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OF LONDON.
FOR THE YEAR
Gee ‘PAPERS READ IN
MAY anp JUNE. ~
‘PRINTED FOR THE SOCIETY,
; AT THEIR HOUSE IN HANOVER-SQUARE.
5 LONDON:
LIST OF CONTENTS.
PART III.—1878.
May 7, 1878.
Page
The Ra i, Report on the additions to the Society’s Menagerie in April 1878......++ 441
1. Note on the Stridulating Organ of Palinwrus vulgaris. By T. Jerrery PARKER, Assoc. !
R.S.M.° (Plate XXX.) 0... cece eee cece ee ee rete eee ce eeees o as ingeratetelaptetb in thas ss 442
2. Contributions to a Knowledge of the Hemipterous Fauna of St. Helena, and Specula-
tions on its Origin. By F. Bucaanan Wuirte, M.D., F.L.S. (Plate ea De reear 4449)
3. Further Remarks on Fuligula nationi. By P.L. Scuater. (Plate XXXII.)...-....+. A174 |
4, On a small pelea. of Lepidoptera from Jamaica. By Arrnur G. Bed E. L.S., an
FDS SC. ins vais che cclge stm oine ones omen sb visvete 2a cg resb ioe cule olole’ ely’ «ate eam eas
5, ee. 7 new Land-Shells from Japan and Borneo, By aoe A. Surru, F.Z.S. 495 —
6. On the Fruit-Pigeons of the Genus Ptilopus. By D. G. Exuior, F.R.S.H, &e. (Plates
OK MTT itis RON IV ci)” cel oinielancte: agri e-o'o, e/siejeie'ereteue oma eigtavs) 6\<l sToneiet ee femme eis ainie-# ate 500
May 21, 1878.
1. Description of a new Genus of Snakes of the Family Calamaride, from Southern India.
By Lieutenant-Colonel R. H. Beppome, O.M.Z.S. «10... cece cece ee ee eet ee tees 576 |
2. Reports on the Collections of Birds made during the Voyage of H.M.S. * Challenger.’ —
No. X. On the Birds of the Atlantic Islands and Kerguelen’s Land, and on the
Miscellaneous Collections. By P. L. Scuarur, M.A., F.R.S......... Fee tetcoie ¢ s0ke
3. On new and little-known Mantide. By J. Woop-Mason. (Plates XXXV, & XXXVI.) 580
4. On new Genera and Species of Geodephagous Coleoptera from Central America. By
HE W. Bares; WLS Ge; «005 las de Oak Gin ps te 2 ee 587
5. Description of a new Species of Tudicula. By Grorcz Frencu ae sj sea ‘
NS palate Soars coe ec ootian SARA NO Mies iricrgLcenpex e's ies sembaraisiaie is: LUM
6. Rosen to the Ornithology of the Philippines.—No. IX. On the Collection made |
Mr. A. H. Everett in the Island of Palawan. By Arrnur, Marquis of TwEEp- 4
; ee FR. S., President of the Society. (Plates XXXVII. & XXXVIIL.) ........ 611
7. On the Trachea of Tantalus loculator and of Vanellus cayennensis. By A. H. Garrop, .
ETA, PROBES. 2S 5 Sa cece aire eaknce a aeoray stata rela tenay cian inary ees oh Sta Se 625°
8. On the Anatomy of the Maleo (Megacephalon mateo). By A. H. Ginned, M.A. B. RS. 6
|
June 4, 1878. #
The Secretary. Report on the additions to the Society's Menagerie in May 1878, and re- BI
marks on Tapirus roulint, (Plate KXXIX.) .c...c cect ees cess ce ereteenscons 631 |
Mr. F. Holmwood. Remarks upon a young specimen of Temminck’s Manis (Manis tem- ‘
miMD Ki) a'ee Bese a agiesv vgheletes rarsake dae hes Lome eRe ete eee ee . 632
Mr. Sclater. Remarks upon the living examples of Ciconia maguari and Ciconia episcopus __
now in the Society’s Gardens ..... ila ota batees ae wee res dijeazs relate. 0s Sveimls SMR TRENT share 633
Mr. Edward R. Alston. Exhibition, on behalf of Dr. Elliott Coues, C.M.Z.S., of two spe- — ~:
cimens of Synaptomys cooper?, BO. .%. ries «jon wicie ole iti vis e's\ele wv aisle one oisieieisisivin 633
Prof. Ray Lankester, M.A., F.R.S. Notice of a memoir on the hearts of Ceratodus, Proto- _
pterus, and Citi 6S is cece 034 sh ow te ded cia eee SNolsle vie 634
1. On the Skull of a ners (R. lasiotis, Scl.?) from India. By WinitAm Henry _
° PTOWEER, ED, Sav Vicks Zishos oe sieeve wile, als\ons ise aie, stabaxatelofehe e¥etmcatel stare py 51s 10 canietete Maieetannvos . 634)
2. Catalogue of Lepidoptera collected by Mr. 8. N. Walter ; in the Island of Billiton.
Rhopalocera by F, D. Gopman aa Ossert Satyrn. Heterocera by H. Druce. x
ea Plate Kai toss ous ay asvds wae dia aah d Sis's, pies So eles Face bale Sea .» 63%
Contents continued on page 3 a bide a. |
Part.
1878.] THE SECRETARY ON ADDITIONS TO THE MENAGERIE. 441
May 7, 1878.
F. Du Cane Godman, Esq., V.P., in the Chair.
The Secretary read the following report on the additions to the
Society’s Menagerie during the month of April 1878 :—
The total number of registered additions to the Society’s Mena-
gerie during the month of April was 76, of which 33 were by
presentation, 26 by purchase, 3 by exchange, 8 by birth, and 6
were received on deposit. The total number of departures during
the same period, by death and removals, was 90.
The most noticeable additions during the month of April were
as follows :—
1. A Squirrel, from Ecuador (Seiwrus stramineus), of a species
new to the Society’s collection.
Mr. Alston, who has been paying special attention to the American
Sciuride, has kindly furnished me with the subjoined note on this
species.
“The Squirrel from Ecuador is certainly the same as Gray’s
Macroxus fraseri* ; but I am convinced that this is a synonym of
Sciurus stramineus, Eydoux et Souleyet®. S. nebouxii of Is.
Geoffroy * will doubtless prove the same; but I hope very shortly
to be able to examine the types of both species. Mr. J. A. Allen*
has lately referred three specimens from Ecuador to S. hypopyrrhus,
Wagl., principally on the strength of their harsh sparse fur and
long narrow tails. The last, however, seems to me to be a very
variable character; and one of these specimens, which Mr. Allen
kindly lent me, agrees very well with our bushy-tailed individual, as
well as with Gray’s types. The irregular white spots and markings
seem characteristic of this species, but are not constant, and, when
present, vary in every example. S. stramineus was described from
Omatope, and S. nebouwi from Payta, Peru, while M. frasert and
Mr. Allen’s specimens were from Ecuador.”
2. A male Beisa Antelope (Orya beisa), presented by H.M. the
Sultan of Zanzibar, and kindly brought home for the Society by
Capt. Pasley of H.M.S. ‘Simoom,’ April 27th.
Our excellent correspondent Dr. Kirk, H.B.M. Consul at Zanzibar,
to whose kind offices we owe this valuable animal, tells me that it
was obtained in the southern part of the Somali country. Along
with it the Sultan sent a specimen of a ‘‘ Dwarf Koodoo, quite
different from the common species.’’ This would probably be
Strepsiceros imberbis, Blyth, of which we have never yet received
an example.
It is much to be regretted that this animal died on the passage
home.
The following papers were read :—
1 Ann. & Mag. N. H., 3rd ser. vol. xx. p. 480 (1867).
2 Zool. Voy. la Bonite, vol. i. p. 37, atlas, Mamm. pl. ix. (1841).
3 Zool. Voy. la Vénus, p. 163, atlas, Mamm. pl. x1. (1855).
“ Mon. N. Am. Rodent. p. 746.
Proc. Zoou. Soc.—1878, No. XXIX. 29
442 MR. T. J. PARKER ON PALINURUS VULGARIS. [May 7,
1. Note on the Stridulating Organ of Palinurus vulgaris.
By T. Jurrery Parxer, Assoc. R.S.M.
[Received March 27, 1878.]
(Plate XXX.)
On the 5th of March I exhibited to the Society a structure,
evidently of the nature of a stridulating organ, which I had found
in the common Palinurus, and which I then believed had hitherto
escaped notice. On the following day, however, Professor Huxley
showed me a reference, which he bad that morning chanced to meet
with, to a paper by Dr. Karl Mobius, in the ‘ Archiv fiir Naturge-
schichte’ for 1867, the title of which, “‘ Ueber die Entstehung der
Tone welche Palinurus vulgaris mit den iusseren Fihlern hervor-
bringt,”’ showed at once that I had been anticipated. I hasten,
therefore, to render to Dr. Mébius such credit as belongs to the
discovery, and take the opportunity, at the same time, of making
some criticisms on his paper, as well as a few additional remarks
upon the organ in question’.
The main anatomical features of the stridulating organ are described
by Mobius, as they could hardly fail to be, with perfect correctness.
He fails, however, to notice either the guiding tubercle (Plate XXX.
figs. 3 and 4, ¢), situated just below the ridged pad (p) on the antenna,
or the groove for its reception on the antennulary sternum (fig. 1, g):
the forme, indeed, is shown in his figure; but no reference is made
to it. As I mentioned in my former communication, these structures
are of great importance, as by their means the apparatus is brought
into gear: when the tubercle does not fit into the groove the pad is
no longer in close apposition with the smooth surface of the anten-
nulary sternum, and the antenna moves noiselessly.
As to the functions .of the various parts of the apparatus, the
account given by Mobius is altogether at variance with my own.
He makes the observation that the lower surface of the flap (figs.
1-4, f) which plays over the lateral ridge (figs. 1 and 2,7") of the anten-
nulary sternum is beset with innumerable close-set minute hairs,
inclined with their points upwards, and that, corresponding to these,
fine scratches are to be seen on the surface of the ridge itself. ~All
this is perfectly correct: there is no doubt about the presence or
the direction of the hairs; and the scratches are quite visible with a
hand lens. But Mobius goes on to say that the scratches are pro-
duced by the hairs, and that it is the friction of the flap against the
smooth edge of the antennulary sternum which produces the sound,
by the upwardly directed hairs catching against the surface, when
the antenna is moved in the same direction. Of course when the
antenna moves downwards the hairs will lie flat and present no ob-
17 find that Leach, in his ‘Malacostraca Podophthalmata Britannix’
(1815), mentions the stridulating of Palinwrus, and correctly ascribes the sound
to the friction of the antenna against the ‘“‘clypeus” (antennulary sternum).
He gives, however, no description or figure of the apparatus.
Mintern Bros imp.
\J.P del. ©. Berjeau lith.
it | STRIDULATING ORGAN OF PALINURUS.
1878. | MR. T. J. PARKER ON PALINURUS VULGARIS. 443
struction ; and in this way Mobius attempts to account for the fact
that no sound accompanies a downward movement.
As to the first of these statements, one finds it rather hard to
believe that hairs so fine’ that the surface they cover has a texture
like that of satin, can produce visible scratches; their effect, one
would rather think, would be to impart a fine polish. I should be
inclined to suspect that the scratches must be due to fine particles
of sand getting in between the flap and the ridge.
With regard to the second statement, that it is the friction of the
flap, and not that of the pad, which produces the sound, I can only
say that I have removed the flap entirely without any sensible dimi-
nution of the noise. The mere observation of the parts while in
action is enough to show the true state of things: when looked at
from the front it is very evident that the flap exerts hardly any
pressure upon the ridge, as, indeed, from the fact that it is a soft
structure supported only along one edge, it could scarcely be expected
to; while the pad, on the other hand, is completely flattened out
against the smooth surface, and in the most perfect contact with it.
It is not easy to see how such a mistake can have arisen; but
Dr. Mobius’s language is sufficiently explicit to leave no doubt of
his meaning. He says:—‘‘ Das gefurchte Feld [the ridged pad]
bringt keinen Ton hervor, wenn es iiber die Fliche des Keiles
[the antennulary sternum] gleitet. Es leistet aber Widerstand gegen
die riickwarts gedringten Haare, da es dicker und steifer als das
behaarte Feld [the flap] ist. Ausserdem geben seine Riefen dem
Gange der ganzen Platte grissere Sicherheit, da sie diejenige Rich-
tung haben, in welcher die Bewegungen ausgefiihrt werden.”
The fact that a downward movement of the antenna gives rise to
no sound seems to me to be explained by the mode of attachment
of the pad. It is connected by its inferior edge (figs. 4 and 5, #2)
with the lower boundary (fig. 5, 4’) of the slit or uncalcified portion
of the basicerite, while at its superior edge it passes insensibly into
the inner wall of the soft flap. ‘The outer wall of the flap, again, is
connected with the upper edge (fig. 5, 6) of the slit in the basicerite.
Thus, while the pad is immovably fixed at its inferior edge, it is
capable of a considerable amount of displacement in the rest of its
extent ; and when pressed upon, by the downward movement of the
antenna, in the direction from y to a (fig. 4), it is foreed inwards
upon the yielding tissue which underlies it, and never comes into
sufficiently close contact with the opposing surface to produce a
sound (fig. 5, A). But when it is pressed upon, in consequence of
the upward movement of the antenna, in the direction from z to y,
the only effect of the pressure is to cause it to bulge out over the
line of attachment i, and so come into closer contact with the
opposing surface the stronger the force applied (fig. 5, B).
In the matter of histological structure, the pad does not differ
from other chitinous membranes, being formed of fine superposed
horizontal laminee, marked by a vertical striation (fig. 5). It is, how-
ever, of unusual thickness ; and its horizontal lamin have, for some
1 They are not more than one seven-hundredth of an inch in length.
29"
444 DR. F. B. WHITE ON THE [May 7
distance down, a wavy appearance, corresponding with the ridges
into which the surface is raised.
The stridulation is almost equally audible in water and air. As
to its production by the living animal, Dr. Mobius states that it was
heard by himself and Mr. Lloyd in the Hamburg Aquarium, and
that it was made especially when the Palinuri were handled; he
compares it to the sound produced by pressing the upper leather of
a boot against a table-leg. I also heard the sound, and observed the
movement of the antenna producing it, in a specimen brought alive
to the Biological Laboratory of the School of Mines shortly after
making my last communication.
As Dr. Mabius observes, the sound cannot be produced in spirit
specimens. These can, however, be again rendered vocal by soaking
them for a longer or shorter time in water.
EXPLANATION OF PLATE XXX.
Fig. 1. The parts concerned in the formation of the stridulating organ, seen
from the right side (nat. size): 0s, ophthalmic sternum; @s, anten-
nulary sternum; 7, one of the projecting ridges which bound it
anteriorly ; g, the groove on its lateral surface ; s, the smooth surface
between the groove and the ridge, against which the pad on the
antenna rubs; @7’, articular cavity for the right antennule ; a7, arti-
cular cavity for the right antenna ; 4, basicerite, and 7, ischiocerite of
the left antenna; /, the flap.
Fig. 2. The same, seen from above (nat. size). Letters as before.
Fig. 3. The basicerite and ischiocerite of the left antenna, seen from the inner
side (nat. size); the dotted line shows the extent of the uncalcified
portion of the basicerite: p, the ridged pad ; ¢, the guiding tubercle.
Other letters as before.
Fig. 4. The pad, with the guiding tubercle and part of the flap (x 4); the
arrow shows the direction taken by the hairs on the inner surface of
the flap: i7, inferior or attached edge of pad; xy, line showing
direction of movement. Other letters as before.
Fig. 5. Diagramatic cross section of the stridulating apparatus, showing the
condition of the ‘parts, (A) when the antenna is moving downwards,
(B) when it is moving upwards, The arrows show the direction of
movement. 4 upper and 4! lower lip of the uncalcified slit in the
antenna. Other letters as before.
Fig. 6. Microscopical section of the pad, taken across the ridges. Drawn under
a z-inch objective.
2. Contributions to a Knowledge of the Hemipterous Fauna
of St. Helena, and Speculations on its Origin. By
F. Bucnanan Waitt, M.D., F.L.S.
[Received April 8, 1878.]
(Plate XXXI.
Part I. SPECULATIVE.
«The extreme isolation of St. Helena—which is nearly 1200 miles
from the nearest point of the African continent, 1800 from that of
South America, and about 700 from even the small and barren island
XXXII
Py
siete.
PZ.5
Hanhart imp
arkiss kth.
HEMIPTERA OF ST.HELENA.
1878.] HEMIPTERA OF ST. HELENA. 445
of Ascension—gives it a degree of importance which it would not
otherwise possess ; for about the faunas of remote islands cluster, in
an especial manner, a variety of problems which, although they may
never be absolutely solved, may yet be brought, by a series of care-
fully conducted observations, within the sphere of discussion, and be
made to throw some additional light, however faint, on the general
questions of geographical zoology. From whatever point of view we
look at them,—and there are many which at once suggest themselves
along the distinct, but ultimately converging, lines of thought,—the
statistics of an oceanic rock, far removed from the ordinary effects of
immigration and change, and bearing more or less of the impress
which was stamped upon it by its aboriginal forms of life, have an
interest about them which it is scarcely possible to overrate. Tow
the organisms, as we now see them, came to occupy their present
areas of distribution,—to what extent they are, or seem to be, ‘re-
lated’ to those of the nearest mainland,—whether there is evidence
for believing that they have changed to any considerable extent, in
their outward configuration, from the types of which they may be
presumed by some naturalists to be the remote descendants,—or
whether there is reason to suspect that the Hand, which originally
placed them where they are, adapted each separate species to the con-
ditions which it was destined to fulfil, subjecting one and all of them
to a law of permanence under which they can never very materially
alter,—are but a tithe of the questions which, if not capable of being
answered positively, we may at least ventilate and probe, not altogether
without profit, in even a small treatise like the present one ; for it
cannot be too carefully borne in mind that, within the limited sphere
where mere speculation (as such) seems likely to have any permanent
value, it is to facts, and not to theories, that we must ultimately
appeal’’’.
The sentences which I have quoted form, I think it will be admitted,
an appropriate introduction to a paper descriptive of one of the last
collections of insects ever made by their eminent and much regretted
writer; but, before proceeding to the special subject of this memoir,
I wish to devote a few words to a consideration of the problems sug-
gested by Mr. Wollaston, not, however, with much hope that I shall
be able to throw any fresh light on a matter that has puzzled many
abler naturalists.
Of what is actually and satisfactorily known regarding the indi-
genous animals and plants of St. Helena, the following is a brief
epitome :—
There are no terrestrial Mammalia, nor any land or freshwater
Amphibia, Reptilia, or Fishes.
Of birds there are at least eight indigenous sea species and one
land bird (the Aygialites sancte-helene, Harting, very closely allied
to the African 4. varius, Vieill.), which is peculiar to the island.
As regards the sea-fishes, Dr. Giinther has, in the ‘ Proceedings’ of
this Society?, given an account of the collections made by Mr. Melliss
1 T. V. Wollaston, Coleoptera Sanctz-Helenzx, pp. vil & viii,
2 March 1868 and April 1869.
446 DR. F. B. WHITE ON THE [May 7,
These included 65 species, of which 17 are peculiar to St. Helena,
31 others, being inhabitants of shallow water, are probably indigenous
(their exotic distribution being for the most part on the American
and African coasts), and nearly 20 are deep-water species and
common in the tropical seas. Much, however, still remains to be done,
not only amongst the fishes of St. Helena, but in the other groups of
marine animals.
The land mollusks seem to have been more looked after than the
marine species. Of the former, 4 (or, including 2 unnamed slugs’, 6)
living, and 20 dead and subfossil native species have been recorded.
Amongst the Insecta, the order Coleoptera has received most
attention ; and, owing to Mr. Wollaston’s researches, the indigenous
species of that order are at least fairly, though, perhaps, not entirely,
known. Of the 203 species that have been found in the island, 57
have without any doubt been accidentally introduced ; 17 others may
have been introduced; and the remaining 129 Mr. Wollaston con-
sidered to be the descendants of the ‘‘ autochthones of the soil,”’ and,
with one exception, absolutely peculiar to St. Helena.
Of the Hemiptera a small, but probably fairly representative col-
lection was made by Mr. Wollaston. In it I find 30 determinable
species; and 2 or 3 more have been recorded from the island. Of
the total number 5 have certainly, and 1 has, probably, been intro-
duced, leaving 26, which, with one exception, seem to be peculiar.
The only other group of animals of which there is (as far as I am
aware) any satisfactory record is that of the Spiders, Mr. Melliss’s
collections’ of which have been reported on by the Rev. O. P. Cam-
bridge. Out of a total of 40, 18 species seem to be indigenous, and
22 probably introduced.
We will now glance at the flora. The great mass of the present
flora consists of introduced species; but of the original flora, as
far as it can now be traced*, 40 flowering plants are absolutely
peculiar, and the 5 other probable natives are species of wide-spread
tropical distribution. Of the 26 ferns, 16 are peculiar. Adding to
. these the other cryptogamic plants, so far as they are known, we have
a total of 77 plants* supposed to be absolutely peculiar to the island.
‘* Whence and by what means came this very peculiar fauna and
flora?’’ is a question that has given rise to more than one suggestion
or theory.
To begin with the flora, Sir J. D. Hooker®, after alluding to the
causes that have assisted in destroying the ancient flora, and indi-
cating the means by which it is possible to distinguish the aboriginal
from the introduced plants, proceeds to say :—‘‘ From such frag-
mentary data it is difficult to form any exact conclusions as to the
affinities of this flora; but I think it may be safely regarded as an
1 J. C. Melliss, ‘St. Helena, p. 118.
2 Proce. Zool. Soc. Nov. 1869 and March 1873.
3 Sir J. D. Hooker, “‘ Lecture upon Insular Floras,” Brit. Assoc. Ady. of Sci.
i866.
4 Melliss, /. c. p. 224.
* «Lecture on Insular Floras,’ quoted in Melliss, /. ec. p. 225, &e
1878. ] HEMIPTERA OF ST. HELENA. 447
African one, and characteristic of Southern extra-tropical Africa.
The genera Phylica, Pelargonium, Mesembryanthemum, Osteosper-
mum, and Wahlenbergia are eminently characteristic of Southern
extra-tropical Africa ; and I find amongst the others scarcely any
indication of an American parentage, except a plant referred to Phy-
salis'. The Ferns tell the same tale: of 26 species, 10 are abso-
lutely peculiar ; all the rest are African, though some are also Indian
and American. The botany of St. Helena is thus most interesting ;
it resembles none other in the peculiarity of its indigenous vegetation,
in the great rarity of the plants of other countries, or in the number
of species that have actually disappeared within the memory of living
man .... Probably 100 St.-Helena plants have thus disappeared
from the Systema Nature since the first introduction of goats on the
island. Every one of these was a link in the chain of created beings,
which contained within itself evidence of the affinities of other species,
both living and extinct, but which evidence is now irrecoverably
lost. If such be the fate of organisms that lived in our day, what
folly it is to found theories on the assumed perfection of a geological
record which has witnessed revolutions in the vegetation of the globe
to which that of the flora of St. Helena is as nothing!”
Mr. Melliss, whose interesting work on St. Helena I have had fre-
quent occasion to refer to, says": —‘“‘ Other theories may be appealed
to in order to account for the presence and position of this wonderfully
curious little flora. Continental land at one time spreading over the
South Atlantic Ocean, with its own peculiar flora and fauna, has been
started as a plausible theory; but the geological investigation of St.
Helena forbids us to look upon it as a remaining portion of some dis-
appearing continent to which the last vestige of a flora, still
struggling for existence, may be clinging; and the great depth of
ocean® around it also seems to deny the possibility of its connexion
at any time with cither African or American land. Still we cannot
tell what geological changes, hundreds or even thousands of centuries
may have witnessed in that portion of the globe, leaving, perhaps,
this unique little floral remnant, now fast disappearing, as almost the
only record of what once was. So far, therefore, the manner in
which this once incandescent mass first received its flora, whether by
the agency of birds or atmospheric and oceanic currents, or direct
from that Hand by which all things were created, still remains un-
fathomed.”
Of the class Arachnida, Mr. Melliss‘ states: —‘‘ Mr. Cambridge says,
in reference to the character of this portion of the island fauna, after
his final examination of the several collections, that ‘the European
stamp observed upon in regard to the spiders of the former collection
is thus equally marked in those now recorded and described.’ It is
worthy of note that the native Spiders are, almost as a rule, least
1 Subsequently referred to a new genus, Mellissia, Hk. f.
2 Pp, 225.
3 « St, Helena is said to be separated from the continents of Africa and America
by a depth nowhere less than 12,000 feet.”
4 Dc. p. 206.
448 DR. F, B. WHITE ON THE [May 7;
abundant now in the island ; in each case where I met with only one
specimen, it turned out to be a new species. It is therefore not at
all improbable that, like the native plants and the Snails, which we
know are fast disappearing, some having gone entirely, the Spiders,
for some cause or other, are also yielding up their native land to
foreign invaders.”
Mr. Wollaston, whose painstaking investigations of the faunas of
the North Atlantic islands, and careful study of the Coleoptera of St.
Helena especially qualified him to give an opinion on this subject,
thus writes regarding the Beetles! :—‘ The whole of the 129 species
to which I have just alluded are, with a single exception (the Chilo-
menes lunata, Fab.), absolutely peculiar to St. Helena; so that the
question of geographical distribution would seem to be well-nigh
‘nipped in the bud.’ Moreover, from all that I know of the South-
African Coleoptera . . . . . it has almost nothing in common with
these 129 aboriginal St.-Helenians, which stand out singly, as it
were, and alone, related more or less inter se, but unrelated for the
most part, to any recognized continental forms. It is true that two
of the most significant of the Rhynchophorous types—namely,
Nesiotes (of the Tanyrhynchide) and Acarodes (of the Anthribiide)
—are allied conspicuously to Echinosoma and Xenorchestes of the
Madeiran archipelago; but if any more successful generalizer than
myself can develop much from these points of quasi-contact, he is
quite welcome to the result. So far as I can understand the evidence
before me, any unprejudiced inquiry into the ‘origin’ (as usually
understood by that term) of these St.-Helenian Coleoptera, does not
elicit, in reply, so much as even an echo; for not only are they en-
demic (in the strictest sense of the word), but an overwhelming
majority of them are attached (or were so originally) to trees and
shrubs which would seem to exist nowhere in the world except on
this remote rock, 1200 miles from the nearest point of the African
coast, surrounded by an all but unfathomable ocean, and which has
every appearance of having been piled up by successive erruptions into
a basaltic mass at no period very considerably larger than that which
we now see. ‘Whence then, came its fauna and flora,’ are enigmas
which I cannot presume to answer on any known principles of deriva-
tion and descent. To a mind which, like my own, can accept the
doctrine of creative acts as not necessarily ‘unphilosophical,’ the
mysteries, however great, become at least conceivable; but those
which are not able to do this may perhaps succeed in elaborating
some special theory of their own, which, even if it does not satisfy
all the requirements of the problem, may at least prove convincing
to themselves. The St.-Helena fauna cannot, I think, be said to
have had much light yet thrown upon it as regards its actual ‘ origin’
(except, perhaps, in so far as my individual opinions on the subject
may be accepted by others who are predisposed to receive them) ;
but its primitive (or at all events remote) stage is another matter, and
appears to be capable of some real elucidation from the facts to
which we have access.”
* Coleoptera Sancte-Helene, p. xix.
1878. ] HEMIPTERA OF ST, HELENA. 449
Regarding the Mollusca, the late Edward Forbes thought that
both the terrestrial and marine species dimly indicated a closer geo-
graphical relationship between the African and American continents
than now maintains. ‘The marine mollusks would seem to point to
the submergence of a tract of land probably linking Africa and South
America before the elevation of St. Helena. Along the sea-coast of
such a tract of land, the creatures common to the West Indies and
Senegal might have been diffused.”” Commenting on this sugges-
tion, Mr. Gwyn Jeffreys’ writes, “I am not quite satisfied with this
hypothesis, and I believe that more information is needed to support
batt Thy) seo A few of the marine shells are Mediterranean, while
the greater number are well-known inhabitants of the Indian Ocean
and the West Indies; all these may have originated anywhere. But
it must be borne in mind that St. Helena is separated from Africa
and South America in every direction and by very deep water, which
is nowhere less than 2000 fathoms or 12,000 feet. It therefore
seems scarcely probable that such an abyssal and extensive tract of
the sea-bed could have been dry land or ‘sea-coast’ in a geologically
recent period, so as thus to account for the diffusion of littoral species
such as Mytilus edulis, M. crenatus, and Littorina striata. 1
should be rather inclined to attribute the present distribution of the
marine fauna of St. Helena (not to a supposed continuity of land be-
tween Africa and South America in that or any other direction, but)
to the action and influence of the great Agulhas current, which
issues from the Indian Ocean and flows round the Cape of Good
Hope northwards towards St. Helena, and thence past Ascension to
the West Indies.”
Mr. Andrew Murray, whose loss to science we have had so recently
to deplore, has given? a considerable amount of attention to the
origin of the fauna of St. Helena. After stating his conviction that
there at one time existed a land communication, now at the bottom
of the Atlantic, between the northern and southern hemispheres, and
arguing that there are only three great Coleopterous faunz or
stirpes (the Indo-African, the Brazilian, and what he calls, for want
of a better name, ‘‘the microtypal stirps’’), he proceeds to say :—
“St. Helena, that great puzzle of naturalists, is a crucial test to my
hypothesis of a communication between the northern and southern
hemispheres by an Atlantic continent; if that link snaps, the whole
chain will fall to the ground... . . . I say that its fauna is cer-
tainly microtypal, and, if so, almost necessarily a branch of the
Atlantic type of that stirps; there is nothing else microtypal for it
to be attached to. Some three years ago Dr. Hooker gave an admi-
rable Lecture on Oceanic Islands, in which he discussed the origin
of the flora of St. Helena, and on the whole seemed inclined to refer
it to Africa. More in the spirit of ‘audi alteram partem’ than from
any settled conviction of my own, I wrote a reply, in which I gave
some reasons for thinking that it might more probably have been
1 Ann. & Mag. Nat. Hist. April 1872; Melliss, 7. c. p. 114.
2 ‘On the Geographical Relations of the Chief Coleopterous Faun,” Journ.
Linn. Soc. x1. no, 49.
450 DR. F. B, WHITE ON THE [May 7,
originally connected with and peopled from Europe, although, also,
possibly connected at some period with Africa. More mature con-.
sideration and subsequent researches have confirmed my opinion.”
Mr. Murray then goes on to point out how the fauna (as then known)
supported his conclusions as to its European origin. Of the flora he
writes :—‘* The general result which I draw from the whole flora is,
that we have here a compound flora, certainly two deep, possibly
three deep. We have, in the first place, I believe, a genuine natural
Atlantic (that is, European) flora; for in the face of the decided
testimony given by the fauna, I cannot accept Dr. Roxburgh’s con-
clusions as to the supposed introduction and naturalization of every
species having a European habitat. If they can be proved to have
been introduced, good and well; but I object to take the thing to be
proved as part of the proof. And in the next place, I believe, we
have the traces of an older African flora (why I call it older I will
explain when I come to speak of an ancient connexion between
Patagonia and South Africa); and I believe that both are due to
actual continuity, however circuitous or interrupted, with the respective
countries the impress of whose floras they bear.. . . . . | .
“Before leaving St. Helena, I have just one other argument to
adduce in support of its former connexion with the other Atlantic
Islands; and that is, the fact (which has only recently been ascertained,
or, at any rate, only recently laid down in our maps) that there is a
long band of elevated submarine bottom running north from St.
Helena to the Cape-Verd Islands, and embracing in its course
Ascension Island and the shoal-ground on the equator.”
Subsequently Mr. Murray explains how, in his opinion, there
might have been a land connexion between Patagonia and the Cape
of Good Hope, “ sending out perhaps arms to Tristan d’Acunha and
St. Helena, which, however, have the option of what, I think, is a
still more ancient union in another direction,” namely from Brazil,
rounding, but not touching, the Cape of Good Hope to Madagascar.
To one or other of these connexions (presumably the first) are owing
‘“‘the score or so of plants left on St. Helena and Tristan d’Acunha,
and a slight sprinkling of microtypal forms which still subsist at the
Cape.”
T have now given the gist of what I have been able to find sug-
gested as to the origin of the flora and fauna of St. Helena, and, be-
fore explaining my own views, will briefly recapitulate.
The facts may be thus stated :—
1, A volcanic island of very ancient origin, very remote from con-
tinental land or other islands, and surrounded by an ocean 12,000
feet in depth on almost every side.
2. A terrestrial fauna and flora, the very great majority of the in-
digenous species comprising which are peculiar, and a marine fauna
with a large proportion of peculiar species.
The various theories suggested for the peopling of this island
arrange themselves as follows :—
1. Special creation and no special affinities.
2. Land connexion with both Africa and America.
1878. ] HEMIPTERA OF ST. HELENA. 451
3. As regards the fauna and part of the flora—land connexion from
the north, and European affinities.
As regards the peculiar part of the flora—land connexion from the
south uniting Africa and America.
4. As regards the marine Mollusca, diffusion by means of the
Agulhas current from the south.
5. Affinities of the flora somewhat suggestive of extratropical
Southern Africa.
Against the first of these theories (special creation) it may be
argued that such a method is contrary to, as far as we can judge, the
whole scheme of creation, and that, while many things can be urged
against it, there is nothing to support such a belief.
Against the theories which necessitate the existence in time past
of a land communication with Africa or America, or both, all the
known facts militate. The non-existence in St. Helena of Amphibia
and terrestrial mammals’, the large number of peculiar species, as
well as the great depth of the surrounding sea, afford reasons for
thinking that it is extremely improbable that the island was ever
connected with either of these continents.
The other theories will be alluded to in giving my own.
A careful consideration of all the known facts leads me to believe
that two, if not three, epochs in the colonization of the island
may be distinguished, and that the great bulk of the inhabitants
came from the north, but not by means of a continuity of land.
The first of these epochs, or that in which the earliest inhabitants
of St. Helena reached the island, occurred possibly early in the last
glacial period, but perhaps and (even probably) in Miocene times ;
and the reasons for thinking so, as well as for believing that the
colonists came from the north, I will now try to prove.
In the first place, all the peculiar species belong either to peculiar
genera, or to genera which, if they have not yet been found in
Miocene formations, are (like the genera which have been found) so
widely distributed that it seems more than probable that they were
Miocene genera. The nearest relations of the peculiar genera are
also of this character.
To begin with the terrestial fauna, viz. the Mollusca, the Coleo-
ptera, the Hemiptera, and the Arachnida.
The molluscan genera are Succinea (with a peculiar subgenus),
Helix, Bulimus, and Achatina—genera which all date from the
Eocene, and which are all very widely distributed.
The Coleoptera, being more numerous, are not so easily disposed
of. Taking Mr. Wollaston’s estimate of what are aboriginal species,
we have 40 genera, of which 25 are peculiar. The remaining 15
genera include 10 actually known as Miocene, 4 of wide distribution
and therefore probably Miocene, and 1 which is only known from
Madeira in addition to St. Helena. Of the 25 peculiar genera, 21
at least belong to families or subfamilies which are Miocene, and 3
are allied to very widely-spread genera.
The Hemiptera are not many in number, nor is our information
1 ©. Darwin, ‘ Origin of Species,’ 6th edition, p. 350.
452 DR. F. B. WHITE ON THE [May 7,
about their geographical distribution so complete as it is for the
Coleoptera. The total number of genera is 21, of which 8 are pecu-
liar. Of the 13 genera not peculiar, 4 have Miocene affinities, 3 are
very widely spread, and 6 are more or less widely distributed, but
there is a want of information regarding them. Of the 8 peculiar
genera, 4 have Miocene affinities, and 3 have relations of very wide
distribution, the remaining genus having African affinities.
As the Hemiptera are specially treated of in the second part of
this paper, I need not discuss them at length just now.
The supposed indigenous Arachnida belong to 13 genera, none of
which are peculiar. All the genera are widely distributed—many
very widely, reaching Australia and New Zealand. One genus is
Miocene, and 5 belong to families known as Miocene.
As already mentioned, there are no freshwater fishes; and even if
the sea-fishes were more extensively known than they are, the conti-
nuity of the medium in which they live affords so many facilities for
the distribution of the species, that it is not very easy to learn much
from them. Still we find that both the geological age and the pre-
sent distribution of the 7 families to which the 17 peculiar fishes be-
long are very great, and that, on the whole, they seem to indicate the
same Palearctic origin as the other classes.
The flora alone remains to be considered. The extratropical
South-African affinities of the flowering plants have been pointed
out by Sir J.D. Hooker. As to the cryptogams, many have a very
wide distribution, either as species or genera; and while many, from
their geological age, have had ample time for diffusion, it would seem
that even at the present day cryptogams have greater facilities for dis-
persion than phanerogamic plants. In addition, more information as
to their geographical distribution is necessary before they can be
considered to throw very much light on the subject we are con-
sidering.
The facts connected with the aboriginal fauna to which I have
just called attention seem to indicate that its probable origin was
somewhat after this manner :-—
1. That the first settlers arrived at a very early date, and that
no very great additions, from outside the island, were made to them
from that period up to the time when man’s agency came into
force.
2. That they did not arrive in a body, but that the colonization
of the island was spread over a considerable period.
3. That the road by which they came was not on a continuous
land-surface.
4, That the colonists came from the Palearctic region.
I shall now endeavour to prove these statements seriatim.
1. The first settlers arrived at a very early date.
As, thanks to Mr. Wollaston, the Coleoptera form the best-known
group of the animals of the island, we will see how they support
my argument. If the colonization did not take place at a very early
1878.] ; HEMIPTERA OF ST, HELENA. 453
period, there ought to be some genera represented (amongst those
which are not absolutely peculiar) which are of comparatively recent
origin ; but the facts are that out of 15 genera not peculiar, 10 are
actually Miocene’, and 4 others are of such wide distribution that,
though they have not yet been detected in Miocene formations, they
must be of Miocene age. The remaining genus may have either
been a later importation, or, what is as likely, originated en route.
The 25 peculiar genera are all, as I have stated, allied to Miocene,
or probably Miocene, families ; and their great number is an addi-
tional argument in favour of the long isolation and consequent early
settlement of St. Helena.
The Hemiptera (so far as our defective knowledge of their geolo-
gical and geographical distribution goes) tell the same tale. Of the
13 genera not peculiar, 4 have at least actual, and 3 probable,
Miocene affinities; of the remaining 6 we know less. Of the 8
peculiar genera, 4 have actual and 3 probable Miocene affinities.
Again, we learn the same thing from the Arachnida. The 13
genera are all probably Miocene, 1 being and 5 belonging to families
known as Miocene.
The 4 genera of terrestrial Mollusca are all Eocene ; and one has a
peculiar subgenus. These quite support the evidence learnt from
the insects. ;
2. The aboriginal fauna did not arrive all at once, but the coloni-
zation was spread over a lengthened period.
This is a question into which I need not enter at length, but
merely refer to Mr. Darwin’s remarks on the inhabitants of oceanic
islands?, and point out how the great number of peculiar species in
St. Helena bears out his arguments. If all the inhabitants had
arrived simultaneously, their mutual relationships would have been
undisturbed, and the liability to modification consequently less-
ened by the absence of new forms of competition. Spread over a
lengthened period, the colonization of a new land must inevitably
upset the preexisting relationships of the colonists, and result in, as
we see in St. Helena, many modifications of form.
3. The road by which the colonists travelled was not on a continuous
land-surface.
This, too, is a question not requiring lengthy consideration here.
The absence of Batrachians and terrestrial mammals is quite in
accordance with what Mr. Darwin has shown is the rule in oceanic
islands; and, moreover, the argument in favour of gradual coloniza-
tion also supports that of want of continuity of land-surface.
4. The colonists came from the Palearctic Region.
This is capable of consideration under two headings :—1, the Pala-
1 There seems reason to believe that many genera of insects known as Miocene
are also of Hocene age.
> L.e, p. 348.
454 DR. F. B. WHITE ON THE [May 7,
arctic character of the fauna; and, 2, the road by which it came ;
but these will be most conveniently treated of together.
St. Helena is, as has been already said, a small island of ancient
voleanic origin, very distant from any continent, and surrounded by
very deep sea. The nearest land is Ascension Island, 698 miles
north-west of St. Helena; and nearly double that distance north of
Ascension lie the Cape-Verd Islands. Improbable as it may
seem to many, this is the route by which, I think, the aboriginal
fauna reached St. Helena.
But it is not necessary to believe that the colonists crossed such
wide stretches of ocean. ‘“‘Stepping-stones,’”’ in the shape of other
islands, doubtless existed ; and, in fact, we can trace the remains of
some of them. For, deep as is the ocean round St. Helena, a some-
what shallower sea (still, however, very deep) extends between it
and Ascension, and is continued north to the equator, where a yet
shallower place is to be found, not more than 700 or 800 miles from
the Cape-Verd Islands. It seems not only possible but extremely
probable that in this equatorial shoal we have the wreck of an island ;
and as the line indicated coincides with an area of depression as well
as of voleanic action (extending more or less from Iceland to Trinidad
and Tristan d’Acunha), there is no reason why other islands should
not have existed, of which (as Mr. Darwin says) “ not a wreck now
remains.”
It may be argued against this theory that, if it were probable, then
Ascension should have a larger native fauna than it at present pos-
sesses; but, as far as I can learn, the present Ascension is of com-
paratively recent origin, and supplies an argument in favour of my
view that volcanic action has made great alterations in the sup-
posed line of route. If (as seems probable) the present Ascension
Is comparatively recent, it by no means follows that an older island
did not once occupy its place and possess a larger fauna than its suc-
cessor, which has been possibly colonized from St. Helena. (I may
here call attention to Mr. Melliss’s remarks’, that Sterna fuliginosa
does not remain all the year at St. Helena, but probably migrates to
Ascension, returning to St. Helena at the end of the year. If there is
any thing in Mr. Wallace’s theory that the lines of migration of birds
often coincide with ancient land-connexion’?—which, however, Colonel
Drummond Hay has shown* to be by no means invariably the case—
then this would seem to point at some closer connexion in former
times between St. Helena and Ascension.)
But even assuming that the supposed requisite intermediate islands
existed, it is necessary to find some means by which the tide of emi-
gration was induced to flow towards St. Helena.
At the present time both the ocean-currents and the prevailing
winds would tend to lessen, if not, as seems more probable, to
entirely prevent, emigration in that direction, even if the “ stepping-
stone ” islands still existed. The S.E. trade is the prevailing wind ;
and the great Agulhas current sweeps past St. Helena from the
LE ep Oe: ° Geographical Distribution of Animals, i. p. 22.
* Scottish Naturalist. 1877-78.
1878.] HEMIPTERA OF ST. HELENA. 455
Cape of Good Hope towards the West Indies, and, while effectually
stopping any emigration from the north, does, in fact, occasionally
(as we shall presently see) bring new inhabitants to the island from
the south. Given, therefore, any cause by which the present direc-
tion of the winds and currents could be reversed, the probability of
colonization from the north would be immensely increased. But
when we find, as in reality we do, that one and the same cause not
only reversed the winds and currents, but induced an immense tide
of migration to flow from the north to the south, then it seems
pretty evident whence St. Helena derived its fauna, if that fauna be,
as I shall presently show it is, of a Palearctic character.
The cause I have alluded to is one or other of the northern glacial
periods, either the last or one in Miocene times. That during such
a period the equatorial currents were reversed, Dr. Croll! has shown
there are very good reasons for believing; and as regards the flow
of migration southwards at the same time, we know that that must
necessarily have taken place’.
But perhaps it may be argued that similar reasons to those which
I have advanced to show that the colonization was from the north
may be brought forward to prove, with as great probability, that the
colonization was from the south. We have seen, however, that the
present configuration of the sea-bottom is much more in favour of
the northern than of the southern theory; we, know, moreover, that
emigration from the north has always been more powerful than from
the south ; and I will try to show that the character of the fauna is
in favour of the northern view.
It is now generally admitted that the Cape-Verd Islands belong
to the Palzearctic region, and have derived their fauna through
the Canaries and Madeira. If, then, St. Helena got its fauna
by that route from the north, the affinities should be Palearctic,
and the imprint of the fauna of the route should be more or less
discernible. That this imprint is not more manifest is explained,
I think, by reason that the route was interrupted at the St.-Helena
end, and that island cut off, at a very remote date; the other islands
being less inaccessible, have had their faunas altered very consider-
ably by later importations, and their original settlers less modified
by the fact that the colonization was more en masse and that fresh
blood has tended to preserve the old types.
Mr. Wollaston, whose opinion must be received with the greatest
respect, arrived at the conclusion, as I have already mentioned, that
St. Helena had little in common with the faunas of the Atlantic
archipelagos to the north ; but even he admits that “‘ two of the most
significant of the Rhynchophorous types—namely Nesiotes (of the
Tanyrhynchide) and Acarodes (of the Anthribiide)—are allied con-
spicuously to Echinosoma and Xenorchestes of the Madeiran archi-
pelago.”
Most of the coleopterous genera not peculiar to St. Helena have
already been cited in evidence of the very ancient peopling of the
1 Quoted in Dr. J. Geikie’s ‘ Great Ice Age,’ p. 107, &e.
* Darwin, I. c. p. 339, &e.
456 DR, F. B. WHITE ON THE [May 7,
island. Most of these genera, in addition to being Miocene and of
very wide distribution, are characteristic Paleearctic genera at the
present time. Amongst the peculiar genera many have their
affinities (and, hence, probable ancestry) with Palearctic genera ;
while in the North-Atlantic islands there seems to me to be more
evidence of alliance with St. Helena than Mr. Wollaston was inclined
to admit. Mr. Murray (whose line of migration, in part, at least,
coincides, as will have been noticed, with mine—he arguing, however,
for continuity of land) was decidedly in favour of affinity with the
North-Atlantic islands; and his knowledge of Coleoptera was, it
must be allowed, so extensive as to require his opinion to be received
with attention. It is true that since he wrote on the subject our
knowledge of the St.-Helenian Coleoptera has been immensely
increased ; but so far as I can see, nothing has been found to invali-
date (but rather to increase) the argument in favour of such alliance.
Taking Mr. Murray’s abstract’ of the Coleoptera of the North-
Atlantic islands, we have in Madeira 266 endemic species against
120 aboriginal European species, the endemic species being all akin
to European forms. Then in the Canaries we find that, out of a
total of 930, 224 species are identical with Madeiran, the peculiar
characters of the Madeiran fauna being there in force. Next come
the Cape-Verd Islands, of which Mr. Wollaston says, ‘‘ Our recent
explorations in the Cape-Verdes have shown their coleopterous
population to be so far more than I had anticipated on the Canarian
and Madeiran type, that I am any thing but certain that it would not
be more natural to regard the whole of these Atlantic islands as
characterized by a single fauna unmistakably the same, even whilst
necessarily differing as to many of its exact details (and through the
fact of mere distance) in the more widely separated groups.”
In fact we find in the three archipelagos just what might have
been expected. As we move southwards the same general character
of the fauna is found to be present, but the particulars. gradually
alter. And this, it seems to me, is apparent even when St. Helena
is reached. Making due allowance for its remoteness and different
latitude, the character of the fauna is the same, though the details
are very considerably altered. For example, in Madeira we have
the Heteromerous genera Hadrus and Hegeter, with 3 and | species
respectively ; in the Canaries Hadrus has vanished, but Hegeter has
no less than 19 species; in the Cape-Verds Hegeter has almost
disappeared, having but a single species, but its place has been taken
by a new genus, Oxycara, with 10 species ; in St. Helena all these
genera have vanished, but are represented by two new and allied
genera—Hadrodes and Tarphiophasis, regarding the first of which
Mr. Wollaston remarks that it has a good deal in common with the
Madeiran Hadrus. Tarphiophasis too seems evidently a develop-
ment of Hadrodes, just as the latter is of Hadrus. Then, again, Mr.
Wollaston remarks of the St.-Helenian Opatrum hadroides that it
is closely allied to species from the Cape-Verd, Canarian, and
Madeiran archipelagos, and is even more akin to one, and probably
1 Lc. p. 12 &e,
1878.] HEMIPTERA OF ST. HELENA. 457
identical with another, Cape-of-Good-Hope species—the significance
of which latter facts will be alluded to in connexion with the flora.
Again, the genus Huwestus was known from Madeira only, till Mr.
Wollaston discovered another species in St. Helena.
But as the most remarkable fact in the coleopterous fauna of St.
Helena is the enormous preponderance of Cossontde—more than a
quarter of the endemic species belonging to that family—which
seem to find there their metropolis, we ought, if the line of migra-
tion is by Madeira &c., to find in the latter a significant development
of this family. And so wedo. ‘In the Madeiran and Canarian
groups,” writes Mr. Wollaston’, ‘‘there is scarcely any fact [the
insular-loving nature of the Cossonide] more distinctly observable—
where every detached rock is tenanted by some one representative,
or more, of this particular department. Nor are trees and shrubs
(which seldom flourish in localities thus weather-beaten and exposed)
by any means essential for their support, the pithy stems of the
ordinary plants being amply sufficient to sustain them; and I have
frequently found the stalks of dead thistles and Umbelliferce to be
perforated through and through by their ravages.” Mr. Wollaston
found 19 species in Madeira and 14 in the Canaries. The Anthri-
btide (which include the next largest number of endemic species in
St. Helena) indicate a like derivation ; but enough has, I think, been
brought forward to show both the Palzearctic origin and probable
route of migration of the Coleoptera of St. Helena.
The Hemiptera are not fitted (from the reasons already given) to
teach us so much as the Coleoptera; but 12 at least, if not all, of the
13 non-peculiar genera are Paleearctic, and many of them Madeiran.
The peculiar genera have also nearly all strong affinities with Pale-
arctic genera. Just as the whole facies of the Madeiran Hemiptera
is European, so that of the St.-Helenian is Madeiran and European.
The characteristics of the Arachnida and of the terrestrial Mol-
lusca have already been pointed out.
But let us now see if there be any thing in the manner of life of
the aboriginal animals of St. Helena which would make their passage
across the sea not only a possible but a probable occurrence. If we
can find that a majority of them are connected with plants, it is not
difficult to imagine how they might have been drifted by sea-
currents to the island; but if, in addition, it turned out that many
inhabited the interior parts of plants, their carriage across the sea
would pass from the region of possibility into that of probability.
Mr. Wollaston has carefully recorded the modus vivendi of the St.-
Helenian Coleoptera; so we will try and prove our case from his
evidence.
At least half of the 12 endemic species of the genus Bembidium
have the very abnormal habit, for that genus, of living within the
dead and rotting stems of the tree ferns. (I may note here that in
the Madeiran group 10, in the Canaries 14 (7 peculiar), and in the
Cape-Verds 5 species of the genus have been found.) The
following St.-Helenian genera are also, amongst others, especially
1 Trans. Entom. Soc, London, 1873, iv. p. 433.
Proc. Zoou. Soc.—1878, No. XXX. 30
458 DR. F. B. WHITE ON THE [May 7,
wood-frequenting—Anchastus, Anobium, the 17 genera of Cos-
sonide, Nesiotes, &c.
Thus 16 out of the 25 peculiar genera, and nearly 80, at. least,
out of the 129 peculiar species, live zx wood at some period of their
existence. The Anthribide, of which 26 species occur in St.
Helena, are not wood-borers, although probably lignivorous as larve,
but cling rather tenaciously to foliage, dead wood, &c. ; and they,
as well as most of the remaining Coleoptera as well as the species of
the other groups which are also attached to plants, must be specially
liable to transportation in or about drift-wood, &c. In the case of
winged species the winds, of course, would assist in the work.
But it is unnecessary to pursue this subject any further, save to
mention that though most of the plant-frequenting beetles are
attached to some one or other of the peculiar plants, it by no means
follows that their ancestors were so restricted ; for, as observed by
Mr. Wollaston, some at the present day devour with apparently
equal relish, the native arborescent Composite and the introduced
Coniferee. . Nor is it necessary to suppose that they were introduced
with the tree Compositze, because it is probable that the progenitors
of the latter were not arborescent when they arrived in the island, but
that that condition was gradually evolved’. Some of the Cossonide
are quite content with the pithy stems of thistles &c., though the
family is essentially wood-loving.
It is also worth while noting the extreme paucity of that section
of the Coleoptera known as the Phytophaga, which seems to show
not only that the lignivorous beetles had more facilities of transport
than those that merely fed upon the leaves of plants, but that, as
Mr. Wollaston remarks, the early flora of St. Helena was essen-
tially a woody one. Now that the forest has vanished, and though
the greater part of the island is suited for the Phytophaga, yet the
number of species remains the same—showing, I think, that coloni-
zation (apart from that brought about by man’s unintentional
agency) is not now going on.
A word now as to the flora. The aboriginal plants have, as has
been already said, most affinity with the flora of Southern extra-
tropical Africa. This affinity can surely only arise from a common
origin ; and if, as I have attempted to prove, the origin of the fauna
is Palzearctic, it seems reasonable to suppose that the origin of the
flora is the same, and that the same agencies which brought its
fauna to St. Helena brought its flora also. Without going into
details of the South-African flora (for which, indeed, I have not the
materials), I may mention that there are one or two genera of plants
common to it and to St. Helena which are strongly suggestive of a
Palzearctic origin and dispersion by the influence of a glacial epoch :—
for example, Stwm, which has an endemic representative in St.
Helena; the very characteristic Cape genus Pelargonium, which has
a straggler in Syria (where, be it noted, the endemic St.-Helenian
coleopterous genus Haplothorax has, according to Lacordaire, its
nearest allies) ; and others.
* Darwin, 7. c. p, 350.
1878. ] HEMIPTERA OF ST. HELENA. 459
Here [ may be permitted to quote a remark by Mr. Darwin
on this point ':—* In Africa, several forms characteristic of Europe
and some few representatives of the flora of the Cape of Good Hope
occur on the mountains of Abyssinia. At the Cape of Good
Hope a very few European species, believed not to have been
introduced by man, and on the mountains several representative
European forms are found which have not been discovered in the
intertropical parts of Africa. Dr. Hooker has also lately shown
that several of the plants living on the upper parts of the lofty
island of Fernando Po and on the neighbouring Cameroon Mountains,
in the Gulf of Guinea, are closely related to those on the mountains of
Abyssinia, and likewise to those of Temperate Europe. It now also
appears, as I hear from Dr. Hooker, that some of these same
temperate plants have been discovered by the Rev. R. T. Lowe on
the mountains of the Cape-Verde Islands. This extension of the
same temperate forms, almost under the equator, across the whole
continent of Africa and to the mountains of the Cape-Verde
archipelago, is one of the most astonishing facts ever recorded in the
distribution of plants”*. Mr. Darwin then proceeds to show how
in a glacial epoch the temperate flora might have invaded the whole
of Africa, and at the return of warmer conditions been driven up the
mountains, or in some cases become gradually acclimatized.
In connexion with this possible, and, as it seems to me, probable,
community of origin of the floras of the Cape (in part) and of St.
Helena, the significance of the occurrence at the former place of
several Coleoptera and Hemiptera closely allied to, if not identical
with, St.-Helenian species, is not to be overlooked’.
Itis not to be wondered at, then, that we do not find, in the parts
of Africa nearest St. Helena, much, if any, relationship to the island
fauna and flora. That such relationship, so far as community of
origin is concerned, once existed, I have little doubt ; but the return
to tropical conditions and the reestablishment of the tropical fauna
and flora have obliterated, except on the mountain-summits, all
traces. And St. Helena by its isolated position and temperate
climate (the mean temperature of the year being only about 61°) is
to all purposes a mountain.
The affinity of the Hemipterous genus Megarhaphis to the
African Macrorhaphis (of which one species is from the Cape, and
the other—rather doubtful as to the genus—is from the Congo)
seems to be an exception; but as we do not know the exact nature of
the locality whence the Congo species was derived, and as it is as
likely as not to be a mountain and not a tropical insect, it may after all
prove our case by being a descendant of one and the same Palearctic
ancestor as the Cape and St.-Helenian species.
There still remain some elements in the fauna and flora of St.
Helena to be accounted for.
1 Origin of Species, p. 337.
2 See also Professor Oliver's ‘Flora of Tropical Africa,’ in which the
occurrence of several species, not only Huropean but even Arctic, is recorded.
3 Oertain European Hemiptera are also natives of the Cape.
30*
“
460 DR. F. B. WHITE ON THE [May 7,
For example, there is the one peculiar land-bird, Ayialites
sancte-helene, which is so closely allied to African species that it
is, in all probability, a comparatively recent introduction. It seems
rather remarkable that there are not more peculiar land-birds ; and
the absence of Bats (none of which are recorded by Mr. Melliss) is
another curious feature in the fauna of the island.
There are also the species of West-Indian fish and marine Mol-
lusca as well as the Indian-Ocean species of the same groups. Not
very much can be made out of our present knowledge of these; but
it seems to me that no great obstacle exists to their introduction by
means of the ocean currents. The present currents might have
brought the southern and eastern species, while the reversal of the
currents during a glacial period would bring the West-Indian and
Brazilian animals. Though I have already invoked the agency of a
reversed current for the purpose of bringing the Palearctic fauna
to the island, I think it is improbable that the arrival of the West-
Indian fish at St. Helena was contemporaneous with that of the
ancestors of the endemic fauna, though, in the case of some of them,
it is quite possible. I would rather suppose that, since they are
species of presumably later origin (or, should their origin be as
remote as that of the endemic species, yet it was elsewhere than at
St. Helena), they were brought to the island during a later (or a
later part of the last) glacial period than the ancestors of the endemic
species.
There are also certain elements in the flora, viz. the five tropical
weeds or sea-side plants of very wide distribution, which Sir. J.
Hooker refers to. These are almost certainly of later introduction
than the endemic and peculiar flora. Mr. Melliss remarks’ of
Entada scandens, L., that the large seeds ‘‘ are cast ashore on the
windward side of the island, having been brought from the Mau-
ritius, or elsewhere, by sea-currents. They have been known to
germinate ; but it is doubtful if one now exists there.” ‘This fact
is suggestive of the origin of the five plants alluded to, as well as of
some others thought by Mr. Melliss to be indigenous.
Part II. Drescriprive.
In his recent work upon St. Helena, Mr. Melliss was able to
record the occurrence of only 8 species of Hemiptera. The names
of only five species had apparently been determined ; and of these 4
were evident introductions.
It was therefore left to Mr. Wollaston to add considerably to our
knowledge of the St.-Helenian species of this order. The list which
I now submit to the Society embraces the results of his six months’
exploration of the island; and though it probably does not include
all the native species, yet there is reason to suppose that the collection
is a very representative one.
A few months before his lamented death, Mr. Wollaston placed
the collection in my hands for determination, with directions that a
1E.¢. ip, 266;
- 1878.] HEMIPTERA OF ST. HELENA. 461
‘set of types” should be placed in the British Museum, and that
the new species should be described in a single paper and not piece-
meal. The first of these requests I have already carried out ; and
the second I now proceed to fulfil. The type specimens of ald the
new species, as well as a larger number of specimens in cases where
the species was represented by more than two individuals, will be
found in the British Museum.
The following is a list of the species in the collection :—-
Order HEMIPTERA.
Suborder Herrrorrera.
Fam. Cimicina, Stil.
Subfam. Asoprna, Stal.
1. Megarhaphis wollastonit, Buchanan White.
Subfam. Penratomina, Stil,
, Nezara viridula, L.
bo
Fam. LyGz1p, Stal.
Subfam. Berytrna, Stal.
3. Metacanthus concolor, Buchanan White.
Subfam. Lycarna, Stil.
Div. Orsituartia, Stal.
4. Nysius sancte-helene, Buchanan White.
5. Nysius thymi, Wolff, var.
Fam. Cimicip, Stal.
Subfam. ANTHOcoRINA, Reut.
Div. ANrHocorARIA, Reut.
. Hapa contorta, Buchanan White.
. Cardiastethus bicolor, Buchanan White.
. Lyctocoris campestris, ¥.
9 Sis or)
Div. Micropuysartia, Reut.
9. Myrmedobia fuliginea, Buchanan White.
Subfam. Capsina, Reut.
Div. Puyrocoraria, Reut.
10. Calocoris (Megacelum) lustratus, Buchanan White.
Div. Cyitocorarta, Reut.
11. Orthotylus mutabilis, Buchanan White.
Div. Capsarra, Rent.
12. Lygus apicalis, Meyer.
462 DR. F. B. WHITE ON THE [ May 7,
Diy. PuacioGnaTHaRiA, Reut.
13. Agrametra ethiops, Buchanan White.
14. Psallus flavosparsus, Buchanan White.
15. Psallus lutosus, Buchanan White.
16. Psallus vinaceus, Buchanan White.
Fam. Nasip4, Stal.
Subfam. Napina (=Corisc1na, Stai).
17. Nabis capsiformis, Germ.
18. Vernonia wollastoniana, Buchanan White.
Fam. SaLprp&.
19. Salda aberrans, Buchanan White.
Suborder Homorprera.
Fam. FuueGorina.
Subfam. Issinz.
20. Issus coleoptratus, F.
Subfam. DeLpuacint.
21. Liburnia (Ilburnia) ignobilis, Buchanan White.
Fam. JASSINA.
Subfam. ByrHoscoprnt.
22. Stonasla undulata, Buchanan White.
23. Stonasla consors, Buchanan White.
24, Nehela vulturina, Buchanan White.
Subfam. ACocEPHALINI.
25. Argaterma alticola, Buchanan White.
26. Argaterma multisignata, Buchanan White.
Subfam. JAssrn1.
27. Grypotes (2) insularis, Buchanau White.
28. Thamnotetti« sancta-helene, Buchanan White.
29. Jassus wollastoni, Buchanan White.
Subfam. TyPHLocysint.
30. Chlorita edithe, Buchanan White.
There were besides in the collection several species in immature
condition and hence undeterminable. One of these species, which
is a native of the high central ridge, appears to belong to the Coreini ;
and any future explorer of the island would do well to search for it.
To the list must also be added Acanthia lectularia, L., a doubtless
introduced species recorded by Mr. Melliss, and Cirius sanctea-
helene, Stil (Eugenies Resa, Entomol. Bidr. 272. 148), a probably
endemic species. Mr. Melliss also mentions the occurrence of several
1878. } HEMIPTERA OF ST. HELENA. 463
Aphides and Cocci. Of the 30 determinable species in Mr. Wol-
laston’s collection 5 species—
Nezara viridula, L., Nabis capsiformis, Germ.,
Lyctocoris campestris, F., and
Lygus apicalis, Meyer, Issus coleoptratus, F.,
have been probably introduced since the island was discovered.
The species which I have referred to Nysius thymi, Wolff, may also
be an introduction ; but I am more inclined to consider it a native.
The remaining 24 species seem to be all peculiar to the island ;
and for the reception of 10 of them I have been obliged to create 7 new
genera and 1 new subgenus. Mr. Wollaston has not, unfortunately,
recorded for the Hemiptera, as he has done for the Coleoptera, the
special modus vivendi of each species; but as the particular part of
the island inhabited by each was briefly noted by him, I have been
careful to give all the information [ can.
i MEGARHAPHIS, gen. nov.
Genus Macrorhaphidi valde affine. Caput apice subtruncatum,
lobis equilongis. Rostrum articulo 2° 4°que subequilongis, sin-
gulatim 3° longioribus. Thorax marginibus lateralibus subcal-
losis, vix tranversim rugosus. Scutellum parte apicali retrorsum
angustata, frenis ultra medium scutelli_ continuatis. Carina
mesosternalis mediocris, nec lata nec sulcata. Venter spina
basali longa compressa armatus, apud marem maculis sericeis
haud instructus. Femora antica subtus apicem versus spina
- parva armata. Tibie antice vie dilatate. Membrana costis 8
instructa.
There is no doubt that Megarhaphis is closely allied to Dallas’s
genus Macrorhaphis ; but it seems to me that the characters pointed
out above are of importance sufficient to necessitate the creation of
a new genus for the species described below. The difference in the
comparative proportions of the joints of the rostrum, the narrow and
unfurrowed keel of the mesosternum, the absence of the two silky
spots on the venter of the male, and other points will at once dis-
tinguish Megarhaphis from Macrorhaphis.
1. MEGARHAPHIS WOLLASTONI, Sp. 0.
M. elongata, rufo-brunnea, rude et dense nigro-punctata ; thoracis
angulis lateralibus prominulis obtusiusculis ; pedibus corpore
concoloribus ; antennis paullo pallidioribus ; capitis lineis 2 lon-
gitudinalibus ocellos includentibus, marginibus tyli, vitta centrali
capitis subtus sternique, maculis marginalibus dorsi abdominis,
nebulisque ventris nigris ; spina basali ventrali ad apicem testacea ;
membrana brunneo-fusca. Capitis lineis 2 longitudinalibus inter
ocellos ad apicem currentibus, margine interiore orbite, tyloque,
pronoti callis irregularibus prope marginem anticum, necnon
carina longitudinali angusta, scutelli linea apicali longitudinali
subelevata, corii spatiis irregularibus (una ad medium, altera prope
apicem), haud vel vir punctatis ; sterno ventreque minus dense
464 DR. F. B. WHITE ON THE [May 7,
punctatis, carina ventrali levigata; scutello fovea parva sed
profunda utrinque ad basin prope pronoti angulos posticos sat
productos instructo.
3. Long. 13, lat. 6 m. m.
Hab. “West Lodge’”’ (exemplar unicum).
M. wollastoni agrees in many points of sculpture &e. with the
description of Macrorhaphis (?) acuta, Dall. (List, i. 88. 2), from
Congo, which species may perhaps pertain to the genus Mega-
rhaphis.
2. NEZARA VIRIDULA, L.
Two examples without note of locality. This almost worldwide
species is probably an introduction.
3. METACANTHUS CONCOLOR, §p. 0.
M. ochraceus ; antennarum articulo 1° annulis, uno prope basin,
altero sat lato ad apicem, articulo ultimo (apice excepto), rostro
ad apicem, oculis, scutello (basi spinaque exceptis), femorum
tibiarumque apicibus tarsisque necnon corti marginibus angus-
lissime exteriore et interiore apicali plus minus piceis aut piceo-
nigris; capitis vertice linea elevata longitudinali instructo ;
pronoti lobo antico rude punctato, sulco sat profundo inter lobos
callis 3 irregularibus instructo, lobo posteriore postice valde
elevato et longitudinaliter carinato; mesosterni spina lateralé
longissima, primum recta, deinde retro curvata.
3. Long. 5 m. m.
Hab. ad “The Barn,” ubi Aster glutinosus (Roxb.) (anglice
““ Scrubwood”’) oceurrit.
I have seen only one specimen.
4, Nysrus SANCTH-HELENA, Sp. 0.
N. oblongus, ochraceus, setulis brevibus concoloribus vestitus ; an-
tennis, rostro, capite, pronoti impressione lineari transversa
anteriore, scutelli basi sternoque plus minus rufo-brunneis ; an-
tennis articulo 4° basi excepto, rostro ad apicem, capite vitta
utringue inter oculos, punctis pronoti maculisque utringue ante
marginem posticum, scutelli punctis, costis et maculis clavo disco-
quecorii, hujus margine apicali plus minus interrupte, membrana
maculis irregularibus, femoribus maculis, apice tarsorum, sterno
ad medium, necnon maculis ventris nigris vel fusco-nigris.
Bucculis basin capitis vie attingentibus, pone medium retrorsum
sensim humilioribus ; articulo 1° rostri pone bucculas extenso et
basin capitis atlingente ; capite cum oculis quam apex pronoti
latiore; antennarum articulo 2° quam 3° longiore ; pronoto et
scutello (illius basi excavata excepta) carina levissima instructis ;
mesosterno distincte sulcato ; margine costali corii antice recto,
dein rotundato-ampliato.
3 9. Long. 53-63, lat. 2-23 m. m.
Hab. in intermediis editioribusque insulee—“ Plantation,’’ “ West
Lodge,” “ Luffkins,”’ et ad “ Diana’s Peak.”
1878. ] HEMIPTERA OF ST. HELENA. 465
Nysius sancte-helene varies considerably both in the number and
intensity of the darker markings. Sometimes the whole of the
clavus and disk of the corium are fuscous with pale spots, and the
apical margin entirely infuscated ; but the widened anterior margin
of the corium is almost always immaculate, except that sometimes
there is a dark streak near the apical margin.
This species belongs to the section aa of the genus Nysius in
Stil’s ‘ Enumeratio ;’ but it does not appear to be very closely allied
to any of the species therein placed.
5. Nysrus Toymi, Wolff.
Hab. in intermediis editioribusque insulee—“ Plantation,”’ “ West
Lodge,” et ad ‘‘ Diana’s Peak.”
I cannot find any points of distinction between British and St.-
Helena specimens of this common and variable European species,
except that the latter seem to be rather smaller and to have more
slender antenne and legs.
It does not appear to have been previously found out of Europe.
Hapa, gen. nov.
Corpus oblongo-ovatum, parce hirsutum. Caput sulca transversa cur-
vata inter oculos anterius preditum, inter oculos latum, modice pro-
ductum, collari mediocri instructum, pone oculos leve nitidumque.
Antenne pilose, articulo 1° apicem capitis superante, 2° sursum
leviter incrassato, 3° 4°que filiformibus. Rostrum medium
mesosterni attingens, articulo 1° oculos attingente. Pronotum
haud vel obsoletissimo collari instructum, transversum, apice
truncato, basi levissime concava, angulis anticis rotundatis et
maxime depressis, lateribus acutis ; discus anterius convexo trans-
verse rugosus, posterius depressus et leviter concavus ante angulos
posticos, ad medium sulco longitudinali plus minus obsoleto
instructus. Scutellum anguste transversum, ad basin convexum
concavum ad marginem utrinque pone basin, apice leviter acumi-
nato, disco leviter concavo et leviter longitudinaliter rugosc.
Elytra macropterorum apicem abdominis paullo superantia et
membrana obsolete 3-venosa. Elytra sepius tantum dimidio ab-
dominis equilonga, membrana haud instructa. Femora antica
maris reliquis crassiora; tibie antice apicem versus distincte
incrassate, margine inferiore serie brevissimarum spinularum
instructo ; tibie postice spinis nonnullis setiformibus instructe.
Pedes femine graciliores, tibiis anticis haud incrassatis nec ut
in 3 formatis. Quartum et sequentia segmenta abdominis maris
sinistrorsum asymmetrica, supra 4°, 5° 6°que abdominalibus et 1°
genitali segmentis sinistrorsum maxime abbreviatis; segmento 1°
genitali fere contecto ad marginem sinistrum ; segmento genitali
ultimo (ut videtur) triangulari, levitur sinistrorsum contorto,
latere sinistro foramine calloso-marginato (quod fere @quilon-
gum est ac segmentum), instructo. Quarti segmenti abdominis
subtus margo posticus ad latus sinistrum profunde sinuatum.
Abdomen femine symmetricum. Apex abdominis 3 et 2 pilis
nonnullis longis apicem versus instructus.
466 DR. F. B. WHITE ON THE [May 7,
Genus Piezostetho affine.
Nomen a Maori “ hapa” (contortus).
Obs. In Piezostethus, to which the genus is allied, the males
have also unsymmetrical hind bodies—a fact, by the way, that I
do not see noticed in any of my books.
6. Hapa contorTa, sp.n. (Plate XXXI. fig. 1.)
H. piceo-nigra, pilis brunneis vestita ; capitis apice, antennis,
pronoti margine postico, elytris, pedibus abdomineque plus minus
piceo-brunneis ; rostro, antennarum articulis duobus ultimis,
cuneo ad basin, trochanteribus, femoribus ad basin, tibiis ad
basin apicemque et tarsis plus minus dilutioribus; abdomine
obscuriore variegato. Macropterorum clavi disco, corti margine
antico basin versus, et macula ad medium basale membrane paullo
dilutioribus ; membrana fusco-luteo-brunnea, venis obscurioribus.
3 Q. Long. 3-33, lat. 15 m. m.
Hab. in excelsioribus insule, ad “ Diana’s Peak.’’
I have seen but a single macropterous specimen, and that rather
mutilated.
7. CARDIASTETHUS BICOLOR, Sp. 0.
C. niger, capillis pallidissime brunneis vestitus; antennis, rostri
apice, elytris pedibusque luteo-brunneis; capite nitido rufo-
brunneo, clypeo dilutiore; rostro (apice excepto), antennarum
articulis 2 ultimis, elytrorum suturis, margine interiore lato
embolii, margine interiore discoque cunei plus minus fusoo-
brunneis ; membrana luteo-fusca. Capite levi; clypeo sat
angusto; pronoto (callo excepto), scutello elytrisque subtiliter
punctatis.
Long. circa 23 m. m.
Hab. in intermediis insulee—“ Plantation.”
Rather closely allied to C. testaceus; but, besides differing in
colour, C. bicolor is a narrower and more slender insect.
9. MyrMEDOBIA FULIGINEA, Sp. n.
M. atra; antennis piceis, articulo 1° dilutiore ad basin, articulo
2° ad basin medium apicemque albido, articulo ultimo ad apicem
rufo-piceo ; pedibus piceo-nigris, femoribus posticis ad apicem,
annulis 2 ad medium tibiarum anticarum intermediarumque, et
dimidio basali tarsorum omnium albidis ; capite convexo ; antennis
subtiliter pubescentibus, articulis 2° 3°que subeaquilongis ; pro-
noto maxime convexo, annulo male definito, sulco transverso,
nullo ; elytris margine antico anguste reflexo.
Q. Long. 1 m. m.
Hab. locos editiores supra ‘‘ West Lodge.”
10. Catocorts (MrGacuLuM) LUSTRATUS, Sp. n.
C. sordide stramineo-testaceus, maculis parvis fuscis rufisque remote
et obscure notatus, capillis tenuibus adpressis pallido-aureis
vestitus ; pronoto obsoletissime punctulato ; oculis, maculis parvis
1878.] HEMIPTERA OF ST. HELENA. 467
in disco et ante marginem posticum pronoti, macula geminata ad
basin scutelli, dimidioque apicali femorum posticorum plus minus
brunneo- vel rufo-fuscis ; apicibus rostri scutellique fusco-nigris ;
antennarum articulo 3° ad apicem et articulo 4° toto, apice
imo tibiarum anticarum, apicibusque tarsorum omnium pallide
Susco-brunneis ; capite, antennarum articulo 1°, dimidio apicali
Semorum anticorum intermediorumque, tibiis omnibus ad basin,
et interdum elytris et lateribus abdominis supra subtusque, sat
conferte rufo-sparsis.
3 2. Long. 6, lat. 2 m. m.
Hab. in intermediis insule—* Plantation.”
Sometimes the apical half of the antenne is vinous-red in colour
and the fuscous markings generally more suffused with red.
11. OxTHOTYLUS MUTABILIS, sp. n.
O. pallide fusco-brunneus, capillis pallidis vestitus ; pronoti margine
postico elytrorumque margine exteriore anguste fusco-nigris ;
elytris lutescenti- vel viridescenti-fuscis ; antennis pedibusque
rufo-fuscis, illarum articulis 1° ad basin, 2° ad apicem, 3° 4°que,
necnon tarsorum articulo ultimo et interdum tibiis ad basin,
obscurioribus ; rostro ochraceo, apice fusco-nigro; membrana
pallide fusca venis dilutioribus. Sat angustus, elongatus ; an-
tennis + corporis equilongis, articulo 3° plus dimidio 2' longiore,
4° = 3' subequilongo ; rostro coxas posticas haud superante ;
pronoti lateribus vix sinuatis, callis parvis; scutello convexo,
ante medium transverse depresso.
3 Q. Long. 4m. m.
Hab. editioribus insulae—“ West Lodge’’ et ‘‘ Diana’s Peak.”
Very variable in the intensity of the coloration. It is allied,
but not very closely, to O. nassatus, and belongs to the same group
of the genus.
12. Lyeus apicauis, Mey., Fieb.
Hab. in intermediis insule—* Plantation.”
A South-European species, probably introduced into St. Helena.
AGRAMETRA, gen. nov.
Corpus oblongo-ovatum, sat rude punctatum, nudum. Caput nutans,
latum, convexum; clypeo modice producto, compresso, a fronte
parum discreto ; vertice immarginato ; gula brevi. Oculi oblongi,
leviter granulati, pronoto contigui. Antenne sat crasse, ad
oculorum medium internum inserte, corporis dimidio subequi-
longe, articulo 1° crasso apicem clypei longe superante. Rostrum
coxas posticas vix attingens, articulo 1° capitis basin vir superante.
Pronotum trapeziforme, convexum, latitudine apicali vir longius,
basi quam apice 4 latius, apice, lateribus et basi ante scutellum
leviter sinuatis. Scutellum ad basin detectum. Hemelytra com-
pleta, corii margine exteriore ad basin subito dilatato-explanato ;
clavo, corio cuneoque coriaceis ; membrana biareolata. Ale hamo
instructe. Pedes sat breves ; femoribus sat incrassatis ; tibiis
468 DR. F. B. WHITE ON THE [May 7,
modice fusco-spinulosis sed haud punctatis; tarsis posticis
longioribus, articulo ultimo duobus primis simul sumptis paullo
breviore.
Generibus Plagiognatho et Sthenaro subaffinis.
13. AGRAMETRA £THIOPS, sp.n. (Plate XXXI. fig. 2.)
A. piceo-nigra nitida; rostro (apice excepto), coxis, femoribus,
tarsorumque articulo ultimo piceo-brunneis ; antennis, tibiis, tar-
sorumque articulis duobus primis sordide ochraceis ; antennarum
articulo 1° ima basi, articulo 2° ad apicem, 3° 4°que piceo-nigris,
imis apicibus 2' 4'gue dilutioribus; membrana fusca, venis
parve cellule exterioris rufis; antennis, pedibus abdomineque
capillis tenuibus pallide fuscis vestitis ; antennarum articulo 1°
paullo supra basin subito incrassato, valido, cylindrico ; 2° sursum
sensim incrassato, quam pronoti basis breviore, 3° 4°que fusi-
formibus, 3° 4 2" paullo longiore, 4° 2 3' subequilongo.
é @. Long. 35 m. m.
Hab. in regione Commidendri robusti DC. (anglice “ Gumwood’’)
—‘ Luffkins”’ et “ Thompson’s Wood.”
14. PsALLUS FLAVOSPARSUS, Sp. 0.
P. testaceus, capillis pallidissime aureis vestitus, capite thoraceque
preterea pilis nigris validioribus instructis ; capite, pronoto
scutelloque ad basin fusco-atris indistincte nigro-guitulosis et
maculis flavidis (una ad marginem interiorem oculi, una utrinque
pone marginem anticam et altera in disco pronoti, necnon una
distinctiore in angulis anticis scutelli) ornatis ; angulis imis posti-
cis pronoti lestaceis ; scutello ad apicem clavoque flavo-guttulosis,
corio cuneoque flavo- et fusco-sparsis, macula orbiculart prope
apicem clavi et macula irregulari oblonga ante mediam basin
cunei fusco-nigris ; membrana dilute fusca, iridescente, basi et
fascia angusta transversa pone albidas venas dilutioribus ; an-
tennis testaceis, annulo prope basin et macula ad interiorem
articuli 1‘, annulis basali apicalique 2', 3° 4°que fusco-nigris ;
femoribus anticis et intermediis ad apicem, margine apicali antico
femorum posticorum, necnon tibiis omnibus e punctis nigris nigro-
spinulosis ; articulo ultimo tarsorum fusco ; antennis mutilatis,
sed, ut videtur, articulis 3° 4°que simul sumptis saltem 2°
equilongis.
2. Long. 3 m. m. .
Hab. in regione Commidendri robusti DC. (anglice ‘“‘ Gumwood ’’)
-—‘* Thompson’s Wood.”
15. PsaLLus LUTOSUS, sp. n.
P. Sat robustus, testaceus, opacus, aureo-pubescens, rufescenti-
brunneo dense irroratus ; signaturis obscuris plus minus confluen-
tibus, presertim in parte antica pronoti, medio apiceque scutelli,
apicibus clavi coriique necnon femoribus ; antennis rufo-brunneis,
articulo 1° ad apicem testaceo, articulo 2° ad basin apicemque
et articulis duobus ultimis fusco-brunneis ; tibiis testaceis, annulis
1878.] HEMIPTERA OF ST. HELENA. 469
plurimis angustis rufescenti-brunneis notatis et e punctis nigris
nigro-spinulosis ; membrana fusca pallide transverso-fasciata,
venis albidis. Antennarum articulis 2 ultimis simul sumptis 2°
brevioribus, 4° 3 3' subequilongo.
Long. 4, lat. 2 m. m.
Hab. in regione Commidendri robusti DC., ad ‘‘ Peak Gut.’’
16. PsaLLus VINACEUS, sp. n.
P. gracilis, obscure vinaceus, nitidus, capillis pallidis sublongis
vestitus ; antennarum articulo 1° et 2° ad apicem, rostro, scutello
ad apicem, clavi disco, guttulis in disco corti, sutura inter corium
cuneumque, cuneo ad apicem, femoribus ad apicem, tibiis tarsisque
plus minus testaceis vel vinaceo-testaceis ; clavi parte pallida,
Semoribus posticis ad apicem et tibiis posticis plus minus vinaceo
maculatis ; apicibus rostri tursorumque necnon membrana fuscis,
hac basi et fascia transversa pone cellulas dilutioribus, venis albidis,
venis cellule exterioris interdum rufis; tibiis e punctis nigris
nigro-spinulosis. Antennarum articulis 2 ultimis simul sumptis
2° aquilongis, 4° quam dimidium 3+ breviore.
3 2. Long. 3-33, lat. 12 m. m.
Hab. in editioribus insule, ad ‘‘ Diana’s Peak.’’
Very like a small dark form of Psallus lutosus, but distinguished
by its slenderer and more shining form, longer and less golden
pubescence, more slender legs and antenne, the more uniform
darker markings, and more convex pronotum with less concave
hind margin.
17. Napis cAapsirormis, Germ.
Hab. in intermediis insule, ad “ Plantation.’’
An African and American species, possibly (probably?) introduced.
VERNONIA, gen. nov.
Corpus elongatum. Caput ante uculos oblongo-productum, parte
anteoculari parti postoculari subequali; clypeo distinctissime
convexo-elevato, angusto, jugis depresso-rotundatis, medium versus
constrictis, ad apicem leviter incrassatis. Oculi sat magni.
Ocelli desunt. Rostrum 4-articulatum, coxas posticas attingens,
gracile, articulo 1° brevi, duplo longiore quam latiore, 2° 3°que
longis, subequalibus. Antenne setacee, gracillime, corpore 3
longiores, articulo 1° quam caput 1 longiore, 2° quam primus j
longiore, 3° 2° equilongo, 4° brevissimo. Pronoti lobus anticus lobo
postico angustior ; lobus posticus lateribus subparallelis obtuse et
callose marginatis instructus, ante marginem posticum truncatum
leviter transverse sulcatus. Scutellum latius quam longius, ad basin
transverse modice elevatum. lytra abbreviata, scutello tantum
duplo longiora, ad apicem rotundata, coriacea, marginata.
Acetabula antica postice aperta, a basi prostethii excisa ; meso-
sternum carina, pone medium distinctissima, instructum ; orificia
subobscura ante coxas posticas sita. Pedes gracillimi et longis-
simi ; cove longiuscule; tibie antice seriebus duabus, tibie
470
DR. F. B. WHITE ON THE [May 7,
intermedie serie unica, dentium nigrorum tenuissimorum instructe ;
femora antica modice incrassata, deorsum sensim angustata,
tibiis paullo longiora ; tibie omnes ad apicem leviter incrassate ;
pedes postici longissimi ; femora postica ad apicem modice in-
crassata ; tarsi omnes sat longi, articulo 1° brevissimo, duobus
ultimis subequilongis. Abdomen elongato-ovale, connexivo dilatato
ad segmenta 3°" 4°™que latissimo, illinc apicem basinque versus
sensim angustato.
Genus Arbele affine, memorize viri celeberrimi beati Thomas
Vernon Wollaston dicatum.
18. VERNONIA WOLLASTONIANA, sp.n. (Plate XXXI. fig. 3.)
V
. rufo-brunnea ( 3 ) vel rufo-lutea (2) rufo-sanguineo variegata ;
capitis parte anteoculari, antennis pedibusque pilis brevibus
pallide brunneis vestitis ; antennarum articulo 1° rufescente
annulis latis obscurioribus, ad apicem basinque distinctioribus,
ornato ; articulo 2° flavescente annulis 5 vel 6 fuscis, quarum 3
in dimidio basali et ad apicem distinctiores sunt, instructo, 3°
4°que fusco-brunneis obscure annulatis; pedibus plus minus
distincte fusco vel rufo fusco-annulatis, ad basin testaceis ;
abdomine supra sanguineo-variegata, subtus testacea vitta angusta
centrali et vitta latiore laterali utrinque ornata; abdominis
segmento 6° ad medium supra subtusque sinuato.
3 Segmento genitali quadrato, ad apicem truncato.
3 2. ‘Long. 10-12, lat. pronoti 13-17; abdominis ¢ 23, ?
4 m.m.
Habitat in excelsioribus centralibus insule, inter Compositas
arborescentes.
19. SALDA ABERRANS, sp.n. (Plate XXXI. fig. 4.)
8. obovata, atra, opaca, pilis brevibus erectis nigris, aliis depressis
aureis intermiztis, vestita; antennarum articulo basilari (apice
excepto), clypeo nitido, rostro, callo utrinque ad medium interne
oculorum, pronoti marginibus lateralibus anguste, marginibus
exterioribus interioribusque (illo basin versus latiore dilutioreque)
elytrorum, necnon maculis nonnullis irregularibus corti pedumque
brunneis vel pallide piceo-brunneis ; clavi maculis triangularibus,
una pone medium, duabus prope apicem, linea ante medium suture
clavalis, corii maculis 5 propemarginem anticum et nonnullisin disco
albidis ; tibiis ad apicem anguste pallidis, apice imo et articulo
ultimo tarsorum piceo-nigris ; unguiculis piceo-brunneis. Late
obovata, pone medium latior ; capite cum oculis quam pronoti apex
latiore ; vertice ante ocellus foveolato, margine antico subobtuse
inerassato ; antennis vix« incrassatis ; rostro longo, coxas posticas
aitingente ; pronoti margine postico quam margo anticus duplo
latiore, marginibus lateralibus anguste reflexis ; callo antico mag-
no, sulco profundo longitudinaliter diviso, pone callum sulco angusto
profundo fovea obliqua ad angulum posticum currente utrinque
conjuncto; scutello convexo, ad medium semicirculariter transverse
sulcato ; elytris totis cortaceis convexis, margine antico rotundato.
1878.] HEMIPTERA OF ST. HELENA. 471
et reflexo precipue ad basin, margine interiore pone apicem scutelli
recto, margini elytri alterius haud incumbente ; (membrana aleque
desunt ;) pedibus modicis, tibiis ad apicem subincrassatis ; tarso-
rum articulo 1° brevi, articulo ultimo longissimo.
3 2. Long 3-5, lat. (pone medium) 2-3 m. m.
Hab. in editioribus insule, juxta “ Diana’s Peak.’’
The absence of the membrane prevents me referring this to
any of the sections given by Dr. Stal in his ‘ Enumeratio.’ It
may perhaps belong to a new genus; but as it has all the facies
of a Salda, I have placed it in that genus.
S. aberrans is very variable in the intensity of the brown mark-
ings. Frequently the corium is more or less of a rich fulvous
brown with anastomosing black lines; the whitish spots also vary
in intensity. There is, moreover, a considerable range of varia-
tion in the size of the amimal.
20. Issus cOLEOPTRATUS, F.
Hab. intermedis insulee, ad “‘ Plantation.’’
A common European species, and doubtless introduced into St.
Helena.
LisurniA, Stal.
Inpurnia, subgen. nov.
Antennarum articulus primus articulo secundo equilongus vel paullo
longior. Tibia postica tarso postico parte quarta longior.
21. LipurniA (ILBURNIA) IGNOBILIS, sp. 0.
L. testaceo-brunnea ; fronte distincte, clypeo, pronoto scutelloque
obsolete (carinis exceptis), sterni maculis nonnullis abdomineque
plus minus brunneis; dorso abdominis utrinque vitta longitu-
dinali pallida, in segmentis 1° ad apicem, 2° et 3° ad basin distinc-
tiore, et in segmentis 1°, 4° , 5°, 6°que interne magis, externe
minus brunneo-marginata ; connexivo brunneo, angulis exterioribus
apicalibus pallidis; tarsorum unguiculis piceo-nigris; elytris
subhyalinis, concoloribus,vix dimidium abdominis tegentibus; oculis
subrufis ; capite desuper viso et pronoto longitudine subequalibus,
illo inter oculos breviter prominulo, vertice singulo oculo subequi-
lato ; carinis omnibus discretis ; fronte longa, deorsum dilatata,
carina centrali ad verticem brevissime furcata; clypei carina
centrali obsoleta; pronoto scutelloque distinctissime carinatis,
illius carinis lateralibus obliquis rectis marginem posticum attin-
gentibus ; alis abbreviatis.
©. Long. 5, lat. 2 m. m.
Habitat in excelsioribus insule, ad “ Diana’s Peak.”
Iiburnia diverges from the characters of Liburnia by the longer
first joint of the antennz, and the shorter hind tarsus. In the
collection are two other Homoptera belonging to this family: one,
which is much broken, is perhaps the ¢ of the above; and the
other seems to be an immature Stiroma ?
472 DR. F. B. WHITE ON THE [May 7,
STONASLA, gen. nov.
Corpus cylindricum, elongatum. Caput obtusissimum, latum, thoraci
aquilatum seu paullo latius, desuper visum brevissimum, suban-
gulato-arcuatum, vertice maximam ad partem declivi; facie
dilatata, transversa, fere subquadrata ; apice obtusissime angulato ;
marginibus lateralibus ante oculos distinctissime sinuatis, dein
valde votundato-dilatatis ; plica supraantennali valde obliqua,
oculos haud attingente. Ocelli inter oculos in parte antrorsum
vergente verticis positi, ab oculis remoti, sed inter se quam ab
oculis longius distantes. Pronotum transversum, sat longum,
antice subangulato-rotundatum. Scutellum mediocre, latius quam
longius. Elytraaleque apicem abdominis longe superantia. Elytra
pone clavum angustissime marginata.” Vene elytrorum alarumque
fere ut in Bythoscopo formate ; membrane cellule apicales 4.
Ale ante apicem emarginate. Tibia postice spinosissime.
Genus Bythoscopo affine, forma faciei &c. divergens. [Typus S.
undulata. |
22. SronasLA UNDULATA, sp.n. (Plate XXXJ. fig. 5.)
St. sordide virescenti-flavescens, subopaca ; capitis macula parva
irregulari inter oculos et ocellos (necnon interdum macula inter
ocellos, maculaque utrinque in margine postico pone ocellos), fovea
utrinque ante antennas, parte antica suture frontalis, et basi (media
et lateribus prope basin clypei exceptis), pronoti macula parva
utringue ad marginem anticum, scutelli basi centrali anguste, macula
triangulari utringue prope angulos basales, maculis parvis rotundis
in disco, sulco transversali ad medium angustissime, necnon apice
imo, elytrorum venarum punctis irregularibus subconfluentibus et
lineis dentato-fractis inter venas (membrane exceptis),alarum venis,
sterno (lateribus exceptis), macula ad apicem superiorem femorum
omnium, tibiarum posticarum margine exteriore superiore, unguicu-
lisque necnon abdomine brunneo-nigris ; oculis, ocellis, frontis serie
linearum irregularium transversarum utrinque, femorum maculis
irregularibus, et tarsis anticis intermediisque plus minus brunneis ;
alis infuscatis. Elytris oblongo-lanceolatis.
3 2. Long. 8-93, lat. 3-33 m. m.
Hab. in editioribus insule, ad ‘‘ Cason’s.”
23. STONASLA CONSORS, sp. n.
St. testaceo-brunnea, opaca, elytra subopaca ; capitis macula rotunda
pone ocellos et prope marginem posticum, fovea ante antennas, fron-
tis sutura antice, basi (media excepta) et lateribus prope basin
clypet, necnon serie utrinque lineolarum transversarum, pronoti
linea longitudinali centrali nec apicem nec basin attingente,
macula parva utrinque prope marginem anticum et macula majore
utringue in disco, scutellim acula parva ad mediam basim et macula
triangular prope angulos basales necnon sulco transverso angus-
tissime, clavi venis irregulariter, corti venis distincte, femorum
intermediorum posticorumque macula superiore apicali, unguiculis
1878. | HEMIPTERA OF ST. HELENA. 473
et abdomine subtus brunneo-nigris ; scutelli dimidio apicali tes-
taceo-flavido ; oculis ocellisque brunneis.
3 Q. Long. 73-9, lat, 23-3 m. m.
Hab. in editoribus insulee, ad “ Diana’s Peak” et ‘‘ Halley’s Mount.”
In addition to the differences of coloration, this species may be
distinguished from the last by the internal sector of the elytra having
three cells exteriorly between the base of the elytron and the apex of
the clavus instead of two, as well as by the somewhat smaller stature.
NEHELA, gen. nov.
Corpus cylindricum, elongatum. Caput obtusissimum, latum, thoract
e@quilatum, desuper visum brevissimum angulato-arcuatum ; ver-
tice magnam ad partem declivi; facie dilatata, subtriangulari,
apice producto, lateribus ante oculos late sinuatis, dein leviter
rotundatis ; plica supraantennali valde obliqua, oculos fere attin-
gente. Ocelli inter oculos in parte antrorsum vergente verticis
positi, ab oculis et inter se fere equidistantes. Clypeus distincte
productus, sulco inter basin et frontis apicem haud instructus.
Pronotum transversum, sat longum, antice subangulato-rotundatum.
Scutellum mediocre, latius quam longius. Elytra apicem abdominis
superantia, pone clavum anguste marginata. Elytrorum alarumque
vene ut in Stonasla formate. Tibie postice spinosissime.
Genus Stonasle valde affine, forma faciei, situ ocellorum, et
elypeo distinctius producto divergens.
24. NEHELA VULTURINA, Sp. 0. (Plate XXXI. fig. 6.)
N. atra, opaca ; capitis vertice (maculis parvis 2 ad ocellos et 4 ad
marginem posticum exceptis), oculis, macula triangulari in medio
frontis, antennis et lateribus, pronoti marginibus lateralibus interne
angulatis, margine postico et maculis 2 parvis in disco (interdum
obsoletis), scutelli macula utrinque ad marginem ante apicem,
elytrorum (clavo excepto) lineolis longitudinalibus latis inter
venas, necnon marginibus antico interioreque corit, pedibus anticis
intermediisque (unguiculis exceptis), coxis posticis, femoribus po-
sticis ad apicem, tibiarum posticarum basi spinisque et articulis 2
primis tarsorum posticorum testaceo-brunneis.
3 Q@. Long. 7-73, lat. 2-25 m. m.
Hab. in excelsioribus insulz, ad “ Diana’s Peak” et “‘High Peak.”
Frequently the veins of the clavus are more or less brown; and
more rarely the whole of the elytra is entirely black.
ARGATERMA, gen. nov.
Corpus oblongum, paullo depressum. Caput breve, pronoto paullo
breviore et latiore, margine antico acuto ; vertice inter oculos quam
singulusoculus duplo latiore, horizontali, concavo presertim antice,
margine antico arcuato, acuto, inter ocellos angustissime reflexo.
Ocelli in margine antico prope oculos positi. Facies convexa,
dilatata, subquadrata, apice obtusissime rotundato, lateribus
ante oculos sinuatis, dein leviter rotundatis ; sutura frontali ocellos
attingente. Pronotum transversum, antice arcuatum, inter oculos
Proc. Zoot. Soc.—1878, No. XXXI. 31
474 DR. F. B. WHITE ON THE [May 7,
prominulum, postice truncatum. Scutellum transversum, latius
quam longius. Elytra subcoriacea, tectiformia, oblonga, apicem
abdominis viv superantia, areis pluribus subquadratis instructa.
Ale areis apicalibus 4 instructe, venis in venam periphericam ter-
minantibus, marginem haud attingentibus. Pedes mediocres,
femoribus posticis paullo compressis, tibiis posticis spinosissimis,
tarsis posticis articulo 1° articulis 2° una cum 3° equilongo,
articulo 2° quam 3°* breviore.
Genus Sive et Selenocephalo affine. [Typus A. alticola. |
25. ARGATERMA ALTICOLA, sp. n. (Plate XXXI. fig. 7.)
3 lete fulvus, nitidus ; verticis limbis, pronoti vitta centrali longi-
tudinali et limbo postico, scutelli apice imo, clavi_ marginibus
interne erosis, et fascia ante medium, corii marginibus (exteriore
ad basin excepto), fascia ante medium et maculis pone medium,
necnon areis apicalibus albidis ; verticis marginibus imis antico
posticoque, macula utrinque ad oculos posterius, maculis 4 ad
marginem anticum (2 intermediis geminatis, 1 lunata utrinque
pone ocellos), facie, pronoti maculis parvis 5 ad marginem posti-
cum, sterno ad medium, corii marginibus macularum albidarum
pone medium et arearum apicalium nigris ; abdominis dorso nigro,
segmentis 4°, 5° 6°que et ventre, necnon segmentis genitalibus et
pedibus brunneo-testaceis ; oculis, alis, tibiarum posticarum margine
postico, maculis marginis antici apiceque, tarsorum posticorum
articulorum apicibus, necnon unguiculis omnibus nigro-brunneis.
Q lete fulva, verticis limbo antico, pronoti limbo postico, scutelli
apice imo, elytrorum areis apicalibus flavescenti-albidis ; verticis
marginibus imis antico posticogue, macula utringue ad oculos po-
sterius, maculis 2 in disco, et 4 ad marginem anticum (2 intermediis
interdum postice conjunctis, 1 lunata utrinque pone ocellos),
pronoti maculis 5 ad marginem posticum, elytrorum arearum
plurium parvarum subquadratarum, areolarum elongatarum pone
medium, necnon arearum apicalium marginibus nigris ; cetera ut
in o.
3 Q. Long. 34-5, lat. 13-2 m. m.
Hab. in excelsioribus insule, ad “ Diana’s Peak.”
I have seen but one ¢ ; and it is possibly a variety. The elytra
of the 2 have numerous black-margined cells or cell-like spots more
or less variable. On the clavus and corium, to beyond the middle,
these are small and vary in size, but are more or less square. Between
these and the apical cells there is a transverse row of 4, more elon-
gate, cells, of which one or more are sometimes subdivided. Beyond
these are 5 marginal cells at the apex of the elytron. The veins of
the elytron are scarcely distinguishable.
26. ARGATERMA MULTISIGNATA, Sp. 0.
A. fulva, nitida 5 verticis limbo antico, pronoti margine imo postico,
elytrorum areis apicalibus flavescenti-albidis ; verticis marginibus
imis antico posticoque, macula utrinque posterius ad oculos, maculis
2 in disco, 3 ad marginem anticum (1 in medio interdum fissa, 1
1878.] HEMIPTERA OF ST. HELENA. 475
utrinque pone ocellos), pronoti margine antico maculis 7, 3 inter-
mediis elongatis magnis transversis in fasciam sepe confluentibus,
disco fascia transversa abbreviata, limbo postico, necnon macula
parva utringue ad angulos laterales, scutelli macula elongata
utringue ad basin, elytrorum maculis plurimis subquadratis arei-
Formibus, sterno abdomineque nigris ; sterni lateribus, abdominis
segmentis 6° et genitalibus (his nigro-signatis) pedibusque brunneo-
testaceis ; femoribus anticis et intermediis supra, posticis totis
(basi apiceque exceptis), tibiis posticis magnam ad partem, tarsis
anticis et intermediis ad apicem, posticorum apicibus articulorum
omnium, unguiculis oculisque nigro-brunneis ; alis nigro-fuscis
nigro-venosis ; elytris areis apicalibus marginalibus nigro-mar-
ginatis 6 instructis.
3 2. Long. 3-43, lat. 13-2 m. m.
Hab. in excelsioribus insule, ad ‘‘ Diana’s Peak.’’
In stature and general colouring rather like the preceding, but dis-
tinguished at first sight by the much more numerous markings.
Vertex more obtusely rounded in front and less prominent. Elytra
with the veins scarcely distinguishable, covered with many somewhat
square-shaped cell-like black markings, arranged in longitudinal
rows. Apex of the elytron with six marginal cells, of which the first
and the last are the largest. Venation of the wing different from
what it isin 4. alticola. There are four apical-cells bounded by a
peripherical vein as in that species ; but the second cell (counting the
apical as the first) is pedunculate, the peduncle joining the first and
second sectors; and at the base of the fourth cell is a smaller cell
formed by a transverse vein cutting off the base of the fourth cell.
The venation is otherwise normal, but rather subject to variation.
The elytra have sometimes obscure pale markings on the inner mar-
gin of the clavus, and vary also in the intensity of the black mark-
ings of the pronotum and scutellum.
27. Grypores (?) INSULARIS, sp. n.
G. aurantio-flavescens, nitidus ; verticis macula subquadrata utringue
anterius et macula posterius prope oculos, frontis lineolis trans-
versis in medio interruptis, clypei vitta centrali ad apicem dila-
tata, lororum marginibus, linea ante et macula pone antennas,
maculaque ante oculos brunneis vel pallide brunneis; vertice
utrinque linea obliqua brevi, pedibus maculis nonnullis, abdominis
dorso, et corpore subtus maculis paucis nigris ; scutello, abdominis
dorsi lateribus, et corpore subtus flavescentibus ; elytrorum mar-
gine antico late et venis, necnon valvulis genitalibus flavescenti-
albidis ; membrana dilutissime fusca.
3 2. Long. 5 m. m.
Hab. in editioribus insule, ad ‘‘ West Lodge.”
Perhaps scarcely correctly placed in Grypotes. Head, seen from
above, angularly rounded and prominent between the eyes; as long
as the hind margin between the eyes ; broadly concave and witha fine
impressed middle line; ocelli near the eyes; face very convex, and
clypeus produced but scarcely curved; pronotuma er —- than
1
476 DR. WHITE ON THE HEMIPTERA OF ST. HELENA. [May 7,
the head. There are sometimes additional brown markings on the
vertex or face ; and the spots on the legs are also occasionally confluent.
28. THAMNOTETTIX SANCTZ-HELEN&, sp. 0.
T. pallide aurantiacus, nitidus ; capitis maculis 2 parvis obsoletis
inter oculos, frontis lineolis obsoletissimis transversis, pronoti
margine postico, scutelli angulis basalibus, clavi nebulis 2 male de-
finitis (1 ante, 1 pone medium), corii nebulis 3 (1 ante, 1 pone
medium) alteraque ad basin membrane indistincte dilutioribus ;
corpore subtus abdomineque supra et sublus flavescentibus ; tibiis
posticis maculis nonnullis et unguiculis omnibus nigris.
3 9. Long. 5 m. m.
Had. in excelsioribus insule, ad “ Diana’s Peak.”
The always ill-defined pale markings are sometimes scarcely
perceptible. The head, seen from above, is prominent and obtusely
angularly rounded between the eyes; the face is very convex, and
the pronotum broader than the head.
29. JassUS WOLLASTONI, sp. n.
J. flavescenti-albidus ; verticis vitta transversa inter oculos, frontis
vittis 2 latis longitudinalibus, elypei dimidio apicali saltem et
macula utringue ante oculos, pronoto (marginibus postico et
lateralibus exceptis), sterni maculis ad latera, scutelli macula
quadrata ad medium basale, clavi vitta ad suturam clavalem et
vitta centrali a basi ad medium, corii vitta subundulata a medio
ad membranam currente, lineis longitudinalibus 2 minoribus (1
inter vittam centralem et basin elytri, 1 inter vittam centralem et
apicem corii), margine antico toto corii, tibiis anticis intermediisque
et tarsis omnibus aurantiacis vel pallide aurantiacis ; clavi linea
juxta apicem marginis antici, membrane margine basali, sterni
maculis nonnullis ad medium, abdomine (marginibus segmentorum
exceptis) nigrescentibus ; membrana, alis fere totis, unguiculisque
Juscis vel brunneo-fuscis ; venis ad apicem corii membraneque
pallide flavidis, venis ceteris obsoletis ; capite pronoto fere equi-
longo, rotundato et levissime prominulo inter oculos ; verticis
margine postico quam singulus oculus paullo latiore ; vertice leviter
dilatato antrorsum, foveola parva utrinque instructo ; fronte haud
carinata ; marginibus oculorum desuper visorum divergentibus ;
ocellis prope oculos sitis ; elytris oblongis.
3 §. Long 6, lat. 2 m. m.
Hab. in editioribus insule, ad ‘ Cason’s,’”’ ‘ Diana’s Peak,” et
“*Vine-tree Gut.”
Sometimes the clavus and inner margin of corium are infuscated,
and the hind tibie have a row of black dots on the outside.
30. CHLORITA EDITH#, sp. n.
C. pallide flavescenti-albida ; verticis maculis 2 elongatis inter
oculos, et pronoti maculis 2 in disco sanguineis ; scutello (macula
quadrata ad medium basale excepta), clavi linea ad marginem
interiorem ab apice scutelli currente, corii margine interiore fere
TOO Te SB Geil Se
1878.] MR. P, L, SCLATER ON FULIGULA NATIONI. 477
a basi ad apicem clavi, vitta dilutiore centrali inter apicem clavi et
margine antico posita, et nebula ad marginem anticum apicem
versus elytri lete aurantiacis ; oculis, unguiculis et macula ad basin
membrane nigrescentibus; membrana pallide fusca, ad basin
obscuriore, venis late pallide flavescenti-albidis, maculas 5 mar-
ginales (4 apicales) et 2 centrales cingentibus ; femoribus tibiisque
ad apicem flavescentibus ; capite obtuse angulariter rotundato-
prominulo inter oculos, longiore supra quam ad marginem posti-
cum inter oculos ; pronoto semicirculari, dimidio marginis postici
@equilongo.
3 2. Long. 3 m. m.
Hab. ineditioribusinsule, ad “ Cason’s,’’“ High Peak,” et “ Diana’s
Peak.”
Species venusta, in honorem Domine Edith Wollaston, quee Lepi-
doptera et Coleoptera insulz Sanctz-Helene diligentissime collegit,
dicata.
EXPLANATION OF PLATE XXXI.
Fig. 1. Hapa contorta, p. 466. Fig. 5. Stonasla undulata, p. 472.
2. Agrametra ethiops, p. 468. 6. Nehela vulturina, p. 473.
3. Vernonia wollastoniana, p.470. 7. Argaterma alticola, p. 474.
4. Salda aberrans, p. 470.
3. Further Remarks on Fuligula nation. By P. L. Scrater.
[Received April 12, 1878.]
(Plate XXXII.)
Professor Nation of Lima, Peru, has now forwarded to me a
second specimen of the remarkable new Fuligula which Mr. Salvin
and I described in the Society’s ‘ Proceedings’ last year (1877, p.
.§22), and dedicated to its discoverer. This is a male, the former
example having been a female ; but, as will be seen on comparison,
there is not a great difference in plumage between the sexes. Prof.
Nation says that this bird is rare in the neighbourhood of Lima, but
that he has lately ascertained that it is common in the north of Peru,
and is found in great numbers on the lakes near the coast in certain
months of the year.
The species may now be diagnosed more correctly as follows :—
FuLIGuLA NATIONI. (Plate XXXII.)
Fusco-atra fere unicolor, in ventre brunnescentior, hypochondriis
et genis in brunneum transeuntibus ; secundariorum parte basali,
fasciam distinctam alarem constituente, alba; subalaribus nigris ;
remigum pagina inferiore obscure cineracea; rostro et pedibus
nigris : long. tota 18-0, ale 8°5, caud@ 2°5, tarsi 1:45, dig. med.
cum ungue 2°6, rostri-a rictu lin. dir. 2°2. Femina omnino
brunnescentior, supra et in ventre tenuissime nigro vermiculata ;
capitis lateribus in brunnescenti-castaneum trahentibus ; et cras-
sitie minore.
Hab. Peruvia occidentalis, prope Lima (Nation).
478 MR. P. L. SCLATER ON FULIGULA NATIONI. [May 7,
Trachea of Fuligula nation g.
Fig. 1. Front view.
Fig. 2. Dilatation at its lower end, seen from the left side.
1878. ] MR. P. L. SCLATER ON FULIGULA NATIONI. 479
This species appears to be a true Fuliguline duck, the only known
representative of the group in South America. In colour it is quite
different from any other species of the genus known to me, being
immediately recognizable by its black dress and the transverse
white bar on the wing.
The characteristic Fuliguline white spot between the rami of
the mandible is small in the male but quite distinct, larger in the
female. The white colour in the secondaries occupies rather more
than the basal two thirds, but does not extend onto the outer elon-
gated feathers ; it likewise pervades portions of both webs of some
of the inmost primaries. The hinder toe has a well-developed lobe,
almost as broad as in F. collaris.
Prof. Nation has also sent to me the dried trachea of the male
specimen, for the preparation of a figure and description of which I
am indebted to Prof. Garrod.
“In the trachea,” Prof. Garrod remarks, “ of this duck, as in the
males of the sea-ducks generally, there is a large dilatation at the lower
end, on the left side, composed of an osseous framework supporting
membranous walls. The outer one of these is traversed by an inter-
vening osseous bar (see fig. 2), which courses backwards and upwards
from its anterior inferior angle, and sends two small extra bars to the
supero-marginal rim, and so forms a pair of oval fontanelles, before
it terminates at the superior and posterior angle of the outer wall.
“The wall of the tracheal box which faces inwards is ossified
through nearly its entire extent, a few small membrane-covered
fontanelles being found not far from its superior margin. In this
respect the species differs from Fuligula rufina, in which the wall
under consideration is almost entirely membranous, whilst it is almost
identical with F. marila.
“The outer aspect of the terminal tracheal box is represented in
fig. 2.
Pa Fig. 1 gives a front view of the trachea, which is seen to be con-
siderably and fusiformly dilated in its middle part, in very much the
same way as in Fuligula rufina, F. marila, Nyroca leucophthalma,
Clangula vulgaris, and Mergus serrator’, except that in the last-
named species the enlargement is situated somewhat nearer the
mouth,
‘In F. rufina the dilatation of the middle of the trachea is rather
more considerable and more localized to the lower portion of the
windpipe, whilst in F. marila it is more extended and not quite so
considerable in breadth. In fact F’. nationi is almost exactly inter-
mediate between the two, tending, if at all, towards the Jatter
species.
“In the South-American Metopiana peposaca the mid-tracheal
dilatation is much more decided and more limited, forming a nearly
globose cavity® like that in Melanitta fusca and Clangula his-
trionica.”
1 Vide Eyton, ‘ Monograph of the Anatide,’ 1838, p. 63.
2 See P. Z. S. 1868, p. 146, and 1875, p. 154.
480 MR. A. G. BUTLER ON [May 7,
4. On a small Collection of Lepidoptera from Jamaica. By
Axrtruour G. Butter, F.L.S., F.Z.S8., &e.
[Received April 12, 1878.]
The Lepidoptera enumerated in the present paper were collected
at or near Kingstown, Jamaica, by Mr. James John Bowrey, who
has kindly permitted a selection to be made from them of all the
species required for the national collection.
Among the Butterflies, as might have been expected, there is very
little new, a species of Pamphila being the only one to which I have
ventured to give a name; but among the more obscure and small
Moths there are about two dozen novelties.
RHOPALOCERA.
1. Danats arcuippus, Fabricius, Ent. Syst. iii. 1, p. 49 (1793).
The males from Jamaica seem to possess a constant peculiarity in
the immaculate black border of the secondaries.
2. Danats JAMAICENSIS, Bates, Ent. Mo. Mag. i. p. 33 (1864).
3. Caisto zAncis, Fabricius, Syst. Ent. p. 486 (1775).
4, Papuia portiA, Fabricius, Syst. Ent. p. 507 (1775).
5. AGANISTHOS ORION, Fabricius, Syst. Ent. p. 485 (1775).
6. HrTEROCHROA ABYLA, Hewitson, Ann. Nat. Hist. ser. 2,
vol, vi. pl. 9. fig. 7 (1850).
7. Evpacis rGma, Fabricius, Syst. Ent. p. 496 (1775).
8. Lucrnta capMA, Drury, Ill. Ex. Ent. ii. pl. 18. figs. 1, 2
(1773).
9. Marpesta ELEUCHA, Hibner, Samml. exot. Schm. ii. pl. 50
(1816-24).
10. Gynzxcra piece, Linneus, Mus. Lud. Ulr. p. 287 (1764).
11. ANARTIA JATROPH#, Linneeus, Mus. Lud. Ulr. p. 289(1764).
12. JunonrA GENOVEVA, Cramer, Pap. Exot. iv. pl. 290.
figs. E, F (1782).
13, CysTrnEURA porcas, Fabricius, Syst. Ent. p. 508 (1775).
14. Puycropes rristA, Poey, Cent. Lep. (1833 ?).
15. Eunica tTatiua, Herrich-Schiffer, Ausl. Schm. figs. 69-72
(1852-58).
In the examples from Jamaica the white spots are smaller than
in the figure.
1878. ] LEPIDOPTERA FROM JAMAICA. 481
16. Everoreta wecesia, Cramer, Pap. Exot. iii. pl. 209.
figs. E, F (1782).
17. Dione vaniLu#, Linneeus, Mus. Lud. Ul. p- 306 (1764).
18. CoLanis pEeixA, Fabricius, Syst. Ent. p. 510 (1775).
19. Hexicontius CHARITHONIA, Linnzeus, Syst. Nat. i. 2
p- 757 (1766).
20. LampipEs cerAunvus, Fabricius, Ent. Syst. iii. 1, p. 303
(1793).
>
21. TmMoLus coLUMELLA, Fabricius, Ent. Syst. iii. 1, p- 282
(1793).
22. Tmoxus pan, Drury, Ill. Ex. Ent. ii. pl. 23. figs 3, 4 (1773).
23. Naruauis roux, Boisduval, Sp. Gén. i. p- 589 (1836).
24, Trrtas GNATHENE, Boisduval, Sp. Gén. i. p. 680 (1836).
Mr. Bowrey has observed this species flying over low bushes.
The flight of Terias is always very low.
25. TertaAs ELATHEA, Cramer, Pap. Exot. ii. pl. 99. figs. C, D
(1779).
26. TERIAS EUTERPE, Ménétriés, Bull. Mosc. p- 299 (1832).
27. Kricocont TErtssa, Lucas, Rev. Zool. p. 429 (1852).
28. KricoGonia LysipE, Godart, Enc. Méth. ix. p- 98 (1819).
29. AMYNTHIA M&ZRULA, Fabricius, Syst. Ent. p. 479 (1775).
30. CaLLIpRYAS SENN#&, Linneus, Syst. Nat. i. 2, p. 764 (1766).
Mr. Bowrey has seen vast clouds of this species passing over the
town of Kingston, sometimes for a week at a time.
31. Appras poryt, Butler, P. Z. S. p. 49 (1872).
32. SyncuLo# sopra, Boisduval, Sp. Gén. i. p. 495 (1836).
33. Papiiio potypamas, Linneus, Mus. Lud.UIr. p. 192 (1764).
34, Papiiio stnon, Fabricius, Syst. Ent. p. 452 (1775).
35. Papixio pexavs, Fabricius, Syst. Ent. p. 444 (1775).
36. Papitio HomeRvs, Fabricius, Ent. Syst. iii. 1, p. 29 (1793).
37. PariLio CRESPHONTES, Cramer, Pap. Exot. ii. pl. 165.
fig. A, 166. fig. B (1779).
38. Papiyio THERsITEs, Fabricius, Syst. Ent. p. 453 (1775).
39. Gonrurus proteus, Linneus, Mus. Lud. Ulr. p. 333
(1764).
482 MR. A. G. BUTLER ON [May 7,
40. GonruRvs CATILLUS, Cramer, Pap. Exot. iii. pl. 260. figs. F, G
(1782).
41. Tetxconvus, sp.? (Olive-brown, dull green at the base.)
This species is quite common in collections, and therefore is sure
to have been described ; but it would be necessary to work out nearly
the whole family in order to identify it, many of the descriptions
of Hesperiide being so bad that their determination is simply
impossible.
42. Prorerrs aMynTAS, Fabricius, Syst. Ent. p. 533 (1775).
43. PAMPHILA ETHLIUS, Cramer, Pap. Exot. iv. pl. 392, figs. A, B
(1782).
44, Pampuita nycTe.ius, Latreille, Enc. Méth. ix. p. 746
(1823).
45. Pampnita pHyivs, Drury, Ill. Ex. Ent. i. pl. 13. figs. 4, 5
(1773).
46. Pameuita uTHA, Hewitson, Descr. Hesp. p. 37. n. 32 (1868).
The form from Jamaica being rather different from that of
St. Domingo (and probably of Cuba), I append a description.
Wings above bright tawny, reddish at the base ; a broad choco-
late brown marginal belt from the end of the cell of each wing
round the outer border to the external or anal angle, where the
fringe becomes tawny; primaries with a large dark brown spot
across the median vein ; head and collar metallic green, bordered and
crossed by testaceous lines of upright scales ; remainder of the body
above dull tawny, slightly olivaceous in front, yellowish below,
with blackish spots down the centre of the venter. Primaries below
with the disk of a yellower tint than above; the blackish median
spot extended to the base, a blackish spot beyond the cell; outer
border broadly brown, becoming ferruginous upon the costa:
secondaries ferruginous, with a spot in the cell and a squamose
angulated discal belt more or less orange ; anal fringe bright orange.
Expanse 2 inches.
I have to thank Mr. Du Cane Godman for the identification of
this handsome species. I think it may be the insect intended by the
description of Hudamus capucinus, Lefebvre; but the Latin of that
description cannot be translated. It runs thus :—“ E. alis paululum
subrotundatis, supra piceo-fuscis ad basin ochraceo-rubescentibus ;
anticis in mare supra arcu minuto subtus macula fere dubia, in
disco luteis (feemine utrinque maculis duabus geminis eodem colore)
nervula sectis ; subtus anticis ad basin et disco inferiori nigris ;
omnibus alis fuscis atomis rubris griseisque omnino rubescentibus.”
The whole of the descriptions of Lepidoptera in Ramon de la
Sagra’s ‘Cuba’ are of this character. LH. trinitad is said to have
three front wings on each side, with transparent unequal and yel-
lowish spots, “ anticis utringue tribus maculis transluctdis inequali-
1878. ] LEPIDOPTERA FROM JAMAICA. 483
’
bus et flavescentibus ; and although in this case it is evident what
is intended, in other cases it is not so’. Surely where a Latin
diagnosis defies all possibility of translation it ought to be ignored.
47. PAMPHILA INSOLATA, N. sp.
3. Primaries bronzy chocolate-brown, with tawny basal area; a
large black patch below the cell, its anterior border embossed and
greyish: secondaries tawny ochraceous; the costal area, outer
border, and veins blackish: body greenish, abdomen dull tawny,
palpi below sulphur-yellow ; legs tawny. Wings below sordid ochra-
ceous ; primaries with the basal half of a brighter tint, the interno-
basal area and a spot below the cell blackish ; internal border dark
greyish. Expanse | inch 8 lines.
Quite unlike any other species known to me.
48. Pampnita oTHo, Smith & Abbot, Lep. Ins. Georg. i.
pl. 11 (1797).
49, AcHLYODES PorRILLO?, Lefebvre, Ramon de la Sagra’s Cuba,
p- 641 (1857).
The single example taken by Mr. Bowrey has only four hyaline
apical dots on each of the primaries; but this may be what is
intended by “ anticis ad apicem 8 punctis translucentibus albis cir-
culatim dispositis,” the only difficulty being the arrangement of four
dots so as to produce a circle.
HETEROCERA.
50. Enyo camertus, Cramer, Pap. Exot. iii. pl. 225. fig. A
(1782).
51. CuzrocaMpa cutRoN, Drury, Ill. Ex. Ent. i. pl. 26. fig. 3
(1770).
52. DeiLerHiLA LINEATA, Fabricius, Ent. Syst. ii. p. 541
(1775).
53. PHILAMPELUS SATELLITIA, Linnzus, Mantissa, i. p. 539
pl ib):
54. Pacuyxia Ficus, Linneus, Mus. Lud. Ulr. p. 352 (1764).
55. AMBULYX sTRIGILIS, Linneeus, Mantissa, i. p. 538 (1771).
56. AMPHONYX CLUENTIUS, Cramer, Pap. Exot. i. p. 124,
pl. 78. fig. B (1779).
57. ANCERYX FASCIATA, Swainson, Zool. Ill. 2nd ser. vol. iii.
pl. 150 (1823).
58. DitorpHonotra ELLO, Linneus, Mus. Lud. Ulr. p. 351
(1764).
1 Examine the description of Hesperia baracoa as an example.
484 MR. A. G. BUTLER ON [May 7,
59. DiLopHonotTa xNoTRUS, Cramer, Pap. Exot. iv. pl. 201.
fig. C (1782).
60. DiLopHONOTA OMPHALE#, Boisduval, Lép. Guat. p. 72
(1870).
61. Prororarce rustica, Fabricius, Syst. Ent. p. 540 (1775).
62. Proroparcre JAMAICENSIS, Butler, Trans. Zool. Soc. ix.
p- 608 (1876).
63. Protoparce crneuLaTa, Fabricius, Syst. Ent. p. 545
(1775).
64. PsEuDOSPHINX TETRIO, Linneus, Mantissa, i. p.538 (1771).
The larva of P. tetrio is well known to Mr. Bowrey, who de-
scribes it as “ black with yellow bands, and with the head, feet, anal
claspers, and horn scarlet.”
65. CosMosSOMA TYRRHENE, Hiibuer, Samml. ex. Schm. Zutr.
figs. 483, 484.
66. CosmMosoMA AUGE, Linneus, Syst. Nat. ii. p. 807. n. 46
(1766).
67. EmpyrEUMA PUGIONE, Linneeus, Syst. Nat. il. p. 807. n. 45
(1766).
68. ARE marorinata, Drury, Ill. Ex. Ent. ii. pl. 22. fig. 2
(1773).
69. ComposiA syBARIs, Cramer, Pap. Exot. i. pl. 71. fig. E
(1779).
70. Evcua#res rnsuuata, Walker, Lep. Het. iii. p. 734. n. 5
(1855).
71. Deropera speciosa, Walker, Lep. Het. ii. p. 568 (1854).
Common in grass. The larva much resembles that of our Euro-
pean Callimorpha jacobee: it is orange, banded with warty-black
bands, and with rather long black bristles projecting from the
warts ; head and legs reddish.
72. CINCIA PALLIDA, 0. sp.
Primaries white, with cream-coloured veins, a black dot at the
base; four oblique parallel equidistant angular series of black dots,
also a marginal series; secondaries pearly grey; head and collar
white; thorax white dotted with black; abdomen pearly grey.
Under surface pale greyish, the spots obsolete; costal margin of
primaries white, with three pale brown spots beyond the middle ;
body whitish. Expanse ] inch | line.
73. Mutona LApiIpARIA, Walker, Lep. Het. Suppl. v. p. 1896
(1866).
1878.] LEPIDOPTERA FROM JAMAICA. 485
74. Guissa VARIEGATA, Walker, Lep. Het. ii. p. 380 (1854).
This beautiful little species seems to be not uncommon. Mr.
Bowrey found it settled on the under surface of the leaves of coffee.
75. LAURONA ERGOLIS, Walker, Lep. Het. ii. p. 335 (1854).
This common species is said to have a hairy caterpillar.
76. EvrHisaNoTia Trmais, Cramer, Pap. Exot. iii. pl. 275.
fig. B (1782).
77. EUGLYPHIA HIEROGLYPHICA, Cramer, Pap. Exot. ii. pl.
147. fig. D (1779).
The larva of this species, which is slightly hairy, feeds on lilies.
78. OcHRIA NIVEOPICTA, N. Sp.
Primaries above deep coppery red; the orbicular and reniform
spots represented by circles of white dots enclosing white spots ;
costal margin white-dotted; a few scattered white dots on basal
area, a discal series and a marginal series; a submarginal series of
white spots: secondaries shining brown, slightly darker at outer
border: body corresponding in colour with the wings. Under sur-
face pinky brown; internal areas of the wings more or less broadly
white ; primaries with pale discocellular spot and outer border.
Expanse | inch 4 lines.
79. PRODENIA IGNOBILIS, 0. sp.
Primaries above shining whity brown; costal margin whitish,
crossed by oblique brown spots ; two black dots and a small brown
spot at the end of the cell; a double discal series of black dots on
the veins; an ill-defined series of internervular bronzy streaks; an
oblique bronzy streak from the second median interspace to near the
base of the inner margin, where it terminates in a black line; a
marginal series of black dots; fringe bronzy, crossed by white lines
from the ends of the veins, and with a white internal margin:
secondaries semitransparent opaline white: body pale brown, the
head, collar, and tegule whitish ; a darker brown patch on each
side of the collar. Wings below white, most of the markings obso-
lete ; primaries sericeous, with bronzy-tinted fringe and costa; a
marginal series of black dots ; body below whitish, varied with pale
reddish brown. Expanse 1 inch 3 lines.
This species and the next have the general aspect of Leucania.
80. PRopENIA PAUPER, 0. sp.
Primaries above shining whity brown, with five or six black costal
dots and three (indicating the reniform spot) at the end of the cell ;
an arched discal series of minute black dots on the veins ; outer
border rather dusky ; a marginal series of minute black dots:
secondaries semitransparent opaline white ; the apex with a brown
marginal line ; costal area pale brown: body whity brown ; front of
the collar brown, with a black posterior edge. Under surface
shining, the markings obsolete. Expanse 1 inch 3 lines.
486 MR. A. G. BUTLER ON [May 7,
81. ConpicA PALPALIS, Walker, Lep. Het. ix. p. 240 (1856).
82. APAMEA INTERMITTENS, Walker, Lep. Het. xv. p. 1686.
(1858).
83. Periteea Mositis, Walker, Lep. Het. x. p. 277 (1856).
84. Hapena asrpa, Felder, Reise der Nov. Lep. iv. pl. 109.
fig. 7 (2).
The example from Jamaica has the apices of the primaries rather
more rounded and the pattern less strongly indicated than in
Felder’s figure ; still it is so like it that it would be unsafe to regard
it as a distinct species.
85. Exousa aLercans, Walker, Lep. Het. xiii. p. 1118 (1857).
This genus is nearly allied to Hrastria; the same species is
described subsequently by Walker under the name of Lrastria
includens.
86. GoNODONTA NuTRIX, Cramer, Pap. Exot. iv. pl. 312. fig. B
(1782).
87. Hypuaa purerA, Cramer, Pap. Exot. ii. pl. 103. figs, D, E
(1779).
88. CosmMoPpHILA EROSA, Hubner, Samml. ex. Schm. Zutr.
figs. 287, 288.
89. DrEcELEA BOWREYI, 0. Sp.
Primaries pale brown mottled with black; a broad central band
(the internal third of which is black and the remainder of the
ground-colour), limited internally by a black line, and externally by
an angulated and slightly irregular white line; a black line across
the base, terminating on interno-median area in a triangular black
patch ; several black costal dashes and an oblique black line from
the costa across the central band; a subcostal greyish lunule partly
bordered by pearly scales beyond the central band; a very irregular
dentate-sinuate whity-brown discal stripe with dark borders ; a sub-
marginal series of black bracket-like markings separated by white
dots upon the veins ; fringe spotted with dark brown: secondaries
reddish brown, darker towards the outer border; a black sinuated
marginal line interrupted on the veins by white dots; fringe pale
brown, bordered and intersected by parallel dusky lunules: body
pale brown speckled with black; collar with an ill-defined central
black line, the border pearly ; a few pearly whitish scales scattered
over the thorax ; abdomen with three very prominent dark brown
dorsal tufts or masses of scales. Primaries below pale olive-brown
slightly sericeous; a black costal spot before the middle, and a
second smaller one beyond the middle; the remainder of the costa
alternately brown and white; disk darker than the rest of the wing,
crossed by a nearly straight dusky line, and limited externally by a
very irregular dentate-sinuate line ; a sinuated black marginal line,
interrupted by white dots upon the veins; fringe with red-brown
1878. ] LEPIDOPTERA FROM JAMAICA. 487
external spots: secondaries pinky brown, crossed by two dark
lines which form a central semicircular area, in the middle of which
is the dark discocellular lunule; a dusky submarginal belt ; margin
and fringe as above: body below pinky-brown; tarsi black, banded
with testaceous or pinky whitish. Expanse 1 inch 6 lines.
This singular species has somewhat the aspect of a Dasychira ; it
seems, however, to be allied to the genus Nenia.
90. Homorrera TERROSA, Guénée, Noct. iii. p. 11. n. 1332
(1852).
Whether Walker has rightly identified this species I cannot say,
but his H. posterior is the same as his H. terrosa.
91. Boxtrna rascioLaris, Hibner, Samml. ex. Schm. Zutr.
figs. 443, 444.
92. Boxtina cunEarts, Guénée, Noct. iii. p. 70. n. 1414 (1852).
93. BoLINA PERPENDICULARIS, Guénée, Noct. iii. p. 65. n. 1404
(1852).
94. BoLINnA EVELINA, 0. sp.
Markings nearly as in B. ochrodes, but the primaries slaty grey,
the large reniform patch of the same colour, the internal two thirds
of the central band whity brown mottled with greyish brown; two
triangular black subcostal spots resting upon the inner margin of
the trisinuate red-brown discal stripe; external area soft silver-
grey; marginal line yellowish; fringe brown, white at external
angle; secondaries with the basal half and fringe white, the external
half black; thorax lilacine, abdomen grey. Wings below with the
basal half and fringe white, the external half blackish; primaries
with a white spot just beyond the cell; fringes tipped with black ;
body white; the palpi and legs fringed with pink. Expanse | inch
9 lines.
Most like B. agrotoides in the pattern and coloration of the
primaries.
95. Botina? conrirmans, Walker, Lep. Het. xiii. p. 1157.
n. 25 (1857).
This species would, in my opinion, be better placed with Biula.
I can find no difference whatever between Syneda limbolaris (a
species much like a Bolina) and Grammodes grandirena of Walker’s
Catalogue. ;
96. Prostna NuMERIA, Drury, Ill. Ex. Ent. i. pl. 23. fig. 5
(1770).
The species from Venezuela is quite distinct, the specimens from
St. Domingo identified by Walker as P. mexicana being referable to
P.numeria. For Walker’s P. numeria I would propose the name of
Hypogramma confusa ; it is nearly allied to Hypogramma sublucida.
488 MR. A. G. BUTLER ON [May 7,
97. THysANIA ZENOBIA, Cramer, Pap. Exot. ii. pl. 115. A, B
(1779).
98. Eresus opora, Linneus, Syst. Nat. ii. p. 811. n. 11 (1766).
The chrysalis of this species is enclosed in a slight cocoon, to
which leaves are attached.
It appears to me that at least five species have been confounded
under this name; of these three are already figured as follows :—
1. E. odora of Linnzus, in which both sexes are alike in general
pattern and both shot with purple; Drury’s figure represents a
specimen in which the purple seems to have faded. Jamaica and
St. Domingo.
2. E. odorata of Clerk, in which both sexes are again alike, but
different from E. odora, a strongly dentate whitish belt intersected
by black lines running through both wings. St. Domingo and New
Granada.
3. E. agarista of Cramer, in which the male has a pale discal
sinuated belt, and the wings from the centre shot with rose-colour ;
whilst the female has the belt snow-white, intersected by black
lines and a feeble Jilacine tint beyond the centre of the wings.
Brazil.
E. agarista is the most richly-coloured species.
We also have a female from Havannah, which is certainly dif-
ferent, and examples of a species which seems to be distinct from St.
Kitts, Nevis, and New Granada.
99. Puurys GARNOTI, Guénée, Noct. iii. p. 307 (1852).
In my opinion this species would be better placed in Trigonodes
than in Phurys.
100. ACHHA INDISTINCTA, 0. Sp.
Primaries red-brown, the base and a diffused belt beyond the
middle paler ; a nearly triangular subapical costal chocolate- brown
patch bounded externally by a white line ; an apical dark-brown spot ;
costa greyish : secondaries smoky brown, crossed at the middle by a
whitish stripe; fringe white; a submarginal sordid white spot near
the anal angle: body whitish. Wings below pale olive-brown, with
the basal area whitish ; primaries with an abbreviated sordid white
discal baud ; a bifid subapical white spot: secondaries with brown
discocellulars, followed by a white central band; a marginal sub-
anal white spot ; fringe snow-white: body creamy-white, Expanse
1 inch 9 lines.
Allied to A. eyllota.
101. PoaPHILA CINEREA, 0. sp.
Ashy grey, wings with diffused chocolate-brown external area; a
faint indication of an irregular central belt, expressed by zigzag
limiting lines; an indistinct grey zigzag discal line across the brown
area; primaries with a white spot at the inferior angle of the dis-
coidal cell. Under surface paler ; the outer edge of the indistinct
1878.] LEPIDOPTERA FROM JAMAICA. 489
central belt relieved by a whitish border; primaries without the
white spot ; secondaries with an ill-defined dusky discal belt ; fringe
white. Expanse | inch 3 lines.
102. Remicia DIssEVERANS, Walker, Lep. Het. xiv. p. 1495
(1857).
This species is identical with R. persubtilis of Walker, and allied
to his Ophiusa delinquens.
103. RuxescrPHa ELEGANS, nD. Sp.
Allied to R. servia, but with narrower wings, the palpi much
more slender; the primaries with less distinctly sinuated margin; a
small white dot on lower median interspace close to the margin ; no
yellow spots ; secondaries pale reddish instead of smoky brown, the
base and costal area white ; thorax burnt-sienna red like the prima-
ries ; abdomen very pale brown, almost white : wings below much
whiter than in R. servia; body below snow-white, palpi and tibize
of anterior and middle pairs of legs brown. Expanse | inch 7 lines.
I find that the genus Rhescipha is most nearly allied to Tetra-
tocera.
104. THERMESIA GEMMATALIS, Guénée, Noct. iii. p. 355. n.
1828 (1852).
105. Azazia MonsrraTuRA, Walker, Lep. Het. xv. p. 1564
(1858).
This species is certainly nearer to Azazia than to Thermesia, with
which genus Walker placed it.
The following genus has been a puzzle to me; but it appears to
be more nearly allied to Zuelidia than to any thing else.
CALLISCOTUS, n. gen.
Form and general structure of Zuclidia, but the body less robust,
the palpi shorter, with very short terminal joint, porrect ; abdomen
shorter ; primaries comparatively longer; subcostal branches of
secondaries emitted from a footstalk; discocellulars concave but
very oblique. Type C. bowreyi.
106. Catuiscotus BowReYI, n. sp.
Primaries pale brown, crossed by numerous irregularly dentate-
sinuate black lines; a broad dark-brown subbasal belt, beyond
which are two large patches a little, paler than the ground-colour,
outlined in black ; in the middle of the upper one is a small ocellus,
representing the reniform spot, and pale brown edged with black
and encircled by a yellowish iris; a few scattered white scales on
the external area; subapical area dusky; a dentate-sinuate pale
line, white, and followed by triangular black spots between the veins;
an interrupted black marginal line: secondaries fuliginous brown,
with dusky external border; a dark angulated discal stripe and a
blackish anal spot: body whity brown. Wings below whity brown,
Proc. Zoou. Soc.—1878, No. XXXII. 32
490 MR. A. G. BUTLER ON [May 7,
with discocellular spots, angulated discal stripes, and external
borders black ; the apices, a triangular spot at the centre of each
outer margin, the fringes, and some spots at the anal angle of
secondaries white; body below whitish. Expanse 1 inch 7 lines.
A pretty species, somewhat calling to mind the Geometrid genus
Scotosia.
107. CapnopEs? CALIDA, 0. sp.
Bright clay-colour ; wings with two very irregular central brown-
bordered whitish dentate-sinuate lines from costa to inner margin,
each dens terminating in a white dot; outer border red-brown,
bounded internally by a pale zigzag line ; a nearly marginal series of
dots black internally and white externally: primaries with a snow-
white spot and one or two dots at the end of the cell ; secondaries
with a black-centred whitish spot at the end of the cell. Wings
below considerably paler and irrorated with smoky brown ; secondaries
whitish towards the base and inner border, the spots at the end of
the cell less strongly defined than above, the lines obsolete. Ex-
panse 1 inch 2 lines.
This species has rather long palpi for a Capnodes, but in general
form, style of coloration, and every thing else agrees with that
genus.
108. Uranta stoanus, Cramer, Pap. Exot. i. pl. 85. figs. E, F
(1779).
109. NycraLazmon acGistus, Fabricius, Mant. Ins. ii. p. 10.
n- 93.
110. CH#RODES TRANSTINCTA, var., Walker, Lep. Het. xx. p. 20
(1860).
111. DrepaNopEs PionariA, var., Walker, Lep. Het. xx. p. 71
(1860).
112. BoARMIA DELICATA, 0. sp.
Wings above chalky white ; the basal area and disk of primaries,
and the external half of secondaries, crossed by parallel undulated
testaceous bands; a well-defined zigzag stripe across the middle of
the disk ; two central angulated slender black lines, between which
is a black discocellular annulus on each wing; a marginal series of
black dots: body white; abdomen banded with testaceous. Under
surface white; wings with black discocellular spots, pale greyish-
brown discal belt, and black marginal dots ; primaries with the costa
black-spotted. Expanse 1 inch 6 lines.
113. AmpHiIDasys ARNOBIA, Cramer, Pap. Exot. iv. pl. 383. fig. I
(1782).
114. JopIs KINSTONENSIS, n. sp.
_ Wings above emerald-green, crossed by a wavy yellowish discal
line; fringes lemon-yellow, tipped with white; discocellular spots
1878. ] LEPIDOPTERA FROM JAMAICA. 491
black, encircled with orange: frons deep red, with white angles and
a central longitudinal green stripe; crest deep red, speckled with
white; antenne deep red, annulated with white ; back of head, collar,
and tegule emerald-green ; thorax and abdomen pale pinky brown
or flesh-colour. Wings below much paler than above, especially at
outer margin, sericeous; primaries with a small red apical spot ;
palpi above dark red tipped with white, below pearly white; pectus
snow-white, legs and venter pearly white. Expanse 1 inch | line.
115. Mecoceras siractariA, Walker, Lep. Het. xxii. p. 607
(1861).
116. Hyria VINACEA, n. sp.
Laky purple, with rather broad ochreous outer border, between
which and the general colour is a belt, reddish externally and slaty
grey internally; abdomen with ochreous margins: wings below
paler than above, the border cream-coloured ; body below cream-
coloured. Expanse 7 lines.
A pretty little species.
117. Erosta INCONGRUA, 0. sp.
Sordid white, reticulated and speckled with brown: primaries
with the base of costa cinereous; an irregularly elbowed testaceous
line across the basal third; an angulated dark brown discal line
(blood-red in certain lights), terminating on inner margin in a semi-
circular brown spot; four submarginal brown spots on the apical
half of outer border: secondaries with a black-edged sagittiform
streak through the cell; a white-edged elbowed blackish line across
the disk, bounded internally near the abdominal margin by an oblong
patch of red-brown; apical area clouded with reddish brown ;
border between the caudal processes greyish, black-edged inter-
nally ; two submarginal black dots on the median interspaces : body
white, with grey head and collar. Under surface sordid white,
speckled with black. Expanse 9 lines.
118, AcrpaLia umMBiLiIcaTA, Guénée, Phal.i. p. 504. n. 872
(1857).
119. Acrpauia apparitartiA, Walker, Lep. Het. xxii. p. 733.
n. 145 (1861).
120. Macaria acmaLiata?, Walker, Lep. Het. xxiii. p. 893.
n. 33 (1861). 4
There is a broken example of what appears to be a species of
Anisodes ; but I hope Mr. Bowrey will send us a good example some
day, and thus enable me to identify it with certainty.
121. Yestrperes? ANOMALA, N. sp.
Primaries above with the basal third and costa stramineous, the
rest of the wing silvery brown; basal area crossed by sinuated red-
32*
492 MR. A. G. BUTLER ON [May 7,
dish ochraceous lines, and the rest of the wing by sepia-brown
lines ; a discal series of white-edged spots, black on the discoidal
interspaces, otherwise brown ; median vein and bases of the branches
black-spotted : secondaries semitransparent sordid white, with opaque
outer border and grey submarginal stripe : body stramineous, streaked
with reddish ochraceous. Wings below cream-coloured, with greyish-
brown submarginal belt ; body white. Expanse 1 inch.
The only example obtained of this singular species is, unfortu-
nately, rather rubbed.
122. Scorosia caTocaLaRiA, Walker, Lep. Het. xxxv. p. 1689
(1866).
123. PrerocyPHA STELLATA, Guénée, Phal. ii. p. 443. n. 1627
(1857).
124, HypostpRA OCHREA, 0. sp.
Ochreous ; wings with a silvery gloss, excepting the outer border,
a discal belt, and a costal dash upon the primaries, which are of a
clearer yellow colour ; primaries with the lower half of the discal
belt edged externally with orange and plumbaginous; edge of
fringe orange: secondaries with two parallel zigzag purplish-brown
streaks across the abdominal half, the upper streak intersected by a
plumbaginous line; edge of fringe orange: head brown. Wings
below bright ochreous, with an oblique discal purplish streak ;
primaries with diffused silvery-whitish external area ; pectus whitish,
legs testaceous. Expanse 1 inch 3 lines.
I have to thank Mr. F. Moore for the identification of this genus,
the type of which is a Javan species, not in the Museum collec-
tion.
125. HyPENA LUNIFERA, 0. sp.
Primaries above lilacine brown, crossed by two slender blackish
zigzag lines, between which is an orange lunule, with its inferior
extremity white and edged with black, representing the reniform
spot; an orange dot at the costal extremity of the second black
line, and two subcostal dots of the same colour close to apex; a
marginal series of black dots: secondaries pale smoky brown, darker
towards the outer margin, crossed at the middle by an arched series
of black spots; a marginal series of black dots: body dark brown,
back of thorax slightly lilacine.- Primaries below greyish brown;
costal area speckled with whity brown; the discocellulars and an
arched discal series of spots dusky ; a marginal series of depressed
triangular spots: secondaries sordid white, speckled with dark brown ;
a black discocellular lunule ; two discal parallel zigzag dark brown
lines, the outer one ill-defined ; a marginal series of black spots, as
in primaries ; pectus smoky brown. Expanse 1 inch.
126. SyNGAMIA FLORELLA, Cramer, Pap, Exot. iv. pl. 348. fig. L
1782).
1878. ] LEPIDOPTERA FROM JAMAICA. 493
127. Dresmta orBa.is, Guénée, Delt. p. 192. n. 128 (1854).
128. Desm1a PROGNEALIS, Walker, Lep. Het. xvii. p. 346. n. 22
(1859).
129. SAMEA CHLOROPHASMA, 0. sp.
Wings hyaline, slightly yellowish, with stramineous borders ;
outer border spotted with rust-red ; primaries with three mustard-
yellow irregular lines across the basal area, followed by a straight
golden brown transverse line, then a stramineous belt margined with
brown, followed by a quadrate spot of the same character at the end
of the cell (its inner edge continued as a single line to internal
margin) ; an angulated brown discal line, terminating at inner margin
in a rusty reddish spot: secondaries with a black-edged stramineous
discocellular spot, from which a brown line runs obliquely to inner
margin ; an angulated brown discal line, terminating at anal angle in
a quadrate rusty reddish spot: thorax stramineous, streaked with
mustard-yellow; abdomen testaceous. Wings below altogether paler
than above, the markings less distinct; body below silvery white ;
palpi yellow; legs cream-coloured. Expanse 10 lines.
A delicate golden-looking little species.
130. AGATHODES MONSTRALIS, Guénée, Delt. p. 209. n. 165
(1854).
131. Hymenta perspecrattis, Hiibner, Ex. Schm. Zutr. fig.
101.
132. CoNCHYLODES DIPHTHERALIS, Hiibner, Ex. Schm. Zutr.
figs. 691, 692.
133. PHAKELLURA HYALINATA, Linn. Syst. Nat. ili. 2, p. 873
(1766).
134, PHAKELLURA NiTIDALIS, Cramer, Pap. Exot. iv. pl. 371.
fig. F (1782).
Mr. Bowrey says that he found the species of Phakellura feeding
upon the flowers of gourds.
135. MARGARONIA QUADRISTIGMALIS, Guénée, Delt. p. 304.
n. 319 (1854).
136. Borys BUTYROSA, 0. sp.
Wings dull lemon-yellow: primaries with the costa sericeous, a
feeble opaline tint beyond the middle, an angular pale browu line
across the basal third, and a sickle-shaped pale brown line across
the disk ; marginal line tawny : secondaries with opaline costal area ;
marginal line tawny: head and prothorax yellow; rest of the body
shining creamy white ; abdomen with a yellow dorsal line ;_project-
ing frontal tuft white; palpi brown above, white below. Body below
pearly white; legs pearly white, with the femora and the anterior
tibiee brown; wings below opaline. Expanse | inch 7 lines,
494 MR. BUTLER ON LEPIDOPTERA FROM JAMAICA. [May 7,
137. Borys LUCILLA, n. sp.
Bright golden yellow, with a very slight pinky spot; wings with
the two usual lines greyish ; primaries with the discoidal spots re-
presented by grey dots; secondaries with whitish abdominal area ;
head pearly white; abdomen pale yellow. Wings below paler than
above, opaline ; body pearly white. Expanse 9 lines.
138. Borys princrpatis, Lederer, Wien. ent. Monatschr. vii.
pl. 10. fig. 17 (1873). _
139. Borys eLaucusauis, Walker, Lep. Het. xvii. p. 576. n. 51
(1859).
140. Borys aGaveatts, Walker, Lep. Het. xviii. p. 574. n. 47
(1859).
141. Borys evryTauis, Walker, Lep. Het. xviii. p. 576. n. 50
(1859).
142. Borys campatis, Guénée, Delt. p. 344. n. 397 (1854).
143. Borys GasTRALIs, Guénée, Delt. p. 346. n. 400 (1854).
144. Borys eratatis, Walker, Lep. Het. xviii. p. 578. n. 54
(1859).
145. Borys oxtvi, n. sp.
Pale olive-brown, with bright bronze reflections; the usual lines
grey; orbicular and reniform spots outlined in grey; body greyish
brown. Wings beiow considerably paler, whitish towards the base,
sericeous ; inner line absent ; body white. Expanse 10 lines.
Nearly allied to B. plebeialis.
146. Borys wexcrrauis, Walker, Lep. Het. xviii. p. 574. n. 46
(1859).
147. Mecyna rREveRSALIs, Guénée, Delt. p. 409. n. 531 (1854),
148. ANERASTIA IGNOBILIS, 0. sp.
Primaries above pale pinky brownish or flesh-colour, the infero-
exterior quarter of the wing slaty grey; the median vein, its two
superior branches, the terminal portions of the subcostal branches,
the extreme ends of the remaining veins, and the centre of the
submedian vein silvery white; an abbreviated longitudinal apical
streak and a subcostal streak black; one or two minute black
dots on the median and submedian veins: secondaries hyaline,
iridescent, with dusky outer border: thorax red-brown, streaked
with pinky white; head and palpi white; abdomen grey. Primaries
and body below pale shining brown; secondaries as above. Ex-
panse 11 lines.
149, ARGYRIA VESTALIS, n. sp.
Silvery white: primaries with a small triangular spot at the
1878. | ON LAND-SHELLS FROM JAPAN AND BORNEO. 495
middle of costa, an oblique dash followed: by a triangular spot near
apex, and a submarginal stripe, with zigzag inner edge dark choco-
late-brown ; outer border and fringe pale brown, traversed by two
straight dusky lines; a minute brown spot at middle of inner
margin: base and sides of palpi chocolate-brown. Below silvery
white; the costal area of primaries pale brownish. Expanse 11
millimetres.
Like A. insons of Felder, but only half as large ; also allied to
A. chrysogyrans, Walker (MS. ’).
150. Cypos1A NOBILITELLA, Cramer, Pap. Exot. iii. pl. 264.
fig. G (1782).
151. Tinea?, sp.
This insect has lost all the fringes of its wings, so that I cannot
recognize its genus. The neuration of the secondaries seems to be
exactly that of Zinea ; but the head is too broad, and the palpi are
too long and robust ; in coloration and the general form of its wings
it is most like Cerostoma vittella.
152. PreROPHORUS ASPILODACTYLUS, Walker, Lep. Het. xxx.
p- 941. n. 43 (1864).
5. Descriptions of new Land-Shells from J apan and Borneo.
By Enear A. Smiru, F.ZS.
[Received April 15, 1878.]
Hexrx (CAMENA) LEWISII.
Testa dextrorsa, conoideo-globosa, subanguste umbilicata, oblique
striata, alba, epidermide tenui luteo-olivacea induta, fascia angusta
fusca ad peripheriam picta; anfract. 64, convewxiusculi, sensim
crescentes, ultimus rotundatus, antice breviter descendens ;
apertura obliqua, intus alba; perist. undique expansum, margine
columellari sordido-roseo tincto, superne late expanso et reflexo.
Diam. max. 35 mill., min. 29; alt. 24.
Hab. Japan (George Lewis).
Perhaps the most closely allied Japanese species to this one is
H. miranda, A. Adams. From it H. lewisii differs in having a
more conical spire, a narrower umbilicus, finer oblique striae, and no
spiral sculpture.
The oblique striz# at the suture are rather deeply incised and
more crowded than on the other parts of the whorls, many of them
extending only about a line from the suture and then gradually
fading away. The first four whorls differ from the last two in
being obliquely punctato-striate, instead of exhibiting an ordinary
striation. The brown band is situated a very little above the middle
of the body-whorl, and passing just above its suture, is visible on two
496 MR. E. A. SMITH ON LAND-SHELLS [May 7,
or three of the preceding yolutions. The umbilicus is very deep
and penetrable to the apex of the shell, but rather narrow, occupying
about one seventh of the smallest diameter.
The peristome rather broadly expanded and white, with the
exception of the columellar and basal margins, which are stained
with a dirty pinkish colour. Over the umbilicus it is broadly ex-
panded, reflexed, and joined to the upper margin by a thin callosity.
This species is quite distinct from H. myomphala, Martens, and, al-
Fig. 1.
/)
a ey
Helix (Camena) lewisii.
though agreeing in the style of coloration, differs in form and the
umbilicus.
The single example in the British Museum was presented by Mr.
George Lewis.
Hexrx (CAMENA) CONGENITA.
Testa suborbicularis; spira breviter conica, subaperte umbilicata,
tenuiuscula, pallide straminea, zona lata interrupta irregulari fusca,
vel strigis latis fuscis infra medium anfr. ultimi ornata et hic
illic supra strigis obliquis paucis indistinctis notata, interdum ad
peripheriam zona angusta fusca cincta, et umbilico aliquando fusco
tincto ; anfractus 53-6, convewiusculi, lineis incrementi elevatis,
filiformibus, obliquis, flecuosis et striis minutissimis spiralibus
inter lineas elevatas sculpti, ultimus angustiusculus, antice prope
labrum breviter subitoque descendens ; apertura subhorizontalis,
intus livida, fusco tincta; peristoma tenue, livido-fusco et albo
diverse pictum, undique reflecum et expansum.
Diam. max. 30 mill., min. 25, alt. 19; apert. 163 long., 14 lat.
Hab. Japan.
This species partakes in a great measure of the form of H. congener,
recently described by me in these ‘ Proceedings’ (1878, p. 105).
The spire, however, is a little more elevated, the last whorl a trifle
less robust, and the aperture narrower.
The sculpture of this species is much coarser, the lines of growth
being developed into thread-like liree, and the spiral striz invisible
to the naked eye, and more conspicuous between than upon the
1878. | FROM JAPAN AND BORNEO. 497
oblique thread-like lines of growth when viewed under a lens. The
descent of the body-whorl is very sudden and brief. The whorl,
owing to the expansion of the peritreme, has a constricted appear-
ance in that part.
The umbilicus is a little more open than in H. congener, and
consequently more perspective. The coloration of this species is not
easily definable. The ground-colour is pale straw, the oblique raised
liree being more opaque and yellower. The body-whorl below the
periphery is stained with brown, or looks as if it were scorched.
The colouring takes the form of a very broad interrupted transverse
band, or, in other words, of wide blotches or stripes. ‘The latter are
sometimes continued upon the upper half of the whorl, and are also
here and there faintly observable on the upper volutions. One
Helix (Camena) congenita.
specimen has a narrow peripherial brown band. The body-whorl is
stained with brown outside the tip; and this, owing to the thinness
of the shell, produces a brownish labrum, which in some places,
particularly at the extreme edge and in the columellar region, is
somewhat whitish. It is rather widely expanded at the base and
columella, and is a trifle reflexed everywhere. The aperture is very
transverse, in fact almost horizontal; it is of a livid white colour
within, exhibiting the brown marking of the exterior.
Subgen. Myxostoma, Troschel.
Testa plus minusve discoidea, late umbilicata ; apertura circularis ;
peristoma duplex, margine interiore superne plerumque leviter
sinuato, externo supra sinum alatim eapanso. Operculum corneum,
crassum, inferne paulo concavum, in medio nucleo prominulo, extus
marginibus anfractuum lamellosis.
This section of Cyclophorus agrees with Pterocyclos in the form
of the shell and its peristome, but differs in having a thick horny
operculum, with the margins of the whorls lamellated exteriorly,
that of Pterocyclos being calcareous and spirally laminated.
CycLopHorvus (MyxosToMA) BATHYRHAPHE.
‘Testa discoidea, apertissime umbilicata, spira parum elevata, sub
epidermide luteo-olivacea sordide vel ce@ruleo-albida; anfr. 53,
rotundati, sutura valde canaliculata sejuncti, ultimus sat magnus
498 MR. E. A. SMITH ON LAND-SHELLS [May 7,
superne ad canalem suturalem carina lamelliformi munitus, lineis
incrementi tenuibus leviter obliquis sculptus ; apertura oblique
circularis, intus ceruleo-alba; peristoma duplicatum, margine
interiort levi vix expanso, superne prope anfractum parum
profunde sinuato, externo anguste expanso, superne ala parva
leviter fornicata instructo. Operculum corneum, concentricum,
multispirale, utringue concavum, extus lamellis brevibus striatis
obtectum.
Diam. max. 31 mill., min. 22, alt. 15, apert. diam. intern. 9.
Hab. Borneo (G. B. Sowerby).
Shell discoidal, openly umbilicated, and clothed with a thin uni-
form yellowish-olivaceous epidermis. Beneath this it is of a dirty
or bluish-white tint. The spire is only slightly raised above the last
whorl. The whorls, about five in number, are convex and bordered
by a deep and broadish channel at the suture, the outer margin of
the canaliculation being produced into a thin lamellar ridge or keel ;
and outside this carina the whorls are a trifle depressed or excavated.
Fig. 3.
Cyclophorus (Myxostoma) bathyraphe.
The sculpture consists merely of the lines of growth, which are fine
and somewhat oblique. The aperture is slopingly circular, bluish
white, and has a double lip. The inner edge is thicker than the
outer, and scarcely at all everted. The sinus is distinct, moderately
deep, and situated at the termination of the sutural canaliculation.
The outer rim is somewhat expanded but not reflexed, narrowest on
the left side where it touches the whorl, and thin and sharp at the
extreme edge. Above it is expanded in the form of a small wing,
which slightly arches over the sinus and has a tendency to
a deflection in front.
This species in the nature of its operculum agrees with the genus
Myzxostoma, which was founded by Troschel (‘ Zeitschrift fiir Mala-
1878. | FROM JAPAN AND BORNEO. 499
kozoologie,’ 1847, p. 44) for the reception of the curious shell
figured by Martyn under the name of Lituus brevis.
It only differs from Myzostoma in having the inner lip of the
aperture sinuated above, and in the wing-like expansion of the outer
rim not leaning upon the penultimate whorl. The canaliculate suture,
I presume, is only a specific character.
In all respects agreeing with this species are two others, Cyclo-
stoma planorbulus, Lamarck, and Pterocyclos albersi, Pfr. The
former has had several localities quoted as its home, among which
are Senegal?, Philippine Islands, Bengal, Java, Borneo, Sumatra (in
Mus. Cuming), and Pulo Condore Island. Which of these is the
true habitat [ cannot say with certainty, nor am I aware that it has
ever been definitely settled; there is, however, some slight evi-
dence to show that the last locality is the correct one. We are also
in the same state of uncertainty respecting Pt. albersi. Pfeiffer
described the species not knowing its locality ; and Benson (‘Annals
of Nat. Hist.’ 1857, vol. xix. p. 208) is wrong in attributing a shell
found at Teria Ghat, Khasia hills, India, to this species ; for it was,
as shown by Hanley (‘Conchologia Indica,’ p. 56), only a variety
of Pt. parvus, Pearson.
Here, then, is a small group of four species, all having a Pterocy-
cloid expansion of the outer rim of the lip, and an operculum of
precisely the same structure. Three of them have channelled sutures
to the whorls, are of a depressed orbicular form, and have the inner
lip sinuated at the termination of the sutural channel. The fourth
(brevis) and the type of Myzostoma lacks the channelled suture,
and has only the slightest trace of a sinus in the lip—both of which
characters, especially the former, I consider more specific than
generic. From Cyclophorus with its simple concentric thin horny
operculum and simple lip to the aperture, the different operculum
and expansion of the lip of Myzostoma warrant, at all events, a
subgenerie separation.
There are two or three Burmese species (Cyclophorus pinnulifer,
Benson, C. calyx, Benson, and C. hispidulus, Blanford) which will
also conveniently range under this genus. They are depressed sub-
discoid shells with a double rim to the aperture, the outer lip with
a small superior wing-like expansion, and the operculum thick, horny,
with the outer margins of the whorls lamellated. For this group
Blanford proposed the name Scadrina (Journ. Asiatic Soc. Bengal,
1863, p. 322).
The museum is indebted to Mr. J. B. Sowerby for a single speci-
men of this interesting species.
500 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
6. On the Fruit-Pigeons of the Genus Ptilopus.
By D. G. Extiot, F.R.S.E. &c.
[Received April 16, 1878.]
(Plates XXXIII. & XXXIV.)
My material for the present paper has been most ample, and was
derived from the following sources :—the collection of the Paris Mu-
seum, rich in the types of the older authors, such as Hombron and
Jacquinot, Lesson, Quoy and Gaimard, Bonaparte, Temminck, Knip,
&e. ; the collection of the British Museum, containing the types of
Gray, Gould, Wallace, &c.; a series of special desiderata from the
Museum at Genoa, collected by D’Albertis, Bruijn, &c., and from
the Museum of Count Turati at Milan; a large number of various
species from different localities brought by Laglaize, and especially
by M. Raffray, now in the Paris Museum ; the species of Péilopus
procured by the ‘Challenger’ Expedition, also those from Duke-of-
York Island sent by Mr. Brown ; specimens in Messrs. Salvin and
Godman’s collection obtained by Mr. Layard in the Fiji Islands ;
and, lastly, the type specimen of Dr. Finsch’s recently described Pt.
ponapensis.
Probably no such complete series, illustrating the species of this
group from all localities, has ever before been at the disposal of any one
wishing to study these birds; and it is very safe to say that, unless I
had had before me the actual specimens described and figured by the
earlier writers, especially of those species belonging to the islands of
the Polynesian subregion, this paper could never have been written
without great danger of adding to the confusion already existing in
these perplexing birds. The descriptions in many instances are so
meagre and unsatisfactory, and the illustrations, in certain cases, so
totally unlike the originals, that it is utterly impossible to recognize
the species by them ; but having had the specimens, in the majority of
instances, before me, I have been enabled to ascertain without diffi-
culty to what species these doubtful birds (made such by their authors)
really belonged. Some of my conclusions may be found to be quite
at variance from those of other authors; but in all cases (unless
otherwise stated), let it be remembered, I have judged from the vast
material at my command, and, having brought to my task a mind to-
tally unprejudiced by any preconceived opinions, it will not perhaps be
deemed too great a request should I ask that, if any of the decisions
arrived at in this paper are rejected, they may be condemned only
after the examination of material at least approximating somewhat
to that which I have consulted, and such as will clearly show my
views to be erroneous.
Probably there are but few groups of birds so liable to lead a
naturalist astray as those species of Ptilopus inhabiting the Pacific
islands ; and, without an ample series before him, no one could
accurately diagnose the forms that really are distinct. In this spe-
cial division ot the genus the collection of the Paris Museum is rich,
1878. ] MR. D, G. ELLIOT ON THE GENUS PTILOPUS. 501
probably beyond all others, through the material brought to France
by the scientific expeditions known as the voyages of the ‘Astrolabe,’
* Coquille,’ ‘Vénus,’ ‘Zélé,’ ‘Pole Sud,’ &. Many types are among
these specimens ; and it has been by their assistance I have worked
out the synonymy of the species of the Polynesian subregion. All
the species figured by Bonaparte in his ‘ Iconographie des Pigeons’
are also in the Museum, and have been invaluable in enabling me to
arrange the synonymy and determine the species to which they be-
long, which has not been always possible from the work itself, some
curious and unaccountable errors having been committed by the
authors.
Man is fallible, and it is not to be supposed that in an extensive
review of some seventy species no errors have been committed ; but,
having endeavoured to verify every statement made in this paper
(taking nothing for granted), I may venture to hope that inaccuracies
have been reduced to a minimum.
The pleasing duty now alone remains for me to express my thanks
to those who have assisted me with material during the time I have’
been occupied with this memoir, without whose aid it certainly never
could have been properly completed. To my friend Prof. A. Milne-
Edwards, to whom I have already so often given my testimony as to
his willingness to accord every facility for the study and examination
of the unequalled collection of birds and quadrupeds under his charge,
I am again under obligations for every assistance rendered to me
during the months I have been engaged upon this group. Not only
was every specimen the Museum contained continually at my dis-
posal, but also many books from his own library were placed at my
service as, from time to time, I had need of them. It is only by
such enlightened assistance and cooperation from those who are at
the heads of great departments in European museums that any satis-
factory work can be accomplished by a naturalist who may not be
himself attached to such institutions. To Dr. Oustalet, Aide-Natura-
liste, I also desire to express my thanks for much assistance continu-
ally and freely given. To my friend Count T. Salvadori I would
acknowledge my indebtedness for a very valuable series of specimens
not otherwise obtainable, procured by the consent of the Marquis G.
Doria from the Museum at Genoa, among which were some types of
Count Salvadori’s lately described species. To Count H. Turati I
am also obliged for specimens from his magnificent museum. To
my friend Dr. Sclater I am indebted for opportunities of examining
the collection of the ‘Challenger’ Expedition, and also those sent to
him from different islands in the Pacific, containing various types of
the species described by him from time to time; and to Messrs. Sal-
vin and Godman for specimens from their collection obtained by Mr.
Layard, and which I brought to Paris for comparison with those in
the Museum collection. To Mr. Sharpe, who placed at my disposal
all the specimens of this genus in the British Museum, among which
were various types of much service in determining some doubtful
points, I am greatly obliged. And, lastly, to the Directors of
the Museum Godeffroy, who sent me the types of Ptilopus pona-
502 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
pensis described lately in this Journal by Dr. Finsch, I am much
indebted.
I commence with a brief review of the literature of the genus :—
1766. Linnzvus, ‘ Systema Nature.’
But one species was known to this author belonging to the pre-
sent genus, and was named by him Columba viridis. Species 1.
1788. GMELIN, ‘Systema Nature.’
In this work three species are given, two of which are described
the first time—C. viridis, C.jambu, and C. melanocephalus. Another
one is mentioned, the C. purpurata, based on Latham’s “ Purple-
crowned Pigeon.” This, however, describes no species known, but
is taken from portions of two different ones, and therefore cannot be
recognized. Species 3.
1813. Temminck, ‘ Histoire Naturelle des Pigeons et des Gal-
linacées.’
Two species are here described belonging to the genus Ptilopus,
viz. Pt. cinctus and Pt. superbus, placed in the genus Columba.
Species 5.
1820-39. Tremminck, ‘ Planches Coloriées.’
Eight species are described in this work belonging to this genus,
all under the term Columba, seven of which are given for the first
time, viz. Pt. magnificus, Pt. monachus, Pt. pulchellus, Pt: hyogaster,
Pt. porphyreus, Pt. perlatus, and Pt. nanus. The Pt. zanthogaster
of Wagl. is renamed diademata. Species 12.
1825. Swainson, ‘ Zoological Journal.’
Pt. regina, from Australia, described as Columba purpurata, var.
regina. Species 13.
1826. Lesson, ‘ Voyage de la Coquille.’
In the volume devoted to the zoology of this expedition one species
is described belonging to this genus, viz. Pt. taitensis as Columba
kurukuru, var. taitensis, from Otaheite. Species 14.
1827. Lesson, ‘ Bulletin des Sciences Naturelles.’
The Colombe amarante of the ‘ Voyage of the Coquille’ is here
named Columba puella. Species 15.
1827. Waacter, ‘Systema Avium.’
Pt. vanthogaster first described as Columba xanthogaster. Spe-
cies 16.
1829. Wacuer, in the‘ Isis.’
In his list of the Columbz several species of this genus are given.
Two are described for the first time, viz. P¢. pectoralis and Pt.
purpuratus, ex Tonga Tabou. Pt. taitensis, Less., is renamed
Columba oopa. Species 18.
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. ° = BOS
1830. Quoy & Garmarp, in the ‘ Voyage of the Astrolabe.’
Pt. gularis is here described and figured as Columba gularis.
Species 19.
1831. Lesson, ‘Traité d’Ornithologie.*
Under the term Ptilonopus eight species are given, one of them
for the first time, Pé. roseicapillus. Four of the remainder are
valid species, viz. melanocephalus, jambu, porphyreus, and taitensis.
Pt. cyanovirens is the female of superbus, Pt. virens is the female of
pectoralis, and Pt. purpuratus is Pt. ewingit. Species 20.
1838? Knipe anp Prevost, ‘ Histoire Naturelle Générale des
Pigeons.’
Ptilopus rivoli described as Columba rivoli. Species 21.
1840. Nrsoux, in the ‘ Revue Zoologique.’
Pt. dupetit-thouarsi first described as Columba dupetit-thouarsii.
Species 22.
1841. HomBron AND JACQuINoT, ‘Annales des Sciences Natu-
relles.’
Pt. luteovirens described in the genus Columba. Species 23.
1842. GouLp, in the ‘ Proceedings of the Zoological Society of
London.’
Pt. ewingii described. Species 24.
1844. Gray anp MircHe t, ‘ Genera of Birds.’
Twenty-five species are here recorded, of which three do not be-
long to this genus, viz. Pt. holosericeus, Temm., maculatus, Gmel.,
and pulcherrimus, Scop. Pt. purpuratus, Lath., is a made-up species,
Pt. virens=pectoralis, Wagl.; Pt. eyanovirens =superbus 9 ,Temm.;
Pt. purpureo-leucocephalus = dupetit-thouarsi, Neboux; Pt. swain-
sont = regina, Swains.; Pt. roseicollis = porphyreus, Temm.; Pt.
viridissimus, Temm. = fasciatus, Peale. The remaining species are
valid, among which P¢. oceipitalis is given for the first time and a
plate published. Species 25.
1848. Praxe, ‘ United-States Exploring Expedition, Ornithology.’
A list of the species belonging to this group, collected by this
Expedition, is given, four in all, among which three are described
for the first time, viz. Pt. coralensis, Pt. perousii, and Pt. fus-
ciatus ; the-fourth, Pt. furcatus, is the Pt. taitensis of Lesson.
Species 28.
1850. Gouxp, in ‘ Jardine’s Contributions to Ornithology.’
Pt. strophium described for the first time. Species 29.
1850. Gov xp, ‘ Proceedings of the Zoological Society of London.’
A race of Pt. magnificus is named Pt. assimilis. Species 30.
1853. Gray, ‘ Proceedings of the Zoological Society of London.’
Pt. chrysogaster described. Species 31.
504. ° = MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
1853, JacaurnoT AND PucHERAN, ‘ Voyage au Pole Sud’ (text).
Young of Pt. luteovirens described as Pt. felicia.
1854. Bonaparte, in the ‘ Comptes Rendus.’
In a paper contributed to this Journal, Bonaparte divided the
then known species of Ptilopus into various genera as follows :—
Leucotreron, Cyanotreron, Rhamphiculus, Omeotreron, Kurutreron,
Thouarsitreron, Lamprotreron, Iotreron, and Chryscena. These
are founded mainly upon the coloration of plumage (a dangerous
character (?) always), and in this case not particularly well selected.
In the case of Rhamphiculus and Omeotreron, the former is created
for the young (!), and the latter for the adult (!), of Ptilopus occipi-
talis ; and in Cyanotreron is placed the female (?) of Péilopus su-
perbus, while the male (!) of the same species is made the type of the
genus Lamprotreron. In Omeotreron, also, is placed the young of
Pt. luteovirens, while the adult is the type of the genus Chrysena!
It does not appear to me necessary, or even advisable, to adopt any
of this author’s genera for this group.
1854. Bonaparte, in the ‘Comptes Rendus.’
The young of Pé. occipitalisis called Pt. batilda, and Pt. apicalis
is first described. Species 32.
1855. DesMurs anp Prevost, ‘ Voyage de la Vénus.’
Pt. merciert and Pt. temminckii first described. Species 34.
1855. Bonaparte, in the ‘ Comptes Rendus.’
Pt. leclancheri described as Trerolema leclancheri. Species 35.
1856. Gray, ‘ List Birds British Museum.’
Among the members of this genus here recorded P¢. greyi is
named but not described. Species 36.
1856. Gouxn, ‘ Proceedings of the Zoological Society of London.’
Pt. eugenie from the Solomon Islands described. Species 37.
1857. Bonaparte, ‘Conspectus Generum Avium.’
In this publication this author made a subfamily of this group,
and divided it into the genera proposed by him in the ‘ Comptes
Rendus’ three years previously. Pt. canthogaster, Wagl., is called
diademata, after Temminck; Drepanoptila holosericea is placed
in the genus Lamprotreron with superba and porphyreus! In Pti-
lopus are purpuratus, Wagl., as porphyraceus, Forst., ex Tonga-
Tabou ; regina, Swains., as swainsoni, Gould; Pt. ewingii, Gould,
from Timor, is separated as flavicollis, after Gray ; Pt. coralen-
sis, Peale, redescribed as viridissimus; the species ex Vanikoro (!)
(errore) called purpuratus, Wagl. [this bird comes from Balaou,
not Vanikoro, and was afterwards named donapartei by Gray];
Pt. roseicapillus, mercieri, fasciatus, Peale (as clementineg, Hombr. &
_ Jacq.); perousit, Peale (as marie, Hombr. & Jacq.); pulchellus and
apicalis. Cyanotreron has cyanovirens (= superbus, Temm., 9)
and monachus, Temm. JTotreron contains viridis, Linn., melano-
cephalus, Gmel., rivoli, Prév. & Knip, iogaster, Temm., and nana,
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 505
Temm. (Kurutreron possesses oopa, Wagl. (=taitensis, Less.),
chrysogaster, Gray, and coralensis, Peale. Omeotreron bas batilda
(=oceipitalis, juv.), virens (=pectoralis, Wagl.), and felicia, Hombr.
and Jacq. (=/uteovirens, Hombr. & Jacq., juv.). In Ohrysena
is placed /uteovirens. Phapitreron is also included in this subfamily,
containing the Columba leucotis, Temm., and PA. amethystina,
Bon.; but these birds have but little in common with those of the
genus Ptilopus.
1858. Gravy, ‘ Proceedings of the Zoological Society of London.’
Five species are here described for the first time, viz. Pé. prasi-
norrhous, Pt. iozonus, Pt. aurantiifrons, Pt. coronulatus, aud Ft.
wallacet. Species 42.
1859. Gray, ‘ List of Birds of the Tropical Islands of the Pacific
Ocean.’
Nineteen species of the genus Péiopus are recorded in this catalogue
of the species of the Pacific islands contained in the British Museum.
One species is described as distinct—P¢. chalcurus. Species 43.
1860. Gray, ‘ Proceedings of the Zoological Society of London.’
Pt. temminckii, Des Murs & Prév., described as Pt. formosus.
1861. Schlegel, ‘ Nederlandsch Tijdschrift voor de Dierkunde.’
Pt. insolitus described. Species 44.
1862. Waxxacg, ‘ Proceedings of theZoological Society of London.’
Pt. humeralis first described. Species 45.
1863. Waxxace, ‘ Proceedings of the Zoological Society of London.’
Pt, albocinctus first named. Species 46.
1863. ScuieGeL, ‘ Nederlandsch Tijdschrift voor de Dierkunde.’
Pt. bernsteini first described. Species 47.
1868. HarrLtaus anp Frnscu, ‘ Proceedings of the Zoological
Society of London.’
Pt. pelewensis described. Species 48.
1867. Finsca anp Hartwavs, ‘Fauna Centralpolynesiens.’
This standard work on the Central Polynesian avifauna contains all
the species of this genus that were known at the time of its publi-
cation. Pt. purpuratus, Wagl., ex Tonga Tabou, is called Pt. por-
phyraceus, Forst.; Pt. purpuratus, Temm., is referred to C. forsteri,
Desm.; and the genus Chrysena is employed for Pé. luteovirens.
Drepanoptila holosericea is also retained in the genus Ptilopus. No
new species are described. Many of the species given had not been
seen by the authors.
1870. Gray, ‘ Hand-list of Birds.’
A list of the known species is here given, without any especial at-
tempt to arrange their synonymy being made. The birds figured
by Bonaparte on pl. xix. of his ‘Iconographie des Pigeons’ are
named Pt. bonapartii. Species 49.
Proc. Zoou. Soc.—1878, No. XXXIII. 33
506 MR. D. G. ELLIOT ON THE GENUs PTILOPUS. [May 7,
1871. Harrtaus anv Finscu, in the ‘ Proceedings of the Zoo-
logical Society of London.’
Pt. rarotongensis described. Species 50.
1871. Scuxecet, ‘ Nederlandsch Tijdschrift voor de Dierkunde.’
Three species are here described, viz. Pt. speciosus, Pt. ornatus,
and Pt. geelvinckianus, the last as Pt. viridis geelvinckianus. Spe-
cies 53.
1871. Goutp, in the ‘Proceedings of the Zoological Society of
London.’
Pt. vietor described. Species 54.
1873. ScHLeGeEL, ‘Muséum des Pays-Bas.’
A catalogue is here given of the species contained in the Leyden
Museum belonging to the genus Ptilopus. Some are included which
should be separated into different genera, such as Drepanoptila holo-
sericea and the members of Funingus. Three species are described
as new—Pt. neglectus, Pt. lettiensis (called Pt. cinctus lettiensis),
and Pt. jobiensis (called Pt. humeralis jobiensis). Species 57.
1873. ScxatTer, in the ‘ Proceedings of the Zoological Society of
London.’
Pt. bellus described. Species 58.
1874. Finscu, in the ‘ Proceedings of the Zoological Society of
London.’
Pt. huttoni described. Species 59.
1875. Satvaporzt, ‘ Annali del Museo Civico Naturale di Genova.’
In different lists of birds inhabiting New Guinea, six species of
Ptilopus are described, viz. Pt. gestroi, Pt. melanospilus, Pt. chry-
sorrhous, Pt. xanthorrhous, Pt. geminus, and Pt. trigeminus. Spe-
cies 65.
1875. Layarp, in the ‘ Proceedings of the Zoological Society of
London.’
Pt. viridis, as Chrysenas viridis, described. This is the Pé.
layardi of the present paper. The name viridis, having been given
by Linneeus to another species of Pti/opus, cannot stand. Species 66.
1876. Sarvaport, ‘ Annali del Museo Civico Naturale di Genova.’
Pt. zonurus described. Species 67.
1876. Ramsay, in the ‘ Proceedings of the Linnean Society of
New South Wales.’
Pt. corriei described. Species 68.
1876. Briaeemann, ‘ Abhandlungen des naturwissenschaftlichen
Vereins zu Bremen.’
Pt. fischeri described. Species 69.
1877. ScLaTER, in the ‘ Proceedings of the Zoological Society of
London.’
Pt. johannis described. Species 70.
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 507
1877. Finscu, in the ‘ Proceedings of the Zoological Society of
London.’
Pt. fasciatus, Peale, described as Pt. ponapensis.
1878. D. G. Exxiot, ‘Annals and Magazine of Natural History.’
Pt. pictiventris described. Species 71.
As the term “ purpuratus” has been applied to so many different
species by authors, it occurs to me that it may be useful if I give a
list of these, referring each one to the species for which it was in-
tended. For the reasons given in the article on Pt. purpuratus of
this paper, I accept for this term the species so designated in 1829
by Wagler, considering that the purpuratus of Gmelin should be re-
jected from the nomenclature of the group as not representing any
known species, but one formed from the characters possessed by two
distinct ones.
(1813) Columba purpurata (prem. var.), Temm. Pig. & Gall.
LOM s Ufa: s camacinaecsws cases = Pt. regina, Swains.
(1813) s ‘A Temm. Pig. & Gall. tom.i. p. 280 = Pt. ewingiz, Gould.
(1821-34) ,, :, Jard. & Selby, Dllustr. Zool.
VOU Tp Oe os dastes seecase soe c= = Pt. regina, Swains.
(1824) Bs zt, Temm. Plan. Col. livr. 43 ...... = Pt. xanthogaster, Wag).
(1826) Ptilonopus purpuratus, Steph. Cont. Shaw Gen.
Zool. vol. xiv. p. 277......s0000 = Pt. xanthogaster, Wag).
(1829) Columba purpurata, Wag). Isis, p. 742 ..-.........008 =Pt. purpuratus, ex Tonga.
(1835) Ptilinopus purpuratus, Selby, Nat. Libr. vol. v.
Bleicher taccncessnoseees =Pt. ewingii, Gould.
(1844) Ptilonopus purpuratus, Gray, List Birds Brit. Mus.
Ob et, (Cel Reem ape cene ebrCrice iar = Pt. ewingii, Gould.
(1855) Kurukuru purpuratus, Des Murs & Préy. Voy. Vé-
mus; ZOOL, P. 222 ‘sscecassscee- o> = Pt. taitensis, Less.
(1857) Pttlopus purpuratus, Bon. Consp. Gen. Ay. vol. ii.
BLO. Sak eee. Jodeci oe ors = Pt. greyi, Gray.
(1857) is, Bon. Iconogr. Pig. pl. xix.adult. =P#. bonapartei, Gray.
(1857) ” ” ” ” ” juy..
(1858) Ptilonopus purpuratus, Cass. U.S. Expl. Exped.
(1859)
Ornith. pl. 30. p. 269 ... =
Gray, List Birds of the
Uy SEN AREER Seoocod: ae
(1867) Pztilinopus purpuratus, Finsch & Hartl. Faun. Cen-
tralpolyn. p.122 .........
(1873) Ptilopus purpuratus, Schleg. Mus. Pays-B. p. 15 ...
” ”
GENERA.
Jie
=r.
purpuratus, Wagl.
. taitensis, Less,
taitensis, Less.
taitensis, Less.
taitensis, Less.
Quite a number of genera have been proposed for these birds ; but
only one is necessary. The term Pti/opus was given to them by
Swainson in 1825, its derivation being zr/Aov, rovs. Swainson wrote
it Ptilonopus, which was afterwards corrected by Strickland in 1841.
Various ways of spelling the name of this genus have been adopted,
viz. Ptilonopus, Ptilinopus, Ptinilopus!, all of which are incorrect,
and should not be used. P#ilopus is an artificial genus, by which I
mean to express that the birds placed together in it do not possess
any characters common to them all, either of form, structure, or
coloration, which will serve to distinguish them from Getter groups
33
508
=
MR. D. G. ELLIOT ON THE GENUS PTiLoPuS. [May 7,
Fig. 1.
6
A 5 \) 3
AY os V/
\ «(
woe
ijititllllle
Be
Ye
WS
SS
1. Pt. purpurutus.
2. Pt. chrysogaster.
3. Pt. mercieri.
4. Pt. dupetithouarsit.
5, Pt. jobiensts.
6. Pt. xanthorrhous. 11. Pé. letliensis.
7. Pc. insolitus. 12. Pt. ornatus.
8, Pé. taitensis, 13. Pt. pectoralis.
9. Pt. aurantiifrons. 14. Pé. bellus.
0. Pt. luteovirens. 15. Pt. victor.
1878. | MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 509
in the family. Some authors have separated them from the rest of
the Pigeons on account of the abruptly attenuated first primary.
This is utterly worthless, either as a generic or a specific character, as
among the species which are included in Péilopus every gradation
of the first primary is procurable, from the most abrupt attenuation
to the normal shape of feather.
T here give (fig. 1), in proof of this assertion, a series of drawings of
first primaries, together with the name of the species to which each
belongs, so that any one can verify it for himself; and I may
state that it would not have been at all difficult to have arranged a
more minute gradation from the species at my disposal if it had been
necessary ; but those given will be found, I think, amply sufficient
for the purpose.
Another so-called character, which has been adopted by some to
distinguish the members of the genus P¢i/opus, and one which is ex-
pressed by the term itself, is the tarsus feathered to the foot. This
proves to be of no more value, generically or specifically, than the
attenuated primary. Species with or without the last-named cha-
racter have the tarsus covered or partly bare, as do also those with the
graduated or normal shape of this feather. The drawings (pp. 510,
511) of the partly bare and partly covered tarsus, together with the
first primary of several species, will show clearly how very unsatis-
factory these characters are.
Relative lengths of wing and tail, as given by some writers for the
separation of some of these birds into different genera, also are not
constant, as the tail can be found shorter than the wing, or nearly equal
to it in length. This last is the main character given by Reichen-
bach for separating the birds placed by him in his genus Meqgalopre-
pia, as all the others are common to different species of Ptilopus.
He says:—‘‘ Schwanz fast so lang wie die Fliigel. Vorderschwinge
nicht verschmilert.” The species included by him, it is true, have
not this last character; but P¢. perlatus has the tail only a little
over half the length of the wing, while P¢. cinctus and Pt. gularis,
neither of which are included in Megaloprepia, have the tail nearly
as long as the wing, with the first primary abruptly attenuated. I
do not see that there is a single character remaining among those
given by Reichenbach which can distinguish the birds included in
his genus from those of the genus Péilopus.
For those who deem colour sufficient to establish a genus, it
might be supposed that among these gaily plumaged birds certain
patterns might be found which would answer for a generic cha-
racter. But this, again, fails; for the variety of the coloration is
so great that it would be necessary to divide the group into
numerous genera (as has, indeed, been attempted), and many of
the species would be left alone to represent a genus. I do not find
any thing better to designate the genus than the following formula :—
Ptilopus. Size moderate; wings usually reaching two thirds the
length of the tail; rectrices fourteen in number, excepting Pt. occi-
pitalis, which has apparently sixteen ; tarsi partly or wholly covered
with feathers ; colours of plumage usually bright, and often strongly
510 MR. D. G. ELLIOT ON THE GENUS PTILOpuS. [May 7,
opposed: distinguished from Treron (with which it constitutes the
subfamily T'reronine) by not possessing the scallop on the inner web
of the third primary.
Ptilopus is well placed between Teron and Carpophaga, leading
Fig. 3.
SS
Ptilopus fasciatus. Upolu, Samoa.
First primary abruptly attenuated ; tarsus partly bare.
up to the latter genus through the different races of Ptilopus
magnificus. It is the absence of the scallop on the third primary
which is the chief distinction between Péilopus and Treron. To any
one who will study this group with ample materials, such as has
fortunately been at my disposal, I think the difficulty, nay, impossi-
bility, of discovering any trenchant generic characters will be readily
acknowledged. Those who have had but a few species before them,
represented by two or three individuals of each, might easily imagine
that some one character of those which I have mentioned is really
1878. | MR, D. G. ELLIOT ON THE GENUS PTILOPUS. 511
of generic value ; and it is only when a great series has been obtained
that it is perceived how these gradually disappear or become evolved
into something quite different. It is from the lack of material to
show this fact that such a genus (among others) as Chrysena has
been accepted for Pt. victor and Pt. luteovirens. First established for
the latter species, chiefly on account of the lanceolate form of the
feathers, it was quickly perceived, on the discovery of P¢. victor,
that the genus could not rest on this character ; for the two species
Ptilopus geelvinckianus. Mafor.
First primary normal in shape; tarsus completely feathered.
Ptilopus coronulatus. Aru.
First primary abruptly attenuated ; tarsus completely feathered.
were generically essentially the same, and yet P¢. victor did not
possess lanceolate feathers. The partly covered tarsus is not suffi-
cient to separate them from Péilopus ; for, as I have shown, many
species of that genus have also the tarsus half nude. The normal
shape of the first primary also fails, as this is also possessed by other
species of Ptilopus ; and therefore no characters remain worthy of
separating these birds generically from the other members of the
group, those that exist being only of specific value. Bonaparte has
proposed most of the generic divisions ; and he founded them mainly
upon the colour of plumage, sometimes placing the two sexes, or
adult and young, of the same species in different genera (vide Pé.
superbus, Pt. luteovirens, &c.). I have not been able to discover
512 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
any reasons (after a careful examination of the species separated
generically by him) for accepting his divisions. The species named
by Schlegel P#. insolitus has been made the type of a genus,
(Edirhinus, by Cabanis and Reichenow. The same peculiarity ex-
hibited by this bird—a bony protuberance upon the base of the
culmen—is also found in certain members of the genus Carpophaga ;
and I see no more reason for separating P¢. insolitus from the
other members of this genus than exists for placing these Carpophage
in a new generic division. The term Megaloprepia, instituted by
Reichenbach for certain birds of this group, was so oddly selected, at
least in one instance (P¢. perlatus) that a doubt arises in my mind if
the author had ever had an opportunity of examining specimens.
No characters were given by him which were not also to be found in
other species of Ptilopus; and although the term has been employed
by some authors, it seems to me to be entirely unnecessary ; for it is
impossible to designate in a large series of Ptz/opi where the genus (?)
should commence, as the species graduate into one another from the
smallest to the largest.
Prof. Garrod, in the ‘ Proceedings’ of this Society for 1874, in
his paper on the “Anatomy of the Columbe,” states that the
members of the genus Ptilopus possess a form of gizzard such as is
not found in any other bird, and gives figures exhibiting its peculi-
arities as compared with Treron. If this is ascertained to be a con-
stant formation in all the species of Péilopus, it may well be deemed
of sufficient importance to constitute a generic character ; but, unfor-
tunately, as yet, it cannot be said to be proven ; for out of the seventy-
one species considered in this memoir as belonging to Péilopus, Prof.
Garrod has only been able to dissect three, a number too insignifi-
cant for it to be decided that all the members of the genus have the
same formation. The Carpophage do not possess a similar gizzard ;
yet certain species of Pétilopus, viz. Pt. magnificus with its races assi-
milis and puella, and Pé. gularis, in their outward form resemble
certain species of Carpophaga ; and it is possible that in some of these
at least a modified form of gizzard may be found toexist. It will be
interesting to learn if such be the case, or if really the species of
Ptilopus are separated from all other Pigeons by an abrupt alteration
in the gizzard’s shape, one not approached by that of any other bird.
A large number of species must be first examined before this can be
definitively ascertained.
The following are the principal generic terms which have been
proposed for these birds.
Type.
(1825) Ptilonopus, err. Swains. Zool. Journ. yol.i. p.473 Pt. regine.
(1841) Pélopus, corr. Strickl. Ann. Nat. Hist. vii. p. 386.
(1854) Leucotreron, Bon. Compt. Rend. vol. xxxix. p.876 Pt. cinctus.
(1854) Cyanotreron, Bon. tom. cit. p. 878 .........0eeeeeeee Pt. monachus.
(1854) Rhamphiculus, Bon. ibid. .........6..620cceeceeeeeeeees Pt. occipitalis.
(1854) Omeotreron, Bon. ibid. ..........ccseceeceeseceseeeeeets Pt. occipitalis,
(1854) 2Rar-verero PBO0 STNG». siae aniclesacniestaed sainnascosecinne Pt. purpuratus.
(1854) Chery sana. Bont a DiGies. s.as; sacuncucpseenscas<arenots Pt. luteovirens.
(1854) Thouarsitreron, Bon. tom. cit. p. 876 .............4. Pt.dupetit-thouarsii.
(1854) Lamprotreron, Bon. ibid. ...........cceccceeeeeeen ences Pt, superba.
1878. | MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 513
T
(1854) Zotreron, Bon. tom. cit. p. 878 ......csecseseesenee eee Pe ngaaeen
(1857) Sylphitreron, Verr. teste Bon. Consp. ii. p. 40 ... Pt. perlatus.
(18 oloprepia, Reich, Tauben ............00.s0seeeeee Pt. magnificus.
6) Gdirhinus, Cab. & Reich. J. f. Orn. p. 326 ...... Pt. insolitus.
The Drepanoptila holosericea, from New Caledonia, has usually
been placed in the genus Ptilopus ; but it does not appear to me that
this is its proper position. With a colouring, it is true, approaching
something the style of these birds, it differs from them in form, and
especially in the shape of the wing-feathers. These (sce fig. 6) are
very peculiar, and, in my opinion, at once remove this species from
Wing of Drepanoptila holosericea.
among the present group of birds into a separate genus. The
drawing will show, better than any description can’ explain, the
unusual formation of the feathers in question’.
The members of the genus Funingus, beyond the feathered tarsus,
1 In the Rev. et Mag. de Zool. 1862, p. 135, pl. 8, MM. J. Verreaux and O.
Des Murs conferred another generic name upon this bird, viz., Calyptomenenas,
and gave a drawing of both it and a portion of the wing. Neither of these, how-
ever, are correctly done, especially that of the wing, which shows only the ends
of the primaries. The illustration here given more faithfully exhibits the
peculiar shape of these feathers.
514 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
have but little in common with the members of Ptilopus, the
peculiarly notched first primary and bare face easily distinguishing
them.
GEOGRAPHICAL DisTrRIBUTION.
Probably no group in ornithology exhibits to a greater degree the
effects upon the coloration of plumage arising from the physical
causes incidental to an insular life than the one here considered as
composing the genus Ptilopus. Some of these birds that have
evidently had a common origin exhibit greater or less variations
from each other, according to the position of their various habitats,
sometimes of sufficient importance to constitute their possessors dis-
tinct species, at others only of that trivial kind that would at
best but cause them to rank merely as varieties, the lapse of time
during which the individuals have dwelt in their separate localities
not having been sufficiently great, or the physical causes of climate,
food, and soil not powerful enough to modify essentially their
appearance from the typical form. It is therefore not surprising
that we find some species inhabiting one island only, others dwelling
upon several, though perhaps separated by miles of sea, while again
in other localities uncertain forms are observed not changed suffi-
ciently to authorize the bestowal upon them of a separate distinctive
rank, but yet differing enough to show that a departure from the
type and towards a distinctive independent form has been com-
menced, and will continue (unless the race should become exter-
minated) until the variety eventually blooms into a separate species
possessing characters not found elsewhere. It is not to be supposed
that the continents of which the islands of the Pacific and those
of the Eastern Archipelago are the sole remains, were broken up
simultaneously or always suddenly throughout their length and
breadth ; but more probably the casualty happened at various periods
and sometimes gradually. Therefore we may not be surprised at
the apparently strange phenomenon that one species should inhabit
various islands, between which are others containing totally distiact
forms of the same genus. This may have been brought about
in two probable ways. A species may have been widely dispersed
over the continent; and when portions of this had disappeared
beneath the waves, the fragments that remained above water at
the outset were all occupied by the same species: but physical
effects at intermediate points were of a different character from
those at the extremes; and in course of time the birds dwelling on
the intervening islands departed entirely from their types, while
those most widely separated retained their original characters. Or
it may, on the other hand, have been that on the breaking-up of the
continent a district inhabited by a strictly local species (but one
surrounded by a more widely disseminated and distinct species) had
not been entirely submerged ; and this, all other circumstances being
equal, would explain the fact that a distinct form should intrude itself
on an island lying between others inhabited by a different one, the
species with the greater range having been preserved at the extremes
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 515
of its habitat, which also had become islands. In considering the
peculiarities of the geographical distribution of these birds, the
great physical changes that have taken place in that portion of the
globe over which they are dispersed must always be borne in mind.
The members of the genus Ptilopus are found in only two of the
zoogeographical regions of the earth, viz. the Oriental and Aus-
tralian. Of these the latter contains by far the great majority of
the species ; and, to judge from the material at present available, the
Papuan group of the Austro-Malayan Subregion is apparently the
head quarters of the genus, The Polynesian Subregion also
possesses many species ; but they are altogether of a different style
of plumage, and constitute almost a division by themselves.
I commence the review of this portion of my subject with the
most easterly group of the Pacific islands (in the Polynesian Sub-
region of the Australian Region) in which any members of this genus
are found, viz. the Marquesas. But two species are met with
among these islands—the Pt. dupetit-thouarsit, on Christine Island,
and the Pé. mercieri, from Nuka-hiva. Both of these are very
distinct from all others known. I have described Pé. pictiventris as
from Nuka-hiva of the Marquesas ; but, for the reasons I give under
the head of this species, I believe that locality to be an error, and that
the bird in question is a native of the Samoan Islands only. To the
south-west of the Marquesas lies the Pomatu archipelago, which is
as yet hardly known in an ornithological sense. But one species of
Ptilopus has been recorded from there, Pt. coralensis, found on
Carlshoff Island. South of the Pomatu group lies the island of
Rapa. Here has been procured probably the most important
member of this genus, Pt. huttoni. To the west of Pomatu
are the Society Islands, upon only two of which have any species
of Ptilopus been procured, viz. P. chrysogaster, upon Huahaine, and
Pt. taitensis in Otaheite. To the south-west of this last is Cook’s
Archipelago, where Pt. chalcurus has been obtained on Harvey
Island. It is still doubtful if this bird is specifically distinct from
Pt. coralensis of the Pomatu group. If the identity of those
two could be established, it would be an interesting and important
fact, since the two clusters of islands are not only separated by miles
of ocean, but several distinct forms of Ptilopus are found upon
the intervening archipelagos, none of which, so far as we know,
are particularly related to Pt. chaleurus. Upon Rarotonga island
has been procured P#. rarotongensis; and these two species are
all that have as yet been found in this group. Savage Island, lying
between Cook’s Archipelago and the Friendly group, contains three
species—Pt. purpuratus, Pt. pictiventris and Pt. rarotongensis.
Passing on to the westward we reach the Tonga Islands, among
which five species of Ptilopus dwell, a number exceeded in only one
of the Pacific groups, viz. the Fijis. On Tonga-Tabou all the
species occur—P¢. chrysogaster, Pt. apicalis, Pt. pictiventris, Pt.
purpuratus, and Pt. perousii. Harpai contains P¢. perousii and
Pi. purpuratus ; and this last has also been procured at Eua and
Ninafou, the last the most northern of them all. North of the
516 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [| May 7,
Friendly Islands lie the Samoan or Navigators’ Islands. These
contain also five species, two of which are not met with in the Tonga
Group. Opalu has four—Pt. perousii, Pt. luteovirens. Pt. picti-
ventris, and Pt. fasciatus, this last being also found on Wallis Island,
while Pt. purpuratus is a native of Fulima, lying to the south
of Wallis. We now come to the Fijis, which possess among the
different islands nine species of Ptilopus. Only one island, however,
contains more than two species, viz. Viti- Levu, which has three—P¢?.
perousii, Pt. luteovirens, and Pt. pictiventris. Schlegel suggests
Viti as the probable locality also of the bird he has called P¢.
neglectus ; but this is very uncertain, and there is no evidence to
support the supposition. Balaou has two species—P¢#. luteo-
virens and Pt. bonapartei. Two species are found upon five islands
which possess no others of this genus. These are Pé. perousii and
Pt. purpuratus, met with at Waikia, Vanua Levu, Mango, Mokani,
and Loma Loma. Bua, Lanthala and Ngamia only have P¢. victor.
Taviuni has Pé. victor and Pt. purpuratus; while Kandavu con-
tains Pt. perousii and Pt. layardi. The New Hebrides possess-
but two species—P¢. greyi on Erromango Island, and Pé. corriet
on Malacolo, On Lifu, of the Loyalty group, Pt. greyi is
found, as well as on the Isle of Pines and New Caledonia to the
west, this last being also recorded by Schlegel as the habitat of Pé.
insolitus—which, however, is probably an error, Passing now to
the northward, Pt. greyi is again met with on Vanikoro of the
Santa-Cruz archipelago; and in the Caroline archipelago, on the
island of Ponapé in the Seniavin group, Pt. fasciatus has been
procured. To the north-west, the furthest north perhaps that any
species of this genus is found, lie the Marianne or Ladrone Islands.
Here is met with the very distinct P¢. roseicapillus, the only repre-
sentative of these birds in this group. To the west are the Pelew
Islands, whence comes Pé. pelewensis; but from which particular
island is not stated by its describers.
We now reach the Austro-Malayan Subregion; and the islands
which come first in this review (proceeding from the east towards
the west) are those contained in the Papuan group, of which New
Guinea is the centre and most important. Beginning with the most
eastern division of this group, we have first the Solomon Islands,
upon one of which, not as yet known, the Pt. eugenie was obtained.
To the westward of this, in the Louisiade archipelago, on the Island
of Duchateau, the type of Pt. strophium was procured. From
New Ireland, to the north-west, come P¢. insolitus and Pt. puella;
while between this last-named island and New Britain lies Duke-of-
York Island, the home of the rare and little-known Pé. rivolii and
also of the Pé. znsolitus mentioned above. From New Hanover Pé.
superbus has been received. To the north-west of this section, on
Wild and D’Entrecasteaux Islands of the Admiralty group, the
beautiful Pt. yohannis was obtained by the naturalists of the ‘ Chal-
lenger’ Expedition. Coming now to New Guinea, the islands in the
great Bay of Geelvinck claim our attention. Krudu, the most eastern
of these, possesses one species, P#. geminus. Next, on the large
1878.] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 517
island of Jobie, eight species of this genus dwell, viz. Pt. superbus,
Pt. coronulatus, Pt. geminus, Pt. aurantiifrons, Pt. perlatus, Pt.
strophium, Pt. puella, and Pt. jobiensis. The small island of
Ansus, to the south of Jobie, contains also one species, Pt. geminus,
being the only one out of those found upon its great neighbour.
Meosnoum or Misnomin, to the west of Jobie, also contains one
species found on that island, Pt. strophium. On the island of
Korido, or Sook, the most northern of those in this bay, two species
have been procured, Pt. pectoralis and Pt. speciosus. Mansinam,
the last island in this bay from which any species of Ptilopus is
known to come, contains one species, the P¢. humeralis. It will thus
be seen that, so far as we are able to determine from our mate-
rial, in the Bay of Geelvinck, Jobie is the central or starting-
point of the members of this genus. The two small islands lying
near are only detached portions of Jobie, each containing one species
found on the last, but none peculiar to themselves. On the other
hand the islands of Korido and Mafor only have one species in
common with Jobie, viz. P¢. superbus, found on Korido, while on
Mafor Pt. pectoralis and Pt. geelvinkianus are met with, the
last-named being also a native of Meosnoum. The relationship of
these islands, so far as regards this genus, are nearer to the western
shores of the bay than to Jobie. Turning now to the mainland, at
Andai and Dorey five species are found— Pt. pulchellus, Pt. pecto-
ralis, Pt. puella, Pt. geminus, and Pt. perlatus ; while on Mount
Arfak Pt. superbus, Pt. bellus, Pt. iozonus, Pt. aurantiifrons, and
Pt. ornatus have been obtained. At Amberbaki, on the north
coast, five species have been procured, viz. Pt. pulchellus, Pt. super-
bus, Pt. perlatus, Pt. pectoralis, and Pt. bellus; while at Sorong
Pt. trigeminus, Pt. humeralis, Pt. coronulatus, and Pt. prasi-
norrhous have been met with. At Triton Bay Pt. nanus has been
obtained; and at Mount Epa, in the south of New Guinea, the
small race of Pt. magnificus, called puella, is found. These are all
that are as yet known to come from the mainland of New Guinea ;
but doubtless others of the described species, as well as those not
yet discovered, remain to be obtained among the dense forests of
this singular country. Passing now to the western islands of the
Papuan group, the most northerly is Waigiou, which contains five
species—L’t. superbus, Pt. pulchellus, Pt. prasinorrhous, Pt. puella,
and Pé. pectoralis, this last being also found upon the near-lying
small islands of Gagie and Giebeh, while P¢. puella is also met
with on Ghémien. Batanta has four, viz. Pt. aurantiifrons, Pt.
superbus, Pt. pectoralis, and Pt. prasinorrhous ; while Salwatty pos-
sesses all these (excepting P¢. pectoralis) and also Pt. coronulatus,
Pt. trigeminus, Pt. humeralis, Pt. pulchellus, Pt. perlatus, and Pt.
puella. Mysol, to the south-west of Salwatty, contains seven
species, viz. Pt. prasinorrhous, Pt. superbus, Pt. aurantufrons, Pt.
pulchellus, Pt. puella, Pt. pectoralis, and Pt. nanus. All of these
save the last two are found upon Salwatty. Far to the south of this
last-named island we reach the most southern ones of the group, viz.
