nov.

Galesus (Schizogalesus ) silvestrii Kieffer, 1913a: 91.

Valid name

Coptera silvestrii (Kieffer, 1913).

Summary of types

Paralectotypes 3 SS-

Remarks

Three male syntypes were found. These are now paralectotypes following the lectotype designation of

Yoder in Yoder & Wharton (2002).

Loxotropa donisthorpei Kieffer, 1913b: 176.

Remarks

The specimen with BMNH number 9.763 is not a type; the lectotype is in the collection of the University

Museum of Natural History, Oxford (Notton 1995).

Aneurhynchus nodicornis Marshall, 1867: 225.

Remarks

A male of this distinctive species was found, originally from the Claude Morley collection and labelled

as a metatype, i.e., a specimen that had been compared with the type by the author of the species. This

specimen has no formal type status within the ICZN Code; however, it was determined by Marshall and

may be of use in the interpretation of this species should the type not be located.

Mimopria campbellorum Masner, 1976: 124,126, figs 1-6.

Valid name

Mimopria campbellorum Masner, 1976.

Summary of types

Paratypes 3 $ $.

Spilomicrus boweni Masner, 1991: 118,124, figs 26, 28, 83-85,108,123-125.

Valid name

Spilomicrus boweni Masner, 1991.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Paratypes 5, S-

Spilomicrus distinctus Masner, 1991: 118,128, figs 27, 81-82,107,114,122.

Valid name

Spilomicrus distinctus Masner, 1991.

Summary of types

Paratypes 5, S-

Spilomicrus exul Masner, 1991: 117,131, figs 13, 77-79,109,112,119.

Valid name

Spilomicrus exul Masner, 1991.

Summary of types

Paratype S-

Spilomicrus inornatus Masner, 1991: 119,136, figs 25, 65-67,102,115.

Valid name

Spilomicrus inornatus Masner, 1991.

Summary of types

Paratypes §, S-

Spilomicrus palustris Masner, 1991: 118,139, figs 86-86a, 110.

Valid name

Spilomicrus palustris Masner, 1991.

Summary of types

Paratype S-

Spilomicrus pumilio Masner, 1991: 119,140, figs 4, 46-48, 94.

Valid name

Spilomicrus pumilio Masner, 1991.

Summary of types

Paratype §.

Spilomicrus sanbornei Masner, 1991: 117,145, figs 5-7, 40-42, 95.

Valid name

Spilomicrus sanbornei Masner, 1991.

European Journal of Taxonomy 75: 1-123 (2014)

Summary of types

Paratype f.

Spilomicrus stepheni Masner, 1991: 118,147, figs 19, 74-76,106.

Valid name

Spilomicrus stepheni Masner, 1991.

Summary of types

Paratype §.

Spilomicrus superbus Masner, 1991: 117,151, figs 3, 43-45, 93,120.

Valid name

Spilomicrus lubomiri Ozdikmen, 2010.

Summary of types

Paratype S.

Spilomicrus sylvicola Masner, 1991: 118,153, figs 11-12, 37-39, 92,116.

Valid name

Spilomicrus sylvicola Masner, 1991.

Summary of types

Paratype §.

Chilomicrus pecki Masner & Garcia, 2002: 42, figs 32, 33.

Valid name

Chilomicrus pecki Masner & Garcia, 2002.

Summary of types

Paratype f.

Apopria coveri Masner & Garcia, 2002: 65, figs 8, 9.

Valid name

Apopria coveri Masner & Garcia, 2002.

Summary of types

Paratype §.

Eladio cruzi Masner & Garcia, 2002: 80, figs 45-47.

Valid name

Eladio cruzi Masner & Garcia, 2002.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Paratype §.

Hansonia pauli Masner & Garda, 2002: 82, fig. 57.

Valid name

Hansonia pauli Masner & Garcia, 2002.

Summary of types

Paratype §.

Leucopria cylindricornis Masner & Garcia, 2002: 86, fig. 61.

Valid name

Leucopria cylindricornis Masner & Garcia, 2002.

Summary of types

Paratype §.

Turripria woldai Masner & Garcia, 2002: 111, fig. 100.

Valid name

Turripria woldai Masner & Garcia, 2002.

Summary of types

Paratype §.

Peckidium enigmaticum Masner & Garcia, 2002: 119, figs 82-83.

Valid name

Peckidium enigmaticum Masner & Garcia, 2002.

Summary of types

Paratype §.

Loxotropa morleii Morley, 1931: 15.

Valid name

Trichopria halterata (Kieffer, 1909).

Summary of types

Holotype §, BMNH number 9.784, by original designation.

Primary type data

3 ix 97; Loxotropa/ Morleyi/ MS. Chitty; det. A.J.C.; probably Paramesius/ $ undescribed TAM; Type/

CM.

European Journal of Taxonomy 75: 1-123 (2014)

Type locality

England, Suffolk, Sproughton Marshes.

Remarks

The name Loxotropa morleii was introduced as a nomen nudum by Morley (1929) and then made

available in 1931. Morley credited the name to Chitty, and had a manuscript description from him

placing this species in Trichopria; however, by placing it in Loxotropa Forster, 1856 and comparing it

to other species of Loxotropa , it appears that Morley has significantly changed Chitty’s manuscript and

was largely responsible for the text, and so is credited here as the author. The specimen is mounted on

a card and is very dirty.

Termitopria sheasbyi Muesebeck, 1965: 190, fig. la-e.

Valid name

Termitopria sheasbyi Muesebeck, 1965.

Summary of types

Paratypes 4 $ ? •

Psilus aquilonius Muesebeck, 1980: 4, 6, 8, figs 1-la, 4, 23, 34, 43-44, 61.

Valid name

Psilus aquilonius Muesebeck, 1980.

Summary of types

Paratype <f.

Psilus dissidens Muesebeck, 1980: 3, 4,10, figs 7, 25, 36, 47, 48, 62.

Valid name

Psilus dissidens Muesebeck, 1980.

Summary of types

Paratype $.

Psilus fords Muesebeck, 1980: 4, 5,11, figs 10, 27, 37, 51.

Valid name

Psilus fords Muesebeck, 1980.

Summary of types

Paratype 3.

Psilus masneri Muesebeck, 1980: 4,12, figs 11, 38, 52.

Valid name

Psilus masneri Muesebeck, 1980.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Paratype §.

Psilus masoni Muesebeck, 1980: 3, 6,13, figs 12, 28, 39, 53, 54.

Valid name

Psilus masoni Muesebeck, 1980.

Summary of types

Paratype <$■

Coptera angulata Muesebeck, 1980: 24, 29.

Valid name

Coptera angulata Muesebeck, 1980.

Summary of types

Paratype S-

Coptera distans Muesebeck, 1980: 23, 26, 34.

Valid name

Coptera distans Muesebeck, 1980.

Summary of types

Paratype S-

Copterapomonellae Muesebeck, 1980: 23, 26, 44, figs 80,100,119.

Valid name

Coptera pomonellae Muesebeck, 1980.

Summary of types

Paratype S-

Coptera strauziae Muesebeck, 1980: 22, 25-26, 46, figs 82,101,120-123,133.

Valid name

Coptera strauziae Muesebeck, 1980.

Summary of types

Paratype S-

Galesus ( Schizogalesus ) quadridens Nixon, 1930: 400, 401, 402.

Valid name

Coptera quadridens (Nixon, 1930).

European Journal of Taxonomy 75: 1-123 (2014)

Summary of types

Syntype 3 , BMNH number 9.669. Syntype §, BMNH number 9.669b. Syntypes §, 8 33.

Primary type data

Type; Natal/ Van Reenen,/ Drakensberg/ 1-22.i. 1927; Africa/ R.E.Turner/ Brit. Mus./ 1927-54; Galesus/

quadridens/ 1930 Nixon/ Type $ (syntype §). Paratype; Natal/ Kloof/ 1500ft/ Sept. 1926; Africa/

R.E.Turner/ Brit. Mus./ 1926-404; Galesus/ quadridens/ 1930 Nixon (syntype $). Type; Natal/ Van

Reenen,/ Drakensberg/ 1-22.i. 1927; Africa/ R.E.Turner/ Brit. Mus./ 1927-54; Galesus/ quadridens/

1930 Nixon/ Type 3 (syntype $). Paratype; Natal/ Van Reenen,/ Drakensberg/ 1-22.i. 1927; Africa/

R.E.Turner/ Brit. Mus./ 1927-54; Galesus/ quadridens/ 1930 Nixon (syntypes 6c?c?). Paratype; Orange

F. State/Harrismith/Feb. 1927; S.Africa/R.E.Turner/Brit Mus./ 1927-117; Galesus/ quadridens/ 1930

Nixon (syntype 3 )- Paratype; Zululand:/ Gingindhloyu/ 15.vi.1926; S.Africa/ R.E.Turner/ Brit. Mus./

1926-277; Galesus/ quadridens/ 1930 Nixon (syntype 3 )-

Type locality

South Africa, Free State, Harrismith; KwaZulu-Natal, Gingindlovu, Kloof and Van Reenen.

Remarks

There are two females and nine males agreeing with Nixon’s description and labelled by him as Galesus

quadridens. Of these, two are labelled as “type” and the rest as “paratype“; however, since neither a

unique holotype or lectotype has yet been fixed, all are still syntypes (Article 73.2). All are mounted on

card points, the females are entire, the male labelled by Nixon as “type” is missing its head, and three

other males are missing parts of their antennae.