Aron, with seven species (Pt. coronulatus, Pt. superbus, Pt. iozonus,
518 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Pt. wallacei. Pt. aurantiifrons, Pt. zonurus, and Pt. perlatus), and
Yule Island, lying to the south-east of the mouth of Fly River, in
the Papuan Gulf, and close to the coast, with two (Pt. coronu-
latus and Pt. gestroi, the last intermediate between Pf. ornatus
and Pt. perlatus). The next group of the Austro-Malayan Islands
that claims our attention is the Moluccas ; and, commencing with the
furthest north, Sanghir is the first to be considered: here is found
a race of Pt. melanocephalus from Java, which has become pos-
sessed of sufficient characteristics to separate it from the type, and
is known as Pé. xanthorrhous. Gilolo and its small neighbours,
Ternate and Batchian, contain four species, found on each, viz. Pé.
togaster, Pt. superbus, Pt. bernsteini, and Pt. monachus. Tidore,
on the west, has three—P¢. monachus, Pt. iogaster, and Pt. prasinor-
rhous; Morty two, Pt. monachus and Pt. iogaster; Obi one, Pé.
bernsteini ; Dammor and Kaisa, each one, Pé. monachus; and Weda
and Dodingo also one, Pt. cogaster. Ceram contains three species—
Pt. chrysorrhous, Pt. superbus, and Pt. viridis, this last being also
found upon the small island of Monavolka, which, together with
Matabello, also possesses Pt. prasinorrhous. Banda, to the south
of Ceram, has Pt. xanthogaster, which is also found upon the
islands of Khoor and Ki, these last having in addition Pt. pra-
sinorrhous. Goram, lying to the east, has two species, the one
last named and P¢. viridis. Bouru has three species—P¢. super-
bus, Pt. viridis, and Pt. prasinorrhous; while Amboyna has also
three species—P¢. superbus, Pt. viridis, and Pt. prasinorrhous ; and
Harouka, east of Amboyna, contains Pt. viridis. Of the Timor
group, I commence with Timor, which possesses two species, P.
ewingtt and Pé. cinctus, this last being also found upon the small
island of Wetta, to the north of Timor; while Lettie, to the
north-east, possesses Pt. lettiensis (a modification of Pt. cinctus)
and Pt. xanthogaster. Flores has three species—P¢. ewingii, Pt.
albocinctus, and Pt. melanocephalus, this last being also found
in Sambawa and Lombock. Celebes has four species, three of
which are peculiar to it, viz. Pt, melanospilus, Pt. temminckii, Pt.
gularis, and Pt. fischeri; while Sula contains Pt. melanospilus and
Pt. chrysorrhous. Australia, the last division of this region, has six
species—L¢. regina and Pt. magnificus in New South Wales, in the
district between the river Hunter and Moreton Bay, Pt. superbus
at Cape York and on the Booby Islands, Pt. assimilis and Pt. mag-
nificus in North-east Queensland at Rockingham Bay, Pt. assimilis
being also found together with P¢. puella at Cape York, and Pt.
ewingii on the Coburg Peninsula near the coast—the distribution
of the three forms of P¢. magnificus being decidedly against the view
that they represent distinct species.
The other zoogeographical region in which these birds are found
is the Oriental; and we here find them restricted to the subregion
containing the Indo-Malay islands. Of these I commence with the
Philippine group, in only one island of which as yet has any species
been found, viz. Luzon, containing three (Pt. occipitalis, Pt. gambu,
and Pt. leclancheri). The great island of Borneo, and also Sumatra,
1878.]
the birds of this genus.
The following key will show the chief differences existing between
I recognize 71 species, and have divided’
them into two great groups, characterized by having the breast-
feathers bifurcate or not bifurcate.
MR. D. G. ELLIOT ON THE GENUS PTILOPUS.
have but one species, Pt. jambu ; while Java, the last locality in
which Péilopus is found, contains two, Pt. melanocephalus and Pt.
porphyreus.
KEY TO THE SPECIES.
A. Breast-feathers bifurcate.
a. Front and top of head purple or deep rose,
margined with yellow or pale green.
a', Tail with a subterminal grey or ash-
coloured band.
a". Abdomen deep rose-colour; under
tail-coyerts yellow, tipped with
rose ; breast bright green............
6", Abdomen and breast washed with
2. Secondaries and tail copper-colour;
sinall dark purple spot on lower
part of abdomen.
a. Front and crown purple .........
8. Front and crown greyish purple
OLaAMemhysh.csrcesc-tecccearecetees
3. Middle of abdomen rufous, upper
edge dark purple, scapulars tipped
NWTUHMLTLAGEZy so Tee eteee ol seNNe
4, Feathers on upper part of breast
with transverse violet spots near
the tips; no spot on lower part of
breast ; abdomen orange; crissum
yellow; under tail-coverts white
at base, then yellow, with rose-
WIGS DS oseeeeysteae oe toe sacceesore
5. Throat white or pale yellow ; breast
pale green, feathers tipped with
light grey ; abdomen orange, with
a lilac-red spot in the centre .....
6. Throat light yellow; breast pale
green, feathers tipped with light
grey ; abdomen orange, witha pale
1.
2.
10.
Pt. grey?, p. 528.
Pt. neglectus, p. 524.
. Pt, bonapartei, p. 524.
. Pt. purpuratus, p. 5265.
. Pt. apicalis, p. 527.
. Pt. coralensis, p. 528.
. Pé. chaleurus, p. 529.
. Pt. pictiventris, p- 530.
. Pt. pelewensis, p. 531.
Pt. regina, p. 581.
lilac band upon the upper part ... 11. P¥, ewingii, p. 533.
519
520 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
7. Abdomen and under tail-coverts
bright yellow; on lower part of
breast a large spot of dark purplish
nO dbs eae asaanlappocmegonaccencansdacawces 12. Pt. rarotongensis, p. 534.
8. Throat, cheeks, and breast yellowish
green; abdomen clear green, with
a purplish black spot in the centre 13. P#. fasciatus, p. 535.
6". Conspicuous deep-rose spot at base
of mandible.
1. Abdomen bright uniform yellowish
green, without spot ............06 14. Pt. mercieri, p. 536.
2. Breast with a deep purple spot on
lower part; abdomen orange;
under tail-coverts bright yellow,
Ups OLATIOO ..0--cataecsagse esac aneee 15. Pt. roseicapillus, 537.
3. Chin pale purplishred; dark purplish
violet spot on breast, with the base
of the feathers dark olive-green ;
under tail-coverts purplish violet-
red, basal portion yellow; purplish
violet of front and crown sur-
rounded by green...........sseeeseee 16. Pt. huttoni, p, 588.
4. Front and top of head greyish or purplish
white, margined with yellow.
a’. Abdomen deep rose; crissum and under
tail-coverts deep yellow ...............0+5 17. Pt. dupetithouarsi, p. 539.
'. Abdomen and under tail-coverts orange-
ry ell OW viteapndoee semen tees eaaascce restate eee 18. Pt. xanthogaster, p. 540.
c' Abdomen and under tail-coverts bright
VCO War tawe sia cnc cep oanea bess acacia 19. Pt. chrysogaster, p. 541.
d', Underparts pale ash-green ; centre of ab-
domen and crissum pale yellow; under
tail-coverts bright yellow ..............- 20. Pt. taitensis, p. 541.
c. Top of head and occiput bright purple;
sides of head light green; nape and sides
of neckrufous-orange; middle of abdomen
and crissum white: flanks and thighs green.
a, Feathers of breast purple at base, broadly
tipped with ashy grey, a broad bluish
black band acrossthe lower partof breast 21. Pt. superbus, p. 542.
6’. Lower part of breast light purple, gra-
duating into bluish black ........000..4 22. Pt. temmincki, p. 544.
d. Front, crown, band across the back, and
under tail-coverts dark purplish red ; fea-
thers of breast purple at base, white at tip;
beneath these an orange band ............ 23. Pt. perousei, p. 545.
e. Front pink, with posteriorly a narrow band
of purple, followed by a broader one of
yellow ; a purple spot on fore part of ab-
domen; middle of abdomen and under
tail-coverts yellow ....c0.es.s.seessssecevsees 24, Pt. coronulatus, p. 546.
f. Front pinkish white, surrounded by a
purple band succeeded by one of yellow ;
purple spot on abdomen, surrounded by
HasTOn=VellOw: \ozssctecsaweenissdgnonseseeastes 25. Pd. geminus, p. 546.
g. Bronut greyish: White. ..cccienwenneawonesatennss 26. Pt. trigeminus, p. 547.
B. Breast-feathers not bifurcate.
a. Middle of abdomen deep orange; vent and
under tail-coverts white varied with yel-
low; tertials violet grey in the centre.
a’. Shoulders greyish violet, mixed with
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS.
green ; tail green, with an apical grey
band above, white beneath............... 27.
8' Shoulders violet-grey, bordered beneath
by deep purple; tail green, with a sub-
terminal grey band on lower surface only 28.
'. Shoulders violet-grey ; tail green above,
beneath black, with an apical white band 29.
d'. Bony protuberance on _ forehead ;
shoulders and patch on back light grey ;
tail bright green, apical third ashy grey 30.
6, Bluish grey band on each side of breast ;
purple spot on centre of abdomen; wing-
coverts, scapulars, and tertials tipped
DEMUVOLLOWHesdeensesiccatsessaecasemsascemcctes 31.
¢. Front, crownand lores, moustaches, and spot
on the breast greyish blue ; superciliary
stripe, chin, throat, lower parts of abdo-
men, and under tail-coverts lemon-yellow 32.
d. Front and crown bright carmine; sides of
head and neck and breast light grey;
chin and throat white ; large purplish red
spot on lower part of breast; abdomen
and under tail-coverts orange .......0.... 3d.
e. Head and neck light grey; occiput and
back of neck black ; throat yellow.
', Crissum and short under tail-coverts yel-
low, long ones carmine ...... ces Caen 34,
4', Throat-mark narrow; crissum yellow,
washed with orange .........ssssseeeeeee 35.
. Throat-mark narrow ; black bar on nape
small; crissum orange-yellow ......... 36.
d'. Lower parts of abdomen and crissum rich
OLANES Fecsveccecessecsnscnssnsscoesnerssneces 37.
f. Head and neck rose- -colour, graduating into
carmine on the breast, beneath which is
a broad white band, followed by another
fof blackish) preen:. ..5... .c.cleseetecavenaleeee 38.
g. Throat, crown, and sides of face in front
scarlet, chin and throat black; middle of
breast rose-colour; under tail-coverts
STRGIETOUAT st See pe a MBB SSeS dar cds ee aS ec 39.
ht, Throat white or pale grey.
a, Scapulars and tertials grey, margined
with green.
a". Top of head bright carmine; mantle,
neck, and breast bluish grey ; lower
part of breast crossed by a white
band, succeeded by a broad one of
deep orange; shoulders deep orange 40.
4". Forehead deep orange; crown and
occiput yellowish green; back and
upper part of breast plumbeous ;
underparts yellowish green......-..... 41.
. Shoulders dark purplish red ; lower
parts of throat dark yellow; breast
dark ochre; tail dark green, with a
light-yellow apical band.
1. Top of head and sides of face pur-
Phish! red!" oo0. lscukes wseeek ee eee 42,
2. Top of head and sides of face olive-
BOUL WAM. Ay dw cee gevaceGolte mance seen 43.
Proc. Zoou. Soc.—1878, No. XXXIV.
521
Pt. iozonus, p. 547.
Pt. humeralis, p. 548.
Pt. jobiensis, p, 548.
Pt. insolitus, p. 549.
Pt. nanus, p. 549.
Pt. monachus, p. 550.
Pt, pulchellus, p. 551.
Pt. melanocephalus, p. 551.
Pt. melanospilus, p. 552.
Pt. chrysorrhous, p. 553.
Pt. xanthorrhous, p. 553
Pt. porphyreus, p. 553.
Pt. gambu, p. 554.
Pt. wallacei, p. 555.
Pt. aurantiifrons, p. 556.
Pt. ornatus, p. 556.
Pt. gestrot, p. 557.
34
522 MR. D. G, ELLIOT ON THE GENUS PTILOPUS. [May 7,
b'. Middle of scapulars and lesser wing-
coyerts rosy purple.
a. Tail uniform dark green ......s0s0+.0. 44, Pt. perlatus, p. 557.
6". Tail dark green, with an apical greyish
Vibe DAMGN Us. c-esac.ce-kecserasenetes ates 45, Pt. zonurus, p. 558.
i. Head, except occiput, light grey ; abdomen
reddish [01 6) eane see eoppBa, sosueceodorbannad 46. Pt. iogaster, p. 558.
j. Forehead and chin greenish grey ; reddish
purple spot on breast ............00cseseeees 47. Pt. pectoralis, p. 559.
k. Forehead and chin grey; tips of tertials
and bar across the shoulders grey.
a', Lower part of throat and upper part of
breast dark reddish purple; tail bronzy
green, tipped with yellow ........+..+... 48. Pt. viridis, p. 560.
b'. Lower part of throat and upper part of
breast bright reddish purple ; tail yel-
lowish green, each feather with a sub-
terminal grey spot on the inner web... 49. Pt. geelvinkianus, p. 560.
1. Head white, secondaries slate-grey ; centre
of throat and chest crimson ............... 50. Pt. eugenia, p. 561.
m. Upper parts, throat, and upper part of
breast green.
a’, Breast crossed by a broad white band ;
forehead purplish red.
a’. Rose spot on abdomen,
1. Crissum and under tail-coverts
bright yellOw <2.........:.0csssceaee 51. Pt. rivolit, p. 561.
2. Crissum and under _ tail-coverts
green, edged with yellow ......... 52. Pt. prasinorrhous, p. 562.
6’. No rose spot on abdomen ............ 53. Pt. strophium, p. 568.
b'. Breast crossed by a broad band, lemon-
yellow above, white beneath.
a''. Front and crown rosy red; purplish
red spot on abdomen ................45 54, Pt. bellus, p. 563.
6". Frontandcrowngreen; abdomenlilae 55. P¢. speciosus, p. 564.
c’, Band on breast, crissum, and under tail-
coverts yellow ; top of head and middle
of abdomen lilac ...... isc. sesecsvsreesese 56. Pt. gohannis, p. 564.
n, Feathers of neck and breast lanceolate, sul-
phur-yellow, tinged with green; abdomen,
vent, and under tail-coverts bright yellow 57. Pt. luteovirens, p. 565.
o. Head and neck olive-green ; plumage green ;
secondaries tipped with yellow; under
tail-coverts yellow, blotched with green . 58. Pt. corriei, p. 566.
p. Head and neck greenish yellow; under tail-
coverts bright yellow; rest of plumage
Fale oes Rapa ar oc bist accueuencon ead poet 59. Pt. layardi, p. 567.
g. Head and throat olive-green ; rest of ae
mage OFANge-CArMINE, ............ee nese eee . 60. Pt. victor, p. 568.
r. Top of head light grey
a'. Sides of head and upper part of breast
ashy white ; a dark purple band across
the breast ; "flanks and underparts ashy
green ; under tail-coverts cinnamon ;
Givin DISK syccos acecdns seeeens . eaeeaeees 61. Pt. leclancheri, p. 568.
b'. Sides of face and occiput purplish red ;
centre of breast ochraceous............... 62. Pt. occipitalis, p. 569.
c'. Sides of face, breast, and flanks grey ;
centre of abdomen yellowish white ;
thighs and under tail-coverts cinna-
mon; chin deep chestnut .......cce00+0 63, Pt. gularis, p. 570.
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 523
d'. Head, neck, and underparts partly grey,
partly golden-ochre ; under tail-coverts
grey and white intermixed; sides of
head bluish red; black band on back
OT WTC epee pth ee Eee eee ES 64. Pt. fischeri, p. 571.
s. Head white.
a’. Abdomen yellowish green, separated
from breast by a black band.
a", Neck and breast light blue ............ 65. Pt. albocinctus, p. 571.
6". Neck and breast white.
1. Tailblack, with apicaldarkgrey band 66. Pt. cinctus, p. 571.
2. Tail greyish black, with apical third
Vellowish/White, gisicena:decssesows 67. Pt. lettiensis, p. 572.
Head and neck greenish grey.
a'. Red spot on lower part of breast ; abdo-
men and flanks yellowish ochre ...... 68. Pz. bernsteini, p. 572.
4’, Centre of throat and neck in front and
underparts to the thighs purplish red 69. P#. puella, p. 574.
ce’. Larger race of the above ..........ss000.0. 70. Pé. assimilis, p. 574.
d', Stilllarger ; centre of throat and under-
parts deepypurple © ...0.2....ss>.0reeereane 71. Pt. magnificus, p. 575.
™
.
1, PriLopus GREYI.
Ptilopus greyi, Grey, List B. Brit. Mus. (1856) p. 4. sp. 8, desc.
nulla—Columbe ; id. List B. Trop. Isl. (1859) p. 38; id. Proc.
Zool. Soc. 1859, p. 165; J. Verr. & Des Murs. Rev. Zool. 1860,
p- 435 ; Bon. Iconogr. Pig. (1857) pl. xx.
Pitilopus purpuratus, Bon. (nec auct.) Consp. Gen. Av. vol. ii.
p. 19. sp. I.
Hab. New Caledonia (Germain); Isle of Pines (Gray); Lifu,
Loyalty Islands (Whitmee)-; Erromaugo, New Hebrides, Vanikoro
(Voy. Astrolabe).
This handsome species was first named by Gray, but not described
(l.c.). Bonaparte gave a description and very good figure of the
bird in his Iconogr. Pig. pl. 20, from the specimen brought, by the
naturalists attached to the expedition of the ‘ Astrolabe,’ from Vani-
koro Island. The species has a rather extensive distribution, from
New Caledonia and neighbouring islands to Vanikoro on the north,
Young and immature birds are greener on the breast and neck than
the adults; and the first primary is not so abruptly attenuated.
Adult. Front and top of head deep rose, margined posteriorly with
_ yellow ; back of head, neck, and breast pale greyish green; throat
yellowish white; flanks and sides of crissum bright green; abdomen
very deep rose-colour, running into orange-yellow oncrissum. Under
tail-coverts rose-colour, but not so dark as on the abdomen. - Upper
parts bronzy green. Primaries black on inner web, dark green on
the outer, and the innermost ones tipped with bronze-green ; first
primary abruptly attenuated, end very narrow. Secondaries shining
green, edged with yellow. ‘Tertials and some of the innermost secon-
daries with a pale lilac spot near their tips. Tail dark green, with a
subterminal grey band. Bill black, with pale tip, Feet brown,
possibly red in life. Total length 8 inches, wing 5, tail 3, culmen 13.
Specimen described is the one figured by Bonaparte, Iconogr, Pig.
pl. 20, ex Vanikoro. 3
4*
524 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Another specimen from New Caledonia, marked as a female,
differs from the male only in having the forehead slightly tinged
with rose, and the abdomen mottled with rose and bright yellow.
The under tail-coverts are deep rose on their apical half, as in the
specimen from Vanikoro. The secondaries are more broadly mar-
gined with yellow, and the primaries tipped with white. I am
inclined to regard this example rather as an immature male than a
female. Three examples are in the British Museum from Erro-
mango Island, one of the New Hebrides, which are in all respects
like the one in the Paris Museum figured by Bonaparte from
New Caledonia.
2. PTILOPUS NEGLECTUS.
Ptilopus neglectus, Schleg. Mus. Pays-Bas (1873), p. 7, Columbe.
Hab. Océanie! Viti? (Schlegel).
Allied to the Pt. greyi, but differs in having the lower part of the
abdomen and under tail-coverts tinged with sulphur, the abdomen
and breast washed with olive-green, the purplish-red spot being
placed upon the dreast and not on the abdomen, and in the neck
and upper part of breast being tinged with light grey (Schlegel).
Wing 5 inches, tail 2 inches 3 lines.
The locality suggested by Schlegel, as given above, “ Viti,” is
without doubt incorrect, as there are no indications to support the
idea, the relatives of this bird having been procured in quite a
different group of islands.
3. PTILOPUS BONAPARTEI.
Ptilopus purpuratus, Bon. (nec auct.) Iconogr. Pig. (1857) pl. xix.
fig. a, adult, ex Balaou (nee Vanikoro).
Ptilopus bonapartei, Gray, Hand-l. B. vol. ii. (1870) p. 225,
descr. nulla, founded on pl. 19, Bon. Iconog. Pig.
Hab. Balaou, Fiji Islands (Hombron and Jacquinot).
This is the species figured by Bonaparte as Pt. purpuratus, fig. a
(J. c.). The specimen is in the Paris collection ; and the plate gives
no idea of the bird; which is reddish in all lights, but in certain ones
exhibits a brilliant coppery-red hue. Bonaparte committed two
strange errors in reference to this bird. He stated that it came
from the island of Vanikoro, and also figured as the young the Pé.
purpuratus, Wagl., from Tonga-Tabou, a very different and distinct
species. Gray (/.c.) bestowed the name of Pé. bonapartei (1. ¢.), ex
Vanikoro!, on the bird represented by Bonaparte on plate 19 of his
work, ‘Iconographie des Pigeons.’ Gray could never have seen
the specimen ; or else he would have corrected Bonaparte’s error, and
stated that it did not come from Vanikoro, but from Balaou, and also
that Bonaparte had confounded two species. Gray gives no de-
scription, which also strengthens the belief that he did not know the
species autoptically. It is a question whether Gray’s name should
be noticed at all; for to name species without seeing them, on the
chance of their proving new, is not a commendable practice ; and, as
he did not designate which figure he bestowed the name of bona-
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 525
partei upon, it is of course impossible to know whether it ought to
be applied to the present or to the Tonga-Tabou bird. As, however,
I now have selected the Balaou species as the type of bonapartei, in
order to avoid giving it a new name (for it is quite distinct from the
Pt. purpuratus, Wagl.), and as the type will be in the collection of
the Paris Museum for future reference, it will not be difficult here-
after to ascertain what the species really is. The type is unique.
Adult. Front and crown rosy purple, faintly margined posteriorly
with yellow; throat white ; occiput, neck, and breast ash-colour,
tinged with green; abdomen and flanks green; an indistinct dark
green spot in the middle of the former. Back and wings green,
with strong coppery-red reflexions. Inner secondaries with a small
dark green spot at tips, and all edged with yellow. Primaries dark
coppery red, edged with white. Tail above green, with bright
coppery-red reflexions, and with a subapical greyish-white band ;
beneath dark grey, with a broad apical white band. Crissum and
under tail-coverts bright yellow. Bill greyish black, with yellow tip.
Feet light red. Total length 9 inches, wing 53, tail 33, culmen 2.
4. Pritorus PURPURATUS.
Columba purpurata, Shaw, Gen. Zool. vol. xi. p- 66 (1819),
partim ; Wagl. Syst. Av. (1827) Columbe, sp. 30.
Columba porphyracea, Temm. Trans. Linn, Soc. vol. xiii. p. 1302
(1822).
Columba porphyrea, Wagl. Syst. Av. (1827) sp. 312, Columbe.
Columba forsteri, Desm. Dict. Scien. Nat. vol. xl. p. 340 (1826)?;
Less. Comp]. Buff. tom. iii. Ois. p. 34 (1837), ex Tonga-Tabou.
Columba purpurata, Wagl. (nec Gmel.) Isis, 1829, p. 742, ex
Tonga-Tabou (1829).
Columba porphyracea, Licht., Forst. Descr. Anim. p- 167 (1844) ;
Finsch & Hartl. Fauna Centralpolyn. (1867) p. 119; Graffe, Journ.
Mas. Godeffr. tab. 7. fig. 3.
Ptilonopus porphyracus, Gray, Gen. B. (1844-49) vol. ii. p. 466;
id. B. Trop. Isl. p. 38 (1839); id. List Columb. p- 3 (1856), partim.
Ptilopus purpuratus, Bon. (nee Gmel.) Iconogr. Pig. (1857)
pl. xix. fig. 6, juv., spec. Paris Mus. ex 'T onga-Tabou (Voy. Astro-
labe), nec Vanikoro.
Ptilinopus porphyraceus, Finsch & Hart. Journ. fiir Ornith. 1870,
p- 131; Finsch, Proc. Zool. Soe. 18775 ppe 7 2557-7557 83.
Ptilonopus porphyraceus, Reich. Taub. p- 95; Layard, Proc, Zool.
Soc. 1876, p. 502. '
Ptilonopus purpuratus, Reich. (nec auct.) Taub. p. 94?
Columba kurukuru, variété, Temm. & Knip, Pig. vol. i. pl. 352
Pilopus fasciatus, Layard (nec Peale), Ibis, 1876, p. 392.
Hab. Tonga-Tabou, Waikai, Mango, Mokani, Loma-Loma, Ta-
viuni, Fiji Islands, Futuna, Friendly Islands (Layard) ; Ninafou
Island, most northern of Tonga group (Finsch); Hapai (Griffe) ;
Bua, Friendly Islands (Hibner) ; Savage Island (Whitmee) ; Vavao,
Friendly Islands.
It is apparently a difficult matter for authors to decide upon the
526 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
correct name for this Fruit-Pigeon ; and, as will be seen by the list of
synonyms given above, the species has been presented with a very
fair number of appellations. In order to explain why I consider
that this bird, of all those given in this paper, should alone bear the
name of purpuratus, it will be necessary to begin at the beginning.
Latham, in his ‘General Synopsis of Birds,’ vol. ii. p. 626. sp. 15
(1783), describes a Dove, which he says came from Otaheite, as the
“« Purple-crowned Pigeon,” the chief characteristics of which were :—
‘“* Forehead to the middle of the crown purple; head, neck, and
underparts of body pale green, inclining to ash-colour; vent and
under tail-cuverts yellow. Tail greenish black, with green edges, and
when spread a greyish bar appears near the end, the colour only
occupying the inner webs of each feather.” To this description
Gmelin gave the name of Columba purpurata. Now there is no
Pigeon known belonging to the genus Ptilopus that will answer to
the above description of Latham. No such bird is found upon the
island of Otaheite, whence, according to Latham, this curiously
compounded specimen came ; for it carries the head of the bird from
Tonga-Tabou and the body of the species found in Otaheite,
distinguished afterwards by Lesson as faitensis. Latham possessed
specimens from both these localities, and remarked upon their dif-
ferences, but did not deem them distinct species; and it is very
evident that when he wrote his description he took his characters
from both forms. The bird from Otaheite does not possess a purple
forehead or crown, these parts being ash-colour with but a faint
tinge of lilac; therefore it could never have been designated as the
‘* Purple-crowned Pigeon.’ Nor does the Tonga-Tabou bird have the
underparts of the body pale green inclining to ash-colour (which is
a characteristic of the Otaheite species), but has the flanks and abdo-
men grass-green, with a conspicuous purplisk-black spot in the centre
of the latter. Therefore the name Col. purpurata, Gmel., having
been founded upon a bird that did not exist, taken from a description
comprising portions of two distinct species, cannot be accepted, and
should be rejected from the nomenclature of the genus.
The next author after Gmelin who mentioned this species was
Shaw (/.c.) (1819). He, however, also confounded two species, and
his name of purpurata must be passed over. Temminck follows in
1822 (l.c.) with Columba porphyracea ; and this name, attributed to
Forster, has been used by various writers when referring to this bird,
especially from the island of Tonga-Tabou. Now the C. porphy-
racea of Temminck was bestowed upon the bird figured by Madame
Knip in their great work on the Pigeons, at plate 35, and there con-
sidered as a variety of the Pt. ewingii, and called Columba pur-
purata. Temminck states (Trans. Linn. Soc. 1822, p. 130) that his
specimen came from New Holland! The figure resembles no species
from that country, nor indeed from any other; and it can only be
through an examination of the type that a correct knowledge of the
species intended can be ascertained. I have therefore placed Tem-
minck’s name, as well as porphyrea, Wagl., founded upon Tem-
minck’s bird, whatever it may have been, among the synonyms of
1878.) MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 527
the present species with a doubt’. Desmarest also published a
name (/.c.) for the bird depicted on Temminck’s plate, viz. forstert.
This, for the same reason which has placed Temminck’s name on
one side, will also have to go among the list of doubtful synonyms.
We now come to Wagler, ‘ Isis,’ 1829, who first diagnosed as distinct
both species, from Otaheite and Tonga-Tabou respectively, and gave
names to each, and properly portrayed their distinctive ckaracters.
To the Tonga bird with the purple crown he applied the term
purpurata, which name will take precedence; and the Otaheite
species he called oopa. Unfortunately he did not appear to be
cognizant of the name ¢attensis previously given to this species by
Lesson, Voy. Coq. tom. i. text, p. 297 (1826); and therefore oopa
will of course become a synonym of Lesson’s term.
Mr. Layard speaks of this species under the name of porphyraceus,
Forst., as not uncommon in the Friendly Islands. In its habits it
resembles the other species of this genus, and is called by the natives
* Kullu kullu.” The female, when in full breeding-dress, resembles
the male, but is not so brightly coloured on the abdomen, the male
at this season having the black abdominal patch shading away into
brilliant purple.
Adult. Front and top of head, extending in a curved line behind
the eyes, rich purple, edged with light yellow. Rest of head, neck,
upper part of back, throat, and breast pale ashy green. Flanks and
abdomen grass-green, with a dark purple spot in the centre of
abdomen. Crissum and under tail-coverts bright yellow, tinged
with orange on the tips of the coverts. Back and upper tail-coverts
dark yellowish green. Wings the same colour, with the inner
secondaries and some of the scapulars deep bluish green near their
tips. Primaries black on their inner webs, green on their outer,
and, together with the secondaries, margined with yellow; first
primary abruptly attenuated and very narrow at the end. Rectrices
bright green on their outer webs, brownish black on their inner, with
a terminal grey or ash-coloured band, this margined with yellow.
Under surface smoke-colour, terminal portion white. Bill black, tip
flesh-colour. Feet brownish. Total length 8? inches; wing 5};
tail 32; bill, culmen 3.
Young. Resembles the adult, except that the top of the head is
ashy like the breast, with only a tinge of purple on the forehead.
The back of the neck is pale green, instead of ash-colour; and the
plumage of the upper part, tail, and wings is bronzy.
5. PTILOPus APICALIS.
Lamprotreron apicalis, Bon. Compt. Rend. vol. xxxix. p. 877
1854).
. nee apicalis, Bon. Consp. Gen. Av. vol. ii. (1857) p. 23 ;
Finsch & Hartl. Centralpolyn. (1867) p. 121.
Ptilonopus apicalis, Reich. Tauben, p. 97.
Hab. Samoa, Vavao ( Bonaparte).
This species, described by Bonaparte (J. c.), while resembling in
? See further remarks upon this plate 35 in the article on Pt. coralensis.
528 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May /,
general appearance the preceding one, differs from all others in the
colour of the spot upon the abdomen. In the type this is mixed
with rufous and yellow, the latter colour perhaps predominating ;
but in a second specimen, also in the Paris collection, the spot is
more rufous, and extends over a greater space. I regard this latter as
a more adult individual. It unfortunately bears no precise locality,
but has merely marked upon the stand, ‘donné par la Société des
Missions Catholique de ! Océanie en 1849.” The type was brought
by Hombron and Jacquinot from the Samoan Islands in their Voyage
au Péle Sud. The exact island is not given on the specimens by
these travellers; but Bonaparte has stated Vavao to be the habitat.
I am not aware how he ascertained this island to be the one in
which the specimen described by him was procured.
Adult. Front and crown rosy purple, faintly margined with
yellow; throat yellowish white; occiput and back of neck pale
green ; sides of neck and breast greenish grey; flanks and abdo-
men pale green, with a rufous spot tinged with yellow in the centre
of the latter ; crissum and under tail-coverts bright yellow, inclining
to a deep orange hue at the tips. Upper part and wings bronzy
green. Secondaries margined on outer webs with pale yellow. Tail
bronzy green, with an apical light-yellow band. Bill green? Feet
dark red? ‘Total length 7$ inches; wing 53, tail 33, culmen 3.
(Type.)
6. PrrLOPUS CORALENSIS.
Ptilopus coralensis, Peale, U.S. Expl. Exped. (1848) p. 190,
pl. 51; Cass. U.S. Expl. Exped. 2nd ed. (1858) p. 272, pl. 32;
Schleg. Mus. Pays-Bas, 1873, p. 14, Columbe.
Columba purpurata, Temm. & Knip, Pig. vol. i. pl. 357:
Ptilopus viridissima, Bon. Consp. Gen. Av. vol. i. p. 20 (1857).
Kurutreron coralensis, Bon. Consp. Av. vol. il. p. 26. sp. 3
(1857); Reich. Taub. p. 93; id. Novit, Suppl. p. 203, t. ii. f. 33.
Ptilinopus coralensis, Cass. Wilkes Expl. Exped. text, p. 272, atl.
pl. 32 (1858).
Ptilonopus coralensis, Gray, List B. Trop. Isl. p. 37 (1859).
Ptilonopus viridissimus, Gray, List B. Trop. Isl. p. 38 (1859),
partim ; Reich. Tauben, p. 95, pl. 236 0. fig. 3367.
Ptilinopus viridissimus, Finsch & Hartl. Faun. Centralpolyn.
(1867) p. 134.
Hab. Island of Carlshoff, Paumotu group (Peale).
The type of Bonaparte’s Péilopus viridissimus is in the Paris
Museum. The example, which was brought home by M. Maugé, the
surgeon attached to the expedition commanded by Captain Baudin,
is marked as from Timor. ‘This locality is evidently erroneous, as
no species belonging to this particular group of the genus Ptilopus
is found in that island, but all are, on the contrary, apparently con-
fined to the islands of the South Sea, from some one of which it is
most probable the specimen described by Bonaparte was brought.
It is in a bad condition, the primaries and tail being worn away to a
considerable degree, as though the bird had been in captivity at
1878.] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 529
some period of its existence. It may possibly have been purchased
by Maugé at Timor. In the coppery-red hue of the wings, as well
as the tail, it answers well to the description given by Peale (J. c.) of
his P. coralensis ; and I have therefore assigned it to that species. The
back of the type is not ‘‘viridissimum immaculatum,”’ as described
by Bonaparte, but has very conspicuous violet spots on the scapulars.
I have given among the synonyms of this species the Columba
purpurata, var., figured by Madame Knip on plate 35 of the work on
Pigeons. Temminck states that there are two specimens in the
Paris Museum, brought by Capt. Baudin, which resemble the type
of his plate 35. I find that the type of Bonaparte’s Pé. viridissimus
is marked as follows :—‘‘Ptilopus viridissimus, Bp. ex Temm. tab. 35,
nec 34.” This unquestionably refers to the work of Temminck and
Knip ; but whether it is also intended to indicate that the example
was the original of Madame Knip’s plate I am unable to ascertain.
I deem it very probable, however, that it is the original from which
the drawing was made. In that case this bird would bear the name
of porphyraceus, Temm. Trans. Linn. Soc. 1822, p. 130, which
antedates coralensis, Peale.
Mr. Peale states that this Dove attracted his attention by its
plaintive call, resembling hoot-hoot-hoot, uttered in rapid succession.
Although not wild, it was very difficult to see them, as their plumage
accorded so well with the leaves. Specimens were subsequently
obtained on the low coral islets of the Paumotu group, but nowhere
else. They always kept in the most wooded and secluded places.
Male. “‘ Vertex having a very pale rubiaceous spot, which does not
reach to the base of the bill; bill yellow; irides red; head, neck,
and breast ashy, the feathers having a very pale greenish-yellow
band near their tips, those on the breast bifurcate ; back and rump
green, inclining to olive; vent, wings, and tail, when viewed in a
positive light, bright copper-colour, barred with golden green;
underparts of the wing and tail cinereous; quill-shafts brown ; tail-
shafts white beneath and black on the upper surface ; tail even, the
inner web having a dark bar near the tip; tertiary feathers edged
with yellow ; primaries and secondaries having a very narrow margin
of white; feathers on the leg cinereous. ‘Total length 9 inches; tail
35), extent of wings 17, bill 3, tarsus 1.”—Peale (J. c.).
7. PTILopUS CHALCURUS.
Piilonopus chaleurus, Gray, List Birds Trop. Isl. Brit. Mus.
(1859) p. 37.
Ptilinopus chaleurus, Finsch & Hartl. Faun. Centralpolyn.
(1867), p. 131.
Hab. Harvey Islands, Cuok’s Archipelago (Gray).
The type of this species (?) is in the British Museum, where I
have had an opportunity of examining it. The colour of the fore-
head is amethyst, a lighter hue than in P¢. viridissimus, Bon. ; but in
their coppery tints the two birds greatly resemble each other. Gray
separated this Harvey-Island bird from Pé. coralensis, Peale, on
account of the light-coloured front and crown. Without the
530 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
type of Peale’s species to compare it with, I cannot say if the
difference mentioned is sufficient to separate the present form as dis-
tinct, and for the present leave it under the designation given to it
by its describer.
Forehead amethyst. Crown, occiput and nape pale green. Throat
yellowish ; sides of neck and breast greyish ash; abdomen pale
green. Back and wings bronzy-green, Primaries and secondaries
with a coppery-red hue in certain lights. Tail coppery red, with
a greenish shade; but in the majority of lights the feathers are
bright copper-colour. Crissum and under tail-coverts pale yellow.
Bill, legs, and feet yellowish in the skin. Wing 53, tail 4in. (Type.)
8. Pritorus pictiventrRis. (Plate XXXIII.)
Ptilonopus fasciatus, Whitmee (nec Peale), Proc. Zool. Soc.
1873, p. 152 (spec. exam.)
Ptilonopus apicalis, Lay. (nec Bon.), Proc. Zool. Soc. 1876, p. 495.
Ptilopus pictiventris, Elliot, Ann. & Mag. Nat. Hist. ser. 5, vol. i.
p- 349 (1878).
Hab. Nukahiva, Marquesas Islands (?) (type Paris Museum) ;
Samoa (Whitmee); Savage Islands (Brenchley) ; Navigators’ and
Friendly Islands (Layard).
This species was sent from Samoa by the Rev. S. J. Whitmee,
and marked by him Pé. fasciatus. I have examined his specimens
now in the British Museum, from one of which the figure (Plate
XXIII.) is taken; and they agree exactly with my type in the
Paris Museum. Mr. Whitmee states (J. c.) that this bird is kept
generally in- captivity by the natives of the Samoan Islands, and
they train it to act as a decoy for the wild bird. I described
this species from a specimen in the Paris Museum, stated to have
been brought by Hombron and Jacquinot from Nukahiva of the
Marquesas. I am inclined to think this locality may be erroneous,
as I find on the bottom of the stand that it is first stated to have
come from “ Upolu,’ Samoan group. This was erased and Nuka-
hiva substituted. It seems to me, in view of the Rev. Mr. Whitmee’s
examples, that Upolu is probably the correct locality of the type.
Mr. Layard, in the ‘ Proceedings’ of this Society, 1876, p. 495,
has erroneously referred this bird to the Pt. apicalis, Bon. This
latter species, the type of which is before me, has a rufous spot
mixed with yellow in the centre of the abdomen, without any black
line about it. The secondaries are also uniform bronzy green without
any lilac spots near their tips; and the under tail-coverts are bright
yellow inclining to an orange hue at their tips. The present species
and Pé. apicalis are readily distinguishable from each other, the pur-
plish-black line above the rufous on the abdomen of the present bird
being at all times sufficient to separate it. Mr. Layard states that he
found this species abundant, feeding on wild berries, and having a
pleasing rolling ‘‘coo.”” The females were not so brightly coloured
as the males ; and the young have the whole of the underparts scaly
with yellow edges to their green feathers. They exhibit no trace
of the bifurcation in the breast-feathers, as in the adults; nor is the
1878.] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 531
first primary so greatly attenuated. Front and crown rosy purple,
with a very faintly indicated yellow margin, almost obsolete. Occi-
put greenish grey. Throat whitish; neck, upper part of mantle,
and breast ashy green. Flanks light green. Centre of abdomen
rufous, bounded above by a deep-purple line. Crissum and under
tail-coverts bright yellow, the latter orange towards their tips. Back
and upper tail-coverts bright green. Scapulars tipped with lilac.
Tail bright green, with a broad apical yellow band. Bill greenish
with a yellow tip. Feet brown in the skin. Total length 83 inches,
wing 53, tail 3, culmen 5%.
9. PriLOoPUS PELEWENSIS.
Ptilinopus pelewensis, Hartl. & Finsch, Proc. Zool. Soc. 1868,
p- 7, 1872, p. 101; Graffe, Journ. Mus. Godeffr. 1873, pt. 1,
pl. 7. fig. 5; Finsch, Vég. Palau-Gruppe, Journ. Mus. Godeff.
1875, Heft viii. p. 24.
Hab. Pelew Islands.
This species, described by Messrs. Hartlaub and Finsch (J. c.)
belongs to the second group into which I have divided the green
Fruit-Pigeons of the South-Sea Islands, viz. the one with the yellow
apical band on the tail. It differs apparently from its relative in
the violet spot on the breast-feathers, and in the colouring of the
under tail-coverts, and also, as Dr. Finsch writes me, in the large
orange vent-spot. It is evidently a very distinct species. In their
first description the above-mentioned authors state that the under
tail-coverts are white at base, then yellow, with the tips rose-violet.
In the Society’s ‘ Proceedings’ for 1872, p. 101, they describe these
coverts as bright purplish red, with narrow bright orange edgings.
From this there would seem to be considerable variation in the hues
of these feathers. I have not seen an example of this species.
Adult. Front and crown rose-violet, margined with yellow ; rest
of head and neck pale greenish yellow, feathers of upper part of
breast crossed transversely with violet spots, tips greenish yellow.
Middle of abdomen deep orange. Crissum yellow. Under tail-
coverts white at base, then yellow, with the tips rose-violet.
Upper parts light green. Primaries black, tipped and margined
on the outer web with light green. Secondaries bright green
edged with yellow. Middle rectrices green; lateral ones black on
inner webs, green on outer, all with an apical yellow band. Bill
lead-colour, feet flesh-colour. Length 93”, wing 8” 2'”, tail 22”,
tarsus 10"", bill 6’” (Hartl. and Finsch).
10. PriLopus REGINA.
Columba purpurata, prem. var., Temm. Pig. et Gall. tom. i.
p- 282 (1813) (2).
Columba purpurata, var.regina, Swains. Zool. Journ. vol. i. (1825)
- 473.
Columba purpurata, Jard. and Selby, Mlust. Ornith. vol. ii. pl.
70 (nec auct.).
Ptilinopus swainsoni, Gould, Proc. Zool. Soc. 1842, p. 18; id.
532 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Birds Austr. vol. v. pl. 55; id. Hand-b. B. Austr. vol. ii. p. 106;
Finsch & Hartl. Faun. Centralpolyn. (1867) p. 125; Graffe,
Journ. Mus. Godeff. 1873, pt. 1, pl. 7. fig. 6; Ramsay, Proc.
Zool. Soc. 1876, p. 114.
Ptilonopus swainsoni, Gray, Gen. B. (1844) vol. ii. p. 466. sp. 6 ;
id. List B. Brit. Mus. (1856) p. 2. sp. 1; Reich. Tauben, p. 94,
pl. 235. figs. 1309-10.
Kurukuru swainsonii, Des Murs & Prév. Voy. Vénus, Zool. (1855)
. 256.
h Ptilopus swainsonii, Schleg. Mus. Pays-Bas, 1873, p. 4, Columbe.
Hab. Australia, river Clarence, between the Hunter and More-
ton Bay (Gould).
Temminck, in his ‘ Pigeons et Gallinacées,’ (/. c.), describes a ‘‘ pre-
miére variété ” of his Columba purpurata (=ewingii, Gould). This
has been referred, by Messrs. Finsch and Hartlaub in their work
on Central-Polynesian birds, to Pt. fasciatus, Peale. I do not
think this is correct ; and although it may be impossible to prove
what really this ‘variety’ of Temminck’s is, yet it seems to me
that his description points more towards the present species. In the
first place Temminck makes no mention of a purplish-black spot upon
the abdomen, which Pé. fasciatus (Peale) possesses ; and he describes
the belly as exhibiting “ plusieurs nuances de jaune et d’orange,”
which certainly are not to be found on Peale’s bird, which has this
part grass-green with a purplish black spot in the centre. As the
Columba purpurata, Temm., is known to be the Pé. ewingii, Gould,
and did not come from Tahiti, it is probable that he had with it
the Pt. regina, also from Australia, and looked upon them as
varieties of the same species. As this cannot, however, be defi-
nitely ascertained, I have placed his name among the synonyms, with
a question. Temminck states that there are two specimens in the
Paris Museum, brought by Capt. Baudin, which are like the variety
described by him. I have examined all the examples brought by
Baudin’s expedition ; and they belong only to the Pé. ewingii and
Pt. viridissimus, Bon. (=coralensis, Peale). This species has been
generally known as the Péilopus swainsoni, Gould; but it was
called regina by Swainson seventeen years before Mr. Gould’s name
was published. In fact it is the type of Swainson’s genus Ptilo-
nopus (Ptilopus) ; and Gould’s name must become a synonym.
Male. Forehead and top of head crimson, margined narrowly
with yellow. Occiput and back of neck greyish green. Throat
white, sometimes tinged with yellow. Breast pale green, each
feather tipped with light grey. Flanks dull green. Abdomen deep
orange, with a spot of deep lilac-red in the centre. Crissum and
under tail-coverts orange-yellow. Upper parts bright green. Pri-
maries slaty grey on inner webs, green on the outer, edged with
yellow. The first abruptly and greatly attenuated. Secondaries bluish
green, becoming deep blue near their tips, and broadly margined with
yellow. ‘Tail deep green, with a broad terminal rich yellow band.
Irides reddish orange. Bill greenish black, horn-colour at the tip.
feet olive-brown. Total length 94 inches, wing 54, tail 33, culmen 3.
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 533
Immature. Front tinged with rose-colour ; loral space and a line
over the anterior half of the eye light yellow. Head, neck, cheeks,
back, and breast green, the feathers of the latter tipped with light
yellow. Throat yellowish white. Abdomen and under tail-coverts
light yellow; the lilac-red spot absent. Rest of plumage as in
the adult.
Specimens described are those figured by Bonaparte, Iconogr. Pig.
pl. 21.
11. Pritopus EWING.
Columba purpurata, Temm. Pig. et Gall. tom. i. p. 280 (1813) ;
Less. Traité Ornith. (1831), p. 472 (nec auct.).
Columba purpurata, Desmar. Dict. Se. Nat. vol. xl. (1826) p.
339; Knip & Prév. Pig. vol. i. pl. 34 (nec auct.).
Le Kurukuru, Less. Compl. Buff. Ois. tom. iii, p- 32 (1837).
Ptilinopus purpuratus, Selby, Nat. Libr. vol. v. pt. 3, p. 103
(1835) (not from Otaheite); Swains. Class. Birds, vol. ii. p. 347
(nec auct.).
Piilinopus ewingii, Gould, Proc. Zool. Soc. 1842, p: 19); id;
B. Austr. vol. v. pl. 56.
Ptilonopus purpuratus, Gray, List B. Brit. Mus. (1844) p. 2,
Galline.
Ptilonopus ewingii, Gray, List B. Brit. Mus. (1844) p. 2, (1856)
p. 3. sp. 2; id. Gen. B. vol. ii, (1844) p. 466; Reich. Tauben,
p- 94, pl. 235. figs. 1307-8.
Kurukuru roseicapillus, Des Murs & Prév. (nec Less.), Voy.
Vénus, Ois. p. 262 (1855), partim.
Ptilinopus flavicollis, Gray, List B. Brit. Mus. (1856) p. 3. sp. 3.
Piilonopus flavipectus, Reich. Tauben, p- 94, pl. 235. fig. 1305
(ex pl. 35, Knip, Pig.), ex Timor.
Ptilonopus flavicollis, Reich. Taub. p. 95; Gray, List B. Brit.
Mas. (1856) p. 3. sp. 3; Wall. Ibis, 1865, p- 380.
Ptilopus flavicollis, Bon. Consp. Gen. Av. vol. ii. p- 20. sp. 4; id.
Compt. Rend. (1855) tom. xl. p. 216. sp. 40; id. Iconog. Pig.
(1857) pl. 25.
Ptilopus ewingii, Bon. Consp. Av. vol. ii. p. 20 (1857) ; id.
Iconog. Pig. (1857) ,pl. 24. fig. 1 ; Schleg. Mus. Pays-B. 1873, p. 5.
Ptilinopus flavicollis, Finsch & Hartl. Faun. Centralpolyn.
(1867) p. 125.
Hab. Coburg peninsula, northern coast of Australia (Gould) ;
Timor, Flores (Semmelink).
The representation of this species given by Madame Knip on
plate 34 of her work is so badly executed that it is impossible to
recognize what is intended ; for it certainly resembles no species that
was ever seen. Therefore Bonaparte was perfectly justified in
giving another figure in his Iconogr. Pig. pl. 25, from which a very
good idea of this species is obtained. It is very safe to say that,
unless the originals of some of Madame Knip’s plates had been
preserved in the Paris Museum to show what the species really
were, they would always have remained a stumbling-block to ornitho-
534 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. _[May 7,
logists. I do not find, after a careful comparison of specimens
from the two localities, that the birds from Timor and Australia
show any characters that should separate them as distinct species.
Desmarest, in the ‘Dictionnaire des Sciences,’ gives a species of
Ptilopus as Columba purpurata. This has been referred to the
Pt. fasciatus of Peale. As, however, the description mentions
yellow and orange colours on the belly, and, moreover, this bird is
referred to plate 34 of Temminck and Kuip’s ‘ Pigeons,’ which re-
presents Pt. ewingii, I have placed Desmarest’s name among the
synonyms given above.
Aduit male. Forehead and crown rose-colour, margined with
yellow. Sides of head greyish. Throat light yellow. Breast pale
green, feathers tipped with yellow. Abdomen orange, with a pale
lilac band crossing the upper part. Flanks green. Crissum and
under tail-coverts orange-yellow. Back of neck and upper parts
bright green. Wings bluish green. Primaries brownish black on
inner webs, dark green on outer, margined with yellow, the first
abruptly and greatly attenuated. Secondaries bluish green; base
of inner webs brownish black, margined with light yellow. Tail
bright green, with a broad terminal yellowish-white bar. Bill
black, with yellow tip. Feet brownish. Total length 9 in., wing 5,
tail 34, culmen }. (Specimen figured by Bonaparte.)
12. PriLOPUS RAROTONGENSIS.
Ptilinopus rarotongensis, Hartl. & Finsch, Proc. Zool. Soc. 1871,
p- 80; Griiffe, Journ. Mus. Godeff. 1873, pl. 7. fig. 1.
Hab. Cook’s Islands (Finsch § Hartlaub); Savage Island
(Whitmee, Brenchley).
A specimen in the British Museum from Savage Island, one of
the Friendly group, is apparently of this species, and answers quite
well to the description given by Finsch and Hartlaub. The spot
on the lower portion of the breast is rosy red in the centre, sur-
rounded by very dark purple. Abdomen and under tail-coverts
are yellow, the latter strongly tinged with orange towards the ends.
** ddult. Front and vertex purplish violet-red, surrounded by a
narrow indistinct line of yellow ; occiput, sides of head, neck, crop,
and upper part of breast delicate grey, the feathers of the latter bifur-
cate and with a pale yellow cross band ; chin and middle of throat
white, washed with pale yellow ; breast and remainder of underparts
yellow, purer on the back and under tail-coverts, paler on the anal
region ; the sides washed with pale greyish green, the same as the
feathered tarsus ; on the middle of the lower portion of the breast a
large spot of dark purplish red; back and upper parts vivid grass-
green; primaries and their coverts on the outer web and end bril-
liant dark green, with a very narrow but distinct white margin, the
inner web black ; secondaries also dark green, but with a somewhat
broader yellow margin along the outer web ; coyverts of the secondaries
dark green, narrowly margined externally and at the end with yellow ;
larger shoulder-coverts brilliant dark green, broadly margined at the
end with grass-green ; wing beneath ashy grey ; under wing-coverts
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUs. 535
delicate grey like the neck ; tail dark green with a broad greyish-
white end ; the feathers, except the two middle ones, are on the inner
web dark grey, crossed above the white end by a blackish cross band ;
the white end is margined narrowly with pale yellow; tail beneath
dark grey, at the end broadly white ; bill plumbeous, tipped with
pale horn-colour ; feet reddish brown, nails dark ; irides yellow ;
bill brownish yellow ; legs purple-red.
“Young. Vertex and hind neck green like the back ; front and
forehead covered with a pale violet-purple patch ; sides of head
and neck grey, washed with green, the crop and upper portion of
breast with greenish yellow ; the underparts are of a less brilliant
yellow ; the sides darker greyish green ; the red pectoral patch is in-
dicated only by some dirty purplish feathers ; the pale yellow mar-
gins at the ends of the coverts of the secondaries are broader and
form a distinct cross line; the white end portion of the tail-feathers
is washed at the outer web and apex with green ; bill and feet brown ;
total length 8 inches, wing 5, tail 3°1.”’—Finsch and Hartl. (1. ¢.).
13. Prinopus rascratus.
Ptilinopus fasciatus, Peale, U.S. Expl. Exped. Ornith. (1848)
p- 193, pl.; Gray, List B. Brit. Mus. (1856) p.4; Finsch & Hartl.
Faun. Centralpolyn. (1867) p. 115; Graffe, Journ. Mus. Godeff.
Heft i. (1872) p. 45, Taf. 7. fig. 2.
Ptilinope de Clémentine, Hombr. & Jacq. Voy. Pdle Sud (1841),
pl. xxix. fig. 3.
Ptilinopus clementine, Bon. Compt. Rend. tom. xl. (1855) p- 216.
sp. 48 ; id. Consp. Gen. Av. vol. ii. p. 22. sp. 9; id. Iconogr. Pig.
pl. xxii. fig. 1 (1857) ; Reich. Taub. p- 96, tab. 240. fig. 2588;
id. Suppl. p. 203, pl. iv. figs. 34, 35 ; Gray, B. Trop. Isl. p. 38 ;
Finsch, Journ. Mus. Godeff. Heft xii. p- 37 (1876).
Columba viridissima, Temm. Pl. Col., text of C. diademata.
Ptinilopus! clementine, Jacq. & Puch. Voy. Péle Sud, vol. iii.
text (1853), p. 117, Atl. pl. xxix. fig. 3.
Kurukuru clementing, Des Murs & Prév. Voy. Vénus, Ois. p-
269 (1855).
Lamprotreron porphyracea, Bon. Consp. Gen. Av. vol. ii. p. 21.
Piilopus clementine, Bon. Iconog. Pig. (1857) pl. xxii. fig. 1.
Ptilonopus fasciatus, Gray, List B. Trop. Isl. (1859) p. 37
(partim).
Ptilonopus clementine, Gray, List B. Trop. Isl. (1859) p. 38.
Ptilopus porphyraceus, Schleg. Mus. Pays-B. 1873, p.8, Columbe.
Ptilopus fasciatus, Schleg. Mus. Pays-B. 1873, p. 6, Columbe.
Ptilonopus ponapensis, Finsch, Proc. Zool. Soc. 1877, p. 779.
Native name Manu-tagi ( Whitmee).
Hab. Samoa (Peale, Hombron and Jacquinot) ; Ponapé Island
(Kuébary).
This is the species first described by Peale (/. c.) from Samoa,
and afterwards by Hombron and Jacquinot as clementine. It is a
well-marked bird, differing from all its allies in the yellowish hue
of its throat, cheeks, and breast. The abdomen is pure green, with
536 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
a purplish-black spot in the centre. Several examples are before
me, including the type of Pt. clementine ; and the species is readily
recognizable from all the others of this especial section of the genus
Ptilopus. Pt. apicalis, Bon., sometimes referred to this species,
is quite distinct. I place with a doubt, among the synonyms given
above, the Ptilinopus fasciatus figured in the Journal of the Museum
Godeffroy (/. c.); for the colouring of the figure is so defective that
it is impossible to arrive at any satisfactory conclusion as to what
species is meant. The purplish-black mark on the abdomen is
apparently omitted. If the specimen does not possess this, it cannot
be the P¢. fasciatus, and is probably Pt. apicalis, Bon. An ex-
amination of the bird itself could alone decide the matter.
Dr. Finsch, in a late number of these ‘ Proceedings’ (1877, p. 779),
described a Ptilopus from the island of Ponapé from specimens sent
to the Museum Godeffroy by Mr. Kubary, as distinct under the
name of Pt. ponapensis. Not being able to make out by the descrip-
tion wherein this bird differed from the Pt. fasciatus, Peale, I wrote
to Dr. Finsch asking if it was not possible to have the type sent to
me. On his making the request to the Directors of the Museum
Godeffroy, they most kindly forwarded me three specimens, among
which was the type of the description published in this Journal.
I compared it with the examples in the Paris Museum ; and, as I
supposed it would be, the species is the same as Pt. fasciatus,
Peale, being identical with those birds in the Paris Museum called
clementine by Hombron and Jacquinot, and procured by them in
the Voyage au Péle Sud, and which do not differ specifically from
the Pt. fasciatus, Peale. The Pt. ponapensis will therefore have to
become a synonym of the present species.
Adult. Top of head and forehead bright purple, margined with
pale yellow. Back and sides of neck, cheeks, upper part of back,
throat, and breast pale yellowish green inclining to ash colour. Flanks
and abdomen grass-green, with a purplish-black spot in the centre
of the abdomen. Back and upper tail-coverts bronzy green.
Primaries and secondaries black on inner webs ; outer bright green,
edged with yellow on the secondaries ; first primary abruptly and
greatly attenuated. Some of the innermost secondaries and tertials
have a pale violet mark near their tips. Tail bright green, with the
terminal fourth bright yellow. Crissum and under tail-coverts
orange-yellow. Bill black, tip yellowish. Feet brown. Total
length 83 inches, wing 54, tail 33, culmen2. (From Hombr. & Jacq.
specimen in Paris Museum.)
14. PriLopus MERCIERI.
Kurukuru mercieri, Des Murs & Prév. Voy. Vénus, Ornith. p.
266 (1855).
Ptilopus mercieri, Bon. Compt. Rend. tom. xl. (1855) p. 216 ;
id. [conog. Pig. pl. xxii. fig. 2 (1857); id. Consp. Gen. Av. vol. ii.
(1857) p. 22. sp. 8.
Ptilinopus mercieri, Finsch & Hartl. Faun. Centralpolyn. (1867)
p. 128; Finsch, Proc. Zool. Soc. 1877, p. 410.
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 537
Ptilonopus merciert, Gray, List B. Trop. Isl. (1859) p. 36;
Reich. Tauben, p. 96.
Hab. Nuka-hiva, Marquesas Islands. (Type.)
A very rare and well-marked species; but one or two examples
besides the type in the Paris Museum are, I believe, all that are
known in European collections. It resembles in its coloration the
last species, but is readily recognized by the red spot on the head at
the base of the mandible.
Adult. Entire upper part of head and occiput and a conspicuous
spot at base of the mandible deep rosy purple. Nape, mantle, sides
. of neck, throat, and breast pale yellowish green, with a tinge of ash
upon cheeks and nape. Abdomen and flanks bright greenish yellow ;
crissum and under tail-coverts bright yellow. Back and upper tail-
coverts bright green, with a yellowish tinge upon the latter. Wings
same colour as the back; primaries black, shaded with very dark
green upon the outer webs, the first abruptly and greatly attenuated ;
secondaries black on the inner webs, dark green on the outer, and
margined with yellow; the innermost ones with a conspicuous blue-
black spot near the tips, and margined broadly with yellow. Rec-
trices smoke-colour at base, the outer webs barred with pale green
in the centre, this colour sometimes extending slightly onto the
inner web; and all the apical third very light yellow. Bill
black with light tip. Feet brown. Total length 9 inches, wing
o4, tail 33, culmen 3. (Description taken from type in Paris
Museum.)
15. Prinorus ROSEICAPILLUS.
Columba roseicapilla, Less. Trait. Ornith. p. 472. sp. 44 (nec
syn.) (1831).
Le Kurukuru des Martannes, Less. Compl. Buff. tom. iii. Ois.
p- 33 (1837).
Ptilonopus purpureicinctus, Gray, Proc. Zool. Soe. 1853, p. 48,
pl. lv; id. List B. Brit. Mus. (1856) p. 4. sp. 5 (partim); id.
Ann. & Mag. Nat. Hist. 2nd ser. vol. xv. (1855) p. 159; id. List
B. Trop. Isl. (1859) p. 37.
Kurukuru roseicapilla, Des Murs & Prév. Zool. Voy. Vénus, Ois.
p. 259 (1855).
Piilopus roseicapillus, Bon. Compt. Rend. tom. xl. (1855) p.
216. sp. 46; id. Iconog. Pig. (1857) pl. xxii. ; id. Consp. Gen: Av.
vol. ii. p. 21. sp. 7.
Ptilonopus roseicapillus, Gray, List B. Trop. Isl. p- 37; Reich.
Taub. p. 96.
Ptilinopus roseicapillus, Finsch & Hartl. Faun. Centralpolyn.
(1867) p. 127.
Hab. Ladrone Islands or Marianne Islands.
This is a very distinct species, differing in certain particulars from
all others known. Besides Lesson’s type there are several examples
in the collection of the Paris Museum. It is a rare species, net
many examples having been brought to Europe.
Proc. Zoo. Soc.—1878, No. XXXV. 33
538 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Adult. Front, crown, and spot at base of lower mandible rich
purple, margined on the posterior part with yellow. Back and sides
of neck greenish grey. Throat white. Upper part of breast greyish
green, the feathers tipped with pearly white. Lower part of breast
crossed by a green band, with a large spot of deep purple in the
centre; abdomen orange, with the centre yellowish green ; under
tail-coverts bright yellow, becoming rich orange towards their tips.
Back and upper tail-coverts bronzy green. Wings like the back ;
primaries black on inner webs, green on the outer, edged with yellow,
the first one abruptly and greatly attenuated ; secondaries bluish
green margined with lemon-yellow. Rectrices pale green on outer
webs, smoke-colour on their inner, with a broad terminal band of
yellowish white. Billblack, tip yellowish. Total length 83 inches,
wing 51, tail 34, culmen 3. (Specimen figured by Bonaparte
l. ec. in Paris Museum. Lesson’s type of this species is also before
me; but not being in so good a condition as Bonaparte’s specimen,
I have preferred not to take my description from it.)
16. PriLopUus HUTTONI.
Plilinopus huttoni, Finsch, Proc. Zool. Soc. 1874, p. 92.
Hab. Island of Rapa or Opara, South Sea. |
I do not know this bird autoptically. It is evidently a very fine
and distinct species, differing in many ways from all others of this
genus. It is quite large, has the first primary attenuated, and also
two thirds of the basal portion of the tarsus feathered. As I
have shown, however, in the discussion of the genera in the earlier
part of this paper, these two characters are not of any especial im-
portance in defining the generic position of the members of the group
now under consideration. The following. description is taken from
that given by Dr. Finsch of the type.
Adult, Front and crown and spot at base of mandible purplish
violet-red ; chin red, but paler than the crown. Red of crown
margined narrowly and indistinctly with green. Occiput, neck,
throat, and crop ashy grey tinged with green ; feathers on the crop
bifurcate, and dull olive-green in the centre. A pale yellowish
longitudinal patch below the chin. A large dark purplish-violet
patch on the breast ; base of feathers dark olive-green. Sides of
breast and flanks dark olive-green. Anal region and back of thighs
dirty yellow. Under tail-coverts purplish violet-red like the crown,
basal portions yellowish. Upper parts dark olive grass-green. Quills
dark green, margined narrowly with yellow, with apical portions
having coppery reflections. Shoulder-coverts with a triangular spot
of dull lilac. Tail-feathers dark green; apical margin yellowish
white, with coppery reflections in certain lights. _ Bill horny grey,
tip yellowish ; feet pale, apparently yellow; eye pink. Total length
12-20 inches, wing 6°70, tail 4°90; bill above *75, at gape 1°15; tarsus
1:20; middle toe *95, nail °35.
1878.] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 539
17. PrtLopus DUPETIT-THOUARSI.
Ptilinope de dupetit-thouars, Hombr. & Jacq. Voy.Péle Sud, pl.
29.4.1.
Columba dupetithouarsii, Neboux, Rev. Zool. 1840, p. 289 ; id.
Voy. Favor. pl. 7.
Columba kurukuru purpuro-leucocephalus, Hombr. & Jacq. Ann.
Se. Nat. xvi. p. 316 (1841).
Ptilinopus emilie, Less. Echo du Monde Sav. (1844) p. 873.
Ptilonopus leucocephalus, Gray, List B. Brit. Mus. p. 2 (1844),
Galline.
Ptilonopus dupetit-thouarsi, Gray, Gen. B. vol. ili. App. p. 23
(1844-49) ; id. List B. Brit. Mus. (1856) p. 3, Columbe ; id. B.
Trop. Isl. p. 36 (1859).
Ptinilopus dupetit-thouarsi, Jacq. & Puch. Voy. Pdle Sud, vol.
ill. (1858), text, p. 114, atl. pl. 29. fig. 1.
Trygon dupetit-thouarsi, Reich. Syst. Av. p. xxvi.
Thouarsitreron leucocephala, Bon. Compt. Rend. tom. xxxix.
(1854) p. 276, and tom. xl. (1855) p. 216 ; id. Iconogr. Pig. (1857)
pl. 17; id. Consp. Av. p. 16 (1857).
Columba dupetit-thouarsi, Des Murs & Prév. Voy. Vénus, Zool.
(1855) p. 241.
Thouarsitreron dupetit-thouarsii, Reich. Tauben, p. 93, pl. 240.
fig. 2587.
Ptilinopus dupetit-thouarsi, Finsch & Hartl. Faun. Centralpolyn.
(1867) p. 129; Finsch, Proc. Zool. Soc. 1877, p. 410.
Ptilopus dupetit-thouarsii, Schleg. Mus. Pays-B. 1873, p. 15,
Columbe.
Hab. Christine Island, Marquesas.
A very handsome species, not likely to be confounded with any
other of this group. It has, I believe, only been obtained on Chris-
tine Island of the Marquesas. The first primary is especially
noticeable for the great length of its attenuation.
Adult. Top of head greyish white, margined faintly with light
yellow. Throat and spot in front of the eye light yellow. Neck
and upper part of back greyish green. Feathers of the breast pale
green tipped with grey. Flanks pale green. Abdomen deep rose-
colour; crissum and under tail-coverts bright yellow. Back and
upper tail-coverts bronzy green. Wings like the back ; primaries
black, with a dark green shade on their outer webs, and the inner-
most ones bronzy green at their tips, the first one abruptly and greatly
attenuated. Secondaries—outer ones dark green on their outer webs,
dark brown on inner with bronze-green tips, innermost ones deep
blue near their tips, and all broadly margined with light yellow.
Rectrices dark green (sometimes strongly tinged with bronze), with
a broad apical yellowish-white band. Bill black, tipped with yellow.
Feet red. Total length 93 inches, wing 6, tail 33, culmen 3.
(Type of the species from Christine Island.)
540 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
18. PriLopus XANTHOGASTER.
Columba purpurata, Temm. (nec auct.) Pl. Col. livr. 43 (1824),
vide text to pl. 254, livr. 95 (1835).
Columba xanthogaster, Wagl. Syst. Av. (1827) sp. 29,
Columba.
Columba diademata, Temm. text to Pl. Col. 254, livr. 95
(1835).
Ptilinopus purpuratus, Steph. (nec auct.) Cont. Shaw’s Gen.
Zool. vol. xiv. p. 277. sp. l.
Ptilinopus flavigaster, Swains. Class. B, vol. ii. p. 347.
Le Kurukuru @ ventre jaune, Less. Compl. Buff. tom. iii. Ois.
p- 36 (1837).
Ptilonopus zanthogaster, Gray, Gen. B. (1 844-49) vol. ii. p. 446 ;
id. List B. Brit. Mus. (1856) p. 5; id. List B. Trop. Isl. p. 36
1859).
; Kurukuru wanthogaster, Des Murs and Prév. Voy. Vénus (1855),
Zool. p. 238. :
Thouarsitreron diademata, Bon. Compt. Rend. tom. xxxix.
(1854) p. 876, tom. xl. (1855) p. 216; id. Iconog. Pig. pl. 18
(1857); id. Consp. Av. vol. ii. p. 16 (1857); Reich. Tauben, p.
93, figs. 1326, 1327.
Ptilonopus diadematus, Wall. Ibis, 1865, p. 380.
Ptilinopus diadematus, Finsch & Hartl. Faun. Centralpolyn.
(1867) p. 124.
Ptilopus diadematus, Schleg. Mus. Pays-B. 1873, p. 11,
Columbe.
Ptilopus wanthogaster, Salvad. Ann. Mus. Civ. Genov. 1876,
vol. ix. p. 199.
Hab. Banda (Reinwardt, Hoedt); Khoor or Kanaloor, Little
Ké, Grand Ké (Von Rosenberg); Lettie, east of Timor ( Hoedé).
This species was first called by Temminck Columba purpurata,
Lath., in the 43rd livraison of the ‘ Planches Coloriées,’ he sup-
posing that it was a female of Latham’s bird. Afterwards he
ascertained, from an examination of the specimens obtained by Quoy
and Gaimard during the voyage of the ‘ Uranie,’ that the bird figured
by him represented a distinct species ; and he therefore, in the 95th
livraison, issued a new sheet of text calling the bird diademata, and
requested that the previous page should be replaced by this one.
However, between the dates of these two livraisons (1824-1835),
Wagler had in 1827, in his ‘ Systema Avium,’ described the species
as xanthogaster, which name will of course take precedence over
Temminck’s later one of diademata.
Adult. Head, neck, and upper part of the breast greyish white.
An orange-yellow line passes over the eye and encireles the occiput.
Throat bright yellow. Lower part of breast, abdomen, and under tail-
coverts orange-yellow ; a large pale grey patch on upper part of abdo-
men. Upper parts dark green. Centre of tertials, scapulars, and
innermost secondaries deep blue, bordered with yellow. Primaries
bordered with white, the first one abruptly and greatly attenuated.
Tail dark green, with an apical band of lighter green. Bill green,
1878.] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 541
tip yellow; feet dark red. Total length 9 inches, wing 54, tail
33, culmen 4.
19. Prinopus CHRYSOGASTER.
Ptilonopus chrysogaster, Gray, Proc. Zool. Soc. 1853, p. 48;
id. Ann. & Mag. Nat. Hist. 2nd ser. vol. xv. (1855) p. 158; id.
List B. Brit. Mus. (1856) p. 5. sp. 10, Columbe ; id. List B. Trop.
Isl. p. 36 (1859); Sclat. Pioc. Zool. Soc. 1864, p. 9.
Kurutreron chrysogaster, Bon. Compt. Rend. (1854) tom, xxxix.
p- 876, (1855) tom. xl. p. 217; id. Iconogr. Pig. (1857) pl. 29.
fig. 2; id. Consp. Gen. Av. vol. ii. p. 26. sp. 2 (1857).
Kurutreron chrysogastra, Reich. Taub. p. 92, tab. 249 5. figs.
3525, 3526.
Ptilinopus chrysogaster, Finsch & Hartl. Faun. Centralpolyn.
(1867) p. 128; Graeffe, Journ. Mus. Godeff. 1873, Taf. 7. fig. 4.
Hab. Tonga-Tabou, Huaheine, Society Islands (Wodehouse).
Although distinct, this species is a close ally of the Pt. xantho-
gaster. It can, however, be readily recognized by its bright yellow
abdomen and under tail-coverts and light-purple ‘crown, those of its
relative being orange and greyish white respectively. It is one of
the two species known to inhabit the Society Islands.
Adult. Front and crown light-purple, margined posteriorly with
dark yellow. Rest of head, neck, mantle, breast, and flanks greenish
grey. Throat pale yellow. Abdomen and under tail-coverts bright
yellow. Back and wings bronzy grass-green. Primaries black, the
outer webs margined with dark green, the first abruptly and greatly
attenuated for one third its length; secondaries bluish green, mar-
gined with pale yellow. ‘Tail dark bronze-green, with a broad ter-
minal band of yellowish white edged with a bright yellow. Bill
black, tip yellow. Feet brown. Total length 94 inches, wing 53, tail
4, culmen 3.
20. PTILOPUS TAITENSIS.
Columba kurukuru, var. taitensis, Less. Voy. Coq. tom. i. text,
p- 297 (1826).
Columba oopa, Wagl. Isis, 1829, p. 742.
Oolumba tattensis, Less. Trait. Ornith. (1831) p. 472.
Le Kurukuru @ Otaiti, Less. Compl. Buff. tom. ii. Ois. p. 35
(1837).
Colombe de Taiti, Neboux, Rev. Zool. 1840, p. 289.
Ptilonopus taitensis, Gray, Gen. B. vol. ii. p. 466 (1844-49).
( Ptilinopus furcatus, Peale, U.S. Expl. Exped. Birds, p. 191
1848).
Kurukuru purpuratus, Des Murs & Prév. (nec auct.) Voy. Vénus,
Zool. p. 222 (1855). Spec. Paris Mus.
Kurukuru dupetit-thouarsi, Des Murs & Prév. Voy. Vénus, Zool.
pl. vii. (1855).
Kurukuru nebouxii, Des Murs & Prév. Voy. Venus, Zool. p. 253
atl, pl. vii.
542 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Kurukuru taitensis, Des Murs & Prév. Voy. Vénus, Zool. (1855)
Poco
Ptilonopus purpuratus, Gray, List B. Brit. Mus. (1856) p. 4
(partim); id. List B. Trop. Isl. (1859) p. 35 (partim); Finsch,
Proc. Zool. Soc. 1877, p. 741.
Kurutreron oopa, Bon. Compt. Rend. (1854) tom. xxxix. p. 878 ;
id. Iconogr. Pig. (1857) (text); id. Consp. Av. vol. ii. p. 26 ; Reich.
Taub. p. 92, pl. 245 6. figs. 3527-28.
Ptilinopus purpuratus, Cass. U.S. Expl. Exped. (1858) Ornith.
pl. xxx. p. 269; Finsch & Hartl. Faun. Centralpolyn. (1867) p-
122; Finsch, Proc. Zool. Soc. 1877, p. 741.
Ptilopus purpuratus, Schleg. Mus. Pays-B.1873, p.15, Columbe.
Hab. Otaheiti.
Peale says this species utters a cov-coo-coo slowly and at regular
intervals. It is not as sprightly as the Coral-Island Dove, and
delights to sit on some dead branch in the shady recesses of the
woods. In my article on Pt. purpuratus I have given my reasons
for considering that this species should not bear that name. The
fact that Latham states that the bird he describes came from
Otaheite, which is the chief reason why his name of purpuratus
was assigned to this species, loses entirely its value and force when
it is seen that no bird of the genus Péi/opus inhabits Otaheite that
answers to bis description. Having specimens from both the Society
Islands and Tonga in his possession, and having confused them in
his description, he probably did not particularly regard his specimen
or eompare it with his deseription when he stated it to have come
from Otaheite. By no stretch of the imagination can the present
species be deemed to have a “‘ purple crown.” It is ashy grey with
a lilac tinge, but in no way does it approach any shade of purple.
A series of specimens now before me from the Paris-Museum collec-
tion clearly substantiates this.
Adult. Top of head ash-grey with a tinge of lilac. Underparts
light ash-green, darkest on the abdomen. Back of neck cinereous,
washed with green. Upper parts bright green. Wings bright green ;
first primary gradually narrowed to tip ; the inner webs of secondaries
and primaries black, outer webs bright green, those of the secondaries
edged with yellow. Centre of abdomen and crissum pale yellow,
becoming bright yellow on the under tail-coverts. The tail has the
rectrices bright green on the outer webs, black on the imner, and a
subterminal interrupted greyish bar, extending only over the inner
webs. Feet violet; eyes orange; bill green at base, tip yellow.
Feet brown. Total length 9 inches, wing 53, tail 33, culmen 3.
21. Pritopus suPERBUS.
Columba superba, Temm. Pig. & Gall. tom. i. p. 277 (1813) ;
Shaw, Gen. Zool. vol. xi. pt. 1, p.64 (1819) ; Desm. Dict. Se. Nat.
vol. xl. (1826) p. 337; Temm. & Knip, vol. 1. pl. 33.
Columba cyanovirens, Less. Voy. Coq. pl. xhi. fig. 1 (1824) ;
Desm. Dict. Se. Nat. vol. xl. (1826) p. 343; Less. Trait. Ornith.
(1831) p. 471; Knip & Prév. Pig. vol. ii. pl. 8,2.
1878. MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 543
Columba superba, Wagl. Syst. Av. (1827) sp. 32, Columba.
La Verte, Less. Compl. Buff. tom. ii. Ois. p. 29 (1837).
La Poukiopou, Less. Compl. Buff. tom. iii. Ois. (1837).
Ptilinopus leucogaster, Swains. Class. B. vol. ii. p. 347.
Ptilinopus superbus, Steph. Gen. Zool. vol. xiv. p. 279. sp. 6;
Gould, B. Austr. vol. v. pl. lvii.
Ptilonopus superbus, Gray, Gen. B. (1844-49) vol. ii. p. 467 ; id.
List B. Brit. Mus. (1856) p. 7; id. Proc. Zool. Soc. 1858, p. 184;
id. List B. Trop. Isl. (1859) p. 39; id. List Mamm. and B. New
Guin. (1859) p. 46; Wall. Ibis, 1865, p. 379; Salvad. Ann. Mus.
Civ. Gen. (1875) vol. vii. p. 786.
Ptilonopus cyanovirens, Gray, Gen. B. (1844-49) vol. ii. p. 467 ;
id. List B. Brit. Mus. (1856) p. 8. sp. 23 ; id. Cat. Mamm. and B.
New Guin. (1859) p. 47.
Kurukuru superbus, Des Murs & Prév. Voy. Vénus, p. 231
1855).
; Lamprotreron superba, Bon. Consp. Gen. Av. (1857) vol. ii. p. 18;
Reich. Tauben, p. 97, pl. 23646. fig. 3366, 9, pl. 236. figs. 1316-17,
¢ juv.; Ibis (1866), p. 333; Ramsay, Proc. Zool. Soc. 1876,
» 114.
; Cyanotreron cyanovirens, Bon. Compt. Rend. (1854) tom. xxxix.
p- 878 ; id. Consp. Av. vol. ii. p. 23; Reich. Tauben, p. 98, pl. 238.
fig. 1324.
Ptilonopus (Lamprotreron) superbus, Gray, Proc. Zool. Soc.
1860, p. 360.
Piilopus superbus, Schleg. Mus. Pays-B. 1873, p. 30, Columbe;
Salvad. Ann. Mus. Civ. Genov. (1877) vol. x. p. 158.
Lamprotreron porphyrostictus, Gould, Ann. & Mag, Nat. Hist.
(1874) vol. xiii. p. 137, 2 ?
Hab. Amboyna, Ceram, Batchian, Gilolo, Waigiou, Mysol, Aru,
New Guinea (Wallace); Mount Arfak (Bryjn.); Halmahera,
Ternate, Batanta, Jobie (Bernstein, Von Rosenberg); Bouru, Sal-
watty (Hoedt) ; Amberbaki, Mafor (Raffray) ; New Hanover (Ca-
banis); Booby Islands (Gould) ; Cape York, Australia (Cockerell).
As will be seen from the list given above, this species has received
many names. The two sexes have been described as distinct species,
and have been assigned to different genera! It has a wide dis-
tribution, having been procured as far to the eastward as New
Hanover, reaching Gilolo on the north, and Cape York, Australia,
on the south. In fact it is found generally throughout the Papuan
group of islands and along the northern part of New Guinea. As
it has been obtained at Mount Arfak and Amberbaki, and also on Sal-
watty on the west coast, it is probable that it will be ascertained to
inhabit the entire northern portion of New Guinea. According to
Mr. Ramsay, this bird is tolerably abundant in the scrub-lands of
the Herbert River and coast-range. The note, which is a rolling
guttural ‘* coo,’ may be heard half a mile off. The young resemble
somewhat those of P¢. swainsoni (regina), but do not show the bifur-
cate breast-feathers before the end of the second year ; but the green *
bands on the flanks are conspicuous even in the nestling. The
544 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Lamprotreron porphyrostictus of Gould seems to be only a female of
this species, having “‘a purplish-blue spot at back of head.”
Adult male. Upper part of head bright purple. Ear-coverts and
sides of face light green. Chin and throat ashy white. Nape and
sides of neck above shoulders rufous-orange. Breast has the feathers
light purple at the base, broadly tipped with ashy grey, this latter
hue giving the predominant colour to the breast. A broad bluish-
black band crosses the lower part of the breast. Middle of abdomen
and crissum white. Flanks and thighs green, the former crossed by
a white band above the thighs. Upper parts bright green. Shoulders
and a spot on the scapulars, wing-coverts, and secondaries deep
purple. Primaries brownish black, the first abruptly attenuated.
Secondaries bright green, edged with light yellow. Tail has the rec-
trices blackish on the inner webs, bright green on the outer, and all
except the central pair with a white apical band. Under tail-coverts
white, tinged with yellow, with broad central green stripes. Bill
olive-green, tip yellowish. Feet pink; claws pale. Iris yellow.
Total length 93 inches, wing 53, tail 4, culmen 8.
Specimen described from Ternate. Amboyna examples seem to
be bright pale orange on the nape and sides of neck.
Female. Uniform green above. A spot of deep greenish blue on
the occiput. Throat greyish white; breast greyish green. Middle
of abdomen white.
Young. Uniform yellowish green, tinged with grey upon the
breast.
22. PrirLorus TEMMINCKI.
Columba superba, Kuip & Prév. Pig. tab. 42, 2 , ex Celebes (1838).
Kurukurutemminchii, Des Murs & Prév. Voy. Vénus, p. 234 (1855).
Ptilonopus formosus, Gray, Proc. Zool. Soc. 1860, p. 360; Wall.
Ibis, 1865, p. 379; Wald. Trans. Zool. Soc. vol. vili. (1872)
p- 82.
Ptilopus superbus, Schleg. Mus. Pays-B. 1873, p. 32, Columba,
ex Celebes.
Megaloprepia formosa, Salvad. Ann. Mus. Civ. Genov. (1876)
vol. ix. p. 199.
Hab. Celebes.
This form, from the island of Celebes, possesses quite sufficient
difference in coloration to entitle it to be considered distinct from
Pt. superbus. The nape and sides of the neck are a much brighter
orange ; and the lower part of the breast is pale purple, graduating
insensibly into the bluish-black band that crosses the body above
the abdomen. An immature bird of this species from Celebes was
figured by Knip & Prévost (/.c.) as the female of Pt. superbus ;
but afterwards, convinced that it represented a distinct species, they
called it temminckii in the Voyage of the Vénus (J. c.), which name
antedates the formosus of Gray. The figure originally published by
these authors does not represent well the differences existing between
this species and Pt. superbus, on account of the specimen not having
yet assumed its perfect livery; but the locality, Celebes, proves
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 545
that their bird cannot belong to the older known form. Gray did
not well indicate the distinctive characters of this species in the
‘Proceedings’ of this Society when he conferred a name upon
it (J. c.) ; but it was properly and completely described by Wallace
afterwards in ‘ The Ibis’ (/.¢.). It can readily be distinguished from
its ally by the purple breast, the general plumage of the two species
being the same, as are also the dimensions.
The female resembles that of Pt. superbus, with rather a deeper
coloration.
23. PTrILOPUS PEROUSEI.
Ptilopus perousei, Peale, U.S. Expl. Exped. (1848) p. 195, pl. 54;
Cassin, ibid. (1858) 2nd ed. p. 274, pl. 33; Finsch & Hartl. Faun.
Centralpolyn. (1867) p. 110; id. Proc. Zool. Soc. 1869, p. 548 ;
Finsch, Journ. fiir Ornith. 1872, p. 44; Schleg. Mus. Pays-Bas
1873, p. 10, Columbe.
Ptilinope de Marie, Hombr. & Jacq. Voy. Pole Sud, pl. 29. fig. 2.
Columba kurukuru superba, Hombr. & Jacq. Ann. Scien. Nat.
(1841) tom. xvi. p. 316.
Ptinilopus marie, Jacq. & Puch. Voy. Péle Sud (1853), tom. iii.
p- 115.
Kurukuru samoensis, Des Murs & Prév. Voy. Vénus, Zool. p. 247.
Type examined.
Ptilopus maria, Bon. Compt. Rend. (1854) tom. xxxix. p. 877 ;
id. Iconog. Pig. (1857) pl. 26, ad. et juv.; id. Consp. Av. vol. ii.
p. 22. sp. 10.
Ptilonopus marie, Gray, List Columb. (1856) p. 4; Reich. Tauben,
p- 96, pl. 240. fig. 2586 ; id. List B. Trop. Isl. p. 37 (1859).
Piilinopus cesarinus, Hart]. Journ. fiir Ornith. 1864, p. 413, juv.
Ptilinopus peroysei, Cass. Expl. Exped. (1858) p. 274, pl. 33 ;
Reich. Tauben, Supp. p. 203, nov. taf. 38-40; Finsch & Hartl.
Faun. Centralpolyn. (1867) p. 110, t. xi. fig. 1; Griffe, Journ. fir
Ornith. 1870, pp. 122 & 408; Finsch, Journ. fiir Ornith. 1872,
. 44,
- Ptilinopus maria, Finsch & Hartl. Journ. fiir Ornith. 1870, p. 131.
Ptilonopus perousei, Whitmee, Ibis, 1875, p. 441.
Hab. Ovalau (Filhol) ; Upolu, Hapai (Griéfe) ; Samoan Islands,
Viti-Levu (Gréfe); Wakaia, Mokani, Vanua Levu, Loma Loma,
Kandavu, Mango (Layard); Fiji Islands; Tonga or Friendly
Islands.
This species was first described by Peale (/.c.) under the name
here adopted, and afterwards by Hombron and Jacquinot (/.c.) as
«* Ptilinope de Marie,”’ still later latinized by Jacquinot and Pucheran
(1. c.) as Ptinilopus! marie. Peale’s appellation, however, has un-
doubted priority. Mr. Peale found it in the mountain-gorges of the
Feejee Islands, but not abundant. The native name, Manu-ma,
means shame or modest bird. It is one of the most beautiful and
curiously marked members of this genus, The young are so dif-
ferently coloured from the adult that they might easily be mistaken
for a different species.
546 MR. D. G. ELLIOT ON THE GENUS PTILoPUS. [May 7,
Adult, Front and crown, and a broad band across the back, dark
purplish red. Head, neck all around, and throat white with a wash
of yellow. Feathers of the breast, split at their ends, have their
basal half rose-red, apical portion yellowish white ; beneath these
feathers is a reddish-orange band. Abdomen, flanks, and crissum
yellowish white. Under tail-coverts dark purplish red. Back and
upper tail-coverts pale greenish yellow. Wings pale grey, feathers
margined with greenish yellow. Primaries and secondaries dark
bronzy green; first primary abruptly attenuated. Tail greyish
white. Bill black, with the tip yellow. Feet black.
Total length 82 inches, wing 53, tail 33, culmen 3.
Immature. Front and crown deep rosy purple. Cheeks grey.
Upper parts shining golden green. ‘Throat white. Breast green,
feathers tipped with white. Abdomen and crissum yellowish white.
Under tail-coverts bright yellow. Wings shining golden green ; pri-
maries and secondaries margined with yellow. ‘Tail golden green,
very bronzy on the edges and tips of the feathers. Bill and feet as
in the adult. (Four specimens before me, all of which lack the
purple bar on the breast shown in the figures of Bonaparte and
Hartlaub and Finsch.)
24. PrILOPUS CORONULATUS.
Ptilonopus coronulatus, Gray, Proc. Zool. Soc. 1858, p. 185,
pl. 138; id. List Mamm. & Birds, New Guinea (1859), p. 45 ; Wall.
Ibis, 1865, p. 378; Schleg. Mus. Pays-Bas 1873, p. 9, Columbe ;
Salvad. Ann. Mus. Civ. Genov. vol. vii. (1875) p. 833.
Ptilopus coronulatus, Salvad. Ann. Mus. Civ. Genov. (1876)
vol. ix. p. 198.
Hab. Aru Islands (Wallace, Rosenberg) ; Salwatty and Sorong,
(Bernstein) ; Jobie (Rosenberg) ; Yule Island (D’ Albertis).
This species and the two next mentioned form a small section of
this genus, related closely to each other in the distribution of the
hues of their plumage. Like so many others of the genus, they are
conspicuous for their brightly-coloured crowns and abdomens.
Male. Front and crown pink, bordered posteriorly by a narrow
line of purple, succeeded by a broader one of bright yellow. A
purple spot on the upper part of the abdomen; middle of abdomen,
crissum, and under tail-coverts bright yellow. Entire rest of plumage
bronzy green. Secondaries and primaries margined with yellow.
First primary abruptly attenuated at tip. Bill greenish yellow. Feet
red. Iris orange.
Total length 73 inches, wing 43, tail 27, culmen 2.
Female like the male.
25. PTILOPUS GEMINUS.
Ptilonopus geminus, Salvad. Ann. Mus. Civ. Genoy. vol. vii.
(1875) p. 786.
Ptilonopus senev, Briigg. Abhandl. natur. Ver. Brem, 1876,
p. 82.
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 547
Ptilopus geminus, Salvad. Ann. Mus. Civ. Genov. vol. ix. (1876)
p- 198.
Hab. Sobie (Von Rosenberg, Bruijn); Krudu (Beccari) ; New
Guinea, near Dorey (Bruijn) ; Ansus (Salvadori).
This is a species apparently entitled to be considered distinct from
the Pt. coronulatus. It differs by having the front and crown pinkish
white, and the upper part of the abdomen around the violet spot saffron-
yellow. The cheeks are also a dark greenish grey. First primary
abruptly attenuated at tip. Size the same as Pt. coronulatus.
Prof. Schlegel, in Mus. Pays-Bas, was the first to point out the dif-
ferences existing between these two birds, but did not deem them of
specific importance; and Dr. Salvadori bestowed the name here
adopted.
26. PriLorus TRIGEMINUS.
Pétilonopus trigeminus, Salvad. Ann. Mus. Civ. Gen. vol. vii.
(1875) p. 787.
Ptilonopus marginalis, Brigg. Abhandl. natur. Ver. Brem.
1876, p. 82.
Ptilopus trigeminus, Salvad. Ann. Mus. Civ. Genov. vol. ix.
(1876) p. 198.
Hab. Salwatty (Von Rosenberg); New Guinea, near Sorong
(Bernstein).
The claims of this form to be considered distinct from the last
are based upon very slight grounds ; and it may well be doubted if it
should be separated from Pt. geminus. The only difference is in the
slightly paler crown. At best it seems to be but a local race of the
preceding species.
27. PTILOPUS IOZONUS.
Ptilonopus iozonus, Gray, Proc. Zool. Soc. 1858, p. 186; id.
List Mamm. & Birds New Guin. (1859) p. 46; Wall. Ibis, 1865,
p- 378.
Ptilopus iozonus, Schleg. Mus. Pays-B. 1873, p. 17, Columbe ;
id. Nederl. Tijdsch. Dierk. tom. iv. p. 25 (1873); Meyer, Rowl.
Ornith. Mise. pt. ix. p. 342.
Ptilonopus itonozonus, Salvad. Ann. Mus. Civ. Gen. vol. vii.
(1875) p. 834.
Piilopus ionozonus, Salvad. Ann. Mos. Civ. Gen. vol. ix. (1876)
p- 198.
Hab. Aru (Wallace, Von Rosenberg) ; New Guinea (D’ Albertis).
We have here, in this species and its two relatives, another
small section of this genus. One of their chief differences is the ar-
rangement of the colours on the margin of the wing, and another
the manner in which the terminal band on the tail is exhibited.
Although closely allied, the species seem to be quite distinct, their
geographieal distribution also giving strength to this view.
Male. Yellowish green. Bend of wing greyish violet, mixed with
green, Middle of the abdomen deep orange. Vent and under tail-
548 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
coverts white, varied with yellow. Greater wing-coverts and tertials
bordered with yellow, the latter grey in the middle of each feather.
Quills shining deep emerald-green, first one gradually narrowed to
the tip. Under surface of wings slate-colour. Tail green, with a
broad band of grey at the tip, which is white beneath, especially on
the inner webs.
Length 8" 3! ; wings 4! 9!" (Gray, 1. c.).
Female with a patch on the abdomen of an orange-yellow.
28. PriLopus HUMERALIS.
Ptilonopus humeralis, Wall. Proc. Zool. Soc. 1862, p. 166.
Ptilopus humeralis, Meyer, Proc. Zool. Soc. Soc. 1862, pl. 21;
Schleg. Mus. Pays-B. (1873) p. 16, Columbe ; id. Nederl. Tijdsch.
Dierk. tom. iv. p. 24 (1873) ; Meyer, Rowl. Ornith. Mise. p. 342,
pt. ix.; Salvad. Ann. Mus. Civ. Genov. (1876) vol. ix. p. 198.
Hab. Salwatty and adjacent coasts of New Guinea (Wallace) ;
Sorong (Bernstein) ; Salwatty and Mansinam (Zeyer).
This bird is easily distinguished from its immediate allies by the
deep-purple band on the shoulder. It apparently ranges across the
northern part of New Guinea, having been procured, as will be
observed, from Mansinam, on the mainland, in the Bay of Geel-
vink, to Sorong, on the west coast, and also on Salwatty, opposite
Sorong.
Adult male. General plumage bright green. Middle of abdomen
rich orange. Crissum and under tail-coverts bright yellow. Shoulders
violet-grey, bounded beneath by deep purple. Wing-coverts violet-
grey, margined with green. Secondaries margined with yellow,
Primaries dark green, the first sharply narrowed towards the tip.
Tail bright green above, dark grey beneath, with an apical whitish
band. Bill greenish, tipped with yellow ; base above red and swollen.
Feet purplish red.
Total length 83 inches, wing 43, tail 25, culmen 2.
29. PriLoPpus JOBIENSIS.
Ptilopus humeralis jobiensis, Schleg. Mus. Pays-B. 1873, p. 16,
Columbe ; Rowl. Ornith. Misc. pt. ix. pp. 340, 348, pl.; Salvad.
Ann. Mus. Civ. Genov. (1876) vol. ix. p. 198.
Hab. Island of Jobie.
This species, a close relative of Pt. humeralis, has as yet only been
found upon the island of Jobie, in the great Bay of Geelvink, New
Guinea. It differs chiefly from the species just named in the colo-
ration of the shoulders of the wing, which are violet-grey, without
any purple; and the tail is uniform green above, without the apical
grey band.
Adult. General plumage yellowish green. Abdomen rich orange.
Crissum and under tail-coverts white, the latter tipped with yellow.
Shoulders violet-grey. Wing-coverts violet in the centre, surrounded
by greenish blue, and margined with green. Primaries and secon-
daries dark green, the latter edged with yellow; first primary nar-
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 549
rowed at tip. Tail bronzy green above, beneath black, with an
apical greyish-white band. Bill black, tip yellow. Feet red.
Total length 7} inches, wing 43, tail 2}, culmen 3.
30. PriLoPpus INSOLITUS.
Ptilopus insolitus, Schleg. Neder]. Tijdsch. (1861) vol. i. p. 61,
pl. 3. fig. 3.
Ptilopus humeralis jobiensis (monstr.?), Schleg. Mus. Pays-B.
1873, p. 16, Columba.
Gidirhinus globifer, Cab. & Reich. Sitzungsb. Gesellsch. nat.
Freunde Berlin 1876, p. 73; id. Journ. fiir Ornith. 1876, p. 326.
CGdirhinus insolitus, Sclat. Proc. Zool. Soc. 1877, p. 110;
Salvad. Proc. Zool. Soc. 1877, p. 196.
Hab. New Ireland (Huesker); Duke-of-York Island (Brown).
This strange and attractive species was first described by Schlegel
(l.c.). He afterwards, on account of the protuberance on the fore-
head, imagined it was only a monstrosity of the Pt. jobiensis. This,
however, is not the fact, as the species is very distinct. The locality
given by Schlegel (New Caledonia) is probably an error, as the bird
has lately been received from both New Ireland and Duke-of-York
Island. Cabanis and Reichenow have made a new genus for the re-
ception of this bird, ‘‘ @dirhinus,” on account of the swelling of the
forehead. I have not adopted this term; for the division does. not
seem to me to be necessary. The same peculiarity exists in species
of Carpophaga, viz. C. tumida and C. rubricera and others ; and if it
is a sufficient cause to separate P¢. insolitus from the members of the
genus Pti/opus, the species above mentioned should also be generically
divided from Carpophaga, a proceeding which is hardly likely to
meet with favour. Nothing is known about the habits and economy
of the present species. In the arrangement of the group it naturally
comes next to the Pé. jobiensis.
Adult. Head, neck, back, breast, and flanks bright bronzy green.
Throat green, slightly tinged with grey. Abdomen deep orange.
Shonlders anda patch on each side of the back at the edge of the mantle
light grey. Wings green; secondaries margined with bright yellow
on their outer webs; inner secondaries light grey, margined with
green. Tail bright green, with the apical third ashy grey. Crissum
and under tail-coverts bright yellow. Bill greenish at base, yellowish
at tip. Forehead and base of culmen covered by a bony protuberance
large and rounded in form, very conspicuous, and of a red colour.
Tarsi and feet red.
Total length 93 inches, wing 5, tail 3, bill at gape 3.
31. PTILoPpUS NANUS.
Columba Naina, Temm. Plan. Col. no. 565; Knip & Prév. Pig.
vol. ii. pl. 59.
La Naine, Less. Compl. Buff. tom. iii. Ois. p. 30 (1837).
Ptilonopus naina, Gray, Gen. B. vol. iii. p. 467 (1844-49),
Totreron nana, Bon. Consp. Gen. Av. vol. ii. p. 25 (1857),
550 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Tonotreron nana, Reich. Taub. p. 100, Taf. 239. fig. 1330.
Ptilonopus nanus, Gray, List Mamm. & B. New Guinea (1859),
p- 46; Wall. Ibis 1865, p. 381.
Ptilopus nanus, Schleg. Mus. Pays-B. (1873) p. 21, Columbe ;
Salvad. Ann. Mus. Civ. Genov. vol. ix. (1876) p. 196.
Hab. Triton Bay, New Guinea (Miiller) ; Mysol (Hoedt).
This is the smallest species of this genus, and one of the mest at-
tractive. It is very rare in collections, but few specimens having
been procured since it was first descrived by Temminck (J. ¢.). It
is conspicuous among its relatives from having all the wing-coverts
tipped with yellow, and from the blue-grey bands on each side of the
breast.
Male. General plumage bronzy green. Large band of bluish grey
on each side of the breast. A purple spot on the centre of the ab-
domen. Wing-coverts, scapulars, and tertials tipped with yellow, a
greenish-blue shade behind the yellow on the last two. Feathers
of the legs whitish. First primary narrowed at tip.
Wing 2 inches 11 lines, tail 1 inch 6 lines.
Female. Green, without abdominal spot.
32. PriLOPUS MONACHUS.
Columba monacha, Temm. Pl. Col. 253; Desm. Dict. Se. Nat.
tom. xl. p. 344 (1826); Knip & Prév, Pig. pl. 53; Wagl. Syst. Av.
(1827) sp. 35.
Ptilinopus monachus, Steph. Shaw, Gen. Zool. vol. xiv. (1826)
p- 277. sp. 2; Swains. Class. B. vol. ii. p. 347 (1837).
La Monache, Less. Compl. Buff. tom. tii. Ois. p. 29 (1837).
Ptilonopus monachus, Gray, Gen. B. vol. ii. p. 466. sp. 8 (1844—
49); Wall. Ibis, 1865, p. 378; Salvad. Ann. Mus. Civ. Genov.
vol. vil. p. 786 (1875).
Cyanotreron monacha, Bon. Consp. Gen. Av. vol. ii. p. 24 (1857) ;
Reich. Taub. p. 98, Taf. 236. figs. 1314, 1315.
Ptilonopus (Cyanotreron) monachus, Gray, Proc. Zool. Soc. 1860,
. 359.
: Ptilopus monachus, Schleg. Mus. Pays-B. (1873) p. 12, Columbe ;
Salvad. Ann. Mus. Civ. Genov. vol. ix. (1876) p. 198.
Hab. Batchian, Kaisa, Ternate, Gilolo, Morty (Wallace) ; Motir,
Dammar (Bernstein) ; Tidore (Raffray).
A series of this bird from different localities brought by M.
Raffray is now before me. 1 notice that those from Ternate differ
from Gilolo specimens, as well as those from Tidore &c., in being of
larger size, and also in having the blue spot of greater extent and
situated upon the abdomen, and not on the breast, as in those from
Gilolo. The type of Pt. monachus was erroneously stated to have
come from Celebes. A specimen in the Paris collection was obtained
in exchange from M. Temminck, and the locality given as Celebes.
This agrees precisely with Ternate examples, and has the blue patch
upon the abdomen. ‘Temminck’s figure in the ‘ Planches Coloriées’
exhibits this large abdominal patch very clearly. The Gilolo birds
may require separation.
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. aod
Adult male. Front, crown, lores, moustache, and a spot on the
breast greyish blue. Superciliary stripe, chin, and throat, lower
part of abdomen, and under tail-coverts lemon-yellow. Rest of
plumage of body bright green. Primaries and secondaries black on
inner, green on outer webs, edged with white or pale green ; first
primary abruptly attenuated at tip. Tail bright green, with a sub-
terminal bar of darker green. Bill greenish, tip lighter ; iris dark ;
feet red.
Total length 7 inches, wing 4, tail 23, culmen 2.
Female differs in having the front and top of head greenish blue,
the throat only slightly tinged with yellow, no spot on the breast,
rest of plumage like the male but all the colours much duller.
33. PriLopus PULCHELLUS.
Columba pulchella, Temm. Pl. Col. 564; Knip & Prév. Pig.
vol. ii. t. 14.
La Mignonne, Less. Compl. Buff. tom. iii. Ois. (1837).
Ptilonopus pulchellus, Gray, Gen. B. vol. ii. (1844) p. 466. sp. 7;
id. Cat. Mamm. Birds New Guin. (1859) p. 44; Reich. Tauben,
p- 96, pl. 236. fig. 1311; Wall. Ibis, 1865, p. 378; Salvad. Ann.
Mus. Civ. Genov. vol. vii. (1875) pp. 786, 833.
Ptilopus pulchellus, Bon. Consp. Gen. Av. vol. ii. p. 22 (1857) ;
Schleg. Mus. Pays-B. 1873, p. 2, Columbe; Salvad. Ann. Mus.
Civ. Genov. (1876) vol. ix. p. 198, & vol. x. (1877) p. 158.
Hab. New Guinea, Salwatty, Waigiou (Bernstein) ; Mysol ( Wal-
lace); Amberbaki, Andai (Ruffray); Monte Epa, New Guinea
(D Albertis).
A very brightly coloured and charming little species. From the
localities in which it has been procured, we should judge its range
was the northern part of New Guinea, with some of the islands lying
off the west coast. Its synonymy is very simple, the species never
having received a second name.
Male. Front and crown bright carmine. Sides of head and neck,
together with the breast, light grey. Chin, throat, and lores white.
A large spot or bar of purplish red on the lower part of the breast.
Flanks green. Abdomen orange, shading into deep yellow upon the
lower parts and crissum. Under tail-coverts orange. Entire upper
parts and back of neck bright green. Wings green; primaries and
secondaries black on inner webs, green on outer, those of the secon-
daries edged with yellow; first primary abruptly attenuated at tip.
Tail bright green. Bill yellow, tip greenish yellow. Feet dull car-
mine. Iris orange, paler within. Eyelids yellow.
Total length 8 inches, wing 43, tail 33, culmen 74.
There is no difference between the sexes in plumage.
34. PrrLoPpUS MELANOCEPHALUS.
Columba melanocephala, Gmel. Syst. Nat. tom. i. p. 781 (1788) ;
Temm. Pig. & Gall. tom. i. (1813) p. 263; Horsf. Trans. Linn.
Soc. 1821, vol. xiii. p. 182; Wagl. Syst. Av. (1827) Columba,
552 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
sp. 33; Temm. Pl. Col. 214; Knip & Temm. Pig. vol. i. pl. 30;
Less. Trait. Ornith. (1831) p. 471.
Tourterelle de Batavia, Buff. Plan. Enlum. 214.
Columba melanocephala, Shaw, Gen. Zool. vol. xi. pt. | (1819).
Ptilinopus melanocephalus, Steph. Shaw, Gen. Zool. vol. xiv.
p- 279. sp. 7; Wall. Ibis, 1865, p. 381.
Le Turgis, Less. Compl. Buff. tom. iii. Ois. p. 26 (1837).
Ptilonopus melanocephalus, Gray, List B. Brit. Mus. (1844) p. 1,
(1856) p. 6. sp. 15; id. Gen. B. vol. ii. p. 467 (1844-49).
Ionotreron melanocephala, Reich. Taub. p. 100, Taf. 237. figs.
1318, 1319.
Totreron melanocephala, Bon. Consp. Gen. Av. vol. ii. p. 24. sp. 2
(1859).
Ptilopus melanocephalus, Schleg. Nederl. Tijdsch. Dierk. vol. it.
p- 207; id. Mus. Pays-B. 1873, p. 28, Columbe.
Ptilopus melanauchen, Salvad. Ann. Mus. Civ. Genov, vol. vii.
(1875) p. 670 (ex Flores).
Totreron melanocephala, id. ibid. p. 671.
Hab. Java (Diard, Temminck); Sumbawa, Flores (Forstein,
Rosenberg) ; Sulabessie (Hoedt); Lombock (Wallace).
Count Salvadori has called the birds from Flores (/. c.) Péelopus
melanauchen; but I really cannot see that this island form differs
sufficiently from typical Pt. melanocephalus, ex Java, to require sepa-
ration as a distinct species. It would seem that four races of this
style of Péilopus are quite sufficient to indicate the varieties that
exist. This species is one of the longest-known in the genus; and
while the typical style is confined to Java and the Timor group of
islands, representative forms, constituting races if not distinct species,
are met with on other islands of the Malay archipelago.
Male. Top and sides of head and neck, and lower part of throat
light grey ; occiput and back of neck black ; chin and throat yellow ;
crissum and short under tail-coverts rich yellow, long ones carmine ;
entire rest of plumage bright green; primaries black on inner web,
green on outer; first primary slightly scalloped and narrowed at
tip; bill yellow, greenish horny at the tip; feet pink-red ; orbits
bare, green.
Total length 93 inches, wing 43, tail 3, culmen 3. (Java.)
Female. Entirely green, with the exception of the longest feathers
of the under tail-coverts, which are carmine, like those of the male.
35. PriLOPUS MELANOSPILUS.
Ptilopus melanocephalus, Schleg. Mus. Pays-B. 1873, p. 29, ex
Celebes.
Totreron melanospila, Salvad. Ann. Mus. Civ. Genov. vol. vii.
(1875) p. 670.
Ptilinopus melanocephalus celebensis, Brigg. Brem. Abhandl.
(1876) p. 80.
Hab. Celebes.
The various forms of Pé#. melanocephalus were pointed out by
1878 .] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 553
Prof. Schlegel (/. c.), and names afterwards given to them by M.
Salvadori. The principal difference between examples from Gelebes
and Java appears to be that the former have a much narrower throat-
mark, which is lemon-yellow washed with orange.
36. PriLopus CHRYSORRHOUS.
Ptilopus melanocephalus, Wall. Proc. Zool. Soc. 1862, p-» 344 ;
Schleg. Mus. Pays-B. 1873, p. 29, ex Sula.
Totreron chrysorrhoa, Salvad. Ann. Mus. Civ. Gen. (1875) vol. vii.
p- 671.
Ptilonopus sulaensis, Brigg. Abhandl. natur. Ver. Brem. 1876.
Hab. Sula, Ceram (Bernstein).
This form has the throat-mark as in specimens from Celebes.
The black band on the occiput is smaller, and the abdomen orange-
yellow. The characters it offers to support its claim to a distinct
rank are not very striking ; and it may be questioned if it should be
really separated from Pt. melanocephalus.
37. PritLopus XANTHORRHOUS.
Ptilopus melanocephalus, Schleg. Mus. Pays-B. 1873, p. 29,
ex Sanghir.
Lotreron «anthorrhoa, Salvad. Mus. Civ. Gen. vol. vii. (1875)
p- 670.
Ptilinopus nuchalis, Briigg. Brem. Abhandl. 1876, p. 80.
Hab, Island of Sanghir.
This well-marked race of Pt. melanocephalus differs, both in size
and colour of plumage, so much that, more than any of the other
styles, it deserves a separate name. The colour of the throat is
of a deeper yellow ; and both abdomen and crissum are orange; and
this colour runs up nearly to the breast instead of being confined to
the short under tail-coverts and crissum. It is much larger than
any of the other races.
Male. Head and neck pearly white. Occiput and back of neck
Jet-black. Throat orange-yellow. Abdomen and crissum bright
orange; long feathers of the under tail-coverts rich carmine. Entire
rest of plumage bright yellowish green. Primaries and secondaries
margined on their outer webs, near the tips, with lemon-yellow-
First primary gradually narrowed towards tip. Bill black, tipped
with yellow. Feet very dark red.
Total length 10 inches, wing 5}, tail 33, culmen 3.
Female. Entirely green. Feathers of the abdomen tipped with
yellow. Some feathers of the under tail-coverts tipped with yellow,
others with carmine. Bill black. Feet dark red.
38. PriLopus PORPHYREUS.
Columba porphyrea, Temm. Pl. Col. 106 (1824); Knip & Préy.
Pig. vol. ii. pl. iv.; Less. Trait. Ornith. (1831) p. 472.
Ptilinopus porphyreus, Swains. Zool. Journ. vol. i. p. 474 (1824—
25); Steph. Shaw, Gen. Zool. vol. xiv. (1826) p. 277. sp. 3.
Proc. Zoo. Sec.—1878, No, XXXVI. 36
5a4 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
Columba roseicollis, Wagl. Syst. Av. (1827), Columba, sp. 30.
La Porphyre, Less. Compl. Buff. tom. iii. Ois. (1837).
Ptilonopus erythrocephalus, Swains. Class. B. vol. ii. p. 347 (1839).
Ptilonopus roseicollis, Gray, Gen. B. vol. ii. p. 467 (1844-49) ;
id. List B. Brit. Mus. (1844) p. 1, (1856) p. 8.
Ptilopus porphyrea, Reich. Taub. p. 97, pl. 237. figs. 1322-23.
Ptilonopus porphyreus, Wall. Ibis, 1865, p. 380.
Ptilopus porphyreus, Schleg. Mus. Pays-B, 1873, p. 33, Columbe.
Hab. Java (Wallace, Diard).
This is a rather large and brightly plumaged species, with a long
square tail, approaching somewhat in form Pt. cinetus, Pt. bern-
steini, &e. The first primary is abruptly attenuated at the tip. It
is apparently restricted to Java, all the specimens that I have seen
having come from that island. There is considerable difference in
the colour of the plumage of the adult and young, the latter almost
entirely wanting the rose or carmine on the head and breast which
makes the adult so conspicuous. There does not appear to be any
difference in the plumage of the sexes.
Adult. Head rose-colour, graduating into carmine upon the neck
and breast. Beneath the red colour on the breast is a rather broad
white band, below which is another of blackish green. Abdomen
and flanks clear grey. Thighs and crissum bright green, feathers
tipped with bright yellow. Under tail-coverts greenish on the inner
webs, white on the outer, with the margins light yellow. Entire
upper parts dark yellowish green, lightest on the rump. Primaries
black, the first abruptly attenuated at tip. Tail dark green, with a
broad dull grey apical band. Bill black, with pale tip. Feet red.
Total length 11 inches, wing 53, tail 43, culmen 2.
Young. Have a generally green plumage, with the feathers mar-
gined with yellow, and the breast mottled with purplish red.
39. PriLoPpus JAMBU.
Columba jambu, Gmel. Syst. Nat. (1788) vol. i. p. 784; Rafi.
Trans. Linn. Soc. (1822) vol. xii. p. 212 ; Wagl. Syst. Av. (1827)
sp. 28, Columba.
Columba jambos, Lath. Ind. Ornith. (1790) vol. i. p. 598. sp.
18; Less. Trait. Ornith. (1831) p. 472.
Columba jamboo, Temm. Pig. et Gall. (1813) p. 257; Temm.
& Knip, Pig. vol. i. pl. 27 9, 28 ¢; Shaw, Gen. Zool. vol. xi.
pt. 1 (1819), p. 49.
Le Jambou, Less. Compl. Buff. tom. iii. Ois. p. 27 (1837).
Ptilonopus jambu, Gray, List B. Brit. Mus. (1844) p. 2, (1856)
p-. 7. sp. 18; id. Gen. B. vol. ii. p. 467 (1844); Sclat. Proc. Zool.
Soc, 1863, p. 221; Wail. Ibis, 1865, p. 378.
Ramphiculus gambu, Bon. Consp. Gen. Av. vol. ii. p. 17 (1857) ;
Moore, Proc. Zool. Soc. 1859, p. 465; Reich. Taub. p. 99,
pl. 239. figs. 1334-35 ¢, pl. 244. figs. 3488-89 9°.
Ptilopus jambu, Schleg. Mus. Pays-B. 1873, p. 36, Columbe.
Native name Punei-chitrah in Borneo (Motley).
Hab. Malacca (Wallace) ; Sumatra (Miller) ; Borneo (Motley) ;
1878.] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 555
Luzon, Philippines (Crockewit) ; Busan and Bintulu, Borneo (Lve-
rett).
This bird, although very plentiful at Banjermassing, is, according
to Mr. Motley, rare in other parts of Borneo. It feeds on fruits,
particularly of the different species of Ficus. Quite different in its
style of coloration, it cannot be confounded with any other species
of Ptilopus. Of late hundreds of specimens have been sent to
Europe to be cut up to decorate ladies’ bonnets. I saw only lately
a box, at least three feet square, full of these birds, which were con-
demned to be torn to pieces in order to supply the demand this un-
fortunate fashion has created.
Adult male. Front, crown, and sides of face in front scarlet, the
top of head in some lights having an amethyst shade. Upper part
of throat and chin black. Sides of neck, from just below the eye,
lower part of throat, and entire underparts pure white, washed with
rose-colour on the centre of the breast. Under tail-coverts deep
chestnut. Entire upper parts and wings dark green. Primaries
black, tinged with green on the outer webs, and edged with rufous,
the first abruptly attenuated at tip. Tail dark green, with a dusky
grey apical band. Bill bright yellow. Inris sienna-orange. Feet
dark red.
Total length 103 inches, wing 53, tail 4, culmen 3.
Female. Top of head greyish purple, sides of face in front deep
rose, centre of throat and chin black. Upper parts and breast
bronzy green, rest of underparts white. Under tail-coverts chest-
nut. Tail green, with a pale rufous apical band.
Young. Plumage almost uniformly green. Chin pale red. Middle
of abdomen white. Under tail-coverts chestnut.
40. Pritopus WALLACEt.
Ptilonopus wallacei, Gray, Proc. Zool. Soc. 1858, p. 185,
pl. 136; id. List Mamm. & B. New Guin. p. 45 (1859); Wall.
Ibis, 1865, p. 380.
Ptilopus wallacei, Schlegel, Mus. Pays-B. 1873, p. 18, Columbe ;
Salvad. Ann. Mus. Civ. Genov. (1876) vol. ix. p. 197.
Hab. Aru Islands (Wallace); Kei Island (Von Rosenberg).
This beautiful bird, one of the most lovely species of this lovely.
group, was procured by Mr. Wallace on the Aru Islands, and de-
scribed by Gray (/.c.). It is still very rare in collections; and for
the examples from which my description was taken I am indebted
to the Marquis Doria, who sent them to me from the Museum of
Genoa. I have also examined the type in the British Museum.
Adult male. Top of head and occiput carmine. Chin, throat,
and sides of head pure white. Nape, neck, and breast pure grey.
A band of white across lower part of breast, succeeded by a broad
one of deep orange. Abdomen and under tail-coverts light yellow,
the latter striped with green. Upper part of back grey, bordered
with orange; scapulars and lesser wing-coverts grey, margined with
green. Shoulders orange. Secondaries bright green, margined with
lemon-yellow. Quills deep green, the first one narrowed gradual];
36*
556 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
to the tip. Back and rump yellowish green. Tail coppery green,
apical half of lateral feathers greenish white. Bill yellow. Feet
red. Iris orange-red.
Total length 10 inches, wing 6, tail 33, culmen 2.
41. PriLopus AURANTIIFRONS.
Ptilonopus aurantiifrons, Gray, Proc. Zool. Soe. 1858, p. 185,
pl. 137; id. List Mamm. & B. New Guin. (1859) p. 45; Wall.
Ibis, 1865, p. 380.
Ptilopus aurantiicollis, Schleg. Mus. Pays-B. 1873, p. 18,
Columbe.
Ptilopus aurantiifrons, var. nove-guinee, Meyer, Sitz. Ak. Wiss.
1874, p. 508.
Ptilopus aurantiifrons, Salvad. Ann. Mus. Civ. Gen.(1876) vol. ix.
p- 197.
Hab. Aru Islands, Mysol (Wallace); Salwatty (Von Rosenberg) ;
Batanta (Laglaize); New Guinea (Wallace, D' Albertis); Jobie
(Meyer, Beccari).
This is a very beautiful and very distinct species. Dr. Meyer
(/.c.) has proposed to separate the bird from the mainland of New
Guinea, procured at Passim, from that of Jobie, on account of some
slight variation in the hues of the plumage, such as the deeper
orange of the forehead and the orange tint of the back of the head
and breast, with also metallic reflections of the top of the head. I
do not deem these slight differences of sufficient importance to give
New-Guinea specimens a distinct specific rank. Examples from
Batanta before me seem to be intermediate between those of Jobie
and Passim, with more orange tints than in individuals from Jobie,
and less than in those of Passim.
Adult. Forehead deep orange, rest of top and sides of head yel-
lowish green. Throat pure white. Neck all around and upper part
of breast leaden grey. Underparts yellowish green. Mantle and
scapulars with some of the lesser wing-coverts grey, margined with
green, some of the feathers of the mantle margined with orange.
Rest of wing-coverts and secondaries yellowish green, margined with
lemon-yellow. Primaries dark green, the first narrowed gradually
to the tip. Back and upper tail-coverts yellowish green. Tail
bronzy green, with a subterminal interrupted grey band; this band
does not extend entirely across the outer web of each rectrix. Under
surface smoky grey, with a light-grey apical band. Bill yellow;
base swollen, red. Feet red. Iris orange.
Total length 94 inches, wing 53, tail 2#, culmen 2.
42. PriLoPUS ORNATUS.
Ptilopus ornatus, Schleg. Nederl. Tijdsch. 1871, vol. iv. p. 52;
id. Mus. Pays-B. 1873 p. 18, Columba; Salvad. Ann. Mus. Civ.
Genov. (1876) vol. ix. p. 197, et (1877) vol. x. p. 158.
Hab. New Guinea, Mount Arfak (Rosenberg, Laglaize).
This species seems to be entirely confined to the mainland of New
1878.] MR D. G. ELLIOT ON THE GENUS PTILOPUS. 557
Guinea. At all events I have not seen a specimen from any of the
islands. To state that it was a beautiful bird would be only equi-
valent to saying that it belongs to the genus Péilopus. It is still
quite a rare species in collections.
Adult. Top and back of head, sides of face, and back of neck
purplish red. Chin and throat pale grey. Upper part of breast
dark ochre, becoming a clearer yellow on the lower part of throat.
Underparts bright green; feathers of abdomen edged with yellow ;
centre of abdomen pale yellow. Under tail-coverts lemon-yellow,
streaked with dark green. Upper parts bright green, darkest on
the mantle. A band of dark purplish red upon the shoulders.
Wing-coverts light grey, margined with green. Primaries black,
tinged with dark green on outer webs, the first narrowed gradually
to tip. Secondaries bright green, edged with lemon-yellow. Tail
bright green, with a broad light yellow apical band. Bill yellow,
dark brownish towards the base in the skin. Feet red. Iris lemon-
yellow, bordered with orange.
Total length 92 inches, wing 6, tail 33, culmen 2.
43. Pritopus Gestror. (Plate XXXIV.)
Ptilinopus gestroi, D’ Alb. & Salvad. Ann. Mus. Civ. Gen. vol. vii:
p. 834 (1875).
Ptilopus gestroi, Salvad. Aun. Mus. Civ. Gen. (1876) vol. ix.
p- 197.
Hab. Yule Island (D’ Aldertis).
This very beautiful species is singularly intermediate between the
Pt. ornatus and Pt. perlatus, possessing as it dues the head and
neck of the latter with the body and tail of the former. I have only
seen one specimen, kindly sent to me by my friend Count Th. Salva-
dori, a male, collected by Signor D’Albertis in Yule Island. _ In size
it is larger than P¢. ornatus and more like Pé. perlatus. It is the
representative of its near allies in the district it inhabits. As yet
Yule Island is the only locality in which it has been obtained.
Adult male. Top and sides of head and upper part of back of
neck olive-yellow. Chin, throat, and a broad line going round the
lower part of neck light lead-colour, nearly white on throat. Lower
part of throat ochraceous yellow. Upper part of breast greenish
ochre, changing in certain lights to purplish. Lower parts of breast;
abdomen, and flanks bright green. Under tail-coverts yellowish
white, striped with bright green. Upper parts bright green. Wings
with a rich dark purple band near the shoulders; the coverts pale
grey, margined with green. Secondaries green, edged with lemon-
yellow. Primaries dark green, the first narrowed gradually at the
tip. Tail bright green, with a broad pale yellow apical band edged
with green. Bill olive-yellow. Iris yellow. Feet carmine. Total
length 10 inches, wing 6, tail 37, culmen 3.
44. PriLopus PERLATUS.
Columba perlata, Temm. Pl. Col. 559, livr. 94; Knip & Prév,
Pig. vol. ii. pl. xxi.
558 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
La Perlée, Less. Comp]. Buff. tom. iii. Ois. p. 31 (1837).
Ptilonopus perlatus, Gray, Gen. B. vol. ii. p. 466 (1844) ; id.
Proc. Zool. Soc. 1858, p. 185; id. Cat. Mamm. & B. New Guin.
(1859) p. 47; Wall. Ibis, 1865, p. 380.
Megaloprepia perlata, Bon. Compt. Rend. (1854) tom. xxxix.
. 1097.
' Sylphitreron perlatus, Bon. Consp. Gen. Av. vol. ii. p. 40.
Ptilopus perlatus, Schleg. Mus. Pays-B. 1873, p. 17, Columbe ;
Salvad. Ann. Mus. Civ. Genov. (1876) vol. ix. p. 197, (1877) vol. x.
p. 157.
Hab. Aru Islands (Wallace); New Guinea (Bernstein) ; Sal-
watty, Jobie (Von Rosenberg); Amberbaki (Laglaize) ; Dorey
(Raffray).
Like the last, this is rather a large species. It was placed by
Reichenbach in his genus Megaloprepia ; but I cannot see that it
possesses any characters to separate it generically from the members
of the genus Péilopus.
Male. Chin and upper part of throat pure white. Moustache,
sides, lower part of throat, and a narrow band separating the neck from
the mantle pale grey. Top and sides of head and back of neck greenish
yellow. Breast and a line beneath the grey band on the neck rufous.
Lower part of breast, abdomen, and flanks yellowish green. Crissum
and under tail-coverts pale yellow; the latter striped with dark green.
Upper parts and wings bright green, with the centre of scapulars
and smaller wing-coverts rosy purple. First primary slightly nar-
rowed at tip. Secondaries edged with light yellow. Tail dark green
above, beneath dark smoke-grey, with an apical greyish-white band.
Bill yellow ; feet red; iris orange-yellow. Total length 92 inches,
wing 63, tail 33, culmen 2. No difference in the sexes.
Immature. Top of head green ; lores and cheeks greenish yellow.
Throat greyish white. Sides and back of neck grey; breast dark
ochre. Rest of plumage like the male. (Spec. ex Dorey, New
Guinea, Voy. Astrolabe.)
45. PriLopus ZONURUS.
Ptilopus zonurus, Salvad. Ann. Mus. Civ. Gen. vol. ix. (1876-77)
p- 197.
Hab. Aru Islands (Wallace, Von Rosenberg).
Like Pt. perlatus, save that the tail has an apical greyish-white
band above and below. Barely distinguishable from Pt. perlatus.
46. PriLopus 10GASTER.
Columba hyogastra; Temm. Pl. Col. pl. 252, livr. 43; Desm.
Dict. Sc. Nat. tom. xl. pl. 344 (1826); Knip & Prév. Pig. vol. ii.
pl. 54.
Columba iogastra, Wagl. Syst. Av. (1827) sp. 34, Columbe.
TP’ Hyogastre, Less. Compl. Buff. tom. iii. Ois. p. 30 (1837).
Ptilinopus hypogaster, Swains. Class. B. vol. ii. p. 347 (1837).
Ptilonopus hyogaster, Gray, Gen. B. vol. ii. p. 466 (1844); Wall.
Ibis 1865, p. 381.
1878.] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 959
Totreron hyogaster, Bon. Consp. Gen. Av. vol. ii. p. 25. sp. 4
(1857).
Ionotreron ionogaster, Reich. Taub. p. 100, pl. 239. figs. 1332-33.
Ptilonopus (Iotreron) iogaster, Gray, Proc. Zool. Soc. 1860,
p. 359.
Ptilopus iogaster, Schleg. Mus. Pays-B. 1873, p. 20, Columba.
Ptilopus ionogaster, Salvad. Anu. Mus. Civ. Genov. (1876) vol.
ix. p. 196.
Hab. Tidore (Bruijn); Batchian, Gilolo (Wallace); Weda,
Dodingo, Ternate, Morotai (Bernstein).
This is a well-marked species, its grey head and purple abdomen
making it very conspicuous among the other species of the genus.
Adult. Head, except occiput, clear grey. General plumage of
body, neck, and occiput brouzy green. Abdomen almost entirely
covered by a spot of deep reddish purple. Crissum and under tail-
coverts rich lemon-yellow. Wings green. Some of the scapulars and
greater wing-coverts violet-grey, margined with green. Primaries
emerald-green, first narrowing gradually. Secondaries yellowish green,
edged with lemon-yellow. Tail bronzy green, with an interrupted
subterminal grey bar, this not reaching the edge of the outer web of
the rectrices. Beneath smoke-grey, apical band white. Bill bluish
white, tip yellow ; feet lilac-purple. Total length 9 inches, wing 54,
tail 3, culmen 2.
47. PriILorus PECTORALIS.
Columba pectoralis, Wag). Isis, 1829, p. 759.
Columba virens, Less. Voy. Coq., descr. 9; id. Trait. Ornith.
(1831) p. 471.
Columba cyanovirens, Less. Voy. Coq. pl. 42. fig. 2, 2.
Omeotreron pectoralis, Bon, Consp. Gen. Av. vol. ii. p. 27.
Ptilonopus roseypectus, Gray, Proc. Zool. Soc. 1861, p. 432 ;
Reich. Tauben, p. 191 ; Wall. Ibis, 1865, p. 381.
Ptilonopus virens, Wall. Ibis, 1865, p. 382.
Ptilopus pectoralis, Schleg. Mus. Pays-B. 1873, p. 23, Columbe ;
Meyer, Rowl. Ornith. Misc. pt. xiii. p. 114 (1878).
Ptilonopus pectoralis, Salvad. Ann. Mus. Civ. Genov. (1875)
vol. vii. p. 785.
Hab. Rubi, Passim, Andai, New Guinea (Meyer); Mysol (Hoedt);
Gagie, Guébeh, Waigiou (Bernstein); Mafoor, Sook (Von Rosen-
berg); Batanta (Laglaize) ; Amberbaki (Raffray).
Examples from Batanta are of larger size, but otherwise do not
seein to differ from those procured in New Guinea and other
localities. The species has a rather wide distribution among the
Papuan Islands.
Male. Forehead and chin greenish grey. A small reddish-purple
spot upon the breast. Middle of abdomen and under tail-coverts
light yellow ; the latter striped with dark green. Rest of plumage
of body yellowish green. A few of the scapulars with grey tips,
Primaries dark green on outer webs, and bronzy green at tips; first
560 MR. D. G. ELLIGT ON THE GENUS PTILOPUS. [May 7,
equal for its entire length. Secondaries yellowish green, edged with
pale yellow. ‘Tail bright yellowish green, with a grey spot at tip of
feathers on inner webs. Underneath smoky black, with apical grey
band. Bill yellow, base red; feet red; iris orange. Total length
84 inches, wing 4?, tail 27, culmen 2.
Female. Like the male, but without the spot on the breast.
48. PrrLopus virInis.
Columba viridis, Linn. (nee Scop.) Syst. Nat. vol. i. p. 283. sp.
23 (1766); Gmel. Syst. Nat. (1788) p. 780. sp. 23; Shaw, Gen.
Zool. vol. xi. pt. 1, p. 65 (1819); Knip & Préy. Pig. vol. ii. pl. xvii.
Tourterelle a gorge pourprée d@ Amboine, Buff. Pl. Enl. no.
142.
Ptilinepus viridis, Steph. Shaw, Gen. Zool. vol. xiv. p. 299. sp.5
1826).
Ionotreron viridis, Reich. Taub. p. 99, pl. 237. figs. 1320-21.
Totreron viridis, Bon. Iconogr. Pig. (1857) pl. 28, juv. 92; id.
Consp. Gen. Ay. vol. ii. p. 24. sp. 1.
Ptilonopus viridis, Gray, Gen. B. vol. ii. p. 467 (1844-49) ; id.
List B. Brit. Mus. (1856) p. 6; id. List Mamm. & B. New Guin.
(1859) p. 45; id. Proc. Zool. Soc. 1863, p. 34; Wall. Ibis, 1865,
p. 381.
Ptilopus viridis, Schleg. Mus. Pays-B. 1873, p. 22, Columbe ;
Salvad. Ann. Mus. Civ. Gen. vol. ix. (1876) p. 195.
Hab. Bouru, Amboyna, Ceram, Goram (Wallace); Harouka,
Monavolka (Rosenberg).
This is the oldest-described member of the genus Ptilopus, and
the only one known to Linnzeus. It is entirely an insular species,
confined to the islands west of New Guinea, upon which land I
have found no record of its ever having been procured. It is repre-
sented in the east (Bay of Geelvink) by the next species.
Male. Forehead and upper part of throat light grey. Lower part
of neck, and upper part of breast very deep reddish purple. Rest
of underparts yellowish green. Crissum and under tail-coverts light
yellow ; the latter with broad dark green stripes on the inner webs of
the feathers. Upper parts, occiput, and back of neck bronzy green.
A grey band near the shoulder, and tertials also grey, edged with
green. Primaries dark green; first about equal in width for the
entire length ; secondaries bronzy green, margined with lemon-yellow.
Tail bronzy green and tipped with yellow, lightest on the edges. Bill
yellow, base red ; orbits yellow; feet pink-red; iris with inner ring
yellow, outer red. Total length 8 inches, wing 43, tail 3, culmen 2.
49. PriLopus GEELVINKIANUS.
Piilopus viridis geelvinkianus, Schleg. Nederl. Tijdsch. vol. iv.
(1871) p. 23; id. Mus. Pays-B. 1873, p. 23.
Ptilopus musschenbreeckii, Rowl. Ornith. Mise. pt. xiii. pl. 95,
p- 114; Salvad. Ann. Mus. Civ. Gen. vol ix. (1876) p. 195.
Hab. Islands of Mafor and Meosnoum (Von Rosenberg, Raffray) ;
Mysore (Meyer).
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 561
This bird, separated from the Pt. viridis by Schlegel, is appa-
rently entitled tc be considered a distinct species. With a large
series of examples before me, mainly brought by M. Raffray, the
difference of the colour on the breast is very perceptible, being very
much brighter and purer than the deep purple of Pé. viridis. In
other respects the two species closely resemble each other. The
term musschenbreckii, being only a MS. name of von Rosenberg’s,
without any description of the bird given, must become a synonym.
Adult. Forehead and chin dark grey ; sides of face grey, washed
with green. Breast and lower part of throat bright reddish purple,
much lighter than the similar spot in Pt. viridis. General plumage
yellowish green, slightly bronzed in certain lights. Centre cf abdomen
and under tail-coverts pale yellow ; the latter striped with dark green.
Across the wing, near the shoulders, a broad band of pale grey. Some
of the innermost secondaries with a spot of light grey near their tips ;
first primary not narrowed. ‘Tail yellowish green, each feather with
a subterminal grey spot on the inner web. Bill yellow, base red.
Iris red; feet red. Total length 8 inches, wing 43, tail 3, cul-
men 2. Sexes alike.
50. Prinorus EUGENI.
Totreron eugenia, Gould, Proc. Zool. Soe. 1856, p. 137 ; Gray,
List B. Brit. Mus. (1856) p. 6; Bon. Iconogr. Pig. (1857), text.
Ptilonopus eugenia, Gray, List B. Trop. Is!. (1859) p.39; Wall.
Ibis, 1865, p. 381; Sclat. Proc. Zool. Soc. 1869, p. 124.
Ptilopus eugenie, Salvad. Aun. Mus. Civ. Genov. vol. ix. (1876)
p- 195.
Hab. Solomon Islands.
The type specimen, which I have examined, of this apparently
distinct species, is in a mutilated condition, and wants entirely the
tail. The species is very close to the two preceding, but differs in the
white head. The type in the British Museum is unique.
Top of head, cheeks, and upper part of throat white. Centre of
lower throat and breast rich deep crimson. Back of neck and upper
parts yellowish green. Across the shoulders a band of grey, and
the tertiaries tipped with the same. Secondaries bright green, edged
on the outer webs with yellow. Under surface greyish greeen. Tail
wanting. Bill apparently greenish at base, yellow at tip. Wing 43
inches, culmen 3. (Type.)
51. Pritopus RIvOLII.
Columba rivoli, Knip & Préy. Pig. vol. ii. pl. 57; Des Murs,
Iconogr. Ornith. pl. 4 (1845).
Lotreron rivolit, Bon. Consp. Av. vol. ii. p. 25 (1857).
Tonotreron rivolii, Reich. Taub. p. 100, Taf. 235. fig. 1306.
Ptilonopus rivoli, Wall. Ibis, 1865, p. 381.
Ptilonopus solomonensis, Gray, Ann. Mag. Nat. Hist. vol. y.
(1870) p. 328.
Ptilopus rivolit, Sclat. Proc. Zool. Soc. 1877, p. 109.
562 MR. D. G. ELLIOT ON THE GENUS pPTiLoPus. [| May 7,
Hab. Duke-of-York Island (Brown), Wanga, San Christoval
(Brenchley).
The type of the Péclopus rivolit was originally in the collection of
the Prince d’Essling, which was purchased by Dr. Wilson, and now
forms part of the Museum of the Academy of Natural Sciences at
Philadelphia. In the catalogue of Prince d’Essling’s collection made
at the time of its sale, the specimen of Pt. rivolii is entered on page
30 as having ‘‘Inde” for the habitat. The exact locality of this
handsome species was unknown until Mr. George Brown, C.M.Z.S.,
obtained it in Duke-vf-York Island, lying off the north-east coast of
New Guinea. I am inclined to believe that the Pé. solomonensis,
Gray, is based on a young bird of this species.
Male. Forehead purplish red ; breast crossed by a broad white
band, a large rose-red spot on the abdomen ; lower part of abdomen,
crissum, and under tail-coverts bright yellow. Rest of plumage
bright green, with some small round bluish-black spots on the
scapulars. In its dimensions this species agrees with the following
one.
Female. General plumage bright green.
52. PriLopus PRASINORRHOUS.
Ptilonopus prasinorrhous, Gray, Proc. Zool. Soc. 1858, p. 185;
id. List B. & Mamm. New Guin. 1859, p. 46; Wall. Proc. Zool.
Soc. 1863, p. 34, ex Bouru.
Ptilopus rivoli, Schleg. (nec Prév.) Nederl. Tijdsch. tom. iv.
p- 21 (1873) ; id. Mus. Pays-B. 1873, p. 24, Columbe ; Salvad.
Ann. Mus. Civ. Gen. vol. ix. 1876, p. 196, ex Bouru.
Ptilopus prasinorrhous, Salvad. Ann. Mus. Civ. Gen. (1876)
vol. ix. p. 196, (1877) vol. x. p. 157.
Hab. Ké Islands, Khoor, Tegoor, Matabello, Monavolka, Pulo-
Pandang, Goram, Salwatty (Von Rosenberg); Bouru, Waigiou
(Wallace) ; Amboina, Mysol (Hoedt); Koffiao (Beccarz); Batanta
(Bruijn) ; Sorong (D’ Albertis).
This species appears to be very variable in its coloration from
different localities; and it is a question if examples might not be
procured leading completely from the typical style of this bird into
that of the Pt. rivolii. It is mainly distinguished from this last by
having the vent and under tail-coverts green, slightly margined with
yellow, instead of these parts being bright yellow asin P¢. rivolit. In
the collection at Leyden are specimens from the same locality (island
of Bouru), with the under tail-coverts green and yellow respectively,
which would seem to show that this character is an indefinite and
unsatisfactory one. I do not consider that the specific distinctness of
this bird from the Pé. rivoliz has as yet been established. A larger
series than I have had at my command, from all the localities known,
will be necessary before the question can be determined. If the two
are distinct, it would be a curious fact that, in the case of such
closely allied species, P¢. rivolit should be restricted to the Duke-of-
York Island, while the present is scattered over a dozen islands and
also found upon the mainland of New Guinea.
1878. | MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 563
Resembling in its coloration P¢. rivolit, but differs chiefly in the
vent and under tail-coverts being green, slightly margined with
yellow; the reddish-purple patch of the abdomen sometimes reaches
the white pectoral band. First primary slightly narrowed at tip.
Bill yellow ; feet red.
53. Pritopus sTROPHIUM.
Ptilinopus strophium, Gould, Jard. Contr. Ornith. 1850, p- 102,
pl. 105 (note).
Ptilonopus strophium, Gray, List B. Brit. Mus. (1856) p-. 6
(partim) ; id. List Mamm. & B. New Guin. (1859) p- 46; Salvad.
Ann. Mus. Civ. Gen. (1876) vol. ix. p. 196.
Tonotreron rivoli, Reich. Taub. p. 100, pl. 236 8. fig. 3364.
Ptilopus miqueli, Schleg. Ned. Tijdsch. (1873) vol. iv. p. 22;
id. Mus. Pays-B. 1873, p. 26, Columbe; Rowl. Ornith. Mise.
part xii. p. 60, pl. 88 (1878) ; Salvad. Ann. Mus. Civ. Gen. vol. ix.
p. 196 (1876).
Piilopus miqueli, Meyer, Sitzungs. d. k. Akad. d. Wiss. vol. lxx.
p- 128 (1874).
Ptilinopus miqueli, Salvad. Ann. Mus. Civ. Gen. (1875) vol. vii.
p- 786.
Hab. Duchateau Isles (MacGillivray, type of Pt. strophium,
Gould) ; Jobie, Meosnoum (Von Rosenberg, Meyer).
Mr. Gould’s type of his Pé. strophium is in the British Museum,
and is undoubtedly the same bird as was afterwards described as
Pt. miquelli by Prof. Schlegel. The type is faded upon the forehead
and is more of a rosy hue than purple-red; but it is plainly
evident, I think, that the original colour has faded away. Another
specimen in the Museum collection, without any locality, has the
forehead purplish red. The name miqueli will have to give way
to the older one of strophium.
Male. Front and top of head purplish red. A broad pure white
band across the breast, beneath which is a stripe of bluish green.
Centre of abdomen and under tail-coverts bright yellow. Entire rest
of plumage bright yellowish green, with some small round bluish-
black spots on the scapulars near the tips. Primaries greenish black,
first narrowed to the tip. Tail same colour as the back. Bill
greenish, tip yellowish; feet red; iris orange. Total length 103
inches, wing 45, tail 2?, culmen?. (Ex type.)
Female. Plumage almost entirely green.
54. PriLoprus BELLUS.
Piilonopus bellus, Sclat. Proc. Zool. Soc. 1873, p- 696, pl. 57 ;
Salvad. Ann. Mus. Civ. Genov. (1875) vol. vii. p. 786.
Piilopus bellus, Salvad. Aun. Mus. Civ. Genov. (1876) vol. ix.
p. 197, aud (1877) vol. x. p. 157.
Hab. Atam, New Guinea (D’ Albertis) ; Mount Arfak (Laglaize);
Amberbaki ( Raffray).
This very fine and distinet species is one of the largest of the
364 MR. D. G. ELLIOT ON THE GENUS pTiLopus. [May 7,
section of the genus to which it belongs. Originally described by
Mr. Sclater froin specimens obtained by D’ Albertis on Mount Arfak,
it has since been found in other parts of the northern coast of New
Guinea. Beside its much greater size, it can always be distinguished
from Pt. speciosus, its uearest ally, by the rosy-red forehead and
crown, together with the purplish-red spot on the abdomen.
Male. Front and crown deep rosy red. Occiput dark bluish green.
Breast covered by a broad, halfmoon-shaped band, pure white on
the lower parts and sides, lemon-yellow on the upper part. Abdomen
has the middle portion covered by a purplish-red patch. Entire
rest of plumage yellowish green, with small round bluish-black
spots on the scapulars, and the feathers of the crissum and under
tail-coverts edged with yellow. Primaries greenish black, first not
narrowed. ‘Tail yellowish green, with a pale apical band of the same
colour. Bill yellow; feet dark red; iris yellow. Total length 93
inches, wings 52, tail 33, culmen 2.
55. PriLopus SPECIOSUS.
Ptilopus speciosus, Schleg. Neder]. Tijdsch. vol. iv. (1871) p. 23;
id. Mus. Pays-B. 1873, p. 27, Columbe; Salvad. Ann. Mus. Civ.
Genov. (1876) vol. ix. p. 197.
Hab. Sook, Mafor (Von Rosenberg, Raffray); Misori (Raffray).
This very peculiarly-coloured and striking species is apparently
confined to the islands in the great Bay of Geelvink, New Guinea.
It was first described by Sehlegel from specimens sent by Von
Rosenberg, and is very rare in collections. Lately a large series
has been received at the Paris Museum, obtained by M. Raffray in
the localities given above. There does not appear to be any variation
among individuals from the different habitats.
Male. General plumage yellowish green, darkest on the head and
neck. A broad bright-yellow band bordered on the lower side with
white crosses the breast. Abdomen beautiful lilac ; lower part of
abdomen and under tail-coverts bright lemon-yellow. A purple spot
in front of the eyes. Primaries grey on their outer webs and tips ;
first not narrowed especially towards tip. ‘Tail yellowish green.
Bill black, tip yellow; feet dark red. Total length 7 inches,
wing 43, tail 23, culmen 3.
Female. Green, with the feathers of the abdomen margined yellow,
and under tail-coverts yellow. Primaries have outer webs and tips
grey.
56. PriLopus JOHANNIS.
Ptilopus johannis, Sclat. Proc. Zool. Soc. 1877, p. 556.
Hab. Wild and D’Entrecasteaux Islands, of the Admiralty group
(‘ Challenger’ Expedition).
This very distinct species was obtained by the ‘Challenger’ Ex-
pedition, and derives its name from Mr. John Murray, one of the
naturalists. By the kindness of Mr. Sclater I have been enabled
to examine the types. With its nearest relationship perhaps to
1875. |] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 565
Pt. speciosus, it differs conspicuously from all the members of this
particular section of the genus Péilopus in having the band which
crosses the breast all yellow—and in this respect differs from all the
others, as it is the only one so marked, the others having this band
white, or yellow and white. It is a very beautiful species.
Male. General plumage green, with small dark blue spots on the
scapulars. A broad band across the chest, which, together with the
lower part of abdomen, crissum, and under tail-coverts, is light yellow.
Top of head and middle of abdomen lilac. Primaries leaden grey,
with the tips light grey, nearly white. Bill black, feet dark red.
Total length 7} inches, wing 43, tail 22.
Female. Nearly uniform green; middle of belly and crissum
yellow.
57. PrILOPUS LUTEOVIRENS.
Colombe jaune, Hombr. & Jacq. Voy. Péle Sud, plaid 2..£,2.
Columba luteovirens, Hombr. & Jacq. Ann. Sc. 2° sér. vol. xvi.
(1841) p. 315.
Colombe de Felicie, Hombr. & Jacq. Voy. Pole Sud, pl. 12. f. 1,
v.
a Columba felicie, Hombr. & Jacq. Aun. Se. 2° sér. vol. xv. (1841)
p. 316, juy. ‘
Columba flava, Gray, Gen. B. vol. ii. p. 470 (1844-49).
Calenas luteovirens, Hartl. Wiegm. Arch. 1852, p. 134.
Ptinilopus! luteovirens, Jacq. & Puch. Voy. Péle Sud, vol. iii.
p. 112 (text) (1853).
Ptinilopus! felicie, Jacq. & Puch. Voy. Péle Sud, (1853), text,
uv.
Ptilonopus luteovirens, Gray, List B. Trop. Isl. (1859) p. 38.
Chrysenas luteovirens, juv. felicia, Reich. Taub. Suppl. p. 203,
Taf. nov. no. iii. fig. 32.
Chryscenas luteovirens, Reich. Taub. p- 90, t. 245. fig. 5594, t.
263. figs. 1469-70.
Chryscena luteovirens, Bon. Compt. Rend. (1854) tom. xxxix. p-
879; id. Consp. Av. vol. ii. p. 28; id. Iconogr. Pig. (1857) pl. 31;
Finsch & Hartl. Faun. Centralpolyn. (1867) p. 134, Taf. ii. fig. 4,
juv. ; Layard, Proc. Zool. Soc. 1875, p: 436; Finsch, Proc. Zool.
Soc. 1875, p. 557; Finsch, Journ. Mus. Godeff. Heft xii. (1876)
p- 10.
Omotreron felicia, Bon. Consp. Gen. Av. vol. ii. p. 27 (1857), juv.
Hab. Fiji Islands, Balaou (Homér. § Jacq., type); Ovalau (Lay-
ard) ; Viti Levou (Godefroy).
This species, according to Layard, dwells in the forest-covered
parts of the island of Ovalau and the Rewa. It breeds in October,
November, and December. On account of the colour of. their
plumage it is difficult to see them in the trees ; but their presence
is betrayed by their voice, which sounds something like a bark. It
feeds on berries, and is fond of the seeds of araliaceous trees and
ereepers of the genus Melastoma.
This species is the type of Bonaparte’s genus Chryscena, the chief
566 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. {May 7,
character being the lanceolate shape of the feathers of the body.
This, however, would seem to indicate more a specific than a generic
value, as the Pt. victor, with which the present bird is generically
identical, does not possess this form of feather. As is shown in my
remarks on genera, in the beginning of this paper, neither of these
two species exhibits any characters that, in my opinion, would separate
them from the genus Ptilopus, those relied upon by authors for
establishing a distinct generic rank not being at all peculiar to, or
possessed solely by, these birds.
Male. Head greenish yellow. Throat and irregular collar on lower
part of neck lemon-yellow. Feathers of breast and back lanceolate
sulphur-yellow, tinged with green. Abdomen, vent, and under tail-
coverts bright yellow. Primaries dark greenish yellow, first not
narrowed especially ; secondaries pale green, both edged with lemon-
yellow. Tail pale green, graduating into light yellow at the tip. Bill
black, tip yellow. Feet black. Total length 8 inches, wing 43, tail
23, culmen 3. (Ex type, Paris Museum.)
Young male. Head and throat pale green. General plumage
bright green, with yellow lanceolate feathers appearing upon the
breast and back. Abdomen mixed with yellow and green. Under
tail-coverts saffron-yellow. Primaries and secondaries brownish
green, broadly margined with yellow. Some feathers of the ramp
and upper tail-coverts tipped and margined with yellow. Tail bright
green. Bill black, tip yellow. Feet black.
Female. General plumage bright green, verging to yellow on the
throat and abdomen. Primaries and secondaries margined with light
yellow. Under tail-coverts pale yellow. Tail bright green, lightest
at the tip. Bill black, tip yellow. Feet black.
58. Pritopus CoRRIEI.
Ptilinopus corriei, Ramsay, Proc. Linn. Soc. New South Wales,
1876, p. 133, vol. i.
Hab. Malacolo Island, New Hebrides.
In the above-named publication Mr. Ramsay describes a specimen
of Ptilopus as representing a distinct species. It is evidently imma-
ture, and, judging from the description, belongs to the group com-
prising Pt. victor, luteovirens, and layardi; and it is not impossible
that it may be one of these species from a new locality, although the di-
mensions given are larger than those of any examples of the above-
named species that I have seen. The type had been in spirits, and, as
stated by Mr. Ramsay, was somewhat discoloured on the sides of the
neck and interscapular region. The texture of the feathers resembled
that of Chryseenas viridis, Layard (Pt. layardi of this paper). Only a
single specimen was procured by Dr. A. Corrie, at Malacolo Island,
during the cruise of H.M.S. ‘ Pearl,’ in August 1875. More speci-
mens will be required before its specific rank can be satisfactorily
determined.
The following is Mr. Ramsay’s description :—
Male. ‘‘The whole of head, chin, and throat olive-green, becom-
ing bright green on the neck and occiput ; the whole of the remainder
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 367
of the upper and under surface bright deep green ; the wing-quills
deep golden green, mesially shaded with black ; the inner webs of
the primaries black towards the middle and basal portion of the
feather. The three inner secondaries and their corresponding coverts
of the larger series only have an oblique oblong blotch of bright
golden yellow at their tips, which on the secondaries is confined to
the outer web only, the smaller wing-coverts at the bend of the wing
adjacent to the scapulars having a roundish, crescentic or cordate
spot of white (?) near the tips of the feathers; some show a green
shaft-line and a margin of green round the tip of the feather ; under
wing-coverts green; the outer series ashy ; under surface of the wings
ashy brown ; under surface of the tail ashy brown, becoming lighter
towards the base, and crossed conspicuously near the tip with a band
of light ashy grey ; upper surface green, the grey band not so con-
spicuous ; the inner portion of the legs and lower part of the abdo-
men yellowish (much faded); under tail-coverts yellow, blotched
with oblong marks of green on inner webs; the thighs, outer
feathered portion of the legs, and the flanks bright green, like the
rest of the body; tail of 14 feathers, square, even ; of the wings, the
third, fourth, and fifth feathers are nearly equal and longest ; bill,
legs, and feet bluish lead-coiour; tarsi not feathered to the toes,
lower portion scaled in front. Total length 9°5 inches, wing 6°3,
tail 3; bill from forehead 0-8, from gape 1; tarsus 95.”
59. PriLorus LAYARDI.
Chrysenas viridis, Layard, Proc. Zool. Soc. 1875, pp. 151, 437 ;
Finsch, Proc. Zool. Soc. 1877, p- 736.
Native name Sohulu (Layard).
Hab. Kandavu, Fiji Islands.
This apparently very distinct Pigeon was described by Layard
(J. ec.) from specimens obtained by himself at Kandavu. He at first
identified it, as he tells us, with P. /uteovirens, but afterwards, having
obtained it in the full breeding-dress, saw that his first impression
was an erroneous one, and then conferred upon the species the
name of viridis. Several specimens of this pretty Dove are in the
collection of the ‘Challenger’ Expedition, which, through the.
kindness of my friend Dr. Sclater, I have had the Opportunity of
examining. ‘hey are all in the adult dress, with greenish-yellow
head, and entirely green body relieved only by the yellow of the
under tail-coverts. The feathers of the back are slightly inclined to the
lanceolate form of those of P. luteovirens ; but the two species cannot
well be confounded together. As I do not consider that the birds
included in the term Chrysenas possess any characters that should
properly separate them generically from Péilopus, it will be impos-
sible to retain the name of viridis given to this species by Mr. Layard,
as there is already, unfortunately, a species of this group so named,
described originally by Linneeus. I have therefore substituted the
name layardi, in justice to my friend, who has, in his researches
among the birds of the Fiji Islands, exhibited the same energy and
568 MR. D. G. ELLIOT ON THE GENUS PTILOoPUS. [May 7,
love for natural history which has always characterized him in other
parts of the world in which his lot has been temporarily cast.
Male. Head and neck greenish yellow ; under tail-coverts bright
yellow. Thighs and lower part of abdomen grey. Rest of
plumage of body and wings dark green, with a golden gloss upon back
and chest. Primaries are edged with yellow on outer webs, and,
together with the secondaries, have nearly all the inner webs golden
yellow; the first primary is narrowed gradually towards the tip.
Bill blue-black, tip livid ; legs dark crimson ; iris yellowish. Length
72 inches, wing 43, tail 25, culmen 2 (skin).
Female. Like the male, but less brilliant generally.
60. PriLopus vicTorR.
Chrysena victor, Gould, Proc. Zool. Soc. 1871, p. 642; Layard,
Proc. Zool. Soc. 1875, p. 437; Finsch, Proc. Zool. Soc. 1875,
p- 557; Rowl. Ornith. Mise. pt. v. pl.; Finsch, Journ. Mus. Godeff.
Heft xii. (1876) p. 10; Lay. Ibis, 1876, p. 151; Finsch, Proce.
Zool. Soc. 1877, p. 736.
Native name Buli ndamu (Layard).
Hab. Fiji Islands, Bua (Filhol); Vanua Levu, Taviuni, Ngami,
Lanthala (Layard).
Layard says that this Dove is the glory of the forest of Taviuni,
and is also found in Lanthala and Rambi Islands, also on Vanua
Levu, about Bua, Ndreketti, &e. The young males would be mis-
taken for females, but that the vent is more orange. They breed
about December or November, making a rude platform of small
twigs for a nest about ten feet from the ground, and lay two eggs,
axis 1" 4, diam. 1”, pure white in hue. The species feeds on
many sorts of small and large berries and fruits.
Male. Head and throat dull olive-green. Entire rest of plumage
bright orange-carmine. Under surface of wings bright yellow.
Primaries greenish yellow, edged with orange; first not narrowed.
Tail, which is very short and almost hidden by the upper-coverts,
brownish orange, graduating into pure orange at the tip. Bill and
feet black. Total length 7 inches, wing 44, tail 23, culmen 3.
Female. Head yellowish green, lightest upon the throat. Entire
plumage of body rich green ; inner webs of primaries and secondaries
broadly margined with orange; outer webs edged with the same
colour. Under tail-coverts orange, with the centre of the feathers
green. Bill black, tip yellow; feet black.
61. PriLopus LECLANCHERI.
Trerolema leclancheri, Bon. Compt. Rend. (1855) tom. xli. p.
247; id. Iconog. Pig. (1857) pl. 16.
Carpophaga leclancheri, Gray, List B. Brit. Mus. (1856) p. 21.
sp. 19.
Leucotreron gironieri, J. Verr. & Des Murs, Ibis, 1862, p. 342,
pl. 12, 2 ; Wald. Trans. Zool. Soc. vol. ix. p. 213 (1875).
Ptilopus geversi, Schleg. Ibis, 1863, p. 120.
Ptilopus hugonianus, Schleg. Neder). Tijdsch. 1863, p. 60, pl. 3,
1878. } MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 569
fig. 2, juv. ex Luevu; Wall. Ibis, 1865, p. 378; Schleg. Mus.
Pays-B. 1873, p. 36.
Ptilopus leclancheri, Salvad. Ann. Mus. Civ. Genov. (1876) vol.
ix. p. 199.
Hab. Philippines, Luzon, and Guimaras (Meyer).
This species was first described by Bonaparte (J. c.) as Trerolema
leclancheri. The type is a female, and resembles in a great degree
the specimen described by Lord Walden (J. ce.) from Guimaras,
brought by Meyer, and called, in Lord Walden’s memoir on Philippine
birds, Leucotreron gironieri. The type was brought to Paris by
M. Leclancher, who was attached as surgeon to the expedition of the
‘ Favourite ;’ but no locality whatever was given to the specimen ; so
that of New Guinea, furnished by Bonaparte (J. c.), is erroneous.
Messrs. J. Verreaux and Des Murs redescribed the species in ‘ The Ibis’
(1. c.) as Leucotreron gironieri, their specimen being also a female or
possibly a young male, bearing, however, a very close resemblance to
Bonaparte’s type. Schlegel (/. c.) also described a young bird of
the same species as Péilopus hugonianus. All these names must be-
come synonyms of that given by Bonaparte.
Male. Head, neck, and upper part of the breast ashy white,
washed on the nape with pale green ; chin and throat black. A broad
dark purple band crosses the breast just below the white. Flanks
and underparts ashy green. Under tail-coverts cinnamon. Upper
parts bright green. Wings green; primaries and secondaries black
on inner webs, dark green on the outer, all edged with yellowish
white. Tail bright green, with a terminal light-yellow band. Bill
yellowish ; feet red. Total length 94 inches, wing 6, tail 4,
culmen 3.
Female. Head and breast ashy white, nape pale green. Throat
and chin black. A purplish band on the breast, which does not ex-
tend across. Rest of underparts and flanks tawny ashy green.
Under tail-coverts chestnut. Upper parts and tail as in the male
described above. (From type of 7’. leclancheri, Paris Museum.)
62. PriLopPus OCCIPITALIS.
Ptilonopus occipitalis, Gray, Gen. B. vol. ii. p. 467, pl. 118 (1844);
id. List B. Brit. Mus. Gall. p. 1 (1844), (1856) p. 7. no. 17;
Wall. Ibis 1865, p. 378; Mart. Journ. fiir Ornith. 1866, p. 22.
no. 124.
Ramphiculus occipitalis, Bon. Compt. Rend. tom. xxxix. p. 878
(1854) ; id. Iconogr. Pig. pl. 14 (1857); id. Consp. Av. vol. ii.
p- 17; Wald. Trans. Zool. Soc. vol. ix. (1875) p. 214.
Omotreron batilda, Bon. Compt. Rend. tom. xxxix. p. 878 (1854);
id. Consp. Av. vol. ii. p. 27; Wall. Ibis, 1865, p. 382 ; Von Mart.
Journ. fir Ornith. 1866, p. 22. no. 125 (juv.).
Tonotreron occipitalis, Reich. Taub. p. 101, pl. 239. fig. 1331.
Columba occipitalis, Schleg. Handl. vol. i. p. 411 (1857).
Lamprotreron porphyria, Bon. (nec Temm.), Iconogr. Pig. (1857)
pl. xv. (juv.), from type of O. batilda.
Proc. Zoou. Soc.—1878, No. XXXVII. 37
570 MR. D. G. ELLIOT ON THE GENUS PTILopus. [May 7,
Ptilopus occipitalis, Schleg. Mus. Pays-B. 1873, p. 35, Co-
lumbe.
Hab. Luzon, Philippines (Meyer).
Bonaparte described a young bird of this species, in the ‘Comptes
Rendus’ (J. c.), as Omotreron batilda; and afterwards, in his ‘ Icono-
graphie des Pigeons,’ he figured this specimen as the young of
Ptilopus porphyreus! The type of O. datilda is in the collection of
the Paris Museum, and proves to be simply a young bird of P. oc-
cipitalis. It is larger than any specimen of porphyreus, of which
there is a series before me, even exceeding the adults in its measure-
ments, and presents quite a different coloration. The type was
brought from the Philippines in 1839 by M. Barrot, but no parti-
cular island is indicated.
Adult. Top and front of head light leaden grey ; sides of face
below the eyes and entire back of head purplish red. Throat yel-
lowish white; sides of neck and breast grey, like the top of head.
Entire centre of breast, from the lower part of the throat to the ab-
domen, ochraceous ; flanks bright green ; a large purplish-red spot in
the middle of the abdomen. Crissum grey mingled with white. Upper
parts bronzy green, darkest upon the rump and upper tail-coverts.
Wings bright green; primaries and secondaries black on their inner
webs, green on their outer and edged with light yellow. Tail bright
green, with an apical greyish band; under coverts white, with a
broad green stripe on their inner webs. Bill red, with light tip ; feet
red. Total length 11? inches, wing 61, tail 43, culmen 2. (Specimen
figured by Bonaparte, Icon. Pig. pl. 14.)
Young. Top of head, back of neck, and entire upper parts bronzy
green. Throat yellowish white. Sides of head yellowish white, in-
terspersed with green. Underparts greyish green, bronze upon the
breast. Crissum and under tail-coverts whitish, the latter striped
with green. Wings like the back, the secondaries and primaries
edged on outer web with yellowish white. Tail bright green, with a
greyish-white apical band, which is observed only on the inner
webs. ‘Total length 11 inches, wing 6, tail 4, culmen 8. (Type of
batilda, Bou.)
63. PriLopus GULARIS.
Columba gularis, Quoy & Gaim. Voy. Astrol. Zool. vol. i. p. 247,
pl. xxix. (1830).
La Mentonniere, Less. Comp. Buff. tom. iii. Ois. (1837).
Columba gularis, Knip & Prév. Pig. pl. xi. -
Laryngogramma gularis, Reich. 'Tauben, p. 102, pl. 233. fig. 1297;
id. suppl. p. 203, nov. Taf. iv. fig. 44.
Leucotreron gularis, Bon. Consp. Av. vol. ii. p. 15 (1857); Wald.
Trans. Zool. Soc. vol. viii. (1872) p. 83; Salvad. Ann. Mus, Civ.
Gen. (1875) vol. ix. p. 670.
Ptilonopus gularis, Wall. Ibis, 1865, p. 377.
Ptilopus gularis, Schleg. Mus. Pays-B. (1873) p. 57, Columba.
Hab. Celebes.—Menado (Wallace); Tondano (Forsten) ; Goron-
talo, Wawou, Bone, Modelido (Von Rosenberg).
1878. ] MR. D. G. ELLIOT ON THE GENUS PTILOPUS. 571
Peculiarly a Celebes species, the present bird has fortunately never
received another name beside the one originally bestowed on it by Quoy
and Gaimard (/. c.). In its long tail it differs from the majority of
the species of Ptilopus, and approaches the birds placed by
Reichenbach in “ Megaloprepia ;” but the first primary is abruptly
attenuated.
Adult. Top and sides of head grey, graduating into light green
upon the occiput and hind neck. Chin and centre of throat very deep
chestnut; rest of throat, neck on sides, and underparts lavender-
colour, with a large spot in the centre of the abdomen yellowish
white washed with rufous. Thighs and under tail-coverts rich dark
cinnamon. Upper parts, wing, and tail bright green, the rectrices in-
distinctly tipped with yellowish green. Primaries and secondaries
black on inner webs, green on outer, and edged with light yellow.
Bill yellow; iris orange-brown; eyelids and orbits bare, blue ; feet
red. Total length 131 inches, wing 63, tail 53, culmen 3. (Type
in Paris Museum.)
64. PriLopPus FISCHERI.
Ptilinopus fischeri, Brigg. Brem. Abhandl. (1876) p. 82, Taf. iv.
flab. Celebes.
I only know this handsome species by the description and figure
published by Dr. Briiggemann (/. c.)._ Is is apparently very distinct
from all the known forms, and cannot be compared with any of them.
The red patch on the side of the head is a very peculiar character.
It may be briefly described as follows -—
“Back green. Head, neck, and breast partly grey, partly golden
ochre, Under tail-coverts grey and white intermixed. Sides of head
in the male bluish red; back of neck with a black band. Total
length 400 millims., wing 168, tail 147, bill 17, tarsus 21.” (Briigg.
de.)
65. PriLopus ALBOCINCTUS.
Ptilinopus albocinctus, Wall. Proc. Zool. Soc. 1863, p. 496,
pl. xxxix.; id. Ibis, 1865, p. 377.
Ptilopus cinetus florensis, Schleg. Nederl. Tijdsch. vol. iv. p. 20
(1873).
Hab. Flores (Wallace).
This species, which seems distinct from Pt. cinctus, was first de-
scribed by Wallace (/.c.). It differs from its ally in having the
neck and breast light blue, and the first primary less attenuated.
The bill, according to Wallace, is greenish at the base, yellow at tip ;
feet bright red. I have examined the type in the British Museum.
In size and all other characters, save those pointed out, it is like
Pt. cinctus.
66. PriLopus crncTUs.
Columba cincta, Temm. Pig. & Gall. vol. i. p. 243(1813); Knip
& Prév. Pig. vol. ij. pl. xxiii.; Shaw, Gen. Zool. vol. xi. p. 50
(1819); Wagl. Syst. Av. (1827) sp. 19, Columba.
37*
572 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7>
Leucotreron cincta, Bon. Compt. Rend. (1854) tom. xxxix.
p- 876 ; id. Consp. Av. vol. ii. p. 15; Reich. Tauben, p. 103, pl. 238.
fig. 1328.
Patong cinctus, Gray, Gen. B. vol. ii. p. 467 (1844); id. List
B. Brit. Mus. (1844) p. 2, (1856) p. 8. sp. 22; Wall. Ibis, 1865,
0. Oi
| Ptilopus cinetus, Schleg. Nederl. Tijdsch. tom. iv. p. 19 (1873) ;
id. Mus. Pays-B. 1873, p. 34.
Hab. Timor (Wallace, Miller) ; Wetter (Hoedt).
This bird is about the same size as the Pé. gularis, but has a long
and square tail, the feathers of which are very broad. It belongs to
the large members of this genus, and should, from its size, be in-
cluded in the genus Megaloprepia, if that were really a valid division;
but then it has an abruptly attenuated primary, which although not
a generic character, as I have shown, still, according to Reichenbach,
prevents its entrance into his genus (?).
Adult. Head, neck, throat, and entire breast white ; rest of under-
parts yellowish green, separated from the white breast by a black
band. Entire upper parts, wings, and tail black; the rectrices with
a broad apical dark grey band. Under tail-coverts dark grey on the
inner webs, white on the outer, margined broadly with bright yel-
low. Bill ocbre-yellow, greenish at base; feet red ; irisred. Sexes
alike. Total length 123 inches, wing 63, tail 53, culmen 2.
67. PrILOPUS LETTIENSIS.
Ptilopus cinctus lettiensis, Schleg. Mus. Pays-B. 1873, p. 35,
Columbe ; id. Neder]. Tijdsch. Dierk. tom. iv. p. 20 (1873).
Hab. Lettie, east of Timor (Hoed¢).
This bird is apparently entitled to be considered distinct from the
Pt. cinctus. I am indebted to my friend Count H. Turati for the
loan of a very fine specimen, exhibiting very clearly the differences
existing between it and P¢. cinctus. The principal one is the colo-
ration of the tail, which for one third of its length from the tip is yel-
lowish white, instead of having merely a dark grey apical band as in
Pt. cinctus. Another difference between the birds is that the last-
named species has the first primary abruptly attenuated, while that
of the present is gradually narrowed to the tip, which is sharply
pointed. The tails of both are long and square, as in P¢é. bernsteini.
Adult. Head, mantle, throat, neck, and breast ivory white; be-
neath the white of the breast is a broad blue-black band. Lower
part of abdomen and crissum greenish yellow. Under tail-coverts
brownish in the centre; rest white, margined with yellow. Back and
wings black ; first primary gradually attenuated to the tip, which is
pointed. Rump greenish grey. Tail greyish black for two thirds of its
basal half, rest yellowish white. Bill green, tip ochre; feet dark
red. Total length 112 inches, wing 63, tail 43, culmen 3.
68, PriLopus BERNSTEINI.
Carpophaga formosa, Gray, Proc. Zool. Soc. 1860, p. 360 (nec
P. formosus, Gray, ex Celebes).
1878. | MR. D, G. ELLIOT ON THE GENUS PTILOPUS, 573
Ptilopus bernsteini, Schleg. Nederl. Tijdsch. 1863, p. 59, pl. iii.
fig. 1.
Ptilonopus ochrogaster, Bernst. Journ. fiir Ornith. 1864, p. 408.
Ptilopus ochrogaster, Bernst. Nederl. Tijdsch. 1865, p. 324.
Carpophaga bernsteini, Wall. Ibis, 1865, p. 388.
Megaloprepia bernsteini, Salvad. Aun. Mus. Ciy. Gen. (1875)
vol. vii. p. 783.
Hab. Halmahera, Ternate, Batchian, Obi (Bernstein); Gilolo
(Wallace).
This and the next three species have been placed by some authors
in Reichenbach’s genus Megaloprepia. It seems very evident that
at least some of the species which he placed in his genus were not
known to Reichenbach autoptically, or else when desiring to sepa-
rate certain members of the genus Péilopus generically from the
others he would probably not have selected Pé. perlatus, with its
short tail and narrow rectrices, as one of those especially fitted to
go with the present and the succeeding ones. I fail to see any cha-
racters whatever that should cause this bird and Pt. magnificus,
with its two races, to be separated generically from Ptilopus. The gra-
dation from Pé. leclancheri, through Pt. gularis, cinctus, albocinctus,
bernsteini, puellus, and assimilis, is complete between the moderate-
sized, comparatively short-tailed species of Ptilopus and the large Pt.
magnificus, which brings the genus into Carpophaga. In his dia-
gnosis of Megaloprepia, Reichenbach gives no character that may
not equally apply to Péilopus; and it seems to me to be a division
highly artificial, and in no way necessary.
This species was first described by Gray (/. c.) ; but as he had
already bestowed the name formosus upon another species of Ptilopus,
his appellation cannot stand; and even for those authors who place
this bird in Megaloprepia, to avoid confusion, it is better to con-
tinue the name bestowed by Schlegel.
Male. Head and neck greenish grey ; breast, back, wings, and tail
bright green. In the middle of the lower part of the breast a large
spot of rich deep red. Abdomen and flanks yellowish ochre ; under
tail-coverts brownish yellow-ochre. Bill black, tip yellowish ; feet
black. Total length 114 inches, wing 5, tail 53, culmen 2.
Female. Like the male, but wants the red spot on the breast, and
has the neck light green.
It may be seriously questioned if the three following races should
be continued as distinct species. They only differ in size; and this
is very variable even among those examples acknowledged as repre-
senting one of the species. The localities of the forms also seem to
have no recognizable bounds; and while the island form puella is
the smallest, yet at Cape York it meets its large relative magnificus,
which here, according to Mr. Ramsay, is very variable in size, and
which, if the bird is distinct, ought not to be found at all in this
part of the continent, which should be reserved for assimilis. I
have no doubt that in a large series of examples from various
localities the small puel/a would graduate directly up to magnificus,
574 MR. D. G. ELLIOT ON THE GENUS PTILOPUS. [May 7,
and leave no characters by which the forms could be distinguished
apart. The specimens of the smallest race, called Pt. puella, which
are found in the island of Jobie and also at Mount Epa in the south
of New Guinea, have the under surface of the tail lighter in colour
than those from other localities, being blackish-grey instead of
blackish-brown. This, however, cannot be considered of any specific
importanee, the geographical distribution alone forbidding the idea
that they could be designated as a fourth race ; for the Pt. puella is
known to dwell between the Jobie and Mount-Epa birds as well as
to the south of the latter, which fact is alone sufficient to preclude
the supposition that two species, or even races, are indicated by the
slight difference observable. For convenience I have kept the
synonymy of the three races distinct ; but they really should only be
regarded as representing but one species.
69. PriLopus PUELLA.
Colomba amarante, Less. Voy. Coq. (text), 1826, p. 711.
Colomba puella, Less. Bull. Sc. Nat. (1827) p. 400; id. Trait.
Ornith. (1831) p. 469; Knip & Prév. Pig. vol. i. pl. i.
Carpophaga puella, Gray, List B. Brit. Mus. (1856) p. 21. sp.
18; Wall. Ibis, 1865, p. 388.
Megaloprepia puella, Bon. Consp. Gen. Av. vol. ii. p. 40; Reich.
Taub. p. 102; id. Suppl. p. 203, nov. Taf. iv. fig. 41; Ramsay,
Proc. Zool. Soc. 1876, p. 118.
Ptilopus puellus, Schleg. Mus. P.-Bas, 1873, p. 38, Columbe.
Megaloprepia puella, Salvad. Aun. Mus. Civ. Gen. (1875) vol.
vii. p. 788, vol. iv. p. 199, (1877) vol. x. p. 158.
Mapouha of the Papuans (Less.).
Hab. Cape York, Australia (Ramsay) ; Mysol, Salwatty, Wai-
giou, Ghémien, New Guinea (Bernstein, Hoedt) ; Jobie ( Von Rosen-
berg) ; Dorey (Lesson) ; New Ireland (Lesson).
The type of this species is in the Paris Museum. There are two
specimens, one from Dorey, New Guinea, and the other from New
Ireland, both of which were procured during the voyage of the
‘ Coquille.’
Adult. Front and chin greyish white; head and neck greenish
grey ; centre of throat and neck in front, and underparts to the
thighs, rich purplish red; lower part of abdomen and crissum rich
yellow. Under tail-coverts greenish grey. Wings, back, and tail
bright grass-green. Upon the outer web of the wing-coverts are
some pale yellow spots forming a line along the wing. Bill yellow,
red at base; iris orange-red ; feet yellow-green; claws dusky. Total
length 133 inches, wing 6, tail 62, culmen ?. Sexes alike.
70. PTILOPUS ASSIMILIS.
Carpophaga assimilis, Gould, Proc. Zool. Soc. 1850, p. 201.
Megatloprepia assimilis, Bon. Compt. Rend. tom. xxxix. p. 1077 ;
Reich. Taub. p. 102; id. Suppl. p. 203, nov. Taf. iv. figs. 42, 43 ;
1878.] MR D. G. ELLIOT ON THE GENUS PTILOPUS. 975
Ramsay, Proc. Zool. Soc. 1876, p. 115; Salvad. Ann. Mus. Civ.
Gen. (1876) vol. ix. p. 199.
Ptilopus assimilis, Schleg. Mus. Pays-B. 1873, p. 38, Columbe.
Hab. Cape York, Rockingham Bay, Australia (Ramsay).
This is a race of Pt. magnificus, representing it in the northern
part of Australia, and differing mainly in being smaller in size.
Whether this is sufficient to constitute a species, I do not propose at
present to discuss, but merely to remark that, as in many groups of
birds it has been very properly rejected as a specific character, it
would seem that, in the present instance, this bird and Pé. magnificus
might with advantage be retained under one specific name.
Pt. assimilis was procured by Mr. Ramsay, together with P¢.
magnificus itself, at Rockingham Bay, North-east Queensland. Ex-
cept in its smaller size, he found that it presented no differences what-
ever from Pt. magnificus. He regards it as a connecting link between
the large form and the smallest (Pt. puedla), of which form he had
received a fine pair collected at Cape York, Australia, by Mr. J. A.
Thorpe, this being the first instance known to me in which Pt. puella
has been obtained in Australia.
Head and throat grey; centre of throat and underparts to middle
of abdomen rich purple. Under wing-coverts, thighs, and under tail-
coverts rich yellow. Back and rump yellowish green. Wing and tail
bright grass-green ; wing-coverts with a conspicuous oblong spot at
the tip. Total length 14 inches, wing 7, tail 6, culmen 3.
The specimen described was obtained at Cape York by the ‘ Chal-
lenger’ Expedition.
71. PriLopus MAGNIFICUS,
Columba magnifica, Temm. Pl. Col. 163; id. Trans. Linn. Soe.
vol. xii. p. 125 (1822); Wagl. Syst. Av. sp. 26 (1827), Columba ;
Knip & Préyv, Pig. vol. ii. pl. 25; Gould, B. Austr. vol. v. pl. lviii.
(1848); Ramsay, Proc. Zool. Soc. 1876, p. 115.
Carpophaga magnifica, Selby, Nat. Libr. Ornith. vol. v. p. 119.
Megaloprepia magnifica, Reich, Tauben, p. 101, pl. 283. figs,
1299, 1300.
Ptilopus magnificus, Schleg. Mus. Pays-B. 1873, p. 38, Columbe.
Hab. South Australia, river Hunter to Moreton Bay (Gould) ;
Rockingham Bay, North-east Queensland (Ramsay).
Mr. Ramsay states in this journal (J. c.) that he found this spe-
cies abundant at Rockingham Bay in North-east Queensland, and
very variable in size.
Adult. Forehead and chin light grey ; head and neck greenish
grey, becoming light green on sides of breast ; centre of throat and
neck in front, breast, and abdomen deep purple. Lower part of ab-
domen orange-yellow. Under tail-coverts yellowish green. Wings,
back, and tail bright green; wing-coverts with large pale yellow
spots, forming a bar across the wing. Bill red at base, tip yellow.
Total length 19 inches, wing 9, tail 7, culmen 2.
or
NI
for}
MR, P. L. SCLATER ON BIRDS [May 21,
May 21, 1878.
F. D. Godman, Esq., F.Z.S., in the Chair.
The following papers were read :-—
1. Description of a new Genus of Snakes of the Family
Calamaride, from Southern India. By Lieutenant-
Colonel R. H. Beppomeg, C.M.ZS.
[Received April 29, 1878.]
XYLOPHIS, n. gen.
Body cylindrical, slender; head short, not distinct from neck,
gradually narrowed forwards and pointed ; eye very small, with round
pupil; tail about one seventh of the total length; maxillary teeth very
numerous, equal; the two palatine rows very conspicuous, and the
teeth slightly longer behind ; upper labials four, the first very minute,
the second and third enter the orbit, fourth in contact with a large
temporal ; rostral very small; an elongate loreal gradually narrowed
behind replaces anteocular and occupies all the space from rostral to
eye; nasals simple, very small; frontals only one pair, large, a very
small superciliary shield and a similar postocular. Scales smooth,
without apical groove, in fifteen rows ; anal single; subcaudals broad,
bifid, or a few occasionally entire.
XYLOPHIS INDICUS, 0. Sp.
General colour of a uniform brown like an earthworm, but beauti-
fully iridescent ; length 93 inches, of which the tail is 12 inch:
vertical, pointed behind, rounded in front; occipitals elongate ;
ventrals about 136; subcaudals about 26, bifid, or a few of them
entire ; tail ending in a blunt point.
Hab. The dense heavy evergreen forests on the mountains at the
south of Cumbum Valley, Madura district; elevation 5000 feet.
Under old logs along with Uropeltide.
The specimen, which is unique, has been forwarded to the British
Museum.
2. Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘Challenger.—No. X. On the
Birds of the Atlantic Islands and Kerguelen’s Land,
and on the Miscellaneous Collections. By P. L. Scrarer,
M.A., F.R.S.
[Received May 2, 1878.]
In this paper, which concludes the preliminary reports on the
terrestrial birds collected during the voyage of the ‘Challenger,’ I
1878.] COLLECTED BY THE ‘CHALLENGER.’ 577
give an account of the specimens obtained on the Tristan-d’ Acunha
group and Kerguelen’s Land, and of a few small collections made at
various other localities which were touched at during the course of
the expedition. I have arranged all these, according to the dates
at which the places were visited, as follows :—
Date. No. of specimens.
A. St. Iago, Cape-Verds ........ July 1873 4
B. Tristan d’Acunha .......... October 1873 12
©. werencien Land 1... ...0<).. January 1874 16
D. New Zealand ..... oie Reb June 1874 4
© Hoene-Keng :23( 50 feve) oe December 1874 7
F. Meangis Islands ............ February 1875 3
Bt SE ao EE a's bn geipiyr ren ne February 22,1875 2
H. Humboldt Bay, New Guinea .. February 24, 1875 2
PE ine Be oi eH ae ates a April 7, 1875 5
LT SPIE amines par Yet i May 1875 4
59
A. St. Iago, Cape-Verds (July 1873).
Four examples of the peculiar King-hunter of this Island, Halcyon
erythrogastra (Temm.), Sharpe’s Kingf. p. 171, pl. 63, were
obtained here. In one specimen the eyes are marked “ black,” and
the contents of the stomach “insects.”
B. Tristan-d Acunha group (October 1873).
Seven examples of the peculiar Thrush of Tristan d’Acunha, de-
scribed by Mr. Gould, P. Z.S. 1855, p. 165, as Nesocichla eremita,
were obtained. The sexes are not marked; but the specimens are all
nearly alike, so that probably the sexes do not differ in plumage. The
irides are noted as ‘‘ black.”
In the adjoining island “Inaccessible,” five specimens of the
peculiar Finch lately described by Dr. Cabanis (Journ. f. Orn, 1873,
p- 154) as Nesospiza acunhe were obtained. All five are alike ;
the eyes are marked “ hazel; ” but the sexes are not recorded.
Of Crithagra insularis, described by Dr. Cabanis at the same
time as the Wesospiza, there is no specimen in the collection, nor of
the Island-Hen (Gallinula nesiotis, Sclater, P. Z. 8. 1861, p. 261).
The latter is said to be now almost extinct’.
The only oceanic birds in the collection from the Tristan-d’ Acunha
group are two Petrels, Pelagodroma marina and Gstrelata mollis,
both from Nightingale Island, the most southern of the three islands
of the group, and one Penguin (Hudyptes chrysocome) from Inac-
cessible ; but Sir Wyville Thomson (Atl. ii. pp. 164, 178) mentions
several others.
C. Kerguelen Land (January 1874).
The only land-birds from Kerguelen Land in the collection are
1 Wyville-Thomson, ‘ Atlantic,’ ii. p. 165,
578 MR. P. L. SCLATER ON BIRDS [May 21,
nine specimens of Chionis minor and seven of Querquedula eatoni ;
and these are the only land-birds included by Mr. Sharpe in his
memoir in the Royal Society’s volume on the Transit-Expedition of
1875.
The female of the Querquedula (of which there is one example de-
termined as such, and two others, evidently of the same sex) differs
from the male not only in the absence of the green alar speculum,
but also in its smaller size, in the fuscous edgings to the feathers
generally, and in the much shorter tail. There is also one chick of
this species in the collection.
D. New Zealand (June 1874).
In New Zealand only four skins, belonging to three species, were
obtained, namely :—
1 Miro albifrons jr., Hardy Bay.
2 Hematopus unicolor jr., Hardy Bay.
1 Botaurus peciloptilus jr., Wellington.
E. Hong-Kong, China (January 1875).
From Hong-Kong seven skins, belonging to four species, all known
Chinese birds, were procured, viz :—
1 Turdus mandarinus, Bp.
1 Halcyon pileata (Bodd.).
2 Alcedo bengalensis, Gm.
2 Coturnix communis, Bonn.
1 Turnix maculosus (Temm. ).
F. Meangis Islands (February 1875).
Four examples of os indica were obtained here (of which three
have arrived home), under the circumstances mentioned in Mr. Mur-
ray’s MS., as follows’ :—
“On the 10th February, 1875, while the ship was dredging, some
natives came off in a boat. They had with them mats and cocoa-
nuts and some Perroquets. The following are those which were
purchased for tobacco :—
“440, 9.
441, Q.| Feet black; bill orange; eyes red, or light brown in
442, 2. the male.
443, ¢.
“We kept the male for several days alive; he used to fly about
the ship and return to the house on deck when shown his food.
‘‘He died from eating some green stuff, it is supposed, but was
perhaps hurt aloft.
“The natives came from the southermost isle of the Meangis
group.”
This is the most northern species of Hos known; its hitherto only
ascertained locality is the neighbouring Sanghir group of islands’.
1 Cf. also Lord G. Campbell’s ‘ Log-letters’ p. 245.
_? Cf. Rowley’s Orn. Mise. p. 123, where a beautiful figure of this species is
given.
1878. } COLLECTED BY THE ‘ CHALLENGER.’ 579
G. At sea (February 22, 1875).
Two examples of Phalaropus hyperboreus in winter plumage
(nos. 445, 446, both “ females’) were ‘‘shot by Mr. Moseley on
February 22, 1875, among the drift wood.” Mr. Murray ‘saw many
the previous day.”
This Phalarope is said to wander occasionally even as far south as
the Aru Islands!. The present examples were obtained the day
before the ship arrived at Humboldt Bay, New Guinea.
H. Humboldt Bay, New Guinea (February 23, 1875).
At this most interesting locality but two birds, unfortunately, were
obtained, namely :—
447. Trichoglossus cyanogrammus, Wag].
448. Arses insularis, Meyer. .
The first of these is a well-known Papuan species. Of the second
(shot by Mr. Murray under the circumstances mentioned in Lord
George Campbell’s ‘ Log-letters,’ p. 253)the only specimens yet known
were obtained by Dr. A. B. Meyer on the island of Jobi’. Its oc-
currence on the mainland is therefore of much interest.
Mr. Murray notes of this specimen, “ Male; eyes brown; ring
round the eye large and of a delicate sky-blue; bill and legs of a
darker blue or violet.
“IT landed in Humboldt Bay from the pinnace in a native canoe ;
and with some of the natives went a short way into the woods. Was
not much over half an hour on shore. It was then that I shot the
above two birds. I saw a Tern and a small Swift, also some
Cockatoos, but all out of shot. I fired at another small bird, but
did not get it. I also saw three Goura Pigeons, very fine large ones,
and after a time got a shot at one, but did not bring it down. I should
have had several more birds, but had to return to the boat on the
signal.”—J. M.
I. Aé¢ sea (April 7, 1875).
Four days before arriving at Yokohama, and nearly due south of
that port, five specimens of Hirundo gutturalis, the eastern form of
our familiar H. rustica, were “ caught round the ship. They were
apparently in an exhausted state.”
J. Japan (May 1875).
In Japan four specimens of birds were obtained—an example of
Buteo japonicus, Bp., in Jedo Bay, and a hen Phasianus versicolor
and two specimens of V'erekia cinerea from the Inland Sea.
! Cf. Dresser, B. Hurope.
° Monarcha insularis, Meyer, Sitz. Ak. Wien, Ixix. p. 395.
580 MR. J. WOOD-MASON ON NEW MANTID#, (May 21,
3. On new and little-known Mantide.
By J. Woop-Mason.
[Received May 7, 1878.]
(Plates XXXV. & XXXVI.)
Genus DerpHosge, Stil.
M. Stil has recently! made the Fischeria ocellata, Sauss. (the
nearest ally of 7’. laticeps), the type of a new genus, Deiphobe, and
restricted, rightly as it seems to me, the name Fischeria to Fischeria
betica and its allies, species in which, as in Hremiophila, structures
ancillary to the ovipositor* are developed upon the two terminal ven-
tral segments of the abdomen. Though Ff. /aticeps differs from
Deiphobe ocellata in the form of the eyes (which are compressed and
produced straight outwards in the form of blunt cones), in its long,
stout, and subfoliaceous cerci (which extend far beyond the extre-
mity of the abdomen), and in its more elongated and slenderer body,
it yet resembles that species so closely in all other characters that its
separation seems unwarrantable.
DEIPHOBE LaTICcEPs. (Plate XXXV. ¢ @.)
Fischeria laticeps, Wood-Mason, Ann. & Mag. Nat. Hist. 1876,
4th ser. vol. xvill. p. 337, 5; op. cit. 1878, 5th ser. vol.i. p. 144, 2.
Hab. 2 2, Bangalore district, Mysore; ¢, Sheargaon, Kolapur
State, India.
The typical specimens are in the Indian Museum, Calcutta.
HieRopuLA (RHOMBODERA) TECTIFORMIS.
Rh. tectiformis, Saussure, Mél. Orthopt. tom. 1. fase. 3, p. 216,
pl. v. fig. 19,9.
In all the numerous specimens of this species which I have exa-
mined the basal or superior angle of the facial shield is produced into
a small projecting spiniform tubercle—a point of structure not men-
tioned by M. de Saussure in his description.
The first branch of the discoidal vein of the tegmina terminates at
the sutural margin in males in two, in females in three branchlets.
Hab. I havea ¢ from Jalra-patan, captured by Mr. Cecil Tem-
pleton of the Indian Topographical Survey, a 2 from Sahibganj by
myself, anda ¢ from Doomarkoonda in the same district, and a °
nymph from Sambalpur by Mr. Valentine Ball.
Obs. The lamellar fore margin of the tegmina is entire.
Hieropuia (RHOMBODERA) BUTLERI, n. sp. (Plate XXXVI.
figs. 3, 3a, 36.)
3 Q. Very closely allied to H. (R.) tectiformis, Sauss., from which
it differs in having no projecting spiniform tubercle in the middle of
’ Bihang till k. Svenska Vet. Akad. Hand. 1877, Bandet iv. No. 10.
? T have observed similar structures in the South-African Chroicoptera vidua,
Stal. All the insects thus provided probably lay their eggs in the earth.
P.A.S: 1878-Pl. SOK.
4 io s
$eham Lal Dos del. Mintern Bros imp.
NEW MANTIDA.
PZ S676 Pl SOCK,
Mintern Bros imp.
NEW MANTIDZ..
eae bP 7 . 1 “
¥ .
1878. | MR. J. WOOD-MASON ON NEW MANTID&. 581
the superior margin of the facial shield, in having the lamellar expan-
sions of the pronotum arcuate in outline, and consequently broader
postero-laterally, and not angulate antero-laterally—in its more pointed
tegmina, the first branch of the discoidal vein of which terminates at
the sutural margin in three (3) or four (2) branchlets, instead of
two and three respectively ; the last branch of the median vein is
undivided ; and the stigma is bounded at either end by a dark blotch,
is larger, and commences as much before the origin of the third
branch of the discoidal vein as it does after it in the allied species,—
in the armature of -the fore coxee, the anterior crest of which is fur-
nished with five blunt and slightly curved conspicuous conical spines,
arranged 3+ 2, and all arising from its inner side and not from its
summit, the outer crest of which is only smoothly and obscurely
denticulate—and in the coloration of the fore femora, which are
transversely trifasciate externally, and have no conspicuous red blotch
at the base of the inner face.
The fore tibie have thirteen teeth (¢ 2) on the imner, and ten
(eleven on one side in one 2 ) on the outer edge.
The pronotum of the male is faintly constricted behind the setting-
on of the fore legs; so that ‘when viewed from above its lamellar
margins appear slightly excised.
Total length, 9 72-82 millims. ; length of pronotum, 2 20°5-
22°25, 3 15:25; greatest breadth of pronotum (at the anterior end
of its posterior lobe), 2 11°75-13°75, 3 7°75; breadth of primitive
prenotum at dilatation, 2 8°5-10°75, 3 6; length of tegmina, 2
62-69, to stigma 2 19-22; breadth of tegmina, 2 22, of marginal
field, 2 5°5; length of stigma, 2 5-6, ¢ 4:7; breadth of stigma,
2 1:3, b 0°9.
Hab. The neighbourhood of Samaguting, Naga Hills, Assam,
whence a specimen of the female was sent to me some years ago
by my late friend Captain John Butler, the Political Agent, after
whom I name it. A second and smaller specimen of the same sex
has recently been obtained by Mr. A. W. Chennell, of the Indian
Topographical Survey, in the valley of the Um-thana, W. Khasi
Hills, at about 15,000 feet elevation. But the only male I have
ever seen is a much mutilated and abnormally small insect from
Sikkim in the collection of Mr. F. Moore.
Obs. The species differs in the form of the pronotum from H.
tectiformis much in the same way as H. deflexa, Sauss., does from
H., laticollis, Burm., Sauss.
The lamellar front margin of the tegmina is entire.
Hieropuia (RHOMBODERA) FRATRICIDA, n. sp. (Plate XXXVI.
fizs. 5, 5 a.)
3. Allied to H. (2.) macropsis, Giebel, which it closely resem-
bles in the form of the pronotum, but from which it differs in its
larger size, in having the primitive pronotum apparently longer and
stouter, the facial shield higher than broad and distinctly bicarinate,
and in having the anterior crest of the fore coxee rather lower and
blunter (than usual) than produced and lamellar.
582 MR. J. WOOD-MASON ON NEW MANTIDE. [May 21,
The organs of flight are long and rather pointed, extending, when
closed, far (nearly a fourth of their length) beyond the extremity
of the abdomen. The tegmina are hyaline everywhere except in the
marginal field and along a very narrow linear space next to the
principal nervure, between the base of the organ and the stigma, in
which parts they are opaque green; the median vein gives off two
branches; and the first branch of the discoidal is forked ; the stigma
is placed just after the origin of the third branch of the discoidal vein.
The wings are throughout hyaline, save the marginal area, which
is a little clouded with green; the discoidal vein is three-branched.
The front crest of the fore coxe armed with ten to eleven low
blunt tubercles, which become successively higher and sharper
towards the distal end of the joint; and the external crest is quite
smooth ; the tibiee have ten to eleven teeth on the outer edge, and
14 on the inner; the superior margin of the femora is slightly
arcuate. The edges of lamellar margins of pronotum quite smooth.
Total length 85 millims. ; length of pronotum 28, greatest breadth
of pronotum 13:25; breadth of primitive pronotum at dilatation 8 ;
length of abdomen 31+6 (last ventral segment)=37 ; length of
tegmina 76, to stigma 20; width of tegmina 19, of their marginal
field 5-25; length of stigma 4, breadth of stigma 0°9 ; length of an-
tenne 50.
Hab. The records of the British Museum state that the specimen
was received in a collection of insects from Malabar.
Obs. The lamellar front margin of tegmina entire.
Hreropuna (RHOMBODERA) ATRICOXIS, n. sp. (Plate XXXVI.
figs. 4, 4 a, 40.)
3 Q. Very closely allied to H. (R.) laticollis, Burm., from which
it differs in having the lamellar expansions of the pronotum abso-
lutely narrower, concave instead of straight postero-laterally, and not
extended quite so far towards the base of the segment, and in having
the whole inner surface of the fore core coloured jet-black, and the
hinder end of the prosternum and the mesosternum symmetrically
marked with the same colour.
The anterior crest of the fore cox armed with minute blunt
denticles and granules ; tibize with eleven teeth ( ¢ 2) on the outer
edge, and 2 15, ¢ 14 on the inner..
Sexual differences the same as in the allied species.
Total length, 2 about 80 millims.; length of pronotum, 2 27,
3 25°5, of which the anterior lobe is respectively, 2 8 and 3 7;
greatest breadth of pronotum, 9 16, ¢ 15; breadth of primitive
pronotum at supracoxal dilatation, 2 9:5, ¢ 9; length of tegmina,
2 50, 3 64, breadth of tegmina, 2 20, ¢ 19°5, breadth of mar-
ginal field, 9 6, 3 5; length of fore coxa, 2? 18, ¢ 17; femur,
2 22, S$ 20; of intermediate femur, 2 19, ¢ 18; tibia, 17,
3 15; of posterior femur, 2 22°5, ¢ 21°5; tibia, 9 23°5, ¢ 23.
Hab. 92, Australia (C. French); 3, one of the islands in Torres
Straits (MacFarlane).
Obs. With the exception of the Indian H. (R.) tectiformis, which
1878.] MR. J. WOOD-MASON ON NEW MANTID2. 583
has a great shining red blotch at the base of each of the femora, this
is the only species of the genus which has any part of the internal
face of the fore legs conspicuously coloured’.
The lamellar fore margin of the tegmina is toothed? in both sexes,
probably so as to act as a stridulating organ by scraping against the
corresponding part of the wings.
Hirrovvuta (RHOMBODERA) PUSTULIFERA, Nl. sp. (Plate XXXVI.
figs. 6, 6 a.)
Q. Allied to H. (R.) major, Sauss., but differing in its stouter
and broader pronotum, which is more broadly rounded off in front,
in the armature of the anterior crest of the fore coxee, which is fur-
nished with eight hemispheroidal yellow tubercles or callosities, all
arising from its inner surface, gradually increasing in size from the
proximal to the middle, and then gradually decreasing to the distal
end of the joint, and constricted at the base so as closely to resemble
the crushing-teeth of certain fossil fishes (Pyenodus).
The discoidal nervure of the wings emits four branches.
The fore tibize have eleven teeth on the outer edge and fifteen on
the inner.
Total length 85 millims.; length of pronotum 29, greatest breadth
of pronotum (just behind coxal groove) 12-5, of primitive pronotum
at supracoxal dilatation 10, at middle of posterior lobe 6; length
of tegmina 57, to stigma 21; breadth of tegmina 24, of marginal
field 8; length of stigma 3, breadth of stigma 0°9.
Described from two alcoholic specimens.
A dried specimen in British Museum obtained at the same time
measures total length 80 millims, and has the coxal callosities
shrunken or less developed.
Hab. One of the islands in Torres Straits. The specimens were
obtained by the Rev. M‘Farlane.
Obs. The lamellar fore margin of the tegmina toothed.
Hreroputa (RHOMBODERA) TAPROBANE. (Plate XXXVI.
figs. 7, 7 a.)
Hierodula taprobane, Wood-Mason, Ann. & Mag. Nat. Hist. 1878,
5th ser. vol. i. p. 146, 2.
Hab. Ceylon.
Obs. The lamellar fore margin of the tegmina is smooth.
ARCHIMANTIS MONSTROSA,n. sp. (PlateXX XVI. figs. 1, 1 a, 1.)
Mantis monstrosus, Bates, MSS., in coll. Brit. Mus.
2. Closely allied to.4.armata, W.-M., 9, differing in its greater
size, in its proportionally longer tegmina (which are fully equal to
the prothorax in length), in the form of the pronotum (which is
dilated at the setting-on of the fore legs, so as to be conspicuously
i ? These coloured femora serve, in all probability, to allure or fascinate the
eA
rey.
* I first met with this curious structure of the tegmina in the Empuside, in
all of which it occurs in both sexes alike.
584 MR. J. WOOD-MASON ON NEW MANTID&. [May 21,
ovate-cordiform in front), and in the form of the cerci, the margins
of the foliaceous terminal joints of which are arcuate.
Fore tibize with sixteen or seventeen teeth on the inner and ten on
the outer edge.
Total length 115 millims.; prothorax 41, of which the neck is
10; width of prothorax at supracoxal dilatation 10; height of head
8, breadth of head 11:75; length of meso- and metanotum taken
together 19, of abdomen 53, of cerci 12°75; breadth of terminal
joint of cerci 2; length of fore coxa 22:5, femur 28, tibia to insertion
of tarsus 11; of intermediate femur 27, tibia 27; posterior femur 33,
tibia 38; tegmina 42 (or reaching to end of middle third of third
abdominal segment); breadth of marginal field of tegmina 3; length
of stigma 6°75; length from base of tegmen to stigma 14; length
of wings 35.
Hab. Victoria River, N. Australia. The type in British Museum
was ‘‘captured on March 10th, 1856, by R. Elsey, Esq., during the
exploring expedition under Mr. Gregory.”
ARCHIMANTIS ARMATA. (Plate XXXVI. figs. 2, 2 a.)
A. armata, Wood-Mason, Ann. & Mag. Nat. Hist. 1877, 4th ser.
vol. xx. p. 76.
Hab. North Australia (C. French).
ANTHALOCHROA ASHMOLIANA.
Vates ashmolianus, Westw. Ann. & Mag. Nat. Hist. vol. viii.
p- 272; Arcana Entomol. 1843, vol. ii. p. 52 (note f).
Popa? aschmoliana, Saussure, Mél. Orthopt. tom. i. fase. 3, p. 309.
Asthalochroa ashmoliana, Wood-Mason, Aun. & Mag. Nat. Hist.
April 1877, p. 308, 3 2.
Arsacia ashmoliana, Stil, Sv. Vet. Akad. Hand. 1877, Bd. iv.
no. 10, 75, 2.
A male (var. simplicipes) from Bombay, in the collection of Mr.
F. Moore, differs from the type and from the specimens from Ben-
gal (Calcutta and Murshidabad) in not having the crests of the four
posterior femora developed into foliaceous lobes at the apex, and in
having the upper crest of the four posterior tibize more developed
and with more regularly arcuate margins, and in having the ridge of
the posterior lobe of the pronotum more convex longitudinally.
A gigantic female (var. insignis) in the India (olim E.-I. Com-
pany’s) Museum, South Kensington, marked “N. India,” has the
lobes at the extremities of the four posterior femora more highly but
less regularly developed ; this specimen measures :—
Total length 132 millims., length of prothorax 48, of which the
neck is 10, abdomen 61—as against 115, 40, 9:5, and 52, the mea-
surements of the same parts in the Calcutta specimen of the same
Sex.
A female in the collection of Mr. W. B. Farr (though not labelled
as to locality, doubtless from Bengal) agrees exactly with the spe-
cimens from Calcutta and Murshidabad,
1878.] MR, J. WOOD-MASON ON NEW MANTID. 585
Genus CREOBROTER’, Serville.
CREOBROTER PICTIPENNIS, n.sp. (Plate XXXVI. fig. 8.)
2. Closely allied to Cr. apicalis, Sauss. (2), from Assam, from
which it differs in the form of the pronotum (which is slenderer and
more distinctly quadrilobular), in its much narrower and less-de-
pressed abdomen (a part which in the species mentioned, as also in
Cr. urbanus 2, is greatly dilated and suborbicular and depressed),
in its less rebust fore legs, and notably in the colour of the wings.
The head also is a littie smaller, its frontal spine much the same.
Organs of flight in repose, in the dried insect, extending by about
a seventh of their length beyond the extremity of the fusiform ab-
domen, which is longitudinally strongly roof-shaped below. 'Teg-
mina bright green, yellow at the very base, furnished with a well-
defined subquadrate or suborbicular brightyellow spot a short distance
from the base, and with a still richer dead-yellow oval ocellus placed
a little obliquely across the middle; this is bordered both on its ex-
ternal and on its internal margin by a curved black line; the black
line that borders the base of the ocellus is but slightly curved or is
a smaller segment of a larger circle, and sends forwards and inwards
at its posterior end a small process, so as to present the appearance
of a curved arrow that has lost one of its barbs; external black
line strongly curved, or a larger segment of a smaller circle; it
is preceded by an opalescent line rather broader than itself, and upon
the veins and veinlets traversing which the yellow of the ocellus is
continued; this is marked with three black dots, all placed in the
same straight or slightly curved line, each upon one of the sectors of
the organ; the yellow of the ocellus flows out, as it were, in an irre-
gular stream from between the black bounding lines to the sutural
margin, accompanied by the translucent white line only ; but at the
opposite side its exit is barred by the polished and slightly paler
yellow oblong stigma, which there quite blocks up the interval be-
tween the ends of the black lines; the anal gusset is spotted with
brown, paling towards the hinder margin, the network being pale
green, very narrowly lined with hyaline, and its membranous meshes
brown. Wings tricolorous, opaque at the base; the basal third of
the posterior and the basal two thirds of the anterior field of a beau- —
tiful semiopaque cream-colour, slightly tinged with green ; this is
succeeded in the former by a broad band of clear brown (with dark
amethystine reflections) lined with hyaline along the transverse vein-
lets, and in the latter by a narrower band of semiopaque dull vina-
ceous, the rest of both fields being throughout hyaline, slightly tinged
and clouded with very pale greenish white, and having the venation
rather distinct and also coloured pale greenish white.
The head, the pronotum, and the outsides of the fore legs bright
green, concolorous with the elytra, variegated (the two first-named)
and banded (the last) with yellow, much, in fact, as in Cr. apicalis.
1 Neither “Creobrota” nor “‘Creobotra,” nor “Creoboter,’ but ‘‘Creobroter,”
from xpéas, flesh, and Bowrhp, an eater, kpeoBpwrhp, so formed on the analogy
of Kpeorwdys, Kpeopayos, K. T. A.
Proc, Zoox. Soc.—1878, No. XX XVIII. 38
586 MR. J. WOOD-MASON ON NEW MANTID#. — [May 21,
The anterior crest of the fore coxe is armed with eight bluntish
spiniform tubercles, between some of which are one or two minuter
ones ; the right fore tibia has fifteen teeth on each edge ; the four
posterior legs are wanting.
Total length about 32 millims. ; length of pronotum 7:6; breadth
of pronotum at supracoxal dilatation 4°3, at constriction behind dila-
tation 2°6 ; length of tegmina 26; length of abdomen 14, breadth
of abdomen 8°5.
Hab. Ceylon. Communicated by Mr. F. Moore. There is also
a specimen in the Hopeian collection at Oxford.
HyMENOPUS BICORNIS.
Mantis bicornis, Stoll, Spectres et Mantes, 1787, pl. xi. figs. 44
(2), 44a (nymph). '
Mantis coronata, Olivier, Encyl. Méth. 1792, Insectes, t. vii.
. 638.
f Empusa coronata, Latreille, Gen. Crust. et Ins. t. i. p. 91.
Hymenopa coronata, Serville, Revue, p. 19. no. 1, et Hist. Nat.
des Orthopt. 1839, p. 163 9.
Hymenopus coronatus, Burm. Handb. d. Entom. 1839, vol. i.
. 549.
‘ Hymenopus bicornis, Saussure, Mel. Orthopt. i. fase. 3, p. 291.
The larve and nymphs, at any rate, of this species simulate blos-
soms, some specimens being rose-pink, others of a beautiful wax-
white colour. The thighs of the four posterior pairs of legs are ex-
panded into huge broad pear-shaped plates ; so that (according to
Mr. S. E. Peal, who has discovered the species in Assam, many
hundreds of miles from any of its hitherto recorded habitats) the
immature insect when seated on a twig with thorax and abdomen
raised at right angles to one another, with the fore legs drawn up
out of sight beneath the thorax, and with the four expanded thighs
spread out two on each side, the tarsi being brought to one spot,
presents in form as well as in colour a most perfect and deceptive
resemblance to a flower. The small Mantis (exactly resembling a
pink orchis-flower) which, according to Mr. Wallace, was shown to
Sir Charles Dilke in Java, and was said to attract and devour butter-
flies, probably belongs to the same species.
Hab. Sibsagar, Assam (S. #. Peal); Moluceas (Serville) ; Sunda
Isles. The perfect insect is represented in one of the beautiful
sketches of foliage and flowers made by Miss North in Java,
PARABLEPHARIS KUHLII.
Blepharis kuhlu, De Haan, Orthopt. Orient. p. 93, tab. 18.
fig. 3, 9.
Se idbiiphlers kuhlii, Saussure, Bull. Eutom. Suisse, iii. p. 223
(1870); id. Mél. Orthopt. i. 1871, fase. 3, p. 320, 9.
Hab. Java (De Haan) ; *‘ les Indes Orientales?” (Saussure). A
larva has been obtained by Mr. A. W. Chennell, of the Topographical
Survey of India, in the Naga Hills, Assam.
1878.] MR. BATES ON NEW GEODEPHAGOUS COLEOPTERA. 587
EXPLANATION OF THE PLATES,
{N.B. All the figures are of the natural size, except where the contrary is stated.)
PuatE XXXV.
Hig. 1. Deiphobe laticeps, p. 580. Male, with wings extended.
2. Deiphobe laticeps. Female.
2a, The right fore femur and tibia, from the inner side.
26. Front view of the head of female, magnified twice nat. size.
Prats XXXVI.
Fig. 1. Archimantis monstrosa, p. 583, 2. The head, pronotum, and fore legs.
1a. Front view of the head of the same.
1. One of the cerci from the side, enlarged.
2. Archimantis armata, p. 584, 2. The head and pronotum.
2a. One of the cerci from the side, enlarged.
3. Hierodula (Rhembodera) butleri, p. 580, 2 d. Head, pronetum, and
left fore leg of a female.
3a. The right fore coxa, from the inside.
3). Head and pronotum of a small male.
4. Hierodula (Rhombodera) atricowis, p. 582, 2 §. Head, pronotum, and
right fore leg of a female.
4a. The left fore coxa, from the inside,
4b. Head, pronotum, and right fore leg of a male.
5. Hierodula (Rhombodera) fratricida, p. 581, g. Head, pronotum, and
left fore leg.
5a. Right fore coxa, from the inside.
6. Hierodula (Rhombodera) pustulifera, p. 583, 2. Head, pronctum, and
left fore leg.
6a. Right fore coxa, from the inside.
7. Hieredula (Rhombedera) taprebane, p. 583, 9. Head, pronetum, and
right fore leg.
7a. Left fore coxa, from the inside.
8. Creobroter pictipennis, p. 585, 2. Head, pronotum, and fore legs,
4. On new Genera and Species of Geodephagous Coleoptera
from Central America. By H. W. Barss, F.L.S.
[Received May 11, 1878.]
The following diagnoses of new Central-American Coleoptera, of
the families Cicindelidze and Carabide (Geodephaga), are preparatory
to the full descriptions which will be given in the work now pre-
paring by Messrs. Godman and Salvin, entitled ‘Biologia Americz
Centralis.’ In that work figures will be given of all the more
characteristic forms. ‘
Family CicinDELIDé.
TETRACHA IGNEA.
T. carolinee forma similis, paullo magis angustata, supra tgneo-cuprea;
epistomate, thoracis et elytrorum lateribus viridi-eneis ; elytris
grosse confertim usque ad apices punctatis, macula apicali pallida,
angusta, antice gradatim paullo dilatata ; antennis articulis 1™°—4"™
subtus nigro-lineatis ; femoribus quatuor BOSC AURCe late
38
588 MR. H. W. BATES ON NEW [May 21,
nigris : corpore subtus viridi-eneo, ventris apice utringue flavo-
fasciato.
Long. 8lin. ¢ Q.
Hab. Chiriqui.
CICINDELA BELTI.
C. roseiventri affinis, apud elytrorum humeros latior, his rectangulis.
Saturate olivacea, elytris utrinque maculis albis quingue, scilicet
una humerali, secunda antico-discoidali paullo transversa, tertia
mediana lineiformi recte transversa, quarta postertore rotundata
prope suturam, quinta prope angulum posteriorem transversa ;
subtus viridi-enea, lateribus cupreis et dense albo-pilosis ; femo-
ribus infra cupreis ; abdomine rufo, bast eupreo-nitida,
Long. 53 lin. oC.
Hab. Chontales, Nicaragua (Belt).
CICINDELA FLOHRI.
Elongata, capite angusto, thoracis lateribus anticis paullo rotundatis ;
elytris elongato-ovatis, apice obtuse rotundatis; supra rufo-
cuprea subopaca, sericeo-micans, thoracis sulcis elytrorumque
punctulis viridibus ; elytris utrinqgue macula alba humerali et
altera post humeros, fascia mediana omnino discoidali obliqua, et
maculis quatuor subapicalibus, quarum tribus marginalibus ;
corpore subtus lete chalybeo, episternis rubro-cupreis, parce
pilosis ; abdomine toto lete rufo.
Long. 5 lin. of Q.
Hab. Mexico; San Angel, Guadalupe, near the capital (Flohr).
CTENOSTOMA SIGMA.
Ad sectionem Procephalus pertinet. Cylindricum, @eneo-fuscum
nitidum, sparsissime griseo-hirtum, antennis pedibusque dilutiori-
bus ; capite impunctato, supra inequali, fronte plana utrinque
sulcata ; elytris disperse punctulatis, apices versus sublevibus,
apice obtuse sinuato-truncatis, utrinque macula parva anteriore
subsuturali, et pone medium fascia valde flexuosa, S-formi, testaceo-
albis.
Long. 7 lin. 3 Q.
Hab. Chontales, Nicaragua (Belt; Janson).
CTENOSTOMA LETICOLOR.
Ad sectionem Procephalus pertinet. Cylindricum, gracile, lete
viridi-auratum, glabrum, antennis (scapo metallico excepto) tibiis
tarsisque piceis ; capite impunctato, fronte tubere magno rotun-
dato, sulcis antice et postice marginato, verticis lateribus etiam
sulcatis ; elytris pone scutellum subumbonatis, disco prope
suturam undulatis punctisque ocellaribus impressis, reliquis
sparsissime, versus basin grosse, punctatis, apice breviter sinuato-
truncatis, utringue macula parva anteriore subsuturali, et pone
medium fascia valde obliqua intus hamata, testaceo-albis.
Long. 53 lin.
Hab. Chontales, Nicaragua (Janson).
1878. | GEODEPHAGOUS COLEOPTERA. 589
Family Caranips.
ASPIDOGLOSSA BRACHYDERUS, D. sp.
In hoc genere brevis; thorace brevi et lato. Nigro-enea, palpis
antennis pedibusque piceo-testaceis, elytris apicem versus diffuse
rufescentibus ; clypei margine haud emarginato, fovea frontali
parva, simplici ; thorace lato, convexo, antice angustato, lateribus
Sereutin A. subangulata angulatis ; elytris oblongo-ovatis, versus
basin paullo angustatis, fortiter striatis, striis passim crenatis,
interstitio tertio tripunctato.
Long. 23 lin.
Hab. Panama.
STENOUs OLIVACEUS.
Hlongato-ellipticus, gracilis, supra eneo-fuscus, nitidus, subtus niger,
pedibus piceis, femoribus obscurioribus ; antennis rufo-piceis,
articulis 1»°—3" pallidis ; elytris striatis ; limbo alutaceo-sub-
opaco, viridi-tincto, striis obsoletis.
Long. 3? lin.
Hab. Mexico (Boucard).
DIC@LUS FLOHRI.
Oblongo-ellipticus, niger vix nitidus ; capite minus robusto, collo
angustiore ; thorace antice puullulum angustato, lateribus vix
rotundatis, omnino levi; elytris striis omnibus obsoletis, seriebus
punctorum suturali et marginali tantum eastantibus, carina
humerali brevi, valde elevata.
Long. 93-10 lin. of Q.
fIab. Mexico, San Angel (FloAr).
PoLPOCHILA MEXICANA.
Angusta, parva, castanea, palpis, antennis basi, pedibus margineque
apicali elytrorum flavo-testaceis ; foveis frontalibus usque ad
oculorum marginem posticum fortiter impressis; thorace late
cordato, lateribus ante basin fortiter sinuatis, angulis posticis
exstantibus ; elytris profunde striatis.
Long. 3 lin. C.
Hab. Mexico, Vera Cruz (Boucard).
Specimens have been distributed under the name I have adopted,
given by M. de Chaudoir.
ANISODACTYLUS ROTUNDANGULUS.
Elongatus, niger, modice nitidus, pedibus piceis, antennis palpisque
piceo-rufis ; thorace lateribus arcuutis, angulis posticis tate
rotundatis, foveis posticis latis, modice impressis ; elytris apice
haud sinuatis, supra simpliciter striatis,; impunctatis.
Long. 7-7} lin. 3 Q.
Hab. Mexico, near the capital (Flohr).
NorIoBiA DISPARILIS.
AGneo-cuprea (elytris 2 cupreo-fuscis), glabra; epistomate, labro,
590 MR. H. W. BATES ON NEW [May 21,
palpis, antennis et pedibus piceo-rufis; thorace valde transverso,
ante medium late rotundato, deinde vix sinuatim angustato, an-
gulis posticis rectis, anticis omnino rotundatis, supra impunctato,
fovea basali utrinque magna irregulari, subrugata: elytris
thorace basi latioribus, apice profunde sinuatis, supra (3)
fortiter striatis, interstitiis dorso subplanis, duobus marginalibus
alutaceo-opacis ; 9 haud striatis, opacis, sutura costisque
tribus elevatis nitidis.
Long. 6-7 lin. 3 Q.
Hab. Chontales, Nicaragua (Belé).
NOTIOBIA PARILIS.
Glabra, capite thoraceque chalybeis, elytris cupreo-violaceis, subtus
nigra, antennis, pedibus antennisque interdum piceo-rufis ; thorace
valde transverso, ante medium late rotundato, deinde sinuatim an-
gustato, angulis posticis rectis, anticis omnino rotundatis ; elytris
utroque sexu politis, sulcato-striatis, interstitiis omnibus valde
convexis, apice sinuatis (nec dentatis).
Long. 63 lin. 3 Q.
Hab. Chontales, Nicaragua (Belt).
NorioBiA LIMBIPENNIS.
Subcupreo-enea, nitida; thorace valde transverso, medio rotundato-
dilatato, postice modice angustato, angulis posticis vix reetangulis ;
elytris oblongo-ovatis, apice fortiter sinuatis et extus dentatis,
profunde striatis, limbo laterali (apicem versus latiore) alutaceo
subtestaceo-opaco, fovea etiam lata antico-discoidali paullo alu-
taceo-opaca, interstitio tertio postice unipunctato; antennis,
palpis et pedibus rufo-piceis.
Long. 53 lin. oC.
Hab. Chontales, Nicaragua (Belt).
NoTIOBIA LEIROIDES.
Latior, supra eneo-olivacea, labro, antennis basi, palpis et pedibus
rufis ; thorace valde transverso, medio rotundato-dilatato, postice
angustato, angulis posticis rectangulis ; elytris apice paullulum
sinuatis, regulariter striatis, interstitiis equalibus, apicem versus
angustioribus, marginalibus subopacis, tertio postice unipunctato,
margine apicali testaceo.
Long. 7 lin. 9.
Hab. Mexico, Vera Cruz (Boucard).
NOTIOBIA CUPREOLA.
Angustior, polita, nigro-enea, elytris cupreo-violaceis ; antennis
(articulis 1™°-3"™ nigris exceptis) labro, palpis et tarsis fulvo-tes-
taceis ; thorace lateribus antice valde rotundatis, postice vix sinu-
atim angustato ; angulis posticis fere rectis, supra impunctato,
foveis basalibus latis vagis ; elytris (3 2) apice oblique leviter
1878.] GEODEPHAGOUS COLEOPTERA. 591
sinuatis, supra striatis, 2 subtilissime alutaceis sed politis,
interstitio 3° postice unipunctato.
Long. 43 lin. ¢ Q.
Hab. Costa Rica, Irazu, alt. 6000-7000 feet (Rogers).
HARPALUS ALIENUS.
Oblongus, modice convexus, niger nitidus ( 2 elytris sericeo-opacis),
pedibus piceis, ntennis palpisque rufo-piceis, illis articulis 2°-4""
basi nigris ; thorace transversim quadrato, postice plusquam
antice angustato, lateribus rotundatis, angulis posticis obtusis,
impunctato, basi coriaceo, fovea utrinque late impressa ; elytris
apice (3b 2) sinuatim subtruncatis, fortiter simpliciter striatis,
interstitiis paullo convexis, 3 postice unipunctato.
Long. 53-6 lin. ¢ Q.
Hab. Mexico, near the capital (Flohr).
SELENOPHORUS AMBLYDERUS.
Oblongus, converus, nigro-eneus, antennarum articulo basali palpis-
que rufis, pedibus picescentibus ; thorace lateribus arcuatis, an-
gulis posticis rotundatis, fovea basali utrinque parva, margine
medio multistriato ; elytris glabris, striatis, striis apice pro-
fundioribus, interstitiis 3°, 5°, et 7"° punctis seriatis parvis.
Long. 34-43 ln. o Q.
Hab. Mexico, near the capital (Flohr).
SELENOPHORUS CALLISTICHUS.
Oblongo-ovatus, elongatus, minus convexus, cupreo-eneus ; antennis
basi pedibusque flavo-testaceis ; thorace antice gradatim, versus
basin citius, angustato, angulis posticis rotundatis, basi tota
haud profunde punctulata ; elytris apice sinuatis, supra fortiter
striatis, interstitiis alternis punctis numerosis rotundatis seriatim
impressis ; tarsis posticis gracilibus, elongatis ; tibiis et tarsis
absque spinis fortibus.
Long. 44 lin. oO.
Hab. Panama (Salvin).
SELENOPHORUS TENUISTRIATUS.
Oblongus, cupreo-eneus, antennis basi, palpis, tibiis et tarsis rufes-
centi-testaceis ; thorace transversim quadrato, antice minus et
citius quam postice modice angustato ; elytris apice haud sinuatis.
tenuiter striatis, interstitiis planis, alternis conspicue seriatim
punctatis.
Long. 23 lin. ¢ Q.
Hab. Mexico, near the capital (Flohr).
SELENOPHORUS MITIS.
Oblongo-ovatus, niger, vix eneo vel chalybeo tinctus ; palpis anten-
nisque fulvo-testaceis, labro et pedibus piceis ; capite levi, foveis
frontalibus obsoletis ; thorace quadrato, levi, foveis basalibus
Jatis, vagis, lateribus paullulum arcuatis, antice gradatim, postice
592 MR. H. W. BATES ON NEW [May 21,
vie angustato, angulis posticis obtusis ; elytris apice leviter
sinuatis, supra striatis, interstitiis alternis punctis parvis seriatis ;
tibiis anticis sparsim spinosis, tarsis dilatatis, 3 subtus pauct-
squamosis.
Long. 33 ln. ¢ 9.
Hab. Mexico, near the capital (Flohr).
ATHROsTICTUS, nov. gen. Selenophoro affine.
Corpus elongato-ovatum, supra (precipue elytris) dense minute
punctatum. Caput ovatum, postice gradatim paullo angustatum.
Oculi magni sed non prominentes, postice orbitu inclusi. Antenne
filiformes, graciles. Mentum profunde emarginatum, sinu fundo
haud dentatum. Ligula angusta, apice libera, longe bisetosa ;
paraglossis latis ea longioribus. Elytra haud conspicue seriato-
punctata. Caeteris ut in generibus Hypolitho et Harpalo.
Ad hoe genus Selenoph. epeciosus (Dej.), Pangus metallicus
(Reich.), Hypolithus puberulus et H. chlenioides (De}j.), et Harpalus
sulcatulus (De}.) pertinent.
ATHROSTICTUS OPALESCENS.
Oblongus, piceus, breviter erecte pubescens ; labro, palpis, antennis
et pedibus rufo-fulvis; capite et thorace nitidis, elytris opales-
centi-micantibus ; capite postice punctulato; thorace quadrato,
lateribus fortiter arcuatis, angulis posticis fere rotundatis,
basi late subrugose punctulatis; elytris apice leviter sinuatis,
exarato-striatis, interstitiis minute crebre granulatis, 3'° et 5
indistincte seriatim punctatis ; corpore subtus nigro, lete opales-
cente ; metasterno punctulato.
Long. 4 lin. of Q.
Hab. Panama (Salvin).
ATHROSTICTUS SERICATUS.
Latior, oblongus, niger, omnino punctulatus, breviter erecte fulvo-
pubescens ; antennis, palpis et tarsis obscure rufescentibus ;
thorace transversim quadrato, postice viv angustato, angulis
anticis haud prominulis, posticis fere rectangulis, latertbus
paullulum arcuatis ; elytris lete awreo-sericeo relucentibus, fortiter
striatis, ‘interstitiis densissime punctato-granulatis ; corpore
subtus nigro nitido, opalescente, punctulato.
Long. 63 lin. 2.
Hab. Mexico, Vera Cruz (Boucard).
BRADYCELLUS LUCIDUS.
Testaceo-rujis ; elytris piceo-fuscis, politis, striis (suturalt, mar-
ginalibusque exceptis) nullis; meso- et metasternis ventrisque
basi nigris; capite latiore ; thorace vix elytris angustiore, postice
valde sinuato-angustato, angulis posticis reetis.
Long. 3 lin.
Hab. Mexico.
1878. ] GEODEPHAGOUS COLEOPTERA. 593
BrADYCELLUS SUBOBSOLETUS.
Castaneo-fuscus, subtus nigricans, antennis basi pedibusque _flavo-
testaceis ; elytris strits suturali et marginalibus profundis, disco
striato-punctatis.
Long. 23-3 lin.
Hab. Mexico, near the capital (Flohr).
ACUPALPUS CIRCUMDATUS.
Elongato-oblongus, castaneo-fuscus ; antennis basi, palpis, pedibus
margineque angusto elytrorum flavo-testaceis ; thorace postice
valde angustato, levi, basi utringue haud foveato, parum punc-
lato, lateribus antice fortiter rotundatis, pustice sinuatis, angulis
posticis prominulis rectis ; elytris striis modice impressis, haud
punctatis, striola scutellari nulla.
Long. 24 lin.
Hab. Mexico, Vera Cruz (Boucard).
ACUPALPUS FLOHRI.
Elongato-oblongus, castaneo-rufus, capite supra, elytris utrinque
disco, abdomineque castaneo-fuscis ; capite fronte foveolata ;
thorace lateribus usque angulos posticos obtusos rotundatis, basi
utringue grosse sparsim punctato ; elytris simpliciter striatis,
striola scutellari nulla.
Long. 2 lin.
Hab. Mexico, near the capital (Flohr).
ACUPALPUS OBESUS.
Latior, oblongo-ovatus, castaneo-fuscus, thorace et elytris indistincte
castaneo-rufo limbatis ; palpis, antennis basi pedibusque fulvo-
testaceis; thorace transverso, sublunato, lateribus omnino rotun-
datis et angulis posticis fere nullis, supra impunctato ; elytris
subpunctulato-striatis, striolu seutellari elongata, obliqua.
Long. 14 lin.
Hab. Mexico, near the capital (Flohr).
ANCHUMENUS MONTEZUM&.
Elongatus, gracilis, piceo-niger ; capite ovato ; oculis haud pro-
minentibus, orbitu posteriore sensim angustato ; thorace elongato,
truncato-cordato, lateribus explanato-reflexis, angulis posticis
rectis ; elytris convexis, elongato-ovatis, punctulato-striatis, apice
oblique sinuatis, marginibus basali et laterali explanato-reflexis ;
pedibus elongatis.
Long. 43-53 lin.
Hab. Mexico, near the capital (Flohr).
ANCHOMENUS TRANSPUNCTATUS.
A. anthracino (Dej.) affinis, oblongo-ovatus, niger, subtus cum
pedibus nitidus; elytris sericeo-nitentibus ; capite et thorace
eyaneo-tinctis ; antennis articulo basali subtus rufo-piceo ; capite
- parvo, pone oculos gradatim angustato ; thorace elytris multo
594 MR. H. W. BATES ON NEW [May 21,
angustiore, rotundato, postice magis quam antice angustato ;
elytris tenuiter striatis, stria tertia 3-, stria secunda 2-punc-
tata.
Long. 4lin. ¢ Q.
Hab. Mexico, near the capital (Flohr).
ANCHOMENUS SCUTIFER.
Elongato-ovatus, niger, nitidus, supra leviter cyaneo-tinctus ;
antennis articulo basali rufo; capite levi, oculis vix prominulis ;
thorace magno, ovato, levi ; elytris thorace vix latioribus, acute
et fortiter striatis, interstitiis planis, tertio 4—5-punetato.
Long. 43-53 lin. ¢ Q.
Hab. Mexico, near the capital (Flohr).
ANCHOMENUS VIXSTRIATUS.
Parvus, gracilis, fusco-eneus politissimus ; palpis, antennis basi,
pedibusque fulvo-testuceis ; thorace subcordato-ovato, angulis
rotundatis ; elytris ovatis, striis vix impressis, interstitiis paul-
lulum convexis 2% bi- vel tripunctato ; metasterni episternis
brevibus, latis, epimeris transversis.
Long. 2? lin. ¢ Q.
Hab. Mexico, near the capital (Flohr).
ANCHOMENUS CONCISUS.
A. vixstriato affinis, metathoracis episternis brevibus. Fusco-
eneus, modice nitidus ; palpis, antennis, pedibus, murgineque
deflexo elytrorum fulvo-testaceis ; capite collo transversim de-
presso ; thorace subcordato, postice sinuatim angustato, angulis
posticis fere rectis, margine laterali explanato-reflexo, fulvo ;
elytris ovatis apice oblique sinuatis, supra acute striatis, inter-
stitits planis, tertio tripunctato.
Long. 3-37 lin.
Hab. Mexico, near the capital (Flohr).
ANCHOMENUS SUFFECTUS.
A. conciso simillimus : differt elytris amplioribus, thorace an-
gustiore marginibus lateralibus anguste reflexo-marginatis,
postice magis sinuatis, antice angulatim rotundatis. Castaneo-
Suscus ; palpis, antennis pedibusque fulvo-testaceis, margine an-
gusto thoracis, epipleurisque elytrorum obscure testaceis ; elytris
acute striatis, apice oblique fortius sinuatis, interstitio 3” tri-
punctato.
Long. 33 lin.
Hab. Mexico (Boucard).
ANCHOMENUS NUGAX.
A. conciso affinis: differt thorace latiore, postice multo mimus at-
tenuato, antice distincte angustato. Castaneo-fuscus, vel ceneo-~
fuscus, nitidus ; palpis, antennis et pedibus prceis, raro fulvo-
testaceis ; thorace quadrato, paulo ante medium subangulatim
1878.]} GEODEPHAGOUS COLEOPTERA. 595
dilatato, deinde postice paulo angustato, angulis posticis rectis,
margine laterali anguste explanato-refleco vix pallidiore ; elytris
minus rotundato-ovatis, apice viv sinuatim oblique truncatis,
supra acute striatis, interstitiis planis, 3% tripunctato ; margine
deflexo interdum testaceo.
Long. 3} lin. 3 2
Hab. Mexico, near the capital (Flohr).
ANCHOMENUS SIMPLICIOR.
A. nugaci provime affnis : differt tantum thorace paullulum
angustiore, lateribus antice equaliter rotundatis nullomodo
angulatis, postice vix conspicue sinuatim modice angustato, an-
gulis fere rectis, margine laterali angustissime explanato haud
pullido : piceo-niger, viv eneo-tinetus, antennis palpis et pedibus
prceo-rufis ; elytris ovatis, subtiliter striatis etc, ut in precedenti-
bus.
Long. 3 lin.
Hab. Mexico, near the capital (Flohr).
GLYPTOLENUS, nov. gen. Anchomeno affine.
Corpus gracile. Caput angustum, ovatum. Mentum sinu dente
magno elongato. Palpi nudi, articulis terminalibus cylindricis.
Thorax angustus, ante medium angulatim dilatatus, dorso grosse
transversim rugatus. Elytra ampla, convewa, apice haud
stnuata, supra fortiter suleata. Prosternum apie marginatum,
prominulum. Metasterni episterna elongata. Pedes graciles ;
tibiee et tarsi fortiter sulcata ; tarsorum articulis 1°-4"™ latitu.
dine equales, plantis pubescentibus, 4 emarginato.
GLYPTOLENUS RUGICOLLIS.
Niger, subnitidus; thorace capite vix latiore, oblongo, postice
sinuatim paulo angustato, angulis posticis subrectis, supra grosse
rugato; elytris latis, convewis, apice rotundatis, swpra grosse
profunde striatis, interstitiis convexis ; antennis, palpis, tibiis
et tarsis piceo-fulvis. 9°.
Long. 33 lin.
Hab. Chontales, Nicaragua (Belt).
COLPODES STRICTICOLLIS.
Hlongatus, niger, nitidus ; palpis, antennarum articulis 49-11“
tarsisque piceo-rufis ; thorace fere orbiculari, basi fortiter con-
stricto, marginibus lateralibus rotundato-explanatis, supra
levi, polito ; elytris convenris, marginibus lateralibus explanatis
levibus, apice vie sinuatis, supra valde convexis, suleato-
striatis.
Tarsi articulo 4% latiusculo, emarginato, subtus sublonge setoso,
articulis 1°-3"" quatuor posteriorum supra bisulcatis.
Long. 6 lin.
Hab. Chontales, Nicaragua (Belt, Janson).
J
96 MR. H. W. BATES ON NEW [May 21,
CoLPODES INTERGENEUS.
Gracilis, piceo-niger ; elytris purpureo-fuscis ; palpis, antennis
pedibusque pallide testaceis; capite collo supra transversim
depresso; thorace angusto, quadrato, antice paulo rotundato-
dilatato, postice vix sinuatim modice angustato, angulis posticis
distinctis subobtusis, margine laterali angusto ; elytris elon-
gato-ovatis, apice oblique (haud sinuatim) truncatis, supra
subtiliter subpunctulato-striatis, interstitus planis, tertio 3-
punctato.
Tarsi 2 antici articulo 4” latiusculo, breviter bilobo, subtus longe
setoso.
Hab. Costa Rica, Irazu, 6000-7000 feet (Rogers).
CoLPoDES GRATUS.
Elongato-ovatus, convexior, subeneo-niger; palpis, antennis,
tibiis et tarsis piceo-rufis; palpis articulo ultimo sublineari ;
thorace ovato, lateribus equaliter rotundatis, paullo ante basin
sinuatim citius angustatis, angulis posticis subobtusis, margine
@qualiter anguste explanato ; elytris apice leviter sinuatis, in-
terstitiis paululum convexis, 3° puncto unico solum conspicuo.
Tarsi articulo 4° subtus longe setoso, quatuor anteriorum breviter
bilobo, lobis inequalibus.
Long. 43-5 lin. ¢ Q.
Hab. Costa Rica, Irazu, 6000-7000 feet (Rogers).
CoLPoDES DUPLEX.
Elongato-ovatus, subeneo-niger, elytris purpureo-fuscis ; palpis,
antennis (art.5°—-11“™ dilutioribus) pedibusque rufo-piceis ; tho-
race quadrato, ante medium rotundato-dilatato, deinde paullo viz
sinuatim angustato, angulis posticis subrectis ; elytris apice
oblique sinuatis, striatis, interstitiis planis (versus apicem con-
vexis), 3° tripunctato, puncto anteriore interdum deficiente ;
elytris raro eneo-fuscis.
Tarsi, articulo 4° subtus sparsim longe setoso, quatuor anteriorum
brevissimo bilobo.
Long. 33-4 lin. ¢ Q.
Hab. Costa Rica, Irazu, 6000-7000 feet (Rogers).
CoLPODES OBSCURELLUS.
Elongatus, vix convecus, nigro-olivaceus cenescens ; palpis apice
flavo-testaceis, antennarum articulis 3 basalibus et tarsis prcers ; ;
thorace parvo, quadrato-cordato, transverso, lateribus antice
valde rotundatis, postice sinuatim modice angustato, angulis
posticis subobtusis ; elytris apice fortiter oblique sinuatis, supra
subpunctulato-striatis, interstitiis paullulum convewis, 3 (in
striis) tripunctato ; metasterni episternis elongatis ; epimeris
postice fortiter rotundatis.
Tarsi, articulo 4 subtus longe setoso, quatuor anteriortbus leviter
tantum emarginais.
a
1878. ] GEODEPHAGOUS COLEOPTERA. 597
Long. 5 lin.
Hab. Costa Rica, Irazu, 6000-7000 feet (Rogers).
COLPODES LACTIPEs.
Nigro-piceus, pedibus albo-testace is, palpis antennisquerufo-testaceis ;
labro et mandibulis piceo-rufis ; thorace cordato, antice fortiter
rotundato, postice valde angustato, ante angulos breviter recto ;
elytris amplis, convexis, apie sinuatis, tridescentibus, striatis,
interstitio 3° tripunctato ; metasterni episternis elongatis.
Long. 5 lin.
Tarsi, articulo 4to latiusculo, 2 anterioribus breviter bilobis, ceteris
emarginatis.
Hab, Chontales, Nicaragua (Belt).
CoLPODES PROCEPHALUS.,
Elongatus, vix convexus, niger ; elytris purpurascentibus 3 capite
elongato, post oculos gradatim angustato, collo supra haud trans-
versim depresso, oculis haud promnulis ; palpis et antennis
preis ; thorace valde elongato, oblongo-ovato, postice paullo
leviter angustato, angulis posticis obtusis vel rotundatis, supra
subtiliter transversim striato ; elytris ellipticis, apice fortiter
sinuatis, striatis, interstitio 3% tripunctato ; metasterni episternis
brevibus quadratis.
Tarsi articulo 4% latiuscuto, profunde emarginato.
Long. 5 lin.
Hab. Guatemala (Salvin).
Cox.popeEs PROLIXUS.
Elongatus, gracilis, pedibus antennisque valde ‘elongatis, castaneo-
piceus, palpis, pedibus et antennis rufo-piceis ; capite elongato-
ovato, post oculos crasso, rotundato, oculis vix prominulis, collo
paullo angustato, supra transversim depresso 3 thorace quam caput
vix latiore, cordato, postice sinuatim angustato, angulis postice
rectis, apice obtusis ; elytris oblongo-ellipticis, apice extus oblique
truncatis, apud suturam triangulariter excisis, angulo apicali
dentiformi, supra punctulato-striatis, interstitio 3% tripunctato ; -
metasternt episternis brevissimis.
Tarsi articulo 4t profunde emarginato, lobulis valde meequalibus,
Long. 6 lin.
Hab, Costa Rica, Irazu, 6000-7000 feet (Rogers).
CoLropEs PARVICEPs.
Ellipticus, ceeneo-olivaceus, politus, antennis tibiis et tarsis rufo-
testaceis; palpis piceis, articulo ultimo apice attenuato rufo ; capite
puro, oculis prominulis, mox pone oculos angustato ; thorace
quadrato, antice longe postice paullulum angustato, marginibus
explanatis, angulis posticis rectis ; elytris oblongis, prope basin
transversim convexis, apice leviter sinuatis, dorso striato-punc-
tulatis, striis prope basin evanescentibus, prope apicem incisis ;
metasternt episternis elongatis, angustis,
598 MR. H. W. BATES ON NEW [May 21,
Tarsi articulo quarto profunde emarginato, lobulis valde inequa-
libus.
Long. 4 lin.
Hab. Chontales, Nicaragua (Belt).
CoLPODES AUROTINCTUS.
Parvus, elongato-ovatus, supra nigro-eneus ; elytris awrato-enes
viridi tinctis, politis ; palpis, antennis, trochanteribus et tarsis
rufo-piceis ; corpore subtus femoribus tibiisque nigro-piceis ;
capite gracili, collo angustato, supra transversim depresso ;
thorace quadrato, lateribus fere requlariter paullo rotundatis,
postice puullulum magis quam antice angustato, angulis obtusis,
margine explanato rufescente ; elytris ovatis, subtiliter acute
striatis, interstitiis planissimis ; metasterni episternis viv
elongatis.
Long. 33 lin.
Tarsi articulo 4 quatuor anteriorum profunde emarginato vel
bilobo, duorum posteriorum modice emarginato, lobulis paullo
ineequalibus.
Hab. Costa Rica, Irazu, 6000-7000 feet (Rogers).
COLPODES PROSTOMIS.
Elongato-oblongus, niger, nitidus, elytris violaceo-tinctis, palpis
gracilibus antennisque rufo-piceis ; capite mox pone oculos an-
gustato, collo cylindrico, supra depresso ; mandibulis valde elon-
gatis ; thorace elongato-quadrato, swpra levissimo ; antice prope
angulos angustato, postice longius et minus, ix sinuatum, angus-
tato, angulis posticis subrectis apice obtusis, margine laterali
subceequaliter explanato-reflewo ; elytris oblongis, valde convewxis,
apice sinuatis, fortiter, prope apicem profundius, striatis, inter-
stitiis paullo conveais, 3° tripunctato ; metasterni episternis
brevibus.
Tarsi, articulo 4© bilobo, lobulis valde inequalibus,
Long. 63-7 lin. 3 Q.
Hab. Costa Rica, Irazu, 6000-7000 feet (Rogers).
CoLPODES CYANOSTOLUS.
C. cyanonoto affinis ; differt thoracis angulis posticis fere rotundatis.
Oblongus, violaceo- vel chalybeo-niger, supra ceeruleus ; thorace
lateribus regulariter rotundatis, angulis posticrs obtusis ; elytris
subtilissime punctulato-striatis, striis apice haud profundioribus.
Tarsi, articulo quarto bilobo, lobulis valde incequalibus,
Long. 5% lin.
Hab. Chontales, Nicaragua (Belt).
COLPODES CHONTALENSIS.
C. cyanonoto affinis, niger; elytris viridi-eneis, fortiter punctulato-
striatis, striis apice haud profundioribus, interstitiis paulo con-
vewxis ; thorace quadrato, lateribus regulariter rotundatis, postice
haud sinuatis ; antennis rufo-piceis.
1878. | GEODEPHAGOUS COLEOPTERA, 599
Long. 53 lin.
Hab. Chontales, Nicaragua (Belt).
CoLPoDEs LEBIOIDES.
C. chalybeo affnis, niger, capite thoraceque eneo-tinctis, elytris
ceruleis ; antennis tarsisque rufo-piceis ; capite parvo, mox pone
oculos angustato ; thorace parvo, quadrato, medio leviter dilatato,
angulis posticis subrectis ; elytris latiusculis et brevibus, puulo
convenis, apice haud sinuatis, acute striatis, interstitiis planis.
Tarsi supra trisulcati.
Long. 3-32 lin.
Hab. Chontales, Nicaragua (Belt).
CoLPoDES PRINCEPs.
C. ceruleomarginato affnis, ceruleus, elytris lete purpureo-
cupreis ; thorace antice magis quam postice angustato, medio sub-
angulatim dilatato, lateribus postice nullomodo sinuatis, angulis
posticis obtusis; elytris striato-punctatis, margine incrassato
ceruleo.
Long. 54 lin.
Hab. Chontales, Nicaragua (Belt).
CoLPopEs suPERBUS.
C. ceruleomarginato affnis, nigro-politus, elytris lete aureo-
cupreis ; palpis, antennis et tarsis rufo-piceis ; capite mox pone
oculos prominentes angustato: thorace medio Sortiter dilatato,
antice gradatim postice citius et valde sinuatim angustato ; angulis
posticis exstantibus, rectis : elytrorum striis vir conspicuis.
Long. 63 lin.
Hab. Chontales, Nicaragua (Belt).
CoLPoDEs VIRIDIAURATUS.
C. cxruleomarginato afinis, subtus chalyheus, supra lete viridi-
eneis, elytris interdum viridi-auratis vel cupreo-auratis ; capite
mox pone oculos prominentes angustato ; thorace valde transverso,
lateribus regulariter rotundatis, postice haud sinuatis, angulis
posticis haud exstantibus, vir rectis ; elytris conspicue striato-
punctatis.
Long. 7 lin.
Hab. Chontales, Nicaragua (Belé).
CoLPoDES CHRYSOPTERUS.
Elongatus, niger nitidus, elytris aurato-eneis ; thoracis margine,
palpis, antennis et tarsis rufescenti-piceis; capite mox pone oculos
magnos subito angustato ; thorace Sere ut in C, ceruleomarginato,
medio rotundato-dilatato, lateribus postice sinuato-angustatis, an-
gulis posticis subrectis 3 elytris elongatis, striato-punctatis, ver-
sus apicem striis acute incisis et minus punctulatis.
Tarsi lati, subtus dense pubescentes, articulo 4 bilobo.
Long. 52 lin.
Hab. Chontales, Nicaragua (Belt).
600 MR. H. W. BATES ON NEW [May 21,
COLPODES PROLONGATUS.
Valde elongatus, elegans, viridi-eneus, elytris lete aureo-cupreis ;
capite pone oculos elongato, gradatim angustato ; thorace ovalt,
angulis posticis rotundatis ; elytris apice sinuatis, longe productis
sed haud spinosis, striatis, striis passim equaliter fortiter incisis,
interstitiis planis.
Tarsi articulo quarto 2 anteriorum bilobo, 4 posteriorum emarginato,
lobulis haud prolongatis.
Articulo unguiculari biseriatim setoso.
Long. 64 lin.
Hab. Chontales, Nicaragua (Belt).
LEPTOTRACHELUS PUNCTICOLLIS,
Fulvo-testaceus, immaculatus, pedibus pallidioribus; capite levi,
post oculos paullo rotundatim angustato, convexo ; thorace antice
prope apicem distincte angustato, ante medium rotundato-dilatato ;
supra passim punctulato, versus angulos posticos grossius punc-
tato ; elytris depressis, punctato-striatis.
Long. 43-43 lin.
Hab. Chontales, Nicaragua (Janson).
LEPTOTRACHELUS PANAMENSIS,
Fulvo-testaceus fuscescens, sutura angusta plagaque apicali nigro-
fuscis ; untennis et pedibus pallidioribus : capite levi; thorace
antice prope apicem angustato, lateribus anticis leviter rotundatis,
ante basin paullo sinuatis, supra punctato et medio dorsi punc-
tato-rugoso.
Long. 43 lin.
Hab. Panama.
CELIA COSTARICENSIS.
Oblongo-ovata, nigra, polita, plus minusve e@nescens vel fusco-
@nea; antennis piceo-nigris, articulis 1° et 2° vel 1°-3“™ rufis ;
thorace convexo, mox pone angulos anticos rotundato-dilatato,
deinde usque ad basin leviter arcuato vel subrecto, basi utrinque
foveolis duabus distantibus levibus, interiore recta, exteriore versus
angulum oblique ducta; elytris prope apicem angustatis et com-
pressis, striatis, striis lateralibus evanescentibus, striola scutel-
lari ad basin secunde oriente suturam haud attingente, interstitiis
paullo convexis; pedibus piceis, tarsis pallidioribus ; prosterni
apice dilatato subtruncato.
Long. 4 lin.
Variat. 1, elytrorum interstitiis planissimis ; 2, striis. subtilissimis
subpunctulatis ; 3, forma abbreviata, ovata.
Hab. Ivazu, Costa Rica, alt. 6000-7000 (Rogers).
CURTONOTUS PUTZEYSI.
Elongato-oblongus, niger nitidus, subtus cum pedibus picescens, an-
tennis et palpis piceo-rufis; thorace lateribus valde rotundatis,
juata basin fortiter subito constricto, angults posticis rectis, limbo
1878. ] GEODEPHAGOUS COLEOPTERA. 601
antice et postice grosse punctato ; elytris thorace vix latioribus,
parallelis, striis conspicue punctatis, omnibus acute impressis.
3 tibiis intermediis medio intus haud dentatis.
Long. 74 lin. ¢ Q.
Hab. Mexico, near the capital (Flohr).
Differs from C. substriatus by its larger size and more robust pro-
portions, and by the stronger and more uniform striation of the
elytra. The ¢ is destitute of the broad tooth on the inner side of
the middle tibize. Immature individuals are reddish piceous with
paler margins.
DIPLOHARPUS EXSTRIATUS.
Oblongo-ovatus, supra levissimus eneo-relucens, subiridescens ; par-
tibus oris, antennis pedibusque fulvis ; mandibulis longissimis,
rectis ; thorace ovato, marginibus postice explanato-elevatis, angu-
lis posticis rotundatis ; elytris absque striis, marginali excepta.
Long. 32 lin.
Hab. Chontales, Nicaragua (Belt).
PERICOMPSUS LONGULUS.
Elongatus, parallelogrammicus, castaneo-piceus, capite margini-
busque thoracis rufis; elytris fulvo-testaceis plaga mediana oblongu
communi ( postice usque ad marginem dilatata) nigro-picea; par-
tibus oris, antennis pedibusque pallidis; thorace transverso,
postice fortiter sinuatim angustato, angulis posticis rectlis, mur-
ginibus paullo explanato-reflexis.
Long. 1 lin.
Hab. Mexico, near the capital (Flohr).
XyYSTOSOMUS BELTI.
Elongato-ovatus, modice converus, politus, viridi-eneus, elytris
aureo tinctis ; antennis fulvis, pedibus rafo-piceis, corpore subtus
piceo ; foveis frontalibus longis ab suiculo oculari utrinque carina
separatis ; oculis maximis ; thorace transverso, antice prope an-
gulos fortiter rotundato-angustato, postice haud angustato, angulis
posticis rectis ; elytris striis punctulatis novem, 1™4¢-34%™ vix im-
pressis (1 versus apicem excepta), 4*-;" apice evanescentibus.
Hab. Chontales, Nicaragua (Belt).
XYSTOSOMUS OLIVACEUS.
X. belti affinissimus ; differt colore supra omnino olivaceo-eneo,
antennis obscure cinereis, articulis 4 basalibus piceis ; thorace
transverso, antice subito rotundato-angustato, postice haud angus-
tato, sed leviter sinuato, angulis posticis acutis; elytris paullo
convezis, ovatis, mox pone humeros rotundato-dilatatis, margineque
explanato et fortiter reflero, punctulato-striatis, striis 17-30"
viv impressis ; pedibus et palpis obscure piceis, articulo subulato
rufo.
Long. 23 lin.
Hab, Chontales, Nicaragua ( Belt).
Proc. Zoou. Soc.—1878, No. XXXIX. 39
602 MR. H. W. BATES ON NEW [May 21,
TACHYS DIPLOHARPINUS.
Oblongus, modice convexus, sericeo-nitens, subopalescens ; partibus
oris, antennis pedibusque fulvis ; mandibulis acute hamatis ; oculis
prominulis ; foveis frontalibus fortiter impressis, post oculos cur-
vatis; thorace lato, postice modice sinuato-angustato, angulis
posticis exstantibus acutis, margine lateral elevato-explanato ;
elytris levibus, stria suturali solum fortiter impresso, striola api-
cali recurva fortiter impressa, apice setifero, foveis irregularibus
duabus submarginalibus, puncto discoidali mediano.
Long. 13 lin.
Hab. Chontales, Nicaragua (Janson).
A species remarkable for its many points of resemblance to the
genus Diploharpus.
Bemsipium (NorTapnvs) FLOURI.
Oblongum, parallelogrammicum, depressum, eneum nitidum, elytris
pallido-testaceis, regione scutellari, gutta utrinque discoidali,
maculaque majore posimediana transversa fuscis ; palpis, antennis
et pedibus fulvo-testaceis ; sulcis frontalibus modice impressis sim-
plicibus ; capite levi; thorace postice angustato, angulis posticis
obtusis fere rotundatis, supra medio basi strigoso, fovea utrinque
vaga, lata, rugulosa, carinaque acuta oblique ad angulum posticum
ducta ; elytris thorace vie latioribus, integriter punctulato-striatis,
interstitiis planis, tertio tripunctato.
Long. 2-23 lin.
Hab. Mexieo, near the capital (Flohr).
The species belongs to an unnamed group near Notaphus, and is
only referred to the latter for general guidance as to its position.
Bempipium (NOTAPHUS) PLACITUM.
Elongato-oblongum, depressum, viridi- vel subcupreo-aeneum, elytris
cupreo-fuscis ( 3 nitidis, 2 subopacis) vitta laterali ab humero
usque medium, intus biloba, fasciaque maculari ante apicem, fulvo-
testaceis ; sulcis frontalibus latis haud profundis ; thorace late
rotundato, postice magis quam antice angustato, angulis posticis
obtusis, fovea media profunda carinaque parva jurta angulum ;
elytris subtiliter punctato-striatis, interstitiis planissimis, tertio
3-punctato; palpis, antennis pedibusque obscure piceis.
Long. 3 lin.
Hab. Mexico, near the capital (Flohr).
This species has been mistaken for N. semifasciatus of Say. His
description, however, suits much better an equally common and
similar, but smaller, Mexican species.
BremsBipium (PERYPHUS) ROGERSI.
Gracile, convexiusculum, nigro-eneum, pedibus piceis, antennarum
articulo basali subtus rufo; thorace anguste cordato, antice ro-
tundato, postice sinuatim angustato, angulis posticis rectis, basi
transversim depresso, vage punctato, foveis profundis carinaque
elevata ad marginem fere parallela; elytris punctulato-striatis,
1878. | GEODEPHAGOUS COLEOPTERA. 603
interstitiis convexis paulo undulatis, tertio foveis duabus magnis
vagis impressis, stria 7" obsoleta.
Long. 23 lin.
Hab. Costa Rica, Mount Irazu, alt. 6000-7000 feet (Rogers).
Bemsipium (Perypuus) suBMACULATUM,
Elongato-ovatum, modice converum, fusco-eneum, elytris utringue
versus apicem macula obliqua flavo-testacea ; antennis piceis, ar-
ticulo 1° et 240_4tum basi pedibusque fiavo-testaceis ; palpis mavil-
laribus articulo penultimo obscuro ; thorace transversim quadrato,
antice modice rotundato, postice leviter sinuatim angustato, angulis
posticis subrectis, fovea basali profunda vage rugosa, carinaque
oblique juxta angulum ; elytris subtiliter striato-punctulatis, striis
interdum magis impressis, 74 abbreviata, exterioribus omnibus
versus apicem obliteratis, interstitio 3% bipunctato.
Long. 2 lin.
Hab. Mexico, near the capital (Flohr).
Specimens received from Paris as “‘submaculatum, Chaud. MSS.,”’
have the strize impressed; in Mr. Flohr’s specimens the rows of
fine clear punctures are superficial. The nearly allied B. meai-
canum, Dej., is much larger, and has-a short curved fold instead of
a distinct carina at the hind angles of the thorax.
LacHNoPHorRus SEMIRUFUS.
Breviter oblongus, erecte pilosus, capite thoraceque rufis, elytris
pectore abdomineque nigro-eneis, palpis antennis pedibusque flavo-
testaceis ; capite postice modice angustato, oculis minus prominu-
lis, vertice grossissime punctatis ; thorace late cordato, versus
basin modice angustato, grossissime intricato-punctato ; elytris
quadratis, fortiter punctato-striatis, interstitiis supra paulo
planatis, crenulatis, tertio 3-foveato,
Long. 23 lin.
#Hab. Chontales, Nicaragua (Beit).
LACHNOPHORUS LEUCOSCELIS.
L. levicolli (Reich.) proxime affinis. Elongatus, nigro-eneus, .
nitidus, supra glaber ; palpis antennarumque articulis 4 basalibus
rufo-testaceis ; tibiis (apice excepto) albo-testaceis 3 capite (cum
oculis) quam thorax latiore, punctis magnis conspersis; thorace
rotundato-cordato, valde convexo, postice Sortiter angustato, basi
constricto et depresso, fere impunctato, polito ; elytris profunde
striatis, striis a basi usque paulo ultra medium grosse punctatis.
Long. vix 2 lin.
Hab. Chontales, Nicaragua (Belt),
Lacu NOPHORUS LONGULUS.
dn hoc genere elongatus, convezior, olivaceo-eneus, breviter setosus ;
elytris sordide fusco-albis, pone medium Suscia irregulari fusca
prope suturam valde dilatata ; antennis piceo-fuscis, articulo basali
infra rufo ; femoribus flavo-testaceis, basi et apice piceo-nigris f
39”
604 MR. H. W. BATES ON NEW [May 21,
tibiis et tarsis piceis, illis dimidio basali rufo-piceis ; capite longitu-
dinaliter vage striato ; thorace angusto, elongato-ovato, lateribus
anticis modice rotundatis, basi angustata, angulis posticis haud
exstantibus, supra viridi-eneo subtiliter alutaceo et punctulato ;
elytris oblongis, apice distincte sinuato-truncatis, fortiter exarato-
striatis, strits dimidio basali punctatis, interstitiis planis, seriatim
punctulatis, disco utrinque 3-foveatis.
Long. 2}, lin.
Hab. Chontales, Nicaragua (Belt, Janson).
LACHNOPHORUS SCULPTIFRONS.
L. elegantulo similis. Gracilis, breviter pallido-hirsutus, viridi-
@neus ; antennarum articulo basali infra, 2°-4“™" basi, femoribus
(annulo basali et genibus exceptis) tibiisque (apice excepto) rufo-
testaceis ; elytris albo- et flavo-testaceis, fascia ultra medium
fusco-enea; capite crebre longitudinaliter strigoso et punctato,
thorace elongato-cordato, paullulum dilatato, alutaceo-opaco et
sparsim punctulato; elytris acute striatis, striis basi punctatis,
interstitiis planatis uniseriatim punctulatis, disco trifoveato, foveola
anteriore solum conspicua.
Long. 2 lin.
Hab. Guatemala, Chinautla, alt. 4100 feet (Sa/vin); Chontales
(Belt).
CHALYBE BELTI.
Gracilis, subeneo-nigra, breviter griseo-setosa, setis longioribus nigris
commixtis ; antennis ut in gen. Ega brevibus extus incrassatis,
brevius hirsutis, articulis tribus basalibus flavis, 4°-7"™" nigro-
fuscis, 8-11" albis ; capite grosse reticulato-punctato ; thorace
gracile cordato, grosse discrete punctato ; elytris pone basin valde
transversim depressis, acute exarato-striatis, interstitiis summis
planatis, distanter lineato-punctatis, utrinque fasciis angustis
brevibus macularibus testaceo-albis duabus ; pedibus albis, femo-
ribus, tibiis tarsisque apice fuscis.
Long. 13 lin.
Hab. Chontales, Nicaragua (Bel¢).
CASNONIA TUBULIFERA.
Subeneo-niger, elytris macula anteriore antice ramum ad basin
emittente, altera apwali marginibusque rufo-testaceis ; antennis
pedibusque rufescentibus ; capite levi, mow pone oculos subito con-
stricto, deinde in collum tubuliforme prolongato ; media fronte
bicallosa ; thorace minus elongato, grosse punctato ; elytris oblon-
gis, politis, nudis, apice leviter sinuatim truncatis, angulo exte-
riore obtuso, supra striato~punctatis, striis interioribus quatuor
post medium interruptis, interstitio tertio trifoveato, calloque
posteriore submarginali.
Long. 3 lin.
Hab. Chontales, Nicaragua (Belt).
1878. ] GEODEPHAGOUS COLEOPTERA. 605
CALOPHZENA CRUCIATA,
C. acuminatee (Oliv.) affinis. Migro-cyanea, nitida; elytris
utrinque maculis magnis quadratis duabus (interstitia sex occu-
pantibus) albo-testaceis, basi et apice anguste cruceque mediana
nigro-cyaneis ; thorace valde elongato, antice magis quam postice
recte angustuto ; elytris punctulato-striatis, interstitus modice
convexis ; antennis articulis 5°--11”™ fulvis.
Long. 6% lin.
Hab. Chontales, Nicaragua (Belt).
CALOPHENA LEVIGATA.
C. acuminata (Ol.) affinis. Oolore letius cyanea; elytris medio
violaceis, striis (apud maculas albas exceptis) nullo modo wm-
pressis; thorace valde elongato, antice subrotundatim, postice
sinuatim angustato ; elytris aculeatis, maculis utrinque duabus
transversis, albo-testaceis, prima interstitia 5, secunda 6 occupante.
Long. 63 lin,
Hab. Panama.
CaTASCOPUS CHONTALENSIS.
Supra lete viridi-ceneus, elytrorum limbo aurato, pectore et pedi-
bus nigro-piceis, abdomine castanco ; capite post oculos gradatim
modice angustato, sed haud elongato, oculis valde eastantibus ;
thorace transverso vel late cordato, angulis anticis modice provectis,
margine exteriore late explanato et valde rotundato usque «ad
summun angulum, postice sinuatim angustato, angulis posticts
eastantibus acutis ; elytris punctato-striatis, interstitirs viv convexis.
“Long. 6 lin.
Hab. Chontales, Nicaragua (Beit, Janson), 4 exempl.
The elytra viewed from the front against the light have a delicate
milky opalescent bloom, as in C. validus, to which the species is
very closely allied.
CATASCOPUS ANGULICOLLIS.
Supra viridi-eneus, corpore subtus pedibusque nigro- vel rufo-pweis,
abdomine castaneo ; capite mox pone oculos subito anqgustato ;
thorace subquadrato, angulis anticis angustis prominulis, margine
explanato, laterali angusto, ante medium angulato, postice sinua-
tim angustato, angulis posticis prominulis acutis, supra distinc-
tius transversim strigato, margine explanato grosse punctato ;
elytris punctato-striatis, interstitiis paullulum convexis.
Long. 6 lin.
Hab. Chontales (Belt).
CopTroDERA SCINTILLANS.
C. acutipenni (Bug.) affinis. Elongato-oblonga, supra capite tho-
raceque aurato-viridibus politissimis, elytris aurato-cupreis ;
partibus oris, epistomate, antennis, corpore subtus pedibusque
fulvo-testuceis, tibiis obscuris ; thorace valde transverso, latertbus
rotundatis nullo modo angulatis, margine late elevate-explanato,
606 MR. H. W. BATES ON NEW [May 21,
impunctato ; elytris amplis, truncaturee angulis acutis sed non
ut in C, acutipenni productis, subtiliter punctato-striatis, stris
via impressis, interstitiis planis.
Long. 5 lin.
Hab. Chontales, Nicaragua (Belt).
EuRYCOLEUS BELTI.
E. 13-punctato (Chaud.) proxvime affinis, differt maculis pauci-
oribus magnitudineque cequalibus. Latus, ovatus, testaceo-rufus ;
elytris flavo-testaceis, maculis utrinque 7 margineque subapicali
nigris; antennis rufescentibus, articulis 1° (apice) et 29-4", palpis
(apice excepto), femoribus upice, tibtis et tarsis nigris ; elytro-
rum maculis 3 suturalibus (apicali communi antice dilatata),
1 subhumerali, 2 medianis, 1 rotunda subapicali, a margine
laterali longe separata.
Long. 5 lin,
Hab. Chontales, Nicaragua (Belt).
APENES COMISs.
Ad sectionem A. purpuripennis pertinet. A. sinuata paulo mayor,
recipue latior, supra plana, subopaca, fusco-wnea, antennis
palpis pedibusque fulvis ; capite fere levi, subtiliter alutaceo, vix
ruguloso ; thorace late cordato, angulis posticis dentiformibus,
basi media arcuata, sublobata, versus angulos subsinuata, lateribus
marginulo explanato, supra passim subtiliter transversim stri-
gato ; elytris oblongis, apice subfortius sinuato-truncatis, punctu-
lato-striatis, interstitvis planis alutaceis, fascia angusta, cur-
vata, maculari, subapical, rufa, interstitia 4-8 occwpante, apud
zmum Srumone maculis versus apicem elongatis, macula altera
subhumerali minus distincta. .
Long. 4 lin.
Hab. Panama.
PINACODERA AMBLYGONA.
P. nigritee (Chaud.) prowime affinis, differt thorace angustiore et
longiore angulisque posticis omnino rotundatis. Nigra, nitida,
antennis tarsisque rufo-piceis ; capite impunctato ; thorace trans-
versim striolato ; elytris subtiliter punctulato-striatis, interstitiis
planissims, sparsim obsolete et minutissime punctulatis,
Long. 5j lin. 9.
Hab. Mexico (Boucard).
PINACODERA ANGULIFERA.
P. nigritee (Chaud.) simillima, differt thorace postice latiore, angulis
acutis exstantibus. Nigra, nitida, antennis tarsisque rufo-piceis ;
capite ruguloso, via punctulato ; thorace antice et postice sub-
crebre punctato ; elytris acute striatis, striis subtiliter punctula-
tis, interstitiis vin convewis, grossius uniseriatim punctulatis.
Long. 5} lin. 2. .
Hab. Mexico.
EO a rt
i". a oe
Lenn et ee ee
1878. ] GEODEPHAGOUS COLEOPTERA. 607
LEBIA CALLIZONA.
L. bifasciate (Dej.) prowime affinis, differt fascia postica metal-
lica apud suturam antice fere divisa. Levis, testaceo-rufa,
capite fusciisque duabus elytrorum latis et macula apicalr
ceruleis, fascia prima postice medio emarginata, fascia secunda
antice ad suturam fere divisa ; antenmis nigris articulo basali
rufo; pedibus ceruleis, femoribus basi rufis; abdomine apice
subtus rufo concolori, supra nigro.
Long. 3 lin.
Hab. Guatemala.
LEBIA CORCULA.
L. nigricipiti (Chaud.) similis, at capite flavo, elytris ovatis differt.
Parva, ovatula, testaceo-flava ; elytris fasciis duabus nigris, prima
basali (latera haud attingente) postice medio angulatim emargi-
nata, secunda pone medium (marginem lateralem haud attin-
gente) antice et postice medio elytro leviter sinuata, iterstitio
suturalt inter fascias autem nigro; thorace valde transverso,
apud angulos posticos valde explanato ; elytris profunde punc-
tulato-striatis, unterstitiis culminatis.
Long. vix 2 lin.
Hab. Chontales, Nicaragua (Janson).
LEBIA CHARINA.
L. quinquenotatee (Chaud.) simillima, differt elytrorum maculis
mimoribus posticisque ad suturam conjunctis. Late ovata, flavo-
testacea ; palpis, articulis 4-11" antennarum, capite, femoribus
apree, tibiis et tarsis, abdominis seqmento anali, et elytrorum
utrinque maculis 3 nigris ; thorace quam in L. quinquenotata
angustiore ; elytris obsolete striato-punctatis ; elytrorum macu-
lis, 1 ovata communi seutellari, 1 virguliformi humerali, et 1
postmediana transversa, antice apud suturam indentata, latera
haud attingente, apice autem anguste nigro.
Long. 3 lin.
Hab. Chontales, Nicaragua (Belt).
LIA QUADRIANNULATA.
In hoe genere convexa, polita, subcastaneo-rufa, subtus et pedibus
testaceo-rufis ; antennis articulis 4°-11”™ piceis; thorace angus-
tiore, antice gradatim angustato ; elytris utrinque annulis duobus
flavis, primo mediano-laterali, secundo versus apicem, macula
autem flava subhumerali.
Long. 5 lin.
Hab. Chontales, Nicaragua (Belt).
OTOGLOSSA C@LESTINA.
O. terminali (Chaud.) forma similis. Supra lete cyanea ; elytris
violaceo relucentibus, fascia apicali flava ; antennis, partibus oris,
pedibus et pectore chalybeis, tarsis abdomineque flavis ; thorace
angusto, pone medium fortiter sinuato-angustato, bast tterum
608 MR. H. W. BATES ON NEW [ May 21,
subito dilatato, angulis apice rotundatis ; elytris supra modice
_ inequalibus, obsolete punctulato-striatis.
Long. 33 lin.
Hab. Chontales, Nicaragua (Belt).
OTOGLOSSA OBSCURELLA.
Minor, nigra ; elytris leviter violaceo-tinctis, wpice fascia obscure
rufa ad suturam angustata ; tarsis rufo-piceis ; abdomine flavo ;
capite post oculos paulo citius rectiusque angustato, occipite minus
convexo ; thorace angusto, pone medium fortiter sinwato-angus-
tato, basi iterum dilatato ; elytris supra modice incequalibus, ob-
solete striatis.
Long. 24 lin.
Hab. Chontales, Nicaragua (Belt).
CALLEIDA SEMIRUBRA.
Llongato-oblonga, nigro-nitida, elytris (basi excepta) rufo-casta-
neis; abdomine rufo; capite post oculos magis quam in C.
metallica incrassato, collo subito angustato; thorace quadrato,
antice cum angulis rotundato, postice sinuato modice angustato,
angulis posticis obtusis; elytris apice fortiter sinuatim trun-
catis, angulis externis denttiformibus, supra paulo inequalibus,
punctulato-striatis, interstitis planis.
Long. 7 lin.
Hab. Chontales, Nicaragua (Belt).
Taken from the stomach of a toad on the summit of a hill near
St. Domingo.
CALLEIDA L/TIPENNIS.
Subgracilis, castaneo-rufa, elytris (epipleuris exceptis) aurato-
@neis ; capite angusto, post oculos gradatim angustato haud
tumido, collo distincto; thorace quadrato-cordato, postice
leviter sinuatim aagustato, angulis posticis subrectis ; elytris
apice leviter sinuatim truncatis, angulis externis valde rotun-
datis, punctulato-striatis, interstitiis planis.
Long. 4$ lin.
Hab. Chontales, Nicaragua (Belt, Janson).
CALLEIDA JANSONI.
C. onypterygoidi (Chaud.) coloribus similis, at minus nitida, for-
maque multomagis parallela. Cyanea; elytris purpureo-cupreis,
subtiliter alutaceis ; capite post oculos citius recte angustato,
haud tumido ; thorace elongato-quadrato, postice vix sinuatim
modice angustato, angulis subrectis, supra subfortiter transver-
sim strigoso; elytris elongatis, parallelis, truncature angulis
externis valde rotundatis, fortiter vix punctulatim striatis,
interstitits paulo convexis, tertio conspicue tripunctato.
Long. 6 lin.
Hab, Chontales, Nicaragua (Belt, Janson).
1878. | GEODEPHAGOUS COLEOPTERA. 609
AGRA LETICOLOR.
Ad sectionem A. ftemorate (K/.) pertinet. Gracilis, viridi-enea,
antennis pedibusque rufis, femoribus basi obscurioribus ; capite
maxime elongato, levi, post oculos gradatim rotundato-angus-
tato, juxta collum haud tumidulo; thorace angusto, elongato-
conico, sulcis 4 longitudinalibus, foveolato-punctatis, impressis,
quorum duobus dorsalibus antice carina abbreviata separatis ;
elytris ante medium sinuatim angustatis, postice iterum. dilatatis,
apice utringue flexuoso-truncatis, angulo exteriore subspinoso,
suturali acute dentato, supra punctato-striatis, interstitiis
inter se lineolis transversis multifariam conjunctis.
3 metasterno medio, femoribusque basi infra rufo-pilosis.
Long. 7 lin.
Hab. Chontales, Nicaragua (Belt).
AGRA CHRYSOPTERYX.
A. rutilipenni proxime affinis et similis. Castaneo-rufa, elytris
(epipleuris exceptis) lete viridi-auratis; capite post oculos
breviter semiovato, punctis nonnullis piliferis impresso ; thorace
medio modice rotundato-dilatato, paulo ante apicem fere subito
angustato, lineatim subsparsim irregulariter foveato-punctato,
longe piloso; elytris apice subrecte oblique truncatis, angulo
exteriore breviter dentato, suturali rotundato, supra acute
punctulato-striatis, stris secunda et quarta 5-6-punctatis, stria
sexta bipunctato.
3 metasterno et abdomine medio longe rufo-barbatis.
Long. 7 lin.
Hab. Chontales, Nicaragua (Janson).
AGRA PANAMENSIS.
Ad sectionem A. ene (F.) pertinet ; at ceteris speciebus dis-
tinctissima. Latior, nigra, partibus oris, antennis, trochan-
teribus, genibus, tibiis et tarsis fulvo-castaneis ; capite lato,
ome pone oculos quadrato, margine posteriore ante collum exacte
recto, supra impunctato; thorace quam caput longiore, pone’
medium rotundato-dilatato, antice usque ad apicem gradatim
angustato, supra medio fere levi, juxta marginem sparse punc-
tato-impresso; elytris apiee urcuatim truncatis, angulo exteriore
Sortiter spinoso, suturali acute dentato, supra vix inequalibus,
punctato-striatis, struis 2% et 4% seriatim rotundato-foveolatis,
6% et 8% magis confuse foveolatis.
© antennis art. 8% multo abbreviato.
Long. 12 lin.
Hab. Panama (received from Mr. E. Bartlett).
610 MR. G, F. ANGAS ON A NEW TUDICULA. [| May 21,
5. Description of a new Species of Tudicula. By Grorce
Frencu Aneas, C.M.Z.S., F.L.S., &e.
[Received May 11, 1878.]
TuDICULA INERMIS, N. sp.
Shell globosely turbinate, solid, white, ornamented with a broad
band and descending flames of an orange-chestnut colour, sculptured
throughout with numerous elevated concentric ridges alternating with
smaller ones, the interstices crossed by very fine close-set descending
strie ; whorls 64, flattened, last whorl large, tumid, flattened above
and angulated at the periphery, suture impressed, apex papillary ;
aperture acuminately ovate; outer lip simple, arcuate, strongly
grooved within ; inner lip with a broad white callus spreading over
the pillar, and with three transverse plaits at the lower portion of the
columella; canal long, straight, somewhat longer than the entire
body of the shell.
Long. | in. 8 lin., lat. 10 lin.
Hab. Singapore ?
Tudicula inernis.
This remarkable shell belongs to the genus Tudicula of H. and A.
Adams, constituting the third species of the genus as yet described.
The genus Tudicula was proposed by the learned authors of the
‘Genera of Recent Mollusca’ (in the Proc. Zool. Soc. 1863) for
the reception of two species of shells ‘ with spiny varices and three
transverse plaits on the columella,”’ viz. 7’. armigera, A. Ad., and T.
spinosa, H. & A. Ad. The peculiar feature of the species now de-
scribed is that it is destitute of spines, although it agrees perfectly
with all the other characters of this well-marked genus, which, in
my opinion, should be placed in the family Turbinellidz.
I am indebted to my friend Dr. W. Newcomb, of Ithaca, New
York, for an examination of this very interesting shell, two specimens
: Ptah at :
Stow
ad _~ a
es
ey
1878. | ON THE ORNITHOLOGY OF THE PHILIPPINES. 611
of which were obtained from a dealer at Singapore ; but the exact
locality of their habitat could not be satisfactorily determined.
Care must be taken not to confuse Tudicula of H. and A. Adams
with Tudicla of Bolton, of which genus 7. (Murex) spirillus of
Linneus is the type.
6. Contributions to the Ornithology of the Philippines.—
No. IX. On the Collection made by Mr. A. H. Everett
in the Island of Palawan?. By Arruur, Marquis of
TweEEDDALE, F,.R.S., President of the Society.
[Received May 16, 1878.]
(Plates XXXVII. & XXXVIII.)
Until visited by Dr. Steere in July 1874, when he remained for a
month at the Spanish settlement of Puerto Princesa, the island of
Palawan, so far as is on record, had not been explored by any
naturalist. Nothing was known of its zoological productions ; and its
exact relationship as a zoogeographical area remained an unsolved
problem. During his stay at Puerto Princesa Dr. Steere collected
examples of 32 species of birds, all of which have been enumerated
by Mr. Sharpe’.
At the end of November 1877 Mr. A. H. Everett arrived at Puerto
Princesa, and remained until the beginning of January, when,
becoming disabled by fever, he was obliged to return to Manilla to
recruit his health. He, however, succeeded in collecting specimens of
52 species of birds, 32 of which are additional to those obtained by Dr.
Steere. This last gentleman discovered 12 Species not sent to me by
Mr. Everett ; so that the total of known Palawan species of birds
(and all are from Puerto Princesa and its vicinity ) now amounts to 64.
This small number cannot exhaust the avifauna of this large island ;
and as Mr. Everett is now happily recovered, and has returned to
Palawan, I propose postponing any remarks on the character of the
Palawan ornis until we are in possession of the wider evidence which.
further collections will doubtless supply. Generally, however, it
may be affirmed from the data before us that the birds of Palawan
represent mostly Bornean genera and species, although a few distinctly
Philippine types also occur.
The following species in the present collection have not hitherto
been described :—
4. Tiga everetti. 22. Brachypus cinereifrons.
12. Dicrurus palawanensis. 25. Oriniger palawanensis.
17. Broderipus palawanensis. 34. Cyrtostomus aurora.
19. Trichostoma rufifrons. 39. Corvus pusillus.
20. Drymocataphus cinereiceps.
* I treat Palawan as being a Philippine Island in a political sense. It remains
to be shown whether it does not in fact belong, zoologically, rather to Borneo.
? Trans, Linn. Soe. ser. 2, Zoology, vol. i.
612 THE MARQUIS OF TWEEDDALE ON [May 2],
1. TANYGNATHUS LUZONENSIS (3).
[P. Princesa, ¢, January 1878. ]
Not separable from all other Philippine examples.
2. BurasturR inpIcus (20).
[P. Princesa, ¢(?), November 28, 1877 : iris bright vellow; cere
light wax-yellow ; bill and nails black ; legs and feet dull yellow. ]
3. CHRYSOCOLAPTES ERYTHROCEPHALUS.
Chrysocolaptes “erythrocephalus, Sharpe, Tr. L. S. ser. 2, Zool.
1. p. 315. no. 13, t. xlvi. f. 1.
[P. Princesa, ¢, December 2, 1877: iris crimson; bill pale
yellow tinged with green; feet and claws grey-green. 2, No-
vember 27: iris crimson; bill pale yellow tinged with green; feet
and claws greyish green. |
The male example is in full adult plumage. That of the female
(hitherto undescribed) resembles the male in all respects excepting
the crown of the head and the crest, the feathers of which are ruddy
brown, each being tipped with a pale rusty-yellow drop or spot.
The cheeks, chin, and throat are washed with dilute crimson, not
intense as in the male.
4, Tica EveRETTI. (Plate XXXVII.)
Tiga javanensis (Ljungh,), Sharpe, ¢.c. p. 315. no. 14?
Three examples, two of the adult male and one of the adult
female, were obtained near Puerto Princesa by Mr. Everett. They
probably belong to the species doubtfully identified by Mr. Sharpe
(J. c.) with TZ. javanensis. From this species, as well as from all the
other species of the genus known to me, the present bird differs in
the female having the head and crest uniform dark brown, the latter
being broadly terminated with crimson. The lower part of the throat
and the upper breast are uniform buffy brown, and not squamate.
Male. Forehead, crown and crest, uropygium, and upper tail-
coverts crimson. Many of the dorsal plumes washed with crimson.
Lores buffy brown. Space behind the eyes creamy white, each
feather narrowly margined with pale brown. A patch commencing
below the posterior angle of the eye and including the ear-coverts
dark brown, this colour being continued as a narrow band round
the occiput. Space below the eye buffy cream-colour, which is
continued as a narrow line on the cheek, and expands into a broad
stripe down the sides of the neck. A brown stripe commencing
at angle of mouth and descending down the sides of the neck; a
few crimson feathers on the inner edge of this stripe. Chin and
upper throat buffy cream-colour, many of the feathers tipped dark
brown. Lower throat and crop uniform buffy brown. — Breast-
feathers pale yellow, very narrowly margined and boldly centred
with dark brown; within the brown centres a pale yellow stripe
of varying dimensions. Abdominal and ventral feathers and the
under tail-coverts pale yellow, with generally two cross bands of
1878. | THE ORNITHOLOGY OF THE PHILIPPINES. 613
dark brown. Back, wing-coverts, outer edges of secondaries, and
exposed surface of tertiaries and scapulars golden. Primaries and
rectrices dark brown.
Female. Differs from the male in having the top of the head
dark uniform brown, almost black, the crest-plumes being termi-
nated with crimson.
Dimensions :—
Wing. Tail. Tarsus. Culmen.
in. in. in. in.
Gees bbe 4:12 1:00 1°37
2 ar Be 4:12 1:00 1°37
In one example (¢) the lower breast-markings are not so well
defined, and the general markings of the lower surface are less
pronounced. ‘The upper tail-coverts are mostly black, some only
being tipped with crimson, others with golden.
[P. Princesa, ¢, November 25, 1877: iris dark brown; bill
black, basal half of mandible grey ; feet greenish, nails grey. P.
Princesa, 2, November 27, 1877: iris crimson; other parts as
in ¢.]
5. EURYSTOMUS ORIENTALIS (37).
|P. Princesa, 2, November 29, 1877.]
6. CACOMANTIS MERULINUs (57)?
[P. Princesa, ¢, December 10, 1877; iris bright salmon-red ;
bill black, mandible light brown ; feet yellow, claws black. ]
A single example, not sufficiently mature to be identified with
certainty.
7. SURNICULUS LUGUBRIS.
Cuculus lugubris, Horsf. Tr. L. 8. xiii. p. 175.
[P. Princesa, 2, December 6, 1877: iris dark brown; bill
black ; feet dark grey. |
8. PHaNnICOPHAES HARRINGTONI.
Dryococcyx harringtoni, Sharpe, t.c. p. 321. no. 34, f. 1.
[P. Princesa, ¢, November 29, 1877: iris light brown ; orbital
and loral space deep crimson; bill light green, tinged with olive ;
feet and legs lead-grey ; nails dark grey. 9, December: iris bril-
liant yellow, with ring of ochreous orange. |
Of other males Mr. Everett notes the iris as being reddish
brown.
This Coucal is a representative form of P. curvirostris and P.
erythrognathus, closely resembling both species, but differing structu-
rally in the shape and position of the nostril. Mr. Sharpe (P. Z.S.
1873, p. 604) has generically separated P. curvirostris from P. ery-
thrognathus, on account of the shape and position of the nostrils in
those two species being different, and for the same reason has esta-
blished (/. c.) the genus Dryococcyx for the reception of this Palawan
614 THE MARQUIS OF TWEEDDALE ON [May 21,
representative form’. That three birds, one inhabiting Java, another
Borneo, the Malay peninsula, and Sumatra, and the third Palawan,
all so closely resembling each other in their colouring and markings
that they are difficult to recognize without careful comparison of
their shades and tints, should possess nostrils structurally differing
in all three is remarkable ; but is it a sufficient reason to place them
in three different genera? I can only regard the character as
being specific.
The plumage of the sexes is alike. The amount of dark chestnut
on the middle pairs of rectrices varies considerably, from three inches
to one inch in depth.
9, CENTROCOCCYX EURYCERCUS.
Centropus eurycercus, A. Hay, Blyth, J. A. S. B. 1845, p. 551.
[P. Princesa, ¢, January 8, 1878: iris bright crimson; bill,
legs, and feet black. < (juv.), December 9, 1877: iris neutral
tint ; bill and legs jet-black. |
Rather smaller than the Malaccan type.
10. Lanrus LUZIONENSIs (72).
[P. Princesa, ¢ 2, December 6 and 11, 1877.]
11. GRAUCALUS SUMATRENSIS.
Ceblepyris sumatrensis, S. Miller, Verh. Land- en Volkenk. p. 191.
LP. Princesa, 2, December 1877 : iris light lemon-yellow. ]
Does not differ from Sumatran, Malaccan, and Bornean examples.
12. DicruRUS PALAWANENSIS.
In the: Philippines three species of Dicruride are known :—
D. balicassius, type of the genus Dierurus; D. mirabilis, its repre-
sentative form; and D. striatus. This last, by its even, almost
unfurcated tail, resembles D. dalicassius in structure, but in its
general colouring and in the distribution of its markings exhibits a
close relationship to the Papuan and Malaccan species associated by
Mr. Sharpe with Chibia hottentotta. These Papuan species seemed
to me to belong to a group distinct from that represented by D. bali-
cassius on the one hand and Chibia hottentotta on the other; and
their geographical range favoured this view. But Mr. Everett has
discovered in Palawan a species which undoubtedly belongs to the
Papuan section of the Dicruridz ; and it would appear that, with
D. striatus as a connecting link, the Papuan and the Philippine
species must be regarded as members of one section of the family, to
which the title of Dicrurus should be applied. Besides this unde-
scribed species, Palawan is inhabited by at least one other member
of the family, belonging to the genus Buchanga ; and Palawan and
Lombock are the only two islands or areas known to me where there
is a second species associated with a true species of Dicrurus, unless,
* The type was from Balabac; but the Palawan bird does not appear to differ,
* Count T. Salvadori has recently (anted, p, 88, note) proposed the generie
title of Dierwropsis for this group.
1878.] THE ORNITHOLOGY OF THE PHILIPPINES. 615
indeed, D. lemostictus is shown to also inhabit New Ireland or D.
megarhynchus New Britain.
Male and female. Upper surface of wings and tail burnished
bronze-green. Glistening scale-like feathers on crown of head bluish
metallic green. Neck-hackles and pectoral and gular spots dark
metallic steel-blue ; these spots, which extend from lower part of the
throat and cover the breast, are lengthened and ovate in form rather
than round, and occupy the tips of the feathers. With their excep-
tion, the entire under surface is dull jet-black. The nape, back, and
uropygium is black with a blue gloss. The tail is forked for about
one inch in depth; and the distance between the extremities of the
outer pair of rectrices is nearly three inches ; upper tail-coverts tipped
with metallic green.
Dimensions :—
Wing. Tail. Tarsus. Culmen.
in. in. in.
oe ne oor ae 3°5 1-0 1°5
{P. Princesa, ¢, November 1877: iris crimson; bill and feet
black. 2, December: iris deep brown. |
All the female birds have the iris noted as being deep brown by
Mr. Everett. On the label of one of the males Mr. Everett has
written :—“ Iris in the living bird deep crimson.”
D. palawanensis most nearly resembles D. leucops, D. pectoralis,
and D. lemostictus ; but, besides being much smaller, the shape and
position of the pectoral and gular spots are different, not being so
round, but narrower.
13. BUCHANGA LEUCOPHAA.
Dicrurus leucopheus, Vieill. Nouv. Dict. d’ Hist. Nat. ix. p. 587.
Buchanga cineracea (Horsf.), Sharpe, ¢. c. p. 324. no. 48.
[P. Princesa, 3, November 29, 1877: iris crimson; bill and feet
black. o, December: iris deep rich orange-brown. @, December ;
iris orange-brown. |
14. CyorNis BANYUMAS.
Muscicapa hanyumas, Horsf. Tr. L. 8. xiii. p. 146.
Muscicapa cantatriz, Tem. Pl. Col. 226.
[P. Princesa, ¢ , December 1877: iris chocolate; bill black ; legs
and feet purplish grey; claws dark grey. Q, January 1877: iris
chocolate ; bill black ; legs lead-grey. |
The female of this species is well described, but badly figured, by
Temminck (J. ¢.).
15. Hyporuymis azurREA (85).
[P. Princesa, 2, December 1877: iris deep brown; bill entirely
black, interior of gape golden; legs and feet dark blue-grey. |
16. HrruNpDO JAVANICA.
Hirundo javanica, Sparrm. Mus. Carls. t. 100.
[P. Princesa, 9, December 7, 1877. |
616 THE MARQUIS OF TWEEDDALE ON [May 21,
17. BRODERIPUS PALAWANENSIS, 0. sp.
[P. Princesa, 2, January 1878. |
A representative form of B. acrorhynchus, but smaller and with a
shorter bill. The yellow space included with the black horseshoe-
mark on the head extends far beyond the eyes, and measures in depth
over one inch. There is no yellow alar spot.
Dimensions :—
Wing. Tail. Tarsus. Culmen.
in. in. in. in.
Oe. OU 4°5 1-0 1:5
18. Or10oLUS XANTHONOTUS.
Oriolus zanthonotus, Horsf. Tr. L.S. xiii. p. 153; Zool. Res. Java,
t. 46; Temm. Pl. Col. 214.
[P. Princesa, ¢, December 30, 1877: iris crimson ; bill burnt
sienna ; legs dark lead-grey. |
In the colouring and markings the single specimen sent by Mr.
Everett (an adult male) is not to be distinguished from Malaccan,
Bornean, and Sumatran individuals; but it greatly exceeds them by
its dimensions—wing 4°63 inches, tail 3°50.
19. TRICHOSTOMA RUFIFRONS, 0. sp. (Plate XXXVIII.)
Mr. Everett has sent three skins of a Timeliine bird which appear
to belong to a somewhat aberrant form of the genus Trichostoma.
The bill, seen in profile, has the exact contour of the bill of T. abbot,
but is somewhat stouter; at the base it is slightly broader ;
and the rictal bristles are longer. The tail is also more graduated,
the outer pair of rectrices being shorter than the penultimate pair,
which, again, are not quite so long as the remainder. Perhaps this
species is congeneric with the two described and figured by Mr. Sharpe
(t.c.) under the titles of Zeocephus cinnamomeus and Z. cyanescens
(species with which I am unacquainted); but if so, they cannot be
classed under the Muscicapide.
Male and female. Forehead bright rufous-brown, contrasting with
the duller rufous-brown of the vertex and occiput. Nape and back
earthy brown, tinged with olive and paler on the nape. Long, lax
feathers of the uropygium bright rufous. Upper tail-coverts pure
chestnut ; rectrices dull chestnut. Lores and supercilium, which
extends behind the eye, pure grey. Cheeks and ear-coverts greyish
brown. Chin, throat, and vent pure white. Breast, abdomen, flanks,
under tail-coverts, and under wing-coverts tawny, with a pale brown
shade on the breast. Primaries brown, edged with pale tawny
chestnut. Remaining quills and the wing-coverts dull chestnut,
tinged with the colouring of the back.
Dimensions :—
Wing. Tail. Tarsus. Culmen.
in. in. in. in.
Bove OrOr 3°50 1°00 0-94
Y oti Ole 3°25 0°95 O87
1878. ] THE ORNITHOLOGY OF THE PHILIPPINES. 617
[P. Princesa, ¢ 2, December 10, 1877: iris pale Naples-
yellow ; dill dark plumbeous ; feet pale plumbeous. ]
20. DryMocATAPHUS CINEREICEPS, Ui. sp.
This species is congeneric with Brachypteryx malaccensis, Hartl.,
which Count T. Salvadori retains under the genus Brachypteryz.
But B. malaccensis is certainly nearer to Drymocataphus than to Bra-
chypteryx, although it may perhaps be advisable to form a separate
genus for its reception, more especially as the species which inha-
bits Palawan, about to be described, is identical in structure with the
Malaccan and Bornean form (which is also said to inhabit Su-
matra), and also as Brachypteryx murinus is congeneric.
Female. Forehead, crown, and occiput dark iron-grey. Ear-
coverts, space before the eye, and nape pale grey. Back, uro-
pygium, scapulars, upper tail-coverts, and exposed surface of wing
and tail ruddy brown, brighter than in D. malaccensis. Dorsal
plumes pale-shafted. Chin, throat, upper breast, abdomen, and vent
pure white. Flanks, a band across the breast, under tail-coverts, and
thigh-coverts, wing-lining, and axillaries pale rufous-brown.
Dimensions :—
Wing. Tail. Tarsus, Culmen,
in.
im. in.
Q.... 2°50 1:00 1-06 0°75
[P. Princesa, 2, January 1878: iris burnt sienna-orange; bill
brownish grey, mandible white ; legs pallid, the front of tarsus tinged
brown. }
Distinguished from D. malaccensis by its dark-grey head, its
light-grey ear-coverts, and the brighter rufous of the dorsal plumage.
21. Mrxornis woont.
Mixornis woodi, Sharpe, t.c. p. 331. no. 69.
[P. Princesa, ¢, December 5, 1877: iris dark ochre-yellow ; bill
black, mandible grey ; legs and feet horn-yellow. 9, January 1878:
iris ochreous orange; bill dark grey, mandible pale grey; legs
greenish olive. |
22. BRACHYPUS CINEREIFRONS, 0. sp.
Male and female. Chin, throat, and breast pale cinereous brown,
palest and whitest on the chin and throat. Abdomen and ventral
region pale yellowish white, in some examples with a certain admix-
ture of the colouring of the breast. Flanks pale earthy-brown.
Thigh-coverts ochreous yellow. Under tail-coverts light yellow, with
a slight ochreous tinge. Wing-lining and axillaries pale yellow ;
carpal edge ochreous yellow. LEar-coverts pale cinereous, with
almost white shafts. Space before the eye uniform dull brown.
Cheeks dull. cinereous brown, with a few white-shafted feathers.
Head and nape olive-brown, tinged with pale grey, the frontal
feathers being distinctly tipped and centred with pale grey. Back,
scapulars, wing-coverts, and uropygium olive-brown, olive predomi-
Proc. Zoou. Soc.—18738, No. XL. 40
618 THE MARQUIS OF TWEEDDALE ON {May 21,
nating, the long, lax, and fluffy uropygial feathers being of a slightly
lighter shade of olive-green. Upper tail-coverts olive-green. Quills
light brown, margined with a brighter olive-green. ectrices light
brown, edged with olive-green, and some with pallid yellowish apices.
Underneath pale brown, washed with very light yellowish green.
Shafts above brown, below yellowish white.
Dimensions :-—
Wing. Tail. Tarsus. Culmen.
m. mm. m. in.
Si W.L1SSP 3°37 0°88 0°88
Oy) as 25 3°12 0°87 0°75
[P. Princesa, ¢, December 14, 1877: iris deep rich burnt-sienna ;
bill blackish brown; legs and feet purple-brown. ¢, November 28:
iris chocolate; bill dark brown ; legs pale brown. |
One example marked 2 equals the ¢ in its dimensions.
Mr. Everett procured a good series of this short-footed Thrvsh.
At first sight it can be easily mistaken for B. plumosus (Blyth), of
which it is a representative form. It is to be distinguished by
its pale abdomen, which in B. plumosus is browner and nearly
uniform with the breast, by its pale greyish breast, by its yellow under
tail-coverts, which in B. plumosus are ochreous-brown, by its pale
yellow, and not ochreous, wing-lining and axillaries, and by the
cinereous or pearl-grey edgings and centres of the frontal plumes.
Nor is the green of the quills and rectrices nearly so developed. The
colouring of the abdominal plumage resembles that of [ole olivacea,
but is not so distinctly yellow.
23. BracHYPODIUS MELANOCEPHALUS.
Lanius melanocephalus, Gm. 8. N. i. p. 309.
[P. Princesa, ¢ 2, January 1878: iris violet-blue ; bill and legs
black. |
Undistinguishable from typical examples.
24. CRINIGER FRATER.
Criniger frater, Sharpe, t.c. p. 334. no. 77.
[P. Princesa, 3, December 1, 1877: iris red-brown; bill horn-
grey, darker on culmen. @, December: iris red-brown; bill dark
grey ; mandible whitish ; legs, feet, and nails pallid brown. |
25. CRINIGER PALAWANENSIS, 0. sp.
Male and female. Chin, lower breast, abdomen, vent, under tail-
coverts, wing-lining, axillaries, and carpal edge bright yellow. Feathers
of the throat, cheeks, and upper breast of the same yellow, the centres
towards the apices being pale grey. These parts have thus a streaked
appearance. Sides of the breast and the flanks clouded with pale
brown ; a few of the nareal plumes bright yellow. Space before
the eye yellowish grey. Head, ear-coverts, and nape reddish brown,
the frontal and coronal plumes being sublanceolate and with pale
central streaks. Back, scapulars, wing-coverts, and uropygium
1878.] THE ORNITHOLOGY OF THE PHILIPPINES. 619
ruddy olive-brown. Upper tail-coverts, which are almost concealed
by the mass of loose and lax uropygial plumes, ruddy brown. Rec-
trices of a darker tone than the tail-coverts. Quills brown, mar-
gined with the colour of the dorsal plumage.
Dimensions :—
Wing. Tail. Tarsus. Culmen.
in. in. in. in,
BUD Weretige B25 3°20 0°75 0°87
[P. Princesa, 3, January 1878: iris lemon-yellow. 2, Decem-
ber 27, 1877: iris lemon-yellow ; bill dark smoky grey, the mandible
whitish ; legs and claws pale brown. |
26. PHYLLORNIS PALAWANENSIS.
Phyllornis palawanensis, Sharpe, ¢. c. p. 333. no. 72, t. 50.
alee?
[P. Princesa, ¢ 9, January 1878.]
The only example marked 2 has the terminal portions of the
moustache deep blue. ‘Two examples marked ¢ have it white.
Mr. Sharpe’s conclusions (/. c.) on the point were the reverse. This
is one of the most beautiful species of the genus. The throat is
bright golden; and the whole colouring of Mr. Keuleman’s figures
(@. ¢.) is much too pallid.
27. ASGITHINA SCAPULARIS.
Tora scapularis, Horsf. Tr. L. 8. xiii. p. 158.
[P. Princesa, ¢ 2, December 12, 1877: iris white; bill lead-
grey, the culmen black ; legs and feet bluish grey ; claws black.]
All the males. are without a trace of black on the upper parts.
They agree with Bornean and Malaccan examples (J. viridis, Bp. ?).
Bornean individuals are stated by Count T. Salvadori to be identical
with Javan (Ucc. Borneo, p. 191).
28. Monrticota souitarivs (103).
[P. Princesa, ¢ 2, December 1877. ]
29. CrTTOCINCLA NIGRA.
Cittocincla nigra, Sharpe, t. c. p. 335. no. 82, t. lil. f. 1, 2.
[P. Princesa, 3, December 14, 1877: iris very dark brown ; bill,
legs, and feet black. @Q, January 1878: iris dark brown; bill
black ; legs lead-grey.]
30. ORTHOTOMUS RUFICEPS.
Edela ruficeps, Lesson, Tr. d’Orn. p. 309.
[P. Princesa, ¢ 2, December 1877.]
31. ANTHUS MACULATUS.
* Anthus maculatus, Hodgs. Gray’s Zool. Mise. p. 83 ; Tweeddale,
P. Z.S. 1877, p. 596. no. 48.
[P. Princesa, 2, January 2, 1878. ] a
620 THE MARQUIS OF TWEEDDALE ON [May 21,
32. MyzanTHE pyGma (121).
[P. Princesa, ¢, January 7, 1877: iris dark brown ; bill and legs
black. |
33. NECTAROPHILA SPERATA (122).
[P. Princesa, 3, January 1878. @, December 29, 1878: iris
warm brown. |
Two males, not adult; the violet throat and plastron bounded on
both sides with a white border, the scarlet breast mixed with white,
the crown dingy dull brown.
34. CyRTOSTOMUS AURORA, Nl. Sp.
Notwithstanding that Captain Shelley, in his beautiful work on
the Sun-birds, includes the large group of Asiatic and Australian
Nectariniidee, of which C.jugularis and C. asiaticus may be taken as
types, along witha number of other species, under Cuvier’s genus
Cinnyris, I still adhere to the view expressed by me (Ibis, 1870,
p- 19) that these Sun-birds forma natural, distinctive, and separate
section of the family. For it I adopted Dr. Cabanis’s generic title
Arachnechthra; but as the type of that genus is C. lotenia, L., which
may be considered generically separable (although not so in my
opinion), and as Dr. Cabanis established his genus Cyrtostomus
(type C. jugularis, L.) before that of Arachnechthra, according to
strict rule, the generic title of Cyrtostomus should be employed; and
I observe that this is the view of Count Salvadori. There are now
twelve species of this group described—namely, C. asiaticus, C. lote-
nia, C. osea, C. brevirostris, C. flammaaillaris, C. andamanicus, C.
rhizophore, C. pectoralis, C. frenatus, C. jugularis, C. zenobia, and
C. solaris. To this list Mr. Everett’s researches in Palawan enable
me to add a thirteenth.
Male. Above uniform dark olive-green. Forehead, chin, throat,
cheeks, and upper breast metallic steel-blue with shades of green
and violet in certain lights. Lower breast fiery orange-red. Re-
mainder of lower surface, under tail-coverts, and wing-lining bright
yellow. Quills brown, faintly edged with olive-green. Rectrices
black; terminal half of outer pair white, a broad white terminal
patch on the penultimate pair; apices of third pair fringed with
white, of fourth pair still less so.
Female. Above and ear-coverts olive-green, slightly lighter in shade
than that of male. Superciliaries, cheeks, and all the under surface
and the wing-lining bright yellow. Wing-feathers brown, margined
with olive-green. Rectrices as in male,
Dimensions :—
Wing. Tail, Tarsus. Culmen.
in. in. in. in.
ee Be 220 1:62 0°62 1:00
Oe siaccsO0 1°62 0°60 0°87
[P. Princesa, ¢, November 30, 1877 : iris deep crimson ; bill and
legs black. Q, Jannary: iris dark brown; bill and legs black. ]
1878.] THE ORNITHOLOGY OF THE PHILIPPINES. 621
The flame-coloured lower breast at once distinguishes this Sun-
bird from all others. Its nearest ally is C. solaris, in which the
flame-colour covers the abdomen also.
35. CHALCOSTETHA INSIGNIS.
Nectarinia insignis, Jardine, Nat. Lib. xxxvi. p. 274.
[P. Princesa, ¢, January 1878: iris warm brown; bill and legs
black. ]
A single example of an adult male, and not to be distinguished
from Sumatran and Malaccan individuals,
36. AETHOPYGA SHELLEYI.
Aithopyga shelleyi, Sharpe, Nature, 3rd August, 1876, p. 297 ;
Tr. L. 8, 2nd ser. Zool. i. p. 343. no. 105; Shelley, Monogr. Cin-
nyridee, pt. iii. t.
[P. Princesa, ¢, December 2, 1877: iris brown; bill black 3 legs
and feet very dark brown. 9 (?), January 4, 1877: bill and legs
dark brown. |
The female has not been described; and it is with some doubt
that I so identify the single skin noted above. The chin, throat,
and upper breast is pale grey ; the lower breast and abdomen and
wing-lining are pale yellowish white. The vent and under tail-coverts
of a more decided shade of yellow. The head is greyish olive-brown.
Remainder of upper surface a clear olive-green, lightest on the upper
tail-coverts. The quills and rectrices are dark brown edged with
olive-green, having a slight ruddy hue. Cheeks and ear-coverts pale
grey with a greenish tinge. The dimensions are sensibly smaller
than those of the male—wing 1°75; culmen 0°56.
37. ANTHREPTES MALACCENSIS.
Certhia malaccensis, Scopoli, Del. Flor. et Faun. Insubr. ii. p. 91.
no. 62.
[P. Princesa, ¢, January 1878: iris Indian red ; bill black ; legs
greyish olive; scales of feet yellow. 2, December 29, 1877: iris
bright Indian red ; bill very dark sepia-brown; legs and feet yel-
lowish olive ; soles of feet yellow; claws pale brown. ¢ (jun.),
November 27, 1877: iris Indian red; bill dark brown; legs yel-
lowish olive. ]
The example of the young male above noted is in the exact plumage
of the adult female, its sex being betrayed by a couple of metallic
plumes on the crown.
38. ARACHNOTHERA DILUTIOR.
Arachnothera dilutior, Sharpe, Nature, 3rd August, 1876, p- 297;
t. c. p. 341. no. 100; Shelley, Cinnyride, pt. iii. t.
[P. Princesa, ¢, December 6, 1877: iris brown; bill black ; legs
and feet dark grey. 2, January 1878: iris dark brown; denuded
orbital skin green-yellow ; bill black ; mandible grey ; legs and feet
blue-grey.]
Bill of female considerably shorter than that of male.
622 THE MARQUIS OF TWEEDDALE ON [May 21,
39. CoRVUS PUSILLUS, n. Sp.
Female. Basal portion of plumage white. Above purple-black, with
aslight greyish shade. Wing-coverts and outer margins of rectrices
deep purple-black. Plumage underneath dull ashy black, with a
purplish gloss on cheeks and throat. Contour of bill as in the C.-
validus group.
Dimensions :—
Wing. Tail. Tarsus, Culmen.
in, in. in,
Dineen Ore 9°25 1°75 2712
[P. Princesa, 2 , December 27, 1877: iris deep brown ; bill, legs,
and claws shining black. ]
This Crow, or rather small Raven, is a diminutive form of C. vali-
dus and its allies. It has no affinity with true C. enca.
40. CALorRNIS PANAYENSIS (128).
Calornis chalybeus (Horsf.), Sharpe, ¢. ¢. p. 343. no. 111.
[P. Princesa, 2, December 1877: iris brilliant red; bill and feet
black. ]
A large series is sent by Mr. Everett, which I refer to the Philip-
pine rather than to the Bornean species.
41. GRACULA JAVANENSIS.
Corvus javanensis, Osbeck, Voy. China & E. Ind. i. p. 157,
** Java”’ (Eng. tr. 1771).
[P. Princesa, 3, January 1878: iris deep brown; bill fine deep
orange ; wattles and legs chrome-yellow; claws dark brown. 9°,
December 3, 1877: iris deep brown; bill red-orange; wattles
orange-yellow ; legs yellow. |
These examples are not to be distinguished from Bornean, although
they are somewhat smaller and have slenderer bills than Javan and
Sumatran individuals.
42. OxyYCERCA EVERETTI.
Oxycerca everetti, Tweeddale, P. Z. 8S. 1877, p. 699, t. Ixxiii. f. 2.
[P. Princesa, ¢, December 1877: iris rich deep orange-brown ;
bill, maxilla black, mandible pale grey; feet, legs, and claws dark
lead-grey. _¢ juv.: iris umber-brown; mandible blackish ; maxilla
black ; angle of gape pure white ; legs, feet, and claws lead-grey. |
Oxycerca everetti is a very closely allied form of O. leucogastra
of Malacca and Borneo, mainly to be distinguished by the crown of
the head being striated in the former and uniform in the latter, and
the breast and general colouring being nutmeg-brown and not sooty
brown. These Palawan examples, although not quite identical with
either species, most nearly resemble O. everetti. The general tone
of colour is nutmeg-brown; but the crown of the head is not
striated.
1878. | THE ORNITHOLOGY OF THE PHILIPPINES. 623
43. OsMOTRERON VERNANS (135).
[P. Princesa, 2, January 2, 1878: iris (in dead bird) orange ;
bill lead-grey, base yellow-olive ; feet carmine ; claws grey. |
44, TRERON NASICA.
Treron nasica, Schlegel, Ned. Tijdschr. Dierk. i. p. 67.
[P. Princesa, ¢ 9, January 1878: iris deep brown, with outer
ring of burnt sienna-orange ; orbital skin bright emerald-green ;
base of bill dull crimson, the rest pale greenish-yellow ; feet carmine. |
45. CARPOPHAGA NEA (141).
[P. Princesa, ¢, January 1878: iris pure crimson; bill bluish
grey ; feet violet-carmine ; nails dark grey. 9, December 28, 1877:
iris deep crimson ; orbital ring crimson; bill light grey ; feet car-
mine ; nails dark grey. |
46. PotypLeCTRON EMPHANES.
Polypectron emphanum, Temm. Pl. Col. 540; Elliot, Phasianide,
vol. i. pl. 9.
[P. Princesa, ¢, January 1878: iris dark rich brown ; orbital
aud loral regions crimson ; bare setigerous skin above the eye livid
brown ; bill, including vaulted base of maxilla, very dark brown, tip
horny ; legs, feet, and nails purple-brown.
Native name “ Tandikan.”’
Q juv.: orbital skin black, otherwise not different from the male. |
The male of this beautiful Pheasant was first described and figured
‘by Temminck on the 14th of May, 1831, from a single example in
the museum of the Prince of Essling (J. ¢.). Its habitat was
unknown, but was believed to be either one of the Sunda Islands
or else one of the Moluccas. In the same year, but earlier, Lesson
(Tr. p. 437) enumerated the species under number 47 of the genus
Polyplectron, and called it P. napoleonis; but as he gave no de-
scription, his title falls.
A second specimen of the species was afterwards acquired from
Messrs. Verreaux by the British Museum; but the habitat still re-
mained unknown, the late Mr. G. R. Gray giving it as the Moluccas
(List Birds B. Mus., Galline, 1867, p.25). Mr. Sclater (P. Z. 8.
1863, p. 124) attributed its habitat, with some doubt, to the island
of Borneo, in which he was followed by Mr. Elliot (/.c.). Mr.
Everett has now cleared up all doubts; for he has found this Phea-
sant living in the island of Palawan near the Spanish settlement of
Puerto Princesa, and has sent me three examples—one of the adult
male, one of the immature female, and one of an immature male.
This is the second instance in which Mr. Everett has established
the correct habitat of a little-known and remarkable bird.
The young male has the lengthened crest-plumes brown, partially
tinged with green. The brilliant bluish-purple wing-coverts and
interscapulars of the fully adult are absent, only a trace presenting
itself on one covert of each wing. Whereas in the adult the nape
and neck are black with brilliant green borders to the feathers, and
624 ON THE ORNITHOLOGY OF THE PHILIPPINES. [May 2],
the whole throat and breast are black with green reflexions, the
immature bird is brown. The quills and underparts are also very
dark brown in the adult, whereas in the immature bird they are
light brown. The ocelli on the rectrices in the immature bird are
not so perfectly formed, and the dark brown ground-colour of the
dorsal plumage and of the rectrices is not so well defined.
The skin of the female (immature) has the chin, space before the
eye, and throat dingy white. The green ocelli only occur on some
of the rectrices, and are small. The remainder of the plumage
resembles that of the young male, the crest being shorter and uni-
form brown.
47. MEGAPODIUS CUMINGII.
Megapodius cumingii, Dillwyn, P.Z.S. 1851, p. 118, t. 39.
[P. Princesa, 3, January 1878: iris warm brown; orbital skin
coral ; auricular region washed with bright red; bill horn-yellow ;
legs reddish, brighter than in female ; tarsal scutellee and feet dark
olivaceous brown; the red of the face brighter than in female.
3 juv., January 1878: character of soft parts as in mature male,
only paler. 2, January 2, 1878: iris warm brown; bill horn-yellow ;
orbital skin coral ; auricular region washed with vermilion-red (not
crimson) ; legs yellowish grey, the tarsal scutelle dark grey.
Note.—In these birds the bill is sometimes horn-yellow, some-
times the same more or less deeply tinged with brown on culmen
and at the base. ‘'Tabun”’ of the natives. |
Dimensions :—
Wing. Tail. Tarsus. Culmen,
in. in. in. in.
(ot ne 9:00 3°50 2°50 125
Oe dugsthia 9°25 3°50 2°50 1°18
The Palawan megapode closely resembles the Labuan (M. cu-
mingii), and can hardly be separated specifically. In this view Mr,
Sharpe concurs.
48. EvpromtiAs GEOFFRoy! (161).
[P. Princesa, ¢, November 29, 1877.]
49. JKGIALITES CANTIANUs:
Charadrius cantianus, Lath. Suppl. Ind. Orn. p. Ixvi.
jP. Princesa, 2, November 29, 1877. ]
50. TRINGOIDES HYPOLEUCUS (183).
[P. Princesa, 2, December 11, 1877.]
51. BuBpuLcus coromanvus (194).
[P. Princesa, ¢ 9, December 1877. ]
Forehead in both examples buffy yellow.
52. BuTorIpEs JAVANICA (197).
[P. Princesa, 2 juv., January 6, 1878. ]
1878.] ON THE TRACHEX OF TANTALUS AND VANELLUS. 625
7. On the Trachea of Tantalus loculator and of Vanellus
cayennensis. By A. H. Garrop, M.A., F.R.S.
[Received May 21, 1878.]
Tn his ‘ Beitriige zur Naturgeschichte von Brasilien,’ Maximilian,
Prince of Wied’, describes briefly and figures the lower end of the
trachea of Tantalus loculator. A male specimen of the species
having recently died in the Society’s Gardens, I take the opportunity
of more minutely pointing out its peculiarities and of comparing it
with 7’. ibis, the windpipe of which, with its elaborate convolutions,
1 have had the opportunity of bringing before the notice of the
Society upon a previous occasion’.
In Tantalus loculator the trachea is not elongated as it isin 7. ibis ;
nevertheless it is peculiarly modified, and differs in detail from that
of any bird with which I am acquainted, although its plan of con-
struction is perfectly Ciconiine.
The seventy-eight lowermost rings of the trachea are those which
are modified, the rings above them being quite typical, of average
depth, notched in front as well as behind, and overlapped to produce
the well-known zigzag markings on the surface.
With the exception of the last one, all the modified rings are much
reduced in depth; and of them the sixty-one upper rings are com-
pressed from side to side and bent sharply in front, whilst the lower
seventeen are somewhat flattened from before backwards and sharply
bent laterally, the general effect of which is to produce a lateral flat-
tening and an anterior carination of the whole tube opposite the
fifty-one rings, as well as an antero-posterior flattening with a lateral
carination in the part below. The change from the superior unmodi-
fied tube to the laterally compressed portion is somewhat abrupt, as
is that between the two differently modified parts. In figure 1 a
(p- 626) the front view of the lower end of the trachea is repre-
sented, figure 4 giving a side view of the same.
The powerful sterno-tracheal muscles leave the wind-pipe opposite
the middle of the laterally flattened portion of the tube; and a few
muscular fibres from their anterior margin are continued downwards
for a short distance, but not nearly to the last ring, they being lost
upon the sides of the trachea,
The arrangement above described is only an exaggeration of what
is found in Ciconia alba, in which species the lowermost nine-and-
twenty tracheal rings are extremely shallow and slender, the fifteen
above the bifurcation of the bronchi being antero-posteriorly flattened,
the fourteen above them being in no wise peculiar except for their
slenderness, In C, aida there is, however, a small prolongation up-
wards of the lateral portions of the three lowermost tracheal rings,
which forms a consolidated triangular process on each side, overlap-
ping the next few rings, and looking extremely like rudiments of the
? Band iv. p. 687, tab. 1, figs 7 and 8,
2 P. ZS, 1875, p. 298.
626
MR. A. H. GARROD ON THE TRACHEZ
Fig. 1.
Lower end of the trachea of Tantalus loculator ¢.
a. View from the front; 6. View from the right side.
[May 21,
1878. | OF TANTALUS AND VANELLUS. 627
similarly situated processus vocales of the passerine tracheophone
syrinx, which resemblance is increased by the thinness of the neigh-
bouring rings and their being flattened from before backwards.
In Tantalus loculator there is no trace of these triangular processes.
Its last tracheal ring, or three-way piece, is not enlarged, as it is in
so many birds; and the rings of the bronchi for some considerable
distance are complete as in the Ciconiide generally, which is so very
seldom found to be the case in the Class. In this last feature the
Storks agree with the Cathartide, and the general arrangement of
the bifurcation of the Stork’s windpipe would require but little change
to pass into a Cathartine type.
The uppermost bronchial rings are thinner on the outer side of
each bronchus than they are internally, which consequently leaves
greater gaps between them along the outer margin of the tubes.
Ring four on one side and ring three on the other are partly redupli-
cated, the extra processes ending freely in the bronchial membrane.
From this description it is evident that these two Tantali differ
greatly in the arrangement of their windpipes, whilst a recent com-
parison of specimens makes it evident to me that what I thought on
seeing 7’. Joculator might have been an error in my account of 7.
ibis, namely the posterior carination of the windpipe, is correct, in
which, as well as in the relative lengths of trachea, the two species
differ so much.
In other anatomical characters Tantalus loculator agrees with T.
ibis, and is perfectly ciconiiform. In both the great pectoralis
muscle is formed of two layers, as in the Steganopodes, Procellariide,
and Cathartide only. The ambiens muscle is slender ; the femoro-
caudal is minute, without any accessorius ; and the semitendinosus, as
well as its accessorius, are not large. There is no great gluteus muscle,
nor any muscular slip from the biceps of the wing running to the
patagium.
The small intestine measures six and a half feet, the czeca half an
inch, and the large intestine nearly three inches. The stomach is
capacious, with but a small muscular development. The tongue is
an inch long and arrow-head shaped. The sub-equilobed liver has
a gall-bladder.
In Vanellus cayennensis there is an exaggerated development of
the intrinsic muscles of the trachea a short way above its bifurcation
in both sexes that is quite worthy of special note, because the amount
of muscular fibre there present is proportionately as much as in any
bird with which I am acquainted.
There is nothing peculiar about the windpipe itself or the bronchi,
which are represented in the accompanying figure (p. 628). The
uppermost two bronchial semirings are not like those which succeed
them, but are like halves of tracheal rings. The third and fourth
semirings are closely united, whilst those which follow are not
modified in any way.
The sterno-tracheal muscles are powerful, and besides springing
directly from the side walls of the windpipe opposite the spot where
628 ON THE TRACHE# OF TANTALUS AND VANELLUS. [May 21,
Fig. 2.
b.
Windpipe of Vanella cayennensis.
a. Anterior aspect; 4. Posterior aspect.
1878.] MR. GARROD ON THE ANATOMY OF THE MALEO. 629
they run off, their upper fibres are continuous up the trachea itself
in front of and in contact with the intrinsic muscles. These latter,
one on each side as usual, meet in the posterior middle line of the
trachea, but are not unusually near in front; they are of considerable
size throughout. Near their lower ends they increase immensely in
bulk to form, combined posteriorly, a large subglobose mass which
is situated opposite the twenty-four lowermost rings of the trachea,
which are considerably shallower than those above them and conse-
quently occupy a much less space than if they were of the same depth,
as is the case in Tantalus loculator. There is a consolidation of the
last few rings in adult birds, with which the first two bronchial semi-
rings fuse to form a compound three-way piece, and it is to the
lower elements of this that the powerful lateral muscles are attached
(as well as to the third and fourth bronchial semirings slightly) by a
broad fibro-tendinous continuation of their muscular substance, which
fixes itself on each side along nearly the whole length of the semi-
rings, especially the second, of which the extremities are alone free.
So far as I can find out by watching the living birds, there is nothing
peculiar in their note to lead one to surmise so large a muscular
supply for their lower larynx. They make a powerful screech, with
no modulation in it; and it can hardly be possible that the extra
muscular development has not some other function to perform.
What that may be it is not easy to surmise,
8. On the Anatomy of the Maleo (Megacephalon maleo).
By A. H. Garrop, M.A., F.R.S.
[Received May 21, 1878.]
Besides being a Megapode, Megacepkalon maleo is interesting as a
peculiar form; I therefore take the opportunity of bringing before
the Society the results of my dissection of three specimens of this
bird which have died in the Gardens.
Péerylosis.—In the distribution of its feather-tracts the Maleo
is somewhat different from the typical Gallinze. Nitzsch briefly
records what he found in a bad specimen of Megapodius rubripes,
mentioning that the tracts were not different from those in allied
birds, and that the oil-gland was tufted. This is all we know of the
pterylosis of the group.
In Megacephalon maleo the anterior surface of the neck is covered,
not thickly, with feathers, which only tend to divide opposite the
furcula into the two pectoral tracts, each of which descends, strong
and uniform in breadth, to opposite the middle of the carina sterni,
where it ceases obtusely. ‘The ventral tract does not exist over the
anterior or upper part of the pectoral region, but commences narrow
close to and opposite the middle of the carina sterni, dilating oppo-
site the abdomen, near the middle line of which it descends parallel
to its fellow, to just above the anus, where the two meet. The skin
over the carina and in the middle line of the abdomen is hard and
630 MR. GARROD ON THE ANATOMY OF THE MALEO. [May 21,
scaly. The hypopteral tract is but feebly represented, and the
termination of the pectoral tract is some distance from the commence-
ment of the abdominal tract, the angle between the direction of the
one and the other being about 25°.
A strong tract traverses the middle dorsal line of the neck, with a
space on each side of it. This does not bifurcate in the scapular
region, but ceases abruptly a little below the level of the shoulder-
joints, undilated. Over the rump and the inferior scapular region
the dorsal tract is diffused and indefinite, ceasing before the nude oil-
gland is reached. There is no ephippial space. There are sixteen
rectrices. The lumbar tract is only well developed over the tibia-
head and for a little way behind it. ‘The humeral tract is not in any
way peculiar. I counted nine primary and sixteen secondary
remiges, the first and second secondary feathers being considerably
shorter than those which follow.
Down-feathers are generally distributed. The after-shaft is weak,
and the rhachis of each feather is not swollen, except in some of the
smallest size.
Lower larynx of Megalocephalon maleo.
a. From the front; . From behind.
Visceral Anatomy.—The tongue is simple and fleshy ; a well-de-
veloped crop is present, situated between the limbs of the very open
furcula; the proventriculus is zonary, the gizzard powerful; the
small intestine is four and a half feet long, the simple ceca five and
a half inches, and the large intestine five inches in length.
There is only one carotid artery, the left, as in all the Megapodiidee.
In its myology the bird is perfectly gallinaceous, the third pectoral
muscle being found beneath the much larger second of the same
name, the femoro-caudal and semitendinosus with their accessories,
the ambiens, gluteus maximus, being all present. The obturator in-
ternus is triangular; a vinculum joins the two deep flexors of the
foot, and the biceps of the arm sends a fasciculus to the patagium.
The expansor secundariorum muscle ends by running to the scapula
at the same time that it sends a slip of tendon to the first rib.
1878.] THE SECRETARY ON ADDITIONS TO THE MENAGERIE. 631
The lower end of the trachea is represented in the accompanying
figures (p.630), from the front and from behind. It differs in arrange-
ment from all other of the Gallinee with which I am acquainted.
To the terminal tracheal three-way ring is fused the first bronchial
semiring by its extremities, its bowed free portion being upturned
with its limbs almost perpendicular. In the middle line posteriorly
the penultimate tracheal ring fuses with the what may be compound
three-way piece, in the posterior aspect of which a small upward-
directed tongue of cartilage develops from its upper border. The
sterno-tracheal muscles are minute and the intrinsic tracheal muscles
form thin broad sheets which almost cover the tube and end on the
outer surface of the first bronchial semiring and in the semicartila-
ginous large membrane between it and the unmodified following
semiring.
June 4, 1878.
Prof. Flower, F.R.S., V.P., in the Chair.
The Secretary read the following report on the additions to the
Society’s Menagerie during the month of May 1878 :—
The total number of registered additions to the Society’s Mena-
gerie during the month of May 1878 was 104, of which 43 were by
birth, 39 by presentation, 8 by purchase, 7 by exchange, and 7 were
received on deposit. The total number of departures during the
same period by death and removals was 120.
. The most noticeable additions during the month of May were as
follows :—
1. Two male Lesser Birds of Paradise (Paradisea minor), pur-
chased of Mr. Leon Laglaize, May 2. On his return from the
Eastern Archipelago in October last, Mr. Laglaize brought with
him four living specimens of this Paradise-bird. All of these passed
the winter safely in the Jardin des Plantes at Paris; and two of them
in full plumage have now been acquired by the Society.
The only other examples of Paradise-bird previously living in the
Society’s Gardens were the two individuals of the same species
brought by Mr. Wallace from Singapore in April 1862’. One of
these lived until 25th December, 1863, the other until the 28th
March, 1864, in the Society’s Gardens.
2. A Copper-head Snake (Cenchris contortrix) presented by Dr.
Frederick Painter, F.Z.S., of South Pittsburg, Tennessee, U.S. A.
This is a well-known venomous snake in the United States; but we
have not previously received living examples of it.
3. A Hairy or Andean Tapir (Tapirus roulini*), obtained in ex-
change from Mr. C. Rice, May 25.
Although we have had in our Gardens examples of all the other
three known species of Tapirs, this is, I believe, the first example
1 See P. Z. 8. 1862, p. 128.
? See, for the reasons for adopting this name, footnote, P. Z. 8. 1870, p. 51.
632 MR. P. L, SCLATER ON MANIS TEMMINCKI. [June 4,
of this rare and little-known animal that has been seen alive in
Europe’.
Mr. Wolf’s sketch (Plate XX XIX.) will give a good idea of the
external appearance of the present specimen, which is a young male,
probably not full-grown, and with the teeth small and imperfectly
developed. The hairs over the whole body are nearly uniformly
thick, but short ; they are barely an inch in length, and of nearly
uniform black colour throughout, with just a shade of brown tinge.
The hairs on the nape of the neck and sides of the body are rather
longer ; those on the sides of the face are brown, and there is a
slight white spot at the corner of the mouth, and some white hairs
at the extremity of the muzzle. There is a slight white external
edging to both ears. There are no white rings round the feet; but
the upper margins of the naked nails have a whitish appearance.
The irides are of a light bluish hazel, quite different from those of
the ordinary Tapirus terrestris, in which they are brown.
The height of the animal at the shoulders is about 26 inches; the
length from the end of the extended snont to the base of the tail
about 54 inches. :
There can be no question, to my mind, that Dr. Gray’s Tapirus
leucogenys (P. Z.S. 1872, p. 483) is merely an individual variety
of the present species. The figure given (J. c. pl. xxi.) is altogether
of too brown a tinge; and in Dr. Gray’s specimen itself, which is
now in the British Museum, the general colour is darker and more
of a greyish black, and, except as regards the white sides of the jaws
and grey face, does not materially differ from that of our specimen.
As regards the white cheeks, upon which Dr. Gray lays so much
stress, it may be remarked that M. Roulin’s “ Tapir pinchaque”
had traces of the same colour’, and there are likewise traces of it in
our living specimen.
The Secretary exhibited a young specimen of Temminck’s Manis
(Manis temmincki), which had been brought from Zanzibar by Mr.
Frederick Holmwood, Assistant Political Agent at Zanzibar, and read
the subjoined extract from a letter of Mr. Holmwood referring to it :—-
“The mother of this little Pangolin came from the coast opposite
Zanzibar, lat. 6° S.; but I have seen what I took to be the same
animal, both in Somali-land under the equator and as far south as
the Makna country opposite Mozambique. They always appeared
to burrow in hard or stony ground; and I saw them always in the
daytime.
“The mother of the specimen I send you lived three months in
Zanzibar. She only fed at night, and remained coiled up in a ball
all day. She regularly retired to the dark corner of my harness-
room at daylight, and left for the garden at sunset. There were
very few ants; but she seemed to get plenty of insects. She bur-
1 Unless it be true, as stated in Mr. Jamrach’s letter to ‘The Times’ of
May 8lst, that an example of the same animal has lately been acquired by the
Zoological Gardens of Antwerp.
2 Vide Aun. des Sci. Nat. xvii. p. 109.
fe) all
1878.} MR. P. L. SCLATER ON CICONIA MAGUARI. 633
rowed at intervals all round the garden-walls ; but this was evidently
only to try and escape, as she never made a hole large enough to
give cover. The day she had the young one she came out during
the day ; but not being quite up to grubbing for insects, she went
into the stable and remained among the horses grubbing in the
dung. After the birth she tried to entice the young Pangolin
to suck (apparently), sitting up like a dog when begging, and coiling
up the moment she got it in her lap. I could not, however, detect
whether she managed to suckle it; indeed I was quite ignorant of
the habits of the animal in its natural state. The first day the
young one had soft scales ; but they hardened the second day, and
it died the same night. The mother wandered about for two days
afterwards, then came into the house and died.”
Mr. Sclater stated that he had, in company with the Superinten-
dent, examined the living examples of Ciconia maguari now in the
Society’s Gardens, and could confirm what Mr. Ridgway had so
clearly described (Bull. U.S. Geol. Surv. vol. iv. p- 250) as to extra-
ordinary mimicry of the true rectrices by the elongated upper
tail-coverts. ‘The same was also the case with Ciconia episcopus, of
which there were also living examples in the Society’s Gardens.
Mr. Ridgway’s generic term Euaenura, however, based on this
peculiar structure, had, it appeared, been anticipated by Dr. Cabanis,
who, in 1850, had proposed the name Dissoura! for this form of
Stork ; so that the two species should stand for the future as Dissura
maguart (Gm.) and D. episcopus (Bodd.).
On behalf of Dr. Elliott Coues, C.M.Z.S., two specimens of
Synaptomys cooperi, Bd., were exhibited by Mr. Edward R. Alston.
This species, Mr. Alston observed, was the type of Synaptomys, pro-
posed in 1857 by Prof. Baird as a subgenus of Myodes*, and accorded
full generic rank by Dr. Coues in 1874%. The present specimens were,
as far as was known, the first typical specimens sent to Europe.
When Mr. Alston wrote his paper “On the Classification of the Order
Glires’’* he was only acquainted with Dr. Coues’s preliminary remarks,
and hesitated at accepting the full generic value of this form ; he was
therefore now desirous of expressing his full recognition of its rank.
Synaptomys was particularly interesting as presenting a connecting
link between the Lemmings and Field-Voles, combining the general
cranial and dental characters of Myodes with the external pecu-
liarities of Arvicola, but differing from both in its grooved upper
incisors. Its range was much less boreal than that of the Lemmings,
extending as far south as Kansas. The specimens exhibited were
destined for the British and Berlin Museums.
* Cf. Reichenow, J. f. Orn. 1877, p- 168. The name is written Dissoura; but
Dissura is more simple and more correct.
* Mamm. N. Amer. pp. xliv, 558.
° P. Ac. Philad. 1874, p. 192. Cf. Mon. N. Am. Rodent. p- 223.
* P. ZS. 1876, pp. 61-98.
Proc. Zoox. Soc.—1878, No. XLI. 41
634 PROF, FLOWER ON A TWO-HORNED RHINOCEROS. [June 4,
Prof. Ray Lankester, M.A., F.R.S., read a memoir on the hearts
of Ceratodus, Protopterus, and Chimera. The structure of the
conus arteriosus and its valves was more particularly described in
this paper. Owen and Hyrtl had shown that the conus of the
Dipnoans differed from that of cartilaginous fishes and Amphibians
in the fact that its walls were devoid of pocket-valves, and presented
instead a long spiral valve and a second short vertical valve. Dr.
Ginther, the only author who had described the heart of Ceratodus,
showed that it possessed in the upper part of the arterial cone
pocket-valves, whilst the spiral valve was shortened so as to be
absent from this upper region. The possession of pocket-valves
served as a very important character to connect the Dipnoans and
the other fishes.
Prof. Lankester now showed that in the lower part also of the arte-
rial cone of Ceratodus there were numerous small pocket-valves, in
addition to those in its upper part ; and further he showed that these
small pocket-valves (so called “ ganoid valves”) were also present
in the lower part of the arterial cone of Protopterus, the African
Mud-fish, which had been generally supposed to be quite devoid of
this kind of valve. The basal fibro-cartilage of the floor of the heart
was described and compared in Ceratodus and Protopterus, and a pos-
sible rudiment of this remarkable structure pointed out in Ceratodus.
This Paper will be published entire, with illustrations, in the
Society’s ‘ Transactions.’
The following papers were read :—
1. On the Skull of a Rhinoceros (R. lasiotis, Scl.?) from
India. By Witt1am Henry Frower, F.R.S., V.P.Z.S.
[Received October 12, 1877.]
Mr. Sclater has put into my hands for examination the skull of a
Rhinoceros, which he had received from Dr. W. D. Stewart, of Cut-
tack, Orissa, being the skull of the two-horned Rhinoceros killed
near Comillah, in Tipperah, as mentioned in P. Z. 8. 1877, p. 269.
Mr. Sclater thinks that the skull may not improbably belong
to the species (at present only known by the living animal in the
Society’s menagerie) which he has named A. Jasiotis.
It is that of a nearly adult animal. All the sutures of the upper
surface of the cranium are consolidated ; and all the permanent teeth
in both jaws are in place except the posterior molars, which are still
concealed in their alveoli.
In size and general conformation it resembles the skull of R.
sumatrensis, and possesses all the essential characters! which distin-
guish that species from R. indicus and R. sondaicus, viz. the sepa-
ration of the postglenoid from the posttympanic processes of the
squamosal below the auditory meatus, the backward position of the
occipital crest (though, perhaps, less marked than usual), and the
* See “On some Oranial and Dental Characters of the existing Species of
Rhinoceros,” P, Z.8, 1876, p. 443,
1878.] PROF. FLOWER ON A TWO-HORNED RHINOCEROS. 635
indication of a second or frontal horn. I have compared it with the
eight skulls assigned to R. sumatrensis at present in London—four
in the British Museum, and four in the Museum of the College of
Surgeons.
These skulls present considerable individual variation in general
conformation, proportional breadth to length, in the development
and form of the nasal bones, number and position of the lachrymal
foramina, form of the posterior margin of the palate, and other
details. The present skull, however, is strikingly different from all
in its superior breadth compared with its length, and especially the
breadth and flatness of the frontal region. The annexed table of
dimensions exhibits the extent of this peculiarity; and, to make it
more manifest, I have added the ratio of the breadth, taken between
the anterior margins of the orbits (where the difference is most cha-
racteristic), compared with the entire length of the skull, the latter
being taken as 100. On looking down this column of the Table, it
will be seen that, though there is a considerable variation among the
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1. Adolescent (all permanent
teeth except milk-molars),
from Tipperah: ...............06. 200 | 12:0 | 68 | 340 | 73 | 80 | 400
2. Aged 9. Malacca, Died in
Zool. Gardens, 1872. B. M...|207 | 11:8 | 60 | 290 | 64 | 65 | 31-4
3. Aged 9. Sumatra. Mus. R.
C.8. No. 2933 .....e0.. cece eee 204 | 111 | 5:2 | 255 | 61 66 | 32:6
4, Aduit.. Pegu. B.M.......... 21:4 | 111 59 | 276 | 66 | 69 | 32-7
5. Adolescent ¢ (all permanent
teeth except last molars). Su-
matra. M.R.C.8. No. 2935 }23:0 | 120 | 60 | 261 | 70 | 76 | 33:0
6. Young (last milk-molar re-
maining). Sumatra. M. R.
BNO 298 In 154-030 casoaades 190 | 106 | 50 | 263 | 65 | 73 | 380
first and second permanent
molars). Sumatra. M.R.C.S.
PO se «ache b gine ddanedan uel 21-:0*?| 10°7 | 52 | 248?|] 64 | 70 | 33:3?
9. Still younger (all milk-molars
and first permanent molars). Teeth nearly
Borneo. B. M...... sc... sees. 181 96 | 50 | 276 | 62 all lost.
* Nasals broken.
4]*
636 PROF. FLOWER ON A TWO-HORNED RHINOCEROS. [June 4,
other skulls, the difference between the narrowest and widest being
almost as great as that between the latter and the present skull, yet
there are intermediate gradations in the former case, whereas the
Tipperah skull is completely isolated from the others. It is curious
that the skull which comes next to it in width is that of the small
dark-coloured animal from Malacca, which died in the Society’s me-
nagerie in 1872, and of which the external characters were certainly
quite unlike those of R. lasiotis.
A second peculiarity by which this skull is distinguished from all
the others, and one to which I am inclined to attach still more im-
portance, is the greater size of the teeth, especially the premolars, both
absolutely and relatively. Thisis also seen in the Table, in which is
given the absolute length of the series of molars and premolars, ex-
cluding the first and last, which are either absent or not developed
in many of the specimens under consideration, and including, there-
fore, the three posterior premolars and the two anterior molars. In
the case of the younger skulls, milk-molars occupy the place of pre-
molars; but this probably does not materially affect the length of
the series. It will be seen that in the present skull this length ab-
solutely exceeds that of any of the others, and, relatively to the
length of the skull (taken as 100), it is 40-0, whereas, of the other
seven, five range between 31°4 and 33°3, and the other two, which
give a higher figure, are both young animals, in which the skull
had not attained its full dimensions, and the teeth therefore appear
relatively larger than they otherwise would have been. It is pro-
bable that sex may affect the size of the teeth, as in the two known
females (Nos. 2 and 3 of the list) they are smaller than in any of
the others. Not only in antero-posterior diameter are the premolars
larger in the present specimen, but still more notably in breadth.
Thus the last premolar in the Pegu adult skull (No. 4) is 18 in
greatest breadth, in the Sumatran female (No. 3) 1°85, in the Ma-
laccan skull (No. 2) 1’*9, in the Sumatran male (No. 5) 2'"05, in the
present skull 2-2. The lower molars exhibit the same superiority
of size; but in other respects the dentition does not differ appreci-
ably from that of the various specimens of R. sumatrensis.
As an individual peculiarity may be noted the single lachrymal
foramen on each side, whereas many of the others have two; but in
both the Malacca and one of the other British-Museum skulls, the
foramen is also single on both sides, and in the Pegu and one of the
Sumatran specimens it is single on one and double on the other side.
Again, the great amount of ossification in the base of the pterygoid
fossa, at the posterior end of the vomer, forming a sort of “ para-
sphenoid”’ mass, is worthy of note; but it is repeated in the Pegu
skull, and partially in the old Sumatran specimen (No. 2), though
not in the equally aged Malacca female.
I have pointed out the peculiarities of this skull, which are in-
teresting in connexion with the fact that it was obtained from a
locality quite beyond the hitherto known range of the Sumatran
Rhinoceros ; but, in the absence of other evidence, will not attempt
to decide whether they should be considered of specific value.
Hanhart .vap
NEW LEPIDOPTERA...
1878. | ON LEPIDOPTERA FROM BILLITON. 637
___ 2. Catalogue of Lepidoptera collected by Mr. S. N. Walter
= in the Island of Billiton. Rhopalocera by F. D. Gop-
MAN and Ospert Satvin; Heterocera by H. Druce.
es [Received May 17, 1878.]
(Plate XL.)
RHOPALOCERA.
That the Rhopalocerous fauna of Billiton should largely partake
of the character of that of the surrounding islands of Sumatra, J ava,
and Borneo, and of the peninsula of Malacca was to be anticipated.
Of the thirty-three species represented in this collection (which was
made in Billiton by Mr. S. N. Walter, and has been sent to us by
Lord Tweeddale for examination), all but one are well-known forms
from one or all of these islands. The single exception, the Myrina,
described below, has, so far as is at present known, its sole repre-
sentative in a nearly allied species from Tenasserim, also described
in the present paper.
The differential elements of the Butterfly faunas of Borneo, Su-
matra, Java, and Malacca have not yet been traced with sufficient
precision to enable us to say to which of the four localities the island
of Billiton has most affinity. With Borneo it is certainly very closely
allied ; but it possesses several species in common with Java, and
others with Malacca. Unfortunately our series of Sumatran Butter-
flies is not very complete; but, so far as we can see, many species
are also found in that island, geographically its nearest neighbour.
Fam. NyMPHALID&.
= Subfam. Danainz.
1. Hestia CLARA.
Hestia clara, Butl. Trans. Ent. Soc. ser. 3, v. p-. 469.
? Hestia leuconoe, Druce, P. Z. S. 1873, Pp. d5/.
A single female specimen, agreeing with Mr. Butler’s types in the
British Museum. The species also occurs in Borneo, whence we have
an example of the same sex. Its occurrence in Java is doubtful.
2. DANAIS PHILOMELA.
Euplea philomela, Zink. Nov. Act. Ac. Nat.-Cur. xv. p. 184,
pl. 16. f. 17; Druce, P. Z.S. 1873, p. 337.
Agrees with Malaccan and Bornean examples.
3. DANAIs SIMILIs.
Papilio similis, Linn, Syst. Nat.
Danais similis, Druce, P. Z. S. 1873, p. 338.
Identical with Malaccan and Javan examples.
4, DANAIs JUVENTA.
Papilio juventa, Cr. Pap. Ex. pl. 188. f. B.
Danais juventa, Druce, P. Z. 8. 1873, p. 337.
Agrees with Bornean specimens.
638 MESSRS. GODMAN, SALVIN, AND DRUCE {June 4,
5. DANAIS HEGISIPPUS.
Papilio hegisippus, Cr. Pap. Ex. t. 180. f. A.
Agrees with Malaccan specimens.
6. DANAIS CHRYSIPPUS.
Papilio chrysippus, Linn. Cr. Pap. Ex. t. 118. f. B, C.
Identical with specimens of this widely dispersed species.
7, EVPL@A MENETRIESIL.
Euplea ménétriésii, Feld. Wien. ent. Mon. iv. p. 398; Druce,
P. Z. 8. 1873, p. 338.
Agrees with specimens thus named in the British Musium.
8. EuPL@A BREMERI.
Euplea bremeri, Feld. Wien. ent. Mon. iv. p. 398; Druce,
P. Z.8. 1873, p. 338.
Found also in Borneo and Malacca.
9. EuPL@A THOOSA.
Trepsichrois thoosa, Hibn. Ex. Schmett. t. viii.
Agrees very well with Hiibner’s figure, being also identical with
Malaccan specimens.
10. EupL@A MULCIBER.
Papilio mulciber, Cram. Pap. Ex. t. 127. f. C, D; Druce, P.Z.8.
1873, p. 338.
Also a Bornean species.
Subfam. ELYMNIINZ.
11. ELYMNIAS NIGRESCENS.
Elymnias nigrescens, Butl. P.Z.S. 1871, p. 520, t. 42. f. 1;
Druce, P. Z.S. 1873, p. 340.
A female specimen agreeing with Bornean examples of the same
sex.
12. ELYMNIAS LAIS.
Papilio lais, Cr. Pap. Ex. t. 110. f. A, B.
Elymnias lais, Druce, P. Z. 8. 1873, p. 340.
Also found in Borneo and Java.
Subfam. Morpuinz.
13. AMATHUSIA PHIDIPPUS.
Papilio phidippus, Linn. Syst. Nat.
Amathusia phidippus, Druce, P. Z.8. 1873, p. 340.
Both sexes of this widely ranging species.
1878.] ON LEPIDOPTERA FROM BILLITON. 639
14. ZEUXIDIA HORSFIELDI.
Zeuwidia horsfieldi, Feld. Voy. Nov. Lep. p. 460, t. 62. f. 4;
Druce, P. Z. 8. 1873, p. 340.
Agrees with Bornean specimens.
Subfam. NyMPHALIN2.
15. CYNTHIA DEIONE.
Cynthia deione, Erichs. Nov. Act. Ac. Nat.-Cur. xvi. Suppl.
G0. t 2,2 a.
Cynthia arsinoe, Druce, P. Z.S. 1873, p- 342 (mec Cramer).
Agrees with this widely-ranging species, of which we have speci-
mens from Java, Malacca, Borneo, and the Philippine Islands.
16. JuNONIA LAOMEDIA.
Papilio laomedia, Linn. Syst. Nat. i. p. 772.
Junonia laomedia, Druce, P.Z. 8. 1873, p. 342.
A widely-distributed species, occurring throughout the Indo-
Malayan subregion.
17, JUNONIA IDA.
Papilio ida, Cr. Pap. Ex. t. 42. f. C, D.
Junonia ida, Druce, P. Z. 8. 1873, p- 342.
Agrees with Javan, Bornean, and Philippine-Islands specimens.
18. NEptis AcERIs.
Papilio aceris, Lepechin, Tageb. d. Reise Russ. Reich. i. p. 203,
(alles Batis
Neptis aceris, Druce, P. Z.8. 1873, p. 344.
A single specimen of this very widely-distributed species.
19. Tanaicta, sp. ?
A single specimen of a species of this genus which we have not
been able to determine.
20. Apottas, sp. ?
Agrees with a Malaccan specimen of an unnamed species in our
collection. Others from the same locality in the British Museum,
have a MS. name of Mr. Butler’s attached to them.
21, SyYMPHZDRA DIRTEA.
Papilio dirtea, Fabr. Ent. Syst. iii. p. 59.
Symphedra dirtea, Butl. P. Z. S. 1868, p. 613; Druce, P. Z.S.
1873, p. 346.
Adolias boisduvali, Bdv. Sp. Gén. t. 8. f. 2.
A male agreeing with Sumatran and Bornean examples.
640 MESSRS. GODMAN, SALVIN, AND DRUCE {June 4,
22. CHARAXES SCHREIBERI. .
Nymphalis schreiber, Godt. Enc. Meth. ix. p. 825.
Charaxes schreiberi, Druce, P. Z.S. 1873, p. 346.
Found also in Java and Borneo.
Fam, Lyc 2NID&.
23. MyRINA NIVEA, sp.n. (Plate XL. figs. 3, 4.)
@. Exp. 2-2inches. Secondaries with projection at the anal angle
and a long tail on the first median branch; outer margin slightly
dentate, pure white ; outer margin of primaries black, blending into a
delicate blue in the apex, the blue colour extending along the costal
margin; a black spot between the median branches of the secon-
daries and a faint black submarginal line on the same wings. Beneath
pure white, both wings cressed with five concentric bands, consisting
of confluent spots, each enclosed in a faint dark line; a submarginal
black line to both wings ; the black spot of the upper surface between
the median branches of the secondaries is mostly blue ; and there are
two blue spots near the margin, one between the submedian nervure
and the first median branch, and one at the anal angle.
Hab. Billiton Island.
Mus. nostr.
This beautiful species, of which there is’ only a single female spe-
cimen in the collection, seems to belong undoubtedly to the genus
Myrina, as defined by Westwood (D. W. & H. Gen. Diurn. Lep. ii.
p- 475). In coloration it is quite distinct from any hitherto de-
scribed member of the genus. We have, however, since received
from Mr. A. O. Hume an example of a closely allied species, which
was taken at Meetan in Burma. This is also a female, and we de-
scribe it as follows :—
Myrina HIEMALIS. (Plate XL. figs. 5, 6.)
Exp. 2°2 inches. In shape exactly like M. nivea, the dark apex of
the primaries more extensive and the inner edge of this dark mark
with two prominent indentations instead of being simply curved ;
beneath, the bands of the wings are better defined, and the dark
marginal lines of each band enclose a greyish space, whereas in M.
nivea the inside of the spots forming the bands is white.
Hab. Meetan, Burma.
Mus. nostr.
Fam. PAPILIONID&.
Subfam. Prerinz.
24. CALLIDRYAS CATILLA.
Papilio catilla, Cr. Pap. Ex. t. 55. f. C, D.
Callidryas catilla, Druce, P. Z. 8. 1873, p. 355.
A widely ranging species found in Borneo and elsewhere.
25. Trertas, sp. ?
A bad specimen.
1878. } ON LEPIDOPTERA FROM BILLITON. 641
26. ‘[eRIAS HECABE.
Papilio hecabe, Linn. Syst. Nat. i. p. 763.
Terias hecabe, Druce, P. Z. 8. 1873, p. 354.
A widely ranging species, also found in Borneo.
Subfam. PaprLioninz.
27. PAPILIO ANTIPHATES.
Papilio antiphates, Cr. Pap. Ex. t. 72. f. A, B; Druce, P. Z. S.
1873, p. 357.
Agrees with Bornean specimens.
28. PAPILIO SARPEDON.
Papilio sarpedon, Linn. Syst. Nat. i. p. 747; Druce, P. Z.S.
1873, p. 559.
A wide-ranging species, found in Borneo and Malacca.
29. PAPILIO EURYPYLUS.
Papilio eurypylus, Linn. Syst. Nat. i. p. 754; Druce, P. Z.8.
1873, p. 357.
Agrees with Bornean examples.
30. PaprLio PAMMON.
Papilio pammon, Linn. Syst. Nat. i. p. 746.
Papilio polytes, L. Druce, P. Z. 8. 1873, p. 357.
The caudal appendage in Billiton specimens is but slightly deve-
loped ; and in this respect they agree best with Javan examples.
31. PApILio IswARA.
Papilio iswara, White, Entom. i. p. 280; Doubl. & Hew. Gen.
D. Lep. t. 3. f. 1.
Agrees with a Malaccan specimen.
32. PAPILIO MEMNON.
Papilio memnon, Linn. Syst. Nat. i. p. 747; Wall. Trans. L. S.
xxv. p. 47, pl. 1. figs. 2, 4.
A female agrees with Javan specimens and with figure 4 of Mr.
Wallace’s plate, and with Cramer’s figure of his P. achates. A
second female resembles figure 2 of Mr. Wallace’s plate, except that
the white band of the secondaries is slightly narrower. This form
is represented by Cramer under the name P. agenor.
Fam. HesPpeRiID2&.
33. CASYAPA THRAX.
Papilio thrax, Linn. Syst. Nat. i. p. 794.
Casyapa thrax, Druce, P. Z. 8. 1873, p. 358.
Agrees with Indian examples.
642 ON LEPIDOPTERA FROM BILLITON. [June 4,
HETEROCERA.
Fam. SPHINGID&.
Subfam. Co #ROCAMPINE.
Cherocampa suffusa, Walk. List Lep. Brit. Mus. Het. pt. viii.
p: 146 (1856).
The type was obtained at Hong-Kong. Mr. Moore tells me he
has specimens from Borneo.
Fam. LiTHosiip&.
HypsiA SILVANDRA, Cr.
Phalena (Bomby:) silvandra, Cram. Pap. Exot. iv. pl. 369. f. D
(1782).
Fam. SATURNIIDE.
ANTHERA BILLITONENSIS, Moore, n. sp.
Male. Upperside pale purplish ferruginous, suffused with grey on
external borders; ocelli oval, defined by a narrow dusky line, blind,
outer half bordered within by a narrow yellow line and a central
transverse line: fore wing with the area within and below the cell
brighter ferruginous; apex yellowish ; an oblique discal, pale ferru-
ginous, narrow lunular band, bordered within by a dusky lunular
line and without by a straight dusky line; a dusky black streak
crossing the middle of the cell, and another from its base to the hind
margin: hind wing with the base brighter ferruginous; a dusky-
bordered, pale yellowish, transverse, discal, sinuous band, the dusky
border encircling the ocellus and terminating on abdominal margin
above end of sinuous band. Body bright ferruginous.
Expanse 47 inches.
Hab. Billiton Island, Malay archipelago. In coll. H. Druce.
Nearest allied to 4. frithii, from Darjiling.
Fam. EREBID.
NyYcTIPAO CREPUSCULARIS.
Phalena (Attacus) crepuscularis, Linn. Syst. Nat. 2811. 13;
Clerck, Icon. pl. 53. f. 1-4; Drury, Ins. i. p. 37, pl. 20. f. 1, 2.
A species frequently sent from Hindostan.
Fam. HyrporyRrip&.
HyPopyra FENISECA.
Hypopyra feniseca, Guén. Noct. iii. 200. 1599 (1852).
Tn the British Museum, from Silhet.
Tribe URANIDES.
Fam. NycTALEMONIDS.
NycTALEMON bocILE, Butler, in litt.
Very closely allied to Nyctalemon hector, White, from Borneo,
1878. ] ON BUTTERFLIES FROM NEW GUINEA. 643
the principal difference being its larger size, broader white bands,
and longer tails to the posterior wing. Mr. Butler’s description has
been in the hands of the Linnean Society for some time.
GEOMETRID&.
Fam. ENNOMID&.
ENNOMOS TESTACEARTA.
Ennomos testacearia, Moore, P. Z. 8. 1867, p. 623.
EXPLANATION OF PLATE Xi,
Figs. 1, 2. Doleschallia comrit, p. 646.
3, 4. Myrina nivea, p. 640,
5, 6.
hiemalis, p. 640.
3. List of the Butterflies collected in Eastern New Guinea
and some neighbouring Islands by Dr. Comrie during
the Voyage of H.M.S. ‘Basilisk. By F. D. Gopman
and OsBert SALvIn.
[Received May 17, 1878.]
(Plate XL.)
The small collection made by Dr. Comrie, of which we here give a
list, was got together under many difficulties and pressure of official
duties. It is therefore not surprising that the number of species it
contains is not greater than it is. We make no apology for pre-
senting this list to the Society, seeing that several of the places
touched at by Captain Moresby’s expedition were absolutely unknown
as to their Butterfly-fauna before Dr. Comrie visited them. Amongst
these we may specially mention the D’Entrecasteaux Islands, which
lie some little way from the eastern shores of New Guinea. The
result of a comparison of the Butterflies of these islands with those
of the New-Guinea mainland shows that, so far as we can see at
present, no difference can be said to exist. In fact these islands are
(as far as their Butterflies are concerned) as essentially a portion of
New Guinea as the Aru Islands are known to be, if not more so.
Family NyMPHALID#.
Subfamily Danarnz.
Danals CITRINA.
Danais citrina, Feld. Voy. Nov. Lep. p. 350, t. 42. f. 5-7.
Danais gloriola, Butl. P. Z.S. 1866, p. 56.
D’Entrecasteaux Islands.
Agrees with specimens from the Aru Islands, whence both Dr.
Felder’s and Mr. Butler’s types were obtained.
644 MESSRS. GODMAN AND SALVIN ON [June 4,
Danals AFFINIS.
Papilio afinis, Fabr. Syst, Ent. p. 511.
Danais affinis, Blanch. Voy. Pole Sud, Zool. iv. p. 389 ; Atl. Lep.
to. fo.
D’Entrecasteaux Islands and mainland of New Guinea.
A widely-ranging species, found in the islands neighbouring New
Guinea and also North Australia, whence it was originally described.
EurL@a CALLITHOE.
Euplea callithoe, Boisd. Voy. Astr. Ent. p. 93; Kirsch, Mitth.
k. z. Mus. z. Dresden, i. p. 115.
D’Entrecasteaux Islands.
A single specimen agrees with one of Dr. Meyer’s examples
obtained in North-western New Guinea.
EvupLa@a IPHIANASSA ?
Euplea iphianassa, Butl. P.Z.S8. 1866, p. 287.
D’ Entrecasteaux Islands and mainland of New Guinea.
These specimens only agree approximately with Mr. Butler’s type,
which came from New Hebrides. The chief difference consists in
the absence in the male of the submarginal row of spots on the
underside of the secondaries.
EvrLa@a nox?
Euplea noz, Butl. P. Z. 8. 1866, p. 278.
D’Entrecasteaux Islands.
An imperfect specimen, probably of this species, the type of which
was obtained in the Aru Islands.
EuPL@A EURIANASSA.
Euplea eurianassa, Hew. Ex. Butt., Hupl. i. f. 2.
D’Entrecasteaux Islands.
Described from a specimen taken on board ship, off the coast of
New Guinea.
EuPL@a TREITSCHKII?
Euplea treitschkii, Boisd. Voy. Astr. Kut. p. 98; Salv. & Godm.
P. Z. 8. 1877, p. 142.
D’Entrecasteaux Islands.
A single female specimen probably belongs to this species, though
it differs somewhat from our specimen from Duke-of-York Island as
regards the arrangement of the spots on the secondary wings. The
specimen is in poor condition.
HAMADRYAS ZOILUS.
Papilio zoilus, Fab. Syst. Ent. p. 480.
D’Entrecasteaux Islands and mainland of New Guinea.
1878. ] BUTTERFLIES FROM NEW GUINEA. 645
Subfamily Saryrinz.
MYCALESIS MEDUS.
Papilio medus, Fabr. Syst. Ent. p. 488.
Mainland of New Guinea.
A single tattered specimen seems to belong to this widely-ranging
species. The white cross bands of the underside are nearly
obsolete.
MYCALESIS DORYCUS.
Satyrus dorycus, Boisd. Voy. Astr. Ent. p. 152.
D’Entrecasteaux Islands and mainland of New Guinea.
Dr. Comrie’s specimens agree with Boisduval’s description in not
having any of the fulvous colour of the secondaries near the anal
angle of the primaries. In this respect they differ from Guérin’s
figure of his S. duponchelit (Voy. Coq. Zool. Atl. Ins. t. 17. f. 3).
The ocelli on both wings are small, and differ much in size from
those of the female figured by Hewitson (Ex. Butt. Myce. t. 2.
£5'7,°10),
Subfamily Moreuinz.
DRUSILLA CATOPS.
Drusilla catops, Westw. Gen. D. Lep. p. 335, note; Trans. Ent.
Soe. ser. 2, iv. p. 181.
D’ Entrecasteaux Islands and mainland of New Guinea.
DRvsILLA MYOPS.
Drusilla myops, Feld. Wien. ent. Monatschr. iii. p. 109, t. 1.
£44
D’Entrecasteaux Islands and mainland of New Guinea.
These specimens agree fairly with Dr. Felder’s figure ; the white
spot on the primaries, however, is more elongated, and the basal
portion of the secondaries much paler.
Subfamily NyMPHALINa.
CETHOSIA CHRYSIPPE.
Papilio chrysippe, Fab. Syst. Fnt. p. 502; Donov. Ins. N. Holl.
24.
D’ Entrecasteaux Islands and mainland of New Guinea.
Agrees with North-Australian and New-Guinea specimens.
RHINOPALPA AMELIA,
Vanessa amelia, Guér. Voy. Coq. Zool. p. 278, t. 14 bis. f. 1.
D’Entrecasteaux Islands and mainland of New Guinea.
Agrees closely with Guérin’s figure. We much doubt whether
R. algina, Boisd., is really distinct. The figure of the latter in the
* Voyage au Pole Sud’ does not seem to be very accurately drawn.
646 MESSRS. GODMAN AND SALVIN ON | June 4,
CYNTHIA ARSINOE.
Papilio arsinoe, Cr. Pap. Ex. t. 160. f. A, B.
Mainland of New Guinea.
A single female specimen.
JUNONIA IPHITA.
Papilio iphita, Cr. Pap. Ex. t. 209. f. C, D.
Mainland of New Guinea.
JUNONIA VELLIDA.
Papilo vellida, Fab. Mant. Ins. ii. p. 35; Donoy. Ins. N. Holl.
6.925) £3:
Mainland of New Guinea.
DoLEsCHALLIA CoMRII, n.sp. (Plate XL. figs. 1, 2.)
Q@. Expanse 4 inches. Outer third of both wings dark blackish
brown, basal portion reddish brown; a subtriangular large white
patch on primaries beyond the cell, its base on the costa, its apex
nearly reaching the inner margin; a black spot at the end of the
cell; between the white patch and the apex are four white spots,
placed transversely in an irregular line, and below them, on either
side of the lower radial, two irregular bluish spots. Beneath reddish
brown ; a straight band crosses the secondaries near the middle and
passes into the primaries, on the inner side margined with white ; and
outside it (running parallel) is a dark brown band ; the white spots
show as on the upperside of the primaries; and on the secondaries
are two ocelli—one between the first and second median branches,
the other between the middle and upper radials.
Mainland of New Guinea (Dr. Comrie).
The sexes in the members of this genus do not differ to a great
extent; so that we may expect the male of this species to be some-
what similar to the female specimen now described. There is no
described member of the genus which the present species is at all
like. This, we trust, will be sufficiently apparent from the figure
on the Plate accompanying this paper. ‘There is only a single spe-
cimen in Dr. Comrie’s collection.
CYRESTES ACILIA.
Nymphalis acilia, Godt. Eneye. Méth. ix. p. 478.
Mainland of New Guinea.
Agrees with other New-Guinea specimens, and with others from
Waigiou.
ATHYMA VENILIA.
Papilio venilia, Linn. Syst. Nat. i. p. 780.
Mainland of New Guinea.
Agrees with Amboina and New-Guinea specimens.
1878. ] BUTTERFLIES FROM NEW GUINEA. 647
ATHYMA LACTARIA.
Athyma lactaria, Butl. Ann. & Mag. N. H. ser. 3, xvii. p. 98.
Mainland of New Guinea.
Agrees with the specimens in the British Museum.
DIADEMA NERINA.
Papilio nerina, Fab. Syst. Ent. p. 509; Donov. Ins. N. Holl.
t. 27. f. 1.
D’Entrecasteaux Islands.
A female specimen, agreeing with others from Australia.
DIADEMA ALIMENA.
Papilio alimena, Linn. Syst. Nat. i. p. 780.
D’ Entrecasteaux Islands.
Agrees with Amboina specimens.
Family PAPILIONIDS.
Subfamily Prerinz.
TERIAS HECABE.
Papilio hecabe, Linn. Syst. Nat. i. p. 763.
Mainland of New Guinea.
Subfamily PariLionIn&.
ORNITHOPTERA POSEIDON.
Papilio poseidon, Doubl. Ann. & Mag. N. H. xix. p. 173; Wall.
Trans. L. 8. xxv. p. 35.
Possession Bay and D’Entrecasteaux Islands.
Dr. Comrie obtained a number of specimens of Ornithoptera
(mostly females) in these two localities. Unfortunately most of
them are in broken condition ; but enough remains to show an in-
teresting range of variation in the females—enough, indeed, still
further to discredit the many species of the O.-priamus group that
have been attempted to be set up at various times.
Some of the females from Possession Bay have the primaries
spotless brown; others show a trace of the light-coloured spot near
the end of the cell. The specimens from D’Entrecasteaux Islands
are strongly marked, indeed as much so as female examples of
O. arruana.
Unfortunately, Dr. Comrie’s collection contains only one male
(from Possession Bay) in a condition to compare with examples from
other localities. This seems to agree closely with typical specimens
of O. arruana, the apical angle of the secondaries being perhaps
rather less acute; but the difference is very slight. Comparing this
with Montrouzier’s description of his O. boisduvalii, from Woodlark
Island (no great distance from Dr. Comrie’s locality), we find it
agree in every respect.
So far, then, as we can see, these specimens tend to confirm the
648 MR. A. G. BUTLER ON A NEW PHYLLOPTERA. [June 4,
accuracy of Mr. Wallace’s views on these Ornithopteree, expressed
in his paper on the Papilionida of the Malayan region.
PAPILio ERECHTHEUS.
Papilio erechtheus, Donov. Ins. N. Holl. t. xiv., xv.
Mainland of New Guinea.
A female, agreeing with specimens from Moreton Bay, Australia.
PAPILIO EUCHENOR.
Papilio euchenor, Guér. Voy. Coq. Zool. p. 273, t. 13. f. 3.
Mainland of New Guinea.
PaPILio POLYDORUS.
Papilio polydorus, Linn. Syst. Nat. i. p. 746; Wall. Trans. | FR
xxv. p. 42.
Mainland of New Guinea.
These specimens have a light spot on the primaries; and the
secondaries are more elongated than in P. leodamas, Wall., the
species we should have expected to find here. They do not differ
from the ordinary type of P. polydorus.
4. Description of a new Species of the Orthopterous Genus
Phylloptera from Madagascar. By Artuur G. Butter,
F.L.S., F.Z.8., &c.
[Received May 23, 1878.]
The genus Phylloptera is one of the numerous interesting groups
amongst the Orthoptera which exhibit a striking resemblance to
leaves, some of them being imitated in a green condition, and others
when faded.
The species now before me is an admirable example of this
adaptation to surrounding conditions, the tegmina being quite like
sear and yellow leaves in which the brown patches of decay have
begun to appear.
PHYLLOPTERA SEGONOIDES, 0. sp.
Tegmina of the normal elliptical form; horn yellow, with the
veins and borders more or less tinted with burnt sienna; the whole
of the areoles, excepting those along the anterior border, with central
irregular dull greyish-brown spots or patches ; under surface paler,
with greenish borders; wings hyaline-white, veins yellowish, the
normal black-dotted horn-yellow cuneiform patch bounded by an
oblique brown line at the very acuminate apex ; frons testaceous ;
eyes (in the dead example) chocolate-brown ; antennze black, with
testaceous scape; back of head sordid whitish, with two divergent
grey lines from behind the antenne; pronotum black, whitish in
front, with the sides, posterior third, and a V-shaped central
1878. ] MR. A. G. BUTLER ON A NEW PHYLLOPTERA. 649
marking pale castaneous; meso- and metathorax glassy greyish,
with two or three opaque whitish spots; abdomen sordid testaceous,
sides of the venter whitish ; pectus and posterior coxze whitish ;
anterior femora with the upper or iaterior surface castaneous at the
proximal end, otherwise greyish, spotted with cream-colour, with
two small tuberculose castaneous dots and a short longitudinal black
dash, three black denticles along the supero-interior inargin, inferior
or exterior surface deeply grooved, paler, the distal half sordid
whitish, with an irregular longitudinal brown dash; tibize with the
proximal half whitish, expanded, compressed, and marked on each
side by two deep pyriform impressed spots, distal half cylindrical,
castaneous in the centre and black at the extremities, interior
margin with three fine spinules; tarsi black, the terminal joint
A
PEs
SS
Lk SN
mis
WRN =
Phylloptera segonoides.
castaneous, with black claws: middle pair of legs longer than the
anterior pair, the tibiz yellowish, banded with black across the
centre and at each end, without the impressed markings of the
anterior pair: posterior legs with the femora compressed, tawny,
with an external longitudinal blackish lateral streak, inferior margin
deeply grooved, its exterior edge with four short black denticles;
tibiz above flattened, slaty grey, with the distal end and a lateral
series of triangular spots cream-coloured, twelve black denticles
along the inner and eight along the outer margin, otherwise casta-
neous, subcylindrical, with about eleven delicate little spmules along
the inferior margin ; tarsi as in the other legs. Length of the body
23 millims., of the tegmina 91 millims., of the wings 96 millims.
This species much resembles the New-World genus Segona, but
differs markedly in the structure of the anterior pair of legs.
Proc. Zoou. Soc.—1878, No. XLII. 42
650 MESSRS. SCLATER AND SALVIN ON BIRDS [June 4
5. Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘Challenger’”—No. XI. On the
Steganopodes and Impennes. By P. L. Scuater, M.A.,
Ph.D., F.R.S, and Ossert Satvin, M.A., F.R.S.
[Received May 24, 1878.]
a. STEGANOPODES.
Of this order the collection contains 33 specimens belonging to 9
species, as follows :—
1. FREGATID.
1. Freeata aquiza (Linn.).
a, adult, }
6, young, j Ascension Island (March 1876).
c, young,
2. Freeata MINOR (Gmn.).
Attagen ariel, Gould, B. Austr. vii. pl. 72.
133, male, Raine Island.
“Legs and feet black; bill grey; skin of throat red ; eyes red.
The skin of the throat is of a lighter red in the male than in the
female. The stomach contained remains of cuttlefish, Spirula, and
a fish.’—J. MW.
134, female,
135, young, female, Raine Island.
136, nestling, female,
“Feet red; eyes red. The biil and fect of the young birds are
of a white colour with a shade of blue; eyes black.
‘‘In the adult birds the male has the eyelids, feet, and eyes
black. The female has these parts red. This holds good in all our
specimens.” —J. M.
144, female, Raine Island.
482, male, Admiralty Island.
«Eyes black; flesh of throat red. Stomach contained fish. I
shot this bird from the pinnace while with several others. It was
hovering over a shoal of fish. A Noddy and Black-headed Terns,
and a Puffin or Petrel, larger than a Cape-Pigeon, were also fishing.”
—ZJ. M.
The series before us. so far as it goes, tends to show that this
smaller Frigate-bird may remain distinct from its larger brother,
although we are not aware of any other obvious point of difference
than that of size.
The specimens measure as follows :—
Wing. Bill from gape.
PEG ANG AGUIAG 7225 eA. ee ZOD 30
ad Rt Yes te sake Cee 4°5
ae Be wllesragreins 2% vis, Sel) 4-4
1878.] COLLECTED BY THE ‘CHALLENGER.’ 651
Wing. Bill from gape.
Fregata minor 3,133.... ... 20°53 3°9
» sei ID 5 ES elo tialie 9205 4:0
» at, Py tees 20°5 4:0
99 Fa LO aeeO 2 oieo isa -F 20°5 4:0
The bill of F. minor is also narrower at the base, and more feeble.
2. PHAETHONTID.
3. PHaiTHon #THEREUs, Linn.
a, Ascension Island (1876, March).
143, female (young), Raine Island.
“Eyes black, legs white, claws black, bill dark brown colour.
The only one seen on the island.”—/J. M.
4. Paatraon riavrrostris, Brandt.
a,
,
ZL ae } Ascension (March 1876).
e, female, Tongatabou.
507, female, at sea, 19th March, 1875.
“Eyes black, legs lake-white. The claws and membrane between
the toes quite black. Bill pale yellow. Came on board the ship on
the morning of the 19th March, 1875, before daylight. The bill of
a cuttlefish was in the stomach.”—J. M.
} Fernando Noronha (Sept. 1873).
3. PELECANID.
5. SuLA LeucoGastra (Bodd.).
Pelecanus sula, Linn. S. N. i. p. 218.
Le Fou de Cayenne, Buff. Pl. Enl. 973, undé
Pelecanus leucogaster, Bodd. Tabl. de Pl. Enl. p. 57.
Sula leucogastra, Salvin, Trans. Zool. Soc. ix. p- 496.
141, male, Raine Island.
“ Hyes grey ; feet light green ; bill bluish towards the base, white
at the tip.’”—J. M.
510, female, at sea.
“Eyes white or light grey. Stomach had cuttlefish; feet yellow
with green tinge; bill flesh-colour, cere greenish. Came on board
the ship on the evening of the 6th April, 1875.”—J. M.
a, female. Ascension (March 1876).
4 } St. Paul’s rocks, Atlantic (August 1873).
3
6. Suxa PiscaTor (Linn.).
Pelecanus piscator, Linn. 8. N. i. p. 217.
Dysporus piscator, Finsch et Hart!. Orn. Centr.-Polyn. p. 255.
123, female, off Cape York, at sea.
“« Eyes brown ; bill and throat light blue-reddish towards the base,
42*
652 MESSRS. SCLATER AND SALVIN ON BIRDS [June 4,
and the tips of both mandibles brown ; feet coral-red. The stomach
contained cuttlefish about 33 and 4 inches long, allied to Gonatus
ameenus, and the caudal portion of a fish. There were a great many
lice on this bird (preserved in a tube).”—J. M.
139, female, Raine Island.
«Byes hazel; feet red; bill light blue, red towards the base.
Stomach contained fish and cuttlefish.’—J. M.
7. SuLA CYANops, Sund.
Sula cyanops, Sund. Phys. Sallsk. Tidsekr, 1837, p. 218.
Dysporus cyanops, Finsch et Hartl. Orn. Centr.-Pol. p. 252.
137 and 138, females, Raine Island.
«Byes yellow, skin of the throat black, legs and feet slate-colour.
Stomach contained fish and cuttlefish.’—J. M.
8. PHALACROCORAX IMPERIALIS.
Phalacrocorax imperialis, King, P. Z. 8. 1831, p. 20.
Phalacrocoraz carunculatus, Scl. et Salv. Ibis, 1870, p. 500.
637, male, Core Harbour, Messier Channel.
*« Byes brown. Stomach had shells.’—J. M.
This Cormorant appears to be different from the P. carunculatus
of New Zealand, with which we have hitherto united it, having a
broad white patch on the middle of the back in the adult plumage,
no crest, and the white extending further over the cheeks. Both
species have the white bar on the upper wing-coverts.
The species of the Falkland Islands which has hitherto been called
Ph. carunculatus (Sel. P. Z.S. 1860, p. 391; Abbott, ‘Ibis,’ 1861,
p- 166) is again different, having a recurved crest and the caruncles
on the front largely developed. It should probably stand as P. al-
biventris, Lesson’s Carbo albiventer (Trait. d’Orn. p. 604) being
apparently based upon the young of this species.
9. PHALACROCORAX VERRUCOSUS.
Haheus (Hypoleucus) verrucosus, Cab. Journ. f. O. 1875, p. 450.
Halieus verrucosus, Cab. et Reich. Journ. f. O. 1876, p. 359,
tab. 1.
Phalacrocorax verrucosus, Sharpe, Zool. of Kerguelen, p. 49.
a, male, )
6, male,
] ae oung), Betsy Cove, Kerguelen Land, January 1874.
e, female,
J, sex ind.,
The series of this Shag is quite sufficient to warrant us in adhering
to the species as distinct. The principal characters are clearly pointed
out by Dr. Cabanis in his original description ; and a good figure is
given of the adult male under the second reference given above. Not
one of the six specimens, of which a, 6, and, apparently, / are adult,
shows any traces of the white line along the upper wing-coverts
QS
1878. ] COLLECTED BY THE ‘CHALLENGER.’ 653
or of the white spot in the middle of the back which distinguish
P. imperialis. The female, d, is in young plumage.
6. IMPENNES.
Of Penguins the collection contains 37 specimens, belonging to 6
species.
1. APTENODYTES LONGIROSTRIS.
Apterodyta longirostris, Scop. Del. Faun. et Flor. Ins. ii. p- 91.
Aptenodytes longirostris, Coues, Pr. Ac. Phil. 1872, p- 193;
Sharpe, Zool. Kerg. p. 52.
a, Kerguelen Land.
6, Christmas Harbour, Kerguelen.
ec, Marion Island.
Zoe, female, |) pik lands.
735, male,
“Eyes brown; the stomach of the female had the remains of
cuttlefish, there being many beaks of these and also a weed-like sub-
stance. Female weighed 20 Ib. ; skin of male weighed 10 lb.”—
J. M.
2. PYGoscELEs THNIATUS.
Aptenodytes papua, Forst. Nov. Comm. Gott. iii. p. 140, tab. iii.
Aptenodytes teniata, Peale, U.S. Expl. Exp. p. 264 (1848).
Pygosceles wagleri, Scl. P. Z.S. 1860, p. 390.
Pygosceles teniata, Coues, Pr. Ac. Phil. 1872, p- 195; Sharpe,
Zool. Kerg. p. 54.
a, \
6,
* Kerguelen Land.
|
f)
3. SPHENISCUS DEMERSUS.
Diomeda demersa, Linn. S. N. i. p. 214.
Spheniscus demersus, Temm. Tabl. Méth. p. 107 (1836).
a, |
I
“4 ads of Good Hope.
ad; )
4, SPHENISCUS MAGELLANICUS.
Aptenodytes magellanicus, Forst. Nov. Comm. Gott. iii. p. 143,
tab. v.
Spheniscus magellanicus, Scl. P. Z. S. 1860, p. 382.
Spheniscus demersus, var. magellanicus, Coues, Pr. Ac. Phil. 1872,
p- 209.
689, male, Port Churrucha.
654 ON BIRDS COLLECTED BY THE ‘CHALLENGER. | June 4,
“<The only one seen. Eyes hazel; the stomach had many por-
tions of fish, some of considerable size ; upper part of the feet white,
bill black or slate-coloured.”—J. M
741, female, Falklands.
«Eyes brown; stomach had fish.” —J. M.
In our opinion this species is quite distinct from S. demersus of
the Cape, having a double band on the chest, as is well represented
in Forster’s figure.
5. EupYPTES CHRYSOLOPHUS.
Aptenodytes chrysocome, Forst. N. Comm. Gott. ili. p. 135.
Catarractes chrysolophus, Brandt, Bull. Ac. Pét. i. p. 315.
Eudyptes chrysolophus, Scl. Ibis, 1860, pp. 338, 432, et P. Z.S.
1860, p. 390; Abbott, Ibis, 1861, p. 163; Sharpe, Zool. Kerg. p. 57.
Eudyptes diadematus, Gould, P. Z.S. 1869, p. 419.
a—c, males,
d, temale, | Christmas Harbour, Kerguelen.
e, pale var.,
On comparison of the Kerguelen specimen with others from the
Falklands we find no reason for considering them otherwise than of the
same species. There is, however, less appearance of the white upper
tail-coverts in the Falklands specimens. Why Mr. Sharpe should
have supposed that Z. chrysolophus (Sclater et Abbott) of the Falk-
lands was his 2. saltator we cannot understand.
Forster evidently had both the ‘‘ Rock-hopper”’ and ‘* Macaroni”
Penguins under his eyes when he described his Apt. chrysocome’.
Brandt first clearly separated the two species, which are quite dis-
tiret and easily recognized by the character which Brandt gives.
The type specimen of EH. diadematus, Gould, for which we have
made every inquiry, is unfortunately no longer to be found. Mr.
Gould has parted with it, he knows not whither. It was probably
only an individual variety cf this species.
§. EupyPTES CHRYSOCOME.
Aptenodytes chrysocome, Forst. N. Comm. Gott. iil. p. 135 (partim).
Catarractes chrysocome, Brandt, Bull. Ac. Pét. ii. p. 314.
Eudyptes chrysocome, Scl. P.Z.8. 1860, p. 390.
Eudyptes nigrivestris, Gould, P. Z.S. 1860, p. 418.
Eudyptes saltator, Sharpe, Zool. Kerg. p. 60.
a—e, Inaccessible Island.
J, male, | ee
ye } Kerguelen.
4, m, males, } Falklands.
n, young,
On comparison of the series from these three localities we cannot
satisfactorily recognize more than one species of ‘‘ Rock-hopper.”
The bird trom Inaccessible Island has the elongated supereiliary .
) He deseribes {le erest “in aliis indiyiduis in fronte unita, in aliis divisa ”
(7.s.c. p. 137).
1878.) MR. P. L. SCLATER ON FELIS LANEA. 655
plumes more produced ; those of the Falklands and Kerguelen have
them rather shorter.
As regards the Australian and New-Zealand bird (2. pachyrhyn-
chus of Gray), to which Mr. Sharpe proposes to restrict the name
chrysocome ot Forster, we likewise doubt its distinctness. The only
differential characters given by Mr. Sharpe consist in the relative
lengths of the black and yellow feathers of the superciliary tufts. At
the same time we should like to examine a series of this form before
pronouncing a decided opinion on the point.
June 18th, 1878.
Arthur Grote, Esq., F.Z.S., V.P., in the Chair.
Extracts were read from a letter addressed to the Secretary by Mr.
E. L. Layard, F.Z.S8., dated British Consulate, Noumea, March
30th, 1878.
Mr. Layard pointed out that Mr. Gould’s Glycyphila fasciata
(Handb. B. Austr. i. p. 499; B. Austr. iv. tab. 30) was quite a dif-
ferent bird from G. fasciata (Forster) of New Caledonia, the latter
being nearly half as large again. Mr. Layard proposed to change
the name of the Australian species to G. gouldi.
Mr. Layard also sent the subjoined note on Petreca kleinschmidti
of Finsch :—
*‘Dr. Finsch has described as new, under the name of Petroica
kleinschmidti, the little Petroica of Fiji (P. Z. 8. 1875, p. 643). I
wish to point out that this species will not stand, and that the Fijian
species is not separable from the Samoan P. pusilla, Peale. Ur.
Finsch says, ‘it differs from P. pusilla, Peale, from the Navigators, in
lacking the white front and the large white mark on the wing-coverts.’
Dr. Finsch’s description is evidently taken from a female; and she
lacks the white marks, which, however, are very visible in the male.
““T have before me two pairs of Fijian birds (¢ and 9), and
three males from Samoa, Unfortunately one of these, from Mr.
Whitmee, is unsexed ; the other two are of my own killing: one is
a young male, and I have no doubt Mr. Whitmee’s bird is also
a male; none are in very good plumage, whereas my Fiian birds
are in splendid order.
“* Now, I can affirm that the males are not to be distinguished
one from the other, except that the Fijians are in brighter plumage ;
so that P. kleinschmidti must sink into a synonym of P. pusilla, with
Peale’s description of which (U.S. Expl. Exped. p. 165) it entirely
agrees.”
Mr. Sclater called the attention of the members present to the
unique specimen of his Felis lanea (P. Z. S. 1877, p. 532), still
living in the Society’s menagerie, and read the subjoined extract
from a letter of Mr. E. L. Layard, F.Z.S., relating to this animal :—
** Tt will interest you to know that there is a second specimen of
656 MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. [June 18,
your Felis lanea in the South-African museum, sent from the same
place (the Beaufort-West Karras) by the late Arthur V. Jackson,
who killed it himself. Unfortunately I received the skin in very
bad condition. The ground-colour is much paler than in your plate,
almost white.
“Jackson and I thought it an albinism (or rather erythrism)
of F. jubata (see Catalogue of S. A. Museum, p. 38, No. 82, Gue-
parda jubata, specimen 6). At p. 39 of the same Catalogue, I
remark that we have had notices of a second species of Maned Leo-
pard, with solid spots and with retractile claws, from Natal. The
claws of your animal are not shown in Smit’s plate. What is
their structure ?”’
On this last point Mr. Sclater stated that, so far as could be told
from examination of the living animals, the claws of Felis lanea
resembled those of Felis jubata, being observable when the feet
were at rest, and being but slightly extensile.
The existence of a second specimen of the animal in the South-
African Museum (of which Mr. R. Trimen had also informed Mr.
Sclater) was a fact of great interest.
Mr. Sclater read some Supplementary Notes on the Curassows
(Cracidze), mainly based on specimens which had been received by
the Society alive since Mr. Sclater’s previous memoir on this subject
had been read five years ago.
This paper will be published in the Society’s ‘ Transactions.’
The following papers were read :—
1. On the Squirrels of the Neotropical Region.
By Epwarp R. Atston, F.L.S., F.Z.8., &e.
[Received June 3, 1878. ]
(Plate XLI.)
No better example of a polymorphic genus can be found than the
almost cosmopolitan Sciurus. Even our commen European Squirrel
assumes such phases of coloration in the north, in the east, and
among the Alps that the extremes would undoubtedly be considered
perfectly distinct species if the intermediate links were not known.
The same variability is found to a still greater extent in many of
the Oriental species; while the polymorphism of some of the North-
American forms was pointed out by Professor Baird more than
twenty years ago.’
It is only lately that similar critical attention has been given
to the Squirrels of the Neotropical Region. Ofthese no fewer than
Jjifty-nine nominal species have been described by various writers.
The late Dr. Gray, in 1867, published a ‘‘ Synopsis of the Species
? Mamm. North Amer. pp. 244, 245.
i
Smit lith
1878.] MR. E. R. ALSTON ON NEOTROPICAL SQUIRRELS. 657
of American Squirrels in the Collection of the British Museum”’’,
in which he recognized twenty-nine Neotropical species as being
represented in the national cabinet, nineteen of them being described
as new. Last year appeared Messrs. Coues and Allen’s exhaustive
volume of ‘ Monographs of North-American Rodentia’ *, in which
the Squirrels are discussed by the last-named zoologist, who includes
the South-American species in his scope. Having had access to
considerably larger series of specimens than were available to most
previous writers, Mr. Allen was led to reduce the number of valid
species of Neotropical Squirrels to ten, with two “ subspecies,”
namely :—
Sciurus carolinensis. Sciurus leucops.
, var. yucatanensis. e@stuans.
colliei. -—— » var. rufo-niger.
aureigaster. tephrogaster.
——- boothie. gerrardi.
—— hypopyrrhus. —— variabilis.
He further remarked that he thought that “on the whole the
number of species will, by future investigations, be further reduced
rather than increased,” and that he had erred in recognizing too
many species rather than tou few.
No one who is acquainted with Mr. J. A. Allen’s recent memoirs
on the geographical variation of species, needs to be reminded of
his breadth of view and extreme carefulness in comparison; and in
the present monograph these qualities are as conspicuous as ever.
In it Mr. Allen has done a great service in reducing the complicated
synonymy of the South-American Squirrels to some order ; and if
some of his identifications prove to be erroneous, the fault lies in
the often totally insufficient or even misleading descriptions of
some previous writers.
Within the last year I have been able to examine in the British
Museum and the Museums of Berlin and Paris, the types of no less
than forty-one nominal species of Neotropical Sciuri. In these
collections I have also been able to compare much more extensive
series of specimens than even Mr. Allen had access to; and, through
his kindness, I have examined typical examples of the species re-
eognized by him. This study has led me to accept many of Mr.
Allen’s identifications (some of which are sufficiently startling at
first sight), and in some instances to carry the reduction of species
still further ; but it has also enabled me to correct a few errors in
his synonymy, and to point out a few apparently valid species
with which he was not acquainted.
Particularly rich in this group are the Paris and British Mu-
seums; and the study of their long suites of specimens leads one
irresistibly to conclusions which must appear strange to those who
only know the extreme links of the chain. Among other things they
1 Ann. & Mag. Nat. Hist. 3rd ser. xx. pp. 415-434.
> Rep, U.S. Geol. Suryey of Territories, vol. xi.
658 MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. [June 18,
have convinced me that Mr. Allen has laid too much stress on the
comparative size of the ears, and length and bushiness of the tail, as
distinctive characters. In both these points, as well as in the quality
of the pelage, every intermediate stage is often to be found; and I
have therefore been obliged to unite Mr. Allen’s S. awreigaster
and S. leucops, his S. boothie and S. hypopyrrhus, and his 8.
gerrardi and S. variabilis. On the other hand, I have felt obliged
to recognize, at least provisionally, the specific rank of 8. stramineus,
S'. griseogenys, S. rufo-niger, and S. pusil/us, and more doubtfully
that of S. griseoflavus,—thus raising the number of species from
ten to twelve.
In facing the intricate and often baffling problem of distinguish-
ing between ‘species’ and “ varieties”’ in such a protean group, I
have endeavoured to act in harmony with Mr. Darwin’s directions :
“When a naturalist can unite by means of intermediate links any
two forms, he treats the one as a variety of the other’, It must
be remembered that many of these “ varieties” apparently breed
true and prevail in certain parts of the range; but all that are here
brought together are united by such intergradations that a sufhi-
cient series at once convinces one of their identity. It is evident,
however, that still more complete material will be required before
every point can be regarded as definitely settled.
Before proceeding to remark on the various species*, I must
return my warmest thanks to all who have assisted me in this in-
vestigation, especially to Mr. J. A. Allen, Professor A. Milne-
Edwards, Dr. Peters, and Dr. Giinther.
I. Scturus CAROLINENSIS.
Sciurus carolinensis, Gmelin, Linn. Syst. Nat. i. p. 148 (1788,
ex Pennant); Allen, Mon. N.-Am. Rodent, p. 700.
Length of head and body averaging 10 inches ; of tail-vertebree
about 8 inches,
Upper parts grey or brownish grey, the middle of the back
brownish, a rufous lateral stripe usually present ; lower parts white.
Tail moderate, washed with white, the hairs yellowish brown or
whitish, with narrow black rings, broader black subterminal bands
and white tips.
Hab. Southern Canada; Eastern United States; Mexico; Yu-
catan ; Guatemala (7).
Concerning this well-known Nearctic form I can add almost
nothing to Mr. Allen’s exhaustive account. He distinguishes three
geographical races or varieties, of which two, his “ var. caro-
linensis”’ and “var. yucatanensis,” are found within the Neotropi-
eal limits. The former, smaller and browner than the northern
leucotis race, is apparently the S. carolinensis which M. de Saus-
sure records from Mexico * and Mr. Tomes from Guatemala’; and
Origin of Species, 6th ed. p. 36.
° The species of which I have been able to examine the types or typical
species are distinguished in the following pages by an asterisk.
3 Rey. et: Mag. de Zool. 1861, p. 4. * Proce. Zool. Soc, 1861, p. 282.
1878.] MR.E.R ALSTON ON NEOTROPICAL SQUIRRELS. 659
there are specimens from Nuevo Leon in the National Museum at
Washington. Mr, Allen’s “var. yucatanensis”’ is based on speci-
mens collected at Merida, which agree in their pale coloration, the
absence of rufous on their flanks, their small size, and the compara-
tive coarseness of their pelage. He considers that this form is
‘possibly specifically distinct,’? but remarks that the Nuevo-Leon
examples are fairly intermediate; and, judging from the single
typical example which Mr. Allen kindly lent me, I think he was
wise in refusing it specific rank, at least for the present.
Of the geographical variations presented by the Grey Squirrel
throughout its very extensive range, Mr. Allen observes they consist
“mainly in decrease in size southward, and an increase in in-
tensity of coloration, the whitish tips of the hairs of the dorsal
surface seen in northern specimens being replaced by yellowish-
brown in the southern ones, except in the case of var. yucata-
nensis.” The decrease in size as we go southwards is of course
strongly in favour of the distinctness of the next species.
II. Scrurvus ARIZONENSIS.
*Sciurus arizonensis, Coues, Amer. Nat. i. p. 357 (1867).
(Sciurus colliei, Allen, Mon. N.-Am. Rodent, p. 7388, nec
Richardson.)
Length about 12°50, of tail-vertebree 11 inches. Upper parts
grey, more or less mixed with yellowish-brown, especially along
the middle of the back; lower parts pure white. Tail long, very
bushy, strongly washed with white, the hairs tawny or fulvous with
two narrow and one broader black band, succeeded by a long white
tip.
PTab. Arizona; California; Mexico.
Misled by imperfect descriptions and a bad figure of Richardson’s
type, Mr. Allen has referred the Arizona Squirrel of Dr. Coues to
Richardson’s 8S. co/liei. He has since kindly intrusted me with a
typical example of S. arizon-nsis ;.and I find that it is quite distinct
from S. colliei (which is Mr. Allen’s S. doothie), being much more
nearly allied to S. carolinensis, from which, however, both Dr. Coues
and Mr. Allen consider that it is ‘thoroughly distinct.” The
superior size, coloration of the long heavy tail, and absence of a lateral
line appear to be constant, but in some specimens in the Paris and
British Museums the grey of the upper flanks is not so clear from
rufous as in the examples described by Mr. Allen. The example
mentioned by that gentleman as having “the lower surface consi-
derably varied with irregular patches and streaks of pale yellowish
rufous’ gives support to the opinion that furthur material will
render it necessary to unite the next species with the present. If
so, Dr. Coues’s name must be retained, as it has priority over Gray’s
title.
660 MR. £E.R.ALSTON ON NEOTROPICAL SQUIRRELS. [June 18,
III. Scrurus GRISEOFLAVUS.
*Macrozus griseoflavus, Gray, Ann. & Mag. Nat. Hist. 3rd series,
xx. p. 427 (1867).
(Sciurus leucops, Allen, Mon. N.-Am. Rodent, p. 753, nec
Gray.)
Length (in skin) about 13°50 inches, of tail-vertebrze 10°25 inches.
Upper parts nearly uniform yellowish grey, the narrow black rings
and minute whitish tips of the hairs merely obscuring the general
tint produced by their broad pale-brown median bands ; lower parts
yellowish red, the hairs in one specimen obscurely ringed with black.
Tail bushy, strongly washed with white; the hairs yellowish or pale
brown, with two or three narrow and one broader black band suc-
ceeded by a long white tip.
Hab. Mexico (?) ; Guatemala.
Mr. Allen considers Gray’s M. griseoflavus to be specifically
identical with his M. leucops (op. cit. p. 753); and the original
diagnosis certainly seems to give countenance to such a view. The
typical specimens (five in number), however, are very different, and
in my opinion are closely allied to the last species, of which I
suspect it will eventually prove to be a southern race. More speci-
mens, however, are required before they can be united; and provi-
sionally I therefore accept S. griseoflavus as a distinct species.
It” appears probable that this is Mr. Tomes’s S. ludovicianus
from Guatemala’; and a Mexican specimen in the Copenhagen
Museum, labelled “ Se. afinis, an sp. nov.? Reinh.,” does not
appear to be separable.
IV. Scrurvus VARIEGATUS.
Sciurus variegatus, Erxleben, Syst. Reg. An. p. 421 (1777, ex
Hernandez).
*Sciurus aureogaster, F. Cuvier, Hist. des Mamm. iii., livr. lix.
(1829).
( Sciurus leucogaster, F. Cuvier, Suppl. de Buff. i. Mamm. p. 300
1831).
Sciurus albipes, Wagner, Abh. Bayer. Ak. ii. p. 101 (1837).
Sciurus ferruginiventris, Audubon & Bachman, P. Ac. Philad.
1841, p. 101.
Seiurus varius, Wagner, Suppl. Schreber Siugeth. iii. p. 168
(1843).
Sciurus socialis, Wagner, tom. cit. p. 170.
* Macroxus morio, Gray, Ann. & Mag. Nat. Hist. 3rd series, xx.
p- 424 (1867, nec Wagner).
* Macroxus maurus, Gray, tom. cit. p. 425.
* Macroxus leucops, Gray, tom. cit. p. 427.
(Sciurus aureigaster et S. leucops, Allen, Mon. N.-Am. Rodent.
pp. 750, 753.)
Hab. Southern Mexico; Guatemala?
Average length about 10°50 inches, of tail-vertebrae 8°75 inches.
' Proc, Zool. Soc. 1861, p. 281.
1878.] MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. 661
Upper parts grey or reddish grey, the hairs black, tipped or
ringed and tipped with white or red. Pelage somewhat soft, with
a good deal of under-fur; lower parts deep red, orange, or white.
Tail black, washed with white, the hairs usually rufous at their base,
with black rings and white tips.
Under this nameI feel myself obliged to bring together two Mexican
Squirrels of which typical specimens are very different in appearance.
Mr. Allen has kept them separate under the names of S. aureigaster
and S. leucops, remarking that the difference in coloration leaves
little doubt of their distinctness, but adding that “more abundant
material may show that they are not specifically separable ”’ (op. cit.
p. 755). The colour-variation is not nearly so great as we shall find
it to be in the next species ; and after a careful examination of a great
number of specimens, especially of the fine series in the Paris Museum,
[ have been unable to find a single distinctive character which is
constant.
Typical specimens of the two forms may be thus described :—
1. The aureogaster type. Upper parts dark grey, the hairs
black, finely tipped with white. Lower parts deep red, which
is usually more or less washed over the grey on the shoulders and
flanks.
2. The /eucops type. Upper parts lighter grey, with a peculiar
golden lustre, the hairs being finely ringed with bright rufous and
tipped with i oe nape and rump more strongly rufous. Lower
parts bright red, orange, or white.
Many specimens, however, which have the general appearance of
aureogaster show the rufous subterminal ring of Jeucops on some
part of the dorsal surface ; and in others we find the red nape and
rump-patches gradually appearing. I am therefore compelled to
return to the view long ago expressed by Isidore Geoffroy ', and to
regard them as varieties of the same species.
With regard to distribution, all the specimens which I have seen
with authenticated localities are from Mexico ; but Mr. Allen mentions
one from Guatemala in the Washington Museum. Specimens in
the British Museum are labelled as being from Columbia; but
there can be little doubt that this is an error. S. aureogaster is stated
by Prof. Sumichrast (as quoted by Mr. Allen) to be ‘‘ the common
species of Sczurus of the tierra caliente of the east coast” of
Mexico; and both forms are found in the States of Oaxaca and Te-
huantepec.
Turning to the question of synonymy, I must briefly explain why
I have retained Erxleben’s name for this species. Mr. Allen con-
siders it to apply rather to the next species, but rejects it altogether,
on the ground that it is based partly on Buffon’s ‘ Coquallin”
(S. niger, L.), and partly on the ‘‘ Coztiocotequallin ”’ of Hernandez.
It is, however, primarily founded on the latter, Buffon’s name being
only quoted as a synonym; and Erxleben’s diagnosis and description
appear to me to be quite characteristic of the leucops form of the
1 Voy. de la Vénus, Zool. pp. 156-163.
662 MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. [June 18,
present species’. By retaining this appropriate name, we are enabled
to escape from F, Cuvier’s barbarous term aureogaster, under which
this beautiful animal has laboured in so many works. S. albipes,
Wagner, afterwards re-named S. varius by its describer, is doubt-
fully referred to the next species by Mr. Allen; but the description,
especially the character of the hairs being ringed with red, black,
and white, appears to point certainly to identity with Gray’s S. lewcops.
Wagner’s S\. socialis also, founded on a single example, seems also to
be a synonym of the present species ; for the characters of coloration
by which he differentiates it from his S. varius are quite insufficient,
and the small size may, as Mr. Allen suggests, be due to the imma-
turity of the specimen described.
V. SCIURUS HYPCOPYRRHUS.
Sciurus hypopyrrhus, Wagler, Isis, 1831, p. 610.
*Sciurus nigrescens, Bennett, P. Z. 8. 1833, p. 41.
*Sciurus colliei, Richardson, Zool. Voy. ‘ Blossom,’ p. 8, pl. i.
(1839}.
*Sciurus variegatoides, Ogilby, P. Z. S. 1839, p. 117.
*Sciurus richardsoni, Gray, Aun. Nat. Hist. x. p. 265 (1842, nec
Bachman).
*Sciurus boothie, Gray, List Mamm. Brit. Mus. p. 139 (1843).
*Sciurus griseocaudatus, Gray, Zool. Voy. ‘Sulphur,’ p. 34,
pl. xin. (1844).
*Sciurus fuscovariegatus, Schintz, Syn. Mamm. ii. p. 17 (1845,
ex Gray).
* Macroxus adolphei, Lesson, Desc. de Mamm. &e. p. 141 (1847).
*Macroxus pyladei, Lesson, tom. cit. p. 142.
*Sciurus dorsalis, Gray, P. Z. 8S. 1848, p. 138, pl. vii.
*Sciurus rigidus, Peters, Monatsb. Ak. Berl. 1863, p. 652.
* Sciurus oculatus, Peters, tom. cit. p. 653.
*Sciurus intermedius, Verreaux, ap. Gray, Ann. & Mag. Nat.
Hist. ser. 3, xx. p. 421 (1867).
* Macroaus nicoyana, Gray, tom. cit. p. 423.
*Macroxus melania, Gray, tom. cit. p. 425.
(Sciurus boothie et S. hypopyrrius, Allen, Mon. N.-Am. Rodent.
pp. 741, 746.)
Hab. Central America, from the Pacific coast of Mexico to the
{sthmus of Panama.
Average length about 11°75 inches, of tail-vertebrz 10 inches.
Pelage close-lying and rather glossy, with little underfur. Upper
parts dark brown or black, often broadly ringed with rufous or white.
Lower parts varying from deep red to creamy white, sometimes
ringed with black. ‘Tail black, generally washed with white, the
hair rufous, pale brown or whitish at their base, with one or two
black or brown rings and a white tip.
In uniting all the very variously marked Squirrelsthe names of which
1 * § corpore supra nigro, albo et fusco variegato. x « *« Magnitudo dupla
S. vulgaris. Auriculz imberbes. Corpus supra nigro, albo et fusco variegatum,
ventre flavescente. Cauda supra corpus reflexa.”
1878.] MR. E. R. ALSTON ON NEOTROPICAL SQUIRRELS. 663
ure given above, 1 have gone beyond Mr. Allen, who divides them
into two species under the titles of 8. hypopyrrhus and S. boothie.
The series of specimens to which he had access Jeft him strongly
impressed with their distinctness; but he adds that further material
may show that they intergrade. After a careful examination of the
large series contained in various Museums, and especially of that at
Paris, I am quite convinced that this is the case, and have even
grave doubts of the propriety of keeping them distinct from the last
species. But as I have uot yet found specimens strictly intermediate
between S. varieyatus and S. hypopyrrhus in the character of the
pelage and the aunulation of the fur, it seems best to keep them
distinct.
The opinion has been expressed above that the differences in com-
parative length of tail and ears, on which Mr. Allen lays most weight
in separating S. boothie from S. hypopyrrhus, cannot be depended
ou; and I have been totally unable to detect any constant difference
in general stoutness of form or, breadth of muzzle. As to colour,
the intergradations of the different varieties are quite complete, so
that it is often difficult or impossible to say to which a given spe-
cimen is to be referred. Each variety, however, has its own type,
and seems to predominate in its own immediate locality. The five
principal phases known to me may be thus arranged :—
1. The hypopyrrhus type. Upper parts dark-grey, the hairs
black, ringed with white or pale fulvous. Lower parts either con-
colorous, with the upper or washed with rufous. To this variety
T agree with Mr. Allen in referring Bennett’s S. nigrescens ; and it
is to the same form that the description of Macroxus boothie
iu Gray’s memoir of 1867 refers, although the same author’s
Seiurus boothie of 1833 was founded on a young example of the
white-bellied colli#i phase. ‘Lhe rufous-beilied specimens lead us
directly into
2. ‘The rigidus type. Upper parts usually marbled with fulvous
and black, tue hairs being glossy brown or black with a median band
of yellowish brown. Lower parts bright rufous (rigidus), or varied
with rufous aud white, either symmetrically (intermedius) or asym-
metrically (xicoyanus). In many specimens, as observed by Mr.
Allen, there is a more or less marked tendency to the development
of a pale grizzled lateral stripe placed high on the tanks and con-
fining the dark dorsal area to a broad median band. From these we
have every gradation into the rufous phase of
3. The dorsalis type. Upper parts with the median dorsa! area
black, flanks and lower parts white, fulvous, or rufous. In many in-
dividuals the hairs of the black dorsal area will be found to have
concealed rings of white or fulvous; so that we have a further tran-
sition into
4. The colliai type. Upper parts much as in the rigidus form,
but usually less rutous ; lower parts white (colliei, adolphet) or pale
fulvous (variegatoides, griseocaudutus, pyladei, oculatus). Usu-
ally the dark coloration of the back is extended onto the shoulders,
hips, and limbs; but in others it is confined to the middle of
664 MR.E.R. ALSTON ON NEOTROPICAL SQUIRRELS. [June 18,
the back, as in the last phase, while the rufous-bellied phases lead
us directly back into the rigidus form.
5. The melania type. Upper and lower parts uniform glossy dark
brown, often with irregular patches of paler brown. This is a very
peculiar phase, and I have not yet seen any direct intergradation with
the other varieties; but uniformity of coloration is so abnormal
among the Squirrels, that it seems impossible to doubt that Mr. Allen
is right in regarding it as a melanism. The colour is exactly similar
to that of the white-bellied dorsalis varieties, in which alse the paler
apparently faded patches are sometimes seen. It appears to be much
rarer than any of the other forms described.
With regard to the synonymy, J may observe that I have been
able to examine the types of all the “species”’ here united, except-
ing that of S. hypopyrrhus, which, however, has been well described
by Wagler and Wagner; it appears to be a dark variety without
the usual wash of white on the tail. That of 8. nigrescens only
differs in having the fur of the lower parts ringed like that of the
back ; it agrees well with specimens in the Paris and Berlin Museums.
S. collizi and 8. adolphet are synonyms for the white-bellied form
above described, from which I am also unable to distinguish 8S.
richardsoni of Gray (not of Bachman), on which both S. doothie and
S. fuscovariegatus were founded. SS. variegatoides, S. griseocauda-
tus, S. pyladei, and S. oculatus are all intermediate links with more
or less fulvous lower parts, leading to the phase with a red or red
and white belly, on which the names S. intermedius, S. rigidus,
and Macrozus nicoyanus have been bestowed.
Of the geographical distribution of the various races we can only
judge from the comparatively few specimens of which the exact
localities have been noted. The hypopyrrhus phase appears to be
the most northern, the colligi to obtain principally along the Pacifie
slopes, and the dorsalis to be the most southern. Each, however,
appears to be found along with others im some part. Thus, I have
seen specimens of hypopyrrhus type from Mexico, Honduras, and
Guatemala, of rigidus from Guatemala, Nicaragua, and Costa Rica,
of dorsalis from Nicaragua, Costa Rica, Veragua, and Panama, and
of collizi from the west coast of Mexico and Guatemala, Nicaragua,
and Costa Rica. The only localities which I know for S. melania
are Nicaragua and Veragua.