Galesus ( Schizogalesus ) bidens Nixon, 1930: 400, 401, 403.

Valid name

Coptera bidens (Nixon, 1930).

Summary of types

Syntype §, BMNH number 9.668. Syntype 3 , BMNH number 9.688b. Syntypes 4 $ §, 5 33.

Primary type data

Type; MosselBay/ Cape Province/ October, 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1921-450; Galesus/

bidens/ 1930 Nixon/ Type $ (syntype §). Paratype; Mossel Bay/ Cape Province/ June, 1921; S.Africa/

R.E.Turner/ Brit. Mus./ 1921-294; Galesus/ bidens/ 1930 Nixon (syntype §)• Paratype; Mossel Bay/

Cape Province/ October, 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1921-450; Galesus/ bidens/ 1930

Nixon (syntype $). Paratype; Mossel Bay/ Cape Province/ April, 1921; S.Africa/ R.E.Turner/ Brit.

Mus./ 1921-210; Galesus/ bidens/ 1930 Nixon (syntype $). Paratype; Mossel Bay/ Cape Province/

Aug. 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1921-353; Galesus/ bidens/ 1930 Nixon (syntype $).

Type; Mossel Bay/ Cape Province/ April, 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1921-210; Galesus/

bidens/ 1930 Nixon/ Type 3 (syntype 3 ). Paratype; Mossel Bay/ Cape Province/ Febr. 1922; S.Africa/

R.E.Turner/ Brit. Mus./ 1922-97; Galesus/ bidens/ 1930 Nixon (syntype 3 ). Paratype; Mossel Bay/

Cape Province/ 18-30.xi. 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-2; Galesus/ bidens/ 1930 Nixon

(syntype 3 )- Paratype; Mossel Bay/ Cape Province/ 5-3l.vii. 1921; S.Africa/ R.E.Turner/ Brit. Mus./

1921-315; Galesus/ bidens/ 1930 Nixon (syntype 3 )- Paratype; Mossel Bay/ Cape Province/ 1-13.

iii.1922; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-129; Galesus/ bidens/ 1930 Nixon (syntypes 2 33 )-

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Type locality

South Africa, Western Cape Province, Mossel Bay.

Remarks

There are five females and six males agreeing with Nixon’s description and labelled by him as Galesus

bidens. Of these, two are labelled as “type” and the rest as “paratype”; however, since neither a unique

holotype or lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). All the syntypes

are mounted on cards and one female has an antenna mounted separately; two males are missing parts

of their antennae.

Galesus ( Schizogalesus ) inquisitor Nixon, 1930: 400, 402, 404.

Valid name

Copter a inquisitor (Nixon, 1930).

Summary of types

Syntype §, BMNH number 9.659. Syntype c?, BMNH number 9.659b. Syntypes 12 12 33 -

Primary type data

Type; Mossel Bay/ Cape Province/ 5-3l.vii. 1921; S.Africa/R.E.Turner/Brit. Mus./ 1921-315; Galesus/

inquisitor/ 1930 Nixon/ Type $ (syntype $). Paratype; Mossel Bay/ Cape Province/ 15-28.iii. 1922;

S.Africa/ R.E.Turner/ Brit. Mus./ 1922-153; Galesus/ inquisitor/ 1930 Nixon (syntype $). Paratype;

Mossel Bay/Cape Province/18-30.xi. 1921; S.Africa/R.E.Turner/Brit. Mus./1922-2; Galesus/ inquisitor/

1930 Nixon (syntype §). Paratype; Mossel Bay/ Cape Province/ Aug. 1921; S.Africa/ R.E.Turner/ Brit.

Mus./ 1921-353; Galesus/ inquisitor/ 1930 Nixon (syntypes $, 4 3c ?)• Paratype; Mossel Bay/ Cape

Province/ May, 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1921-248; Galesus/ inquisitor/ 1930 Nixon

(syntype §). Paratype; Mossel Bay/ Cape Province/ October, 1921; S.Africa/ R.E.Turner/ Brit. Mus./

1921-450; Galesus/ inquisitor/ 1930 Nixon (syntypes $, 4 33 )- Paratype; Mossel Bay/ Cape Province/

5-3l.vii. 1921; S.Africa/R.E.Turner/Brit. Mus./ 1921-315; Galesus/ inquisitor/ 1930 Nixon (syntypes

5, 2 33 )- Paratype; Aliwal North/ Cape Province/ 4350ft/ 1-134.1923; S.Africa/ R.E.Turner/ Brit.

Mus./ 1923-70; Galesus/ inquisitor/ 1930 Nixon (syntype $). Paratype; Port St. John/ Pondoland/ April

5-30. 1923; S.Africa/ R.E.Turner/ Brit. Mus./ 1923-286; Galesus/ inquisitor/ 1930 Nixon (syntype

5 ). Paratype; Natal/ Kloof/ 1500ft/ Sept. 1926; S.Africa/ R.E.Turner/ Brit. Mus./ 1926-404; Galesus/

inquisitor/ 1930 Nixon (syntype $). Paratype; Natal/ Van Reenen/ Drakensberg/ Nov. 1926; S.Africa/

R.E.Turner/ Brit. Mus./ 1926-499; Galesus/ inquisitor/ 1930 Nixon (syntypes 2 §?)• Paratype; Orange

F. State/Harrismith/ March 1-20. 1927; S.Africa/R.E.Turner/Brit. Mus./ 1927-147; Galesus/ inquisitor/

1930 Nixon (syntype §). Type; Cape Province/ Ceres/ March 1925; S.Africa/ R.E.Turner/ Brit. Mus./

1925-161; Galesus/ inquisitor/ 1930 Nixon/ Type $ (syntype $). Paratype; Mossel Bay/ Cape Province/

Dec. 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-25; Galesus/ inquisitor/ 1930 Nixon (syntype $).

Paratype; Port St. John/ Pondoland/ May. 1924; S.Africa/ R.E.Turner/ Brit. Mus./ 1924-289; Galesus/

inquisitor/ 1930 Nixon (syntype $).

Type locality

South Africa, Eastern Cape Province, Aliwal North and Port St Johns; Free State, Harrismith; KwaZulu-

Natal, Van Reenen and Kloof; Western Cape Province, Ceres and Mossel Bay.

European Journal of Taxonomy 75: 1-123 (2014)

Remarks

There are thirteen females and thirteen males agreeing with Nixon’s description and labelled by him

as Galesus inquisitor. Of these, two are labelled as “type” and the rest as “paratype”; however, since

neither a unique holotype or lectotype has yet been fixed, all of the type series is syntypic (Article

73.2). All syntypes are mounted on cards, except the four from Natal and Orange Free State, which are

mounted on card points.

Galesus (Schizogalesus) distinctus Nixon, 1930: 400, 405.

Valid name

Copter a distincta (Nixon, 1930).

Summary of types

Holotype §, BMNH number 9.655, by monotypy.

Primary type data

Type; S.Africa/R.E.Turner/Brit. Mus./ 1921-315; Mossel bay/ Cape Province/ 5-3l.vii. 1921; Galesus/

distinctus/Nixon/ $ type 1930.

Type locality

South Africa, Western Cape Province, Mossel Bay.

Remarks

The holotype is mounted on a card and is entire.

Galesus {Schizogalesus) rectangularis Nixon, 1930: 400, 402, 406.

Valid name

Copter a rectangularis (Nixon, 1930).

Summary of types

Syntype $, BMNH number 9.656. Syntype BMNH number 9.656b. Syntypes 3 $ §.

Primary type data

Type; Port St. John/ Pondoland/ Nov. 1923; S.Africa/ R.E.Turner/ Brit. Mus./ 1924-6; Galesus/

rectangularis/ Nixon/ § type 1930 (syntype $). Paratype; Port St. John/ Pondoland/ Aug. 15-31 1923;

S.Africa/ R.E.Turner/ Brit. Mus./ 1923-463; Galesus/ rectangularis/ Nixon/ $ paratype 1930 (syntype

5 ). Paratype; Port St. John/ Pondoland/ Jan. 1924; S.Africa/R.E.Turner/Brit. Mus./ 1924-97; Galesus/

rectangularis/Nixon/ $ paratype 1930 (syntype $). Paratype; Mossel Bay/ Cape Province/ Dec. 1921;

S.Africa/ R.E.Turner/ Brit. Mus./ 1922-25; Galesus/ rectangularis/ Nixon/ $ paratype 1930 (syntype

5 ). Type; Mossel Bay/ Cape Province/ June 1921; S.Africa/R.E.Turner/Brit. Mus./ 1921-294; Galesus/

rectangularis/Nixon/ $ type 1930 (syntype $).

Type locality

South Africa, Eastern Cape Province, Port St Johns; Western Cape Province, Mossel Bay.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

There are four females and one male agreeing with Nixon’s description and labelled by him as Galesus

rectangularis. Of these, two are labelled as “type” and the rest as “paratype”; however, since neither a

unique holotype or lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). All the

syntypes are mounted on card points and are entire.

Galesus (Schizogalesus) abdominalis Nixon, 1930: 400, 402, 406.

Valid name

Coptera abdominalis (Nixon, 1930).