VI. Scrurus STRAMINEUS.
Sciurus stramineus, Eydoux & Souleyet, Voy. de ‘La Bonite,’
Zool. i. p. 37, Atlas, pl. ix. (1841),
* Sciurus nebouwii, Is. Geoffroy, Voy. de ‘ La Vénus,’ Zool. p. 163,
Atlas, pl. xii. (1855).
*Macrovus fraseri, Gray, Ann. & Mag. Nat. Hist. ser. 3, xx.
p. 430 (1867).
(Sciurus hypopyrrhus, Allen, Mon, N.-Am. Rodent. p. 747.)
Hab. Ecuador; Peru.
Average length about 12°50 in., of tail-vertebree 12°50. Upper
parts nearly uniform grizzled grey, ramp and lower parts more or
1878.] MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. 665
less washed with rufous, the hairs blackish towards their base, with
long white fulvous or rufous ends ; feet blackish ; tail black, washed
with rufous and whitish; pelage very sparse and harsh, with almost
no under-fur.
This curious Squirrel, which appears to be the sole representative
of the genus in Western Peru, is rare in collections ; and though the
few specimens which have been sent to Europe differ little in colour,
they have received the above names. Mr. Allen referred three
specimens from Guayaquil to S. hypopyrrhus (tom. cit. p. 747) ; and
certainly some varieties of that species bear a strong general resem-
blance to the present. But so far as can be judged from the small
series in the Paris and British Museums, S. stvamineus seems to be
fairly entitled to specific recognition; and intermediate forms are
hardly likely to be found, as no similar Squirrel appears to be known
from any part of Columbia. It is only fair to Mr, Allen to add, that
Gray’s description of M. fraseri is so imperfect that it is not sur-
prising that the American zoologist should have doubtfully referred
it to S. tephrogaster (tom. cit. p. 763).
A remarkable peculiarity of this species is its tendency to the
development of irregular tufts of pure white hairs, rather longer than
the rest of the fur, and sometimes uniting in larger patches. These
asymmetrical markings are present in the majority of the individuals
examined.
A fine specimen of S. stramineus from Ecuador is now living in the
Society’s Gardens’.
VII. Scrurvs VARIABILIs.
*Sciurus variabilis, Is. Geoffroy, Mag. de Zool. 1832, pl. iv.
Sciurus langsdorfi, Brandt, Mém. Ac. St. Pétersb. 6° sér. iii. p.425,
1835).
; Sciurus igniventris, Natterer ap. Wagner, Wiegm. Arch. 1842,
360.
~ Sciurus pyrrhonotus, Natterer, loc. cit.
Sciurus tricolor, Poppig ap. Tschudi, Faun. Peruan. p. 156, pl. xi.
(1844-46).
Sciurus morio, Wagner, Abh. Bayer. Ak. v. p. 275 (1850, nec
Gray),
*Sciurus gerrardi, Gray, P. Z. 8. 1861, p. 92, pl. xvi.
*Sciurus brunneo-niger, Castelnau ap. Gray, Ann. & Mag. Nat.
Hist. ser. 3, xx. p. 429 (1867).
*Sciurus fumigatus, Gray, tom. cit. p. 428 (1867).
(Sciurus gerrardi et S. variabilis, Allen, Mon. N.-Amer. Rodent.
pp- 766, 768.)
Hab. Panama; Venezuela; Columbia; Ecuador; Peru; Brazil;
Bolivia.
Average length about 12 inches, of tail-vertebrze 11 inches. Upper
parts red, either pure or more or less grizzled with black, lower parts
white, fulvous, or light rufous, sharply defined from the dorsal colour-
1 See above, p. 441.
Proc. Zoou. Sec.—1878, No. XLII. 43
666 MR. E.R. ALSTON ON NEOTROPICAL sQuirRELs. [June 18,
ing; tail black, washed with bright-red orange, often very bushy.
Pelage rather harsh and sparse, ears high and narrow.
Here, again, the greater amount of material at my disposal
compels me to go beyond Mr. Allen in the identification of nominal
species. Most of the above synonyms were brought together
by him. under the name of S. variabilis; but S. gerrardi and S.
brunnre - niger . sfeniger were kept separate under the former title. The principal
points on which he rested were the smaller size and shorter ears
of 8. gerrardi; but on examination of a sufficient series, I have
not been able to find any constancy in the proportions of the ears,
while the difference in size totally disappears, as is shown by the
following series of measurements—9°50, 9°75, 10°00, 10°25, 16°75,
11, 11°50, 11°75, 12°00, 12°75, 13°00. The smaller specimens
(S. variabilis, S. gerrardi, &c.) appear to prevail towards the north :
but this is not constant ; for a Santa-Martha example measures over 12
inches, while others from Brazil are under 10 in., although evidently
adult. Nor is it constantly connected with any of the numerous
varieties of coloration—rufous, grizzled, and melanistic specimens
occurring of all sizes.
The range of these variations of colour is not nearly so great as
in S. hypopyrrhus; and they seem to resolve themselves into three
primary groups, namely :—
1. The morio type. Upper and lower parts black, only slightly
ringed with rufous or fulvous, tail black, either entirely or slightly
washed with red. Of this melanistic form no two specimens agree
exactly in the proportion of red ; and every step is present leading
into
2. The variabilis type. Upper parts red, more or less varied with
black—but the rufous tint prevailing, especially on the flanks ; lower
parts pure white, fulvous, or pale rufous; tail (either narrow or bushy)
black, strongly washed with bright rufous or orange. Here we have
an endless series of minor variations, from the very brilliant black
and red specimens sent from the Rio Napo to the British Museum,
and the bright rufous Columbian form, through more and more
greyish red specimens, till we find ourselves in
3. The langsdorffi type. Upper parts reddish or yellowish griz-
zled, the hairs being pretty uniformly ringed with black and red or
black and yellow ; lower parts and tail as in the last.
Each of these styles appears to prevail in certain localities: thus
the melanistic form has been found on the Upper Amazous by Mr.
Bates and by Castelnau and Deville, the rufous type on the Rio
Negro and Rio Madeira by Natterer, and the grizzled in North-
western Brazil by Spix. But there seems to be no regularity in their
distribution ; and I have seen both grizzled and red specimens from
Panama, Venezuela, Columbia, and North-eastern Peru.
The oldest name, and therefore the one here adopted, is S. varia-
bilis, given in 1832 by Isidore Geoffroy to small red specimens from
Columbia, and strictly synonymous with Gray’s S. gerrardi; while
S. igniventris and S. pyrrhonotus were MS. names of Natterer’s,
applied by Wagner to other varieties of the rufous type. 8. langs-
1878.] MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. 667
dori and S. tricolor both refer to the forms in which the upper
parts are more or less uniformly grizzled; while S. morio, S. #4fe5
niger, and M. fumigatus, are all more or less melanistic varieties.
VIII. Scrurvus GRISEOGENYs.
*Sciurus estuans, var. hoffmanni, Peters, Monatsb. Ak. Berl.
1863, p. 654.
*Sciurus hyporrhodus, Gray, Ann. & Mag. Nat. Hist. ser. 3, xx.
p- 419 (1867).
*Macroxus xanthotus, Gray, tom. cit. p. 429.
* Macroaus griseogenu, Gray, tom. cit. p. 429.
(Sciurus estuans, var. rufo-niger, Allen, Mon. N.-Am. Rodent.
p: 797.)
Hab. Ecuador; Columbia; Venezuela; Panama; Veragua; Costa
Rica; Nicaragua.
Length about 9 inches, of tail-vertebrae 8 inches. One upper pre-
molar. Upper parts dark olive, the hairs dusky, very minutely an-
nulated with rufous or fulvous ; lower parts bright orange or rufous ;
tail black, more or less washed with bright red or deep orange, the
hairs reddish brown banded with black and tipped more or less
broadly with rufous.
Mr. Allen, in his monograph, regards this Squirrel as a “ variety ”
or geographical race of the next species, differing in its uniformly
larger size and strikingly in the coloration of its tail. In a sub-
sequent letter to me, he says :—‘“‘ It would perhaps be just as well
to recognize it as entitled to specific rank, although I still feel sure
of their intergradation.”” That such connecting links may yet be
found seems very probable ; but I have not been able to find such
in the very large series which I have examined, and am consequently
compelled to keep them provisionally distinct.
Unfortunately Mr. Allen has identified this species with Pucheran’s
S. rufo-niger, which, as will be seen presently,is a much smaller and
quite distinct species’. Dr. Peters described it only as a variety of
S. estuans; and though specimens in the Berlin Museum are la-
belled ‘‘ Sciurus hoffmanni,” the name remains a manuscript one.
Of Gray’s three titles I have adopted griseogena (more correctly gri-
seogenys) as being simultaneous in date with the others, and as in-
dicating the typical form. His S. hyporrhodus is founded on an
example from Santa Fé de Bogote, and is characterized by its
long soft fur and hairy ears, peculiarities easily explained when
we remember that that city stands at an elevation of over 8800
feet, and is overlooked by lofty mountains. Macroxus xanthotus
is undoubtedly an immature animal, as is proved by the dentition
of the type specimen. By some curious error Gray’s account of
this last has been printed after that of M. brunneo-niger, instead of
after M. griseogena: so that the remark, “very like the former” &c.,
naturally led Mr. Allen to refer the synonym to 8. gerrardi.
S. griseogenys varies very little in colour: in some examples the
1 Cf. infra. p. 669.
43*
668 MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. [June 18,
middle of the back is considerably darker than the flanks; and there
is some variety in the exact tint of the red wash on the tail. I can
confirm Mr. Allen’s statement that this species (as well as S. varia-
bilis and S. wstuans) have normally only one upper premolar. Dr.
Peters states, it is true, that the type of his “ var. hoffmanni” had
two; but the first was minute and only attached to the gum; and
I have not been able to find more than one, either in dried skulls or
in spirit-specimens.
IX. Scrurvus £STUANS.
Sciurus estuans, Linneeus, Syst. Nat. i. p. 88 (1766).
Myoaus guerlingus, Shaw, Gen. Zool. ii. p. 171, pl. elvi. (1801).
Sciurus gilvigularis, Natterer ap. Wagner, Abh. Bayer. Ak. v.
p- 283 (1850).
*Macroxus leucogaster, Gray, Ann. & Mag. Nat. Hist. ser. 3,
xx. p. 430 (1867, nec F. Cuv.).
* Macroxus irroratus, Gray, tom. cit. p. 431.
* Macroaus flaviventris, Castelnau ap. Gray, tom. cit. p. 432.
(Sciurus e@stuans, var. estuans, Allen, Mon. N.-Am. Rodent.
p- 796, 1877).
Hab. Guiana; Brazil; Eastern Peru; Bolivia.
Average length about 7:25 inches, of tail-vertebree 7 inches. One
upper premolar. Upper parts olive, rather lighter and more fulvous
than in the last species; lower parts fulvous, greyish, or white; tail
black, more or less washed with pale fulvous, the hairs pale brown,
banded with black and tipped with pale faded yellow.
I have little to add to Mr. Allen’s critical notes on this well-known
Squirrel. Gray states, in his description of his M. leucogaster (not
to be confounded with FP. Cuvier’s species of that name’), that the
hairs of the lower parts are “‘white to the base;” this is not the
case, though the dusky colour at the roots is little developed on the
chest. M.irroratus must also be placed here, although the original
description is such that Mr. Allen unhesitatingly referred it to the
last species ; and M. flaviventer is to me quite undistinguishable from
the ordinary type of S. estuans. S. pusillus, Geoffr., and Macrotus
kuhlit, Gr., are considered by Mr. Allen to be the young of the
present species; but I believe them to be quite distinct”.
X. ScruRUS DEPPEI.
*Sciurus deppei, Peters, Monatsb. Ak. Berl. 1863, p. 654.
*Macroxus tephrogaster, Gray, Ann. & Mag. Nat. Hist. ser. 3,
xx. p. 431.
* Macroxus teniurus, Gray, loc. cit.
* Macroxus medellinensis, Gray, op. cit. ser. 4, x. p. 408 (1872).
(Sciurus tephrogaster, Allen, Mon. N.-Am. Rodent. p. 763 (1877)).
Hab. Mexico ; Honduras; Guatemala; Columbia.
Average length nearly 9 inches, of tail-vertebrze 7°25 inches. Two
upper premolars. Upper parts dark olive, often darker along the
1 Cf. antei, p. 660. 2 Cf. infra, p. 670.
1878.] MR. E. R. ALSTON ON NEOTROPICAL SQUIRRELS. 669
middle of the back ; lower parts white, greyish white, or light ful-
vous ; tail black, lightly washed with white, the hairs fulvous, with
black bands and short pure white tips.
Mr. Allen was led to admit the specific validity of this Squirrel,
on finding that it differed from the last two in having normally two
well-developed upper premolars ; and the specimens which I’ have
been able to examine confirm his observation. He failed, however,
to recognize this species in Dr. Peters’s description of his S. deppei,
which he doubtfully referred to the southern form of S. carolinensis',
An examination of the types of S. deppei, M. tephrogaster, M. teni-
urus and M. medellinensis convinces me that they are all strictly
synonymous, the species presenting but little variation in coloration.
In S. deppei and M. teniurus the lower parts are washed with ful-
vous ; in M. tephrogaster they are greyish white; while the types of
M. medellinensis ave smaller and have the middle of the back
nearly black, exactly as in many specimens of 8. griseogenys. In-
termediate examples occur ; and the whole range of variation between
the extremes is comparatively trifling.
As already observed”, M. fraseri, Gr., was so insufficiently de-
scribed that Mr. Allen was led to identify it with the present species,
which is about half its size and totally different in coloration.
XI. Scrurus RUFO-NIGHR. cht us Osayus-
*Sciurus rufo-niger, Pacheran, Rev. de Zool. 1845, p. 336{q 20
*Sciurus chrysosurus, Pucheran, tom. cit. p. 337.
*Macroxus tephrogaster minor, Gray, MS. (Mus. Brit.).
Hab. Columbia; Panama; Veragua.
Average length about 5°50 inches, of tail-vertebre 3°75 inches.
Two upper premolars. Upper parts dark olive, the hairs very mi-
nutely tipped with fulvous ; breast bright rufous, rest of lower parts
like the upper, but paler ; tail nearly uniform with the back, the
hairs reddish fulvous, barred with black and minutely tipped with
pale yellow or white.
On examining the type of Pucheran’s S. rufo-niger in the Paris
Museum, I found that it was not identical with S. griseogenys, as
Mr. Allen supposed, but rather allied to 8. deppei; and I soon recog-
nized in it a small Squirrel of which I had seen several specimens
from Panama, and which I had begun to fear would require a new
name. These examples prove to agree further with S. deppei in
having two upper premolars, but differ in being more than one third
smaller, in the colour of the lower parts (which are only paler than
the upper, save on the breast), and in the tail being nearly uniform
in colour with the back (the hairs having only very minute white or
yellow tips). Specimens in the British Museum are labelled M.
tephrogaster minor ; but I cannot doubt the distinctness of the form.
The type of S. rufoniger has the middle of the back nearly black ;
while that of MW. chrysosurus appears to be a variety, merely differing
in the tail being more rufeus.
1 Tom. cit. p. 710. > Cf. ante, p. 665.
‘=>
670 MR. P. L. SCLATER ON BIRDS [June 18,
XII. Scrurus pusttius. (Plate XLI.)
Sciurus pusillus, Geoffroy apud Desmarest, Mamm. p. 337,
pl. Ixxvii. fig. 2 (1820).
*Macrozus kuhlii, Gray, Ann. & Mag. Nat. Hist. ser. 3, xx.
p- 433 (1867).
Hab. Guiana; Brazil.
Length about 4°50 inches, of tail-vertebree.3 inches. ‘Two upper
premolars. Upper parts greyish olive, the hairs dusky, very mi-
nutely tipped with pale fulvous ; lower parts only slightly paler ; tail
black, washed with whitish or pale yellow.
I was not able to ascertain the type of Geoffroy’s S. pusillus in
the Paris Museum ; but there can hardly be a doubt of its identity
with specimens standing under that name ia the British museum, and
with the types of Gray’s Macrovus kuhlit. All these are supposed by
Mr. Allen to be merely very young individuals of S. @stuans’, in spite
of Buffon’ s observations on the development of the reproductive organs
in the type of S. pusillus*. The British- Museum specimens, however,
agree in size with Buffon’s and Desmarest’s descriptions, and areset
et quite adult, as shown by their skulls and teeth ; moreover the two
specimens examined differ from S. estuans in the possession of two
upper premolars. ‘Their very small size and the nearly uniform
greyish olive of both their upper and lower parts prevent their being
confounded with any other species known to me. The type of
M. kuhlii, collected in Brazil by Castelnau, merely differs in having
a white spot at the root of each ear, and in the extreme tips of the
hairs of the tail being white instead of pale fulvous.
2. On a third Collection of Birds made by the Rey. G. Brown,
C.M.Z.S., in the Duke-of-York Group of Islands and its
vicinity. By P. L. Scrarer, M.A:, Ph.D., F.RS.,
Secretary to the Society.
[Received June 11, 1878.]
(Plate XLII.)
T have now the pleasure of submitting to the Society a third® col-
lection of birds made by our Corresponding Member the Rev. J.
Brown, in the group of islands known generally as the Duke-of- York
Islands, and on the adjacent islands of New Britain and New
Treland.
The collection was forwarded from Port Hunter, the missionary
station in the Duke-of-York group, on the 24th of February last by
a small trading-steamer bound for Hongkong. It contains a series
1 Tom. cit. p. 759. 2 Hist. Nat. Suppl. vii. p. 264.
3 For notices of two previous collections, see P. Z. 8. 1877, p. 96, and 1878,
p. 289.
Fu pg
Hanhart
1878.] FROM THE DUKE-OF-YORK ISLANDS. 671
of birds obtained while coasting along in a boat amongst the islands
of the group. Ima letter of the same date Mr. Brown states that
he was hoping to be able shortly to spenda week at Spacious Bay or
some other place in New Britain, where he would be able to make a
more extensive collection.
The present collection contains thirty-five skins referable to thirty
species. The exact localities being marked in every case, I think it
advisable to give a perfect list of the species. ‘ Duke-of-York
group”? means one of the group of twelve islands which form together
the land marked in our chart “‘ Duke-of-York Island ’’?.
See P. Z. 8. Locality. Date.
if.” Gisticola ruficeps’... .20....20..-.-92- 1877, p. 98 | Duke-of-York group. | 1, 78
2. | Sauloprocta melaleuca..............: x 99 of y ee
3. | Rhipidura setosa ... ........:..cceees 3 99 is 33
4. | Monarcha verticalis ................+: + 99 = re
5. | —— chrysomelas ............00ecseees »» LOO| New Ireland.
Ore f—-— aL eCtO = aint... .ce cercnceotap base » 100| Duke-of-York group.
ui AIEOP DEGRA cae bode cetecaunecacl at’ b Geened 5 -
8. | Artamus insignis ..........00....00065 » 101] Spacious Bay, N.B. | 1,’78
9. | Hdoliosoma, sp. ine. ( Q).......++4-. » 10] | New Ireland. ules
RU ualaceskarw: 21.085. tubes seiawainn tens ,, 101) Duke-of-York group.
11. | Dicranostreptus megarhynchus ...| _,, 101 | New Ireland.
ier | Gracia KYEMUL ..ccccesectsccteeresss ‘ 104| Duke-of-York group.
13. | Alcedo moluccensis .......06...se0e8- i 105 » 5 5777
14. | Halcyon albicilla ..................+++ 58 105 a 4
V5.5 jBuceros ruficollis .......c.s<secveeses] | -cevecees Spacious Bay, N.B.
16. | Eurystomus crassirostris............ » 106} Duke-of-York group. |10, 77
17. | Centropus ateralbus ............+06... ,, 106} New Ireland.
i8. | Cacomantis insperatus............++ » 106} Duke-of-York group.
19. | Hudynamis picatus ...........ee es 3) 06 ” ”
20. | Geoffroius heteroclitus............2..) cesses New Ireland. .
21. CYABICEPS- .2.8-..:cervesecceonee » 106| New Britain.
22. | Trichoglossus subplacens..........-. » 108} Duke-of-York group. |10, 77
23. | Carpophaga rubricera ............... - 109 of
24. NDEMEV ORM ear atecttecsact sheets re 109 A, ;
255 melanochroa, Sp. NOV.......+22| sseeeeees “3 :,
26. | Gidirhinus insolitus ...............00. ‘s 110 a, is 10, 77
27. | Macropygia browni .............+0++- - 110 3 a
28. | Chaleophaps stephani ............... seis is - -
29. | Nycticorax caledonicus.............06] sereeees- bs im
30. | Megapodius eremita...............04+ », 113 | New Ireland.
I now give some notes on the species new to the locality, or on
which I have additional information to offer.
4. Monarcua verticais, Scl. P. Z. 8. 1877, p. 99, pl. xiv.
fig. 1.
A second specimen of this interesting species has an oblong white
spot at the extremity of the outer rectrix on both sides.
1 Of. footnote, P. Z. 8. 1877, p. 98.
672 ON BIRDS FROM THE DUKE-OF-YORK ISLANDS. [June 18,
7. Monarcua 1nornatus (Garnot); Scl. P. Z.8. 1877, p. 552.
This bird has not occurred in the former collections of Mr. Brown,
but is a well-known and rather widely extended Papuan species.
15. Bucreros rvuFIcoLiis, Vieill.; Temm. Pl. Col. 577; Sel.
P. Z. S. 1869, p. 122.
Apparently an adult female of this Hornbill from Spacious Bay,
New Britain.
20. GEOFFROIUS HETEROCLITUS.
Psittacus heteroclitus, Hombr. et Jacq. Ann. Se. Nat. xvi. p. 319.
Pionus heteroclitus, id. Voy. au P. 8. Atl. t. 25 bis (¢).
Pione heteroclitus, Puch. ibid. Zool. ii. p. 103.
A fine male of this species from New Ireland agrees with the ori-
ginal figure and description of that sex. One of the Solomon-Islands
skins referred to, P. Z.S. 1869, p. 122, appears to be the female of
this species, the other to belong to the next.
21. Grorrroius CYANICEPS, Pucheran.
A skin from New Britain marked “ male,’? which has had the
wings cut away, and has apparently been in a cage, is probably of
the same species as that previously referred (P. Z. 8. 1877, p. 107)
to Geoffroius cyaniceps. ‘This is decidedly smaller than G. hetero-
clitus, and has the top of the head of a pale greenish grey with the
sides yellowish.
Dr. Salvadori, who examined the specimen in the previous col-
lection of Mr. Brown, remarks that it comes very near the female of
G. simplex, Miller. A better series of this bird is necessary before
it can be made out decidedly.
25. CARPOPHAGA MELANOCHROA, sp. nov. (Plate XLII.)
Nigricanti-ardesiaca fere unicolor, alis cauda et dorso inferiore
in plumbeum transeuntibus ; alarum tectricibus omnibus margi-
nibus angustissimis albis preditis ; subalaribus et crisso castaneo
tinctis ; rostro nigro; pedibus purpureo-rubris.
Long. tota 15°5, alee 9°5, caudee rectr. med. 6°0, ext. 5°5.
Hab. Ins. Duke of York.
This fine Fruit-pigeon is quite new to me, and, I believe, unde-
‘scribed, though I am not acquainted with C. pistrinaria, Bp.’, of the
Solomon Islands, to the neighbourhood of which it would appear to
belong.
Mr. Brown’s single specimen is marked “male; eyes bright
scarlet; legs purple.’’ As will be seen from the measurements, the
tail is slightly rounded; there are fourteen rectrices, as is always,
I believe, the case in Carpophaga. The under-surface of the tail is
rather ashy white, that of the wings grevish black.
The uniform dark colour, varied only by the very narrow margins
of the wing-coverts, will render this species easily recognizable.
1 Consp. Ay, ii. p. 36.
PZ SAGTSPESLE
18 owt em
~s
ieee ea a
JSmit hth
AH Young MB. del
ANATOMY OF CHLAMYDOPHORUS TRUNCATUS.
1878.] DR. M. WATSON ON CHLAMYDOPHORUS TRUNCATUS. 673
27. Macroryera Brown], Scl. P. Z. 8. 1877, p. 110.
I am glad to receive a second example of this fine species from
the Duke-of-York group.
Dr. Salvadori writes me that he suspects that Turaccena crassi-
rostris, Gould (P. Z.S. 1856, p. 136), may be the young of Macro-
pygia browni. Ihave examined Mr. Gould’s typical specimen of the
former species now in the British Museum ; but although I believe
it is, no doubt, the young of an allied species of Macropygia, I do not
think it can belong to JZ. drowni, as it is so much smaller in size.
29. Nycrocorax CALEDoNICcUs (Gm.); Gould, Handb. B. Austr.
i. p. 311.
An adult of this Night-Heron from the Duke-of-York group is
marked :—“‘Trides yellow ; legs dusky reddish pink ; bare space about
eyes lavender.—G. B.”
30. MEGAPODIUS EREMITA.
Megapodius eremita, Hartl. P. Z. 8. 1867, p. 830.
M. hueskeri, Cab. et Reich. J. f. O. 1876, p. 326; Scl. P. Z.8.
1877, p. 113.
M. rubrifrons, Scl. P. Z.S. 1877, p. 556.
Prof. Salvadori has recently examined the typical specimens of M.
eremita from the Echiquier Islands, M. hueskeri from New Hanover,
and M. rubrifrons from Admiralty Island, and pronounces them
to be identical.
es
3. On the Male Generative Organs of Chlamydophorus trun-
catus and Dasypus sexcinctus. By M. Warson, M.D.,
Professor of Anatomy, Owens College, Manchester.
[Received June 11, 1878.]
(Plate XLIIL)
1. CHLAMYDOPHORUS TRUNCATUS.
The anatomy of Chlamydophorus truncatus has formed the subject
of elaborate monographs by Hyrtl!, Macalister’, and others*?; and
little remains to be said regarding such parts of the animal as have
been fully investigated by these anatomists. On looking over their
essays, however, I find that, with regard to the male genera-
tive organs, we possess almost no reliable information. Professor
Hyrtl’s specimen was a female ; and although that anatomized by
Professor Macalister was of the opposite sex, his reference to the
male organs is sufficiently meagre to justify me in giving a detailed
description of these from two male specimens of this rare mammal
which have lately come into my hands.
1 Denkschriften der kais. Akad, Wien, ix. 1855.
? Trans. Roy. Irish Acad. 1875, p. 219.
* Harlan, Lond. Zool. Journ, 1825; Atkinson, Journ. of Anatomy, vol. v.
674 DR. M. WATSON ON THE MALE ORGANS OF _ [June 18,
The specimen measured 43 inches in length, exclusive of the tail.
Penis..—The penis is large for the size of the animal, and, so
far as one can judge from the examination of specimens preserved
in alcohol, appears, in the natural condition, to be strongly curved,
the convexity of the curve being directed forward, whilst the extre-
mity of the penis is directed downward and backward toward the
anus. The organ js slender, and tapers gradually from base to apex,
measuring 3 of an inch in length and 7; of an inch in diameter.
The base is surrounded by a small prepuce consisting of little more
than a narrow ring of integument, and is entirely destitute of a frenum
preeputii. The presence of two powerful retractores penis, however,
justifies us in concluding that in the Chlamydophorus, as in Da-
sypus sexcinctus, the penis is, in the flaccid condition, retracted
within this apparently rudimentary prepuce. The intermediate por-
tion of the penis presents an appearance of irregularly disposed cir-
cular constrictions, apparently due to the annulated character of the
integument, and corresponding exactly to that described below in the
six-banded Armadillo. The glans penis is not distinguishable as a
separate portion from the rest of the organ; and the absence of it,
together with that of the bulb, would appear to indicate that a corpus
spongiosum is altogether wanting, as it apparently is in the Armadilio,
the penis in that animal being formed exclusively by the two cor-
pora cavernosa. I could not distinguish any patch of transverse
ridges on the ventral aspect of the organ, such as is seen in that of
Dasypus seacinctus ; but the small size of the parts, combined with
their immersion in alcohol, may have obliterated these (under any cir-
cumstances) very minute elevations.
Internal Organs.
Kidney.—The kidneys are large in comparison with the size of
the animal, and are situated in the lumbar region. Each is almost
globular in form, measuring 2 of an inch in length and 3 in breadth,
and is surmounted by asuprarenal capsule of large size and pyramidal
form. The ureter measures 5° of an inch in length, and enters
the bladder close to the orifice of the urethra, being crossed on its
inner side from before backward by the corresponding vas deferens.
Bladder,—The bladder, globular in form, small, and provided
with walls of considerable thickness, lies altogether in front of the
transverse ligament which unites the two pubic bones together, and
which corresponds in position to the pubic portion of the brim of
the pelvis in the majority of mammals. The viscus is partly invested
by peritoneum, which forms a well-marked inferior ligament attach-
ing it to the middle line of the anterior abdominal wall. In the spe-
cimen examined the bladder was contracted, and measured } of an
inch in length and the same in greatest breadth. The urethral ori-
fice is on a level with the base of the viscus.
Urethra.— The urethra is divisible into two portions—an intra-
pelvic or muscular, and an extrapelvic or spongy portion. The
intrapelvic portion, from the neck of the bladder to the point of en-
trance of the Cowperian ducts, measures 3°; of an inch in length, Its
1878. | CHLAMYDOPHORUS TRUNCATUS. 675
walls are thick, being surrounded by a layer of circularly disposed
muscular fibres. Resting upon the upper surface of the proximal
half of this portion of the urethra is the prostate gland, the pos-
terior border of which is attached to the tube. On slitting open
the canal, no trace of a vesicula prostatica could be distinguished ;
but it is well to bear in mind that the small size of the parts, together
with the immersion of these for some time in alcohol, may have
served to prevent the recognition of what, under any circumstances,
must be a very minute organ. Professor Macalister’ refers to the
presence of a “Weberian organ, of which, however, he gives no de-
scription, merely statiug that it is “long, and not distinetly bifid ;”
but inasmuch as he makes no mention ofa prostate gland, it appears
to me to be not improbable that he may have mistaken that gland
for the organ in question, an error which need occasion no surprise
when the small size of the parts is taken into consideration. If the
Weberian organ be present, it certainly does not project beyond the
wall of the urethra. The extrapelvic or spongy portion of the
urethra lies within the penis and measures { of an inch in length.
At its commencement it receives the ducts of Cowper’ s glands, whilst
at its termination it opens upon the summit of the rounded extre-
mity of the penis.
Testicles and vasa deferentia.—Professor Macalister? states that the
testicles lie within the ‘‘ wide abdominal ends of the inguinal canals;”
but this was certainly not the case in my specimen. In it the tes-
ticles were situated immediately posterior to the kidneys, and in close
relation to the superior abdominal wall, to which each was closely
attached through the yefection of the peritoneum. The testicle of
the right side was posterior in position to that of the left. The gland
itself was of the same size and form as a canary-seed, and, together
with the epididymis, measured 1 of an inch in length. The latter
was of large size as compared with the testicle, to the outer side
of which it was attached at both its anterior and posterior extremi-
ties. The anterior extremity, or globus major, is continuous with the
testicle, the separation between them being indicated by a slight
constriction. From this it passes forward for +1; of an inch, and,
suddenly curving upon itself, turns backward to the same extent,
and becomes constricted to form the body of the epididymis. The
latter extends along the outer side of the testicle as far as its
posterior extremity, where it again expands into a glandular mass, or
globus minor, which, however, is of larger size than both limbs
together of the globus major. From this body the vas deferens
passes off. The entire testicular mass is placed obliquely, its ante-
rior extremity being directed obliquely forward and outward, the
posterior, which is in contact with the lateral aspect of the apex of
the bladder, in the opposite direction. The vas deferens measures 4
of an inch in length, and crosses the ureter in its course backward.
The two vasa at their termination are almost in contact, and disap-
pear from view by sinking into the anterior border of the prostate
gland, the substance of which they traverse in their course to the
1 Loc, cit. 2 Loe. cit.
676 DR. WATSON ON CHLAMYDOPHORUS TRUNCATUS [June 18,
urethra. As Professor Macalister observes, the vasa are not convo-
luted, neither do they present any trace of lateral diverticula or vesi-
culze seminales.
Prostate gland is well developed, and forms a quadrilateral mass
of lobulated glandular tissue lying upon the upper surface of the intra-
pelvic urethra. It measures + of an inch in length, and the same
in breadth, and is attached by its posterior border to the urethra,
its anterior border, which is free, extending between the vesical
openings of the ureters. The vasa deferentia sink into the latter
in their course to the urethra. Prof. Macalister does not refer to
this gland.
Cowper’s glands.—These, which are not mentioned by Macalister,
are two innumber. Each lies in close relation to, and slightly under
cover of, the inner side of the crus penis. They are regularly pyri-
form; and each measurss ;; of an inch in length. Their ducts pass
forward, and open at once into the commencement of the spongy
portion of the urethra.
Penis.—The penis is formed by the junction of the two crura
exclusively, the third element or corpus spongiosum being entirely
absent. The crus penis of each side is attached to the ramus of the
ischium, as well as to the posterior half of the free border of the pubic
bone, the latter bone not uniting with its fellow of the opposite side.
Each measures } an inch in length, and lies along the inner side of
the adductor muscle of the thigh. The two crura unite opposite
the anterior border of the pubic bones, and together form, as it
were, a bridge, underneath which the rectum passes backward to the
anus. In consequence of the absence of a corpus spongiosum, the
penisis devoid of both bulb and glans. The penis itself measures
4 of an inch in length, of which the basal 2 are concealed by the inte-
gument, the remainder being free.
Muscles of penis.—Of these there are three on each side of the
middle line:—(1) The ischio-cavernosus is strong and covers the
crus. Its fibres are attached posteriorly to the ischium, whilst ante-
riorly they are inserted into the crus behind the junction of the
latter with its fellow. A small portion of the crus between this and
the next muscle is uncovered by muscular fibres. (2) Bulbo-caver-
nosi. These two muscles are apparently continuous across the middle
line, and together form an investment of transversely arranged mus-
cular fibres which covers the lower surface of the basal portion of
the penis. No median raphé could be distinguished between them.
(3) Retractores penis. These are placed on the dorsal aspect of the
organ. ach is a delicate riband-like muscle which, arising from
the upper wall of the pelvic cavity, crosses the outer side of the de-
pressor caudz muscle, and passing down parallel to the crus penis,
and along the dorsal aspect of that organ, is inserted into the corpus
cavernosum immediately behind the tip of the penis. These muscles
doubtless act in the same manner as in Dasypus, and serve to retract
the penis within the rudimentary prepuce. Beyond the statement
that the crus penis is invested by muscular fibres, Prof. Macalister
gives no account of the muscles just described.
1878. | AND DASYPUS SEXCINCTUS. 677
2. Dasypus SEXCINCTUS.
If now we compare the male organs of Chlamydophorus with those
of Dasypus sexcinctus, we shall find that the resemblance between
them is very striking. A full description of the male organs of D.
seacinctus after that already given of those of Chlamydophorus trun-
eatus would be superfluous, seeing that the one would to a large extent
be a mere repetition of the other; and I shall therefore confine my-
self to the indication of some points of interest in the anatomy of
Dasypus which appear to have been overlooked’. The penis is
enormous, and in a specimen measuring 13 inches in length, exclu-
sive of the tail, is 4 incheslong. It closely resembles that of Chla-
mydophorus in form, its extremity being simple and not bifid as in
the allied genus Vatusia. The integumental covering of the penis is
annulated in character, the annulation extending from the base to the
extremity of the organ on its dorsal or abdominal aspect, but inter-
rupted on its lower surface toward the tip of the organ by a cutaneous
patch presenting an altogether different appearance. This patch,
which is situated immediately behind the extremity of the penis, is
oval in form, and measures 14 inch in length. It surface is charac-
terized by the presence of numerous closely placed transverse ridges
of a hard and corneous nature. These ridges give to the patch
somewhat of the appearance of a fine file, and, in the absence of a
distinct glans, in all probability fulfil the function of the recurved
spines so frequently seen on the latter in many mammals. The pre-
puce is very small, and, when the penis is fully extended, disappears
entirely, being stretched-so as to form the cutaneous covering of the
latter. In the flaccid condition, however, and when the organ is re-
tracted, the penis is withdrawn entirely within the prepuce and almost
concealed from view. Improbable as it may appear that an organ of
the length of the penis can be really retracted within so small a pre-
puce, I am ina position to know that such is the case, having exa-
mined two specimens, in one of which the penis was exserted, as in
fig. 4, whilst in the other it occupied the position shown in fig. 5.
The retraction is accomplished by two retractor muscles, which,
arising from the lower surface and outer border of the sacrum, pass
forward along the whole length of the dorsum penis even to the tip,
fibres being inserted into every part of that surface. These muscles,
which exactly repeat those of Chlamydophorus, when contracting,
have the effect of throwing the penis into a series of spiral curves, its
extremity being bent inwards, as shown in fig. 5, so as to complete
the concealment of the organ under its sheath. That these muscles
are really the agents in producing this retraction is proved by the
fact that the concavity of the terminal curve of the penis is formed
by the dorsal aspect of the organ, on which these muscles are placed,
whilst the convewity of the curve is formed by its ventral surface
which is readily distinguished by the presence of the file-like patch
already referred to. The penis is formed, as in Chlamydophorus,
? Thaye been unable to consult Prof. Alexandrini’s monograph of Dasypus,
in Mem. Accad. d. Sci. Bologna, tom. vii. 1856.
678 DR. WATSON ON DASYPUS SEXCINCTUS. [June 18,
by the junction of the two crura alone, and possesses neither bulb
nor glans. With regard to the other muscles of the penis, these
closely resemble those already described in Chlamydophorus, with
this difference, that the single mass formed by the bulbo-cavernosi
muscles in the latter are here represented by two distinct and
powerful fleshy bands which envelop the under surface of each
crus previous to its junction with its fellow. The testicle differs
in position in Dasypus from that which it occupies in Chlamydo-
phorus, being attached to the inguinal region of the anterior ab-
dominal wall in the former, whilst in the latter it lies behind the
kidney. The globus major in Dasypus is closely applied to the
anterior extremity of the testicle, and does not form the loop-
like arrangement seen in Chlamydophorus. The vasa deferentia in
both are non-convoluted; nor is there any trace of vesicule semi-
nales in either. In both, a prostate and Cowper’s glands are
present, the former being traversed by the vasa deferentia in their
course to the urethra, whilst the latter, although occupying the same
position, are relatively considerably smaller in Dasypus than in Chla-
mydophorus. The bladder and urethra are similar in both genera,
the only difference being that the walls of the intrapelvic portion of
the latter are much more muscular in Chlamydophorus than in Da-
sypus, and that in the latter a well-marked caput gallinaginis, of a
pyramidal form, is present, which is not represented in the former.
In neither is there any trace of a vesicula prostatica.
It will thus be seen that in respect of the male sexual organs the
resemblance between Chlamydophorus and Dasypus seacinctus is very
striking—a resemblance which, in respect of other organs, has been
already pointed out by Prof Garrod’. As shown by that anato-
mist, Chlamydophorus agrees also with Tolypeutes and Xenurus ia
the absence of vesicule seminales and the simple condition of the
glans penis, in both of which respects it differs from Tatusia. The
presence of Cowper’s glands constitutes a further element of agree-
ment between Chlamydophorus on the one hand, and Dasypus and
Xenurus on the other. These glands are also present, according to
Dr. Murie’, in Tolypeutes ; but whether in Tatusia I cannot say, as
with regard to the latter we possess no precise information. So far,
then, as the observations above recorded go, they tend to strengthen
the conclusions already arrived at by Mr. Garrod upon other grounds
with regard to the close relationship which exists between Chlamydo-
phorus and Dasypus. Professor Macalister, as the result of his
elaborate investigation into the muscular anatomy of the Edentata,
arrives at the conclusion that ‘“‘the position of Chlamydophorus will
be seen..... to be plainly among the Dasypodidze, and very close
to Tatusia.”’ The arrangement, however, of the sexual organs,
including the nipples, together with that of the alimentary canal,
which in Dasypus and Chlamydophorus is provided with two colic
ceca, these last being absent in the other genera above referred
to*, appear rather to point to a closer relationship between Chla-
1 P. ZS. 1878, p. 222. 2 Trans. Linn, Soe. vol. xxx,
3 See Prof. Garrod’s paper.
1878.] PROF. A. H. GARROD ON THE ANATOMY OF PLOTUS. 679
mydophorus and Dasypus than between Chlamydophorus and Ta-
tusia.
I am indebted to Dr. Young for the preparation of the accom-
panying illustrations.
EXPLANATION OF PLATE XLITI.
Fig. 1. Male generative and urinary organs of Chlamydophorus truncatus, seen
from the front. From above downwards are seen the suprarenal cap-
sules, the kidneys and ureters, the testicles, the bladder and urethra, and
the penis. The penis is hooked back to show the retractor muscles on
its dorsal aspect.
2. Male organs of Chlamydophorus truncatus, perineal view. From
above downwards are seen:—the penis, its basal portion being covered
by the fibres of the bulbo-cavernosi muscles; below this the Cowperian
glands lying in the interspace between the crura penis, these last being
covered by the fibres of the ischio-cavernosi muscles,
3. Male organs of Chlamydophorus dissected, and seen from behind. K,
kidney and suprarenal capsule; T, testicle; P, prostate gland; ©,
Cowper’s gland; Pe, penis; Cp, crus penis.
4. Penis of Dusypus sexcinctus, showing the file-like structure on the lower
aspect of its extremity.
5. Penis of Dasypus sexcinctus retracted within the prepuce.
4. Note on Points in the Anatomy of Levaillant’s Darter
(Plotus levaillanti). By A. H. Garrop, M.A., F.R.S.
[Recgived June 14, 1878.]
In a former communication? I had the opportunity of bringing
before the Society several facts with reference to the anatomy of
Plotus anhinga, and of confirming Mr. Macgillivray’s account of its
most peculiar proventriculus. Several specimens of the species have
since passed through my hands which differ in no way from that first
described.
On the 9th of March last the Society obtained for the first time,
by purchase, a male specimen of Levaillant’s Darter (Plotus levail-
lanti) from Senegal. It unfortunately died on the 7th of this
month (May) from peritonitis, the result of a perforating ulcer in
the stomach.
The severity of the peritonitis caused all the abdominal viscera
to be agglutinated into a single mass, and rendered them particularly
soft. Nevertheless I was able to disentangle most of the alimentary
canal for examination; and it has proved of more than ordinary in-
terest, as the following description will serve to show.
The tongue, as a free organ, is obsolete. The cesophagus is capa-
cious, without any crop. The oesophageal epithelium ceases abruptly
by a transverse line where the gastric portion of the canal com-
mences, below which it is replaced by the tough yellow epithelium
so characteristic of the situation.
The proventriculus is composed of two circular areas of deep
glands, which latter are of considerable size and do not come into con-
£ P’ 7.8. 1876, p. 335,
PROF. A. H. GARROD ON [June 18,
680
tact, being situated laterally upon opposite sides of the gullet. These
areas are flat, except at their upper margins, where a small transverse
ridge is developed, which may be the slight foreshadowing of the
special cavity developed in Plotus anhinga for the reception of the
secretion of the gastric glands. But in P. levaillanti it is to be
noticed that there are two gland-surfaces, whilst in P. anhinga it
will be remembered there is but one gland-cavity.
5
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View of the inner surface of the posterior wall of the stomach of Plotus levail-
lanti, with the cesophagus slit up anteriorly so as to show the whole of its
proventricular surface, with the two gland-areas and the U-shaped elevation
on its anterior wall (on the left of the figure), as well as the pyloric
infundibuliform plug.
The calibre of the cessphagus must be somewhat diminished by
the presence of a curious U-shaped ridge upon the mucous mem-
brane of its anterior wall, which is situated between the antero-
lateral margins of the gland-areas, and has a small pit in the part
is J
1878.] THE ANATOMY OF PLOTUS LEVAILLANTI. 681
corresponding to the surface between the limbs of the U, apparently
not glandular in nature. All these structures are covered with the
tough gastric epithelium, which ceases just above them.
The first true gastric cavity is larger than the second, though not
much so. In the second the peculiar hairy covering of its pyloric
portion is largely developed, and in a different manner from what it
is in P. anhinga, where, as [ have shown in my paper on the anatomy
of that bird, it forms a kind of sieve to prevent large solid particles
from entering the duodenum. In P. devaillanti a more elaborate
arrangement obtains ; the hairy epithelium surrounding the pyloric
orifice, near the lower margin of the gastric surface of which it is
developed, is produced into a considerable conical hair-covered pro-
cess, projecting into the second stomach, and evidently acting as a
valve to close the pylorus wheu necessary. In general appearance
it much resembles the operculum of the Cheilostomatous Polyzoa,
and is very striking at first sight, the hirsute conical plug when re-
tracted, fitting exactly into the equally hirsute conical pyloric end of
the second stomach-cavity. All the rest of the second stomach is
lined with a non-hirsute epithelium, which ceases abruptly where
it meets the hairy surface. I can find no trace of this operculum in
Plotus anhinga, upon re-examination.
The small intestine measured two feet, and the large three inches ;
but they may have been contracted by the inflammation of their
surfaces. Two minute caeca were clearly seen, one a little larger
than the other. In P. anhiaga there is no indication of a second
czecum.
As in P. anhinga, P. levaillainti possesses but one carotid artery,
the left. In their myology the two species agree in every respect,
as far as I can see. In P. levaillanti the ambiens is large, groov-
ing the patella, the femoro-caudal is present without an accessorius,
as is the semitendinosus. There is a slip from the biceps of the
arm, which traverses the patagium; and the temporal muscles run
back beyond the skull, being separated by a median fibrous raphé,:
which is not ossified into a separate bony style. The great pectoral
muscle is formed of two layers.
Donitz’s bridge is ossified, as in the specimen described by the
author after whom it is named: it is developed on the ninth, and
not on the eighth cervical vertebra, as I predicted would be the case.
The lower larynx is indistinguishable from that of P. anhinga.
It is interesting to notice that the Manatee and Dugong have
special gastric gland-structures, the method of arrangement of which
differs in exactly the same way as does that of the two species of
Plotus under consideration, the peculiar flat gland-area found in
Halicore and Plotus levaillanti being converted into a glandular
eavity in Manatus and Plotus anhinga.
Proc. Zoou. Soc.—1878, No. XLIV. | 44
682 MESSRS. GARROD AND TURNER ON [June 18,
5. On the Gravid Uterus and Placenta of Hyomoschus
aquaticus. By Professors A. H. Garrop, M.A., F.R.S.,
and Wixtuiam Turner, M.B., F.R.S.
[Received June 11, 1878.]
(Plate XLIV.)
An adult female of Hyomoschus aquaticus having come into our
hands, it was with no small pleasure that on eviscerating it we found
it far advanced in pregnancy ; for it enables us to give an account
of the placenta, the nature of which has, till now, only been surmised
from what is found in Tragulus.
In his valuable memoir on the Tragulide’, M. Alphonse Milne-
Edwards briefly describes and also figures the foetus with the placenta
of Tragulus stanleyanus. He makes no mention of the uterus, of
which, in an allied species, John Hunter tells us” that it ‘soon
divides into two horns, which are pretty large and not long, having
none of the buttons for the cotyledons.”
In his paper on the visceral anatomy of Hyomoschus aquaticus *,
Prof. Flower describes the female generative organs in the following
words :—‘‘ The vagina was 5 inches in length; the‘uterus 3°5 inches
to the point of bifurcation, sharply bent back on itself near the upper
end, and terminated in a pair of rather short, closely curled
cornua.”
In our gravid specimen the single hairless foetus which, from tip
of nose to end of tail, measures 8°5 inches, the tail being an inch
long, is lodged on the left side.
The uterus consists of two horns communicating with a common
corpus uteri. The horns are united together in the greater part of
their extent, not more than about 1°5 inch of the tip of each horn
being free. The line of union is marked externally by a groove, and
internally by a broad partition, the septum uteri, which extends
longitudinally backwards and terminates in a well-defined semilunar
free border, behind which the two horns are fused together into the
common corpus uteri. The free ends of the cornua are curled back-
wards, and together with the Fallopian tubes and ovaries are situated
upon the anterior part of the superior wall of the uterus. Owing to
the foetus being developed in the left horn, this cornu is much more
dilated than the right ; but the latter is considerably more capacious
than in the non-gravid uterus. The corpus uteri communicates by
a constricted os with a passage which may perhaps be regarded as a
cervix, though some might look on it as only the specially modified
anterior end of the vagina. This part of the genital passage is 1?
inch long and very much constricted. Its mucous lining is longitu-
dinally folded; and the folds are at intervals so projecting as to give
the appearance of transverse constrictions. The passage and the os
1 Annales des Sciences Naturelles, 5th series, vol. ii. 1864, pp. 49-167.
2 «Wssays and Observations,’ edited by Prof. Owen, 1861, vol. ii. p. 135.
* P.Z.8 1867, p. 960.
HUSOWNOAH 40 AWOLVNY
(ueH NW UA yume '¢
.
1878. | THE PLACENTA OF HYOMOSCHUS AQUATICUS. 683
are blocked up by a whitish viscid mucus. Behind the most poste-
rior transverse constriction the vagina undergoes a considerable dila-
tation, and the mucous lining exhibits faint longitudinal folds.
The uterine walls are slightly thinner than those of the human
stomach. The cavity of the two cornua and of the corpus uteri is
lined by a well-defined mucous membrane, from which the foetal
chorion can readily be separated. This mucous membrane forms
the maternal portion of the diffused placenta characteristic of Hyo-
moschus. The mucosa of both cornua is not elevated into folds,
except in close proximity to the openings of the corresponding Fal-
lopian tubes; and the mucous lining of the corpus uteri is longitudi-
nally folded only in proximity to the os uteri. The free surface of
the mucous membrane, both in the cornua and corpus and on both
surfaces of the septum uteri, is soft and velvety and pitted with
multitudes of minute depressions just visible to the naked eye. These
depressions are the crypts in which the villi of the chorion are lodged
when the chorion is ia situ. The erypts are distributed with almost
equal regularity over the surface of the mucosa in the several divi-
sions of the uterus ; but on the more convex part of the impregnated
left cornu the mucosa is not quite so thick, so that the crypts are
shallower, and over a limited arga.the free surface of the mucous
membrane is almost, if not quite, free from crypts. We did not,
however, see any depressed circumscribed smooth areas surrounded
by crypts such as one of us has. described elsewhere * in the Pig and
Lemurs, or polygonal areas occupied by crypts and bounded by ridges
free from crypts, such as are to be seen in the gravid uterus of the
mare. In the regular diffusion of the crypts over the surface of the
mucosa, the gravid uterus of Hyomoschus much more closely re-
sembled what has been described in Orca gladiator? than it did the
uterine mucosa of the Pig, Mare, and Lemurs.
We then carefully stripped portions of the mucous membrane off
the subjacent muscular coat, and soaked them in glycerine for some
days, in order to render the membrane as translucent as possible.
When the mucosa thus prepared was examined microscopically, the
openings of the uterine glands on the surface of the membrane could
be seen. Sometimes these openings were found on the slender raised
folds of mucosa separating adjacent crypts from each other ; at other
times they opened into the crypts, and at other times on smoother
portions of the membrane where the crypts were shallower or almost
absent ; but in no case were the mouths of the glands specially
localized in smooth circumscribed areas of the mucosa, as is the case
in the Pig and in the Lemurs. ‘The gland-orifices were directed
obliquely to the plane of the free surface of the membrane ; and it
was not uncommon to see an epithelial plug projecting through the
mouth.
Additional views of the relation of the glands to the crypts were
obtained by making vertical sections through the mucosa. This
1 Turner, Lectures ‘“‘On the Comparative Anatomy of the Placenta,” Edin-
burgh, 1876, and Trans. Roy. Soc. London, 1876.
? Turner, Trans. Roy. Soc. Edinburgh, 1871.
44*
684 MESSRS. GARROD AND TURNER ON [June 18,
membrane consisted of a gland layer and a crypt layer. The gland
layer was next the muscular coat, and consisted of elongated tubular
glands, somewhat tortuous and occasionally bifureating. In the
vertical sections the glands were cut across so that the tubes were
sometimes transversely, at others obliquely, at others longitudinally
divided, and here and there the stem of a gland could be seen pass-
ing obliquely through the crypt-layer to open on the surface in the
manner already described. The glands were lined by a columnar
epithelium, and possessed a central lumen. The glands were neither
so numerous nor so distinct, neither did they bifureate so frequently
as do the utricular glands in the Pig and the Cetacea.
The crypt layer contained the numerous depressions already re-
ferred to for the lodgment of the villi of the chorion. The epithe-
lium lining the crypts had, as arule, disappeared ; so that it was only
in exceptional localities that it could be seen im sttu, where it ap-
peared to consist of cells, the type form of which was columnar,
though modifications of that shape occurred. The subepithelial
connective tissue contained a large proportion of corpuscles, some of
which were fusiform, others polygonal, others of the rounded form
of white blood-corpuscles. This tissue was more compact where it
formed the walls of the crypts; but deeper in the mucosa, as it ap-
proached the glandular layer and the muscular coat, it had an areo-
lated character. The vessels of the uterus were not injected ; but
there can be no doubt that, if they had been so, the walls of the
crypts would have been seen to contain an abundant freely anasto-
mosing network of capillaries, such as exist in the corresponding
crypts in the Cetacea, the Mare, the Pig, and the Lemurs. In
sections through the wall of the uterus, that had been stained with
heematoxylin, a well-defined band, coloured with the blue pigment,
marked the junction of the deep surface of the mucous membrane
and its glands with the muscular coat. This band in all probability
was the muscularis mucose. In Hyomoschus, as in other animals
possessing a diffused placenta, the uterine glands have no relation, as
regards numbers or termination, to the crypts. The crypts are in-
finitely more numerous than the glands, and are not to be regarded
as formed by a dilatation of their mouths, but are new formations
during pregnancy, due to hypertrophy and folding of the mucous
membrane so as closely to adapt it to the irregular villous surface
of the fcetal chorion.
The chorion extended from the tip of the left uterine cornu, through
the corpus uteri, to the tip of the right uterine cornu. The left horn
of the chorion, which contained the foetus, was longer and much
more capacious than the right horn. The tip of each horn of the
chorion was in close relation to the orifice of each Fallopian tube; and
close to the tip the free surface of the chorion was over a very limited
area smooth and non-villous. That part of the chorion situated in
the corpus uteri, immediately opposite the os uteri, presented a cir-
cular non-villous surface about the size of a shilling. This surface,
though without villi, was folded so as to adapt it to the correspond-
ing folds of the uterine mucosa in the.same locality. A portion of
1878. ] THE PLACENTA OF HYOMOSCHUS AQUATICUS. 685
the chorion corresponding to the more convex part of the gravid
uterine cornu was thinly covered with villi, and, indeed, in one or two
very limited areas was non-villous—these bare or thinly covered
patches being in contact with those portions of the uterine mucosa
(already described) where the crypts are either shallow or absent.
In all other localities the free surface of the chorion was as thickly
studded with villi as the uterine mucosa was with crypts; so that it
furnished an excellent and characteristic example of a diffused pla-
centa. In the absence of villi from those parts of the chorion which
were situated in relation to the three uterine orifices, 2. e. the os uteri
and the two Fallopian tubes, the chorion of Hyomoschus corresponded
with what one of us has elsewhere described in Orca, the Narwhal,
and the Mare.
The villi were arranged in small tufts, separated from each other
by very narrow intervals. The tufts varied in size; and the villi of
which they were composed were short and branched, usually in the
form of filamentous processes. The basal substance of each villus and
of the chorion itself consisted of a delicate corpusculated connective
tissue. The epithelium on the free surface of the chorion was partly
shed ; but considerable patches of it were seen in many localities.
The amnion formed a capacious ag, which extended to within half
an inch of the tip of the left horn of the chorion. It occupied the
rest of this horn and the part of the chorion situated in the corpus
uteri, but did not extend into the right horn of the chorion.
The sac of the allantois occupied the whole of the right horn of
the chorion, extending as far as its tip ; and its wall was in close ap-
position by its attached surface with the deep surface of the chorion.
The sac of the allantois was also prolonged into the left horn of the
chorion to within half an inch of its tip: but in other respects its
distribution in this horn was limited ; for instead of being in contact
with the whole extent of the deep surface of the chorion, it formed
only a circumscribed tubular prolongation attached to that part of
the chorion which was opposite the belly of the foetus. The remain-
ing part of the deep surface of the chorion in this horn was in con-
tact with the attached surface of the amnion, which membrane ex-
tended as far as the edge of the non-villous circular spot opposite
the os uteri. The amnion was also prolonged over the tubular pro-
longation of the allantois which extended into this horn. The
amnion enveloped the umbilical cord up to the abdominal wall of the
foetus. The amniotic investment of the cord was studded with
numbers of small stunted whitish sessile corpuscles, which did not,
however, project from the free surface of the amnion, where it was
in contact with the chorion and allantois. These bodies resembled
in form and size the bodies projecting from the umbilical cord in the
Cetacea. ‘hey had not, however, the yellowish-brown colour of the
corresponding corpuscles in Orea and the Narwhal, but like them
they were developed in relation to the deep surface of the amnion
and caused an elevation of that membrane by their growth.
The umbilical cord was 32 inches long. It contained two arteries
and two veins; and a slender tubular prolongation of the sac of the
686 MR. F. MOORE ON NEW HESPERIDZ. [June 18,
allantois could just be traced into the chorionic end of the cord.
Some injection was passed into the vessels of the cord, which filled
the principal vessels of the chorion ; but as the specimen had been
for some time in spirit before the injection was made, it did not
enter the vessels of the villi. Slender vessels, however, traversed
the non-villous spot opposite the os uteri, and some also were seen
subjacent to the amnion where it was in contact with the wall of the
tubular prolongation of the allantois situated in the left horn of the
chorion.
EXPLANATION OF PLATE XLIV.
Fig. 1. Foetal membranes of Hyomoschus aquaticus, opened into and foetus re-
moved; reduced from natural size. Ch. Villous chorion of the
foetal horn. Ck!, Villous chorion of the opposite horn. Al. Sac of
allantois in the non-fecundated cornu, the communication of which
with the tubular prolongation in the fetal horn is seen to the right
of the letters. Am. Sac of the amnion; the letters are placed on the
amnion where it is in contact with the chorion. U. Vhe umbilical
cord ; the letter is placed on the amnion where itis in contact with the
allantois. o. The non-villous surface of the chorion opposite the os
uteri. 8.8. The smooth non-villous parts of the chorion in relation
to the Fallopian tubes.
2. Magnified view of the free surface of the uterine mucous membrane of
Hyomoschus, showing the crypts in which the villi of the chorion are
lodged. At @ the mouth of one of the uterine glands is shown.
The drawings have been kindly made for us by Mr. J. D. Dunlop.
6. Descriptions of new Asiatic Hesperide.
By F. Moors, Assistant Curator, India Museum, London.
[Received June 11, 1878.]
(Plate XLV.)
IsMENE SUBFASCIATA, Ni. Sp.
Male. Upperside dark velvety umber-brown ; costal edge of fore
wing slightly ochreous; cilia of both wings ochreous. Underside
paler: fore wing with a pale pink triangular costal patch before the
apex ; posterior border ochreous, adorned with a large hairy tuft:
hind wing with a transverse pink fascia across middle of the wing.
Eyes red. Legs beneath and anal tuft ochreous.
Expanse 2 inches.
Hab. Ceylon (In coll. Sir W. H. Gregory and Dr. Thwaites).
Allied to I. aria, Moore, from which it may be distinguished by
the markings on the underside.
CALLIANA, D. g.
Wings ample, broad. Fore wing trigonal ; cell long, broad ; costal
vein extending to one third from apex ; subcostal vein curved at end
of the cell, first, second, third, and fourth branches arising at equal
distances before end of cell, fifth from its end, the three former ter-
NEW ASTATIC
Mintern Bro? i
Ee SPE FD AS.
ths
1878.Pl. XLV.
1878. ] MR. F. MOORE ON NEW HESPERID. 687
minating before the apex, fourth at the apex, and fifth below it; dis-
cocellulars angled close to the upper end; upper radial from the
angle, lower from their middle ; median vein three-branched, widely
apart, the upper from lower end of cell, middle and lower branches at
equal distances, the lower from near base of cell ; submedian slightly
recurved. Hind wing broadly ovate ; subcostal two-branched, second
before end of cell; discocellulars slightly angled; radial from their
angle; median three-branched, middle branch from immediately
before end of cell. Body small, robust, abdomen short. Palpi short,
thick, pilose, ascending, third joint short, conical. Antennee slender.
Legs pilose.
CALLIANA PIERIDOIDES, n. sp. (Plate XLV. fig. 2.)
Male. Upperside creamy-white, glossy ; fore wing with the apex
broadly vinous-brown, the end of the median veins also witha
vinous-brown-speckled spot: hind wing with a curved upper discal
decreasing series of five purple-black spots, the upper one large and
situated between the costal and subcostal veins; the end of the
veins also with a small vinous-brown-speckled spot. Underside
white; fore wing with the costal base, a patch beyond end of
the cell, and the outer border pale vinous-brown : hind wing with
a short black narrow streak at end of costal border, four large
subbasal and a curved discal series of seven purple-black spots, a
marginal series of broad vinous-brown spots, one at end of each vein.
Thorax and abdomen white ; collar, top of head, palpi, and tip of
abdomen pale ferruginous; tip of palpi and antenne black ; legs
pale ferruginous above, purple-brown beneath.
Expanse 22 inches.
Hab. ?N.E. Bengal (4. Grote). In coll. F. Moore.
HESPERIA NAROOA, n. sp. (Plate XLV. fig. 4.)
Male and Female. Upperside dark olive-brown: fore wing with
two prominent semidiaphanous yellowish well-separated spots at end
of the cell, a subapical series of three smaller spots, and an oblique
discal series of five spots: hind wing with two very small discal
spots in male, three in female. Underside dark olive-brown; fore
wing marked as above: hind wing with a prominent white spot at
upper end of the cell, and a curved discal series of four spots. Body
olive-green.
Expanse 1,%; inch.
Hab. Bombay (Wilkinson); Ceylon (Mackwood). In coll. F.
Moore and F. M. Mackwood.
Allied to H. contigua, Mabille ; markings similar, but both sexes
one third less in size.
HESPERIA KUMARA.
Hesperia kumara, Moore, Catal. Lep. E.I. C. i. p. 255 (1857).
Female. Upperside dark olive-brown ; fore wing with a transverse
discal series of seven yellowish semidiaphanous spots from before the
apex ; cilia pale brownish-yellow. Underside deep ochreous-brewn :
688 MR. F. MOORE ON NEW HESPERIDZ, [June 18,
fore wing marked as above, the lowest spot being yellow: hind wing
with a single discal indistinct yellow spot between the two lower
median branches. Palpi and body brown; tarsi ochreous.
Expanse 12 inch.
Hab. Canara (S. N. Ward). In coll. India Museum, London.
This species is also found in Ceylon, the male differing from the
female above and beneath in the first upper and lowest discal spots
being obsolete.
HESPERIA SERIATA, i. sp.
Allied to H. kumara. Male differs from the same sex of that
species in the discal row of spots being slightly smaller, and in having
an additional lower small spot (which is present only in the female
of H. kumara). Underside greenish brown; fore wing marked as
above, the lowest spot suffused.
Expanse 12 inch.
Hab. Ceylon. In coll. F. M. Mackwood and F. Moore.
Hesperia BEVANI, 0. Sp.
Male. Upperside dark olive-brown; cilia pale brownish-cinereous ;
fore wing with a single small, pale white, semidiaphanous spot at upper
end of the cell, three contiguous subapical spots, another below
these, and two larger spots below obliquely on the disk, a small spot
also very indistinctly visible on middle of submedian vein: hind
wing without spots. Underside greyish olive-brown ; spots slightly
more prominent than above: hind wing with a discal series of five
small somewhat indistinct white spots.
Expanse 12 inch.
Hab. Salween, Moulmain (Lieut. Bevan). In coll. F. Movre.
A male specimen of this species from Calcutta, collected by the
late Mr. Atkinson, is in the collection of Dr. O. Staudinger.
HESPERIA BADA, Ni. sp,
Allled to H. mangala.
Male and Female. Upperside dark brown, base of wings olive-
brown; cilia pale cinereous-brown: fore wing with two (in some
specimens three) contiguous subapical small white semidiaphanous
spots, and three spots below obliquely on the disk, the upper one the
smallest: hind wing with a discal irregular linear series of three or
four white semidiaphanous spots, more or less indistinct. Under-
side greyish-brown ; both wings marked as above, the spots on the
hind wing being more prominent.
Expanse, d 15%, 2 153; inch.
Hab. Ceylon. In coll. F. Moore.
Also in the collection of Dr. O. Staudinger from Malacca.
HESPERIA FARRI, 0. sp.
Male and Female. Upperside ferruginous-brown, base of both
wings olive-brown ; cilia yellowish-cinereous : fore wing with two pale
yellow semidiaphanous spots at end of the cell, and a discal recurved
1878. } MR. F, MOORE ON NEW HESPERID&. 689
series of seven spots, the four upper and the seventh smallest, the
upper three being contiguous and obliquely before the apex, the
sixth below end of the cell and the largest. Underside greyer-
brown in the female: fore wing marked as in male, except that the
lowest spot is more diffused : hind wing without spots.
Expanse, ¢ 14, ¢ 12 inch.
Hab, Calcutta (Farr); Cherra Punji (Atkinson). In coll. F.
Moore and Dr. Staudinger.
HESPERIA TOONA, nD. sp.
Allied to H. eltola, Hewits. Exot. Butt. Hesp. pl. 4. f. 40.
Male. Differs in the fore wings being slightly more elongate, the
hind wing more convex exteriorly, and the anal angle less lobed ;
markings above similar, those on the fore wing narrower, the spot
between the. median branches elongated and extending to their basal
angle ; those on the hind wing very small, the outer spot crossed by
a vein. Underside brownish-ochreous ; base of fore wing slightly
dusky-ochreous ; markings as above.
Expanse 15%; inch.
Hab. N.K. Bengal (4. Grote). ~in coll. F. Moore.
Hesperia SUBGRISEA, n. sp. (Plate XLV. fig. 5.)
Allied to H. gremius, Fabr. (Butler, Catal. Fabr. Lep. B. M.
psn; pl. 3..f.'7),
Male. Differs above in having the three oblique discal spots
smaller, the lowest appearing only as a few greenish-grey scales, the
upper subapical series of two and a single spot at upper end of the
cell very minute. Underside less grey in colour, the fore wing
marked as above; hind wing with an indistinct black cell-spot and
a discal series of spots.
Expanse 14 inch.
Hab, Ceylon. In coll. F, M. Mackwood and F. Moore.
PITHAURIA, 0. g.
Fore wing elongated, narrow ; apex pointed ; exterior margin very
oblique; hind margin short ; hind wing convex externally, lobular at
anal angle. Head and thorax very broad, robust; abdomen not so
long as hind wing. Antenne with a slender club and very long
whip-like tip. Venation similar to Pamphila.
PirHavuRIA MuRDAvA. (Plate XLV. fig. 13.)
Hesperia murdava, Moore, P. Z. 8. 1865, p- 764.
Hab. Darjiling.
HALPE, n. g.
Allied to Pamphila (P. sylvanus). Antennze with a more slender
club and longer hook at tip. Fore wing shorter; exterior margin
more convex; the discal oblique series of raised scales in male
shorter and broader, Head and thorax smaller; abdomen slender.
Veins similar, the lower median branch being nearer end of the cell.
690 MR. F. MOORE ON NEW HESPERID. [June 18,
Harr Crytonica, n. sp. (Plate XLV. fig. 9.)
Upperside dark brown; base of wings and body olive-brown ;
fore wing with two contiguous subapical small white spots, two
oblique discal small conical spots with deeply excavated outer border,
and a smaller spot at upper end of the cell; the male with an
oblique discal streak composed of broad raised darker brown scales ;
cilia brownish-cinereous, alternated with dark brown. Underside
dark brown, speckled with ochreous scales; fore wing with white
spots as above, and a small spot also below the discal series; an
upper submarginal row of indistinct pale ochreous spots; hind wing
with a broad transverse median and a maculated discal greyish-
ochreous band. Palpi, body beneath, and legs greyish-ochreous.
Expanse, ¢ 1,%;, 2 12 inch.
Hab. Ceylon. In coll. F. M. Mackwood and F. Moore.
HALPE BETURIA.
Hesperia beturia, Hewitson, Descr. Hesp. p. 36 (1868).
Hab. Calcutta district (Atkinson and Farr); Andamans (Rep-
storff ).
HALPE LUTEISQUAMA.
Pamphila luteisquama, Mabille.
Hab. Luzon.
HALPE DOLOPIA.
Hesperia dolopia, Hewitson, Descr. Hesp. p. 27 (1868); Exot.
Butt. Hesp. pl. 6. f. 60, 61.
Hab, Darjiling (Atkinson).
Havre RADIANS, n. sp. (Plate XLV. fig. 1.)
Male. Upperside luteous-brown, basal hairy scales yellow. Cilia
cinereous-white : fore wing with a pale yellow constricted spot at end
of the cell, and an irregular transverse centinuous discal band of
spots with their lower angles continued outward along the veins ;
hind wing with a yellow streak at end of the cell, and a short discal
band with outer rays. Underside paler, minutely speckled with
yellowish-white; fore wing as above, the hind margin being also
broadly yellow ; hind wing with a subbasal spot, all the veins, and
two (a median and a discal) transverse sinuous bands pale yellow.
Palpi, body beneath, and legs yellowish-white.
Expanse 13 inch.
Hab. Dhurmsala, N.W. Himalaya. In coll. F. Moore.
PAMPHILA PALMARUM, 0. sp. (Plate XLV. figs. 6 & 7, 2 3.)
Differs from typical Java specimens of P. augias, Linn., in its much
larger size, the male having the markings above paler yellow and
broader, the impressed oblique discal streak on the fore wing being
obsolete, the outer border of the discal band much less sinuous, and
the yellow colour not extending along the veins to the exterior mar-
gin of the wing.
1878. ] MR. F. MOORE ON NEW HESPERID&. 691
Female. Paler brown above, the yellow bands on the upperside also
paler, less prominent, and there being no yellow along the costal
border of fore wing, and no median streak from base of the hind
wing. Underside pale vinous-brown, this colour pervading the upper
portion of the discal on the fore wing and the entire discal band on
the hind wing. ‘
Expanse, ¢ 14, ? 14 inch.
Hab. Calcutta (Grote and Atkinson). In coll. F. Moore and Dr.
Staudinger.
“ Larva feeds on the date and cocoa-nut palms” (Atkinson, MS.
note).
PAaMPHILA BAMBUSA, n. sp. (Plate XLV. figs. 11 & 12,3 9).
Allied to P. augias, Linn., from typical Java specimens of which
it differs in its somewhat broader and less pointed wings. Markings
above similar, but more defined, the borders of the wings blacker,
the basal yellow streak on hind wing confined to a terminal spot at
the end of the cell, and the abdominal border black. On the under-
side the markings are also more clearlydefined and the interspaces
blacker.
Expanse, ¢ 13, 2 14 inch.
Hab. Caleutta (Frith and Atkinson). In coll. F. Moore and Dr.
Staudinger.
“ Larva feeds on the bamboo” (Atkinson, MS. note).
PAMPHILA SUBOCHRACEA, 0. sp.
Upperside glossy luteous olive-brown ; cilia yellowish-cinereous.
Male. Fore wing with two pale semidiaphanous spots at end of the
cell, three contiguous spots obliquely before the apex, three upper
discal spots, below which is a narrow white oblique streak or brand:
hind wing with three small yellow upper discal spots, the two lowest
small. Female. Fore wing with a lower or fourth discal spot, anda
small dot below the third spot; the spots angled outward: hind
wing as in male. Underside greenish-ochreous, brown on hind
border of fore wing and anal lobe ; marginal line brown and promi-
nent : fore wing with the lower spot diffused and white: hind wing
with the upper discal white spot large and quadrate; four spots
below in a slightly linear position, the upper spot indistinct ; a white
spot also at upper end of cell, and a smaller indistinct spot above it.
Expanse, ¢ 13, 2 18 inch.
Hab. Caleutta (Atkinson). In coll. Dr. Staudinger.
Allied to P. mathias, Fabr., but of larger size and more promi-
nently marked.
PAMPHILA BRAHMA, n.sp. (Plate XLV. fig. 8.)
Male. Upperside deep ochreous-brown, basal area ochreous. Cilia
cinereous-yellow: fore wing with a black-bordered white streak ob-
liquely below the cell, a semidiaphanous yellowish-white constricted
spot at end of cell, a small subapical streak of three spots, and two
large discal spots, the one above hind margin being ochreous: hind
692 MR. F. MOORE ON NEW HESPERIDZ. [June 18,
wing with an indistinct cell-spot and a discal series of three pale
yellowish, semidiaphanous spots. Underside greenish-ochreous ;
costal base of fore wing ochreous-red, hind margin broadly dusky
black and centred by a white streak ; markings as above, but less
distinct.
Female brown above, with less ochreous at base, the spots more
prominent. Underside as in male.
Expanse 12 inch.
Hab. Masuri, 7000 feet, N.W. Himalaya. In coll. Major Lang
and F. Moore.
Nearest allied to P. subhyalina, Ménétr., from China.
PAMPHILA SIVA, N. Sp.
Male. Allied to preceding (P. brahma). Darker brown above,
the base of fore wing and large lower discal spot only being ochreous,
upper semidiaphanous spots brighter yellow, the one between the
lower median branches broader; spots on hind wing prominent.
Underside brighter greenish-ochreous, base of costa not ochreous red :
spots on hind wing clear white, very prominent and with dark border.
Expanse 13 inch.
Hab. Khasia hills (G. Austen). In coll. F. Moore.
ASTICTOPTERUS OLIVASCENS, 0. Sp-
Allied to 4. unicolor, Brem. (Ménétr. Catal. Mus. Pet. Lep. i. t. 5.
f. 6), from Pekin and Hong Kong, but of larger size. Upperside
uniform glossy olive-brown. Female with a white, semidiaphanous,
narrow subapical spot crossed by the fourth and fifth subcostal
branches. Underside: male uniformly brown; hind wing sparsely
grey-speckled : female with subapical spot on fore wing as above,
the exterior margin grey-speckled ; hind wing with three ill-defined,
sparsely grey-speckled, transverse bands. Palpi, body, and legs
beneath grey.
Expanse 1,6; inch.
Hab. Salween, Moulmain (Lieut. R. C. Bevan); Darjeeling (At-
kinson and Dr. Staudinger). In coll. F. Moore.
CYCLOPIDES SUBVITTATUS, 0. sp.
Male and Female. Upperside dark vinous-brown ; extreme edge
of both the costa and cilia chrome-yellow. Female with an oblique
subapical small chrome-yellow streak, composed of three connected
spots crossed by two branches of subcostal vein. Underside: fore
wing vinous-brown, with the costal vein, subcostal veins along their
marginal end, and a narrow outer marginal band chrome-yellow :
hind wing chrome-yellow, with partly interrupted black streaks be-
tween the veins.
Expanse +4 inch.
Hab. Salween district, Moulmain (Lieut. R. C. Bevan), In
coll. F. Moore.
Note. Also in Mr. Atkinson’s collection, labelled Darjiling.
1878. | MR. F. MOORE ON NEW HESPERIDS. 693
CYCLOPIDES SUBRADIATUS, N. Sp.
Male and Female. Allied to C. subvittatus. Differs in being
larger, the fore wing above having a very indistinct yellow streak,
which is composed of two well-separated spots only. Underside
brighter-coloured, and the streak between the veins more clearly
defined.
Expanse-1 to 12 inch.
Hab. Khasia hills (Godwin-Austen). In coll. F. Moore.
ISOTEINON ATKINSONI, n. sp. (Plate XLV. fig. 10.)
Male. Upperside dark glossy olive-brown; cilia brownish cine-
reous, with a brown inner line and indistinct bars: fore wing with a
small yellow semidiaphanous spot at end of the cell, three smaller
contiguous spots obliquely before the apex, and two contiguous spots
obliquely on the disk. Underside speckled with ochreous-green :
fore wing marked as above: hind wing with a median discal curved
series of eight small prominent white spots, and a spot at end of the
cell.
Expanse 1,1}, inch.
Hab. Darjiling (Atkinson). In coll. Dr. Staudinger.
_—
IsOTEINON KHASIANUS, 0. sp.
Male. Upperside glossy ochreous-brown ; cilia brownish-cine-
reous, with a brown inner line and indistinct bars: fore wing with a
small silky-white semidiaphanous spot at end of the cell, three con-
tiguous spots before the apex and two on the disk. Underside
rufous-brown : fore wing marked as above; a small tuft of black
hairs on middle of hind margin: hind wing grey-speckled, a small
white spot at end of the cell, and a median discal curved series of
white dots, each surrounded by dark brown.
Expanse 1,); inch.
Hab. Khasia hills (Godwin-Austen). n coll. F. Moore.
A specimen of what I believe to be the female of this species
is in the collection of Dr. Staudinger, but without a locality (though
probably Indian). .It differs from the male above described
in having the spots on the fore wing slightly larger, the cell-spot
transversely elongated, and in there being an additional spot below the
two on the disk. It isalso greyer on the underside; the hind margin
on the fore wing has a very pale border; and the hind wing has no
perceptible white dots.
IsOTEINON MASURIENSIS, n. sp. (Plate XLV. fig. 3.)
Male and Female. Upperside bluish purple-brown. Cilia white,
alternated with brown: fore wing with a large semidiaphanous white
quadrate spot at end of the cell, two contiguous elongated spots
below it (and in the male a smaller spot below these), three very small
spots before the apex. Underside dark brown, numerously speckled
with ochrey-brown scales: fore wing marked as above: hind wing
with a discal series of three small indistinct white spots, one being
694 MR. F. MOORE ON NEW HESPERIDZ. [June 18,
between the subcostal branches, the others between the upper median
branches. Body and legs beneath speckled with ochreous-green.
Expanse 1} inch.
Hub. Masuri, N.W. Himalaya. In coll. Major Lang and F.
Moore.
“On clover, June, 7000 feet’ (MS. note).
THANAOS STIGMATA, 0. sp.
Male and Female. Upperside glossy olive-brown: fore wing with
a short black bar or brand of raised scales obliquely above middle of
hind margin, and a very indistinct upper discal slightly curved row
of six small pale spots; cilia whitish-cinereous, with slight brown
bars. Underside paler; costal border of fore wing, veins, and basal
interspaces of hind wing speckled with greenish-grey ; fore wing with
whitish discal maculated band as above, but more distinct, a spot
at end of the cell, and a marginal row of lunules less distinct ; hind
wing with a distinct whitish cell-spot and a submarginal and marginal
lunular band. Female without the raised bar, and the discal band
above less distinct.
Expanse, ¢ 1 inch, 2 5% inch.
Hab. Masuri (7000 feet), N.W. Himalaya. In coll. Major Lang
and F. Moore.
“Common in clover-fields ”’ (Lang).
THANAOS INDISTINCTA, N. Sp.
Upperside uniform olive-brown ; cilia edged with cinereous: fore
wing with a very indistinct grey-speckled submarginal and marginal
line : hind wing with indistinct grey-speckled veins, basal interspaces,
and two outer indistinct lunular bands. Palpi and body greyish-
white beneath.
Expanse 5, inch.
Hab. Salween, Moulmain (Lieut. R. C. Bevan). In coll. F.
Moore.
THANAOS OBSOLETA, N. sp.
Allied to 7. stigmata. Differs above in the absence of the short
black oblique bar or brand on the fore wing ; the maculated band
being slightly more prominent. Underside similarly speckled, the
bands on hind wing not lunular, but composed of a slightly broader
series of spots ; some spots also present around the cell-spot.
Expanse 14 inch.
Hab. Cherra Punji, Assam (Atkinson). In coll. Dr. Staudinger.
EXPLANATION OF PLATE XLV.
. Halpe radians, n. sp., p. 690.
. Calliana pieridoides, n. sp., p. 687.
. Isoteinon masuriensis 3, n. sp., p. 693.
. Hesperia narooa 8, n. sp., p. 687.
subgrisea 8,0. sp., p. 689.
—
Fig.
Ore 9 bo
1875. | MR. F. MOORE ON HAINAN LEPIDOPTERA, 695
Fig. 6. Pamphila palmarum 3, n. sp., p. 690.
palmarum Qn. sp., p. 690.
brahma, n. sp., p- 691.
9. Halpe ceylonica, n. sp., p- 690.
10. Lsotetnon atkinsoni, n. sp., p. 693.
11. Pamphila bambuse 3, n. sp., p. 691.
12. —— bambuse 9, n. sp., p. 691.
13. Pithauria murdava, p. 689.
7. List of Lepidopterous Insects collected by the late R.
Swinhoe in the Island of Hainan. By F. Moors,
Assistant Curator, India Museum, London.
[Received June 11, 1878.]
RHOPALOCERA.
Danaip#.
=
DANAIS LIMNIACE.
Papilio limniace, Cram. Pap. Exot. i. pl. 59. f. D, E.
Danals AGLEA.
Papilio aglea, Cram. Pap. Exot. iv. pl. 377. f. E.
DANAIS PLEXIPPUS.
Papilio plexippus, Linn. 8. N. i. 2, p. 767.
DANAIS CHRYSIPPUS.
Papilio chrysippus, Linn. 8S. N. i. 2, p. 767.
EUPL@A FELDERI.
Euplea felderi, Butler, P. Z. S. 1866, p. 275.
Allied to E. layardi, Druce, P. Z. S. 1874, pl. xvi. f. 1, from Siam ;
differing above in having all the spots on the fore wing larger, those
on the hind wing being smaller, and the inner row nearly obsolete
towards the anal angle.
SALPINX MINORATA, N. sp.
Male. Upperside dark brown: fore wing glossed with blue; two
small narrow bluish-white streaks beyond the cell, a less distinct
streak above the submedian vein, a submarginal series, and a mar-
ginal row of indistinct speckled spots: hind wing with pale canescent
subbasal patch and two marginal rows of small indistinct white spots.
Underside paler brown : fore wing with three purplish-white discal
spots, a smaller costal spot, and two marginal series of white dots:
hind wing with two marginal series of white dots.
Expanse 23 inches.
Hab. Hainan. In coll. H. Druce.
Allied to S. Aluyi, Moore, but of much smaller size; it is also di-
stinct from S. hobsoni, Butler, P. Z.S. 1877, p- 811, from Formosa.
696 MR. F. MOORE ON THE [June 18,
SATYRID.
LETHE EUROPA.
Papilio europa, Fabr. Syst. Nat. p. 500.
ZOPHOESSA MUIRHEADI.
Lasiommata muirheadi, Felder, Wien. ent. Monats. vi. p. 28.
MELANITIS ISMENE.
Papilio ismene, Cram. Pap. Exot. i. pl. 26. f. A, B.
MYCALESIS PERSEUS.
Papilio perseus, Fabr. Syst. Ent. p. 488.
MYCALESIS RUNEKA.
Mycalesis runeka, Moore, Cat. Lep. E.I. C. i. p. 234.
MycaLEsIs CHARAKA.
Mycalesis charaka, Moore, P. Z.S8. 1874, p. 566.
YPHTHIMA PANDOCUS.
Yphthima pandocus, Moore, Cat. Lep. E.I.C. i. p. 235; Hewits.
Trans. Ent. Soe. 1865, p. 290, pl. 18. f. 12.
YPHTHIMA ZODIACA.
Yphthima zodiaca, Butler.
ELYMNIAS HAINANA, 0. sp.
Male. Upperside brownish purple-black: fore wing with narrow
bright blue marginal spots: hind wing with outer border broadly
ferruginous.
Female. Dark ferruginous-brown ; disk of both wings indistinctly
brighter ferruginous ; narrow marginal spots on fore wing pale pur-
ple; small whitish submarginal spots on hind wing. Underside
with the basal portion numerously covered with dark brown strigze,
the outer borders paler and with less and narrower strigz ; the
apical grey patch on fore wing prominent; and costal spot on hind
wing small but distinct.
Expanse 2% inches.
Hab. Hainan. In coll. H. Druce and F. Moore.
Allied to the Bornean species Z. nigrescens, Butler, P. Z. 8. 1871,
p- 520.
PAPILIONIDZ.
PapriLio PAMMON.
Papilio pammon, Linn. 8. N. i. 2, p. 746, 3.
P. polytes, Linn, ib. p. 746, 2.
PAPILIO HELENUS.
Papilio helenus, Linn. 8. N. i. 2, p. 745.
a
1878.] LEPIDOPTERA OF HAINAN. 697
PAPILIO ACHATES.
Papilio achates, Cram. Pap. Exot. pl. 182. f. A, B.
PAPILIO MALAYANUS.
Papilio malayanus, Wallace, Trans. Linn. Soc. 1865, p. 59.
PaPILIo SWINHOET, 0. sp.
Allied to P. nomius, Esper, from S. India. Differs on the upper-
side in the fore wing having the costal bands broader, the submar-
ginal row of spots smaller and narrower; the hind wing is also
shorter, the upright black bands broader, the two pale spots beneath
them widely separated, and the marginal lunules much narrower.
On the underside the hind wing has the discal red maculated band
more heavily margined with black.
Expanse, ¢ 28, 9 3 inches.
Hab. Hainan. In coll. H. Druce and F. Moore.
PAPILIO AXION.
Papilio axion, Felder, Verh. nuplhoteGes xiv. p. 305 (1864).
P. euryphilus, Hiibner, Samm. exot. Schm.
PAPILIO MEGARUS.
Papilio megarus, Westw. Arc. Ent. ii. pl. 79. f. 2.
PAPILIO SATURATA, N. sp.
Distinguished from Indian specimens of P. panope by its intense
and uniform dark colour throughout the upperside, being of a much
darker tint than any specimens from India ; the marginal spots on
the fore wing small but very prominent. In the hind wing there is
an outer marginal series of broad quadrate ochreous spaces, a sub-
marginal row of very narrow brown-speckled ochreous angular lunules,
and an inner row of acutely angular, scarcely perceptible ochreous
marks. Underside also very dark-coloured ; the base of fore wing
black-tinted ; quadrate ochreous spots on outer margin of hind wing
very broad ; submarginal lunules white.
Expanse 4 inches.
Hab. Hainan. In coll. H. Druce.
MorpuHip2.
CLEROME EUMEUS.
Papilio eumeus, Drury, Ill. Exot. Ent. i. pl.. 2. f23:
NYMPHALIDS.
ATHYMA LEUCOTHOE.
Papilio leucothoé, Linn. S. N. i. 2, p. 780.
NEPTIS HAINANA, N. sp.
Near to N. eurynome, Westw. (Donov. Ins. of China, 2nd edit.
p. 66, pl. 35. f. 4), from China. Differs above in its more compactly
Proc. Zoo. Soc.—1878, No. XLV. 45
698 MR. F. MOORE ON THE [June 18,
disposed maculated bands, the fore wing having the discoidal streak
comparatively broader and its terminal portion more elongated at the
point ; in the hind wing the subbasal band is straighter and of equal
width throughout, the submarginal band being narrower. The un-
derside is of a much paler orange-yellow colour, and the bands less
black-bordered, being similar in both these respects to the Assam
species V. kamrupa, Moore.
Expanse, ¢ 27 inches.
Hab. Hainan. In coll. H. Druce.
NEPTIS RIHODONA, 0. sp.
Allied to N. hordonia, Stoll; differs above in the bands being
broader, those on the hind wing considerably so, in the male these
latter also having a prominently waved outer border. Both wings
in the male have a well-marked outer marginal band.
Expanse, ¢ 18, 2 2 inches.
Hab. Hainan. In coll. H. Druce.
ERGOLIS ALTERNUS, 0. sp.
Nearest allied to the typical #. ariadne from Java. Differs in
its more deeply excavated margins, the upperside being of a more
yellowish-ferruginous colour between the discal and marginal bands ;
the broad basal portion, the median discal band, and marginal band
of a deeper dusky ferruginous, the discal band also being distinetly
and darker maculated. Underside with broad dark dusky chestnut-
brown suffused bands.
Expanse 2 inches.
Hab. Hainan. In coll. H. Druce and F, Moore.
CYRESTIS THYODAMAS.
Cyrestis thyodamas, Boisd., Cuv. Rég. An., Ins. ii. pl. 138. f. 4,
Identical with Indian specimens.
CyYRESTIS COCLES.
Papilio cocles, Fabr. Mant. Ins. i. p. 7; Donov. Ins. Ind. pl.
Oe. ees
JUNONIA LEMONIAS.
Papilio lemonias, Linn. S. N. i. 2, p. 770.
JUNONIA LAOMEDIA.
Papilio laomedia, Linn. 8. N. i. 2, p. 772; Drury, Ill. Exot. Ent.
pl. 5. f. 3.
JUNONIA NONE.
Papilio enone, Linn. 8. N. i. 2, p. 770; Cramer, Pap. Exot. i.
pl. 35. f, A-C.
One specimen from Ho Ching Shang, agreeing in size with those
from India, but smaller than the Chinese.
1878. ] LEPIDOPTERA OF HAINAN, 699
JUNONIA ALMANA.
Papilio almana, Linn. S.N, i. 2, p- 769; Cramer, Pap. Exot. i.
pl. 58. f. F, G.
SYMPHZDRA PARDALIS, n. sp.
Allied to S. dirtea, Fabr. Differs in the male having somewhat
shorter wings, the fore wing more convex on the costa, the outer
margin being slightly concave, and the antennz are shorter and less
robust. Distinguished on the fore wing above by the broad greenish
grey marginal band, which is uniformly coloured and extends to the
apex ; the band on the hind wing is also comparatively broader, the
snbmarginal spots larger, and itrorated with greenish-grey anally.
The underside is more uniformly ferruginous, but with the spots
aler.
j Female with the spots above of a greenish-yellow tint.
Expanse, ¢ 33, 9 35 inches. “Ti
Hab, Hainan. In coll. H. Druce.
DIADEMA Avia.
Papilio avia, Fabr. Ent. Syst. iii. 1, p. 111.
Papilio iacintha, Drury, Til. Exot. Ent. ii. ple 21. f. 1, 2.
ATELLA PHALANTA. :
Papilio phalanta, Drury, Ill. Exot. Ent. i. pl. 21. f..1, 2.
P. columbina, Cram. Pap. Exot. iii. pl. 238. f, A, B.
MESSARAS ERYMANTHIS.
Papilio erymanthis, Drury, Ill. Exot. Ent. i. pi 15. 7.3, 4
CETHOSIA BIBLIS.
Papilio biblis, Drury, Ill. Exot. Ent. i. pl. 4. f, 2:
Pirrip2.
NYCHITONA XIPHIA.
Papilio xiphia, Fabr. Spec. Ins. ii. p. 43.
P. nina, Fabr. Ent. Syst. iii. 1, p. 194.
CATOPSILIA GNOMA.
Papilio gnoma, Faby. Syst. Ent. App. p. 808.
TERIAS Esiore.
Lerias esiope, Ménétr. Cat. Lep. Mus. Petr. p. 85, pl. 2. f. 3.
TERIAS SUBDECORATA, n. sp.
Upperside deep yellow: fore wing with a moderately broad blackish
border, the inner margin of which forms a slightly sinuous curve in
the male, but a more angled curve in the female: hind wing with a
slight speckled spot at end of the veins. Underside similarly but
more prominently marked, as in 7. @siope, Ménétr.
Expause, ¢ 18, 9 13 inch.
8
Hab. Hainan. In coll. H. Druce.
700 MR. F. MOORE ON THE [June 18,
TERIAS ATTENUATA, 0. Sp.
Male. Upperside with a very narrow black border, its inner mar-
gin but very slightly sinuous; costal margin blacker than in the
other species: hind wing with a very narrow indistinct speckled
border. Underside with markings similar to T. esiope, but smaller
and narrower.
Expanse 14 inch.
Hab. Hainan. In coll. F. Moore.
TERIAS ARCUATA, 0. Sp.
Male. Allied to T. subdecorata. Differs above in having a more
regularly curved inner margin to the black border of the fore wing,
a nearly eontinuous speckled border on hind wing, and in the mark-
ings beneath being nearly obsolete.
Expanse 1? inch.
Hab. Hainan. In coll. F. Moore.
TERIAS HAINANA, N. Sp.
Male and Female. Allied to T. drona, Horsf., fram Java. Differs
on the fore wing in the marginal band being narrower and slightly
broken at its termination on posterior angle ; the hind wing has the
band very narrow and sinuous, being nearly obsolete at the anal end
in the female ; cilia pinkish. Underside paler yellow ;. two: promi-
nent small black spots at end of the cell on fore wing and dusky
speckled markings on hind wing.
Expanse 1,4) inch.
Hab. Hainan. In coll. H. Druce.
A distinct species from 7’. vagans, Wallace, P. Z. 8. 1866, p. 357.
THESTIAS SESIA.
Papilio sesia, Fabr. (Donov. Ins. China, pl. 31. f. 2).
Papilio evippe, Drury, Ill. Ins. pl. 5. f. 2, 3.
Papilio pyrene, Cram. ii. pl. 125. f: C, 3.
APPIAS ZELMIRA.
Papilio zelmira, Cram. iv. pl. 320. f. C, D, ¢.
APPIAS AMASENE.
Papilio amasene, Cram. Pap. Exot. i. pl. 44. f. A.
APPIAS COPIA.
Pieris copia, Wallace, Trans. Ent. Soc. 1867, p. 340.
One specimen (male) froni Ho Ching Shang.
APPIAS INORNATA, N. sp.
Allied to 4. vacans, Butler, from Sikkim.
Male. Upperside with the markings similar to 4. hippo (Cramer,
pl. 195. f.B,C). Female as in A. vacans, having the costal border of
fore wing, and outer borders of both wings only, black. Underside
of male with the apical spot on fore wing and the hind wing deep
1878. ] LEPIDOPTERA OF HAINAN. 701
yellow, the former absent and the latter paler in the female; outer
border of hind wing narrow, and of a golden-brown colour.
Expanse 2} inches.
Hab. Hainan. In coll. H. Druce.
This is quite a distinct species from the Formosan (4. formosana),
described by Mr. Wallace in P. Z. §, 1866, p. 356.
SyNCHLOE CANIDIA.
Papilio canidia, Sparrm. Ameen. Acad. vii. p. 504 (1768).
P. gliciria, Cramer, pl. 171. f. E, F.
Tuyca PAsiITHO:.
Papilio pasithoé, Linn. 8. N. i. 2, p. 759.
P. dione, Drury, Ul, Exot. Ent. ii. Bl, Sto
eo
ERyYcINID&,
SospirTa saTURATA, n. sp.
Form of outline as in S, lydda, Hewits. Exot. Butt. iii. Sosp.
pl. fig. 13, from Hong Kong. Upperside dark purple-brown, basai
half darkest: hind wing with very indistinct small black white-
bordered apical and anal spots. Underside brighter ; pale bands
distinct, the discal band broad, nearly straight on fore wing, and
its inner border darker-coloured ; three small white-bordered black
apical and two anal spots on hind wing.
Female brighter-coloured.
Expanse 1 inch.
Hab. Hainan. In coll. H. Druce and F. Moore.
ZEMEROS CONFUCIUS, n. Sp.
Z. confucius, Wallace, MS.
Male. Smaller than Javan or Indian specimens of Z. flegyas.
Markings above similar; tlie subapical spots larger and indistinct,
two only being visible above, these being white and prominent be-
neath,
From Z. albipunctata, Butler, from Malacca, it may be distin-
guished by the much less prominent and smaller number of subapical
spots both above and beneath, and in the inner series of spots on
both wings being white-bordered as in Z. Jlegyas.
Expanse 1,4; inch.
Hab. Hainan. In coll. H. Drace.
LycznIpD&.
MILETUS CHINENSIS.
Miletus chinensis, Felder, Verh. zool.-bot. Ges. xii. p. 488 ( 1862).
Lucia pinama, n. sp.
Male. Similar to L. evius (Westw. Gen. D. L. p. 902, pl. 76.
f. 5) from India, Underside of a browner grey, the transverse zig-
702 MR. F. MOORE ON THE {June 18,
zag markings less defined, and the white interspace beyond the end
of the cell much wider.
Expanse 1 inch.
Hab. Hainan. In coll. H. Druce.
PITHECOPS NIHANA, N. Sp.
Upperside dusky brown; cilia white: fore wing with a slight
white brown-speckled longitudinal central patch, and hind wing with
a similar streaky subapical patch. Underside white: fore wing with
a submarginal series of six very narrow, dusky, short, recurved marks,
and a marginal series of narrow dusky lunules bordered within by
an indistinet Iunular line: hind wing with a moderately large black
costal spot before the apex, and a smaller spot on abdominal margin ;
a series of very indistinct dusky submarginal and marginal streaks.
Expanse +} inch.
Hab. Hainan. In coll. H. Druce and F. Moore.
PoLYO@MMATUS LAIUS.
Papilio laius, Cramer, Pap. Exot. iv. pl. 319. f. D, E.
POLYOMMATUS VARUNANA.
Polyommatus varunana, Moore, P. Z. 8. 1865, p. 772, pl. 41. f. 6.
POLYOMMATUS SIMILIS, 0. sp.
Allied to P, argia, Ménétr. Cat. Lep. Mus. Petr, p. 125, pl. x. f. 7.
from Japan. Differs above in being of a pale lilac-blue, the margin
of the fore wing less dusky, and the hind wing with the marginal
spots searcely perceptible (both these characters in P. argia are very
prominent), On the underside the colour is of a brownish-grey ;
markings similar, but those of fore wing less prominent, the spots
on the hind wing having the e¢entre of the same tint as the ground-
colour of the wing.
Expanse 1,'5 inch.
Hab. Hainan. In coll. H, Druce and F. Moore.
LAMPIDES BA&TICUS.
Papilio beticus, Linn. 8. N. i. 2, p, 789.
LAMPIDES KANDARPA.
Lycena kandarpa, Horsf. Cat. Lep. E.I. C. 1829, p- 82.
APHN ZUS LOHITA.
Amblypodia lohita, Horst. Cat. Lep. E.I. C. 1829, p, 106,
ILERDA BELENA.
Heliophorus belenus, Hiibner, Zutrage, f. 785 (1832).
MAHATHALA, n. g.
Wings broad: fore wing somewhat short, exterior margin nearly
erect and slightly scalloped ; third subcostal branch biftieate; hind
1878. ] LEPIDOPTERA OF HAINAN. 703
wing short ; costa abbreviated, lobed at the base, concave towards
the end, the apex produced and pointed upward ; exterior margin
slightly concave below the apex, and very convex in the middle ;
anal anzle lobed ; a spatulate tail extending from end of lower me-
dian vein; palpi long, stout, apical joint broad laterally ; antennz
thickened to apex.
Differs from typical Amblypodia (A. narada, Horsf.) in the en-
tirely different form of wings ; the fore wing having the third branch
of the subcostal vein bifurcate instead of trifurcate, as in male of
A, narada.
MawaTHALa Ameria.
Amblypodia ameria, Hewits. Cat. Lyc. B. M. p. 14, pl. 8. f. 85.
(Harr BETURIA. a
Hesperia beturia, Hewits. Dese. Hesp. p. 36 (1868).
Hesperip2.
Hesperia GREMIUs. ;
Hesperia gremius, Fabr. Ent. Syst. Suppl. p. 433; Butler, Cat.
Fabr. Lep. B. M. p. 271, plan fy.
HI. divodasa, Moore, P. Z.S. 1865, p. 791.
Hesperia matutas, Fabr. Ent. Syst. Suppl. p. 433; Butler,
Cat. Fabr. Lep. B. M. p. 275, pl. 3. f. 8.
The specimens from Island of Ho Ching Shan.
Hesperia MANGALA, Moore, P. Z. S, 1865, p. 792.
Hesperia QUINIGERA, n. Sp.
Male and Female. Upperside olive-brown; cilia pale cinereous-
brown: fore wing with five small pale semidiaphanous spots, two
only being subapical, and three obliquely discal : hind wing with a
single indistinct pale semidiaphanous spot. Underside pale ochre-
ous-brown ; fore wings marked as above.
Expanse, ¢ 1,2, 2 1545 inch.
Hab. Hainan. In coll. H. Druce and F. Moore.
GEGENES HAINANUS, n. sp.
Male. Allied to the European species G. nostrodamus, but smaller
in size, also much paler above and less grey beneath.
Expanse 1} inch.
Hab. Hainan. In coll. H. Druce.
PAMPHILA CONFUCIUS.
Pamphila confucius, Felder, Wien. ent. Monat. 1862, p. 29.
ISMENE RIA.
Ismene aria, Moore, P. Z. S. 1865, p. 784; Hewits. Exot. Butt.
iv. Hesp. pl. 2. f. 24,
704 MR. F. MOORE ON THE {June 18,
CYCLOPIDES CAMERTES.
Cyclopides camertes, Hewits. Desc. Hesp. p. 43 (1868).
HETEROCERA.
BoMBYCES.
SYNTOMID&.
RHAPHIDOGNATHA HAINANA.
Artona hainana, Butler, Journ. Linn. Soe. xii. p. 357.
CHALCOSIID&.
Ptworvs GLAUCOPIS.
Phalena glaucopis, Drury, Exot. Ins. ii. p. 11, pl. 6. f. 4.
CHALCOSIA NYMPHA, 0. sp.
Mate. Fore wing silvery-white; a circular purple-black spot in
middle of the cell, and a speckled spot obliquely below it; apical
band purple-black, traversed with white spots: bind wing with a
black-streaked pale blue apical band, the extreme margin and cilia
being white. Head and collar crimson; body metallic green ; an-
tenne black ; legs grey.
Expanse 144 inch.
Hab. Hainan. In coll. F. Moore.
Allied to OC. diana, Butler, P. Z.S. 1877, p. 815, from Formosa.
EvuscHEMA MILITARIS.
Phalena militaris, Linn. 8. N. 1. 2, p. 811.
LiITHOSIID.
UrerHeEsiA PULCHELLA.
Phalena pulchella, Linn. S. N, i. 2, p. 884; Guér.-Meén. Icon.
Rég. An. pl. 88. f. 8.
BoMBYCID.
ARISTHALA, Noy. gen.
Fore wing elongate, trigonate, falcate ; costa moderately arched
at the end; apex convex; exterior margin oblique, concave below
the apex, slightly produced and angled in middle; posterior margin
nearly straight, angle acute; costal and subcostal with its first and
second branches running close together along the extreme margin,
third branch trifureate at nearly equal distances, the two upper
ascending to the costa, the first before the second to the apex, the
third to below the apex ; discocellulars bent inward at the middle,
upper shortest ; upper radial as a subcostal branch beyond the cell,
lower radial from angle of discocellulars ; a discoidal veinlet emitted
within the cell paraltel to lower radial ; median vein three-branched
at equal distances, upper from end of the cell; a submedian and
internal vein close to the margin. Hind wing elongated, narrow, apex
1878. } LEPIDOPTERA OF HAINAN. 705
and exterior margin very cunvex ; anal angle concave in male, angled
in female ; abdominal marginal long, nearly straight, fringed; two
subcostal branches emitted beyond the cell; discocellulars very
oblique, bent outward ; one radial from their angle; median three-
branched, two upper from end of cell ; a submedian and internal
vein. Body stout, with a dorsal crest and anal lateral tuft ; antennze
short, bipectinate to near end; palpi very small; legs very hairy
aud tufted to end of tarsi.
This genus is allied to Ocinara and Ernolatia, Walker, to Tri-
locha, Moore, and to Bombyx (B. mori, Linn.).
ARISTHALA HAINANA, 0. sp.
Female. Fore wing dark ferruginous ; posterior border brighter
ferruginous ; discal area speckled with buff and-purple-brown scales ;
an oblique curved subbasal and two discal sinuous darker lives; a
small bright-coloured ferruginous spot at end of the cell; exterior
border speckled with chalybeous-white : hind wing yellowish-ferru-
ginous, with short basal, median, and narrower discal purple-brown
bands; a subanal lunular white streak bordered above with purple-
brown. Thorax dark ferruginous; abdomen greyish-ferruginous ;
head, legs, and body beneath brighter ferruginous. Underside
bright ferruginous: fore wing palest at base: exterior border
dusky and white-speckled; one blackish transverse discal line:
hind wing with outer curved discal line, two discocellular spots, and
indistinct median fascia ; anal angle speckled with dusky-brown and
white.
Expanse 2 inches.
Heb. Hainan. In coll. F. Moore.
A closely allied species of this genus, from which the generic
characters of the male have been taken, is found in the Sikkim
Himalayas.
Liparip2.
PANTANA AMPLA.
Pantana ampla, Walker, Cat. Lep. Het. B. M. iv. p. 820.
PyYRALEs.
LEPYRODES GEOMETRALIS.
Lepyrodes geometralis, Guén. Spec. Gén. Lép. Delt. et Pyr. p. 278.
706 MR. F, MOORE ON THE [June 18,
Tabular List showing Geographical Distribution.
l
Sb :
2 | 4 4
1 6 |
B i Far:
Sieh e a |=
colic Cam He os S|
g § Sa S| z= ica]
“| altel lPSilaae hes
a) Ss )\Ola ts |e
DaANAID#. | |
Danais limniace * ye Be
apleai: <3 wgeaseeedas *
—— plexippus...... * | * | * | *
chrysippus ... * | * | # | ®
Euplea Felderi * | *
PalPierWHNOALA. weedoses os se cesses. cones *
SavyrIDz.
DEBISVEUO PaAwasoss- seca slewlsen se waceccioaise ones % tude al tate meckall ge *
Zophoessa muirheadi .....-...s++seeeeeee ees Tia Zeb TH af
MGiAmttiS ISMONG Toc eccatss sacnpendandescersne * | « | x | x | *
Mycalesis perseus..... ooh x | *
WUDGKS s.h03:. 0002 * *
———charaka ......... sant | G8
Yphthima pandocus *
zOdiaca..........- *
Hlymnias hainana ............00-s6000-
PAPILIONIDA,
Papilio pammon ............... Petceeebe atic Sa ba Sal ee
WPGVONIUB Se oes ee eoes Pete teen eae ee) iE
ere BCH ALES SA iiis voc Rae nled secanechoeeelees * * | x
———— NALAVALNIS! \seeciseecuwoscesceccupcemsaterec cl) oar oe ns cael! os
swinhoei *
PRION. fies ashes ay -U9e oH coz, csiewi eine, Meee aE
—— megarus * ¥
saturata ... * |
MorrpHip%. |
Olerome OumMOUS. 4... sd0c..c0nseeeeoaee ech BA Scan} 36
NyMpPHALID2£.
Athyma leucothoé ............ Ae ye 2 x | «| «| « | x | x
Neptis hainana...... *
TINA OWA ei aatascwavseocles «ceccsccoree.s *
Ergolis alternus ................ a eacosse * }
Cyrestis thyodamas ...............0.-0s0s0008- WD SET nes, Tee geen re
Cote Aa ride aac deecte oeki as Hakee TS « * | Se
Junonia lemonias.......... x | « | « | * | # | x
Taomiedis.....ace-sssce * | * | * * | *
—— None ..........0e eee * co | nce eee ee
—— almana............. Re Meetaaeaceimeet veo * * * * *
Sympheedra pardalis ..............:.00eeee *
IAC GRAN AVIRA: Siaceccchins soaee sv oncateres tee 3 1] gael aecall ae
Atellambhalantaly 7: 0t:nsctlte oscastsvaSsescs et doe [re |) Se gee
Tabular List (continued.)
78.] LEPIDOPTERA OF HAINAN.
rs
3s
a
a
an
S
io}
q
&
3
NympuaLipe (continued).
Messaras erymanthis. ...............002+ee0e *
Cethosia biblis *
Pinrips.
Nychitona xiphia *
Catopsilia gnoma ‘ *
Terias wsiope ...........- WASTES tn eepe 12 sae *
AUIDUCCOLALA a. codedtac cache sso ves seace ene *
BIGOHU A Goce. t note deamancene.(iesseeer eae *
BRC Ub os tec neces cee *
hainana .......- Srrerse *
PHeRHAS BESIG: ”ss..s demawclageces *
Appias zelmira................6- *
amasene *
copia........ *
inornata *
Synchloé canidie .............ceceeeeseeeneteee *
Thyca pasitho€...........c020ceseseeeeeasceeee %
Erycinip&.
Sospita saturata ........sccleecceesceeceee ees *
Zemeros CONFUCIUS .........0.eeeereeeereee ees *
Lyce&nip4,
Val obus (Chin engi sc .vese clus anne aicceeush eres *
Lucia dilama .......... *
Pithecops nihana...... *
Polyommatus laius . *
yarunana........ *
similis .......... 1%
Lampides baeticus ..........0..0sseeneeeeeees *
Pee Aes sera tease ra se aavawanes es See (ee
Aphneus lohita ....... Saithhec net wn doeaea tots *
lord belenas « pye.csjc-spnsconsapeeess seer ensc« *
Mahathala ameria ..,...........+ BRT soecee® *
Hesparip£.
Hesperia gremius...... oO
beturia....... *
— mathias *
mangala *
quinigera ... eae + a
Gegenes hainanus.......00...cseeeeceeceeeeeees *
Pamphila) confuctus......5-5.02.---..5-+25e+20- | *
HRMIENOPAMIG cesses -asscertecsseesses | *
| Cyclopides camertes ............. raccehice wal
07
Z é
B r=)
(=) n
fe ‘=|
ie & | 8
° ; Aa iE
a=} a . >| a
Sh) te ele fetes ;
3 ‘a =I &
a | | es - gs
= je) wm = A
% * *
* * *
ean We: * * *
% *
* *
*
* * *
*
*
* * *
*
*
A huis * *
* | * * *
wae da oe * *
Fhe 2 Spee *
| x | *
*
*
* *
* * * *
*
* *
* *
708 THE MARQUIS OF TWEEDDALE ON [June 18,
Tabular List (continued).
Sg a
a |g i=
ale A
cm] & a | a
&
© || co a |S
ei || ere] os i bl
@/s/-8/8/3/a
coche 2
SE SNS) ea) |p
Bompyces.
Rhaphidognatha hainana ............+++... *
IPIGOUS PIAMCOPIS! ioeccse<geecmsee ce ser osama * * *
Chalcosia nympha ...............s0000ceerees *
Huschema militaris ......,.....0.....s0eeesee % [oe | * * | *
(Utethesia pulchella: oo... ice. sesceeesessens ai ee wel aeRO alee (83
APISG Mele DAINATIAl A -Sc.nescctseccscesnecs eee ®
IPanitanayamiplaressnactasdarsccctcessvocecesasee * *
PyRALEs.
Lepyrodes geometralis .................442- x [-* | % x | *
8. Contributions to the Ornithology of the Philippines.—
No. X. On the Collection made by Mr. A. H. Everett
in the Island of Bohol. By Arrnur, Marquis of
Twreppate, F.R.S., President of the Society.
[Received June 14, 1878.]
The island of Bohol lies between Leyte and Zebu, and is situated
about seventy miles N.W. of Mindanao. It has a length of about
forty miles and a breadth of thirty miles. After leaving Leyte, Mr.
Everett stopped for a week at Talibon, on the north coast of the
island, and then left for Palawan, while his brother proceeded to the
interior of the island to collect. There he found a country covered
with grass 12 feet high, and with no forest except on the tops of a
few hills. Birds were scarce; and he had to return stricken with
fever. The collection obtained in North Bohol contains representa-
tives of 47 species; and although all belong to previously known
birds, seven of them have not been hitherto recorded as being
inhabitants of the Philippines. These are :—
Cisticola cursitans. Limosa lapponica.
Alauda wattersi. Tringa albescens.
Terekia cinerea. Strepsilas interpres.
Limicola platyrhyncha.
The rest of the species are generally distributed throughout
the archipelago, with perhaps the exceptions of Lanius nasutus,
which appears to be a northern form, and Orthotomus frontalis and
Loriculus hartlaubi, which are southern species.
1878. ] THE ORNITHOLOGY OF THE PHILIPPINES. 709
1. Lorrcutus wartiavusi (7) ?
[Bohol, 2, October. ]
Only one example of a female, and not in full dress. It agrees
with individuals of true Z. harélaubi in a similar stage of plumage,
but has the maxilla shorter by one tenth of an inch.
2. HaLiAsTuR INTERMEDIUs (17).
[N. Bohol, g 2, October. |
3. Burastur rnpicus (20).
[N. Bohol, 2, October : iris golden-yellow ; bill black; cere and
base of bill bright chrome-yellow; legs and feet dirty chrome-
yellow ; nails black. ]
~~"
4. Merops pHILIPPiNus (35).
[N. Bohol, 3, October.]
5. ALCEDO BENGALENSIS (38).
[N. Bohol, 2, October. ]
6. SAUROPATIS CHLORIS (47).
[N. Bohol, $ 2, October. ]
7. CACOMANTIS MERULINUS (57).
[N. Bohol, 3, October: iris light red ; bill biack; base of man-
dible reddish ; nails black ; feet and legs waxy Naples-yeliow. ]
The above note pertains to the adult. Two other examples are
in rufous and brown-banded plumage.
&. CenTRococcyx vrripis (64).
[N. Bohol, ¢ 2, October.]
The male is immature; the female adult.
9. Lanrus nasutus (70).
[N. Bohol, ¢ 2, October and November. ]
10. Lanrus LucroneEnsts (72).
[N. Bohol, ¢ 2, October and November. |
11. ARTAMUS LEUCORHYNCHUS (73).
[N. Bohol, ¢ 9, October and November. |
12. LaLacGe pominica (76).
[N. Bohol, ¢, November. ]
13. Leucocerca nicRitorauis (83).
[N. Bohol, ¢ 2, October. |
14. HrrunpDo JAVANICA,
Hirundo javanica, Sparrm. Mus. Carls. t. 100.
[N. Bohol, ¢ 2, October. ]
710 THE MARQUIS OF TWEEDDALE ON [June 18,
15. BRopDERIPUS ACRORHYNCHUs (90).
[N. Bohol, 2, October and November. |
16. Ixus GorAvIER (99).
[N. Bohol, 2, October. |
17. Pratrncoua cCapRATA (104).
[N. Bohol, ¢, November. Locust in gizzard. }
18. CisTICOLA CURSITANS.
Prinia cursitans, Franklin, P. Z. S. 1831, p. 118; Dresser, B. of
Europe, parts I]xi., Lxii.
[N. Bohol, 3, October: iris yellow-brown. |
Four examples identical with Indian individuals, but distinct from
the Negros example referred to anted, p. 285. no. 31.
19. ORTHOTOMUS FRONTALIS.
Orthotomus frontalis, Sharpe, Ibis, 1877, p. 112, t. i. f. 1;
T. L.S. ser. 2, Zool. i. p. 336. no. 85.
{N. Bohol, ¢ 9, October. |
20. Bupyres viripis (114).
[N. Bohol, ¢ 2, November. ]
21. CoryDALLA LUGUBRIS (117).
[N. Bohol, ¢ 2, October and November. |
22. CyrTostomus JUGULARIS (123).
[N. Bohol, 2, October. ]
23. Corvus PHILIPPINUS (125).
[N. Bohol, ¢ 2, November. ]
Female smaller than the male.
24. CALORNIS PANAYENSIS (128).
[N. Bohol, ¢ 2, October. ]
25. Munia sacort (132).
[N. Bohol, 3, October. |
26. ALAUDA WATTERSI.
Alauda wattersi, Swinhoe, P. Z. S. 1871, p. 389.
[N. Bohol, 3, October. |
A single example of the genus Alauda, which most nearly agrees
with the South-Formosan race of A. celivox, separated under the
above title by Mr. Swinhoe. The occurrence in the Philippines of
any member of the genus has not before been recorded,
27. OSMOTRENON VERNANS (135).
[N. Bohol, ¢ 9, October. |
1878.] THE ORNITHOLOGY OF THE PHILIPPINES. 711
28. Turtur pussumiert (147).
[N. Bohol, ¢ 2, October and November. }
29. CHARADRIUS FULVUS (159).
[N. Bohol, ¢ 2, October. |
30. EupromiAs GEOFFROY! (161).
[N. Bohol, 3 2, October and November. |
31. EupRoMIAS MONGOLICUS (163).
[N. Bohol, ¢ 2, October and November. |
32. ANGIALITIs DUBIA (162). aii
[N. Bohol, 2, October. |
33. ANGIALITIS CANTIANA.
[N. Bohol, 2, October. ]
34. ANGIALITIS PERONII.
Charadrius peronii, Temminck, Schlegel, Mus. P.-Bas, Cursores,
p- 33.
ARGIALITIS, sp.?
{N. Bohol, ¢, November. ]
35. STREPSILAS INTERPRES.
Tringa interpres, Linn. 8. N. i. p. 248.
[N. Bohol, ¢, November. 9, November: iris chocolate ; ne”
and bill black ; legs and feet dirty orange. |
New to the Philippines.
36. LIMOSA LAPPONICA.
[N. Bohol, ¢, November. ]}
37. RHYACOPHILUS GLAREOLA (182).
[N. Bohol, ¢, November: iris dark brown ; bill and nails Black ;
legs and feet ‘yellow-green. ]
38. TRINGOIDES HYPOLEUCOs (183).
[N. Bohol, ¢ 2, October and November. |
39. ToTaANUS INCANUS.
Scolopax incanus, Gm, 8. N. i. p. 658.
{N. Bohol, ¢ 2, November : iris chocolate-brown; legs and feet
dirty chrome ; nails black; bill dark dull grey, base yellow. ]
40. TeREKIA CINEREA.
Scolopax cinerea, Guldenstadt, N. Comment. Ac. Sc. Imp.
Petropol. xix. p. 473, t. 19 (1775).
[N. Bohol, ¢ 2, October. }
New to the Philippines.
712 PR. 0. FINSCH ON A NEW STURNUS. [June 18,
41. LimicoLa PLATYRHYNCHA.
Tringa platyrhyncha, Temm. Man. d’Orn. p. 398 (1815).
[N. Bohol, 2, October. }
The specimens have been submitted to Mr. Dresser, who is of
opinion that they belong to this species and not to L. sibirica.
42. TRINGA ALBESCENS.
Tringa albesecens, Temminck, Pl. Col. 41, f. 2.
[N. Bohol, ¢ 2, October and November.]
New to the Philippines.
43. GALLINAGO SCOLOPACINA (187).
[N. Bohol, ¢, November.]
44. Busuxcus coromannvs (194).
[N. Bohol, 2, October. }
45. HERODIAS GARZETTA (195).
[N. Bohol, ¢, October. |
46. DENDROCYGNA vaAGANS (203).
[N. Bohol, ¢ 2, October and November. ]
47, STERNA BERGII.
Sterna bergii, Licht. Verz. d. Doubl. Berl. Mus. p. 80.
[N. Bohol, ¢ 2, November. |
9. On a new Species of Starling. By O. Fivscn,
Ph.D. C.M.Z.83
[Received June 17, 1878.]
This species, which I propose to name after General Poltaratzsky,
Governor of Semipalatinsk, is easily distinguishable from our common
Starling by having the back green instead of purplish-violet, and
the underparts below the neck of a deep purplish-violet instead of
green as in the remaining species.
I found this Starling breeding on Lake Marka-Kul, in the Chinese
High Altai, about 5000 feet above the sea-level. A careful compa-
rison of it with the rich series of Séurni in the British Museum has
convinced me that it belongs to a distinct species hitherto confounded
with S¢. vulgaris and its allies. The British Museum contains
fifteen specimens of tne same bird from North-Western India,
Persia, and Baluchistan. Specimens from the two last-named
localities are marked S¢. unicolor and St. vulgaris by Mr. Blanford.
The single specimen of a Starling brought home by Mr. Seebohm
from the Yenisey belongs also to this species. It is the St. humei,
Gould (1877), but not of Brooks (1876), and most probably also
the S¢. vulgaris of Radde and Schrenck from Eastern Siberia. The
1878. | MR. H. W. BATES ON NEW COLEOPTERA. 713
group to which our common Starling belongs contains the following
Species :—
St. vulgaris, L.; Sé. indicus, Hodgs. (Nepaul); Sé. humei,
Brooks (= xifens, Hume, = ambiguus, Hume) ; St. purpurascens,
Gould ; and St. poltaratskyi, mihi (= humez, Gould).
All these species are well marked, and distinguishable even in
winter plumage and in the young state, as J intend to show in a
more extended description of them.
10, On new Species of Coleopterous Insects (Geodephaga
and Longicornia) taken by Dr. Stoliczka during the
Forsyth Expedition to Kashgar in 1873-74. By H.
W. Bates, F.Z.S.
[Received June 17, 1878.]
The following are preliminary diagnoses of new genera and spe-
cies of Coleoptera, of the groups above mentioned, taken during
the mission of Sir T. Douglas Forsyth to the Amir of Kashgar in
1873-4. Complete descriptions will hereafter appear in the general
work on the results of the expedition, now in course of publication
at Calcutta.
GEODEPHAGA.
CICINDELA STOLICZKANA.
C. burmeisteri (Fischer) affinis, sed minor. Nigra; corpore sub-
tus, pedibus antennarumque basi chalybeo-violaceis ; elytris
lunula humerala et apicali fausciaque mediana, lata, abbreviata,
recta flavo-albis ; thorace brevi, lateribus fere rectis, supra sub-
talisseme granulatim strigoso; elytris minute, haud confertim
granulaties ; palpis nigris, albo-setosis ; labro albo, convexo, antice
medio rotundato producto, unidentato ; corpore subtus pedibusque
sparsim albo-pilosis.
Long 6-73 lin.
North of the Kuen-lun. Exact locality not given.
CARABUS STOLICZKANUS.
C. caschmirensi (Kollar) affinis. Maxime elongatus et angustus,
niger subnitidus ; thorace late subcordato-quadrato, angulis pos-
ticis retrorsum productis, acutis ; elytris angustis, post medium
perparum rotundato-dilatatis, dorso tuberculorum triplict serie,
inter se carina uniea separatis; menti dente verticaliter ex-
stante, valde compresso,
Long. 14-15 lin.
Murree.
DicHIROTRICHUS ALTICOLA.
Oblongus, supra testaceo-fuluus, capite (maculis rufis exceptis),
Proc. Zoox, Soc.—1878, No, XLVI. 46
14 MR. H. W. BATES ON NEW COLEOPTERA. [June 18,
thoracis macula discoidal alteraque elytrorum postico-discoidalt
nigro-ceeneis ; palpis apice acuminatis ; capite et thorace grosse
subsparsim punctatis, hoc postice angustato, angulis posticis
oblique rotundatis, margine postice arcuato ; elytris striatis,
interstitiis medio leviter culminatis, biseriatim punctatis ; corpore
subtus nigro ; antennis fuscescentibus. 3. Tarsi duo antici ar-
ticulis 1°-3"™" ovatis, 4° bilobo.
Long 2+ lin.
Pamir, ‘between Sirikol and Panga.
HARPALUS CHRULEATUS.
Elongato-oblongus, glaber ; thorace transversim quadrato, postice
distincte angustato, lateribus arcuatis, angulis posters rotundatis,
bast utringue late subcrebre punctato, margine basali bisinwato ;
elytris pice Sortiter sinuatis, supra striatis, interstitis planis,
umpunctatis, tertio unipunctato.
d supra capite thoraceque nagr is, politis ; elytris cwruleis, subviola-
ceis, nitidis ; antennis nigris, articulo basali rufo ; corpore sub-
tus nigro, pectore medio pedibusque rufopicers ; abdomine minus
nitido. Immaturus toto corpore castaneo-rufo, elytris violaceis.
2 nigro- vel rufo-castanea, raro obscure nigra ; elytris opacis, inter-
dum violaceo tinctis, wpice fortius (sicut in H. eeneo Q ) sinuatis.
Long. 53-6 lin.
Yangi Hissar.
HARPALUS MELANEUS.
H. calceato (Dufts.) forma coloreque similis, at thorace fere wm-
prnctato, angulisque posticis obtusis. Oblongus, modice elongatus,
niger nitidus, elytris x opacis ; antennis et palpis nagris, arti-
culs omnibus apie piceo-rufis ; thorace transversim quadrato,
postice leviter angustato, angulis posticis obtusis, lateribus antice
modice arcuatis, basi utrinque vage vie punctato, fovea linear
impresso ; elytris apice paullulum sinuatis, supra convexis,
simpliciter striatis, interstitiis modice convexis, tertio unipunctato.
Long. 6 lin.
Sind valley; Murree; near Leh.
HARPALUS TURCULUS.
Oblongus, niger, 3 nitidus, Q sericinitens ; antennis articulo primo
rufo, palpis apice flavis ; capite modice angusto, levi ; thorace
quadrato, antice prope angulos rotundato-angustato, postice latert-
bus exacte parallelis, elytris multo angustiore, angulis posticis
rectis, supra impunctato ; elytris utroque sexu apice fortiter
subrecte sinuatis, supra subtiliter striatis, interstitus planis,
tertio unipunctato ; metasterno grosse sparsim punctato,
Long. 4 lin.
No locality. Probably near Yarkand.
HARPALUS INDICOLA.
Elongato-oblongus, angustior, nigerrimus, g magis, 2 minus niti-
- 1878.] MR. H. W. BATES ON NEW COLEOPTERA. 715
da, palpis et antennis fulvis ; thorace quadrato, latertbus leniter
arcuatis, postice longe angustato, angulis posticis obtusis, base
tota subsparsim punctata et paulo rugosa, fovea utringue obliqua ;
elytris convexis, apice modice sinuatis, supra striatis ( 3b fortius),
interstitiis vix convexis, tertio puncto conspicue impresso ; sternis
et ventre lateribus grosse haud profunde punctatis.
* Long. 5 lin.
Murree.
“—
HARPALUS MASOREOIDES.
Parvus, niger subsericeus, levis, partibus oris antennisque flavo-
testaceis, pedibus magis rufescentibus ; thorace transverso, elytris
vie angustiore, antice gradatim paullulum angustato, angulis
posticis rotundatis, levi, foveola basali utrinque oblonga, margi-
nibus rufescentibus ; elytris oblongis, apice late obtusis, leviter
sinuatis, supra striatis, striis minutissime punctulatis, interstitirs
via convexis, marginibus reflexis et epipleuris piceo-rufis.
Long. 23 lin.
Pamir, between Sirikol and Panga.
HARPALUS LIODES.
Ovatus, latus, modice conveaus, niger, gS sericinitens, Q serici-
opacus, antennis palpisque piceo-rufis, wllis nigro maculatis, tarsis
piceo-rufis ; capite lato, levigato, oculis minus prominulis ;
thorace valde transverso, antice angustato, postice multo latiore,
angulis posticis rectis, fere impunctato; elytris ovatis, apice
paullulum sinuatis, supra subtiliter striatis, interstitiis planis,
tertio minute unipunctato.
Long. 5-53 lin.
No locality. Probably near Yarkand.
HYPOLITHUS PERLUCENS.
Piceo-niger, lete iridescens, glaber, antennis, palpis e pedibus
fulwo-testaceis ; capite levissimo, post oculos angustato, fovea
frontali lineari versus oculum curvata ; thorace quadrato, lateribus
leniter fere equaliter arcuatis, angulis postics valde obtusis,
margine postico late sinuato, supra limbo toto crebre subtiliter
punctulato, polito, marginibus rufescentibus ; elytris fortiter
striatis, interstitiis paulluium conveais, politissimis, tertio (prope
striam secundam) multipunctato,
Long. 4} lin. 2.
Jhelam valley.
HyPpsINEPHUS, nov. gen,
A genere Selenophoro differt pedibus elongatis, validis, tarsisque 3
articulis dilatatis cordatis, primo basi gracili, apwe subito dila-
tato, 4" breviter bilobo. Mentum edentatum ; elytra «nterstitirs
alternis pluripunctatis.
46*
716 MR. H. W. BATES ON NEW COLEOPTERA. [June 18,
HypsINEPHUS ELLIPTICUS.
Piceo-niger vel castaneus ; partibus oris, antennis pedibusque fulvo-
testaceis ; thorace quadrato, elytris angustiore, lateribus postice
explanatis, angulis posticis obtusis, supra impunctato ; elytris
elongato-ovatis, striatis, interstitio tertio punctis quinque parvis,
quinto versus basin duobus, septimo versus apicem pluribus
umpressis.
Long. 6 lin.
Pangong valley.
AMARA BAMIDUNY.
A. triviali (Dufts.) afinis. Ovata, subtus viridi-cenea, supra enea,
antennis articulis 2 basalibus rufis ; pedibus nigro- vel ceneo-
piceis ; thorace quam in A. triviali breviore, bast impunctato ;
elytris striis subtilibus, apice haud profundius unpressis, distincte
punctulatis, interstitiis planis.
Long. 33-4 lin.
Pamir, between Sirikol and Panga.
AMARA AMBIGENA.
Breviter ovata, nigro-cenea polita, ventris apice rufo, palpis, anten-
nis basi, pedibus elytrorumque epipleuris rufis ; elytris interdum
castaneis ; thorace brevi, antice gradatim rotundato-anqustato,
apud basin elytris paulo angustiore, margine basali flecuoso,
angulis posticis subacutis, foveolis basi utrinque duabus latis,
sparsim grosse punctatis ; elytris brevibus, punctulato-striatis,
interstitus planis ; menti dente elongato triangulari, apice’ an-
guste fisso.
Long. 33 lin.
Between Tanktze and Chagra, Pangong valley.
LEIOCNEMIS HIMALAICA.
Elongato-ovata, rufo-picea, supra eneo tincta ; partibus oris, an-
tennis pedibusque flavo-testaceis ; thorace brevi, transverso, lateri-
bus fere cequaliter arcuatis, antice paulo magis quam spostice
angustato, angulis posticis obtusis, lateribus paullulum explanatis,
foveolis basalibus grosse sparsim punctatis, interiore rotundata,
exteriore vage impressa, carinaque obsoletissima ; elytris acute
et simpliciter striatis, interstitiis planis.
3. Tibi intermedi subtus medio sinuate, deinde paullo dilatate
et denticulate.
Long. 35 lin.
Ladakh.
LEIOCNEMIS TARTARI.
Oblongo-ovata, modice convexa, nigra, polita, supra enescens ; par-
tibus oris, antennis pedibusque piceo-rufis ; thorace valde trans-
verso, lateribus fortiter arcuatis, antice et postice fere cequaliter
angustato, angulis posticis distinctis sed obtusis, bast utrinque
foveis duabus modice impressis, fortiter punctatis ; elytris
punctato-strratis, interstitiis planis ; prosterno apice late rotun-
1878.] MR. H. W. BATES ON NEW COLEOPTERA. 717
dato, marginato ; meso- et metastérnis punctatis ; menti dente
lato, magno, bifido.
Long. 33 lin.
Between Yangi Hissar and Sirikol.
LEIOCNEMIS FRIVOLA.
Parva, oblonga, subtus rufo-castanea, supra nigro-cenea vel cenea,
elytris interdum castaneis ceneo tinctis ; partibus oris, antennis
pedibusque rufo testaceis ; thorace transversim quadrato, lateribus
leniter arcuatis, postice minus quam antice angustato,-mox ante
basin paullulum sinuato, angulis posticis fere rectis, foveis
basalibus grosse punctates ; elytris punctulato-striatis ; menti
dente bifido, plano.
Long. 23 lin.
No locality. Probably near Yarkand or eastern slopes of Pamir.
AMATHITIS BADIOLA.
A. rufescenti (Dej.) prowime affinis, at angustior corporeque infra
nigro. Subtus nigra, supra capite thoraceque rufo-castaneis (illo
obscuriore) ; elytris subfusco-badiis, interdum ceneo-nitidis, strits
obscurioribus ; oculis multo minus quam in A. rufescente promi-
nulis ; thoracis angulis posticis via rectangulis (sed apice acutis),
basi tota discrete punctata, carina via elevata ; elytris punctulato-
striatis, interstitus planis ; menti dente prominulo, triangulari.
Long 4 lin.
North of Kuen-Lun. One example ticketed “ Sanju.”
AMATHITIS KUENLUNENSIs.
Valde elongatus, pallide ferrugineus, antennis pedibusque gracilibus,
dlarum articulo 3° ceteris multo longiore ; thorace elytris multo
angustiore, late cordato, lateribus antice fortiter arcuatis, postice
sinuatim angustato, angulis posticis acutis, supra impunctato,
foveis basalibus latis, vagis, carinaque prope angulum indistinete
elevata ; elytris elongato-ovatis, apice paulo sinuatis, striis levi-
bus, modice impressis.
Long. 5 lin. 9.
Sanju.
BRrabDYTUS COMPACTUS.
Breviter oblongo-ovatus, latus, subceneo-niger, convexus ; capite brevi
et crasso, oculis parum convexis, epistomatis margine antico
transversim sulcato; thorace elytris haud angustiore, postice
modice angustato, angulis posticis acutis, foveolis basalibus parvis,
punctatis, carinaque obsoleta ; elytris simpliciter striatis, inter-
stitus plans; palpis, antennis pedibusque rufo-piceis ; menti
dente magyno, apie inconspicue emarginato.
Long. 4 lin.
Murree.
CURTONOTUS PAMIRENSIS,
Hlongato-oblongus, angustus, rufo-castuneus, supra olivaceo-ceneus,
718 MR. H. W. BATES ON NEW COLEOPTERA. [June 18,
thoracis elytrorumque marginibus reflexis rufescentibus ; capite
levi, mox pone oculos angustato ; thorace transversim quadrato,
ante basin subfortiter constricto, ibique lateribus parallelis et
margine laterali haud interrupto, angulis posticis rectis, base
grosse subsparsim punctato, foveis utrinque linearibus ; elytris
striatis, striis (versus apicem exceptis) punctatis.
3 tibiis intermediis post medium angustatis et subtus acute breviter
bidentatis.
Long. 43-5 lin.
Pamir, between Sirikol and Panga.
BreMBIDIUM (PERYPHUS) PAMIRENSE.
Oblongum, depressum, capite thoraceque viridi- vel awrato-ceneis,
politis ; elytris fulvo-testaceis, vitta suturali (ante apicem abbre-
viata) fasciaque pone medium, interdum quoque margine et apice
eneo-fuscis ; antennis, palpis pedibusque flavo-testaceis ; thorace
breviter cordato, antice fortiter rotundato, basi rugato, angulis
posticis rectis, fovea utrinque profunda carinulaque obliqua ; ely-
tris striato-punctatis, interstitiis planis.
Long. 23 lin.
Pamir, between Sirikol and Panga.
Bemeipium (PERYPHUS) PUNCTULIPENNE.
Subdepressum, eneo-nigrum, politum, mandibulis piceo-rufis; thorace
antice leniter rotundato, postice usque ad angulos angustato, his
fortiter reflexis, obtusis, margine basali utrinque prope angulum
valde obliquo, supra basi et margine rugulosis, fovea oblonga ;
elytris punctato-striatis, striis 1°-4"™ solum impressis, 6% et 74
obsoletis, interstitiis planis, minutissime sparsim punctatis,
Long. vix 2 lin.
No locality. Pamir or near Yarkand?
MOL.uopPs PILIFERUS.
Niger, nitidus ; thorace late cordato, post medium subsinuatim
angustato, angulis posticis rectis, antice juxta marginem lateralem
punetis decem longe piliferis lineatim disposites ; elytris elongato-
ovatis, convexis, prope apicem fortiter sinuatis, supra exarato-
striatis, striis 7¢ et 8% valde approximatis, 7" continuo punctatis,
punetis longe piliferis, interstitus dorsalibus planis, 3° et 5° apice
pilifero-punctatis.
Long. 6-7 lin.
Murree.
ANCHOMENUS LADAKENSIS.
A. parumpunctato (Linn.) proxime affinis, sed gracilior, thorace
longiore, etc. Supra subfusco-cupreus, capite thoraceque magis
eneis, interdum toto viridi-eneus ; femoribus nigro-piceis, tibiis
tarsisque rufopicers ; antennis prceo-fuscis, articulo basali rufo ;
thorace subquadrato, lateribus leniter arcuatis, angulis posticis
rotundatis, ibique margine explanato-reflewo, toto limbo alutaceo ;
1878. } MR. H. W. BATES ON NEW COLEOPTERA. 719
elytris elongatis, parallelis, acute striatis, interstitis planis, tertio
5-punctato.
Long. 34 lin.
Between Tangtze and Chagra, Pangong valley ; also Pamir, be-
tween Sirikol and Panga.
ANCHOMENUS POLITISSIMUS.
A. fuliginoso (Panz.) forma subsimilis, nigro-cneus, politissimus¢
capite breviter ovato, oculis vix prominulis; palpis art. ult.
acuminatis ; thorace postice angustato, angulis posticis oblique
rotundatis, margine prope angulum valde refleao ; elytris versus
apicem valde sinuatis, swpra obsolete striatis, disco utrinque haud
conspicue bipunctato ; pedibus wneis, tibiis rufo-testaceis.
Long. 23 lin.
Murree.
COLPODES OVALICEPS.
Minus elongatus, nigro-chalybeus, nitidus ; elytris ampliatis, ovatis ;
capite parvo, ovato, oculis haud prominulis ; menti dente apie
sulcato-emarginato ; thorace ovato, quamcaput dimidio latiore,mar-
gine laterali equaliter explanato, subreflexo, angulis posticis subro-
tundatis ; elytris convexrs, late ovatis, apice via sinuatis, humeris
rotundatis. striatis, interstitis planis, tertio tripunctato ; meta-
stern episternis brevibus ; antennis, palpis pedibusque rufo-picers,
Ffemoribus nigris.
Long. 5 lin.
Murree.
METABLETUS TARTARUS.
M. truncatello (Linn.) paullo major, magis elongatus, subeneo-
niger, nitidus, antennis et pedibus fusco-piceis ; thorace quam in
M. truncatello magis angustato, angulis posticis obtusiortbus,
deinde usque ad basin magis obliquis ; elytris elongatis, apice
obtuse subsinuatim truncatis, obsolete striatis, umpunctatis.
Long. 12 lin.
ebweon Vane Hissar and Sirikol.
CyYMINDIS GLABRELLA.
C. andree (Ménétr.) affinis, at gracilior, oculis minus prominulis,
elytrisque fusco-castaneis flavo marginatis. Gracilis, glaberrima,
castaneo-rufa, abdomine margine picescente, capite obscuriore,
partibus oris, antennis, pedibus elytrorumque margine fulvotesta-
ceis ; capite angusto, sparsim punctulato ; thorace quam caput haud
latiore, anguste cordato, lateribus postice leviter sinuatis, angulis
posticis fere rotundatis, subpunctulato-striatis, interstitiis sparsim
unctulatis.
Long. 4-43 lin.
Ladakh.
720 MR. H. W. BATES ON NEW COLEOPTERA. [June 18,
LONGICORNIA.
PRIONUS CORPULENTUS.
Q magna, elongata, nigro-castanca, supra omnino coriacea via
nitida ; thorace parvo, utrinque acute trispinoso ; elytris versus
basin parallelis, compressis, deinde modice dilatato-rotundatis,
apice late obtusis ; pectore toto dense fulvo-hirto ; abdomine poli-
tissimo ; antennis 12-articulatis, grosse punctatis, articulis
39-11” serratis. b
Long. 24 lin.
Murree.
HEsSPEROPHANES CRIBRICOLLIS.
Cylindricus, fulvus, pubescentia ineumbente cinereo-fulva apud
elytra maculatim vestitus ; thorace rotundato, elytris multo
angustiore, inequali, lateribus medio subtuberoso, supra erebre
alveolato-punctato ; scutello cinereo ; elytris omnino discrete
punctatis, haud conspicue bicarinatis, apice gradatim angustatis,
subacuminatis.
Long. 8-9 lin.
Murree.
TRINOPHYLUM, nov. gen.
Gen. Hesperophani affine, sed femoribus abrupte clavatis, oculisque
minus fortiter granulatis. Corpus elongatum, subdepressum, brevi-
ter suberecte pubescens, crebre punctatum. Caput brevissimum :
tubera antennifera haud elevata; palpis brevissims. Thorax
rotundatus, inermis. LElytra apice obtusa. Acetabula antica
extus haud elongata. Prosternum angustum ; mesosternum latum,
subconverum ; metasterni episterna elongata, angusta, postice
attenuata. Antenne filiformes, corpore paullo breviores, arti-
culo 4° paulo abbreviato.
TRINOPHYLUM CRIBRATUM.
Castaneo-fuscum, subnitidum, omnino suberecte fusco-pubescens,
crebre sed discrete punctatum ; thorace supra antice et postice
paullo depresso, inva dorsali levi; elytris linets duabus indi-
stinctis, lavibus.
Long. 63 lin.
Murree.
LEPTURA RUBRIOLA.
L. sanguinolente(Linn.)afinis. Nigra, subtus sparsim fulvo-pilosa,
thoracis plaga magna discoidea elytrisque rufo-opacis ; capite et
thorace reticulato-punctatis, illo paullo post oculos subito et for-
titer constricto ; thorace medio haud conspicue dilatato, modice
convexo, angulisque posticis modice productis ; elytris subcrebre
punetulatis, apice recte truncatis, angulisque breviter dentatis, 2 .
Long. 6 lin.
Murree:
1878.] DR. G. HARTLAUB ON A NEW NOTAUGES. 721
CLYTANTHUS IGNOBILIS.
Cl. 4-punctato (Fabr.) prowime affinis. Nigro-fuscus, tomento
cinereo-flavo vestitus ; elytris utrinque maculis 5 nigris, quam in
Cl. 4-punctato majoribus, scilicet 1 curvata post scutellum, 1
parva humerali, 1 antico-discoidali, 1 mediana majore transversa
et 1 huic proxima longitudinali oblonga.
Long. 6 lin.
Murree. a
11. On anew Species of Notauges (N. hildebrandti, Cab.).
By Dr. G. Harrravs.
[Received June 17, 1878.]
The Museum of Natural History of Bremen has been enriched of
late with a fine adult specimen of a new typical Notauges, discovered
by the famous traveller Mr. Hildebrandt, in Eastern Africa. The
exact place where he collected this interesting bird is “Tkanga in
Ukamba.”
Mas ad. capite, gutture, dorso, tergo, uropygio, supracaudalibus,
tectricibus alarum minoribus margineque alari chalybeis, nitore
purpurascente ; collo postico late et circumscripte «neo-vire-
scente; alis duabus seriebus macularum holosericeo-nigrarum
bifasciatis ; tectricibus alarum majoribus viridi-eneis ; remigibus
primariis in pogonio externo eneo-cerulescentibus, obscure macu-
latis, interno nigris ; tectricibus minoribus in pogonio externo
unicoloribus eneo-virentibus; pectore, abdomine, cruribus et sub-
caudalibus cinnamomeo-rufis ; subalaribus majoribus rufis, minori-
bus chalybeis ; rectricibus cerulescenti-eneis, minus distincte
undulatis ; rostro et pedibus nigris ; iride aurantiaco-rubra.
Long. 19 centim., rostr. afr. 18 mill., al. 123 mill., caud. 83 mill.,
tars. 30 mill.
The steel-blue colour of the head is less brilliant on the crown;
there appear some rufous feathers on the uropygium; and some of
the upper tail-coverts have aslight rufous terminal edging; the first
primary is of a uniform blackish colour.
In Notauges superbus (Abyssinia, Shoa) the colouring is very
different: the upper parts are brown-green, the vent and under
tail-coverts are white, and there are other differences.
Mr, Hildebrandt told me that during his many excursions in East
Africa he never met with Notauges albicapillus, Blyth. This is cer-
tainly one of the rarest birds, the Calcutta specimen collected by
Speke and Burton being as yet the only one known.
722 LIEUT.-CUL, BEDDOME ON A NEW BATRACHIAN. [June 18,
12. Description of a new Batrachian from Southern India,
belonging to the family Phryniscide. By Lieut.-Col.
R. H. Bepvomg, C.M.ZS.
[Received June 17, 1878.]
MELANOBATRACHUS, gen. Nov.
Toes cylindrical, not dilated at the ends ; no maxillary or vomerine
teeth; ear imperfectly developed, no parotids; sacral vertebra
dilated; toes webbed; body slender, elongate, uniform in width ;
muzzle short, obtuse; fingers and toes short, metatarsus with a
slightly swollen inconspicuous tubercle.
MELANOBATRACHUS INDICUS, 0. Sp.
Body with nearly parallel sides as in Ansonia ; no canthus rostralis,
no sign of a tympanum or cavum tympani; cleft of mouth extends
nearly as far back as the eyelid, the space between the nares yy of
an inch; fingers 4, free, short, the third double as long as the others ;
toes 5, slightly webbed, the fourth much longer than the third.
Eustachian tubes closed, tongue rounded and free behind, surface of
the head and body tubercled, arms smooth, the humerus short, hind
limbs tubercled, the thigh shorter than the calf; belly and sides
smooth. Colour black, with minute white dots on the back and larger
ones on the belly, the thighs near their junction with the body are
girt with a brilliant scarlet band, j inch in breadth, either continuous
all round or interrupted above ; and there are sometimes a few scarlet
blotches on the anterior portion of the belly between the fore legs
and sometimes on the lower portion of the humerus (the scarlet
colour of these markings fading to cream-colour in spirits). Length
of body 13 to 12 inch, length of hind limb to end of fourth toe
14 to 1¢ inch.
Hab. The Anamallays and the Ghat range to the south of those
mountains, very rare, in moist evergreen forests of 4000 feet
elevation, under old rotten logs. The specimens found were all in
guite a torpid state and curled up almost into a ball, but became
very active when put in spirits.
Dr. Anderson, who has kindly prepared the skeleton of one of
these Batrachians, informs me that its osteology much more resembles
that of Bufo than Rana; the skull is short and broad, and the
frontals and parietals have considerable lateral extension; it is also
edentulous, and there are no vomerine eminences; a careful exami-
nation of the sternum does not reveal the existence of any omoster-
num ; clavicles are present. Dr. Anderson was not able to detect
any tympanum under the skin; nor could he trace a tympanic
cavity. He mentions that the stem of the stapes is not developed,
and moreover that he could not detect any Eustachian openings in
the mouth.
There are only from 8 to 9 vertebrze in Bufo melanostictus; and the
1878. ] SIR V. BROOKE ON GAZELLA GRANTI. 723
first vertebra, which has considerable length and breadth, is marked
on its under surface by a prominent ridge which does not occur in
the above-mentioned Indian toad.
The transverse processes of the sacral vertebra are dilated as in
Bufo.
13. On Gazella granti. By Sir Victor Brooks,
Bart., F.Z.S.
[Received June 17, 1878.]
Some years ago (vide P. Z.S. 1872, p. 601. pl. xli.) I had the
pleasure of bringing before the Society’s notice a species of Gazella
which I supposed to be new, and upon which I conferred provi-
sionally the name Gazella granti. My original description of the
species was based on small water-colour sketches of the head and flat
skins of a male and female obtained by Capt. Speke and Col. Grant
in Ugogo, which were executed by them previous to the dispatch of
the specimens to England. When writing my first notice, I believed
that these specimens had been lost, there being no record of their
arrival at the Society’s rooms, where Capt. Speke had expressed a
wish that they should be kept until his return; but Col. Grant has
since informed me that they reached England safely and are still
preserved along with some other objects of natural history collected
by Capt. Speke, in Capt. Speke’s father’s house. ;
In 1875 Dr. Kirk presented to the Society (vide P. Z. S. 1875,
p- 527, pl. lix.) the first living specimen of the species which
had ever been exhibited in Europe. Unfortunately the animal,
a very young female, arrived in very delicate health, and died
shortly afterwards. It is now preserved in the British Museum
(spec. 75. 12. 29. 4), whither it was transferred after Mr. Sclater’s
exhibition of it to the Society. It was not, however, till towards
the end of last year that the first specimen of an adult male of this
magnificent gazelle came under my personal observation. The
animal was shot by Mr. Charles Arkwright, about 8 miles from the
village of Tubugwe, which is situated, as nearly as Mr. Arkwright
could ascertain, about 50 miles N.E. of Ugogo. It was one of
a herd composed of about 20 individuals, amongst which there
was one other old male. Mr. Arkwright has most kindly intrusted
his valuable trophy to my care ; so that I am enabled to exhibit it
to the Society this evening, and to base upon it and upon the young
female in the British Museum a fuller and more accurate description
of the species than was possible at the time of my original notice.
GAZELLA GRANTI.
1863. Antilope semmerringii? Speke, P. Z.S. 1863, p. 3, sp. 15.
1872. Gazella granti, Brooke, P. Z. 8. 1872, p. 601, pl. xli.
1873. Gazella granti, Brooke, P. Z.S. 1873, p. 550.
1875. Gazella granti, Sclater, P. Z. 8S. 1875, p. 535.
General colour very rich fawn tinged with purple, the neck and
724 SIR V. BROOKE ON GAZELLA GRANTI. [June 18,
body in certain lights mottled like watered silk ; this pattern is gained
by indefinitely bounded bands of darker hair. Central facial band *
darker than the neck and body, and strongly tinged with rufous on
the forehead. A large black patch above the nostrils. Light facial
streaks very definite, white. Dark facial streak black, extending
Fig. 1.
>
Gazella granti.
from the base of the horn to the eye, which it almost encircles, and
from the eye to above the corners of the mouth. Posterior to the
eye is a small white patch. External surface of the ear-conchs
streaked with dark brown. Dark lateral and pygal bands distinctly
marked; the former grizzled, the latter very broad and jet-black.
1 For a detailed definition of the terms used in this description, wide P.Z.8.
1873, p. 436.
1878. ] SIR V. BROOKE ON GAZELLA GRANTI. 725
Lower two thirds of the tail black, the upper third white. The tip
of the tail is tufted ; for the remainder of its length it is furnished
with coarse hairs which project at right angles to its axis. Chin, lower
surface of the head, breast, belly, inside of limbs and rump white,
the white of the rump extending forwards in an angle into the
darker colour of the body. Knee-brushes of moderate length,
Fig. 2.
Gazella granti.
grizzled. Horns of great length in both sexes, much compressed
from side to side, marked by strong annulations which completely
surround the horn for the proximal half of its length, but become
obsolete on the posterior surface of its distal or upper half. The
horns diverge at first gradually, then more boldly, their tips con-
726 SIR V. BROOKE ON GAZELLA GRANTI. [June 18,
wereing and giving to the set of the horns a remarkably perfect lyrate
orm.
The skull is in every respect typically Gazelline.
Dimensions of Mr. Arkwright’s head and horns (figs. 1 and 2,
pp- 724, 725).
English Inches. Metre.
Length of horns round the curve ...... 240 0610
Length of horns in a straight line ...... 23°0 0°580
Circumference of horns at base ......... 66 0-170
Distance between the tips.................. 13-0 0-330
Wien tiMon Cars is.04.kvcsss kecswoweesseraae 6:5 0165
Dimensions of the young female in the British Museum, of which
the dentition is d. ig, d. e¢, d. m.3, m.4.
—
English Inches. Metre.
Height at shoulder ...................000e- 28°0 0:710
Bength*of Horns «.; 00. csesatees sss eiseeetee 75 0-190
Tength’ of ars | ....2...tecottyctsy-ss0ecses 5.5 0-140
Trenptl of Gail)....¢- 0-0: «c+ Scesmtere- sens c's
Length of tail, exclusive of hair ......... 65 0165 |
The horns of the adult female shot by Capt. Speke and Col. Grant
in Ugogo measured 15 inches round the curve.
Comparison with allied Species.
Gazella granti belongs, as pointed out in my former description,
to the group of long-limbed large Gazelles which are distinguished
by the white of the rump projecting in an angle into the fawn-colour
of the back and sides (vide P. Z.S. 1873, p. 537, B'). The allied
species are G. semmerringi, G. mohr, and G.dama. From all these
it differs in the very much greater size, and in the form of the horns.
It is also peculiar in the presence of the dark lateral band, which is
wanting in the three above-named species, and in the remarkably defi-
nite black pygal band, which is only shown in very young specimens
of Gazella semmerringi, is absent entirely in Gazella dama, and ouly
indefinitely shown. in Gazella mohr. In the watered-silk mottling
of the neck and back Gazella granti differs from all other known
Gazelles.
P Z.S. 1878. Pl. XII.
Df eet:
Peestuermenene
a Se te
wy PTT esee iON
a
NEW AND OTHER SHELLS FROM VARIOUS LOCALITIES.
C, Berjeoat ith
/antern 7
1878.] ON SHELLS FROM CHINA AND JAPAN. 727
14. Descriptions of two new Species of Shells from China
and Japan. By Cart Bock, F.G.S. &e.
[Received June 18, 1878.]
(Plate XLVI.)
TELLINA WROBLEWSKYI. (Plate XLVI. figs. 1, 2.)
Shell subovate, slightly inequilateral, gaping at both ends, marked
with coarse and rather distant concentric lines of growth, covered
with fine radiating strie, pellucid at intervals, with concentric opaque
layers, the posterior region also being somewhat opaque. Colour
bluish white stained with grey, especially at the posterior end. The
latter portion is 4 longer than the anterior, acuminately produced,
and terminates in a somewhat obtuse angle. Anterior margin
rounded, ventral but slightly arcuate, almost rectilinear near the
middle ; posterior dorsal slope deeply excavated, the valves being
strongly carinate from the umbo towards the posterior extremity,
with a deep excavation between the keel and the extreme margin.
Cardinal teeth 2 in the left valve, 1 in the right. Anterior muscular
impression very elongated near the margin, posterior irregularly
quadrate. Ligament situated on a narrow ledge of the posterior
dorsal slope, marked off by an incised line, and extending from the
umbo nearly 20 millims. In both valves (principally towards the
umbones) there are rather large and irregular pittings. Pallial sinus
very deep, forming a subacute angle, which extends anteriorly beyond
the umbo.
Breadth 112 millim., length 61 millim., diameter, with the valves
closed, 39 millim.
Hab. China.
The magnificent species here described is one of the largest of the
family (Tellinide), and resembles in outward appearance certain
species of the genus Anatina.
I propose to associate with this beautiful shell the name of Dr.
Wroblewsky of Copenhagen.
Trocaus (THaxoriA) yoxonamensis. (Plate XLVI. fig. 3.)
Shell imperforate, acutely pyramidal, dull white, marked at in-
tervals with irregular dark brown blotches, and the base ornamented
with brown lines radiating from the centre (where they are paler,
almost orange) to the periphery. Whorls 7, quite flat, spirally
ridged ; ridges crenulate or subgranular through being crossed by
oblique raised strize or lirulee ; on the upper whorls there are about
six ridges, and on the last, which is acutely angled at the middle,
they are about fourteen in number, that at the carina consisting of
two or three small approximated ones, and hence broader than
the rest. Aperture slightly oblique, subquadrate, upper half above
the carina shallowly sulcate, lower portion smooth; columella
white, callous, terminating in an oblique slight truneation. Greatest
diameter 15 millim., length 18.
Hab. Yokohama, Japan.
728 MR. E. A. SMITH ON NEW SHELLS FROM [June 18,
15. Descriptions of five new Shells from the Island of For-
mosa and the Persian Gulf, and Notes upon a few known
Species. By Encar A. Smiru, F.Z.S.
[Received June 18, 1878.]
(Plate XLVI.)
ME.aniA FoRMOSENSIS. (Plate XLVI. figs. 4 & 5.)
Shell elongate, subulate, olivaceous yellow, purer yellow beneath
the whorls, marked with longitudinal, undulating, more or less in-
terrupted and broken flammules of a dark brown colour, at the top of
the whorls forming conspicuous blotches, and at the base of the
body-whorl with a broad blackish-brown band. Whorls 13, a little
constricted at the upper part and submarginated and somewhat
convex beneath, the first 9 or 10 obliquely costate and deeply
transversely sulcate, the lowermost sulcus at the suture being par-
ticularly wide, forming a canaliculation, the last three whorls only
spirally sulcate and striated by obliquely flexuous lines of growth ;
sulci narrower than the interstices, about 10 in the penultimate
whorl and 24 in the last. Aperture ovate, acute superiorly, within
of the same colour as the exterior, but clouded by a white enamel,
the brown flammulation and spiral basal band being visible ; colu-
mella white, oblique, but slightly arcuate.
Length 33 millim., diameter 93; aperture 93 long, 5 wide.
Var. In this variety the longitudinal plicze extend as far as the
penultimate whorl, and are even partially developed on the last.
Hab. Formosa (Dickson).
This species, which I feel much pleasure in naming after Mr.
Matthew Dickson, has been liberally presented to the Museum by
him, together with the other Melaniz here described, and a nu-
merous series of insects,
Me tania picksoni. (Plate XLVI. fig. 6.)
Shell elongate, acuminate, shining, olivaceous yellow, purplish
brown at the apex, with or without spiral brown bands. Whorls 8
or 9, moderately convex, the six upper ones almost smooth, only
finely striated longitudinally by lines of growth, and also trans-
versely, the last two with a few faint transverse keels or ridges..
Aperture ovate, acuminate above, occupying nearly 2 of the entire
length, white within, the brown bands of the exterior (when present)
also visible. Lip thin, almost even, not prominent or sinuated ;
columella rather thickly callous in the umbilical region, and united
to the lip above by a very thin callosity.
Length 21 millim., diameter 8; aperture 8} long, 5 wide.
Hab. Formosa (Dickson).
This is a very remarkable species, and looks more like a North-
American than an Eastern form.
The brown transverse bands are present in five out of the six spe-
1878. ] FORMOSA AND THE PERSIAN GULP. 729
cimens presented to the Museum by Mr. Dickson. There are two
on the penultimate whorl, one at its base, and one a little above the
middle. On the body-whori there are three, the two upper ones
being continuous with those on the preceding volution, and the third
at the base.
MELANIA OBLIquIGRANOSA. (Plate XLVI. figs. 7 & 8.)
Shell elongately ovate, acuminate above, of a dirty yellowish
colour, and usually coated with a black earthy deposit ; spire acutely
conical, with straight outlines; whorls 9-10, turreted, quite flat,
obliquely plicated and spirally ridged, with nodules at the point of
contact of the plicee with the ridges; on the whorls of the spire the
ridges are four in number, the granules on the uppermost one a little
larger than those on the three lower ones. Body-whorl almost flat
above the middle; at that point and beneath somewhat convex,
encircled by 11 or 12 transverse ridges, whereof 5 or 6 of the upper
ones are granuliferous, the rest either uninterrupted or displaying
an indication of oblong granulations. Aperture pyriform, acute su-
periorly, within of nearly the same colour as the exterior, but coated
with a thin smooth callous deposit occupying about 2 of the entire
length of the shell; columella considerably arcuated, at the middle
white, united above to the lip by a very thin enamel on the whorl ;
lip thin, sinuated above and prominent below the middle. °
Length 25 millim., diameter 8; aperture 10 long, 5 wide.
Var. monstrosa. Shell ovate; spire short; aperture more than
half as long as the shell.
Hab. Formosa (Dickson).
This species may be recognized among the several allied forms
from the Philippine Islands and other places by its very rectilinear
acute spire and perfectly flat whorls. ‘The uppermost series of gra-
nules is at a little distance beneath the sutural line; and from this
circumstance the whorls have a turreted appearance. The nodules
of this series are larger and fewer in number than those of the other
series, and consequently do not quite regularly terminate the ob-
solete plicee, upon which the other three granules are situated in an
oblique direction. Some of the granules haye a squarish form,
others are transversely somewhat oblong; and most of them have
the upper and lower margins rather straight, with the lateral edges
less defined, but gradually blending off into the transverse ridges.
The colour of this species is pretty constantly uniform ; one spe-
cimen, however, is marked with a few reddish dots towards the base,
and these are most conspicuous within the aperture.
MeE;aniA TUBERCULATA, Miiller. (Plate XLVI. fig. 9.)
Hab. Formosa ( Dickson).
The shell here figured appears to be inseparable from this re-
markably variable and widely distributed species.
MELANIA suBPLICATULA. (Plate XLVI. fig. 10.)
Shell rather small, elongate, acuminate, eroded or truncate at the
Proc. Zoou. Soc.—1878, No. XLVII. 47
730 MR. E. A. SMITH ON NEW SHELLS FROM [June 18,
apex, olive-brown, more or less coated with a black earthy deposit.
Whorls probably about 9 in number, the remaining four or five
slightly convex, spirally suleate, and longitudinally sculptured with
coarsish flexuous lines of growth; ridges between the sulci about 10
in the penultimate whorl and 18 in the last, those near the base
separated by wider sulci than above. Aperture oval, acuminate
superiorly and very slightly effuse at the base, occupying about 3 of
the supposed entire length of the shell, sordid bluish within; lip
very thin, broadly sinuated above and somewhat prominent below
the middle. Columella whitish, moderately arcuate.
Supposed total length 23 millim. ; actual length of five remaining
whorls 19, diameter 74; aperture 74 long, 44 wide.
Hab. Formosa (Dickson).
This species resembles M. dolorosa, Gould, to some extent ; how-
ever, it is distinguishable on account of more elongated whorls and
coarser spiral sulci. Some of the lines of growth here and there are
well marked, and give the whorls a subplicated appearance.
Buta persica. (Plate XLVI. fig. 11.)
Shell ovate-acuminate superiorly, greyish or lavender-colour,
whitish at the upper part of the whorls, with the callous band at their
base white, bordered below by a brown line at the suture; and the
base or cauda of the last whorl is also white. Whorls 6, the first
two smooth, convex, the rest spirally sulcated, the ridges between
the sulci 8 in number in the upper whorls and about 16 in the last,
whereof 12 are above and 4 below the acute ridge or keel, which
winds round the lower portion of the whorl. Of these ridges the
four immediately below the suture are more or less white, granulous
through being crossed by oblique sulci, which only extend from the
suture a short distance. Aperture deep brown within, with a paler
or white border at the outer lip and over the basal sinus, and a little
longer than the spire. Columella moderately arched, and coated
with a thinnish white enamel. Basal sinus rather broad and deep.
Length 24 millim., diameter 10; aperture 12 long, 7 broad.
Hab. Bushire, Persian Gulf.
Two specimens of this very interesting species have recently been
presented to the Museum by Mr. A. S. Betts. The peculiar form
and colouring, the granular ridges at the upper part of the whorl,
and the brown aperture with the white margin to the labrum and
basal sinus are the most obvious distinguishing characters of the
species.
OsrreEA, sp., adhering to Trochus (Polydonta) maculatus, Linn.
(Plate XLVI. fig. 12.)
This is a remarkable instance in which a shell has assumed, to a
certain extent, the sculpture of another to which it has adhered.
This assumption or mimicry of the surface of other shells and foreign
substances has already been noticed in other species of Ostrea and
other genera, and is common to most forms of the geuus Myochama.
The same process which effects so remarkable a change in the normal
1878. | FORMOSA AND THE PERSIAN GULF. 731
aspect of the latter, namely the moulding of the edges of the upper
valve by the animal to the surface of the substance to which the
shell is attached, no doubt, was employed by the Oyster here figured ;
for I do not think that this alteration of outward appearance has
been in any way effected by contact of the tentacular cirri which
proceed from the edge of the mantle of the Trochus, because if this
were the case some alteration in colour would have been expected.
The lower valve is white both externally and within, and, being
thin, does not conceal the granular surface of the Trochus upon
which it rests. The upper valve is olive-green within, and smooth,
with the exception of the pear-shaped muscular scar and a few
small granules scattered all round the margin. The exterior is dark
lilac, with six obscure brownish rays, most observable towards the
margin, and terminating at the digitations or prolongations at the
edge.
The lamellated surface, so common to the upper valves of most
Oysters, is almost altogether obliterated, and only here and there
traces of it are noticeable. The exterior is rather smooth, sub-
angular, the indistinct granules arranged in series corresponding to
those on the Trochus; and the suture dividing the last and the pen-
ultimate whorls, and a depression or sulcus round the middle of the
former, are also reproduced. The specimen is in the collection of
Mr. Carl Bock.
Cyprma pease, Sowerby. (Plate XLVI. figs. 13 & 14.)
Cyprea peasei, Sowerby, Thesaurus Conchyl. iv. pl. 311. figs.
167, 168.
Hab. Mauritius. Coll. Carl Bock.
The shell here figured appears to be a monstrous growth of this
species. The alteration of form and the obtuse ridges on the dorsal
‘surface are suggestive of certain forms of the genus Ovulum.
Convus PASTINACA.
Much uncertainty and confusion exists with regard to the deter-
mination of this Lamarckian species. The original description of it
is so brief that it is absolutely impossible ever to know with cer-
tainty what species Lamarck really had before him, more especially
as the actual type (which has been lost) was never figured by him,
nor does he refer to any previously published figure. The shell de-
scribed and delineated by Kiener (Coq. Viv. pl. 26. fig. 2) certainly
cannot be the true C. pastinaca. The species there represented
(unknown to me) is decidedly distinct. The character of the spire
does not at all agree with Lamarck’s description, which is as fol-
lows :—‘‘spira obtusa, immaculata, submucronata;” and again in
French: ‘‘a spire non tachée.’’ Kiener’s shell has a rather con-
cavely elevated spire and fulvous blotches upon it.
Reeve’s idea of C. pastinaca, figured (Conchol. Icon. i. pl. 46.
fig. 257, copied by Sowerby, Thesaurus Conchyliorum, iii. pl. 201.
fig. 353) from a specimen in the British Museum, is also an erro-
neous one. This shell I have very carefully examined, and unhesi-
732 ON SOME KNOWN SPECIES OF SHELLS. [June 18,
tatingly pronounce it to be a bleached example of C. virgo; for in
form and sculpture it is absolutely identical with certain specimens
of that species, and differs only in colour. This variation may be
due to bleaching; or it may possibly be an albino form. The
purple base, which is so constant a character in this species, is trace-
able in a faint degree in Reeve’s shell. He does not refer to this in
his description; yet in the figure it has been represented by the
artist.
The species figured by Weinkauff in Kuster’s Conchilien-Cabinet,
pl. 32. figs. 1 and 2, apparently is the same as or very closely allied
to that depicted by Reeve.
In order to come to some decision in the matter, I commuicated
with Dr. Brot, of Geneva, asking him for information respecting
Lamarck’s species, as his types in Delessert’s collection have recently
been obtained by the museum of that city. Unfortunately, it ap-
pears that Lamarck’s Cones are not marked as in other genera, but
placed on tablets bearing the designation L. According to a cata-
logue of Delessert’s collection, Dr. Brot informes me that there
ought to be five specimens of C. pastinaca; but only four are now
to be found; the fifth (possibly that figured by Kiener) had disap-
peared before the collection reached Geneva. Of the remaining
four; three existed in Lamarck’s cabinet. None of these, how-
ever, has the dimensions indicated by that author. Two are con-
sidered by Dr. Brot small specimens of Conus quercinus ; for they
are ornamented (indistinctly, however) with the fine thread-like
brown lines which are characteristic of that species. ‘The third shell
he considers a diminutive specimen of that figured by Kiener ; for it
so resembles the figure, that at first he thought that it had been en-
larged for Kiener’s plate. Finally, the fourth specimen of C. pasti-
naca, which is not from Lamarck’s collection, resembles the form
figured by Reeve under this name. Thus it appears that, under the
name of C. pastinaca, the Delessert collection contains three species,
viz. the C. pastinaca of Kiener (young), that of Reeve, also young,
and, lastly, two small specimens of Conus quercinus. The question
which now has to be settled is this: —Is the specimen in Delessert’s
collection, which is similar to Reeve’s figure, really the same species
as that delineated? if not; may it not be the true pastinaca?
In the British Museum the shell which accords most closely with
the Lamarckian diagnosis is a worn specimen of C. tabidus, Reeve,
figured in the ‘Conchologia Iconica’ under the name of C. hepaticus,
Kiener (Conch. Icon. Suppl. pl. viii. fig. 278).
EXPLANATION OF PLATE XLVI.
Figs. 1, 2. Tellina wroblewskyi, p. 727.
. Trochus (Thalotia) yokohamensis, p. 727.
. Melania formosensis, p. 728.
Sormosensis, var., p. 728.
. —— dicksoni, p. 728.
obliquigranosa, p. 729.
obliquigranosa, var., p.
—— tuberculata, p. 729.
72
Wel
OH IG Sum go
1878.] MESSRS, GODMAN AND SALYIN ON NEW BUTTERFLIES. 733
Fig. 10. Melania subplicatula, p. 729.
11. Bullia persica, p. 730.
12. Ostrea, attached to a Trochus, whose character of sculpture it has
assumed, p. 730.
13, 14. Cyprea peasei, var. monstrosa, p.'731.
16. Descriptions of some apparently new Species of Butter-
flies from New Ireland and New Britain, sent by the
Rey. G. Brown. By F. D. Gopman and O. Satyin.
[Received June 18, 1878.]
The five species described below form part of a collection just
received from Mr. G. Brown. We are preparing a full report of
this collection, which we hope to lay before the Society in the
autumn. In the mean time we have thought it desirable to publish
the following descriptions.
EUPLa@a ERIMAS.
3. Exp. 3:7. Upper surface rich steel-blue, a large patch of
bluish white on the primaries cut by the dark median nervure and
its branches and by the nervules at the end of the cell; there is a
dark triangular spot in the acute angle between the median nervule
and its first branch; the inner margin of the primaries is strongly
convex, aud covers a brown patch on the secondaries; the secon-
daries have a whitish median band cut by the nervules, one section
falling inside the cell; the inner edge of this band is deeply sinuated,
as also is the outer edge to a less extent.
Beneath greenish black, the spots of the upper surface are much
more broken up, that on the primaries being divided into eight
separate portions, the largest of which lies just within the cell; that
on the secondaries is broken into eleven portions, one of which oc-
cupies the end of the cell.
Hab. New Ireland.
Oés. Allied to ZH. jessica from the Fiji Islands, and more remotely
to E. eupator, Hew., from Celebes. From the former it differs in
having the markings on the upper surface bluish white instead of
yellow, and the spot on the upper wing is not broken into two as
shown in Mr. Butler’s figure (Lep, Ex. pl. 8. fig. 3).
PIERIS MADETES.
d. Exp. 3°5. Above uniform orange, apex and costa of prima-
ries black.
Primaries beneath black, a large spot at the end of the cell and
seven large elongated submarginal spots yellow; a pale yellowish
patch spreads from the inner margin over the median branches to just
within the cell; secondaries orange, base and outer margin black,
the latter including a row of six large yellow spots; a large red spot
between the costal and subcostal nervures near their origin.
2. Exp. 3°8. Above, primaries black; a large yellowish spot at
734 GODMAN AND SALVIN ON NEW BUTTERFLIES. [June 18,
the end of the cell and six submarginal spots running from the costa
to the anal angle, those nearest the latter nearly white; the central
portion and inner margin of secondaries dull yellowish white, the
base and outer margin broadly black, the latter including six obso-
lete yellowish spots.
Beneath like the male, except that the primaries have no yellowish
spot spreading from the inner margin over the median branches.
Hab. New Ireland.
Obs. Belongs to the Delias group. The male resembles on the
upperside the same sex of P. aruna, Boisd., as figured by Hewitson,
and D. inferna, Butl. The female on the upperside, and both
sexes on the under, more nearly resemble P. descombresi, Boisd.
PizeRIS EURYGANIA.
3. Exp. 3:2. Above white, the apex of the primaries chrome-
yellow, margined with the black costa and outer margin and a curved
band running from inside the cell to near the anal angle, the outer
margin of the secondaries broadly black.
Beneath, primaries chiefly black, yellow apical spot as on the
upperside, the proximal end of the cell yellow, and the inner margin
white ; the secondaries at the basal half yellow, black towards the
distal half, across the black a distinct band of crimson, which becomes
orange where it meets the yellow near the inner margin.
@. Like the ¢, except that the basal portion of the primaries is
black, the outer margin of the secondaries is more broadly black,
and the inner portion of the secondaries, together with inner margin
of the primaries, pale yellow.
Beneath same as the male.
Hab. New Ireland.
Obs. This is a very distinct species, apparently belonging to the
Delias group.
PIERIS LYTHA.
Q. Exp. 2°5. Above black, a large subtriangular spot on the
inner margin, extending to just within the cell, three apical spots of
the primaries and the basal half of the secondaries white.
Beneath, black arranged as above, a submarginal row of large
white spots on both wings; the base of the primaries and the greater
portion of the base of the secondaries sulphur-yellow ; the rest of the
secondaries (not occupied with black) is white.
Hab. New Britain.
A female, apparently belonging to the Belenois group; we have
no nearly allied species with which to compare it.
PApILio SEGONAX.
3. Exp. 4°6. Primaries slightly faleate, secondaries produced,
Upper surface rich metallic green, a median row of five nearly cir-
cular spots starting from the apex to the space between the third
and fourth median nervules, a similarly coloured subtriangular spot
near the middle of the inner margin, and a very small one close
1878.] ON BIRDS COLLECTED BY H.M.S, ‘CHALLENGER. 735
inside the submedian nervure. The proximal half of the secon-
daries greyish white, rather darker towards the anal angle.
Beneath umber-brown, the outer portion of both wings rather
darker, the spots on the primaries as above.
Hab. New Ireland.
Obs. Allied to P. codrus, from which it differs in having the
band of spots of the primaries interrupted, there being no spots in
the spaces between the first and second, and second and third median
branches, that between the first median branch and the submedian
nervure being wanting. On the other hand, there is a spot on the
inner margin of the primaries not usually seen in P. codrus. The
single specimen sent by Mr. Brown has the secondary wings slightly
injured, so that the prolongation of the second median branch,
common to all the allied species of the P.-codrus group, is broken off.
17. Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘ Challenger..—No. XII. The Pro-
cellariide. By Ossert Savin, M.A., F.R.S., &c.
[Received June 18, 1878.]
Eighty specimens of Procellariidee are comprised in the ‘ Chal-
lenger’ collection, belonging to 22 species and 13 genera. The
greater part of these were obtained during the visit to Kerguelen
Island ; others were secured at sea in the South Pacific Ocean, a few
near the Straits of Magellan, at Nightingale Island, and elsewhere.
None of the species present any difficulty as regards their nomencla-
ture; but I take this opportunity of adding a few notes on this sub-
ject gathered during recent study of the species to which they
belong.
OCEANITES OCEANICUS.
Procelluria oceanica, Kuhl, Beitr. p. 136, pl. x. f. 1.
Oceanites oceanica, Coues, Pr. Ac. Phil. 1864, p. 82; Salv. Orn.
Mise. ii. p. 227.
a-c. Males.
d, e. Females.
} Ice Barrier.
‘Obtained on the 14th February, 1874.”’
FREGETTA GRALLARIA.
Procellaria grallaria. Vieill. N. Dict. d’Hist. N. xxv. p. 418.
Fregetta grallaria, Bp. Consp. ii. p. 197; Coues, Pr. Ac. Phil.
1864, p. 86.
Thalassidroma leucogastra, Gould, Ann. & Mag. N. H. xiii. p.
367, et B. Austr. vii. pl. 63.
568-571. Females. :
Bb, 578. Males. _f South Pacific.
“Eyes brown. Shot on the 11th November, 1875, at sea. Their
736 MR. 0. SALVIN ON THE PROCELLARIIDE [June 18,
stomachs were filled with a yellow oil, and mixed with it some pieces
of Crustacea.”
I am strongly disposed to add to the above synonyms Thalassi-
droma gracilis, Elliot, Ibis, 1859, p. 391, and T. segethi, Phil. &
Landb. Arch. f. Nafurg. 1860, p. 282, names admitted by many
authors to apply both of them to a species found on the Chilian coast.
I have seen specimens of the present species in the Paris Museum
from the same locality ; and the Challenger specimens now before
me were obtained at sea at no great distance from the same shore ; so
that on geographical grounds their identity is a reasonable supposi-
tion. The chief, if not the only discrepancy I can find between the
description of F. gracilis and F. grallaria is one of dimensions, the
former being considerably smaller (as shown by Mr. Elliot’s measure-
ments) than the ‘Challenger’ birds. Thus we have :—
Wing. Tarsus. Tail.
in. in, in.
F. gracilis ..... 5°22 2°40 1:20
F. grallaria .... 5°9-6°3 2°9-3°0 1-3-1°4
T. segethi seems to be intermediate in dimensions ; but so far as
the description goes, its identity with /. grallaria is obvious.
Mr. Sharpe (Zool. Kerg., Aves, p. 31) suggests that P. leuco-
gastra may prove to be a stage of plumage of P. melanogaster; but
I cannot think that this will ever turn out to be the case.
FREGETTA MELANOGASTRA.
Thalassidroma melanogastra, Gould, Ann. & Mag. N. H. xiii.
p. 367, et B. Austr, vii. pl. 62.
a. Betsy Cove, Kerguelen.
Mr. Sharpe (Zool. Kerg., Aves, p. 30) unites 7. tropica, Gould,
with this bird, and calls it by the last-mentioned name. They may
be identical ; but in any case the adoption of the name ¢ropica in
place of melanogastra cannot be justified, as the latter, having been
in use for over thirty years, ought not to be supplanted by the
former, the two having been published simultaneously on opposite
pages of the same publication !
PELAGODROMA MARINA.
Procellaria marina, Lath. Ind. Orn. p. 410.
Pelagodroma marina, Salv. Orn. Mise. ii. p. 228.
a-c. Nightingale Island.
_ “Byes black; anight-bird. These were taken out of holes in the
ground during the day by help of the dogs.”
THALASSGECA GLACIALOIDES.
Procellaria glacialoides, Smith, Ill. Zool. S. Afr. pl. 51.
Thalasseca glacialoides, Coues, Pr. Ac. Phil. 1866, p. 30.
a. Male. Ice Barrier.
‘Obtained on the 26th February, 1874.”
1878. | COLLECTED BY H.M.S. ‘ CHALLENGER.’ 737
THALASSGCA ANTARCTICA.
Procellaria antarctica, Gm. Syst. Nat. i. p. 565.
Thalasseca antarctica, Coues, Pr. Ac. Phil. 1866, p. 31.
a. Male. Ice Barrier.
Obtained on 14th January, 1874.
6. Male. Fors Banc:
c, d. Females, f “°° ?@™™er-
Obtained on the 14th January, 1874.
ADAMASTOR CINEREUS.
Procellaria cinerea, Gm. Syst. Nat. i. p. 563.
Adamastor cinereus, Coues, Pr. Ac. Phil. 1864, p- 119.
ope. Pemale- | South Pacific.
“Byes hazel; feet flesh-colour: the stomach of one was full of
the beaks of Cuttlefish ; stuff from the ship in the other, and small
crustacea.”’
“Obtained on the 5th November, 1875.
MAJAQUEUS ZQUINOCTIALIS.
Procellaria equinoctialis, Linn. Syst. Nat. i. p. 213.
Majaqueus equinoctialis, Coues & Kidder, Bull. U.S. Nat. Mus.
No. 2, p. 25; Sharpe, Zool. Kerg., Birds, p. 19.
-d. Males.
e Bede. } Kerguelen,
OssIFRAGA GIGANTEA,
Procellaria gigantea, Gm. Syst. Nat. i. p. 563.
Ossifraga gigantea, Coues, Pr. Ac. Phil. 1866, p. 32.
567. Female. South Pacific.
** Bill yellowish green, feet black.”
Obtained on the 5th November, 1875.
a. Betsy Cove, Kerguelen.
PAGODROMA NIVEA.
Procellaria nivea, Gm. Syst, Nat. i. p. 562.
Pagodroma nivea, Coues, Pr. Ac. Phil. 1866, p. 160.
a, 6. Females. Ice Barrier.
“Obtained on the 14th January, 1874.”
DAPTION CAPENSIS.
Procellaria capensis, Linn. Syst. Nat. i. p. 213.
Daption capensis, Coues, Pr. Ac, Phil, 1866, p. 163.
a. Antarctic Sea.
“Shot in April 1874.”
CESTRELATA LESSONI.
Procellaria lessoni, Garn, Ann. Se. Nat. 1826, vii. p. 54.
Cistrelata lessonia, Coues, Pr. Ac. Phil. 1866, p- 142; Coues &
Proc. Zoou. Soc.—1878, No. XLVIII. 48
738 MR. 0. SALVIN ON THE PROCELLARIID# [June 18,
Kidd. Bull. U.S. Nat. Mus. No. 2, p. 27; Sharpe, Zool. Kerg.
Birds, p. 26.
a, 6. Males. Betsy Cove, Kerguelen.
c-f. Females. Betsy Cove, Kerguelen.
“ Eyes black.”
During a recent visit to the Paris Museum I had an opportunity
of examining the type of @. sericea, concerning which I had occa-
sion to publish some notes in ‘The Ibis’ for 1875 (p. 373). On
seeing this type I at once recognized a specimen of @. lessoni! GH.
sericea (Less.), placed in the genus ddamastor by Bonaparte and
Coues, has therefore no existence as a species distinct from @.
lessoni.
CEsTRELATA MOLLIS.
Procellaria mollis, Gould, Ann. & Mag. N. H. xiii. p. 363.
CEstrelata mollis, Coues, Pr. Ac. Phil. 1866, p. 150.
a, 6. Nightingale Island.
“Eyes hazel. Night bird. Obtained on the 17th of October,
1875.”
CEsTRELATA BREVIROSTRIS.
Procellaria brevirostris, Less. Man. d’Orn. ii. p. 611.
Estrelata brevirostris, Salv. Orn. Misc. ii. p. 235; Sharpe, Zool.
Kerg., Birds, p. 24.
Cstrelata kidderi, Coues, Bull. U.S. Nat. Mus. No. 2, p. 28.
a. Young. Kerguelen.
The species included in Cabanis and Reichenow’s paper on the
Birds collected by the ‘ Gazelle’ (J. f. Orn. 1876, p. 329) under the
name Procellaria mollis, belongs here. Last year I examined in
Berlin the specimen brought home by the ‘ Gazelle,’ and found it to
belong to this species.
The ‘Challenger’ specimen is quite young, but interesting, as
showing that these birds moult at once from the downy plumage into
the feathering of the adult bird.
PRION DESOLATUS.
Procellaria desolata, Gm. Syst. Nat. i, p. 562.
Prion desolatus, Gray, H.-list. iii. p. 108 ; Sharpe, Zool. Kerg.,
Birds, p. 37 (excl. syn.).
Pseudoprion desolatus, Coues, Bull, U.S. Nat. Mus. No. 2, p. 32.
a. Male. Betsy Cove, Kerguelen.
Obtained 11th January, 1874.
** Eyes black.”
b-e. Males. Betsy Cove, Kerguelen.
‘Eyes black.”
f. Female. Ice Barrier.
Obtained on the 14th February, 1874.
g, h. Females. Betsy Cove, Kerguelen.
Of the eight specimens of this species brought home by the ex-
1878. | COLLECTED BY H.M.S. ‘CHALLENGER.’ 739
pedition, five are males and three females. They thus offer an
opportunity of testing the view advanced by Mr. Sharpe as to the
supposed sexual difference in the size of the bill existing in these
birds’.
In these specimens there is no tangible difference between the bills
of the sexes; and as the opposite sexes of P. banksi, as shown by the
two ‘ Challenger’ specimens, are also alike as regards the size of the
bills, doubts may fairly be raised whether the great difference obser-
vable in the size of the bills in these birds is a sexual character at all.
PRION BANKSI.
Pachyptila banksi, Smith, Ml. Z. 8. Afr. Birds, pl. 55.
Pseudoprion banksi, Coues, Pr. Ac. Phil. 1866, p. 166.
a. Male. Marion Island.
«yes brown.”
6. Female. ‘Caught at sea near Crozets.”’
Without attempting to decide the question as to how many species
of Prion exist, I use the name P. banksi for these birds, as their
bills agree most nearly with that of the type of P. banksi in the
British Museum.
PELECANOIDES URINATRIX.
Procellaria urinatriz, Gm. Syst. Nat. i. p. 560.
Pelecanoides urinatrix, Coues, Pr. Ac. Phil. 1866, p. 190; Sharpe,
Zool. Kerg., Birds, p. 14.
a. Male. Kerguelen.
“‘ Eyes dark grey.”
6. Male. Betsy Cove, Kerguelen. Obtained January 1874.
e. Male. Christmas Harbour, Kerguelen.
d. Male. Betsy Cove, Kerguelen.
e-h. Females. Kerguelen.
2. Kerguelen.
j- Young. Kerguelen?
PELECANOIDES GARNOTI.
Puffinuria garnoti, Less. Voy. Coq. Zool. pl. 46.
Pelecanoides garnoti, Coues, Pr. Ac. Phil. 1866, p- 190.
ne Aa } Cove Harbour, Messier Channel.
** Eyes hazel. These birds and another one which was put in spirit
were found dead. Two were picked up by the boat in which Cap-
tain Maclear and I went, floating on the surface of the water; and
the third was picked up by the galley on the rocks. They were quite
fresh ; the bodies were very thin.
“Their stomachs were filled with small leaves. It looks very much
1 Latham seems to have been the first author who mentioned this supposed
sexual difference in Prion, After describing P. vittatus (Syn. iii. p.414), he adds:—
“ The female has the same plumage ; but the bill, though greatly exceeding that
of any other Petrel, is scarcely more than half the breadth of that of the “ male.”
740 ON BIRDS COLLECTED BY H.M.S. ‘CHALLENGER. [June 18,
as if these birds had got out of their region, Mr. Moseley says he
saw what he supposed to be one of these birds on Penguin Island.”
DIOMEDEA EXULANS.
Diomedea exulans, Linn. Syst. Nat. i. p. 214; Coues, Pr. Ac.
Phil. p. 866, p. 175; Coues & Kidder, Bull. Nat. Mus. No. 2, p. 11;
Sharpe, Zool. Kerg., Birds, p. 45.
574. Kerguelen.
DioMEDEA BRACHYURA.
Diomedea brachyura, Temm. Pl. Col. 554; Coues, Pr. Ac. Phil.
1866, p. 177.
517-519. Males. ) 1 .
590. Female. North Pacific.
‘These were all caught with the hook from the ship while at sea
in June and the first half of July 1874, between Japan and Honolulu ;
they followed the ship every day in numbers till we got into the
trade-winds, when no more were observed.”
509. Female. North Pacific.
‘Eyes brown, bill black, stomach empty. Shot on the Ist April,
1875, by Lord Campbell, with the Henry rifle, while on thewing. We
were just north of the tropic; but this bird, as well as another Alba-
tross, were seen some days before we had passed out of the tropics.”
511. Male. North Pacific.
«Eyes brown, feet and bill dark or nearly black ; stomach had
cuttlefish. Caught with a hook by Percy on the 7th "April, Pia.”
DIoMEDEA MELANOPHRYS,
Diomedea melanophrys, Temm. PI. Col. 456 ; Coues, Pr. Ac. Phil.
1866, p. 181; Sharpe, Zool. Kerg., Birds, p. 46.
Male.
Bele. } Shot at sea, 8th July, 1874.
Sar } Christmas Harbour, Kerguelen.
DIOMEDEA FULIGINOSA,
Diomedea fuliginosa, Gm. Syst. Nat. i. p.595 ; Sharpe, Zool. Kerg.,
Birds, p. 48.
Phebetria fuliginosa, Coues, Pr. Ac. Phil. 1866, p. 186; Coues &
Kidder, Bull. Nat. Mus. No. 2, p. 2].
Male. See Barnier, 16th February, 1874.
Female. Royal Sound, Kerguelen, 17 January.
1878. | M. DU BOCAGE ON ANGOLAN ANTELOPEs. 741
18. Liste des Antilopes d’Angola. Par J. V. Barpoza pu
Bocace, F.M.Z.S.
[Received June 17, 1878.]
La présente liste des espéces d’Antilopes observées en Angola a
éte dressée d’aprés les exemplaires qui existent au Muséum de
Lisbonne. La plupart de ces individus ont été recueillis dans
Pintérieur de Mossamedes et au Humbe, sur Ja rive droite du Cunene,
par notre infatigable explorateur M. de Anchieta.
1, GazeLLA EUCHORE (Forst.).
Une paire d’individus adultes envoyés de Huila par M. de Anchieta.
Le blane de la téte, de abdomen et des membres est remplacé chez
la femelle par une teinte uniforme d’un fauve-jaunatre pale.
2. AEPYCEROS PETERSI, sp. nov.
Nous possédons également une paire d’individus adultes de cette
espéce, qui nous semble nouvelle. Le male, que nous avions recu
dabord, nous avait paru identique & ! Zpyceros melampus daprés
un premier examen ; mais ayant recu plus tard la femelle, nous avons
été frappé de quelques particularités de coloration qui nous semblent
autoriser leur distinction spécifique. Nous commencerons done par
donner un rapide apercu des caractéres de la femelle.
Pelage d’un roux-fauve, plus pale sur les flancs et Ja face externe
des membres; menton, gorge, poitrine, ventre et partie interne des
cuisses et de l’avant-bras d’un blanc presque pur. La téte porte sur
un fond de la couleur du dos quelques marques trés-caractéristiques :
le dessus du chanfrein, une bande au travers de l’ceil et une tache
en fleur-de-lys sur le front d’un noir profond; au devant de
Poel une bande blanche bien distincte. Oreilles longues, arrondies
A la pointe, d’un roux-fauve en dehors, revétues de poils blancs en
dedans, avec un espace noir 4 l’extrémité, plus large en dehors qu’en
dedans. Queue longue, de la couleur du dos a la base, le reste
blanc, marquée en dessus d’une bande longitudinale noire, a laquelle
vient aboutir de chaque cété une autre bande étroite de la méme
couleur qui s’étend sur les fesses. Un trait noir coupant verticale-
ment la face antérieure du genou, une tache de la méme couleur a
la pointe du jarret. Aux membres postérieurs, de chaque coté de
Pextrémité inférieure du canon, une brosse de poils noirs. Point de
mufle et point d’onglons. Deux paires de mamelles.
Dimensions. métre.
Du bout du museau 4a la base de la queue ........ 1°30
ReMECUL MU OMTOE: ooo nag oe eo aw 2s ca ew avers, OBO
Longueur de la téte .-...... Se a es a
Biperciye Ge VOLEMIG: or Oo noe whit +) + eiroretsl se aha
Longucur de la queue | ht Rie palitens A axtoopy Pi
Proc. Zoou. Soc.—1878, No. XL ix 49
742 M. DU BOCAGE ON ANGOLAN ANTELOPES. [June 18,
Chez notre individu male, plus jeune que la femelle, nous
remarquons des curnes en forme de lyre fortement annelées sur leur
tiers inférieur. II présente quelques légéres différences de coloration.
La teinte des parties supérieures est d’un bai-roux plus foncé et le
blanc des parties inférieures moins pur, plus lavé de jaune-isabelle;
le dessin de la téte ressemble 4 celui de la femelle, quoique le noir
y soit plus rembruni. Malheureusement une portion de la peau
de la région frontale 4 été détruite par suite du coup qui l’a tué, de
sorte qu’on ne peut pas constater le présence de la tache noire si
nettement indiquée sur la région frontale de la femelle.
Les cornes portent trois bourrelets étroits mais saillants sur
leur tiers inférieur, supérieurement elles sont lisses et arrondies ;
leur courbure est simple, 4 convexité en dehors et a pointes
convergentes.
Dimensions du male. métre.
De l’extrémité du museau a la base de la queue.... 1°20
Hautenr ai garrot. ois. aj ais cere ait « 1) Ls eee
Sionpnenmde da tGtey: ii. 25 sung aye vag es a ae
Bonedeur der oreile tric. wok. vy + cn ceneic denne. = ae eeetee
Longneur We Taqnene sca) ayayetn «oe diate Se oak eee
Hoveneur Ues/eormess os). i ees weet s bie oy. onmeliee
Le male nous vient de Capangombe, la femelle du Humbe, tous
les deux par M. d’Anchieta’.
3. NANOTRAGUS TRAGULUS (Forst.) ?
Nous rapportons avec hésitation 4 cette espéce une jeune femelle
que M. d’Anchieta nous a envoyée du Humbe en 1874. Voici ses
principaux caractéres :—
Pelage d’un roux-fauve, plus pale sur les flanes et les membres ;
gorge et ventre d’un blanc pur; front et chanfrein de la couleur du
dos ; les jones d’une teiute plus pile ; tour des yeux blanchatre ; une
tache brune en forme de fer 4 cheval sur le front ; une autre petite
tache triangulaire brune sur le chanfrein immédiatement au-dessus
du mufle, qui est petit. Oreilles trés-longues, légérement acuminées,
grises en dehors, blanches en dedans avec un liséré brun sur les
bords. Point de brosses aux genoux; point d’onglons; queue,
incomplete, de la couleur du dos. Quatre mamelles.
Dimensions. metre.
De l’extrémité du museau a la base de la queue .... 0°75
IEVATCC UT ALN ATTOb Sees ee - «ee oretel's oe apes ws ores 0°45
Peoneteur de letéte «2254. fs. os ara sah ips 0-14
5 detlagliovenlem ss free. eee ie ens ae 011
* La figure qui accompagne cette note permettra de bien juger les caractéres
de coloration de la femelle. Notre ami le Prof. Peters, 4 qui nous l’ayons com-
niuniquée, penche a croire, comme nous, qu’il s'agit d’une espéce nouvelle.
1878. ] M. DU BOCAGE ON ANGOLAN ANTELOPES. 713
4, NeorraGus SALTIANUs (Blainv.).
Des individus de Capangombe et du Humbe, sur la rive droite du
Cunene, par M. d’Anchieta.
CrePHALOPHUs GRiIMMivs (Linn.).
Une paire d’individus adultes et un jeune male de Capanugombe
par M. d’Anchieta.
Il y a méme au Muséum d’autres individus de cette espéce
proveuant d’ Angola, mais dont on ignore la localité of ils auraient
été pris ; quelques uns de ces individus ont vécu pendant quelques
années au Jardin du Palais de Necessidades appartenant a sa
Majesté le Roi Ferdinand, et s’y sont méme reproduits.
Tous nos individus nous sembleut appartenir par leur syst¢me de
coloration a la variété dont le Dr. Gray a fait une espéce distincte
sous le nom de Grimmia splendidula (Proc. Zool. Soc. Lond. 1871,
p. 590). Nous les avions d’abord rapportés au C. burchellii (Journ.
Acad. Sci. Lisboa, ii. p. 222).
6. CerpHaLoruus monticoxa (Thunb.).
Deux individus, male et femelle, rapportés de Benguella par M.
Freitas Branco; un individu male envoyé de Loanda par M.
Toulson.
CEPHALOPHUS ANCHIETA, Sp. lov,
Une paire d’individus adultes et une jeune femelle capturés a
Capangombe par M. d’Anchieta,
Caractéres :—Brun-ceudré ; les flanes, l’avant-bras et les cuisses
d’une teinte plus pale et plus cendrée; dessus de la téte d’un cendré
noiratre, avec une touffe de poils noirs entre les cornes ; cdtés de la
téte et du cou de la couleur des flanes; raie Joureilived roussatre ;
menton, gorge, poitrine, veutre et face interne des avant-bras et des
cuisses d’un blanc pur ; face antérieure et externe des inembres d’un
cendré lavé de fauve ; queue trés-courte, d’un noir profond en dessus ;
la région anale de cette méme couleur. Cornes etfilées.
Dimensions. metre.
De lextrémité du museau a la base de la queue.... 0°58
eomemenr della tGte® is. 005. tt Syd tahoe be de 2 4 Orl4
Pinsitenr auearrotycils its ase sian cedatliow- doavsts . 0°33
iomsenn de, Ppreilles sc. alice, ists stews aes. tyes Sheers 0:07
5 BGM HACE YriGt nis ebhima tid sehen saw sls elOOZ
La teinte générale est plus rembrunie chez la jeune femelle, les
membres sont plus lavés de fauve 4 leur partie inférieure, le dessus
de la queue et la région anale d’un brun foncé.
Nos individus doivent ressembler sans doute a C’. melanorrhous,
49*
744 M. DU BOCAGE ON ANGOLAN ANTELOPES. [June 18,
Gray, de Fernéo do Pé, que nous connaissons 4 peine d’aprés la
planche publiée dans Knowsley Menagerie (Gray, Knowsl. Ménag.
p. 11, pl. 10); ils en différent cependant par le moins d’étendue
du noir, qui couvre a peine le dessus de la queue et la région
anale, tandis que , chez le C. melanorrhous la croupe est aussi de
cette couleur. A juger d’aprés les figures du Dr. Gray, cette
espéce aurait une queue plus longue que la nétre et des cornes
moins effilées. Malheureusement le Dr. Gray dans ses diverses pub-
lications sur les Antilopes a presque toujours oublié de nous donner
les dimensions des individus qu'il décrit.
8. CEPHALOPHUS RUFICRISTA, Sp. Nov.
C. longiceps, Bocage, Jorn. Acad. Sci. Lisboa, ii. p. 220.
Cette espéce se trouve a peine representée au Muséum de Lisbonne
par une téte, recouverte de sa peau, envoyée en 1869 de Loanda par
M. Toulson, sans s’accompagner d’aucune indication au sujet de sa
provenance.
Nous avions d’abord pensé qu’elle pourrait bien étre identique a
O. longiceps, Gray, espéce établie exclusivement d’aprés les caractéres
ostuologiques d’une téte recueillie au Gabon par Du Chaillu (Proc.
Zool. Soe. 1865, p. 204); aujourd’hui nous revenons sur nos pas, car
notre premicre affirmative nous semble trop hasardeuse.
Le spécimen du Muséum de Lisbonne ressemble aussi par ses
couleurs a la figure du C. niger, publiée par le Dr. Gray dans
Knowsley Menagerie, pl. 7. Cependant, méme sous ce rapport, leur
identité specifique ne nous parait pas évidente, et a cela vient encore
s’ajouter une différence sensible dans les dimensions d’apres les
chiffres publiés par Temminck. Nous trouvons dans les Esquisses de
Mammalogie de cet auteur que la distance du bord antérieur de
Yoeil A la pointe du nez chez C. pluto (=C. niger, Gray) est de 4
pouces et 4 lignes, et en mesurant cette distance sur la téte d’Angola
nous constatons qu’elle dépasse 5 pouces; elle est de 14 centi-
métres.
La face supérieure le la téte est d’un brun clair sur le chanfrein
et d’un brun foncé, couleur chocolat, sur le front ; les faces latérales,
en haut d’un gris brunatre, deviennent blanchatres en se rapprochant
du menton, ot le blanc domine ; une raie sourciliére brun-jaunatre ;
une tache allongée de la méme couleur au-dessous de l’ceil se pro-
longeant vers le chanfrein ; tour des yeux brun-foncé ; ligne muqueuse
Jégérement courbe et horizontale. Sur le front un toupet abondant
de poils bruns et d’un roux ardent, en partie couchés sur les cornes,
en partie sélevant verticalement au milieu de ces appendices; la
touffe verticale est composée de poils longs de 7 centimétres d’un
roux vif, les deux touffes latérales sont formées de deux ordres de poils,
les uns courts et bruns, les autres longs et roux. Les oreilles sont
petites, mesurant moins de la moitié de la longueur totale de la téte ;
elles sont arrondies au bout, couvertes en dehors de poils trés-courts
et clair-semés, nues en dedans avec une large bande marginale de
poils blanchatres. Les cornes arrondies, a peine rugueuses a la
PZS 1878.F
re}
real
NGALENSI
ar
MEY N CHASA-BE
fs
PRACIISA “O
HH Godwin Ansten. del et Lith
ita
1878. | ON THE- TRACHEA OF RHYNCIIHA CAPENSIS. 745
base, sont couchées en arriére et placées sur le plan du front; elles
ont 10 centimetres de longueur. Le mufle est régulier.
Dimensions. :
metre,
Longueur de l’occiput 4 l’extrémité du museau.... 0°26
Du bord de l’ceil 4 Vextrémité du museau ........ 0°14
Longueur de l’oreille..................00.0..-. 0°95
am Gesvearhes) .,..Jgeuatics Amctiodt dawn. Or0
uy de la touffe frontale.................. 0:07
9. HELEorrRaAGus REDUNCUS (Pall.).
Un male adulte de Huilla par M. d’Anchieta.
10. HirpporraGus nicer (Harris).
La téte d’un individu male envoyée de l’intérieur de Mossamedes
par Welwitsch. Les cornes de cet spécimen ont 130 centimetres
de longueur.
11. SrRepsicERos kuDU, Gray.
Deux individus adultes, mile et femelle, envoyés vivants d’ Angola
et ayant vécu longtemps dans la ménagerie du Jardin de Neces-
sidades.
Un jeune, de quelques jours 4 peine, né des précedents, tué par
le male, qui était d’une grande férocité.
Une jeune femelle de Capangombe, par M. d’Anchieta,
12. Orgs CANNA, Gray.
Un male jeune d’ Angola.
19. On the Structure and Development of the Trachea in
the Indian Painted Snipe (Rhynchea capensis). By
J. Woop-Mason.
[Received June 18, 1878.]
(Plate XLVIL.)
During the cold season of 1876-77, Lieut.-Colonel Godwin-
Austen and I paid almost daily visits to the Calcutta bazaar for the
purpose of making collections of the skins and skeletons of the
numerous migratory and other birds which are at that season of the
year caught and carried to market in such enormous numbers. On
one of these visits my attention was attracted to a bird that lay con-
spicuous amongst its fellows of the same species by its greater size
and more richly coloured plumage, by the low and regular, hoarse
746 MR. J. WOOD-MASON ON THE [June 18,
but rich, purring call which, with breast puffed out, it was uttering—
a call perceptible to the hand no !ess than to the ear. I at once re-
cognized the smaller and less conspicuously coloured birds as speci-
mens of one sex; and there seemed very little doubt, from its perfect
correspondence in structure and general similarity in plumage, that
the larger and handsomer one was an individual of the other sex of
the Indian Painted Snipe (Rhynchea capensis, sive bengalensis);
but which of the two was the female and which the male, it must be
confessed, I was at the time ignorant. In order that I might be
enabled to determine the precise relation of the two forms to one
another, and to ascertain whether any structural differences in their
vocal organs accompanied the observed differences in their vocal
powers—merely that I might know these facts of my own knowledge,
certainly without the slightest hope or thought that I should be able,
from such a cursory examination as alone I could give to it, to glean
any thing new about so common an animal, I purchased a pair of
the species.
Before killing the birds for examination, I referred to Darwin’s
‘Descent of Man’'; and what I read therein served but to increase
my interest in the matter; for I soon saw that I hadit in my power
to corroborate or to contradict a statement which had been made
about the trachea of this very species—the very part, curiously
enough, the sounds issuing from which had drawn my attention to
the bird.
It is well known that in many birds the windpipe, instead of
taking a straight course from the rima glottidis through the inter-
clavicular membrane to the point where it bifurcates to form the
bronchi, is bent upon itself or convoluted, and that often to an ex-
traordinary extent. The position of such flexures is very variable:
‘they may lie outside the thorax under the integument (as in Te¢rao
urogallus, some species of Craxzx and Penelope, aud, I may add, the
Manucodias’ and the Rhyncheeas), in the cavity of the thorax (as in
some Spoonbills), on the exterior of the sternum (as in some Swans
and Cranes), or even in a sort of cup formed by the median process of
the furcula (as ina species of Guinea-fowl)’’*. ‘The increase in the
length of the tracheal column implied by these convulutions, and
other modifications of the windpipe, such as the swollen tympanum of
Ducks and Geese, the air-sac of the Emu, may be dependent wholly
upon sex; aud the males may have the trac! amore or less looped or
more complex, whilst the female: jsaye +: s.:g*ht or simple, or only
partially so, in which case the pee a eae modification may be
more marked in males than in femzi##:* Gut however this may be,
it is a general rule that whenever ‘ the trachea differs in structure
in the two sexes it is more developed and complex in the male than
in the female’’*; and it is a fact familiar to all that in the vast
majority of instances it is the male which, in point of richness of
plumage, vocal powers, and ornamental appendages, is the more highly
1 Op. cit. p. 476. ? ‘Nature,’ yol. xv. p. 127.
“ Huxley, ‘Anatomy of Vertebrated Animals,’ ». 315,
* «Descent of Man,’ l. supra cit.
1878. ] TRACHEA OF RHYNCH#A CAPENSIS. 747
endowed of the two sexes; but to this there are a few interesting
and remarkable exceptions (Phalaropus, Casuarius, Dromeus, Mil-
vago, Climacteris, Eurostopodus, and the Rhynchzas), of which
the last, the ones we are here concerned with, are not the least
conspicuous. In these birds in general, and in the Painted Snipes
in particular, it may be taken as established that we have, to use
the words of Mr. Darwin, a complete reversal not only of the
secondary sexual characters, but also of the parental and incubating
instinects—the females being not only larger and much more richly
coloured than the males’, but having the trachea more or less
tortuous instead of straight and simple, deputing the duty of incuba-
tion to the other sex, and reserving the business of courting to them-
selves.
In Rhynchea australis, according to Gould?, the trachea, which is
simple in the males, in the females passes down between the skin
and the muscles of the breast for the whole length of the body,
making four distinct convolutions before entering the lungs; but
Mr. Darwin states, on the authority of Blyth, who had examined
many specimens, that “it is not convoluted in either sex in Rh.
bengalensis, which species resembles R. australis so closely that it
can hardly be distinguished except by its shorter toes.” This is the
statement which seemed to me to stand in need of corroboration,
especially when I called to mind the peculiar call of the female* and
the sharp squeak jerked out only at long and irregular intervals by
the male, and then apparently only in answer to the female.
On opening the necks of the two birds by a longitudinal incision
extending to the middle of the breast or thereabouts, and carefully
turning aside the skin on either hand so as not to disturb the natural
rélations of the underlying parts, I found that the trachea of the
adult male (ascertained to be such by subsequent examination of the
genital organs) was straight and simple throughout, whilst that of
the female had a distinct loop lying between the integument and the
interclavicular membrane on the left side, and was not only an abso-
lutely but apparently also a relatively stouter tube than that of the
male.
As the contrary of what I have found has been stated by so good
and usually trustworthy an observer as Mr. Blyth, I put in evidence
two sketches (figs. 1 and 2, p. 748) showing the course of the trachea
in the two sexes. For these sketches I am indebted to Col. God-
win-Austen, who was with me at the time.
The numerous birds belonging to this species examined by me may
be divided according to sex and age into the following groups :—
1. Adult and probably old females, remarkable for the extreme
richness of their plumage. In all the birds of this group which
* Jerdon, ‘ Birds of India,’ vol. iii. p. 677.
2 «Handbook to the Birds of Australia,’ vol. ii. p. 275.
* Of R. australis J. Gould says (op. cit. p. 276), “The use of this convoluted
trachea, so exclusively confined to the female, I could not in any way discover
or surmise; no note whatever was heard to proceed from either sex while on
the wing or when flushed ”—times at which a call such as that of the female of
R. capensis, like the coo of a doye, would be least likely to be heard.
748 MR. J. WOQD-MASON ON THE {June 18,
were purchased alive and killed and examined before muscular con-
traction had set in, the trachea was found to be looped as in fig. 1 ;
a specimen living when obtained, but which died unexpectedly and in
which decomposition had set in before I had had an opportuuity of
Rhynchea capensis 3, dissected, so as to show its straight and simple trachea
in situ.
opening it, also had its windpipe looped ; several, however, that were
purchased dead had it retracted, almost straight, with the sterno-
tracheal muscles strongly contracted.
2. Females not nearly so richly and deeply ccloured as the pre-
ceding. In these there was only a slight superficial sinuosity in
re
1878.] TRACHEA OF RHYNCH4A CAPENSIS. 749
the trachea, and the remarkable modification of the intrathoracic
rings, to be presently described, had not proceeded so far.
3. Immature females, indistinguishable in point of plumage, though
already larger than males. In all of these, without exception, the
windpipe was found quite straight and simple throughout, though
even at this stage females are infallibly to be distinguished from males
by their stouter trachea, by the more powerful musculature of this,
and by the more inflated condition of the delicate membranes
connecting the bronchial half-rings with one another and with the
three-way piece.
4. Young, adult, and apparently old males, all agreeing together
in plumage and in the straight and simple condition of the trachea.
We shall see that the superficial loop which is invariably to be found
in birds belonging to group 1 (old females) is the outward expres-
sion, so to speak, of a modification of the intrathoracic tracheal rings
that takes place pari passu with those external changes which, when
they are completed, mark the adult. . But in order to make my
description of this curious modification more intelligible, a few
words, by way of preface, about the unmodified trachea of the male,
or, better, of an immature female. If such a trachea be drawn
through the fingers from end to end, a broad and shallow constric-
tion will be felt near its posterior end, twenty rings or so from the
compound three-way piece; the rings composing it are cylindrical
instead of flattened, more than thrice as numerous as they are in an
equal length of any other part of the tube, and so closely packed
and firmly bound together as to possess little or none of that power
of expansion and contraction which, by reason of their peculiarly
bevelled ends, so eminently distinguishes the rest; it occupies a
position as much within as without the thorax ; and the great extrinsic
muscles which pass between the sternum and the trachea, serving
amongst other purposes as ‘‘guys’’ to keep the latter in place,
expand and unite sheath-like over it, being inserted into it at nume-
rous points, but especially at its anterior extremity ; it is, in fact, the
part of the trachea upon which the sterno-tracheal muscles directly
pull when they contract, and thereby approximate the rings of the
extensible intrathoracic portion of the windpipe in the adult female,
to which we may now return.
On more closely examining the extrathoracic portion of the trachea
in a bird in which that part is in the condition represented in fig. 1,
p- 748, it can be seen that the loop is almost wholly composed of the
constricted portion above described, and that the 2 or 3 rings that
immediately succeed in order from before backwards (those situated
at the point where the tube disappears within the cavity of the chest)
suddenly get coarser and more prominent, and at the same time
separated from one another by perceptible membranous intervals.
On cutting away the sternum so as not to sever the sterno-tracheal
muscles from their attachments, and so as to leave the furcula together
with the membrane: included between its two arms in place, the rest
of the tube is displayed in a completely extended condition ; it is
then seen that the 12 or 13 rings immediately succeeding the coarser
750 MR. J. WOOD-MASON ON THE | June 18,
ones which follow upon the fine-ringed constricted portion become
suddenly still coarser and more prominent and convex, have lost
their typical bevelled form, and are separated from one another by
far longer and subequal membranous intervals. These are thin and
transparent, and each is so constricted in the middle that any two
consecutive rings form, with the membrane that connects them, a
short hourglass-shaped figure. Six or seven coarsish rings, sepa-
rated by much narrower membranous intervals, but otherwise un-
modified, complete this portion of the tube, which is twisted spirally
to the left, carrying spirally entwined with it the elongated sterno-
tracheal muscles. The rings have come to be bent, and, instead
of appearing as regular parallel bands as in other parts of the column,
to be arranged obliquely, by adaptation, no doubt, to the spiral form
taken by the tube whenever it is extended, so’as, in fact, to present
somewhat the appearance of having resulted from the breaking up
into rings of what was primitively a hard spiral thickening of the
walls of a membranous tube. Inside the thorax the spiral is neces-
sarily very open, from the tube being restricted to a middle position
by the membranous bands which sling it, together with the anterior
moieties of its great contractor muscles, from the dorsal wall of the
body, so as to form a sheath for it; whilst outside, in the neck,
where is more room and more lateral freedom, it becomes closer,
there constituting the well-marked superficial loop, the concave cur-
vature of which is the true ventral surface of that fine-ringed portion
of the tube over which the muscles spread, twisted out of its natural
pesition.
If an adult female be killed with ehloroform and rapidly opened,
the sterno-tracheal muscles may be seen slowly to contract, and
thereby gradually to take out the superficial spiral ‘‘turn”’ from the
trachea, and the intrathoracic rings of this to close up, until at last
all that remains of the loop is a slight sinuosity visible near the
point where the tube passes into the thorax, the constricted and
close-ringed portion, which occupied that position in the unmodified
trachea, also having acquired an ineffaceable crook.
Unfortunately, we possess no more complete description of the
highly convoluted trachea of the Australian species than that quoted
above ; but the two species R. capensis and R. australis are so very
closely allied that we may feel tolerably confident that the tracheal
modification is of the same kind in the two, only carried to a much
greater extent in the latter, the constricted many-ringed part of the
trachea of the former containing two or three potential convolutions.
In conclusion, I have to thank Col. Godwin-Austen for aid
rendered to me in the matter of the illustrations.
EXPLANATION OF PLATE XLYVII.
Fig. 1. The complete trachea of an adult female of Rhynchea capensis, nat.
size, showing the modified intrathoracic portion of the tube in an
expanded condition. The compound three-way piece is seen to be
formed by the partial fusion of the last tracheal ring with the modi-
fied first pairs of bronchial half-rings; these have their ventral ends
1878. | TRACHEA OF RHYNCH&A CAPENSIS. 751
oO
greatly expanded and produced backwards, so as to form by their
fusion in the middle line a broad, flat, and squarish plate of bone
strongly bilobed and tipped with cartilage at the hinder extremity,
into which ossification extends with advancing age, rendering the
posterior angles of the plate prominent, and bringing them into very
loose relation of apposition with the much less expanded ventral
ends of the second pair of bronchial half-rings; the ovoid saccular
dilatations seen in figs. 5 and 4 result partly from the inflation of the
membrane interposed between the ends of these half-rings and the
posterior angles of the three-way piece, but principally from that of
the ventral halves of the membranous inner walls of the second and
third pairs of bronchial half-rings. The spatulate dorsal ends of the
first pair of bronchial half-rings do not meet in the middle line, but
curve inwards and backwards so as to leave between them a membra-
nous interval, into which a narrow tongue of bone projects from the
middle of the posterior margin of the last tracheal ring. In so small
a figure no distinction between bone and cartilage in the three-way
piece was possible. Drawn from a fresh specimen by Behari Lal Dés.
Fig. 2. A much enlarged view of a portion of the same, to show the form the
modified part of the tube assumes when it is naturally expanded ;
the constricted portion (a) presents a singularly finely and regularly
ribbed appearance, being composed of about forty very fine and closely
packed cylindrical rings, all firmly bound together so as to form a
stiff but still somewhat elastic mass.
Fig. 3. A much magnified ventral view of the posterior: end of the same, to
show the inflated condition of the membrane connecting the compound
three-way piece with the second (apparent first) bronchial half-ring on
each side, and also the two egg-shaped saccular dilatations (¢, e) of the
membranous inner walls of the bronchi.
Fig. 4. The same, from the left side, to show the egg-shaped dilatations (e, e) in
profile, and the thin and narrow lateral slip of muscle (7) which is
attached to the three-way piece at m, whence some of its fibres pass on
to the second bronchial half-ring (7).
(All the three preceding figures were drawn under the microscope
by the aid of a camera lucida, immediately after the death of the
animal.)
Fig. 5. The complete trachea of an immature female, nat. size. The two
sterno-tracheal muscles (s¢.t, st.¢) are seen to be blended on the
ventral surface of the constricted portion of the tube at a; J, 1, are the
lateral muscles, somewhat exaggerated in the drawing. Drawn by
B. L. D.
Fig. 6. The posterior portion of the unmodified windpipe of an adult male,
nat. size. The lateral muscles (/, /) are here so pale and transparent
as to be all but undistinguishable in the fresh state.
Fig. 7. The same, much enlarged.
(With the two exceptions above mentioned, the figures of this
plate have been obligingly drawn for me by Lieut.-Col. H. H. God-
win-Austen, by whom also the plate has been lithographed.
The following paper was read on June 4th, but was necessarily
omitted from its proper place in consequence of the illustrations not
having beeu finished in time :—
752 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
On the Classification and the Distribution of the Cray-
fishes. By T. H. Huxzey, Sec. R.S., V.P.Z.S.
[Received May 23, 1878. ]
I. Introduction, p. 752.
Il. The Modifications of the Branchis in the Crayfishes, p. 756.
. The branchiz of Astacus, p. 756.
The branchiz of Cambarus, p 763.
. The branchiz of Astacopsis, p. 764.
The branchiz of Cheraps, p. 768.
The branchie of Engeus, p. 769.
. The branchix of Paranephrops, p. 770.
. The branchix af Parastacus, p. 771.
. The branchi of Astacoides, p. 773.
III. The Classification of the Crayfishes, p. 775.
IV. The Distribution of the Crayfishes considered in relation
to their morphological ditferences, p. 786.
OID oP cote
I. INTRODUCTION.
The dismemberment of the genus Astacus of the older naturalists,
down to the time of Fabricius, was commenced by Leach, who sepa-
rated the Norway Lobster as the type of a new genus, Nephrops’.
Milne~Edwards advanced a step further by establishing the genus
Homarus for the Lobsters, and leaving only the freshwater Astacz,
or the proper Crayfishes, in Astacus”.
The later proposal of Leach, to use Astacus for the Lobsters, and
to give a new generic name (Potamobius) to the freshwater Cray-
fishes, would have had the advantage of retaining the primitive
signification of doraxés. But Potamobius had already been used in
another sense; and the change introduced by Milne-Edwards is so
generally adopted that it would be confusing to attempt any further
alteration.
Guérin * next proposed to distinguish the Astacus madagascari-
ensis of Audouin and Milne-Edwards, as Astacoides, from the other
Crayfishes ; and Erichson, in his valuable Monograph of the
group‘, adopts Astacoides for the Madagascar and some of the Aus-
tralian forms, and establishes the new genera Cambarus, Cheraps,
and Engeus. In Cambarus and Cheraps the number of the branchiz
is taken into account as an important generic character.
In 1842° Mr. Adam White described some Crayfish from New
Zealand, for which he constituted a new genus, Paranephrops,
under the impression that the New-Zealand form approximated
to the genus Nephrops. Mz. Wood-Mason® has since “ denied
the existence of any special relationship between the New-Zealand
1 Leach, Trans. Linn. Soc. xi. 344.
2 «Histoire Naturelle des Crustacés,’ 1837.
3 Revue Zoologique, 1839.
4 “ Uebersicht d. Gattung Astacus,” Archiv fiir Naturgeschichte, Bd. 6.
5 Gray’s ‘Zoological Miscellany.’ See also Dieffenbach’s ‘ New Zealand,’ 1843,
yol. ii. p. 267.
§ Proc. Asiat. Soc. Bengal, 1876, p. 4.
1878. ] AND DISTRIBUTION OF THE CRAYFISHES. 753
species of freshwater Astacide and the marine genus Nephrops ;”
and “as the species referred to Paranephrops differed less from [some
of] those of Astacoides than these latter did from one another, and
as, moreover, the latter name had the priority, he proposed provi-
sionally to refer the New-Zealand species of Astacide to it.’
Mr. Wood-Mason is unquestionably right both in denying any
special relationship between Paranephrops and Nephrops, and in
asserting that the New-Zealand Crayfish differ less from some of the
species of the genus Astacoides, as its limits are at present under-
stood, than these do from one another. But I shall have occasion
to show that the type of the genus Astacoides, the Madagascar
Crayfish, differs so widely from the other Crayfishes of the southern
hemisphere, that the latter cannot be included in the same genus ;
while Paranephrops is sufficiently different from the Australian and
Tasmanian Crayfishes to render its recognition as a distinct generic
type desirable’.
The distribution of the Cra fishes, so far as it is hitherto ascer-
tained, is not a little remarkable. Astacus fluviatilis occurs in various
parts of England and in Ireland ; but I cannot find any record of it
in Scotland. Dr. M‘Intosh, who has been kind enough to look into
this point in aid of my inquiries, assures me that Crayfishes are not
indigenous to that part of Britain, that they do not exist in the
Tweed and the Teviot, and that an attempt to introduce them into
the island of Mull failed ; they were placed in various streams,
_ but none were ever seen again. Even in England, Crayfishes appear
to be restrieted to certain rivers, They abound, for example, in the
Thames ; but I cannot hear of any in the Cam or the Ouse, though
their absence in the latter rivers cannot be ascribed to any want
of calcareous matter in the districts through which those rivers
flow.
Astacus fluviatilis, however, extends all over the western half of
Europe, as far south as the Pyrenees and the northern shores of the
Mediterranean ; while, eastward, it reaches Sicily, Northern Greece,
and the western shores of the Black Sea. In Spain there appears
tu be no doubt that it occurs about Barcelona; but whether it is
found in the rest of the Spanish peninsula is uncertain 2.
Northwards and eastwards, Astacus Jluviatilis extends to Sweden
and the Baltic provinces of Russia, and through Western Russia, by
the basins of the Dniester and the Bug, to the Black Sea.
Over this vast area, marked local varieties appear to be not un-
common ; and most authors agree to regard a Crayfish which occurs
in Southern Europe, France, Switzerland, and Germany, and which
? See on this subject the remarks of Mr. Miers, “Note on the genera Asfa-
coides and Paranephrops” (Ann. & Mag. Nat. Hist. ser. 4, vol. xviii, 1876).
* See Gerstfeldt, “ Ueber die F lusskrebse Europa’s” (Mém. presentés al’ Acad.
Twp. des Sciences de St. Petersbourg, 1859, p. 587). As far back as 1675,
Sachs 4 Lewenbeimb wrote :—*Sic in Hispania licet flumina, negant inveniri
ibi Cancros fluviatiles, contra in Gallia frequentissimi, item in Pannonia, in
Helvetia, in Germania; etiam in fluviis Americanis inveniri referunt navigatores,
In fluvio Lydiz Haly cancros invenit Busbequius, Ep. Ture. i. p. 89.” But
were these Lydian “ cancri” crayfishes or fluyiatile crabs ?
794 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
is known in the latter country as the ‘Steinkrebs,’”’ as a distinct
species, Astacus torrentium or A. saxatilis *.
Eastward of the region inhabited by Astacus fluviatilis, from the
Arctic to the Black and Caspian Seas, another species, A. leptodac-
tylus, ranges, associated with the allied but possibly distinct forms
A. pachypus and A. angulosus, in the southern part of the area ;
and it is remarkable that these Crayfishes not only frequent the
rivers which debouch into the Black Sea and the Caspian, but are said
to thrive in the salt waters of those seas.
No Crayfishes are known in the Ob, Jenisei, Lena, or other rivers
which flow into the Arctic Ocean?; but the Amur has one or two
species (A. dauricus). There is a species in Japan (A. japonicus) ;
and Dr. Hagen* enumerates no fewer than six species from British
Columbia, Oregon, and California.
East of the Sierra Nevada, all the Crayfishes at present known
belong to the genus Cambarus, of which Dr. Hagen distinguishes as
many as thirty-two species. They extend from the Great Lakes to
Mexico, Guatemala, Cuba, and probably other of the West-India
Islands. Sloane, in his ‘Natural History of Jamaica’ (vol. i.
p. 271) describes two species in that island. According to the figure,
one of these attains a length of 12 inches.
No Crayfishes are known to occur in the whole continent of
Africa, in Syria, the Euphrates valley, Persia, Hindostan, and India
beyond the Ganges, nor in China as far as the Corea, nor in the
Philippines, nor in any island of the Malay or Papuan archipelagos *.
The late Prof. Agassiz, though he sought for Crayfishes in the
1 Jn 1560, Gesner was acquainted with this distinction :—‘ Astacus fluviatilis
talis apud Helvetios et Germanos est, major silicet et simpliciter dictus Krebs
vel Edelkrebs; eo enim minor est, et colore diversus qui saxatilis cognominatur
Steinkrebs.” (‘Nomenclator Aquatilium,’ p. 374). Heller (Die Crustaceen des
siidlichen Europa, p. 217) refers our English Crayfish to this species ; but no
specimens I have seen agree with his diagnosis. Whether there is any differ-
ence between 4. sawatilis and the Crayfishes which have been named A. pallipes
and A. fontinalis by Lereboullet and Carbonnier; and whether our English
Crayfish is more similar to these than to the form which is commonly known
as A. fluviatilis on the Continent, is more than I am able to say at present. A
critical comparison of large series of specimens from different localities would
probably yield results of great interest to the theory of the origin of species.
2 Kessler, “Die russischen Flusskrebse ” (Bull. de la Soc. Imp. des Nat. de
Moscou, 1874).
3 ‘Monograph of the North-American Astacidx,’ Illustrated Catalogue of
the Museum of Comparative Zoology at Harvard College, 1870.
4 T have been favoured by Sir Henry Barkly with the opportunity of ex-
amining specimens of two kinds of “ Cammarons,” or so-called Crayfishes, from
the rivers of Mauritius. They are large Prawns.
I must confess myself to be in a state of hopeless perplexity respecting the
Crayfish or Lobster which is said to occur at the Cape of Good Hope, Cancer
(Astacus) capensis of Herbst. At the beginning of his description (Naturge-
schichte der Krabben und Krebse,’ Band ii. p. 49) Herbst says :—‘‘ Dieser schone
Krebs halt sich auf den Kap in solchen Flissen auf, die sich auf den Bergen
befinden;” and at the end he states, ‘‘die Fiisse haben insgesammt scheeren-
formige Spitzen, da bey dem gemeinen Flusskrebs nur die ersten zwey Paare
dergleichen haben.” Moreover, the diagnosis runs, ‘ pedibus omnibus cheli-
feris.” It is impossible to suppose that Herbst should have made a mistake on
such a point as this; aud therefore it must be concluded that his Cancer ca-
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 759
Amazons, could find none. Two species from Southern Brazil have
been described by Dr. von Martens’ as Astacus brasiliensis and A.
pilimanus ; but Von Martens recognizes the affinity of these forms
with the Astacoides of Erichson.
Several species of Paranephrops have been described from New
Zealand ; and the Fijian Crayfish belongs to the same genus.
Crayfishes occur all over Australia ; and the species have been re-
ferred to the genera Astacoides and Cheraps. The only Tasmanian
species which have been described constitute the genus Engeus of
Erichson.
Thus it appears, from what is already published on this subject :—
1. That the Crayfishes of the northern hemisphere are generically
distinct from those of the southern hemisphere.
2. That the American Crayfishes, east of the Sierra Nevada, are
generically distinct from those west of that range, as well as from the
South-American species ; and that,while the western North-American
Crayfishes belong to the same genus as those of the Old World, the
South-American forms are more closely allied with those of Mada-
gascar and Australia.
3. That the New-Zealand species are distinct from the Australian
forms ; and that the latter are to be placed in the same genus as the
Madagascar and South-American species.
4. That there is a negative fact of distribution, not to be accounted
for by any apparent difference of climate or other physical conditions—
namely, the entire absence of Crayfishes in Equatorial South America,
Africa, and the rest of the Old World south of the northern escarp-
ment of the great Asiatic highlands.
The problem thus offered is one of the most remarkable among
the many presented by the facts of Geographical Distribution ; and
it appeared to me that one of the first steps towards attempting its
solution was to obtain some more definite conception, than is fur-
nished by extant descriptions, of the actual amount of resemblance
and difference between the Crayfishes which are found in the dif-.
ferent areas of distribution.
For the most part the Crayfishes are so similar in their general
structure, that the characters by which the genera have been distin-
guished are almost trivial. LErichson, however, has drawn attention
pensis is neither a Crayfish nor a Lobster, and that, unless he was wrongly in-
formed, it is an inhabitant of fresh water.
Milne-Edwards (Hist. Nat. des Crustacés, ii. p. 335) identifies his Homarus
capensis with the Cancer capensis of Herbst ; but, as it is stated in the defini-
tion of the genus Homarus a c, p. 333) that the Lobsters “ne se trouvent que
dans la mer,” and as Homarus has only three pairs of chelate limbs, the identi-
fication presents difficulties. Krauss (Siidafrikanische Crustaceen, p. 5+), under
the head of ‘‘ Homarus capensis,” refers to Herbst and Milne-Edwards, and,
apparently on the authority of the former, merely says:—‘ In den Bergflussen
des Kaplandes. Ich habe ihn in Natal nie gesehen.” Elsewhere (p. 20) he
ives “‘ Thelphusa perlata and T. depressa and Homarus capensis” as the only
outh-African freshwater Thoracostraca.
1 “ Siidbrasilische Siiss- und Brackwasser-Crustaccen,” Archiy fur Natur-
geschichte, 1869.
796 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
to the diminution of the number of the branchize in Cambarus and
Astacus, and to an important difference in the structure of those of
Engeus; and Hagen has pointed out some important peculiarities
of these organs in Cambarus; while the remarkable fact, that the
appendages of the first somite of the abdomen are absent in many of
the Crayfishes of the southern hemisphere, has been duly noted by
Erichson and several other zoologists.
Having recently had occasion to make a careful reexamination of
the structure of Astacus fluviatilis, 1 found two minute filaments
attached to the epimera of the penultimate and antepenultimate
thoracic somites. The structure and the position of those filaments
led me to suspect that they must be rudimentary branchize '; and as
the Australian Crayfishes appeared to me to be, on the whole, less
specialized forms than the European species, I thought that I should
probably find in them fully-formed functional branchiz oecupying
the place of these rudiments. Through the kindness of my friend and
former pupil, Mr. J. Wood-Mason, a specimen of “ Astacoides”’
Sranklini was placed at my disposal ; and on examination, I not only
found the functional branchiz I sought, but discovered a number of
other interesting differences between the respiratory organs of this
Crayfish and those of dstacus.
Following up the line of inquiry thus suggested, I have examined
examples of all the chief forms of Crayfishes at present known, with
the result of establishing some remarkable parallel relations between
the morphology and the distribution of these animals.
In order to make these points clear, [ must premise a fuller and
more precise description of the branchial apparatus of the common
Crayfish than has yet been given, in order that it may serve as a
standard of comparison for the branchie of the other Cray fishes.
II. Tae Moprirications oF THE BRANCHIE IN THE CRAYFISHES.
The Branchie of Astacus fluviatilis.
When the branchiostegite of a Crayfish is removed, seven
branchize are seen, running from the base towards the apex of the
branchial cavity, parallel with one another, and disposed in curved
lines, which are concave forwards and convex backwards. The length
of the branchiz gradually increases from the first to the sixth; the
seventh ascends as high as the sixth, but is rather shorter, in conse-
quence of the attachment of its base lying at a higher level.
In each of the six anterior branchiz, a basal portion, a stem, an
expanded damina, and an apical plume may be distinguished. The
basal portion (fig. 1,1, B) is broad, with a convex posterior and
inferior free edge, beset with Jong setze ; and it is articulated by its
1 T have met with no allusion to these structures, unless the following pas-
sage in Brandt's and Ratzeburg’s description of the Crayfish (Med.-Zoologie, ii.
p: 61) refers to them :—‘ Ueber jeder der obersten Kiemen der beiden mittlern
Fusspaare steht etwa 1'” entfernt ein kleiner fadenformiger, unten breiterer,
bartelabnlicher Theil.” Ido not quite see the applicability of “ unten breiterer,”
unless “unten” applies to the attached ends of the filaments; but in other
respects the description fits the rudimentary branchie very well.
1878.] AND DISTRIBUTION OF THE CRAYFISHES. 757
Tig. 1.
WIZ
—
or----~---
Astacus fluviatilis.
I. The outer face of the podobranchia of the antepenultimate thoracic limb of the
left side (x 4): B, the basal portion ; s¢, the stem; /, the lamina; br, the
branchial filaments ; ap, the apical plume. II. The inner face of the same
podobranchia ( x 4): the letters as before, except g, the decurrent fringe,
continued from the inner lobe of the lamina. III. A transverse section
: of the podobranchia, taken a little above the level of 4 in II. (x 4): @, the
+ inner lobe of the lamina ; 3, its outer lobe; s¢, the stem; 27, the branchial
; filaments, IV. One of the hooks of the lamina; and V The extremity
of one of the sets of the base of the podobranchia : the vertical line indi-
cates the scale to which these figures are drawn, and represents 45 of an
¢ inch. VI. The coxopoditic sets of the same limb (x 4). VII. The free
end of one of these set, magnified to the same degree as IV. and V,
Proc. Zoou. Soc.—1878, No. L. 50
798 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
truncated anterior end with the coxopodite of one or other of the
thoracic limbs, from the second maxillipede to the penultimate am-
bulatory limb inclusively. I shall therefore term these gills podo-
branchie. A fold of the integument rises over the outer face of the
basal portion of the branchia, which is freely movable, inwards and
outwards, upon its articulation.
The basal portion of the branchia curves slightly upwards, and is
continued into the comparatively narrow stem (fig. 1, sé), which
bends up at right angles to the base, and lies nearly parallel with
the inner wall of the branchial chamber. Rather beyond the middle
of its length, the stem expands into the broad lamina (fig. 1, 1), the
two lobes of which are folded together like the leaves of a partly
open book, their free rounded edges being turned backwards and
the uniting fold forwards. The lamina of each branchia, from the
second to the sixth, is received between the lobes of the lamina of
that which precedes it, and, from the first to the fifth, receives the
lamina of the branchia which follows it.
The edges of the laminz are beset with minute hooked spines,
seated upon low tubercles (fig. 1, 1v.), and are flat; but, a short
distance within the edge, each lamina is folded longitudinally in such
a manner as to give rise to ten or twelve plaits. From the mode of
their formation, these plaits or folds are, of course, as distinctly
marked on the outer as on the inner faces of the lamine (fig. 1, 111.).
Small elevations, terminated by minute hooked setze, like those on
the edges of the laminze, are observable on both faces, particularly
on the ridges.
The inner lobe of the lamina is continued down the stem asa
decurrent gradually narrowing fringe, which terminates on the inner
face of the basal portion (fig. 1, 1. g). The outer lobe ends more
abruptly, in a thin ridge which lies external and posterior to the last.
Between this ridge and the fringe there is a longitudinal groove,
which occupies the inner and posterior face of the stem.
A short distance from the upper edge of the lamina, and pro-
ceeding, as it were, from the anterior face of the median fold of the
latter, the apical plume (fig. 1, 1. ap) is given off. At its base this
is beset with cylindrical branchial filaments; but at its free extre-
mity it becomes simple and filiform; at its attached extremity, it
passes into the stem of the gill, of which it is seen to be the direct
continuation. The whole of the outer and anterior face of the stem,
as far as the basal fold, is beset with branchial filaments, of which
those in the region of the lamina are the longest, while towards the
base they are shorter and more closely set.
Close to the attachment of the podobranchia, the upper surface of
the coxopodite presents a rounded elevation, from which a great
number of long, flexible and tortuous sete proceed (fig. 1, VI.).
When undisturbed, they ascend vertically between the gills on the
sides of the branchial chamber. The apex of each of these cowopo-
ditic sete is acute; and at a short distance from its extremity it is
beset with numerous short foliaceous scales, which gives it the aspect
of a minute catkin (fig.1,vm.). Further down these scales become
1878. ] AND DISTRIBUTION OF THE CRAYFISHES. 759
more widely separated, and at length disappear, the base of the seta
being smooth. A canal traverses the whole length of the seta.
These bundles of setee were regarded as branchise by Brandt and
Ratzeburg ; but they certainly have no direct relation to the respi-
ratory function. They may aid in keeping parasites out of the
branchial chamber.
The sete which fringe the base of the podobranchia are straight,
stiff, hollow, and composed of a cylindrical basal portion, and an
acuminated acicular terminal part, which often appears articulated
with the foregoing; the junction of the two is marked by an in-
flexion of the wall of the seta. The acicular part (fig. 1, v.) is fringed
with minute pointed scales, which, in the sete: of the anterior part of
the base, pass into long lateral processes, and give the seta a penni-
form appearance. None of these setz are hooked at the apex.
The only differences, except those of size, which are observable in
the six podobranchiz, are, that the external lobe of the lamina,
which is not nearly so large as the internal, in the hindermost
Fig. 2,
Cc
K4 X2 K4
A, The podobranchia (epipodite) of the first maxillipede of Astacus fluviatilis,
viewed from the outer side (xX 4). B. The podobranchia of the first
maxillipede of Astacoides madagascariensis, outer side (X 2): &, branchial
filaments. C. The podobranchia (epipodite) of the first maxillipede of
Astacus fluviatilis, viewed from behind (x4). D. The podobranchia of
the first maxillipede of Parastacus brasiliensis, from behind (X 4); br,
branchial filaments. In each figure a marks the pedicle by which the more
or less modified podobranchia is attached to the coxopodite.
branchia increases in relative size, until, in the most anterior
branchia it becomes larger than the internal lobe. The penniform
Setge are more numerous on the bases of the anterior podobranchiee.
The first maxillipede is said, and, in a physiological sense, rightly,
to possess no branchia ; but it is provided with an appendage (fig.
2, 4, C) which is undoubtedly the homologue of the podobran-
chia of the other thoracic limbs. This is a soft membranous
50
760 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
Fig. 3.
sarod}
iopdb ° pdb
Astacus fluviatilis.
The branchiostegite, all the podobranchix, except the epipodite of the first
maxillipede, and all the arthrobranchiz of the right side are removed (X 4).
en | Za
3. arb. x
spd
K \ )
a
1 r 1 mee Tt \ \
garb; uarbi warl\~,, 7.
pdb updb iopdb opdb
Astacopsis franklinii.
The branchiostegite, all the podobranchiz, except that of the first maxillipede,
and all the anterior arthrobranchis, except the first, of the right side are
removed (X 2).
scg, scaphognathite (not shown in fig. 4); 7 ep, the podobranchia of the first
maxillipede; 8 pdb to 13pdb, the other podobranchix; Sard to 13aréb, the
arthrobranchiz or their attachments to the arthrodial membranes of the second
and following thoracic appendages; 11 plb to 14 plb, the pleurobranchiz ; 14,
the coxopodite of the hindermost ambulatory limb; exp, the coxopoditie sets
of that limb in Astacus,
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 761
plate, which broadens at its upper extremity, and sends a short
process downwards beyond its articujation with the coxopodite of
the maxillipede. The plate is slightly folded upon itself longitu-
dinally, but in such a manner that it is concave forwards instead of
backwards. It bears no branchial papille, and has no longitudinal
plaits ; but, on its posterior face and along its inner edge, it presents
hooked tubercles, like those of the lamine of the podobranchiz.
It is obvious that this structure, which lies immediately behind and
parallel with the scaphognathite of the second maxilla (but, as I have
ascertained, does uot share its function of scooping the water out of
the branchial cavity’), is a modified podobranchia, reduced, as it
were, to the part which, in the other podobranchiz, is represented
by the base, stem, and lamina.
Thus every thoracic limb, except the last, is provided with the
representative of a podobranchia—though, in the case of the first
maxillipede, this structure, if it plays any part in the respiratory
process, does so simply in virtue of its thin and soft texture, and not
by means of any special branchial filaments. The podobranchia of
the first thoracic appendage is, in fact, reduced to a mere epipodite.
When the podobranchiz are removed, six other gills come into
view. They are attached (fig. 3, ard) to the flexible membrane which
unites the coxopodites of all the thoracic limbs to the thorax, save
the first and last, and may be termed anterior arthrobranchie.
Like the foregoing, they are disposed vertically, and increase in
size from the first, which belongs to the second maxillipede and is
hidden behind the epipodite of the first maxillipede, to the last.
The apex of each of these gills is exactly like the apical plume of
one of the podobranchize ; and the branchial filaments are set upon
the outer and anterior face of the stem in the same way. The
inner face is flat and free from filaments ; and there is no trace of a
lamina or of a basal dilatation.
Above and behind these, more directly above in the posterior,
more behind in the anterior limbs (fig. 3, a7), are five other branchiz
of similar character, attached to the arthrodial membranes of the
third maxillipede and the anterior four ambulatory limbs. These
may be termed the posterior arthrobranchie.
After the removal of all these functional branchize, there will be
found, immediately above the bases of the penultimate and antepenul-
timate thoracic limbs (fig. 3, 12 plb, 13 plb), two minute filamentous
processes, the longer of which was not more than one sixth of an inch
in length in any specimen I have examined, while both are so deli-
cate as to be invisible except under a simple lens. The posterior
of these is the larger: it has the structure of an ordinary branchial
filament, with a somewhat swollen base, which is attached to the
margins of a foramen in the lower part of the epimeron of the pe-
nultimate thoracie somite, just below a transverse depression which
separates this from the upper part of the epimeron. The position
1 Tf the branchiostegite of a living Crayfish is carefully removed, the rapid
rhythmical motion of the scaphognathite is readily seen; but the modified
podobranchia of the first maxillipede remains quiescent.
762 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
of this filament is sometimes vertical, but more frequently horizontal.
The anterior filament is sometimes a mere papilla; it is attached to
the margins of a small foramen which occupies a similar position in
the antepenultimate epimeron—namely, close to the anterior edge and
just below the transverse depression. These are two rudimentary
gills, of the same order as that next to be described.
The seventh, and most posterior branchia of those which become
visible when the brachiostegite is removed (fig. 3, 14 plb), has yet to
be considered. It resembles one of the arthrobranchiz in all essen-
tial characters, but it is not attached to the arthrodial membrane ;
on the contrary, the base of its stem is fixed to the margins of a
circular aperture situated close to the edge of a peculiar shield-
shaped plate, the posterior and outer surface of which is covered
with strong setee. Immediately behind and below the attachment
of the gill there is an oval space, occupied by a soft and flexible
portion of the cuticle, like a tympanic membrane. By its lower
margin this plate furnishes an articular surface to the outer condyle
of the coxopodite of the last thoracic limb, while its anterior and upper
angle, bending sharply upwards, passes into a curved prolongation,
which extends upwards and backwards in the soft integument of the
flank, and articulates with a slender process of somewhat similar
form sent forward from the first abdominal somite. Internally this
shield-shaped branchiferous plate is continuous with the sternum
of the last thoracie somite. It is obvious that this plate, with its
anterior process, represents the epimeron of the last thoracic somite,
which is thus adherent to the penultimate somite only by the slender
anterior and superior process and the soft integument. Hence, the last
thoracic somite moves easily upon its predecessor, though, in strict-
ness, the usual statement that the last thoracic somite in Astacus
is ‘‘ free’’ is not altogether exact.
It follows from this determination of the nature of the shield-
shaped plate, that the gill which it bears is attached to the epimeron,
or side-wall, of the last thoracic somite; and it may be termed a
pleurobranchia. The similarly attached filaments (12 plé and 13 plb)
represent reduced or rudimentary pleurobranchiz.
We may suppose that the total number of branchize which a
thoracic somite can possess is eight, four on each side, namely :—one
podobranchia, connected with the coxopodite of the appendage; two
arthrobranchie, fixed to the articular membrane; and one pleuro-
branchia, attached to the epimeron. And if four places for branchize
are assigned to each somite, the extent to which the hypothetically
complete scheme or formula is actually filled up will be readily seen,
and the branchial arrangements of different Crayfishes will be easily
compared.
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 763
The Branchial formula of Astacus fluviatilis’.
Somites Arthrobranchize.
and their ra oa ——_~ cepa
appendages. “* Anterior. Posterior. anchiz.
WITS: oO (ep) 0 0 0 0g (ep)
VIET oL, 1 ] 0 0 = 5
EXG2A 1 l | 0 a
X... I | 1 0 = 3
Ry,. 53% 1 l 1 OY = 3
MB t6:. DIAN SED) snails
XIII..... 1 1 l , = 34
XIV. 0 0 0 eh 3
6+tep+6 + 5 + 1+2r=18+ep+2r
“ep” here signifies a podobranchia which has lost its branchial
filaments and become completely metamorphosed into an epipodite,
while r indicates that a rudiment of a branchia exists.
It will be observed that, in this species of Crayfish, no somite
possesses its hypothetically full complement of branchiz except XII.
and XIII. ; and even in them the pleurobranchiz are rudimentary.
The representatives of eleven possible branchie are altogether
wanting.
2. The Branchie of Cambarus.
The principal distinction between this genus and Astacus, as it
was established by Erichson, lies in the absence of the single pleuro-
branchia of the latter, and the consequent reduction of the number
of the branchize to seventeen on each side.
In his elaborate monograph of the genus, Dr.-Hagen observes,
«But there is also another difference, not before noticed?. In
Astacus each pair of gills, except the single one on the fifth set of
legs, has a broad deeply-folded membrane, closely fixed behind the
most external gill-lobe. In Caméarus, this membrane is always
wanting in the gills of the fourth pair of legs, but exists, as in
Astacus, in all the others. ;
“In the true Astacus, all the gills with a folded membrane behin
have a basal external bundle of shorter but broader and irregularly
placed gill-tubes ; these are never to be found in Cambarus.”
In a species of Cambarus from Guatemala, of which a number of
specimens have been presented to the British Museum by Mr.
Salvin’, I find Dr. Hagen’s first remark fully borne out. The last
1 In this, as in all other cases, it is to be understood that the branchial
formula gives the branchiz of only one side of the body, and that the total
number of branchiz is therefore double that given in the formula.
2 Dr. Hagen appears to have overlooked De Haan’s definition of the distine-
tive characters of the American Orayfishes known to him :—
“ Branchiz 17 ; nulla supra pedes quintos; externe supra quartos tantum e
tubulis liberis, externse supra sequentes infra e tubulis, supra e laminis tuber-
culatis composite” (Fauna Japonica, Crustacea, p. 164).
* Mr. Salvin informs me that they were obtained in a river near Coban,
in Vera Paz, at an elevation of about 4300 feet above the sea.
764 PROF. T. H. HUXLEY ON THE CLASSIFICATION — [June 4,
podobranchia is devoid of even a trace of a lamina ; in the five which
precede it, on the other hand, the lamina is very large, and folded
into two longitudinally plaited lobes, as in Astacus. The edges and
the surfaces of the laminz present tubercles, which are more pro-
minent than in Astacus, and bear similar hooked spines.
The inner decurrent prolongation of the lamina is wider than in
Astacus, particularly in the anterior podobranchiz. It is also beset
with hooked setee mounted on low tubercles.
The sete of the upper part of the base are relatively shorter.
As in Astacus, their apices are straight and not hooked.
The coxopodites bear bundles of twisted setae, which are similar
to those of Astacus, but are more obtusely pointed.
The arthrobranchiz are similar in number and in form to those of
Astacus. Those of the posterior series are proportionately larger.
I cannot discover a trace of the hindermost pleurobranchia, nor of
the rudiments of the anterior ones, in this species; but it is not im-
probable that they may be discovered in larger forms.
The modified podobranchia (epipodite) of the first maxillipede is
soft, and folded longitudinally in such a manner as to present a broad
and shallow anterior groove. It bears no branchial papillee, nor any
setee ; but there are a few short hooks here and there.
The Branchial formula of Cambarus.
Somites Arthrobranchie.
Sn Saeantiieaie AGE eae. and Miche
appendages. ~“" Anterior, Posterior i
WEES 0 (ep) 0 0 Ch = Oe
VII... ] 1 0 0 = 2
| ib. ae ] l 1 0) a
met 1 1 1 0 S45}
pF 1 ] ] 0 = 3
XII. 1 ] l Ce =o
XIIl. l I ] 0 —
XIV. 0 0 0 0 = 0
6+ep+6 + .5 + O = 17 + ep.
In comparison with Astacus, there is obviously a reduction of the
branchial system, arising from the complete disappearance of all the
pleurobranchize in Caméarus.
3. The Branchie of Astacopsis! franklinii.
When the branchiostegite is removed, seven branchiz are seen,
having the general appearance and disposition of those of Astacus ;
and, as in Astacus, the six anterior are podobranchiz, attached to the
coxopodite of the second maxillipede and the five following append-
ages, while the seventh is a pleurobranchia, fixed to the epimeron of
? Since, as will be shown below, those Australian Crayfishes which neither
belong to Cheraps nor to Engeus are distinct from Astacoides, as represented
by the Madagascar speeies, I propose to apply the generie name of Astacopsis
to them.
or
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 76
Fig. 5.
Astacopsis franklinit.
I, The outer face of the podobranchia of the antepenultimate thoracic limb of
the left side (x34). II. The inner face of the same podobranchia: 37,
branchial filaments; /, rudimentary lamina. IIa. The apex of the same,
more highly magnified. III. A transverse section of the podobranchia at
about the junction of its middle and upper thirds. IV. The extremity of
one of the hooked filaments; and V, that of one of the hooked sete. In
these two figures the vertical line represents =}, of an inch magnified to
the same extent. VI. The coxopoditic sete (x33).
766 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
the last thoracic somite. There is also an anterior series of six
arthrobranchie, and a posterior series of five (p. 760, fig. 4, ard),
disposed as in Astacus. But instead of the two rudimentary pleuro-
branchiz borne by the twelfth and thirteenth somites, in Astacus,
there are three perfect branchiz attached to the eleventh, twelfth,
and thirteenth somites (fig. 4, 1-13 p/b), in positions which, in the
case of the two latter, exactly correspond with those of the rudi-
mentary pleurobranchiz of Astacus.
The pleurobranchiz and the arthrobranchiz are very similar to
those of Astacus ; but the inner surface of the stem is channelled by
a sort of groove, the edges of which are pretty sharp. I could find
no hooks, nor spines, nor hooked papillz on any part of these
branchiz.
The podobranchie (fig. 5, 1, 11, 11) resemble those of Astacus
in their general form; but the basal portion is relatively larger and
the stem stouter, while, at first sight, the lamina appears to be
altogether wanting. Close examination, however, shows that the
apex of the branchia is distinguishable into an apical plume and a
lamellar appendage, but that the latter is relatively very small; in
fact, it is a mere outgrowth of the inner wall of the stem, and is
neither bilobed nor plaited. Moreover its surface is beset with
numerous filamentous prolongations, which are altogether similar to
ordinary branchial filaments, except that the extremity of each,
instead of being smooth and rounded, is provided with a short
recurved hook-like seta (fig. 5, 11a, 1v). The groove on the inner
or posterior face of the stem is broader than in Astacus. It is
bounded on the inner side by a decurrent fringe, which runs down
onto the base and stops there, as in the foregoing genera (fig. 5, 11).
This fringe is provided with long slender curved sete, each beset
with imbricated scales and terminated by a recurved hook (fig. 5, v).
The external ridge is wider than in dAstacus, and is provided with
similar hooked sete. It may be said, in fact, that the whole lower
half of the posterior margin of the stem in Astacopsis has the character
of the posterior margin of the base, while in Astacus there is a
sharp demarcation between the base and the stem.
In principle, therefore, the podobranchiz of Astacopsis are similar
to those of Astacus ; and the main difference between the two is that
the lamina in the former is represented by a slight expansion of the
stem, which is neither bilobed nor plaited, while its surface is covered
with cylindrical filaments terminated by hooked spines. In Astacus,
as in Cambarus, on the other hand, the lamina is large, bilobed,
plaited, and the place of the filaments is taken by mere papille
terminated by similar hooks. Moreover, in Astacus and Cambarus
the sete of the base are not hooked.
The appendage of the first maxillipede is similar to that of
Astacus in form; but on the outer surface of the outer lobe there
are sixteen or eighteen short branchial papille, some of which, but
apparently not all, are terminated by hooks; in fact, except in size,
they quite resemble the filaments of the other branchiz.
In this Crayfish, therefore, the first podobranchia is not reduced
1878. ] AND DISTRIBUTION OF THE CRAYFISHES, 767
Fig. 6.
Cheraps (?).
I. The outer face of the podobranchia of the antepenultimate thoracic limb of
the left side(x3). II. The inner face of the same podobranchia : B, basal
portion ; sz, stem ; al, ala; br, branchial filaments. IIT. A transverse sec-
tion of the middle of the podobranchia (x3). IV. A sickle-shaped hook of
a branchial filament xiz Of an inch in Iehgth. V. One of the coxopoditie
setze more highly magnified : a, the circumferential inflexion of the wall of
the seta. The central canal does not stop at this point, but is continued
to the end of the seta. VI. A bundle of coxopoditic sete (x 3). WIT,
The extremity of one of the long sete of the posterior edge of the stem.
The vertical line represents x30 Of an inch magnified to the same extent.
768 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
to a mere epipodite, but retains true branchial characters in the
scanty respiratory filaments of its outer lobe.
There are only small tufts of short, straight or slightly curved sete
in the position of the bundles of long coiled coxopoditie setz of
Astacus and Cambarus (fig. 5, v1).
The Branchial formula of Astacopsis.
Somites Pade: Arthrobranchiz. Pinus
and their ~ branchize ag at aS branchiz.
appendages. * Anterior. Posterior.
‘ O(epr) 0 0 0 .= OCepr)
VE 1 l 0 0 = 2
|. 1 ] ] 0 =e
ae ] ] 1 0 = 3
bc eee 1 ] 1 1 = 4
ATG 2.29 1 1 1 1 = 4
D2 Ne 1 1 1 1 = 4
RAV. i *. 0 0 0 1 = |
6+epr+6 + 5 Be = 21-Fep7
Thus Astacopsis presents a much nearer approximation to the
hypothetically complete branchial formula than Astacus, inasmuch
as the epipodite of the first maxillipede is an imperfect branchia,
and there are four complete pleurobranchiz: only the hindermost
podobranchia, the first and last anterior arthrobranchiee, the first
two and the last posterior arthrobranchiz, and the first four pleuro-
branchiz are wanting. In fact, this is the most complete branchial
formula with which I am at present acquainted, among the podo-
phthalmous Crustacea.
4, The Branchie of Cheraps (?).
_I have examined a single specimen of a species attributed to this
genus, from the Yarra- Yarra river, in the collection of the British
Museum’.
The second maxillipede and the five following thoracic limbs bear
podobranchize, which increase in length from the first to the last,
and have a close general resemblance to those of Astacopsis; but
they differ in the production of the upper part of the anterior lip of
the groove of the stem into a broad limb or ala, wider at the upper
end than below, which must not be confounded with the lamina of
Astacis, as it corresponds only with part of the inner lobe and the
decurrent fringe in that genus (fig. 6,1, 1, 111 @/). Both faces,
as well as the free posterior margin of this fringe, are beset with
cylindrical branchial filaments, the apex of each of which is termi-
nated by a strong sickle-shaped hook (fig. 6, rv). The majority of
the branchial filaments of the stem are also terminated either by
smaller hooks or by short straight spines. Long sete, hooked at
1 My best thanks are due to Dr. Gimther, F.R.S., for the readiness with
which he has aided my investigations by giving me free access to the Crusta-
cean collection under his charge.
1878. ] AND DISTRIBUTION OF THE CRAYFISHES. 769
the ends, and otherwise similar in structure to those of Astacopsis,
are attached to the posterior edge of the stem of the podobranchia
(fig. 6, vit).
The ala is widest, and extends furthest towards the summit of the
branchia, in the most anterior podobranchia, while in the hinder-
most it is reduced to little more than a longitudinal ridge. This
branchia, in fact, is very similar to one of those of Astacopsis.
There are six anterior arthrobranchiz, which are not more than
half as long as the podobranchiz, and five still smaller posterior ar-
throbranchiz, the hindermost of which is almost rudimentary.
As in Astacopsis, there are four large pleurobranchiz, the hinder-
most of which is only slightly larger than the other three, which are
nearly equal in size.
Small branchial filaments are scattered over the whole posterior
face of the epipodite of the first maxillipede; and the apex of each
is provided with a strong sickle-shaped hook.
The coxopoditic setz are relatively short, as in Astacopsis, but
slender and curved, and even slightly undulating (fig. 6, v1). Each
is hooked at its free extremity (fig. 6, v).
If the distinctive character of Cheraps, as given by Erichson, is
correct, this is not a true Cheraps; but the branchiz of these
animals are so readily detached, that I can give no opinion on this
point without comparison with the original specimens. In any case,
the structure of the podobranchiz shows the distinctness of this form
from Astacopsis.
The Branchial formula of Cheraps (?).
A Bee Podo- eee sheen Pletro-
appendages. branche. "Anterior, Posterior. branchive.
0 (ep r) 0 0 = 0 (ep r)
VERA... . 1 iu 0 0 eantih,
{DCR an 1 uf il 0 heck
ONE © a. ] ] 1 0 a
5.10 a ee ] l 1 1 |
XIII.,... ] 1 1 1 ded)
XIV.. ae 0 0 0 ] — ]
Gtepr+6 + 5 + 4 =2 tepr.
5. The Branchie of Engeus.
The number, the general disposition, and the structure of the
branchize are the same as in Astacopsis and Cheraps.
The podobranchiz have no vestige of a lamina. The first podo-
branchia has a small ala, much as in Cheraps; but only a few of the
branchial filaments have terminal hooks.
The arthrobranchie are very small, those of the posterior series
being the smaller; and the three anterior pleurobranchix are much
smaller than the hindermost.
Inall the branchiz, and especially in the arthrobranchiz and pleuro-
-branchiz, the terminal filament is exceptionally long and thick.
770 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
There are only a few very small hooked papillary elevations on
the epipodite of the first maxillipede; in fact the podobranchia is
reduced to nearly the same condition as in Astacus. The hooks of
the setz are very slender.
_The branchial formula is the same as in Astacopsis and Cheraps,
Viz. :—
sr ae Arthrobranchie. ahh tn:
and their j pe a c
appendages. branehise. Anterior. Posterior. branchiz.
VIl..... 0 (ep r) 0 0 0 = 0 (ep r)
Mie | 0 gag
X.. eae 1 l ] 0 aa
DO age 1 1 ] 1 —) 4
C0 ie lin 1 1 jsut og
>: I aroma 0 0 1 5 4)
6+ epr +6 es Cae! Seeiany. Gaia Ye
6. The Branchie of Paranephrops.
In Paranephrops planifrons I find the branchie to have the same
general character and disposition as in Astacopsis and Engaeus.
The podobranchize are devoid of any trace of a lamina. The
branchial filaments on the posterior faces of these branchiz are, for the
most part, provided with terminal hooks, while the rest have smooth
and rounded apices.
The anterior arthrobranchie have not half the size of the podo-
branchie, while the posterior arthrobranchiz are very minute, and the
hindermost is rudimentary, being a mere simple filament, like one of
the rudimentary pleurobranchie of As/acus. The pleurobranchiz
are larger, but the three anterior ones are small. None of these
branchiz have hooked papille.
The external and posterior face of the epipodite of the first maxilli-
pede is beset with short hooked branchial filaments.
The coxopoditic sete are few, slender, short, and hooked at the
extremity.
The Branchial formula of Paranephrops.
Somites ea Arthrobranchiz. Plauen:
and their jranchie PED DE a: branchiz
appendages. “Anterior. Posterior. rs
Rib cosy OO hep) yO 0 0 = O(epr)
VF e027 1 1 0 0 = 2
IXSe as ol l sag 0 = 43
XO. # 1 1 1 0 jo
pC bares 1 1 1 1 = 4
SLD eee 1 1 ] = 4
XI at 1 r 1 = 34r
XIV.. 0 0 0 1 = |
6+epr+6 + 4 +7r 44 = 20+epr+r
1378. ] AND DISTRIBUTION OF THE CRAYFISHES. 771
The branchiz of two specimens of a Paranephrops, from the Fiji
Islands, like the foregoing in the British Museum, are in such bad
condition, the specimens having apparently got dry before they were
placed in spirits, that I cannot make out all the details of their
structure ; but, so far as it can be ascertained, they agree with those
of the preceding species.
7. The Branchie of Parastacus.
By the kindness of Prof. Peters, I have been able to examine two
well preserved males of the Astacus brasiliensis and A. pilimanus of
Von Martens; and the results are very interesting.
The branchiz of the two species are so much alike that they may
be described together. In many points they resemble those of
~ Cheraps ; but the structure is by no means identical ; and as these
Crayfishes are peculiar in other respects, I think it will be most
convenient to consider them as members of a distinct genus, Par-
astacus.
There are six ordinary podobranchiz, of which, as usual, the first
is the smallest, the next two are longer, and the last three are
longest of all and nearly equal. In the podobranchia of the second
maxillipede, the inner lip of the groove of the stem of the branchia
is produced into a broad ala, as in Cheraps ; but the ala becomes
broader towards the apex, and is there abruptly truncated. The
truncated edge is fringed by a single series of branchial papille. The
posterior lip of the groove is beset with long hooked setz. In the
four following podobranchiz the ala is a little narrower, especially
at its apex, but it has essentially the same characters. In the last
the ala is present in the basal half of the stem, but narrows to a mere
ridge in the apical half.
The modified podobranchia of the first maxillipede bears from
ten to sixteen longer or shorter branchial filaments on the outer half
of the posterior surface of its apical end (fig. 2, p, p. 759).
The six anterior arthrobranchize are full-sized, and increase in
length from before backwards. The five posterior arthrobranchiz
are much smaller ; and the last is rudimentary, consisting of a very
short slender stem, with from one to three lateral filaments.
There are four pleurobranchiz, all well developed, but the hinder-
most the longest.
In the podobranchiz, many of the posterior branchial filaments
are terminated by hooks. As usual, these are absent in the other
branchiz.
The coxopoditic tubercles give origin to bundles of long and
tortuous sete, with hooked apices. These are neither so long, nor
so numerous, as in Astacus and Cambarus, but are more like them
than are those of any other Parastacide.
772 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
Fig. 7.
Astacoides madagascariensis.
I. The outer face of the podobranchia of the antepenultimate thoracic limb of
the left side (x23). II. The inner face of the same: 07, branchial fila-
ments. III. A transverse section of the middle of the podobranchia: s¢,
the grooved inner face of the stem, IV. The terminal hook of a branchial
filament; and V. The apex of a coxopoditic seta. The straight line repre-
sents ;}4? of an inch magnified to the same extent as these,
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 773
The Branchial formula of Pacastacus.
Somites Bites Arthrobranchiz. Pia.
et nchie. 7. ce, branchiz.
appendages. Anterior. Posterior.
Milos 0eps)e. O 0 0 = O(epr)
ft} Serene | 1 0 Osnneeiin 2
| > ae ] 1 | Os = «3
Rett cberfiy 1 ] I 0 =
oe | 1 l 1 =e Pe!
Meliss airs, (2 ] 1 l ee
pO |) ee | 1 r 1 = 3+4+r
UE darcia! 90 0 0 1 —
6 +ep7+ 6 + 44744 20 +epr-+r.
8. The Branchie of Astacoides.
I am indebted to the courtesy of Prof. Alphonse Milne-Edwards
for the opportunity of examining the branchiz of a male specimen
of the Crayfish of Madagascar, Astacoides madagascariensis. On
account of the rarity of this species, it is desirable to describe its
branchial apparatus in some detail. The length of the specimen
was 5°7 inches.
The branchiostegite of the left side being carefully removed, the
six large podobranchiz were seen. ‘The first, 0°9 inch long (mea-
sured from its attachment to its apex), was directed upwards and
backwards in the cervical depression. The second, | inch long,
took a similar direction, but was concave forwards. The third,
slightly shorter, lay parallel with the second. The fourth (fig. 7, 1,
11) was much longer ; bent round the third, its summit touched the
apex of the second ; the total length of the branchia when straight-
ened out was about 1°3 inch. The fifth branchia was still longer,
curving round the posterior edge of the last, so that its apex touched
the front boundary of the branchial cavity. The sixth branchia
started from a higher level than any of the others, in consequence of
the great size of the coxopodite of the penultimate thoracic limb ; its
length was 1:05 inch; and it was nearly straight, its apex fitting
into the summit of the branchial cavity. The base of each podo-
branchia is elongated upwards, as in Astacopsis ; and there is a soft
fold of integument over its attachment. There is no trace of any
lamina.
On the removal of the podobranchie the arthrobranchiz came
into view. But, in contradistinction to all other known Crayfishes,
there are only five of them fully developed, and even these are re-
markably small in comparison with the podobranchie. The first
is the shortest (0°5 in.); and they increase in length to the hinder-
most, which is 0°35 in. long. Each is concave forwards and con-
vex backwards; and the apex of the hindermost comes within 0°15
inch of the’ nearest part of the anterior superior boundary of the
branchial cavity. In all these branchie the branchial filaments
are very numerous, stiff, relatively short and close-set ; so that they
Proc. Zoou. Soc.—1878, No. LI. 51
774 PROF. T. H. HUXLEY ON THE CLASSIFICATION —__[ June 4,
approach the characters of the branchie of the Lobster. The groove
on the inner face of the stem seems, at first, to be reduced to nothing
in the upper part of the podobranchie. However, a transverse
section (fig. 7, 111) shows that it exists and is bounded by two lips,
the anterior of which is produced into a narrow ala. But the
numerous hooked branchial filaments, which arise from the posterior
face of the stem, fill the groove and appear to obliterate it.
The hindermost of the arthrobranchie is attached close to the
articulation of the coxopodite with the epimeron, and is therefore
in the same position as one of the anterior series of arthrobranchie
in other Crayfishes. here is not the slightest trace of the attach-
ment of another branchia on the arthrodial membrane behind this.
But on the epimeron, just above the margin of the articular cavity,
and 0:1 inch behind the root of the anterior branchia, a minute
filament, 0-05 in. long, which I take to be a rudiment of the posterior
branchia, enclosed by the downward extension of the epimeron,
projects.
The attachment of the next arthrobranchia is like that of its pre-
decessor, but not quite so far forward. At a distance of 0°5 in.
behind and above its root, an exceedingly minute papilla, not more
than a fifth of the length of the foregoing, is enclosed in the arti-
cular edge of the epimeron. This is doubtless the rudiment of the
posterior arthrobranchia. The three remaining arthrobranchiz are
fixed into the arthrodial membranes of the other ambulatory and
prehensile limbs, in a position corresponding with that occupied by
the fourth. In the case of the hindermost of these, or that of the
eleventh pair of appendages, there is, above and behind the root of
the gill, and 0°U8 inch distant from it, a filamentous rudimentary
branchia, rather less than 0:1 inch long, attached within the articular
margin of the epimeron. Three or four minute unequal processes
are given off from the posterior edge of this filament. There is
another very small papillary rudiment immediately above and behind
the root of the next arthrobranchia; but none could be discovered
above that of the most anterior arthrobranchia.
In addition to the five conspicuous and functional outer arthro-
branchize, there was another very small one, only 0°18 inch long, and
almost rudimentary, attached to the second maxillipede.
The pleurobranchia of the last thoracic somite was only 0°3 inch
long, and was hardly visible at first, from being overlapped by the
hindermost podobranchia. I could not discover the slightest trace
of any other pleurobranchize.
The upper part of the modified podobranchia of the first maxillipede
is sharply folded upon itself longitudinally ; and its outer lobe is beset
with numerous short branchial filaments, most of which have ter-
minal hooks. Similar hooked filaments are to be found about the
base and the posterior region of the other podobranchiz ; but a large
proportion of the filaments of these branchie, and all of those of the
arthrobranchiz, appear to be hookless.
The tubercles of the coxopodites give rise to small tufts of short
and straight setee, which are hooked at the end, like those of Asta-
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 775
copsis (fig. 7, v). The hinder and upper sete of the bases of the
podobranchiz are similarly hooked; but, as in the other species,
the anterior sete are straight, or only slightly curved at the ex-
tremities.
The Branchial formula of Astacoides madagascariensis,
Somites Arthrobranchiz.
ori iegy Lo ag caeegeean a ares
appendages. ’ Anterior. Posterior. "
VI O(epr) 0 0 0 = (epr)
VIII... 1 r 0 0 a= | + r
EX... l 1 0 0 Sey;
ao lets 1 r 0. = 2+ 7
XI... ] 1 r 0 = 2+ ?
XII 1 1 r 0 = Best ?
XIII l 1 r 0 = 2+ r
XIV. 0 0 0 ] = i
6+epr4+i5+r+4r 4+ 1 12+ep r+ ir
In Astacoides, therefore, the branchize have suffered more reduc-
tion than in any other known Crayfish ; and this reduction is, as it
were, a continuation of the process already commenced in Engeus
and Paranephrops, in which the anterior pleurobranchiee and the
posterior arthrobranchize are small, or even rudimentary.
I
IIT. Tue Crasstrication or THE CRAYFISHES.
Whatever may be the variation in the structure of the branchize of
the different species of Crayfish, it will be observed that they all agree
in possessing podobranchiew, or branchie attached to the coxo-
podites, of the six middle thoracic appendages, and that these are
either not at all, or incompletely, differentiated into a branchial and
an epipoditic division. Moreover Astacopsis, Cheraps, Engeus,
Paranephrops, Parastacus and Astacoides, in which the apices of
the podobranchie are not separated into a branchial plume and a
well developed lamina, present a less-differentiated type of branchial
structure than that which obtains in Aséacus and Cambarus.
Thus the structure of the branchie in the Crayfishes separates
them into two groups, of which I propose to term the latter the
PoramMosirp&, and the former the PARASTACID2.
In the Parasracip the podobranchie are devoid of more than
a rudiment of a lamina, though the stem may be alate. The podo-
branchia of the first maxillipede has the form of an epipodite; but,
in almost all cases, it bears a certain number of well-developed
branchial filaments.
The first abdominal somite possesses no appendage in either sex;
and the appendages of the four following somites are large.
The telson is never completely divided by a transverse suture.
More or fewer of the branchial filaments of the podobranchiz are
terminated by short hooked spines ; and the coxopoditic ae as well
5
776 PROF. T. H. HUXLEY ON THE CLASSIFICATION = [June 4,
as those which beset the stems of the podobranchie, have hooked
apices.
In the Poramontip the podobranchie of the second, third,
fourth, fifth, and sixth thoracic appendages are always provided with
a large plaited lamina. The pcdobranchia of the first maxillipede is
converted into an epipodite, entirely devoid of branchial filaments.
The first abdominal somite invariably bears appendages in the male,
and usually in both sexes. In the male these appendages are styli-
form, and those of the second somite are always peculiarly modified.
The appendages of the four following somites are relatively small.
The telson is usually completely divided by a transverse suture.
None of the branchial filaments are terminated by hooks; nor are
any of the coxopoditic sete or the longer sete of the podobranchie
hooked, though hooked tubercles occur on the stem and on the lamina
of the latter.
The coxopoditie setee are always long and tortuous.
It is worthy of notice that the Parastacidee agree with the
Palinuridze and Scyllaride in the abortion of the appendages
of the first abdominal somite, and in possessing hooked sete, while
in the Potamobiidee, as in the Lobsters, the sete are not hooked,
and, as in almost all the Podophthalmia, except the Palinuride and
Scyllaridsc’, the appendages of the first abdominal somite are present,
and are specially modified in the males.
Of the six genera of the Parastacide, Astucoides is widely sepa-
rated from all the rest by the reduction of the number of its func-
tional branchiz to twelve, while all the other genera at present known
have, at fewest, twenty, and usually twenty-one, branchie.
Astacopsis, Cheraps, Engeus, and Parastacus have, in common,
a long epistoma, the surface of which is flattened, the basal joint
of the antenne fixed by the overlapping edge of the cephaloste-
gite, the posterior thoracic sterna narrow, the coxopodites of the
hinder thoracic limbs large and approximated in the middle line.
The rostrum and the antennary squame are short. Engeus is dis-
tinguished among these, not only by the narrowness of the first ab-
dominal somite, but by the form of the anterior part of the head,
with its short and deflexed rostrum, and very small antennary squame.
In Cheraps the podobranchiz are alate, in Astacopsis they are
not. Parastacus somewhat resembles Cheraps in its alate podo-
branchiz, but differs from all the rest by possessing long and tortu-
ous coxopoditic sete.
I have nothing to add to the distinctive characters of the two genera
of the Potamobiide, Astacus and Cambarus, already given by
Erichson, Hagen, and others.
All the branchize of the Crayfishes consist of a stem beset with nu-
merous cylindrical filaments. In fact, they are typical examples of
what are termed by Milne-Edwards “ branchies en brosse,” and may
be called “ trichobranchiz,’’ in contradistinction to the lamellar
1 In Gebia, Calliavis, and Porcellana, the first abdominal appendages are
yudimentary or abortive in the male sex.
1878.] AND DISTRIBUTION OF THE CRAYFISHES. 777
gills or “ phyllobranchiz,’” which are met with in a large number of
other Crustacea. The whole of the Macrurous Podophthalmia,
excepting the genera Gebia and Callianassa, the Prawns, the
Shrimps, and the Myside, have trichobranchie.
In the Mysidz the branchize are rudimentary or absent, and in
the Euphausidee and Penzeidze they are peculiarly modified. In the
Prawns and Shrimps, in Geéia and Callianassa, in all the Auomura
and Brachyura, the gills are phyllobranchiz.
Thus the Podophthalmia or Thoracostraca (to use the convenient
name proposed by Prof. Claus) are divisible in respect of the structure
of their respiratory organs into three groups, which may be termed
Abranchiata, Trichobranchiata, aud Phyllobranchiata.
Among the trichobranchiate Podophthalmia, the Euphauside
possess no other than podobranchiz ‘. These are mere respiratory
plumes presenting no differentiation even into base and stem. Ail
the rest of the Trichobranchiata have arthrobranchiz, either with or
without functional podobranchiz and pleurobranchie. Among
those which possess all three kinds of branehiz, the Parastacide and
the Palinuride are highly exceptional among the Thoracostraca in the
absence of the appendages of the first abdominal somite in both sexes.
They further, as a rule, possess 21 branchize ( pdd. 6, arb. 11, pld. 4),
though the number is, in some cases, reduced by the suppression of
more or fewer of the arthrobranchiz and pleurobranchie.
In most, if not all the other Trichobranchiata, the first abdominal
appendages of the males are present and specially modified. Among
these, the Potamobiidze are characterized by the imperfect division
of their podobranchiz into a proper branchial and an epipoditic
portion.
In Homarus and Nephrops, Awius and Thalassina, the podo-
branchiz are completely differentiated, from their bases onward, into
a proper branchial and an epipoditic portion. Jn this condition the
podobranchia is usually described as an epipodite, to the base of
which a branchia is attached.
In Homarus the branchial filaments are numerous and multiser nt
and the branchial formula is :—
Somites Peay: Arthrobranchiz. ee
and their branchize eis ee . ~~ branchize
appendages. * Anterior. Posterior, a:
\'0 0 ee (Gi (ep) 0 0 On ee 0s(en)
VIII..... 1 0 0 9 = 1
ae 1 l ] 0 = $8
>. ae ] ] ] 0 = 3
> Cs ie ] ] l 1 = 4
0) ESP ee ] l l 1 = 4
XIII..... 1 1 1 | a — are:
MOEV ee 0 0 0 a |
6-Hey si ey on. +i, 4-, = 20 hep.
1 Possibly some of the branchial plumes in Sergestes may be attached to the
arthrodial membranes. A critical examination of the species of Sergestes in
reference to this point would probably yield interesting results,
778 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
In Nephrops the branchie have the same structure, but the
branchial plume of the podobranchia of the second maxillipede is
absent or rudimentary; hence the number of the branchiz is re-
duced to 19.
In Azius the podobranchia of the penultimate thoracic limb is
reduced to an epipodite by the disappearance of the branchial plume,
and the hindermost pleurobranchia is also wanting. The arthro-
branchia of the second maxillipede is represented by a mere ru-
diment.
The Branchial formula of Axius stirhynchus.
da the Poilo- Weg seats Pleuro-
appendages, branchie. Antarior. Posterior, renchi=.
Villiiox l = 0 Oe
16.6 . ] ] ] 0 =,3
Xda i ] 1 jy RsSes
Xl 2 ] 1 1 ene,
XII... 1 l l ec
XI... 0 (ep) 1 1 rapes die
XIV. 0) 0 0 Oni =a0
5+2 ep ot, =) 15 aie a. 1842 ep +r.
The branchial filaments in this genus, again, are arranged only in
two series on the stem, though, as they remain filiform, the ap-
proximation to the phyllobranchiate type is but slight.
In Homarus, Nephrops, and Avius the epipoditie divisions of the
podobranchiz are very large; but in Thalassina, Stenopus, and
Peneus they are much reduced in size, and the branchial element
disappears in more than the hindermost.
In Thalassina scorpioides the last thoracie somite is provided with
no branchizw of any kind, nor have its limbs any epipodite, and
there are no pleurobranchie. The podobranchize of the twelfth and
thirteenth somites are reduced to stout curved setose epipodites. In
the eleventh there is a similar epipodite, but a branchial plume
springs from its base. The tenth had a small epipodite, without a
branchia in the specimen examined ; but I am inclined to think that
the branchia may have become detached ; for the similar epipodite of
the external maxillipede bore a mutilated, or rudimentary, small
branchia. The small epipodites of the second maxillipedes figured
by Milne-Edwards were broken off; but the places to which they
were attached were discernible. The first maxillipede had no
epipodite, in which respect Thalassina approaches Callianassa.
There are twelve arthrobranchiz attached in pairs, from the second
maxillipede to the penultimate thoracic limb inclusively. All these
branchize are remarkable from the fact that, for a greater or less
distance from the base of the gill, the stem is provided with broad
imbricated foliaceous expansions, which are traversed by ramified
1878. ] AND DISTRIBUTION OF THE CRAYFISHES. 779
vessels, and take the place of the branchial filaments, which are
sometimes very few, and confined altogether to the basal region of
the branchia. These branchial plates differ from those of the true
phyllobranchize in their small number and in their disposition, in-
asmuch as they are directed obliquely to the stem and not at right
angles to it. Nevertheless it is interesting to find, in both Awius
and Thalassina, a certain approach to the phyllobranchiate type,
which is completely reached in Gedia and Callianassa.
The Branchial formula of Thalassina.
Somites Arthrobranchie.
Be ectiel) aces se bee
appendages. ‘Anterior. Posterior. te
Red 0 0 0 0 = 0
VIII... 0 (ep) 1 ] 0 = 2+ep
ax? 1 l 1 r= 3
er: 1 1 1 0 = 3
pC 1 ] 1 Oe = 98)
XII. 0 (ep) iI ] 0 = 2-+¢ep
XIII. 0 (ep) l l 0 = 2-+ep
XIV. 0 0 0 ae
B-Pe ep a) AE 6 ese Oe 15-25 ep:
In Stenopus hispidus the branchiz resemble those of Astacopsis
in structure, the filaments being loose, slender, and multiserial. But
the disappearance of the branchial plumes from the podobranchize
has gone still further than in Thalassina; and all these organs are
reduced to delicate, almost vesicular epipodites entirely devoid of
any proper branchial filaments, except in the case of the second
maxillipede, in which a few such filaments are attached to the base
of the epipodite. Moreover the number of the pleurobranchiz is
increased to seven.
The Branchial formula of Stenopus.
Somites Pein Arthrobranchiz. ner ae
% ———-_-—_—~ =
ere aes Anterior. Posterior. oa
WV inns, -'.0,(ep) ? 0 De ty Gen)
WEA 54 3.3.) 0,(ep:1) 1 1 1 = 3+epr
eg... O(ep) Los 1 I= 3-tep
Baas.c, OED) 1 1 1 = 3+ep
XI,...... ,0(ep) l 1 1 = 3+ep
mit... 0 (ep) 1 1 fr = 3+ep
Si; ... . O (ep) 1 ] P= Sep
a 0 0 io
0+7ep+r4+6 4+ 6 + 7 = 19+7/ep+r
780 PROF. T. H. HUXLEY ON THE CLASSIFICATION — [June 4,
I am inclined to suspect the existence of a rudimentary anterior
arthrobranchia in VII.; but I could not make sure of it.
By the structure of its branchize Stenopus is sharply separated
from Peneus, with which it has hitherto been associated, although
it approaches Peneus in the almost complete abortion of the bran-
chial element of the podobranchize. In the Penzida, in fact, each
branchia consists of a stem which is pointed at both ends and gives
off two series of opposite lateral branches. This stem is attached
by a pedicle near its lower end. Near each end of the stem the
lateral branches are very short, and stand straight out; but they
rapidly become longer; and as they do so they curve outwards to-
wards one another, and eventually meet in the middle line. The
middle of the branchia consequently assumes the form of a hollow
cylinder.
The outer face of each lateral branch gives off a close-set series
of secondary branches, which diminish in size towards the free end
of the lateral branch, and at the free end are simple undivided.
filaments. But towards the attached end of the branch the secon-
dary branches are themselves dichotomously subdivided in the direc-
tion of their length; so that the most complicated of these secondary
branches presents a short stem whence two branches proceed, each
of these again gives off two, and these may terminate in yet other
two. Hence each secondary branch is like a flat triangular plate
slit by fissures of varying depths, and attached by its apex to a
lateral branch. All these secondary branches are directed upwards
and outwards.
A detached lateral branch closely resembles one of the branchiz
of Thysanopoda ; while, so far as I can judge from the figures given
by Kréyer! and Sars®, the branchiz of Sergestes and Lophogaster
still more closely approach those of Peneus.
A similar structure was described by Duvernoy in Aristeus, and
was supposed by him to be characteristic of that genus; but Dana
has already justly expressed a doubt whether, in this respect, Aris-
teus differs from the other Penzeidz.
In Penaeus, the last and the penultimate thoracic limbs present no
trace of podobranchiz; and inall the rest the podobranchia is reduced
to a small epipodite, which, in the middle of the series, is bifurcated
atitsfreeend. The latter lies between the arthrobranchiee of its own
somite and the next’following. There are seven pleurobranchiz, of
aoe the hindermost is the largest, while the most anterior is very
small.
ae ee til en monographisk Fremstilling af Kraebsdyrslagten Sergestes :
2 Beskrivelee over Lophogaster typicus: 1862.
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 781
The Branchial formula of Penzeus brasiliensis.
abn Pada Arthrobranchiz. Pivare.
an eir : —_—_—~—_~ z
appendages. branchise. Anterior. Posterior. branchiee.
‘34 ae 0 (ep) ] (small) 0 0 = I+ep
SAAT... 0 (ep) 1 1 1 = 38+ep
EXs sls 0 (ep) 1 l 1 = 3+ep
: a 0 (ep) ] ] 1 = 3-+ep
Le 0 (ep) 1 1 1 = 3+ep
Bt Ree 0 (ep) l 1 1 = 3-+ep
MTS 0 1 1 legs ao
>. 0 0 0 1S
0+6ep +7 + 6 + 7. == .20=56 ep.
The arthrobranchia of VII. is very small; in fact, it is a mere
tuft of dichotomously divided branchial filaments.
The results of this examination of the principal forms of those
Trichobranchiata which possess more than one kind of branchiee may
be thrown into a tabular form, as follows :—
I. No appendages to the first abdominal somite in either
sex. Astyla.
f. Podobranchize undivided: .....0..2:+0+.c0ssd0-sesesecaess PARASTACIDE.
2. Podobranchiz divided into branchial and epipoditic
OEUIONSmNes. jeacucseece tic toteas eee sateen inane names PALINURID,
I. Appendages of the first abdominal somite almost always
present, and specially modified in the males. Stylophora,
1. Podobranchix partially divided.............0..00ccceecaee PoramoBuDs.
2. Podobranchiz completely divided or reduced to epi-
podites.
a. Pleurobranchiz not more than four.
a. Posterior pleurobranchia present, and the
most posterior of the podobranchix com-
[SGLGHPAALE RRR pose anes eter te scare eae Homaripz,
Homarus, Nephrops.
B. Posterior pleurobranchia wanting, and more
or fewer of the posterior podobranchiz
reduced to epipodites.
a. Branchize with only filamentous pro-
COBRGB A. sade siedenmacya-cahiussinekWon Saactaes Axius,
6. Branchiz with foliaceous as well as
filamentous processes ...............+4- Thalassina.
b, Pleurobranchiz more than four.
a. Branchiz filamentous ...............seccceccouee Stenopus.
Bo cBranchies ramose sins esswanereanaanceees spain Peneus.
All the Trichobranchiata are Macrura, in the sense of having the
abdomen and its appendages well developed; but, in Iéacus and
Thenus, the abdomen becomes shortened and the cephalothorax wide,
while the antennary and the ocular regions are modified in a manner
very similar to that which gives rise to the peculiar “face” or
““metope”’ of the typical Brachyura. A very little further modifi-
cation would convert Thenus, for example, into a trichobranchiate
Crab. Such forms as these, which simulate the Brachyura, and yet
differ profoundly from them, may be termed ‘“ pseudo-carcinoids.””
782 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
If the branchial filaments of Avius were shortened and widened,
the structure of the branchize would approach that which obtains in
Gebia and Callianassa, which are truly phyllobranchiate. But in
other respects there is a wide interval between these genera, on the
one hand, and Awius and Thalassina, which are ordinarily associated
with them among the Thalassinide, on the other hand; for the
podobranchize have entirely disappeared on the six hinder thoracic
limbs, and even on the first or second maxillipede they are repre-
sented only by rudimentary epipodites.
There are no pleurobranchie ; and the total number of gills is re-
duced to five pairs of arthrobranchiz on each side.
The Branchial formula of Gebia and Callianassa.
Rene | odlo- Arthrobranchie. Pleuro-
appendages. branchue. Anterior. Posterior. Braviehice.
Vile 20 0 0 On — ao
WDE se. 40 0 0 0; 5=.50
XC eA) ] 1 OF pate
Geen eesti!) 1 1 5 ieee
xe. 0 ] 1 0 = 2
ME 80 1 1 Ng ehh
See AO 1 1 Sn
XIV... 0 0 0 a
0 + 5 ae 5 + 0 10
In the almost complete abortion of the podobranchiz, and in the
presence of ten arthrobranchiz attached in pairs to the middle tho-
racic somites, Gebia and Callianassa agree with Porcellana, Gala-
thea, Lithodes, Pagurus, and Remipes. But in Galathea and
Porcellana the four hindermost pleurobranchize are present, making
fourteen gills on each side; in Lithodes and Pagurus the penul-
timate pleurobranchia exists, making eleven ; in Remipes there is no
pleurobranchia, and only nine arthrobranchiz, viz. one on the ninth
and two for each of the four following somites, are present.
In this group, which nearly corresponds with the Anomala of De
Haan, and which I propose to term the ‘‘ Anomomorpha,” there is
every degree of modification—from such typically Macrurous forms
as Gebia and Galathea to such pseudo-carcinoids as Lithodes and
Porcellana. It is interesting to remark, however, that, while in
Thenus and Ibacus the process of modification has chiefly affected
the head, in the Anomomorpha the characteristic changes are more
marked in the abdomen. In none of the latter are the basal joints
of the antennee fixed, nor are distinct orbits formed.
It is easy to understand the possibility of the derivation of the
Anomomorpha from some form allied to Axius and Thalassina (but
with four pleurobranchiz) by the further reduction, and final almost
complete disappearance, of the podobranchiz, while the biserial
filaments of the other gills flattened out and became lamellar.
The Prawns and Shrimps (‘‘Salicoques”’ of Milne-Edwards, Carides
1878.] AND DISTRIBUTION OF THE CRAYFISHES. 783
of De Haan), if we exclude the Penzeidee, constitute a natural assem-
blage, to which I will apply the name of “ Caridomorpha.” They are
all eminently Macrurous; and the characteristic feature of their
branchial system is the predominence of the pleurobranchiz, and the
concomitant diminution in the number and the importance of the
arthrobranchize and of the podobranchiz. In fact, so far as I am
aware, there are never any traces of the latter except upon the
maxillipedes.
In both Palemon and Crangon I find five pleurobranchiz attached
to the posterior thoracic somites, from the tenth to the fourteenth
inclusively. In Palemon, two arthrobranchize, one of which is very
small, are attached to the arthrodial membrane of the external maxil-
lipede, which has a very short and rudimentary epipodite. The second
maxillipede bears a podobranchia divided into a small branchia and
an oval epipoditic plate. In the first maxillipede the place of the
podobranchia is occupied by a rounded bilobed lamella.
In Crangon none of the maxillipedes bear gills. The epipodite
of the first maxillipede is relatively much larger and triangular ;
that of the second is tongue-shaped and almost vesicular; that of
the third is a mere rounded process.
I can find only one arthrobranchia on the ninth somite.
The Branchial formula of Palemon.
Somites Pod Arthrobranchiz. Piece
E odo- euro-
and their branchiz. nd Suara: branchiz.
appendages. Anterior, Posterior.
MIE onc O(ep) 0 0 0 = O-ep)
WEEE... I 0 0 He
TX.sa.s 70. (ep) 1 1 0 = 2-+ep
KM =O 0 0 Is 22.21
Pligeices oa 0 0 Lem Des ey
MN... “0 0 0 Leo el
2.11) Cee 0) 0 0 eee |
XEVE ace O 0 0) | pn — len |
14+2ep +l + 1 + 5 = 842 ep.
From the number of their pleurobranchize the Caridomorpha can-
not be regarded as a reduced modification of any of the Tricho-
branchiata, except the Penzide and Stenopus. But it is easy to
derive them from a Stenopus-like primitive form by the reduction
of the podobranchiz and the arthrobranchiz, and the conversion of
the five posterior pleurobranchiz into gills of the lamellar type.
In the Brachyura of Milne-Edwards the disposition of the bran-
chial apparatus is well known to be definite and characteristic. In
Cancer pagurus, for example, there are nine branchie ; seven of these
are pyramidal in form, and take a general direction from the base
towards the apex of the branchial chamber, to the inner walls of
which they are applied. The two posterior of these gills are pleuro-
branchize, being attached respectively to the epimera of the eleventh
and the twelfth somites. The fifth and fourth, the third and second,
784 PROF. T. H. HUXLEY ON THE CLASSIFICATION [June 4,
are fixed in pairs by a common pedicle to the arthrodial membrane
of the appendages of the tenth and ninth somites—that is, the great
forceps and the third maxillipede. The most anterior gill, slenderer
than the others, is attached to the arthrodial membrane of the
second maxillipede. ‘There are therefore five arthrobranchiz.
The podobranchia of the first maxillipede is represented only by
the large curved epipodite, which sweeps over the surface of the
arthrobranchiz and the pleurobranchize. The podobranchia of the
second maxillipede is divided into a branchial plume, which lies
horizontally under the bases of the four anterior arthrobranchiz, and
an epipodite, which ascends between the arthrobranchia of its somite
and those of the next, and lies internal to the latter, close to the inner
wall of the branchial cavity.
In the third maxillipede the epipodite is very long, and forms the
valve to the afferent aperture of the branchial cavity. Attached to
its base is a short truncated branchia, which fits in between the bases
of the second and the third arthrobranchie.
7 * > . ‘ . _
The Branchial formula of Cancer pagurus.
Somites Polo: Arthrobranchie. eine:
and their jranchive. Aap PRG Tr, branchiz.
appendages. Anterior. Posterior.
Wane ks 0 (ep) 0 0 0... 0 (ep)
Wise 1 1 0 Oly t= sae
1 ree l ] 1 0.5= ° 3
Dicer 0 1 1 Oe
pa fen se 0 0 0 een
m4 Ieper 0 0 0 baie Sat yl)
b.¢ fi Fs 0 0 0 0. = 0
bh eae 0 0 0 0 = 0
2+ep+3 4+ 2 + #2 = YQ+ ep.
A considerable reduction of the branchial apparatus occurs in some
Brachyura, especially the Catometopa. Thus, in Gelasimus the
hinder pleurobranchia and the most anterior arthrobranchia have
disappeared, and the two podobranchiz are so small as to be almost
rudimentary.
In the Raninide and in Laéreillia, the Brachyurous metope is
incompletely formed; but the branchial system is essentially that
of the true Crabs. The same may be said of Dromia, although the
branchial apparatus of this crustacean presents some very interesting
approximations to the less-modified type from which it cannot be
reasonably doubted that the Brachyura have proceeded.
The epipodites of. the three maxillipedes are very similar in form
and disposition to those of the ordinary Crabs ; and that of the second
bears a small horizontal branchial plume. That of the third some-
times bears a small branchia. The coxopodite of the fourth thoracic
limb has a small epipodite ; but I find no trace of such an appendage
on the rest of the thoracic limbs. Moreover there are five pairs of
arthrobranchiz attached to consecutive somites from the ninth to
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 785
the thirteenth—and four pleurobranchize, one for each of the four
posterior thoracic somites.
The Branchial formula of Dromia.
ea ih. Podo- Pola cata Pleuro-
appendages. oar Anterior. Posterior, branchie.
VIT. oewe 0 (ep) 0 0 0 me 0 (ep)
sy 9 ] 0 0 ‘ee a |
Meme): 8 1 1 ] Ge 2s
2. - 0 1 ] 0 2% 2
XII 0 1 1 1 Ne
Vee: 0 0 0 ] ba l
witty te cdl spe 5 vas a aC
On comparing this branchial formula with that of Homarus, the
relation between the two is obvious. In fact, if the three posterior
podobranchize of the Lobster are suppressed, and the next is reduced
to an epipodite, the branchial formula becomes the same as that of
Dromia, and the remaining differences between the respiratory organs
of the two result from the modification in form and structure of the
branchial elements which remain. Thus it is a permissible, if not a
probable, suggestion that, just as the Anomomorpha may have been
derived from the modification of some form allied to Avius, and the
Caridomorpha from some form allied to Stenopus and Peneus, so
the Carcinomorpha (= the Brachyura and the Raninide, with
Homola and Dromia) may have proceeded from some Homarine
stock.
However this may be, the actual morphological relations of the
Thoracostraca appear to me to be represented with tolerable accuracy
by the following scheme :——
CRUSTACEA.
THORACOSTRACA OR PODOPHTHALMIA.,
Ill. PHYLLOBRANCHIATA.
CARIDOMORPHA. ANOMOMORPHA. CARCINOMORPHA,
II. TRICHOBRANCHIATA,
Palinuride,
Peneide. Stenopide, Thalassinide. Axiide. Homaride, Potamobiide. Parastacida,
— ~~
=> — -—y- 7 — a=
Euphauside,
I. ABRANCHIATA,
Myside.
———— a a
seas
STYLOPHORA. ASTYLA,
—— -—,~-—_—
786 PROF. T. H. HUXLEY ON THE CLASSIFICATION | June 4,
IV. Tur DistriBuTion OF THE CRAYFISHES CONSIDERED IN
RELATION TO THEIR MorPHOLOoGICAL DIFFERENCES.
From what has been said above, it will be obvious that there is a
remarkable correspondence between the morphological and the geo-
graphical divisions of the Crayfishes. Thus, all the Crayfishes of the
northern hemisphere are Potamobiide, and all those of the southern
hemisphere are Parastacide. In the northern hemisphere, again,
the Astaci are Eurasiatic and West-American, while the Cambari
are characteristic of the North-American region east of the Sierra
Nevada—in other words, of the river-basins which flow into the Gulf
of Mexico and the West Atlantic.
The Astacine region nearly answers to the Palearctic province of
Mr. Sclater, minus the southern shores of the Mediterranean, and
plus Western North America; while the Cambarine region takes
in most of the Palearctic region, with the Neotropical region as far
as Guatemala and the West Indies.
In the southern hemisphere, Astacopsis, Cheraps, and Hngeus
are confined to the Australian region, Paranephrops to New Zea-
land and the Fijis ; while the South-American Parastacus is distinct
from either of these, though closely allied with the Australian
forms ; and the peculiarity of the Madagascar fauna is vindicated
by Astacoides.
Thus, if we were to establish provinces of distribution on the
Crayfishes alone, they would bear only a partial resemblance to those
based on the association of terrestrial animals. On the other hand,
if we compare the distribution of the Crayfishes with that of the
freshwater fishes, there are, as might be expected, some curious
points of resemblance. The distribution of the Salmonide, for ex-
ample, corresponds pretty closely with that of the Potamobiide,
though the range of the Salmonidee extends less far to the south
in North America, and a little further, namely, as far as Algeria in
the Old World. Again, the Salmonide to the east of the Rocky
Mountains are, for the most part, distinct from those to the west,
while the genus Onchorhynchus is, like Astacus, common to both
the Asiatic and the American shores of the North Pacific.
With the singular exception of Retropinna, there is no true Sal-
monoid in the southern hemisphere; but, as Dr. Giinther has
pointed out, the Haplocbitonidee and the Galaxiadee, which stand in
somewhat the same relation to the Salmonide as the Parastacide do
to the Potamobiidee, represent the Salmonide in the fresh waters of
New Zealand, Australia, and South America.
It is-worthy of remark that the Salmonide, the Haplochitonide,
and the Galaxiadz are singular among Teleostean fishes for the em-
bryonic character of their female reproductive organs, which have
no oviducts—just as, among the Podophthalmous Crustacea, the
Crayfishes are distinguished by the undifferentiated character of
their podobranchiz.
With the exception of one or two species in Algeria and Asia
Minor, the Salmonoids and their allies are wanting in the whole of
1878. | AND DISTRIBUTION OF THE CRAYFISHES. 787
Africa and Asia south of the great Asiatic highlands, just as the
Crayfishes are. It will be very interesting to learn, from the thorough
investigation of the fauna of Madagascar, which is now being carried
out, whether the Salmonoids or their allies are in any way repre-
sented there.
The broad similarity in distribution between the Salmoniform
fishes and the Crayfishes is doubtless due to the likeness of the
conditions under which the two groups have reached their present
development. I do not think that there can be any reasonable
ground for questioning the assumption, that both the freshwater
fishes and the freshwater Crustacea are modifications of a marine
prototype, which has more or less completely adapted itself to fresh-
water conditions. In the case of the Crayfishes, at any rate, there
is abundant analogical evidence in support of this hypothesis. It is
well known that, in many parts of the world, the Prawns ascend
rivers, and become fluviatile. The Palemon lacustris (Anchistia
migratoria, Heller) of the Lago di Garda is identical with a Prawn
now living in the Mediterranean. Again, the 1, ysis relicta of the lakes
of Norway, Sweden, Western Russia, and North America (Michigan
‘and Superior) is only a variety of the Mysis oculata of the Arctic
seas’. Nor do I think it can be seriously questioned that the
fluviatile and the land Crabs are modified descendants of marine
Brachyura.
Let it be supposed that, at some former period of the earth’s
history, a Crustacean, similar to Paranephrops or Astacopsis in its
general characters, but with the first pair of abdominal appendages
fully formed, which we may call provisionally Protastacus, in-
habited the ocean, and that it had as wide a distribution as Palemon
or Peneus have at the present day. Let us suppose, further, that
the northern form of the genus tended towards the assumption of
the Potamobiine, and the southern towards that of the Parastacine
type. Under these circumstances, it is easy to understand how such
rivers as were, or became, accessible in both hemispheres, and were
not already too strongly tenanted by formidable competitors, might
be peopled respectively by Potamobiine or Parastacine forms, which,
acquiring their special characters in each great river-basin, would
bring about the distribution we now witness. As time went on, the
Protastacus stock might become extinct, or might be represented
only by rare deep-water forms, as the Homaridee are represented in
the Indian Ocean only by Nephropsis.
Some such hypothesis as this appears to me to be fully justified by
the present state of knowledge; and though it cannot as yet be said
to be directly supported by palzeontological facts, these facts agree with
the hypothesis very well as faras they go. For the Mesozoic marine
1G. O. Sars, ‘Histoire Naturelle des Crustacés d’eau douce de Norvége.’
In the British Museum there is a species of that especially marine genus
Peneus, which is affirmed by the Messrs. Schlagintweit to have been obtained
from an affluent of the Sutlej, at the foot of the Himalayas. Peneus brasiliensis
ascends the North-American rivers for long distances (Smith, in Prof. Baird’s
Report, 1872-72).
788 THE SECRETARY ON ADDITIONS TO THE MENAGERIE. [Nov. 5,
strata abound in Crustacea, such as Glyphea and Hoploparia, which
are evidently closely allied to the Crayfishes.
The great difficulty is, not to account for the Crayfishes where we
find them, but to understand their absence over so large a part of the
Old World and of intertropical America. Whether this is to be
explained by extensive alterations in geographical conditions since the
extinction of the Protastacus stock, or by the competition of Prawns
and freshwater Crabs, or by some other circumstance at present
unknown, is a very interesting subject for further inquiry.
Postscript, Oct. 24, 1878.
Since this paper was read, my friend Mr. Moseley, F.R.S., has
written for and obtained specimens of the ‘‘ Crayfish”’ said to exist
in the fresh waters of the Cape-Verd Islands. They belong to the
genera Atya and Palemon. Moreover, by the intervention of Sir
Joseph Hooker, I have procured a considerable number of fresh-
water Crustacea from Jamaica. But these also all belong to Atya
and Palemon. I suspect that all Sloane’s ‘‘Crayfishes” are
simply Prawns. The largest example of one of the species sent to
me measures sixteen inches in length when the great chelate limbs
are fully extended.—T. H. H.
November 5, 1878.
Arthur Grote, Esq., V.P., in the Chair.
The Secretary read the following reports on the additions to the
Society’s Menagerie during the months of June, July, August,
September, and October 1878 :—
The total numiber of registered additions to the Society’s Mena-
gerie during the month of June was 159, of which 35 were by birth,
75 by presentation, 29 by purchase, 14 were received on deposit,
and 6 by exchange. The total number of departures during the
same period, by death and removals, was 73.
The most noticeable additions during the month of June was as
follows :— }
A Japanese Wolf (Canis hodophylax, of the ‘Fauna Japonica,’
Mamm. t. 9, p. 38), presented by H. Heywood Jones, Esq., F.Z.S.,
June 26th, being the first example of this little-known animal we
have ever received alive.
Judging from the present specimen the Japanese Wolf, although
nearly allied to Canis lupus, would seem to be a distinct species, to
be recognized by its smaller size and shorter legs.
The Japanese Dog sent to us by Mr. Pryer (see-P. Z.S. 1878,
p- 115) is quite a different animal, and is, I have no doubt, only a
variety of the domestic dog or a hybrid,
1878.] THE SECRETARY ON ADDITIONS TO THE MENAGERIE. 789
The registered additions to the Society’s Menagerie during the
month of July were 154 in number ; of these 68 were acquired by
presentation, 45 by purchase, 7 by exchange, 18 were bred in the
Gardens, and 16 were received on deposit. The total number of
departures during the same period by death and removals was 108.
The most noticeable additions during the month were :—
1. A male Beatrix Antelope (Oryza beatrix), presented 10th July,
1878, by Commander F, M. Burke, of B. I. S. N. SS. ‘Arcot.’
Capt. Burke writes me that he obtained this animal at Jeddah
from a friend of his, who received it as a present from the Shereff
of Mecca. It had been originally captured in the neighbourhood of
Tyeff or Tayf, some 150 miles E.S.E. from the Red Sea, in the
Hedjaz passes.
I am sorry to bave to add that this interesting animal (so far as
I know, the third’ specimen of its species known) reached the Gar-
dens in a very poor state of health, and died shortly afterwards.
2. A fine young female Orang (Simia satyrus) received in ex-
change from the Zoological Gardens of Calcutta, July 27th.
The registered additions to the Society’s Menagerie during the
month of August were 117 in number. Of these 40 were acquired
by presentation, 12 by purchase, 13 were bred in the gardens, 1
were obtained by exchange, and 38 were received on deposit. The
total number of departures during the same period, by death and
removals, was 116.
The most noticeable additions during the month were :—
1. One Spanish Blue Magpie (Cyanopolius cooki), presented
August 2nd, by Lord Lilford, F.Z.S.
z. A young example of the Prairie-Wolf of North America
(Canis latrans), presented by Prof. C. M. Vincent, F.G.S., F.R.G.S.,
August 21st.
Both these species are new to the Society’s collection.
3. Three hybrid Boas, bred between a male of Chilobothrus inor-
natus and a female Epicrates angulifer, and born on the 30th of
August,
There can be no question about the paternity of these curious
hybrids, as there is no male Hpicrates in the collection, and the
Chilobothrus was observed in copulé with her in March or April last.
The three young ones were brought forth alive on the 30th of
August ; and six eggs, more or less decomposed, came away subse-
quently.
So far as I know, hybridism among the Ophidians in captivity has
not yet been recorded, and in this instance is the more singular as
occurring between species of different genera.
The total number of registered additions to the Society’s Mena-
gerie during the month of September were 85 in number; of these,
43 were acquired by presentation, 4 by purchase, 13 by birth, and
1 For a notice of two previous specimens, see P. Z.8. 1857, p. 158, Mamm.,
pl. ly., and P. Z. 8. 1872, p. 608.
Proc. Zoou. Soc.—1878, No. LIL. 52
790 THE SECRETARY ON ADDITIONS TO THE MENAGERIE. [ Nov. 5,
25 were received on deposit. The total number of departures during
the same period, by death and removals, was 78.
The most noticeable additions during the month were as follows :—
1. An Oriental Eagle-Owl (Budo orientalis), from Karennee, in
the interior of Siam, presented September 16th by Charles Fowler,
Esq., being the first specimen of this remarkable Owl that has been
received by the Society.
2. Two fine Ostriches (Struthio camelus), presented by the Hon.
H. C. Vivian, H.B.M. Consul-General for Egypt; and two Seere-
tary Birds (Serpentarius reptilivorus), presented by C. Rivers
Wilson, Esq., C.B.; received September 19th. These birds are
from the collection of the Khedive of Egypt, at Cairo.
The total number of registered additions to the Society’s Mena-
gerie during the month of October 1875 was 93, of which 4 were
by birth, 43 by presentation, 21 by purchase, 20 were received on
deposit, and 5 by exchange. ‘The total number of departures
during the same period, by death and removals, was 123.
The most noticeable additions during the month of October were
as follows :—
1. A female Sumatran Rhinoceros (Rhinoceros sumatrensis),
deposited by Mr. C. Jamrach, October 4th. This Rhinoceros,
which is full-grown, seems to agree in every respect with the two
females of the same species previously purchased (in 1872 and 1875’).
2. Two Leopard Tortoises (Testudo pardalis), presented by the
Rev. G. H.R. Fisk, C.M.Z.S., of Capetown, October 9th. Mr. Fisk
writes to me as follows respecting these Tortoises :—
*<T am informed by Mr. Foster, of Clanwilliam, that in the year
1846 two full-grown Leopard Tortoises were transferred from Kaffir-
land to Clanwilliam, where they remained for many years without
progeny, but that in the year 1857 these two young ones were
hatched from the eggs of the female. We arrive, then, at the inter-
esting fact, speaking of a Tortoise after the manner of men, which,
perhaps, we may do of a Vertebrate, that these Tortoises are now of
age, having lived just twenty-one years, and yet, according to the
nature of Tortoises, are only children; for I am assured that they
are in size scarcely two thirds of that of their parents, which died
early in the year 1877, and within three months of each other.
**T wish to bestow on this Tortoise and its fellow the popular
names of Kreli and Sandilli, in remembrance of the land inhabited by
their ancestors, aud of the war just ended between the Colony and
those chiefs who will ever find a place in the history of the Cape.”
3. A young male Tufted Deer (Zlaphodus cephalophus*), pur-
chased October 15th, being the second example of this rare and
little-known animal received alive. Having been caught in a trap, it
is unfortunately deficient in one of its fore feet.
4. A female of the new Fallow-deer lately described by Sir
Victor Brooke in the Society’s ‘ Proceedings’ (P. Z.S. 1875, p. 261)
1 See Trans. Zool. Soe. ix. p. 659, et P.Z.S, 1876, p. 694.
2 See P. ZS. L876, p. 275 et pp. 757, 758.
1878.] MR. J. H. GURNEY ON ASTURINULA MONOGRAMMICA. 791
as Cervus mesopotamicus, purchased October 31st. This makes a
pair of this fine Deer now in the Society’s Gardens. Like the male
obtained in 1877, the present specimen was obtained at Bussorah
and brought to England by our energetic correspondent Capt.
Phillips, of the 8.8. ‘ Mesopotamia.’
I take this opportunity of recording the following facts relating to
some hybrid Monkeys lately born in the Society’s Menagerie, which
Prof. Garrod has kindly put together for me :—
During the earlier months of the present year (1878) there were,
together in one cage, three Monkeys along with others of smaller size
and less developed. The three were :—(1) a male of Macacus cyno-
molgus, or of one of the allied local forms, from Upper Burmah
(presented on May 19th, 1875), a particularly fine specimen; (2) a
female adult Cercocebus fuliginosus (presented on the 3rd of April,
1876); and (3) a female Mandrill, Cynocephalus mormon (presented
on the 5th of September, 1877), not adult.
The keepers of the House assert that they repeatedly observed the
male Macaque in copulation with both females.
On October 2nd the Mangabey fell down from a high perch in
her cage dead. All organs, including the brain, appeared quite
healthy on postmortem examination. The uterus contained a foetus
far advanced in growth, apparently lacking about three weeks or a
fortnight of full development.
On October 14th the Mandrill gave birth to a live young one,
which still survives.
There seems no reason to doubt that the Macaque was the father
of both the young ones, there being no male Mandrill nor Mangabey
which could have had access to them, and the Mandrill’s young
one having a short tail’.
The following note was read from Mr. J. H. Gurney, F.Z.S. :—
“Northrepps Hall, Norwich,
July 30, 1878.
‘In the ‘ Proceedings’ of the Society for the present year, mention
is made at page 354 of two specimens of Asturinula monogrammica,
procured by Mr, E. C, Buxton at Darra-Salam, on the eastern
coast of Africa.
“One of these skins was presented by Mr. Buxton to the Norwich
Museum, accompanied by the following memorandum :—‘ Sings like
any thing of an evening and, I believe, morning.’
“IT am not aware that this habit of 4. monogrammica has been
previously noticed; and as it appears to indicate an affinity between
this species and the genus Melieraz (which Asturinula also resembles
when adult, in the remarkable red coloration of the cere, tarsi, and
feet), I think Mr. Buxton’s short note on the subject ought to be
recorded. “J. A. Gurney.”
1Cf. Blyth’s notice of a hybrid Monkey between Macacus nemestrinus and
Cynocephalus porcarius in J. A. 8. B. xxxii. p. 455 (1863).
92*
792 MR. A. H. EVERETT ON ANOA DEPRESSICORNIS. [ Nov. 5,
The Secretary read the following extract from a letter addressed
to him by Dr. A. B. Meyer, C.M.Z.S., of the Royal Zoological
Museum, Dresden :—
‘*Mr. Van Musschenbroek, the well-known discoverer of Diphyl-
lodes gulielmi I11., tells me that one of his colleagues in the
Moluccas received about ten years ago from the west coast of New
Guinea (vid Aru Islands) a Bird-of-Paradise as large as P. apoda,
but in general like P. minor, with this difference, that the whole
body was covered with orange-coloured feathers. This bird was
presented to the king of Holland ; but never was any thing heard of
it again. In the year 1875 Mr. Van Musschenbroek saw in the
hands of Prof. Steere, at Menado, among a collection of bird-skins
which the latter had bought at Amboina, a specimen of P. minor
which reminded him of the orange Bird-of-Paradise mentioned
above. This specimen was spotted with yellow all over; but it was
a bad skin, and gave Mr. Van Musschenbroek the impression that it
had belonged to a sick bird.
“Are these individual varieties of P. minor? or have we to deal
with a new and unknown Bird-of- Paradise?”
A communication was read from the Marquis of 'Tweeddale, Pre-
sident, pointing out that a correction made im the proof of his last
paper on Philippine Birds, anted, p. 611, had been through some
mistake overlooked. ‘The correct specific name of the Polyplectron
there mentioned, page 623, was xapoleonis, Lesson, not emphanes,
Temminck, Lesson having given a description of his bird in the
additions and corrections to his ‘Traité d’Ornithologie,’ page 650.
The followmg extract was read from a letter addressed to Lord
Tweeddale by Mr. A. H. Everett, dated Singapore, July 21, 1878 :—
“ Just before leaving Manilla I paid a visit to the small museum
attached to the College of St. Thomas. There were a few bird-skins ;
but the room was very dark, and, being with a party of friends, I
could not make any satisfactory notes. My visit was, however,
repaid by finding a specimen of the Wild Ox of Celebes (dAnoa
depressicornis), which had been brought from Mindoro. I had
been led to suspect its existence in that island, but had hitherto
failed to get any tangible evidence. I cannot say whether the
Philippine animal is identical with that found in Celebes or not.
The Professor of Natural History, who had no idea of the interest
attaching to his specimen, although he had labelled it correctly, said
that he had never heard of the 4noa being found at that part of the
world, except in the rugged interior of Mindoro, where, however, it
abounded. It is not easy to weigh the full import of the occurrence
of this singular animal in the Philippines until something more is
kuown about the faunaof Mindoro. 1 had previously been disposed
to doubt the existence of another Celebesian animal, the black
Cynopithecus, in the islands; but I now feel less confidence on this
point, and it is quite possible that it may turn up in Mindoro.”
1878. ] MR. R. B. SHARPE ON A NEW INDICATOR. 793
Professor Newton, M.A., F.R.S., V.P., exhibited the skin of a
bird supposed to be a hybrid between the Red Grouse (Lagopus
scoticus) and the Ptarmigan (LZ. mutus), observing :—
“This remarkable specimen was lately given to me for the museum
of the University of Cambridge by Capt. Houston, of Kintradwell,
in Sutherland, having been shot there out of a covey of Grouse on
the Ist of September, 1878. As will be seen, it bears some con-
siderable resemblance, above, to a hen Ptarmigan in summer
plumage; but its general appearance is much darker. Beneath,
there is a greater resemblance to the young of the Red Grouse ; and
the primaries are much as in that bird, being, however, partially
edged with white to a much greater extent than is commonly found
in the latter. I have shown the skin to several ornithological
friends, none of whom have been able to offer any other suggestion
concerning it than that originally made by the donor, namely that
it is a hybrid between the two species named; and in confirmation
thereof, Capt. Houston told me that the part of his ground on
which it was shot is close to a locality frequented by the Ptarmigan.
Without having made an exhaustive search, I may say that I am
not aware of any record of such a hybrid as this is supposed to be,
though infcrmation received from several quarters induces me to
believe that other examples have before now occurred; and my
chief object in exhibiting the present specimen is to call attention to
the subject.”
The following papers were read :—
1. On a new Species of Indicator, with Remarks on other
Species of the Genus. By R. Bowpter SuHarpes,
F.L.S., F.Z.S., &c.
[Received July 18, 1878.]
In an article which I sent to Mr. Dawson Rowley’s ‘ Ornitholo-
logical Miscellany’ (vol. i. part iii.) I gave a revision of the Indica-
toridee, or family of Honey-Guides ; and I am now able to add some-
what to the information there recorded.
INDICATOR conrRosTRIS (Cass.) : Sharpe in Rowley’s Orn. Mise.
vol. i. p. 196.
A few specimens of Honey-Guide having in the hasty packing
away of my African collection become mislaid in another box, were
not available when I wrote my paper on Indicatoride. Now that
my collection has been added to the British Museum they have
been discovered ; and I was pleased to find among the missing birds
a fine adult specimen of the present species from the interior of Fan-
tee. It was sent to me by Governor Ussher in one of his last collec-
tions, and is the first example known from the Gold Coast. I find,
794 MR. R. B. SHARPE ON A NEW INDICATOR. [Nov. 5,
on comparing the two birds, that I. conirostris is a very distinct
species from JI. minor, having the head and back strongly washed
with deep golden yellow, the centres of the feathers being blackish
instead of pale brown as in J. minor. This contrast is especially
seen in the wings, where the golden edgings to the black feathers stand
out in bold relief, and make them very different from the appearance
of the South-African species. The Fantee bird measures—total
length 6 inches, culmen 0°45, wing 3°6, tail 2°6, tarsus 0°6.
INDICATOR ARCHIPELAGICUS (Temm.): Sharpe, ¢. c. p. 197; id.
Ibis, 1877, p.8.
As already recorded by me (/.c.) Mr. Alfred Everett procured a
female specimen of this rare species at Bintulu, in North-western
Borneo. ‘Temminck’s description is not very good ; but the species
is unmistakable from its yellow shoulder-spot.
InpicaTor Exits (Cass.): Sharpe, ¢.c. p. 198.
A specimen of this little Honey-Guide was sent recently from
Landana, on the Congo, by M. Petit.
INDICATOR MALAYANUS, sp. n.
General colour above ashy, washed with olive-yellow, dimly on
the head and mantle, the feathers of the back and wings more dis-
tinctly edged with olive-yellow, the primaries scarcely perceptibly
margined ; tail-feathers dark brown, edged indistinctly with olive-
yellow, all but the two centre tail-feathers internally whity brown,
increasing in extent towards the outermost, which is almost entirely
whity brown, obliquely dark brown towards the tip; sides of face
and ear-coverts dingy ash-colour with a very faint olive tinge;
under surface of the body pale yellowish, creamy white on the chin
and upper throat, abdomen, thighs, and under tail-coverts; the
feathers of the breast and fore neck with mostly concealed greyish
bases, imparting asomewhat mottled appearance ; lower flanks mesially
streaked with dark brown; under wing-coverts yellowish white, as
also the edge of the wing; quills dark brown below, whitish along
the edge of the inner web.
Total length 5-7 inches, culmen 0°55, wing 3°9, tail 2°45, tar-
sus 0°55.
The specimen from which the above description is taken was
received by Mr. Henry Whitely, of Woolwich, in a large collection
sent direct from Malacca along with the usual Malaccan birds.
It cannot be the Himalayan species, I. wanthonotus, because the
latter has the back and rump yellow; and it differs from the Bornean
Honey-Guide (I. archipelagicus) in wanting the yellow shoulder-spot,
grey breast, and flanks of the last-named bird, and in being alto-
gether larger. This latter fact seems to prevent the possibility of its
being considered a young bird of I. archipelagicus,
I propose the following amended key for the genus Indicator :—
P78 1878s
GBSowerby hth
NEW MARINE SHELLS.
1878. ] MR. G. B. SOWERBY, JUN., ON NEW SHELLS.
a. Uropygio et supracaudalibus dorso concoloribus, flayo nee albo
marginatis,
a’, Tectricibus alarum minimis interioribus reliquis concolori-
bus; plaga cubitali nulla.
a, Subtus fere concolores, haud squamati.
a", Pectore cineraceo, fascia mystacali nigricante ; rectri-
cibus extimis pure albis brunneo terminatis.
a‘, Majores; ala 3:5-3°7 une.
a’, Pallidus: mento summo albicante ; tergi alarum-
que plumis brunneis flayo-olivascenti marginatis minor.
%. Saturatior: mento summo nigricante; tergi ala-
rumque plumis nigris aurato-flayo marginatis... conirostris.
deeManor: sala: A Diier . scenes desa-s tases tit,- sah datos exilis.
6". Pectore olivascenti-flavido ; fascia mystacali nulla ;
genis et regione parotica saturate cineraceis olivaceo
yix layatis ; rectrice extima albicanti-brunnea, versus
apicem oblique brunnea, reliquis saturate brunneis
intus albicanti-brunneis ............... BY ako er eaeenan se malayanus.
6". Subtus squamati, haud concolores; gule plumis fuscis
albo marginatis quasi squamatis.
e'”, Pectore tantum squamato; abdomine fere concolori
albo ; hypochondriis albis brunneo striatis ............ variegatus.
d'". Pectore et abdomine squamatis ; hypochondriis brun-
MAISMOLE CON COLOTIDUS Fic seaeias daue's «pe. sinaic'ap crea dunes ad aeons maculatus,
b'. Tectricibus alarum minimis interioribus leete flavis, plagam
Conspicuam exhibentibus.............cc0c2 ceescsceeceesssecsesees archipelagicus.
yperag
6. Uropygio et supracaudalibus albicantibus, his albo marginatis.
e'. Tectricibus alarum minimis interioribus lete flavis, plagam
formantibus ; hypochondriis brunneo striatis .............. sparrmant.
d'. Plaga cubitali nulla; uropygio pure albo, haud brunneo
striato ; hypochondriis concoloribus ; pileo flayo lavato.
ce". Regione parotica pallide brunnea; rostro pallide brunneo
d'’. Regione parotica nigricante ; rostro nigrO ...........468
major.
barianus.
c. Uropygio et dorso postico flavis ............ceeceeceeee att atetere xanthonotus,
By G. B. Sowzrsy, Jun.
[Received July 18, 1878. ]
(Plate XLVIII.)
Conus MELVILLI, n. sp. (Plate XLVIII. fig. 1.)
2. Descriptions of ten new Species of Shells,
C. testa abbreviato-subcylindracea, solida, obtuse angulata, levi,
antice crenulato-sulcata, griseo-albida, cinnamomeo-fusco longi-
tudinaliter strigata, lineis fuscis undulatis transverse cincta ;
interstittis lineis curvatis longitudinalibus hic illic irregulariter
divisis ; spira obtusa, fusco-strigata ; anfractibus 7, convezis,
levibus ; apertura modica, fauce castanea.
Long. 20, maj. lat. 11 mill.
Hab. Key West (J. Cosmo Melvill).
A small obtuse Cone, with characteristic markings, somewhat re-
sembling those of C. /wcidus (Mawe), on a smaller scale; the trans-
-verse brown lines are more numerous, and the short longitudinal
ones crossing the interstices lessso. The light-brown streaks almost
796 MR. G. B. SOWERBY, JUN., ON NEW SHELLS. __[Nov. 5,
cover the shell, leaving small white patches between the transverse
lines.
The shell described was picked up on the beach by Mr. Melvill
at Key West (south of Florida) in 1872, and is at present the only
specimen known of the species.
LaTirus CAYOHUESONICUS, n. sp. (Pl. XLVIII. fig. 4.)
L. testa elongaio-turrita, solidula, atro-fusca, antice contracta, sub-
elongata; spira elongata; anfractibus 9, convexiusculis, spira-
liter liratis, longitudinaliter costatis, costis latis rotundatis, liris
alternis minoribus, interstitiis striatim cancellatis ; apertura
ovata, parva, fauce levi, purpureo-fusca; columella bicostata ;
canali subelongato, recto.
Long. 16, maj. lat. 53 mill.
Hab. Key West (J. Cosmo Melvill).
A very dark little Zaéirus, almost black within and without, with
general characters common to the genus, and of the ordinary form
of the larger species, such as L. infundibulum.
Among the shells found by Mr. Melvill at Key West, and sub-
mitted to me, the two above described are all that I can consider
new species ; but the following are worth mentioning :—
Conus japonicus (Hwass), which is identical with C. largillierti
(Kiener) of Reeve’s Conch. Icon. Suppl. pl. 2, species 275. It is
a question whether this species, which in all probability never in-
habited Japan, should retain the name given by Hwass, or whether
that of Kiener should be adopted.
Conus nebulosus: a young specimen with a very sharp elevated
spire.
Hemifusus corona (Gmel.). A number of specimens of a very
small variety of this species, adult and well developed, but ranging
from an inch to an inch anda half in length (Plate XLVIII. fig. 13).
From the collection of the late Sir E. Belcher :-—
Conus CARNALIS, n. sp. (Plate XLVIII. fig. 2.)
C. testa oblonga, subpyriformi, solida, obtuse angulata, obsolete
striata, antice leviter costata; pallide carnea, late pallide fus-
cescenti bifasciata ; spira obtuse-exserta ; anfractibus 9, levi-
bus, in medio depressis ; apertura modica, carneo tincta.
Long. 50, maj. lat. 25 mill.
Hab.?
A yery pale pink, smooth shell, very faintly banded with buff,
of the form of C. scadriusculus.
A single specimen from the collection of the late Sir E. Belcher,
without any note of locality.
CONUS CATENATUS, n. sp. (Plate XLVIII. fig. 3.)
C. testa subfusiformi, subtiliter granoso-lirata, postice acute angu-
lata, antice attenuata, medio leviter contracta ; albida, fusco ir-
regulariter grandimaculata, liris transversis fusco eaximie arti-
1878. ] MR. G. B. SOWERBY, JUN., ON NEW SHELLS, 797
culatis; spira elata, conica, gradata; anfractibus 11, levibus,
leviter coronatis ; apice acuto, fusco; apertura angusta, fusco
tincta.
Long. 31, maj. lat. 14 mill. ; spira alt. 10 mill.
A very neatly marked Cone, allied to C. mahogani, C. ximines,
and C. tornatus, but perfectly distinct, both in form and markings.
Hab. Panama? (ea coll. Sir HE. Belcher).
Mirra BERTHS, n. sp. (Plate XLVIII. fig. 11.)
M. testa angulato-fusiformi, solidiuscula, cinerea, longitudinaliter
striata et valide plicata, transversim leviter sulcata, antice at-
tenuata, recurva, transversim valide costata; spira acuminato-
turrita; anfractibus 10, superne angulatis, ad angulum bicarina-
tis, supra angulum concavis, in medio balteolis nigris duobus cin-
gulatis, linea unica fusca superne circumeinctis ; apice nigro-
Susco ; apertura elongata, intus fusco cincta ; columella quadri-
plicata, plicis duplicatis; labio externo leviter sinuato, levi,
crassiusculo,
Long. 32, maj. lat. 13 mill., spira alt. 13 mill.
Hab. China Sea (Sir E. Belcher).
Several specimens of this pretty shell were found in the above
collection, and seem to have been hitherto unnoticed, although the
species is very distinct and easily distinguishable from its con-
geners.
MITRA FULVOLIRATA, D. sp. (Plate XLVIII. figs. 9, 10.)
M. testa fusiformi, solidiuscula, cinerea, fusco-purpureo late fas-
ciata, fulvo tincta, utrinque fuscata, antice contracta, leviter
recurva; spira acuminato-turrita ; anfractibus 10, convexiuscu-
lis, superne obtuse angulatis, transversim leviter sulcatis, longi-
tudinaliter costatis, costis subangulatis, obtusis, fulvescentibus ;
apertura elongata, fauce levi, fusco-purpurea ; columella quadri-
plicata, plicis obtusis, quadratis ; labio externo simplici.
Long. 27, maj. lat. 9 mill., spira alt. 12 mill.
Hab. China Sea (Sir E. Belcher):
Several specimens presenting scarcely any variation were found
in the above collection. Although a species of no very striking cha-
racter, I cannot identify it with any hitherto described.
Mirra acouTa, n. sp. (Plate XLVIII. figs. 7, 8.)
M. testa angulato-fusiformi, albida, interdum purpureo tincta, fusco
punctata, castaneo unifasciata, vel flammulata; spira acutissima ;
anfractibus 10, transversim carinatis ; carinis numerosis, eleva-
tis, acutissimis ; interstitiis concavis, interdum levibus, plerum-
que cancellatis ; apertura subangusta, intus alba, levi ; columella
quingueplicata, plicis acutis ; labio externo via arcuato, crenulato.
Long. 20-23, maj. lat. 8-9 mill.
Hab. China Sea (Sir E. Belcher).
A number of specimens of this species were found in the above
collection. They vary considerably in sculpture, some being smooth
798 MR. G. B. SOWERBY, JUN., ON NEW SHELLS. __[ Nov. 5,
between the keels, and others cancellated ; but they all agree in
being much more sharply angular than the specimens of M. annu-
lata from Ceylon and Mauritius, the keels being more acute and
prominent, and the body-whorl being shorter in proportion to the
spire.
From other sources :—
MITRA DEBURGHI4, nl. sp. (Plate XLVIII. fig. 12.)
M. testa cylindraceo-conica, solidula, cinerea, hic illic sparsim
fusco punctata, maculis albidis distantibus triseriatim dispositis
cingulata, postice albida, ubique transversim sulcata, sulcis pro-
fundis, valide cancellatis ; spira brevissima, conica, albida; an-
fractibus 9, obtuse angulatis ; apice fusco; apertura oblonga,
antice subexpansa, fauce cinereo-fusca ; columella quinqueplicata,
plicis acutis, labio externo serrato.
Long. 16, maj. lat. 8 mill.
Hab. Tahiti.
A very pretty and distinct species of the Imbricaria section.
Mirra PuNCTURATA, n. sp. (Plate XLVIII. fig. 5.)
M. testa angusta, solida, antice leviter contracta, carneola, rufo-
fusco maculata et flammulata, ubique longitudinaliter costata ;
costis numerosis, subconferlis, planulutis, interstitiis profunde
punetatis ; spira elongata ; anfractibus 9, planato-convezis ;
sutura impressa; apertura oblonga, intus costata, carneola; colu-
mella 5-plicata.
Long. 20, maj. lat. 5, spira alt. 10 mill.
A single specimen without locality, but very strikingly different
from every known species.
ZIZIPHINUS JUCUNDUS, nl. sp. (Plate XLVIII. fig. 6.)
Z. testa acute sublate conica, tenui, fulvo-carneola, lineis fuscis
minutis confertis interruptis spiraliter cingulata, versus apicem
cinereo-fasciata ; anfractibus 10, vix convexis, primis grano-lira-
tis, sequentibus levibus, ultimo juxta suturam iterum subconspicue
grano-lirato, liris deinde fere obsoletis ; sutura impressa ; mar-
ginibus obtuse carinatis, maculis rubidis subdistantibus ornatis ;
basi planato-convexa ; apertura oblique quadrata, fauce margari-
tacea ; columella callosa, obliqua, vie intorta.
Alt. 30, maj. diam. 33 mill.
Hab. Japan.
A very neatly formed and delicately marked shell, of which only
one specimen is at present known.
EXPLANATION OF PLATE XLVIII.
Fig. 1. Conus melviili, p. 795. Figs. 7, 8. Mitra acuta, p. 797.
2. carnalis, p. 796. 9, 10. fulvolirata, p. 797.
3. catenatus, p. 796. ll. berthe, p. 797.
4. Latirus cayohuesonicus, p. 796. 12. deburghie, p. 798.
5. Mitra puncturata, p. 798. 13. Hemifusus corona, var.
6. Ziziphinus jucundus, p. 798. | minor, p. 796.
1878.] MR. BUTLER ON A NEW SPIDER FROM MADAGASCAR. 799
3. Description of a Remarkable new Spider from Mada-
gascar. By Artuur G. Burier, F.L.S., F.Z.S., &e.
[Received July 24, 1878.]
Amongst a series of insects of various orders recently collected
in Madagascar by the Rev. William Deans Cowan, I found two
Epeirids which especially interested me. The first of these I at
once recognized as a fine example of the Epeira tuberculosa of Vin-
son (Aran. de Réunion, Maurice, et Madagascar, pl. xiv. He 2)
specimens of which were previously in the collection of the British
Museum.
The second spider struck me as so extraordinary that I wrote to
the Rev. O. P. Cambridge respecting it. He tells me that it isa
Cerostris, allied to C. mitralis, Vinson (loc. cit. pl. ix. figs. 2-4),
but it appears to be perfectly distinct. “Epeira tuberculosa”? be-
longs also to Cerostris.
Fig. 1. Cwrostris avernalis (enlarged). Fig. 2. Profile of abdomen.
Fig. 3. View of abdomen from behind.
Ca@RoOsTRIS AVERNALIS, 0. sp.
Cephalothorax and falces dark castaneous, clothed at the sides
and behind with whity-brown pile; palpi and legs above bright
reddish castaneous, hairy at the sides, the femora cylindrical, with
blackish lateral hairs, the tibiee and tarsi flattened and longitudinally
bisuleate, with whitish lateral hairs; tibie and tarsi below with
their basal or distal extremities broadly belted with whitish, and their
proximal extremities blackish ; abdomen dark clay-coloured, covered
(excepting the tubercles and odontoid processes) with whitish pile ;
centre of ventral surface blackish.
Falces large, rugulose, subcylindrical ; palpi broad, flattened, bi-
800 - LIEUT.-COL. R. H. BEDDOME ON [Nov. 5,
sulcated, hairy at the sides; cephalothorax broader than long, with
a small central projection in front, so that its anterior margin is
——-shaped; a broad gradually formed transverse ridge almost
uniting the three oculiferous tubercles; the central oculiferous
tubercle forming a transverse oval, with a central longitudinal de-
pression, its highest point just within the eyes, and the uppermost
surface of the eyes themselves flattened (as if filed smooth), the an-
terior pair of eyes much nearer to each other than the posterior
pair ; lateral oculiferous tubercles standing out obliquely from the
antero-lateral angles of the cephalothorax, the eyes much smaller
than those of the central group ; a long cylindrical process emitted
from each side of the cephalothorax at its postero-lateral angles and
parallel to the lateral oculiferous tubercles; two widely separated
conical tubercles standing perpendicularly upon the posterior berder
of the cephalothorax: abdomen irregularly subscutiform, with two
terminal tubercles, between which it is sinuated; anterior margin
with six conical tubercles, followed by a large unequally bidentate
process on each side, these processes being widely divergent and
oblique; a very minute tubercle on a bare spot between them ;
second superior abdominal fold with an acute conical tubercle on
each side; pectoral shield heptagonal, its margins swollen.
Length 16 millims., including cephalothorax, of abdomen alone
12, width at widest part 15; length of lateral odontoid processes 6,
width from tip to tip of processes 15; relative length of legs 1, 2,4, 3.
Fianarantsoa, Madagascar. Type in Coll. B. M.
Readily distingnished from C. mitralis by the much more promi-
nent, more unequally bidentate, and far more divergent odontoid
processes on the abdomen, the almost entire absence of the large
central conical tubercle, the different arrangement of the tubercles
on the margins of the abdomen, and the absence of bands on the
upper surface of the legs.
4, Description of six new Species of Snakes of the Genus
Silybura, Family Uropeltide, from the Peninsula of India.
By Lieut.-Colonel R. H. Beppome, C.M.Z.S.
[Received August 2, 1878. ]
SILYBURA.
Sect. 1. Scales in nineteen rows.
* Caudal disk flat.
SILYBURA BROUGHAMI, 0. Sp.
Scales in 19 rows round middle of body, and neck; ventrals 224.
Rostral large, high and sharp, nearly but not quite separating the
nasals. Caudal disk flat, the scales prominently 3-5-keeled;
terminal scute slightly bicuspid, the points side by side. Subcaudals
6. Colour brown, with a series of transverse wavy black blotches
1878.] NEW INDIAN SNAKES. 801
in which are often ocellated white spots; sides with yellow blotches :
abdominals dark brown.
Hab. Sirumullay hills (Madura district), 5500 feet elevation.
SILYBURA LEVINGII, 0. sp.
Scales in 19 rowsround body, ventrals196. Rostralsharply pointed;
nasals not quite separated by the rostral. Caudal disk very flat, the
scales prominently 3-10-keeled; subcaudals 8-9. Upper parts
purplish-brown, with narrow transverse black bands, in which are
arranged yellowish white spots; sides with large yellow blotches ;
belly purplish brown, the blotches of the sides sometimes extending
more or less across, particularly in the posterior portion. Tail with
several yellow blotches on each side.
This species has much the coloration of S. melanogaster.
Hab. Lower Puluey hills (Madura district), 4000 feet elevation.
Sect. 2. Scales in seventeen rows.
A. Number of scutes exceeding 160.
b. Caudal disk convex.
SILYBURA OCHRACEA, 0, sp.
Seales in 17 rows (neck and body), ventrals 225-231, rarely only
214. Rostral obtusely conical, moderate; nasals forming a suture
behind the rostral; caudal distinct, more or less convex, the scales
3-8-keeled. Terminal scute much broader than long, ending in two
points side by side; anal bifid ; subcaudals 6—8-10. Back yellowish
green in life (yellowish-brown in spirits), in the young and half-grown
purplish brown, with irregular cross bands of ocellated spots, which
are yellow with a black ring; sides and belly yellow, but the latter
much marbled and blotched irregularly with the ground-colour ; tail
black beneath, surrounded with a bright yellow band; tail and head
blackish olive. Length of a full-grown specimen 20 inches.
Hab. Anamullays, at Nelliamputty, 3000 feet, aud at Ponachi,
4500 feet, and on the Bolumputty hills, 2000 feet.
SILYBURA DUPENI, 0. sp.
Scales in 17 rows (19 round neck) ; ventrals 233. Snout obtuse ;
nasals not separated by the rostral; caudal disk more or less convex,
the scales rough, with 3-7 keels; terminal scute very sharply
bicuspid and very rugose, the points side by side; subcaudals 10.
Back uniform brown; belly and sides (2-3 outer rows of scales)
yellow.
Hab. Anamullays at Nelliamputty, 3000 feet.
This may be only a variety of the last ; but it has quite a different
coloration.
SILYBURA GUENTHERI, 0. sp.
Scales in 17 rows, ventrals 168. Rostral obtuse; nasals entirely
separated by the rostral; no supraorbital ; caudal disk convex ; tail
laterally compressed (as in Plectrurus); the scales perfectly smooth ;
802 MR. R. B. SHARPE ON P@OPTERA LUGUBRIS. [Nov. 5,
the terminal scute hard, yellow, and almost square at the end, the
inconspicuous points being side by side, not superposed ; subcaudals
13. Back uniform black nacreous, belly with two rows of the scales
on each side of the ventrals yellow; chin and the anterior first inch
or so of the belly black; tail black beneath, but with a yellow band
on each side; upper labials yellow.
Hab. “High Wavy” mountain, Madura district, 5000 feet eleva-
tion.
B. Number of scutes less than 160.
a. Caudal disk flat; snout obtuse.
SILYBURA MADURENSIS, 0. Sp.
Scales in 17 rows, abdominals 142-149. Rostral very small,
obtusely conical; nasals large, forming a suture behind the rostral ;
vertical distinctly six-sided ; caudal disk flat, each scale with 1—2 very
prominent keels ; terminal seute sharply bicuspid, the points side by
side ; subcaudals 7-8. Back black, very iridescent, each scale girt
with a narrow white rim; belly orange-yellow, with very irregular
broad black markings, which sometimes meet and form cross bars.
Hab. “ High Wavy”? mountain, Madura district, elevation 5500
feet.
5. A Note on Peoptera lugubris, Bp.
By R. Bowpier Suarpre, F.L.S., F.Z.S.
[Received August 15, 1878.]
(Plate XLIX.)
This species was first described by Prince Bonaparte, in the
‘Comptes Rendus’ for 1854, as Paoptera lugubris, as he thought it
was the Abyssinian Muscicapa lugubris of Miiller (Beitr. Orn. Afr.
Taf. ii.) ; and it was on account of this confusion of names that Dr.
Briiggemann wrote his note in the ‘ Annals of Natural History’ for
1877 (xx. p. 244), and suggested that the species should be called
Peoptera eryptopyrrha (Cabanis). When he states, however, that the
species had no name of its own until 1876, he is mistaken ; for the
title ‘ /ugubris, Bonap.’’ was adopted for the species by Dr. Hart-
laub (Orn. Westafr. p. 69), which was sufficient to fix the specific
name. But on reading Bonaparte’s description again, I find that I
missed the point of it, and Dr. Briiggemann does not appear to have
consulted the original description, having been incited to write his
notice probably by the footnote to the genus Peoptera in my
‘Catalogue,’ wherein I hinted that Bonaparte had so confused his
species with Miiller’s Muscicapa lugubris that the title ought per-
haps to be rejected. I am sorry to have led Dr. Briggemann wrong,
for I had overlooked the fact that when Bonaparte described the
Peoptera he added :—“ Nous appellerons l’espece lugudris, qu elle
JGKeulemans lith
Hamhart 1 mp
PQ@OPTERA LUGUBRIS.
E:
=
)
Tee rita
ths
'eee,
1878. ] MR. R. B. SHARPE ON P@OPTERA LUGUBRIS. 803
soit ou non la dugubris, Miiller;” and I am glad that I followed
Hartlaub, and did not alter the name, as I had felt inclined to do.
Dr. Briiggemann was also mistaken in supposing that he was the
first to inform me of the probable identity of the species described
by Cabanis with Bonaparte’s Peoptera lugubris; for I had finished
my synonymy of this bird for my ‘Catalogue’ long before the
casual conversation to which he alludes.
After all, however, it appears that both Dr. Briiggemann and
myself were wrong in our facts, and Myiopsar eryptopyrrhus of
Cabanis is no¢ synonymous with Peoptera lugubris of Bonaparte.
Dr. Peters having very kindly sent me from Berlin some of the new
species of birds lately described by Drs, Cabanis and Reichenow for
examination, has added the type of Myiopsar, and asked me to com-
pare it with our examples of Pawoptera lugubris. I am surprised to
find it specifically distinct; and so the subjoined modification must
take place in the synonymy of the genus as given by me in the
‘Catalogue.’
Genus Paorrerra.
Paoptera, Bp. Comptes Rendus, xxxviii. p. 381 Type.
eS (nn oi ee Se P, lugubris.
Myiopsar, Cabanis, J. f. O. 1876, p. 93 ...... P. eryptopyrrha.
Range. West Africa, from the Gold Coast to the Congo.
Key to the Species.
a, Purple; edges of secondaries whity brown, form-
ing a light band down the wing ; inner webs of
quills dark brown, scarcely lighter than the
G iTV SYS.) Uo ere anc oh 55- cendeoe De SO ABRs EME aces EEC Ae P. lugubris.
6. Violet-blue with somewhat of a greenish gloss ;
no whity-brown edges to the wing; quills
rufous on the inner web, forming a red lining
RGRLMO WAL. sccaatentanie ess + va.ncusmuasbébce ce aptcekes P. eryptopyrrha.
1. PaworreraA tucusRis. (Plate XLIX.)
Peoptera lugubris, Bp. C. R. xxxviii. p. 381 ; Hartl. Orn. W.-Afr.
p- 69 (1857) ; Sharpe, Cat. B. ii. p. 281 (1877).
Thamnobia lugubris, Gray, Hand-l. B. i. p. 212 (1869).
Peoptera erptopyrrha, Briggs. Ann. N. H. (4) xx. p. 244 (1877,
nec Cabanis).
Hab. From the Gold Coast to Gaboon.
Since the publication of the Catalogue, I have discovered in the
British Museum a third specimen, a male, put away by mistake
along with the species of Thamnobia, according to Gray’s classifi-
eation. It was only discovered this month (August 1878), during a
re-arrangement of the specimens of Chats.
2. P&oprera CRYPTOPYRRHA.
Myjiopsar eryptopyrrhus, Cab. J. f. O. 1876, p. 93.
Adult male (type of species).—General colour above glossy
violet blue, with here and there a slight shade of steel-green; least
804 MR. E, A. SMITH ON MARINE SHELLS [Nov. 5,
and median wing-coverts like the back, the remainder, as well as the
quills, dark brown, with slightly paler edges ; the primaries rufous for
the greater portion of their length, this red colour concealed when
the wing is shut; tail-feathers black, edged with dull violet-blue ;
sides of face and entire under surface ot body glossy violet-blue, a
little duller than the upper surface; under wing-coverts like the
breast ; quills dusky brown below, the inner webs broadly rufous for
a great portion of their extent. Total length 7°5 inches, culmen
0-6, wing 3°5, tail 3°3, tarsus 0°75. (Mus. Berol.)
Hab. Congo district.
6. Ou a Collection of Marine Shells from the Andaman
Islands. By Encar A. Suitu, F.Z.8., Zoological De-
-partment, British Museum.
[Received August 16, 1878.]
(Plate L.)
The shells treated upon in this paper form part of a series
recently presented to the British Museum by Capt. L. Worthington
Wilimer, by whom they were dredged whilst stationed at the Andaman
Islands. All, with one or two exceptions, were obtained at the depth
of a few fathoms off Port Blair. The collection contains about half
a dozen new forms, the most interesting of which is a remarkable
shell which I have provisionally located in the genus Fusus.
After each species I have quoted the locality first assigned to it, and
others have been added on the authority of specimens in the Museum,
in order to give at a glance the known geographical distribution,
which is always interesting, and calculated to assist materially in the
identification of the various forms.
1, Conus ANDAMANENSIS. (Plate L. figs. 1, 1 a.)
Shell subcylindrical, with somewhat convex lateral outlines, pale
pinkish white, marked irregularly with small brown spots and lines,
and covered with a very thin, smooth, greyish epidermis: spire
elevated concave, composed of ten whorls; the two nuclear ones
subglobose, semidiaphanous, smooth, the four following slightly
turreted, and the rest obliquely planulated, spirally sculptured with
two to three fine revolving striae; the markings on the spire are in
the form of short brown lines following a radiating direction (that is,
across the whorls): the last whorl has the upper angle rather
obtuse, and is sulcated at the base with about twelve transverse
grooves. Aperture rather narrowed above, and a little widened to-
wards the base, white within. Lip thin, with a small sinuation at
the upper extremity. Length 22 mill., diam. 11.
This pretty little species, which I am unable to place as the young
state of any larger form, is remarkable on account of the fine brown
dots or lines, which are irregularly scattered over its surface of a
PZ.54878: Pris
22%
Mintern Bros. imp.
SHELLS OF THE ANDAMAN ISLANDS.
1878. ] FROM THE ANDAMAN ISLANDS. 805
delicate pale pink. The spire too is peculiar, on account of the few
first whorls being suddenly elevated into a little prominent cone.
Since writing the above description I have seen a larger specimen
from the collection of Dr. Hungerford. It measures 31 millims. in
length and 15 in diameter; and the markings are rather larger,
certain spots a little above the middle of the last whorl taking the
form of a transverse interrupted band (fig. 1 a).
2. Conus ceyLanensis, Hwass.
Hab. Red Sea, Ceylon, Rodriguez Island ; varieties at Sandwich
Islands, Lord Hood’s Island, &e.
3. PievrRotoma (Driti1a) vartasiuis, Smith, Annals & Ma
Nat. Hist. 1877, vol. xix. p. 495. (Plate L. figs. 2 & 3.)
There is a non-adult variety of this species, which does not ex-
hibit at this stage of growth any indication of the slight umbilical
rimation which is characteristic of the mature shell. It is whitish
dotted with brown beneath the suture, and stained with a darker tint
above it in the spaces between the short coste ; and the spiral lire
are also dotted with brown. The body-whorl is encircled around
the middle by a broad brownish band, which is darker at the edges,
and below this it is entirely white.
s=
4. Prevroroma (Driniia) wiemert. (Plate L, fig. 4.)
Shell subfusiform, not rimate, white, dotted with brown beneath
the sutural line, and with a series of brown dots or (in other words)
an interrupted line around the middle, the dots being situated just
above the interspaces between the nodules, which form a series a
little below the middle of the whorls. The lower half of the body-
whorl dark brown; and a spot of the same colour stains the shell at
the superior sinus; the apex is also brown. Whorls ten, broadly
concave above, and somewhat acutely noduled below; nodules eleven
in number on the penultimate whorl, the last volution transversely
striated, the basal strize being stronger than elsewhere ; aperture
rather small, white, stained with dark brown at the short basal canal ;
labrum thin, arcuate when viewed laterally, deeply and broadly
sinuated at the suture, with a second small sinus or indentation near
the base; columella but little curved or sinuous, covered with a thin
brown callosity, terminating above in a large white tubercle. Length
16 mill., diam. 6.
This shell is very distinguishable from its congeners, on account of
the peculiarity of its painting, and the series of sharpish tubercles
encircling the whorls. I have much pleasure in naming: it after
Capt. Wilmer, its discoverer,
5. TEREBRA AFFINIS, Gray.
Hab. Philippines, Fiji, Tahiti.
6. TEREBRA Ex1GuA, Deshayes.
Hab. East Australia.
It is interesting to obtain authentic examples of this species from
Proc. Zoot. Soc.—1878, No. LILI. 5S
806 MR. E, A. SMITH ON MARINE SHELLS [Nov. 5,
the Andaman Islands, as the type shells were described as Kast-
Australian. The specimen before me, which is apparently adult, has
a length of 21 millims., and the last whorl is 4 in diameter. The
coste (about 18 on a whorl) are thickest at the upper extremity,
above the sulcus which parts off the infrasutural belt, gradually
attenuating downwards, and at length become quite obsolete at the
extreme base of the body-whorl.
7. Murex (MuricipgA) crrrosvus, Hinds.
Hab. Straits of Macassar (Hinds).
Some small specimens of this charming shell were dredged by
Capt. Wilmer, one of which possesses an unusually long canal.
8. Murex (Muricipea) Rusticus, Reeve. (Plate L. fig. 5.)
Hab. ?
I feel uncertain whether the Andamanese shell is without doubt the
same species as that figured by Reeve. It agrees very well with the
description, but exhibits some difference from the figure, which is
very coarsely executed. The number of transverse ridges is greater,
the mouth a little smaller ; and the fronds, although a little broken
and worn, would not I think, if perfect, be as long as those repre-
sented in the figure.
9. Murex (MuricrpEa) BARCLAYANA, H. Adams.
Coralliophila barclayana, H. Ad. P. Z. 8. 1873, p. 205, pl. 23.
fig. 1.
Hab. Mauritius (H. dd.).
The Andamanese specimen of this species, the type of which is
now in the British Museum, is only 21 millims. in length, yet appa-
rently full-grown. Its coloration is much less brilliant than the
representation of the described shell, being pale pinkish white, with
the lateral varix and the termination of the canal tinged with brown,
the aperture being light purple.
10. Murex (OcINEBRA) BREVICULUS, Sowerby, Proc. Zool. Soc.
1840, p. 146; Conchol. Illustr. fig. 37.
M. tetragonus, Reeve (non Broderip), Conch. Icon. iu. fig. 118.
Hab. - ?
A single, much worn, yet undoubted example of this species is
valuable as indicating the habitat of so interesting a form. Reeve
considers it a short variety of Broderip’s M. tetragonus, the
figure he gives of the latter representing a typical breviculus. M.
tetragonus proper is well figured by Sowerby in his ‘ Conchological
Illustrations,’ fig. 25. Judging from the specimens of the two species
which I have examined, all of them unfortunately in a more or less
worn and faded condition, I am of opinion that they are specifically
distinct. Besides the much stouter and less-produced form of MZ.
breviculus, its aperture is larger and more circular, white or slightly
blotched with brown far within; the transverse coste, especially
-on the obtuse varices, are also brown. M. tetragonus, on the other
1878. ] FROM THE ANDAMAN ISLANDS. 807
hand, is white, or very pale purple, and the aperture, the interior of the
basal channel, and the columella, are of a uniform light purple. It
is true, however, that the sculpture in both is remarkably similar.
11. CotumBetta (Anacuis) niGRIcostara. (Plate L. fig. 6.)
Shell acuminately ovate, subturreted ; beneath a thin dirty yellowish
epidermis, white, with black ribs, with a series of white spots on the
cost a little above the middle of the last whorl, and with a black
elongate spot or line in each of the interstices between the ribs, those
towards the labrum being produced downwards to the base in a
rather zigzag manner. Whorls 8, a little convex, with about 12
thinnish ribs on each. Body-whorl a little flattened or even concave
at its broadest part; the ribs in front are produced somewhat below
the middle ; and on the back four or five nearest the lip are developed
only a short distance from the suture, leaving the whorl below them
smooth. At the base it is transversely grooved, the eight or nine
ridges or lire between the sulci being spotted with black. These
spots are the continuation of the colouring of the longitudinal costz.
Aperture small, narrow, bluish white within; labrum exteriorly
thickened, thin at the edge and bearing about 8 tubercles within, of
which the three or four upper ones are the largest ; columella sub-
erect, slightly convex at the middle, covered with a thin callosity,
with a free margin, through which the transverse ridges which wind
round the end or cauda of the whorl appear somewhat in the form
of nodules. Length 124 mill., diam. 6.
This species is well characterized by its coloration, the black cost
contrasting conspicuously with the pale interstices. It is consider-
ably like C. terpsichore, Sowerby, in form and sculpture, but different
in colour.
12. ConumBEeLLA (ATILIA) PuELLA, Sowerby ; Reeve, Conch.
Icon. xi. sp. 65.
Hab. Philippines.
The plicz on the back of the body-whorl are almost obsolete ; but
immediately behind the labrum three are well developed and situated
very close together, thus forming a very strong postlabral varix.
The denticulation within the aperture consists of about eight short
fine lire. The columella is coated with a thickish enamel, and is
armed with about seven fine, short, transverse lirations.
13. ENGINA AsTRICTA, Reeve-
Ricinula astricta, Reeve, Conch. Icon. iii. fig. 31.
Hab. ? (Ree).
14. Nassa ELEGANS, Kiener; Reeve, Conch. Icon. viii. pl. 2.
fig. 10 (not good) ; Kiener, Coq. Viv. pl. 24. fig. 97.
Hab. Indian Ocean.
15. NassA CRENULATA, Bruguiére; Reeve, Conch. Icon. sp. 2;
Kiener, Coq. Viv. pl. 23. fig. 90.
Hab. Philippines.
53*
808 MR. E. A. SMITH ON MARINE SHELLS [Nov. 5,
16. Nassa Tanta, Gmelin, Syst. Nat. p.3493; Knorr, Vergnig.
vy. pl. 10. fig. 3.
N. olivacea, Bruguiére, Reeve, fig. 19; Kiener, pl. 15. fig. 53.
Hab. “West Indies’? (Lamarck and Reeve) ; Ceylon and Chili
(Kiener).
“West Indies,” which is cited by some authors as the home of this
species, is doubtless erroneous; for it is scarcely possible that any species
of Mollusca living on the shore between tide-marks could be found
inhabiting such remote localities as the Andaman Islands and the
West Indies. It was collected by Cuming on mud banks in the
Bay of Manilla; and the specimens obtained by Capt. Wilmer were
found on the mud in mangrove-swamps.
17. Nassa LuripDA, Gould, Proc. Boston Soc. Nat. Hist. 1849,
vol. iii. p. 153; Atlas, Wilkes’s Explor. Exped. pl. 19. fig. 325 a, ¢ ;
Otia, p. 68.
N. dispar, A, Adams, P. Z. 8. 1851, p. 96; Reeve, Conch. Ic, fig.45.
Hab. Samoa Islands (Gould) ; Philippines (Reeve).
18. Nassa BrrartA, Baird, Cruise of the Curagoa, p. 436, pl. 38.
figs. 1, 2. (Plate L. fig. 7.)
Hab. New Caledonia
The type of this species was brought from New Caledonia. The
Andaman specimens differ from that form in a few respects: the
plicze on the body-whorl are more numerous and more developed
towards the base; and the transverse sulci on the central portion are
generally more pronounced.
19, NassA MONILE, Kiener, Coq. Viv. pl. 11. fig. 40; Reeve,
Conch. Ie. fig. 38.
Hab. Philippines.
20. NASSA MARGARITIFERA, Dunker.
Buceinum margaritiferum, Dkr., Philippi’s Abbild. vol. iii. pl. 1.
fig. 12. (Non Nassa margaritifera, Dkr., Reeve, figs. 59 a, 6.)
N. costellifera, A. Adams, P. Z. 8. 1851, p.113; Reeve, figs.58 a, 6.
Hab. Rocks below Ross Island ; Port Blair (Wilmer).
The shell figured by Reeve as margaritifera of Dunker certainly
is distinct from that species, and may possibly be a closely ribbed
variety of Nassa morginulata, Lamarck.
21. Nassa stiemartA, A. Adams, P. Z.8. 1851, p. 96; Reeve,
Conch. Icon. sp. 42.
Hab. “Dredged in deep water, Port Blair’ (Wilmer). Other
localities are Japan, Philippines, and New Caledonia.
The specimens from the Andaman Islands are very much smaller
than that figured by Reeve, having only a length of 10 to 13 millims.
In all other respects they agree very well, varying one from another
1878. j FROM THE ANDAMAN ISLANDS. 809
in the length of the spire, the proximity of the series of granules, and
the size of them.
A peculiarity of this species consists in the manner in which some
of the yellowish tubercules are as it were set in an irregular darker-
coloured ring, or, in other words, are surrounded by a dark line at
their base.
22. Nassa GEMMULIFERA, A. Adams, P. Z. S. 1851, p. 99 ; Reeve,
Conch. Icon. viii. figs. 132 a, b.
Hab. Philippines (Cuming).
23. Nassa atsescens, Dunker.
Buccinum albescens, Philippi’s Abbild. neuer Conch. iii. pl. 2. fig.
15.
Nassa albescens, Reeve, Conch. Icon. viii. fig. 100.
Hab. West Indies (Dunker and Reeve) ; Sandwich Islands.
This species is undoubtedly an East- and not West-Indian species.
It has been found at the Sandwich Islands by Pease, and also at other
localities in the Pacific.
24. Nassa (ARcULARIA) GLOBosA, Quoy and Gaimard.
Buccinum globosum, Q. and G. Voy. Astrolabe, pl. 32. figs. 25-27.
Nassa globosa, Reeve, Conch. Icon. viii. figs. 62 a, 6.
Hab. Vanikoro Island and New Ireland (Quoy).
25. Nassa MARRATHU, Smith, Journ. Linn. Soe. xii. p. 543, pl. 30.
fig. 4, (Plate L. fig. 8.)
Hab. Solomon Islands.
This species I described from specimens collected at the Solomon
Islands. The shells from the Andaman Islands are a trifle larger,
having a length of 19 millims; with the exception of this difference,
they agree perfectly with the typical shells. In the above-quoted
figure of this species the form of the aperture is not at all correctly
drawn. The columella has not such a long curve; and the labrum
should be more expanded at basal end.
26. Nassa (Hira) sisrroipea, G. and H. Nevill, Journ. Asiatic
Soc. Bengal, 1874, xliii. part 2, pl. 1. fig. 6.
Hab. “Found in one spot below the rocks near the barracks, Ross
Island, very local” (Wilmer). ‘
In their description of this interesting species, Messrs. Nevill do
not remark upon the curious way in which the coste on the body-
whorl are trinodose through the prominence of some of the transverse
lire at intervals. The uppermost row of nodulous projections borders
the depression or elevation at the top of the whorl ; the median series
is situated about the middle, and the lowermost alittle below that.
This I take to be a specific character, as it is present in all the adult
specimens which I have seen, and the above-named figure of their type
also exhibits this peculiarity. The whitish band which is seen within
the aperture runs between the uppermost and median series of nodules.
810 MR. E. A. SMITH ON MARINE SHELLS [Nov. 9,
Besides the two or three “‘plaits,’”” which perhaps should rather be
termed elongate tubercles, at the base of the columella, there is a
small one at the upper part, usually present in most species of Nassa.
N. trinodosa of Smith has the same nodulation on the last whorl,
but, however, is quite distinct from this species. It differs in colour,
has fewer longitudinal cost, which are less granulous ; and the spiral
thread-like liree which cross them are fewer and less prominent.
N. subspinosa of Lamarck is another allied species of much the
same form, and having the same character of sculpture; only the
nodules are much more prominent and the ribs less numerous.
27. Nassa (ARCULARIA) GRANIFERA, Kiener, Coq. Viv. pl. 27.
fig. 111; Reeve, Conch. Ie. viii. fig. 72.
Hab. Philippines (Reeve).
28. Nassa (Arcutaria) Brmacutosa, A. Ad. P.Z.S. 1851,
p- 102; Reeve, Conch. Icon. viii. sp. 61.
Hab. Philippines (Cuming) ; ‘Andaman Islands, on sand banks
at low tide ; very active animal” (Wilmer).
29. Nassa (ArcuLARIA) caLuospirA, A. Ad. P.Z.S. 1851, p.
102; Reeve, Conch. Icon. viii. figs. 66 a, 6; G. and H. Nevill, Journ.
Asiatic Soc. Bengal, 1874, vol. xliii. pt. 2, pl. figs. 5, 5 a.
Hab. Andaman Islands (Nevill and Wilmzr); Philippines (Cu-
ming).
On comparing an Andaman specimen with the type of the species,
I do not hesitate for a moment with regard to its identity.
30. Nassa HoRRIDA Dunker.
Buceinum horridum, Dkr., Philippi’s Abbild. ueuer Conch. iii. pl.
2. fig. 8.
Nassa horrida, Dkr., Reeve, Conch. Icon, viii. figs. 69 a, 6.
N. curta, Gould, Otia, p. 69; Atlas, Wilkes’s Explor. Exped. pl.
19. figs. 326 a, 0.
Hab. Samoa Islands (Gould).
31. Nassa ecuinaTA, A. Adams, P. Z. 8. 1851, p. 101; Reeve,
Conch. Icon. viii. fig. 131. (Plate L. fig. 9.)
Hab. Philippines.
The Andaman specimens are all shorter than the type, which is
figured by Reeve ; and not one of them has the fifth or lowermost series
of nodules on the body-whorl, which is very closely approximated to
the fourth row. In other respects they agree perfectly with the typical
shells said to have been found at the Philippine Islands. This species
is allied to N. muricata of Quoy and Gaimard, yet is perfectly distinct.
It differs from that species in being almost quite smooth in the inter-
stices between the nodules, there being only the faintest indication of
spiral strize in some specimens. ‘The aperture, too, is less elongate,
and the margin of the basal canal is stained with dark brown. This
1878. | FROM THE ANDAMAN ISLANDS. 811
is very characteristic, and, although not referred to by Reeve or Adams,
is constant in all the specimens which I have seen.
The colour is whitish, with the oblique longitudinal cost usually
pale brown; and the tips of the rather acute nodules are white.
Nassa gruneri, Dunker, is another closely allied form, yet readily dis-
tinguished by its colour and sculpture.
32. Pos senticosuM, Linn.
Hab. Philippines.
This beautiful and well-known species varies very considerably in
colour, from white with dark transverse bands to uniform rich purple-
brown, with some of the spiral thread-like lirze yellow.
33. PHos rosEATUM, Hinds.
Hab. Philippines, Moluccas, &e.
34. Puos Textum, Gmel.
Buceinum blainvillei, Deshayes.
B. cancellatum, Quoy & Gaimard.
Phos cyanostoma, Sowb. (not of A. Ad.), Thes. fig. 46.
Hab. Philippines.
The shell figured by Sowerby as P. cyanostoma of A. Adams is
a slight variety of P. ¢ewtum, differing in having a somewhat elongated
spire and the body-whorl a trifle shorter than usual, and constricted
just above the raised ridge which encircles the extreme base or cauda
of the whorl. The painting is precisely the same in both forms. The
true cyanostoma (described in the Proc. Zool. Soc. 1850, p. 155) is
very like this variety in form and sculpture, but is distinguished by
the margins of the columella and labrum and the tip of the body-
whorl being stained with violet.
35. Fusus? apnormis. (Plate L. fig, 10.)
Shell imperforate, oblong, turreted, pale yellowish-brown ; whorls
9, very convex, divided by a deep channelled suture, obliquely plicated ;
plicze or costz rounded, swollen, gradually increasing in size from the
apex downwards and becoming less prominent, crossed by numerous
slender finely-scaled liree, which are alternately coarser and finer,
numbering about 15 on the penultimate and about 40 on the last
whorl; aperture small, irregularly pyriform, about one third as long as
the entire shell, dirty white within, with the outer lip simple, with 10
fine thread-like lirze, which extend within as far as can be observed,
but do not attain quite to the edge of the lip: basal canal short,
narrow, oblique, and a little recurved. Length 23 mill., diam. 113.
This species is totally unique in form and sculpture ; and it is with
considerable doubt that I have placed it among the species of usus.
It also exhibits some affinity with Coralliophila, which genus, as
generally accepted, contains a group of species which are usually
inhabitants of Madrepores, and are characterized by the white or
purplish colour, and the scaly nature of the transverse ridges, and
the shortish open channel at the base of the aperture, which is
812 MR. E. A. SMITH ON MARINE SHELLS [Nov. 5,
lirate within. The present species is remarkable for the great
convexity of the whorls, and the fineness of the spiral scaly liree.
The aperture also is unusually short in proportion to the whole length ;
and the liree within are very fine, and produced as far as the eye can
trace inwards.
36. SisTRUM MARGARITICOLA, Bruguiére.
Hab. Singapore ; Lord Hood’s Island ; Natal.
37. Latirus pecoratus, A. Adams, P.Z.S. 1854, p. 316;
Kuster, Con. Cab. Ad. 2, pl. 25. figs. 12, 13 (as Turbinella).
(Plate L. fig. 11.)
Hab. New Zealand (Adams).
The Andaman variety of this species has the Jongitudinal coste
much thicker, and consequently fewer in a whorl, than the New-
Zealand form ; and the colour isa little different. The general tint is
whitish with a blush of rose, the two granulous lirze beneath the
suture light brown, interstices between the coste dark brown, the
body-whorl with a rose-coloured band bordered on each side by a
white liration around the middle, the tip of the canal blackish, the
columella and outer lip pinkish, and the apex of the spire pink. The
colour of this species (“alba castaneo varie picta’’) given by Adams
does not at all describe the variety of painting. There are two speci-
mens in the Cumingian collection, both pinkish white with a rosy
apex and a pinkish aperture; and one, probably the type, has the dark
brown tip to thecanal. The shell figured in the Conchylien-Cabinet
is larger than the type or the Andaman specimens. The former is
18 mill. long, 17 in diameter, and the largest of the latter 20 in length
and 8 in width.
38. Larirus rasticiuM (Reeve). (Plate L. fig. 12.)
Turbinella fastigium, Reeve, Conch. Ic. iv. figs. 72 a, 6.
Hab. Ceylon.
Of this species the British Museum possesses three specimens—
one from Ceylon, the type (the habitat of which is unknown), and the
third presented by Capt. Wilmer, from the Andaman Islands. Two of
them have six longitudinal cost, whilst the type has only five. The
latter is an immature shell; and consequently the figure of it in the
‘Conchologia Iconica’ does not convey a true idea of the aperture in
the adult. When perfect the canal is a little longer and the columella
is covered with a callosity, the edge of which is produced, and at the
base of the shell forms an umbilical fissure.
Reeve does not mention that one or two of the spiral liree just be-
neath the suture are conspicuously scabrous; nor does he ‘point out
that 4 or 5 of them around the cauda of the body-whorl are also
scabrous and thicker than the rest, which are frequently alternately
coarser and finer. The plaits on the columella are somewhat in-
distinct and about five in number ; and the aperture is finely lirated
risen the liree terminating at some distance from the margin of the
abrum.
1878.] FROM THE ANDAMAN ISLANDS. 813
39. Latrrus INCARNATUS (Deshayes), var.
Turbinella incarnatus, Desh., Kiener, Coq. Viv. pl. 18. fig. 3;
Reeve, Conch. Icon. iv. fig. 55.
Hab. Philippines; Australia.
The shell from the Andaman Islands, which I consider a variety of
this species, is light brown and darker between the ribs; the apex of
the spire is pinkish; and the body-whorl has two spiral whitish or
yellowish lirze, one above and the other below the middle of it, the
former being visible on the penultimate and antepenultimate whorls.
The aperture and columella are pinkish, the outer lip finely lirate
within ; and the columella has one or two rather indistinct plice at
the base.
40. Mirra masta, Reeve, Conch. Icon. ii. sp. 323. (Plate L.
fig. 13.)
Hab. Philippine Islands.
Shell fusiform, turreted, olive-brown, subrimate. Whorls 10 or 11,
shortly tabulated above, but very slightly convex at the sides, spirally
grooved with three narrow subequidistant sulci, the uppermost one
at the top of the whorls, dividing off a thickened edge to the tabula-
tion, which is channelled ; this thickened belt is of a brighter brown
colour than the rest of the shell. Last whorl transversely sulcated
throughout; sulci about 11 in number; a few of the superior ones,
with the exception of the, uppermost, more or less obsolete; those
below the middle broader and deeper, leaving raised ridges or lire
between them. Aperture narrow, brownish within, a little less in
length than half the entire shell ; columella furnished with a thickish
callosity, with a raised free edge forming in the umbilical region a
slight fissure; plaits 4, the uppermost thickest, the others in turn
by degrees finer; basal channel and tip of the whorl dark brown.
Length 19 mill., diam. 63.
This species is remarkable on account of the turreted character of
the spire. The tabulation of the whorls, however, is narrow and
shallowly channelled. The Andaman specimen differs from the
type in having the whorls more distinctly turreted ; the spiral sulci
are not so pronounced ; and the colour is somewhat different. In the
type (which is a worn shell), the upper part of the whorls is white,
whilst in the specimen above described it is light brown.
41. Mirra maria, A. Adams, Proc. Zool. Soc. 1851, p. 140.
(Plate L. fig. 14.)
Hab. Eastern Seas.
In describing this species Adams writes, “ labro intus sulcato.”’
I have carefully examined the type specimens in Cuming’s collection
and also the Andamese shell, and do not find a trace of sulcation
within the labrum of any of them, and therefore conclude that the
observation of Mr. Adams was incorrect.
The body-whorl has two broad purplish-brown bands indistinctly
separated by a faint light zone occupying the greater part of the whorl.
Beneath the suture there is a greyish-white band. The entire surface
814 MR. E. A. SMITH ON MARINE SHELLS _ [Nov. 5,
is variegated with fine white zigzag lines and small brownish spots,
principally on the upper edges of the transverse liree. The superior
margin of the upper purplish-brown band is interrupted by opaque
white blotches edged on one side by dark brown. In the Monograph
of this genus (Thesaurus Conchy]l. iv.) Sowerby cosiders this species
a variety of M. incisa, Ad. & Reeve. However, I think there are
sufficient differences to separate them. The form of the latter is
narrower, the whorls proportionally higher, the transverse ridges on
the spire flatter, and the spire itself is less acutely conical.
42. Mirra (Canciuia) ANNULATA, Reeve, Conch. Icon. ii. sp.
103.
M. nitens, Kiener, Coq. Viv. pl. 29. fig. 96.
Hab. Zanzibar (Reeve).
The plaits on the columella are not invariably five in number, some
specimens being furnished with a sixth. The shells from the Anda-
man Islands have less of the brown markings than appear in Reeve’s
figure, and in this respect resemble more the specimen delineated by
Kiener.
43. Mirra (CanciLia) PHILIPPINARUM, A. Adams, P.Z.S.
1851, p. 141.
M. flammea, Reeve (not of Quoy & Gaimard), Conch. Icon. ii.
fic. 120.
Hab. Philippine Islands.
This species, when in fine condition, has the transverse costze prettily
marked with small, close-set, olive-brownish dots. ‘These are not
mentioned by Reeve or Adams, owing to the fact that the specimens in
Cuming’s collection which they described are somewhat worn or over-
cleaned ; however, they are traceable when closely searched for.
44. Mirra (CuHrYsAME) TABANULA, Lamk., Kiener, Coq. Viv.
pl. 9. fig. 27; Voy. Astrolabe, pl. 45 bis, figs. 10-13 ; Reeve, Conch.
Tcon. ii. fig. 332.
Hab. Philippines (Reeve) ; Vanikoro (Quoy).
The specimen of this species from the Andamans is rather small,
and of a uniform deep brown colour. The three plaits on the colu-
mella are whitish, the aperture olivaceous ; and the outer lip has
three or four conspicuous tubercular teeth at a short distance from
the upper extremity.
45. Mirra (Cytinpra) unputosa, Reeve, Conch. Icon. species
193, fig. 192.
Hab. Philippines (Cuming).
A very short stunted variety was obtained by Capt. Wilmer. It
has eight plaits on the columella, whereof the uppermost is much
thicker than the rest.
46, Mirra (CauurTHEA) acupicta, Reeve, Conch. Icon. fig. 76.
The Andaman shell agrees precisely in form with the type from
Zanzibar ; but the longitudinal coste are less numerous.
1878.] FROM THE ANDAMAN ISLANDS. 815
47. Mirra (CaLuirHEa) oBexiscus, Reeve, Conch. Icon. ii.
fig. 107.
A small specimen, not quite adult, from the Andaman Islands,
agrees with those from the Philippines in all respects, but has the
second white band below the middle of the body-whorl more con-
spicuous than usual.
48. Mirra (CosreLiaria) EXASPERATA, Chemnitz; Reeve,
Conch. Icon. fig. 162.
M. torulosa, Lamk., Kiener, Coq. Viv. pl. 25. fig. 77.
Hab. Java; Philippines.
49. Mirra (CosTeELLARIA) pEsHayest1, Reeve, Conch. Icon.
fig. 170.
Hab. New Caledonia (Mus. Cuming).
Capt. Wilmer tells me that the orange-scarlet markings on this
pretty species fade considerably after death. One of the specimens
he collected is 25 millims. long. The mouth is purplish black
within, with a single white zone ; and the lip is white within at the
margin.
50. RANELLA PusILLA, Broderip, P. Z. S. 1832, p. 194; Sowerby,
Conchol. Illustr. fig. 1*, fig. 1a, var.; Reeve, Conch. Icon. ii. figs. 44
a, 6 (both varieties). (Plate L. fig. 15).
Triton laciniatum, Mighels, fide Pease, Amer. Journ. Conch. iv.
. 107.
: Var. =R. rosea, Reeve, /. c. fig. 46.
Var. = Bursa concinna, Dkr. P. Z.S. 1862, p. 239.
Hab. Lord Heod’s Island and Philippine Islands ; Red Sea.
R. rosea, Reeve, and concinna, Dkr., which I consider varieties
of this species, are distinguished from the normal form almost solely
on account of a difference in colour. If this were tenable, the Anda-
man shell would constitute a third species ; for it certainly varies in
this respect either from R. pusilla, concinna, or rosea. The
principal colour is reddish brown; the lowermost series of granules on
the upper whorls, the fourth and fifth series from the top of the
body-whorl, and the second rib from the suture on each varix, on all
the whorls, are yellow. The columella is purple, with three or four
transverse whitish nodules, and within the outer lip near its edge there
are about eight white bead-like granules on a dark-purple ground.
The interior of the aperture is purple-brown, with the exterior yellow
zone visible.
The normal form of the species described by Broderip is white in
every part. In the Museum series of nearly 40 specimens there are
various intermediate and connecting gradations between this and the
pink form (rosea) or the brown variety (concinna). In fact,
the differences are so subtle, that I feel it impossible to draw a line
of distinction any where between these varieties. The difference in
specimens apparently adult is very striking. The largest example of
816_ MR. E. A. SMITH ON MARINE SHELLS [Nov. 5,
the variety (rosea) measures 22 millims. in length, whilst a very
small one is only 9 millims. long.
The tuberculation is also subject to considerable variation. The
number of the transverse series of granules appears to be very con-
stant, there being invariably three principal rows in the upper whorls,
and seven in the body-whorl, but the size and number of the gra-
nules differ in various examples; and this difference is not peculiar
to each form, but exists in shells belonging to the same variety.
Reeve describes the tubercle-like nodules on the angle of rosea as
bipartite. This is scarcely correct ; for there certainly are two series
at this point; and being situated rather closely together, they might
be considered single nodules subdivided.
51. Trrron (StmpuLUM) sTRANGEI, A. Adams & Angas, P. Z.S8.
1864, p. 35. (Plate L. fig. 16.)
Hab. Moreton Bay, East Australia.
The whorls in this species are seven in number, and not four as
described by the authors of it. The four apical or nuclear volutions
are smooth, convex, and horn-colour ; the remaining three are some-
what concave beneath the suture, of a pale brown colour; and the
varix on the labrum between the spiral ridge is stained with bright
red, and the canal with brown.
The transverse costz are described by Adams and Angas as being
trisulcate. This, however, is not the case; for there are only two
sulci, making each of the ridges trilirate, the central liration being
most prominent. The coste are prettily undulated, owing to their
being prominent at intervals where they-cross the longitudinal sub-
plications ; there are two of them round the middle of ‘the penulti-
mate whorl, and the one preceding it, and five on the last. The aper-
ture is white, and the columella furnished with a tubercle at both ex-
tremities ; the Andaman specimen, besides these, has a few interme-
diate ones. The labrum is strongly varicose, and has six coarse
lirations within.
52. ToRINIA PERSPECTIVIUNCULUS, var. (Plate L. figs. 17—
17 b.)
Hab. Red Sea, Seychelles Islands, Philippines, Pacific Islands, &e.
The Andaman and Seychelles specimens of this widely distributed
and very diversely shaped species are very depressed in form, similar
to the variety (planulata) figured by Hanley in Sowerby’s ‘The-
saurus Conchyliorum,’ vol. ii. pl. 254. fig. 63.
The operculum is very elongate-conical, having a marked resem-
blance in form to a small Triphoris.
53. CypR#%A GANGR2ZNOSA, Solander.
Hab. Zanzibar, Mozambique.
54. Lirorina unpuLata, Gray, Philippi’s Abbild. i. pl. 5.
figs. 17, 18; Reeve, Conch. Icon. figs. 67 b-d (? fig. 67a).
Hab. North Australia, Philippines, and Society Islands.
1878.] FROM THE ANDAMAN ISLANDS. 817
55. Lirorina ventricosa, Philippi, Abbild. iii. pl. 6. fig. 19 ;
Reeve, Conch. Icon. fig. 93.
Hab. Pulo Penang.
L. nove:-zealandie, Reeve, and L. melanacme, Smith, have a some-
what similar ventricose form, but differ in colour and the absence
of the granulation on the transverse ridges between the sulci.
56. Ceriraium (Vertacus) Kocut, Philippi, Abbild. iii. pi. Ts
fig. 3; Conch. Icon. figs. 26 a, 6. (Plate L. fig. 18.)
Hab. East Africa, Philippines, Japan.
The Andaman specimens are small, but agree very closely in de-
tail of sculpture with more typical ones, yet differ rather in colour.
They are pale horn-colour, with the eranules, varices, and a band at
the top of the whorls whitish. Behind the varices they are blotched
with dark brown; and their entire surface is irregularly dotted and
spotted with brown, and the transverse thread-like striz are also of
the same colour ; the liration below the middle of the body-whorl,
which is more prominent than the rest, is white, ornamented with
brown dots.
57. CerirH1um (VERTAGUS) TURRITUM, Sowerby, Thesaurus,
ii. pl. 180. fig. 101; Reeve, Conch. Icon. xv. fig. 58 ; Smith, Annals
& Mag. Nat. Hist. 1875, xvi. p. 106. (Plate L. fig. 19.)
Hab. Philippines, Japan.
The shells in the present collection are in colour and sculpture
exactly like those which I described in the Annals and Mag. Nat.
Hist., from Japan, but a trifle smaller.
58. TripHoris scutptus, Hinds, Voy. Sulphur, pl. 8. fig. 3.
Hab. Straits of Malacca (Hinds).
59. TURRITELLA INFRACONSTRICTA. (Plate L. fig. 20.)
Shell subulate, subturreted, brownish white, ornamented with
countless minute reddish-brown dots. Whorls about 20, somewhat
convex at the top and sides, and slopingly narrowed at the base, thus
making the width at the suture considerably less then a little distance
above it, transversely finely lirated ; lire varying in thickness, a few
being a trifle coarser than the rest, two especially—the upper, situated
very near the middle of the whorl, and the lower one at the broadest
part of it, being conspicuous and forming indistinct angles; they are
all minutely dotted with brown. Last whorl excavated beneath,
finely lirated, and of a uniform brownish white or pale pink tint,
with a rather acute ridge circumscribing the base. Aperture irregu-
larly obliquely quadrate, of the same colour as the exterior ; but the
flattened base is coated with a thin pale rose-coloured enamel. Colu-
mella slightly oblique and a little arcuate. Length 50 millims.,
diam. 12.
This species, like most others in the genus, is chiefly distinguished
by the form of the whorl, the character of the markings, and the
spiral liration. The dottings on the ridges are very small, and are
818 MR. E. A. SMITH ON MARINE SHELLS [Nov. 5,
arranged under one another in oblique and more or less flexuous lines.
In other words, the whorls are ornamented with fine oblique or
flexuous lines, which are interrupted by numerous spiral sulci, thus
producing series of dots. Through the prominence, although slight,
of the two principal lire, the whorls between them appear somewhat
concave, and the ridge around the base of the last whorl passes
just above the labrum and winds up the spire at the sutural line.
60. AMALTHEA AUSTRALIS, Lamarck.
Hab. Kangaroo Island, Australia, Philippines.
61. Guaprus (RimELLA) CANCELLATUs, Lamarck ; Kiener, Coq.
Viv. (Rostellaria), pl. 3. f. 3.
Hab. “ Chili and Moluccas ”’ (Kiener).
62. XENOPHORA SOLARIOIDES, Reeve.
Phorus solarioides, Reeve, Conch. Icon. i. fig. 8.
Hab. Philippine Islands.
ess 63. Trocaus (CLANcuLUS) MiIcRopon, A. Adams, P. Z.S.
1851, p. 162. (Plate L. figs. 21-210.)
Hab. ig
This species, the habitat of which was not hitherto known, is not
very fully described by Adams. The colour is subject to consider-
able variation. The series of granules are usually lightish brown,
and the interstices almost black. This simplicity of colour is de-
stroyed by broad, irregular, white patches radiating from the suture ;
and many of the granules, especially on the base, are white. Two
or three of the series of the interstices between them around the
umbilicus are white, only a few of the granules being brown. The
whorls are six in number and rather convex.
64. Trocuus (MoniLea) CaLycuLvus, Wood, Index Test. Suppl.
pl. 6. f. 44; Adams, Gen. Rec. Moll. iii. pl. 49. figs. 3-38.
Trochus (Monilea) masoni, G. & H. Nevill, Journal Asiat. Soc.
Bengal, 1874, vol. xliii. part 2, pl. 1. figs. 1, la.
Hab. Eastern Seas (A. Adams, P. Z. 8. 1851).
On comparing Andaman specimens of this species with the type
presented to the British Museum by the late Dr. Gray, I have not
the slightest doubt of their specific identity.
65. Trocuus (ForsKAaLIA) PULCHERRIMUS, A. Adams, Gen.
Rec. Moll. i. p. 432. (Plate L. figs. 22, 22a.)
Gibbula pulcherrima, A. Ad. P. Z.8. 1854, p. 39.
Hab, China Sea.
The colour of this beautiful species varies in some examples, but
appears to be constant in its distribution. The oblique arcuate
pink or scarlet stripes on the upper part of the body-whorl, which
in some specimens are black, generally terminate a little above the
middle. At this part the whorl is encircled by two close-set gra-
1878. ] FROM THE ANDAMAN ISLANDS. 819
nulous liree, with a furrow on each side, whereof the lower is the
more conspicuous. Immediately below this the colour of the inter-
rupted scarlet stripes changes to black, and they assume the form of
somewhat quadrate spots, but towards the aperture become more
flame-like and merge into scarlet.
66. Hatrotis varia, Linné.
H. varia, Linn., teste Reeve, Conch. Icon. ili. sp. 4.
Hab. Philippines.
67. Atyscytinprica, Helbling; Sowerby, Thesaurus, ii. pl. 125.
f. 114.
Var. = Bulla solida, Brug. Ency. Méth. pl. 360. f. 2; Thesaurus,
pl. 124. figs. 112, 113.
Var.=Atys elongata, A. Ad.; Sowerby, Thesaurus, ii. pl. 125.
fig. 121.
Hab. Philippines, Ceylon.
68. Carprum uysTrix, Reeve, Conch. Icon. ii. figs. 40 a, 6.
Hab. Philippines (Cuming).
The Andaman specimen of this lovely shell belongs to Reeve’s var.
B, being pure white, with the exception of a pale tint of pink on the
posterior side near the ligament.
69. Carpium (L&vicARDIUM) AUSTRALE, Sowerby, Proc. Zool.
Soe. 1840, p. 105 ; Conch. Illust. figs. 12, 12* (overcoloured) ; Reeve,
Conch. Icon. ii. fig. 97.
C. pulchrum, Reeve, Conch. Icon. fig. 98.
Hab. Philippines, China, Ceylon.
The differences of form, sculpture, and colouring which Reeve con-
sidered sufficient to distinguish his C. pulehrum from C. australe are, in
my opinion, hardly sufficient to constitute a variety. The form in
almost every species of Cardium is subject to considerable variation,
and likewise the colour. Reeve does not mention that the anterior
dorsal area in his type is blotched with purple at the extremity, as
is the case in C. australe. The sculpture of both forms is identical :
both have the groove on the posterior side, which, within the valves,
forms a raised oblique ridge ; but in some specimens it is more con-
spicuous than in others. The Andaman examples are rather young
and bear the remains of their epidermis, which takes the form of very
thin, short, irregular yellowish lamellz arising from the radiating
strie. Their colour is whitish or yellow, marked with subcencentric
irregular yellowish brown bands, which are more purple-brown within
the valves and towards the umbones. The latter at the tips and the
hinge-margin are purple ; and within, a line of the same colour extends
from the umbo to about the middle of the valves.
70. CorBULA FORTISULCATA. (Plate L. figs. 23-236.)
Shell very inequivalve, inequilateral, somewhat triangular, dirty
white, solid, rounded at the posterior extremity and beaked anteriorly :
820 ON MARINE SHELLS FROM THE ANDAMAN ISLANDs. [Nov. 5,
upper valve slightly carinate from the umbo to the posterior margin,
the keel terminating a little above the middle, rather smooth, finely
concentrically striated by the lines of growth and suleated at the umbo;
covered with a thin greyish epidermis, a trifle laminated towards the
margin ; and from the umbo about four rather indistinct ridges radiate
down the posterior half of the valve, becoming obsolete about the
middle of it, and again conspicuous in some specimens at the margin :
lower valve vastly larger than the other, very deeply and widely con-
centrically sulcated; sulci fine and close together at the umbo;
ridges between the’sulci very thick and prominent, thicker anteriorly
than at the opposite extremity, and terminated in front by a sulcus or
depression extending from the umbo downwards at a little distance
from the margin, which is raised, taking the form of a stout marginal
ridge which terminates the beaked end of the valve; transverse
grooves finely striated in the same direction : one tooth in each valve,
that of the upper bifurcate; pallial sinus small; anterior scar sub-
rotund, posterior more elongate. Diam. 14 mill., length 10, thick-
ness 84; the lower valve of another larger specimen is 233 mill. in
diameter and 16 long.
For many years several lower valves of this species have been in
the British Museum with the locality ‘‘ Jack’s Island, South Sea,”
attached to them. Others were presented by J. B. Jukes, Esq., in
1845, who dredged them at Port Essington in 7 fathoms, from a
muddy bottom ; and two specimens in the Cumingian Collection are
from the same place. The form, the coarse costze (which, however,
are sometimes rather close together), and the anterior raised and
beaked margin of the lower valve well distinguish the species.
71. CRASSATELLA RADIATA, Sowerby; Reeve, Conch. Icon.i. fig. 12.
Hab. Singapore.
72. Arca (ScapHarca) Myrist1ca, Reeve, Conch. Icon. ii. sp. 42.
Hab. Philippine Islands (Cuming).
73. PECTEN ALBOLINEATUS, Sowerby, Thesaurus Conch. pl. 14.
figs. 69, 70.
Hab. Philippine Islands.
74. IsoGNOMON SAMOENSIS, Baird.
Perna (Isognomon) samoensis, Baird, ‘Cruise of the Curagoa,’
p- 454, pl. 42. fig. 8.
Hab. Tutuila, Samoa Islands.
BRACAIOPODA.
75. LinGuLa HIANS, Swainson ; Sowerby, Thesaurus, i. pl. 67.
fig. 4.
Hab. China.
Capt. Wilmer’s notes on this species are as follows :—
“It lives in mud or sandy clay at low-water mark, the shell being
buried about a foot from the surface. It is very easily alarmed, and
P Z.S.1878.PP1
Mintern Broj?
Yantern del.
LEPIDOPTERA FROM UPPER TENASSERIM.
{
-_,
P Z. 3.1878. Pl t
|
\
|
R.Mintern del.
LEPIDOPTERA FROM UPPER TENASSERIM.
BR Mantern del
LEPIDOPTERA FROM UPPER TENASSERIM.
1878. } MR. F. MOORE ON LEPIDOPTEROUS INSECTS. 821
retreats rapidly downwards. In order to collect specimens, I
searched for oval orifices in the mud ; and having found one, a spade
was plunged very deeply in and the mud turned up, the hands then
being used to go deeper; yet, still, in many cases, the creature was
too quick for the diggers.’’
EXPLANATION OF PLATE L.
Fig. 1, la. Conus andamanensis, p- 804.
2. Pleurotoma (Drillia) variabilis, p. 805.
3. (——) variabilis, var. p. 805.
4. (——) wilmeri, p. 805.
5. Murex (Muricidea) rusticus, p. 806.
6. Columbella ( Anachis) nigricestata, p. 807.
7. Nassa bifaria, p. 808.
8. marratii, p. 809.
a: echinata, p. 810.
10. Fusus abnormis, p. 811.
11. Latirus decoratus, p. 812.
Fastigium, p. 812.
13. Mitra mesta, p. 813.
: marié, p. 813.
15. Ranella pusilla, var., p. 815.
16. Triton (Simpulwm) strangei, p. 816.
17. Torinia perspectiviunculus, front view, p. 816.
17a. Ditto, basal view.
176. Ditto, operculum.
18. Cerithium (Vertagus) kochi, p. 817.
19. —— (——) turritum, p. 817.
20. Turritella infraconstricta, p. 817.
21. Trochus (Clanculus) microdon, p. 818.
21a. Ditto, variety.
21 4. Ditto, base.
22. Trochus (Forskihlia) pulcherrimus, p. 818,
22a. Ditto, base.
23. Corbula fortisulcata, upper view, p, 819.
23a. Ditto, lower valve.
23 6. Ditto, top view.
7. A List of the Lepidopterous Insects collected by Mr.
Ossian Limborg in Upper Tenasserim, with Descriptions
of new Species. By F. Moors, F.Z.S., Assistant Cura-
tor, India Museum, London.
[Received September 17, 1878.]
(Plates LI—LIII.)
This list comprises the Lepidopterous portion of the collection
made on and around the Moulayet range in Upper Tenasserim by
Mr. Limborg, who was accompanied by Mr. Wood-Mason’s native
collector, Motiram. The party proceeded, about the middle of
December 1876, from Moulmein up the Houngduran, collecting as
they went along to the Hills, on and around which, and along the
banks of the river leading to them, the whole collection was made.
Proc. Zoou. Soc.—1878, No. LIV. 54
822 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
The expedition was organized and the funds provided by the
Marquis of Tweeddale, President of the Zoological Society, Mr. W.
T. Blanford, Lt.-Col. Godwin-Austen, Sir E. Clive Bayley, Presi-
dent, and Mr. J. Wood-Mason, Natural-History Secretary of the
Asiatic Society, Calcutta, Mr. R. Lydekker, Dr. J. Anderson, Di-
rector of the Indian Museum, Calcutta, and others.
PAPILIONES.
Fam. NYMPHALID.
Subfam. DANAINz.
DANAIS SEPTENTRIONIS.
Danais septentrionis, Butler, Ent. Mo. Mag. xi. p. 163 (1874).
Ahsown ; Meetan, 3000 feet; Hatsiega ; Houngduran source.
DaNais AGLEA.
Danais aglea, Cramer, Pap. Exot. iv. tab. 377. f. E (1782).
Ahsown; Hatsiega; Naththoung to Paboga; Moolai, 3000-
6000 feet.
DaANAts VULGARIS.
Danais vulgaris, Butler, Ent. Mo. Mag. xi. p. 164 (1874).
Ahsown; Moulmein to Meetan ; Houngduran source.
DANAIS MELANEUS.
Papilio melaneus, Cram. Pap. Exot. i. tab. 30. f. D (1775).
Ahsown; Moulmein to Meetan.
DANAIS TYTIA.
Danais tytia, Gray, Lep. Ins. Nepal, p. 9, pl. 9. f. 2 (1846).
Moolai to Moolat, 4500 feet.
DANAIS PLEXIPPUS.
Papilio plexippus, Linn. Syst. Nat. i. 2, p. 767 (1767).
Ahsown; above Ahsown ; Moulmein to Meetan; Meetan, 3000
feet ; Taoo, 3000-5000 feet.
DANAIS CHRYSIPPUS.
Papilio chrysippus, Linn, Syst. Nat, i. 2, p. 767 (1767).
Ahsown.
SALPINX RHADAMANTHUS.
Papilio rhadamanthus, Fabr. Ent. Syst. iii. 1, p. 42 (1793).
Ahsown, 2000 feet; Meetan, 3000 feet; Hatsiega; Houngduran
source; Naththoung to Paboga.
SALPINX CRASSA.
Euplea crassa, Butler, Proc. Zool. Soc. 1866, p. 278.
Ahsown; Moulmein to Meetan; Hatsiega; Naththoung to Paboga;
Taoo, 3000-5000 feet.
1878. ] INSECTS FROM UPPER TENASSERIM. 823
SALPINX MASONI, 0. sp.
Male. Allied to S. crassa, Butler (Proc. Zool. Soc. 1866, p. 278),
but differing in its darker colour, in the basal area of the fore wing
being bright glossy blue, and the marginal spots on this wing con-
fined more to the apex; hind wing less convex along the exterior
margin ; otherwise similarly marked.
Expanse 3 inches.
Taoo, 3000-5000 feet ; above Ahsown. In coll. Wood-Mason.
From S. klugii, Moore, this species may be distinguished by the
blue gloss being confined to the basal area, whereas in S. Alugii it is
more brilliant and suffuses the entire wing.
SALPINX MARGARITA.
Euploea margarita, Butler, Proc. Zool. Soc. 1866, p. 279.
Hatsiega; Houngduran source.
TREPSICHROIS MIDAMUS.
Papilio midamus, Linn. Syst. Nat. i. 2. p. 765 (1767).
Ahsown, 2000 feet; Moulmein to Meetan; Hatsiega; Houng-
duran; Naththoung to Paboga.
CrastTia CUPREIPENNIS, Nl. Sp.
Allied to C. modesta, Butler, P. Z. S. 1866, p. 273, from Siam.
Upperside cupreous brown, the outer borders broadly paler, basal
area of fore wing blue-glossed; cell and contiguous area of hind
wing also blue-glossed ; the two marginal series of spots larger.
Expanse 33 inches.
Hatsiega; Houngduran source. In coll. Wood-Mason.
EvurLa@a LimporGil, n. sp. (Plate LI. fig. 2.)
Male. Differs from EF. deione, Westw., in the fore wing having
some very indistinct submarginal white spots, and the hind wing
having two marginal rows of prominent white spots similar to but
smaller than those in #. margarita, Butler.
Expanse 3? inches.
Ahsown, 2000 feet; above Ahsown; Hatsiega; Houngduran,
In coll. J. Wood-Mason.
EupPLa@a ALCATHOE.
Luplea aleathoé, Godt. Enc. Méth. ix. p. 178 (1819).
Ahsown, 2000 feet.
EUPLG@A SUBDITA, N. sp.
Allied to #. core, Cram. Pap. Exot. pl. 266. figs. E, F.
Male. Fore wings shorter, broader, and the posterior margin more
convex : upperside paler, basal area slightly blue, glossed in the fore
wing ; markings similar, smaller and paler, those on the fore wing
indistinct.
Expanse 3} inches.
Hatsiega. In coll. J. Wood-Mason.
54*
824 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
EuPL@a GODARTII.
Euplea godartii, Lucas, Rev. Zool. 1853, p. 319.
Ahsown; Moulmein to Meetan; Hatsiega; Houngduran ; Nath-
thoung to Paboga.
STICTOPL@A GROTEI.
Euplea grotei, Felder, Reise Nov., Lep. ii. p. 339, tab. 49.
‘An
Ahsown ; above Ahsown; Hatsiega to Houngduran.
Subfam. SaTyRInz&.
LETHE ARCADIA.
Papilio arcadia, Cram. Pap. Exot. ii. tab. 116. f. E, F (1779).
Ahsown.
LETHE MEKARA.
Debis mekara, Moore, Catal. Lep. Mus. E.I. C. i. p. 219 (1857 )
Ahsown, 2000 feet; Taoo, 3000-5000 feet.
LETHE ROHRIA.
Papilio rohria, Fabr. Mant. Ins. ii. p. 45 (1787).
Hatsiega; Naththoung to Paboga; Taoo, 3000-5000 feet.
LETHE VERMA.
Satyrus verma, Kollar, Hiigel’s Kasch. iv. p. 447, tab. 16. f. 1,
(1844).
Moolai, 3000-6000 feet.
MELANITIS ISMENE.
Papilio ismene, Cram. Pap. Exot. i. tab. 26. f. A, B (1775).
Ahsown, 2000 feet; Moulmein to Meetan; Meetan, 3000 feet ;
Naththoung to Paboga; Moolai, 3000-5000 feet.
MELANITIS BELA.
Melanitis bela, Moore, Catal. Lep. Mus. E.I. C. i. p. 223 (1857).
Ahsown; Meetan, 3000 feet, April; Naththoung to Paboga ;
Moolai, 3000-6000 feet ; Moolai to Moolat, 4500 feet.
ZETHERA DIADEMOIDES, n. sp. (Plate LI. fig. 3.)
Male and female. Upperside dark brown: fore wing with a sub-
marginal series of seven small bluish-white spots, which decrease in
size to the costa; a marginal series of smaller less distinct spots, two
between each vein : hind wing with a series of six large broad oval
spots; a submarginal series of small reversely triangular spots, and
a marginal series of smaller narrow spots, the two latter series with
two of these spots disposed between the veins. Underside as above.
Expanse, ¢ 3, Q 3% inches.
Taoo, 3000-5000 feet, March ; Moulai, 3000-6000. In coll. J.
Wood-Mason.
1878. ] INSECTS FROM UPPER TENASSERIM. 825
MYCALESIS MINEUS.
Papilio mineus, Linn. Syst. Nat. i. 2, p. 768 (1767).
Moulmein to Meetan; Meetan, 3000 feet; Taoo, 3000-5000
feet, March.
MYycALESIS PERSEUS.
Papilio perseus, Fabr. Syst. Eut. p. 488 (1775); Butler, Proc.
Zool. Soc. 1867, p. 719, fig. 2.
Moulmein to Meetan.
MYCALESIS BLASIUS.
Papilio blasius, Fabr. Ent. Syst. Suppl. p. 426 (1798) ; Butler,
Proc. Zool. Soc. 1867, p. 718, fig. 4.
Moulmein to Meetan.
MYCALESIS ANAXIAS.
Mycalesis anaxias, Hewitson, Exot. Butt. iii. Myc. pl. 4. f. 25,
26 (1862).
Ahsown; Moolai, 3000-6000 feet.
MycALEsIS RUNEKA.
Mycalesis runeka, Moore, Catal. Lep. E.I. C. i. p. 234 (1857).
Ahsown; Moulmein to Meetan; Hatsiega; Naththoung to
Paboga.
CuLaPa, 0. g.
Differs in form from Mycalesis, having a greater length of fore
wing, less convexity of the costa, the apex being more produced and
the exterior margin more oblique ; the hind wing is somewhat acutely
angled at the apex, and the exterior margin produced before the anal
end, thus giving this wing a quadrate appearance. In form it more
resembles the American genus Anchiphlebia.
CULAPA MNASICLES.
Mycalesis mnasicles, Hewits. Exot. Butt. iti. Myc. pl. 5. f. 32,
33 (1864).
Ahsown, 2000 feet; Meetan, 3000 feet, April.
YPHTHIMA BALDUS.
Papilio baldus, Fabr. Syst. Ent. App. p. 809 (1775); Donov.
Ins. Ind. t. 36. f. 2.
Moulmein to Meetan.
YPHTHIMA METHORA.
Yphthima methora, Hewitson, Trans. Ent. Soc. 1865, p. 291,
pl. 18. f. 20, 21.
Naththoung to Paboga.
ERITES ANGULARIS, N. Sp.
Male. Wings longer than in 2. madura, Horsf. (from Java) ;
826 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
the fore wing being more produced at the apex and the hind wing
posteriorly. Underside similarly marked, excepting that the yellow-
bordered black spot on the fore wing is more oval, and the spots on
the hind wing smaller, this wing also having the inner yellow band
acutely angled in the middle.
Expanse 23 inches.
Taoo, 3000-5000 feet. In coll. J. Wood-Mason.
ELYMNIAS TINCTORIA, 0. sp.
Allied to £. undularis, of India.
Male. Upperside dark blue-black, the marginal blue spots on
fore wing larger; border of hind wing dusky purple, with a series of
small white spots. Female with prominent blue-black borders and
white markings.
Expanse 3 inches.
Meetan, 3000 feet, April; Moolai, 3000-6000 feet. In coll. J.
Wood-Mason.
ELYMNIAS VASUDEVA.
Elymnias vasudeva, Moore, Catal. Lep. E.I. C. i. p. 238 (1857).
Taoo, 3000-5000 feet.
Dycris LEUCOCYMA.
Biblis leucocyma, Godt. Enc. Méth. ix. p. 326 (1819).
Ahsown.
Subfam. NyMPHALIN2E.
AMATHUSIA PHIDIPPUS.
Papilio phidippus, Linn. Syst. Nat. i. 2, p. 752 (1767); Cramer,
Pap. Exot. i. tab. 69. f. A, B.
Meetan, 3000 feet, April.
ZEUXIDIA MASONI, 0. sp.
Allied to Z. amethystus, Butler, P. Z. S. 1865, p. 485, from
Sumatra.
Female. Differs in the paler colour of the wings, and in the
greater breadth of the yellow oblique band, the band entire and ter-
minating at the middle median branch, below which are two similar-
coloured spots; a small pale patch before apex of the wing; hind
wing pale cinnamon-brown broadly along outer border.
Expanse 43 inches.
Meetan, 3000 feet, April. In coll. J. Wood-Mason.
DIsCOPHORA TULLIA.
Papilio tullia, Cramer, Pap. Exot. i. tab. 81. f. A, B (1779).
Above Ahsown.
DiIscoPHORA ZAL.
_ Discophora zal, Westw. Gen. D. Lep. p. 331 (1851); Trans.
Ent. Soc. 1858, p. 188, pl. 21. f. 5, 6.