Summary of types

Syntype §, BMNH number 9.667. Syntype c?, BMNH number 9.667b. Syntypes 2 $?,(?•

Primary type data

Type; Port St. John/Pondoland/ July 10-31, 1923; S.Africa/R.E.Turner/Brit. Mus./ 1923-398; Galesus/

abdominalis/ Nixon; $ type, 1930 (syntype §). Paratype; Port St. John/ Pondoland/ Sept. 1923;

S.Africa/ R.E.Turner/ Brit. Mus./ 1923-510; Galesus/ abdominalis/Nixon; $ paratype, 1930 (syntype

5 ). Paratype; Port St. John/ Pondoland/ 6-25 Feb. 1923; S.Africa/ R.E.Turner/ Brit. Mus./ 1924-136;

Galesus/ abdominalis/Nixon; $ paratype, 1930 (syntype $). Type; Port St. John/Pondoland/ July 10-31,

1923; S.Africa/ R.E.Turner/ Brit. Mus./ 1923-398; Galesus/ abdominalis/Nixon; $ type, 1930 (syntype

$). Paratype; Port St. John/Pondoland/ July 10-31, 1923; S.Africa/R.E.Turner/Brit. Mus./ 1923-398;

Galesus/ abdominalis/Nixon; $ paratype, 1930 (syntype $)■

Type locality

South Africa, Eastern Cape Province, Port St Johns.

Remarks

There are three females and two males agreeing with Nixon’s description and labelled by him as Galesus

abdominalis. Of these, two are labelled as “type” and the rest as “paratype”; however, since neither a

unique holotype or lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). All the

syntypes are mounted on cards and are entire.

Galesus {Schizogalesus) magnificus Nixon, 1930: 401, 408.

Valid name

Coptera robustior (Silvestri, 1913).

Summary of types

Syntype $, BMNH number 9.660. Syntype §.

Primary type data

Type; Natal/ Kloof/ 1500ft./ Sept. 1926; S.Africa/ R.E.Turner/ Brit. Mus./ 1926-404; Galesus/

magnificus/ $ type Nixon/ 1930 (syntype §). Paratype; Port St. John/ Pondoland/ 6-25.Feb. 1924;

S.Africa/R.E.Turner/Brit. Mus./ 1924-136.; Galesus/ magnificus/ $ paratype Nixon/ 1930 (syntype §).

Type locality

South Africa, Eastern Cape Province, Port St Johns; KwaZulu-Natal, Kloof.

European Journal of Taxonomy 75: 1-123 (2014)

Remarks

There are two females agreeing with Nixon’s description and labelled by him as Galesus magnificus. Of

these, one is labelled as “type” and the other “paratype”; however, since neither a unique holotype or

lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). Both syntypes are mounted

on card points and are entire.

Galesus (Schizogalesus) modestus Nixon, 1930: 401, 408.

Valid name

Copter a modesta (Nixon, 1930)*

Summary of types

Syntype §, BMNH number 9.665. Syntype §.

Primary type data

Type; Port St. John/Pondoland/ July 10-31, 1923; S.Africa/R.E.Turner/Brit. Mus./ 1923-398; Galesus/

modestus/ Nixon $ type 1930 (syntype §). Type; Port St. John/ Pondoland/ Jan. 1924; S.Africa/

R.E.Turner/Brit. Mus./ 1924-97; Galesus/ modestus/Nixon $ paratype 1930 (syntype §).

Type locality

South Africa, Eastern Cape Province, Port St Johns.

Remarks

There are two females agreeing with Nixon’s description and labelled by him as Galesus modestus. Of

these, one is labelled as “type” and the other “paratype”; however, since neither a unique holotype or

lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). Both syntypes are mounted

on card points and are entire.

Galesus {Schizogalesus) turned Nixon, 1930: 401, 402, 409.

Valid name

Copter a turned (Nixon, 1930).

Summary of types

Syntype $, BMNH number 9.670. Syntype S, BMNH number 9.670b. Syntypes 2 $$,(?.

Primary type data

Type; Port St. John/ Pondoland/ Oct. 1923; S.Africa/ R.E.Turner/ Brit. Mus./ 1923-547; Galesus/

turneri/ Nixon/ $ type 1930 (syntype $). Paratype; Port St. John/ Pondoland/ 18-31 Mar. 1924;

S.Africa/ R.E.Turner/ Brit. Mus./ 1924-191; Galesus/ turneri/ Nixon/ $ paratype 1930 (syntype $).

Paratype; Mossel Bay/ Cape Province/ Dec. 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-25; Galesus/

turneri/Nixon/ § paratype 1930 (syntype §). Type; Mossel Bay/ Cape Province/Dec. 1921; S.Africa/

R.E.Turner/ Brit. Mus./ 1922-25; Galesus/ turneri/Nixon/ <f type 1930 (syntype $). Paratype; Mossel

Bay/ Cape Province/ 18-30.xi. 1921; S.Africa/R.E.Turner/Brit. Mus./ 1922-2; Galesus/ turneri/Nixon/

S paratype 1930 (syntype S).

Type locality

South Africa, Eastern Cape Province, Port St Johns; Western Cape Province, Mossel Bay.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

There are three females and two males agreeing with Nixon’s description and labelled by him as Galesus

turneri. Of these, two are labelled as “type” and the rest as “paratype”; however, since neither a unique

holotype or lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). The two syntypes

from Port St John are mounted on card points, the three syntypes from Mossel Bay are mounted on cards;

all are entire except for the male labelled as a paratype, which has the metasoma mounted separately.

Galesus (Scltizogalesus) turneri var. carinaticeps Nixon, 1930: 401, 402, 410.

Valid name

Copter a carinaticeps (Nixon, 1930).

Summary of types

Syntype §, BMNH number 9.671. Syntype BMNH number 9.671b. Syntype $.

Primary type data

Type; Mossel Bay/ Cape Province/ October 1921; S.Africa/R.E. Turner/Brit. Mus./ 1921-450; Galesus/

turneri Nixon/ v. carinaticeps/ Nixon/ $ type 1930 (syntype $). Type; Mossel Bay/ Cape Province/

15-31.vii.1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1921-315; Galesus/ turneri Nixon/ v. carinaticeps/

Nixon/ $ type 1930 (syntype $). Paratype; Port St. John/Pondoland/ Sept. 1923; S.Africa/R.E.Turner/

Brit. Mus./ 1923-510; Galesus/ turneri Nixon/ v. carinaticeps/ Nixon/ $ paratype 1930/ G.Nixon det.

(syntype $).

Type locality

South Africa, Western Cape Province, Mossel Bay, but evidently including Eastern Cape Province, Port

St Johns.

Remarks

There are one female and two males agreeing with Nixon’s description and labelled by him as Galesus

turneri v. carinaticeps. Of these, two are labelled as “type” and the third as “paratype”; however, since

neither a unique holotype or lectotype has yet been fixed, all of the type series is syntypic (Article 73.2).

The specimen labelled as “paratype” is from Port St John, not Mossel Bay; however, this appears to be

a lapsus by Nixon. It appears he included it in the description since he referred to two males, so it is not

excluded here from the type series. The two syntypes labelled as “type” are mounted on card points, the

other is mounted on a card.

Galesus {Scltizogalesus) longiceps Nixon, 1930: 400, 402, 410.

Valid name

Copter a longiceps (Nixon, 1930).

Summary of types

Syntype $, BMNH number 9.666. Syntype S, BMNH number 9.666b.

Primary type data

Type; Port St. John/ Pondoland/ Sept. 1923; S.Africa/ R.E.Turner/ Brit. Mus./ 1923-510; Galesus/

longiceps/ Nixon/ $ type 1930 (syntype §). Type; Mossel Bay/ Cape Province/ April 1921; S.Africa/

R.E.Turner/Brit. Mus./ 1923-510; Galesus/ longiceps/Nixon/ $ type 1930 (syntype $).

European Journal of Taxonomy 75: 1-123 (2014)

Type locality

South Africa, Eastern Cape Province, Port St Johns; Western Cape Province, Mossel Bay.

Remarks

There are one female and one male agreeing with Nixon’s description and labelled by him as Galesus

longiceps , both of which are labelled as “type”. Since neither a unique holotype or lectotype has yet been

fixed, both specimens are syntypes (Article 73.2). The female syntype is mounted on a card point, the

male syntype is mounted on a card, and both are entire.

Galesus (Schizogalesus) silvestrii var. nigricornis Nixon, 1930: 401, 412.

Valid name

Coptera mosselensis nom. nov.

Summary of types

Syntype 3 , BMNH number 9.658. Syntypes 3 33 -

Primary type data

Type; Mossel Bay/ Cape Province/ Dec. 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-25; Galesus/

silvestrii Kieff./ var. nigricornis/ Nixon (syntype 3). Paratype; Mossel Bay/ Cape Province/ 18-30.

xi. 1921; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-2; Galesus/ silvestrii Kieff./ var. nigricornis/ Nixon

(syntypes 233 )- Paratype; Mossel Bay/ Cape Province/ 5-31 .vii. 1921; S.Africa/ R.E.Turner/ Brit. Mus./

1921-315; Galesus/ silvestrii Kieff/ var. nigricornis/Nixon (syntype 3 ).

Type locality

South Africa, Western Cape Province, Mossel Bay.

Remarks

There are four males agreeing with Nixon’s description and labelled by him as Galesus silvestrii v.

nigricornis. Of these, one is labelled as “type” and rest as “paratype”; however, since neither a unique

holotype or lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). All syntypes

are mounted on cards and are entire. This taxon is not conspecific with Coptera silvestrii ; as well as

the characters given by Nixon 1930, it differs also in the shape of the frontal projections (outer angle

forming a longer, more acute projection in silvestrii ) and in the size of the lateral fovea on the scutellar

disc (larger in nigricornis , approaching the size of the basal scutellar fovea) and should be given the

rank of species. Galesus ( Schizogalesus) silvestrii var. nigricornis Nixon, 1930 is permanently invalid

as a junior primary homonym of Galesus (G.) foersteri var. nigricornis Kieffer, 1911 and no other name

appears to be available for it, so it is given a replacement name here based on the name of the type

locality, Mossel Bay.

Galesus {Schizogalesus) difficilis Nixon, 1930: 401, 402, 412.

Valid name

Coptera difficilis (Nixon, 1930).

Summary of types

Syntype §, BMNH number 9.657. Syntype 3 , BMNH number 9.657b. Syntypes 2 § §, 12 33 -

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Type; Mossel Bay/ Cape Province/ Febr. 1922; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-97; Galesus/

difficilis/ 1930 Nixon/ type $ (syntype $). Paratype; Mossel Bay/ Cape Province/ Febr. 1922; S.Africa/

R. E.Turner/ Brit. Mus./ 1922-97; Galesus/ difficilis/ 1930 Nixon (syntype $). Paratype; Zululand/

Mtunzini/ 7.vi.l926; S.Africa/ R.E.Turner/ Brit. Mus./ 1926-277; Galesus/ difficilis/ 1930 Nixon

(syntype $). Type; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-129; Mossel Bay/ Cape Province/ 1-13.

iii.1922; Galesus/ difficilis/ 1930 Nixon/ type $ (syntype $). Paratype; Mossel Bay/ Cape Province/

1-13.iii. 1922; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-129; Galesus/ difficilis/ 1930 Nixon (syntypes

4c? c?)- Paratype; Mossel Bay/ Cape Province/ Febr. 1922; S.Africa/ R.E.Turner/ Brit. Mus./ 1922-97;

Galesus/ difficilis/ 1930 Nixon (syntypes 4c? c?). Paratype; Mossel Bay/ Cape Province/ 15-28.iii. 1922;

S. Africa/ R.E.Turner/ Brit. Mus./ 1922-158; Galesus/ difficilis/ 1930 Nixon (syntypes 3c? c?)- Paratype;

Umatata/ Transkei/ 18.ii-18.iii. 1923; S.Africa/ R.E.Turner/ Brit. Mus./ 1923-189; Galesus/ difficilis/

1930 Nixon (syntype $).

Type locality

South Africa, Eastern Cape Province, Transkei; KwaZulu-Natal, Mtunzini; Western Cape Province,

Mossel Bay.

Remarks

There are three females and thirteen males agreeing with Nixon’s description and labelled by him as

Galesus difficilis. Of these, two are labelled as “type” and the rest as “paratype”; however, since neither

a unique holotype or lectotype has yet been fixed, all of the type series is syntypic (Article 73.2). All the

male syntypes and the female syntype from Mossel Bay labelled as “paratype” are mounted on cards and

are entire. The other female syntypes are mounted on card points and are entire.

Trichopria lewisi Nixon, 1940: 59, fig. la-d.

Valid name

Trichopria lewisi Nixon, 1940.

Summary of types

Holotype $, BMNH number 9.706, by original designation. Paratypes 24$ $, <f?.

Primary type data

Type; Kenya Colony/ Kabete/ Ex Glossina/ brevipalpis &/ G. fuscipleuris/ Dr E.A.Lewis; Trichopria/

lewisi Nixon/ $ type 1940; Pres, by/Imp. Inst. Ent./B.M. 1940-181.

Type locality

Kenya, Kabete.

Remarks

Although Nixon mentioned 24 $ $ and 1 $ as paratypes, there are 63 $ $ and 3 $$ specimens labelled

as paratypes, so it is not clear which are Nixon’s original paratypes. While the error in labelling paratypes

is unfortunate, it is not likely to be a problem, as the holotype is certainly recognised and all the type

specimens appear to be the same species. Nixon’s preparation of the male genitalia is missing. The

holotype is mounted on a card point and is entire. It is worth noting a singular feature of the male of this

species which was not recorded by Nixon: the male head is truncate behind, and the genal pubescence

is long, and, unlike other Trichopria , turned back to form a row from the top to the bottom of the gena.

European Journal of Taxonomy 75: 1-123 (2014)

defining a vertical channel behind each gena, between the back of the head and the pronotum. By

analogy with the observed mating behaviour of other Trichopria it is predicted that this channel is used

to constrain the female antenna during courtship.

Spilomicrus myrmecophilus Nixon, 1947: 787, figs 1-3.

Valid name

Spilomicrus myrmecophilus Nixon, 1947.

Summary of types

Holotype §, BMNHnumber 9.646, by original designation. Paratypes 3

Primary type data

Type; 93/ Mauritius/ 26.x. 1945/ R.Mamet; In nest of Solenopsis/ mameti/ Donis.; Spilomicrus/

myrmecophilus/Nixon/ type §, 1947; Pres, by/Imp. Inst. Ent./B.M. 1947-417.

Type locality

Mauritius, Corps de Garde Mountain.

Remarks

The holotype is on a card point and is entire.

Entomacis penelope Nixon, 1980: 12.

Valid name

Entomacis penelope Nixon, 1980.

Summary of types

Holotype $, BMNH number 9.809, by original designation.

Primary type data

Eire/ Co. Sligo/Bundoran/ vii.1933/ G.E.J.Nixon; Entomacis/ penelope/Nixon/ type §, 197.

Type locality

Ireland, County Sligo, Bundoran.

Remarks

The holotype is mounted on a card point and is entire.

Entomacis laertes Nixon, 1980: 13.

Valid name

Entomacis graeffei Kieffer, 1909.

Summary of types

Holotype §, BMNH number 9.810, by original designation. Paratypes 5 $ §.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Sweden, Sk./ Ring sjo/ 27.vi.1938/ D.M.S.R & J.F.P./ B.M. 1938-414; Entomacis/ laertes Nixon/ type

¥, 197.

Type locality

Sweden, Skane County, Ring Sjo.

Remarks

The holotype is mounted on a card point and is entire.

Spilomicrus pelion Nixon, 1980: 17, figs 33, 40.

Valid name

Spilomicrus crassiclavis Kieffer, 1911.

Summary of types

Holotype S, BMNH number 9.811, by original designation. Paratype f.

Primary type data

Surrey/Horsley/ 14.vi.1930/ G.Nixon; Spilomicrus/ pelion Nixon/ type S, 197.

Type locality

England, Surrey, Horsley.

Remarks

The holotype is mounted on a card point and is entire.

Aneurhynchus fragilis Nixon, 1980: 23, fig. 60.

Valid name

Aneurhynchus fragilis Nixon, 1980.

Summary of types

Holotype ?, BMNH number 9.814, by original designation.

Primary type data

England:/ Glos. E./ Badgeworth/ field mushroom/ 18.ix. 1963; Pres, by/ A.Muncaster/ B.M. 1963-697;

Aneurhynchus/ fragilis Nixon/ type ¥ 1969.

Type locality

England, Gloucestershire, Badgeworth.

Remarks

The holotype is mounted on a card point, one antenna and the left mid leg are disarticulated, and the tips

of three tarsi are lost.

European Journal of Taxonomy 75: 1-123 (2014)

Aneurhynchus ariadne Nixon, 1980: 23, 24, fig. 67.

Valid name

Aneurhynchus ariadne Nixon, 1980.

Summary of types

Holotype BMNH number 9.815, by original designation. Paratypes 3 $ §.

Primary type data

France, Fi./ Morlaix/ 10.vi.1954/ .1F.Perk in s/ B.M. 1954-396; Aneurhynchus/ ariadne/ Nixon/ holotype

¥, 197.

Type locality

France, Morlaix.

Remarks

The holotype is mounted on a card point and is entire.

Aneurhynchus mese Nixon, 1980: 23, fig. 70.

Valid name

Aneurhynchus mese Nixon, 1980.

Summary of types

Holotype ?, BMNH number 9.816, by original designation. Paratypes 3 ¥ ¥•

Primary type data

Eng: Northants/ Spratton/ viii. 1975/1. & PGauld; Aneurhynchus/ mese Nixon/ type ¥, 197.

Type locality

England, Northamptonshire, Spratton.

Remarks

The holotype is mounted on a card point, with the right antenna disarticulated.

Aneurhynchus coryphe Nixon, 1980: 25, fig. 69.

Valid name

Aneurhynchus coryphe Nixon, 1980.

Summary of types

Holotype 3 , BMNH number 9.817, by original designation. Paratypes 3 33 -

Primary type data

France, Fi./ Huelgoat/ 3.vi. 1954/ J.F.Perkins/ B.M. 1954-396; Aneurhynchus/ coryphe Nixon/ type 3, 197.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Type locality

France, Huelgoat.

Remarks

The holotype is mounted on a card point and is entire.

Basalys orion Nixon, 1980: 26, fig. 81.

Valid name

Basalys orion Nixon, 1980.

Summary of types

Holotype §, BMNH number 9.818, by original designation. Paratype §.