Taoo, 3000-5000 feet ; Moolai, 3000-6000 feet.
1878. ] INSECTS FROM UPPER TENASSERIM. 827
DiscOPHORA NECHO.
Discophora necho, Felder, Reise Nov., Lep. iii. p. 462 (1867).
D. celinde, Horsf. Catal. Lep. E.1. C. pl. 6. f. 6.
Houngduran source.
CLEROME ARCESILAUs.
Papilio arcesilaus, Fabr. Mant. Ins. ii. p. 28 (1787); Donov.
Ins. Ind. pl. 30. f. 2.
Above Ahsown.
JEMONA LENA.
Aimona lena, Atkinson, Proc. Zool. Soc. 1871, p. 215, pl. 12.
1
Moolai, 3000-6000 feet.
XANTHOTANIA BUSIRIS.
Xanthotenia busiris, Westw. Trans. Ent. Soc. 1858, p. 187.
Moolai, 3000-6000 feet.
THAUMANTIS LOUISA.
Thaumantis louisa, Wood-Mason, Proc. As. Soe. Beng. 1877,
p- 163; id. Journ, As. Soc. Beng. 1878, pl. xii.
Taoo, 3000-6000 feet.
Allied to 7. howqua, but differs in having the upper surface white
instead of fulvous, and in having five red spots instead of ocelli on
the underside of the fore wings, and only two well-developed ocelli
(the anterior and posterior) on the hind wings, instead of three and
five ocelli respectively.
CIRROCHROA SURYA, D. Sp.
Allied to C. lanka (Moore, P. Z.S. 1872, p. 557) of Ceylon.
Male differs above on the fore wing in the narrower marginal band,
a single sinuous line extending with fulvous interspaces to near the
apex ; hind wing with the submarginal line more sharply sinuous ;
other markings similar. Underside purplish fulvous grey; the
discal transverse band broader, and pale bluish purple on the fore
wing, bluish purple-white on hind wing, its outer border being dark
blue, and its inner border red and waved; other markings red.
Female darker above, the discal black line sinuous at the costal end;
a double sinuous marginal line; other markings and underside as in
male.
Expanse, ¢ 14 to 22, 2 24 inches.
Moulmein to Meetan; Taoo, March, 3000-5000 feet; Moolai,
3000-6000 feet. In coll. J. Wood-Mason.
MESSARAS ERYMANTHIS.
Papilio erymanthis, Drury, Ill. Exot. Ent. i. pl. 15. f. 3, 4
(1778).
Moolai to Moolat; Hatsiega.
828 MR. F. MOORE ON LEPIDOPTEROUS [ Nov. 5,
ATELLA SINHA.
Terinos sinha, Kollar, Hiigel’s Kasch. iv. p. 438 (1844).
A melanoid variety. Moulmein.
JUNONIA LAOMEDIA.
Papilio laomedia, Linn. Syst. Nat. i. 2, p. 772 (1767); Drury,
Il. Exot. Ent. pl. 5. f. 3.
Ahsown, 2000 feet ; Moulmein to Meetan; Naththoung to Paboga.
JUNONIA LEMONIAS.
Papilio lemonias, Linn. Syst. Nat. i. 2, p. 770 (1767).
Ahsown, 2000 feet; Moulmein to Meetan; Meetan, 3000 feet ;
Taoo, 3000-6000 feet (March) ; Naththoung to Paboga.
JUNONIA ORITHYIA.
Papilio orithyia, Lion. Syst. Nat. i. 2, p. 770 (1767); Cram.
Pap. Exot. i. tab. 19. f. C, D, tab. 32. f. BE, F.
Moulmein to Meetan.
JUNONIA ALMANA.
Papilio almana, Linn. Syst. Nat. i. 2, p. 769 (1767); Cram.
Pap. Exot. i. tab. 58. f. F, G.
Ahsown, 2000 feet; Moulmein to Meetan; Taoo, 3000-5000
feet.
PRECIs IPHITA.
Papilio iphita, Cram. Pap. Exot. iii. tab. 209. f. C, D.
Moolai, 3000-6000 feet.
DOLESCHALLIA PRATIPA.
Doleschallia pratipa, Felder, Wien. ent. Mon. iv. p. 399 (1860) ;
Reise Nov., Lep. ii. p. 206.
Hatsiega.
KALLIMA LIMBORGII, 0. sp.
Differs from K. inachis, from Sikkim. Male and female of a
deeper and more uniform steel-blue colour above; the fore wing is
not angled on the hindward part of the exterior margin; the apex
more produced in the male and less so in the female, the discal
hyaline spot small, and almost circular; the hind wing also is less
convex on the exterior margin, and has a much shorter tail.
Expanse, ¢ 32, 2 4 inches.
Moolai, 3000-6000 feet. In coll. J. Wood-Mason.
CyYRESTIS THYODAMAS.
Cyrestis thyodamas, Boisd. Cuv. Reg. Anim. Ins. ii. t. 138. f. 4
(1836).
Amathusia ganescha, Kollar, Hiigel’s Kasch. iv. p. 430, tab, 7.
foBAe
Moolai, 3000-6000 feet.
1878. ] INSECTS FROM UPPER TENASSERIM. 829
CYRESTIS COCLEs.
Papilio cocles, Fabr. Mant. Ins. ii. p. 7 (1787) ; Donov. Ins. Ind.
pl. 23. f. 2.
Hatsiega.
CYRESTIS RISA.
Papilio risa, Doubleday, Gen. D. Lep. pl. 32. f. 4 (1850).
Moulmein to Meetan; Moolai, 3000-6000 feet.
HERONA ANGUSTATA, 0. Sp.
Female. Similar to H. marathus from the Khasia hills. Differs
from same sex of that species in having all the maculated bands
narrower, these being but half the width of those in the former
species.
Expanse 3 inches.
Moolai, 3000-5000 feet. In coll. J. Wood-Mason.
PENTHEMA DARLISA, 0. sp.
Male and Female. Differs from P. lisarda, Doubleday (Gen. D.
L. pl. 39. f. 3), on the fore wing in all the markings, except the
streak along posterior margin, bemg bluish white aud smaller, these
within the cell oval, two at the base and two near the end; the discal
series also are short and conical (not continuing outward to the inner
transverse series), the streak between the lower median and submedian
vein only being elongated. On the hind wing the pale luteous inter-
spaces between the veins are deeply excavated at their end and partly
coalesce with the inner transverse series of spots (the spots being
larger), the outer row are also larger, conical, and deeply excavated
externally ; a black streak traverses the cell.
Expanse 5 inches.
Meetan, 3000 feet (April). Iu coll. J. Wood-Mason.
PARTHENOS APICALIS, Nl. Sp.
Distinguished from P. gambrisius by the fore wing having the
white band quite diaphauous, without any borders to the veins, and
extending broadly and uninterruptedly to the costal margin, thence
continuing along it to the extreme apex.
Expanse 33 inches.
Above Ahsown ; Taoo, 3000-5000 feet. In coll. J. Wood-Mason.
LEBADEA ATTENUATA, 0. Sp.
Male and Female. Differ from L. ismene, Doubleday (Gen. D. Lep.
pl. 34. f. 2), in being smaller and of a brighter colour, the white
medial transverse band narrower, the apical portion on fore wing
more angular in position and extending to the costa, the outer
series of narrow lunules being present on the hind wing in both
Sexes.
Expanse, ¢ 2, 2 23 inches.
Hatsiega; Naththoung to Paboga. In coll. J. Wood-Mason.
830 MR. F. MOORE ON LEPIDOPTEROUS [ Nov. 5,
NEPTIS ADARA, 0. sp. “
Allied to N. varmona, from S. India. Differs above in the macu-
lated bands being narrower, the discoidal terminal spot broader and
less pointed, the discal series of spots more oval in shape, and the
submarginal series not. so prominent. On the underside the ground-
colour is much brighter, and the markings are less black-bordered,
Expanse, ¢ 2, 2 2,%; inches.
Moulmein to Meetan; Ahsown; Naththoung to Paboga ; Moolai,
3000-6000 feet. In coll. J. Wood-Mason.
NEPTIS MEETANA, 0). Sp.
Allied to the preceding (NV. adara), but with the markings above
more like thosein NV. andamana. It is asomewhat narrower-winged
insect, the spots of the discal band relatively narrower in both sexes,
the underside of a brighter red, and the markings less black-bordered.
Expanse, ¢ 2, 2 23 inches.
Meetan 3000 feet, March; Taoo, 3500 feet. In coll. J. Wood-
Mason.
NEPTIS JUMBA.
Neptisjumba, Moore, Catal. Lep. Mus. E.I. C. i. p. 167 (1857).
Moulmein to Meetan.
NEPTIS ADIPALA.
Neptis adipala, Moore, Proc. Zool. Soc. 1872, p. 563, pl. 22.
8.
Moulmein to Meetan ; Hatsiega.
NEPTIS PLAGIOSA, N. Sp.
Allied to N. hordonia, Stoll, Cram. Pap. v. pl. 33. f. 4.
Differs on the upperside in all the ferruginous bands being broader,
the fore wing having the lengthened discoidal band spreading over
the median vein, the marginal line also broader and very distinct ;
hind wing with the submarginal band nearly as broad as the discal.
On the underside the strigz are more numerous and darker, covering
both bands on the hind wing.
Expanse, ¢ 14, 2 14 inch.
Naththoung to Paboga. In coll. J. Wood-Mason and F. Moore.
ADOLIAS LEPIDEA.
Adolias lepidea, Butler, Aun. Nat. Hist. 1868,1. p. 71.
Meetan, 3000 feet, April.
ADOLIAS SATROPACES.
Adolias satropaces, Hewitson, Entom. Monthly Mag. 1876, p.
150; id. Dese. Lep. Coll. Atk. Asiat. Soc. Bengal, p. 1, pl. 1. f. 6,
7, 8 (1878).
Meetan, 3000 feet, April.
1878. ] INSECTS. FROM UPPER TENASSERIM. 831
ADOLIAS PARVATA, n. sp. (Plate LII. fig. 3.)
Female allied to 4. sedeva, Moore, Trans. Ent. Soc. 1859, p. 68,
pl. 4. f. 3. Differs above on the fore wing in the discal band being
narrower, the costal portion only partially whitish, and the lower
portion without the whitish inner border. Underside with the band
on fore wing only partially white anteriorly, the lower portion with
a slight white lunule to the inner border, and white point with black
tip to outer border.
Expanse 2? inches.
Hab. Meetan, 3000 feet, April. In coll. J. Wood-Mason.
ADOLIAs DISCIsPILOTA, n. sp. (Plate LII. fig. 2.)
Allied to 4. kesava, Moore, Trans. Ent. Soc. 1859, p. 67, pl. 3. f#5.
Female. Smaller in size: upperside with the borders uniformly
greyish brown ; the transverse discal band with less sinuous dusky
outer border; the whitish spot between the lower subcostal branches
and the spot between the upper median branches prominent.
Underside paler ; markings similar, with less blue along abdominal
border.
Expanse 2,2, inches.
Moolai, 3000-6000 feet. In coll. J. Wood-Mason.
ADOLIAS JAHNU.
Adolias jahnu, Moore, Catal. Lep. Mus. E.I. C.i. p. 192 (1857) ;
Trans. Ent. Soc. 1859, p. 74, pl. 7. f. 1.
Hatsiega ; Taoo, 3000-5000 feet.
ADOLIAS TAOOANA, 0. sp.
Allied to A. iva (Moore, Trans. Ent. Soc. 1857, pl. 8 f. 2). Fore
wing with the oblique discal maculated band with shorter lower por-
tions, the penultimate spot quite oval, the last being short and
conical ; the two contiguous spots beneath are also smaller, and the
two before the apex closer together; on the hind wing the three sub-
apical spots are contiguous, the two upper being large, quadrate, and
excavated externally.
Expanse 4 inches.
Taoo, 3000-5000 feet. In coll. J. Wood-Mason.
CHARAXES SAMATHA, N. Sp.
Smaller than Indian examples of C. athamas, the yellow band on
both wings one third less in width, the subapical spot smaller, and the
apical either minute or obsolete.
Expanse 22 inches.
Moolai, 3000-6000 feet. In coll. J. Wood-Mason.
CHARAXES MARMAX.
Charaxes marmax, Westwood, Cab. Orient. Ent. p. 43, pl. 21.
f. 3-5 (1848).
Moolai, 3000-6000 feet.
832 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
CHARAXES DESA, 0. sp.
Male. Allied to C. marmax, Westw. Cab. Orient. Ent. pl. 21. f.
3. Differs on fore wing above in the marginal black band being
broader and extending at the apex further along the costa, the inner
submarginal lunular line is closer to the band; the black maculated
band on the hind wing is formed of larger angled spots. Underside
purplish ochreous ; hind wing with a transverse black lunular fascia,
which is also further from the outer margin.
Expanse 33 inches.
Moolai, 3000-6000 feet. In coll. J. Wood-Mason.
Also distinct from C. lunawara, Butler.
CHARAXES AGNA, 0. sp.
Male. Allied to C. harpav, Feld. Nov. Voy., Lep. iii. p. 444. Dif-
fers in its larger size, broader marginal black band on the fore wing,
and in the hind wing having a less black apical patch, the marginal
series of spots also being much smaller.
Expanse 337 inches.
Moolai, 30U0—6000 feet. In coll. J. Wood-Mason.
CHARAXES HARPAX.
Charaxes harpaz, Felder, Reise Nov., Lep. ui. p. 444 (1867).
Moulmein to Meetan; Moolai, 3000-6000 feet.
PROTHOE FRANCKII.
Nymphalis franckii, Godt. Enc. Méth. ix. Suppl. p. 825; Horsf.
Catal. Lep. E.I. C. (1829), pl. 5 f. 4, 4 a.
Meetan, 3000 feet, April.
Subfam. LipyTH4xINz.
LiBYTHEA MYRRHA.
Libythea myrrha, Godt, Enc. Méth. ix. p 171; Boisd. Sp. Gén.
Lép. i. t. 10. f. 8.
Moulmein to Meetan ; Hatsiega.
Fam. Erycrnip&.
ZEMEROS FLEGYAS.
Papilio flegyas, Cram. Pap. Exot. iii. pl. 280. f. E, F (1782).
Ahsown, 2000 feet.
TAXILA FASCIATA, n. Sp. (Plate LII. fig. 1.)
Allied to 7. drupadi, Horsf. Catal. Lep. Mus. E.I. C. 1828, pl. 2.
f. 3, from Java.
Male. Differs above in the apex of the fore wing being less suffused
with red, and having an oblique subapical indistinct whitish fascia.
Underside similarly marked, with the subapical fascia distinct, well-
defined, and pure white.
Expanse 13 inch.
Above Ahsown. In coll. J. Wood-Mason.
ae
1878. ] INSECTS FROM UPPER TENASSERIM. 833
ABISARA NEOPHRON.
Sospita neophron, Hewitson, Exot. Butt. ii. Sosp. pl. i. f. 3
(1861).
Moolai, 3000-6000 feet.
ABISARA ANGULATA, 0. Sp.
From 4. kausambi, Felder, of the Malay peninsula, this species
differs in the male above being paler plum-colour, the fore wing show-
ing the transverse discal fascize, and the hind wing the apical and
anal black spots ; the female above is also paler purple-brown, the
inner discal-band is angled at the middle of its inner border on both
wings, and is only slightly paler at its costalend. On the underside,
the discal bands are angled in the middle, the inner band being
narrowly bordered with brownish white at its costal end only in the
male, its entire length in the female.
Expanse 143 inch
Above Ahsown; Hatsiega; Naththoung to Paboga. In coll. J.
Wood-Mason.
An entirely different insect from the Andaman A. bifusciata,
Moore, P. Z.S. 1877, p. 587, pl. 58. f. 1.
Fam. Lyca&nip&.
CASTALIUS ROXUS.
Lycena roxus, Godt. Ene. Méth. ix. p. 659 (1823); Horsf.
Catal. Lep. E.I. C. 1828, p. 70, pl. 2. f. 4.
Moulmein to Meetan.
LAMPIDES ALEXIS.
Papilio alexis, Stoll, Suppl. Cramer’s Pap. Exot. pl. 38. f. 3
(1790).
Moulmein to Meetan; Naththoung to Paboga.
LAMPIDES ZLIANUS.
Papileo zlianus, Fabr. Ent. Syst. iii. 1, p. 280 (1793).
Moulmein to Meetan.
« LAMPIDES ELPIS.
Lycena elpis, Godt. Enc. Méth. ix. p. 654 (1823); Horsf, Catal.
Lep. E.I. C. 1828, p. 96, pl. 1. f. 4.
Ahsown ; Moulmein to Meetan.
LAMPIDES KANDARPA.
Lycena kandarpa, Horsf. Catal. Lep. Mus. E.I. C. 1828, p. 82.
Moulmein to Meetan
HyYpoLyCNa LISIAS.
Papilio lisias, Fabr. Mant. Ins. p. 65 (1787); Donov. Ins. Ind,
pl. 40. f. 1°; Boisd. Spec. Gén. Lép. i. t. 22. f. 2.
Ahsown; Meetan, April; Moolai 3000-6000 feet.
834 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
Myrina FREJA.
Papilio freja, Fabr. Ent. Syst. iii. 1, p. 263 (1793); Butler,
P. Z.8. 1867, pp. 34, 36, f. 1.
Ahsown ; Moulmein to Meetan; Taoo, 3000-5000 feet.
THAMALA, N. g.
Male and Female. Fore wing short, broad, trigonate ; costa convex
at base, apex moderately acute, exterior margin slightly oblique and
recurved, posterior margin nearly straight, angle acute ; cell broad,
short ; subcostal vein four-branched, first, second, and third arising
before end of the cell, fourth at its end ; discocellulars slightly curved
outward ; radial from their middle; median vein three-branched, two
upper contiguous at base from end of the cell: hind wing elongated
posteriorly, narrow and quadrate below anal margin ; with two narrow
lengthened tails, one at each angle; exterior margin slightly waved.
Body moderate; antennz stout, uniformly thickened to apex ; palpi
long, squamous, second joint projecting half its length beyond the
head, apical joint slender; legs squamous, femora slightly pilose
beneath.
Differs from Deudorix (D. melampus (jarbas, Fabr.) and D. epi-
jarbas) in its shorter and broader fore wings and longer hind wings,
the fore wings having only four subcostal branches (there being five
in both the above species), and in the antennze being more robust.
THAMALA MINIATA, n. sp. (Plate LIT. fig. 6.)
Male and Female. Upperside deep scarlet-vermilion: fore wing
with the base, costal and the outer border broadly black, confining
the red colour to a circular area ; veins also black-lined, the median
prominently so and forming a slight streak at the base of its branches :
hind wing black at the extreme base, along abdominal border and
slightly along median veins ; exterior border narrowly black, some
black speckles ascending between lower median branch and submedian
vein; tails red, black-margined and white-tipped in female. Underside
luteous-yellow in male, ochreous-yellow in female: both wings with
indistinct dusky discal zigzag-lunular line terminating above anal angle
in a white-speckled patch: fringe of hind wing black, the margin
also black-lined from the angle; space above the tails black- and
white-speckled. Body beneath, legs, and palpi white, speckled with
black, tip of palpi black.
Expanse, $13, 2 154, inch.
Taoo, 3500 feet. In coll. J. Wood-Mason.
Devuporix surrusa, n. sp. (Plate LIL. fig. 8.)
Allied to D. xenophon, male. Wings slightly broader: upperside
of fore wing with broader outer band, the red colour suffused with
dusky-brown ; hind wing also duller red. Underside dull sulphur-
yellow ; transverse lunular line more curved, the zigzag end at anal
angle black and with an additional similar line above its end.
Expanse 1,4; inch.
Taoo, 3000-5000 feet. In coll. J. Wood-Mason.
1878. ] INSECTS FROM UPPER TENASSERIM. 835
LoxuURA ATYMNUS.
Papilio atymnus, Cram. Pap. Exot. iv. t. 331. f. D, E (1782).
Ahsown ; Meetan, 3000 feet, April; Moolai, 3000-6000 feet.
AMBLYPODIA TAOOANA, 0. sp.
Differs from A. narada and A. anita in its larger size, the upper-
side being of a very brilliant blue colour as in A. si/hetensis, and the
outer black marginal band twice the width of that of those species.
Underside purplish-ochreous, the transverse black-speckled band
and basal speckled markings prominent, the band on fore wing being
much curved, the outer markings also prominent.
Expanse 2 inches.
Taoo, 3500 feet. In coll. J. Wood-Mason.
NARATHURA, 0. g.
Wings broad: fore wing very convex at base of costa, exterior
margin rounded, even: hind wing very convex externally, with even
margin; a very slender tail at end of lower median vein (so delicate
that in most cabinet specimens it is broken off); anal angle not
lobed.
Type N. hypomuta (Amblypodia hypomuta, Hewits. Catal. Lyc.
B. M. p. 11, pl. 6. figs. 63, 64, 1862).
NARATHURA MOOLAIANA, 0. Sp.
Amblypodia epimuta, Hewitson, Catal. Lyc. B.M. p. 11, pl. 6. f.
59, 60 (1862).
Taoo, 3000-5000 feet. Moolai, 3000-6000 feet. In coll. J.
Wood-Mason.
This is a distinct species from N. epimuta, Moore, Catal. Lep.
Mus. E.I.C. i. p. 42, the type of which is from Borneo.
ARHOPALA NAKULA, Feld. Wien. ent. Monats. iv. p. 395 (1860).
Taoo, 3000-5000 feet.
ARHOPALA viHARA, Feld. Wien. ent. Monats. iv. p. 395 (1860).
Taoo, 3000-5000 feet.
SURENDRA, 0. g.
Allied to Thaduka. Sexes dissimilar in colour above: fore wing
short, broad, costa slightly arched, apex acute, exterior margin in
female very convex in the middle, less so in male; hind margin
nearly straight: hind wing short, somewhat quadrate; anterior
margin nearly straight, apex angled; exterior margin nearly straight
anteriorly, truncate posteriorly, and in male with one tail situated
at end of lower median vein, in female with two tails, one at the
lower, the other at the middle median vein, anal lobe large. Vena-
tion similar to Thaduka. Palpi long, slender; legs short; antennse
uniformly thickened.
836 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
SURENDRA QUERCETORUM.
Amblypodia quercetorum, Moore, Catal. Lep. Mus. E.I. C. i. p.
42, pl. la. f. 7 (1857).
THADUKA, 0. g.
Allied to Mahathala. Fore wing short, broad, costa very convex
at base, apex acutely angled; exterior margin erect, scalloped ;
posterior angle lobular ; hind margin same length as costal, concave
in the middle: hind wing short, broad, anterior margin convex, apex
and exterior margin very convex, sinuous, with three prominent tails,
the middle one longest, and large anal lobe ; abdominal margin very
concave above anal lobe. Venation similar to Mahathala(Amblypodia
ameria, Hewitson). Body short, stout; antenne uniformly
thickened to end; palpi slender ; legs short.
THADUKA MULTICAUDATA, n. sp. (Plate LII. fig. 7.)
Female. Upperside purple-black, basal area bright smalt-blue.
Body and abdominal border greyish, thorax blue. Underside dark
vinous-brown ; fore wing with short subbasal, medial, and an entire
discal, purple-black maculate bands, and an outer marginal series
of lunules: hind wing with three irregular curved transverse
purplish-black maculate bands, and indistinct marginal lunules ;
a narrow metallic-green lunule above the tails.
Expanse 12 inch.
Taoo, 3000-5000 feet. In coll. J. Wood-Mason.
Fam. PAPILionIp&,
Subfam. Prerinz&.
TERIAS HECABE.
Papilio hecabe, Linn. Syst. Nat. i. 2, p. 763 (1767).
Moulmein to Meetan ; Hatsiega; Naththoung to Paboga.
TERIAS SUAVA.
Terias suava, Boisd. Spec. Gén. Lép. p. 670 (1836).
Moulmein to Meetan.
TERIAS SILHETANA.
Terias silhetana, Wallace, Trans. Ent. Soc. 1867, p. 324.
Moulmein to Meetan ; Hatsiega to Houngduran; Naththoung
to Paboga.
TERIAS FORMOSA.
Eurema formosa, Hiibn. Zutr. exot. Schmett. f. 979 (1837).
Hatsiega.
TERIAS LATA.
Terias leta, Boisd. Spec. Gén, Lép. i. p. 674 (1836).
Moulmein to Meetan.
_
1878. ] INSECTS FROM UPPER TENASSERIM. 837
NYCHITONIA XIPHIA.
Papilio xiphia, Fabr. Spec. Gén. Lép. ii. p. 43.
P. nina, Fabr. Ent. Syst. iti. 1, p. 194.
Ahsown; above Ahsown.
CALLIDRYAS CATILLA.
Papilio catilla, Cram. Pap. Exot. iii. t. 229. f. D, E (1782);
Butler, Lep. Exot. i. p. 24, pl. 9. f. 7-10.
Hatsiega to Houngduran ; Naththoung to Paboga; Moolai, 3000-
6000 feet.
CALLIDRYAS CROCALE.
Papilio crocale, Cram. Pap. Exot. i. t. 52. f. C, D (1779);
Butler, Lep. Exot. i. p. 22, pl. 9. f. 1, 2.
Moolai to Meetan.
HEBOMOIA GLAUCIPPE.
Papilio glaucippe, Linn. Syst. Nat. i. 2, p. 762 (1767).
Houngduran source.
Ix1As PALLIDA, Nl. Sp.
Nearest to I. andamana, Moore, P. Z. 8. 1877, p. 590. Upper-
side of the same pale colour, the orange band on fore wing slightly
wider, and having a more convex outer margin, the inner black border
also broader, and the marginal band on hind wing wide and of equal
width throughout. Underside differs in having a uniform series of
delicate black strigze and a cell-spot on both wings.
Expanse 22 inches.
Moolai, 3000-6000 feet. In coll. J. Wood-Mason.
From J. latifasciata, Butler, this species differs in its much paler
colour above and beneath, and in the absence of the dark blotches
on the underside.
IXIAS CITRINA, N. Sp.
Male. Allied to the preceding; the orange band comparatively
narrower and shorter hindward, its black inner border also narrower,
and the marginal band on hind wing only half its width. Underside
darker, with more prominent black strigee and cell-spot, and with a
submarginal series of brown-speckled spots with whitish centre.
Expanse 2 inches.
Houndurang source. In coll. J. Wood-Mason.
This also may be distinguished from J. latifasciata, by the colour
above and by the narrow band on the hind wing.
Ix1AS MOULMEINENSIS, N. sp.
Male and Female. Near to I. pirenassa, Wallace, from Madras.
Upperside of the same bright yellow, the orange band in male much
broader anteriorly ; pale blackish band on hind wing narrow and not
extending to posterior angle. Underside dull ochreous-yellow, with
Proc. Zoot. Soc.—1878, No. LY. 55
838 MR. F. MOORE ON LEPIDOPTEROUS [ Nov. 5,
delicate dark strigze, and submarginal brown spots centred with white.
Female with moderately broad orange band.
Expanse, ¢ 2, 9 12 inch.
Moulmein to Meetan. In coll. J. Wood-Mason,
Eronia Ltutescens, Butler.
Moulmein to Meetan.
CaTorHAGA NEOMBO.
Pieris neombo, Boisd. Spec. Gén. Lép. i. p. 539 (1836); Moore,
Catal. Lep. Mus. E.I. C. i. pl. 2a. f. 3.
Moulmein to Meetan (a single female).
CATOPHAGA PAULINA.
Pieris paulina, Cram. Pap. Exot. ii. t. 110. f. E, F (1779).
Moulmein to Meetan; Hatsiega to Houngduran; Naththoung
to Paboga.
(All males.)
CATOPHAGA LAGELA, 0. sp. (Plate LII. fig. 4.)
Allied to C. /alage, Doubleday, D. Lep. pl. 6. f. 3.
Male and Female. Smaller. Upperside differing on the fore wing
in the black apical border terminating in both sexes before reaching
the posterior angle, and the medial portion partly excavated outside the
lower end of the cell and thence extending across the end to its base ;
hind wing with a broad marginal continuous band (as in C. pandione,
Hiibn.). Underside—fore wing with the black band terminating
as above; the apex and hind wing greyish-yellow, speckled with
purple in male, and brownish-grey with darker speckles in
female, the speckles numerous across the disk, and forming zigzag
fasciz.
Expanse 23 inches.
Moolai, 3000-6000 feet. In coll. J. Wood-Mason.
APPIAS LEA.
Pieris lea, Doubleday, Aun. Nat. Hist. 1846, p. 23.
P. clemanthe, Doubleday, Gen. D. Lep. pl. 6. f. 3.
Moolai, 3000-6000 feet.
APPIAS NAMA.
Pieris nama, Moore, Catal. Lep. Mus. E.I. C. i. p. 76 (1857) ;
Proc. Zool. Soc. 1857, p. 102, pl. 44. f. 1, 2.
Houngduran source; Taoo, 3000-5000 feet (March); Moolai,
8000-6000 feet; Moolai to Moolat, 4000-5000 feet.
APPIAS DAPHA, Nl. Sp.
Allied to dA. zeuzippe, Cram.
Male differs on the upperside in being white ; the apical band on
the fore wing more like that of 4. nama, narrower, deeply scalloped
within; anterior spots obsolete; and the contiguous discal spots
1878.] INSECTS FROM UPPER TENASSERIM. 839
between the two upper median branches scarcely perceptible;
marginal spots on hind wing indistinct, speckles disconnected.
Underside white; apex of fore wing slightly dusky grey; subcostal
and median vein within the cell, and all the veins of hind wing with
dusky borders.
Female also white ; apical band on fore wing without anterior spots ;
other markings similar but less prominent.
Expanse 17 inch.
Moulmein to Meetan. In coll. J. Wood-Mason.
APPIAS ZELMIRA.
Papilio zelmira, Cram. Pap. Exot. iv. t. 320. f. C, D (1782).
Moulmein to Meetan.
APPIAS AMBA.
Pieris amba, Wallace, Trans. Ent. Soe. 1867, p. 340.
Moulmein to Meetan ; Houngduran source.
APPIAS VACANS.
Appias vacans, Butler, Trans. Ent. Soc. 1870, p. 490, 9°.
Moulmein to Meetan; Hatsiega to Houngduran.
DELIAS PASITHOE,
Papilio pasithoé, Linn. Syst. Nat. i. 2, p. 755 (1767).
Ahsown, 2000 feet ; Taoo, 3000-5000 feet (March).
DELIAS INDICA.
Thyca indica, Wallace, Trans. Ent. Soc. 1867, p. 351.
Above Ahsown ; Moulmein to Meetan. Moolai, 3000-6000 feet.
DELIAS DESCOMBESI.
Pieris descombesi, Boisd. Spec. Gén. Lép. i. p. 465.
Ahsown ; Moolai, 3000-6000 feet.
PRIONERIS WATSONI.
Prioneris watsoni, Hewitson, Trans. Ent. Soc. 1868, p. 100.
Moulmein to Meetan ; Houngduran source.
PRIONERIS CLEMANTHE.
Pieris clemanthe, Doubleday, Ann. Nat. Hist. 1846, p. 23.
P. helferi, Felder, Reise Nov., Lep. ii. p. 161, t. 25. f. 10.
Houngduran source; Naththoung to Paboga.
Subfam. PAariLioninz.
PAPILIO TELEARCHUS.
apilio telearchus, Hewitson, Trans. Ent. Soc. 1852, p. 22, pl.
6. f. 3.
Hatsiega.
55*
840 MR. F. MOORE ON LEPIDOPTEROUS { Nov. 5,
PAPILIO XENOCLES.
Papilio xenocles, Doubleday, Gray’s Zool. Mise. p. 74; Gen. D.
Lep. pl. 1*. fig. 2; Westw. Arc. Ent. ii. pl. 79. f. 2.
Hatsiega.
PAPILIO MEGARUS.
Papilio megarus, Westwood, Arc. Ent. ii. pl. 72. f. 2 (1845).
Houngduran source.
PAPILIO ONPAPE, 0. Sp.
Differs from specimens of Himalayan P., panope Linn. (Cram.
Pap. Exot. iv. pl. 295. f. E, F) in having comparatively shorter
wings, smaller and less-prominent spots on the fore wing, smaller
marginal lunules, and much shorter dentate submarginal streaks
on the hind wing.
Expanse 34 inches.
Hatsiega ; Houngduran source; Naththoung. In coll. J. Wood-
Mason and F. Moore.
PAPILIO PAMMON.
Papilio pammon, Linn, Syst. Nat. i. 2, p. 746 (1767).
P. polytes, Linn. Syst. Nat. i. 2, p. 746.
Ahsown, 2000 feet; Meetan, 3000 feet (March) ; Taoo, 3000-
5000 feet (March).
PAPILIO ARISTOLOCHIZ.
Papilio aristolochiea, Fabr. Syst. Ent. p. 443 (1775).
P. diphilus, Esper, Ausl. Schmett. pl. 40 3. f. 1.
Moulmein to Meetan ; Hatsiega ; Naththoung to Paboga.
PapILio PHILOXENUS.
Papilio philowenus, Gray, Zool. Mise. p. 32 (1831); Lep. Ins.
Nepal, p. 5, pl. 2.
Moulmein to Meetan; Moolai to Moolat, 4500 feet.
PAPILIO DOUBLEDAYI.
Papilio doubledayi, Wallace, Trans. Ent. Soc. 1865, p. 42.
Moulmein to Meetan.
PAPILIO HELENUS.
Papilio helenus, Linn. Syst. Nat. i. 2, p. 745; Cram. Pap. Exot.
Met. tod. 1, A,B.
Hatsiega ; Moolai to Moolat, 4500 feet.
PAPILIO MAHADEVA,n. sp. (Plate LI. fig. 1.)
Male. Allied to P. castor, but with shorter hind wings, which are
of the same form as in P. panope. Upperside dark brown,
nemerously covered with minute ferruginous scales; fore wing
with a marginal series of very small white lunular spots, one between
_
1878.] INSECTS FROM UPPER TENASSERIM. 841
each vein; hind wing with a discal series of seven short conical
yellowish-white spots, a submarginal series of narrow dentate lunules,
and a marginal very narrow lunular line, one between each vein.
Underside paler, more numerously covered with ferruginous scales ;
the markings as above; the fore wing also having a second marginal
series of very small white spots, and a spot at the end of the cell.
Expanse 4 inches.
Moolai to Moolat, 4500 feet. In coll. J. Wood-Mason.
A specimen, labelled ‘‘ Eastern Bengal,” from Mr. A. Grote’s col-
lection, of what I believe to be the female of P. mahadeva, differs from
that sex of P. castor in the fore wings having very small and less-
distinct submarginal white spots, no spot at the end of the cell, and
no white speckles between the discal interspaces ; the hind wings on
both sides also have the discal streaks shorter and confined to the
middle of the wing, there being none either at the base or in the
cell, thus forming a transverse medial band similar in position to
that of the male above described ; the submarginal dentate marks are
less defined.
PAapIL{0 ANDROGEOS.
Papilio androgeos, Cram. Pap. Exot. i. pl. 91.
Ahsown, 2000 feet ; above Ahsown.
PAPILIO ZELEUCUS,
Papilio zeleucus, Hewitson, Exot. Butt. iii. Pap. pl. 8. f. 24, 25
(1865).
Moulmein to Meetan.
PAPILIO SARPEDON.
Papilio sarpedon, Linn. Syst. Nat. i. 2, p. 747 (1767).
Hatsiega ; Houngduran source.
PaPILIO AGAMEMNON.
Papilio agamemnon, Linn. Syst. Nat. i. 2, p. 748 (1767).
Moulmein to Meetan ; Hatsiega ; Houngduran source.
PAPILIO ANTIPHATES.
Papilio antiphates, Cram. Pap. Exot. i. t. 72. f. A, B (1779).
Houngduran source.
LEPTOCIRCUS VIRESCENS.
( 2 eaabae virescens, Butler, Catal. Fabr. Lep. B. M. p. 259
1870).
Ahsown ; Moulmein to Meetan; Houngduran source; Moolai,
3000-6000 feet.
Fam. HEsPERID&.
IsMENE BENJAMINI.
Hesperia benjamini, Guér. Deless. Voy. Inde,‘ii. p. 79, pl. 22. f. 2
(1843).
Above Ahsown.
842 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
IsMENE DRUNA.
Ismene druna, Moore, Proc. Zool. Soc. 1865, p. 784; Hewitson,
Exot. Butt. iv. Hesp. pl. 2. f. 26.
Taoo, 3000-5000 feet.
TAGIADES PRALAYA.
Tagiades pralaya, Moore, Proc. Zool. Soc. 1865, p. 779.
Ahsown.
TAGIADES MEETANA, n. sp. (Plate LIII. fig. 1.)
Male. Allied to T. obscurus, Mabille, from Java: fore wing above
similar: hind wing differs in the marginal area of the anal angle
being only slightly white-speckled between the veins; the cilia below
it dark brown with a very narrow whitish inner line; on the under-
side there is less white, and the black border more lengthened and
prominent, the cilia also being brown.
Expanse 1% inch.
Meetan, 3000 feet (April). In coll. J. Wood-Mason.
ASTICTOPTERUS DIOCLES.
Nisoniades diocles, Moore, Proc. Zool. Soc. 1865, p. 787.
Meetan, 3000 feet.
ASTICTOPTERUS SUBFASCIATUS, 0. Sp.
Male and Female. Upperside uniform vinous-brown. Underside—
fore wing with a short vinous-grey streak and two small ochreous-grey
spots from the apex, three small ochreous-grey spots obliquely from
the costa before the apex ; hind margin pale ochreous-brown: hind
wing with a short, broad, medial and a subbasal vinous-grey band ;
abdominal margin broadly pale ochreous-brown; some indistinct
spots between the bands and at the base also pale ochreous-brown.
Palpi, body, and legs beneath pale ochreous-brown.
Expanse 14 inch.
Ahsown. In coll. J. Wood-Mason.
A specimen of this species from S. India is also in the collection
of the British Museum,
PAMPHILA BAMBUS&.
Pamphila bambuse, Moore, Proc. Zool. Soc. 1878, p. 691, pl. 45.
fe dl, 12s
Hatsiega.
PAMPHILA MASONI, n. sp. (Plate LII. fig. 5.)
Male. Upperside dark oliye-brown; cilia brownish ochreous,
palest on hind wing ; fore wing with a yellow semidiaphanous sinuous
spot at end of the cell, two spots before the apex, and a band of three
oblique discal spots, the lower one elongated; hind wing with an
opaque yellow discal patch, the basal hairs green. Underside dark
ochreous-yellow ; fore wing with the basal area black ; spots as above,
1878. ] INSECTS FROM UPPER TENASSERIM. 843
the lower discal portion broad: hind wing with an indistinct series of
marginal and submarginal brown spots. Front of head, palpi, legs,
and body beneath yellow.
Expanse 1,4 inch.
Hatsiega. ‘In coll. J. Wood-Mason.
PAMPHILA MATHIAS.
Hesperia mathias, Fabr. Ent. Syst. Suppl. p. 433 (1798) ;
Butler, Catal. Fabr. Lep. B. M. pl. 3. f. 6.
Moulmein to Meetan.
HEsPERIA MOOLATA, 0. sp.
Male and Female. Upperside dark vinous-brown, slightly ative
brown at the base; cilia cinereous-yellow : fore wing with a single
small semidiaphanous white spot at lower end of the cell, two smaller
spots obliquely before the apex, and three below on the disk, the
lowest being the largest and angular; the female with two spots at
end of the cell, and a small opaque yellowish spot between lower
median branch and submedian vein. Underside brighter-coloured,
marked as above, the yellow discal lower spot in female larger.
Expanse 14 inch.
Ahsown (2000 feet), March. Moolai to Moolat.
This species is allied to H. kumara, Moore, from South India, and
to H. cahira from the Andamans.
PLESIONEURA AURIVITTATA, n. sp. (Plate LIII. fig. 2.)
Male and Female. Upperside dark golden olive-brown : fore wing
with a broad oblique golden-yellow discal band curving from middle
of the costa to posterior angle, the band semiopaque from the costal
vein to lower median branch ; a small curved yellow streak composed
of three vein-crossed spots before the apex. Cilia of both wings
brown. Underside duller brown: fore wing as above: hind wing
slightly yellow-speckled, and with a very indistinct yellowish streak
at the end of the cell.
Expanse 1,/, inch.
Hab. Above Ahsown. In coll. J. Wood-Mason and F. Moore.
Allied to P. dhanada, Moore, P. Z. 8. 1865, p. 789. Distinguished
by the oblique transverse band being very broad throughout and ex-
tending to the posterior angle, also in the cilia of hind wing being
entirely brown.
PLESIONEURA ALBIFASCIA, 0. sp. (Plate LIII. fig. 3.)
Male. Differs from P. alysos, Moore, in its narrower fore wing
and smaller hind wing; the oblique semidiaphanous white band is
shorter and straight, the portion beneath the lower median branch
being very small, this part on the underside showing only as a suf-
fused white patch ; and there are no subapical spots.
Expanse |4 inch.
Hatsiega. In coll. J. Wood-Mason.
A specimen of this species, collected by Mr. Buxton, probably in
Sumatra, is in my own collection.
844 MR. F. MOORE ON LEPIDOPTEROUS [ Noy. 5,
TIsoTEINON SUBTESTACEUS, Nl. Sp.
Male and Female. Upperside dark olive-brown ; cilia cinereous-
brown, with a brown inner line and indistinct bars: fore wing with
a semidiaphanous spot at end of the cell, a curved series of three
subapical contiguous spots, and two larger contiguous spots on the
disk. Underside brownish ochreous, grey-speckled : fore wing with
the basal area below the cell and the disk fuliginous black ; spots
as above; male with a black tuft on hind margin: hind wing with
a curved discal series of six sinall white spots, and four subbasal
spots, the two lower of the latter being contiguous, the third at the
end of the cell, and the fourth above the cell. Palpi and body
beneath ochreous-white ; legs brownish ochreous. Allied to I. a¢-
kinsoni, from Darjiling.
Expanse 12 inch.
Ahsown. In coll. J. Wood-Mason.
SPHINGES.
Fam. SpHINGID.
Subfam. MacroGLossin&.
MACROGLOSSA ORIENTALIS.
Macroglossa orientalis, Butler, Trans. Linn. Soc. x. p. 528
(1876).
Ahsown.
MacroGuLossa LUTEATA.
Macroglossa luteata, Butler, Proc. Zool. Soc. 1875, p. 241, pl.
hs: de Dp
Ahsown.
Subfam. AMBULYCINE.
AMBULYX SUBSTRIGILIS.
Ambulyx substrigilis, Westwood, Cab. Orient. Ent. pl. 30. f. 2.
Taoo, 3000-5000 feet.
BoMBYCEs.
Fam. ZYG&NID&z.
Subfam. SynroMINz.
SYNTOMIS GROTEI.
Syntomis grote, Moore, Proc. Zool. Soc. 1871, p. 245, pl. 18.
f.4.
Taoo, 3000-5000 feet, March.
SYNTOMIS SLADENI.
Syntomis sladeni, Moore, Proc. Zool. Soc. 1871, p. 245, pl. 18.
f. 5.
Moolai to Moolat, 4500 feet. —
1878. | INSECTS FROM UPPER TENASSERIM. 845
SYNTOMIS ATKINSONI.
sepsis atkinsoni, Moore, Proc. Zool. Soc. 1871, p. 245, pl. 18.
Baas Ahsown; Naththoung to Paboga.
SYNTOMIS LIBERA.
Syntomis libera, Walker, Catal. Lep. Het. B. M. Suppl. p. 78.
Above Ahsown; Meetan, 3000 feet, April.
SYNTOMIS MASONI, n. sp. (Plate LIII. fig. 4.)
Allied to S. vitreata, H.-Sch. Lep. Spec. Nov. f. 247. Distin-
guished by having the hyaline spaces broader, by the space between
the lower subcostal and upper median branch being black, and the
front of the head white.
Expanse, 5 1,5, 2 1,%; inch.
Moolai, 3000- 6000 Ba Moolai to Moolat, 4500 feet. In coll.
J. Wood-Mason.
SYNTOMIs DISRUPTA, n. sp. (Plate LIII. fig. 5.)
Similar to S. divisa, Walker. Differs in its more pointed fore
wing, the hyaline spaces extending throughout to the exterior mar-
gins, the black borders being narrower.
Expanse 14 inch.
Moolai, 3000-6000 feet ; Moolai to Moolat, 4500 feet. In coll.
J. Wood-Mason.
SYNTOMIS BERINDA, n. sp. (Plate LIII. fig. 8.)
Allied to S. fenestrata (H.-Sch. Lep. Spec. Noy. f. 270). Smaller,
the hyaline spots on fore wing shorter, those on the hind wing nar-
rower and pale orange-colour. Front of head, collar, a spot on
tegulze, and a spot on hind part of thorax orange-red; the orange
bands on abdomen narrow ; anal segment black ; tarsi with a white
band.
Expanse, gd 1,4, 2 1,3; inch.
Ahsown, 2000 feet ; above Ahsown.
SYNTOMIS ALBIFRONS, n. sp. (Plate LIII. fig. 6.)
Allied to S. diaphana, Kollar, and S. melas, Walker. Dark
purple-black : fore wing with the basal area of the cell, five large
elongated discal spots, a small spot between the two lower median
veins (obsolete in some specimens), and a narrow indented space from
the base above the submedian vein hyaline: hind wing with three
short hyaline spots, the outer small, and in some specimens obsolete :
front of head, collar above, and four streaks down the thorax white;
an orange band on hind part of thorax ; abdomen glossy greyish
blue, with an orange band on the first and fifth segments, and a
lateral band on third and fourth segments ; legs slaty blue.
Expanse 2,2; inches.
Above neon Moolai, 3u00-—6000 feet. In coll. J. Wood-Mason.
846 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
Fam. CHALCOSIIDZ.
RETINA RUBRIVITTIA.
Retina rubrivitta, Walker, Catal. Lep. Het. B. M. ii. p. 439
(1854).
Moolai te Moolat, 4000-5000 feet.
HIsTIA PAPILIONARIA.
Gynautocera papilionaria, Guérin, Mag. Zool. 1836, p. 12.
Moolai, 3000-6000 feet.
Fam. EuscHEMID#.
EvusCHEMA MILITARIS.
Phalena militaris, Linn. [Cram. Pap. Exot. i. pl. 29. f. B}.
Moolai, 3000-6000 feet ; Moolai to Moolat, 4500 feet.
EvusCHEMA EXCUBITOR, 0. sp.
Male and Female. Differs from £. militaris in its much smaller
size and more angular outer margin of hind wing; the basal markings
are linear and very narrow, those on the hind wing smaller, the
discal zigzag band and marginal marks also very narrow, the sub-
marginal spots much smaller, and continuing in a separated series to
the subcostal branch.
Expanse 22 to 22 inches.
Taoo, 3000-5000 feet, March ; Houngduran source. In coll. J.
Wood-Mason.
This species is also an inhabitant of the Khasia hills.
EvuscHEMA AURILIMBATA, 0. Sp.
Allied to E. malayana, Guér. Deless. Voy. l’Inde,’ pl. 23. f. 2,
from the Malay coast.
Male pale blue: fore wing with the bands broad and partly con-
fluent : hind wing with the marginal area beyond the zigzag band,
and the abdominal border, bright chrome-yellow. Abdomen at base
above, tufted sides, body beneath, and legs chrome-yellow.
Expanse 3} inches.
Ahsown. In coll. J. Wood-Mason.
EusCHEMA HORSFIELDI.
Euschema horsfieldi, Moore, Catal. Lep. Mus. E.I. C. ii. p. 334,
pl. 8a. f. 7.
Ahsown, 2000 feet; Moolai, 3000-6000 feet.
Fam. NycTeMERIDA.
NyYcTEMERA LATISTRIGA.
Nyctemera latistriga, Walker, Catal. Lep. Het. B. M. ii. p. 397
(1854).
Ahsown.
1878.] INSECTS FROM UPPER TENASSERIM. 847
PirastLa MOOLAICA, n. sp. (Plate LIII. fig. 10.)
Male and Female. Smaller than P. varians (Nyctemera varians,
Walker); markings similar, those on fore wing narrower and dis.
connected, the transverse bands broken ; and there is also a small
white spot at the extreme apex of the wing; those on the hind wing
are also smaller.
Expanse 2 inches.
Moolai, 3000-6000 feet. In coll. J . Wood-Mason.
Fam. Liruosiip.
ANAGNIA SUBFASCIATA.
Anagnia subfasciata, Walker, Catal. Lep. Het. B. M. ii. p. 446
(1854).
Ahsown, 2000 feet ; Moolai to Moolat, 4500 feet.
PERIDROME ORBICULARIS.
Peridrome orbicularis, Walker, Catal. Lep. Het. B. M, ii. p- 445
(1854).
Aganopis subquadrata, H.-Sch. Lep. Spec. Noy. p- 70, f. 501
(1856).
Above Ahsown ; Taoo, 3000-5000 feet.
DaMALIs PLAGINOTA.
Damalis plaginota, Butler, Trans. Ent. Soc, 1875, p. 320.
Above Ahsown.
DaMatis ALcipHRon,
Phalena alciphron, Cramer, Pap. Exot. ii. pl. 133. f. E,
Moulmein to Meetan ; Moolai to Moolat, 4500 feet.
Hypsa suBsiMILIs.
Hypsa subsimilis, Walker, Catal. Lep. Het. B. M. Suppl. p. 212
(1864).
Ahsown.
NEOCHERA MARMOREA.
Neochera marmorea, Walker, Catal. Lep. Het. B. M. vii. p- 1674
(1856).
Above Ahsown; Taoo, 3000-5000 feet; Moolai, 3000-6000 feet.
CEonIsTIs ENTELLA.
Phalena entelia, Cramer, Pap. Exot. iii. pl. 208. f. D,
Moulmein to Meetan.
Fam. Arctiip 2».
ATTATHA, n. g,
Allied to Euplagia (2. hera of Europe). Fore wing elongated,
narrow ; costa straight ; apex bluntly angular ; exterior margin ob-
848 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
lique, angle conxex ; posterior margin convex at base. Hind wing
oval, exterior margin very convex. Venation similar to that of Z.
hera. Body elongated ; abdomen extending beyond hind wing: an-
tennee slender, minutely setose. Legs long, femora and tibia slightly
pilose. Palpi long, slender, porrect.
ATTATHA REGALIS.
Hypercompa regalis, Moore, P. Z.S. 1872, p. 575, pl. 33. f. 7.
Meetan, 3000 feet, March.
Fam. Liparipz.
ARTAXA VARIANS.
Artaxa varians, Walker, Catal. Lep. Het. B. M. iv. p. 796
(1855).
Ahsown.
PERINA BASALIS.
Perina basalis, Walker, Catal. Lep. Het. B. M. iv. p. 966 (1855).
Above Absown.
Fam. LastocaMPIp&.
DREATA TAOOENSIS, n. sp. (Plate LIII. fig. 7.)
Allied to D. petola, Moore, Catal. Lep. Mus, E.I. C. ii. pl. 10a.
f. 2, from Java.
Male. Upperside ochreous-grey ; three medial transverse very in-
distinct dusky lunular lines, a slightly darker discal line and a narrow
outer sinuous line, with brighter interspace between the two lines, the
point between the two lower median branches only with dusky centre.
Underside as above.
Expanse 37 inches.
Taoo, 3000-5000 feet. In coll. J. Wood-Mason.
TRABALA VISHNU.
Gastropacha vishnu, Lefebvre, Zool. Journ. iii. p. 207 (1827).
Taoo, 3000-5000 feet.
NocrTueEs.
Fam. OPpHIDERID.
PoOTAMOPHORA MANLIA.
Phalena manlia, Cram. Pap. Exot. pl. 92. f. A.
Above Ahsown.
Fam. ErResip2.
ARGIVA HIEROGLYPHICA.
Phalena hieroglyphica, Drury, Exot. Ins. ii. pl. 2. f. 1.
Moulmein to Meetan.
ARGIVA CAPRIMULGUS, Fab.
Phalena caprimulgus, Fab. (Guén. Noct. iii. p. 180).
Moolai, 3000-6000 feet.
1878. ] INSECTS FROM UPPER TENASSERIM. 849
NycTIPAO CREPUSCULARIS.
Phalena crepuscularis, Linn. (Clerck, Icones, pl. 53. f. 1).
Meetan, 3000 feet, April.
Fam. Hypopyripz&.
SprrRAMA HELICINA.
Speiredonia helicina, Hiibn. Zutr. f. 437.
Moolai, 3000-6000 feet.
ENTOMOGRAMMA FAUTRIX.
Entomogramma fautriz, Guén. Noct. iil. p. 204.
Naththoung to Paboga.
Fam. Opuiusip£.
ARTENA SUBMIRA.
Artena submira, Walker, Catal. Lep. Het. B. M. xiv. p. 1389.
Above Ahsown.
FopiNa SToLa.
Fodina stola, Guén. Noct. iii. p. 275.
Ahsown ; Taoo, 3000-5000 feet.
CaLEsIA FLABELLIFERA, N. Sp.
Smaller than C. comesa, Guén. Noct. iii. p. 258 ; wings somewhat
brighter-coloured, the lappet of hairs on fore wing also smaller and
composed of shorter hair; fore tarsi densely clothed with black hair.
Expanse 12 inch.
Ahsown. In coll. J. Wood-Mason.
CALESIA GASTROPACHOIDES.
Calesia gastropachoides, Guén. Noct. iii. p. 258.
Above Ahsown.
PYRALES.
Fam. SprLoMELID.
ZEBRONIA PLUTUSALIS.
Zebronia plutusalis, Walker, Catal. Lep. Het. B. M. xvii. p. 478.
Ahsowa.
Fam. MarRGARONID.
GLYPHODES BIVITRALIS.
Glyphodes bivitralis, Guén. Delt. et Pyral. p. 293.
Houngduran source.
GLYPHODES STOLALIS.
Glyphodes stolalis, Guén. Delt. et Pyral. p. 293, pl. 3. f. 11.
Moulmein to Meetan; Houngduran source; Naththoung to
Paboga.
850 MR. F. MOORE ON LEPIDOPTEROUS [Nov. 5,
CypDALIMA CONCHYLALIS.
Margarodes conchylalis, Guén. Delt. et Pyral. p. 303, pl. 8. f. 9.
Hatsiega.
PacHYARCHES VERTUMNALIS.
Margarodes vertumnalis, Guén. Delt. et Pyral. p. 309.
Houngduran source ; Naththoung to Paboga.
PaCHYARCHES MARTHESIUSALIs.
Margaronia marthesiusalis, Walker, Catal. Lep. Het. B. M.
Xvill. p. 531.
Houngduran source.
SisyYROPHORA PFEIFFER.
Sisyrophora pfeiffere, Lederer, Wien. ent. Monat. vii. p. 399,
pl. 13. f. 13.
Moulmein to Meetan; Houngduran source; Naththoung to
Paboga.
EuUGLYPHIS PROCOPIALIS.
Phalena procopialis, Cram. Pap. Exot. iv. pl. 368. f. E.
Ahsown, 2000 feet.
Fam. Botyp.
Borys JOPASALIS.
Botys jopasalis, Walker, Catal. Lep. Het. B. M. xviii. p. 652.
Houngduran source.
Borys CALDUSALIS.
Botys caldusalis, Walker, Catal. Lep. Het. B. M. xviii. p. 650.
Naththoung to Paboga.
Borys VINACEALIs.
Botys vinacealis, Moore, Proc. Zool. Soc. 1877, p. 619.
Moulmein to Meetan; Houngduran source.
Borys UNITALIS.
Botys unitalis, Guén. Delt. et Pyral. p. 349.
Moulmein.
GEOMETRES.
Fam. URAPTERIDZ.
URAPTERYX PODALIRIATA.
Urapteryx podaliriata, Guén. Phal. i. p. 32 (1857).
Above Ahsown.
1878. ] INSECTS FROM UPPER TENASSERIM. 851
Fam. ENNomID&.
HyprryYTHRA LIMBOLARIA.
Hyperythra limbolaria, Guén. Phal.i. p. 101, pl. 3. f. 3, 4.
Ahsown.
HyYPeRYTHRA ANGULIFASCIA, n. sp. (Plate LIII. fig. 11.)
Male. Upperside purple-grey, traversed by short black strige; a
broad greenish-grey band crossing both wings, its outer border brown,
and angled before the costa and at the middle of hind wing ; inner
border curved; apex of fore wing with a brown and black patch,
and a blackish discal patch on hind wing below angle of the band.
Underside pale yellow traversed by several short brown strigz, crossed
by a narrow discal brown line, bordered outwardly with purple-
brown ; posterior angles of both wings greyish-white ; body beneath
and legs yellow.
Expanse 12 inch.
Moolai to Moolat, 4500 feet. In coll. J. Wood-Mason.
Omiza SCHISTACEA, n. sp. (Plate LIII. fig. 12.)
Male. Upperside vinous-grey: fore wing with two zigzag black
lines from the costa, terminating across end of the cell; a black
speckled spot below the cell: hind wing with a black wavy discal
line. Underside bright ochreous-yellow ; apex of fore wing purple;
antennee and palpi grey ; legs grey above, yellow beneath.
Expanse 13 inch.
Above Ahsown. In coll. J. Wood-Mason.
Fam. Fiponip.
CoryYMICA ARNEARIA.
Corymica arnearia, Walker, Catal. Lep. Het. B. M. xx. p. 231
(1860).
Ahsown.
TINOLEUS EBURNEIGUTTA.
Tinoleus eburneigutta, Walker, Catal. Lep. Het. B. M. iii. p. 621.
Moolai, 3000-6000 feet.
Fam. Patyap2.
EUMELEA ROSALIA.
Phalena rosalia, Cramer, Pap. Exot. iv. pl. 368. f. F.
Above Ahsown.
Fam. Macarip&.
MACARIA NORA.
Macaria nora, Walker, Catal. Lep. Het. B. M. xxiii. p. 934.
Moolai, 3000-6000 feet.
852 MR. F. MOORE ON LEPIDOPTEROUS [ Nov. 5,
Fam. ACIDALID&.
BITHIA EXCLUSA.
Bithia exclusa, Walker, Catal. Lep. Het. B. M. xxii. p. 320.
Ahsown.
ACIDALIA ATTENTATA.
Acidalia attentata, Walker, Catal. Lep. Het. B. M. xxii. p. 734
(1861).
Ahsown.
Fam. Micronip2.
Micronia GRAMMEARIA.
Micronia grammearia, Geyer, Hiibn. Zutr. p. 36, f. 761.
Hatsiega ; Taoo, 3000-5000 feet ; Moolai, 3000-6000 feet.
MICRONIA ACULEATA.
Micronia aculeata, Guén. Phal. ii. p. 26, pl. 13. f. 8 (1857).
Moolai, 3000-6000 feet.
MicRONIA VAGATA.
Micronia vagata, Moore, Proc. Zool. Soc. 1877, p. 622, pl. 60.
f. 18.
Ahsown; above Ahsown ; Moolai, 3000-6090 feet.
Fam. ZERENID2.
Porera, n. gen.
Male. Fore wing somewhat trigonal; costa arched at base; apex
convex ; exterior margin oblique ; costal vein extending to near apex ;
subcostal vein four-branched, first and second branches before the end
of the cell, second bitid, fourth from the end of the cell; discocellulars
bent inward, radial from their middle; median vein three-branched ;
submedian extending to above posterior angle. Hind wing oval ;
costal vein extending to apex; subcostal vein two-branched from
the end of the cell; other veins as in fore wing. Body slender ;
antennz minutely pectinated ; palpi very small, slender; legs slender,
squamose.
PoTeRA MARGINATA, n. sp. (Plate LIII. fig. 9.)
Male. White; cilia black: fore wing with the costal and exterior
borders narrowly and irregularly black-bordered ; a very large black
spot also at the end of the cell, a smaller spot. on the disk beyond,
another near base of posterior margin, and a curved streak at the
posterior angle: hind wing with black marginal line and spots.
Body and legs yellow, spotted with black; antennee brown.
Expanse 12 inch.
Moolai, 3000-6000 feet. In coll. J. Wood-Magon.
— =
Se
1878. } INSECTS FROM UPPER TENASSERIM. 853
Tabular List showing the Geographical Distribution of the Lepidoptera
of Tenasserim.
(Silhet, Khasias).
| Sikkim, Himalayas.
| Western Yunan.
| Siam.
alay Peninsula
or Penang.
cutta district).
| M
| Java.
| astern Bengal
S. India (W. Ghats).
Sumatra.
| Andamans.
| Lower Bengal (Cal-
| Ceylon.
| Upper Tenasserim.
| Pegu.
PAPILIONES.
Nymphalide.
DANAIN#.
ae
*
Danais plexippus ...............:02+000+-
chrysippus ......... peeecneseneneue
—— melaneus .............000e000s piace
Ok OK OK
—— aglea............ ekvecsopessariobsesir
MUU HIIS ewee act cany gun oMpe wena
Septentrionis .......seceessesseeee
Salpinx rhadamanthus......... peonesas
PPASGHE NERS e 1 ecs'edccvaed- iis esleenges
—— MASONI, N. HP. «00... eee eeeeseeeees
—— Margarita ...........ceeceseeseeee
Trepsichrois midamus_ ...............
Crastia cupreipennis, n.sp. ......-..
Stictopleea grotei ...... ae ee ee
Euplea godartii
— aicathoé ..............
—— limborgii, n. sp. ......-.........6.
—— subdita, n. sp. ............seeeeee ee
*
*
*
*
*
*
*
*
*K KKK EHH HK KKK KK KK K
; . ar SED er neta
*
SaTyRINZ.
Lethe arcadia ...... Lecce Sere Reread alee
—— mekara ...... Bates een eee (bg ee
Sa FIAL tc ccivnpjes'vs ce coo ecdaeovnccsae eats ae
WBEHIEY 2. ccceseacdssnqoxsevs
eK OG
x KK HOKE
rm z
——— perseus ........00.. wef fee]
— blasius ......
—— anaxias ...
runeka ......
Oulapa mnasicles :
Yphthima methora ...................5. cal ee (eee a
— baldus .................. eeeinseden =A {PRE 9 allenic nsdless|<oa]eatle v6
Erites angularis, n. sp. ............... *
Elymnias tinctoria, n. sp. .......... .|*
—— VASUCOVA oe. eeeeeeseeeecseeeee eomeaeiecafe soul) 3 4
Dyctis leucocyma............-.. sees 5 Heel aera ial
Proc. Zoou. Soc.—1878, No. LVI. ‘ee 56
854 MR. F. MOORE ON LEPIDOPTEROUS [ Nov. 5,
Tabular List (continued).
d| fe | @g 2
€| Pe elele] fs E
S| ja 8/eSlals| |a. co)
2|-\Gsl/eels15| |e :
s| |e elo lalp ad al =
© 8-5 1 si |" o 8 ay St on
a aa = gd 3 n I Ay o s oo ae rc)
| (SESS (E(Slcle™| ls llsie
BSE Sle2e|BlEla Seles leis
E\Sio Pla “IH ho sig “lals|sloym
DidiS © BES Blal|dla
NyMPHALINE.
Amathusia phidippus ...... “chedadsocns | * Bare ve % |e]...
Zeuxidia Masoni, N. SP. ...eeereeeeeeee| *
Discophora tullia...... 303 .000B80 relat * | %
DOM weseubais icsseeeneespieess Seb ee|t wee) |! ami o®
—— necho ......-e-seee enc: I... woe | elacelee * *
PARTON Ay LOA eee ss ieseescceneasae se pee line) | Ree eco |isstena tee lO
Clerome arcesilatis .........:..eseeenres 6, ball acs. ee Ae le as eae
Xanthotenia busiris ......... | HJeee] cee * *
Thaumantis louisa .......... eas .|*
Cirrochroa surya, ni. Sp. «++... | *
Messaras erymanthis ........c.ssceeese| ¥ || eee | % [oselee % *
Atella simb@ .....cecsrecscesceseeceesessns nal Relocate ia ick] oer hia?
Junonia laomedia........sscoccsscvsceess|* | %*| * | * [¥/H)...| # 1 | % |
—— lemonias ...........seeeeee ees ecGeee| Ki [oen| HR A eee | #| *
orithyia ...... Deceasiceketsestairsinas Pe -|% | *
ALIA ech iaesece sco oseesneas laa SF [ee caer | sees eal pet eae .| > |
Precis iphita ......... nccideticnibi ic Sec oe H [one] woes 13°21 Fe bse 1 LYS |e oeatesel Fe ae
Doleschallia pratipa......cscseseeseeeeee| [eee] eee | vee fovfee[eee] He Jeee[eee]
Kallima limborgii, n. sp....---.-..s0005| ¥
Oyrestis thyodamas ......0.-...sessee eee ¥|e..| x [*)#]... *
cocles . > Ae aie ie ae *
WISE Jeeves wedavasstseecues .| * |. * | %
Herona angustata,n.sp. . *
Penthema darlisa, n. sp... .|%*
Parthenos apicalis, n. sp. Al
Lebadea attenuata, n. sp. . | *
Neptis adara, n. Sp. ... sss. |
—— meetana, . 8p. +--+. .| *
—— jumba ......eseeeeee By) fe bance eee (eed Bed ere peat %| x
adipala . | * *
—— plagiosa, n. sp. | accel Wi
Adolias lepidea............ Pet aaa posres |. % |*|%].. = *
satropaces ....... Bei cose saceee|
Parvata, ND. SP.........-eeeseeneeeee *
— discispilota, n. sp. ......... ere *
{PRISER cae oocoehoddudnceeneeraae..ce * |. x |*
taoocana, 0. Sp....... eaeceesecsoenes *
Charaxes marmaX ...........-44+ Bete e bec}} coor |! ce
Warpak ..ce.,eeccosessceseecesee ven] H [awe Cale
——— APN, D. BPs coerseseseceseraccereess *
dlesa, N. BP. .....-+-.- _Aeb0 SOB * ;
samatha, 1. BP. sscereseeseecerees * :
Prothoo tranChitteecrscnccr<cconee+..: * |. * |%
LipytuZina. :
Taibythea miyrrhe, .!...s00ese-ceceossees| W'loe.| coe PSH [H [ecefese] 6 beeefoeefene *
1878. ] INSECTS FROM UPPER TENASSERIM. 855
Tabular List (continued).
zie Ata a
#) 6 .| 3/8 | [2 g
oS Sere 81a 3 B=
@| |a-=| G4 /3) 5 ms oS
a| |e@S5/ay|8|5)] |. ee : :
8| |B3la sme! |8s g| |B
S| facies jel msl lslsl le
Se a 2 a :|.8
B| [85/55 /8/3| [eo] JS] 8l sis
SISIE Sie mlalalgiS als alse q
Be 5|2Cta| 8) sia SEE res
Plaid ff wea hlald|ols
Erycinide.
Zemeros flegyas ...........00c008 a eee |p * [el x *
Taxila fasciata, n. sp. Ree ood his
Abisara neophron .............seseeeee *|. H 1% | #
angulata, 1. BP. ......ceeeeeeeeees *
Lycenide.
Oastalius TOXUS.c:....2.0c00cevecseerenses DEER ar: | Gente tae’ */% x
Lampides alexis .......... etal [Maecae al vessel if ol Ged ica RCM [ace esd ocd A Ng
kandarpa .......... Hie] H | He foe * |e | *
—— el]pi8...........ceee eee DEB] rcieet | eer thd | ¥ | scl oe.| |
—— M]LANUS.....ccscrcceceeersseceresenee| * [eee] % | * | *]. -| | |
Hypolyczena lisias .......0.........006- *
WV MAE OFAS tic ae- Joe. soncerce ne tases * |. % | %
Deudorix suffusa, n. sp. ..........0.08- *
Thamala miniata, n. sp. ........-....0- *
Woxura, abyYMNUS ..........ceeeerescecee- ¥|%| %
Amblypodia taooana, n. sp. ......... *
Narathura moolaiana, n. sp. ......... *
Arhopala nakula, ..............0s-s00¢-: *}. % |e
vihara ...... Scnbe hoaoce Bacane see * meh Neate cletahlt oh
Surendra quercetorum ............064| * [++ | * || % * | %
Thaduka multicaudata, n. sp.......... “211 taal |
|
Papilionide. |
PIERINA. |
Nychitonia xiphia ... eal eb sae pte ceed | ag be eT ED [9
Terias hecabe #) oe | * Le] e- % |e fee] %
Bh etaNA <.05s.000060. 0.000000 CRoSEE 1.2] Bel hae neat fod fe
—— formosa ....... op se acectivctnaaeses' * |. taal ies | Bec] od piece Hea foe *
sebaiiyesceaesseasess : * |. *
Hronia lutescens ................0000000 Derlsh| hscewlb aso eee [eae we] #
Callidryas catilla ................c.eeeees a ee eo ee a ee a
BEDCALO ere itech cle slerdacaucicsesys-oss Hl *| | we [HLH] He [HL HL]
Hebomoia glaucippe «| % * | %*|%|). SA nee eee i
Ixias citrina, n. sp. .........., | *
pallida, ns Sp. ...0.c...00.sseceeeee] #
— moulmeinensis, n. sp. ......... *
Catophaga neombo ..................44. FB. bees: | Seal * |. *
HULIW Biancsseeseessnesnsscesd-ca%s Belge | e0F * |. coe |
agela, n. sp.
Appias lea .............
TAMAR. “ho «cacscoeros * | *| *]. bot
— dapha, n. sp.
amba ..... nigdpangcooe ae
RICE NINE Dc cceccaun ace ce secretes. ellie |r ey
MBGRHE incosscsawascopeae |
56*
856 MR, F. MOORE ON LEPIDOPTEROUS
Tabular List (continued).
Delias pasithoé ............4s-seeeeeee
indica. ...... eee don weicosistes Scetune
descombesi
Prioneris watsoni
——clemanthe ...... Pere cle shade wee.nets
PapmLioniIné.
Papilio telearchus ...........--..060++
Se HTH LN nga dhaodoeaneecadanaucene
—— MEGarus ......seceveees %.
ONPape, N. SP. ..2..--0e-
pammon .......0...
——aristolochie .....
— philoxenus .........,
doubledayi -........
helenus...
— alcmenor
— mahadeva, 0. 8p........-2..2.0000
—— androgeos...............
ZOICUCHIN Pectredtetact cece
SarPedOn 22... ce se0- ;
APAINEMINOM crscetcchas a teas+s6 446
— antiphates ........s.ccecesesseeeee
Leptocircus virescens ...... eeertateeree
Hesperide.
Ismene benjamini............6......s000
druna
Tagiades pralaya .......e0sses00.
—— meetana, 1. SP. ....seveeeee
Astictopterus diocles ................+-
subfasciatus, n. sp. .......
Pamphila bambuse ...............
masoni, 0. sp. .
mathias
Hesperia moolata, n. sp..............4.
Plesioneura aurivittata, n. sp. ......
albifascia, NW. SPs coecesscercecces--
Isoteinon subtestaceus, n. sp. .........
SPHINGES.
Macroglossa orientalis................45
= Tuiteatia: oes tevav dp acdesentcsosoose
x x x x x | Upper Tenasserim.
| Pegu.
mo] ee
Pes S
Leas
°
So | Wag
O05 BMY
Bela
Behe
a\e 4
Hela
o+v\jon
Beinn
6°? a
+H
*
*
*
*
*
*
*
*
* *
* *
*
*%
*
*
*
*
*
*
eee *
*
*
* *
*
*
x * * | Sikkim, Himalayas.
*
* | Western Yunan.
| Siam.
alay Peninsula
or Penang.
| M
[Nov. 5,
a
=
a
[oo
a| |F
- —
€/8| ./3
2/8] |
a/Sis\els
oil ae A
EIEVEIBIR
=} 3
Inq jO ln
aH * | %
* %/¥|*
ey Le (a La *
* i.) ele]
% |...| *
* |. Nie
et ed oe ¥
-
*
set ler *
ios |
Kile .| *
1878.] INSECTS FROM UPPER TENASSERIM.
Tabular List (continued).
Be
3
a
5
aS
Pa
ey
pis
BOMBYCES.
Syntomis grotei ...........ccesseeseeeee %, [ses
sladeni .......... muna emaeddas ssenm Ec a
HURUMBODT i) -ccancidstecneccun'sd -=ene] 36s om
Beee I IDEEA) sancise aVinsidnvite dese swoon dancin % |...
——— MASONI, 1. BP. ....cecsessseoeseeeee| ¥
—— disrupta, n. sp. ............c000 *
erINd as Ns. SP. s<c.csc-veccvessucess *
albifrons, D. Sp. ........-ss0ee+0.| ¥
Retina rubrivitta .......... Paeeqcnddenges % Joes
Histia papilionaria .............c00c00 Calis
Euschema militaris ................2006- %/...
CXCUMON; Ty BPs ..tccaveuconseddes *
aurilimbata, 1. Sp. .......eeeeeees *
—— horsfieldi ........5........:seceeeees * | %*
Nyctemera latistriga ...............66. *|%
Pitasila moolaica, n. sp. ...........066 ¥
Anagnia subfasciata.................0068 *|%
Peridrome orbicularis............... * |
Damalis plaginota .................506 ral ne
——— aleiphron .............00.ceseeneesee # |...
Hypsa subsimilis ...............:000se00s Cane
Neochera marmorea ...........6...5+. * | *
CHonistis entella ..............ceeeseeees *}...
Attatha regalis ...........0...cesssssceree *
Werina pasalis: ...3./3.2..-525.-Saaagty a * I...
PATON VALIBDS 5 cscecccccoesssosceet secve Co Bee
Dreata taooensis, 0. Sp. .........-0005 *
Trabala vishnu............. iste # Jee.
NOCTUES.
Potamophora manlia .................. eal
Nyetipao hieroglyphica ............... * |e.
CapTIMUlgus.......0.000secsereseees % [eo
Grepuscularis: .6.......c.c0e0..: 50 * |
Spiraea HElMCiHA,<z...ccasnc0sssseedeeees # |...
PALOMA AUPIITA ...0...2..saceccecaress # fee.
Entomogramma fautrix ............... ¥ |...
Fodina stola.............+ pedeante gt testes * |...
Calesia flabellifera, n. sp..............-5 *
gastropachoides .................- % [owe
PYRALES.
Zebronia plutusalis .................066 26
Glyphodes bivitralis .................. % |...
BEOUMIIS sau scncn athe <pg'eansn cee o0% #0.
Lower Bengal (Cal-
cutta district).
Kastern Bengal
eK KK
(Silhet, Khasias).
Sikkim, Himalayas.
| Western Yunan.
| Siam.
M
* *
* KK OK
aslewelone
alay Peninsula
or Penang.
ES
Q
2
a
a
o
lé|
Blglg}.8
é/SiSiSle
Pao ele
Qaisi/e;a;.,.
|n\4/O ja
*
*
.| *
*|...] %
#{...]...| # |
| ¥*
Nag * | *
Beales * | *
Seles va *
* |...) * || *
% |...] 4 | 9¢ |
5 eel Bee *
% |...) |
enfece] oe |
| *
oo |
*
sl |
crores
. | le
858 MR. F. MOORE ON LEPIDOPTEROUS INSECTS. [Noy. 5,
Tabular List (continued.)
' a |S =
e) 6 Zle.| (3 g
= Waa S| 81a = 4g
oOo 3s a) =!
a ra.-2| 8/5 e a, o
3 ws Bt 8/5 A 80 4
= Bm | Ot TS Ib ag a =
oO Qs > as} os -|q mot
A] Mele sicle| as a\s3! Ja
x] |e Sl\eS/gi8 pA] (8/8) sig
oO x O/C LAS S13 :|/Sia}o
Be|ems\a|/8/S x\s =)
&/S8 S|e cm )siale S|ESIS BH
PIAA Ff Beas Sas|oa
[ach dE ie Se a a ee
Pachyarches vertummalis ............ =a Fer | ecm | Ronee Geel eee 6 | *
marthesiusalis................-.++- H Jace] | coe leeefove 26 *
Sisyrophora pfeifferze ...........-...6+- Hl...| eos | ene | ¥
Euglyphis procopialis ..............66++ ok | fil Mere ea wl Ae es [ema ere *
Botys jopasalis ............ceesereeeees [ace H | enweilvee|emol oes] oeelm|atelelateat ane ME
PALGUBANISM Wpceantetterne ts nesarle H fe ee] cow | osandlit
ViNaCeAlIS © oo 2c. esseseesenses 5000] face|| teen! |nasac| sec fewelan oftldeatl| se) esha
————= UNIAN, veceescececscerscceseecsess |e] *
GEOMETRES.
Urapteryx podaliriata ............... |...) * | ¥
Hyperythra limbolaria ............... ME [bse | es en | cease [e's b soveeayl pina Mere Paea| cet ie
— angulifascia, n. Sp. ...........666 *
Corymica arnearia ...............0008+- lecall cos J | [aetless| weal rset hace tie
Omiza schistacea, n. 8p. ......0e00e-+0 *
Tinoleus eburneigutta .............6605- HE |ecalh cco.) wwet{e |nsslwaal igen estate ie
Humelea rosalia .......00.s.cssesssereee SEVEN cee 9/598 [owa|aGaleec) lanes leas eneleeebsce *
Miacaria Ot yreen.cas: oc) cede -ianenn acme Kilenal| cos |’ He ailves|ooel's oe} temmaleel Uae ise Ne
BEST OXC MINA iar ela ens ches cevae ses sane Mesa |) 'ecay [owes [wee|eeelcoslp eeetalens eels
Acidalia attentata <.............2s0.006 ME [Esclb reser || oe> ||seeleos|ens |aemeadl reat per nel |e
Micronia grammearia ................+- [baci cae, |) Seo aE scetess|| omeel oe
——— ACERT .....00eereevecvonceveosees Hleee|) H [ees [ees [occ|sce|l aden | el een ie lite
VELADA wen ceddeeseasehaevenspessecss- 2 Oe | ac (ie cead ee Bod Fase [ime esth Gos) sit 22
Potera marginata, n. sp. .........++ *
EXPLANATION OF PLATES LI.-LIII.
Puate LI.
Fig. 1. Papilio mahadeva, n. sp., p. 840.
2. Euplea limborgii, n. sp., p. 823.
3. Zethera diademoides, n. sp., p. 824.
Puare LI.
Fig. 1. Tavila fasciata, n. sp., p. 832.
2. Adolias discispilota, n. sp., p. 831.
8, Adolias parvata, nu. sp., p. 831.
4. Catophaga lagela, n, sp., p. 888.
5. Pamphila masoni, n. sp., p. 842.
6. Thamala miniata, un. sp., p. 834.
7. Thaduka multicaudata, n. sp., p. 836.
8. Deudorix suffusa, n. sp., p. 834.
Prats LITT.
Tagiades meetana, n. sp., p. 842.
. Plesioneura aurivittata, n. sp., p. 843.
albifascia, n. sp., p. 843.
. Syntomis masoni Jd, u. sp., p. 845,
disrupta, n. sp., p. 845.
Fig.
Sup co bor
fs
ry
4 Ft eine GT
P 2Z.5.1878-PLLIV.
GBSowerby hth Hanhart imp.
NEW SHELLS.
1878.] MR. G. F. ANGAS ON NEW BIVALYE SHELLS. 859
Fig. 6. Syntomis albifrons, n. sp., p. 845.
6
7. Dreata taooensis, n. sp., p. 848.
8. Syntomis berinda, n. sp., p. 845.
9. Potera marginata, n. sp., p. 852.
10. Pitasila moolaica ¢,n. sp., p. 847.
11. Hyperythra angulifascia, n. sp., p. 851.
12. Omiza schistacea 3, n.sp., p. 891.
8. Descriptions of six Species of Bivalve Shells in the Collec-
tion of Mr. Sylvanus Hanley, F.L.S., and of a Helix
from the Solomon Islands. By Grorer Frencu Ane@as,
C.M.Z.S., F.L.S., &c.
[Received September 19, 1878.]
(Plate LIV.)
SEMELE HANLEYI, un. sp. (Plate LIV. fig. 1.)
Shell transversely ovate, subequilateral, solid, cream-coloured,
tinged with violet below the umbones, with narrow irregular de-
scending white rays towards the base, with several zigzag purplish
markings near the centre of the valves; sculptured all over with
narrow prominent close-set, somewhat flattened, transverse ridges,
which become rather sharp and irregular posteriorly and erectly
scabrous at the margin, whilst radiating from the umbones close to
the anterior dorsal margin are six close-set ribs, upon which the
transverse ridges become nodulous at the points of intersection ; um-
bones prominent, sharp, approximate ; posterior dorsal margin nearly
straight, anterior one very slightly incarved, ventral margin acute ;
dorsal area stained with deep purple; valves slightly gaping and
flexuous posteriorly ; interior of the valves yellow in the middle,
stained with rays of violet and rose-colour.
Long. 11, alt. 8, lat. 4 lines.
Hab. Japanese seas (Belcher). Collection of Hanley.
A very charming shell; the sculpture of the anterior portion re-
minding one of certain species of Psammobia.
SEMELE APHRODITE,n. sp. (Plate LIV. fig. 2.)
Shell orbicular, nearly equilateral, rather thin, white, with the
posterior half of the umbones tipped with rose-colour, from which
descend four or five irregular bundles of more or less narrow, faint
pink rays, which become a little deeper in colour at the lines of
growth and near the base; concentrically sculptured with narrow,
thread-like, somewhat distant lamella, the spaces between which are
crossed with exceedingly fine, close-set, regular hair-like*descending
strize ; umbones rather tumid, sharp, approximate ; dorsal margin
posteriorly very slightly arched, anteriorly a little incurved ; dorsal
area freckled with a few reddish spots; ventral margin strongly
arcuate.
Long. 7, alt. 6, lat. 3 lines.
860 MR. G. F. ANGAS ON NEW BIVALVE SHELLS. [Nov. 5,
Hab. China seas? (Belcher). Coll. Hanley. :
An exquisitely sculptured species of great beauty, unique in the
collection of Mr. Hanley.
SEMELE ASPASIA, n. sp. (Plate LIV. fig. 3.)
Shell orbicularly ovate, nearly equilateral, subequivalve, mode-
rately thin, yellowish white, concentrically very finely and regularly
sculptured with close-set erect frilled laminz, the interstices being
crossed by fine thread-like descending strize that correspond to the
undulations of the transverse frills; concentric ridges smooth and
more distant at the umbones; interior of the valves very faintly
tinged with flesh-colour.
Long. 123, alt. 10, lat. 6 lines.
Hab. ? In coll. Hanley.
This species differs from S. phryne in being thinner, more ovate,
and less tumid in form, and in the transverse sculpture consisting of
elegantly undulating frilled erect laminz, whilst the interstices
between the descending strie are not squares but narrow paral-
lelograms.
V SEMELE PHRYNE, n. sp. (Plate LIV. fig. 4.)
Shell suborbicular, subequilateral, equivalve, somewhat tumid,
solid, white, very finely regularly concentrically sculptured with
close-set thread-like strize, the interstices of which are crossed
throughout by similar thread-like descending striz, giving the surface
of the valves a beautifully neat and regular reticulated appearance,
excepting at the umbones, where the concentric ridges are more
distant and the descending striz finer and nearly obsolete ; posterior
side very slightly truncate and sinuate below, with a nearly obsolete
flattened ridge extending upwards towards the umbo on the right
valve ; interior slightly tinged with flesh-colour towards the base.
Long. 15, alt. 14, lat. 8 lines.
Hab. ? In coll. Hanley.
A solid, white, somewhat tumid species, very regularly and finely
latticed throughout, the interstices taking the exact form of minute
squares.
LucINA CiTRINA, n. sp. (Plate LIV. fig. 5.)
Shell quadrately orbicular, very inequilateral, equivalve, rather
thin, primrose-yellow, paler towards the base, concentrically sculp-
tured with very fine thread-like striee, that become more distant as
they approach the base; lunule very small, depressed; umbones
tumid ; beaks small, incurved ; anterior side produced and somewhat
flexuous above ; interior of valves crenate at the edge, pale yellow.
Long. 4, alt. 33, lat. 24 lines.
Hab. ? Coll. Hanley.
A small species, of a peculiar primrose-yellow colour.
Lucina ROSEA, n. sp. (Plate LIV. fig. 6.)
Shell elongately orbicular, equivalve, inequilateral, a little tumid,
1878.] MR. G. F. ANGAS ON NEW MARINE SHELLS. 861
moderately solid, whitish tinged with pale rose-colour towards the
umbones ; encircled by close-set, subgranular, spreading, flattened
ridges, that become broader and closer towards the base; lunule
large, triangular, convex ; umbones tumid ; beaks small, pointed, in-
curved ; interior of valves pink, the edges crenate within.
Long. 8, alt. 8, lat. 5 lines.
Hab. Natal. In coll. Hanley.
In this pretty shell the sculpture of the valves approaches that of
L. pennsylvanica; and, as in that species, small fragments of grit
and shell are to be found lodging between the overlapping ridges.
HELIX BRENCHLEYI, un. sp. (Plate LIV. fig. 7.)
Shell imperforate, depressedly trochiform, rather thin, obliquely,
irregularly, and obscurely striated, yellowish white, with two dark
chocolate bands, thickly crossed with white diaphanous zigzag
markings encircling the last whorl, the uppermost band the broadest ;
spire obtusely conical, apex white ; whorls 5, flatly convex, sutures
impressed, last whorl descending in front, base a little tumid ante-
riorly ; aperture oblique, subrhomboidal; peristome very slightly
thickened and expanded, and but little reflexed, the right margin
moderately sinuated; columellar margin somewhat flattened and
dilated, with a straight abrupt callus; the umbilical region and the
inner edge of the lip with the reflected portion brown, immediately
behind which is a band of pigment-like deposit of a brilliant orange-
colour.
Diam. maj. 11, min. 9, alt. 63 lines.
Hab. Ysabel Island; Solomon Group. Collected by the late Mr.
Brenchley.
A singular shell, having the orange colouring of H. doivini behind
the outer lip, but differing from that species in its other characters.
9. Descriptions of ten Species of Marine Shells from the
Province of South Australia. By Grzorcr FRrencu
Aneas, C.M.Z.S., F.L.S., &c.
[Received September 19, 1878.]
(Plate LIV.)
Mirra TATE!, n. sp. (Plate LIV. fig. 8.)
Shell ovately fusiform, solid, shining, ochraceous, with a broad
deep-chocolate-coloured band next below the sutures, and a second
similar band towards the lower part of the last whorl; whorls 53,
stoutly and distantly longitudinally ribbed, the ribs becoming
obsolete towards the base of the last whorl, which is encircled by three
or four transverse ridges ; aperture narrowly ovate ; outer Jip simple ;
columella 4-plaited.
Long. 33, lat. 14 lines.
862 MR. G. F. ANGAS ON NEW MARINE SHELLS. __[ Nov. 5,
Hab. Living, in two fathoms water, Surveyor’s Point ; among
shell-sand, Holdfast and Aldinga Bays and Salt Creek.
PARTHENIA GRACILIS, n. sp. (Plate LIV. fig. 9.)
Shell elongately turreted, moderately thin, white, regularly longi-
tudinally ribbed, the interstices crossed with very fine hair-like striz ;
whorls 73, slightly convex, contracted above and below the sutures ;
sutures strongly impressed; aperture quadrately ovate; outer lip
subarcuate, slightly angled above ; inner lip with a single sharp tooth
about the middle of the columella.
Alt. 24, diam. 3 line.
Hab. Shell-sand, Glenelg, St. Vincent’s Gulf.
CycLosTREeMA TATEI, n. sp. (Plate LIV. fig. 10.)
Shell widely and deeply umbilicated, orbicular, depressedly turbinate,
moderately thin, under the lens very finely transversely striated,
shining pearly white; whorls 4, rounded, flattened and slightly
excavated next below the suture, with one, or sometimes two, narrow
thread-like keels at the upper part, and strongly keeled round the
umbilical region ; sutures distinct ; apex rounded, obtuse ; aperture
subcircular ; lip simple.
Diam. maj. 13, min. 1, alt. 2 line.
Hab. Shell-sand, Holdfast Bay ; found alive in the estuary of
the Sturt.
The above is the normal condition of the species ; but examples
occur which are thinner and have a greater number of keels, some-
times as many as seven or eight. At first I was inclined to regard
the many-keeled variety as specifically distinct ; but on the exami-
nation of a large series by Professor Tate, he assures me that the
number of keels varies so greatly that it would be impossible to
separate them. The thin hyaline examples with many keels are
probably younger shells.
BuccINULUS INTERMEDIUs, n.sp. (Plate LIV. fig. 11.)
Shell elongately ovate, solid, shining, white, painted with two bands
of irregular descending brown flames and spots; spire acuminate,
pointed at the apex, the same length as the aperture; whorls 6,
encircled by numerous grooved and finely punctured strie, that
become obsolete on the centre of the last whorl; sutures strongly
impressed ; outer lip simple, thin, non-arcuate ; columella with a
strong bilobed fold near the base, and a smaller projecting plate
above it; inner lip with a broad spreading callus.
Long. 43, lat. 13 line.
Hab. Aldinga Bay.
This, the only species of Buccinulus as yet discovered in South
Australia, is allied to B. afinis, A. Ad., from New South Wales,
from which it differs somewhat in form, and also in the style of
coloration.
NACELLA PARVA, n. sp. (Plate LIV. fig. 12.)
Shell depressedly conical, cap-shaped, semipellucid, nearly smooth,
1878.] MR. G. F. ANGAS ON NEW MARINE SHELLS. 863
or with very fine concentric lines of growth, pale horn-colour, with
a single row of pale blue spots and crescent-shaped opaque markings
extending from the apex centrally, more or less along the outer are
of the shell; apex anterior, recurved, submarginal ; aperture nar-
rowly ovate, margin simple, entire ; interior pearly white, shining.
Diam. maj. 3, min. 13, alt. 1 line.
Hab. Holdfast Bay and Aldinga Bays; parasitic on seaweed.
MYSELLA DONACIFORMIS, n. sp. (Plate LIV. fig. 13.)
Shell quadrately cuneate, moderately solid, equivalve, very inequi-
lateral, subventricose, compressed towards the base, white, shining,
finely concentrically ridged ; dorsal margin slightly arcuate posteriorly,
short and abruptly descending anteriorly ; ventral margin a little
convex ; umbones somewhat tumid; beaks distinct, incurved,
approximate.
Long. 3, alt. 2, lat. 1 line.
Hab. Holdfast and Aldinga Bays, St. Vincent’s Gulf.
In the ‘Proceedings’ of this Society for 1877, page 176, I
gave the diagnosis of the genus Mysel/a, for the reception of a
small bivalve from Port Jackson, which I described as Mysella
anomala. I have subsequently received from Professor Tate one or
two examples of the same species from South Australia, together
with a second form of the genus, of a more cuneate shape than the
type, now described above as M donaciformis.
LEPTON AUSTRALIS, n. sp. (Plate LIV. fig. 14.)
Shell quadrately orbicular, compressed, equivalve, nearly equi-
lateral, thin, very slightly gaping at the sides, white, very finely
concentrically striated ; dorsal margin arched, ventral margin nearly
straight ; beaks small, produced and acute.
Long. 4, alt. 3, lat. 3 line.
Hab. Glenelg, and Port Creek.
Belonging to the typical form of the genus, with the surface of
the valves finely striated but not shagreened.
Lucina (CopaxktA) TATEI, n. sp. (Plate LIV. fig. 15.)
Shell quadrately ovate, subventricose, moderately solid, subequi-
lateral, white, concentrically ridged, the ridges closer near the
umbones, and more distant towards the base, and crossed on both
sides with radiating ribs, that become slightly nodulous at the inter-
sections ; dorsal margin deeply excavated in front, nearly straight
and rapidly descending behind; ventral margin arcuate, rounded in
front, forming an obtuse angle where it joins the dorsal excavation ;
umbones very acute, incurved and approximate.
Long. 4, alt. 3, lat. 1? lines.
Hab. East side of St. Vincent’s Gulf, and Cape Northumberland.
This little species is quite distinct from Z. guadrata, Angas
(P. Z.S. 1877, p. 176), from Port Jackson, although of a somewhat
similar form.
864 MR. G. F. ANGAS ON THE [Nov. 5,
Nucuta micans, n. sp. (Plate LIV. fig. 16.)
Shell minute, obliquely and triangularly ovate, subventricose,
very inequilateral, moderately solid, white, shining, very finely con-
centrically striated, the strize occasionally running into each other ;
dorsal margin somewhat arched posteriorly, short and descending in
front ; ventral margin moderately arcuate ; umbones tumid, smooth,
pearly, and approximate.
Long. 1, alt. 2, lat. $ line.
Hab. Shell-sand, Salt Creek; Glenelg, St. Vincent’s Gulf.
Somewhat ailied to Nucula pusilla, Ang. (P. Z. 8. 1877, p. 177),
from Port Jackson, but possessing a different style of sculpture.
EXPLANATION OF PLATE LIV.
Fig. 1. Semele hanleyi, p. 859. Fig. 9. Parthenia gracilis, p. 862.
2. aphrodite, p. 859. 10. Cyelostrema tatet, p. 862.
3 aspasia, p. 860. 11. Buccinulus intermedius, p. 862.
4, phryne, p. 860. 12. Nacella parva, p. 862.
6. Lucina citrina, p. 860. 13. Mysella donaciformis, p. 863.
6. rosea, p. 860. 14. Lepton australis, p. 863.
7. Helix brenchleyi, p. 861. 15. Lucina (Codakia) tatet, p. 863.
8. Mitra tatei, p. 861. 16. Nucula micans, p. 864.
10. A List of additional Species of Marine Mollusca to be in-
cluded in the Fauna of the Province of South Australia ;
with Notes on their Habitats and Local Distribution.
By Grorcr Frencu Aneas, C.M.Z.S., F.L.S., &c.
[Received September 19, 1878.]
In the year 1865 I published in the ‘ Proceedings’ of this Society
a list of all the species of marine Mollusks that had been met with on
the shores of the Province of South Australia up to that date, num-
bering about 231 Gasteropods and 97 Conchifera. Since that period
other species have come to light, several of them hitherto only known
as inhabiting the New-South-Wales and Tasmanian seaboards ; whilst
others have proved to be new to science, and have been described by
me recently. I am indebted to Professor Ralph Tate and Mr. W.
T. Bednall, of Adelaide, for several interesting additions to my list—
which includes 54 Gasteropoda, and 21 Conchifera, &c., hitherto
unrecorded as occurring in South Australia, thus making a grand
total of 403 species.
GASTEROPODA.
1, Murex ANGASI.
Typhis angasi, Crosse, Journ. de Conch. 1863, p. 86, pl. 1. fig. 2,
Port Lincoln (Bednall/).
2. Triton (EPIpROMUS) BEDNALLI.
Triton bednalli, Brazier, Proc. Linn. Soc. N. S. W. 1875, p. 6.
Guichen Bay (Bednal?).
1878. } MARINE MOLLUSCA OF AUSTRALIA. 865
3. CoMINELLA LACTEUM.
Buccinum lacteum, Reeve, Conch. Icon. pl. 14. fig. 117.
Port Elliot and Port Wallaroo (Stansbury).
4, NASSA JACKSONIANA.
Buccinum jacksonianum, Kiener, Mon. Buce. pl. 19. fig. 73.
Holdfast and Aldinga Bays (Bednall) ; also Port Jackson.
5. NASSA INTERMEDIA.
Nassa intermedia, Dunker, ‘ Novara’ Exp. Moll. tab. 1. fig. 2.
Port Lincoln (Bedna/l) ; also Port Jackson.
6. AMALDA MARGINATA.
Ancillaria marginata, Lam., Reeve, Conch. Icon. pl. 3. fig. 8.
Port Lincoln; South Coast (Bednall). Also Tasmania and New
South Wales.
7. MELO MILTONIs.
Melo piltonis, Gray, Grif. An. King. (1833).
Fowler’s Bay, to the westward of Port Lincoln.
8. Mirra PICA.
Mitra pica, Reeve, P. Z.S. 1845 ; Conch. Icon. pl. 31. fig. 247.
Port Elliot ; Kangaroo Island (Bednall).
9. MiTRA LINCOLNENSIS.
Mitra lincolnensis, Angas, P. Z.S. 1878, p. 313, pl. 18. figs.
10, 11.
Port Lincoln, Semaphore Beach ; and Salt Creek.
10. Mirra SCHOMBURGKI.
Mitra schomburgki, Angas, P.Z.S. 1878, p. 313, pl. 18. figs.
12, 13.
St. Vincent’s Gulf.
11. Mrrra TATEI.
Mitra tatei, Angas, P. Z.S. 1878, pl. 54. fig. 8.
Found alive in two fathoms of water, Surveyor’s Point; among
shell-sand, Holdfast and Aldinga Bays, and Salt Creek (Tate).
For Columella dermestoides, Kien. (No. 65 in my first list), read
C. lineolata, Pease. Port Lincoln and Port Elliot.
For Columella interrupta, Angas (specific name preoccupied),
read C. angasi, Brazier. York’s Peninsula.
12. MARGINELLA VOLUTIFORMIS.
Marginella volutiformis, Sow. Thes. Conch.
Macdonnell Bay (Tate).
866 MR. G. F. ANGAS ON THE [ Nov. 5,
13. MARGINELLA TURBINATA.
Marginella turbinata, Sow. Thes. Conch.
Macdonnell Bay ; also New S. Wales.
14. CryprosPIRA CYMBALUM.
Cryptospira cymbalum, Tate, Proc. Adelaide Phil. Soc. 1878.
Aldinga. Like C. ovulum, with eight plaits on the pillar.
15. HyaLiIna ALBIDA.
Hyalina albida, Tate, Proc. Adelaide Phil. Soc. 1878.
Marino. Very narrow and white.
16. Erato BIMACULATA.
Erato bimaculata, Tate, Proc. Adelaide Phil. Soc. 1878.
Minute, yellowish white, with a red spot at each end of aperture.
17. NATICA MAROCHIENSIS.
Natica marochiensis, Lam., Reeve, Conch. Icon. pl. 13. fig. 52.
Very small specimens, living in St. Vincent’s Gulf. Subfossil in
the Port-Adelaide-Creek limestone (Tate). -
18. SCALA FRIABILIS.
Scalaria friabilis, Sow., Reeve, Conch. Icon. pl. 7. fig. 52.
Port Lincoln ; also Swan River.
19. Scata (OPALIA) AUSTRALIS.
Scalaria australis, Lam., Sow. Thesaurus, pl. 35. fig. 135.
Aldinga Bay (Bednall); also New S. Wales.
20. TEREBRA BRAZIERI.
Terebra brazieri, Angas, P. Z. S. 1871, pl. 1. fig. 15.
Port Elliot (Bednall); Brisbane Water, New S. Wales.
21. TeEREBRA USTULATA.
Terebra ustulata, Desh. P. Z. S. 1859, p. 294.
Fowler’s Bay ; Port Elliot (Stansbury). Also Tasmania.
22. OBELISCUS BRUNNEA.
Obeliscus brunnea, A. Ad., Sow. Thesaurus, pl. 171. fig. 35.
Syrnola brunnea, A. Ad. P. Z. 8. 1862, p. 233.
Port Lincoln.
23. SYRNOLA TINCTA.
Syrnola tincta, Angas, P. Z. 8. 187], pl. 1. fig. 11. °
St. Vincent’s Gulf; also Port Jackson.
24. ODOSTOMIA EBURNEA.
Odostomia eburnea, Metcalfe, M.S. in Brit. Mus.
Holdfast Bay (Tate).
A very small, ovate, ivory-white species.
1878. ] MARINE MOLLUSCA OF AUSTRALIA. 867
25. PARTHENIA GRACILIS.
Parthenia gracilis, Angas, P. Z. 8. 1878, pl. 54. fig. 9.
Shell-sand, Holdfast Bay (Tate).
26. TURBONILLA ACICULARIS.
Turbonilla acicularis, A. Adams.
St. Vincent’s Gulf (Tate).
27. TuRBONILLA FUSCA.
Turbonilla fusca, A. Adams.
Holdfast and Aldinga Bays; Port Lincoln (Tate).
28. TuRBONILLA HOFMANI.
Turbonilla hofmani, Angas, P. Z. S. 1877, p. 183.
-T. nitida, Angas, P. Z. 8. 1867, p. 112 (name preoccupied by A.
Adams: T.. nitida, from Japan, Ann. Nat. Hist., vi. p. 419 (1860).
T. angasi, Tenison-Woods, Proc. Roy. Soc. Tasmania, Mar. 13,
1877.
Holdfast and Aldinga Bays (Tate) ; also Port Jackson.
29. TRIVIA AUSTRALIS.
Cyprea australis, Lam.; Sow. Conch. Illus. fig. 29.
Fowler’s Bay ; Cape Northumberland (Tate and Bednall). Also
New-South-Wales coast.
30. DiaALA IMBRICATA.
Diala imbricata, A. Adams, Ann. & Mag. Nat. Hist. Oct. 1862.
Shell-saud, Glenelg and Aldinga Bay (Tate).
31. Dia VARIA.
Diala varia, A. Adams, Ann. & Mag. Nat. Hist. Oct. 1861.
Between tide-marks, under stones, Aldinga Bay (Tate).
32. D1aua PICTA.
Diala picta, A. Adams, Ann. & Mag. Nat. Hist. Oct. 1861.
St. Vincent’s Gulf; and Wallaroo, Spencer’s Gulf (Tate).
33. RIsSOINA NIVEA.
Rissoina nivea, A. Adams, P. Z. S. 1851, p. 265.
Port Lincoln.
34. TROCHITA CALYPTREFORMIS.
Trochus calyptreformis, Lam. Anim. sans Vert. vii. p. 627.
Crepidula tomentosa, Quoy et Gaim.
On Pinne, Holdfast and Aldinga Bays (Bednall). Also NewS.
Wales.
35. TRIPHORIS SCITULUS.
T. scitulus, A. Adams.
St. Vincent’s Gulf ( Bednall).
868 MR. G. F. ANGAS ON THE [Nov. 5,
36. AMATHINA TRICARINATA.
Amathina tricarinata, Gray, Woodward’s Manual, pl. 11. fig. 19.
Found dead on the beach at Port Lincoln.
37. CyCLOSTREMA TATEI.
Cyclostrema tatei, Angas, P. Z. S. 1878, pl. 54. fig. 10.
Shell-sand, Holdfast Bay ; alive in the estuary of the Sturt ee
38. ScHISMOPE ATKINSONI.
Scissurella atkinsoni, Tenison-Woods, Proc. Roy. Soc. Tasmania,
1876.
Holdfast and Aldinga Bays (Tate).
39. FisSURELLIDHA SCUTELLA.
Fissurella scutella, Gray, Brit.-Mus. Cat. (Fissurella), No. 42;
Sow. Thes. Conch. pl. 9. fig. 207 (Trapezina).
Guichen and Holdfast Bays (Bednall). Also Tasmania, New South
Wales, and Cape of Good Hope.
40. MAcROSCHISMA TASMANLE.
M. tasmania, Gray.
South-east coast (Bednall); Tasmania (JT. Woods).
Emarginula tasmaniez, Sow., and E. tenuicostata, Sow., are re-
garded, and, I believe, rightly, by Rev. Tenison-Woods as varieties
of EZ. rugosa of Quoy and Gaimard.
41. EMARGINULA DILECTA.
E. dilecta, A. Adams, P. Z. 8. 1851.
St. Vincent and Spencer’s Gulfs.
42. NacELLA PARVA.
Nacella parva, Angas, P. Z.S. 1878, pl. 54. fig. 12.
Holdfast and Aldinga Bays, parasitic on seaweed (Tate).
43. DENTALIUM OCTAGONUM?
D. octagonum, Lam.
Henley Beach.
44. CaDULUS ACUMINATUS?
C. acuminatus, Desh., M.S. in coll. Cuming.
Holdfast and Aldinga Bays (Tate); also Port Jackson.
45. LEPIDOPLEURUS ANTIQUUS,.
L. antiquus, Reeve, Conch. Icon. pl. 25. fig. 169.
Aldinga Bay, and Marina (Bednall); also Port Jackson.
46. CHITON SICULOIDES.
Chiton siculoides, Carpenter.
Port Lincoln (Bednall).
1878. ] MARINE MOLLUSCA OF AUSTRALIA. 869
47. BucctNuLUs INTERMEDIUS.
B. intermedius, Angas, P. Z. S. 1878, p. 869, pl. 54. fig. 11.
Aldinga Bay (Tate).
48. TorNATINA FUSIFORMIS.
T. fusiformis, A. Ad., Sow. Thes. Conch. pl. 121. fig. 37.
Glenelg Beach (Bednall); also Port Jackson and Port Stephen,
New S. Wales.
49. VoLVULA ROSTRATA.
V. rostrata, A. Adams, Sow. Thes. Conch. pl. 125. fig. 154.
Holdfast Bay (Tate).
50. CyLICHNA PYGMZA.
C. pygmea, A, Adams, Sow. Thes. Conch. pl. 125. fig. 150.
Holdfast Bay (Tate).
51. ATYS EXIGUA.
Atys exigua, A. Adams, Sow. Thes. Conch. pl. 125. fig. 129.
Holdfast Bay (Taée).
52. ALEXIA MERIDIONALIS.
A. meridionalis, Brazier, M.S.
Port-Adelaide Creek (Bednal/).
53. MARINULA XANTHOSTOMA.
M. xanthostoma, H. & A. Adams, P. Z.S. 1854, p. 35.
York’s Peninsula, Aldinga, &c. (Bednall),
54. SIPHONARIA ALBIDA, >)
if.
Stphonaria albida, Angas, P. Z. 8S. 1878, p.869, pl. 18. figs. 14, 15.
Port Lincoln.
CONCHIFERA.
55. HuMPHREYIA STRANGEI.
Aspergillum strangei, A. Adams, P. Z. 8. 1852, p. 91, pl. 15. fig. 5.
St. Vincent’s Gulf (Bednall). Also Port Jackson, New S. Wales
(Angas); Tasmania (7. Woods).
56. GASTROCHANA TASMANICA.
G. tasmanica, Tenison- Woods.
Burrowing in limestone forming the bed of Port-Adelaide Creek ;
also on Chame; and in the incrusted growth of Pinna, St. Vincent’s
Gulf (Tate).
57. CORBULA NASUTA.
Corbula nasuta, Sow. P. Z.S. 1833, p. 35.
Holdfast and Aldinga Bays (Tate); also New S. Wales.
‘Proc. Zoo. Soc.— 1878, No. LVII. 57
870 ON THE MARINE MOLLUSCA OF AUSTRALIA. [Nov.5,
58. Donax soRDIDUS.
D. sordida, Reeve, Conch. Icon. pl. 4. fig. 32.
Port Lincoln.
59. MySELLA ANOMALA.
Mysella anomala, Angas, P. Z.S. 1877, p. 176, pl. 26. fig. 22.
Holdfast and Aldinga Bays (Tate); also Port Jackson, New S.
Wales.
60. MysELLA DONACIFORMIS.
Mysella donaciformis, Angas, P. Z. 8. 1878, p. 863, pl. 54. fig. 13.
St. Vincent’s Gulf (Tate).
61. Tapes FLABAGELLA.
Tapes flabagella, Desh., Reeve, Conch. Icon. pl. 13. fig. 66,
Aldinga and Holdfast Bays (Tate); also New South Wales.
62. Carone (Marcia) L&VIGATA.
Venus levigata, Sow. Thes. Conch. pl. 159. figs. 156-8.
Port-Adelaide Creek (Bednall). New S. Wales, Victoria, and
Tasmania. :
63. TIvELA UNDULOSA.
Venus undulosa, Lam. Anim. sans Vert. vi. p. 370.
Venus variabilis, Sow. Thes. Conch. Venus, pl. 158. figs. 142-6.
Semaphore Beach, Guichen Bay, and Port Lincoln (Bednald).
64. CARDIUM PULCHELLUM.
C. pulchellum, Reeve, Conch. Icon. pl. 8. fig. 42.
Dredged in St. Vincent’s Gulf (Tate); also Port J ackson and Tas-
mania. (Very small specimens.)
65. Lucina TATE.
Lucina (Codakia) tatei, Angas, P, Z. 8. 1878, p. 863, pl. 54. fig. 15.
East side of St. Vincent’s Gulf, and Cape Northumberland (Ta¢e).
66. LorIPEs OvUM.
Lucina ovum, Reeve, Conch. Icon. pl. 5. fig. 21.
Holdfast Bay and Salt Creek (Zate); also Botany Bay, New 8.
Wales.
67. LASEA AUSTRALIS.
Poronia australis, Souv. Journ. de Conch, 1863, pl. 12. fig. 8.
Under stones, Aldinga and Holdfast Bays, Cape Northumberland,
and Wallaroo. Common in Tasmania, and also in New S. Wales.
68. LEPTON AUSTRALIS.
Lepton australis, Angas, P. Z. S. 1878, p. 863, pl. 54. fig. 14,
Glenelg and Port Creek (Tate).
1878. ] MR. G. E. DOBSON ON MYXOPODA AURITA. 871
69. CRENELLA BARBATA.
Modiolaria barbata, Angas, P. Z. S. 1867, p. 911, pl. 44. fig. 12.
St. Vincent’s Gulf &c.; also Botany Bay, New S. Wales.
70. TRIGONIA MARGARITACEA.
Trigonia margaritacea, Lam., Reeve, Conch. Icon. pl. 1. fig. 3.
Port Lincoln; dead valves washed upon Semaphore Beach (Bednal/).
Also Tasmania.
71. PecrUNCULUS LATICOSTATUS.
P. laticostatus, Quoy et Gain. Voy. de l’Astrol.; Reeve, Conch.
Icon. pl. 2. fig. 8 a, 8.
Cape Jervis; and Hallet’s Cove, St. Vincent’s Gulf (Tate); Badger
Island, Bass’s Straits (Tate). New Zealand.
72. Limopsis MACGILLIVRAYI.
Limopsis macgillivrayi, A. Ad. in Coll. Cuming.
Guichen Bay (Bednall). Semifossil at McDonnell Bay (Tate).
73. NUCULA MICANS.
Nucula micans, Angas, P. Z.S. 1878, p. 864, pl. 54. fig. 16.
Shell-sand, St. Vincent’s Gulf (Tate).
74. Puacunanomia (Monta) Ione.
Placunanomia ione, Gray, P. Z.S. 1849, p. 123.
Henley Beach ; Marino (Bednall). Common in New South Wales.
75. KRAUSSIA LAMARCKIANA.
Kraussia lamarckiana, Davidson, P.Z.S. 1852, p. 80, pl. 14.
figs. 22, 23.
St. Vincent’s Gulf (Tate). Common in N. S. Wales and Tasmania,
also New Zealand.
11. Notes on Myxopoda aurita, Miln.-Edw.
By G. E. Doxsson, M.A., M.B., &c.
[Received October 21, 1878.]
Among the many valuable specimens lately added to the collection
of the Muséum d’Histoire Naturelle at Paris, perhaps the most
remarkable is that of a very peculiar species of Bat from Madagascar,
described by M. Alphonse Milne-Edwards’ under the appropriate
name of Myzopoda’* aurita.
During a short visit to Paris last month I was enabled, through
the kindness of M. Milne-Edwards, to examine the type of this most
interesting species, of which the following is a description.
1 In Bull. Soc. Philom. de Paris, June 22, 1878.
2 uwvEa, mucus; zrods, pes.
57*
872 MR. G. E. DOBSON ON Myxopopa auriTA. __[Noy. 5,
Crown of the head but slightly raised above the face-line ; muzzle
obliquely truncated, in general form closely resembling that of the
species of the genus Chilonycteris; for the nostrils open widely
apart by similar circular sharply defined margins, and the lower lip
is also papillate and reflected outwards, though not so broadly, and
it has not a thin free margin; the obtuse extremity of the rather
long muzzle projects in front considerably beyond the lower lip.
Ears very large, much longer than the head, in general outline like
those of Vespertilio murinus; but the inner margin commences in a
small lobe projecting downwards ; in the usual position of the tragus,
or slightly in front of it, there is an irregularly square lobe con-
tinuous above with the keel of the ear-conch; opposite this, on the
outer side of the concave surface of the conch, is a mushroom-shaped
process consisting of a short stalk supporting a broad flat reniform
expansion; the outer margin of the conch terminates near the angle
of the mouth.
Thumb with an ill-developed claw; but the whole of the inferior
surface of its metacarpal and phalangeal bones supports a large flat
horseshoe-shaped pad, more than 0':2 inch in diameter, of which
the circular margin is directed forwards and slightly notched in
front; this pad, though considerably larger*than that occupying
the same position in Thyroptera tricolor, does not form nearly so
perfect a clinging-organ as in that species; for it is quite sessile,
without a trace of a peduncle, and its surface is flat, like the thumb-
pad of Vesperugo pachypus, with which it is evidently homologous.
The feet have also adhesive pads; but, while resembling closely
those of the thumbs in structure and external form, they differ in
being much smaller, and so agree rather with those of Thyroptera
tricolor than with Vesperugo pachypus.
The metacarpal bone of the index finger is well developed, being
nearly as long as that of the third finger, but there are no distinct
phalanges; the third finger has ¢iree phalanges, of which the first
and second are nearly equal in length.
The tail projects beyond the posterior margin of the interfemoral
membrane as in Thyroptera tricolor, but to a much greater extent,
the free portion being nearly equal to the tibia in length; the
calcanea are almost as long as the tibiee; the very narrow post-
calcaneal lobe is notched or toothed near the foot.
As in Thyroptera tricolor, the toes are united as far as the base of
the claws, and have each two phalanges, and the wing-membrane
extends almost to the base of the claws.
aA 2-2)) Jal 33 3—3
Dentition.—Inc. —}, c. > pm. x= M. 5.
Upper incisors short, in pairs, placed close to the canines; the
outer incisor, on each side, small, conical, and acutely pointed, but
much larger than the inner one, which lies close to it and is hardly
visible without the aid of a lens; lower incisors short and blunt, in
the direction of the jaws; first and second upper premolars very
short, the third exceeding the molars in vertical extent; second
lower premolar minute, in the tooth-row, the first premolar slightly
1878. ] ON A COLLECTION OF CHIROPTERA. 873
smaller than the third; molars acutely tubercular, with W-shaped
cusps.
Length (of the type specimen, an adult male), head and body
2-3; tail 1-9, tail free from membrane 0’"6; head 0'°85; ear
1"°3, tragus 0°25; forearm 1°85; thumb 0:3; third finger—
metacarp. 1'°5, lst ph. 0'*7, 2nd ph. 0'°75, 3rd ph. 0°55; fifth
finger—metacarp. 1"°5, Ist ph. 0'°5, 2nd ph. 0'°5; tibia 0'*7;
calcaneum 06; foot 0'°3.
Certain peculiarities in the structure of this very remarkable
species recall similar peculiarities in Thyroptera tricolor, and have
evidently resulted from adaptation to the same purpose. Thus in
these two species alone are the toes united to the base of the claws, and
in them alone, among all known species of Bats (except the Phyllo-
rhinine), have the toes an equal number of phalanges ; they also, in
the possession of a third phalanx in the middle finger, differ from
all the species of Vespertilionide, and from those of the allied
families. This species, however, differs remarkably from Thyroptera
tricolor in the structure of the adhesive disks, in the presence of a
well developed metacarpal bone of the second finger, in the form of
the head and ears, and in dentition, and must undoubtedly be con-
sidered the type of a distinct genus of Vespertilionide.
As remarked above, the adhesive pads are evidently less perfect as
clinging-organs than the corresponding parts in Thyroptera tricolor,
and occupy, in this respect, an intermediate position between those
of that species and of Vesperugo pachypus.
It is probable that this species (in common with the few other
species of Bats provided with such accessory clinging-organs) uses
the adhesive pads in sustaining its hold on the smooth hard stems
and leaves of palms and of other hard-wooded trees'.
12. Notes on recent Additions to the Collection of Chiroptera
in the Muséum d’Histoire Naturelle at Paris, with
Descriptions of New and Rare Species. By G. E.
Dosson, M.A., M.B., &e.
[Received October 28, 1878. ]
Through the kindness of M. Alphonse Milne-Edwards I have
lately been permitted to examine and describe the valuable additions
to the collection of Chiroptera in the Muséum d’ Histoire Naturelle,
made by various collectors since my last visit to Paris in 1876.
The collections, from which the specimens which form the sub-
ject matter of this paper were derived, were made chiefly in the
islands of New Guinea and New Caledonia, in Siam, in Africa and
Madagascar, and in Central America. The species may therefore
1 See my ‘Catalogue of the Chiroptera in the Collection of the British
Museum,’ p. 444; also ‘ Nature,’ yol. xviii. p. 589.
874 MR.G.E. DOBSON ON RECENT ADDITIONS TO THE (Nov. 5;
be conveniently arranged for description and remarks under the heads
of the four zoological regions thus represented.
1. Australian Region.
PTEROPUS GERMAINI, N. Sp.
Ears shorter than the muzzle, concealed by the long fur of the
head, triangular, obtusely pointed, thinly clothed throughout with
soft hairs. Fur long and woolly, like that of Pt. aneiteanus ; on the
back long like that of the head, directed backwards. Humerus
and forearm rather thinly covered with straight fur, like that of the
back. The legs are clothed with long fur which extends to the backs
of the feet; the margin of the wing membrane above, almost as far
outwards as the extremity of the fifth finger, is clothed with straight
appressed hairs; the posterior margin of the narrow interfemoral
membrane is quite concealed.
Face in front of and immediately above the eyes light greyish
brown; head and the whole inferior surface of the body dark
blackish brown, interspersed with several shining greyish hairs, the
shoulders and back darker, the rump and legs greyer; upper surface
of the neck and shoulders pale yellow, with reddish extremities.
Teeth simple, like those of Pt. medius ; the first upper premolar
small, scarcely raised above the level of the gum, and occupying the
centre of the small space between the canine and second premolar ;
last upper molar slightly larger than the first lower premolar, and
about the size of the last lower molar.
Length (of a not quite adult 9), head and body about 6";
head 2"-3; ear 0-8; forearm 4-7; thumb 2/3; third finger—
metacarp. 3", Ist ph. 2'"5, 2nd ph. 3! 5; fifth finger—metacarp.
3", Ist ph. 1/5, 2nd ph. 1-35; tibia 2/2; foot 1!'*7.
Hab. New Caledonia.
This species resembles externally, to some extent, Pé. aneiteanus ;
but the very different form of the teeth at once distinguishes it. From
Pt. vetulus, inhabiting the same islands, it is distinguished by the
completely different colour of the fur, as well as by the absence of
transverse basal ridges in the molars and premolars.
The food of this species appears to consist, in part at least, of
figs, as I found portions of these fruits in the mouth of the typical
specimen. ‘
At the request of M. Milne-Edwards, who pointed out to me this
new species, I have much pleasure in connecting with it the name of
its discoverer, M. Germain.
PTEROPUS VETULUS, Jouan.
Besides the type, the Museum possesses several specimens of this
species, both male and female, from New Caledonia. These all more
or less resemble the type specimen in the colour and distribution of
the fur; and the few differences observable are unimportant.
Hab. New Caledonia ; Loyalty Isles,
1878.] CHIROPTERA IN THE MUS. D’HIST. NAT. PARIS. 875
Preropus KERAUDRENI, Quoy & Gaimard.
The specimens in the collection from New Caledonia have the
face, back, and inferior surface of the body of a much darker colour
than is usually observed in specimens of this species from other loca-
lities, and they agree in this respect closely with the types of Ptero-
pus vanicorensis, Q. & G., which, as I have indicated in my ‘ Cata-
logue of the Chiroptera,’ is evidently a variety only of Pt. kerau-
dreni.
Hab. New Caledonia, and most other islands of the Polynesian
subregion. (See my ‘ Catalogue of Chiroptera,’ p. 64.)
Preropus HYPOMELANUS, Temminck.
Several specimens of this species were obtained by M. Laglaize
at Amberbaki, New Guinea.
CYNOPTERUS SCHERZERI (Fitzinger).
Several specimens of this species, labelled “ N. Guinée, M. La-
glaize.” With the exception of that very aberrant form Cynopterus
latidens, described by me from a specimen from Morty Island, this
is the first instance of any species of this genus extending beyond
the Oriental region. It is strange that this species should extend
from the Nicobar Islands to New Guinea ; and it is just possible
that some mistake may have been made by the collector when label-
ling the specimens.
Harpyia CEPHALOTEs, Pallas.
An adult specimen from New Guinea, collected by M. Raffray.
This is the first example of this species obtained in the above-named
locality. ;
CEPHALOTES MINOR, n. sp.
Not half the size of adult specimens of C. peroni, but closely re-
sembling that species in general structure. The wing-membrane is
similarly attached only to the central line of the back along the spine,
and is also naked from the shoulders backwards; the head and
muzzle are of the same shape as in the larger species; but the ears
are less pointed.
e feet are much smaller than in very young specimens of CO. pe-
vroni; and the wing-membrane is attached to the outer toe, not to the
space between the toes as in that species ; it also extends further out-
wards, terminating opposite the second joint of the next toe.
The teeth are also slightly different : the upper incisors are wider
apart; the second upper premolar has not the prominent antero-in-
ternal basal cusp observed in C. peront; and the first lower premolar
scarcely rises above the gum.
The following Table exhibits the comparative measurements of
this species and of C. peroni :-—
876 MR.G.E. DOBSON ON RECENT ADDITIONS TO THE [Nov. 5,
C. minor. C. peronii.
cli head and body 0.06 ss eeyeen fs ANB 6:0
a (INL ake Ca eer ete 0°6 iss}
5 CACM kites Catt a3. 3a 6.2. aaseagerea nace ries 1°6 2°0
za CAT ee raya ceetk, 3, spect ape eeepc a ama eiieed 0°7 1°0
53 RONCHI Mcgee here dot nc. gis. cteaaale hee Ba) 53
as PUBS UOC on a oid Feed Beco ats 1°3 15
- third finger, metacarp. .......... 2°0 Shull
AS es Wer GBC. Diller cum epee 2 cule 1°5 2°35
33 eae aie ETE] Ril ge is Shea 1°9 3°4
me fifth HBBCT, AEtACAL ps nef ec. ane 2°8
os Se COUSE eee ccna near ae 1'] Mery
a “A My, CCM teed cal von eae 113 | 1:8
Ph tibia .. 2. Soke tees 1] 2°3
ae caleaneum Le ees Soi om AOR2G 0°5
53 HOOT ee Meee oe Ae Rs erence 0°8 12
Hab. Amberbaki, New Guinea.
PHYLLORHINA TRICUSPIDATA, Temminck.
Several specimens obtained by M. Raffray in New Guinea. This
is the first recorded instance of this species having been found in
that country, which, however, it might be expected to inhabit, as it
has been noted from Batchian, Amboyna, Morty, and Duke-of-
York Island.
PHYLLORHINA CERVINA, Gould.
Examples of this species from New Guinea, collected by M. La-
glaize.
NYCTOPHILUS TIMORIENSIS.
_ Nyctophilus timoriensis (Geoffroy), Dobson, Catal. Chiropt. p. 172
(1878).
Specimens from New South Wales collected by M. de Castelnau.
ScoTOPHILUS TEMMINCKII.
Vespertilio temminckii, Horsfield, Zool. Researches in Java
(1824).
Scotophilus temminckii, Gray ; Peters 5 Dabsan; Monograph Asiat.
Chiropt. p. 258 (1876).
Specimens of this very abundant and widely distributed species
from Celebes, collected by M. de la Saviniere.
VESPERTILIO ADVERSUS, Horsfield.
Specimens of this species, which extends from Siam to South
Australia, have been sent from New Guinea by M. Laglaize.
EMBALLONURA. RAFFRAYANA, 0. Sp.
Slightly larger than #. nigrescens from the same region, and
1878.] | CHIROPTERA IN THE MUS. D’HIST. NAT. PARIS. 877
agreeing with that species in the comparatively widely separated
nostrils, but resembling the species of the other section of the genus
in the projecting extremity of the muzzle, which extends consider-
ably beyond the lower lip; the ears also are much broader, and the
upper third of the outer margin of the conch is convex, not concave ;
the tragus is comparatively shorter and much broader, attaining its
Emballonura raffrayana.
greatest breadth above, where it is so broadly rounded off so as to
appear abruptly truncated ; the outer and inner margins are straight
or faintly concave.
Wings from the ankles or from the tarsi; feet much larger than
in Z. nigrescens ; calcanea about two thirds the length of the tibiz.
Fur above dark-brown, paler at the base; beneath paler through-
out. Wings nearly naked ; upper surface of the interfemoral mem-
brane thinly clothed as far as the extremity of the tail.
Teeth as in LZ. nigrescens, except that the first upper premolar is
smaller, and scarcely raised above the level of the gum.
Length (of an adult 3), head and body 1''-65; tail 0-5; head
0"-65; ear O58, tragus 0-2; forearm 1-55; thumb 0/25 ;
third finger—metacarp. 1-3, Ist ph. 0-4, 2nd ph. 0-65; fourth
finger—metacarp. 1'-1, Ist ph, 0-3, 2nd ph. 0-2; fifth finger—
metacarp. 1", Ist ph. 0'-38, 2nd ph. 0-15; tibia 0'-6; caleaneum
0-45; foot 0-3.
Hab. Gilolo Island.
At the request of M. Milne-Edwards, I have much pleasure in
connecting with this species the name of its discoverer, M. Raffray.
2. Oriental Region.
CYNONYCTERIS AMPLEXICAUDATA, Geoffr.
Cambodja (M. Harmand).
CYNOPTERUS MARGINATUS, Geoffr.
Singapore (M. de Castelnau).
EonycTEeRIs SPELZA, Dobson.
Java (M. de la Saviniére).
878 MR.G.E. DOBSON ON RECENT ADDITIONS TO THE [Nov. 5,
RHINOLOPHUS ACUMINATUS, Peters.
Siam, Laos (M. Harmand).
This species has been previously recorded from Java only.
Ruino.torus ArFFinis, Horsfield.
Cochin China (M. Harmand).
PHYLLORHINA DIADEMA, Geoffr.
Cochin China (M. Harmand); Sanghir Island (J. Laglaize).
The specimens from the latter locality differ from all others exa-
mined by me in the great development of the central projecting ridge
of the sella, which, in one instance, projects almost as far forwards
as the corresponding process of the same part of the nose-leaf in Ph.
cyclops (see my ‘Catalogue of Chiroptera,’ pl. ix. fig. 16); the
blunt projection in the centre of the margin of the upper transverse
nose-leaf is also much more defined than in other specimens of this
species, and in one of the specimens from Sanghir Island corresponds
to a large cell in the back of the leaf.
PHYLLORHINA ARMIGERA, Hodgson.
Cochin China (M. Harmand).
One of the specimens, an adult male, has the glandular elevations
between the nose-leaf and the frontal sac as much developed as in
the type of this species in the Calcutta Museum, described and
figured by me in the ‘ Proceedings’ for 1873 (p. 241).
PHYLLORHINA BICOLOR, Temminck.
Cochin China (M. Pierre).
PHYLLORHINA FULVA, Gray.
Cochin China (M. Harmand).
PHYLLORHINA LARVATA, Horsfield.
Cochin China (M. Harmand).
These specimens have the same brilliant golden-yellow fur as
those collected by Dr. Anderson at Prome, Upper Burma, and de-
scribed by me in my ‘ Monograph of the Asiatic Chiroptera.’
Cators rritui, Blyth.
Laos, in the roof of the Great Pagoda at Lakhon, collected by M.
Harmand.
The well-preserved specimens of this most remarkable Bat form
a valuable addition to the collection of the Muséum d’ Histoire Natu-
relle. Hitherto this species was known only from the type in the
Indian Museum, Calcutta (a dilapidated skin from the Bengal Sun-
derbuns), and a single specimen, in alcohol, from Java, in the collec-
tion of the Leyden Museum. Examination of these specimens per-
mits me to add a few additional notes on the structure of the species.
The caleaneum is weak, but distinct, nearly one fifth of an inch
_ in length, and projects at its extremity slightly beyond the membrane;
1878.] | CHIROPTERA IN THE MUS. D’HIST. NAT. PARIS. 879
there is no trace of a tail externally ; the wing-membrane extends to
the proximal extremity of the metatarsus; the female has pubic
teat-like processes, as in other species of Rhinolophide ; the ter-
minal phalanx of the fourth finger ends in a large T-shaped process.
The measurements agree closely with those of the specimen in the
Leyden Museum.
MEGADERMA SPASMA, L.
Laos (M. Harmand).
ScoOTOPHILUS TEMMINCKI, Horsf.
Cochin China, and Laos (M. Harmand).
VrESPERUGO ABRAMUS, Temminck.
Cochin China and Laos (M. Harmand).
VESPERUGO PACHYPUs, Temminck.
Laos (M. Harmand).
VESPERTILIO MURICOLA, Hodgson.
Laos (M. Harmand) ; Sanghir Island (M, Laglaize)
TAPHOZOUS MELANOPOGON, Temminck.
Cochin China (M. Harmand).
NyctTinomvs PLicatus, Buch.-Ham.
Laos (M. Harmand).
3. Ethiopian Region.
EroMOPHORUS CompPTUs, Allen.
Ogoné (M. Marche).
EromMoruHorvus monstrosus, Allen,
Ogoné (M. Marche).
PHYLLORHINA COMMERSONI, Geoffr.
Bugamojo, Zanzibar (Mission du St.-Esprit).
VESPERUGO NANUS, Peters.
Ogoné (M. Marche).
Myxoropa aurita, A. Milne-Edwards.
Madagascar (vide anted, p. 871).
CoLEURA SEYCHELLENSIS, Peters.
Seychelle Islands (M. Lantz).
4. Neotropical Region.
Mo .ossus RuFus, Geoffr.
Cayenne (M. Mélinon).
Motossus aBrasus, Temm.
Cayenne (M. Mélinon).
880 MR. DOBSON ON A COLLECTION OF CHIROPTERA. [Nov. 5,
RHYNCHONYCTERIS NASO, Wied.
Cayenne (M. Mélinon).
NOcTILIO LEPORINUS-
Cayenne (M. Mélinon).
ScHIZOSTOMA BRACHYOTE, 0. Sp.
Muzzle as long, or slightly longer than the distance between the
eyes ; side of the face deeply grooved beneath the eye. Ear much
shorter than the head, but broad at the base, owing to the strong
convexity of the inner margin of the conch in its lower two thirds ;
at the upper third the inner margin becomes abruptly concave, and
the ear terminates in a short, obtusely pointed tip: tragus simple.
Nose-leaf small, the lancet very acute, but much narrower than the
horse-shoe, which is slightly notched on either side, and confluent
with the muzzle in front. Centre of the upper lip slightly grooved ;
lower lip with a shallow narrow groove between two naked promi-
nences, each of which is divided in its upper third by a small oblique
roove.
: Wings from the ankles or from the extremities of the tibice ; inter-
femoral membrane and tail as in other species of the genus; calca-
neum slightly longer than the foot. Fur brown, above and beneath.
Dentition.—luce. 3 c. = pm. a3 m. =
Upper incisors as in Sch. megalotis ; lower equal, grooved, not
crowded ; first upper premolar with a very oblique cusp, touching
the canine, second straight ; first lower premolar much exceeding the
second in antero-posterior diameter ; second premolar well developed,
in the tooth-row ; third premolar nearly equal to the first in vertical
extent, but less than it in cross section at the base.
Length (of an adult ¢), head and body 2''-2; interfemoral mem-
brane 0-9; head 0-9; ear 0-65 X0'-5, tragus 0'-28 ; nose-leaf
0'-3 x 0-2; forearm 1-6; thumb 0'"35 ; third finger—metacarp.
1-4, Ist ph. 0-55, 2nd ph. 0:7, 3rd ph. 0-35; fifth finger—
metacarp. 1!35, 1st ph. 042, 2nd ph. 0-45; tibia 065; calca-
neum 0!'-45; foot 0!-4.
Hab. Cayenne (M. Meélinon).
This is a very aberrant form, differing from all the other species
in the development of the chin-warts, and in the shallowness of the
central groove between them, as well as in the shortness and peculiar
shape of the ears. In the second phalanx of the middle finger
being conspicuously longer than the first, it agrees with Sch. behni;
but, unlike that species, it has the calcaneum longer than the foot.
CAROLLIA BREVICAUDA, Wied.
Darien (Dr. Viguier) ; Cayenne (M. Mélinon).
ARTIBEUS QUADRIVITTATUS, Peters.
Cayenne (M. Meélinon).
1878.] DR. A. B. MEYER ON ANOA DEPRESSICORNIS. 881
November 19, 1878.
Arthur Grote, Esq., V.P., in the Chair.
Mr. Sclater read the following extract from an article in ‘ Science-
Gossip’ of October 1, 1878, by Mr. R. Davenport, of Bury, Lanca-
shire :—
“Tt is a pleasing duty to me to record the taking of a very
beautiful specimen of what I consider an exceedingly rare bird in our
neighbourhood (Saxicola stapazina). The specimen was shot by a
friend of mine, about the middle of May this year, on the margin of
the Bury and Radcliffe Reservoir ; and though much mangled with
number-6 shot, it has been very well mounted indeed by my friend
Johnson, of Prestwich. Considering the condition it was in from
being killed with such large shot, I really doubted at one time
whether it could be mounted ; however, it has been ; and a valuable
addition to our list of birds it is.”
Mr. Sclater exhibited the specimen in question, which had been
kindly sent up for exhibition by Mr. Davenport. It appeared to be
an adult in full plumage of Sawicola stapazina (called by Mr.
Dresser ‘‘ S. rufa” B. of Eur. pt. xxv.). The species had not been
previously recorded as occurring in the British Isles, and was an
interesting addition to the list of ‘‘ Accidental visitors.”
The following letters from Dr. A. B. Meyer, C.M.Z.S., and Mr.
Bartlett were read, in reference to the communication at the last
meeting from Mr. Everett respecting the supposed existence of the
Anoa ( dnoa depressicornis) in the Philippines.
**R. Zool. Mus., Dresden,
: November 16, 1878.
“Dear Dr. SciraterR,—
‘Having seen in the report of the Society’s meeting of the 5th
November that Mr. Everett had stated, in a letter to the Marquis of
Tweeddale, that the Anoa of Celebes (dnoa depressicornis) or an
allied species is found in the island of Mindoro, I beg leave to
send you a few notes from my diary on this subject.
“Mr. Wood of Manilla, an American gentleman, now dead, known to
many travellers as a amateur naturalist, and to many European
Museums as a collector, told me that Mr. Cuming (in whose company
he had formerly collected) had sent home an ‘Antelope’ from
Mindoro, similar to Anoa depressicornis from Celebes, which was
called ‘ Tamarao’ by the natives. Mr. Wood had not only seen
the animal in the Philippines, but also subsequently in the British
Museum, where it had received a specific name of its own.
** Wishing to know something more of the ‘ Tamarao’ in Manilla,
I succeeded in finding a stuffed animal in the Museum of the
Dominicans, which they told me was the animal in question. This
was apparently, as far as I could make out (it was in a bad state, and
882 MR. A.D. BARTLETT ON ANOA DEPRESSICORNIS. [Nov. 19,
could not be closely inspected), a Buffalo, small in size, with rather
straight horns bent backwards, with not much hair on the skin,
which was of a light greyish colour. There was no similarity to
the Anoa from Celebes, which I had seen the year before at Menado,
as regards colour, skin, size, or horns.
«The ‘Tamarao’ is said to live only on Mindoro. The wild Buffalo
of Luzon does not differ much from the tame one. The name for
Buffalo in the Tagaloe language of Luzon is Anooang, the same as
for Anoa depressicornis in South Celebes. (In the Pampanga
language of Luzon, ‘Damulag’ means buffalo. For a wild Buffalo
they say on Luzon ‘ Karabao cimarron,’ ‘ cimarron’ being a
Spanish expression for wild.)
«* When back in Europe in the same year, I looked together with
the late Dr. Gray through the galleries of the British Museum for
this ‘ Tamarao,’ but in vain; we cculd not find it.
“Tn the gallery of the Jardin des Plantes of Paris, I saw this
autumn a skeleton of a small Buffalo from Timor, brought home by
the ‘Astrolabe,’ and labelled Bubalus seligniceros, which reminded
me of the ‘T'amarao’ of Mindoro, as I mentioned to Prof. Gervais,
who kindly accompanied me, telling him also of its supposed affinity
to the dnoa.
‘“‘Of course I cannot depend on the alleged identity of the stuffed
Buffalo in the Museum of the Dominicans in Manilla with the
‘amarao’ of Mindoro; and, also, my inspection of this specimen
was scientifically not sufficient; but I hope to be able to throw
some light on this question very soon, as a skull of the real
‘Tamarao’ from Mindoro will be in my hands in a few days, as I
bought some time ago for the Dresden Museum Professor Semper’s
Mammalian collection from the Philippines, which contains such a
skull. I therefore shall soon recur to the subject.
“I take this occasion to remark that there exists on the island of
Balabac, Philippines, a species of Tragulus, specimens of which I
have seen living at the Government House in Cebti; but, having
no books with me, I could not make out whether it differs from Tra-
gulus kanchil or not. « Yours very sincerely,
E “A. B. Meyer.”
“* Zoological Society’s Gardens,
Regent's Park, London, N.W.
November 18, 1878.
“Dear Sir,—About thirty years since, a collector of natural-
history specimens, named Napper, wrote to me from the Philippines
to say that there existed on the island of Mindanao or Mindoro, a
small kind of Buffalo extremely wild and difficult to obtain. I
engaged him to obtain specimens ; and after much trouble and expense
he sent me an adult bull, a cow, and calf that he had shot and
skinned. They were offered by me to Dr. Gray for the British
Museum, who declined them, as he was of opinion they were only
small varieties of the common Manilla or Water-Buffalo; and this
was my own opinion, and that also of every one who saw them. I
P. 2.5.18 78. Pig
JSmit lth
CLASSIFICATION OF THE CERVIDA:.
1878. ] ON THE CLASSIFICATION OF THE CERVIDZE. 883
kept them a long time; and not finding any one that would have
them, I at last sent them to Stevens’s sale-rooms where they were sold
for a few shillings. I can assure you that these animals were not like
the Anoa, but much larger, and had all the appearance of a common
Indian Buffalo of small size. Yours faithfully,
“A.D. BartLett.”
The following papers were read :—
1, On the Classification of the Cervide, with a Synopsis of
the existing Species. By Sir Vicror Brooks, Bart.,
F.Z.S,
I. Introduction, p. 883.
II. On the division of the Cervide into sections larger than genera, p. 889.
TII. On the subdivision of the sections into minor groups, p. 889.
IV. Geographical distribution, and remarks thereon, p. 893.
V. Synopsis of the existing Cervide, p. 897.
VI. Explanation of Plate LV., p. 928.
(Plate LY.)
I. Introduction.
Some years ago I brought before the notice of the Society certain
pedal characters in the Cervidee (vide P. Z. S. 1874, p. 33) which ap-
peared to me to be of some importance; but at that time, not having had
the opportunity of extending my observations to all the forms of ex-
isting Cervidee, a generalization, which I believe to be of considerable
taxonomic value, deducible from these characters, escaped my notice.
A reconsideration of the characters themselves, and an attempt to work
out the deductions which may be drawn from these and other esta-
blished facts relating to the Cervidee, is the object of the present paper.
Before entering into the subject immediately before us, it may be
well to consider briefly the peculiarities common to all Deer, the
possession of which distinguishes them from all other mammals, and
which may therefore be called their family characters.
The Cervide, together with Moschus, the Giraffide, and the
Bovide, constitute the Pecora, one of the four distinct sections into
which the Artiodactyla, or even-toed Ungulates, is divisible. In having
(1) the odontoid process crescentic, (2) the left axillary rising from
the innominate artery, (3)the placenta cotyledonous, (4) the psalterium
added to the stomach, and (5) the outer bones of the metapodium
_Tudimentary, the inner confluent, all the members of the Pecora, as
Professor Flower in his exhaustive paper on Moschus (P. Z. S. 1875, p.
159) has taught us, agree together, and differ from the three remaining
sections of the Artiodactyla, viz. the Tragulina, Suina, and Tylopoda.
So far there is little difficulty in defining the zoological position
of the Cervide. That they form part of the Pecora there can be no
doubt ; but when the subdivision of the Pecora is attempted difficuties
arise which bid fair to set at defiance all systems of classification. An
estimate of the nature and relative value of the characters which dis-
tinguish the Cervide from the Bovidz will be most clearly shown by
placing them in tabular form. With Moschus and the Giraffidee we
884
SIR V. BROOKE ON TUE
[Nov. 19,
need not at present occupy ourselves; in Professor Flower’s paper,
above referred to, the characters in which they differ from the
typical Cervidee, of which Professor Flower regards them as early
offshoots, are distinctly defined.
Cervide.
(1) Osseous deciduous out-
growths from cylindrical pro-
cesses of the frontals in the males
of all excepting Hydropotes, in
which genus they are wanting
in both sexes. These appen-
dages, the antlers, are absent in
the females excepting Rangifer.
(2) Two orifices to the lacry-
mal duct, situated on or inside the
rim of the orbit.
(3) A depression in the facial
plate of the lacrymal, called the
lacrymal pit or fossa.
(4) An anteorbital vacuity of
such dimensions as to exclude the
lacrymal from articulation with
the nasals.
(5) In all species the first
molar in both jaws is brachy-
odont, or short-crowned, so that
when the tooth is in place the
neck, which is much constricted,
is on a level with or a little above
the alveolar border, and in the dry
skull the upper ends of the roots
are visible.
(6) Upper canines present in
both sexes. Exceptions :—Ca-
preolus, Axis, Dama, Cariacus,
Blastocerus, Pudu, Alces, and
some of the smaller Rusine Deer.
(7) Distal ends of the lateral
metacarpals present in some.
(8) The first and second
Bovide.
(1) Epidermal persistent (exc.
Antilocapra) sheaths envelop
processes of the frontals in both
sexes, with the exception of the
females of the following genera,
in which these appendages, the
horns, are wanting :—Portaz,
Tragelaphus, Procapra, Anti-
lope, Aipyceros, Saiga, Kobus,
Cervicapra, Pelea, Nanotragus,
Neotragus, Tetraceros.
(2) One orifice to the lacry-
mal duct, situated inside the rim
of the orbit. Exceptions :—Tra-
gelaphus sylvaticus, Tragela-
phus decula, Tragelaphus an-
gast, Tragelaphus spekii; in
these species there are two ori-
fices, situated as in the Cervidee.
(3) Absence of this depression.
Exceptions :—Gazella, Antilope,
Nanotragus, Neotragus, Cepha-
lophus, Tetraceros, Alcelaphus.
(4) Lacrymal articulates with
the nasals, except in Gazella and
Oryx.
(5) The first molar in both
jaws is hypsodont, or long-
crowned, so that a large part of
the crown is buried in the socket
when the tooth is first in place,
and there is no constriction
between the long crown and the
very short roots. Exception, ‘
Tragelaphus.
(6) Canines absent in both
sexes. Exception, Nanotragus
pygmeus has sometimes the
upper milk-canine.
(7) Distal ends of the lateral
metacarpals wanting in all genera.
(8) The first and second pha-
1878.]
Cervide (continued).
phalanges of the lateral digits
present. Exception, Cervulus.
(9) Parieto-squamosal suture
nearer the upper than the lower
border of the temporal fossa'.
(10) Navicular, cuboid, and
ectocuneiform united in some.
(11) Placenta with few coty-
ledons (Oligocotyledontophora,
Garrod 2).
CLASSIFICATION OF THE CERVID&%.
885
Bovide (continued).
langes of the lateral digits wanting
in all.
(9) This suture is nearer the
lower than the upper border of
the temporal fossa. Exception,
Antilocapra.
(10) These bones never all
united,
(11) Placenta with many
cotyledons (Polycotyledonto-
phora, Garrod).
(12) Gall-bladder absent. 12 Gall-bladder present. Ex-
ception, Cephalophus.
From this table it becomes apparent that, with the exception of the
eleventh, none of these characters can be considered exclusively
characteristic of the Cervidee ; and even with regard to it, more exten-
sive observation is required before it can be laid down as a fixed and
exclusive Cervine character. But although, with this exception, none
of the above characters can be taken singly as distinctive of a Cervine
from a Bovine animal, the coexistence of the second, third, fourth,
and fifth in all the Cervide and their absence in combination in
the Bovide will, I think, be found universal.
Deer may therefore for the present be characterized as Pecora
having two orifices to the lacrymal duct, situated on or inside the
orbit, lacrymal fossee, an anteorbital vacuity, cutting off the lacrymal
from articulation with the nasals, the first molar brachyodont, the
parieto-squamosal suture nearer the upper than the lower border of
the temporal fossa, and the placenta with few cotyledons,
II. On the Division of the Cervidee into Sections larger than Genera,
I will now pass on to the consideration of the pedal characters
above alluded to.
The fore limbs of all existing species of Cervide exhibit one of
the two following conditions.
(1) Two slender, short, detached, splint-like bones, the rudiments
of the second and fifth metacarpals (Plate LV. figs. 2, 2’ and 5'),
situated posterior to and one on each side of the proximal extremity
of the metacarpal cannon (or confluent third and fourth metacarpals) ;
the phalanges of the second and fifth digits separated from their
rudimentary metacarpals by a very wide interval ; the proximal
phalanx of these digits (Plate LV. fig. 2 p) inferior in size to the
central and ungual phalanges. ~ In Cervudus the phalanges of thie
second and fifth digits are undeveloped.
1 Vide Prof. Garrod, P. Z.8. 1877, p. 15,
2 Garrod, loc, cit. p. 12.
Proc. Zoou. Soc.—1878, No. LYIII. 58
886 SIR V. BROOKE ON THE [Nov. 19,
(2) Two slender pointed bones, the rudiments of the second and
fifth metacarpals (Plate LV. figs. 1, 2’ and 5’), situated posterior to
and one on each side of the distal extremity of the metacarpal cannon.
These bones articulate with their respective phalanges, but are
separated from the carpus by a wide interval. The proximal phalanx
of these digits (Plate LV. fig. 1, p) is superior in size to the central
and ungual phalanges.
For these forms Professor Flower has kindly assisted me in finding
convenient names. For the former, that in which the proximal (or
near) extremities of the lateral metacarpals remain, I would prepose
the term Plesiometacarpal (hycioy, near, and peraxaprioy, metacar-
pus); and the latter, that in which the distal (or more remote) extremity
of these bones remains, I would designate Telemetacarpal (rnXe, far,
pleraxapmiov, metacarpus).
With the view of ascertaining whether the myology of the fore
limbs of the Cervide confirms or confutes the importance which I
would assign to their osteological characters, I have been lately en-
gaged in the examination of the muscles of the fore limbs of the
Artiodactyles in general. My materials have been, however, as yet
far too meagre to justify me in attaching any great weight to the cha-
racters afforded by myology; but I am encouraged in expecting that
my observations may not be altogether unimportant, in which case
I shall hope to lay them before the Society upon some future oc-
easion.
I will only here mention that in the telemetacarpal limb of the
Common Roe, the phalanges of each lateral digit are supplied with
two tendons from the extensor muscles of the forearm—a condition
precisely similar to that which obtains in the tetradactyle Tragulus.
On the other hand, in the plesiometacarpal limb of the Red Deer and
Fallow Deer, the lateral phalanges are entirely without muscular at-
tachment. In neither the Roe, Red Deer, or Fallow Deer are the
lateral phalanges supplied with tendons from the flexors. A similar
condition is seen in Tragulus, in which form the lateral phalanges are
but slightly functional ; but in the telemetacarpal limb of Moschus
the lateral phalanges are (as in Sus) supplied with strong tendons
from the flexor profundus digitorum. In Moschus, the lateral pha-
langes are highly functional, The special office performed by the
lateral phalanges is the prevention of the animal slipping when
rushing at speed down slopes cr sinking in swampy ground ; and this
function is effected without danger of the displacement of these deli-
cate structures, by means of strong ligamentous bands developed in
the aponeurosis surrounding them, which bind them firmly in their
normal position. In the telemetacarpal limbs which I have ex-
amined, in addition to these bands, distinct round ligaments pass from
the sides of the distal articular surface of the metapodium to the
centre of the metacarpal splint on each side (Plate LV. fig. 1, r-r’),
and aid in preventing its displacement upwards, upon undue pressure
being applied from without.
In the annexed list of all the known species of existing Deer, ar-
ranged according to their geographical distribution, the names of
1878. ] CLASSIFICATION OF THE CERVIDE. "Be
those with telemetacarpal limbs are printed in italics, and those with
plesiometacarpal limbs in ordinary type. From this a remarkable
parallelism becomes at once apparent between the distribution of the
Cervidee in the two main divisions of the earth’s surface, and the
condition of the rudimentary external digits of their fore limbs. Of
the 39 species confined to the Old World, 36 are Plesiometacarpi,
the exceptions being Hydropotes inermis and the two species of
Capreolus. Of the 22 species confined to the New World, 21 are
Telemetacarpi, Cervus canadensis being the sole exception. Both
species of circumpolar range are T'elemetacarpi.
Northern regions of
Old World. both Hemispheres. New World,
1. Ceryulus muntjac. 1. Alces machlis. 1. Cariacus(Cariacus)virginianus.
2. lacrymans. 2. Rangifer tarandus. 2. (——) leueurus ?
3. —— reevesil. 3. —— (——) mewxicanus,
4, Elaphodus cephalophus. 4. —— (——) similis?
5, —— michianus ? 5, —— (——) savannarum.
6. Cervus (Rusa) aristotelis. 6. —— (——) peruvianus,
7. — ( ) equinus. 7. —— (——) gymnotis.
8. ——( ) swinhoii. 8. —— (——) toltecus.
9. —— (—_) philippinus. 9. —— (——) macrotis,
10. —— (——) mariannus ? 10. —— (——) columbianus.
11, —— (——) nigricans. 11. —— (Blastocerus) paludosus.
12, —— (——) alfredi. 12. —— (——) campestris.
13, —— (——) kuhli. 13. —— (Furcifer) chilensis.
14. —— (——) porcinus. 14. —— (——) antisiensis.
15. —— (——) lepidus? 15, —— (Coassts) rufus.
16. —— ( ) hippelaphus. 16, ——( ) stmplicicornis.
17. —— (Rucervus) duyauceli. 17. —— (——) rujinus.
18. —— (——) schomburgki. 18. —— (——) nemorivagus.
19. —— (——) eldi. 19. —— (——) supereiliares.
20. —— (Elaphurus) davidianus. 20. (——) whitelyi.
21, —— (Axis) axis, 21. Pudua humilis.
22, —— (Pseudaxis) sika. 22. Cervus canadensis.
23, —— (-——) euopis?
24, —— (——) mantchuricus.
25. —— (——) dybowskii?
26. —— (——) kopschi?
27. —— (——) taévanus.
28. —— (—— ?) caspicus.
29. —— (Cervus) elaphus.
30. —— ( ) xanthopygus ?
31. —— (——) eustephanus.
32. —— (——) maral.
33 cashmeerianus.
34. —— affinis.
85. —— (Dama) dama.
36. —— (——) mesopotamicus.
37. Aydropotes inermis,
38. Capreolus caprea,
39. —— pygargus.
The fact of the plesiometacarpal and telemetacarpal limb-charac-
ters so closely corresponding with the distribution of the Cervidee
in the Old and New Worlds, would in itself have convinced me of
their fundamental importance ; but a certain osteological cranial cha-
58*
888 SIR V. BROOKE ON THE [Noy. 19,
racter lately discovered by Professor Garrod (see P. Z. S. 1877, p. 13,
fig. 24) has confirmed me in this opinion.
Professor Garrod has observed that in all the American Deer, with
the exception of Cervus canadensis, the vertical plate developed from
the lower surface of the vomer is prolonged sufficiently downwards
and backwards to become ankylosed to the horizontal plate of the
palatals, and to form a complete septum dividing the nasal cavity
into two distinct chambers. The posterior edge of the vertical plate
of the vomer is plainly visible upon looking into the posterior nares
in the dry skull. In none of the Deer confined to the Old World is
the vertical plate of the vomer sufficiently developed to reach the
horizontal plate of the palatals; and therefore in none is there a sep-
tum dividing the nasal cavity into two chambers separated from the
anterior to the posterior nares.
This cranial character agrees with the pedal characters in placing
the Reindeer, one of the species of cireumpolar distribution, with the
Deer of the New World. Alces, along with Hydropotes and Ca-
preolus, all of which are Telemetacarpi, Prof. Garrod’s cranial
character places with the Plesiometacarpi of the Old World.
So long ago as 1836, Dr. Gray’ pointed out characters afforded by
the tufts of hair observable on the tarsus and metatarsus of most
Deer, and expressed his opinion that they constitute ‘‘a means of sub-
dividing them in natural sections.’ In his Catalogue of the Ruminant
Mammalia in the British Museum (1872), Dr. Gray divides the
Cervide into two primary groups:—(1) that in which tuft of hair on
the outside of the metatarsus is above the middle of that bone, and
(2) that in which this tuft is below the middle of the metatarsus.
Now I cannot but think it isa highly significant fact, and one
that should make us very cautious in condemning any character as
trivial, to find that a classification based on these apparently super-
ficial characters coincides exactly, as far as it goes, with that founded
on Prof. Garroa’s cranial characters, and, with the exceptions referred
to, with that based on the osseous structure of the limb.
There are some species both in the Old and the New World which
are devoid of either metatarsal or tarsal tufts. When present, the
metatarsal tuft is always above the middle of the metatarsus; and
there is never any tuft on the inuer surface of the tarsal joint in the
Deer confined to the Old World. In the Deer confined to the New
World, the metatarsal tuft when present is always below the middle
of the metatarsus, and there is frequently a distinct tarsal tuft. Cer-
vus canadensis constitutes the solitary exception.
There is yet another character which lends some weight to the
teaching of those already mentioned ; and for the observation of it we
are again indebted to Prof. Garrod*. Prof. Garrod has observed that
the ascending rami of the preemaxillee articulate with the nasals in
all the Old-World Deer, with one or two exceptions, whilst in the
Deer of the New World, again with one or two exceptions, the rami
of the preemaxillze do not reach the nasals.
I shall not now attempt to decide the relative value of these charac-
Vide P. Z. 8S. 1886, p. 66. 2 P.Z.8. 1877, p. 18.
1878. ] CLASSIFICATION OF THE CERVID2. 8389
ters, a task which can only become possible when our knowledge of
the extinct forms of Deer shall be very much more complete than it
is at present. Accepting the facts as we find them, the existing Cer-
videe may be divided into three primary sections.
A, Proximal ends of the lateral metacarpals remaining.
(PLESIOMETACARPI.)
Posterior portion of the nasal cavity not divided by the vomer
into two distinct chambers.
Tuft of hair on the external surface of the metatarsus, when pre-
sent, above the centre of that bone. Tuft of hair on the
inside of the tarsus always absent.
Ascending rami of the preemaxillee articulating generally with the
nasals.
Genera :—Cervulus, Elaphodus, Cervus (subgen. Rusa, Rucer-
vus, Elaphurus, Axis, Pseudaxis, Dama).
B. Distal ends of the metacarpals remaining. (TELEMETACARPI.)
Posterior portion of the nasal cavity not divided by the vomer
into two distinct chambers.
Tuft of hair on the external surface of the metatarsus, when pre-
sent, above the centre of that bone.
Genera :—Alces, Hydropotes, Capreolus.
C. Distal ends of the metacarpals remaining. (TELEMETACARP!).
Posterior portion of the nasal cavity divided by the vomer into
two distinct chambers.
Tuft of hair on the external surface of the metatarsus when pre-
sent, below the centre of that bone. Tuft on the inside of
the tarsus frequently present.
Ascending rami of the preemaxille generally not reaching the
nasals.
Genera :—Cariacus (subgen. Cariacus, Blastocerus, Furetfer,
Coassus), Pudua, Rangifer.
Of the inequivalency and heterogeneous nature of section B I am
most fully aware. It can only be considered a temporary receptacle
for species whose natural relationship to sections 4 and C cannot as
yet be finally decided.
III. On the Subdivision of the Sections into minor Groups.
The definition of the minor groups into which the three primary
sections are ultimately divisible in such a manner as to render their
boundaries readily appreciable, is, I believe, almost impossible. And
yet that a solid bond of affinity unites the members of each natural
minor group, that they possess in common certain peculiarities which
distinguish them from species contained in other minor groups, no
one who has studied the Cervidee can for a moment doubt. This
being so, I think that the distinct recognition of the various clusters
of species, into which the Cervidee naturally group themselves, is a
matter of the first importance in an attempt to understand their
entire life-history.
890 SIR V. BROOKE ON THE [Noy. 19,
The question as to whether these minor groups should be re-
garded as genera or subgenera, or merely marked as nameless sections,
appears to me to be of wholly secondary importance.
Actuated by this belief, I have retained the names originally given
to these subgroups whenever the grouping has appeared to me to
be natural, and have spoken of them as subgenera. The characters
upon which the subgenera are based are derived principally from the
form of the antlers, the skull, the rhinarium’, external coloration,
position and degree of development of the cutaneous glands, and
general external form.
As is the case with the family characters, it is seldom that any
one subgeneric peculiarity can be taken singly as exclusively charac-
teristic of a certain subgenus; but the existence in different fixed
combinations of these characters in the several subgenera is an
empirical certainty. For example, the form of rhinarium peculiar to
Rusa (vide infra) is shared by other groups; but never in combina-
tion with the form of antler and immense lacrymal pit typical of
that subgenus; and no species of Rusa is without this combination.
Again, the form of the rhinarium in the subgenus Cervus differs
from that of Rusa, but resembles that of Dama &c.; but in
Cervus this rhinarium is always associated with the antlers typical
of that subgenus, and an anal disk of paler colour than the adjacent
parts surrounding the tail; whilst in Dama it is combined with the
palmated form of antler and tuft of long hair surrounding the
prepuce. It is needless here to give further examples of this law:
it will be found fully illustrated in the diagnoses of the subgenera
(imperfect as these doubtless are) given in the synopsis.
The objection that antlers are absent in the females of all existing
species with the exception of Rangifer, whilst in Hydropetes they
are wanting in both sexes, has been frequently urged against the
value of characters based on these appendages. The stability of
this objection strikes me as very questionable. There can be no
doubt that it would be highly convenient if such prominent and
easily observed features as those afforded by the antlers were univer-
sally attainable; and the lack of their invariable occurrence is
doubtless a source of some slight confusion to the systematist.
But surely the value of a character as a test of genetic affinity
cannot be judged upon utilitarian principles ; and if, as I think, I
can show reason for believing the form of the antlers is one of the
clearest proofs of the blood-relationship of the species contained in
the various subgenera, characters derived therefrom must be regarded
as the most trustwothy for taxonomic purposes.
In order to account for the phenomena presented by the antlers
in the males of the existing Deer, four factors are necessary :—
(1) The gradual evolution of the antlers from very simple to complex
forms; (2) Their constant tendency to vary; (3) Variation extend-
ing far enough to induce the partial atrophy of one part of the antler
1 T follow Tiger and Sundevall in the adoption of this term for the naked
moist skin between the nostrils. It is decidedly preferable to muffle, which is
French, or muzzle, which signifies something quite different in every-day speech.
i878.] CLASSIFICATION OF THE CERVID&. 891
to compensate for the extra development of some other part ; (4)
The transmission and fixation of such variations by heredity.
(1) The gradual evolution of the antlers from very simple to more
complex forms.
The earliest Cervine animal of which we have any knowledge is
Dremotherium from the early Miocene of Europe. This species was,
so far as is at present known, without antlers. In the middle Mio-
cene of France and Germany, and in a somewhat similar horizon in
North America, the earliest antlered Deer as yet discovered have
been found. These species, belonging to the very closely allied
genera Dicrocerus and Cosoryz, possessed very short antlers with a
single tine projecting forwards ; in fact they closely resembled the
antlers of the existing species of Furcifer (fig. 17, p. 923).
From this period to recent times a gradual increase in the luxu-
riance of growth and diversity in the form of the antlers is evidenced
even by the miserably imperfect materials as yet at the command of
naturalists. Of this the following forms bear witness :—
CERVUS MATHERONI, Gerv.
Cervus matheroni, Gaudr. An. foss. Mont Léb. p. 66, pl. 13.
Horizon. Upper Miocene.
Cervus cusanvs, Croiz. & Job.
Cervus cusanus, Boyd Dawk. Quart. Journ. Geol. Soc. 1878, p. 405,
fig. 2,
Horizon. Pliocene (Lower ?)
Cervus perriert, Croiz. & Job.
Cervus perrieri, Boyd Dawk. Quart. Journ. Geol. Soc. 1878, p. 488,
figs. 3-5.
Horizon. Pliocene.
CreRVUs CyLINDRICORNIS, Boyd Dawk. Vide loc. cit. p. 415,
figs. 11, 12.
Horizon. Upper Pliocene.
Cervus TETRAcERUS, Boyd Dawk. loc. cit. p. 417, figs. 13-17.
Horizon. Upper Pliocene.
MEGACEROS HIBERNICUS.
Horizon. Post-tertiary Deposits.
CERVUS ELAPHUS.
Horizon. Post-tertiary deposits, and existing epoch.
I need hardly say that these species have no direct affinity to each
other ; they are simply chosen as illustrating the general fact that the
antlers of the Cervide have gradually increased in complexity from
the Upper Miocene period to recent times. :
It might at first sight appear as though the occurrence of existing
Species with very simple antlers militated against the more compre-
892 SIR VY. BROOKE ON THE [Noy. 19,
hensive fact of the gradual augmentation of complexity in these
organs through geological time; but as the converse, namely the
existence of highly complicated antlers in the Miocene and Phocene
periods, finds no support in the Geological record, the simple antlers
of some existing species may be, I think, safely regarded as the
result of arrestation, dependent ona less severe struggle for existence,
or possibly upon some hitherto undiscovered influence.
(2) A constant tendency to vary.
Of this, any one can satisfy himself by the inspection of large
collections of antlers. No two antlers are ever exactly alike; and
the variations to which the antlers are subject in some species (e. g.
Cervus elaphus, vide infr. p. 910) is so great, that in the absence of a
large series they would be held as indicative of several distinct species.
(3) Variation extending far enough to induce the partial atrophy
of one part of the antler to compensate for the extra development of
some other part.
To illustrate this subject exhaustively would occupy more space
than can be well devoted to it in the present paper. An attentive
examination of any large collection will yield abundant evidence that
such is invariably the case. It will, perhaps, be sufficient for my
present purpose to bring forward one special example. The normal
form of the upper half of the antlers of the common Fallow Deer
(fig. 9, p. 914) may be described as broadly palmated, with several
rudimentary tines produced from the posterior margin of the palm,
the tine situated lowest on the palm exceeding the others slightly in
length (fig. 9,d@). In my cousin Mr. J. Bloomfield’s place, Castle
Caldwell, on the shores of Lough Erne, there is a herd of Fallow
Deer which have lived at large in the woods, and under perfectly
natural conditions, for upwards of 50 years. The antlers of these
Deer are so remarkable in form, and present the peculiarities in
which they are remarkable so constantly, that they could be recog-
nized amongst a large collection of Fallow-Deers’ antlers without the
slightest difficulty. The tine growing from the inferior and pos-
terior border of the palm (Plate LV. fig. 3, d) is in the Castle-Cald-
well Deer immensely developed, and the palm itself is much con-
tracted and seldom bears more than 3 or 4 tines. Variation is still
actively at work on the antlers of these Deer, and reversion towards
the normal form occasionally asserts itself; but, notwithstanding, a
fixed type of antler has stamped itself indelibly upon the males in
this herd in the short space of 50 years, and the peculiarities cha-
racteristic of that type are unquestionably the result of the extra
development of one of the posterior tines and the consequent par-
tial atrophy of the palm.
(4) The transmission and fixation of such variations by heredity.
That this is the case has been in a large measure proved by what
has been already said; but the following fact places the matter be-
yond all possibility of doubt. The Castle-Caldwell Deer are the
descendants of about one or two males and twice as many females.
A single antler of one of the fathers of the herd has been fortunately
preserved in the Museum at Castle Caldwell. This antler (Plate
1878. | CLASSIFICATION OF THE CERVIDZ. 893
LV. fig. 4) I_have the pleasure of exhibiting this evening. It will
be seen that it presents the same peculiarities that are characteristic
of the distant descendants of its original owner, and that we have
here a direct proof of heredity transmitting, and to a very great
extent fixing, a definite and prominent variation’.
Did space permit I could give many more illustrations corrobo-
rating the evidence of the above-mentioned facts. Innumerable in-
stances of the power of heredity to transmit and fix variations in the
antlers until some stronger influence interferes, are enacted annually
before my eyes, amongst about 600 Deer of several different species
preserved in my parks. Enough has, however, I think, been ad-
duced to leave the onus probandi upon those who deny to characters
derived from the antlers the right to be considered one of the surest
land-marks of affinity in the Cervide.
IV. Geographical Distribution of the Cervidee, and Remarks thereon.
ae ; i |
ewe | = = i ee
° | - = 5)
Genera Number zs Sots 2 § ‘Ep Fa
and of H ep wae &) ia] 3
Subgenera. | species.| 3 § 5 a a 2 =
=o 8s = ars iS
BE | asi 4 Z A
No. of sp.No. of sp.|No. of sp.|No. of sp.|No. of sp.
Cervulus ... 3 aa Bes 3
Elaphodus... 2 Soc sp 2
DBTBA iia ore Sb 05 11 % bait 11
Ruceryus ... 3 Ree Sen 3
Elaphurus ... 1 45 1
WAGs Pet eee S. 1 i mm 1 |
Pseudaxis ... ae NG, 4 2
Cervus ...... i 2 4 1
Hama asf... 3 2 2
Hydropotes . 1 1
Capreolus ... 2 1 1
Cariacus...... 10? $e 7 3
Blastocerus.. 2 ee 2
Furcifer ...... 2 Z 2
Coassus ...... 6 6
Pu 2. cece 1 1
Motalty ..ic. 61 6 11 22 8 14
? The production of so marked a yariation from the form of antler typical
of Dama vulgaris in the short space of fifty years as that exhibited by the
Castle-Caldwell Fallow Deer might be considered as tending to prove the
instability of the original specific characters. It may, however, be observed
that the general form of antler typical of the species has not been obliterated
by the evolution of the characters now peculiar to the variety, but that, on the
contrary, heredity has preserved a record of the earlier, or specific, alongside of
the later, or varietal, modification. Essentially analogous phenomena are pre-
sented by the antlers typical of the species and subgenera of the Cervide, the
distinctive specific characters being, in eyery instance, an evident modification
of the form of antler characteristic of the subgenus.
894 SIR V. BROOKE ON THE [Nov. 19,
In this Table, showing the geographical distribution of the Cer-
videe exclusive of the Elk and Reindeer, which have a cireumpolar
range, the following are the most important facts observable. Out
of a total of 61 species 39 are confined to the Old and 22 to the
New World. Of the 39 Old-world species 33 are found in the
Eastern Palearctic and Indian regions, the latter containing 22
species, a much larger number than that contained in any one of the
remaining four regions.
Of the 22 species confined to the New World, 8 are found to the
north and 14 to the south of the Isthmus of Tehuantepec, the
division adopted by Mr. Sclater between the Nearctic and Neotro-
pical regions. These figures cannot, however, be taken as afford-
ing a just estimate of the relative degree of specialization of the
Cervidee in the Nearctic and Neotropical regions respectively. On
the one hand, the three species of Cariacus in the latter are undoubt-
edly immigrants from the former region; and on the other hand, the
species contained in the subgenera Coassus and Furcifer are, as
far as the antlers are concerned, amongst the least-specialized
of existing species.
Out of the eleven genera into which the Old-world Deer are di-
visible, there is only one (Dama) which has not a representative in
either the Palearctic or Indian region.
Owing to the exceeding paucity of Palzeontological material, any
attempt to account for the present differentiation and distribution
of the Cervidee must be considered as purely tentative; and I venture
to put forward the following hypothesis, more to provoke investiga-
tion and criticism, which may possibly bring new and more correct
ideas to light, than from any great confidence that I feel in its
approximation to completeness.
In order to avoid unnecessary recapitulation, and to indicate as
clearly as possible the interdependence of the facts and hypothesis,
I shall place the facts and theory in two columns, numbering the
former, and merely introducing the number attached to each fact
before the deduction or deductions which are drawn from it.
Facts.
(1) So many characters are
common to the Cervide and
Bovide that it is very difficult
to lay down definitions distinc-
tive of the families.
(2) Dremotherium, the earliest
known Cervine form, was with-
out antlers and tetradactyle.
Hypothesis.
(1) A very high degree of
specialization had been attained
by the common ancestor of the
Cervide and Bovide, especially
in the structure of the skull,
dentition, and limbs; hence the
possibility of divergence of cha-
racter in its descendants was
much restricted. (2) Fora long
period the early ancestors of the
Cervide, which were tetradactyle,
1878. ]
Facts (continued).
(3) The antlers of the Euro-
pean Procervulus and of some
species of the North-American
Cosoryx appear to have been
persistent.
(4) The antlers of Dicroceros
and Cosory# possessed one, or
at most two tines.
(5) The increase in the com-
plexity of the antlers of extinct
species accompanies their chro-
nological sequence in geological
time.
(6) In a conversation which I
have lately had with Prof. Marsh,
he informed me that some of the
specimens of Cosoryx collected
by him in North America were
tetradactyle. The lateral meta-
carpals in these specimens are
excessively attenuated in the
centre third of their length. In
other specimens Prof. Marsh
tells me a natural separation had
taken place between the proxi-
mal and distal ends of the bones.
(7) The condition of the late-
ral metacarpals separates exist-
ing Deer into two great groups,
the Plesiometacarpi and Tele-
metacarpt.
CLASSIFICATION OF THE CERVID&X.
895
Hypothesis (continued).
remained unantlered; and spread-
from the centre of their geogra-
phical area (13), which was pro-
bably in the Eastern Palearctic
and Indian regions, they passed
westward into Europe, and east-
ward into North America. (3)
Processes then became developed
from the frontals, which gradu-
ally elongated and in some in-
«stances branched. At first these
outgrowths from the frontals
remained persistently attached ;
but eventually the great advan-
tage enjoyed by individuals who
through necrosis lost, and through
an inherited tendency to produce
frontal processes renewed their
antlers, over individuals who
retained antlers broken and
rendered useless by frequent
combats, caused the natural se-
lection of the former in the
struggle for existence. (4, 5)
These first deciduous antlers
were exceedingly simple; but as
time rolled on the advantage of
large and complex antlers as a
means of offence and defence
established an ever-increasing
tendency towards complexity in
their form. (6) A diminution
in the size of the lateral digits of
the early forms of Deer accom-
panied the increase in the size of
their antlers. The centre part
of these bones, after attaining an
extreme degree of atrophy, at
last ceased to ossify. (7,8) In
some species the default in ossi-
fication took place nearer the
distal than the proximal extre-
mity of the bone. In others the
converse obtained. The reduc-
tion of the rudiments steadily
continued, resulting eventually
in the disappearance of the
shorter rudiment in both forms,
Hence the origin of the plesiome-
896
Facts (continued).
(8) In some of the Telemeta-
carpt the distal extremities of
the lateral metacarpals is two
thirds the length of the meta-
carpal cannon. In some of the
Bovidee (e. g. Gazella), of which
family all the species are plesio-
metacarpal, the proximal extre-
mity of these bones is two thirds
the length of the metacarpal
cannon.
(9) With three exceptions all
the Old-world Deer are plesio-
metacarpal ; and with one excep-
tion all the New-world Deer are
telemetacarpal.
(10) In none of the Old-
world Deer is the vomer suffi-
ciently prolonged backwards to
divide the nasal cavity into two
chambers. In all the New-
world Deer the nasal cavity is
so divided.
(11) The vomer of Megaceros
hibernicus exhibits an interme-
diate condition. The limb in
this form is plesiometacarpal,
and as highly specialized as in
the existing Plestometacarpi.
(12) The Telemetacarpi of
the Old World resemble the
Plesiometacarpi in the form of
their vomer and in the position
of the metatarsal tufts.
(13) Thirty-three out of the
thirty-nine species of Old-world
Deer are inhabitants of the In-
dian or East Palearctic regions ;
and of the eleven subgenera into
which they are divisible there is
only one (Dama) that has no
representative in these regions.
SIR V. BROOKE ON THE
[Nov. 19,
Hypothesis (continued ).
tacarpal and telemetacarpal con-
dition of limbs in existing Cer-
vide. (9) It is probable that
the separation between the upper
and lower extremities of the me-
tacarpals took place long after
the first Dremotherium-like Cer-
vides had found their way into
North America. In these the
default in ossification began in
the upper third of the lateral
metacarpals; (12) whilst in the
Cervide remaining in the Old
World both modes of reduction
obtained a footing, the Plesiome-
tacarpi gaining ultimately the
preeminence. (10) The vomer
in the early forms of Deer was,
as in the existing less-specialized
Artiodactyles (e. g. Hippopota-
mus, Sus, Tragulus), prolonged
backwards, dividing the nasal
cavity into two distinct chambers.
(11) As the competition he-
tween rapidly multiplying and
allied forms became severe in the
Old World, specialization exten-
ded to this part of the cranium,
and a reduction in the backward
extension of the vertical plate
of the vomer, and consequent
modification of the surrounding
parts, set in. It is not impro-
bable that the limbs of the Plesto-
metacarpi had nearly reached
their present stage of specializa-
tion before the form of their
vomer departed from the primi-
tive type. The Deer of the
New World, being less numerous
both in individuals and species,
and not being subjected to such
severe competitions as the Old-
world Deer, retained the primi-
tive form of vomer. The greater
development of the antlers in the
larger majority of Old-world
Deer, as compared with those
of the New-world Deer, may be
1878. ]
Facets (continued).
(14) There are only twenty-
two species in America; and the
South-American species have
remarkably simple antlers,
(15) The antlers of the Cer-
videe are subject to great varia-
tion; and such variations are
transmissible by heredity.
(16) Two species have a cir-
cumpolar range, and Cervus
canadensis, a species allied to the
common Red Deer of Europe, is
found in North America.
CLASSIFICATION OF THE CERVIDA.
897
Hypothesis (continued).
attributed to the same cause.
(14) The Deer of South Ame-
rica, which, according to the
theory under consideration, have
been freer from contact with al-
hed forms than species inhabit-
ing districts nearer to, and more
in direct continuity with, the
centre of the family area of dis-
tribution, are remarkable for the
great simplicity of their antlers.
(15, 16) For long periods there
was no possibility of communi-
cation between the Deer of the
Old and New World ; and during
this time the perpetuations by
heredity of the more favourable
varieties which arose from time
to time resulted in the diver-
gence of the Cervide into the
clusters of species which we now
term genera. (16) At length
the means of dispersal was for a
second time opened between
Asia and North America; an
interchange of species took place ;
the ancestors of Cervus canaden-
sis passed over into North Ame-
rica from the Old World, and the
Elk and Reindeer extended their
range till it was cireumpolar.
In the following synopsis I have admitted every species of whose
invalidity I am not absolutely certain ; and frequently, even where I
have felt grave doubts as to the soundness of the claims of a species
to specific recognition, I have left it distinct, if by so doing I have
thought it probable that future investigation of its claims may throw
light on the important problems of geographical variation and dis-
tribution. Where colour is not mentioned in the diagnoses it affords
no generic or subgeneric character. Supplementary columns are
absent in the molars, and the central incisors are much expanded,
where the contrary is not stated. Where the metatarsal and tarsal
tufts are not specially mentioned, they are present.
V. Synopsis of the existing Cervide.
A. Proximal ends of lateral metacarpals remaining.
(PLESIOMETACARPI. )
Posterior portion of the nasal cavity not divided by the vomer
into two distinct chambers.
898 SIR V. BROOKE ON THE [Nov. 19,
Tuft of hair on the external surface of the metatarsus, when
present, above the centre of that bone. Tuft of hair on the
inside of the tarsus always absent.
Ascending rami of the premaxille articulating with the nasals.
(Exception, species 3, in genus 1.)
1. CeERVULUS.
1816. Cervulus, Blainv. Bull. Soc. Phil. p. 74.
1827. Stylocerus (subgen.), Ham. Sm., Griff. An. Kingd. vol. v.
p- 319.
1836. Prox, Ogilby, P. Z.S. 1836, p. 135.
Antlers haif the length of the head, and placed on pedestals
which nearly equal (and sometimes surpass) them in length. Brow-
antler (fig. 1, @') very short, inclined inwards and directly up-
wards. Terminal extremity of the beam (fig. 1, 7), which is un-
branched, curved downwards and inwards. Lacrymal pit of great
depth and extent, the facial plate of the malar taking part in its
floor. Anteorbital vacuity of moderate extent. Ascending rami of
the preemaxille equalling or exceeding the nasals in length. A
strong ridge passes from the root of each pedestal over the orbit,
lacrymal pit, and anteorbital vacuity, serving to protect frontal cuta-
neous glands which lie on its inner surface. Auditory bulle very
Fig. 1
Cervulus muntjae,
slightly inflated, the tympanohyal abutting against the paroccipital
process.
Canines strongly developed in the males, and curved downwards,
backwards, and outwards ; in the females they are small; and in
neither are the pulps persistent.
Rhinarium extensive, completely surrounding the nostrils, the
upper border of its internarial portion slightly convex. Preenarial
portion of the rhinarium broad, its lateral borders passing sub-
vertically downwards to the upper lip. No metatarsal tufts. Tarsus
with the navicular, cuboid, and ectocuneiform united. Phalanges of
lateral digits absent in both fore and hind extremities. Young
spotted (exe. species 12). Stature small.
Distribution. Indian and South-eastern Palzearctic regions.
1 Prof. Garrod, in his valuable paper to which I have so frequently alluded
(P. Z.8. 1877, p. 16), has drawn attention to a law of which the various modifica-
tions of the antlers in the Cervid are, in his opinion, the result. I have ven-
tured to extend, and in some slight degree to modify, Prof. Garrod’s suggestion.
The lettering in the woodcuts indicates the parts of the antlers which appear
to me to be homologous,
1878.] CLASSIFICATION OF THE CERVID2. 899
1. CERVULUS MUNTIAC.
1780. Cervus muntjac, Zimm. Geogr. Gesch. Band ii. p. 131.
1785. muntjak et vaginalis, Bodd. Elench. Anim. vol. i.
p. 136.
1867. Cervulus aureus, Jerd. Mamm. p. 264.
1873. amostylis et tamulicus, Gray, Hand-list Rum. Mamm.
p. 165.
1874. muntjac, Brooke, P. Z.S. 1874, p. 38, fig. 3.
Range. British India, Burmah, Malay peninsula, Sumatra, Java,
Hainan, Banka, Borneo.
This species appears to attain a larger size in Java, Sumatra, and
Borneo than it does on the mainland ; and I think it not improbable
that persistent race-characters may eventually be found, distin-
guishing the Muntjac of these islands from that of British India.
2. CERVULUS LACRYMANS.
1871. Cervulus lacrymans, Alph. M.-Edw. Nouv. Arch. du Mus.
t. vii., Bull. p. 93; and Rech. s. 1. Mamm. p. 348, pls. 63, 64.
1872. scelateri, Swinh. P. Z. S. 1872, p. 814. ,
1874. , Brooke, P. Z. S. 1874, p. 40, pl. 8 & fig. 4.
Hab. Moupin (David); hills near Hangchow, China (Swinhoe).
Type, Mus. d’ Hist. Nat. Paris.
3. CERVULUS REEVESII.
1838. Cervus reevesii, Ogilby, P.Z. 5S. 1838, p. 105.
1862. Cervulus reevesi, Swinh. P. Z.S8. 1862, p. 361.
1874. , Brooke, P.Z. 8S. 1874, p. 41, fig. 5.
Range. Southern China from latitude of Canton as far north as
Ningpo ; Formosa (Swinhoe).
2. ELAPHODUS.
1871. Elaphodus, Alph. M.-Edw. Nouv. Arch. du Mus. p. 93,
pl. 7; and Rech. Mamm. p. 353, pls. 65-67.
1874. Lophotragus, Swinh. P. Z.8. 1874, p. 452.
Antlers very small, unbranched, supported on slender, long, con-
verging pedestals. Ascending rami of the przemaxille very strong,
and articulating broadly with the nasals, to which they are inferior
in length. No supraorbital ridges or frontal cutaneous glands.
Canines in the male massive and long, curved downwards, and not
everted. A well-developed frontal tuft. In other respects resembles
Cervulus, with which genus Hlaphodus may be united to form the
subfamily Cervuline, as proposed by Professor Garrod (P. Z. S.
1876, p. 765).
Distribution. South-Eastern Palearctic region.
1. ELarpHopus CEPHALOPHUS.
' 1871. Elaphodus cephalophus, Alph. M.-Edw. (loc. cit. supra).
Hab. Moupin (David). Type, Mus. d’ Hist. Nat. Paris.
900 SIR V. BROOKE ON THE [Nov. 19,
2. ELAPHODUS MICHIANUS.
1874. Lophotragus michianus, Swinh. P. Z. 8. 1874, p. 452, pl. 59.
1876. » Garr. P. Z. 8. 1876, p. 757; pl. 76.
Hab. Neighbourhood of Ningpo (Swinhoe). ‘Type, Mus. Berlin.
Although I feel sure that Professor Garrod is correct in uniting the
genera Elaphodus and Lophotragus, I have been unable to satisfy
myself that the types of the two genera are referable to the same
species, as he supposes. I have therefore refrained from amalga-
mating the Moupin and Ningpo specimens in a single species, and
have left the question for the present open.
3. Cervus.
(Rusa, subgen.)
1827. Rusa (subgen.), Ham. Sm., Griff. An. Kined. vol. y. p. 309.
Fig. 2.
Cervus (Rusa) aristotelis.
Antlers twice or thrice the length of the head, supported on
pedestals of moderate length. Brow-antler (fig. 2, a) strong, curved
boldly upwards, its axis forming an acute angle with that of
the beam. The beam (fig. 2, x) bifureates, giving rise to two
strong tines (fig. 2,6 & c). Lacrymal pit of very great depth and
extent, its antero-posterior diameter exceeding that of the three
upper molars. Anteorbital vacuity very large. Auditory bullz but
slightly inflated and rugose externally (exe. species 8 and 9). Rudi-
mentary canines present in both sexes (exe. species 6, 7, 8, 9).
Molars with small supplementary columns.
Rhinarium resembling that of Cervulus, but less extensive, its
upper internarial margin being slightly concave, and its infranarial
portion narrower. Tail of moderate length. Neck maned (exc.
species 7 and 9). Colour uniform brown (exc. 7 and 9). Stature
diverse, the largest species being amongst the largest, and the smaller
almost amongst the smallest of the Cervide. Young unspotted
(exc. species 7 and 9).
Distribution. Indian region.
1878. ] CLASSIFICATION OF THE CERVID&. 901
(a) The upper tines (6 & e, fig. 2) of nearly equal length in adult
animals ; in young males the outer and anterior tine (b, fig. 2)
the longer.
1. CeRVUS ARISTOTELIS,
1835-7. Cervus aristotelis, Cuv. Oss. foss. (ed. 4) tom. vi. p. 84.
Range. British India, Ceylon, Burmah, Siam, Hainan (Swinh.).
(8) The outer tine (6, fig. 2) always surpassing the inner
tine (ce) in length.
2. CERVUS EQUINUS.
1825. Cervus equinus, Cuv. Oss. foss. (ed. 3) tom. iv. p. 454;
ib. (ed. 4) pl. 201. fig. 10.
1839-44. Cervus equinus, Mill. Verh. Nat. Gesch. Ned. Bez.
Zool. p. 213, pls. 42-45. figs. 7-11.
Range. Borneo, Sumatra (Miill.) ; Singapore (Brooke).
I have seen several living specimens of this species, all of which
in their much darker colour, more bushy tail, short slightly curved
antlers, presented a tout ensemble very unlike the common Sambur.
On the other hand, I have received skulls and antlers from Siam
which are strikingly intermediate between typical specimens of Cervus
aristotelis and C. equinus. It appears to me therefore probable
that the existence of these intermediate forms may ultimately render
it impossible to maintain the distinction of the species.
3. CERVUS SWINHOII.
1862. Rusa swinhoii, Sclat. P. Z.S. 1862, p. 152, pl. 17.
1862. , Swinh. loc. cit. p. 364.
1871. ——,, Sclat. Trans. Z. 8. 1871, p. 333, pl. 39.
Hab. Formosa. Type, Brit. Mus.
This species closely resembles Rusa equinus.
4, CERVUS PHILIPPINUS.
1827. Cervus philippinus, Ham. Sm., Griff. An. Kingd. vol. iv.
p- 147, and vol. v. p. 319.
21877. —— , Brooke, P. Z.S. 1877, p. 51, pl. 8.
Hab. Luzon (Philippines). Type, Mus. d’Hist. Nat. Paris (No.
409 in Cat.).
5. CERVUS MARIANNUS.
1820. Cervus mariannus, Desm. Mamm. sp. 669, p. 436.
1877. ——— , Brooke, P. Z. S. 1877, p. 53.
Hab. Island of Guam (Marianne Islands). Type, Mus. d’ Hist.
Nat. Paris (414 a in Cat.).
As Ihave stated in my paper above referred to, I feel pretty
certain that this and the preceding species are identical. I am, how-
Proc. Zoou. Soc.—1878, No. LIX. 59
902 SIR V. BROOKE ON THE [Nov. 19,
ever, now, as then, unable to substantiate this opinion satisfactorily,
owing to insufficiency of material.
6. CERVUS NIGRICANS.
1877. Cervus nigricans, Brooke, P. Z. 8. 1877, p. 57, pls. 9, 10.
Had. Philippines (exact locality’). Type, in my own collection.
7. CERVUS ALFREDI.
1870. Cervus alfredi, Sclat. P. Z.S. 1870, p. 383, pl. 28.
1877. , Brooke, P. Z. S. 1877, p. 59.
Hab. Philippines (Luzon’?). Type, Brit. Mus.
8. CERVUS KUHLI.
1839-44. Cervus kuhlii, Mull. Verh. Nat. Gesch. Ned. Bez. Zool.
p. 223, pls. 44, 45. figs. 12-14.
Hab. Bavian Islands. Type, Mus. Leyden.
This species was originally described by Miiller from specimens
collected by Diard in the Bavian Islands, to which locality the
species, as far as is at present known, is confined. The auditory
bullze in Cervus kuAli are largely inflated. By this character the
skull may be distinguished from that of any of the Rusine deer with
the exception of C. porcinus. The skull of C. kuhii differs from
that of C. porcinus in having the facial portion anterior to the orbits
much more compressed, and the orbits much more prominent.
C. kuhli has proportionally much longer limbs than C. porcinus ;
and all the hairs of the back and sides are annulated in the former
and not in the latter species. The young of C.kuhli are unspotted,
which is not the case with those of C. porcinus’. Both species are
devoid of upper canines.
9. CERVUS PORCINUS.
1777. Cervus porcinus, Zimm. Spec. Zool. Geogr. Quadr. p. 552,
sp. 6.
Range. British India, Ceylon, Burmah.
10, Cervus LEPIDUS.
1844. Cervus lepidus, Sund., Pec., Vetensk. Akad. Handl. p. 180,
sp. 14.
Hab. Java? Type, Mus. Frankfort ?
This is a very doubtful species. Some years ago I saw the type
in the museum at Frankfort; and it appeared to me to closely
resemble Cervus sika. Upon the occasion of my last visit to
Frankfort I sought in vain for the specimen, and much fear that it
has been destroyed.
1 Vide Selater, P. Z. 8. 1863, p. 224.
1878. ] CLASSIFICATION OF THE CERVID. 903
(y) The tuner tine (fig. 3, c) always surpassing the outer tine
(fig. 3, b) in length.
11. Cervus HIPPELAPHUS.
1825. Cervus hippelaphus, Cuv. Oss. foss. (ed. 3) tom. iv. p. 40;
ib. (ed. 4) tom. vi. p. 77, pl. 166. figs. 31-34.
1839-44. Cervus rusa, Mil. Verh. Nat. Gesch. Ned. Bez. Zool.
p. 217, pl. 43, pl. 44. figs. 1-6.
Hab. Java, Borneo (introduced). I
Deer resembling Cervus hippelaphus in every particular excepting
size are found in many of the islands in the Indo-Malayan and
Austro-Malayan regions. How far these are the result of natural
distribution, or how far their occurrence is attributable to artificial
transportation is now most difficult to decide. The best authori-
ties on the fauna of the East Indies, Wallace, Schlegel, and Miller,
Cervus (Rusa) hippelaphus.
are inclined to lean towards the latter alternative; and it is I think
certain that the introduction of many of the larger mammals (e. g.
Monkeys, Pigs, and Deer) into the islands of the Austro-Malayan
region has been effected by the Malays, who, according to Wallace,
are much given to taming animals and conveying them from island to
island. At the same time, the existence of a species of Melis peculiar
to Timor, and of the singular Anoa depressicornis on Celebes, seems
to me to necessitate caution in accepting this solution as of universal
application.
I shall therefore, for the present, keep the references to Cervus
timortensis of Timor and C. moluccensis distinct.
12. (a) CERVUS TIMORIENSIS.
1822. Cervus timoriensis, Blainv. Journ. de Phys. vol. xciv. p. 267.
1825. peronii, Cuv. Oss. foss. (ed. 3) tom. iv. p. 46.
59*
904 SIR V. BROOKE ON THE [Nov. 19,
1833. Cerf de Timor, F. Cuv. Mamm. pl. 361.
1852. Cervus peronii, Puch. Cerf, Arch. du Mus. vol. vi. p. 409.
1839-44. Cervus rusa, var. timoriensis, Schl. et Mull. Verh. Ned.
Bez. Zool. pp. 212, 220.
Hab. Timor (Peron, Dussumier), Samao, and Cambing near
Timor (Schlegel and Miiller).
Specimens examined :—
Type of F. Cuvier’s and Pucheran’s descriptions, Mus. d’ Hist. Nat.
Paris.
Miiller’s types, Mus. Leyden.
Specimen from Leyden, British Museum.
Living specimens in Zool. Gardens of Paris and Rotterdam.
The Timor Deer is a short-legged, thickset animal. The facial
profile is remarkably concave, a character which I have frequently
noticed in Red Deer and Fallow Deer living under unfavourable cir-
cumstances. C. ¢imoriensis is not more than half the size of C. hippe-
laphus.
(3) CeRvUS MOLUCCENSIS.
1830. Cervus moluccensis, Quoy et Gaim. Zool. de | Astrolabe,
tom. i. p. 133, pl. 24.
1839-44. Cervus rusa, var. moluccensis, Schl. et Mull. Verh. Ned.
Bez. Zool. pp. 212, 220.
1836. Cervus moluccensis, Eyd. et Gerv. Mag. de Zool. tom. vi.
“26.
1839. —— ——, Eyd. et Gerv. Zool. de la Favorite, tom. v.
Mamm. p. 26.
Hab. Bouru (Quoy and Gaimard), Amboina (Schlegel and
Miiller), Celebes (Beecar%).
Specimens examined :—
Schlegel and Miiller’s types, Mus. Leyden.
Specimens collected by Beccari, Museo Civico, Genoa.
Cervus moluccensis isa much more delicately built, graceful animal
than C. timoriensis.
The following are the dimensions of one of Beccari’s specimens
CS: ,
Eng. inches. Metre.
Total length of skull in a straight line .. 11 0-280
From ant. rim of orbit to free extremity of
Pramaxiles COP PS ee, MI 57 =—-0"145
Length of upper molar series 72 SER
Length of upper premolar series ........ 14 0°035
Greatest length of nasals .............. 3°5 —0°088
Lengti-of premaxillee).¥ 0020 0 2 Le ODD (OOS
Across the lower rims of orbits at the
widest place .. 475 0-114
Length of humerus .... 79 0°200
Length of radius 72 —O' 186
1878.] CLASSIFICATION OF THE CERVID#. 905
Eng. inches. Metre.
Length of metacarpal cannon ......-... 6°71 0°155
Hhenetlt Of femurs) 66698 eo Se sa seeinah « 9-7 0°245
easth Or sibia wigs ts Gt Se coco aiid - 10°65 =: 0°265
Length of metatarsal cannon.. ... ) ee 7 0°178
(Rucervus, subgen.)
1838. Rucervus (subgen.), Hodgs. Ann. Nat. Hist. vol.i. p. 154;
J. As. Soc. Beng. vol. x. pt. ii. p. 914 (1841).
Antlers on short stout pedestals. Brow-antlers (fig. 4, a) very
powerful, projecting forwards at a right angle from the beam,
sometimes forked, and frequently developing rudimentary tines from
their upper surface. Beam curving boldly forwards along the upper
half of its length, and bifurcating into two upper tines, which in their
relative dimensions present specific differential characters.
Skull narrower in proportion to its length, in other respects re-
sembling that of Rusa: the bulle are moderately inflated, and the
upper and under molars have small supplementary columns. Tail
short. Neck maned. Stature large. Young spotted.
Distribution, Indian region.
Fig. 4.
Cervus (Rucervus) duvauceli,
13. CeRvVUS DUVAUCELI.
1835-37. Cervus duvaucelli, Cuv. Oss. foss. (ed. 4) tom, vi.
p- 89, pl. 201. figs. 6-8,
1871. —— ——, Sclat. Trans. Zool. Soc. vol, vii. p. 346, pl. 36.
Hab. Nepal; Assam; Denwa valley, Mahadeo hills, Central
India (Forsyth).
14. CeRVUS SCHOMBURGKI.
1863. Rucervus schomburgki, Blyth, P. Z. S. 1863, p. 155, fig.
1867. , Blyth, P. Z. 8. 1867, p. 835, figs. 6-12.
1876. Cervus schomburgki, Brooke, P. Z. 8. 1876, p. 304,
figs. 1, 2: Scl. P. Z. S. 1877, p. 682.
Hab. Siam. Types, Brit. Mus,
906 SIR V. BROOKE ON THE [Nov. 19,
15. Cervus ELDI.
1842. Oervus eldi, auct. anon. Cale. Journ. vol. ii. p. 417+
1867. , Beavan, P. Z. 8. 1867, p. 759.
1871. , Sclat. Trans. Z. S. vol. vii. p. 348, pls. 37, 38.
Hab. Burmah; Siam ; Hainan (Swinh.).
(ELapHuRus, subgen.)
1866. Elaphurus, Alph. M.-Edw. Compt. Rend., and Ann. Sc. Nat.
ser. 5, tom. v. p- 380, and Nouv. Arch. du Mus. 1866, vol. il. p. 27-
Antlers dichotomous, about twice the length of the head, pedestals
short. The anterior branch of the antlers (fig. 5, 6) more strongly
developed than the posterior branch (fig. 5, c-), and in adult specimens
bifurcating and carrying many strong tines. Facial portion of the
skull much elongated. Ascending rami of the praemaxillee of great
length, articulating broadly with the nasals. Nasals long, and ex-
Fig. 5.
Cervus (Elaphurus) davidianus.
panded posteriorly between the large anteorbital vacuities, of which
they form the larger part of the superior border. Lacrymal pit
deep and large. Upper molars with supplementary columns. Ears
small and narrow. Eye small. Tail long. Neck maned. Stature
large. Young unspotted ?
Distribution. North-eastern Palearctic region ?
16. CERVUS DAVIDIANUS.
1866. Elaphurus davidianus, Alph. M.-Edw. Nouy. Arch. du
Mus. 1866, vol. ii. p. 27, pls. 4, 5, 6.
1871. , Sclat. Trans. Z. S. vol. vii., p. 333, pl. 28.
Hab. Mantchuria. The specimens which have reached Europe
1878.] CLASSIFICATION OF THE CERVID. 907
were brought from the imperial park, south of Pekin. Types,
Mus. d’Hist. Nat. Paris.
(Axis, subgen.)
1827. Axis (subgen.), Ham. Sm., Griff. An. Kingd. vol. v. p. 312.
Antlers about three times the length of the head, supported on
slightly elongated pedestals. Brow-antler (fig. 6, a) rising at some-
thing less than a right angle from the beam (fig. 6, a), which bifur-
cates at a little above the half of its entire length. Of the tines
thus formed, the external and anterior tine (fig. 6, 6) is much the
longer. Skull and rhinarium as in Rusa. Upper canines wanting.
Molars with small supplementary columns. Both sexes at all ages
and seasons distinctly spotted with white. Stature medium.
Distribution. Western portion of the Indian region.
Fig. 6.
Cervus (Axis) axis.
17. CERVUS AXIS.
1877. Cervus axis, Erxl. Syst. Reg. An. p. 312.
1843. Axis maculata, Gray, Spec. Mamm. Brit. Mus. p. 178.
Range. British India; Ceylon.
(PsEuDAXIS, subgen.)
1872. Pseudaxis, Gray, Cat. Rumin. Mamm. Brit. Mus. p. 70.
Antlers about twice the length of the head, supported on short
stout pedestals. Brow-antler (fig. 7, @, p. 908) rather short, directed
upwards at a rather acute angle with the beam (fig. 7,7). A
strong tine (fig. 7, 6) is developed from the anterior surface of the
antler at about half its entire length, and a short tine (fig. 7, d) from
the posterior surface of its upper third.
Lacrymal pit of moderate depth and extent, its antero-posterior
diameter being less than that of the three upper molars. Ante-
orbital vacuity moderate. Auditory bull moderately inflated, com-
pressed, and smooth externally. Rudimentary upper canines pre-
sent in both sexes. Rhinarium as in Rusa, with the exception of
the internarial portion, which is slightly less prolonged upwards.
908 SIR V. BROOKE ON THE [Nov. 19,
Metatarsal tuft whitish. Tail moderate. Neck maned. In summer
spotted ; in winter uniform brown. Black bands on each side of the
pure white anal disk form a cross with the narrow black streak along
the dorsum of the tail. Stature medium. Young indistinctly spotted.
Distribution. Eastern Palearctic region.
18. CERVUS SIKA.
1847. Cervus sika, Temm, et Schl. Jap. Mamm. p. 44, pl. 17.
1860. , Sclat. P. Z.S. 1860, p. 377.
1871. ——, Sclat. Trans. Z.S. vol. vii. p. 346, pl, 35,
Hab. Japan. Type Mus. Leyden.
Cervus (Pseudaxis) sika.
19. Cervus evorts?
1874. Cervus euopis, Swinh. P. Z.S. 1874, p. 151.
Hab. North China? Type, ¢, Zool. Soc. Menag.
The type, which is still living, differs in no appreciable external
characters from ordinary specimens of Cervus sika. It was ori-
ginally received from Tientsin, China. I have not included C.
euopis among the synonyms of C@. sika,in the hope of attracting
further attention to the subject, as it would be of considerable in-
terest should C. stka be found inhabiting the mainland as well as the
islands of Japan.
20. CERVUS MANTCHURICUS.
1861. Cervus pseudazis, Gray, P. Z.S. 1861, p. 236, pl. 27 (nec
Eydoux et Souleyet ?)'.
1864. hortulorum et mantchuricus, Swinh. P. Z.8. 1864,
p- 169.
1865. mantchuricus, Swinh. in lit. P. Z. 8. 1865, p. 1.
1871, -—— ——,, Sclat. Trans. Z. S. vol. vii. p. 344, pls. 31, 32.
mandarinus, Alph. M.-Edw. Rech. Mamm. p. 184, pls,
22; 22 Aas
Hab. Neighbourhood of Newchwang. Type, Mus. d’Hist. Nat.
Paris,
1 Cf. infra, p. 909.
1878. ] CLASSIFICATION OF THE CERVID &. 909
The Society has within the last few years received living speci-
mens of a Pseudaxis from Japan, which are intermediate in size &e.
between P. sika and P. mantchuricus. These have, in my opinion,
with excellent judgment, been labelled by the Secretary as Cervus
mantchuricus minor. I think it probable that, when a larger series,
of Pseudaxine Deer are brought together, it will be found impos-
sible to separate them into definite species, but that it will be found
necessary to regard them as one species of wide geographical range,
endowed with a constitution sufficiently elastic to enable it to support
very varied conditions.