Primary type data

Berks:/Windsor Forest/ ix,1931/H.St.J.Donisthorpe; Basalys/ orion Nixon/ type §, 197.

Type locality

England, Berkshire, Windsor Forest.

Remarks

The holotype is mounted on a card point and is entire.

Basalys cymodes Nixon, 1980: 27, 29, fig. 74.

Valid name

Basalys cymocles Nixon, 1980.

Summary of types

Holotype §, BMNH number 9.819, by original designation. Paratypes 3 $ §.

Primary type data

Surrey/ Ashtead/ 30.vii.1932/ G.Nixon; Basalys/ cymocles/Nixon/ type $, 197.

Type locality

England, Surrey, Ashtead.

Remarks

The holotype is mounted on a card point and is entire.

Basalys semele Nixon, 1980: 27.

Valid name

Basalys semele Nixon, 1980.

Summary of types

Holotype §, BMNH number 9.820, by original designation.

European Journal of Taxonomy 75: 1-123 (2014)

Primary type data

White Downs, near/ Dorking Surrey/ England/ lg. Boucek 29.8.1970; Basalys/ semele Nixon/ type §,

197.

Type locality

England, Surrey, White Downs near Dorking.

Remarks

The type is mounted on a card and is entire.

Basalys insignificans Nixon, 1980: 28.

Valid name

Basalys insignificans Nixon, 1980.

Summary of types

Holotype §, BMNH number 9.821, by original designation. Paratype §.

Primary type data

Wales: Glamorgan/Pontypridd/ 25.V.1975; grassland/ J.S.Noyes/B.M. 1975-265; Basalys/ insignificans/

Nixon/ type §, 197.

Type locality

Wales, Glamorgan, Pontypridd.

Remarks

The holotype is mounted on a card point and is entire.

Basalys singularis Nixon, 1980: 28.

Valid name

Basalys singularis Nixon, 1980.

Summary of types

Holotype $, BMNH number 9.823, by original designation. Paratype §.

Primary type data

Furry Glen/ Co. Du., A.W.S./ 26.9.37; Ireland/A.W.Stelfox coll./BMNH 1949-110; Basalys/ singularis/

Nixon/ type §, 197.

Type locality

Ireland, County Dublin, Furry Glen.

Remarks

The holotype is mounted on a card and is entire.

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Basalys euterpe Nixon, 1980: 30.

Valid name

Basalys semele Nixon, 1980.

Summary of types

Holotype S, BMNH number 9.822, by original designation. Paratypes 4 SS-

Primary type data

Co. Sligo/Trawallua/ 24-29.7.1933/ G.Nixon; Basalys/ euterpe/Nixon/ type c?, 197.

Type locality

Ireland, County Sligo, Trawallua.

Remarks

The holotype is mounted on a card point and is entire.

Basalys iphicla Nixon, 1980: 30.

Valid name

Basalys macroptera (Kieffer, 1911) syn. nov.

Summary of types

Holotype S, BMNH number 9.824, by original designation. Paratype S-

Primary type data

Inverness:/ Loch Morlich/ 24.vi. 1934/ R.B. & J.E.Benson; Pres, by the/ Trustees of the/ Godman Fund./

B.M. 1934-429; Basalys/ iphicla/Nixon/ type §, 197.

Type locality

Scotland, Inverness, Loch Morlich.

Remarks

The holotype is mounted on a card point and has al2-14 and both hind tarsi missing. This species is

evidently the same as Basalys macroptera , with which it shares a number of characters, most notably its

relatively large size, hairs on anterior margin of large tergite, straight carina just below apex of antennal

shelf, dark colour and northern distribution.

Corynopria aphrodite Nixon, 1980: 32, fig. 9.

Valid name

Monelata aphrodite (Nixon, 1980).

Summary of types

Holotype §, BMNH number 9.825, by original designation. Paratypes 3 $ §.

Primary type data

S.E.London/Norwood/ vii.1929/ G.Nixon; Corynopria/ aphrodite/Nixon/ type $, 1979.

European Journal of Taxonomy 75: 1-123 (2014)

Type locality

England, London, Norwood (as Tulse Hill in original description).

Remarks

The type series is from Norwood, not Tulse Hill as Nixon stated, although these places are very close;

evidently this was just an error in recording the locality by Nixon and should not prevent them from

being types. The holotype is mounted on a card point and is entire.

Trichopria isis Nixon, 1980: 33.

Valid name

Trichopria aequata (Thomson, 1858).

Summary of types

Holotype $, BMNH number 9.826, by original designation. Paratypes 2 $ §.

Primary type data

Farley, S.H./ 12.vi. 1938/ R.B.Benson/B.M. 1938-190; Trichopria/ isis Nixon/ type §, 1977.

Type locality

England, Hants, Farley.

Remarks

The holotype is mounted on a card point and is entire.

Trichopriaprema Nixon, 1980: 34.

Valid name

Trichopria prema Nixon, 1980.

Summary of types

Holotype §, BMNH number 9.827, by original designation.

Primary type data

Boston Manor/ Mx., ix.1938; Trichopria/ prema/ Nixon/ type §, 197.

Type locality

England, Middlesex, Boston Manor.

Remarks

The holotype is mounted on a card point, and is entire.

Trichopria bifoveata Nixon, 1980: 34, 38, figs 92,102,104,124.

Valid name

Trichopria nixoni Notton, 1995.

100

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Summary of types

Lectotype $, BMNH number 9.894, designated by Notton (1995). Paralectotypes 7 $ $, 2 <$< 5 -

Primary type data

Devon/ Torquay dist./ Aug. 1929/ G.Nixon; Trichopria/ bifoveata K./ G.Nixon det. 1975; Lectotype §/

Trichopria/bifoveata/Nixon, 1980/ det. D.G.Notton,1992.

Type locality

England, Devon, Torquay district.

Remarks

Although Nixon attributed the name to Kieffer, there is no species of that name described by Kieffer

and Nixon (1980) must be considered the author, the description provided in the key being sufficient

to make the name available (Notton 1995). The lectotype is mounted on a card point, has the right fore

wing folded and the left hind tarsus broken off and stuck to the right fore wing.

Trichopria credne Nixon, 1980: 34, fig. 119.

Valid name

Trichopria credne Nixon, 1980.

Summary of types

Holotype $, BMNH number 9.828, by original designation. Paratypes 2 $ §.

Primary type data

Surrey/ Ashtead/ 6.viii. 1932/ G.Nixon; Trichopria/ credne Nixon/ type $, 197.

Type locality

England, Surrey, Ashtead.

Remarks

The holotype is mounted on a card point and is entire.

Trichopria alifera Nixon, 1980: 34, fig. 118.

Valid name

Trichopria incrassata (Jansson, 1955).

Summary of types

Holotype §, BMNH number 9.829, by original designation. Paratypes 2 $ §.

Primary type data

England/ Surrey/ Weybridge/ viii. 1944/ G.E.J.Nixon; Trichopria/ alifera Nixon/ type $, 197.

Type locality

England, Surrey, Weybridge.

101

European Journal of Taxonomy 75: 1-123 (2014)

Remarks

The holotype is mounted on a card point, the left fore leg, right hind leg and apex of the left hind tarsus

are lost and the right fore wing is chipped.

Trichopria nana Nixon, 1980: 36.

Valid name

Trichopria paludicola Notton, 1995.

Summary of types

Holotype §, BMNH number 9.830, by original designation.

Primary type data

Cambs:/Wicken Fen/ 10.ix.l934/H.St.J.Donisthorpe; Trichopria/ nana Nixon/ type §, 197.

Type locality

England, Cambridgeshire, Wicken Fen.

Remarks

The holotype is mounted on a card point and is entire.

Trichopria sequester Nixon, 1980: 37, fig. 97.

Valid name

Trichopria subimpressa (KiefFer, 1911).

Summary of types

Holotype 3 , BMNH number 9.831, by original designation. Paratypes 4 33 -

Primary type data

England, SE:/ Epping Forest/ 14-15.viii.1954/ J.A. & D.J.Clark/ B.M. 1954-521; Trichopria/ sequester/

Nixon/ type 3, 197.

Type locality

England, Essex, Epping Forest.

Remarks

The holotype is mounted on a card point and is entire.

Trichopria crassifemur Nixon, 1980: 38, fig. 91.

Valid name

Trichopria crassifemur Nixon, 1980.

Summary of types

Holotype 3 , BMNH number 9.832, by original designation. Paratypes 18 33 -

102

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Surrey:/ Ashtead/ 10.viii.1930/ G.Nixon; Trichopria/ crassifemur/ Nixon/ type S, 197.

Type locality

England, Surrey, Ashtead.

Remarks

The holotype is mounted on a card point and its right antenna is lost.

Trichopria stelenes Nixon, 1980: 38.

Valid name

Trichopria credne Nixon, 1980.

Summary of types

Holotype S, BMNH number 9.833, by original designation. Paratype S-

Primary type data

Burnham Beeches/ Bucks., England/ Boucek 14.ix.74; Trichopria/ stelenes Nixon/ type S, 197.

Type locality

England, Buckinghamshire, Burnham Beeches.

Remarks

The holotype is mounted on a card and is entire. The synonymy of Notton (1995) is confirmed here based

on similarities in the morphology of the head, scutellum and petiole, and the coincident distribution and

simultaneous collection of males and females.

Phaenopria miron Nixon, 1980: 39: 40, fig. 7.

Valid name

Trichopria fucicola (Walker, 1834).

Summary of types

Holotype §, BMNH number 9.834, by original designation. Paratypes 7 $ §.

Primary type data

S.E.London/ Norwood/ ix.1929/ G.Nixon; from rotten/ fungus; Phaenopria/ miron Nixon/ holotype $,

1979.

Type locality

England, London, West Norwood.

Remarks

The holotype is mounted on a card point and is entire. Nixon says there were nine paratypes; however,

only seven were found.

103

European Journal of Taxonomy 75: 1-123 (2014)

Trichopriapolita Notton, 1993: 139, figs 1-13.

Valid name

Trichopria polita Notton, 1993.

Summary of types

Holotype §, BMNH number 9.884, by original designation.

Primary type data

UK: Cambs: Chippenham/Fen: carr-reedbed:/TL650693: Malaise/ 22.8-5.9.1985/ J.Field; Holotype §/

Trichopria/ polita/ D.G.Notton.

Type locality

England, Cambridgeshire, Chippenham Fen.

Remarks

The holotype is mounted on a card and is entire.

Trichopria striata Notton, 1993: 141, figs 14-21.

Valid name

Trichopria striata Notton, 1993.

Summary of types

Holotype §, BMNH number 9.885, by original designation. Paratype f.

Primary type data

NCC fen survey/ water/ trap; UK: W.Norfolk/ Brancaster/ TF768443/ 2-16.9.1988: A.Foster; $

Holotype/ Trichopria striata/ Notton, D.G., 1992.

Type locality

England, Norfolk, Brancaster.

Remarks

The holotype is mounted on a card and is entire.

Trichopria melanopa scilloniensis Notton, 1993: 143, figs 22-28.

Valid name

Trichopria subimpressa scilloniensis Notton, 1993.

Summary of types

Holotype §, BMNH number 9.889, by original designation. Paratypes 9 $ $, 8 SS-

Primary type data

England: Scilly/ Tresco/ 21.ix. 1975; J.S.Noyes/ Brit. Mus./ 1975-479; $ Holotype/ Trichopria/

melanopa/ scilloniensis n. ssp./ det. D.G.Notton 1991.

104

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Type locality

England, Isles of Scilly, Tresco.

Remarks

The holotype is mounted on a card and is entire.

Diapria luteipes Notton, 1993: 145, figs 33-42.

Valid name

Diapria luteipes Notton, 1993.

Summary of types

Holotype §, BMNH number 9.886, by original designation.

Primary type data

NCC survey/pitfall/ trap; UK: W.Norfolk: Middle/Haring: TT989856/ 1-15.9.1988/ A.Foster; Holotype

5/ Diapria/ luteipes/ D.G.Notton.

Type locality

England, Norfolk, Middle Harling.

Remarks

The holotype is mounted on a card and is entire.

Diapria cava Notton, 1993: 145,147, figs 43-47.

Valid name

Diapria cava Notton, 1993.

Summary of types

Holotype S, BMNH number 9.887, by original designation.

Primary type data

NCC code - water/ trap: RSPB 3.A/ Open Pits; UK: East Kent/ Dungeness: TR0718/ 22.8-5.9.1988/

R.K.A.Morris NCC.

Type locality

England, Kent, Dungeness.

Remarks

The holotype is mounted on a card and is entire.

Trichopria quadrifida Notton, 1994: 201, fig. la.

Valid name

Trichopria quadrifida Notton, 1994.

105

European Journal of Taxonomy 75: 1-123 (2014)

Summary of types

Holotype §, BMNH number 9.890, by original designation. Paratype §.

Primary type data

Kent/Eynsford/ Sl.viii. 1929/ G.Nixon; Phaenopria/ incrassata Janss./ G.Nixon det. 1978; $ Holotype/

Trichopria/ quadrifida/ n. sp./D.G.Notton, 1992.

Type locality

England, Kent, Eynsford.

Remarks

The holotype is mounted on a card point, and the left hind leg is missing.

Diapria drosopliilae Perkins, 1910: 629.

Valid name

Trichopria drosophilae (Perkins, 1910).

Summary of types

Syntype $.

Primary type data

Honolulu/ Oahu 6-19-05; Diapria/ foreign; R.C.L.Perkins coll./ B.M. 1942-95/ Hawaii.

Type locality

Hawaii, Oahu, Honolulu.

Remarks

The syntype is mounted on a card point with a small fly and is entire.

Diapria modesta Ratzeburg, 1848: 144.

Valid name

Trichopria modesta (Ratzeburg, 1848).

Summary of types

Neotype §, BMNH number 9.891, designated by Notton (1995).

Primary type data

46-16; Diapria/ modesta/ Foerst./ Aachen; Neotype; Neotype/ Diapria/ modesta/ Ratzeburg 1848/ des./

D.G.Notton, 1994.

Type locality

Germany, Aachen.

106

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Remarks

This species was described by Ratzeburg from an unspecified number of specimens, probably female,

sent by A. Forster, most probably collected in Germany. The original type(s) is lost and a neotype was

designated by Notton (1995). The neotype is mounted on a card point and the point has been mounted

on a card. The tip of the left hind tarsus is disarticulated.

Spilomicrus quadriceps Smith, 1878: 6.

Valid name

Spilomicrus quadriceps Smith, 1878 stat. rev.

Summary of types

Syntype BMNH number 9.682.

Primary type data

New/ Zeal.; Type; New/ Zeal./ 77-100; Spilomicrus/ quadriceps/ type Sm.

Type locality

New Zealand, Otago.

Remarks

This species was described from an unspecified number of males, of which one was found. The syntype

is mounted on a card, generally dirty, the right a9-13 are disarticulated, and the tip of the left hind tarsus

is missing. This species was placed in Malvina by Masner (1965); however, Malvina is here considered

a derived species group of Spilomicrus and the original combination is reinstated.

Psilus fucicola Walker, 1834: 117.

Valid name

Trichopria fucicola (Walker, 1834).

Summary of types

Lectotype BMNH number 9.799, designated by Masner (1965). Paralectotypes 2 $$.

Primary type data

Brit. Mus./ Walker coll.; Type; Psilus?/ fucicola/ see MS/ stood under this/ name in old B.M./ coll.;

fucicola; Lecto-/type/L.M.; Selected as/ lectotype by/L.Masner 28.xi.61; Lectotype c?/Psilus fucicola/

Walker, 1834: 117/ des. L.Masner, 1965.

Type locality

England, Devon, Torquay.

Remarks

Three syntypes were found, of which one had been designated lectotype by Masner (1965). Nine more

syntypes (now paralectotypes) are known to be in Dublin (Notton & O’Connor 2004). The lectotype is

mounted on a card and is entire.

107

European Journal of Taxonomy 75: 1-123 (2014)

Diapria apicalis Walker, 1860: 359.

Valid name

Trichopria walkeri (Dalla Torre, 1890) comb. nov.

Summary of types

Syntype §, BMNH number 9.895.

Primary type data

63-52; apicalis; Diapria.

Type locality

Sri Lanka (as Ceylon).

Remarks

The syntype is mounted on a card and is entire. The number 63-52 refers to Natural History Museum

Entomology register number 1863-52, which includes a group of insects described as “Type specimens

described by Mr Walker in the Annuals & Magazine for 1858-59-60”. Diapria apicalis was renamed

Diapria walkeri by Dalla Torre (1890) because of homonymy with Diapria apicalis (Say 1836). The

type of D. apicalis is a Trichopria so Trichopria walkeri is a new combination.

Monelata nigra Whittaker, 1930: 133.

Valid name

Monelata solida (Thomson, 1858).

Summary of types

Holotype §, BMNH number 9.688, by original designation. Paratypes 3 $ §.

Primary type data

Type; Type; Hollybum/B.C. 8.ix.29/ coll. O.W.; Canada:/ O.Whittaker coll./per W.H.Storey/B.M. 1947-

212; 4379/ Monelata/ nigra, Whitt./ §/ Det. O.Whittaker.

Type locality

Canada, British Columbia, Hollybum.

Remarks

The description was based on 12 $ ? • Whittaker referred to paratypes, implying that there was a holotype.

The holotype is mounted on a card and is entire.

Galesus fissus Wollaston, 1858: 25.

Valid name

Copterafissa (Wollaston, 1858).

Summary of types

Syntype §, BMNH number 9.652. Syntypes 3 $ §.

108

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Primary type data

Type; Madeira/ Wollaston/ 55.7; Galesus/ fissus W. (syntype §). Rib. da/ Janella; Madeira/ Wollaston/

55.7 (syntype §). 1462; Madeira/Wollaston/ 55.7 (syntypes 2 §§).

Type locality

Madeira.

Remarks

This species was described from at least five syntypes, since Wollaston mentions five different localities:

S. Antonio da Serra; Ribeira da Janella; Feijaa de Corte; another locality at intermediate altitude; and

another locality at high altitude. Not all the syntypes could be assigned to these localities with certainty;

although two of them have the locality code 1462, a key to localities codes for Wollaston’s Hymenoptera

has not been found. The four syntypes are mounted on cards: one is entire; one has the right hind

telotarsus disarticulated; one has the right al2 missing; and the last one has the wings much torn.