24. Cervus pysowsxk11?
1876. Cervus dybowskii, Tacz. P. Z. S. 1876, p. 123, fig.
axis? Radde, Reise im Siid. von Ost-Sib. Bandi. p- 286.
Hab. Upper Ussuri (Mantchuria).
It is highly probable that, when we know more of this form, it will
be found inseparable from C. mantchuricus.
22. Cervus KopscuHt ?
1873. Cervus kopschi, Swinh. P. Z. 8. 1873, p. 574.
Hab. Department of Kienchang, Kiangse, China (Swinh.). Types,
Brit. Mus.
Also a very doubtful modification. had an opportunity of exa-
mining the types upon their arrival, and could perceive no particulars
in which they differ from O. mantchuricus.
23. Cervus TARVANUs.
1862. Cervus taévanus, Sclat. P. Z. S. 1862, p. 152, pl. 16.
, Swinh. loc cit. p. 362.
1871. , Sclat. Trans. Z. S. vol. vii. p- 345, pls. 33, 34.
Hab. Mountains of Formosa.
24. Cervus CaAspicus.
1874. Cervus caspicus, Brooke, P. Z. &. 1874, p. 42.
Hab. Mountaing §.W. of the Caspian.
Since my original notice of this supposed species, I have received
no further information respecting it. The antlers are in my private
collection ; and I am still unable to refer them to any known species.
Although I have placed the species provisionally amongst the Pseud-
axine Deer, I am very far from satisfied that this is its proper loca-
tio. I am chiefly influenced in doing so by an antler I have
lately received from Mr. Robertson (H.B.M. Vice-Consul, Busra)
from the Karun, which is of decided Pseudaxine character, and may
possibly belong to the same species. It is to be hoped that, through
the indefatigable exertions of Mr. Robertson, we may ere long have
the advantage of seeing this species living in the Society’s Mena-
erie.
4 I have hesitated in referring the Cervus pseudaxis of Eydoux and
Souleyet (‘ Voy. de la Bonite,’ vol. i. p. 64, pl. 12) to any species of
this subgenus. The specimen is still preserved in the Muséum @’ His-
910 SIR V, BROOKE ON THE [Nov. 19,
toire Naturelle at Paris; but though I have often carefully examined
it, the absence of the skull and the great uncertainty of the locality
where it was procured render it impossible to form a decided
opinion.
(Cervus, subgen.)
1766. Cervus, Linn. Syst. Nat. (ed. 12) p. 92.
Antlers differ from those of Pseudaxis in having a second brow-
antler (fig. 8, a’) developed above the brow-antler (fig. 8, a), in the
brow-antler being curved forwards at a right angle with the beam
(fig. 8, x), and in the posterior coronal tine (fig. 8, @) being more
strongly developed than the anterior coronal tine (fig. 8, ce). Skull as
in Pseudaxis. Molars sometimes with small supplementary columns.
Cervus (Cervus) elaphus.
Upper border of the internarial portion of the rhinarium forming
an acute angle, with apex directed forwards. Infranarial portion of
rhinarium not developed ; the prenarial portion much constricted,
but spreading out before joining the upper lip. Tail short. Neck
maned. A patch of lighter colour than that of the body surrounds
the tail. Stature large. Young spotted.
Distribution. Paleearctic and Nearctic regions.
25. CERVUS ELAPHUS.
1766. Cervus elaphus, Linn. Syst. Nat. (ed. 12) p. 93.
1837. barbarus, Benn. List An. Gard, Zool. Soe. 13th pub.
p- 31 (sine descr.).
Range. Co. Kerry (Ireland) ; Cumberland; Devonshire ; high-
lands of Scotland, and following islands—Harris, Skye, Rum, Mull, ,
Jura, and Arran ; island of Hitteren (Norway); Southern Sweden ;
1878.] CLASSIFICATION OF THE CERVIDZ, 911
France; Spain; Germany; Austria; Turkey; Greece; Corsica ;
Sardinia; Province of Constantine (Algeria) ; Asia Minor; Cau-
casus.
A comparison of the gigantic antlers of the Red Deer of the 15th,
16th, and 17th centuries preserved in the old hunting Schloss at
Moritzburg (built 1540), near Dresden, with the antlers of Red Deer
from the islands of Harris or Sardinia, shows in a most striking
manner the great variations to which this species is subject. Some
of the antlers at Moritzburg measure 50 inches along the outside
curve, are 10 inches in circumference round the smallest part of the
beam, and the two antlers carry from 24 to 50 points. The spread
between the coronal tines of one specimen is 74 inches. Antlers from
Harris and Sardinia rarely exceed 30 inches in length, their cireum-
ference being about 4 inches; they very rarely carry a larger num-
ber than 12 points; and their span seldom exceeds 28 inches.
I have considered the African Deer, Cervus barbarus, specifi-
cally identical with Cervus elaphus. Specimens from Corsica and
Sardinia completely bridge over the characters which have been ad-
vanced as distinctive of the African and European races. The second
brow-antler is rarely developed in the African form ; but I have known
instances of its occurrence.
26. Cervus XANTHOPYGUS.
1831. Cervus elaphus, Pall. Zoogr. Rosso-Asiat. vol. i. p. 217.
1853. ——, Midd. Sib. Reis. Wirb., Band ii. Th. 2, p. 120.
1859. ———- ——, Schrenck, Reis. u. Forsch. Amurlande, Band
i. p. 170.
1862. ——- ——, Radde, Reis. im Siid. v. Ost-Sib. Band i. p. 284.
1867. wanthopygus, Alph. M.-Edw. Ann. Sc. Nat. sér. 5,
vol. viii. p. 376; Rech. s.1. Mamm. p. 181, pl. 21.
1876. —— elaphus, Prejevalsky, Mongolia (Engl. trans.), vol. i.
p- 164.
Range. From the Caucasus to the Altai, and thence round
Baikal through Dauria, as far as the Lena and Witim (Pallas) ;
thence to the mountains surrounding the sources of the Silimdsha
and Bureja (Middendorf’) ; along the Amoor, as far eastwards as
the Gorin and the Chelasso, and thence southwards to the sea-
coast, a few days’ journey south of the Bay of Kastries (Schrench) ;
Chinghan Mountains (Radde) ; Munniula Mountains (China, lat.
of Pekin) (Prejevalsky).
Type, Mus. d’Hist. Nat. Paris.
This species was founded provisionally by Professor Alphonse
Milne-Edwards upon the examination of a single specimen, certain
cranial peculiarities and the large size of the anal disk deciding him
‘that it was expedient for the time being to keep the form separate
from Cervus elaphus. It is by no means certain that Cervus xantho-
pygus (should the species ultimately prove distinct from Cervus
elaphus, which I greatly doubt) is the form found in all the localities
above mentioned. The exact limits of the distribution eastwards of
Cervus elaphus, Cervus eustephanus (Blanf.), and Cervus maral
912 SIR V. BROOKE ON THE [Nov. 19,
(Ogilby) remain as yet unknown; and it is fully possible that the
Deer met with by some of the celebrated travellers above quoted may
have appertained to one or more of these species.
27. CERVUS EUSTEPHANUS.
1875. Cervus eustephanus, Blanf. P. Z.S. 1875, p. 638, fig.
Hab. Thian-Shan mountains.
This species is only known from its antlers, which are of immense
size. Specimens which I have seen, in their great size and flattened
crowns so closely resemble antlers of Cervus canadensis that it would
be impossible to decide to which species they had belonged.
28. CERVUS MARAL.
1840. Cervus maral, Ogilby, Rep. Counc. Zool. Soc. 1840,
. 22.
‘ 1871. , Sclat. Trans. Z. 8. vol. vii. p. 336, pl. 29.
Hab. Circassia, Persia.
Cervus maral differs in a marked manner from the allied species
C. elaphus and C. cashmeerianus in the much greater length of its face,
as shown by the table given below. The Red Deer of which the
measurements are there given was a remarkably large old male. A
pair of Cervus maral, which lived at large in one of my parks for
some years, kept entirely apart from the Red Deer inhabiting the
same park. They bred together; and during the rutting-season the
species never showed the faintest desire to cross. This was the more
remarkable as the old stag Maral, though considerably larger in size,
lived in great fear of the Red Deer stags, which during that season
roamed incessantly through the park in search of hinds, but at all
times treated the female Maral with sovereign disdain, although at
any moment they could have taken possession of her had they so
desired.
The limits of the eastern range of Cervus maral are as yet un-
defined.
Cervus maral. | Cervus elaphus. | C. cashmeerianus.
Eng. in. | metre | Eng. in. | metre| Eng. in. | metre
16
Total length of skull ...| 18 0-457 0-405) 16 0-405
From oce. prominence to
ant. rim of orbit ... .. 76 | 0-193 7-5 | 0-190 7 0-178
From ant. rim of orbit
to free extremity of the
preemaxille..............- 11 0:280 9 0229 88 | 0:223
Length of upper molar
and premolar series ...| 5 0°127 45 |0114 45 |0114
29. CreRVUS CASHMEERIANUS.
1839. Cervus cashmeerianus, Falconer, MS. (non vidi).
1871. , Sclat. Trans. Z.S. vol. vii. p. 339, pl. 30.
Hab. Cashmere.
The voice of the Cashmere Stag in the rutting-season differs greatly
1878. ] CLASSIFICATION OF THE CERVIDZ. 913
from that of the Red Deer in the same season. In the former it isa
loud squeal ending in a more guttural tone; in the latter it is a
distinct roar resembling that of a Panther. The voice of the Wapiti
resembles that of the Cervus cashmeerianus. The second brow-
antler (fig. 8, a’) in Cervus cashmeerianus, with very rare excep-
tions, exceeds the brow-antler (fig. 8, a) in length, a peculiarity
by which the antlers of this species may be distinguished from those
of its allies.
30. CERVUS AFFINIS.
1835. Cervus affinis, Hodgs. Journ. As. Soc. Beng. vol. iv. p-
648, pl. 53. fig. 5.
1871. , Sclat. Trans. Z. 8. vol. vii. p. 343.
Cervus wallichii, Cuv. Oss. foss. (ed. 4) tom. vi. p. 88; and F. Cuv.
Hist. Nat. Mamm. pl. 356.
Hab. Choombi valley (Eastern Himalayas).
Should Cervus wallichii, Cuv., prove to be specifically identical
with Cervus affinis, Hodgs., the former name, having priority, must
stand. The antlers of the type of Cuvier’s original description are,
I believe, still preserved in the Museum of the Asiatic Society of
Calcutta ; F. Cuvier’s plate shows them to be of decidedly abnormal
growth.
31. Cervus CANADENSIS.
1777. Cervus canadensis, Erxl. Syst. Régn. An. p. 305.
1857. Cervus canadensis, Baird, N. Am. Mamm. p- 638.
ftange. Alleghany regions of Pennsylvania and Virginia, Northern
Wisconsin ?, Minnesota, Dakota, Nebraska, Washington, Oregon,
and California ; 57° N. is its northern limit (Baird).
(Dama, subgen.)
1827. Dama(subgen.), Ham. Sm., Griff. An. Kingd. vol. v. p.306.
Antlers about twice the length of the head, palmated, the tines on
the upper half of the antler rising from their posterior surface (figs.
9and 10). Skull resembling that of Cervus, but more depressed,
broader proportionally, and the orbits more prominent. Auditory
bull subglobular; no upper canines; rhinarium as in Cervus.
Tail moderate. Prepuce furnished with a tuft of long hair. Both
sexes spotted in summer, unspotted in winter. Black markings on
the rump and tail as in Pseudavis. Stature medium. Young
spotted.
Distribution. South-western Palearctic region.
32. CeERVUS DAMA.
1766. Cervus dama, Linn. Syst. Nat. (ed. 12) i. p. 93.
1843. Dama vulgaris, Gray, Spec. Mamm. Brit. Mus. p. 181.
1874. Cervus dama, Jeitt. Zool. Gart. 1874, p. 288, (translated by
Sclater) Nature, vol. ii. p. 71 ; vide also Boyd Dawkins, tom. cit. pp-
112, 210, 226, and Brooke, tom. cit. p. 210, figures.
Range. Greece, Spain, Anatolia, island of Rhodes, Sardinia, N.
Palestine ; frontier of Tunis in Algeria? (fide Loche). ;
914 SIR V. BROOKE ON THE [Nov. 19,
The semidomesticated life which the Fallow Deer has lead for so
many generations in the parks of Europe has given rise to many
permanent variations of colour. The natural colour of the species in a
wild condition in summer is a brilliant fawn with white spots irregu-
larly distributed over the upper parts of the sides and haunches, this
spotted region being bounded inferiorly on the sides and posteriorly
on the haunches by indefinite white lines. In winter the spots en-
tirely disappear.
Cervus (Dama) mesopotamicus.
33. CERVUS MESOPOTAMICUS.
1875. Cervus (Dama) mesopotamicus, Brooke, P. Z. S. 1875,
p. 265, pl. 38. figs. 1, 2.
1876. , Brooke, P. Z. S. 1876, p. 298, figs. 1-4.
Hab. Luvristan hills (Persia).
1878. ] CLASSIFICATION OF THE CERVID&. 915
Since my description of the species, the Society has received two
living specimens( ¢ and 2) from Mr. Robertson, to whom I was in-
debted for my original knowledge of its existence. Cervus mesopo-
tamicus is a very much larger animal than C. dama.
B. Distal ends of metacarpi remaining. (TELEMETACARPI.)
Posterior portion of the narial cavity not divided by the vomer
into two distinct chambers.
Tuft of hair on the external surface of the metatarsus when
present above the centre of that bone. Tuft on inside of the
tarsus absent.
4. ALcEs.
1827. Alces (subgen.), Ham. Sm., Griff. An. Kingd. vol. v. p. 303.
Antlers dichotomous, both main branches palmated, the posterior
branch (fig. 11, 6) much surpassing the anterior branch (fig. 11, a) in
dimensions. Anterior nares of great extent, owing to the shortness
of the nasals. Lacrymal pit and anteorbital vacuity moderate.
Fig. 11.
=>
S
—S
Se =
Alces machlis.
Ascending rami of the premaxillz long, not articulating with the
nasals. Incisors decreasing gradually in size from the central to the
exterual pair.
A very small triangular rhinarium. Head very long, neck and
body short, legs very long, stature great. Young unspotted.
Distribution. North Palearctic and Nearctic regions.
916 SIR V. BROOKE ON THE [Nov. 19,
ALCES MACHLIS.
1836. Alces machlis, Ogilby, P. Z.S. 1836, p. 135.
1834-7. americanus, Jard. Nat. Libr. xi. p. 125.
Range. Sweden, Lithuania? Northern Russia, Altai Mountains ?
mountains to the west of Lake Baikal. Apple Mountains (west of
Chita), Khinghan Mountains ; forests of the Lower Amoor as far
south as the mouth of the Ussuri, in lat. 48° (Radde, &c.). ‘ West
coasts of America from the shores of the Arctic Ocean nearly to the
Columbia River. Further east the northern limit is about 65° and
thence through Canada to Maine, New Hampshire, Vermont, and
the north parts of New York” (Baird).
5. Hypropores.
1870. Hydropotes, Swinh. P. Z.S. 1870, p. 90.
Without antlers in either sex. Lacrymal pit deep, small, its
antero-posterior diameter about equalling that of m. 3. Anteorbital
vacuity moderate. Ascending rami of the premaxille strong.
Orbits small and but slightly prominent, their greatest diameter less
than the extent of the three upper molars. Auditory bulle con-
siderably inflated, a portion intervening between the tympanohyal
and the paroccipital process. Angle of lower jaw much produced
backwards, forming a compressed semicircular prominence. Rami
of lower jaw between the dental foramen and the incisors compressed
and raised into sharp ridges, the edges of which are everted. Ca-
nines of immense size, in the male curved downwards, and gently
convergent, pulp non-persistent. Rhinariumample. No metatarsal
tufts. Interdigital glands in hind feet deep, in fore feet slightly
developed and shallow. Spigelian lobe of liver Rusiform (jide Garrod).
Young spotted distinctly, from 3 to 6 atabirth. Stature small.
Distribution. Eastern Palearctic region.
HyDROPOTES INERMIS.
1870. Hydropotes inermis, Swinh. P. Z. 8. 1870, p. 89, pls. 6, 7.
1871. , Hams? ZS. 1871, p. 258.
1877. 5 Gart. PAS: 1877, p.'789.
Hab. Islands of the Yang-tse-Kiang.
6. CAPREOLUS.
1827. Capreolus (subgen.), Ham. Sm., Griff. An. Kingd. vol. v.
p- 313.
Antlers less than twice the length of the head. Normal number
of tines 6. Brow-antler (fig. 12, a) developed from the an-
terior surface of the upper half of the antler, directed upwards.
Lacrymal pit very shallow and of small extent. Anteorbital vacuity
small. Ascending rami of the premaxille articulating generally
but not invariably with the nasals. Auditory bulle very slightly
inflated, their external surface rugose. Angle of lower jaw as in
Hydropotes, Crowns of central incisors moderately expanded.
1878. ] CLASSIFICATION OF THE CERVID. 917
Upper margin of the internarial rhinarium running straight across
from the upper angle of the nostrils. No infranarial rhinarium.
Tail very short. In the female the clitoris is surrounded by a long
tuft of hair. Interdigital glands in the fore feet rudimentary, in the
hind feet deep, opening by a narrow orifice into a capacious round
pouch. Stature small. Young spotted.
Distribution. Central and Western Palearctic region.
Capreolus caprea.
1. CAPREOLUS CAPRA.
1843. Capreolus caprea, Gray, Cat. Spec. Mamm. Brit. Mus.
p- 176.
1844. Capreolus europeus, Sund. Pec., Vetensk. Akad. Handl.
1844, p. 184.
Range. Scotland, Southern Sweden, France, Germany, Austria,
Hungary, Spain, Tuscany, Greece, Turkey, Northern Palestine,
Elburz Mountains (South of Caspian).
I have received a specimen from the last-mentioned locality which
differs in no respect from the European specimens.
2, CAPREOLUS PYGARGUS.
1771. Cervus pygargus, Pall. Reise Russ. Reichs, vol. i. p. 97,
Append. p. 453
1831. Cervus capreolus, Pall. Zoogr. Rosso-As. vol. i. p. 219.
1853. , Midd. Sib. Reis. Wirb. Band ii. Th. 2, p. 118.
~ 1859. ——- ——, Schrenck, Reis. u. Forsch. Amurlande, Band i.
p- 163.
1862. , Radde, Reise Siid. v. Ost-Sib. Band i. p. 277.
Range. Suitable localities in the mountains forming the watershed
between the Russian and Chinese empires, in Central Asia ; Chingan
Mountains (Mantchuria).
Most authors have considered C. pygargus to be a large variety of
the common Roe. There are, however, in the British Museum two
Specimens purchased from Brandt and said to be from Siberia,
which, in their very much larger size, larger anal disk, and much
more hairy ears, differ widely from all specimens of the common
species. Specimens of Roe from Mantchuria in the British and
Paris Museums are very much smaller than the above-mentioned
Proc. Zooxu. Soc.—1878, No. LX. 60
918 SIR V., BROOKE ON THE [Nov. 19,
specimens; they also differ from the larger specimens and the common
Roe in some details of coloration. It is not improbable that the
Mantchurian Roe may prove to be a third modification of the form.
C. Distal ends of the lateral metacarpals remaining. (TELEMETA-
CARPI.)
Posterior portion of the nasal chamber divided by the vomer into
two distinct chambers.
Tuft of hair on the external surface of the metatarsus, when pre-
sent, below the centre of that bone. Tuft on the inside of
the tarsus frequently present.
Ascending rami of the preemaxillze not reaching the nasals. (Ex-
ception, subgen. Furcifer.)
7. Cartacvs.
(Cartacus, subgen.)
1827. Mazama (subgen.), Ham. Sm., Griff. An. Kingd. vol. v.
p- 314 (nec Rafinesque)’.
1850. Cariacus, Gray, P. Z.S. 1850, p. 237.
Antlers not greatly exceeding the head in length, on short
pedestals. A short upright brow-antler (fig. 13, a) rises from the
inner surface of the lower third of the antler some little distance
above the burr. The antlers are curved boldly forwards, down-
wards, and inwards, one or more tines being developed from their
Fig. 13.
Cariacus (Cariacus) virginianus,
convex or posterior surface. Skull elongated and narrow. Laery-
mal pit of moderate extent, owing to the very large anteorbital vacuity,
which encroaches on the region usually occupied by the facial plate
of the lacrymal. WNasals forming the larger part of the superior
border of the anteorbital vacuity. Auditory bulle slightly inflated.
No upper canines; molars sometimes with small supplementary
columns; central incisors bnt slightly spatulate. A tuft of long
2 Mazama, of Rafinesque (1817), is too ill-defined to be used for either the
Rocky-Mountain Sheep or the Pronghorn ; but its publication invalidates its
subsequent use in any other sense. Cf, Baird, N.-Am. Mamm. p. 665.
1878. ] CLASSIFICATION OF THE CERVIDE. 919
hair on the inner surface of the limb at the tarsal joint. Rhinarium
as in Cervus. Tail long. Stature medium. Young spotted.
Distribution. Nearctic and northern portions of the Neotropical
region.
(a) Antlers non-dichotomous. Lacrymal of moderate depth.
1. CARIACUS VIRGINIANUS.
1774. Cervus virginianus, Bodd. Elench. An. vol. i. p. 136 (non
vidi).
1788. —— ——, Gm. L. Syst. Nat. p. 179.
1857. , Baird, N.-Amer. Mamm. p. 643, figs. 12, 13.
Range. From Maine all over the United States east of the Mis-
souri (Baird).
2. CARIACUS LEUCURUS.
1829. Cervus leucurus, Dougl. Zool. Journ, vol. iv. p. 330.
1857. , Baird, N.-Amer. Mamm. p. 649, figs. 14-18.
1873. —— , Walsingham, P. Z.S8. 1873, p. 561.
Range. N. California, Oregon principally east of the Cascade
Mountains (Walsingham), Washington, Dakota west of the Mis-
souri, Nebraska (Baird).
3. CARIACUS MEXICANUS.
1827-1834. Cervus mexicanus, Licht. Darst. d. Thiere, Taf. 18.
1827. Cervus nemoralis, Ham. Sm., Griff. An. Kingd. vol. iv.
p- 137, plate.
1857. , Baird, N.-Amer. Mamm. 653.
Range. Texas, Mexico, Sonora (Baird); Guatemala, Honduras,
Nicaragua (Salvin) ; Costa Rica (Saussure) ; Panama (Salvin).
Lichtenstein’s types, ¢ et 2 Mus. Berlin.
The name Cervus mexicanus was originally given by Gmelin (Syst.
Nat. p. 179) to the Mexican deer of Pennant, described in his
‘Quadrupeds,’ p. 110. The description and dimensions given by
Pennant are reconcilable with the Cervus meaicanus of Lichtenstein ;
but the antlers figured by him (plate xi. figure 3) are certainly not
referable to the same form. They appear rather to represent ab-
normal antlers of Cariacus macrotis. The specimens described by
Lichtenstein were obtained by Graf von Sack in Mexico, and are in
all particulars thoroughly typical of the Deer inhabiting the localities
I have given in the range of this species.
I give the dimensions of a female specimen collected by Mr. Salvin
in Guatemala, and of the antlers of a male collected by the same
gentleman in the same locality. The latter resemble in every par-
ticular antlers collected by Mr. Salvin in Nicaragua and Panama.
There are not more than six points, inclusive of the brow-antler, on
both antlers in any of the specimens collected between Guatemala
and Panama; but I have seen specimens from Mexico and Texas
with eight and ten tines; and one splendid specimen collected by
Mr. Dresser near Friedrichsburg in Texas has as ee fifteen.
0
920 SIR V. BROOKE ON THE [Nov. 19,
The Texan deer are decidedly larger than those inhabiting Guate-
mala, Nicaragua, and Panama.
Dimensions of Cariacus mexicanus.
Engl. inches. Metre.
© Heicht at shoulder 0... ceca en 27°5 0°700
Lengthvof ears «0253... . sah 4:3 0°110
Length of tail exclusive of hair...... 5:5 0°140
Total length of skull in a straight line 90 0°230
Length from ant. rim of orbit to free
extremity of preemaxille ...,.... 4°5 0-114
Length of molar and premolar series
(Chi st eee es aaine tees Lain 2°7 0:067
@Lengthormntlers .°.).% . sane secww,.» 79 0°190
Greatest span........ Avis ovRodg © guvte aReRe 8:2 0°210
4. CaRIACUS SIMILIS ?
1852. Cervus similis, Puch. Nouv. Arch. du Mus. 1852, p. 357,
pl. 26.
Hab. Unknown. Type, ¢ Mus. d’Hist. Nat. Paris,
5. CARIACUS SAVANNARUM.
1848. Cervus savannarum, Cab. & Schomb. Reis. Brit. Guian.
vol. ili. p. 785.
1852. Cervus gymnotis, Puch. Nouv. Arch. du Mus. 1852, p. 348,
pl. 25, pl. 23. figs. 2-10.
Hab. Guiana, Venezuela? Type, g Mus. Berlin.
6. CARIACUS PERUVIANUS.
1874. Coassus peruvianus, Gray, Ann. & Mag. Nat. Hist. (ser. 4)
vol. xiii. p. 332,
Hab, Peru. Types, 2 Brit. Mus., 2 in my own collection.
The number of specimens from well-authenticated localities at
present existing in European collections is far too small to enable me
to form any decided opinion respecting the degree of persistency of
modification in the above-mentioned forms. Notwithstanding, I
think it highly probable that when we possess a sufficiently large
series we shall find that C. peruvianus and the four preceding species
are nothing more than climatic varieties of C. virginianus, connected
inseparably by every shade of intermediate forms. Mr. Allen, in
his valuable paper on the geographical variation of North-American
Mammals, especially in respect to size (Bull, U.S. Survey, 1876,
p- 304), has conclusively proved that a decrease in size and charac-
teristic development takes place in all mammals the centre of whose
area of distribution is in North America, as they spread southwards ;
and, so far as the materials at my command enable me to judge, I
think it probable that the characters upon which C. leucurus, C.
mexicanus, C. similis, C. savannarum, and C. peruvianus have been
established as species may be in large measure the result of this law.
1878.} CLASSIFICATION OF THE CERVIDH. 921
7. CARIACUS GYMNOTIS.
1833. Cervus gymnotis, Wiegm. Isis von Oken, p. 965.
Hab. New Granada. Type, ¢ Mus. Berlin.
This is, I think, a distinct species. The large drooping ears
(naked externally), the very narrow head, and the remarkably slender,
delicate form of the animal render it easily distinguishable from
specimens of the ailied species,
8. Carracus ToutEcus.
1860. Cervus toltecus, Sauss. Rey. et Mag. Zool. (ser. 2) tom. xil.
p. 247.
Hab. Vera Cruz.
M. Saussure gives 6-9 inches as the total length of the skull of
his type specimen, which was a male of at least two years of age, as
evidenced by his antlers, which are not those of thé first year. If
this measurement is correct, Cariacus toltecus is a very small species,
one third smaller than C. mexicanus from the same locality,
(8) Antlers dichotomous, Lacrymal pit deep.
9. Carracus MAcRoTIs.
1823. Cervus macrotis, Say, Narr. Long’s Exp. vol. ii. p. 88.
1857. » Baird, N.-Amer. Mamm. p- 656.
1873. , Walsingham, P. Z. 8. 1873, p. 561.
Range. Oregon, Nebraska, Dakota west of the Missouri, Colorado,
Kansas. Fie. 14,
Cariacus (Cariacus) macrotis,
10. Cartacus COLUMBIANUs.
1829. Cervus macrotis, var. /3. columbianus, Rich, Faun, Bor.-Am.
p- 257,
922 SIR V. BROOKE ON THE [Nov. 19,
1848. Cervus lewisii, Peale, Mamm. Birds U.S. Expl. Exp. p. 39
(fide Baird).
1850. Cariacus punctulatus, Gray, P. Z.S. 1850, p. 239, pl. 28.
1857. Cervus columbianus, Baird, N.-Amer. Mamm. p. 659.
Range. California, Oregon, and Washington west of the Cascade
Mountains.
This species is about one third smaller than C. macrotis, which
it otherwise closely resembles. The two species are found on the
same ground in Washington.
(Biastocerus, subgen.)
1850. Blastocerus, Gray, P. Z.S. 1850, p. 237.
Antlers slightly exceeding the head in length, on short pedestals.
They are dichotomous, the anterior branch (figs. 15 and 16, a) infe-
rior to the posterior branch (0) in development. In adult speci-
mens the former sometimes, the latter always bifurcates. Lacrymal
Cariacus (Blastocerus) paludosus.
pit deep. Skull in all essential particulars resembles that of Ca-
riacus. Central incisors very slightly spatulate, and exceeding the
pair next to them but slightly in size. Tail short. No metatarsal
tuft. Tarsal tuft present.
Distribution. Central and Southern Neotropical region.
11. CarIACUS PALUDOSUS.
1820. Cervus paludosus, Desm. Mamm. p. 443.
, Licht. Darst. pl. 17 (1827-1834).
1872. , Hensel, Beitr. Kenntn. Saug. Siid-Bras. p. 95.
Range. South Brazil, Paraguay, Rio Grande do Sul, Uruguay.
1878.] CLASSIFICATION OF THE CERVID®. 923
12. CARIACUS CAMPESTRIS.
Cervus campestris, F. Cuv. Dict. Se. Nat. tom. vii. p. 484.
, Licht. Darst. pl. 19 (1827-1834).
» Hensel, Beitr. Kenntn. Siug. Siid-Bras. p. 96 (1872).
Range. Paraguay, Uruguay, Buenos Ayres, N. Patagonia.
Fig. 16.
Cariacus (Blastocerus) campestris.
(Furcirer, subgen.)
1850. Furcifer, Gray, P. Z.S. 1850, p. 236.
Antlers the length of the head, with a strong brow-antler (fig. 17, a)
curved forwards and upwards at a right angle to the simple beam
(fig. 17, «), which is gradually attenuated toa sharp point. The
entire beam is curved gently forwards. lLacrymal pit deep, but
of moderate antero-posterior extension. Auditory bulle very slightly
.
Fig. 17.
Cariacus (Furcifer) antisiensis.
inflated, their external surface rugose. Ascending rami of the pre-
maxille articulating with the nasals. Free extremities of the nasals
forming together a single point. Central incisors very slightly
exceeding the pair next to them in size, and but slightly spatulate.
Upper canines in both sexes. Rhinarium as in Cariacus. Stature
medium.
Distribution. South-western Neotropical region.
13. CARIACUS CHILENSIS.
1846. Cervus chilensis, Gay et Gerv. Ann. d. Sc. Nat. 1846, p. 91.
1849. Capreolus leucotis, Gray, P.Z.S.1849, p.64, Mamm. pl. 12.
924 SIR V. BROOKE ON THE [Nov. 19,
1875. Cervus chilensis, Sclat. P. Z. S. 1875, p. 45, figure.
Range. Andes “from Magellan to near Santiago, but far more
rare in the north than in the southern portion of its range” (Reed,
P. Z. S. 1875, p. 44). Type, 2 Mus. d’Hist. Nat. Paris.
14. CARIACUS ANTISIENSIS.
1834. Cervus antisiensis, D’Orb. Nouv. Arch. du Mus. iii. p. 91.
1869. Anomalocera huamel, Gray, Scient. Opin. 1869, p. 384.
1869. Xenelaphus huamel, Gray, P. Z.S. 1869, p. 497.
1872. Xenelaphus leucotis, Gray, Cat. of Rum. Mamm. p. 89.
1872. Xenelaphus anomalocera, Gray, Aun. Nat. Hist. (ser. 4)
vol. x. p. 445.
1873. Xenelaphus chilensis, Gray, Ann. Nat. Hist. (ser. 4) vol. xii.
Bees is
y 1874. Furcifer chilensis, Gray, Ann. Nat. Hist. (ser. 4) vol. xiii.
. 332.
i 1875. Cervus antisiensis, Sclat. P. Z.8. 1875, p. 46.
Range. Peruvian Andes. Type, ¢ Mus. d’Hist. Nat. Paris.
Naturalists owe a debt of gratitude to Mr. Sclater for the care and
clearness with which he has unravelled the meshes of confusion in
which the synonymy of the two very distinct species, Cariacus chi-
lensis (Gay) and C. antisiensis (D’Orb.), was formerly enveloped.
There is not the smallest foundation for a third species of Furcifer.
The type of Xenelaphus leucotis, &c. of Dr. Gray is preserved in
the British Museum, and is undoubtedly nothing but a very fine old
male C. antisiensis with strongly developed abnormal antlers.
(Coassus, subgen.)
1827. Subulo (subgen.), Ham. Sm., ‘Griff. An. Kingd. vol. y.
. 318%.
' 1850. Coassus, Gray, P. Z.S. 1850, p. 240.
Antlers simple spikes not exceeding half the length of the head
(fig. 18). Skull resembling that of Cariacus, the auditory bulle
less inflated and the facial profile more arched. Molars sometimes ~
Fig. 18.
<i5
Cariacus (Coassus) rufus.
with supplementary columns. Canines of uncertain occurrence.
Rhinarium ample, resembling that of Cervulus. No metatarsal tuft.
Tarsal tuft present. Stature small. Form heavy, with the back
much arched. Young spotted.
Distribution. Neotropical region.
1 Tf this name, which has been totally unused, were to be revived, it would
take precedence of Cariacus as the title of the whole genus. But it is too close
to Subula (Mollusca) of Schumacher, 1817.
1878. CLASSIFICATION OF THE CERVID®, 925
15. CARIACUS RUFUS.
1817. Cervus rufus, F. Cuv. Dict. Se. Nat. vol. vii. p. 485.
Hab. Surinam. Types, Mus. d’Hist. Nat. Paris, 2 ad. (No. in
Cat. 532), young (No. in Cat. 527).
16. CarIACUS SIMPLICICORNIS.
1827. Cervus simplicicornis, Ham. Sm., Griff, An. Kingd. vol. v.
p- 318.
Hab. Brazil (St.-Hilaire, Lalande), New Granada (Brooke).
Types:—Mus. d’Hist. Nat. Paris, 2 (No. 525 in Cat.); skull of same,
Mus. d’Anat. Comp. Paris (in Cat. 2208); skull of ¢ collected by
St.-Hilaire, in Mus. Anat. Comp. (in Cat. 2225).
17. CARIACUS RUFINUS.
1851. Cervus rufinus, Puch. Arch. du Mus. p. 491, pl. 30; id. Rev.
Mag. Zool. 185§, p. 561.
Hab. Ecuador (Pucheran), Guatemala (Salvin). Type, ¢ Mus.
d’ Hist. Nat. Paris.
18. CartIacus NEMORIVAGUS.
1817. Cervus nemorivagus, F. Cuv. Dict. Se. Nat. vol. vil. p. 485.
Hab. Surinam. Types, stuffed head of ¢ and skull of same
(Cat. 2223), Paris, in Mus. d’Hist. Nat. and Mus. d’ Anat. Comp.
I append a table of the comparative measurements of the four
last-mentioned species. The cranial measurements are taken from
adult females.
C. simplici- C. nemori-
| rufus, : C. rujfinus.
C. rufa cornis. fi vagus,
t |
Engl. Engl Engl. Engl
: metre.| - "| metre.|- | metre.} - * | metre.
inches. t inches. ure inches,| inches.) ™¢'re
Height at shoulder ...) 27 0-685 | 21 0535/19 | 0485) 19 | 0-485
Total length of skull
in a straight line... 9:2 | 0235] 7:10] 0-200) 65 | 0165) 62 | 0-157
From anterior rim of|
orbit to free extre- |
mity of premaxille| 49 | 0:124| 310! 0-098} 3:3 | 0-083) 3-1 | 0-078
Length of upper molar
and premolar series} 2°7 | 0-068) 23 | 0058} 21 | 0-053; 2 0-050
Greatest width be-
tween the orbits, |
measuring from the
0:076| 2-7
LOwenrIM fe5<s 50-03. 36 | 0:090| 32 | 0-080) 3: 0-068
|
From this it will be seen that Cariacus rufus is considerably the
largest, and C. nemorivagus the smallest and most delicately formed
species. C. rufus is red at all ages, and very heavily built. C. sim-
plicicornis is brown when young, never attains so brilliant a colour as
926 SIR V. BROOKE ON THE [Nov. 19,
C. rufus, and is much more gracefully shaped. C. rufinus, when
adult, is of a beautiful shining red, with the face and fore limbs strongly
shaded with bluish brown, as shown in exaggeration in Pucheran’s
figure (wide supra) ; and C. nemorivagus is of a dull pepper-and-salt
brown, and never becomes red at any age.
19, CARIACUS SUPERCILIARIS.
1850. Coassus superciliaris, Gray, P. Z.S8. 1850, p. 242, pl. 25.
Hab. Brazil (fide Gray).
20. CARIACUS WHITELYI.
1873. Coassus whitelyi, Gray, Ann. & Mag. Nat. Hist. (ser. 4)
xii. p. 163; id. Hand-list Edent. &c. 1873, p. 162, pl. 32. fig. 2.
Hab. Peru. Founded on a single skull of an immature animal in
Brit. Mus. (No. in Cat. 73. 6. 27. 2).
It is now many years since I commenced the study of this difficult
group of Cervide; but although I have examined the specimens
contained in nearly all the continental museums, and made a private
collection of some importance, I must confess that I am still far
from a satisfactory understanding of the subject. The complete
absence of cornual and cranial characters renders it exceedingly
difficult to grasp the characteristic peculiarities of the different mo-
difications of the form, six or seven of which are, I think, probably
persistent, and worthy of specific recognition by naturalists. In the
above list I have enumerated all the species (or supposed species) of
which the types are extant, and must leave it to the future to decide
whether they are valid or not. There is still much valuable work
to be done amongst the Cervidee by collectors in South America,
well-authenticated specimens from Rio Grande do Sul, Paraguay,
South Brazil and Bolivia being almost a blank in European museums.
The type of Coassus auritus, Gray, not having been preserved, I have
omitted the name from my list, as without the type it will be
impossible to decide with any certainty upon what species Dr. Gray’s
name was conferred.
8. Pupva.
1850. Pudu (subgen.), Gray, P. Z. 8. 1850, p. 242.
Antlers minute simple spikes. Lacrymal pit oval, very deep, its
antero-posterior extent slightly exceeding m. 3. Ascending rami of
the preemaxille reaching the nasals, which are considerably expanded
between the moderate anteorbital vacuities. Auditory bullz as in
Coassus. Molars without supplementary columns. Canines absent.
Central incisors slightly expanded, but exceeding those external to
them very much in size. Muzzle as in Cervulus, but with the
internarial and infranarial portions less extensive. Neither tarsal
nor metatarsal tufts. Ectocuneiform, navicular, and cuboid bones
in tarsus united. Stature very small.
Distribution, Chilian Andes.
So
bo
N
1878. ] CLASSIFICATION OF THE CERVID2.
PupvUA HUMILIS.
1831. Cervus humilis, Benn. P. Z.S. 1831, p. 27,
1871. Cervus pudu, Sclat. P. Z.S. 1871, p- 238, pl. 17.
Hab. Chilian Andes.
9. RANGIFER.
1827. Rangifer (subgen.), Ham. Sm., Griff. An. Kingd. vol. v.
p- 304.
Antlers developed in both sexes; a strong brow-antler (fig. 19, a) is
developed sometimes on both, but more frequently on only one antler,
that on the other being a mere rudiment. The brow-antler, when fully
developed, is directed downwards parallel with the face and palmated
anteriorly. A short distance above the brow-antler another similar
tine (fig. 19, 6) is developed. Above this the antler forms a bold
curve, the concavity forwards, its extremity (fig. 19, ¢) is palmated ;
and a short tine (d) is developed from its posterior surface at half
Rangifer tarandus.
its length. Portion of the skull anterior to the horns elongated,
that posterior to it remarkably short. Lacrymal pit and anteorbital
vacuity of moderate dimensions, the former shallow. Nasals much
expanded posteriorly. All the teeth remarkably small in proportion
to the skull; the incisors decrease very gradually in size from the
central to the external pair, they are none of them spatulate. Vomer
produced backwards nearly as far as the basilar suture. Frontal
928 ON THE CLASSIFICATION OF THE CERVID#&. [Nov. 19,
process for the support of the antlers carried back into the parietals,
this region of the skull being much restricted. No rhinarium,
Ears and tail short, neck maned, stature large. Young unspotted.
Distribution. Northern Nearctic and Palzearctic regions.
RANGIFER TARANDUS.
1766. Cervus tarandus, L. Syst. Nat. (ed. 12) p. 93.
1829. Cervus tarandus et var. artica et sylvestris, Rich. Faun.
Bor.-Am. pp. 238-251 (1829).
1857. Rangifer caribou et R. grenlandicus, Baird, N.-Amer.
Mamm. pp. 633, 634; American Reindeer, Hardy, ‘ Forest Life in
Acadie,’ pp. 120-163.
Range. Norway, Lapland, Spitzbergen, Nova Zembla, Western
Siberia from the Arctic Sea, about as far south as lat. 60° in Eastern
Siberia, to lat. 55° in the Yablonoi Mountains, to lat. 49° on the
Japanese sea-coast, and to lat. 46° on the island of Saghalien; Russian
America from the Arctic Sea, somewhat further south than the
Arctic circle? British America from Melville Island to Great Slave
Lake in the west, and in the east from Labrador to Nova Scotia,
Greenland, Newfoundland.
There are many well-marked varieties of Reindeer in this wide
and varied range; but the study of a considerable series has only
shown me the difficulty, if not impossibility, of defining them; and
consequently I have been constrained to include all under one title
in this synopsis, although I think the differentiation of the form
according to latitude and locality merits further attention.
EXPLANATION OF PLATE LY.
Fig. i. Left fore limb of Capreolus caprea, selected as an example of the TeLE-
METACARPI.
3. Metacarpal cannon, seen from the inside. 4. Ditto, seen from the
outside. 3/,4’, Phalanges of the 5rd and 4th metacarpals,
which coalesce to form the metacarpal cannon. 2-5. Distal
extremities of the 2nd and 5th metacarpals. 2'-5'. Their
respective phalanges. 7-7'. Place of origin and insertion of
the round ligament which prevents the upward displacement
of the 2nd and 5th metacarpals. ¢ Trapezium. @r and o,
Confluent trapezoid and os magnum. w. Unciform.
ii. Left fore limb of Cervus elaphus, selected as an example of the Puzsio-
METACARPI.
2,5. Proximal extremities of the 2nd and 5th metacarpals. s.
Metacarpo-phalangeal sesamoids. p. Proximal phalanges of
ae and 5th digits. (The remainder of the lettering as in
g. i.)
iii. Antler of the present Castle-Caldwell Fallow Deer, showing the exces-
sive development of the tine, d, and consequent contraction of the
alm, ¢.
iy. ious of one of the males from which the Castle-Caldwell herd of
Fallow Deer are descended.
1878. | SIR V. BROOKE ON A NEW GAZELLE. 929
2. On anew Species of Gazelle from Western Africa. By
Sir Victor Brooke, Bart., F.Z.S.
[Received November 19, 1878. ]
(Plate LVI.)
I have the pleasure of bringing before the notice of the Society a new
and exceedingly interesting form of Gazelle. As is well known, the
different species into which the genus Gazella is divisible (vide P. Z.S.
1873, p. 535) resemble each other so closely that it is often very
difficult to define them. The differences exist in nature, let their
taxonomic value be what it may; but in many cases they absolutely
baffle verbal description. __
In the form at present under consideration there is no such
difficulty. Although the materials at my command consist only of
the skulls of three males, with a small portion of the skin of the head
attached to one of them, characters are apparent which separate the
form from all known Gazelles. Indeed my only doubt is as to
whether the species should not constitute the type of a new subgenus.
The specimens belong to Mr. Gerald Waller, and it is through his
courtesy that I have the opportunity of exhibiting them this evening.
Mr. Waller tells me that the skulls were procured on the mainland
of Africa, north of the island of Zanzibar, about lat. 3° S. and long.
38° E. He is unfortunately unable at present to give any further
particulars respecting the species.
The new Gazelle differs from all hitherto described species in the
very great backward prolongation of the occiput. This extension is
is principally gained by the great size of the supraoccipital bone
and the prominence of the occipital crest. The skull is much more
depressed, the facial axis forms a more obtuse angle with the cranial
axis, and the rami of the lower jaw are more slender than in any
known Gazelle. The ascending rami of the preemaxille are remark-
ably thick, and do not nearly reach the nasals. In all other Gazelles
the molars and premolars are decidedly hypsodont; in the new
species they are as decidedly brachyodont, and, in comparison with
the skull, remarkably small. The horns do not differ greatly from
those of Gazella dorcas; they are, however, more massive and more
laterally compressed. ‘The skin of the head is of a uniform foxy
red, with white marks under the eye. Mr. Waller has expressed a
desire that the species should be named after his brother, who lost
his life in Africa. I have therefore much pleasure in proposing the
name Gazella walleri for the new form.
The following table of the comparative dimensions of Gazella
walleri and Gazella dorcas, and the Plate (LVI.), will render further
description unnecessary.
930 PROF. A. H. GARROD ON INDICATOR MAJOR. [Nov. 19,
G. walleri 3. G. dorcas &.
English | | English |
| inches, | metre. | inches | metre.
Total length of skull...................2. 9:2 0-234 a5 0190 |
From anterior rim of orbit to free
extremity of the premaxille ...... 4:3 0-108 36 0-090
From anterior rim of orbit to the
occipital protuberance ............... 60 | 0152 40 0-101 |
From centre of the external audi-
tory meatus to occipital protu-
eYaNCe | vit. bccddeet size ss ostea tees. 2-4 0-061 10 0°025
From the centre of external auditory
imeatus to posterior rim of orbit...) 1:9 0-046 16 0-041
Length of the premaxille ..... ...... 21 | 0:053 2:2 0-055
Length of basicranial axis ............ 37 0-093 2:8 0-071
From inferior rim of orbit to alyeolus| |
of third upper molar ............... 9- | 0:022 11 | 0-028
Greatest depth of the rami of the
lower jaw in the portion under-|
lying the molars ...............0000+ 6 0-015 8 0-020
Length of diastema of the lower jaw) 2-1 0-053 1-4 0-035
Length of upper molar series ......... 1°15 | 0:029 1-4 0035
99 9y_-~-Premolar series ...| 8 0-020 3 0-020
»» 5, lower molar series ......... 13 | 0032 15 0:038
1» 9) 3, premolar series ...| 7 0:018 8 0-020
Length of horns round the curve ...) 12:0 0°305 9-0 0°230
Their circumference................c000 | 44 0-111 38 0-096
3. Notes on the Anatomy of Indicator major. By A. H.
Garrop, M.A., F.R.S., Prosector to the Society.
[Received October 25, 1878.]
Through the kindness of Mr. R. B. Sharpe I have had the oppor-
tunity of examining a spirit-preserved specimen of Indicator major,
from Fantee, which enables me to lay before the Society some fresh
facts in its anatomy confirmatory of its non-Cuculine affinities.
In his contributions to Orr’s edition of Cuvier’s ‘Animal Kingdom’
(1840), the late Mr. E. Blyth referred the Honey-guides to the Wood-
peckers as their nearest allies’; and this idea was expanded by him two
years later in the Journal of the Asiatic Society of Bengal®. In‘ The
Ibis’ for 1870 *, Mr. Sclater brought forward fresh facts in proof of
the non-Cuculine affinities of the genus, and proposed to place it next
to the Capitonidee, with which, or with the Coliide, Mr. Blanford
also in the same year showed it had its nearest relationships‘.
The following observations tend to prove the correctness of the con-
clusions arrived at by the two last-mentioned ornithologists, and
the error of imagining that Indicator is related to the Cuculide.
1 Loe. cit. p. 215. * Vol. xi. p. 167, 1842. Sp. des
* Observations on the Geology and Zoology of Abyssinia, 1870, p. 308.
—
1878. | PROF. A. H. GARROD ON INDICATOR MAJOR. 931
Pterylosis.—This has been recorded by Nitzsch in his ‘ Pterylo-
graphy ;’ and it seems more than strange that the characteristically
Picine distribution of its feather-tracts did not lead that able orni-
thologist to recognize its true relationships. He retained it among
the Cuculide.
My study of Pterylography has led me to look upon the nature
of the dorsal tract as all-important in determining to which great
group of Birds, the Homalogonate or Anomalogonatz', any
doubtful family belongs. When the dorsal tract develops a fork
between the shoulder-blades a bird is homalogonatous ; when the tract
runs on unenlarged to near the lower ends of the scapulz, then it is
anomalogonatous. Again, among the Anomelogonatz, when the
pectoral tract bifurcates into an outer and an inner branch just after
commencing on the chest, then the bird is one of the Piciformes,
and has a tufted oil-gland; when the pectoral tract does not bi-
furcate at all, or only at the lower end of its pectoral portion, but is
only increased in breadth instead, then the bird is Passeriform *,
and has a naked oil-gland. Exceptions to these rules scarcely
exist.
In that the dorsal tract of Indicator does not form a fork, but
remains narrow, between the shoulder-blades, it is anomalogonatous
—the Cuculide being homalogonatous ; in that it possesses a free
outer pectoral band to the anterior tract, it is a Piciform bird. The
oil-gland is also tufted.
A careful examination of the skin proves that the genus agrees
more closely in its feather-tracts with the Picide, Capitonide, and
Ramphastidz than Nitzsch’s figures would tend to prove. To me it
is evident that there is a communication between the thoracic ex-
tremity of the inferior neck-tract and the upper extremity of the
humeral tract, as in the Picide, Capitonide, and Ramphastide.
Again, in the three groups just named there is a great weakness or
an entire disappearance for a short distance of the dorsal tract
towards the lower extremity of the interscapular region. In Indi-
cator this same weakness exists; so that, with the exception of a
single feather in the middle line, the appearance of the region in
question is much more like the arrangement in Ramphastos than in
Nitzsch’s figure of the genus. On the ventral surface of the fleshy
tail I find a mediau subcircular space surrounded by feathers, as in
the Picide and Ramphastide, as well as in the Capitonide appa-
rently. The caudal termination of the dorsal tract agrees with the
account given by Nitzsch. I could find no trace of a duplication
of the lumbar tract. In the possession of twelve rectrices Indicator
differs from the Capitonidee and Ramphastide, which have ten,
like the Picidzx.
Skeleton—The specimen of the skull of Indicator examined by
Professor Huxley * at Mr. Sclater’s request was too imperfect for
exact description. The considerable interval between the maxillo-
palatines is recorded by him. I am now able to add that the vomer
1 Vide P. Z. 8. 1874, p. 116, for definition of these terms.
2 Vide P. ZS. 1874, p. 119. * Ibis, 1870, p. 179.
932 PROF. A. H. GARROD ON INDICATOR MAJOR. [Noyv. 19,
is but little different from that of the Capitonide. The palate of
Megalema asiatica is described and figured by Professor Parker in
the ‘ Transactions of the Linnean Society’*. It is truncated in front
and strongly bifid, the cornua running forward to blend with the
maxillo-palatines. These last-named inward-directed processes of
the maxillary bones blend with the mid-nasal septum in some
Fig. 1.
Palate of Indicator.
specimens of Megalema asiatica, whilst in others they remain free
from one another, separated by an inconsiderable interval. In
Pogonorhynchus bidentatus and Tetragonops remphastinus they
completely blend across the middle line, without the nasal median
septum persisting in front of the junction. So these two last-
named species, and most probably all the species of the genera,
are genuinely desmognathous.
The point in which the truncated vomer of the Capitonide most
differs from that of the order Passeres, is that in the former the
truncation occurs behind the line joining the posterior angles of the
maxillo-palatines, whilst in the latter the truncation occurs some
way in front of the same transverse line. The limbs of the forked
vomer in the Capitonide run forward to meet the posterior angles
of the maxillo-palatines; in the Passeres they continue, often in
cartilage alone, to the nasal labyrinth.
' 2nd series, Zoology, vol. i. p. 122, and pl. 23.
1878. ] PROF, A. H. GARROD ON INDICATOR MAJOR. 933
In Indicator the vomer is less ossified than in the Capitonidee
above mentioned, and is smaller ; the fork is slenderer and has longer
limbs, which, however, quite typically join the well separated
maxillo-palatines (which advance but slightly beyond the inner
margins of the palatine bones) at their posterior angles.
As the Ramphastidz have to be mentioned so frequently in con-
nexion with the genus under consideration, it may be useful to
refer to the vomer of this family. By Prof. Huxley it is included
among his Coccygomorphee, in which the vomer, if present, is pointed
anteriorly. Prof. Huxley farther remarks * that in Ramphastos “ the
antero-internal angles of the palatines not only meet, but are united
by bone.” But close examination demonstrates a large tabular
truncation of the anterior extremity in the Ramphastine vomer,
which I cannot help thinking Prof. Huxley interpreted as a median
osseous bridge formed by the (supposed) blending of the antero-in-
ternal angles of the palatines. Figure 2 represents the vomer of
Fig. 2,
Palatal aspect of the truncated vomer of Ramphastos ariel, with the posterior
parts of the palatine bones retained in union with it.
Ramphastos ariel, freed from its surroundings. It does not send
forward limbs to join the maxillo-palatines [which are those of the
desmognathous Capitonide inflated], but helps by its terminal ex-
pansion to complete, by contact or ankylosis with the palatine on
either side, the posterior wall of a cavity in the dried skull, bounded
laterally and superiorly by the inflated and infused maxillo-palatines,
anteriorly by the nasal septum together with ossifications in the nasal
cartilages associated with it.
In the ‘ Transactions’ of the Linnean Society* Prof. Parker de-
scribes the vomer of Ramphastos toco as double, it being composed
of a smaller posterior and a larger anterior bone, the truncated
nature of which I am not able to infer from the account given.
The Capitonide, the Ramphastide, and Indicator are intimately
associated, therefore, so far as the vomer is concerned. Nevertheless
the proportionally great length of the limbs of the vomerine fork
in the last-named form, and the considerable separation of its small
? Pp. Z.8. 1867, p. 444. 2 Zinn, Trans. 2nd ser, vol. i. p. 127.
Proc. Zoou. Soc.—1878, No. LXI. 61
934 PROF. A. H. GARROD ON INDICATOR Masor. [Nov. 19,
maxillo-palatines, are characters which tend to bring it nearer than
either of the others to the Picide.
The pterygoid bones of Indicator are much flattened from above
downwards, with thin outer and inner margins, which are curved, a
triangular groove on the palatal surface running from end to end.
In the Capitonidee and Ramphastide these bones are much more
cylindroid, the superior surface alone being thin-edged, whilst in the
Picidze they are thin, as in Indicator, but differ in possessing a
large anteriorly directed process springing from the superior surface
of each.
In Indicator there is a small notch in the middle of the superior
margin of the osseous orbit, no trace of which exists in any of the
other birds above referred to. In its external osseous nares, also,
there is no tendency towards the Ramphastine position of those
orifices, such as is so well marked in T'eéragonops ; the alinasal ossi-
fication that tends to divide each of the nares into an anterior and
a posterior moiety is likewise far less considerable than in Megalema.
As is known, and well illustrated in Mr. Sclater’s figure of the
bone (Ibis, 1870, p. 178), the sternum agrees most closely with that
of the Capitonidee and Ramphastidee ; and this is especially the case
in the imperfect development of the posterior extremity of the me-
dian xiphoid process, which in the Picidz continues further onward
to reach the level of the ends of the lateral xiphoid processes, at
the same time that the manubrial rostrum of the last-named family
only of the group is bifid. As to the posterior sternal notches, the
inner is the deeper ; and the same is the case in Gecinus viridis, whilst
in Picus, the Capitonide, and Ramphastide the outer is the deeper.
In its soft parts Indicator agrees with the Capitonidee, Ramphas-
tide, and Picide, and differs from the Cuculide, in the following
particulars :—There is only one caroti¢ artery, the left ; the ambiens
and the accessory femoro-caudal muscles are absent (the latter of
these is wanting in the Tree-Cuckoos) ; there are no colic ceca,
The femoro-caudal, semitendinosus and accessory semitendinosus are
present, as is the large gluteus. The tensor patagii brevis muscle
of the wing is inserted into the extensor metacarpi radialis longus
exactly as in the Capitonide, Ramphastide, and Picide, and as in no
other birds'. As in these three groups also (and in the Galbulide,
but not in the scansorial Cuculide and Psittaci), the deep plantar
tendons are distributed peculiarly—the flexor profundus digitorum
supplying the third digit only, whilst the flexor longus hallucis
sends slips to digits r. 11. and rv., as well as a vinculum to its com-
panion muscle *. The trachea at its lower end (fig. 3, p. 935) conso-
lidates into a bony box, formed by the fusion of the lowermost rings.
To the enlarged uppermost bronchial half-ring (a a) the single slender
intrinsic muscle of each side fans out to be attached at its middle.
Summing up the results of the above analysis, it may be stated
that, among the Piciform birds, pterylosis, osteology, myology, and
visceral anatomy place the Picide, Indicator, the Capitonide, and
1 Vide P. Z. 8. 1876, p. 508, pl. xlviii. fig. 1.
2 Vide P.Z. 8. 1875, p. 346.
1878. ]} PROF. A. H. GARROD ON INDICATOR MAJOR. 935
the Ramphastide in one great group of subordinal importance (if
the peculiar hammer skull of the Picidze be omitted from considera-
tion). In this suborder the Picidze constitute one main division—
a family ; whilst I, for one, cannot Separate off the Capitonide from
the Ramphastide by any well marked differences, the two sub-
families graduating into one another.
Syrinx of Indicator major (enlarged); anterior aspect.
Indicator must, in my opinion, also be placed in this family, from
all members of which it differs in possessing an extra pair of rectrices.
No one, however, objects to keeping all the Momotidx together
because of a similar difference in some of its genera ; why remove
Indicator therefore from its allies? Nevertheless Indicator is not
exactly like a Capitonine bird in certain details, so may be placed
as a subfamily by itself, the Indicatorine ; and the whole series may
be thus tabulated.
Order. Suborder. Family. Subfamily.
Pe Indicatorine.
ae ot Picidz. Ne
Piciformes, Pici. { aera Oapitonine.
Capitonide, Ramphasting,
6L*
936 MARQUIS OF TWEEDDALE ON THE [Nov. 19,
4. Contributions to the Ornithology of the Philippines.—
No. XI. On the Collection made by Mr. A. H. Everett
at Zamboanga, in the island of Mindanao. By
Artrnur, Marquis of Twerppate, F.R.S., President of
the Society.
[Received October 26, 1878.]
(Plates LVII.-LIX.)
The Spanish settlement of Zamboanga, situated at the southern
extremity of the long south-western limb of the large Philippine
island of Mindanao, is classical ground to the ornithologist. More
than a century ago Sonnerat collected birds there ; and in the year
1839 D’Urville’s second Expedition remained a couple of months at
Zamboanga. Yet only 19 species in all were recorded from Minda-
nao when in 1875 I published my List of Philippine Birds. Since
that date Dr. Steere and the naturalists of the ‘ Challenger’ Expe-
dition have collected in the vicinity of Zamboanga, and added 40
species, thereby increasing the total of known species to 59.
Mr. Everett arrived at Zamboanga last March, and remained
through April and part of May. During his stay he obtained
examples of 98 species of birds. Mr. Everett writes that these were
all procured “ within a radius of ten miles of Zamboanga, chiefly in
the hilly country some five miles distant at the back of the town.”
Of these 98 species only 33 had been previously known to inhabit
this part of Mindanao; so that 65 species have been added by
Mr. Everett, of which 11 are new to the Philippines, the following
6 being also new to science—
Pseudoptynx gurney, Chetura picina,
Ninox spilocephalus, Lyncornis mindanensis,
Scops everetti, Folvocivora mindanensis,—
the remaining 5 having been previously unknown in the Phi-
lippines—
Accipiter stevensoni, Cacomantis sepulchralis,
Ninox lugubris, Ptilopus melanocephalus.
Coccystes coromandus,
Examples of 6 species previously said to have a Philippine habitat
Mr. Everett found at Zamboanga, thus reducing the number of un-
certain Philippine species to 29.
Twenty species authentically recorded from Zamboanga and its
vicinity are not contained in Mr. Everett’s collection; so that the
now known total of Zamboanga species identified from specimens
amounts to 118.
Mr. Everett writes :—‘‘ I enclose a note of the species observed by
me here, which I think were not represented by skins in my col-
“> ¢
Smit lith.
| ah
Hanhart imp
AC CIPITER.STEVENS ONI
|
RH
JSmit lith
PoRUDOP TYNX
Pee .2. 1878 Pl Lin
Hanhart imp
GURNEYI.
aAnnary tmz
ICINA.
CHA TURA P
1878. ] ORNITHOLOGY OF THE PHILIPPINES. 937
lection. I have only included in these lists those birds the names
of which I am absolutely certain I can give with accuracy.”
Cacatua hematuropygia. {New to Zamboanga. |
Haliastur intermedius. {Already recorded. ]
Merops bicolor. {Already recorded. }
Entomobia gularis. {Already recorded. |
Sauropatis chloris. {Already recorded. |
Artamus leucorhynchus. {Already recorded. |
. Anthus, sp.? [?]
Carpophaya, sp.? [Probably Ptilocolpa griseipectus, and new
to Zamboanga. |
Turtur dussumieri. [New to Zamboanga. |
Charadrius fulvus. Throat and belly jet-black. [Already re-
corded. |
Tringoides hypoleucus. | Already recorded. ]
Sterna, sp. Bill chrome; crown of head black ; wing delicate
brownish-grey ; underparts snowy white. Rather large. [Probably
Sterna bergii, and already recorded. }
Three of the above species (omitting the Anthus as undetermin-
able) not having previously been recorded from Zamboanga, when
added to the number of species represented by specimens, raise the
total number of birds known from this part-of Mindanao to 121.
1. CAcaTUA HEMATUROPYGIA (1).
| Zamboanga. |
T include this species on Mr. Everett’s authority.
2. PRIONITURUS DISCURUs (2).
[Zamboanga, 2 3, April.]
3. TANYGNATHUS LUZONENSIS (3).
[Zamboanga, 2, May. |
4. CYCLOPSITTA LUNULATA (4).
[Zamboanga, ¢ 2, March, April, and May. ]
Examples with lunated collars are marked female by Mr. Everett,
those with blue collars as being males.
5. LoricuLus HARTLAUBI (7).
[Zamboanga, ¢ 9, April and May.]
6. HyporriorCHIs SEVERUS (9).
[Zamboanga, g, April: iris dark brown; bill black; cere and
orbital skin greenish yellow ; legs bright yellow ; claws black.]
In the plumage described by Mr. Sharpe (Cat. Accipitres, p. 397)
as belonging to the young bird.
7. MicrowieRax ERYTHROGENYS (10).
[Zamboanga, ¢ 9, April.]
938 MARQUIS OF TWEEDDALE ON THE [Nov. 19,
8. ACCIPITER STEVENSONI. (Plate LVII.)
Accipiter stevensoni, Gurney, Ibis, 1863, p. 447, t. xi.
[Zamboanga. a. 2, April: iris golden; bill black; legs greenish
ochre. 4. 9, April: iris golden ; orbital skin greenish-yellow ; bill
leaden, apex and cere blackish ; legs greenish-yellow, but pale brown
on the toes; claws dark brown. c. ¢, pullus, April: iris yellow;
bill black; legs light green; sole of foot yellow; claws black.
d. (sex ?), pullus, April : iris yellow ; cere greenish; beak and claws
black ; legs light greenish; sole of foot yellow. |
Dimensions :-—
Wing. Tail. Tarsus. Middle toe.
in. in. in. in.
@: Qa tise nice ROO 5°75 1°87 1:16
‘es Sods Sa oe tae 6°75 3°79 1°75 1°25
e. 6 juv.( 92). 5°75 4:00 1°81 125
Aire JAVe-— 21, . An Oeee 4°00 1°62 1:06
Example a is slaty brown above, darkest on the head. Rectrices
pale greyish slate-colour, with five dark-brown bands on middle
pair. The under surface is white; the pectoral, abdominal, and
flank-plumes being traversed with two, three, or more pale-brown
bands. The pure white throat has a dark brown and extremely
narrow median line formed by the white feathers having a very fine
dark edging to the shafts. The quills underneath are regularly
banded with white and brown, above brown banded with dark
brown. The innermost tertiaries are at most pure white with a pale
brown margin. Axillaries pure white with a series of narrow brown
cross bars.
Example 6 (which is figured in the accompanying plate) has the
breast and abdomea uniform pure bright rufous. When the feathers
are discomposed they appear white banded with bright rufous.
Vent and under tail-coverts are pure white. Thigh-coyerts white,
with faint pale rufous cross bands. Middle of throat white, with
terminal pale earthy marks. Sides of head dingy pale rufous-brown.
Upper plumage brown, mixed with rufous in some of the feathers.
Middle pair of rectrices with six transverse brown bands. Quills
barred as in example a, pale rufous occupying the interspaces be-
tween the brown bands to a great extent. Avxillaries white, banded
with pale rufous.
Examples ¢ and d are of nestlings in rufous dress. Underneath
creamy-fulvous,with broad ruddy brown stripes. A central gular stripe
of brown. Middle pair of rectrices with three brown bands. Upper
surface of body and head dark brown, with rusty margins and bases
to the feathers. Thigh-coverts spotted with light ruddy brown.
Mr. Gurney observes on these specimens as follows :—The hawk
“‘ marked a on the ticket is a female of Accipiter stevensoni in the
ordinary adult dress ; that marked 4 I believe to be the same ; but
I have never before seen a female with so rufous a breast, though it
much resembles the figure of the male type specimen (now somewhat
faded) in the ‘ Ibis’ (/. s. c.). The two nestlings may be either A. ste-
1878. ] ORNITHOLOGY OF THE PHILIPPINES. 939
vensoni or A. virgatus. We have a very similar one from Java; but
as both species are found there, I am not sure to which it belongs,
but probably to 4. virgatus, as I suspect that 4. stevensoni is only
a winter visitor to Java. Judging from the size of the tarsus and
foot, I think the larger nestling, marked c, is a female though marked
male, and that the smaller, marked d, is a male.”
Having expressed to Mr. Gurney some doubts about the correct-
ness of identifying example 4 with the female of 4. stevensoni, that
gentleman kindly favoured me with the following remarks :—
“Your rufous-breasted hawk from the Philippines, which I
marked 6, is certainly nearer to A. stevensoni than to any other
species that I am acquainted with; but since returning it to you
I have felt some doubt as to whether it will not prove to belong to a
. distinct and undescribed species.
“Your bird differs in having the wing shorter than any of our
females, and larger than any of our males. Our three males have
but four transverse bars on the tail. Of our four females three have
five bars, and the fourth (an immature bird from Java) four, like the
males, whereas your specimen 6 has six, as mentioned in your letter.
“None of our females have any rufous on the breast ; and in all
our males it is hardly more than a tinge of buff. Our type speci-
men (a male, figured in the ‘Ibis’ for 1863, pl. 11) is perhaps
somewhat faded ; but our Curator is confident that it was never so
rufous as is represented in the ‘Ibis’ plate ; and in this I believe he
is right.
“Your bird is more like the male of 4. stevensoni than it is to
the female ; but as your collector has marked it 2, it probably is so.
‘Specimens of A. stevensoni are scarce, and we hardly know the
limits of variation to which it is liable ; and on this account I should
be glad if additional specimens like your skin 4 could be obtained,
before venturing to publish it as a species distinct from A. stevensont ;
but at the same time, if it should ultimately prove distinct, it would
by no means surprise me.
*‘ | ought toadd that as 4. stevensoni has not yet been met with
when paired, we have no positive proof that the bird which I suppose
to be its female (the type being a male) is so, though I do not
myself entertain any doubt of such being the case.”’
9. SPILORNIS HOLOsPILUS (16).
[Zamboanga, ¢ 9, April and May. |
10. ELaANus HYPOLEUCUs (18). ?
[Zamboanga, ¢ juv., May: iris light yellow-brown; bill black ;
cere greenish yellow; feet pale chrome; claws black. ]}
11. Nrnox sPILOCEPHALA, sp. n.
(Zamboanga, ¢, March: iris golden; bill greenish leaden ; feet
pale yellow. 9, April: iris golden; bill-greenish lead, tinged
yellow on culmen and tip; feet wax-yellow. 2, April: iris light
greenish-yellow. |
940 MARQUIS OF TWEEDDALE ON THE [Nov. 19;
Were it not that the large series of examples sent by Mr. Everett
(six males and thirteen females) all agree in having spotted heads,
I should not have ventured to separate the Zamboanga species from
N. philippinensis of Luzon. Every variety of markings and colora-
tions is exhibited in this series, from dark brown to tawny-rufous
brown above, and stripes and spots of the same colours below ; but
all the nineteen examples have the frontal and coronal feathers
brown, spotted with rufous—in some bright rufous, in others pale
tawny rufous. Some have the whole under plumage, from throat to
vent, dark-centred; others have the breast almost uniform rich
rufous, without pale margins, and with a subterminal brown trans-
verse narrow band.
Every specimen has its sex noted on its label; and the length of
the wing in the six birds marked ¢ is greater than im those marked
Q. The wing of the six males ranges in length from 6°50 to 7:0,
of the thirteen females from 6°25 to 6°50. Two Luzon males have
the wings 6°37 and 5:25. Mr. Sharpe (Cat. ii. p. 168) states 2
inches as the length of the tarsus of an example of NV. philippinensis
in the British Museum: none of these Zamboanga birds has the
tarsus longer than 1°12.
12. ‘Nrnox LUGUBRIS.
Striv lugubris, Tickell, J. A. S. B. ii. p. 573.
[Zamboanga. a. 2, March: iris golden; bill blackish ; cere, cul-
men, and mandible greenish’ yellow; feet dark chrome-yellow ;
claws black. 6. 2, April: ims deep brown; bill greenish; feet
chrome-yellow. |
Dimensions :—
Wing. Tarsus. Culmen. ‘Tail.
in. in. in. in,
Ctr eae 1°12 0°62 4°75
hoe 2 Oe 1:12 imperfect. 4°82
Six caudal bands. Virst two primaries without bars. The wings of
a ruddy hue. The outer webs of most of the primaries rusty-red.
The remarks Mr. Gurney has been good enough to favour me
with about this Zamboanga race of Ninow lugubris are as follows :-—
“I think the outer webs of the primaries unusually rufous. We
have only one similar specimen, which is from Formosa and is
described in Sharpe’s volume (Catalogue, ii. p. 161); and with
this specimen the Zamboanga bird seems to me to agree gene-
rally. If N. japonica be admitted as distinct from N. lugubris
(though I doubt whether it ought to be), the Zamboanga specimen,
I think, ought to be referred to NV. japonica.”
Mr. Sharpe has suggested (¢. c. p. 166) that the large Japanese
and North-China form is migratory. These Zamboanga examples
favour his hypothesis.
13. PsreupopTyNx GURNEYI, sp. n. (Plate LVIII.)
[Zamboanga, 3, pairing, April : iris warm brown; bill greyish
white; feet pale grey ; claws white tipped with dark grey.]
1878. | ORNITHOLOGY OF THE PHILIPPINES. 941
Sides of face bright pure unmarked rufous. Lengthened stiff
plumes springing from base of maxilla rufous mixed with tawny,
and many with dark-brown shafts and tipped with dark brown.
Space above front of eyes, uniting on forehead and extending back
over the eyes, pale tawny rufous. Crown and occiput dark rufous,
many of the feathers with a dark brown broad mesial stripe. Nape
and sides of neck pure rufous. Lengthened ear-tufts the same,
some with very narrow brown mesial linear markings near their
apices. Plumes bordering the facial disk albescent tawny ; some
almost pure white, tipped with dark brown. Chin and upper throat
pale tawny rufous. Middle of throat white. Breast and remainder
of lower surface pure rufous, more dilute on lengthened tibial
plumes and under tail-coverts. A few pectoral plumes, with
dark-brown large terminal drops. Many abdominal plumes, with
dark-brown elongated central stripes. Back rufous, minutely
freckled with brown, each feather with a bold, irregular, dark-brown
central stripe. Scapulars like the back, but some of the shorter
and outer albescent tawny on outer webs. The dark brown central
marks are so arranged that the back, together with the scapulars,
appears to have three parallel dark-brown stripes running down it.
Uropygium and upper tail-coverts rufous-brown, with darker shafts,
Rectrices brown, minutely freckled with pale rufous, and with eight or
nine narrow pale rufous cross bands. Minor and median wing-coverts
brown, freckled with rufous, and each with a dark narrow central
brown line. Major coverts brown on inner web, freckled with
rufous on outer. Quills brown, alternately banded with freckled
brown and pale rufous. Tertiaries pure rufous, with traces of dark
brown along the shafts. Carpal edge white; wing-lining yellowish
white ; some of the under carpal coverts rufous, Thigh and tarsal
coverts pale rufous and tawny white.
Mr. J. H. Gurney writes te me :— “The Pseudoptynx is cer-
tainly distinct from P. philippinensis, and, so far as I know, is un-
described. Besides its very much smaller size, it is very much more
rufous below, and of a much darker rufous above; and the dark
central marks of the mantle are altogether of a different character,
and very much less coarse than those in P. philippinensis. As to the
difference of size, we have two specimens of P. philippinensis here ;
and the comparison between their dimensions and those of the
Zamboanga bird is as follows :-—
Culmen
Wing. Tarsus. Middle without
toe cere,
in. In. in In.
“P. philippinensis, No.1.. 15°54 1°6 1-4
4; a> Niozs28 : imperfect 2°5 1.5 1°5
See WUIHCYE A... ms cess 9°3 1'8 1-4 Lis?
The dimensions of the type in the British Museum given by Mr,
Sharpe (Cat. ii. p. 43) also much exceed those of the Zamboanga
bird.
I name this fine Owl after Mr. Gurney, to whom for many years
942 MARQUIS OF TWEEDDALE ON THE [Nov. 19,
I have been largely indebted for great assistance in determining the
obscurer species of Rapaces.
14. Scops EVERETTI, sp. n.
{ Zamboanga, 2, April: iris warm brown; bill pale greenish leaden,
nearly white; feet whitish grey ; claws dark grey. ¢, April: iris
deep brown; feet whitish. |
Mr. Everett has sent three examples of a species of Scops, which
may be described as being a large form of Scops lempiji as it
occurs in Java. Above it has the deep rich brown colouring of the
Javan species; the same broad dark-brown mark in the form of
a parallelogram on the head and nape; the same pale-coloured
forehead, stripe above each eye and round the nape, and the broad
dark brown stripe leading from behind the eye, and including most
of the ear-tufts. In the Zamboanga species (as represented in Mr.
Everett’s series) there are no pale uniform tawny or fulvous sca-
pulars. Underneath, the markings differ from those of the Javan
bird by being more confused, and by the absence of regular dark-brown
cross markings on the abdominal plumes. The general colouring
of the under surface is of a ruddier brown,
Dimensions :—
Wing. Tarsus. Tail. Culmen.
in. in. in. in.
GC eaeene Gho0 1-20 3°30 0°62
Oo Osho 1°20 3°62 0°68
Mr. Sharpe, to whom I submitted two of the specimens, has
obliged me with the following observations :—‘‘ Your Owl is of the
S. lempii group, having that peculiar light streak on the sides of
the crown, running onto the ear-tufts ; but underneath it is quite
aberrant, and verges more towards the Moluccan S. leucospilus sec-
tion. At the same time I expect it will come very near Scops
umbratilis, Swinhoe.””
Mr. Gurney writes, “‘ The Owl is new to me, and different to any
that we have here.”
15. THRIPONAX JAVENSIs (28).
[Zamboanga. ¢, March: iris yellow. ]
The white at the tips of the primaries is unusually developed in
the specimens from Zamboanga.
16. CarysocoLaPTEs Lucipus (32).
Chrysocolaptes maculiceps, Sharpe, Tr. L. 8. ser. 2, Zool. i.
p. 314, t. xlvi. fig. 2.
[Zamboanga, 2 , May : iris crimson. |
An example (marked a female) wears the plumage of C. macu-
liceps, as described and figured by Mr. Sharpe.
1878. ] ORNITHOLOGY OF THE PHILIPPINES. 943
17. YUNGIPICUS VALIDIROSTRIS (33 partim).
Picus nanus, Vigors, Blyth, J. A. 8. B. 1845, p. 197.
Picus validirostris, Blyth, Cat. Cale. Mus. p. 64. no. 305;
J. A. 8. B. 1849, p. 805.
[Zamboanga, ¢ 2, May: iris crimson?; bill black ; mandible
lead-grey ; feet olive. |
When writing on Picus maculatus, Scopoli (Tr. Z. 8. ix. p. 148),
I stated that the titles I then brought together were treated as
synonyms on the assumption that the islands of Luzon, Panay, and
Mindanao possessed but one and the same species of Yungipicus.
I had had no opportunity of examining an example from any one of
the Philippine Islands. Since then Mr. Everett has sent me ex-
amples of a species of the genus from Luzon ; and these I identified
(P. Z.S. 1877, p. 689) with P. maculatus, rather than create a new
title, while their dimensions were too small for P. validirostris,
Blyth. The birds from Zamboanga differ specifically from the
Luzon species. They are larger; the uropygium and upper tail-
coverts are unspotted tawny-white ; and the rectrices are tawny buff
banded with dark brown, and not dark brown, for the most part,
as in the Luzon bird, with narrrow albescent bands or marks. In
both, the lower throat and upper breast are spotted, and uot streaked,
as in Y. fuscoalbidus of the Sunda Islands and Malacca. Until
typical examples of P. maculatus from Panay are compared, it cannot
be affirmed whether the type of P. maculatus belongs to the Luzon
or the Mindanao species, or whether it may not be a species distinct
from either. In the meantime I adopt Blyth’s title, the dimensions
he gives being exactly those of the Zamboanga species :—bill to
forehead 0°75 ; wing 3:25.
I may mention that Y. aurantiiventris, Salvadori, is an excellent
species, quite distinct from Y. fuscoalbidus.
18. HaRPACTES ARDENS (34).
[Zamboanga, 2, March: coloration of soft parts as in male. |
The coloration of the soft parts is not mentioned on the labels of
the males sent.
19. ALCEDO BENGALENSIS (38).
[Zamboanga, 2, March. |
20. PELARGOPSIS GIGANTEA.
Pelargopsis gigantea, Walden, Ann. & Mag. Nat. Hist. ser. 4,
xiii. p. 123.
{[Zamboanga, ¢ 9, March and May. |
21. Creyx ARGENTATA.
Ceyx argentata, Tweeddale, anted, p. 108. no. 7, t. vi.
[Zamboanga, 3, May.]
22. Enromosia GuLaris (44).
{ Zamboanga. |
944 MARQUIS OF TWEEDDALE ON THE [Nov. 19,
23. HALCYON WINCHELLI.
Haleyon winchelli, Sharpe, Tr. L. 8. (2) Zool. i. p. 318, t. xvii.
(Zamboanga, 9, April: iris dark brown; bill black; the basal
half of mandible white ; feet light greyish-green. |
I have not been able to compare the Zamboanga birds with the
Basilau type; but the female agrees well with Mr. Sharpe’s descrip-
tion and plate. A second example (sex unrecorded) differs in
having the whole under surface pure white.
24. CALIALCYON COROMANDA (46).
[ Zamboanga, 2, March. |
The discovery of this species by Mr. Everett at Zamboanga re-
moves all doubts of its being an inhabitant of the Philippines.
25. ACTENOIDES HOMBRONI (48).
[Zamboanga, $, April: iris brown; bill orange-red ; the culmen
blackish ; feet dirty greenish orange. |
In the perfectly adult male the crescentic pectoral markings dis-
appear.
26. XANTHOLEMA H2&MACEPHALA (50).
[| Zamboanga, ¢, March and April. |
27. MacropTeryx CoMATUus (52).
[Zamboanga, ¢, April.]
28. CH&TURA PICINA, sp.n. (Plate LIX.)
[Zamboanga, 9, April. ]
Black with a blue gloss, greenish in some lights. Chin, throat,
and larger under wing-coverts pure white. Wing 6°37; tail 1°25.
29. LyNCORNIS MINDANENSIS, Sp. 0.
[Zamboanga, ¢, April: iris warm brown; feet purplish brown. ]
Typical Z. macrotis (ex Luzon) has the crown, forehead, ear-
tufts, and nape rufous brown, of almost the same shade as in
L. temminckii. The three examples of the species of Lyncornis
which inhabits the vicinity of Zamboanga, sent by Mr. Everett,
differ from Z. macrotis by having the crown, nape, forehead, and
ear-tufts dark greyish-brown, and not rufous, by all the browns of
the plumage being much darker, and by the wing being shorter.
Wing 9°90.
30. CAPRIMULGUS MANILLENSIS (55).
[Zamboanga, 2, April.]
Compared by Mr. Sharpe with the type.
31. BATRACHOSTOMUS SEPTIMUs.
[Batrachostomus septimus, Tweeddale, P. Z. S. 1877, p. 542,
no. 13.
[Zamboanga, ¢, April: iris orange-yellow; bill light horn-
1878. } ORNITHOLOGY OF THE PHILIPPINES. 945
brown ; feet pale ochreous yellow ; nails grey. Interior of gape,
3 2, lemon-yellow. |
Two examples of the adult male and one of an adult female are
sent by Mr. Everett. The white nuchal band of the female is not
sO conspicuous as in the type, nor are the white markings of the
throat-band and pectoral plumes. The general hue of the male-
plumage agrees with that of the female; but in the males the
elongated scapulars have the outer webs pure white, with a black
subterminal spot on the inner web. Four examples of this species
are now known, two of males and two of females; and all four are in
the rufous phase. Mr. Everett, in his remarks about one of the
males is very explicit ; he writes :—‘ I have one Batrachostomus, a
good skin ; it is in rufous plumage, andis ¢. I took especial care
in the sexing; and luckily the parts had not been touched by shot.
Judging from the size of the testes, I think the bird is rather im-
mature.”
32. CACOMANTIS SEPULCHRALIS ?
Cuculus sepulchralis, 8S. Miller, Land- en Volkenk. p. 177.
[Zamboanga, ¢ 9, April. ]
Mr. Everett obtained five examples of a Plaintive Cuckoo, two of
which are in adult plumage, and so closely resemble the Javan
C. sepulchralis, that I do not venture to separate them. The dimen-
sions of the wings and tail are slightly less, while the chin and
throat are of the same rufous as the breast, and not grey. S. Miller
described (/. c.) the chin of the Javan and Sumatran type as being
grey ; and so it is in my Javan specimens ; but it is not certain that
in Javan birds this grey does not change into rufous. An example
obtained by Mr. Everett at San Mateo, Luzon (C. merulinus, a,
P. Z.8. 1877, p. 691. n. 19), belongs, without doubt, to C. sepul-
chralis, and has the chin and upper throat grey, the others being
examples of true C. merulinus. This last species can hardly be kept
separate from the grey-breasted species of Continental Asia, C. thre-
nodes, &c.
33. SURNICULUS VELUTINUS.
Surniculus velutinus, Sharpe, Tr. L. S. ser. 2, Zool. i. p. 320.
[Zamboanga, 2, April. }
34, CHALCOCOCCYX MALAYANUS.
Cuculus malayanus, Raffles, Tr. L. 8. xiii. p. 286, “ Malay Pe-
ninsula’’ (March 1821).
? Cuculus basalis, Horsfield, Tr. L. 8. xiii. p. 179, “ Java”
(April 1820).
[Zamboanga. a.d, April: iris burnt-sienna colour ; orbital ring
fine vermilion; bill blackish; feet very dark lead-grey. 6. April:
iris light yellowish brown; bill black; legs lead-grey; nails
black. |
Mr. Everett sends three Golden Cuckoos, which, although some-
what smaller than typical examples, differ in no other respect.
946 MARQUIS OF TWEEDDALE ON THE [Nov. 19,
Wing 3°62. I adopt the more recent specific title of malayanus, as
it has not as yet been absolutely demonstrated that the Malayan-
Peninsula bird is the same as the Javan. The only Javan example
I have seen is pure white underneath, without any transverse bands.
Above, the whole back is very dark green without any coppery gloss,
the head and nape coppery greenish-brown. This individual is
fairly represented on plate 102. f. 2, of the ‘ Planches Colori¢es,’
under the title of Cuculus chalcites, Iliger, exact habitat unknown.
It may be only a phase of plumage; but Horsfield and Moore
(Cat. Mus. E. I.C. ii. pp. 706, 707) keep the Malaccan species
distinct from the Javan.
35. H1erococcyx PECTORALIS (60).
[Zamboanga, ¢, April: iris pale yellow; orbital skin bright
chrome-yellow ; bill black’; mandible and base of maxilla greenish ;
feet bright wax-yellow ; claws horn-yellow. |
A single example, which agrees in every detail with the descrip-
tion given by Dr. Cabanis. It may also belong, as suggested by me
(J. ¢.), to Cuculus hyperythrus, Gould ; but it differs from the figure
of that species in wanting the black chin, in not being so intensely
rufous underneath, in the rufous colouring not extending below the
breast, and in the upper plumage not being of so dark a shade of
grey. It also has four, distinct, transverse, narrow, dark caudal bands,
and not merely two as described by Mr. Gould.
36. CoccysTES COROMANDUS.
Cuculus coromandus, Linn. 8. N. i. p. 171. no. 20.
[Zamboanga, 2, March: iris brown; bill jet-black ; feet bluish
grey ; interior of gape salmon-red. |
; New to the Philippines, and a most unexpected addition to their
auna.
37. EUDYNAMIS MINDANENSIS (61).
[Zamboanga, 2, March: iris bright crimson; bill and legs
greenish plumbeous. 4, ¢, March.]
These are typical examples of the species, and do not differ from
North-Mindanao examples. They are somewhat smaller than birds
from Guimaras and Zebu. Maximum length of wing 7°50. In
the female the spots and bands appear in this species to be always
rufous.
38. CeNTROCOCCYX vIRIDIS (64).
[ Zamboanga, March. |
39. PyrRHOCENTOR MELANOPS (65).
[Zamboanga, ¢ 9, March.]
Sexes alike.
4(). Bucrros MINDANENSIS.
Buceros mindanensis, Tweeddale, P. Z. 8.1871, p. 543.
[Zamboanga, ¢ 2 2 juv., March. ]
1578. | ORNITHOLOGY OF THE PHILIPPINES. 947
41. PEnrLopipeEs AFFINIs.
Penelopides affnis, Tweeddale, P. Z. 8. 1877, p. 824. n. 29. f. 1.
[Zamboanga, ¢ Q, March. ]
A new onter rectrix has the central band pure white, the central
bands on all the other rectrices being bright rufous.
42. Lanius nasutus (70).
[Zamboanga, ¢ 9, April.]
Mr. Everett sends a nestling killed in April.
43. Lanius Luctonensts (72).
[Zamboanga, ¢, April.]
44. Gravcatus srriatus (73).
[Zamboanga, ¢, April: iris pale yellow. |
Two examples marked ¢ have uniform plumbeous breasts and
banded underparts and uropygium. One marked 9 is banded
from the chin downwards as well as on the uropygium,
45. VoLvocivora MINDANENSIS, sp. n.
[Zamboanga. a. 3, April: iris dark brown ; bill and legs black.
6. 2, April: iris dark brown. e. 2, April: iris dark chocolate-
brown. |
Adult male (a). Chin, cheeks, sides of head, space before the eyes,
a broad supercilium, ear-coverts, forehead, throat, and upper breast
jet-black. Remainder of breast, crown, and whole under surface of
body dark leaden-grey, paler near the vent and on the under tail-
coverts. Wing-coverts, nape, and axillaries a slightly lighter shade
of leaden-grey, palest on the uropygium and upper tail-coverts,
which are slightly mixed with white. Quills above black, with
faint grey margins. Sixth and remaining primaries tipped with greyish
white. Secondaries and tertiaries leaden-grey on their outer webs,
Part of inner webs of quills, seen from underneath, pale grey. This
colour commences at the base of the first primary, occupying but a
small part of it, but is more extended on each succeeding quill, and
reaches to the extremities of the inner quills. Rectrices jet-black ;
outer pair with a large grey terminal mark, the same colour being
slightly indicated on the next pair. Carpal edge and wing-lining
almost black.
Adult female (6). The same as the male, but the grey colour of
the whole plumage of a paler shade, the uropygium more albescent,
and no black about the head, throat, and breast. Carpal edge and
axillaries pale grey.
Young female (¢). Like the adult, but the grey colour not so
pure. The chin, central part of the throat, a few scattered feathers
on the breast and flanks, the wing-lining, and axillaries finely banded
with white. Upper tail-coverts nearly pure white. Outer margins
of some of the secondaries and carpal edge inclining to white.
948 MARQUIS OF TWEEDDALE ON THE [ Nov. 19,
Dimensions :—
Wing. Tail. ‘Tarsus. Culmen.
in. in. in. in.
Giendiyire) 44562 4:0 0°88 0°88
Oa Seek ALO 4:0 0°88 0-88
Q juve... 4:48 3°88 0:88 0°87
A representative form of VY’. morio ex Celebes, but altogether of a
darker shade of grey, with a black forehead, and with the inner
margins of the quills grey, and not pure white.
46. LaLaGe pomrinica (76).
[Zamboanga, 9, April. }
47. DICRURUS STRIATUS.
Dicrurus striatus, Tweeddale, P. Z. S. 1877, p. 545. no. 20.
[Zamboanga, 2, Marchi: iris crimson. |
48. LEUCOCERCA NIGRITORQUIS (83).
[Zamboanga, 9, April. ]
49. CyoRNIs PHILIPPINENSIS (84 partim).
Cyornis philippinensis, Sharpe, Tr. L. S. ser. 2, Zool. 1. p. 325.
[Zamboanga, 2, March. ]
50. HyporHymMis AzUREA (85).
(Zamboanga, 9, May: maxilla black, mandible light blue; legs
dark grey slate ; interior of gape yellow. ]
51. Broprripus ACRORHYNCHUS (90).
[ Zamboanga. |
52. ERyYTHROPITTA ERYTHROGASTRA (94).
[Zamboanga, ¢ 2, March, April, May. |
These examples all belong to ZL. erythrogastra, and not to the
Balabae race named B. propinguus by Mr. Sharpe (Tr. L. 8. (2)
Zool. i. p. 330). The young Dumalon (Mindanao) example ob-
tained by Dr. Steere, and doubtfully identified with B. propinquus
by Mr. Sharpe (J. c.), in all probability belongs to 2. erythrogastra.
An immature bird, killed in April near Zamboanga, is dirty —
rufous brown above, with dashes of green; rectrices blue; wing-
feathers without a trace of blue; under surface dirty white,
feathers margined with pale rufous brown ; some scarlet feathers on
the breast ; vent and under tail-coverts washed with brownish red.
The females differ in the scarlet colouring of the abdominal region
not being so intense and vivid, and in the throat not being so dark
brown, almost black. The blue colouring of the breast is not
exhibited until complete maturity, that part remaining green. In
fully mature birds of both sexes the green pectoral band is almost
entirely replaced by blue.
1878. ] ORNITHOLOGY OF THE PHILIPPINES, 949
53. MELANOPITTA SORDIDA (95).
[Zamboanga, 3, March : iris chocolate ; bill black ; legs brownish
grey. o 9, March, April, and May.]
This species has the black central abdominal patch much more
largely developed than in M. muelleri ex Borneo. In no other
respect does it appear to differ. The extent of pure white on the
quills is dependent on sex. Of fourteen individuals with the sex noted
by Mr. Everett, nine are males and five are females. The primaries
of the male have a much greater white surface than is found in the
primaries of the female. The 6th, 7th, and 8th primaries are
scarcely tipped with black ; and the first five primaries have double
the amount of white that exists on the primaries of the female. On
examining Luzon and Negros examples with the sexes noted by
Dr. B. Meyer, the same character differentiates the sexes. In
Bornean individuals of M. muelleri (sex determined by Mr. Everett)
the sexes can be discriminated by the amount of white on the
remiges. In M. cucullata some of my examples exhibit a maximum
of white on the quills and the others a minimum ; but as none
have had their sexes determined by dissection, I am unable to
affirm that in that species the same law holds good. In two
Zamboanga skins the green of the breast and flanks is assuming the
blue tint so prominent in the large representative form M. steerii,
54. MELANOPITTA STEERII.
Brachyurus steerii, Sharpe, ‘ Nature,’ August 1876, Tr. L. S,
ser. 2, Zool. i. p. 329, t, xlix.
[Zamboanga, 3, March and April. ]
55. MACRONUS STRIATICEPS.
Macronus striaticeps, Sharpe, Tr. L. S. ser. 2, Zool. i. p. 331.
{[Zamboanga, ¢, March: iris white: bill black; legs purplish
grey. ]
56. Ixus Gotavier (99).
[Zamboanga, ¢ 2, March. }
57. PoLioLoruus uRostictus (101).
[Zamboanga, ¢, April: orbital ring light yellow.]
58. HypsipeTes RUFIGULARIS.
Hypsipetes rufigularis, Sharpe, Tr. L. S. ser. 2, Zool. i. p. 335.
(Zamboanga, 2, April; iris crimson; bill blackish brown ; legs
brown. |
59. Monticoxa souitarivs (103).
[Zamboanga, 9, April. ]
60. CopsycHus MINDANENSsIs (106).
[Zamboanga, ¢, March. |
Proc. Zoo. Soc.— 1878, No. LXII. 62
-
950 - MARQUIS OF TWEEDDALE ON THE [Nov. 19,
61. PHYLLOSCOPUS BOREALIS.
Phyllopneuste borealis, Blasius, Naumannia, 1858, p. 313.
[Zamboanga, ¢ 9, April.]
62. ORTHOTOMUS CINEREICEPS ?
Orthotomus cinereiceps, Sharpe, Ibis, 1877, p. 113, t. 11. f. 2,
* Basilau.”
[Zamboanga, ¢, March: iris clay-orange; 2, April. }
Mr. Sharpe’s title is here adopted with doubt for this Zambo-
anga species of Tailor-bird, because it does not quite agree with the
description and figure, and I have not been able to compare it with
the type (now in America).
In the example marked as being of a male, the cheeks, chin, and
entire throat are jet-black. In that of the female the chin and
upper part of the throat are white, the middle of the breast,
the abdomen, and vent are silky white, whereas in the male these
parts are bright olive-green. ‘The rectrices of the male are devoid
of a subterminal bar; but in those of the female brown subterminal
bars are conspicuous. It is not improbable that the male example
with entire throat black is O. einereiceps in full breeding-dress.
63. ORTHOTOMUS FRONTALIS.
Orthotomus frontalis, Sharpe, Ibis, 1877, p. 112, t.ii. f. 1, “Zam-
boanga.”
(Zamboanga, ¢ 2, March and April. |
The examples noted as belonging to males have no subterminal
bar on the rectrices. Those marked as being of females have the
subterminal bar well pronounced.
64. BupyrTes vrripis (114).
[Zamboanga, ¢, April. |
65. ZosTEROPS EVERETTI.
Zosterops everetti, Tweeddale, P.Z.S. 1877, p. 762. no. 42.
[Zamboanga, 5 9, April. ]
66. Dic#uM CINEREIGULARE.
Diceum cinereigulare, Tweeddale, P. Z.S. 1877, p. 829. no. 50,
“ N. Mindanao.”
(Zamboanga, Q, April: iris dark brown; bill black; legs very
dark grey, nearly black. }
The female differs from the male in wanting the orange-red inter-
scapular patch, in the abdomen, vent, and under tail-coverts be‘ug
pure yellow, and in the upper plumage being deeply tinged with
olive-green.
67. Dic£UM RUBRIVENTRE (120 partim).
Pipra papuensis, Gm. 8. N. i. p. 1004.
(Zamboanga, 2, April: iris dark blood-red ; bill and legs black. ]
1878.] ORNITHOLOGY OF THE PHILIPPINES. 951
68. NECTAROPHILA JULIA.
Nectarophila julie, Tweeddale, P. Z.S. 1877, p. 547. no. 32,
‘* Melanipa.”’
Cinnyris julia (Tweeddale), Shelley, Cinnyride, pts. vii. & viii.
[Zamboanga, ¢ 9, April: iris brown; bill and feet black. }
In some examples of the male the breast is suffused with orange-
red ; in others little or no red is perceptible.
69. Cyrrosromus sJUGULARIS (123).
[Zamboanga, ¢ 9, March and April.}
70. ANTHOTHREPTUS CHLOROGASTER.
Anthreptes chlorigaster, Sharpe, Tr. L. S. ser. 2, Zool.i. p. 342 ;
Shelley, Cinnyride, pt. vi. pt. vil. t.
[Zamboanga, Q, April: iris indian-red ; bill light umber-brown ;
legs greenish leaden. ]
71. Raaspornis MystacaLis (124).
{Zamboanga, ¢, April. ]
A single example, which differs from the Luzon species in its
much shorter bill. Culmen barely 0°62, in the Luzon bird fully 0°75.
- The wing is also shorter ; and the colouring of the wings and back
is warm brown, and not greyish olive-brown.
72. Corvus PHILIPPINUS (125).
[Zamboanga, ¢, March. ]
73. CALORNIS PANAYENSIS (128).
{[Zamboanga, ¢, March. |
74. SaARcops CALvus (129).
{Zamboanga, 2, March. |
7d. Munia sacori (132),
{[Zamboanga, ¢ 2, March.]
76. OSMOTRERON VERNANS (135).
{Zamboanga, ¢ 2, March, April, May. |
77. PTILOPUS MELANOCEPHALUS.
Columba melanocephala, Forster, Zoologia Indica, p. 16. no. 7,
t. viil., ex Java (1781).
[ Zamboanga. a. 3, March: iris, orbital skin, and bill yellow, with
more or less of a green tinge; feet carmine, nails dark grey. 6.¢,
April: iris yellow; bill yellow-green. ]
Hardly separable from the Javan species. Wing 4°50. The
tint of the throat-patch resembles that of P. xanthorrhous ; but the
abdomen and crissum are yellow, and not orange. New to the
Philippines.
62
952 MARQUIS OF TWEEDDALE ON THE [Nov. 19,
78, RAMPHICULUS OCCIPITALIS (138).
[Zamboanga, ? (juv.), March: iris dark brown; bill light brown,
the basal half dull red ; feet coral-red. |
Sexes alike.
79. PHABOTRERON BREVIROSTRIS.
Phabotreron brevirostris, Tweeddale, P. Z. S. 1877, p. 549.
no. 38.
{Zamboanga, ¢ 2, March and April.]
The pale, and not rufous, chin and throat distinguishes this
species from P. leucotis.
80. CARPOPHAGA £NEA (141).
[Zamboanga, 2, March. |
81. PriLocoLPA GRISEIPECTUS (142)?
Mr. Everett remarks in epist.:—‘‘ The Carpophaga of the Zam-
boanga list is a bird markedly smaller than C. enea; and those
parts which in the latter are coloured fine bronzy green, are in the
smaller species dark iron-grey.” The above-given description, so
far as it goes, agrees with Pézlocolpa griseipectus, a rare species in
museums, and the exact habitat of which in the Philippines remains
to this day (unless this identification is correct) undetermined.
82. MyristiIcrvora BICOLOR (143).
[Zamboanga, 2, May. ]
83. HemipHaGa POLIOCEPHALA (144).
(Zamboanga, @ (not quite adult): bill black; orbital skin
crimson. |
Sexes alike. The margin of the inner web of the first primary is
- scooped out, and that of the second a little less so.
84. MacropyGIA EURYCERCA.
Macropygia eurycerca, Tweeddale, anted, p. 288, no. 49.
[Zamboanga, ¢ 9, March and April. |
Considerably smaller than the Negros type, but otherwise undis-
tinguishable. Wing 6°75, as against 7°50; tail 7-00, against 8-00.
85. TuRTUR DUSSUMIERI (147).
{ Zamboanga. |
Included on the authority of Mr. Everett.
86. PHLOG@NAS CRINIGERA.
Phlogeenas crinigera (Pucheran), Tweeddale, P. Z. S. 1877,
p. 833. no. 66.
[Zamboanga, ¢ 2, March and April. |
Sexes alike. An example marked ‘“¢ juv.” by Mr. Everett
has the claret-coloured pectoral plastron only indicated by a few red
1878. ] ORNITHOLOGY OF THE PHILIPPINES. 953
plumes, and the terminal margin of the major coverts fulvous instead
of pale grey, the remainder of the plumage being marked and
coloured as in the adult.
_ 87. Cuatcornaps 1npica (150).
[Zamboanga, ¢ Q, March. |
88. Gaius Banxrva (153).
[Zamboanga, ¢ 9, March.]
89. Mrcapopius pILLWwyNl.
Megapodius dillwyni, Tweeddale, P. Z. S. 1877, p. 766.
[Zamboanga, 9, March: iris burnt sienna-brown; bill horn-
brown; orbital skin dirty crimson ; legs reddish-brown ; tarsi! dark
brown. 6, March: orbital skin dark purplish brown. |
90. GALLINULA CHLOROPUs (169).
(Zamboanga, 9, May: iris light warm brown; bill greenish
yellow; frontal plate orange-red ; legs grass-green. }
91. GaLticrEex CINEREA (170).
[Zamboanga, ¢, March and April.]
92. ERYTHRA PH@NICURA (171).
[Zamboanga, 2, March. ]
93. ORTYGOMETRA CINEREA (172).
[Zamboanga, ¢, April. |
94. Porzana Fusca (174).
[Zamboanga, ¢, April: iris brick-red ; legs dull wax-red.]
95. GALLINAGO MEGALA (188).
[Zamboanga, 9, March. ]
96. Ruyncw#a CaPENsis (189).
[Zamboanga, ¢, May.]
97. ARDETTA CINNAMOMEA (192).
- [Zamboanga, 2, March. |
98. HrropiaAs INTERMEDIA (196).
[Zamboanga, 3, March. ]
99. MELANOPELARGUS EPISCOPUS.
Ardea episcopus, Bodd. Tabl. d. Pl. En. p. 54.
[ Zamboanga, 2, March. ]
In full dress.
* So written on label; perhaps a slip of the pen for the word “toes” or
claws.”
954 MR. E.R. ALSTON ON NEOTROPICAL SQUIRRELS. [Nov. 19,
100. DenpRocyGNna vaGAns (203),
[Zamboanga, ?, April.]
101. Dysporvus suta (214).
[Zamboanga, 9, May: iris white; bill dirty white; orbital and
gular space pale green; feet very light greenish yellow ; nails grey. |
A young female in dirty-brown plumage, darkest on breast, throat,
neck, head, and back.
5, Supplementary Note on the Neotropical Squirrels.
By Epwarp R. Atston, F.L.S., F.Z.S., &e.
[Received November 8, 1878.]
I regret to say that several errors have crept into my paper “On
the Squirrels of the Neotropical Region,” ! owing in parttomy absence
from town when the sheets were passing through the press. Two
or three of these appear sufficiently serious to require correction.
Scrurus varrasiuis. In the remarks on this species 8. rufo-
niger is a clerical error for S. brunneo-niger, at p. 666, seventh line,
and at p. 667, second line.
Scrurus pusittus. At p. 670 I remark that I was not able to
ascertain Geoffroy’s type in the Paris Museum. It appears that I
had overlooked it ; for Professor A. Milne-Edwards has kindly in-
formed me that it is still there. Unfortunately the skull is wanting,
so that its age cannot well be ascertained. The want, however, is sup-
plied by the British-Museum specimens, of which the skulls and teeth
show them to be fully adult, although an unhappy misprint (p. 671,
eighteenth line) makes me state the exact contrary. M. Milne-
Edwards has sent me the following measurements of the types of
S, pusillus and 8. rufo-niger :—
S. pusillus.
millim. inches.
Length of head ........ 0°039 1°56
RS head and body 0°089 3°56
> tail essence (imperfect)
S. rufo-niger.
millim. inches,
Length of head........ 0°048 1°92
e head and body 0:178 “12
“ tail (with hair) 0°139 5°56
This last specimen appears to me to have been considerably over-
stretched in mounting.
In conclusion I may add that I have had the pleasure of receiving
a letter from Mr. J. A. Allen, in which he accepts almost all-the
determinations at which I arrived in the paper in question.
* Supra, pp. 656-670,
PZ: 3.16 76P ie
=
m ’
3 Ey AE AA : 4
gu: [LA ZOELHCIEL, LZ, A a
op EF 2-325 PR ‘
ME: ea Ro
C2 == Ch = 2 |
A - | 7 =
I =a — =
! [L. aN é a 3 ;
ESD ONe -_
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RO del J Smit ith.
CHAMBERED SHELLS OF CEPHALOPODS.
1878.] PROF. OWEN ON THE SHELLS OF CEPHALOPODS. 955
6. On the Relative Positions to their Constructors of the
Chambered Shells of Cephalopods. By Prof. Owen,C.B.,
F.R.S., F.Z.S., &e.
[Received October 4, 1878.]
(Plate LX.)
The relations of the chambered and siphonated shells to their con-
structors can now, owing to the extent to which they have been
subject to extinction, only be fully elucidated by a study of species
or varieties of two of the genera, Nautilus and Spirula.
The fossil shells of Cephalopods, as is well known, exhibit a pro-
gressive uncoiling from Nautilus to Orthoceras and from Ammonites
to Baculites, with various modifications of the process ; but that this
had been carried out to a coiling in the reverse direction, required
anatomical evidence for its demonstration.
The insight, however, gained into the organization of Spirula
peronit at the date of the publication of the ‘ Zoology of the Voyage.
of the Samarang’!, added to that previously obtained on the organi-
zation of Nautilus pompilius®, led me to express a conviction of their
shell-relations in the following terms :— “ These shells (Nautilus and
Ammonites) are revolutely spiral or coiled over the back of the
animal, not involute like the Spirula”*, And, if the direction of the
coils be determined by their relation to the back and belly of the
framer of the shell, no other interpretation can be given of such rela-
tion as it is exhibited in Spirula (Plate LX. fig. 4) and Nautilus
(ib. fig. 3) respectively.
It is, however, to the exposition and characters of the extraordinary
number and manifold variety of the extinct Cephalopods, now known
only by their polythalamous and siphonated shells, that an exact
and accepted determination of their relative position to the body of
the framer is most needed.
In the year 1829 Leorotp von Bucu initiated, in his notable
memoirs on the Ammonites‘, the definition and nomenclature of the
shell-characters by which the species, genera, and families might be
defined.
Assuming, and correctly in my judgment, that the shells of an
Ammonite and a Nautilus were coiled in the same direction, he
premises :—“ Le caractére distinctif entre ces deux genres de Cépha-
lopodes, consiste en ce que le syphon des Ammonites est toujours
dorsal, et qui'il ne Vest jamais dans les Nautiles”®. Next, calling
attention to the lobed and foliaceous sutures of the Ammonitic shells,
he defines the parts which he calls “lobes” and “selles” (saddles )—
specifying of the former, ‘‘Ze lobe dorsal,” “le lobe ventral,’ and “les
lobes latéraux”’*. Von Buch’s descriptions and figures admit of no
1-“ Mollusca,” Part 1, 4to, p- 6, pl. iv.
? Memoir on the Pearly Nautilus, &c. 4to, 1832.
$ Palxontology, 8vo, 1861, p. 97.
+ Annales des Sciences Naturelles, 8yo, tome xvii. p. 267, tome xyiii. p: 417.
* Tb. p. 268. 8 Tb. ib.
956 PROF. OWEN ON THE [Nov. 19,
doubt as to hisconclusion that the outer convex curve was the ‘‘dorsal ”’
one, the inner concave curve the “ ventral’ one, in both Nautilus
and Ammonites.
Such, indeed, seemed the obvious relations to back and belly of
the Nautilus-shell before the structure and position of its framer
were made known. And it should be remembered that my conelu-
sions on the latter point were inferential, a mere fragment only of
the shell having been left attached to the unique specimen submitted
to my scalpel in 1832.
Accordingly, in 1835, M. pe BLainviLte’, adopting Von Buch’s
view of the ‘‘ dorsal’ position of the siphuncle in Ammonites, and con-
ceiving the shell of Spiru/a to be convoluted in the same direction as
in Nautilus, characterizes its siphuncle as ‘‘ ventral,’ and that ofthe
Ammonite as “ dorsal.”
In his applications of these views of relative position to the Ammo-
nites, with little change of the families into which these fossils had
been arranged by Von Buch, he proposes new names for them, and
adopts the shape and proportions of the outer curve or border of the
shell as the family characters. Thus, when such outer, and in my
view ventral, curve of the shell is broad, asin Ammonites dilatatus,
a family of ‘‘ A. latidorses”’ is diagnosed; a reverse proportion, as
shown in Ammonites discus, characterizes the family ‘“ 4. compressi-
dorses ;’ but neither these families, nor those of the ‘ cavidorses,”’
“cristidorses,” ‘ lzvidorses,” &c, have gained currency or accept-
ance.
ALcipE p’OrpiGNny’ adcpts the view of relative position, the ter-
minology, and in the main the classification proposed by Von Buch.
By Morris* and Anstep‘ the aspects of the Nautiloid and
Ammonitoid shells propounded by Von Buch are retained. Prof.
Ansted associates Spirula with Nautilus in his family NauTILacea,
and writes :—‘ The next point of difference to be attended to is in the
siphuncle; and it is one both of position and magnitude. In the
genera of the first family, Nauriiacea, this important part is some-
times ventral, or on the inner margin, more frequently central, and
is very rarely observed to approach the dorsal or outer margin. On
_ the other hand, it is almost always very near the dorsal margin in
the Ammoneara, and sometimes is actually placed outside, in a
channel opened for it, and projecting from the back of the shell in
the shape of a keel ’’*.
In fact, to have propounded that the siphuncle in the Ammonitidze
was ventral, as in Spirula, would have implied that the shells were
coiled in reverse directions—an assumption seemingly held to be too
? Prodrome d’une Monographie des Ammonites, 8vo, 1840.
? Paléontologie Francaise, 8vo, 1842, p. 185:—* Les lettres suivantes, les
mémes que celles qu’emploie M. de Buch, indiquent toujours les mémes parties
dans les figures—e. g. D. lobe dorsal, V. lobe ventral,” &c.
8 “On some new Species of the Genus Ancyloceras” (“The ribs ornamented with
two conical tubercules on the dorsal part”), Ann. & Mag. of Nat. History, 1845,
vol. xv. p. 82, pl. vi. figs. 3, a, d.
* Observations on the Animals inhabiting multilocular Shells, 8yo.
5 Ib. p. 278,
1878. ] SHELLS OF CEPHALOPODS. 957
extravagant for adoption. And yet, if the Ammonite was tetra-
oe like the Nautilus, no other conclusion could logically be
rawn.
Accordingly, such view is formally repudiated by the experienced
zoologist Dr. J. E. Gray, in his paper “On the Animal of the
Spirula”!, “The examination,’’ he writes, ‘“‘of this animal confirms
me in the opinion which I expressed in the ‘Synopsis of the British
Museum ’ (1840, p. 149), that the Ammonites, from their texture
and the small size of the last chamber, are internal shells and should
be arranged with the decapodous Cephalopods, being chiefly dis-
tinguished from the Spirula by the siphon being always on the dorsal
margin of the whorls and the septa foliated on the edge. I am
aware,”’ he adds, ‘that this opinion is not in conformity with the ideas
of many zoologists and comparative anatomists ; for Mr. Owen, in the
last arrangement of these animals (Todd’s Encyl. Comp. Anat.), though
he places the Spirule with the Dibranchiate Cephalopods, places the
Ammonites with the Tetrabranchiate next to Nautilus” (p. 259).
Before recapitulating the grounds on which it was inferred that the
Ammonitide were external shells, I will, finally, cite the valuable
Memoirs of the Geological Survey of India, published under the
direction of the accomplished and lamented Superintendent of the
Survey, THomas Otpuam, LL.D., F.R.S., in which the sanction
given to the views of Gray and before-cited authors as to the aspects
of the shells of the Ammonites has mainly induced me to offer the
present elucidation of the grounds on which I still hold the contrary
opinion, viz., that the siphuncle in the Ammonitide is ‘‘ ventral ”’ or
‘** margino-ventral,”’ as it is in Spirula, but thatit is “‘ external” in the
one and “internal ”’ in the other, through the reversed direction of the
spiral whorls.
In the “ Monograph of the Cretaceous Cephalopoda of Southern
India,” the preparation of which was confided to the accomplished
naturalist F. SroriczKa, too early lost to science, he premises the
following characters of the Ammonitide :—
“Animal not known: shell spiral, more or less involute with
numerous regularly (?) and gradually increasing whorls in the same
plane, many-chambered, the last or body-chamber extending generally
over about two thirds of the last whorl. The margins of the septa
are deeply divided into lobes and saddles, the first having their sub-
divisions always pointed, the latter more rounded. The dorsal lobe
is divided by a small saddle into two parts, corresponding to the
siphunele: in the regular forms of Ammonites this is always placed
in the middle of the back; the siphuncle is also similarly placed
inside the shell.”
It is to be observed, however, that the author associates the
Ammonitide in the same order ‘“‘TETRABRANCHIATA”? with the
Nautilide ; and it would seem therefore that he entertained the
doubts originally expressed by Gray* and supported by Grant® as
1 Annals and Magazine of Natural History, vol. xv. 1845.
? Phil. Trans. 1833, p. 774. ‘ ;
3 My description and figures were called in question, with more detail, by
958 PROF. OWEN ON THE [Nov. 19,
to the correctness of the position assigned by me to the soft parts
of Nautilus in the shell’; but the sole reason assigned by Stoliczka
for adhering to Von Buch’s view of the position of the shell and his
consequent denomination of its parts is the following:—‘* Our de-
scriptions are of the shell (Am. ¢esta), not of the animal which was
living in it, and which we know not’’’.
We shall never know the organization of that animal ew visu.
But there are, in respect of the Ammonite as of most extinct ani-
mals, other sources of knowledge in kind and degree sufficient to
decide such a question as the relations of soft parts to shell in a
Molluscous species.
With regard to those relations in Nautilus I may merely cite the
subjoined works on its anatomy confirmatory in the main of my
own, and fully so in regard to the relative position of the soft parts
to the shell’.
the accomplished Professor of Comparative Anatomy in University College,
Dr. Rosert E. Grant, F.R.S., and begat a general belief, in 1833, that I had
placed the animal of the Pearly Nautilus in a position the reverse of the real
one, in the ‘Memoir’ of 1832. In the report of the Lecture on the Shells of
the Cephalopoda, given in the ‘ Lancet, vol. i. 1833, with the sanction and
revision of the Professor, Dr. Grant remarks :—
“The exact position of the animal in the recent Nawtili, Spirule, and other
polythalamous spiral shells, although important in the interpretation of the
fossil forms, has not yet been satisfactorily observed by naturalists. In our
present uncertainty, therefore, regarding the position of the living Cephalopods
in these convoluted shells, we can only be guided by analogy.”
That which guided Dr. Grant to his conclusion he illustrated by figures of
sections of the external shell of Nautilus and of the internal shell of Sepia (p.
506), and explained them as follows. Comparing the layers of the ‘“cuttle-
bone” to the septa of the Nautilus shell, he states :—‘‘ These layers begin in the
Sepia by forming a small hollow shell, which receives into its interior another
hollow shell; this receives within it a third and a fourth hollow shell; and so
they go on in the first stages of its growth. If these cones continued thus to
extend outwards, and with an oblique direction, turning on the same vertical
plane, they would have formed a convoluted suborbicular shell, like that of the
Ammonite or the Nautilus. Where, then, in this so convoluted shell of the
Sepia, would have been the convex outer border of the shell, or the upper lip,
with relation to the body of the Sepia? The Sepia would haye looked for-
wards over the spire from the last-formed chamber, keeping the exterior convex
part of the chamber” (answering to B in fig. 3) “or the upper lip extending across
its back, as in all other known orbicular shells of Pteropodous and Gasteropo-
dous Mollusca.” (P. 509.)
1 Memoir, ut supra, pp. 12, 44, pl. i.
* Op. cit. p. 44. So, likewise, affirms a later writer :—‘‘ We have only the
shells preserved to us. We know nothing of the animals.”—H. Woodward,
Geol. Mag. vol. v. p. 497.
As well might it be affirmed of Anoplotherium, e. g.:—‘It must always be
borne in mind, when arguing from these early fossil remains, that we have only
the skeleton preseryed to us. We know nothing of the soft parts.”
3 Vavencrennes, M. A. “Nouvelles Recherches sur le Nautile flambé
(Nautilus pompilius, Lam.),” Archives du Muséum d'Histoire Naturelle, Paris,
4to, tome 11., 1841, p. 275, pls. viii., ix., x.
Van ver Horyrn, J. “Contributions to the Knowledge of the Animal of
Nautilus pompilius,” Trans. Zool. Soc. vol. iv. p. 21, pls. 5, 6, 7, 8.
Vroxik, W. “ Lettre sur quelques points de lorganisation de V’animal du
Nautile flambé,” Mémoires de la Soc. Linnéenne de Normandie, tom, x., 4to,
1855, pp. 1-18, pls. i. and ii. 3
1878. | SHELLS OF CEPHALOPODS. 959
A colleague of Stoliczka, accepting the recorded structure and
relative position of Nautilus, frames his nomenclature of the parts
and aspects of the fossil shells of the Nautilide accordingly', the
grounds for such being those of positive or direct evidence from
dissection. I am led, therefore, to submit remarks on the guiding
knowledge in regard to the structure and shell-relations of the extinct
Ammonites, which may be gained from negative or inductive testi-
mony.
J. E. Gray, we have seen, deemed that testimony to be such in
kind and amount as to warrant his rejection of the affinity of Ammo-
nites to Nautilus, and his adoption of the association of Ammonites
with Spirula in the Dibranchiate order of Cephalopoda.
Examples of unmutilated Ammonite shells are, indeed, very rare ;
seldom is a specimen acquired with any considerable proportion of
the last chamber. Yet some such were to be seen in an accessible
museum in London long anterior to 1845.
A bisected specimen of the Ammonites odtusus, Sow., in the
Hunterian collection (No. 188), shows well the extent of the last or
inhabited chamber of the shell, and the effects of the influence of
the animal matter of the decaying Cephalopod upon the petrifactive
processes after death. The liassic clay has penetrated as far as the
retracted soft parts of the Ammonite permitted; the decomposing
mollusk has been partially replaced by crystals of spar discoloured
by the pigmental or carbonized parts of the animal; the spar which
has more slowly infiltrated through the pores of the shell into the
air-chambers is of a much lighter colour.
In the same collection may be seen exemplifications of injury and
repair of the cell. In No. 195, Ammonites goliathus, D’Orb., from
the Oxford Clay, a portion of the shell, at the period when it formed
the dwelling-chamber, ‘‘had been broken away during the lifetime
of the animal, and has been repaired by fresh material, wanting the
ribbed structure of the originally formed shell”’ *.
The reparation closely resembles that which recent Nautilus-shells
occasionally present, and which we know was effected by the forma-
tive border of the mantle reflected over the last chamber and applied
to the fractured part. No such process could take place in Spiruda,
the mantle of which is muscular, and inapplicable to the last
chamber. There is no need of a living Ammonite to assure us that
its mantle, like its porcellano-nacreons shell, had the same structure
as that of the living Nautilus.
1 Memoirs of the Geological Survey of India, 4to, 1866. “The Fossil
Cephalopoda of the Cretaceous Rocks of Southern India: Belemnitide—Nauti-
lide,” by Heyry F, Buanrorp. ‘‘ Throughout the following descriptions I
have employed the terms ventral and dorsal strictly with reference to the posi-
tion of the animal, and therefore in an opposite sense to that in which they
were used by palxontologists before the anatomy of the animal was known,”
p. 7. (And Mr. Blanford might have added ‘‘ long after.’—R. O.)
2 Catalogue of Fossil Invertebrata, Mus. College of Surgeons, London, 4to,
1856, p. 43, in which work I described upwards of 350 specimens, illustrative
of the different families and genera of Ammonitide, collected by Joun Hunrer
in the last century.
960 PROF. OWEN ON THE [ Nov. 19,
The perfect specimen (Plate LX. fig. 1) of the last chamber of the
Ammonite was derived from that eminently conservative matrix, the
lithographic slate of Solenhofen, Bavaria. The extent of the outer
curve of the dwelling-chamber to the outer border of its floor (the
last septum, a to y) is 4 inches; the extent of the outer curve of
the chambered part of the shell to the inner border of the cham-
ber-floor (last septum, y to z) is 6inches. In the Nautilus pom-
pilius transmitted in its perfect shell to Prof. Vrolik the corre~
sponding admeasurements are 7 inches 9 lines and 11 inches 6 lines.
The correspondence in the proportion of the dwelling-chamber to
the camerated part of the shell in the Ammonite and Nautilus is
thus, as in numerous other instances, shown to be instructively
close.
In the last chamber of Ammonites lingulatus, moreover, as if to
proclaim to the most sceptical its function of lodging its constructor,
is preserved the only fossilizable part of such (ib. fig. 1, 0).
Since the publication of the ‘ Paléontologie’ of Pictet’, who refers
the Trigonellites to the Cirripeds, abundant evidence has been ob-
tained of the accuracy of the opinion of Volz’, that the T'rigonellites of
Parkinson* (Aptychus of v. Meyer*) were parts of the animal of the
Ammonite, and stood in opercular relation to the shell of that extinct
Tetrabranchiate. Pictet takes no note of the confirmation contri-
buted by Morris to the Volzian examples of Trigonellites within the
last or dwelling-chamber of Ammonites, in the portion of an Ammonites
walcotti in which “the Aptychus, of a corneo-calcareous nature,
was found imbedded in the matrix filling the last chamber about
6 inches from the aperture”’’.
This common, though not constant, position led some palzeonto-
logists to surmise that the Trigonellites might be parts of the
Ammonites’ gizzard, like the triturating plates in Bulla hgnaria,
M. Valenciennes believed them to be lateral supports of the funnel °.
Van der Hoeven “hazards the opinion that the two juxtaposed fossil
shells, known by paleographs as Aptychus, were two shelly sup-
ports of the hood of Ammonites” (Trans. Zool. Soc. vol. iv. p. 22).
In this opinion I concur; but it has been rejected by some experi-
enced students of the extinct polythalamous shells. Keferstein and
Waagen’, e. g., deem the Trigonellites to be sexual characters, and
to have served as protective plates of the nidamental glands of the
female Ammonite. Waagen adduces, in support of this view, the
' Traité de Paléontologie, &e., vol. ii. p. 551, 8vo (1854).
2 «Sur le Belopeltis,’ in Mémoires de la Soc. d’Hist. Nat. de Strasbourg,
tome iii. (1836).
3 ‘Organic Remains of a Former World,’ 4to, 1811, p. 184, pl. xiii. figs, 9-12.
4 Nova Acta Acad. Nat. Cur. t. xv. pt, 2, p. 125, pl. lviii. (1831).
> “Note on Aptychus,’ in Ann. & Mag. Nat. Hist. vol. x. (2nd ser.), p. 356,
pl. v. D. fig. 1.
6 « Je crois qwil faut admettre que l’entonnoir de l’ammonite, s'il était formé
de deux valves, ne contenait pas de cartilage interne; mais que cette piéce était
remplacée par un organe extérieur composé de deux piéces paires symétriques
comme le sont les aptychus.”—Ib. p. 504.
7 “ Ueber die Ansatzstelle der Haftmuskeln beim Nautilus und Ammoniten,”
Paleontographia, 4to, p. 185, taf. xxxix., 1871.
1878. ] SHELLS OF CEPHALOPODS. 961
correspondence of the relative position of the Aptychus', in the body-
chamber, to that of the nidamental glands of Nautilus pompilius? to
the shell ; and he deems each valve of Aptychus to have been applied
to a lateral lobe of the nidamental gland*. But these lateral lobes
are divided from each other by a part of the middle lobe *, while the
valves of the Aptychus are usually in contact (as in Plate LX. fig. 1, 0),
or may be suturally united along the mid line (as in ib, fig. 2, 0).
But to return to Waagen’s argument from position in the fossil shell.
In the course of decomposition after death the calcareous plates would
be likely to gravitate or sink deeper into the body-chamber than
their natural position in the living Ammonite. So sinking, they
would rather lodge or settle in the hollow of the outer (ventral)
wall of the body-chamber (8) than upon the involute convexity on
the opposite (dorsal) wall (a). Moreover there are examples (as in
Plate LX. fig. 2) in which the Trigonellites (0, 0) have been found
in an opercular relation at the mouth of the shell. And, considering
the movements to which an Ammonite must have been subject from
the time of its death to the solution of the soft parts and final im-
bedding of the shell in the matrix or seat of its petrifaction, one is
prepared for the rarity of the conservation with the shell of its loose
operculum, and for the still more rare retention of the Aptychus in
its original position. Waagen admits that there are five Specimens
of Ammonite in the Munich collection exhibiting this position®, He,
however, contends that the breadth of the aperture of the body-
chamber is less, in certain Ammonites (A. steraspis, e. g.), than the
united breadth of the aptychal plates. But so, likewise, if the side
lobes of the hood of Nautilus® were outstretched horizontally, it
would exceed the breadth of the outlet of the dwelling-chamber in
N. pompilius. But the side lobes of the hood are bent back
obliquely in order to close the dorsal side-curved borders, or notches,
of that part of the shell-aperture, just as the aptychal lobes or
valves are bent down in Plate LX. fig. 2, 0, 0.
It may be further remarked, in respect to the nidamental glands,
that they are subject to seasonal changes, and gain the relative bulk
with which the size of the aptychal plates accord only at the period
of discharge of the impregnated ova, for which they have to furnish
the protective coat or nidus. Such seasonal change is exemplified
in the figure of these glands given in the ‘Memoir on the Nautilus’
of 1832, and in that which is shown in Taf. xix. of Waagen’s
Treatise, 1871. Moreover, in not one of the existing genera or
species of Cephalopod, Nawéilus included, in which these glands are
superadded to the more essential organs of generation, are they en-
cumbered in any way or degree with such calcareous plates as Kefer-
stein’s hypothesis applies to them in the Ammonite.
* Paleontographia, 1871, taf. xl. fig. 4.
? See Memoir on the Nautilus, 1882, pl. i, e.
* Ib. pl. viii. fig. 10, 2, d. * Tb. ib. a.
* “Unser Museum besitzt gegen 100 exemplare yon Ammoniten mit erhal-
tenen Aptychus; unter dieser ganzen Anzahl sind 5, welche den Aptychus in
senkrechter Stellung am Ende der Wohnkammer liegend haben” (op. cit, p. 192),
° Memoir on the Nautilus, 1832, pl, iii. fig. 1, 9, 9.
962 PROF. OWEN ON THE [Nov. 19,
In the application of the anatomy of the constructor of the Pearly
Nautilus to the solution of the problem of the nature and function
of the Trigonellites, [ was led to regard them as the homologue of the
organ, or a portion of the organ, in Nautilus, which is “ of a fibrous
texture, resembling dense corium,”’ called from its shape and posi-
tion the “hood” (Plate LX. fig. 3, e), and which, when the animal
had withdrawn into its dwelling, ‘‘ would serve as a rigid defence at
the outlet of the shell”’’. It needed only that this part should be
more or less calcified to form the preserved portions of an operculum
like that ascribed to the Ammonite. If, for example, calcification
had commenced in each half of the symmetrical ‘‘ hood,” and had
stopped at the mid line (where the hood is thinnest), the pair of the
there-often-suturally-joined symmetrical pieces of the Aptychus would
have resulted. The relative size of Aptychus agrees with that of the
shell. It has been found to measure 7 inches 6 lines in length and
6 inches in breadth in gigantic Ammonites’. It may be doubted
whether the nidamental glands ever increased in the same ratio ; and
it is still less likely that they needed such defensive plates in their
season of rest and attenuation.
If, therefore, my homology of the symmetrical halves of the
Nautilus’ hood with the parial Trigonellites (Aptychus, v. M.) be
preferably accepted, the supposition that these parts are calcifica-
tions of an Ammonite’s hood may be deemed reasonable. That the
fibrous basis of the hood was retained in different degrees in the
Ammonites is indicated by the simply corneous or chitinous condi-
tion of the Aptychus which has been preserved in some examples of
Ammonites falcifer, Sow., and its allies. In other species, as in
the Ammonites lingulatus (Plate LX. fig. 1, 0), the lateral calcifica-
tions have partially met and joined at the mid line; in a third series
these opercular plates are thicker and are there suturally united.
This is the case in the small or young specimen of Ammonites sub-
radiatus, Sow., in the British Museum, which is described and figured
by S. P. Woodward, F.G.S., in ‘The Geologist’*. As the view of
the specimen there given is an oblique side one, I here append a
direct view of the aperture of the dwelling-chamber as closed and
protected by its operculum (Plate LX. fig. 2, 0). The correspond-
ence of general shape with that of the “hood” of the Nautilus
pompilius, as figured in plate iil. fig. 1 of my ‘ Memoir,’ is close*.
The conjoined plates of the Aptychus (ib. fig. 2, 0, 0) form a tri-
angular disk, of which the base is backward, excavated to receive the
involute part of the shell, p, with the sides of the base, like the
corresponding lobes of the ‘“‘ hood,” bent down to cover the laterally
extended parts of the wider terminal coil of the shell, g. Even in
the contrast between the papillose wrinkled outer surface and the
1 Memoir on the Pearly Nautilus, 1832, p. 12, pl. i. 2, and pl. iii. fig. 1.
2 See de Zigno, Memorie del R. Istituto Veneto, tom. xv. tab. viii., 1870.
2 8yo, 1860, p. 328.
* Woodward gives the following description of its ammonitic homologue :—
“The operculum is flat in the middle, with a slight furrow along the suture,
and is much bent down at the hinder corners, where it abuts against the inner
whorl of the shell. It is sculptured externally with about twelve angular con-
centric furrows; the inner surface is smooth,”—ZB, 2b,
s
1878. ] SHELLS OF CEPHALOPODS. 963
smooth inner surface of the “hood”? of Nautilus the resemblance
to the “‘Aptychus”’ of the Ammonite is carried out. Anterior to
the apex of the triangular operculum in the Ammonites subradiatus
a small portion of the dwelling-chamber (ib. v) is left uncovered,
through which a slender stream of water might pass from the retracted
funnel.
As the soft parts of the Ammonites lingulatus decayed and were
dissolved the calcareous or opercular parts of the hood (ib. fig. 1, 0)
have subsided to near the bottom of the dwelling-chamber, probably
with such change of their original relative position, as exemplifies the
value of the demonstration given in the specimen described by
Woodward. If an independent centre of calcification were set up in
the mid region of the “hood” (Mem. cit. pl. ii. fig. 1, f), a part
corresponding to the dnaptychus might result.
In further ventilation of the mooted affinity of Ammonites to
Spirula, it may be remarked that in not one of the examples of
Ammonites in which the dwelling-chamber has been in any propor-
tion preserved has there been any trace of an ink-bag. Yet fossili-
zation of this or of its secretion is abundantly exemplified in the
extinct Belemnites’. Hence the inference may as confidently be
drawn, as from a dissection of the animal of the Ammonite, that
this Cephalopod iacked, like the Nautilus, the singular defensive
contrivance with which the more active Dibranchiate Cephalopods
were endowed, and that the animal of the Ammonite was compen-
sated, like that of the Nautilus, by having an external protective
shell into which it could retreat and close the entry against the
assaults of an enemy. Moreover, admitting the homology of the
“Aptychus”’ with the “hood,” we further learn that the defensive
door of the house was “dorsal,’’ and that the relative position of
_ the soft parts to the external shell was the same in Ammonites as in
Nautilus.
It cannot be averred, therefore, in excuse of a nomenclature im-
plying a different and opposite relative position of the soft parts to
the shell, that “the animal of the Ammonite is unknown.”
To the composite porcellano-nacreous structure by which the
Ammonitic agree with the Nautiloid series of shells, and their diffe-
rence in this character from the simply nacreous structure of the
Spirula-shell, reference is here made in illustration of the ‘ Law of '
Correlation.” ‘The conformity, in this respect, with the Nautiloid
series is maintained under every modification of shape from straight
to convolute.
But the persistence with which monographers of these numerous
and beautiful fossils, notwithstanding the appeals of Pictet? and
M‘Coy*, and the practice of Barrande, adhere to the erroneous views
of Von Buch as to which was the dorsal and which the ventral aspect
of the shells, has moved me to supplement the original grounds of
1 Buckland, Bridgewater Treatise, vol. i, 1835; Owen, “On certain Be-
lemnites from the Oxford Clay,” Phil. Trans. 1844.
2 Paleontology, vol. ii. p. 618.
3 Ann, & Mag. Nat. Hist., 2nd ser. vol. viii. p. 481.
964 PROF. OWEN ON THE [Nov. 19,
my conclusions on that question by additional or confirmatory evi-
dence from autopsy of the fabricators of the still existing polythala-
mous and siphoniferous Cephalopods.
Having prevailed upon Mr. Cuming, subsequently to my Mono-
graph on the anatomy of Spirula in the ‘Zoology of the Sama-
rang,’ to permit me to test that anatomy by dissection of his unique
specimen, I proceed to point out such structural facts, confirmed by
that dissection, as demonstrate the ventral position of the marginal
siphon of the internal shell of Sperula—and to combine therewith
corresponding anatomical observations on Nautilus, which, with the
demonstrated affinity thereto of Ammonites, equally prove the
ventral position of the marginal siphon of the external shell of that
extinct genus and its allies.
The dorsal aspect of a Cephalopod is determined by the position
of the brain and eyes, i.e. by that predominating part of the brain
which sends off the optic nerves. The ventral aspect is shown by the
respiratory funnel (Plate LX. figs. 3 and 4, 7, and gills, m). No mala-
cologist has questioned these conclusions. The proposition might be
simplified by stating that the funnel shows the “ventral side” of
the animal, and that the opposite one is the “dorsal side.”
Accordingly, all who have occupied themselves with the organiza-
tion of the Cephalopods have pointed out the singular reversed posi-
tions of the mandibles as compared with those in such vertebrate
animals as repeat the cephalopodic condition of a “beak,” as, e. g.
Chelonia and Aves.
Instead of the dorsal or upper mandible being the largest and
longest, so as to overlap the ventral or under mandible, as in birds,
the dorsal mandible, A, figs. 3 and 4, is the smaller and shorter one,
and is underlapped by the larger and longer ventral mandible, i, in
all Cephalopods.
So, likewise, the branchize, 70. figs. 3 and 4, m, m, lie in the ven-
tral part of the pallial cavity ; and, in short, the several viscera
occupy similar relations to back and belly in both Tetrabranchiate
and Dibranchiate Cephalopods’.
The aspects of the body being thus abundantly and unmistaka-
bly determined, they can be as unequivocally predicated of the shell
under whatever shape or proportion it may be present.
The portion of the shell of Nautilus which extends from the
fundus of the last chamber, along the dorsal aspect of the mantle,
as from A to A’, fig. 3, is the ‘dorsal wall” of such shell, and so
must be its continuation backwards.
The portion of the shell of Spirwla which bears the same relation
to the dorsal aspect of the body is the “ dorsal wall” of the shell,
A, Al, fig. 4; while that part of the last chamber which protects the
ventral side of the muscular and visceral mass which it contains, small
though it may be, is part of the ‘‘ ventral wall”’ of the shell, B, and
so must be its continuation as far as it extends, viz. to the ‘‘ proto-
conch”’ (fig. 5, @) or incipient nuclear chamber.
1 See ‘‘ Supplementary Observations on the Anatomy of Spirwla,” in Annals
and Magazine of Natural History for January 1879.
1878.] SHELLS OF CEPHALOPODS. 965
Accordingly, the dorsal wall, A, of the spiral shell of Spirula
describes a convex curve, the ventral wall, B', a concave curve. In
Nautilus the curves are reversed. If the shell of Orthoceras or
Baculites were curved so as to present the same relative positions
of convexity and concavity which exist in that of Spirula, they must
have been coiled in a reverse direction to that which is presented in
the shells of Nautilus and Ammonites.
Admitting that the siphon in Ammonites and Spirula are both
“ventral” or “ventro-marginal,”’ yet the relative position to the
shell is so contrasted as to call imperatively for terms indicative of
such contrast. And such terms are obvious. Of a convolute shell
the convex curve is the outer one, the concave curve the inner one,
whatever may be the relative position of its constructor.
Had Von Buch been content to call those curves in a convolute
Ammonite “ external’? and “internal,” and to define the position of
the siphon as “externo-marginal” and ‘interno-marginal” respec-
tively, his terminology would have stood and been unquestioned to
the present day. But he proceeded to a conclusion as to their
relations to the constructors of the shells; and, as no specimen of
either a Nautilus or a Spirula had been dissected in 1829, such
conclusion could only be a guess.
So probable, however, seemed the guess, that his most experienced
contemporaries, De Blainville, Grant, and Gray, after the organiza-
tion of the Pearly Nautilus had been made known, preferred, as we
have seen, the opinion of the conchologist to that of the anatomist,
and deemed the latter to have reversed the true position of the animal
of the Nautilus'. And, with most?, the nomenclature of the parts of
the shell of the Ammonitide has continued in concordance with that
opinion to the present day.
If, however, the facts and inferences now submitted to the
Zoological Society should be accepted with their logical applica-
tions in conchology, and the siphon in both Ammonites and Spirula
be acknowledged to be “ ventral,” or “ ventro-marginal,’’ nevertheless
the different positions of the siphon in these shells and in that of
Nautilus demand to be defined by distinctive terms. And these
are easy, obvious, and incontrovertibly applicable. The siphon in the
Ammonitide is “ external,” “externo-marginal,” or “ ecto-marginal.”’
The siphon in Spirula is “internal,”’ ‘ interno-marginal,” or “ ento-
marginal.” The siphon in the Nautilide may be “central,” or
*‘subcentral ;”’ and if the latter, either ‘‘ectocentral’’ or “‘entocentral,”
according as it deviates from the typical central position toward the
1 See notes 2 and 3, p. 957.
2 Even in 8. P. Woodward’s classical ‘Manual of Mollusca,’ 12mo, “ dorsal”
and “ventral,” are used as synonyms of “ external” and “internal.” Thus the
genus Ammonites is characterized by “siphuncle dorsal” (p. 94, and p. 197, ed.
1868). The genus Gonéatites is characterized by “ siphuncle dorsal” (p. 93).
The experienced Editor, in a later edition of the ‘ Manual of the Mollusca,’ in
his ‘ Appendix’ (12mo, 1868) approximates to the conclusion here advocated, by
amending the character of Fam. iii. Ammonitide, thus:—‘ Siphuncle—convexo-
marginal?” (p. 10). If the present paper should serve to dissipate Mr, Tate’s
remaining doubt it will be an acceptable return for my labour,
Proc. Zoou. Soc.—1878, No. LXIIL. 63
966 PROF. OWEN ON THE [Nov. 19,
external or internal part of the shell : in some rare cases (Cryptoceras)
the siphuncle is ectomarginal, as in Ammonites; and in still rarer
instances (C/ymenia) it is entomarginal, as in Spirula. There are
also species in which it begins by being marginal, and gradually shifts
to the more typical position as the shells grow and the number of
the septa increases.
In the Nautilus imperialis, e. g., the siphuncle is at first, i. e.
along the first twenty chambers, entomarginal, or near the concavity
of the shell-curve, as in Spirula; but in opposite relations to the
back and belly of the animal. After the twentieth chamber the
siphuncle gradually gains the central or excentral position’. Nautilus
ztezac shows a similar structure. The immature position of the
siphon in the existing Nautilus was longer retained in the old
Tertiary species.”
Various suggestions have been made as to the efficient as well as
final cause of the successively vacated parts called ‘ chambers,”’ with
their partitions and connecting siphon, in the polythalamous and
siphoniferous shells of Cephalopods.
In elucidation of this question, much mooted by different writers *
Vermetus gigas.
Section of chambered part of shell.
after the publication of my Memoir on the Pearly Nautilus, I
adduced the instances of such vacuities or chambers in the Mollusca,
? Catalogue of Hunterian Cephalopoda, 4to, no. 137, p. 82.
? See the just remarks of Barrande on the variation of the siphuncular cha-
racter in certain nominal species (Céphalopodes Siluriens de la Bohéme, 8vo,
1867, p. 24).
3 Buckland, ‘Geology and Mineralogy considered with reference to Natural
Theology.’ 1835, vol. i. p. 329. De Blainville, “Sur l'usage du siphon des
coquilles polythalames,” Annales frangaises et étrangéres d’Anatomie et de
Physiologie, t.i. 1837.
1878. | SHELLS OF CEPHALOPODS. 967
as, e.g., Fistulana clava, Vermetus giyas, Helix decollata, the Ostree,
Estheria, and other Bivalves, more especially the Spondylus varius’.
In Fistulana and the gastrepodal Vermetus, the animal periodically
withdraws itself from its dwelling-chamber ; the growth of the walls
is continuous and uninterrupted; but a thin new floor is formed at
some distance from the old one left behind, and a series of chambers
fig. 1, aa, results. If the calcareous deposit had been continuous in
every part of the shell, a solid tract would have been left behind, as in
Magilus. The successive floors or “septa’’ in Vermetus (fig. 1, 6 6),
extend freely across and are concave toward the outlet; they are
entire and adherent only by their marginal circumference to the
shell-wall (A, B). The contents of the chambers in the living
Vermetus are unknown. Both chambers and partitions are the con-
sequence of a mode of shell-growth ; physiological ken stretches not
beyond this.
In Spondylus varius (fig. 2) the “septa’’ are not continued freely
across the shell, but are united together near the middle or centre
of their extent, at the position of the impression of the adductor
Spondylus varius.
Section of chambered part of shell.
muscle. This, in the forward movement of the mollusk, does not
quit its attachment to the nacreous layer of the valves; whilst the
pallial lobe, except at its circumference and where it adheres to the
adductor, can and does detach itself from the surface of the valve
about to be abandoned, in the progressive growth of the visceral
mass. The mantle at each period of repose, then secretes on the
fluid occupying the deserted part of the shell, the new septum or
1 Proc. Zool. Soc. 1837, p. 63. See also H. Woodward, “On the Struc-
ture of the Shell of the Pearly Nautilus,” Report of the British Association,
Liverpool, 1870, Trans. of Sections, p. 128. ae
968 PROF. OWEN ON THE [Nov. 19,
basis of support of the mollusk. Thus the septa, as they are succes-
sively formed, adhere, not only to the circumference of the growing
valve, but to a central part of the preceding septum, and for an
extent more or less corresponding to the circumference of the corre-
spondingly advancing adductor muscle.
If the adductor were a tube instead of a solid mass, the central
confluent part of the septa would be perforated, and a siphon would
result, the calcareous walls of which would be continuous, as in
Spirula, Nautilus striatus, the Orthoceratites, &e.
The contents of the deserted chambers in Spondylus varius are
sea-water with an increased proportion of the saline ingredients.
The efficient cause of the forward movement of the Spondylus varius
appears to be the need of a shell of a size suitable to the growing
bulk of the animal, coupled with the frequeut fixation of the lower
valves of the young shel! to an overcrusting mass of coral, in advance
of which the growing shell must increase. Such increase and the
testaceous provision for it are not, therefore, attributable to special
expansion of one organ, but to the concomitant growth of the whole
of the soft parts of the Palliobranch.
It has been suggested that the periodical increase of the ovarium
or testis might initiate and constrain the forward movement of the
soft parts in the Cephalopods with chambered shells, and that a
polythalamous structure is related conditionally to the generative
function’.
But it will be observed, in both Spirula and Nautilus, that the
formation of the chambers commences from the embryonal cup (proto-
conch, fig. 5a, p. 973), and continues through an early period of growth
antecedent to the acquisition of the procreative function, or the adult
stage of existence—and, moreover, that those early chambers are re-
latively deeper® than the succeeding ones, indicative of a more extensive
forward movement of the soft parts, in accordance with the more
rapid growth of the animal which characterizes the period of nonage,
when all the assimilative functions are concentrated on general
increase and no degree of that power is diverted to the development
of special organs, such as the testis or ovarium.
The last or open chamber of Nautilus, and, by analogy of size and
certain known contents, in Ammonites, was occupied by the entire
soft parts. In Spirula it contains only the hind end of the liver
and portions of the origins of the retractor muscles of the head and
funnel. It has been stated to contain the ink-bladder in Spirula® ;
but in my dissections of that Cephalopod made subsequently to that
detailed in the ‘ Zoology of the Voyage of the Samarang,’ I find the
same positions and relations of the ink-bag as are described and figured
in that monograph‘.
1 Prof. Seeley, Proceedings of the British Association, Bath, 1864, Section
Zoology, ‘‘ Nautiloid shells,” 8yo, p. 229.
* By depth is meant the diameter from septum to septum; by breadth that
between wall and wall.
* Woodward, ‘Manual of Mollusca,’ p. 77.
4 “A very minute pyriform ink-bag, z, is situated close to the rectum; and
its duct opens within the verge of the anus” (p. 10, pl. iv. fig. 11).
1878. ] SHELLS OF CEPHALOPODS. 969
As to the function or “ final cause”’ of the chambers, I hold by
the opinion expressed in my original memoir! and in the ‘ Catalogue of
the Fossil Cephalopoda in the Hunterian Museum,’ 4to, ed. 18567, viz.
that they so affect the specific gravity of the active, highly organized,
cephalopodous mollusk, as to enable it with little effort to rise, in the
case of the Nautilus, from its habitual position at the bottom of the
sea—and in the case of the Spirula, to sink from its more usual zone
at or near the surface,—such vertical movements being executed, like
the horizontal ones, by means of the hydrostatic mechanism worked
by the muscular forces of the mantle and funnel.
The contents of the vacated chambers in Nautilus pompilius are
stated to be nitrogenous gas. Neither the contents nor the vital
properties of the siphuncle are yet known; an artery and vein are
assigned for its life and nutrition, and to extend a low degree of the
vivifying influences to the shell. Vrolik confirms the existence of
the siphonal artery described and figured in pl. vi. fig. 1, 14, of my
‘Memoir,’ and repeats this illustration in his pl. i. fig. 2, “2, artére
allant au siphon” *. .
The siphonic artery sends off, according to Keferstein, branches to
the mantle which lines the bottom of the body-chamber before penetra-
ting the siphon. Waagen figures the impression of these arterioles on
the shell-surface*; and it has been suggested that these vessels may
supply, by secretion, the chamber-gas which [ inferred might occupy
the space left free on the recession of the visceral sac from the
chamber-floor prior to the formation of a fresh septum’.
Thus, in the analysis of the structure of chambered shells, we
find :—septa simple, distinct, attached only by their circumference
(woodcut fig. 1, p. 966); septa attached, subcentrally, to each
other, as well as by their circumference to the shell-walls (fig. 2,
p- 967) ; septa (fig. 3 6, p.-907) attached marginally to the shell-
walls, A, and also to each other by tubular prolongations, c, with
an organized, vascular, membranous canal, d, traversing such tes-
taceous tube; septa with a calcareous siphuncle consisting of a
series of superimposed, elongate, funnel-shaped tubes, with the
wide end directed toward the aperture of the shell, as in Spirula
(fig. 4, p. 971), or in the opposite direction, as in Bathmoceras.
The more complex siphons of Orthoceratites will be presently
referred to. Finally, we see in the existing Nautilus the shelly tube
interrupted, forming the “collar of the siphon,” and the septa and
chambers traversed by a lime-coated membranous canal running
through the interrupted shell-tube (Plate LX. fig. 3, ¢, d).
1 “From the adhesion of the entire circumference of the mantle to the shell
by means of the ‘horny girdle’ (‘annulus, Waagen, op. cit.), I am inclined. to
suppose that the whole of the chambers are excluded during the life-time of
the animal from external influence, and are filled only by exhalations or secretions
from the animal.”—Memoir on the Nautilus (1832), p. 47.
2 «The proportion of the air-chambers to the occupied dwelling-chamber of
the Nautilus is such as to render the whole animal of nearly the same specific
gravity as the surrounding water,”—Catalogue, p. 29.
3 Mémoires de la Soc. Linnéenne de Normandie, tom, x, 1855, p, 17,
4 Waagen, op. cit. Taf. xxxix. fig. 4, ,
5 Op, cit, 4to, 1832, p. 47,
970 PROF. OWEN ON THE [Nov. 19,
For the different views which have been propounded as to the
nature and function of this complex siphon, reference may be made
to the authors cited, p. 966. Some of these views were based on the
partial knowledge of its structure at the date of the first dissection
of the Pearly Nautilus.
The true structure of the siphon in Nautilus pompilius is rarely
preserved ; the somewhat loose calcareous matter by which the
membranous part is incrusted is commonly lost with that part in the
dry cast-off shells. The calcareous incrustation is apt to be dis-
solved, like that of the mandibles in Valenciennes’s specimen, by the
acetous change of the alcohol when charged with soluble parts of the
animal during a prolonged transit to a European museum.
Nautilus striatus.
Section of part of shell.
When cataloguing, in 1854 and 1855, the Hunterian Cephalopods,
I saw sufficient to supplement the description of the siphon in the
‘Memoir on the Nautilus,’ as follows :—“ An artery and vein are
assigned for its life and nutrition, and to extend a low degree of the
same influences to the shell; but the structure of the membranous
siphuncle presents, beyond the first chamber, an inextensible and
almost friable texture, apparently unsusceptible of dilatation and
contraction ; it is also coated beyond the extremity of the short
testaceous siphuncle with a thin calcareous deposit”’. The fact of
this incrustation has been ascertained, independently, by Prof..
Vrolik. The subject of his memoir in the volume cited in note 1, was
1 Descriptive Catalogue of the Fossil Organic Remains of the Invertebrata
contained in the Museum of the Royal College of Surgeons of England, 4to,
1856, p. 29. This volume was not issued until all the invertebrate fossils were
described ; and the first sheets were printed off before the ‘ Mémoires de la
Société Linnéenne de Normandie,’ yol. x. 1855, came to my hands,
1878. ] SHELLS OF CEPHALOPODS. 971
in such a favourable state of preservation that the calcareous incrusta-
tion of the membranous siphon was entire, and formed the subject of
the continuous inflexible composite tube represented in pl. i. fig. 5, a, b,
in that accomplished naturalist’s memoir. The notion of the dilating
and contracting action of the siphon of the Nautilus upon its con-
tents, whatever these may be proved to be, could be no longer enter-_
tained. Vrolik, confirming by his dissection the existence of the
siphonal artery, infers, like myself, the coexistence of a returning
channel, although no vein with definite tunics was demonstrable in
either dissection. .
Evidence of the capillary ramifications of the siphonal artery upon
the pallial membrane lining, as periostracum, the interior of the
shell-chambers has been adduced by the careful observers Keferstein
and Waagen ; but such channels of vitality are not supposed to pene-
trate the shell itself. Molluscous shells, like avian feathers and
mammalian hairs and teeth, do not receive the terminal divisions of
the blood-vessels supplying their several pulps or formative organs.
Ordinary or hard dentine, like conchine, piline, and plumine, is extra-
vascular, but not, therefore, extravital. The percolation by cellular
passages and intervals of a rarer, plasmal exudation from the vital
Spirula australis.
Section of part of shell, magnified:
fluid renders intelligible the change and movements of pigment in
the same hair and in the same feather. As the dentist distinguishes
dead from living teeth, so the conchist regulates his estimate of
the value of a “ dead” as contrasted with a “living” shell. The
estimable researches of Carpenter on the modifications of micro-
scopic texture in shells parallel those that have demonstrated as
many modifications of the microscopic channels by which the
plasma percolates the dentinal as it does the chonchinal tissues!.
The high organization of the Cephalopods compared with other
* Laborious studies of this kind in quest of truth beget a modest reticence
and an abstaining from such dogmatic utterances as those of the writer who
“denies the possibility of the siphuncle maintaining the vitality of the shell,
=
because it is certainly a non-vascular structure.”—“ Recent and Fossil Cepha-
lopods,” Geological Magazine, yol. y. p. 490.
972 PROF. OWEN ON THE [Nov. 19,
mollusks suggests to the physiological mind the probability that
some corresponding step in advance of the ordinary condition of a
testaceous defence would be likely ; and the retention of the parts of
such defence vacated in the course of growth, with the superaddi-
tion of a vascular by-way running through the whole, verifies such
anticipation of the means whereby the camerated and siphoniferous
shell is thus brought into closer harmony with the rest of the organic
structure.
When the first simple, smgle-chambered, nuclear dwelling is added
to and, in part, abandoned, a closer connexion is therewith preserved
than in Vermetus, e.g.; and, as the artery penetrating the mem-
branous siphon continued from the last-formed dwelling-chamber
was demonstrated by mercurial injection’, there is no reason to
doubt that such organic comexion was maintained between the
fabricator of the second chamber after it had advanced from and
vacated the first or nuclear one.
The constancy of this siphuncular connexion running through all
the chambers of the largest and most complex of the polythalamous
shells, with the great size and singular complexity of the siphunele
in several extinct species, form the grounds on which I still hold te
my original belief in the function of the siphuncle as related to a
maintenance of the vitality of the shell. But this relation may
be connected, alsu, with a greater share assigned to the siphon in the
protection of the soft parts of the Cephalopod at the earlier stages of
its existence.
The chief charaeter of the tetrabranchiate chambered and sipho-
nated shell is its affording, besides a sheath or case to the whole
animal, a speeial protection to a part of the animal.
Such twofold office is performed by the shell in certain Gastro~
pods, conspicuously in the genus Calyptrea, in which an accessory
“cup,” springing from the concavity of the larger “ saucer,” lodges
part of the muscular system’.
Every chambered and siphonated shell begins in this simple fashion.
The protoconch is cup-shaped or flask-shaped, and includes a similar
bat smaller blind beginning of the siphon.
The proportion of the inner partially protecting shell to the outer
wholly encasing shell is greatest in the Silurian Orthoceratites ; and
with the large proportional siphon are associated complexities cha-
racteristic of the genera Ormoceras, Huronia, &c.°
' Modifications of the contents of such siphons in the Silurian Vagi-
1 «he lesser aorta” sends off a small branch (a4, pl. 5 & 6) “which, winding
round to the ventral aspect of the ventricle (to which it is connected by a pro-
cess of membrame), passes through a foramen in the septum dividing the peri-
cardium from the cayity at the bottom of the pallial sac, is then continued
through that cavity, passing between the ovary and gizzard, and lastly enters,
without diminution of size, the membranous tube that traverses the parti-
tions of the shell.”—Memoir on the Pearly Nautilus, p. 36.
* “Tts cavity is filled by what maybe termed the apex of the foot, which here
loses its muscular character, and assumes a gelatinous texture.”—“ Anatomy of
the Calyptreide,” Trans. Zool. Soc. vol. i. p. 208, pl. 30. figs. 2, 6f, +85 /f35-
3 Charles Stokes, in Geol. Trans. 2nd ser, yol. y. p. 706, 1837.
1878. ] SHELLS OF CEPHALOPODS, 973
nati of Bohemia led the exemplary and acute observer Barrande to
conceive that a larger proportion of the soft parts had been therein
lodged than is the case in the slender simple siphons of other Tetra-
branchiates, and in all that are of later date, when the partial office
of the siphon is reduced to reception of a trace of the vascular system.
Produce both the *‘saucer” and cup” of Calypeopsis, with the
resultant ‘false bottoms” or septa, as in Vermetus (fig. 1, p- 966),
and an analogue, if not parallel, of Endoceras would ensue.
Hyatt has noted the resemblance of the cup-like internal chambers
of Beatricea to a line of siphonal czca. The protoconch (primal
shell, nucleus, ovisac, embryo, &e., fig. 5, a) has been determined,
Spirula australis.
Protoconch and protosiphon, with following chambers, magnified.
described, and figured by De Blainville (in Spirula’) ; also by
Sandberger’, by Barrande’, and, with exemplary care and patience, by
Alpheus Hyatt’, in various species of tetrabranchiate shells. The
results by no means support any doubt as to the externality of the
shells of the Ammonitidee, or at all support the notion of their closer
affinity to Spirula than to Nautilus. :
The supposed well-marked distinction between the protoconch of
Ammonites and that of Nautilus is mainly due, as Hyatt has shown,
to its decadence in Nautilus after the deuteroconch (fig. 5, 6), or suc-
cessional shell-chamber, is formed—the primal communication with
the protoconch being then indicated by a linear cicatrix on the first
septum’. On this indication of the original existence and attach-
ment of the protoconch I may remark, that when the young Nau-
tilus, becoming too big for its first shell, moved on to make another,
part of that “making” encroached upon the space which the young
had traversed, and reduced it more or less to the shape of a chink
or scar. But it must not be concluded that this chink preexisted
to the progress, and that the young crept or squeezed through such
chink, as when Mr. Hyatt writes of ‘the central scar which marks
the former aperture through which the animal probably passed into
the fundus of the first whorl” ® It is only under such impression
that it becomes “ difficult to account for the passage of the large
1 Op. cit.
: Cpiesieche Gesellschaft fiir Natur und Heilkunde, 1858.
~ CU.
4 Fossil Cephalopods of the Museum of Comparative Zoology.—Embry-
ology (Svo, Cambridge, Mags.), vol. iii, no. v. 1872.
5 Ibid. p. 64. § Ibid. p. 73.
974 PROF, OWEN ON THE SHELLS OF CEPHALOPODs. [Nov. 19,
body of the embryo through the narrow aperture thus made’ !.
The difficulty merely depended upon the “time of the making” the
communicating aperture so narrow.
A true ‘‘embryology”’ of the Tetrabranchiates will be the welcome
result and reward of the fortunate and equal student of the living
Nautilus and Spirula, on the shores or in the seas they still fre-
quent, or in the aquarium adapted to conserve and exhibit the male
and female of these representatives of the almost extinct order.
I cannot conclude the present paper without expressing the
pleasure with which may be confidently expected the determination
of important points in the physiology of the Tetrabranchiate and
Dibranchiate constructors of chambered and siphonated shells by
observations made on the living specimens, under the favourable
opportunities afforded by their capture to the accomplished Natu-
ralists of H.M.S. ‘Challenger,’ whose attention had no doubt been
directed to the following needful observations and experiments sug-
gested in previous works on the subject.
“It would be advisable in the event of another fortunate capture
of the Nautilus, to lay open the chambers under water, when the
presence of gas in any of them would be ascertained ; and it might
be received and analyzed ; the contents also of the central tube, if
gaseous, would at once be detected ”’ ?.
Prof. Vrolik records similar appeals on more than one point in
the structure and physiology of the constructor of the chambered
shell. =
““M. Owen a eu raison de dire que l’orifice 2 (pl. ii. fig. 2) est en
communication avec le péricarde.”’.,.‘« D’aprés ce que j’ai vu, le siphon
s’ouvre dans la cavité abdominale, comme la pl. i. fig. 5, ¢c, le dé-
montre, et dans laquelle j’indique aussi la communication entre la
cavité branchiale et le péricarde en d, et entre celui-ci et la cavité
abdominale ou splanchnique en e.”...“En ouvrant la paroi postéri-
eure du péricarde, je me suis assuré qu’au fond de celui-ci il y a une
ouverture, que M. Owen a deja reconnue, et dans laquelle pénétre
une petite artére (pl. i. fig. 2, /). Cette ouverture est mise en rap-
port, prés de lovaire et du gésier, avec la cavité abdominale.”’
With regard to the course of sea-water from the branchial to the
pericardial or peritoneal cavity and thence to the cavity of the siphon,
or the reverse course, Vrolik appeals to the fortunate possessors of
the living Nautilus :—‘* L’observation directe seule pourrait les dé-
cider’’®.
And again, in regard to the circulation, especially of the venous
blood :—‘‘Serait-il possible que le sang veineux du canal intestinal fut
versé dans ces interstices ou dans ces /acunes, pour passer plus tard,
par les orifices cités, dans la veine cave ; et le sang veineux des autres
1 Op. cit. p. 74.
? Memoir on the Nautilus pompilius, 4to, 1832, p. 47. The same might be
urged in the case of the capture of a living Spirula.
* “Sur quelques points de V’organisation de l'animal du Nautile Flambé.”
Mémoires de la Société Linnéenne de Normandie, tom. x. (1885), p. 7, pls. i.
and il.
1878.] THE SECRETARY ON ADDITIONS TO THE MENAGERIE. 975
viscéres abdominaux serait-il regu dans des vaisseaux veineux qui se
réuniraient en suite pour former les trones que M. Owen représente
planche vi. 3,3? Je n’ai pas de réponse A donner A ces questions.
L’injection du Systeme vasculaire d’un individu a létat frais, et
observation d'un Nautile vivant pourraient seules les éclaircir.?—
VROLIK, op. cit. p. 11.
EXPLANATION OF PLATE LX.
Fig. 1. Section of Ammonites lingulatus, showing the opercula, 0, displaced but
retained in the dwelling-chamber,
2. Ammonites subradiatus, with the operculum, 0,72 st¢u, closing the aperture
of the shell, save the space, v, for the passage of the respiratory
currents.
3. Section of Nautilus pompilius, with diagrammatic outlines of the muscles
of the shell and of the soft parts indicative of the relative position of
the animal.
4. Section of Spirula australis.
In the figures 3 and 4 the parts are indicated by the same
letters, as follows :—
Shell.
A. Dorsal wall. B. Ventral wall. a. Chambers. 4, Septa. c. Hard siphon.
d. Soft siphon.
Mantle.
e. Dorsal wall. f£ Ventral wall.
Head,
g. Tentacles or prehensile organs,
Beak.
h. Dorsal mandible. 7. Ventral mandible. 7. Funnel. %. Retractores infun-
dibuli. 7 Retractores capitis. mm, Branchie, 7, Liver.
December 3, 1878.
Robert Hudson, Esq., F.R.S., V.P., in the Chair.
The Secretary made the following report on the additions to the
Society’s Menagerie during November 1878 :—
The total number of registered additions to the Society’s Mena-
gerie during the month of November was 66, of which 2 were by
birth, 26 by presentation, 28 by purchase, 1 received in exchange,
and 9 received on deposit. The total number of departures during
same period, by death and removal, was 89.
The most noticeable additions during the month were :—
1. Two examples of Horsfield’s Tortoise, Testudo horsfieldi, from
the neighbourhood of Tschina in Turkestan, presented by Dr. A.
Strauch, C.M.Z.S., of the Imperial Zoological Museum of St. Peters-
burg, November 15th.
The species having been established by Dr. Gray upon a single
young specimen ', said to have come from Afghanistan, Dr. Strauch
was anxious to have examples of the Central-Asiatic species (which
1 Gray, Cat. Tort. Brit. Mus. p. 7 (1844), and Cat. Shield Rept. in Brit. Mus,
p: 7, tab. i. (1855),
=
976 MR. P. L. SCLATER ON SAXICOLA STAPAZINA. [Dec. 3,
he has excellently described in his ‘ Chelonologische Studien,’ p. 86)
compared with the original type. This, through Dr. Ginther’s
kindness I have now been able to do; and I have no doubt of the
identity of the species, in which opinion Dr. Giinther fully coneurs.
Besides the original type, there are two small examples of this Tor-
toise in the British Museum, obtained in Persia, during the Survey
of the Persian Boundary Commission’; so that the species would
appear to range from Afghanistan through Persia onto the shores
ot the Caspian and Aral seas and so into Turkestan, whence the
present specimens were forwarded to the Imperial Zoological Mu-
seum of St. Petersburg by their conservator, von Russow.
Testudo horsfieldi in general appearance much resembles Testudo
greca, but has only four toes on the front limbs.
2. A small blue Macaw, apparently referable to Spix’s Macaw
(Ara spivi), purchased of the Jardin d’Acclimatation of Paris,
November 15, and new to the Society’s collection.
This bird seems to agree in every respect with the species as de-
scribed by Finsch*, except that the bare space round the eye and
lores are black, and not yellow as described by Finsch and as also
described and figured by Spix.
I have examined the single specimen of this species in the British
Museum (obtained by purchase from a dealer in 1859), and believe it
to be the same as our bird. Our specimen, of which I exhibit a
figure by Mr. Smit (Plate LXI.) is generally of a bluish grey, the
head of a paler light grey, a narrow naked ring round the eye;
naked lores and bill black; total length about 18 inches, wing 10,
tail12. The iris is of a pale yellow.
This is the first example of this rare species I have ever seen alive.
It is immediately distinguishable from Ara hyacinthina and A.
glauca (of both of which we have living specimens in the collection)
by its small size and small bill.
Mr. H. Seebohm exhibited a large series of specimens of the
Hooded and Carrion Crows (Corvus corniz aud C. corone) and of
the intermediate forms between these species, and made remarks on
their geographical range and on the origin of the hybrids.
Col. Irby, F.Z.S., exhibited and made remarks on some nests,
eggs, and young of Cypselus pallidus taken at Gibraltar.
Mr. Howard Saunders exhibited and made remarks on some
Indian eggs, especially those of Sterna bergii and Larus hemprichii,
on behalf of Capt. Butler, H.M. 83rd Regiment. These specimens
had been taken on the island of Astola, on the Mekran coast.
Dr. F. Day, F.Z.S., exhibited and made remarks on three jaws
of Indian Sharks—one belonging to the genus Galeocerdo, and two
belonging to the genus Carcharias,—and pointed out that one of the
1 See Blanford’s ‘ Zoology of Persia,’ p. 308.
2 Sittace spixi, Finsch, Papag. vol. i. p. 393,
Nanharbt im
JSmit lith
Mute song Us qayyy
JAR BLS “S 2d
1878. ] DR. A. GUNTHER ON MIDIAN REPTILES. 977
latter had the teeth of Odontaspis, whereas the other had the teeth
in the upper jaw furnished with cross serrations, approaching in
this respect Galeocerdo, while the teeth in the lower jaw were more
like those of Odontaspis.
Mr. Sclater read a letter with enclosures from Mr. R. Davenport,
of 124 Georgiana Street, Bury, whereby it appeared that there could
be no doubt of the specimen of the Black-throated Wheatear
(Sazicola stapazina) exhibited at the last meeting having been
obtained in Lancashire, as there stated’. The bird was shot by Mr.
David Page, of 103 Spring Street, Bury, on or about the 8th May,
1875, whilst sitting on the ridge of the outbuildings belonging to the
Bury Angling Association near the reservoir. It was taken in the
flesh to Mr. Wright Johnson, of Prestwick, to be mounted. Mr.
Johnson had ascertained the sex by dissection to be male.
The Secretary called attention to the error that had accidentally
taken place in reference to the Collection of Butterflies from Billiton,
on which Messrs. Salvin and Godman had reported in the last
number of the Society’s ‘ Proceedings’ (antea, p. 637). The col-
lection had been made and forwarded to England by Hr. J. G. F.
Riedel, of Koepang, Resident of Timor and its dependencies.
The following papers were read :—
1. On Reptiles from Midian collected by Major Burton. By
Dr. Atsert Gtntuer, F.R.S., F.Z.S.
[Received October 31, 1878.]
(Plate LXIT.)
Major Burton has kindly presented to the Trustees of the British
Museum the zoological specimens collected by him in Midian*. The
reptiles are five in number, viz. Gongylus ocellatus, Zamenis clif-
fordi, Zamenis elegantissimus (sp. u.), Echis carinata, and Echis
colorata (sp. n.),—all of which are characteristic of the Cireum-
mediterranean fauna. A toad proved to be the common Bufo vul-
garis.
Major Burton does not seem to have had the means of preserving
any mammals and birds ; and it is to be hoped that travellers who
may succeed him in the exploration of this interesting country will
be more fortunate in this respect, as, at present, we are entirely
ignorant of this part of the Midianitish fauna.
ZAMENIS ELEGANTISSIMUS. (Plate LXII.)
The body is moderately slender, head rather narrow, eye of
moderate size. _ Rostral shield convex, reaching the upper surface
of the head. The scutes of the upper surface of the head are not
1 Above p. 881.
? See Burton’s ‘Gold-mines of Midian,’ London, C, Kegan Paul & Co., 1878.
978 MR, H. SEEBOHM ON SYLVIA BLANFORDI. [ Dec. 3,
subdivided : anterior frontals much smaller than the posterior ;
vertical large, broad in front, with the lateral margins concave ;
occipitals rounded behind. Loreal nearly square; three ante-
oculars, the upper of which is large, concave, extending onto the
vertical shield ; the two lower are small, and may be regarded as
detached portions of the fourth upper labial. Two postoculars ;
eight upper labials, of which the fifth enters the orbit. Temporals
seale-like, 2+3+3, the lower of the anterior pair being larger
than the upper. The scales smooth, in 19 rows. Ventrals 197 ;
anal bifid; subcaudals 79. Abdomen with a slight ridge on each
side.
The ground colour of this most beautiful snake is olive; a bright
orange streak begins on the vertical shield and runs along the entire
median line of the back and tail. It is interrupted by broad black
cross bands, of which the two anterior occupy the crown of the head,
twenty-one encircle the trunk, and eight the tail. The bands on
the trunk do not reach across the middle of the abdomen; but the
rings on the tail are complete.
The length of the entire specimen is 26 inches, that of the tail
being 64. It was found on the mountains east of El Muwaylah.
ZAMENIS CLIFFORDII, Schleg.
A young specimen from the sandy coast-region of Tihamat Midian.
EcuIs CARINATA, Merr.
One specimen from the sandy coast-region.
EcHIS COLORATA.
This specimen approaches the true Vipers in general habit ; but
the scales on the side are arranged in the same peculiar manner
which is characteristic of Echis. In the thickest part of the body
the scales form about 35 longitudinal series. There are four series
of small scales between the eye and the upper labials. Greyish with
large pinkish spots on the upperside; they are rounded on the
front part of the body, but more irregular and broken up on the
posterior. Lower parts whitish, speckled and powdered with
greyish.
Ventrals 208 ; subcaudals 48.
The single example, which is much injured, was caught on Jebel
Sharr, at an altitude of 4500 feet above the level of the sea. It is
27 inches long, the thin tail measuring 3 inches only.
2. On a new Species of Sylvia from Abyssinia, and on some
other Abyssinian Sylvians. By Henry Szxsonm, F.Z,S.
[Received November 7, 1878.]
The only species of the genus Sylvia obtained by Jesse on the
Abyssinian expedition, was the common Whitethroat. This species
was also obtained by Blanford ; and in addition he records (Geol. and
cll il
1878. ] MR. H. SEEBOHM ON SYLVIA BLANFORDI. 979
Zool. of Abyss. p. 379) a single specimen of Sylvia melanocephala,
Gm., shot at Rairo in Habab. We learn from the personal narrative
that Blanford was at Rairo between the 10th and 15th of August.
The skin, doubtless obtained between these two dates, is in the
British Museum, and appears to me to belong to a hitherto unde-
scribed species, which I propose to call
SyLVIA BLANFORDI.
The general colour of the upper parts is brown, the innermost
secondaries, the quills, and the wing-coverts being narrowly
margined with brownish white. The cheeks, head, and nape are
brownish black. The tail is very dark brown, the outside tail-
feathers (which are much abraded) showing traces of having been
tipped with white. The general colour of the underparts is white,
shading into brown on the sides of the breast, flanks, axillaries, and
under tail-coverts. The bastard primary projects °3 inch beyond the
primary-coverts ; and the second primary is between the eighth and
ninth in length. The bird is moulting some of the primaries
between the third and the eighth. Both mandibles of the bill are
dark, and the tarsus and feet are dark slate-grey. The wing
measures 2°52, and the tail 2°62. The culmen, which is slightly
injured, measures about °51 when perfect.
The only species with which this bird can be confounded are S.
eurruca (Linn.), S. melanocephala, Gm., and S. rubescens, Blanf.
From S. curruea it is easily distinguished by its head being brownish
black instead of pale slate-grey, and by its tail being longer instead
of shorter, than its wing. From S. melanocephala it may be dis-
tinguished by the length of wing being 2°52, instead of varying from
2°15 to 2°35, and the colour of the tarsus and feet being dark slate-
grey instead of brown. Besides being a larger bird with darker feet,
it has a larger bastard primary, a shorter second primary, and has
less white on the outside tail-feathers. From S. rubescens it may be
distinguished by its tail being longer, instead of shorter, than the
wing, by its feet being dark slate-grey instead of palish brown,
and by its larger size, the less amount of white on the outside
feathers of its tail, its longer bastard primary, and more rounded
wing.
Sylvia blanfordi appears to be quite distinct from any of the birds
described by Riippell in his ‘ Neue Wirbelth. Abyss.’, or by Heuglin
in his ‘Orn. Nordost-Afrika’s,’ and also from the types of Hemprich
and Ehrenberg, in the Berlin Museum, described by Dresser and
Blanford in the Ibis (1874, p. 335).
Another error of identification in Blanford’s ‘ Abyssinia’ will be
found on page 358. The skin from Senapé in the British Museum,
labelled Ruticilla phenicura, Linn., does not belong to that bird, but
to the nearly allied species Ruticilla mesoleuca, Ehr. I have also exa-
mined the Pratincole from the Abyssinian collection in the British
Museum ; and Mr. Sharpe has pointed out to me that Pratincola
semitorquata, Heugl., is undoubtedly the breeding-plumage of
P. albofasciata, Ripp., and that Blanford’s skins labelled Pratincola
980 MR. H. SEEBOHM ON HORORNIS FoRTIPES, Etc. [Dec. 3,
pastor, Strickland, are undoubtedly P. indica, Blyth. This species,
distinguished, amongst other characters, from P. rudicola (Linn.), by
its pure white unspotted rump, and its almost entirely black axil-
laries, was first discovered in Europe by Harvie-Brown and myself
in the valley of the Petchora; Drs. Finsch and Brehm found it on
the Obb; and I brought home several skins from the Yenesay.
Although Severtzoff records both species from Turkestan, there can
be scarcely any doubt that Pallas’s birds from the Irtish and the
Tobol belong to the eastern species, which must therefore stand as
Pratincola maura, Pall. (Reise Russ. Reichs, ii. p. 708).
3. On the Identity of Horornis fortipes, Hodgs., Neornis
assimilis, Gray, Horeites robustipes, Swinhoe, Horeites
pallidus, Brooks, and Horeites brunnescens, Hume. By
Henry Seresoum, F.Z.8S.
[Received November 7, 1878.]
CeTTia ForTIPES (Hodgson).
Salicaria ?, Hodgson, icon. ined. nos. 900 & 923.
Horornis fortipes, Hodgson, J. A. S. Beng. xiv. p. 584 (1845).
Drymeca brevicaudata, Blyth, J. A.S. Beng. xvi. p. 459 (1847).
Horornis assimilis, Gray, Cat. Mamm. &c. Nepal coll. Hodgson,
p- 30, no. 143 (1863, ex Hodgson).
Horeites robustipes, Swinhoe, Ibis, 1866, p. 398.
Neornis assimilis, Gray, Hand-list of Birds, no. 3096 (1869).
Horeites pallidus, Brooks, J. A. S. Beng. xli. p. 78 (1872).
Horeites brunnescens, Hume, Ibis, 1872, p. 109.
An examination of thirty-two skins of Horornis fortipes, Hodgson,
H. assimilis, Gray, Horeites robustipes, Swinhoe, and Horeites pal-
lidus, Brooks, leads me to the conclusion that they all belong to one
species.
They agree in having the general colour of the upper parts russet
(not olive) brown, somewhat yellower on the rump. The wings are
brown, fringed externally with russet-brown, and margined inter-
nally with white. The tail is brown, fringed with russet-brown. An
indistinct eyebrow and the underparts are buffish white, shading
into ochraceous brown on the flanks, thighs, and under tail-coverts.
Axillaries and under wing-coverts pale yellow. Young birds have
the underparts yellower. The bill is moderately stout, dark horn-
colour above, pale horn-colour below, but darker towards the tip.
Rictal bristles slender. The wing is very rounded, the first primary
rather more than half the length of the second, and the fifth, sixth,
and seventh are nearly equal and longest. The tail consists of ten
feathers only, and is rounded, its outside feathers being about 0°45 inch
shorter than the longest. The foot and tarsus are robust, pale brown
in colour ; and the latter is very indistinctly scutellated in front. The
length of wing varies in adults from 1°9 to 2°28, the tail being about
one-twentieth shorter. The culmen measures about 0°5.
1878.] MR. 4. SEEBOHM ON HORORNIS FORTIPES, ETC. 981
Brooks obtained this species in Cashmere ; Mandelli has sent skins
from Darjeeling; Hodgson found it in Nepal, and Godwin-Austen
in Assam. Swinhoe’s type from Formosa seems less remotely
situated when we remember that Pére David obtained the nearly allied
species (Horeites major and H. brunneifrons) in China, where the
species under consideration doubtless also occurs.
I place this species in the genus Cettia, because it agrees with the
type of that genus in having only ten tail-feathers, somewhat simi-
larly graduated, and because both species have a somewhat similar bill,
and a rounded wing, not flat like that of a Thrush, but twisted like a
plough-plate to fit the body, evidently adapted less for extended
flight than to be out of the way when the bird is creeping through
dense foliage. Both species agree in having the feathers of the
rump considerably developed, and in laying eggs of a uniform dark-
red colour in a cup-shaped nest.
The position of this genus is somewhat intermediate between the
Turdine and the Timeliine. So far as I can see at present, I feel
disposed to restrict the Turdine to birds with a comparatively flat
wing, in which the first primary is almost obsolete, whilst the second ©
is lengthened with the other primaries, forming a long pointed
wing, adapted for the extended flight of species whose winter-home
may be thousands of miles away from their breeding-stations. This
scheme would include the Chats, Thrushes, Redstarts, Warblers,
and Accentors in the Turdine, and leave the short-winged Warblers,
such as Prinia, Cisticola, &c., to take their place along with the
Babblers and Bulbuls in the Timeliine. This latter group of birds
is characterized by having strong legs and feet, adapted for creeping
through tangled foliage, whilst their wings, instead of being flat, are
moulded to fit the body of the bird and to occupy as little room as
possible. The conformation of the wing is ill-adapted to extended
flight. The first primary is large and takes its place naturally beside
the shortened second primary, so that it no longer deserves to be
called a bastard primary. The other primaries are also short and
graduated in length, making a short rounded wing, sufficient for birds
whose annual migrations are confined to such narrow limits that they
can often look down from the mountains where they breed onto the
plain or into the valleys, where they find an abundant supply of
winter food.
It must be admitted, however, that in many genera of the Tur-
dine we find approaches to the Timeliine, so as to make the two
subfamilies not only to come into contact with each other but some-
times to overlap, so that we may have occasionally a turdine species
of the Timeliine more turdine than the most timeliine species of the
Turdine. These little difficulties are very puzzling to the systematic
ornithologist ; but possibly they may be evidence that his system is
a natural one rather than otherwise.
The following list of skins examined will show how impossible it
is to draw any distinction in respect of size between the four reputed
species which I propose to unite :—
Proc. Zoou. Soc.—1878, No. LXIV. 64
982 MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. [ Dec. 3
Length
of
wing.
in.
2:28 Sikkim. G. Austen. Horornis fortipes.
2°24 Naga hills. a Neornis assimilis¢.
2:2 Sikkim. Brooks. Horornis fortipes.
2:2 Darjeeling. Hume. Horeites brunnescens.
2:2 Nepal. Hodgson. Neornis assimilis.
2:2 Naga hills. G. Austen. =
2:18 Shillong. 3 ¥
2:18 Nepal. Hodgson, .
2°15 Darjeeling. Brooks. Horeites pallidus.
2:15 Khasa. G. Austen. Neornis assimilis.
2:14 Sikkim. : Horornis fortipes.
2-14 Shillong. o Neornis assimilis ¢.
2:14 Cashmere. Brooks. Horeites pallidus.
2:13 Shillong. G. Austen. Neornis assinilis ¢.
2°13 Darjeeling. Blyth. Drymeca brevicaudata.
2-11 Nepal. Hodgson. | Neornis assimilis.
2-11 ” ” %
21 ” ” ”
271 ” ”? ”
2:1 Shillong. | G. Austen. + d.
2:07 Naga hills. Fe “i 2.
2-05 Munipur 3 i
2:0 Formosa. Swinhoe. Horeites robustipes.
2:0 Nepal Hodgson. Neornis assimilis, juv.
196 ” 22 ”
1:95 sf a Neornis assimilis.
1:95 Khasi. G. Austen. 2
19 Formosa. Swinhoe. Horeites robustipes.
19 Darjeeling. Brooks. Horeites pallidus.
ig Nepal. Hodgson, Neornis assimilis, uy.
1:88 ” ” ”
1-88 ” ” ”
1:81 ” ” ”
4. Description of new Species of Phytophagous Coleoptera
from Central and South America. By Martin Jacosy.
[Received November 13, 1878. ]
1, LEMA SUFFRIANI, Sp. nov.
Oblong, light testaceous. Head and breast black ; elytra striate-
punctate, testaceous, a short sutural and submarginal stripe at the
base and two small spots, one before, the other behind the middle of
each elytron, violaceous black.
Length 23-33 lines.
Head shining black, impunctate, not constricted behind the eyes,
the lateral grooves very deep; antenne nearly as long as half the
body, of a uniform light rufous or testaceous colour, only the basal
joint stained with piceous ; thorax scarcely longer than broad, deeply
1$78.] MR. M. JACOBY ON PHYTOPBRAGOUS COLEOPTERA. 983
constricted behind the middle, smooth, impunctate, testaceous ; scu-
tellum subquadrate, black. Elytra strongly and very regularly
punctate-striate, the ninth stria entire, the interstices slightly raised
towards the apex; a short longitudinal sutural stripe at the base,
another of equal length covering the shoulder and pointed posteriorly,
where both stripes finish a little before the middle, a small round
spot between these stripes at their end, and another spot at a little
distance from the apex of each elytron, shining violaceous black.
Abdomen and the legs testaceous.
Hab. Voleano of lrazu, Costa Rica.
Collected by Mr. Rogers.
This species bears a close resemblance to L. bifida, Oliv., and
L. semisepta, Lac., in coloration, but is distinguished from the
first by the elytra haviag two spots instead of one, the uniformly
coloured antenne and the smooth thorax, while the same differences
distinguish it from the latter species. I have seen about a dozen
specimens, all of which agree exactly with each other in having the
light-coloured antennz and legs and the two spots on each elytron.
2. MasrosteTHUS SALVINI, Sp. Noy.
Elongate, parallel, testaceous, the sides of the breast and the
upper part of the head shining black. Thorax rufous, nitidous.
Elytra light testaceous, a large semiquadrate patch from the base to
the middle on each, and a broad transverse fascia behind the middle,
common to both elytra, black.
Length 43 lines.
Head distinctly punctured round the eyes, the other part smooth
and shining, lower part of face flavous, upper part black; antennz
reaching to the base of the thorax, testaceous, the first to the eighth
joint above stained with black, but diminishing towards the apex,
and entirely disappearing from the ninth joint. Thorax narrowed
from the base to the apex; surface very minutely and widely punc-
tured, impressed near the hinder angles with a distinct fovea;
scutellum black. Elytra subdepressed above, narrowed towards the
apex, surface strongly but not very closely punctured ; pale testa-
ceous, a large spot on each elytron from the base to the middle,
narrowed towards the suture, and a transverse fascia (neither of
which extends quite to the lateral margin), the latter only separated
from the former by a narrow interval, with the posterior margin
irregular, black ; the apex of the elytra are of the ground-colour to
about the same extent as the width of the transverse band. The
underside, with the exception of the sides of the breast, the dorsal
surface of the four anterior femora, and all the tibia, which are
black, testaceous ; hinder femora and inner side of tibize of the same
colour, the former with a distinct tooth near the apex,
Hab. Cache, Costa Rica.
Collected by Mr. Rogers.
3. LAMPROSOMA CHAPUISI, sp. nov.
Ovate, convex, dark violaceous ; head, lateral and anterior margins
64*
684 MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Dec. 3,
of thorax, apex of the elytra, outside of legs brilliant aureous or
copper-coloured. Head distinctly punctured.
Length 2 lines.
Head finely granulose, rather deeply but not closely punctured,
bright aureous ; first and second joint of the antenne of the latter
colour, the rest black. Thorax with its sides straight, anteriorly
much deflexed, the surface very convex and obtusely punctured,
much less distinctly visible than the punctures of the head, of an
almost black violaceous, with the anterior margin very narrowly and
the sides to a greater extent, but diminishing in width towards the
base, ornamented with bright metallic reddish-golden colour; the
inner margin of this colour at the sides of the thorax is deeply den-
tate. Elytra very convex near their base, distinctly narrowed
towards the apex ; each elytron with ten rows of rather coarse and
very regularly placed punctures, which scarcely diminish in size
towards the apex, the interstices smooth and impunctate ; the colour
is of the same dark violaceous as that of the thorax ; and at the apex
of each elytron is an elongate-oval patch of the same metallic colour
as the head and the thorax. Underside black, the outer sides of all
the legs metallic aureous ; tarsi black.
Hab. Costa Rica.
In my collection.
This species is very closely allied to L. ignicaudatum, Lac., and
L. coruscum, Lac.; but differs from the former by the deeply
punctured head, and the shape of the golden lateral band of the
thorax, while it is distinguished from the latter species by the shape
of the elytral spot, the colour of the thorax, and the colour of the legs.
4, LAMPROSOMA INORNATUM, Sp. lv.
Ovate-globose, black, shining, above brilliant violaceous blue.
Head finely, thorax distinctly punctate; elytra striate-punctate ;
prosternum smooth,
Length 23 lines.
Head rather convex, with a small roundish fovea in the middle,
widely and finely punctured, violaceous blue, bordered anteriorly
with metallic green ; antennz black, the second joint fulvous. Thorax
transverse, the anterior angles much deflexed, the sides rounded ;
posterior margin sinuate on each side, with an obsolete short de-
pression on each side of the median lobe; the surface much more
distinctly punctured than the head, and increasing in size towards
the base. Elytra very convex, their greatest convexity lying very
near the middle viewed sideways, they are gradually narrowed towards
the apex ; and each elytron has ten rows of rather deep punctures,
which diminish slightly towards the apex in size; the colour is a
brilliant violaceous blue with a greenish tint. Underside and legs
black, shining, finely alutaceous and punctured; prosternum elongate,
quadrate, perfectly smooth, with a few rows of short whitish hairs.
Hab. Chiriqui.
In my collection.
Although this species is nearly allied to Z. testudineum, Lac., as
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 985
well as to L. cyaneum, Lac., it is from both distinguished by its smooth
prosternum, from the former by the punctured head, from the latter
by the want of the depression at the head.
5. NODA LEFEVREI, sp. nov.
Broadly ovate, uniformly greenish wneous. Head coarsely,
thorax less strongly punctured. Elytra semipunctate-striate, costate
near the apex.
Length 12 line.
Head strongly and closely punctate, especially between the eyes,
where the punctures are of an oblong shape. Eyes very prominent ;
clypeus also strongly punctured, separated from the face, its anterior
margin crescent-shaped ; antenne as long as half the body, the last
five joints much thickened, and the terminal one very robust and
strongly pointed at the apex, the lower six joints shining greenish
geneous above, fulvous below, the rest black and opaque. ‘Thorax
very narrowed towards the apex, with the posterior angles acutely
produced ; surface moderately punctured, the punctures not stronger
near the sides than those on the disk, and not extending to the an-
terior or the posterior margin. Scutellum with a few punctures.
Elytra convex, with a very obsolete depression below the base ;
surface very deeply and regularly punctate-striate near the outer
margin, finer and more irregularly towards the suture ; the intervals
towards the apex raised in longitudinal costs, of which the third
and fourth (counting from the outer margin) are united posteriorly
sending off a short costa towards the apex. Underside and legs
geneous, extreme apex of the tibize and the tarsi darkish rufous.
Hab. Costa Rica (volcano of Irazn) ; Guatemala.
Closely allied to Noda semicostata, Lefévre ; but distinguished by
the equal punctuation of the thorax, which, in the former species, is,
according to the description, strong and dense at the sides.
6. Nopa BALYI, sp. nov.
Broadly ovate, eneous, shining ; antenna, palpi, and legs ferru-
ginous. Elytra punctate-striate, with a short round tubercle on
the disk of each elytron.
Length 13 line.
Head rather deeply but widely punctured, the clypeus not sepa-
rated from the face ; antenne with the third, fourth, and fifth joints
of equal length, entirely ferruginous. Thorax of the same shape as in
N. viridis, distinctly punctate throughout, greenish bronze-coloured.
Elytra convex, a little more strongly punctured than the thorax, the
punctures arranged in regular striw, the intervals scarcely raised
except near the lateral margin and the apex, where they are semicos-
tate; the humeral callus prominent ; and each elytron is furnished at
a little distance before the middle with a short round tubercle, which is
placed nearer the lateral margin than the suture ; a very shallow trans-
verse depression is also visible in certain lights immediately above
this tubercle. Underside greenish black, opaque ; legs ferruginous.
Hab. Guatemala.
986 MR. M.JACOBY ON PHYTOPHAGOUS COLEOPTERA. [ Dec. 3, |
Collected by Mr. Salvin. Collection Jacoby.
Allied to N. propinqua, Lefévre,. but differing in the stronger
punctuation of the thorax and the sculpture of the elytra.
7. NopA vVIRIDIS, sp- Nov.
Broadly ovate, shining metallic-green above, below greenish black,
the first six or seven joints of the antennz and the legs entirely fer-
ruginous.
Length 14-13 line.
Head coarsely but widely punctate, the clypeus distinctly sepa-
rated, also strongly punctured, with the anterior margin emarginate ;
Jabrum dark ferruginous or brown; vertex of head with an obsolete
longitudinal groove. Antenne of half the length of the body, very
robust, with the exception of the short second joiut, gradually
thickened towards the apex, the three penultimate joints of nearly
equal length and cylindrical, the terminal joint the longest of any,
and pointed at the apex ; the first six or seven joints ferruginous,
the rest black, opaque. Thorax transverse, the anterior margin
nearly straight, the posterior one evenly rounded and produced in
the middle, the sides nearly parallel at the base, but narrowed
towards the apex, all the angles acute; surface shining metallic
green with a slight bluish tint, not very closely but rather deeply
punctate. Scutellam smooth. Elytra slightly wider at the base
than the thorax, very gradually and evenly rounded towards the apex ;
disk rather convex, without any basal depression ; the humeral callus
produced and smooth ; surface finely punctured a little more strongly
near the base, the punctures arranged in rather regular lines, partially
running in pairs, and forming one stria near the suture towards the
apex. Underside opaque blackish green. Legs and tarsi ferruginous.
Hab. Guatemala.
Collected by Mr. O. Salvin. In my collection.
8. NopA BOUCARDI, sp. nov.
Broadly ovate, above violaceous blue, shining, below greenish
black. Tibi, tarsi, and base of the antenne rufous. Elytra
geminate-punctate-striate.
Length 13 line.
Tiead strongly punctured, especially towards the lower part,
shining greenish blue ; antennz with the first six joints light rufous,
the rest darker. Thorax of the same shape as WN. /efevrei of this
paper, also equally punctate throughout, the puncturing extending
at the same time to all the margins. Elytra not differmg in form
from the above species, the punctuation on the disk, however, united
in pairs, and only the extreme apex distinctly costate. Underside
semiopaque, greenish black, all the tibize and tarsi bright rufous.
Hab. Valin, Guatemala.
Collected by Mr. Salvin.
Allied to Noda rufipes, Lefév. (Mittheil. Minch. Ent. v. 1878),
but differing in the equal punctuation of the thorax, the double rows
of punctures of the elytra, and the costate apex of the latter.
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 987
9. NODA VIOLACEIPENNIS, sp. nov.
Oblong-ovate, blackish green below, violaceous blue above ; thorax
moderately punctured, elytra semi-punctate-striate, without coste.
Legs and antenne greenish eneous.
Length 2 lines.
Head and clypeus with some widely dispersed deep punctures,
the former besides with an obsolete fovea in the middle; antennze
more than half the length of the body, robust, uniformly greenish
zneous, with the exception of the two basal joints, which are stained
with piceous. Thorax of the same shape as the other species of this
paper, but rather finely punctured, the punctures on the disk
assuming an oblong shape here and there, while the punctuation
near the lateral margins is very obsolete. Elytra more oblong than
in the other species, shining violaceous blue, with the punctures
stronger than those of the thorax, and rather confusedly placed, only
towards the apex more distinctly geminate-punctate-striate ; the
intervals not raised, only an obtuse tubercle visible near the extreme
apex; the suture on each side accompanied by an impressed line,
commencing about the middle of the elytra, to the apex. Underside
and legs greenish black. Tarsi and claws obscure piceous.
Hab, Guatemala.
My collection. Collected by Mr. Salvin.
10. MeraxyONYCHA RUFOLIMBATA, Sp. nov.
Oblong, broadly ovate. Head aureous, deeply and closely punc-
tured. Thorax laterally obsoletely tridentate, greenish, coarsely
punctured. Elytra punctate-striate, the interstices longitudinally
costate, greenish; the sides and the apex broadly rufous-coloured,
abdomen flavous.
Length 4% lines.
Head deeply and closely punctured, with a deep longitudinal
middle groove, bright aureous or copper-coloured ; antennex black,
the first six joints shining. Thorax transverse, distinctly triden-
tate at the sides, the disk covered with rather deep punctures in
the middle and along the anterior margin; the rest of the surface
minutely punctate, of a greenish opaque colour. Scutellum elongate,
smooth, purplish. Elytra not much wider at the base than the
thorax, rather finely punctate-striate, the interstices broadly costate
from the base to the apex, where their ends are joined, of the same
green opaque colour as the thorax, the sides (narrowly at the base,
gradually widened to the apex, the latter broadly) of a rufous colour.
Breast and legs purplish green, abdomen light rufous or flavous.
Hab. Venezuela.
Collection Jacoby.
11. PRIONODERA ELEGANS, sp. nov.
Elongate, dark purplish. Head and thorax irregularly punctured
and finely transversely wrinkled, the latter at the sides obsoletely
bidenticulate. Elytra longitudinally costate, the intervals punctate-
988 MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. | Dee. 3,
striate, purplish blue, a large triangular sutural patch from the base
to the middle reddish copper-coloured.
Length 43 lines.
Head and clypeus deeply punctured, with a deep longitudinal
groove in the middle, of a greenish semiopaque colour, here and
there with a purplish tint ; labrum and jaws nearly black; first
joint of the antennee very thick and subcylindrical, the second short
and globose, the rest cylindrical, equal in length ; the first six joints
shining greenish aureous, the rest black, opaque. Thorax sub-
quadrate, slightly broader than long, all the angles acute and pro-
duced externally, the sides with a distinct tooth behind and another
very obsolete one before the middle; disk irregularly transversely
wrinkled, visible only when viewed sideways, and covered with large
and small punctures ; of a greenish colour, sometimes with purplish
patches. Scutellum elongate, finely punctate. Elytra much wider
at the base than the thorax, a little widened posteriorly ; each elytron
with eight longitudinal cost, the intervals subgeminate-punctate-
striate; bluish purple, a large sutural triangular patch, extending
from the base to below the middle, of a reddish copper, surrounded
by light green colour. Underside and legs dark green, with a
purplish hue.
Hab. Columbia.
Two specimens in my collection.
12. CHALCOPHANA SEMIRUFA, Sp. nov.
Elongate, subparallel, piceous below. Head, thorax, first six
joints of the antennz, and thelegs rufous. Elytra deeply and closely
punctate-striate and partly costate, shining bronze-colour, extreme
margin and the apex rufous.
Length 33 lines.
Head deeply punctured, with a median groove, labrum yellowish,
apex of mandibles black ; antennz with the first six joints rufous,
the rest black. Thorax transverse, the posterior margin round and
widened in the middle, the anterior one straight, all the angles pro-
duced into a short acute tooth, the surface rather coarsely punctate
towards the sides, less so on the disk. Scutellum rufous. Elytra
with a more or less distinct oblique depression below the base, closely
punctate-striate, the punctures not united in pairs, and regularly
placed only towards the suture, the rest confused and irregular ; the
apex as well as the’sides of each elytron furnished with three or four
more or less distinct cost, those at the sides commencing below the
humeral callus; the surface of a bronze or metallic green colour,
the extreme lateral margins and the apex rufous. Underside dark
piceous, as well as the tarsi.
Hab. Costa Rica.
Of the six or seven similarly-coloured described species, the pre-
sent one will be easily recognized by the colour of the first six joints
. of the antenne, which is constant in all the specimens before
rte as well as by the strong punctuation of the thorax and the
elytra,
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 989
13. CHALCOPHANA UNIFORMIS, sp. Nov.
Entirely dark rufous, with the exception of the last five joints of
the antenne, the apex of the tibiw, and the tarsi. Thorax finely,
elytra deeply geminate-punctate-striate, and more or less costate.
Length 3-3$ lines.
Head exactly punctured as in semirufa, the antenne also coloured as
in that species. Thorax a little more finely punctate. Elytra distinctly
and closely punctured, the punctures running together in pairs, the
interstices either raised over the whole disk in costz, or only distinctly
visible towards the apex. The underside rufous; extreme apex of
tibiz and the tarsi black.
Hab. Costa Rica.
This species may possibly be only a variety of C. semirufa ; but
the uniform coloration, the punctuation of the elytra approached in
pairs distinguish it sufficiently from that species, while the colour of
the first six joints of the antenne will separate it from species in
other respects similarly coloured.
14. DoRYPHORA BRUNNEIPENNIS, Sp. noy.
Broadly ovate, narrowed behind, below black ; antenne with the
last two joints yellowish white. Head, thorax, and elytra rufous,
opaque.
Length 6 lines.
Head strongly but not closely punctate, brown with a greenish
tint ; antennze brownish zneous, last two joints whitish. Thorax
with the sides parallel, the anterior angles acute; surface with the
same-sized punctures as the head, but closely and uniformly dis-
tributed. Scutellum smooth, broadly triangular. Elytra broader at
the base than the thorax, very convex at their first third, from there
strongly declining to the apex, the latter pointed ; surface finely and
irregularly punctured and longitudinally aciculate, the intervals finel
alutaceous, opaque, of a uniformly rufous colour. Beneath black ;
mesosternal process robust, long, slightly curved. Legs rufous.
Hab. Columbia.
Collection Jacoby.
15. DoRYPHORA FLAVIPENNIS, sp. nov.
Greenish black, shining. Head and thorax strongly punctate.
Elytra sparingly irregularly punctured, flavous, a sutural stripe
narrowed towards the apex, greenish black.
Length 53 lines.
Head more strongly punctured near the apex than at the vertex ;
antennze with the last five joints flattened and widened, the first five
joints testaceous, stained with greenish zeneous above, the rest
fuscous. Thorax more strongly punctured than the head, very
coarsely towards the sides, dispersedly on the disk. Scutellum
smooth. Elytra rotundate towards the apex, parallel near the base,
sparingly and rather finely punctate without order, the intervals
finely aciculate ; the extreme outer margin has a row of punctures,
and is accompanied, at some distance, with another row of double
990 MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Dec. 3,
punctures ; the suture has also at its posterior part a deeply im-
pressed line running parallel with it at each side. The elytra are
of a light flavous, shining, the entire suture greenish or bluish
black, which colour is gradually narrowed towards the apex. Under-
side and legs black with a bluish tint.
Hab. Columbia.
In colour this species approaches close to D. limbatipennis, Stal ;
but the want of the outer marginal vitte, together with the roundish
shape, will distinguish the species before us from the above insect.
16. DoryPHORA VITTATIPENNIS, Sp. nov.
Black. Head and thorax opaque. Elytra strongly semipunctate-
striate, black, shining ; each elytron with five narrow longitudinal
flavous stripes.
Length 6 lines.
Head finely punctate ; antennz black, shining. Thorax deeply
and irregular punctured, the intervals partly convex; sides narrowed
at the base, from there rounded to the apex. Scutellum smooth,
triangular, black. Elytra parallel, convex, deeply punctate, striate
near the suture, more irregularly towards the sides ; black, shining,
each elytron with five narrow flavous vittee at equal distances fron
each other, of which the outer one is the broadest, all of which are
confluent at the apex and extending to the base. Underside and legs
black.
Var. Elytra with only the outer vittze entire, the sutural one only
visible at the posterior third, the other stripes entirely absent or
only partially visible near the apex.