Synonymic List

Subfamily Diapriinae Haliday, 1833

Tribe Psilini Fallen, 1812

Aneurhynchus Westwood, 1832

A. ariadne Nixon, 1980

A. coryphe Nixon, 1980

A. firagilis Nixon, 1980

A. indicus Dodd, 1920

A. mese Nixon, 1980

A. nodicornis Marshall, 1867

A. obliquus Kieffer, 1911

Aneuropria Kieffer, 1905

= Mantara Dodd, 1920 syn. nov.

A. bifurcata (Dodd, 1920) comb. nov.

Copter a Say, 1836

C. abdominalis (Nixon, 1930)

C. angulata Muesebeck, 1980

C. bidens (Nixon, 1930)

C. carinaticeps (Nixon, 1930)

C. cratocerus (Cameron, 1912) comb. nov.

C. difficilis (Nixon, 1930)

C. distans Muesebeck, 1980

C. distincta (Nixon, 1930)

C. fissa (Wollaston, 1858)

C. hispanica (Kieffer, 1911)

C. inquisitor (Nixon, 1930)

C. longiceps (Nixon, 1930)

C. modesta (Nixon, 1930)

C. mosselensis nom. nov.

= C. nigricornis (Nixon, 1930) preocc.

C. muscidorum (Dodd, 1920)

109

European Journal of Taxonomy 75: 1-123 (2014)

C. pijiguaorum Notton, Montilla & Garda nom. nov.

= C. sexpunctata Montilla & Garcia, 2008 preocc.

C. pomonellae Muesebeclc, 1980

C. punctulata (Kieffer, 1907)

C. quadridens (Nixon, 1930)

C. rectangular is (Nixon, 1930)

C. robustior (Silvestri, 1913)

= C. magnifica (Nixon, 1930)

C. sexpunctata (Ashmead, 1893) comb. nov.

= C. bipunctata (Ashmead in Riley, Ashmead & Howard, 1894) syn. nov.

C. silvestrii ( Kieffer, 1913)

C. strauziae Muesebeclc, 1980

C. turneri (Nixon, 1930)

Labolips Forster, 1856

L. innnpta Hal id ay, 1857

Psilus Panzer, 1801

P. aquilonius Muesebeck, 1980

P. cornutus Panzer, 1801

= P. gracilipes (Kieffer, 1907)

P. dissidens Muesebeck, 1980

P. fords Muesebeck, 1980

P. fuscipennis (Curtis, 1831)

= P. walkeri (Kieffer, 1907) syn. nov.

= P. nigricornis (Kieffer, 1911) syn. nov.

= P. bispinosus ( Kieffer, 1911)

= P. cameroni (Kieffer, 1911)

P. masneri Muesebeck, 1980

P. masoni Muesebeck, 1980

P. parvulus (Kieffer, 1911)

Tribe Spilomicrini Ashmead, 1893

Bruchopria Kieffer, 1921

= Aulatopria Brethes, 1927

B. tucumana (Brethes, 1927)

Chilomicrus Masner & Garcia, 2002

C. pecki Masner & Garcia, 2002

Doddius Masner & Garcia, 2002

D. rugosus (Dodd, 1920)

Entomacis Forster, 1856

E. curdcornis Kieffer, 1912

E. fallax (Kieffer, 1912)

E.filiformis (Ashmead in Riley, Ashmead & Howard, 1894)

E. flaviclava (Kieffer, 1912)

E. graeffei Kieffer, 1909

= E. laertes Nixon, 1980

110

NOTTON D.G., Diapriinae types at The Natural History Museum, London

E. latipennis (Ashmead in Riley, Ashmead & Howard, 1894)

E. longicornis Kieffer, 1912

E. penelope Nixon, 1980

E. subaptera Early, 1980

Hemilexomyia Dodd, 1920

H. spinosa Early, 1980

Idiotypa Forster, 1856

I. nigriceps Kieffer, 1909

= /. nigriceps Kieffer, 1911 preocc., syn. nov.

I. pallida Ashmead, 1893

= I. pallida Ashmead in Riley, Ashmead & Howard, 1894 preocc., syn. nov.

I. tinctipennis (Cameron, 1888)

Neurogalesus Kieffer, 1907

N. carinatus Kieffer, 1907

Paramesius Westwood, 1832

P. brachypterus (Thomson, 1858)

= P. angustipennis Kieffer, 1911

= P. earner wn Kieffer, 1911

P. laetus Fouts, 1927

P. longior Dodd, 1920

P. rufipes (Fonscolombe, 1832)

= P. inchoatus Kieffer, 1911

= P. subinermis Kieffer, 1911

= P. nigricornis Kieffer, 1911

= P. dentatus Kieffer, 1911

= P. minor (Kieffer, 1911)

P. thoracicus Ashmead in Riley, Ashmead & Howard, 1894

Pentapria Kieffer, 1905

P. chiriquensis (Cameron, 1888)

Rostropria Early & Naumann, 1990

R. simplex Early & Naumann, 1990

R. spiniventris Early & Naumann, 1990

Spilomicrus Westwood, 1832

= Malvina Cameron, 1889 syn. nov.

= Antarctopria Braes in Tillyard, 1920 syn. nov.

S. affinis (Dodd, 1920)

S. aneurus Ashmead in Riley, Ashmead & Howard, 1894

S. annulicornis Kieffer, 1911

S. aterrimus (Dodd, 1920) comb. nov.

S. atriclavus Ashmead, 1893

= S. kiefferi Fouts, 1925

S. barnesi Early & Horning, 1978

S. boweni Masner, 1991

S. campbellanus (Yoshimoto, 1964) comb. nov.

Ill

European Journal of Taxonomy 75: 1-123 (2014)

S. canaliculatus (Cameron, 1888)

S. coelopae (Early, 1978) comb. nov.

S. crassiclavis Kieffer, 1911

= S. pelion Nixon, 1980

S. curvispina (Cameron, 1913)

S. diomedeae (Early, 1978) comb. nov.

S. distinctus Masner, 1991

S. exul Masner, 1991

S. fasdatipennis (Cameron, 1888)

S. helosciomyzae (Early & Horning, 1978) comb. nov.

S. hemipterus Marshall, 1868

= S. pedissequus Kieffer, 1911

= S. pedisequus Kieffer, 1916 unjustified emend.

S. inornatus Masner, 1991

S. insulae (Early, 1980) comb. nov.

S. integer Thomson, 1858

= S. variicornis Kieffer, 1911

S. kozlovi nom. nov.

= S. punctatus Kozlov, 1978 preocc.

S. latigaster (Brues in Tillyard, 1920) comb. nov.

S. lubomiri Ozdikmen, 2010

= S. superbus Masner, 1991 preocc.

= S. masneri Ozdikmen, 2011

S. maculipennis (Cameron, 1888)

S. myrmecophilus Nixon, 1947

S. obsoletus (Dodd, 1920)

S. painstris Masner, 1991

S. paucisetis (Dodd, 1920)

S. picicorms (Cameron, 1913)

S. pilgrim I Early, 1978

S. pumilio Masner, 1991

S. punctatus (Cameron, 1889) comb. nov.

S. quadriceps Smith, 1878 stat. rev.

S. rekohua (Early, 1978) comb. nov.

S. sanbornei Masner, 1991

S. stepheni Masner, 1991

S. stigmaticalis Westwood, 1832

= S. tripartitus Kieffer, 1911

S. sylvicola Masner, 1991

S. vulgaris Ashmead in Riley, Ashmead & Howard, 1894

Tribe Diapriini Haliday, 1833

A canthopria Ashmead, 1896

A. crassicornis Ashmead, 1896

A. triangularis (Ashmead in Riley, Ashmead & Howard, 1894)

Apopria Masner & Garcia, 2002

A. coveri Masner & Garcia, 2002

112

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Asolenopsia Kieffer, 1921

A. schwarzmaieri Borgmeier, 1939

Basalys Westwood, 1833

= Microgalesus Kieffer, 1912 syn. nov.

B. abruptus Thomson, 1858

= B. convexus (Kieffer, 1911)

B. bifoveatus (Kieffer, 1911)

= B. unifoveatus (Kieffer, 1911)

B. ciliatus (Kieffer, 1911)

B. clavatus (Ashmead, 1893)

= B. clavatus (Ashmead in Riley, Ashmead & Howard, 1894)

B. collaris Kieffer, 1911

B. cymocles Nixon, 1980

B. exsul (Kieffer, 1912)

B. flavidipes (Kieffer, 1912)

= B. flavipes (Ashmead, 1896) preocc.

B. formicarum (Kieffer, 1911)

= B. rufiscapus Kieffer, 1911 preocc.

= B. mill lens is (Masner, 1965)

B.fumipennis Westwood, 1833

= B. atricrus ( Kieffer, 1911)

B. grenadae (Kieffer, 1912)

= B. grenadensis (Ashmead, 1896: 807) preocc.

B. grenadensis (Ashmead, 1896: 803)

B. insignificans Nixon, 1980

B. longipennis (Kieffer, 1911)

B. luctuosus (Kieffer, 1911)

B. macroptera (Kieffer, 1911)

= B. sulcatus (Kieffer, 1911)

= B. iphicla Nixon, 1980 syn. nov.

B. nigrescens (Fouts, 1927)

B. orion Nixon, 1980

B. parvus Thomson, 1858

= B. pedes ter (Kieffer, 1907)

= B. alacer (Kieffer, 1911)

B. pedisequa (Kieffer, 1911)

= B. cursitans (Kieffer, 1911) syn. nov.

= B. fuliginosi (Box, 1921)

B. pleuralis (Ashmead, 1896)

B. quadrideas (Kieffer, 1912) comb. nov.

B. scoticus (Kieffer, 1911)

B. semele Nixon, 1980

= B. euterpe Nixon, 1980

B. semirufus (Kieffer, 1912)

B. singularis Nixon, 1980

B. thoracicus (Ashmead in Riley, Ashmead & Howard, 1894)

Diapria Latreille, 1796

D. am/Notton, 1993

113

European Journal of Taxonomy 75: 1-123 (2014)

D. luteipes Notton, 1993

Doliopria Kieffer, 1910

D. magniclavata (Ashmead, 1896) comb. nov.

Eladio Masner & Garcia, 2002

E. cruzi Masner & Garcia, 2002

Hansonia Masner & Garcia, 2002

H. pauli Masner & Garcia, 2002

Lepidopria Kieffer, 1911

L. lloydi (Ferriere, 1935)

Leucopria Masner & Garcia, 2002

L. cylindricornis Masner & Garcia, 2002

Mimopria Holmgren, 1908

M. barbata Borgmeier, 1939

M. campbellorum Masner, 1976

M. comes Borgmeier, 1939

Mimopriella Masner & Garcia, 2002

M. splendens (Borgmeier, 1939)

Monelata Forster, 1856

M. aphrodite (Nixon, 1980)

M. solida (Thomson, 1858)

= M. nigra Whittaker, 1930

Neivapria Borgmeier, 1939

N. penicillata Borgmeier, 1939

Notoxoides Ashmead, 1903

N. pedissequa (Borgmeier, 1939)

N. pronotalis (Borgmeier, 1939)

Platymisch oides Ashmead, 1901

P. molokaiensis Ashmead, 1901

Platymisch us We s two o d, 1832

P. dilatatus Westwood, 1832

= P. pedester (Kieffer, 1911)

Termitopria Muesebeck, 1965

T. sheasbyi Muesebeck, 1965

Tetramopria Wasmann, 1899

T. cincticollis Wasmann, 1899

= T. donisthorpei Kieffer, 1911

= T. femoralis Kieffer, 1911

114

NOTTON D.G., Diapriinae types at The Natural History Museum, London

Townesella Huggert & Masner, 1983

T. marjoriae Huggert & Masner, 1983

Trichopria Ashmead, 1893

= Xyalopria Kieffer, 1907 syn. nov.

T. aequata (Thomson, 1858)

= T. inaequalis (Kieffer, 1911)

= T. variipes (Kieffer, 1911)

= T. isis Nixon, 1980

T. angulifera (Ashmead, 1896)

T. ashmeadi Kieffer, 1912

= T. grenadensis Ashmead, 1896: 808 preocc.

T. atriceps Ashmead in Riley, Ashmead & Howard, 1894

T. balthazari (Ashmead, 1896)

T. basalis (Thomson, 1858)

= T. ciliaris Kieffer, 1911

T. bifoveata Ashmead, 1896

T. cameroni (Kieffer, 1909)

= T. cameroni (Kieffer, 1911)

T. capensis robustior Ferriere, 1935

T. confusa Masner, 1965

= T. grenadensis (Ashmead, 1896: 809, 810) preocc.

T. conotoma (Kieffer, 1911)

= T. bouceki (Masner, 1959) comb, nov., syn. nov.

T. eras si femur Nixon, 1980

T. credne Nixon, 1980

= T. stelenes Nixon, 1980

T. drosophilae (Perkins, 1910)

T. ensia Huggert, 1979

T. euvulgaris Huggert, 1979

T. fluminis nom. nov.

= T. nigriceps (Kieffer, 1907) comb, nov., preocc.

T. fucicola (Walker, 1834)

= T. miron (Nixon, 1980)

T. grenadensis (Ashmead, 1896: 805)

= T. grenadicola Kieffer, 1916

T. halterata (Kieffer, 1909)

= T. halterata (Kieffer, 1911) preocc., syn. nov.

= T. atricornis Kieffer, 1911

= I morleii (Morley, 1931)

T. hawaiiensis (Ashmead, 1901)

T. hyalinipennis (Thomson, 1858)

= T. tetratoma (Kieffer, 1911) preocc.

T. incrassata (Jansson, 1955)

= T. a l if era Nixon, 1980

T. insularis Ashmead in Riley, Ashmead & Howard, 1894

T. lewisi Nixon, 1940

T. mahensis (Kieffer, 1912)

T. melanopleura (Ashmead, 1896) stat. rev.

T. mellea (Ashmead in Riley, Ashmead & Howard, 1894)

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European Journal of Taxonomy 75: 1-123 (2014)

T. modesta (Ratzeburg, 1848)

T. myrmicae Huggert & Masner, 1983

T. neotropica Masner, 1965

= I affinis Ashmead, 1896 preocc.

T. nigra (Nees, 1834)

= T. inermis Kieffer, 1909

= I fimbriata Kieffer, 1911

T. nigriceps (Ashmead in Riley, Ashmead & Howard, 1894) comb. nov.

T. nigriclavata (Ashmead, 1896)

T. nigricornis (Marshall, 1868)

= T. subterranea (Kieffer, 1911)

= T. donisthorpei (Kieffer, 1913)

T. nirva Huggert, 1977

T. nixoni Notton, 1995

= T. bifoveata Nixon, 1980 preocc.

T. obscura Masner, 1965

= T. nigricornis (Ashmead, 1896) preocc.

T. occidentals (Fouts, 1927)

T. oogaster (Thomson, 1858)

= T. nocticolor (Kieffer, 1911)

T. pallida ( Ashmead in Riley, Ashmead & Howard, 1894)

T. paludicola Notton, 1995

= T. nana Nixon, 1980 preocc.

T. peraffinis (Ashmead, 1896)

T. percyi Huggert, 1979

T. pleural is Ashmead in Riley, Ashmead & Howard, 1894

T. polita Notton, 1993

T. prema Nixon, 1980

T. pseudosaxatilis Huggert, 1977

T. quadrifida Notton, 1994

T. saxatilis (Kieffer, 1912)

T. scotti (Kieffer, 1912)

T. seychellensis (Kieffer, 1912)

T. simillima (Ashmead in Riley, Ashmead & Howard, 1894)

T. smithii (Ashmead, 1896)

T. sociabilis Masner, 1965

= T. formicaria Kieffer, 1911 preocc.

T. spinosiceps (Dodd, 1920) comb. nov.

T. stomoxydis Huggert, 1977

T. striata Notton, 1993

T. subclavata (Ashmead in Riley, Ashmead & Howard, 1894)

T. subimpressa (Kieffer, 1911)

= T. melanopa (Kieffer, 1911)

= T. clavatipes (Kieffer, 1911)

= T. sequester Nixon, 1980

T. subimpressa scilloniensis Notton, 1993

T. tabanivora Fouts in Cameron, 1926

T. tachinidarum Ferriere, 1933

T. thermarum nom. nov.

= T. nigriceps (Kieffer, 1913) preocc.

116

NOTTON D.G., Diapriinae types at The Natural History Museum, London

T. tricarinata (Cameron, 1912)

T. unicolor (Ashmead, 1896)

T. vulgaris (Kieffer, 1912)

T. walkeri (Dalla Torre, 1890) comb. nov.

= T. apicalis (Walker, 1860) preocc.

Turripria Masner & Garcia, 2002

T. woldai Masner & Garcia, 2002

Zacranium Ashmead, 1901

Z oahuense Ashmead, 1901

Subfamily uncertain

Calogalesus Kieffer, 1912

C. parvulus Kieffer, 1912

Peckidium Masner & Garcia, 2002

P. enigmaticum Masner & Garcia, 2002

Discussion

The nature of this paper is such as not to require a lengthy closing discussion, except to say that the

location and recognition of types and their placement in currently valid genera takes the study of

Diapriinae further towards a modern understanding and is intended as a basis for future revisionary

studies by all. The discovery and analysis of types must continue until all are accounted for. The author

is grateful for notification of any lost or poorly known types which can be included in future catalogues.

Acknowledgments

Sincere thanks are due to Lubomir Masner (Canadian National Collection of Insects, Ottawa, Canada);

Theresa Howard, Andrew Polaszek, Goulven Keineg, Dez Mendoza, Sharon Touzel (The Natural History

Museum, London); Miguel Alonso Zarazaga (International Commission on Zoological Nomenclature);

an anonymous reviewer; Rafael Montilla (Instituto Nacional de Investigaciones Agricolas (INIA),

Trujillo, Venezuala); Jose Luis Garcia Rodriguez (Universidad Nacional Experimental Simon Rodriguez,

Maracay, Venezuala).

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Manuscript received: 13 February 2013

Manuscript accepted: 24 October 2013

Published on: 26 February 2014

Topic editor: Koen Martens

Desk editor: Charlotte Thionois

Printed versions of all papers are also deposited in the libraries of the institutes that are members of

the EJT consortium: Museum National d’Histoire Naturelle, Paris, France; National Botanic Garden

of Belgium, Meise, Belgium; Royal Museum for Central Africa, Tervuren, Belgium; Natural History

Museum, London, United Kingdom; Royal Belgian Institute of Natural Sciences, Brussels, Belgium;

Natural History Museum of Denmark, Copenhagen, Denmark.

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