Hab. Columbia.
Collected by Mr. Salmon.
17. DoryPnuORA POROSA, Sp. Nov.
Broadly ovate, very convex, piceous below. Head aud disk of
thorax black; sides of latter testaceous. Elytra deeply foveolate,
the intervals irregular transversely reticulate, dark brown.
Length 7 lines.
Head opaque, with a few very minute punctures; antenna long,
scarcely thickened towards the apex, fuscous. Thorax with the
anterior and posterior margin parallel in the middle, the anterior
angles acute and much produced, the sides much flattened and
slightly thickened ; disk minutely punctate, of the same colour as
the head, the sides light testaceous, impunctate. Scutellum
broad, smooth, piceous. Elytra convex at their first third, thence
suddenly declining and subdepressed towards the apex, uniformly
dark brown; the suture shining, piceous, smooth ; surface deeply
punctate, striate near the suture, foveolate towards the sides, the
intervals raised and connected partially by transverse short costz,
forming a kind of network towards the sides. Femora greenish
zneous ; tibize piceous.
Hab. West Columbia.
Collection Jacoby.
1878.| MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 991
18. DoryPHORA HYBRIDA, Sp. Ov.
Ovate, black below. Head and thorax greenish black, distinctly
punctured. Elytra geminate punctate-striate, testaceous; the
suture, the outer margin, and two transversely placed large irre-
gularly shaped spots, one placed before, the other behind the middle,
as well as the base narrowly, greenish black.
Length 8 lines.
Antenne rufous. Thorax not very strongly or closely punctured ;
the intervals also minutely punctate. Scutellum black. Elytra
convex, geminate-punctate-striate, each puncture surrounded by a
piceous spot ; the extreme base, a spot on the shoulder, the outer
margin, the suture (which is widened below the base into a square-
shaped spot), and a transverse patch, consisting of two united spots
before the middle as well as a similar shaped one-(formed by three
united spots) behind the middle, greenish black. Underside and
legs black.
Hab. Peru.
Collection: Jacoby.
The insect before me is so exactly similar in markings to the
Doryphora imperialis of Stal, that it may be only a variety of that
species. The colour, however, instead of being rufous, is greenish
black. The elytra, besides having the sutural stripe broader, are
also marked with regular rows of small piceous spots which surround
the puncturing everywhere ; and the underside is shining black.
19. DoryPHORA MILITARIS, Sp. nov.
Greenish neous, shining. Elytra closely punctate-striate, four
roundish spots placed transversely at the base, two others behind
the middle, with two submarginal longitudinal vittee from the middle
to the apex, light flavous, margined with rufous.
Length 6 lines.
Head closely punctured, dark brownish green, labrum fulvous ;
antennze gradually thickened towards the apex, the basal joints
shining fulvous, the rest greenish e#neous. Thorax closely and
deeply punctured towards the sides, much more sparingly punctate
on the disk, the intervals also minutely punctured. Scutellum
smooth. Elytra moderately convex, closely and rather regularly
punctate-striate, the punctures diminishing towards the apex; of a
greenish geneous colour; each elytron with two spots near the base,
of which one is placed on the shoulder, another near the scutellum,
an elongate spot behind the middle near the suture, and a longitu-
dinal stripe near the outer margin, from immediately behind the
middle to the apex, flavous, all the markings surrounded by a
narrow rufous border. Underside shining greenish black ; legs more
brownish ; mesosternal process slightly curved and pointed.
Hab. Columbia.
My collection.
20. DorypHORA OCELLATA, sp. nov.
Ovate, very convex, testaceous. Thorax with four piceous spots.
992 MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Dec. 3,
Elytra geminate-punctate-striate, fuscous; two large round spots
near the scutellum, four others placed transversely, and the apex
broadly flavous, the latter with a small fuscous spot.
Length 5 lines.
Head minutely punctured, flat, with an indistinct piceous mark
at the vertex ; antennee testaceous, slightly thickened towards their
apex, terminal joint scarcely longer than broad. Thorax narrow,
sides rounded in front, anterior angles scarcely produced ; above
much more strongly (but not closely) punctured than the head, of
a rather indistinct testaceous colour ; four spots placed transversely
on the disk, the middle ones, which are in shape of two short
longitudinal vittee, of a greenish piceous colour. Scutellum testa-
ceous. Elytra slightly broader at the base than the thorax, very
convex on the disk, their apex regularly rounded; the surface
rather finely punctate-striate, consisting of double rows of punctures,
which are more deeply impressed on the disk than at the base or
the apex ; the flavous patches on the disk of each elytron arranged
as follows, viz. one in the middle of the base, two transversely, of
which the outer larger one is placed immediately below the shoulder,
the inner one on the middle near the suture, the fourth occupying
the apex to nearly a third of the length of the elytra, having its
inner margin sinuate, and in its interior a small fuscous square spot.
Underside and legs testaceous. Mesosternal spine rather short,
slightly curved. ;
Hab. Costa Rica?
My collection.
Closely approaching in colouring to D. estuans, Lin., the species
described here is distinguished by its much larger size and by the
four spots on the thorax.
21. DoryPHORA DECORATA, Sp. nov.
Ovate, convex, testaceous. Head with the base, thorax with
two vitte black. Elytra geminate-punctate-striate, testaceous, with
three transverse metallic green fasciee and a longitudinal vitta between
the second and third bands.
Length 53 lines.
Head minutely punctate, with a shallow transverse depression and
an obscure longitudinal groove, testaceous, with an irregular trans-
verse bifurcate patch on the vertex ; antennz longer than the thorax,
gradually thickened towards the apex, as well as the palpi, testaceous;
apex of mandibule black. Thorax with a longitudinal depression
parallel with the lateral margins, the latter regularly rounded ;
surface very sparingly but rather deeply punctate, shining testaceous,
with two longitudinal black vittee from the apex to the base, slightly
curved outwards, and connected at the base by a very narrow band.
Scutellum impunctate, testaceous. Elytra regularly geminaie-punctate-
striate near the suture, more confusedly towards the sides ; shining
testaceous, the suture, an obliquely placed transverse band from the
shoulder to the suture a little below the seutellum (with its posterior
margin denticulate and connected by a narrow stripe at the lateral
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 993
margin anteriorly with another transverse band of a 4-shaped form,
and placed in the middle of the elytra), as well as a short rounded
band near the apex (this latter band being also connected with the
former one bya short vitta near the lateral margin), shining metallic
green; the inflexed margin of the elytra, as well as the underside
and the legs, testaceous.
Hab. Costa Rica?
Collection Jacoby.
Allied to D. vittaticollis, Stal, but differing in the colour of the
head, antennz, and palpi, and the longitudinal stripe between the
second and third bands.
22. SCELIDA BALYI, Sp. nov,
Elongate, parallel, testaceous; vertex of head, last six joints of
antennz, and the coxze black. Elytra metallic green, closely rugose-
punctate.
Length 43 lines.
Head impunctate, with two distinct tubercles between the an-
tenn, limited behind by a transverse groove, lower part of face
with a raised triangular ridge ; last joint of maxillary palpi piceous ;
vertex of head black, shining ; lower part and the basal five joints
of the antennz flavous, the latter about three fourths the length of
the body, with the three terminal joints slightly curved. Eyes pro-
minent, black. Thorax subquadrate, widened from the apex towards
the middle, from there straight to the base, the anterior and
posterior margins slightly sinuate in middle; surface entirely im-
punctate, the disk depressed, flavous, with indistinct irregular
blotches of a piceous colour. Scutellum semiovate, dark piceous,
with a light-coloured margin. Elytra much wider at the base than
the thorax, sides entirely parallel, the surface covered throughout
with crowded and rather deep punctures, the intervals of which are
finely rugose, of a metallic, moderately shining green colour, which
assumes towards the apex and near the suture a purplish hue.
Under surface and legs flavous, the base of the coxee and a spot at
the base of the anterior femora piceous or black ; the metathorax
on each side raised in the shape of a triangular-pointed elevation.
This species is the second one belonging to this genus, the first
of which was published in diagnosis by Mr. Chapuis in his eleventh
volume of the ‘Genera of Coleoptera.’ From thisspecies it is sufficiently
distinguished by its smaller size, the black vertex of the head,
together with the spots on the coxze of the legs, and by the peculiar
shape of the metathorax.
23. DIABROTICA WATERHOUSEI, sp. nov.
Elongate, convex, widened behind, flavous. Head, breast, and
the extreme base of the elytra pitchy black, the apex of the latter
brownish fuscous.
Length 3 lines.
Head shining black, with a distinct fovea between the eyes; an-
tennz ferruginous, the basal joint flavous. Thorax quadrate,
994 MR.M.JACOBY ON PHYTOPHAGOUS COLEOPTERA. [ Dec. 3,
rather convex, smooth, narrowed towards the base, surface shining,
flavous. Scutellum black. Elytra widened towards the apex,
rather convex, light flavous, a narrow transverse band at the base
undulated at its posterior edge, and extending to the humeral callus,
dark fuscous or black; the posterior third of the elytra fuscous
brown, surrounded by an indistinct crescent of a darker colour; the
rest of the surface covered with minute piceous spots, visible only
under a lens. Underside black ; abdomen brownish; legs flavous,
the hinder tibize on their cuter side blackish.
Var. Thorax at the anterior margin with a triangular black spot ;
the apical markings of the elytra obsolete.
Hab. Cache, Costa Rica.
Collected by Mr. Rogers.
24. DIABROTICA JANSONI, Sp. nov.
Elongate, convex, widened behind, fulvous. Thorax and elytra
closely punctate, a large oval patch near the apex of the latter
violaceous blue.
Length 43-5 lines.
Head with a deep transverse groove, closely punctured, fulvous ;
antenne: robust, as long as half the length of the body, fuscous,
the first joint below fulvous, the fourth joint nearly double the
length of the third. Thorax transverse, sides nearly parallel, with
three indistinct shallow depressions on the disk, of which two
smaller ones are situated on each side and a large one near the
posterior margin at its middle; surface distinctly but not very
closely punctate, fulvous. Scutellum broad, triangular, its surface
very finely alutaceous, and slightly excavated near the base. Elytra
more than four times the length of the thorax, much widened
towards the apex, and distinctly margined; the surface crowded
everywhere with distinct punctures of the same colour as the head
and thorax, the whole of the posterior third occupied by a large
rounded spot of violaceous blue, which sometimes terminates at some
distance from the apex. Underside and the femora testaceous ;
tibize and tarsi fuscous.
Hab. Chontales, Nicaragua. Collected by Mr. Janson.
In my collection.
25. DIABROTICA FUSCOMACULATA, Sp. NOV.
Testaceous below. Head, thorax, and elytra olive-green, the
latter obsoletely costulate, with the base and two spots on each
elytron dark fuscous, margined with pale yellow.
Length 24 lines.
Head impunctate, with a fovea in the middle, shining green ;
labrum dark brown; antennee nearly reaching to the end of the
body, with the second and third joint very short, the fourth three
times as long as the second joint, of an indistinct brownish colour,
the first joint pale greenish, Thorax bifoveolate, impunctate, very
shining vlive-green. Scutellum flavous. Elytra subdepressed,
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 995
parallel, closely punctate, with two or three obsolete costz on the
disk, of which the outer one is more distinctly raised, of the same
colour as the thorax; a transverse band at the base narrowed towards
the suture and extending to the shoulders, a transverse spot in the
middle and another at a little distance from the apex of each elytron,
dark fuscous; all the spots, as well as the lateral margin at its
anterior part, surrounded with yellowish white. Underside pale
testaceous, the tibize darker, and the femora olive-green.
Var. The middle spot on the elytra absent, as well as the apical
one, which is replaced by a yellowish patch.
Hab. Nicaragua, Bogota. Var. Guatemala.
Collected by Mr. Salvin.
26. DIABROTICA NIGROVITTATA, Sp. NOV.
Elongate, parallel, black. Thorax bifoveolate. Elytra yellowish
white, with the suture and a submarginal longitudinal stripe on
each elytron not reaching the apex, black.
‘Length 23 lines.
Head very shining black, smooth, with a fovea above the antenne;
the latter of about half the length of the body, entirely black ;
second and third joints short, equal. ‘Thorax quadrate, narrowed
near the base, surface deeply bifoveolate, smooth, very shining black.
Scutellum black. Elytra about three times the length of the
thorax, with a longitudinal depression below the shoulders and ex-
tending a little below the middle; surface very minutely punctured
and finely granulose, of a yellowish white, a longitudinal stripe from
the base, extending toa little distance from the apex and narrowed
at each end, black. Body and the legs shining black.
One specimen from Mexico and another from Guatemala, in my
collection.
27. DiaABROTICA NOVEMMACULATA, sp. NOV.
Elongate, widened behind, flavous. Head and breast black.
Elytra finely punctate, flavous, five spots near the base and four
near the apex black.
Length 4 lines.
Head black, impunctate, with a fovea and two flat tubercles in
front of the antennz ; the latter more than half the length of the
body, with the third joint a little longer than the second, and the
fourth the longest, of a flavous or ferruginous colour, with the
middle joints stained more or less with fuscous. Thorax impunctate,
with two shallow depressions on its surface; sides much deflexed
and margined, parallel near the base, widened above the middle.
Scutellum fuscous. Elytra but little wider at the base than the
thorax, gradually widened towards the apex, closely and irregularly
punctured, flavous, a triangular spot extending a little below the
base at the suture and surrounding the scutellum, an elongate
short stripe on each shoulder, a roundish spot between the latter
and the sutural one on each elytron, as well as four spots placed
996 MR.M.JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Dec. 3.
transversely at a little distance from the apex, of which the outer
one is elongate, black. Underside and legs flavous, breast black.
Hab. Costa Rica.
Var. The round spots on the elytra replaced by short longitudinal
stripes.
28. DiaBROTICA COSTATIPENNIS, Sp. nov.
Elongate, uniform rufo-flavous. Thorax deeply bifoveolate, shining.
Elytra closely rugose, punctate, with five or six elevated more or
less distinct longitudinal coste.
Length 3 lines.
Head perfectly smooth and shining, with a deep fovea above the
antennee ; the latter of half the length of the body, the second and
third joints very short, the fourth joint more than twice as long as
the preceding one, ferruginous, the first three joints shining, the
rest opaque. ‘Thorax distinctly margined, narrowed at the base, the
surface with two deep fovez, very shining and impunctate, rufo-
flavous. Elytra scarcely widened behind, with a distinct margin °
which is wider in the middle than on either end; surface closely
punctate and finely rugose; each elytron with about six longitudinal
raised cost, which are a little more obsolete near the apex and the
base, of a little lighter rufous colour than the thorax. Entire
underside and legs flavuus, the breast covered with whitish
hairs.
Hab. Costa Rica.
29. CEROTOMA NIGROFASCIATA.
Oblong, widened behind, black. Thorax and legs flavous. Elytra
dark red, with three broad transverse black bands.
Length 33 lines.
Head black, vertex punctate; lower part of face coarsely and
deeply punctate, with a few flavous hairs ; antennze closely approximate,
of half the length of the body, with the first joint very long, the
second short, the third longer than the fourth, of a flavous or fer-
ruginous colour. Thorax transverse, the sides much deflexed, the
lateral margins much widened before the middle, anterior angles
obsolete, posterior ones produced but rounded; the surface rather
convex, smooth, indistinctly depressed on the disk, of a light flavous
colour. Scutellum black. Elytra much widened towards the apex,
deeply and closely punctate, the intervals raised in irregular longi-
tudinal cost and transverse wrinkles, of a dark red colour ; a trans-
verse band common to both elytra at the base, another band slightly
arched in the middle not reaching the suture, and a large triangular
patch near the apex black ; none of these bands reach quite to the
lateral margins. Underside black, shining ; legs uniformly flavous.
Hab. Costa Rica, Guatemala.
APPENDIX.
LIST OF ADDITIONS TO THE SOCIETY’S MENAGERIE
DURING THE YEAR
1878.
Jan. 1. 1 Japanese Wild Dog (Canis familiaris, var.), 3. Presented by
Harry Pryer, Esq. See P. Z. S. 1878, p. 115.
1 Bristly Ground-Squirrel (Xerus setosus), ¢. Purchased.
1 Robben-Island Snake (Coronella phocarum). Presented by
Messrs. Rice and A. Jamrach.
. 1 Macaque Monkey (Macacus cynomolgus), 3. Presented by
J. F. Wood, Esq.
1 See Hyena (Hyena striata). Presented by Captain F.
otton.
1 Red-and-Yellow Macaw (Ara chloroptera). Presented by
Captain King. From Carthagena.
1 Common Sole (Solea vulgaris). Presented by G. H. King,
Esq.
. 1 Mexican Deer (Cervus mexicanus). Presented by Cyril Graham,
Esq. From Venezuela.
. 1 White-handed Gibbon (Hylobates lar), 9. Presented by
W. H. Newman, Esq. From the forests of Laos, Northern
co
STi Or
Siam.
1 Brown Monkey (Macacus arctoides), $. Presented by W. H.
Newman, Esq.
9. 2 Black Francolins (Francolinus vulgaris). Presented by Major
Newton Pauli.
1 Chukar Partridge (Caccabis chukar). Presented by Major
Newton Pauli.
1 Grooved Tortoise ( Testudo sulcata). Purchased.
2 Rough Terrapins (Clemmys punctularia). Purchased.
1 Scorpion Mud-Tortoise (Cinusternon scorpioides). Purchased.
10. 4 Common Marmosets (Hapale jacchus). Deposited.
14, 1 Arabian Gazelle (Gazella arabica), §. Presented by Mark
Whyley, Esq.
2 Violet Tanagers (Huphonia violacea), Purchased.
1 Superb Tanager ( Calliste fastuosa). Purchased.
1 Taherculated Lizard (Iguana tuberculata). Purchased.
3 Summer Ducks (Atv sponsa),3 2. Presented by the Lord
Braybrooke, F.Z.S.
15. 1 Stanleyan Chevrotain (Zragulus stanleyanus), 3. Presented
by Mrs. Leslie Walker.
1 Javan Chevrotain (Zragulus javanicus). d. Presented by
Mrs. Leslie Walker.
Proc. Zoou. Soc.—1878, No. LXV. 65
31.
Feb. 4.
APPENDIX.
. 2 Mandarin Ducks (Aix galericulata), § and 9. Purchased,
2 White-bellied Storks (Abdimia sphenorhyncha). Purchased.
. 1 Broad-banded Armadillo (Xenwrus unicinctus). Purchased.
. 1 Grivet Monkey (Cercopithecus grisev-viridis), Q. Presented
by Madame Patey.
. 2 Black Swans (Cygnus atratus). Presented by Captain W. H.
Kecles.
. 1 Common Fox (Canis vulpes). Presented by Mr. George
Fredericks.
2 West-India Rails (Aramides cayennensis). Purchased.
. 1 Hog Deer ( Cervus porcinus). Born in the Menagerie.
. 1 Upland Goose (Bernicla magellanica), 2. Purchased. From
hili.
1 Humboldt’s Penguin (Spheniscus humboldti). Purchased. See
P.Z. 8. 1878, p. 116.
5. 1 Wood-Owl (Syrnium aluco). Presented by J. E. Liardet,
Esq.
1 Common Magpie (Pica caudata). Presented by G. E. Lad-
bury, Esq.
1 Jackdaw (Corvus monedula). Presented by G. E. Ladbury,
Esq.
. 1 Hoary Snake (Coronella cana). Presented by the Rev.G. H.R.
Fisk, C.M.Z.S.
1 Derbian Opossum (Didelphys derbiana). Purchased.
. 1 Common Puffin (Fratercula arctica), Received in exchange.
From the Banks of Newfoundland.
. 1 Brown Bear (Ursus arctos). Presented by J. Mason Allen,
Esq.
1 Mieaqae Monkey (Macacus cynomolgus), 2. Presented by
L. E. Lewis, Esq.
1 White-cheeked Capuchin (Cebus lunatus). Presented by John
Ikin, Esq.
. L Yaguarundi Cat (Felis yaguarundi). Purchased.
2 Yarrell’s Curassows (Crav carunculata), ¢ and 2. Pur-
chased.
1 Lesser Razor-billed Curassow (Mitua tomentosa). Purchased.
2 White-bellied Guans (Ortalida albiventris). Purchased.
1 White-fronted Guan (Penelope jacucaca). Purchased.
1 Common Trumpeter (Psophia crepitans). Purchased.
1 Guira Cuckoo (Guira pirtrigua). Purchased.
1 Sun-Bittern (Zwrypyga hehas). Purchased.
1 Cayenne Lapwing (Vanellus cayennensis). Purchased.
1 Double-striped Thicknee (Ctdicnemus bistriatus). Purchased.
1 Garden’s Night-Heron (Wycticorax gardenr). Purchased.
1 Maguari Stork (Ciconia maguarz). Purchased.
1 Black Vulture (Cathartes atratus). Purchased.
2 Silky Cowbirds (Molothrus bonariensis). Purchased.
1 American Kestrel (Tinnunculus sparverius). Purchased.
3 Chilian Pintails (Dajila spinicauda). Purchased.
1 Common Badger (Meles taxus). Presented by the Lord Sal-
toun, F.Z.S.
2 Lesser Black-backed Gulls (Larus fuscus). Presented by the
Rey. 8. R. Wilkinson, F.Z.8.
7 Bankiva Jungle-fowls (Gallus bankiva), Presented by Mrs.
Kendall.
1 Azara’s Fox (Canis azare). Purchased.
Feb, 5,
28.
Mar. 1.
ADDITIONS TO THE MENAGERIE, 999
1 Brazilian Tortoise (Testudo tabulata), Presented by Captain
ing.
. 4 Reeves’s Terrapins (Clemmys reevesi). Presented by Mr. A.
Thomson.
» 1 Common Swan (Cygnus olor). Presented by John Colam,
Esq., F.Z.8,
. 2 Crested Guinea-fowls (Numida cristata), Presented by Mrs.
Collingwood.
2 Canadian Geese (Bernicla canadensis). Presented by Edw, J.
Philpot, Esq.
- 1 Wood-Owl (Syrnium aluco). Presented by J. Torpey, Esq.
1 Hey’s Partridge (Caccabis heyi). Purchased.
» 2 Macaque Monkeys (Macacus cynomolgus). Presented by
Lieut.-Colonel Feilden.
» 2 Chimpanzees (Troglodytes niger), § and 2. Deposited.
1 Black-footed Penguin (Spheniscus demersus). Purchased.
1 Garden’s Night-Heron (Nycticorax gardeni). Presented by
Henry Bottrell, Esq.
- 1 Grivet Monkey (Cercopithecus griseo-viridis), Presented by
E. H. Lockley, Esq.
- 1 Collared Peccary (Dicotyles tajagu). Purchased.
1 Black-faced Spider Monkey (Ateles ater), 9. Purchased.
1 Globose Curassow (Crax globicera), $.° Purchased.
- 1 Chimpanzee (Troglodytes niger). 3. Deposited.
. 2 Leopards (Fevis pardus). “Purchased. From Persia. See
P. Z. S, 1878, p. 289.
. 2 Red-vented Bulbuls (Pycnonotus hemorrhous). Presented by
Colonel A. L. Annesley, F.Z.S.
- 1 Black-billed Parrakeet (Palzornis melanorhyncha). Presented
by Mrs. Barthorp. From Muttra, N.W. India.
2 Black-necked Swans (Cygnus nigricollis). Received in ex-
change.
. 2 Tigers (Felis tigris). Born in the Menagerie.
2 Barbary Wild Sheep (Ovis tragelaphus), 3 and 2. Received
in exchange.
. 2 Black-winged Peafowls (Pavo migripennis), 2. Presented
by the Hon. A. 8. G, Canning, F.Z.S.
2 Pale-headed Parrakeets (Platycercus pallidiceps). Purchased.
4 Turquoisine Parrakeets (Euphema pulchella). Purchased.
» 1 Black-billed Sheathbill (Cheonis minor). On approval.
. 1 Red Kangaroo (Macropus rufus), 2. Born in the Menagerie.
» 1 Red-and-Yellow Macaw (Ara chloroptera). Deposited.
1 Variegated Sheldrake (Tadorna variegata), §. Received in
exchange,
. 2 Rock-Sparrows (Petronia stulta). Presented by D’Arey
Thompson, Esq.
1 Indian Muntjac (Cor thaaas muntjac), 3. Born in the Menagerie.
1 Common Snipe (Gallinago media), Presented by WK
Stanley, Esq.
1 ee Adder (Vipera berus). Presented by Colonel Irby,
E.Z8.
1 Four-lined Snake (Coluber quadrilineatus). Deposited.
2 Brown Coatis (Nasua nasica). Presented by the Hon. C. H.
Wynn.
1 Palm-Squirrel (Sciurus palmarum). Presented by Miss Bar-
clay.
65*
1900
Mar. 2.
10.
ie
13.
14,
APPENDIX.
1 Ocelot (Felis pardalis), 9. Presented by H. B. Whitmarsh, Esq.
1 Yellow Snake ( Chilobothrus inornatus). Purchased.
2 Black-capped Bitterns (Butorides atricapilla). Received in
exchange.
1 Red-backed Buzzard (Buteo erythronotus). Received in ex-
change.
] Yellow-checked Amazon (Chrysotis autumnalis), Purchased.
. 1 Macaque Monkey (Macacus cynomolgus), 3. Presented by H.
Wright, Esq.
1 Green Monkey (Cercopithecus callitrichus),2. Presented by
G. H. Garrett, Esq.
2 Common Chameleons (Chameleon vulgaris). Presented by
G. H. Garrett, Esq.
. 1 Macaque Monkey (Macacus egnomolgus), $. Presented by
Captain Pole Carew.
1 Herring-Gull (Larus argentatus). Presented by J. W. Cap-
stick, Esq., F.Z.S.
. | Green Lizard (Lacerta viridis). Deposited.
2 Undulated Grass-Parrakeets (Melopsittacus undulatus). Pre-
sented by Mrs. Hylton Jolliffe.
. 2 Sambur Deer (Cervus aristotelis), G and 9. Received in ex-
change.
1 Isabelline Bear (Ursus isabellinus). Received in exchange.
See P. Z. 8. 1878, p. 378.
1 Javan Adjutant (Leptoptilus javanicus). Received in ex-
change.
1 Siren (Stren lacertina). From South Carolina. Received in
exchange.
. 2 Long-legged Buzzards (Buteo ferox). Presented by W. J. E.
Cartwright, Esq. From India.
1 White-cheeked Bulbul (Pycnonotus leucogenys). Purchased.
1 Levaillant’s Darter (Plotus levaillanti). Purchased. See P. Z.S.
1878, p. 378.
2 Dace (Leuciscus vulgaris). Presented by Mr. Parsons.
1 Eel (Anguilla vulgaris). Presented by Mr. Parsons.
1 Blue-bonnet Parrakeet (Psephotus hematogaster). Purchased.
1 Lion (Felis leo), 3. Received in exchange.
1 Three-striped Paradoxure (Paradoxurus trivirgatus) Presented
by Captain Dalrymple.
3 Black Swans (Cygnus atratus). Bred in the Gardens.
1 Zebu (Bos indicus), 2. Born in the Gardens.
2 Common Marmosets (Hapale jacchus). Presented by Robert
Donaldson, Esq.
1 Green Glossy Starling (Lamprocolius chalybeus). Presented
by Mrs. Arabin, F.Z.8.
1 White-eared Bulbul (Pycnonotus leucotis), Presented by Mrs.
Arabin, F.Z.S.
1 Californian Quail (Callipepla californica). Presented by Mrs.
Arabin, F.Z.8.
1 Common Kestrel (Timnuneulus alaudarius). Presented by A.
Blumenthal, Esq.
5. 2 Common Badgers (Meles taxus). Born in the Gardens.
5. 1 Secretary Vulture (Serpentarius reptilivorus). Presented by
Messrs. W. Rigg and J. Curtiss.
1 Variegated Sheldrake (Tadorna variegata). Purchased.
. | Savigny’s Eagle-Owl (Bubo ascalaphus). From Bushire, Per-
sian Gulf. Presented by Dr. J. Huntly.
ADDITIONS TO THE MENAGERIE. 1001
Mar. 19. 2 Darwin’s Pucras Pheasants (Pucrasia darwint), § and 9.
Apr.
Purchased.
20. 2 Reindeer (Rangifer tarandus). Received in exchange,
21.
10.
1 Jay (Garrulus glandarius). Presented by J. Young, Esq.
1 Malayan Bear (Ursus malayanus), Q. Presented by 8, Palmer,
E
sq.
1 Spotted Eagle-Owl (Bubo maculosus). Received in exchange.
2 Plantain-Squirrels (Sciwrus plantan). Received in exchange.
1 Beceari’s Cassowary (Caswarius beccarii). Received in ex.
change.
4 Pied Grass-Finches (Spermestes Sringilloides). Received in
exchange.
1 Nvaioartia (Eunectes murinus). Received in exchange.
2 Matamata Terrapins (Chelys matamata). Received in ex-
change. See P.Z.S. 1878, p. 378.
1 One-streaked Hawk (Melierax monogrammicus). Received
in exchange.
2 Hog Deer (Cervus porcinus), 2 2. Received in exchange.
1 Egyptian Ichneumon (Herpestes wchneumon). Received in
exchange,
1 Short-toed Eagle (Circaétus gailicus). Presented by H. M.
Upcher, Esq., F.Z.S.
23. 1 Rhesus Monkey (Macacus erythreus), ¢. Presented by Mrs.
Baxter.
1 Green Monkey (Cercopithecus callitrichus). Presented by
James Bennet, Esq.
1 Vervet Monkey (Cercopithecus lalandii). Presented by W. B.
Westwood, Esq.
25. 1 Black-faced Spider Monkey (Ateles ater). Deposited.
28. 1 Philippine Paradoxure (Parudoxurus philippensis). Received
in exchange.
. 1 Arabian Gazelle (Gazella arabica), 2. Presented by W.
Webb, Esq.
- 2Pudu Deer ( Cervus humilis), 3 and 2. Purchased. From Chili.
1 Naked-eared Deer (Cervus gymnotis). Purchased. From
Keuador.
1 Macaque Monkey (Macacus cynomolgus). Presented by Francis
Pym, Esq.
2 Persian Gazelles (Gazella subgutturosa). Presented by R. W.
Inglis, Esq.
1 Maned Goose (Bernicla jubata), $. Purchased.
. 1 Egyptian Gazelle (Gazella dorcas), 3. Deposited.
1 Vulpine Phalanger (Phalangista vulpina). Presented by Capt.
T. Ayling.
. 1 Fraser’s Squirrel (Seiurus stramineus). Received in exchange,
See P.Z. 8. 1878, p. 441.
1 Black Sternothere (Sternotherus niger). Received in ex-
change.
- 1 Common Squirrel (Sciwrus vulgaris). Presented by Madame
Harte.
» 1 Arabian Baboon ( Cynocephalus hamadryas), 2. Presented liy
Dr. A. P. Woodforde.
2 Chaema Baboons (Cynocephalus porcarius). Presented by
Capt. W. L. Coke.
1 Short-eared Owl (Otws brachyotus). Presented by W. K.
Stanley, Esq.
1002
Apr. 11.
APPENDIX.
1 Yellow-footed Rock-Kangaroo (Petrogale sxanthopus), 3.
Born in the Menagerie.
. 1 Great Kangaroo (Mucropus giganteus). Presented by Crawford
Caffin, Esq., Lt. R.N.
2 Laughing Kingfishers (Dacelo gigantea). Presented by Craw-
ford Caffin, Esq., Lt. R.N.
1 Golden-winged Parrakeet (Brotogerys chrysoptera). Received
in exchange.
. 1 Green Monkey (Cercopithecus callitrichus), §. Presented by
George Milward, Esq.
1 South-American Rat-Snake (Spilotes variabilis), On ap-
proval.
15. 1 Stanley Crane (Tetrapteryx paradisea). Deposited.
16,
26.
1 Indian Leopard (Felis pardus), 3. Presented by Robert
Tubbs, Esq., F.Z.S,
1 Red Deer (Cervus elaphus), 3. Presented by Carrol W.
Ansdell, Esq.
1 Common Fox (Canis vulpes). Presented by Carrol W. Ansdell,
sq.
2 Spotted Ichneumons (Herpestes auropunctatus). Presented by
J. McIntosh, Esq.
1 Leadbeater’s Cockatoo (Cacatua leadbeaterr), Presented by
W. Ruston, Esq.
. 1 Surieate (Suricata zenik), 2. Presented by Percy Howard,
Esq.
. 1 Azara’s Fox (Canis azare). Presented by Dr. A. Stradling.
. 4 Common Foxes (Canis vulpes). Born in the Menagerie.
. 1 Collared Fruit-Bat (Cynonyeteris eollaris). Born in the Mena-
gerie.
. 2 Snow-Buntings (Plectrophanes nivalis). Purchased.
. 1 Bennett's Wallaby (Halmaturus bennett?). Born in the Mena-
erie.
1 Hybrid Duck (between Mareca chiloensis and Dajila spini-
cauda). Presented by I. Charlton Parr, Esq.
2 Common Carp (Cyprinus carpio). Presented by Richard
Seyd, Esq., F.Z.S.
5. 1 Black Wallaby (Halmaturus ualabatus), d. Presented by
D. W. Barker, jun., Esq.
1 Laughing Kingfisher (Dacelo gigantea). Presented by D. W.
Barker, jun., Esq.
1 ee Finch (Sycalis flaveola). Presented by Mrs. Hylton
oliffe.
1 Cariama (Cariama cristata). Purchased.
1 Crested Curassow (Crav alector). Purchased.
1 Guira Cuckoo (Gara piririgua). Purchased.
1 laoreet Tree-Duck (Dendrocygna autumnalis). Pur-
chased.
2 Red-billed Tree-Ducks (Dendrocygna autumnalis). Purchased.
1 Common Seal (Phoca vitulina). Purchased.
4 Short-tailed Wallabies (Halmaturus brachyurus),1 g and 3 9°.
Purchased.
1 Long-tailed Cuckoo (Eudynamis taitensis). Purchased.
1 Dominican Gull (Larus dominicanus), Purchased.
1 Blue-bonnet Parrakeet (Psephotus hematogaster). Purchased.
1 Stanley Parrakeet (Platycercus icterotis). Purchased.
1 Bar-headed Goose (Anser indicus). Purchased.
27. 1 African Leopard (Felis pardus), 3, Presented by Mrs. Kirk,
a
Apr. 27.
29.
30.
May 1.
. 2 Black-faced Spider, Monkeys (Ateles ater), 29. Purchased,
ADDITIONS TO THE MENAGERIE, 1003
1 Beisa Antelope (Oryx besa), 3. Presented by ELH. the
Sultan of Zanzibar.” See P. Z. 8. 1878, p- 441,
1 Black-crested Eagle (Spizaétus oceipitalis). Purchased.
1 Sand-Lizard (Lacerta agilis). Presented by Master W. L.
Sclater.
1 Smooth Newt (Ziiton teniatus). Presented by Master B. L.
Sclater,
1 Great Kangaroo (Macropus gganteus), 2. Born in the Mena-
gerie,
2 Black Storks (Ciconia nigra). Purchased,
1 Brazilian Tree-Poreupine (Sphingurus prehensilis), Purchased,
Long-nosed Crocodile (Crocodilus americanus). Purchased,
1 Macaque Monkey (Macacus cynomolgus). Presented by J. M.
Neil, Esq.
1 Lion (Felis leo). Presented by J. D. Massey, Esq.
1 Black Ape ( Cynopithecus niger), Deposited,
1 Eland (Oreas canna), 2. Born in the Menagerie.
1 Common Cassowary ( Caswarius galeatus). Purchased.
1 Greek Partridge (Caccabis savatilis), white var., 3. Presented
by F. Papillon, Esq. From Beyrout,
1 Golden-winged Woodpecker ( Colaptes auratus). Purchased.
2 Lesser Birds of Paradise (Paradisea minor). Purchased. See
P. Z.8. 1878, p. 631,
- 1 Vervet Monkey ( Cercopithecus lalandii), Presented by G. W.
Twining, Esq.
1 Common Cormorant (Phalacrocorax carbo). Presented by
Lord Braybrooke, F.Z.S.
4 Green Lizards (Lacerta viridis). Presented by F. E. Lawder,
Esq.
i Bibhiecues Marmoset (Huapale penicillata). Presented by
Walter M. St. Aubyn, Isq.
- I Bonelli’s Eagle (Misaétus Sasciatus). Deposited,
1 Hawk’s-billed Turtle ( Chelone wmbricata). Presented by J. W.
Cossback, Esq.
1 Common Snake ( Tropidonotus natrix). Presented by Henry
Smith, Esq.
1 Leadbeater’s Cockatoo (Cacatua leadbeateri). Presented by
Mrs. Tennent.
- 1 Macaque Monkey (Macacus cynomolgus). Presented by W.
Mackie, Esq.
. 1 Toque Monkey (Macacus pileatus), 3. Presented by Master
R. C, Heyworth.
1 Black-backed Jackal (Canis mesomelas), Q. Presented by
Richard Seyd, Esq., F.Z.S.
1 Common Badger (Meles tarus). Presented by Messrs, J. H.
and B. H. Thompson.
1 White-backed Piping Crow (Gymnorhina leuconota). Pre-
sented by John Ritchie, Esq.
- LSlow Loris (Mycticebus tardigradus), 2. Received in ex~
change,
1 Bonnet-Monkey (Macacus radiatus). Presented by R. Bryant,
Esq.
1 Two-spotted Paradoxure (Nandinin binotata), Presented by
Capt. E. J. Howes, ;
1 Goffin’s Cockatoo (Cacatua goffnt). Presented by Mrs, Pitt,
1004
May
9.
10.
We
12.
15.
14,
15.
live
bo
Or
LN)
“I
APPENDIX.
1 Wonga-wonga Pigeon (Leucosarcia picata). Bred in the
Gardens.
1 Spotted Turtle Dove (Turtur suratensis). Bred in the
Gardens.
1 Jaguar (Felis onca). Presented by Major Wood.
1 Blue-fronted Amazon (Chrysotis estiva). Presented by Dr.
Byron Whitehead.
9 Chilian Pintails (Dajila spinicauda). Bred in the Gardens.
1 Darwin’s Pucras Pheasant (Purcrasia darwint). Bred in the
Gardens.
3 Water-Ouzels (Cinclus aquaticus). Presented by R. J. L.
Price, Esq., F.Z.S.
1 Reindeer (Rangifer tarandus). Born in the Menagerie.
1 Collared Fruit-Bat (Cynonycteris collaris). Born in the
Menagerie.
1 Wood-Owl (Syrnium aluco). Presented by C. B, Wharton,
Esq., F.Z.S.
1 Rattlesnake (Crotalus durissus). Deposited.
1 Reindeer (Rangifer tarandus), 2. Born in the Menagerie.
1 European Bearded Vulture (G: es barbatus). Deposited.
1 Nilotic Fox (Canis niloticus), 3. Presented by Commr. F.
Cotton. From Djeddah, Arabia.
1 Hog-nosed Snake (Heterodon platyrhinos). Presented by
Dr. Painter, F.Z.S.
1 Chinchilla ( Chinchilla lanigera). Born in the Menagerie.
4 Variegated Sheldrakes (Yadorna variegata). Bred in the
Gardens.
. 1 Great Kangaroo (Macropus giganteus), 2. Born in the
Menagerie.
22. 2 2 Bar-headed Geese (Anser indicus), ¢ and 2. Purchased.
23. 2 Chimpanzees (Troglodytes niger), 2 3. Deposited.
4 Water-Ouzels . Cinclus aquaticus). Presented by R. J. L.
Price, Esq., F.Z.S.
1 Vacated Sheldrake (Tadorna variegata), ¢. Received in
exchange.
2 Great-billed Rheas (Rhea macrorhyncha). Received in ex-
change.
2Sulphury Tyrants (Pitangus sulphwratus). Received in ex-
change.
2 Geoffroy’ s Doves (Peristera geoffrow). Bred in the Gardens.
7 Chilian Pintails (Dajila spinicauda). Bred in the Gardens,
. 1 Bonnet-Monkey (Macacus radiatus), §. Presented by Henry
Willatt, Esq.
1 White-fronted fe uchin rey: albifrons), 2. Presented by
Dayrell Davies, Esq., R.N.
2 Wild Cats (Felts catus). Born in the Menagerie.
8 Wild Boars (Sus scrofa). Born in the Menagerie.
1 Yellow-legged Herring-Gull (Larus leucopheus). Bred in
the Gardens.
1 Hairy Tapir (Tapirus roulint), 3. Received in exchange.
See P. Z.S. 1878, p. 681, pl. xxxrx.
83 Common Rheas (hea camer wcana). Presented by Frank
Parish, Esq.
. 4Temminck’s Tragopans (Certornis temmincki). Bred in the
Gardens.
1 Brazilian Caracara (Polyborus brasiliensis). Presented by Miss
Amslie, _
Oa a arn
ADDITIONS TO THE MENAGERIE. 1005
May 29. 1 Macaque Monkey (Macacus cynomolgus), ¢. Presented by
J. Farmer, Esq.
1 Axis Deer (Cervus axis), $. Born in the Menagerie.
30. 1 Geoftroy’s Cat (Felis geoffroi), 3. Presented by Ronald
Bridgett, Esq.
1 Yellow-footed Rock-Kangaroo (Petrogale xanthopus). Born
in the Menagerie.
31. 1 Tamandua Anteater (Tamandua tetradactyla). Purchased.
June 3. 8 Common Sticklebacks (Gasterosteus aculeatus), Presented by
Master G. L. Sclater.
4, 2 Indranee Owls (Syrnium indranee). Deposited.
5. 1 White-fronted Amazon (Chrysotis leucocephala). Purchased,
6. 1 Mouflon (Ovis musimon). Born in the Menagerie.
2 Greater Spotted Woodpeckers (Picus major), Presented by
J. A. Cooper, Esq.
1 Greater Sulphur-crested Cockatoo (Cacatua galerita). Pre-
sented by N. Partocalis, Esq.
- 2 Horned Tragopans (Ceriornis satyra). Bred in the Gardens.
1 Impeyan Pheasant (Lophophorus impeyanus). Bred in the
Gardens.
2 Oystercatchers (Hematopus ostralegus). Purchased.
12 Green Tree-Frogs (Hyla arborea). Purchased.
8. 2 Mandrills (Cynocephalus mormon). Presented by G. H.
Garrett, Esq.
1 Ocellated Monitor (Monitor ocellatus). Presented by G. H.
Garrett, Esq.
2 Black Rats (Mus rattus). Presented by W. H. Hatcher,
N
sq.
1 Gnaeap Chameleon (Chameleon vulgaris). Presented by
W. W. Spicer, Esq.
9. 1 Rhesus Monkey (Macacus erythreus). Born in the Mena-
erie,
2 oot Seals (Phoca vitulina), Purchased.
10. 1 Horned Lizard (Phrynosoma cornutum). Presented by J. C.
Witte, Esq.
1 White Ibis (Ics alba). Purchased.
1 Spotted Cavy (Cewlogenys paca). Purchased.
1 Ocelot (Felis pardalis). Presented by Capt. J. Moir.
1 Black-faced Spider Monkey (Ateles ater). Presented by Capt.
J. Moir.
1 Common Boa (Boa constrictor). Presented by Capt. J. Moir.
1 West-India Rail (Aramides cayennensis). Presented by Capt.
J. Moir.
1 Black Tortoise (Testudo carbonaria). Presented by Capt. J.
Moir.
2 Black-crested Cardinals (Gubernatrix cristatella). Presented
by Mrs. Arabin, F.Z.S,
1 American Thrush (Turdus miugratorius). Presented by Mrs.
Arabin, F.Z.S.
11. 1 Spotted Turtle Dove (Turtur suratensis). Bred in the
ardens,
2 Brazilian Teal (Querquedula brasiliensis). Bred in the
Gardens.
1 Bay Antelope ( Cephalophus dorsalis), 2. Purchased.
1 Black Saki (Pithecia satanas). Purchased.
12. 1 Chimpanzee (Troglodytes niger), g%. Deposited.
1006
June 13.
14,
165.
Wi
19.
20.
21.
bo
bo
APPENDIX.
1 Himalayan Bear (Ursus tibetanus), d. Presented by Com-
mander J. S. Murray.
1 Indian Crow (Corvus splendens). Presented by Commander
J.S. Murray.
6 Herring-Gulls (Zarus argentatus). Presented by Arthur
Clarke, Esq.
1 Rufous Rat-Kangaroo (Hypsiprymnus rufescens). Presented
by T. Wickenden, Esq.
1 Reeves’s Muntjac (Cervulus reevest), Born in the Gardens,
6 Upland Geese (Bernicla magellanica). Bred in the Gardens.
1 Indian Gazelle (Gazella bennetti) g. Presented by Miss A.
Statter.
1 Vulpine Phalanger (Phalangista vulpina). Presented by E.
J. Beagle, Esq.
1 Common Cormorant (Phalacrocorax carbo). Presented by W.
Thompson, Esq.
. 2 Cereopsis Geese (Cereopsis nove-hollandie), § and 9.
Purchased.
2 Australian Sheldrakes (Zadorna tadornoides), 3 and Q.
Purchased.
4 Smooth Newts (Triton teniatus). Presented by Master B.
L. Sclater.
2 Mantchurian Crossoptilons (Crossoptilon mantchuricum), 3
and 9. Received in exchange.
2 Black-billed Tree-Ducks (Dendrocygna arcuata). Received
in exchange.
2 Prairie Marmots (Cynomys ludovicianus). Presented by
Mons. J. W. Cornély, C.M.Z.8,
1 Yellow-billed Sheathbill (Chzonts alba). Purchased.
1 Barbel (Lota vulgaris). Presented by Earl Brownlow, F.Z.S,
1 Green Monkey (Cercopithecus callitrichus), 2. Presented by
Samuel Curtis, Esq.
1 Red-billed Tree-Duck (Dendrocygna autumnalis). Presented
by R. M. Hyde, Esq.
1 Scarlet Ibis (Ibis rubra). Presented by R. M. Hyde, Esq.
1 Pig-tailed Monkey (Macacus nemestrinus), 3. Presented by
W. H. Gedge, Esq.
1 Hybrid Ass (between Equus onager 3 and Equus hemippus),
Born in the Menagerie.
1 Mueller’s Parrot (Tanygnathus muellert). Purchased.
1 Talpacoti Ground-Dove (Chalcopelia talpacoti). Bred in the
Gardens.
. 2 Argus Pheasants (Argus giganteus). Bred in the Gardens.
4 Summer Ducks (Aix sponsa). Bred in the Gardens.
4 Chilian Widgeon (Mareca chiloensis). Bred in the Gardens.
3 Australian Wild Ducks (Anas superciliosa). Bred in the
Gardens.
2 Common Cormorants (Phalacrocorax carbo).. Presented by-
Edward Banks, Esq.
. 1 Pine-Marten (Martes abietum). Presented by Robert Walters,
Esq., F.Z.8.
1 Green-necked Peafowl (Pavo spicifer). Bred in the Gardens.
. 1 Rhesus Monkey (Macacus erythreus), 9. Presented by Mrs.
Walcot.
10 Herring-Gulls (Larus argentatus). Deposited.
1 Brown Capuchin (Cebus fatuellus). Presented by A. Warm-
ington, Hsq.
June 25.
26.
ADDITIONS TO THE MENAGERIE. 1007
1 Daubenton’s Curassow (Crax daubentoni). Presented by A.
Warmington, Esq.
1 Common Boa (Boa constrictor). Purchased.
lL Garden’s Night-Heron (Nycticorax gardeni), Purchased.
1 Japanese Wolf (Canis hodophylav), Q. Presented by H.
Heywood Jones, Esq., F.Z.5. See P. Z. 8. 1878, p. 788.
1 Common Seal (Phoca vitulina). Deposited.
6 Goldfinches (Carduelis elegans). Presented by C. F. Johnson,
Esq.
1 Panndn Chameleon (Chameleon vulgaris). Presented by C.
F. Johnson, Esq.
A Geoffroy’s Doves (Peristera geoffrow). Bred in the Gardens.
4 Cunningham’s Skinks (Eyernia cunninghami). Presented by
J. G. Pearson, Esq.
. 1 Bonnet-Monkey (Macacus radiatus), 2. Presented by Alex.
C. Forbes, Esq.
. 1 Short-tailed Capromys (Capromys brachyurus). Presented by
the Hon. H. J. Burford Hancock.
1 Indian Antelope (Antilope cervicapra), 2. Received in ex-
change.
1 Stump-tailed Lizard (Zrachydosaurus rugosus), Presented by
Philip Saville, Esq.
vee como ta Monkey (Cercopithecus campbelli), g. Presented
by Capt. C. F. Filliter.
1 Common Marmoset (Hapale jacchus), Q. Presented by
Edward Clayton, Esq.
1 Pinche Monkey (Midas edipus), g. Presented by Edward
Clayton, Esq.
. 1 Green Monkey (Cercopithecus callitrichus), 9. Presented by
Mrs. George Yeomans.
. 1 Reeves's Muntjac (Cervulus reevest), 2. Born in the
Menagerie.
. 2 Black Wallabies (Halmaturus ualabatus), GandQ. Received
in exchange.
1 Australian Monitor (Monitor gouldi). Purchased.
. 6 Amherst Pheasants (Zhaumalea amherstie). Bred in the
Gardens.
1 Fork-tailed Jungle-fowl (Gallus furcatus). Bred in the Gar-
dens,
1 Demeraran Cock of the Rock (Zupicola crocea), Purchased.
1 Prince Albert’s Curassow (Crax alberti), ¢. Purchasd.
2 Black-faced Spider Monkeys (Ateles ater), ¢ and 9. Pur-
chased.
. 1 Persian Gazelle (Gazella subgutturosa), §. Presented by Col.
Nixon.
1 Goliath Heron (Ardea goliath). Presented by Col. Nixon.
1 Spotted Ichneumon (Herpestes awropunctatus). Presented by
Irs, Payne.
. 1 King Parrakeet (Aprosmictus scapulatus). Presented by
Ebenezer Carr, Esq.
. 1 Malbrouck Monkey (Cercopithecus cynosurus), 2. Presented
by Edward Wright, Esq.
2 Red-faced Weaver-birds (Foudia erythrops). Presented by
Miss M. Jackson.
1 Chestnut-bellied Finch (Munia rubro-nigra). Presented by
Miss M. Jackson,
1008
July 6.
co
10,
ll.
14.
APPENDIX.
2 Common Redpoles (Linota rufescens). Presented by Miss M.
Jackson.
2 Canary Finches (Serius canarius). Presented by Miss M.
Jackson.
2 Aisculapian Snakes (Coluber esculapit). Presented by Lord
Lilford, F.Z.S. From Dalmatia.
2 Lacertine Snakes (Celopeltis lacertina). Presented by Lord
Lilford, F.Z.S. From Tunis.
1 Horse-shoe Snake (Zamenis hippocrepis). Presented by Lord
Lilford, F.Z.S. From Tripoli.
1 Leopardine Snake (Coluber leopardinus). Presented by Lord
Lilford, F.Z.S. From Dalmatia.
1 Vivacious Snake (TZachymenis vivaz). Presented by Lord
Lilford, F.Z.S. From Montenegro.
1 Dark-green Snake (Zamenis atrovirens, var. carbonarius).
Presented by Lord Lilford F.Z.8. From Italy.
1 Four-rayed Snake (Hlaphis quaterradiatus). Presented by
Lord Lilford, F.Z.5. From Dalmatia.
. 4 Vulturine Guinea-fowls (Nwmida vulturina). Deposited.
. 1 Hybrid Fallow Deer between Dama mesopotamica $ and
Dama vulgaris 9. Born in the Menagerie.
1 Hairy Tree-Porcupine (Sphingurus prehensilis). Born in the
Menagerie.
2 Water- Vipers ( Cenchris piscivorus). Purchased.
5 Seven-banded Snakes ( Zropidonotus leberis), Purchased.
4 Red-headed Buntings (Huspiza luteola). Purchased.
4 Green-winged Doves (Chalcophaps indica). Purchased.
1 Plantain-Squirrel (Sezwrus plantani). Purchased.
1 Water-Chevrotain (Hyomoschus aquaticus). Purchased.
1 Electric Silurus (Malapterurus beninensis). Purchased,
1 Beatrix Antelope (Oryx beatriz), §. Presented by Comman-
der Burke, 8.8. ‘Arcot.’ From Arabia. See P.Z.S. 1878,
. 789.
1 BaneOwl (Strix flammea). Presented by Commander Wyatt,
8.8. ‘Deccan.’ From Mesopotamia.
3 Bullfinches (Pyrrhula rubicila). Presented by Miss A.
Sargeant.
4 Paradise Whydah-birds (Vidua paradisea). Presented by
Archibald Brown, Esq.
1 Pin-tailed Whydah-bird (Vidua principalis). Presented by
Archibald Brown, Esq.
3 Grenadier Weaver-birds (Zuplectes oryx). Presented by
Archibald Brown, Esq.
2 Hooded Finches (Spermestes cucullatus). Presented by Archi-
bald Brown, Esq.
2 Crested Poreupines (Hystrix cristata). Presented by Dr. G.
S. Badger.
1 Banded Ichneumon (Herpestes fasciatus). Presented by Dr.
G. S. Badger.
1 Hawk’s-billed Turtle (Chelone imbricata). Presented by Capt.
Henderson.
1 Green Tree-Frog (Hyla arborea). Presented by J. Green-
wood, Esq.
1 Green Tree-Frog (Hyla arborea). Presented by Lord Arthur
Russell, F.Z.8.
. 1 Macaque Monkey (Macacus cynomolgus), g. Presented by
Mr. Enoch.
-
July 15,
16.
19,
20.
22,
23.
29.
31,
ADDITIONS TO THE MENAGERIE. 1009
1 Golden-headed Marmoset (Midas chrysomelas), 3. Purchased.
From Para.
1 Horrid Rattlesnake (Crotalus horridus). Purchased. From
Bahia.
1 Thick-billed Tanager (Zuphonia crassirostris), 9. Purchased.
From Para.
3 Alligator Terrapins ( Chelydra serpentina). Presented by J. H.
Thompson, Esq., C.M.Z.S.
1 Eland (Oreas canna), 6. Born in the Menagerie.
1 Rhesus Monkey (Macacus erythreus), $. Presented by Miss
Davis.
2 Cape Zorillas (Ictonyx zorilla). Presented by Mrs. J. J.
Monteiro,
1 Argus Pheasant (Argus giganteus). Bred in the Gardens.
5 Amherst Pheasants (TZhaumalea amherstie). Bred in the
Gardens.
1 Common Cuckoo (Cuculus canorus). Presented by G. D.
Careless, Esq.
2 Smooth Snakes (Coronella levis). Purchased. From Hamp-
shire.
1 Black-necked Swan (Cygnus nigricollis). Bred in the Gar-
dens.
1 Nightingale (Daulias luscinia). Presented by — Gee, Esq.
1 heart Raa igey aluco). Presented by H. T, Wharton,
Esq., F.Z.8.
it Bed-nndeblne Macaw (Ara macao), Presented by H. T.
Wharton, Esq., F.Z.S.
1 Blue-and-yellow Macaw (Ara ararauna). Presented by H.
T. Wharton, Esq., F.Z.S.
1 Common Gull (Larus canus). Presented by F. Lancelott,
Esq.
q
. 2 Macaque Monkeys (Macacus cynomolgus), ¢ andQ. Presented
by Dr. Adcock.
2 American Flying Squirrels (Sciwropterus volucella), g and 2.
Received in exchange.
1 Coati (Nasua nasica). Received in exchange.
. 1 Orang-outang (Simia satyrus),Q. Received in exchange.
See P.Z.S. 1878, p. 789.
1 Green-winged Dove (Chalcophaps indica), Presented by Capt.
W. Otho N. Shaw.
1 Burrowing Owl (Speotyto cunicularia). Presented by Geo. E.
P. Nixon, Esq., M.D.
6 Common Guillemots (Uria troile). Presented by Sir Hew
Dalrymple, Bart.
1 Greater Sulphur-Crested Cockatoo (Cacatua galerita). Pre-
sented by Mrs, A. A. Hole.
1 Ocelot (Felis pardalis), Purchased.
1 Black Vulture (Cathartes atratus), Purchased.
2 Chilian Sea-Eagles ( Geranoaétus melanoleucus). Purchased.
1 Brazilian Caracara (Polyborus brasiliensis), Purchased.
2 Rufous Tinamous (Rhynchotus rufescens). Purchased.
9 Spotted Tinamous (Nothura maculosa). Purchased.
1 Razor-billed Curassow (Mitwa tuberosa). Purchased.
1 Mexican Deer (Cervus mexicanus), 9. Presented by A.
Serutton, Esq.
1 Common Fox (Canis vulpes). Presented by Athelston Riley,
Esq.
1010
July 31.
Aug. 1.
APPENDIX.
1 Brown Coati (Nasua nasica), §. Deposited.
4 Indian Rat-Snakes (Ptyas mucosa). Received in exchange.
3 kong-snouted Snakes (Passerita nycterizans). Received in
exchange,
4 Long-banded Snakes (Zropidonotus vittatus). Received in
exchange.
1 Slow Loris (Mycticebus tardigradus) 3. Presented by The-
odore Hance, Esq.
3 Common Cranes (Grus cinerea). Presented by Theodore Hance,
Esq. From Chinkiang China.
1 Copper-head Snake (Cenchris contortrir). Presented by Dr.
F. Painter, F.Z.8.
. 8Great Bustards (Otis tarda), 1 g and 2 9. Presented by
Lord Lilford, F.Z.S.
1 Tiger Bittern (Tigrisoma brasiliense). Presented by G. H.
Hawtayne, Esq., C.M.Z.S.
1 Passerine Owl ( Glaucidium passerinum). Presented by W. E.
Tait, Esq., C.M.Z.S.
1 Spanish Blue Magpie (Cyanopolius cooki). Presented by
ord Lilford, F.Z.S. See P. Z.S. 1878, p. 789.
1 Caspian Terrapin (Clemmys caspica), Presented by T. Wilson,
Esq.
2 Wass Pheasants (Argus giganteus). Bred in the Gardens.
. 2 Beisa Antelopes (Oryx beisa), g and 2. Presented by Capt.
F. M. Hunter, B.C.S.
1 Bacha Eagle (Spilornis bacha). Presented by J. 8. Jameson,
Esq. From Labuan, N.W. Borneo.
. 1 Axis Deer ( Cervus axis), 2. Born in the Menagerie.
. 2 European Lynxes (Felis lynx), d and 9. Presented by Major
Chadwick. From Norway.
1 Hobby (Hypotriorchis subbuteo). Presented by W. H. Scratton,
Es
2 Common Kingfishers (Alcedo ispida). Purchased.
1 Wapiti Deer (Cervus canadensis), 2. Born in the Gardens.
. 1 Common Boa (Boa constrictor). Presented by Arthur Strad-
ling, Esq.
1 Coati (Nasua nasica). Received in exchange.
3 Bluish Finches (Spermorphila cerulescens). Deposited.
1 Thick-billed Seed-Finch (Oryzoborus crassirostris). Deposited.
1 Red-and-blue Macaw (Ava macao). Deposited.
1 Yellow Snake (Chilobothrus inornatus). On approval.
. 1 Weeper Capuchin (Cebus capucinus). Deposited.
1 Black-crested Cardinal (Gubernatrix cristatella). Bred in the
Gardens.
2 Talpacoti Ground-Doves (Chamepelia talpacoti). Bred in the
Gardens. r
1 Solitary Thrush (Monticola cyanus). Presented by W. Verner,
Ks
. 1 Black-footed Penguin (Spheniscus demersus). Purchased.
. 1 Black Rat (Mus rattus). Presented by R. Morton Middle-
ton, Esq.
7 Greek Land-Tortoises (Testudo greca). Deposited.
2. 3 Guinea-Pigs (Cavia caprera). Presented by Mr. J. Gould.
3. 1 Lion (Felis leo), §. Presented by W. H. Wylde, Esq.
1 Patas Monkey (Cercopithecus ruber). Presented by W. H.
Wylde, Esq.
Aug. 13.
14,
15.
Liye
19,
21,
22,
23.
24,
26.
3l.
ADDITIONS TO THE MENAGERIE. 1011
2 Common Buzzards (Buteo vulgaris). Presented by Valentine
Marks, Esq.
2 Herring-Gulls (Larus argentatus). Presented by Thomas
Landseer, Esq.
1 Crested Ground-Parakeet (Calopsitta nove-hollandig). Pre-
sented by Mrs. Parker,
2 Barbary Apes (Macacus inuus). Deposited.
1 Bladder-nosed Seal ( Cystophora cristata). Purchased,
1 Common Paradoxure (Paradoxurus typus). Presented by
Edwin Etty Sass, Esq.
1 Macaque Monkey (Macacus cynomolgus), 3. Deposited.
2 Beautiful Parrakeets (Psephotus pulcherrimus). Deposited.
1 Many-zoned Hawk (Melierax polyzonus). Presented by C. H.
Fisher, Esq. From Mogador.
6 Common Kingfishers (Alcedo ispida). Purchased.
13 Greek Land-Tortoises (Testudo greca). Deposited.
2 Saffron Finches (Syealis flaveola). Bred in the Gar-
dens.
1 Prairie-Wolf (Canis latrans). Presented by Prof. C. M.
Vincent, F.G.S., F.R.G.S. °° From the Rocky Mountains.
See P. Z. S. 1878, p. 789.
1 Common Jackal (Canis awreus), 3. Presented by Mr. James
Smith.
1 Spotted Ichneumon (Herpestes auropunctatus), 2. Presented
ry Mr. W. Pyne.
1 Bronze Fruit-Pigeon (Carpophaga @nea). Presented by Mr.
A. H. Jamrach.
1 White-eared Bulbul (Pycnonotus leucotis). Presented by J.
W. Wooler, Esq.
1 Bonnet-Monkey (Macacus radiatus), 3. Presented by Capt.
Clarke.
11 Spinose Lizards (Agama colonorum). Received in ex-
change.
1 Cape Bucephalus (Bucephalus capensis). Received in ex-
change.
1 Smooth-bellied Snake (omalosoma lutriz). Received in ex-
change.
1 Crossed Snake (Psammophis crucifer). Received in ex-
change.
. 1 Common Marmoset (Hapale jacchus). Deposited.
- 1 Grey Parrot (Psittacus erithacus). Deposited.
» 1 Common Crow (Corvus corniv). Presented by F. Sydney B.
Brough, Esq.
3 Hybrid Snakes (between Chilobothrus inornatus ¢ and Epi-
crates angulifer?). Born in the Menagerie. See P.Z, S.
1878, p. 789. ;
1 White-lipped Peccary (Dicotyles labiatus). Purchased.
1 Golden Agouti (Dasyprocta aguti). Presented by Geo. Ham-
mond Hawtayne, Esq., C.M.Z.S.
1 Bahama Duck (Dajila bahamensis). Bred in the Gardens.
1 Red Deer (Cervus elaphus). Born in the Menagerie,
- 1 Caspian Ouaran (Psammosaurus caspicus). Presented by F. A.
Floyer, Esq., F.Z.S. From Khan Yunis, Egypto-Syrian
frontier.
. 1 Turquoisine Parrakeet (Euphema pulchella). Bred in the Gar-
dens.
16.
19.
20.
21.
APPENDIX.
1 Goffin’s Cockatoo (Cacatua goffini). Deposited by F. A.
Rogers, Esq.
1 Sykes’s Monkey (Cercopithecus albigularis),3. Presented by
Capt. Benson.
. 1 Hybrid Kaleege,¢. Presented by A. F. Wiener, Esq.
2 Tayras (Galictis barbara). Born in the Menagerie.
1 Banded Ichneumon (Herpestes fasciatus), §. Presented by the
Rey. H. F, B. Lawrence.
1 Japanese Warbler (Cettia cantans). Deposited by E. L. Hyde,
Esq.
. 1 Banded Ichneumon (Herpestes fasciatus), 2. Presented by
F. T. Blackley, Esq.
. 2 Vinaceous Turtle Doves (Twrtw- vinaceus). Presented by
Miss Harris.
1 Greek Land-Tortoise (Zestudo greca). Presented by Miss
Harris.
. 1 Common Adder (Vipera berus). Presented by the Viscount
Mandeville.
. 1 Spotted Turtle Dove (Turtur aurita). Bred in the Gardens.
1 Oriental Eagle-Owl (Bubo orientalis). Presented by Charles
Fowler, Esq. From Karennee, Siam. See P.Z.S. 1878,
p- 790.
1 Passerine Owl (Glaucidium passerinum). Presented by Al-
bert Momber, Esq.
. 1Common Adder ( Vipera berus). Presented by F. Buckland, Esq.
. 2 Prairie-Marmots (Cynomys ludovicianus). Presented by Wil-
fred G. Marshall, Esq.
2 Leconte’s Rattlesnakes (Crotalus lecontii). Presented by
Wilfred G. Marshall, Esq. From Colorado.
1 Macaque Monkey (Macacus cynomolgus),3. Presented by
Miss Emily Palmer.
2 Ostriches (Struthio camelus), ¢ and 2. Presented by the
Hon. H. C. Vivian, H.B.M. Consul General, Cairo, See
P.Z.S. 1878, p. 790.
2 Secretary Vultures (Serpentarius reptilivorus). Presented by =
C. Rivers Wilson, Esq., C.B
2 Ecyptian Gazelles (Gazella doréas), § and 9. Presented by
Thos. Moss, Esq.
3 Reddish Finches (Spermophila 7% mtia). Presented by
R. C. Batterbee, Esq. Ni
1 Lineated Finch (Spermophila lineata), Presented by R. C.
Batterbee, Esq.
1 Guttural Finch (Spermophila gutturalis). Presented by R. C.
Batterbee, Esq.
1 Tropical Seed-Finch (Oryzoborus torridus). Presented by R.
C. Batterbee, Esq
1 Plumbeous Finch (Spermophila plumbea). Presented by R.
C. Batterbee, Esq.
3 Rufous Tinamous (Rhynchotus rufescens). Presented by J.
A. Tliffe, Esq.
1 Leonine Monkey (Macacus leoninus), 3. Deposited.
1 Purple-faced Monkey (Semnopithecus leucoprymnus). Pur-
chased. :
1 Molucea Deer (Cervus moluccensis), ¢. Born in the Mena-
erie.
1 Motking Bird (Mimus polyglottus). Deposited.
1 Common Hangnest (Icterws vulgaris). Deposited.
ADDITIONS TO THE MENAGERIE. 1013
Sept. 22. 2 Lesser Black-backed Gulls (Larus fuscus). Presented by A.
Hi. Cocks, Esq., F.Z.S.
23, 2 White-crested Touracous (Corythaix albocristata). Presented
by W. Wormald, Esq.
25. 1 Bonnet-Monkey (Macacus radiatus). Presented by Frederick
Carter, Esq. -
1 Chestnut-backed Weaver-bird (Zyphantornis castaneofusca).
Deposited.
4 Po see Weaver-birds (Hyphantornis textor). De-
posited.
2 Grenadier Weavyer-birds (Zuplectes oryx). Deposited.
1 Barbary Turtledove ( Turtur risorius). Deposited.
1 Vinaceous Turtledove ( Zurtw vinaceus). Deposited.
1 Turquoisine Parrakeet (Zuphema pulchella). Deposited.
2 Undulated Grass-Parrakeets (Melopsittacus wndulata). De-
posited.
7 Crested Ground-Parrakeets (Calopsitta nove-hollandie).
Deposited.
26, 1 Tabuan Parrakeet (Pyrrhulopsis tabuensis), Purchased.
1 Masked Parrakeet (Pyrrhulopsis personata). Purchased.
2 Barbel (Barbus vulgaris). Presented by G. H. Jones, Esq.,
F.Z.5.
8 Mocassin Snakes ( Tropidonotus fasciatus). Born in the Mena-
gerie.
27. 1 Green Turtle (Chelone viridis). Presented by Capt. J. Smith.
28. 1 Yellow Wagtail (Motacilla flava). Purchased.
29, 1 Black-faced Spider Monkey (Ateles ater). Presented by W.
C. Kenny, Esq.
30. 1 Variable Squirrel (Sciurus variabilis). Purchased.
1. 1.Gommén Fox (Canis vulpes), g. Presented by F. B.
~~ Toogood, Esq.
2. 1 Mona Monkey (Cercopithecus mona), 3. Presented by the
Rev. W. N. Ripley.
1 Chinchilla (Chinchilla lanigera). Born in the Menagerie.
3. 1 Macaque Monkey (Macacus cynomolgus), g. Presented by
the Rey, E. L, Marrett.
4, 1 Macaque Monkey (Macacus cynomolgus), 2. Deposited.
1 Anubis Baboon ( Cynocephalus anubis). Deposited.
1 Kinkajou (Cercoleptes caudivolvulus). Deposited.
1 Sumatran Rhinoceros (Rhinoceros sumatrensis), 2. Deposited.
See P. Z. S. 1878, p. 790.
1 Laughing Kingfisher (Dacelo gigantea). Presented by D. L.
Broughton, Esq.
5, 1 Bonnet-Monkey (Macacus radiatus), 2. Deposited.
1 Macaque Monkey (Macacus cynomolgus), 9. Received in
exchange,
1 Red-beaked Weaver-bird ( Quelea sanguinirostris). Presented
by W. H. Simmonds, Esq.
1 Passerine Owl (Glaucidiwm passerinum). Presented by Miss
Turner.
1 Red-crested Touracou (Corythaix erythrolophus). Presented
by J. Colman, Esq., C.M.Z.S.
1 Cape Zorilla (Ictonyx zorilla). Presented by J. Colman, Esq.
7. 1 Red-sided Eclectus (Eelectus polychlorus), 9. Purchased.
1 Brown Mynah (Acridotheres fuscus). Presented by Capt. J.
Murray.
Proc. Zoou. Soc.—1878, No. LX VI. 66
6.
~
1014
APPENDIX.
Oct. 7. 1 Pied Mynah (Stwrnopastor contra). Presented by Capt. J.
11.
13,
15,
16.
17.
19.
21.
Murray.
1 Indranee Owl (Syrnium indranee). Presented by Capt. J.
Murray.
1 Black-footed Penguin (Spheniscus demersus). Purchased.
. 4 Egyptian Geese (Chenalopex egyptiaca). Presented by Mr.
Justice Denyssen, of the Cape colony.
2 Leopard Tortoises (Testudo pardalis). Presented by the Rey.
G. H. R. Fisk, C.M.Z.S. See P. Z. S. 1878, p. 790.
1 Collared Peccary (Dicotylestajacu). Presented by J. Lloyd, Esq.
1 Common Otter (Letra vulgaris), Presented by W. H.
Baylis, Esq.
4 Chinese Turtledoves (Turtur chinensis). Purchased.
1 Cross Fox (Canis fulvus). Presented by Wilfred G. Marshall,
Esq. From Colorado.
1 Hybrid Mandrill Monkey (between Cynocephalus mormon Q ,
and Macacus cynomolgus 3, large variety). Born in the
Menagerie. See P.Z.S. 1878, p. 791.
1 Indian Muntjac (Cervulus muntjac), 2. Born in the Me-
nagerie.
1 Philantomba Antelope (Cephalophus maxwelli), 2. Received
in exchange.
1 Cinereous Opossum (Didelphys cinerea), Received in exchange.
1 Squirrel Monkey (Saimaris sciwrea), $. Presented by Kd.
althrop, Esq.
1 Michie’s Tufted Deer (Elaphodus cephalophus), 3. Purchased.
See P. Z.8. 1878, p. 790.
2 Ruddy Sheldrakes (Tadorna rutila). Purchased.
1 Naked-throated Bell-hird (Chasmorhynchus nudicollis), 3. Pur-
chased.
3 Blue-bearded Jays (Cyanocorax cyanopogon). Purchased.
From Bahia.
1 Saturnine Mocking-bird (Mimus saturninus). Purchased.
From Pernambuco.
2 Lined Finches (Spermophila lineola). Purchased. From
Pernambuco.
2 Guttural Finches (Spermophila gutturalis). Purchased, From
Pernambuco. ;
1 Pileated Song-Sparrow (Zonotrichia pileata). Purchased.
From Rio de Janeiro.
1 Palm-Tanager (Zanagra palmarum), 9. Purchased. From
Monte Video.
1 Dark-winged Buzzard (Leucopternis scotoptera), Purchased.
From Bahia.
1 Patas Monkey (Cercopithecus ruber). Purchased.
1 Emu (Dromeus nove-hollandie). Presented by C. Hampden
Wigram, Ksq. ,
2 Radiated Tortoises (Testudo radiata). Presented by the Rev.
G. H. R. Fisk, C.M.Z.S.
1 Water-Rail (Rallus aquaticus). Presented by Capt. H. F.
Salvin.
1 Macaque Monkey (Macacus cynomolgus), 3. Presented by
Capt. E. Waterhouse.
1 Macaque Monkey (Macacus cynomolgus), 3. Presented by
Mr. Samuel Thompson.
22.1 Common Seal (Phoca vitulina). Presented by Messrs.
Thompson and Gough.
ae
Oct. 22.
23,
24,
31.
Noy. 1.
2.
ADDITIONS TO THE MENAGERIE. 1015
1 Bornean Fireback (Zuplocamus nobilis), 2. Presented by A.
Dent, Esq. From Northern Borneo.
2 Mandarin Ducks (Aw galericulata), and 2. Presented by
Edward Trelawny, Esq.
1 Collared Fruit-Bat (Cynonycteris collaris). Born in the Me-
nagerie,
1 Common Jackal (Canis aureus). Presented by Capt. Easson.
1 Common Roe (Capreolus caprea), 3. Presented by Edward
Jones, Esq.
1 Mandarin Duck (Aix galericulata), 2. Purchased.
1 Merrem’s Snake (Liophis merremi). Purchased.
1 Plumbeous Snake (Oxyrrhopus plumbeus). Purchased.
1 Black-headed Snake (Homalocranion melanocephalum). Pur-
chased.
1 D’Orbigny’s Snake (Heterodon d’orbignyi). Purchased.
1 Anaconda (Hunectes murinus). Purchased.
1 Tuberculated Lizard (Iguana tuberculata). Purchased.
1 Teguexin Lizard ( Zeius tegueain). Purchased.
1 Common Marmoset (Hapale jacchus). Purchased.
5. 2 Ruddy Sheldrakes (Zadorna rutila). Received in exchange.
» 1 Wood-Owl (Syrnium aluco). Presented by F. W. Jaedecke.
. 1 Grison (Galectis vittata), 9. Presented by H. Potier, Esq.
. 1 Toque Monkey (Macacus pileatus), 3. “Presented by Mrs.
Tranchell.
1 Rhesus Monkey (Macacus erythreus). Deposited.
1 Horse-shoe Snake (Zamenis hippocrepis). Presented by W.
C. Verner, Esq. From Gibraltar.
. 2 South-American Rat-Snakes (Spilotes variabilis). Presented
by G. H. Hawtayne, Esq., C.M.Z.S. From Demerara.
1 Chequered Elaps (Elaps lemniscatus). Pvesented by G. H.
Hawtayne, Esq., C.M.Z.S. From Demerara.
1 Mesopotamian Fallow Deer (Dama mesopotamica), 2. Pur-
chased. See P. Z. 8. 1878, p. 790.
2 Small-scaled Mastigures (Uromastia microlepis), Presented
by Capt. Phillips.
1 Green Monkey ( Cercopithecus callitrichus), §. Presented by
Henry Hands, Esq.
1 Common Boa (Boa constrictor), Presented by W. D. Bell,
Esq.
2 Japanese Pheasants (Phasianus versicolor), g and 2. Pur-
chased. .
1 Gold Pheasant ( Thaumalea picta), $. Purchased.
1 Lineated Pheasant (Euplocamus lineatus), 3. Purchased.
2 Barred-tailed Pheasants (Euplocamus reevesr), ¢ and 2. Pur-
chased.
1 Siamese Pheasant (Euplocamus prelatus), 3. Purchased.
1 White-crested Kaleege (Euplocamus albocristatus), 9. Pur-
chased.
2 Swinhoe’s Pheasants (Euplocamus swinhoii), g and 9. Pur-
chased.
1 Bewick’s Swan (Cygnus bewicki), 9. Purchased.
2 Macaque Monkeys (Macacus cynomolgus), 9. Presented by
Mr. ©. Loveless.
1 Weasel (Mustela vulgaris). Purchased.
1 Three-quarter-bred_ Zebu (between Bos indicus 3, and hy-
brid B. frontalis 2). Born in the ee
Od Or
N
ie}
24.
25,
APPENDIX.
1 Green Tree-Frog (Hyla arborea). Presented by Lord Arthur
Russell, M.P., “EZS.
1 Spotted Salamander (Salamandra maculosa). Presented by
Lord Arthur Russell, M.P., F.Z.i
1 Common Hedgehog (Erinaceus eur opaus). Presented by Mr.
E. Sass.
3 Green Tree-Frogs (/yla arborea). Presented by Mr. E. L.
Boyd.
. 1 Sykes’s Oriole (Oriolus kundoo). Received in exchange.
. 2 Squirrel-like Phalangers ichdlets sciureus). Presented by
K. S. Waller, Esq.
. 1 Blue-and- -yellow Macaw (Ara macao). Deposited.
1 Yellow-shouldered Amazon (Chrysotis ochroptera). Depo-
sited.
. 2 Arabian Baboons (Cynocephalus hamadryas), g and 2. Pre-
sented by C. Wood, Esq.
1 Burchell’s Zebra (Equus burcheili), 2. Received in exchange.
1 Sulphury Seed-eater (Crithagra sulphurata). Purchased.
2 Hooded Crows (Corvus cornix). Presented by Capt. F. H.
Salvin.
. 1 Nisnas Monkey (Cercopithecus pyrrhonotus). Purchased.
1 Red-fronted Lemur (Lemur rujifrons). Purchased.
1 Mantchurian Deer ( Cervus mantchuricus), 9. Purchased.
1 American Tantalus ( Tantalus loculator). Purchased.
2 Hudson-Bay Squirrels (Seirus hudsonius). Purchased.
. 1 Woodcock (Scolopax rusticola). Purchased.
1 Sun-Bittern (Lurypyga helias). Deposited.
. 2 Egyptian Jerboas (Dipus egyptius). Purchased.
1 Cananina Snake (Spilotes pecilostoma). Purchased.
1 Bonnet-Monkey (Muacacus radiatus), §. Presented by Mr.
F, Hinde.
. 2 Horsfield’s Tortoises (Zestudo horsfieldi). Presented by Dr. A
Strauch, C.M.Z.S.. From Turkestan. See P. Z.8. 1878,
p- 975.
. 1 Wanderoo Monkey (Macacus silenus), 9. Deposited.
. 1 Green Monkey (Cercopithecus callitrichus), Q. Presented by
Miss G. E. Marryat.
1Spix’s Macaw (Ara spizi). Purchased. See P. Z.S. 1878,
976.
Fae Maveieeit Monkey (Macacus cynomolgus). pepasyed:
. 1 Macaque Monkey (Macacus cynomolgus).
resented by T
Beck
Na Yellow-bill ed Grosheaks (Pheucticus chrysogaster), g and 9.
Purchased. From Cuzco.
1 Black-crested Cardinal (Gubernatria cristatella). Purchased.
2 Red-crested Cardinals (Paroaria cucullata). Purchased.
1 Equine Antelope (Hippotragus equnus), g. Received in
exchange.
1 Brazilian Tree-Porcupine (Sphingurus prehensilis), §. Pre-
sented by Dr. J. F. Chittenden, C.M.Z.S. From inidaad.
1 Hybrid Vulturine Guinea-fowl (between inte vulturina
and IV. meleagris). Presented by Mons. J. M. Cornély,
C.M.ZS.
1 Red Kangaroo (Macropus one ©. Born in the Menagerie.
1 Black Lemur (Lemur macaco), ¢. Purchased.
. 1 Water-Rail (7tallus aquaticus). Presented by W. Thompson,
Esq.
Sige 75,
Noy. 26.
28
29
ice)
11.
14,
ADDITIONS TO THE MENAGERIE. 1017
1 Macaque Monkey (Macacus cynomolgus), 2. Presented by
Dr. Whately, R.N.
- 2 Common Cormorants (Phalacrocorag carbo). Presented by
F. Buckland, Esq., F.Z.S.
- 1 Common Peafowl (white variety) (Pavo cristatus), g. Pre-
sented by Mrs. Russ,
. 1 Rufous-vented Guan (Penelope cristata). Purchased.
3 Snow-Buntings ( Plectrophanes nivalis), Purchased.
1 Kinkajou (Cercoleptes caudivolvulus). Purchased.
- 2 Black-faced Spider Monkeys (Ateles ater), S and Q. Pre-
sented by Mr. A. Gonzalez Carazo.
2 Rufous-vented Guans (Penelope cristata). Presented by Mr.
A. Gonzalez Carazo. :
2 Horsfield’s Tortoises (Testudo horsfieldi), Presented by Dr.
A. Strauch, C.M.Z.S. From Turkestan.
- 1 Green Monkey (Cercopithecus callitrichus). Presented by Mr.
A. G, Lytton Squires.
2 Black-eared Marmosets (Hapale penicilata). Presented by
the Countess of Cottenham.
2 Laughing Kingfishers (Dacelo gigantea). Presented by Mr.
Edwin A. B. Crochett.
1 Rhomb-marked Snake ( Psammophylax rhombeatus), Pre-
sented by the Rev. G. H. R. Fisk, C.M.Z.S.
% Rufescent Snakes (Leptodira rufescens). Presented by the
Rey. G. H. R. Fisk, C.M.Z.S.
1 Aurora Snake (Zamprophis aurora). Presented by the Rev.
G. H. R. Fisk, C.M.Z.S.
- 1 Ceylon Jungle-Fow! (Gallus stanleyi), 2. Presented by Mr.
G. Lyon Bennett.
2 Ring-necked Pheasants (Phasianus torquatus), Q. Pre-
sented by Mr. G. Lyon Bennett.
1 Grey Francolin (Francolinus ponticerianus). Presented by
Mr. G. Lyon Bennett.
. 1 Mayotte Lemur (Lemur mayottensis), 3. Purchased.
1 Ocelot (Felis pardalis), Purchased.
1 Globose Curassow (Crax globicera), Purchased.
3 Yellow-winged Blue Creepers (Ceereba cyanea). Purchased.
1 Common Swan (Cygnus olor). Presented by Capt. Marx.
1 Yellow Baboon (Cynocephalus babouin), 3. Esssaited by
My. G, A. Shaw.
2 Ring-tailed Lemurs (Lemur catta), gd and 9. Presented by
Mr. G. A. Shaw. From Betsileo, Madagascar.
1 Ring-tailed Lemur (Lemur catta), 3. Deposited.
1 Broad-nosed Lemur (Hapalolemur stmus). From Betsileo,
Madagascar. See P.Z. 8. 1879, p. 2.
1 Brown Mouse Lemur ( Chetrogaleus mitii ?). From Betsileo,
Madagascar. See P,Z.S, 1879, p. 2.
9 Smith’s Dwarf Lemurs (Mierocebus snuthii). From Betsileo,
Madagascar. See P.Z.S. 1879, p. 2.
1 Common Fox (Canis vulpes), 3. Presented by Sutton Sharpe,
Esq.
1 WabHooelt (Scolopax rustieula). Presented by Messrs. E. and
W. H. Davis.
1 Green Monkey (Cercopithecus callitrichus),?. Presented by
J. Williams, Esq.
1 Ceropsis Goose ( Cercopsis nove-hollandie). Purchased.
17.
18,
APPENDIX.
. 1 Pine-Martin (Martes abietum), 2. Presented by Robert
Walters, Esq., F.Z.S.
1 Ring-hals Snake (Sepedon hemachates). Presented by the
Rev. G. H. R. Fisk, C.M.Z.8.
1 Hissing Sand-Snake (Psammophis sibilans). Presented by the
Rey. G. H.R. Fisk, C.M.Z.S,
2 Green-cheeked Amazonsi( Chrysotis viridigenalis). Purchased.
2 Guatemalan Amazons (Chrysotis guatemale). Purchased.
1 Golden-naped Amazon (Chrysotis auwipalliata). Purchased.
1 Crowned Eagle (Spizaetus coronatus). Presented by J. da
Costa Andrade, Esq.
. 1 Grey Ichneumon (Herpestes griseus). Presented by J. B.
Jameson, Esq.
.. | Macaque Monkey (Macacus cynomolgus), 3. Deposited.
. 1 Common Barn-Owl (Strix jflammea). Presented by W.
Davies, Esq.
. 2Philantomba Antelopes (Cephalophus maaxwelli), § and 2.
Purchased.
2 Egyptian Jerboas (Dipus egyptius). Purchased.
. 1 White-whiskered Paradoxure (Paradoxurus leucomystax).
Presented by G. Wilson, Esq.
1 Common Coot (Fulica atra). Presented by T. H. O'Donoghue,
Ksq.
. 8 River-Jack Vipers (Vipera rhinoceros). Received in exchange.
. 6 Brown Tritons (Geotriton fuscus), Presented by Prof. Giglioli,
C.M.Z.S. From Italy.
. 1 Feline Douroucouli (Nyctipithecus felinus), §. Purchased.
1 Punjaub Wild Sheep (Ovzs cycloceros), 9. Presented by Col.
W. R. Alexander. From Upper Sind. See P.Z.8. 1879,
p: 2.
Abisara
angulata, 833, 859.
bifasciata, 833.
kausambi, 833.
neophron, 833, 855.
Acanthia
lectularia, 462.
Acanthias
blainvillei, 117.
Accipiter
etorques, 92.
poliocephalus, 91.
stevensoni, 936, 938,
939.
virgatus, 938, 939.
Achzxa
cyllota, 488.
indistincta, 488.
melicerte, 298.
Achlyodes
potrillo, 483,
Acidalia
apparitaria, 491.
attentata, 852, 858.
umbilicata, 491.
Aclesia
glauca, 277.
Acrocephalus
fasciolatus, 430.
insularis, 430.
Actenoides
hombroni, 944.
Actitis
hypoleuca, 138, 359.
Adamastor
cinereus, 737.
Adelarus
belcheri, 182.
heermanni, 183.
hemprichii, 193.
leucophthalmus, 193.
INDEX.
Adelarus
melanurus, 183.
neptunus, 184,
Adelpha
cocala, 368.
diocles, 270.
falcata, 270.
iphicla, 270, 368.
syma, 271.
Adolias
boisduvali, 639.
discispilota, 831, 854.
858.
iva, 851.
Jahnu, 831, 854.
kesava, 831.
lepideg 830, 854.
parvata, 831, 854, 858,
satropaces, 830, 854.
sedeva, 831.
taooana, 831, 854.
JEgialitis
cantiana, 624, 711.
dubia, 344, 711.
geoffroyt, 82.
inornata, 121, 127.
peronti, 711, 344.
sancte-helene, 445,
460.
varius, 445.
ABgithina
scapularis, 619.
/&gotheles
wallacei, 94.
/Elurcedus
arfakianus, 99.
melanotis, 99.
/Hmene
guttulosana, 34.
maculifascia, 33, 37.
modesta, 34.
sinuata, 34, 37.
sordida, 34.
subcinerea, 34.
taprobanis, 34.
/Hmene
tenebrosa, 84.
Atmona ;
lena, 827, 854.
Alpyceros
melampus, 741.
peverst, 741.
/Xthalochroa
ashmoliana, 584.
Aithopyga
bella, 112, 113.
dubia, 107, 112.
magnifica, 287.
shelleyi, 621.
Aganisthos
orion, 480.
Aganopis
subquadrata, 847.
Agathodes
monstralis, 493.
era
chrysopteryx, 609.
leticolor, 609.
panamensis, 609.
Agrametra
ethiops, 462, 468, 477.
Alauda
celivox, 710.
watterst, 708, 710.
Alcedo
bengalensis, 341, 578,
709, 943.
diops, 79.
moluccensis, 671.
Alces
americanus, 916.
machlis, 887, 915, 916.
letis
druryi, 386.
Sascelis, 386.
helcita, 386.
libyssa, 386.
macularia, 886.
tenuis, 385.
variabilis, 386.
1020
Alexia
meridionalis, 869.
oa
rhodophea, 382.
Alpenus
purus, 382.
Amadina
sharpii, 130.
Amalda
marginata, 865.
Amalthea
australis, 818.
Amara
ambigena, 716.
bamidunye, 716.
Amathina
tricarinata, 868.
Amathitis
badiola, 717.
kuenlunensis, 717.
Amathusia
ganescha, 828.
phidippus, 638, 826,
854.
Amaurornis
olivacea, 845.
Amazilia
cinnamomea, 71.
Amblynura
kleinschmidti, 440.
Amblypodia
ameria, 703.
anita, 855.
epimuta, 835.
hypomuta, 835.
lohita, '702.
narada, 703, 835.
quercetorum, 836.
taooana, 835, 855.
Ambulyx
strigilis, 483.
substrigilis, 844, 856.
Ammonites
dilatatus, 956.
diseus, 956.
falcifer, 962.
goliathus, 959.
lingulatus, 960, 962,
963.
obtusus, 959.
steraspis, 961.
subradiatus, 962, 963.
walcotti, 960.
Amphidasys
arnobia, 490.
Amphonyx
eluentius, 483.
Amynthia
merula, 481.
Anabas
scandens, 216.
Ana
INDEX.
Anabazenops
variegaticeps, 59.
Anzeretes
Sernandezianus, 432.
parulus, 432.
gnia
subfasciata, 847, 857.
Anartia
jatrophe, 480.
Anas
boschas, 350,
obscura, 350.
superciliosa, 350.
wyvilliana, 350.
| Anceryx
fasciata, 483.
Anchista
migratoria, 787.
| Anchomenus
concisus, 594.
ladakensis, 718.
nuntezume, 593.
nugax, 594.
politissimus, 719.
seutifer, 594.
simplicior, 595.
suffectus, 594.
transpunctatus, 593.
viestriatus, 594.
Ancillaria
marginata, 865.
Anerastia
ignobilis, 4946
Anisodactylus
rotundangulus, 589.
Anoa
depressicornis, 792,881,
882, 903,
Anomalocera
huamel, 924.
Anomia
caput-serpentis, 401.
decollata, 409.
detruncata, 409.
ephippium, 41}.
pera, 410.
rostrum-psittact, 413.
scobenata, 410.
terebratula, 407.
truncata, 411.
turbinata, 415.
vitrea, 403.
Anous
albivitta, 273.
albivittatus, 212.
ceruleus, 211, 212, 271,
272, 273.
cinereus, 211, 212,
272.
gracilis, 272.
leucocapillus, 273.
| Anou s
parvulus, 211, 272.
stolidus,
tereticollis, 273.
Antennarius
multiocellatus, 222.
| Antherza
billitonensis, 642.
Srithii, 642.
Anthocephala
castaneiventris, 69.
Anthodieta
zambesiana, 356.
Anthothreptus
chlorogaster, 287, 951.
griseigularis, 113.
Anthreptes
chlorigaster, 951.
malaccensis, 621.
Anthus
chit, 390, 391.
maculatus, 430, 619.
gustavi, 430.
peruvianus, 390, 391.
raaltent, 356.
rufus, 391.
Antilope
semmerringii, 723.
Apamea
intermittens, 486.
Apenes
comis, 606.
a
meridionalis, 819, 821.
Aphnzus
lohita, '702, 707.
Aplonis
tavuensis, 352.
vitiensis, 352.
Appias
amasene, 700, 707.
amba, 839, 855.
copia, 700, 707.
dapha, 888, 855.
formosana, 701.
hippo, 700.
inornata, 700, 707.
lea, 838, 855.
nama, 838, 855.
poeyt, 481.
vacans, 700, 839, 855.
zelmira, 700, 707, 839,
855.
zeuzippe, 838.
Aptenodytes
chrysocome, 654.
longirostris, 653.
magellanicus, 635.
papuus, 658.
Apterodyta
longirostris, 658,
Ara
glauca, 976.
hyacinthina, 976.
macao, 38, 46.
militaris, 46.
spixt, 976.
Arachnechthra
Jugularis, 112, 286,
343.
Arachnothera
dilutior, 621.
flammifera, 840, 343.
Aramides
calopterus, 439.
saracura, 440.
Area
(Scapharea) smyristica,
820.
Archimantis
armata, 583, 584, 587.
monstrosa, 588, 587.
Arctictis
binturong, 142.
Ardea
episcopus, 953.
javanica, 83.
sacra, 271.
Ardetta
cinnamomea, 845, 953.
Are
marginata, 484.
Argaterma
alticola, 462, 474, 475,
477.
multisignata, 462, 474.
Argiope
biplicata, 410.
capsula, 399, 410,
411.
cistellula, 399, 410.
cuneata, 399, 409, 410,
412.
decollata, 399, 409,
410.
forbesi, 409.
neapolitana, 309, 409,
410.
schrammi, 410.
Argiva
caprimulgus, 848.
‘hieroglyphica, 848.
Argyria
chrysogyrans, 495.
insons, 495.
vestalis, 494.
Arhopala -
nakula, 835, 855.
vikara, 835, 855
Aricoris
gelasine, 364,
myrtis, 364,
INDEX.
Arinia
boucardi, 71.
Aristeus
antennatus, 809.
edwardsianus, 308, 809,
310.
Aristhala
hainana, 705, 708,
Arremon
aurantiirostris, 56.
rujfidorsalis, 56.
Arsacia
ashmoliana, 584.
Arses
insularis, 579.
Artamus
insignis, 671.
leucopygialis, 122.
leucorhynchus, 709,
937.
leucorynus, 283, 342.
Artaxa
varians, 848, 857.
Artena
submira, 849, 857.
Artibeus-
quadrivittatus, 880.
Artocarpus
incisa, 30.
Artona
hainana, 704.
Asio
accipitrinus, 348.
galapagoensis, 348.
Aspergillum
stranget, 869.
Aspidoglossa
rachyderus, 589.
Astacoides
Sranklini, 756.
madagascariensis, 759,
772, 778, 'F75.
Astacopsis
franklinii, 760, 764,
765.
Astacus
angulosus, 754.
brasiliensis, 755, 771.
dauricus, 754.
Jluviatilis, 753, 754,
756, 757, 759, 760
763.
fontinalis, 754.
japonicus, T54.
leptodactylus, 754.
madagascariensis, 752.
pachypus, T54.
pallipes, 754.
pilimanus, 755, 771.
saxatilis, 754.
torrentium, 'Td4.
1021
Astarte
suleata, 403.
Aster
glutinosus, 464.
Astictopterus
diocles, 842, 856.
olivascens, 692.
subfasciatus,
856.
unicolor, 692.
Astur
contumax, 92.
hiogaster, 80.
poliocephalus, 91.
tachiro, 354.
trivirgatus, 340.
Asturinula
meridionalis, 354.
monogrammica, BA,
791.
Atella
phalanta, 699, 706.
sinha, 828, 854.
Athene
capensis, 354.
variegata, 290.
Athrostictus
opalescens, 592.
sericatus, 592.
Athyma
lactaria, 647.
leucothoé, 697, 706.
venilia, 646.
Atolmis
rubricollis, 9.
Atretia
gnomon, 400, 412, 413,
41d.
842,
Atricilla
catesbyi, 194.
megalopterus, 194.
micropterus, 194.
Attacus
cynthia, 388.
insularis, 388.
pryeri, 388.
walkeri, 388,
Attagen
ariel, 650.
Attatha
regalis, 848, 857.
Atticora
cyanolenca, 67.
Attila
torridus, 139.
Atys
cylindrica, 819.
elongata, 819.
exiqua, 869.
Awacorhamphus
ceruleogularis, 47.
1022
Axius
stirhynchus, 778.
Azazia
monstratura, 489.
Barbatula
pusilla, 359,
subsulphurea, 131.
Barsine
cruciata, 29.
flammealis, 28, 37,
flavivenosa, 30, 37.
gloriosa, 29, 37.
inflexa, 29, 37.
lineata, 29.
mactans, 29.
punicea, 29.
Barsinella
mirabilis, 385,
Basileuterus
culicivorus, 52.
mesochrysus, 52.
uropygialis, 52.
Batrachostomus
septimus, 944,
Belideus
ariel, 274.
Bembidium
mexicanum, 603.
(Notaphus) flohri, 602.
( ) placitwm, 602.
(Peryphus) pamirense,
718.
(——) punctulipenne,
718
(—) rogersi, 602.
(——) submaculatum,
603.
Bernicia
antarctica, 436.
Bias
musicus, 129.
Biblis
leucocyma, 826.
Bithia
exclusa, 852, 858.
Bizone
alba, 28.
bellissima, 27, 37.
coccinea, 28, 37.
Blasipus
bridgesi, 183.
crassirostris, 183.
heermanni, 188.
polios, 183.
Blepharis
kuhlii, 586.
Boarmia
delicata, 490.
Bolborhynchus
monachus, 77.
INDEX.
Bolborhynchus
murinus, 77.
rubrirostris, 77.
Bolina
agrotoides, 487.
confirmans, 487.
cunearis, -
evelina, 487.
fasciolaris, 487.
ochrodes, 487.
perpendicularis, 487.
Bomb
mori, 705.
Boreogobius
stuvitzii, 319.
Boroa
punetivaga, 28.
08
americanus, 392.
Botaurus
peciloptilus, 578.
Botys
agavealis, 494,
butyrosa, 493.
caldusalis, 850, 858.
campalis, 494,
eratalis, 494.
eurytalis, 494.
gastralis, 494.
glaucusalis, 494.
helcitalis, 494.
jopasalis, 850, 858.
lucilla, 494.
olivia, 494.
plebeialis, 494.
principalis, 494.
unitalis, 850, 858.
vinacealis, 850, 858.
Brachypodius
melanocephalus, 618.
Brachypteryx
malaccensis, 617.
murinus, 617.
Brachypus
cinereifrons, 611, 617.
plumosus, 618.
Brachyurus
steerti, 949.
Bradycellus
lucidus, 592.
subobsoletus, 593.
Bradyornis
pallidus, 357.
subalaris, 357.
Bradytus
compactus, 717.
Broderipus
acrorhynchus, 110, 285,
342, 380, 616, 710,
948.
palawanensis, 711, 616.
Brotogerys
tovi, 46.
Bruchigayia
corallinus, 186.
gouldi, 186.
Jamesonti, 186.
melanorhynchus, 190.
pomare, 186.
Brunia
antica, 15, 36.
Buarremon
brunneinuchus, 56.
capitalis, 55.
castaneiceps, 439.
chrysopogon, 38, 56.
leucopis, 439.
tibialis, 56.
Bubalus
seligniceros, 882.
Bubo
orientalis, 790.
Bubulcus
coromandus, 624, 712.
Buccinulus
afinis, 862.
intermedius, 862, 864,
869.
Buceinum
albescens, 809.
blainvillei, 811.
cancellatum, 811.
globosum, 809.
horridum, 810.
jacksonianum, 865.
lactewm, 865.
margaritiferum, 808.
Buceros
hydrocorax, 277, 278.
melanoleucus, 358.
mindanensis, 277, 278,
946.
ruficollis, 671, 672.
semigaleatus, 279, 280,
339, 341, 379, 380.
Buchanga
assimilis, 357.
cineracea, 615.
leucophea, 615,
Budytes
viridis, 82, 710, 950.
Bufo
melanostictus, "722.
vulgaris, 977.
ulimus
balstoni, 811.
clavator, 311.
iodostylus, '74.
trazuensis, 73, 74.
Josephus, 78, 74.
moreleti, 311.
navarrensis, 73, 74.
Bulimus
obtusatus, 311.
watersi, 311, 314.
(Eurytus) eros,
314.
(Otostomus) xapo, 312,
314
ae) quadrifasciatus,
312,
312, 314.
Bulla
lignaria, 960.
solida, 819.
Bullia
persica, 730, 733.
ursa
concinna, 815
Butalis
grisola, 129.
latirostris, 280, 284.
Butastur
indicus, 612, 709.
Buteo
desertorum, 349.
japonicus, 579.
latissimus, 44.
pennsylvanicus, 44.
solitarius, 346, 348,
350.
vulgaris, 349.
Butorides
javanica, 83, 114, 345,
624.
Cacatua
hematuropygia,
107, |
281, 340, 379, 937. |
Cacomantis
flabelliformis, 125.
insperatus, 671.
merulinus, 613, 709.
sepulchralis, 936, 945.
Cadulus
acuminatus, 868.
Calamoherpe
fumigata, 480.
subflavescens, 430.
Calesia
comesa, 849.
flabellifera, 849, 857.
gastropachoides, 849,
857
Calialeyon
coromanda, 944.
Calisto
zangis, 480.
Calleida
jansoni, 608.
letipennis, 608.
semirubra, 608.
Calliana
pieridoides, 687, 694.
INDEX.
Callidryas
catilla, 640, 788,
855,
crocale, 837, 855.
senne, 481
Calligrapha
violaceo-maculata, 147,
Callimorpha
jacobee, 484.
Calliscotus
bowreyi, 489.
Calliste
dowii, 54,
francisce, 5A.
Srantziti, 54,
gyroloides, 5A.
zeterocephala, 5+.
larvata, 54.
Callorhynchus
antarcticus, 117.
Calobates
melanope, 343.
Calocoris
(Megaceelum) lustra-
tus, 461, 466.
Calcenas
luteovirens, 565.
nicobarica, 289.
Calophzxna
cruciata, 605.
levigata, 605.
Calornis
amboinensis, 88.
chalybea, 622.
gularis, 88, 89.
metallica, 88, 89, 97.
nitida, 88, 289.
panayensis, 113, 287,
343, 622, 710, 951.
purpurascens, 88, 89.
viridescens, 88.
Calpenia
khasiana, 5.
Camarhynchus
cinereus, 137.
Campephaga
melanops, 95.
Campephilus
magellanicus, 434.
Campias
sedulus, 140.
Cam pylopterus
henileucurus, 68.
Campylorhynchus
zonatus, 51.
Cancer
pagurus, 785, 784.
(Astacus) capensis, 754,
755.
Cancros
fluviatilis, 758.
1023
Canis
antarcticus, 376.
anthus, 376.
aureus, 376.
azare, 376.
cancrivorus, 376.
famelicus, 376, 392.
Jamiliaris, 373, 374,
376
376.
fulvus, 376.
hodophylax, 116, 788.
lagopus, 376.
laniger, 376.
latrans, 789.
lupus, 116, 374, 376,
788.
vulpes, 376,
Capissa
auriflava, 19.
JSasciata, 20, 36.
flavens, 20.
innotata, 19, 36.
insolita, 20.
pallens, 19, 36.
vagesa, 19.
Capito
bourciert, 47,
richardsoni, 141.
steerit, 140.
Capnodes
calida, 490.
Capreolus
caprea, 887, 917,
928.
europeus, 917.
leucotis, 924.
pygargus, 887, 917.
Capenaitana
brachyurus, 94, 95.
europ@us, 249.
manillensis, 944.
Carabus
stoliczkanus, 713.
Caranx
hippos, 218.
Carbo
albiventer, 652.
Cardiastethus
bicolor, 461, 466.
testaceus, 466.
Cardium
hystrix, 819.
pulchellum, 870.
pulchrum, 819.
(Levicardium) aus-
trale, 819.
Cariacus
antisiensis, 924.
campestris, 923.
chilensis, 923, 924.
columbianus, 887, 921.
1024
Cariacus
gymnotis, 887, 921.
leucurus, 887, 919, 920.
macrotis, 887, 919, 921.
mexicanus, 887, 19,
1
nemorivagus, 925, 926.
paludosus, 922.
peruvianus, 887, 920.
punctulatus, 922.
rujinus, 925, 926.
rufus, 925, 926.
savannarum, 887, 920.
similis, 887, 920.
simplicicornis, 925.
superciliaris, 926.
toltecus, 887, 921.
virginianus, 887, 918,
9195950: on:
whiteleyi, 926.
(Blastocerus) campes-
tris, 887, 925.
( ) paludosus, 887,
922,
(Coassus) nemorivagus,
887.
( ) rufinus, 887.
( ) rufus, 887, 924.
(——) simplicicornis,
87.
(——) supereiliaris,
( ) whitelyi, 887.
(Furcifer) wntisiensis,
887, 925.
(——) chilensis, 887.
Carollia
brevicauda, 880.
Carpodectes
nitidus, 39, 40, 65.
Carpophaga
aie 113, 288, 344,
628, 982.
assimilis, 126, 574.
bernsteini, 573:
chalybea, 85.
chalybura, 85.
concinna, 8d, 89.
formosa, 572.
latrans, 102, 103, 104,
105.
leclancheri, 568.
magnifica, 575.
melanochroa, 671, 672.
pacifica, 104, 274,
pistinaria, 672.
puella, 574.
roscinucha, 8.
rubricera,. 289, 549,
671.
tumida, 549.
INDEX,
Carpophaga
ieaby aR 289, 671.
zoee, 99.
Casnonia
tubulifera, 604.
Cassicus
prevosti, 58.
Castalius
roxus, 838, 855.
Casuarius
australis, 214.
becearii, 214.
bennetti, 390.
edwardsi, 389, 390.
galeatus, 80, 214.
kaupi, 389.
papuanus, 389, 390.
salvadorit, 218.
tricarunculatus, 214.
wniappendiculatus, 389,
390.
westermanni, 389.
Casyapa
thraz, 641.
Catarractes
chrysocome, 654.
chrysolophus, 654.
Catascopus
angulicollis, 605.
chontalensis, 605.
validus, 605.
Catharistes
atratus, 45.
Cathartes
aura, 435.
papa, 45.
Catharus
Srantati, 40, 50.
gracilirostris, 40, 50.
melpomene, 38, 50.
Catophaga
lagela, 838, 855, 858.
neombo, 838, 855.
paulina, 838, 855.
Catopsilia
gnoma, 699, 707.
Ceblepyris
karu, 87.
plumbea, 95, 96.
rupfwentris, 87.
sumatrensis, 614.
Celia
costaricensis, 600.
Cenchris
contortria, 631.
Centrites
niger, 432.
Centrococeyx
eurycercus, 614.
viridis, 283, 341, 709,
946.
Centropus
ateralbus, 289, 290,
671.
eurycercus, 614,
phasianus, 125.
superciliosus, 359.
Centurus
hoffmanni, 49.
pucherani, 49.
Cephalophus
anchiete, 743.
burchellii,.743.
grimmius, T48.
longiceps, "744.
melanorrhous, 743, 744.
monticola, 743.
niger, T44.
pluto, 744.
ruficrista, 744.
Cephalopterus
glabricollis, 66.
Cephalotes
minor, 875.
peronti, 316, 875.
Cerchneis
sparveria, 45.
Cercocebus
fuliginosus, 791.
Cercomacra '
tyrannina, 61.
Cerithium
(Vertagus) Kochi, 817,
821
(
821
Cerostoma
vittella, 495.
Cerotoma
nigrofasciata, 996.
Certhia
malaccensis, 621.
sanguinea, 347.
Cervulus
amostylis, 899.
aureus, 899.
lacrymans, 887, 899.
muntjac, 887, 899.
reevesit, 887, 899.
sclateri, 899.
tumulicus, 899.
Cervus
affinis, 887, 913.
alfredi, 902.
antisiensis, 924.
aristotelis, 900, 901.
axis, 907, 909.
barbarus, 910, 911.
campestris, 923.
canadensis, 887,
897, 912, 913.
capreolus, 917.
.
turritum, 817,
885,
Cervus
cashmeerianus, 887,912,
918.
caspicus, 909.
chilensis, 928, 924.
columbianus, 921, 922.
cusanus, 891.
eylindricornis, 891.
dama, 913, 915.
davidianus, 906.
duvaucellii, 905.
dybowskii, 909.
elaphus, 887, 891, 910,
911, 912, 928.
eldi, 906.
equinus, 900, 901.
euopis, 908.
eustephanus, 887, 911,
912.
gymnotis, 920, 921.
hippelaphus, 903, 904.
hortulorum, 908.
humilis, 927.
kopschi, 909.
kuhlii, 902.
lepidus, 902.
leucurus, 919.
lewisit, 922
macrotis, 921,
mandarinus, 908.
mantchuricus,
909.
minor, 909.
maral, 887, 911, 912.
mariannus, 901.
matheroni, 891.
mesopotamicus,
914, 915.
mexicanus, 919, 921.
moluccensis, 903, 904.
muntjac, 898, 899.
nemoralis, 919.
nemorivagus, 925,
nigricans, 902.
paludosus, 922.
peronit, 904,
perriert, 891.
philippinus, 901.
porcinus, 902.
pseudaxis, 908, 909.
pudu, eae a
argus
Poa 899,
rufinus, 925,
rufus, 925.
rusa, 903, 904.
savannarum, 920.
schomburgki, 905.
sika, 908.
similis, 920.
simplicicornis, 925.
908,
791,
|
INDEX,
Cervus
swinhowi, 901.
taévanus, 909,
tarandus, 928.
tetracerus, 891.
timoriensis, 903, 904.
toltecus, 921.
virginianus, 919.
wallichii, 918.
wanthopygus, 887, 911.
(Axis) axis, 887,
(Dama) dama, 887,
914.
(——) mesopotamicus,
887, 914.
(Elaphurus) davidia-
nus, 906.
(Pseudaxis) caspicus,
887.
(——) dybowskii, 887.
(——) euopis, 887.
(——) kopsehi, 887.
ere, mantchuricus,
( ss 887, 908.
( ) taévanus, 887.
(Rucervus) duvaucelli, |
887, 908.
(——) eldi, 887.
(——) schomburgki,
887.
(Rusa) alfredi, 887.
(——) aristotelis, 887,
901.
(ae) equinus, 887.
(——) hippelaphus,
887, 903.
(——) kuhlit, 887.
(——) lepidus, 887.
(—-) mariannus, 887.
(——) nigricans, 887.
(——) philippinus,
887.
(——) porcinus, 887.
( ) swinhoii, 887.
Ceryle
cabanisi, 48.
rudis, 35D.
stellata, 484.
Cestracion
philippi, 117.
Cethosia
biblis, 699, 707.
chrysippe, 645.
Cetosparactes
eburneus, 162.
Cettia
fortipes, 980.
Ceyx
argentata,
341, 948.
107, 108,
|
|
|
1025
| Ceyx
lepida, 81.
Chaca
lophioides, 220.
Cheerocampa
chiron, 483.
suffusa, 642.
Cherodes
transtincta, 490,
Cheetopterus
norvegicus, 338.
Cheetura
picina, 936, 944.
Chaleocoecyx
malayanus, 945,
Chaleophana
costatipennis, 144,
discolor, 144.
rufipennis, 144.
semirufa, 988, 989.
uniformis, 989.
Chaleophaps
indica, 953.
stephani, 289, 671.
Chaleopsitta
rubrifrons, 93.
Chalcopsittacus
seintillatus, 93.
| Chalcosia
diana, 704.
nympha, 704, 708.
Chaleostetha
insignis, 621.
| Chalybe
belti, G04.
Chamepelia
passerina, 43.
rufipennis, 43.
Chameepetes
unicolor, 42,
Charadrius
cantianus, 624.
fulvus, 82, 344, 711,
937.
geoffroyi, 82.
peronit, 544, 711,
veredus, 82.
Charaxes
agna, 832, 854.
athamas, 831.
desa, 832, 854.
harpax, 832, 854.
lunawara, 832.
marmax, 831, 832,
854.
samatha, 831, 854
schreiberi, 640.
Charis
crocea, 364,
holostieta, 364.
iris, 364,
1026
Charis
ochrias, 365.
perone, 364, 365.
peciloptera, 365.
sulphurea, 365.
velutina, 364.
Chasiempis
sandvicensis, 346.
Chasmorhynchus
tricarunculatus, 40,
6
Cheimonea
tridactylus, 164.
Chelys
matamata, 378,
Chibia
assimilis, 96.
hottentota, 88, 614.
megalornis, 88.
Chilobothrus
inornatus, 789.
Chilomenes
lunata, 448.
Chiloscyllium
ocellatum, 117.
Chione
(Marcia) levigata,
870.
Chionis
minor, 578.
Chiromacheris
cande@i, 66
Chiroxiphia
linearis, 66.
Chiton
siculoides, 868.
Chlamydophorus
truncatus, 228, 673,
677, 679.
Chloéphaga
poliocephala, 436.
Chlorita
edithe, 462, 476.
Chlorolampis
caniveti, 71,
osberti, 38.
salvini, 71.
sedulus, 140.
Chlorophanes
guatemalensis, 54.
spiza, 54.
Chlorophonia
callophrys, 39, 54.
Chlorospingus
alhitemporalis, 55.
Chroicocephalus
atricilla, 194.
bonapartii, 207.
brunneicephalus, 198.
brunniceps, 198.
capistratus, 200.
INDEX,
| Chroicocephalus
cirrhocephalus, 204.
cucullatus, 195.
Franklini, 195.
gelastes, 193.
glaucodes, 204,
glaucotes, 203.
whthyaétus, 198,
kitlitzii, 195, 205.
leucophthalmus, 193.
maculipennis, 202.
melanocephalus, 200,
minutus, 206.
personatus, 196.
philadelphia, 207.
pileatus, 200.
plumbiceps, 204.
ridibundus, 200.
saundersi, 205.
schimperi, 195.
serranus, 194.
subulirostris, 207.
tibetanus, 198.
Chroicoptera
vidua, 580.
Chryseglia
ferrifasciata, 13, 36.
magnified, 13.
Chrysococcyx
klaasi, 131.
Chrysocolaptes
erythrocephalus, 612.
lucidus, 379, 942.
maculiceps, 942.
Chrysena
luteovirens, 505, 565.
victor, 568,
viridis, 506, 566, 567.
Chrysomitris
bryantit, 56.
Chrysorabdia
viridata, 19, 36.
Chrysotis
Farinosa, 38, 46.
pulverulenta, 46.
Churinga
rufifrons, 10, 36.
Cicindela
belti, 588.
Slohri, 588.
stoliezkana, 718.
Cicinnurus
regius, 98.
Ciconia
alba, 625.
episcopus, 633.
maguari, 633.
Cincia
pallida, 484.
Cinclodes
patachonicus, 433.
Cinnyris
aspasioides, 81.
auriceps, 80.
clementie, 81.
Srenata, 79.
Suliginosa, 129.
gutturalis, 355.
Jardinii, 355.
julia, 951.
microrhyncha, 855.
splendida, 129.
venusta, 130.
zenobia, 81.
Cirrhocephalus
potocephalus, 204.
Cirrochroa
lanka, 827.
suryd, 827, 854.
Cisthene
niveata, 384.
Cisticola
cursitans, ‘708, 710.
grayt, 286
tsodactyla, 356.
ruficeps, 286, 671.
semirufa, 286.
Cittocinela
nigra, 619.
Cixius
sancte-helene, 462.
Clangula
histrionica, 479.
vulgaris, 479.
Clerome
arcesilaus, 827, 854.
eumeus, 697, 706.
Clupea
harengus, 338.
sprattus, 338.
Clupeilarus
antipodum, 181.
Suscus, 173.
Clytanthus
ignobilis, 721.
Coassus
auritus, 926.
peruvianus, 920.
superciliaris, 926.
whitelyi, 926.
Coccystes
coromandus, 936,
946.
Jacobinus, 359,
Coccyzus
minor, 47.
Ceeligena
clemencia, 68.
Cceelomera
atro-cerulea, 152.
Ceelops
Srithii, 878.
Cceerostris
avernalis, 799.
mitralis, 799, 800.
Colznis
delila, 481.
Colaptes
rupicola, 140,
Colaspoides
viridicollis, 144.
Coleura
seychellensis, 879.
Colius
leucotis, 358.
Oollita
griseola, 16.
lilacina, 16.
parva, 16, 36.
vetusta, 16.
Collocalia
esculenta, 283.
francica, 280, 282, 283.
troglodytes, 283.
Colluricincla
megarhyncha, 96.
parvissima, 122.
rufigaster, 122.
Colpodes
aurotinctus, 598.
chontalensis, 598.
chrysopterus, 599.
cyanostolus, 598.
duplex, 596.
gratus, 596.
intergencus, 596.
lactipes, 597.
lebioides, 599.
obscurellus, 596.
ovaliceps, 719.
parviceps, 597.
princeps, 599.
procephalus, 597.
prolivus, 597
prolongatus, 600.
prostomis, 598.
stricticollis, 595.
superbus, 599.
viridiauratus, 599.
Columba
albilineata, 43.
amarante, 574.
cyanovirens, 542, 559.
diademata, 540.
dupetit-thouarsii, 503,
539.
felicie, 565.
flava, 565.
forsteri, 505, 525.
gularis, 503, 570.
Aypogastra, 558.
INDEX,
| Oolumba
iogastra, 558.
jambu, 502, 554.
kurukuru, 502, 525,
541
purpuro-leucoce-
phalus, 539.
superba, 545,
leucotis, 505.
luteovirens, 565,
magnifica, 575.
melanocephala, 502,
551, 552, 951.
monacha, 550.
nana, 549.
occipitalis, 695.
enas, 437,
oopa, 502, 527, 541.
pectoralis, 559.
perlata, 557.
porphyracea, 525, 526.
orpyrea, 525, 553.
puella, 502, 574.
pulchella, 551.
purpurata, 502, 507,
25, 526, 528, 529,
531, 582, 533, 534,
540.
regina, 502.
rivoli, 503, 561.
roseicapilla, 537.
roseicollis, 554.
subvinacea, 43.
superba, 542, 548,
ix
544.
taitensis, 502, 541.
virens, 559.
viridis, 502, 560.
viridissima, 535.
zanthogaster, 502, 540.
zoée, 99.
Columbella
terpsichore, 807.
(Anachus) nigricostata,
807, 821.
(——) puella, 807.
Columbula
campestris, 141,
Columella
angasi, 865.
dermestoides, 865.
interrupta, 865.
lineolata, 865.
Cominella
lactea, 865.
Composia
sybaris,
Conchylodes
diphtheralis, 493.
Oondica
palpalis, 486.
| Contopus
virens, 64.
| Conurus
astec, 38, 46.
hilaris, 75.
hotfmanni, 40, 46.
murinus, 75.
nanday, 77.
patagonus, 75.
smaragdinus, 434.
Conus
andamanensis, 804,821.
articulatus, 796, 798.
carnalis, 796, 798.
ceylanensis, 805,
hepaticus, 732.
japonicus, 796.
largillierti, 796.
lucidus, 795.
mahogani, 797.
melvilhi, 795, 798.
nebulosus, 796.
pastinaca, 731, 732.
quercinus, 732.
scabriusculus, 796.
tabidus, 732.
tornatus, 797.
virgo, 732.
wimines, 797.
Copaxa
expandens, 387.
gemmifera, 387.
Copsychus
mindanensis, 285, 342,
Coptodera
scintillans, 605.
Coracias
caudatus, 354.
Coralliophila
barclayana, 806.
Corbula
fortisulcata, 819, 821.
nasuta, 869.
Corvus
cornix, 976.
corone, 976.
enca, 622.
Javanensis, 622,
orru, 289.
hilippinus, 113, 287,
4 343 381. 710, 951.
pusillus, 611, 622.
validus, 622.
Corydalla
gustavi, 430.
lugubris, 286, 380, 710.
Corymica
arnearia, 851, 858.
Cosmophila
erosa, 486.
1028
Cosmosoma
auge, 484.
tyrrhene, 484.
Cossa
basigera, 21.
brunnea, 22, 37.
pallida, 21.
quadrisignata, 21, 37.
Cossypha
verticalis, 129.
Coturnix
conununis, 578.
Cracticus
cassicus, 97.
Crambomorpha
splendens, 8.
Crania
anomala, 400, 414.
lamellosa, 414.
personata, 414, 415.
ringens, 415.
rostrata, 415,
suessia, 415,
Craspedophora
magnifica, 125.
Crassatella
, radiata, 820.
Crastia
cupreipennis, 825, 853.
modesta, 823.
Crateropus
kirki, 556.
Orax
globicera, 42.
Creagrus
furcatus, 210,
Crenella
harbata, 871.
Creobroter
apicalis, 585.
pictipennis, 585, 587.
urbanus, 58D.
Crepidula
tomentosa, 867.
Criniger
everetti, 110, 380.
frater, 618.
palawanensis,
618.
simplex, 129.
Crioceris
australis, 152.
nigripes, 152.
Criopus
Simbriatus, 414.
orcadensis, 415.
Crithagra
chrysopyga, 130,
insularis, 577.
Crotophaga
sulcirostris, 47.
611,
INDEX,
Cryptospira
cymbalum, 866.
ovulum, 866.
Crypturus
bartletti, 142.
boucardi, 39, 40, 41,
42.
sallei, 142.
transfasciatus, 141.
variegatus, 142.
Crystallogobius
nilssoni, 318, 329, 331,
332, 334, 335, 337,
338.
Ctenolabrus
rupestris, 329.
Otenostoma
leticolor, 588.
sigma, 588.
Cuculus
basalis, 945.
chalcites, 946.
cineraceus, 125.
coromandus, 9AG.
hyperythrus, 946.
malayanus, 945, 946,
merulinus, 945.
sepulchralis, 945,
threnodes, 945.
Culapa
mnasicles, 825, 853.
Cuncuma
leucogaster, 91.
Curzeus
aterrimus, 432.
Curtonotus
pamirensis, 717.
putzeysi, 600.
substriatus, 601.
Cuscus
orientalis, 274, 275.
Cyanaleyon
diops, 79.
macleayt, 122.
Cyanocitta
pulchra, 439.
Cyanocorax
bellus, 138.
mystacalis, 138.
ortoni, 138.
Cyanopolius
cooki, 789.
Cyanospiza
ctris, 57.
cyanea, 57, 58.
Cyanotreron
eyanovirens, H04, 543.
monacha, 504, 550.
Cyclophorus
calyx, 499,
hispidulus, 499.
Cyelophorus
pinnulifer, 499.
(Myxostoma) bathy-
rhaphe, 497, 498.
Cyclopides
camertes, 704, 707.
subradiatus, 693.
subvittatus, 692, 693.
Cyclopsitta
lunulata, 937.
Cyclopsittacus
melanogenys, 92.
Cyclorhis
flavipectus, 88, 53.
subflavescens, 53.
Cyclostoma
crocewm, 31.
planorbulus, 499.
Cyclostrema
tate, 862,
868.
Cyclotus
boucardi, 73, 74.
mexicanus, 73.
864,
| Cyenia
punetivaga, 28.
Cydalima
conchylalis, 850, 857.
Cydosia
nobilitella, 495.
Cylichna
pygmed, 869.
Cymindis
glabrella, 719.
Cynocephalus
gelada, 1.
mormon, 791.
Cynonycteris
amplexicaudata, 877.
brachyotis, 316.
Cynopterus
latidens, 875.
marginatus, 877.
scherzeri, 875.
Cynthia
arsinoé, 689, 646.
deione, 639.
Cyornis
banyumas, 615.
philippinensis, 109,284,
342, 948.
Cypriea
australis, 867.
gangrenosa, 816,
peasei, 731, 733, ~
| Cypselus
mystaceus, 81.
pallidus, 976.
parvus, 131.
Cyrenia
martia, 362.
a
Eo
Cyrenia
pyrippe, 362.
Cyrestis
acilia, 646. |
cocles, 698, 706, 829,
854. .
risa, 829, 854.
thyodamas, 698,
828, 854.
Cyrtostomus
andamanicus, 620.
asiaticus, 620.
aurora, 611, 620.
brevirostris, 620.
flammacillaris, 620.
Srenatus, 79, 97, 620.
Jugularis, 620, 710,
951.
lotenia, 620.
melanogaster, 81.
osea, 620.
pectoralis, 620.
rhizophore, 620.
solaris, 620, 621.
zenobia, 81, 620.
Cystineura
dorcas, 480.
706, |
Debis
INDEX,
Danais
septentrionis,
853.
similis, 637.
tytia, 822, 853.
vulgaris, 822, 853.
Daption
capensis, 737.
Dasycrotapha
822,
speciosa, 114, 280, 281,
285.
Dasypus
minutus, 228, 229.
sexcinetus, 226, 227,
228, 229, 673, 674,
677, 678, 679.
vellerosus, 228, 229.
villosus, 228, 229.
europa, 706.
mekara, 824.
Decelea
bowreyi, 486.
Deilemera
artemis, 386.
pellex, 386.
signata, 386.
+ Deilephila
Dacelo
gaudichaudi, 94.
tyro, 94. |
Dacnis i
venusta, 54. |
Dama
mesopotamica, 887, |
9l4. .
vulgaris, 895, 913.
Damalis
alciphron, 847, 857.
concana, 4.
plaginota, 847, 857.
Damophila
amabilis, 71.
Danais
affinis, 644. .
aglea, 695, 706, 822, |
853.
archippus, 480
chrysippus, 638, 695,
706. 822, 853.
citrina, 6438. |
eleutho, 297.
gloriola, 643.
hegisippus, 638.
jamaicensis, 480.
Juventa, 637.
limniace, 695, 706. |
melaneus, 822, 853.
a
322,853.
lineata, 483.
_ Deiopeia
pulchella, 298.
speciosa, 484.
Deiphobe
laticeps, 580, 587.
ocellata, 580.
Delattria
henrici, 68.
- Delias
descombest, 839, 856.
indica, 839, 856.
inferna, 734.
pasithoé, 839, 856.
Dendrobiastes
basilanicus, 114.
| Dendrochelidon
mystacea, 289, 290.
| Dendrocolaptes
puneticollis, 60.
sancti-thome, 60.
Dendrocygna
autumnalis, 44.
vagans, 346, 712, 954.
Dendreeca
estiva, 52.
blackburnia, 52.
pennsylvanica, 52,
vieilloti, 52.
; Dendropicus
goerte, 131.
Dendrornis
susurrans, 60.
Proc. Zoox. Soc.—1878, No. LXVII.
1029
Dendrortyx
leucophrys, 40, 42.
Dentalium
octagonum, 868.
Desmia
orbalis, 493.
prognealis, 493.
Deudorix
epijarbas, 834.
Jarbas, 834.
melampus, 834.
suffusa, 834, 855, 858.
xenophon, 834.
Diabrotica
costatipennis, 996.
fuscomaculata, 994.
Juscomarginata, 149.
gracilis, 150, 151.
jansoni, 994.
marginata, 151.
mexicana, 149.
multipunctata, 149.
nigrolineata, 149.
nigromaculata, 150.
nigrovittata, 995.
novemmaculata, 995.
peruana, 151.
sexplagiata, 151.
sexpunctata, 148.
tripunctata, 148.
variolosa, 151.
ventricosa, 148.
viridimaculata, 150.
waterhousei, 993.
| Diadema
alimena, 647.
auge, 297.
avia, 699, 706.
nerina, 297, 647.
otaheite, 297.
Diala
imbricata, 867.
picta, 867.
varia, 867.
Dias
longiremis, 330,
| Diceum
cinereigulare, 950.
everett2, 107, 111.
hematostictum, 286.
hirundinaceum, 125.
modestum, 380, 381.
rubriventre, 111, 950.
schistaceum, 107, 111.
vulneratum, 82.
Dichirotrichus
alticola, 713.
Dicelus
flohri, 589.
Dicranostreptus, 671.
megarhynchus, 289.
67
1030
Dicrocercus
hirundinaceus, 355.
Dicruropsis
assimilis, 96.
megalornis, 88.
Dicrurus
assimilis, 96.
balicassius, 614.
divaricatus, 357.
lemostictus, 615.
leucopheus, 615.
leucops, 615.
marginatus, 339,
megalornis, 88.
megarhynchus, 615.
mirabilis, 284, 614.
palawanensis, 611, 614,
615.
pectoralis, 88, 615.
striatus, 109, 380, 614,
948.
Dictyopicus
Jjardinei, 49.
Digama
Jigurata, 5.
hearseyana, 4, 5.
insulana, 5.
similis, 5.
Diglossa
plumobea, 54.
Dilophonota
enotrus, 484.
omphalee, 484.
ello, 483.
Diomedea
brachyura, 740.
demersa, 658.
exulans, 740.
fuliginosa, 740.
melanophrys, 740.
Dione
vanille, 481.
Diphyllodes
gulielmi tertii, 792.
Diploharpus
exstriatus, 601.
Discina
atlantica, , 415,
416. rk:
fallens, 415.
norvegica, 415.
ostreoides, 415.
Discophora
celinde, 827.
necho, 827, 854.
tullia, 826, 854.
zal, 826, 854.
Dissura
episcopus, 633.
maguari, 633.
INDEX
Doleschallia
comrii, 643, 646.
pratipa, 828, 854.
| Dolgoma
angulifera, 20.
brunnea, 20, 37.
reticulata, 20.
Domicella
atricapilla, 79.
eyanauchen, 79.
lory mysorensis, 79.
rubra, 86.
schlegeli, 86.
scintillata, 93.
Dominicanus
antipodum, 180.
antipodus, 180.
azare, 181.
fritzei, 180.
Fuscescens, 173.
Suscus, 173.
marinus, 180.
pelagicus, 180.
verreauxtt, 181.
vetula, 180.
vociferus, 180.
Donax
sordidus, 870.
Dorycha
bryante, 40, 70.
Doryphora
estuans, 992.
brunneipennis, 989.
decorata, 992.
dorsomaculata, 146.
flavipennis, 989.
flavoguttata, 145.
flavomarginata, 146.
hybrida, 991.
imperialis, 991,
insularis, 147.
limbatipennis, 990,
militaris, 991.
ocellata, 991.
porosa, 990.
punctipennis, 145.
vittaticollis, 993.
vittatipennis, 990.
Dreata
petola, 848.
taooensis,
859.
Drepanis
coccinea, 347.
flava, 348.
sanguinea, 347.
Drepanodes
pionaria, 490.
Drepanoptila
|
506, 513.
848, 857, |
holosericea, 504, 505,
Drusilla
catops, 645.
myops, 645.
Drymacedus
beccarii, 91, 97.
Drymocataphus
cinereiceps,
617.
Drymeca
affinis, 356.
brevicaudata,
982.
isodactyla, 356.
Drymophila
cinerascens, 83.
Dryococcyx
harringtoni, 613.
Dryoscopus
affinis, 357.
cubla, 357.
gambensis, 128.
major, 128.
salime, 357.
Dyctis
leucocyma, 826, 853.
Dysithamnus
semicinereus, 60.
striaticeps, 60.
611,
980,
| Dysporus
eyanops, 652.
piscator, 651.
sula, 114, 954.
Echis
earinatus, 977, 978.
coloratus, 977, 978.
Eelectus
grandis, 78.
linnei, 92.
megalorhynchus, 78.
personatus, 86.
polychlorus, 79, 92, 93,
289, 290.
rhodops, 81.
roratus, 78.
Edela
ruficeps, 619.
Edoliosoma
sp. ine., 671.
muelleri, 95.
schisticeps, 96.
tenuirostre, 95.
Hilema
caniola, 17.
Elainea
albiceps, 433.
frantzii, 63.
pagana, 38, 63.
subpagana, 63.
Elanoides
Surcatus, 45.
Elanus
hypoleucus, 939.
Elaphodus
' cephalophus, 790, 887,
899.
michianus, 887, 900.
longicauda, 129.
Elousa
albicans, 486.
Elymnias
hainana, 696, 706.
lais, 638.
nigrescens, 638, 696.
tinctoria, 826, 853.
undularis, 826.
vasudeva, 826, 853.
Emarginula
dilecta, 868.
rugosa, 868.
tasmania, 868.
tenuicostata, 868.
Emballonura
nigrescens, 876, 877.
Embernagra
raffrayana, 876.
striaticeps, 56.
Empidonax
aneaced 64.
Empusa
coronata, 586.
Empyreuma
pugione, 484.
Engina
testacearia, 643.
Entomobia
gularis, 282, 341, 379,
937, 943.
Entomogramma
Sautriz, 849, 857.
Enyo
cyanogenys, 86.
indica, 578.
riciniata, 86.
rubra, 86.
ira
tuberculosa, 799.
Epicrates
angulifer, 789.
Epitelarus
neglectus, 184,
Epomophorus
comptus, 879.
INDEX.
Epomophorus
monstrosus, 879.
Erastria
includens, 486.
Erato
bimaculata, 866.
| Erebus
agarista, 488.
odora, 488.
odorata, 488.
Eresia
drypetis, 269.
eunice, 269.
mechanitis, 269.
Ergolis
alternus, 698, 706.
ariadne, 698.
Erites
angularis, 825, 853.
madura, 825.
Eronia
lutescens, 838, 855.
Erosia
incongrua, 491.
Erythra
phenicura, 953.
Erythraucheena
umeralis, 126.
Erythropitta
erythrogastra, 948.
Kochi, 430.
Estrelda
cyanogastra, 358.
Etroplus
énsulata, 484.
Eudamus
capucinus, 482.
trinitad, 482.
Eudromias
geoffroyi, 281, 288, 344,
24, 711.
co
modestus, 438.
mongolicus, 344, 711.
veredus, 82.
Eudynamis
mindanensis, 946.
picata, 671.
Eudyptes
chrysocome, 577, 654,
655
asp
chrysolophus, 654.
diadematus, 654,
ee = :
pach nchus, 655.
viltator, 654,
Hugenes
falas 68.
1031
| Eugenes
spectabilis, 40, 68.
| Euglyphia
hieroglyphica, 485.
Euglyphis
procopialis, 850,
858.
Eumelea
rosalia, 851, 858.
Eumomota
superciliaris, 49, 100.
Eunectes
murinus, 378.
Eunica
tatila, 480.
Eupherusa
egregia, 71.
Euphonia
crassirostris, 54.
elegantissima, 54.
gouldi, 54.
| Euplagia
hera, 847, 848.
Euplectes
capensis, 358.
Sranciscana, 130.
nigriventris, 358.
Eupleea
alcathoé, 823, 853.
angasii, 297.
bremeri, 638.
callithoé, 644.
core, 823.
crassa, 822,
deione, 823.
distincta, 296, 297.
eleutho, 296, 297.
erimas, 738.
eupator, 733.
eurianassa, 644.
felderi, 695, 706.
godartit, 824, 853.
helcita, 297.
tphianassa, 644.
Jessica, 733.
layardi, 695.
limborgii, 823, 853,
858.
margarita, 823.
ménétriésti, 638.
mulciber, 638.
nox, 644,
philomela, 637.
subdita, 823, 858.
thoosa, 688.
treitschkii, 644,
EHuptoieta
hegesia, 481,
Kuptychia
argentella, 265.
ithama, 266.
10382
Euptychia
philodice, 264, 265.
phocton, 265.
pyracmon, 265.
rogersi, 265.
vetones, 265.
Eurema
formosa, 836.
Eurycoleus
belti, 606.
Eurygona
amphidecta, 361.
hypophea, 360.
inconspicua, 36).
leucorrhoa, 360.
russata, 361.
Euryleemus
javanicus, 143. .
steerti, 110.
Eurystomus
afer, 354.
crassirostris, 671.
orientalis, 108, 282,
341, 379, 613.
Euscarthmus
squamicristatus, 62.
zosterops, 138,
Huschema
aurilimbata, 846, 857.
excubitor, 846, 857.
horsfieldi, 846, 857.
malayana, 846.
militaris, 704, '708, 846,
857.
Eusemia
africana, 381.
incongruens, 381.
Euspiza
americana, 58.
Eustephanus
Ffernandensis, 434.
galeritus, 433.
Euthisanotia
timais, 489.
Falco
babylonicus, 2.
barbarus, 2.
henderson, 262.
hiogaster, 80.
Felis
jubata, 656.
lanea, 655, 656,
leopardus, 239.
uncia, 289.
Fischeria
batica, 580.
laticeps, 580.
ocellata, 580.
Fissurella
scutella, 868,
INDEX.
Fissurellidsea
scutella, 868.
Fistulana
’ elava, 967.
| Fodina
stola, 849, 857.
Formicarius
hoffmanni, 62.
Formicivora
boucardi, 61.
Francolinus
granti, 359.
rovuma, 359.
Fregata
aquila, 273, 650.
minor, 650, 651.
| Fregetta
grallaria, 735, 736.
melanogastra, 736.
Frenula
Jeffreysti, 407.
Fringilla
minuta, 287.
| Fulica
alai, 351.
armillata, 291.
gallinuloides, 291.
leucoptera, 291.
Fuligula
collaris, 479.
marila, 479.
nationt, 477, 478,
479.
rufina, 479.
Funchalia
woodwardi, 298, 309.
Furcifer
chilensis, 924.
Furnarius
Jigulus, 139.
pileatus, 139,
Fusus
abnormis, 811, 821.
Gabianus
bathyrhynchus, 182.
georgti, 182.
pacificus, 181.
Gadus
esmarkti, 330.
merlangus, 330.
morrhua, 330, 338.
Galbula_ -
melanogenia, 47,
Gallicrex
cinerea, 953.
Gallinago
megala, 345, 953.
paraguaia, 438.
scolopacina, 345,
712.
| Gallinula
stenura, 83.
chloropus, 344, 953.
euryzonoides, 288,
nesiotis, 577.
Gallus
bankiva, 953.
Gambetta
pulverulenta, 127.
Gampola
Ffasciata, 27, 37.
Gandhara
serva, 15, 16, 36.
Gasterosteus
aculeatus, 330.
Gastrochena
tasmanica, 869.
Gastropacha
vishnu, 848.
| Gavia
| andersoni, 186, 187.
atricilla, 194.
audouini, 179.
bonapartii, 207.
capistrata, 200.
cirrhocephala, 204.
citrirostris, 164.
eburnea, 162.
gelastes, 192.
glaucotis, 204.
gouldi, 186.
hartlaubi, 188.
Jamesoni, 185.
| kittlitzi, 205.
maculipennis, 202.
melanocephala, 200.
minuta, 206.
nivea, 162.
personata, 196.
pomarre, 185, 186, 187,
190, 191.
ridibunda, 200.
roseiventris, 204.
sabint, 209.
serrana, 196.
subulirostris, 207.
tridactyla, 164.
Gavina
audouini, 179.
bruchi, 177.
cana, 177.
heinei, 177.
kamtschatchensis, 177.
| zonorhynchus, 177.
| Gazella
| dama, 726.
| doreas, 929, 930.
euchore, 741.
granti, 723, 724, 725,
726.
Gazella
mohr, 726.
semmerringi, 726.
walleri, 929, 980.
Gecinus
viridis, 934.
Gegenes
hainanus, 703, 707.
nostrodamus, 703.
Gelastes
andersoni, 186.
columbinus, 192.
corallinus, 186.
gouldi, 186.
lambruschini, 192.
leucocephalus, 192.
rubriventris, 122.
Geoffroius
aruensis, 80.
cyaniceps, 671, 672.
heteroclitus, 671,
672.
keyensis, 86.
pucherani, 81,
rhodops, 80.
schlegeliz, 80, 81.
simplex, 672.
Geopelia
humeralis, 126.
tranquilla, 126.
Geothlypis
poliocephala, 52.
trichas, 52.
Geotrygon
sae belo 43.
chiriquensis, 43.
costaricensis, 39, 40,
montanus, 48.
Geranoaétus
melanoleucus, 434.
Ghoria
albocinerea, 13, 36.
sericeipennis, 13.
Gibbula
pulcherrima, 818.
Ginglymostoma
cirratum, 117.
Gladius
(Rimella) cancellatus,
818.
Qlaucidium
capense, 354.
Glaucus
argentatoides, 167.
argentatus, 167.
audouini, 179.
borealis, 179.
canus, 177.
consul, 165.
glacialis, 166.
| Glissa
| Glyptolenus
Gobiosoma
INDEX,
Glaucus
glaucescens, 167.
glaucopterus, 167.
lacrymosus, 177.
leucopheus, 169.
leucopterus, 166.
michahellesii, 169.
occidentalis, 172, 176.
zonorhynchus, 176.
variegata, 485.
Glycyphila
Jfasciata, 655.
gould, 655.
Glyphodes
bivitralis, 849, 857.
stolalis, 849, 857.
Glyphorhynchus
cuneatus, 60.
rugicollis, 595.
Gnathospiza
raimondit, 137.
Gnophria
ceramensis, 14.
nilssonit, 331.
stuvitzit, 319.
Gobius
albus, 319, 3820, 321,
$22.
linearis, 332.
nilssonit, 331.
i peitak 319, 321,
22.
stuwitzui, 319, 320.
Gonatus |
amenus, 652. |
Goniurus |
catillus, 482.
proteus, 481.
| Gonodonta |
nutria, 486.
Gongylus
ocellatus, 977.
Gracula
javanensis, 622. /
kreffti, 289, 671. '
nobilis, 347.
Grallaria |
perspicillata, 62.
Grammodes |
grandirena, 487.
Graucalus |
caledonicus, 87.
hypoleucus, 122.
melanops, 95.
personatus, 87.
pollens, 87.
striatus, 108, 283, 380, | /
947.
1033
Graucalus
sumatrensis, 614.
temmincki, 87.
Grimmia
splendidula, 743.
Grypotes
insularis, 462,
475.
Guiraca
cinerea, 137.
ygis
candida, 273.
Gymnotus
electricus, 221.
| Gynecia
dirce, 480.
Gynautocera
papilionaria, 846.
Hadena
abida, 486.
Hematopus
ater, 438.
leucopus, 437.
unicolor, 578.
Hemophila
stolzmanni, 137.
| Haleyon
albicilla, 671.
chelicutensis, 354.
diops, 79.
erythrogastra, 577.
macleayi, 122.
pileata, 578.
sancta, 94, 289.
senegalensis, 131.
sordida, 94.
striolata, 354.
winchelli, 944.
| Haliaétus
girrenera, 91.
leucogaster, 91.
leucoryphus, 393.
' Haliastur
girrenera, 91.
intermedius, 282, 840,
709, 987.
Halieus
(Hypoleucus) verruco-
sus, 65:
| Halhotis
varia, 819.
Halpe
betaieay 690, 703.
ceylonica, 690,
95.
dolopia, 690.
luteisquama, 690.
radians, 690, 694.
Hamadryas
zoilus, 644,
1034
Hapa
contorta, 461, 466.
477.
Harpactes
ardens, 107, 948.
Harpalus
alienus, 591.
ceruleatus, 714.
indicola, 714.
liodes, 715.
masoreoides, 715.
melaneus, 714.
suleatulus, 592.
turculus, 714.
Harpyia
cephalotes, 875.
Hebomoia
glaucippe, 837, 855.
Hedymeles
ludoviciana, 58.
Heleotragus
reduneus, 745.
Heliastes
lepidurus, 216.
Heliconius
charithonia, 481.
Heliodoxa
henryi, 69.
jacula, 69.
Heliomaster
constanti, 69.
Heliophorus
belenus, 702.
Helix
adela, 72, 74.
esopus, 72, 'T4.
auriculina, 73.
boivini, 861.
boucardi, 72, 74.
brenchleyi, 861, 864.
congener, 496, 497.
cornu-gigantewm, 312.
decollata, 967.
guestieriana, 312.
herklotsi, 106.
luhuana, 106.
miranda, 495.
myomphala, 496.
peliomphala, 105,
106.
quadridentata, 73.
simode, 106.
vesicalis, 312.
(Camena) congener,
(5 congenita, 496,
497.
(——) lewisii, 495,
6,
Helminthophaga
peregrina, 52.
|
INDEX.
Hemifusus
corona, 796, 798.
Hemiphaga
poliocephala, 113, 952.
Hemipodius
melanotus, 127.
Hemiprocne
zonaris, 67.
Henicorhina
leucophrys, 51.
leucosticta, 51.
Hermotimia
aspasioides, 81.
auriceps, 80.
Herodias
garzetia, 345, 712.
intermedia, 958.
Herona
angustata, 829, 854.
marathus, 829.
Hesperia
bada, 688.
baracoa, 483.
benjamini, 841.
beturia, 690, 703, 707.
bevani, 688.
cahira, 845.
contiqua, 687.
divodasa, 703.
dolophia, 690.
eltola, 688.
Sarri, 688.
gremius, 689, 703, 707.
kumara, 687, 688, 843.
mangala, 688, 703, 757.
mathias, ‘03, ‘707,
843.
moolata, 848, 856.
murdava, 689.
narooa, 687, 694.
quinigera, 703, 707.
seriata, 688.
subgrisea, 689, 694.
toona, 688.
Hesperophanes
cribricollis, 720.
Hesudra *
divisa, 12, 36.
Hestia
clara, 637.
leuconoé, 687.
Heterochroa
abyla, 480.
Heteroclitus
tartaricus, 261.
Hiaticula
inornata, 127.
| Hierococe
pectoralis, 946.
Hierodula
deflera, 581.
Hierodula
laticollis, 581, 582.
(Rhombodera) afri-
coxis, 582, 587.
(
) butleri, 580,
587.
(——) fratricida, 581,
587.
( ) macropsis, 581.
(——) pustulifera, 583,
587
3 taprobane, 583,
587
(——) tectiformis, 580,
581, 582.
Hippotragus
niger, 745.
Hirundo
ethiopica, 129.
Ffraneica, 282.
frontalis, 95.
fuciphaga, 429.
guituralis, 579.
javanica, 95, 109, 342,
615, 709.
meyent, 452.
nigricans, 95.
rustica, 579.
Histia
papilionaria, 846, 857.
Homarus
capensis, 755.
Homoptera
posterior, 487.
terrosa, 487.
Horeites
brunneifrons, 981.
brunnescens, 980, 982.
major, 981.
pallidus, 980, 982.
robustipes, 980, 982.
Horornis
‘fortipes, 980, 982.
Humphreyia
strangei, 869.
Hyena
brunnea, 422, 425, 426,
427.
crocuta, 416, 417, 419,
421, 422, 423, 424,
425, 426, 427, 428.
striata, 422, 423, 425,
426, 427, 428.
Hyalina
albida, 866.
Hybleea
puera, 486.
| Hydrocolceus
minutus, 206.
| Hydropotes
inermis, 887, 916.
Hydrornis
nipalensis, 430. |
Hylochelidon
nigricans, 95.
Hylomanes
gularis, 100,
Hylophilus
decurtatus, 53.
Hyloterpe
philippinensis, 109.
Hymenia
perspectalis, 493.
Hymenopa
coronata, 586.
Hymenopus
bicornis, 586.
Hyomoschus
aquaticus, 682, 686.
ena
lunifera, 492.
Hypercompa
regalis, 848,
Hyperythra
angulifascia, 851, 858,
85
59.
limbolaria, 851, 858.
Hyphantornis
brachyptera, 130.
nigriceps, 308.
textor, 130.
Hypochera
chalybeata, 130.
Hypogramma
confusa, 487.
sublucida, 487. |
Hypolais |
Lega 129. |
Hypolithus
chlenioides, 592. |
perlucens, 715.
puberulus, 592.
Hypolycena
lisias, 833, 855.
Hypopyra
stan, 642.
Hyposidra
ochrea, 492.
Hypotenidia
striata, 345.
torquata, 114, 288,
345.
Hypothymis
ee, 109, 284, 615,
948.
celestis, 107, 109.
Hypotriorchis
severus, 937.
H
ypsa
aleiphron, 4.
canaraica, 3.
heliconia, 3, 4, 6.
| Icterus
INDEX,
Hypsa
sericea, 3.
siluandra, 642.
subsimilis, 847, 857.
Hypsinephus
ellipticus, 716.
Hypsipetes
philippinensis,285, 342,
380
rufigularis, 949,
Hyria
vinacea, 491.
Tanthenas
griseigularis, 288.
Ichthyscopus
inermis, 220,
baltimore, 59.
grace-anne, 137.
prosthemelas, 59.
Tonotreron
ionogastra, 559.
melanocephala, 552.
nana, 550.
occipitalis, 569.
rivolii, 561.
viridis, 560.
Tole
olivacea, 618.
Tora
scapularis, 619.
viridis, 619.
lotreron
chrysorrhoa, 553.
eugeni@, 561.
hyogaster, 559.
dogaster, 504.
melanocephala, 504,
552.
melanospila, 552.
nani, 504, 549.
rivoli, 504, 561.
viridis, 504, 560.
wanthorrhoa, 553."
Tlerda
belena, 702, 707.
Indicator
archipelagicus, 794,
795.
barianus, 795.
conirostris, 793,
795,
exilis, 794, 795.
maculatus, 795.
major, 795, 930, 935.
malayanus, 794, 795.
minor, 794, 795.
sparrmani, '795.
variegatus, 795. —
xanthonotus, 794, 795.
794,
10385
Tris
leucophthalmus, 193.
Irrisor
erythrorhynchus, 355.
Ithomeis
eulema, 362, 363.
imitatria, 362.
Txias
andamana, 837.
citrina, 837, 855.
latifasciata, 837.
moulmeinensis, 837,
855.
pallida, 837, 855.
pirenassa, 837.
gotavier, 110, 285, 342,
710, 949.
Ixus
nigricans, 356.
Ismene
aria, 686, 703, 707.
benjamini, 841, 856.
druna, 842, 856.
Jelfreysi, 406, 407.
subfasciata, 686.
Tsognomon
samoensis, 820.
Tsoteinon
atkinsoni, 698, 695,
844
khasianus, 693.
masuriensis, 693, 694.
sudtestaceus, 844, 856.
Ispidina
picta, 131.
Tssus
coleoptratus, 462, 463,
471.
Jassus
wollastoni, 462, 476.
Jodis
kinstonensis, 490.
Junonia
almana, 699, 706, 828,
854.
genoveva, 480.
ada, 639.
iphita, 646.
laomedia, 639, 698, 670,
828, 854.
lemonias, 828, 854, 698,
706.
enone, 698, 706.
orithyia, 828, 854,
villida, 297, 646.
Kallima
limborgii, 828, 854.
Katha
apicalis, 17.
1036
Katha
eucullata, 17.
nigrifrons, 17, 36.
terminalis, 17, 36.
Klais
guimeti, 69.
merriti, 69.
Korawa
pallida, 12.
Kraussia
lamarckiana, 871.
Kricogonia
lyside, 481.
terissa, 481.
Kurukuru
clementine, 585.
dupetit-thowarsi, 541.
mercieri, 536.
nebouxit, ee
wratus, 507, 541.
pase abe 9 Hadael.
samoensis, 540.
superbus, 543.
swainsonii, 532.
taitensis, 542.
temminckii, 544.
axanthogaster, 540.
Kurutreron
chrysogaster, 505, 541,
coralensis, 505, 528.
oopa, 505, 542.
Lachnophorus
leucoscelis, 603.
longulus, 603.
sculptifrons, 604.
semirufus, 603.
Lagopus
Pie. 793.
scoticus, 793.
Lalage
INDEX,
| Lamprosoma
ignicaudatum, 984.
inornatum, 984.
testudineum, 984.
' Lamprotornis
metallicus, 88, 89.
_ Lamprotreron
apicalis, 527.
porphyracea, 535.
porphyreus, 504.
porphyria, 569.
porphyrostictus, 543,
544,
superba, 126, 504,
543.
Laniarius
affinis, 357.
salime, 357.
' Lanius
dominica, 108, 284,342,
709, 948.
karu, 87, 671.
polygrammica, 87.
Lamna
cornubica, 117.
Lampides
elianus, 838, 855.
alexis, 833, 855.
beticus, 702, 707.
ceraunus, 481.
elpis, 833.
kandarpa, 702,
833.
Lamprolema
rhami, 68.
Lamprosoma
chapuisi, 983.
coruscum, 984.
eyaneum, 985.
707,
cucullatus, 79.
karu, 87.
lucionensis, 342, 380,
614, 709, 947.
melanocephalus, 618.
nasutus, 341, 708, 709,
947.
smithii, 128.
Laroides
americanus, 175.
argentaceus, 167.
argentatoides, 167.
argentatus, 167.
argenteus, 167.
audouint, 179.
borealis, 169.
cachinnans, 169.
californicus, 175.
canescens, 177.
chalcopterus, 166.
cinerea, 164,
Ffuscus, 173.
glaciatis, 166.
glaucescens, 167.
glaucoides, 166.
glaucus, 165.
harengorum, 173.
leucopheus, 169.
leucopterus, 166.
major, 167.
melanotus, 173.
michahellesii, 169.
occidentalis, 172.
177.
procellosus, 177.
rissa, 164.
subleucopterus, 166.
tridactylus, 164.
Larus
affinis, 171, 172, 174, |
175.
albipennis, 208.
arus ‘
albus, 162, 163, 180,
206
antipodus, 180.
arabicus, 193.
arcticus, 166.
argentatoides, 167, 169,
175, 176.
argentatus, 166, 167,
168, 169, 170, 171,
172, 178, 174, 175,
176, 180.
argenteus, 167.
ag ce 160, 194, 195,
atricilloides, 206.
audouini, 158, 160,177,
79:
bathyrinchus, 181.
belcheri, 160, 182, 183.
bonapartiz, 196, 206.
brachyrhynchus, 164,
178, 179.
brachytarsus, 162.
brehmii, 192.
brevirostris, 165.
bridgesi, 183.
bruchi, 177.
brunneicephalus, 197,
200.
bulleri, 187, 190, 191.
cachinnans, 160, 161,
168, 169, 170, 171,
172, 173.
cahiricus, 201.
cahirinus, 201.
californicus, 175, 176,
177, 178.
candidus, 162.
canescens, 200.
canus, 160, 163, 175,
176, 177, 178, 179.
canus major, 177.
capistratus, 158, 200,
206.
chalecopterus, 166, 167.
cinerarius, 163, 200.
cinereo-caudatus, 195.
cinereus, 167, 173, 177.
cirrhocephalus, 197,
202, 204, 205.
collaris, 208, 209, 210.
columbinus, 192.
consul, 165.
corallinus, 187.
crassirostris, 160, 174,
183, 193.
cucullatus, 195.
cyanorhynchus, 177.
delawarensis, 175, 176,
177, 178.
—
Larus
dominicanus, 161, 180, |
181 |
@ orbignyi, 206.
eburneus, 158, 162.
epargyrus, 169.
erythropus, 200.
fabricit, 180.
Alavipes, 173.
Franklini, 195, 205.
Fritzei, 181.
Ffrobeni, 182.
Frontalis, 181.
fuliginosus, 156, 157,
160, 182, 184.
furcatus, 160, 210.
fuscescens, 170, 173.
fuscus, 158, 161, 167,
170, 171, 172, 173,
174, 175, 181, 192, |
183.
gavia, 164. H
gelastes, 160, 191, |
192,
Larus
. leucophthalmus, 193,
194,
genei, 192.
georgii, 181. |
giganteus, 165. |
glacialis, 165. |
glaucescens, 167.
glaucodes, 203.
glaucoides, 165.
glaucopterus, 167. |
glaucotes, 203. |
glaucotis, 196, 197.
glaucus, 158, 165, 166,
16
hematorhynchus, 184.
hartlaubi, 188, 189,
205.
heermanni, 160, 182, |
183, 184, 185.
heinei, 177.
hemprichi, 193, 194,
976.
heuglini, 171.
hutchinsii, 165, 166.
hybernus, 177.
ichthyaétus, 159, 170,
198.
minor, 198.
tslandicus, 165, 166.
jamesonii, 185.
kitlitzi, 195.
kotzebui, 164.
kroicocephalus, 198.
lacrymosus, 198.
lambruschini, 193.
leuceretes, 165. |
leucocephalus, 191.
leucomelas, 181.
leucopheus, 169, 170.
INDEX,
|
i
leucopterus, 166.
littoreus, 180.
maculatus, 180.
maculipennis, 187, 197,
201, 202, 203, 204.
major, 177, 185, 198.
marinus, 158, 161, 167,
179, 181
masauanus, 193.
maximus, 180.
medius, 173.
melanocephalus, 199.
melanorhynchus, 190,
206
méelanurus, 174, 183.
mexicanus, 177.
minor, 166.
minutus, 159, 206.
modestus, 160, 183, 184.
milleri, 180.
nevius, 163, 179, 200.
nigrotis, 206.
niveus, 162, 175, 177,
178.
nove-hollandie, 185,
186, 187, 188, 189,
190
occidentalis, 171, 172,
173, 174.
pacificus, 159,161, 181.
payraudei, 179.
personatus, 196.
pheocephalus, 189, 204.
philadelphia, 206, 207.
pipixcan, 195.
plumbiceps, 159.
poiocephalus, 189, 204.
pomare, 190, 191
procellosus, 177.
guadricolor, 179.
ridibundus, 158, 190,
193, 194, 198, 200,
201.
riga, 163.
rissa, 163.
roseiventris, 204.
roseus, 159, 208.
rossii, 208.
sabini, 209.
saundersi, 195, 205.
schimperi, 195, 205.
scopulinus, 185, 187,
188, 189, 191, 195.
—— major, 186.
scoresbiz, 160, 161, 184,
185.
serranus, 196, 197, |
02,
202.
Proc. Zoou. Soc.—-1878, No. LXVIII.
1037
Larus
smithsonianus, 168.
subroseus, 192.
suckleyt, 177, 178.
tenuirostris, 192.
torquatus, 164.
tridactylus, 158, 163,
164 +
verreauxi, 161, 181.
vetula, 181.
vociferus, 180.
warnecki, 164.
zonorhynchus, 176,
177
(Blasipus) heermanni,
183
(Bruchigavia) melano-
rhynchus, 190
(Pagophila) eburneus,
162
(Rissa) brevirostris,
165,
Laryngogramma
gularis, 570.
asea,
australis, 870,
Lasiommata
muirheadi, 696.
Latiaxis
elegans, 74,
Latirus
cayohuesonicus, 796,
798
decoratus, 812, 821.
fastigium, 812, 821.
incarnatus, 813.
infundibulum, 796.
Latrunculodes
nilssoni, 331.
Latrunculus
albus, 318, 319, 320,
821, 323.
nilssoni, 331.
pellucidus, 318, 319,
321, 322, 323, 330,
332, 333, 334, 337.
338.
stuvitzi, 318, 319, 320,
321, 322, 323.
Laurona
ergolis, 489.
Lebadea
attenuata, 829, 854.
ismene, 829.
ebia
callizona, 607.
charina, 607.
corcula, 607.
Leda
(Adrana)
314.
68
newcombi,
1038
Legatus
Fnicollis, 63.
Leiocnemis
frivola, 717.
himalaica, 716.
tartarie, 716.
Lema
bifida, 985.
semisepta, 983.
suffriani, 982.
Lemonias
debilis, 366.
hypoglauca, 367.
orpheus, 366.
parthaon, 366.
pelarge, 366.
theages, 366.
Lepidopleurus
antiquus, 868.
Leptocircus
virescens, 841, 856.
Leptodon
uncinatus, 45.
Lepton
australis, 863, 864,
ae
Leptoptila
Paa 43.
verreauzi, 43.
Leptotrachelus
panamensis, 600.
puncticollis, 600.
Leptura
rubriola, 720.
Lepyrodes
geometralis, 705,
708
Lestris
antarctica, 181.
fusca, 181.
scopulina, 188.
Lethe
arcadia, 824, 853.
europa, 696,
mekara, 824, 853.
rohria, 824, 853.
verma, 824, 853.
Leucocerca
nigritorquis, 109, 284,
709, 948.
Leucophzeus
belcheri, 182.
fulrginosus, 184.
hematorhynchus, 184.
heermanm, 183.
modestus, 183.
scoresbii, 184.
Leucotreron
cincta, 572.
gironiert, 287, 568,
569.
INDEX.
Leucotreron
gularis, 570.
Leucus
arcticus, 166.
chaleopterus, 166.
fuscus, 173.
glaucescens, 167.
glaucus, 165.
leucopterus, 166.
marinus, 180.
Lia
quadriannulata, 607.
Liburnia
(Ilburnia) zgnodilis,
462, 471.
Libythea
myrrha, 882, 854.
Limicola
platyrhynchus, 708,
12
sibirica, 712.
Limnaétus
philippensis, 281.
Limnas
bryaxis, 362.
ceruleata, 362.
melanochlora, 362.
Limopsis
macgillivrayt, 871.
Limosa
egocephala, 280, 288.
fedoa, 141.
hudsonica, 141.
lapponica, 708, 711.
Lingula
hians, 820.
Lipaugus
holerythrus, 65.
Lithosia
angulifera, 20.
antica, 15.
apicalis, 17.
aureola, 18.
beema, 10.
brevipennis, 17.
chilomorpha, 23.
complanula, 16.
distorta, 25.
dives, 14.
igas, 7, 8.
gilveola, 18.
griseola, 16.
helveola, 17,
innotata, 19.
insolita, 19, 20.
intermiata, 17.
magnifica, 13.
muscerda, 27.
natara, 15.
nigrifrons, 17.
nigripars, 19,
Lithosia
nivea, 15.
quadra, 14.
remelana, 10.
reticulata, 20.
rotundipennis, 23.
sambara, 19.
sarawaca, 15.
serva, 15, 16.
stramineola, 16.
vagesa, 19.
varana, 15.
vetusta, 16.
viridata, 19.
Litorina
melanacme, 817.
nove-zelandia, 817.
striata, 449.
undulata, 816.
ventricosa, 817.
Lituus
brevis, 499.
Lophius
piscatorius, 220.
Lophogaster
typicus, 780.
Lophosalea
anamallayana, 153.
Lophostrix
stricklandi, 45.
Lophotragus
michianus, 900.
Loriculus
hartlaubi, 107, 340,
708, 709, 937.
regulus, 281.
Loripes
ovum, 870.
Lorius
cyanauchen, 78, 79.
domicella, 79.
hypenochrous, 289.
sguamatus, 86.
Loxia
psittacea, 347.
Loxioides
bailloni, 346.
Loxura
atymnus, 835, 855.
Lucia
dilama, 701, 707.
epius, 701.
Lucinia
cadma, 480.
citrini, 860, 864.
pennsylvanica, 861.
quadrata, 863.
rosea, 860, 864.
tate?, 870.
(Codakia) Zatet,
864, 870.
863,
Lycena
elpis, 833.
kandarpa, 702.
roxus, 833.
picta, 373, 374, 375,
376.
Lyclene
artocarpi, 30.
assamica, 33, 37.
cuneigera, 33.
delicata, 31.
discistriga, 32.
euprepioides, 32.
humilis, 30.
inconspicua, 32.
indistincta, 33, 37.
interserta, 32, 37.
obsoleta, 32, 37.
palmata, 31, 37.
prominens, 31, 37.
radians, 30, 31, 37.
rubricosa, 30, 37.
spilosomoides, 33.
strigipennis, 32.
terminata, 38.
undulosa, 33.
zebrina, 31, 37.
Lyctocoris
campestris, 461, 463.
Lygus
apicalis, 461,
467.
Lymanopoda
euopis, 266.
Lyncornis
macrotis, 944.
mindanensis,
944,
temminckii, 944.
936,
Macacus
cynomolgus, 791.
Macaria
acidaliata, 491.
nora, 851, 858.
Machzeropterus
pyrocephalus, 139.
Macherorhynchus
xanthogenys, 95.
Macotasa
biplagella, 25, 37.
Macrobrochis
gigas, 7, 8.
interstitialis, 7.
leucospilota, 8.
nigrescens, 8.
Macrocercus
macao, 77.
Macroglossa
luteata, 844, 856.
orientalis, 844, 856.
463,
INDEX.
Macrones
vittatus, 218.
Macronus
striaticeps, 110,
115, 949.
Macronyx
croceus, 356.
striolatus, 356.
Macropodus
pugnax, 215.
Macropteryx
comata, D44.
mystacea, 81, 95.
Macropygia
amboinensis, 82, 90.
browni, 671, 673.
doreya, 90, 99.
eurycerca, 280, 288,
952.
keyensis, 89.
phasianella, 89, 99.
tenuirostris, 288.
turtur, 89.
Macrorhaphis
acuta, 464.
Macroschisma
tasmanie, 868.
Macroxus
adolphet, 662.
boothia, 663.
chrysosurus, 669.
flaviventris, 668.
fraseri, 441, 664, 665,
669.
griseoflavus, 660.
griseogena, 667.
irroratus, 668.
kuhlii, 668, 670.
leucogaster, 668.
leucops, 660.
maurus, 660.
medellinensis,
669.
melania, 662.
morio, 660.
nicoyana, 662, 664.
pyladei, 662.
teniurus, 668, 669.
tephrogaster, 668, 669.
minor, 669.
canthotus, 667.
Mepha
ditus, 385.
Mahathala
ameria, 703, 707.
(Amblypodia) ameria,
836.
114,
668,
Mahavira
Alavicollis, 11, 36.
| Majaqueus
equinoctialis, 773.
1039
Malacoptila
panamensis, 47.
Mangalura
dobsoni, 303.
Manis
temmincki, 632.
Mantis
bicornis, 586.
coronatus, 586.
monstrosus, 583.
Manucodia
atra, 98.
gouldt, 125.
keraudreni, 123.
Manulea
calamaria, 18.
gilveola, 18.
Mareca
sibilatrix, 436.
Margarodes
conchylalis, 850,
vertumnalis, 850.
Margaronia
marthesiusalis, 850.
quadristigmalis, 493.
Margarornis
rubiginosa, 60.
Marginella
turbinata, 866.
volutiformis, 865.
Marinula
xanthostoma, 869.
Marpesia
elucha, 480.
Mascaria
crocea, 311.
Mastostethus
salvini, 983.
Mecoceras
bitactaria, 491.
Mecyna
reversalis, 494.
Megacephalon
mileo, 629, 630.
Megaceros
hibernicus, 891.
Megaderma
spasma, 879.
Megalema
asiatica, 932.
Megalomastoma
litteratum, 311.
Megaloprepia
assimilis, 126, 574.
bernsteint, 573.
formosa, 544.
magnifica, 575.
perlata, 558.
puella, 574.
Megalopterus
plumbeus, 211, 272.
1040
Megalurus
ruficeps, 285.
Megapodius
cumingi, 624.
dillwyni, 953.
duperreyz, 100.
eremita, 289, 290, 671,
6
73.
hueskeri, 290, 673.
reinwardtii, 100.
rubrifrons, 290, 673.
rubripes, 100, 629.
rufipes, 100.
tumulus, 126.
Megarhaphis
wollastoni, 461,
464.
Megarhynchus
priangua, 63.
Megerlia
jefireysi, 406, 407,
415.
463,
monstruosa, 399, 411.
sanguinea, 407.
truncata, 399, 405, 406,
410, 411.
Melsenornis
edolioides, 128.
Melagayia
melandcaplals: 200.
Melanerpes
formicivorus, 49.
Melania
dicksoni, 728, 732.
dolorosa, 730.
formosensis, 728, 732.
monstrosa, 729.
obliquigranosa, 729,
732.
subplicatula, 729,
43a.
tuberculata, 729, 732.
Melanitis
bela, 824, 853.
asmene, 696, 706, 824,
853.
Melanitta
fusca, 479.
Melanobatrachus
indicus, 722.
Melanopelargus
episcopus, 344, 953.
Melanopitta
cucullata, 949.
muelleri, 949.
sordida, 949.
steerii, 949.
Mellisuga
coccinea, 347,
Melo
miltonis, 865.
INDEX.
Melonycteris
melanops, 316.
Melozone
leucotis, 40, 56.
Melursus
labiatus, 378.
Mergus
serrator, 479.
Merops
albicollis, 355.
bicolor, 282, 340,
937.
hirundineus, 355.
minutus, 355.
miger, 347.
ornatus, 121.
philippinus, 107, 282,
340, 709.
pusillus, 355.
superciliosus,
355.
Mesene
@rope, 363.
ignicauda, 363.
pyrippe, 363.
sagaris, 363.
silaris, 363.
tyriotes, 363.
Messaras
erymanthis, 699, 707,
27, 854.
Metabletus
tartarus, 719.
Metacanthus
concolor, 461, 462.
Metaxyonycha
rufolimbata, 987.
Metopiana
peposaca, 479.
Micrastur
semitorquatus, 44.
Microchera
parvirostris, 69.
Microglossum
alecto, 92.
aterrimum, 92, 125.
pps ees "
erythrogenys, 937.
Micronia
aculeata, 852, 858.
grammearia, 852,
858.
vagata, 852, 858.
Micropterus
cinereus, 437.
Miletus
chinensis,
707.
Milvago
chimachima, 849.
chimango, 435.
131,
701,
!
|
'
Milvulus
forficatus, 65.
Mimeta
flavocinctus, 123.
viridis, 123.
Mino
dumonti, 98.
Mionectes
oleaginus, 63.
iro
albifrons, 578.
Mithuna
quadriplaga, 21, 37.
Mitra
acuta, 797, 798.
annulata, 798.
berthe, 797, 798.
deburghie, 798.
flammea, 814.
fulvo-lirata, 797, 798.
incisa, 814.
lincolnensis, 865.
marie, 813, 821.
mesta, 813, 821.
nitens, &! 4.
pica, 865.
puncturata, 798.
schomburgki, 318, 314,
865.
tatei, 861, 864, 865.
(Callithea) acupicta,
814.
( ) obeliscus, 815.
(Cancilla) annulata,
814.
(——) philippinarum,
814
(Chrysame) tabanula,
814.
(Costellaria) deshayest,
815.
(——) exasperata,
815.
(——) lincolnensis, 313,
(Cylindra) undulosa,
814
Mitrephorus
auranttiventris, 64.
Mixornis
capitalis, 107, 110, 114,
115, 342, 380.
woodi, 617.
Mniotilta
varia, 2.
Modiolaria
barbata, 871.
Mohoa
nobilis, 347.
| Molops
pileferus, 718.
Molossus
abrasus, 879.
rufus, 879.
Molothrus
eneus, 59.
Momotus
equatorialis, 100.
brasiliensis, 100.
lessoni, 48, 49, 100,
101.
Monarcha
alecto, 289, 671.
chrysomelas, 671.
fulviventris, 83, 84.
inornatus, 83, 671,
672.
insularis, 579.
verticalis, 671.
Monticola
solitarius, 342, 380,
619, 949.
Morasa
lorimeri, 387.
modesta, 387.
Morococcyx
erythropygus, 48.
Morrisia
davidsoni, 412.
gigantea, 411, 412.
Moschus
moschiferus, 1.
Mulona
lapidaria, 484.
Munia
jagori, 287, 348, 710,
951.
malabarica, 348.
nisoria, 348.
punctularia, 348.
Murex
angast, 864.
tetragonus, 806.
(Muricidea) barclay-
anus, 806.
(—) eirrosus, 806.
(——) rusticus, 806,
821
(Ocinebra) breviculus,
806.
Muscicapa
banyumas, 615.
cantatrix, 615.
inornatus, 83, 84.
lugubris, 802.
megarhyncha, 96.
obscura, 347.
sandvicensis, 346.
Muscipipra
vetula, 339.
INDEX.
Muscivora
mexicana, 63.
| Mustelus
antarcticus, 116.
Mycalesis
anaxias, 825, 853.
blasius, 825, 8538.
charaka, 696, 706.
dorycus, 645,
medus, 645.
mineus, 825, 858.
mnasicles, 825.
perseus, 696, 706, 825,
853.
runeka, 696, 706, 825, |
853. |
Mycetes
palliatus, 38.
Myiadestes
melanops, 51.
Myiarchus
cinerascens, 64.
crinitus, 38, 64.
lawrencei, 64.
nigricapillus, 64.
panamensis, 64.
semirufus, 138.
Myiobius
erythrurus, 64.
sulphureipygius, 64.
Myiodynastes
lutewventris, 63.
Myiolestes
aruensis, 96.
phajonotus, 84.
Myiopsar
eryptopyrrhus, 803.
Myiozetetes
texensis, 63.
Myodioctes
pusillus, 52.
Myoxus
guerlingus, 668.
Myrina
freja, 834, 855.
hiemalis, 640, 6438.
nivea, 640, 643.
Mpyristicivora
bicolor, 89, 113, 952.
luctuosa, 89.
melanura, 89.
spilorrhoa, 89.
Myrmeciza
immaculata, 61.
Myrmedobia
fuliginea, 461, 466.
Myrmotherula
melena, 39, 61.
menetriesi, 61.
Mysella
anomala, 863, 870.
-
}
1041
Mysella
donaciformis, 863, 864,
870.
Mysis
oculata, 787.
relicta, 787.
| Mytilus
crenatus, 449.
edulis, 449.
Myxopoda
aurita, 871, 879.
Myzanthe
pygmed, 343, 381, 620.
Myzomela
botei, 84.
cardinalis, 353.
chermesina, 852.
erythrocephala, 97.
nigrita, 97.
obscura, 125.
Nabis
capsiformis, 462, 463,
469.
Nacella
parva, 862, 864, 868.
Nandinia
binotata, 142.
Nanotragus
pygmeus, 884.
tragulus, 742.
Napothera
eleoides, 96.
Narathura
hypomuta, 835.
moolaiana, 835, 855.
Nasiterna
pusio, 289.
Nassa
albescens, 809.
bifaria, 808, 821.
costellifera, 808.
crenulata, 807.
curta, 810.
dispar, 808.
echinata, 810, 821.
elegans, 807.
gemmulifera, 809.
gruneri, 811.
horrida, 810.
intermedia, 865.
jacksoniana, 865.
lurida, 808.
margaritifera, 808.
marginulata, 808,
marratiz, 809, 821.
monile, 808.
muricata, 810.
olivacea, 808.
stigmaria, 808.
subspinosa, 810.
1042 ‘
Nassa
tenia, 808.
trinodosa, 810.
(Arcularia) bimaculosa,
810
(——) callospira, 810.
(——) globosa, 809.
( ) granifera, 810.
(Hima) sistroidea, 809.
Nathalis
iole, 481.
Natica
marochiensis, 866.
Nautilus
imperialis, 966.
pompilius, 955, 958,
961, 962, 969, 970,
974.
striatus, 968, 970.
Nectarinia
aspasia, 289.
aspasioides, 81.
auriceps, 80.
botei, 84.
collaris, 356.
frenata, 79, 125, 289.
fuliginosa, 129.
gutturalis, 355.
insignis, 621.
jardinii, 355.
splendida, 129,
venusta, 130.
Nectarophila
julie, 951.
sperata, 112, 286, 343,
620.
Nehela
vulturina, 462, 473,
477.
Neochera
heliconioides, 6.
marmorea, 847, 857.
Neomorphus
radiolosus, 439.
rufipennis, 439.
Neornis
assimilis, 980, 982.
Neotragus
saltianus, 743.
Neptis
aceris, 639.
adara, 830, 854.
adipala, 830, 854.
andamana, 830.
eurynome, 697.
hainana, 697, 706.
hordonia, 698, 830.
jumba, 830, 854.
kamrupa, 698.
meetana, 830.
plagiosa, 830, 854.
INDEX,
Neptis
rihodona, 698, 706.
varmona, 830.
Nesocichla
eremita, 577.
Nesospiza
acunhe, 577.
Nezara
viridula, 461, 463, 464.
Nigrita
canicapilla, 130.
emilie, 130.
Ninox
boobooks 121.
Japonica, 940.
lugubris, 936, 940.
philippinensis, 940.
solomonis, 290.
spilocephala, 936, 939.
Nishada
flabrifera, 23.
Nisoniades
diocles, 842.
Nisus
poliocephalus, 91.
tachiro, 354.
Noctilio
leporinus, 880.
Noctua
variegata, 290.
Noda
balyi, 985.
boucardi, 986.
lefevrei, 985, 986.
propingua, 986.
rufipes, 986.
semicostata, 985.
violaceipennis, 987.
viridis, 985, 986.
Notaphus
semifasciatus, 602,
Notauges
albicapillus, 721.
hildebrandti, 721.
superbus, 721.
Nothocercus
bonapartii, 41.
Notidanus
cinereus, 116.
Notiobia
cupreola, 590.
disparilis, 589.
leiroides, 590.
limbipennis, 590,
parilis, 590.
Nucula
micans, 864, 871.
pusilla, 864.
Nudaria
Sasciata, 36.
margaritacea, 36.
Numenius
pheopus, 90,
uropygialis, 90, 100.
Nychitona
wiphia, 699, 707, 887,
855.
Nyctaleamon
egistus, 490,
docile, 642,
hector, 642.
Nyctemera
latistriga, 846, 857.
separata, 386.
varians, 847.
Nyctereutes
procyonides, 378, 374,
875, 376
Nycticorax
caledonicus, 671, 673.
manillensis, 288, 345.
obscurus, 436.
Nyctidromus
albicollis, 67.
Nyctinomus
plicatus, 879.
Nyctipao
crepuscularis, 642, 849,
Nyctophilus
timoriensis, 876.
Nymphalis
acilia, 646.
Sranckii, 832.
schreiberi, 640.
Nymphidium
adelphinum, 368.
agle, 368.
hematostictum, 367.
ictericum, 367.
lycorias, 368.
phliasus, 368.
sicyon, 368.
velabrum, 368.
Nyroca
leucophthalma, 479.
Nysius
sancte-helene, 461, 464,
465.
thymi, 461, 463,
Obeliscus
brunneus, 866.
Oceanites
oceanicus, 730.
Ochria
niveopicta, 485.
Ochthodromus
inornatus, 127.
Oculina
prolifera, 401.
ee
Ocyalus
wagleri, 58,
Odontophorus
guttatus, 42.
Odostomia
eburnea, 866.
(cophora
biplagella, 25.
C£dirhinus
globifer, 549.
msolitus, 289, 290, 549,
671.
CEinas
arenaria, 234,
bicincta, 234, 256, 257.
cata, 234, 250.
indica, 2384, 254.
namaqua, 234, 252.
Ginops
aura, 45.
falklandica, 435.
Cnistis
dives, 14.
entella, 138, 14, 847,
857.
Gstrelata
brevirostris, 738.
kidderi, '738.
lessoni, 737, 738.
mollis, 577, 738.
sericea, 738,
Omeotreron
batilda, 505,
570.
felicie, 505, 565.
pectoralis, 559.
virens, 505.
Omiza
schistacea, 851, 858,
859.
569,
Oncocarpus
vitiensis, 102.
Opatrum
hadroides, 456.
Ophiocephalus
striatus, 217.
Ophiusa
delinquens, 489.
Orbicula
lamellosa, 415.
norvegica, 415,
rea
gladiator, 683.
Orchilus
ecaudatus, 138.
Oreas
canna, 745,
Oreo
say ae 69.
cinereicauda, 40, 68.
hemileuca, 40, 69.
INDEX.
Oreothraupsis
arremonops, 439.
riolus
Alavocinetus, 123.
larvatus, 856.
viridis, 123.
wvanthonotus, 616.
Ornithoptera
arruana, 647.
boisduvalii, 647.
poseidon, 647.
priamus, 647,
Orthis
anomioides, 411.
bifida, 409.
eusticta, 411.
lunifera, 410, 412.
Orthotomus
castaneiceps, 286.
cinereiceps, 950.
frontalis, 110,708, 710,
950.
ruficeps, 619.
Orthotylus
mutabilis, 461, 467.
nassatus, 467.
Ortygometra
cinerea, 114, 344, 953.
Ortyx
leylandi, 42.
ryx
beatriz, 789.
beisa, 441.
Oryzoborus
atrirostris, 136.
erassirostris, 136, 137.
maximiliant, 137.
melas, 137.
occidentalis, 137.
Osmotreron
axillaris, 118, 287.
vernans, 623, 710,
951.
Osphromenus
olfax, 217.
Ossifraga
gigantea, 737,
Ostinops
montezume, 58,
Otaria
stelleri, 371.
ursina, 371.
Otocyon
lalandii, 376.
Otoglossa
celestina, 607.
obscurella, 608.
Otus
brachyotus, 348, 434.
Oxeoschistus
phaselis, 267.
1043
Oxeoschistus
rogersi, 267.
Oxycerca
everetti, 622.
leucogaster, 622.
Oxyechus
vociferus, 44.
Oxyurus
spinicauda, 433.
Pachyarches
marthesiusalis, 850,
858.
vertumnalis, 850, 858.
Pachycephala
sp., 121, 122.
albicollis, 79.
brunnea, 84.
flavifrons, 351.
melanura, 79.
mentalis, '79.
nigrimentum, 79.
icterotides, 351.
pheonota, 84.
senex, 84.
Pachylia
Jicus, ar
Pachyptila
Kaa kei 739.
Pachyrhamphus
cinereiventris, 65.
versicolor, 65.
Pachythone
gigas, 364.
Pagodroma
nivea, 737.
Pagophila
Geckapinl 162,
eburnea, 159, 162, 175.
nivea, 162.
Palemon
canaliculatus, 298.
lacustris, 787.
Palinurus
vulgaris, 292, 442.
Pamphila
augias, 690, 691.
bambuse, 691, 695, 842,
856.
brahma, 691, 692, 695.
confucius, 703, 707.
ethlius, 482.
insolata, cage
luteisquama, 3
masoni, 842, 856, 858.
mathias, 691, 843, 856.
nyctelius, 482.
otho, 483.
palmarum, 690, 695,
phylus, 482,
stva, 692,
1044
P
P
P
P
P
P
Paphia
portia, 480.
Paphiosa
helene, 70.
Papilio
abderus, 271.
aceris, 639.
amphila
subhyalina, 692.
subochracea, 691.
sylvanus, 689.
utha, 482.
andion
leucocephalus, 86.
solitarius, 348.
anglima
narcissus, 7.
angus
metallicus, 592.
antana
ampla, 705, 708.
‘anterpe
insignis, 40, 71.
achates, 641, 697, 706.
elianus, 833.
affinis, 644.
agamemnon, 841, 856.
agenor, 641.
aglea, 695.
alexis, 833.
alimena, 647.
almana, 699, 828.
amasene, 700.
androgeos, 841, 856.
antiphates, 641, 841,
856. H
arcadia, 824.
arcesilaus, 827.
aristolochie, 840, 856.
arsinoé, 646.
asclepius, 271.
atymnus, 835.
avia, 699.
axion, 697, ‘706.
beticus, 702.
baldus, 825.
biblis, 699,
blasius, 825.
canidia, 701.
castor, 840, 841.
catilla, 640, 837.
chrysippe, 645.
cheeeeee 638, 695,
cocles, 698.
codrus, 735.
columbina, 699.
eresphontes, 481.
crocale, 837.
dione, 701.
diphilus, 840.
dirtea, 639.
INDEX.
Papilio
doubledayi, 840, 856.
erechtheus, 648.
erymanthis, 699, 827.
euchenor, 648.
ewmeus, 697.
europa, 696.
euryphilus, 697.
eurypylus, 641.
evippe, 700.
flegyas, 832.
Freja, 834. |
glaucippe, 837.
gliciria, 701.
noma, 699. |
hecabe, 641, 647, 836.
hegisippus, 638.
helenus, 696, 706, 840, |
856. |
homerus, 481.
iacintha, 699.
ida, 639.
iphita, 646, 828.
ismene, 696, 824. |
iswara, 641.
Juventa, 637.
lais, 638.
laius, 702.
laomedia, 639, 698,
828.
lemonias, 698, 828.
leodamas, 648.
leucothoé, 697.
limniace, 695.
lisias, 883.
mahadeva, 840, 841,
856, 858.
malayanus, 697, 705.
medus, 645. !
megarus, 697, 706, 840, |
856.
memnon, 641.
midamus, 825.
mineus, 825. |
mulciber, 638.
nerina, 297, 647.
nina, 699, 837.
nomius, 697.
enone, 698.
onpape, 840, 856.
orithyia, 828.
pammon, 641, 696, 706, |
840, 856.
panope, 697, 840.
pasithoé, 701, 839.
pelaus, 481.
perseus, 696, 825.
phalanta, 699.
phidippus, 638, 826.
philoxenus, 840, 856.
plexippus, 695.
Papilio
polydamas, 481.
polydorus, 648.
polytes, 641, 696,
840.
poseidon, 647.
pyrene, 700.
rhadamanthus, 822.
risa, 829,
rohria, 824.
sarpedon, 641, 841,
856
56.
saturata, 697, 706.
segonax, 734.
sesia, 700.
similis, 637.
sinon, 481.
syedra, 271.
telearchus, 839, 856.
thersites, 481.
thrar, 641.
tullia, 826.
vellida, 297, 646.
venilia, 646.
xenocles, 840, 856.
xiphia, 699, 837.
zeleucus, 841, 856.
zelmira, 700, 839.
zoilus, 644.
| Parablepharis
kuhlii, 586.
Paradisea
apoda, 98, 792.
minor, 681, 792.
Paranephrops
planifrons, 770.
Paraona
splendens, 8, 36,
Parastacus
brasiliensis, 759.
| Parthenia
gracilis, 862, 864,
867.
Parthenos
apicalis, 829, 854.
gambrisius, 829.
Parula
gutturalis, 39, 40, 52.
| Parus
diffusus, 358,
domesticus, 57.
elegans, 286.
swainsoni, 358.
Patella
anomala, 414.
distorta, 415.
Pecten
albo-lineatus, 820.
Pectunculus
laticostatus, 871.
_—
a
Pedaliodes
cremera, 267.
praxithea, 266, 267.
triaria, 266, 267.
Pelagodroma
marina, 577, 736.
Pelargopsis
gigantea, 108, 3541,
943.
Pelecanoides
garnoti, 739.
urinatrixc, 739.
Pelecanopus ~
pelecanoides, 212, 272.
Pelecanus
leucogaster, 651.
piscator, 651.
sula, 651.
Pelidnota
chrysargyrea, 294.
Pelosia
muscerda, 27.
Penzus
affinis, 304, 308.
antennatus, 309.
avirostris, 304, 308.
barbatus, 304.
brevicornis, 308.
canaliculatus, 298,
306.
caramote, 299, 306.
carinatus, 300.
constrictus, 304, 308.
crassicornis, 302.
eurrirostris, 300, 307.
distinctus, 302.
dobsoni, 298, 302, 303,
305, 307, 310.
edwardsianus, 298, 308,
g is, 308.
nhs 300, 306,
310.
indicus, 301, 307.
kroyeri, 305, 307.
lamellatus, 308.
longirostris, 301.
marginatus, 299.
membranaceus, 301,
307.
monoceros, 301, 307.
monodon, 299, 300,
307.
plebejus, 299.
odophthalmus, 308.
pubescens, 304.
INDEX.
Penzeus
sculptilis, 300, 303,
304, 306
sem isiloatus, 299, 300,
306,
setiferus, 307.
siphonocerus, 301, 302.
stenodactylus, 308.
styliferus, 804, 305,
307
styliretrés 301.
tahitensis, 300.
tenuis, 308.
velutinus, 304.
Penelope
purpurascens, 42.
Penelopides
affinis, 108, 341, 947.
manille, 341,
panini, 233, 341.
Penthema
darlisa, 829, 854.
lisarda, 829.
Peosina
mexicana, 487.
aumeria, 487.
Perdix
aragonica, 245.
indica, 254.
Pericompsus
longulus, 601.
Peridrome
orbicularis, 847, 857
Perigea
mobilis, 486.
Perina
basalis, 848, 857.
Periophthalmus
schlosseri, 218.
Peristera
tympanistria, 131.
Perna
(Isognomon) samoensis,
Dy
Petasophora
cabanisi, 69.
cyanotis, 69.
thalassina, 68.
Petreeca
kleinschmidti, 655.
pusilla, 655.
Pezopetes
capitalis, 39, 40,55.
Pezophaps
solitaria, 291.
Phabotreron
amethystina, 118, 381,
505.
brevirostris, 118, 344,
952.
leucotis, 952.
Proc. Zoou. Soc.—1878, No. LXIX.
1045
Pheenorhina
goliath, 104, 105.
Phaéthon
ethereus, 651.
flavirostris, 651.
Phaéthornis
enilie, 67.
longirostris, 67.
Phainoptila
melanoxantha, 39, 40,
35.
Phakellura
hyalinata, 493.
nitidalis, 493.
Phalacrocorax
albiventris, 652.
carunculatus, 652.
imperialis, 652, 653.
VEITUCOSUS, 652.
Phalena
alciphron, 847.
caprimulgus, 848.
crepuscularis, 849.
entella, 847.
glaucopis, ‘704.
manlia, 848.
militaris, 704, 846.
procopialis, 850.
pulchella, 704.
rosalia, 851.
(Attacus) erepuscularis,
642.
pe) silvandra,
Soa Os
melicerte, 298,
Phalaropus
hyperboreus, 579.
Phaos
fascinula, 385, 384.
lacteatum, 384.
nexum, 384.
nigriceps, 383.
notatum, 388.
vigens, 383.
Pharomacrus
costaricensis, 40,
mocinno, 48.
Phasianus
- versicolor, 579.
Pheucticus
tibialis, 58.
Philampelus
satellitia, 483.
Philemon
buceroides, 124.
cockerelli, 289.
plumigenis, 88, 289.
Philentoma
albiventris, 284.
69
1046
Philentoma
cyaniceps, 284.
Philyaues
rufo-brunneus, 59.
Phlogeenas
crinigera, 952.
Phlogothraupis
sanguinoienta, 5d.
Pheebetria
fuliginosa, 740.
Pheenicophaes
curvirostris, 613.
erythrognathus, 613.
harringtoni, 613.
Pheenicothraupis
Suscicauda, 55.
Pholidauges
leucogaster, 130, 357.
verreauat, 357.
Phonipara
pusilla, 58.
Phorus
solarioides, 818.
Phos
cyanostoma, 811.
roseatum, 811.
senticosum, 811.
textum, 811.
Phrygilus
gayt, 432.
melanoderus, 432.
Phurys
garnoti, 488.
Phyciodes
ardys, 268, 269.
boucardi, 268.
cyneas, 269.
drymed, 268.
elada, 268.
frisia, 480.
leanira, 269.
nebulosa, 269.
orseis, 267.
subota, 268.
thebais, 267.
Phyllopneuste
borealis, 342, 950.
Phylloptera
segonoides, 648, 649. _
Phyllorhina
armigera, 878.
bicolor, 878.
calearata, 317.
cervind, 317, 876.
commersont, 879.
coronata, 318.
cyclops, 878.
diadema, 878,
fulva, 878.
galerita, 317.
larvata, 878.
INDEX.
Phyllorhina
tricuspidata, 317, 876.
Phyllornis
palawanensis, 619.
Phylloscopus
borealis, 342, 950.
Piaya
wehleri, 48.
ica
mystacalis, 138.
Picolaptes
compressus, 60.
Picumnus
sclateri, 140.
Picus
maculatus, 943.
nanus, 943.
validirostris, 943.
Pidorus
glaucopis, 704, 708.
Pieris
amba, 839.
aruna, 734.
clemanthe, 838, 839.
copia, 700,
descombesi, 734, 839.
eurygania, 734.
helferi, 839.
lea, 838.
lyted, 734.
madetes, 733.
nama, 838.
neombo, 838.
paulina, 838.
Piezorhina
cinerea, 137.
Pinacodera
amblygona, 606.
angulifera, 606.
Pinarolestes
megarhynchus, 9G,
rufigaster, 122.
Pione
heteroclitus, 672.
Pionia
calopteridia, 381.
Pionias
Fuscicapillus, 359.
rhodops, 80, 86.
Pionus
heteroclitus, 672.
senilis, 38, 46.
Pipra
leucorrhoa, 66.
mentalis, 66.
papuensis, 950.
Pitasila
moolaica, 847, 857,
varians, 847.
Pithauria
murdava, 689, 695.
Pithecops
nihana, 702, 707.
Pithys
bicolor, 62.
Pitta
kochi, 430.
Placunanomia
(Monia) tone, 871.
Platalea
ajaja, 44.
Platydia
anomioides, 399, 409,
410, 411, 414.
Platyrhynchus
albigularis, 62.
Platystira
cyanea, 129.
Plautus
glaucus, 165.
Plectrurus
hewstoni, 155.
madurensis, 155.
pulneyensis, 155.
sanguineus, 155.
trilineatus, 155.
Plesioneura
albifasciata, 843, 856,
858.
alysos, 843.
aurivittata, 848, 856,
858
dhanada, 843.
Pleurobranchus
nove-zealandie, 276,
277.
ornatus, 275, 277.
Pleuronectes
flesus, 329.
Pleurotoma
(Drillia) variabilis, 805,
1
821.
( Jwilmeri,805,821.
Plotus
anhinga, 378, 679, 680,
68
i
levaillanti, 378, 679,
680, 681.
Plusiotis
auripes, 295.
badeni, 294, 296.
batesi, 293, 294.
boucardi, 296.
chrysargyred, 293, 294.
costata, 295.
gloriosa, 295, 296.
lacordairei, 295.
leta, 294.
mnizechii, 296.
prasina, 295.
Plusiotis
psittacina, 294, 296.
rodriguezt, 295, 296.
sallei, 295.
victorina, 296.
Poaphila
cinerea, 488.
Podargus
papuensis, 121.
Peeocephalus
Fuscicapillus,
359.
Pceoptera
eryptopyrrha, 802,
803.
358,
lugubris, 802, 803.
Pogonorhynehus
bidentatus, 932.
melanopterus, 359.
Polioaétus
solitarius, 348.
Poliolophus
wrostictus, 110, 3880,
949.
Polpochila
mexicana, 589.
Polyborus
tharus, 231, 232,
3D.
Polyodon
foliosus, 117,
Polyommatus
argia, 702.
laius, 702, 707.
similis, 702, 707.
varunana, 702, 707.
Polyplectron
emphanes, 623, 792.
napoleonis, 623, 792.
Polypterus
bichir, 117.
Pompelon
ampliatum, 387.
marginatum, 387.
Popa
aschmoliana, 584.
Poronia
australis, 870.
Porzana
erythrothorax, 344.
fusca, 344, 953.
hauaswelli, 135.
Potamophora
manlia, 848, 857.
Potera
marginata, 852, 858,
859.
Prabhasa
costalis, 26.
flavicosta, 26, 27.
zenosa, 26, 37.
INDEX.
Pratincola
albofasciata, 979.
caprata, 710.
indica, 979,
maura, 980.
pastor, 979.
rubicola, 979.
semitorquata, 979.
Precis
iphita, 828, 854.
Prinia
cinereocapilla, 370,
371.
cursitans, 710.
hodgsoni, 370.
poliocephala, 370, 371.
stewarti, 370, 371.
Prion
bankst, 739.
desolatus, 738.
vittatus, 739.
Prioneris
clemanthe, 839, 856.
watsom, 839, 856.
Prionirhynchus
carinatus, 39, 49.
Prioniturus
discurus, 379, 937.
Prionochilus
olivaceus, 107, 111.
Prionodera
elegans, 987.
Prionops
graculinus, 357.
Prionus
corpulentus, 720.
Pristiophorus
japonicus, 117.
Pristis
cuspidatus, 117.
Procellaria
equinoctialis, 737.
antarctica, 737.
brevirostris, 738.
capensis, 737.
cinerea, 737.
desolata, 738.
gigantea, 737.
glacialoides, 736.
grallaria, 735.
lessoni, 737.
leucogastra, 736.
marina, 736.
melanogaster, 766.
mollis, 738.
nived, 737.
oceanica, 730.
urinatria
Procellarus
heermanni, 184.
neglectus, 161, 184.
1047
Procelsterna
albivitta, 212, 272.
cinerea, 272.
tereticollis, 211, 272.
Prodenia
ignobilis, 485.
pauper, 485.
Proteides
amyntas, 482,
Prothoé
franckii, 832, 854.
Protoparce
cingulata, 484.
Jamaicensis, 484.
rustica, 484.
Psallus
flavosparsus, 462,
468.
lutosus, 462, 468,
469.
vinaceus, 462, 469.
Psarisomus
dalhousieg, 148.
Pseudaxis
euopis, 908.
sika, 902, 909.
Pseudocolaptes
boissoneauti, 59.
Pseudoprion
banksi, 739.
desolatus, 738.
Pseudoptynx
gurneyt, 936, 940,
941.
philippensis, 941.
Pseudosphinx
tetrio, 484.
Psilorhinus
morio, 59.
Psittacula
melanogenia, 92.
Psittacus
alecto, 92.
capistratus, 86.
cyanauchen, 79.
domicella, 79.
fuscicapillus, 80, 359.
grandis, 78.
heteroclitus, 672.
megalorhynchus, 78.
polychlorus, 92.
rhodops, 80.
roratus, 78.
seintillatus, 93.
(Hos) bernsteini, 86.
Psittirostra
psittaced, 347.
Pterocles
alchata, 287, 288,
239, 244, 247, 250,
201.
1048
Pterocles
arenarius, 238, 234, 238,
239, 242, 244, 245,
246, 251.
bicinctus, 234, 237, 240,
255, 257, 258, 259.
caspius, 251,
coronatus, 234, 237, 238,
239, 242, 244.
decoratus, 234, 237, 238,
240, 260.
exustus, 234, 287, 288,
239, 243, 248.
fasciatus, 238, 238, 239,
254, 256, 259.
gutturalis, 234, 237,
239, 241, 243, 256.
lichtensteini, 234, 237,
238, 240, 258.
maculosus, 245,
namaqua, 237, 239, 241,
252.
pallasi, 234,
personatus, 234, 289,
240,
quadricinctus, 234, 237,
238, 240, 254, 257,
258, 260.
senegalensis, 243, 248.
senegalus, 233, 237, 238,
239, 248, 244, 247.
sepultus, 236.
setarius, 234, 250.
simplex, 252.
tachypetes, 234, 252.
tricinctus, 234, 237, 256,
257, 258, 260.
variegatus, 284, 237,
239, 244.
Pteroclurus
alchata, 251.
exustus, 248.
namaqua, 253.
senegalus, 243.
Pterocyclos
albersi, 499.
parvus, 499.
Pterocypha
stellata, 492.
Pteroglossus
frantzit, 47.
torquatus, 46.
Pterophorus
aspilodactylus, 495.
Pteropus
aneiteanus, 874.
capistratus, 315.
germaini, 874.
hypomelanus, 875.
keraudreni, 875.
medius, 874.
INDEX,
Pteropus
melanopogon, 315.
i ie 315.
vanicorensis, 875.
vetulus, 874.
(Cheiropteruges) albo-
scapulatus, 316.
Ptilinopus
albocinetus, 571.
apicalis, 527.
cesarinus, 545,
chaleurus, 529.
chrysogaster, 541.
clementine, 535.
coralensis, 528,
corriei, 566.
cyanovirens, 543,
diadematus, 540.
dupetit-thouarsi, 539.
emilie, 589.
ewingit, 533.
fasciatus, 535.
felicie, 565.
Jischeri, 571.
flavicollis, 533.
flavigaster, 540,
Surcatus, 541.
gestrot, 557.
huttoni, 588. Q
hypogaster, 558.
pep) 543.
luteovirens, 565.
marie, 545.
melanocephalus, 552.
melanocephalus celeben-
sis, 552.
mercieri, 536.
miquelt, 563.
monachus, 550.
nuchalis, 558.
occipitalis, 569.
pelewensis, 581.
perouset, 545.
porphyraceus, 525.
porphyreus, 553, 554.
purpuratus, 533, 540,
542,
rarotongensis, 534.
roseicapillus, 537.
strophium, 568.
superbus, 543.
swainsoni, 531.
viridis, 560.
viridissimus, 528.
Ptilocolpa
griseipectus, 937, 952.
Ptilogonys
caudatus, 39, 53.
Ptilonopus
apicalis, 527, 530.
aurantiifrons, 556.
Ptilonopus
bellus, 563.
chaleurus, 529.
chrysogaster, 541.
cinctus, 572.
clementine, 535.
coralensis, 528.
coronulatus, 99, 546.
diadematus, 540.
dupetit-thouarsi, 539.
erythrocephalus, 554.
eugenia, 561.
ewingit, 53d.
fasciatus, 5380, 5385.
flavicollis, 533.
flavipectus, 533.
Sformosus, 544.
geminus, 546.
gularis, 570.
humeralis, 548.
hyogaster, 558.
tonozonus, 547.
zozonus, 99, 547.
gambu, 554.
luteovirens, 565.
marginalis, 547.
marie, 54d.
melanocephalus, 552.
merciert, 537.
monachus, 550.
nana, 549.
nanus, 550,
ochrogaster, 573.
pectoralis, 559.
perlatus, 558.
perousei, 545.
ponapensis, 535.
prasinorrhous, 562.
pulehellus, 551.
purpuratus, 525, 588,
542.
purpureicinctus, 587.
rivoli, 561.
roseicapillus, 537.
roseicollis, 554.
roseipectus, 559,
senex, 546.
solomonensis, 561.
strophium, 568.
suluensis, 553.
superbus, 543.
swainsoni, 532.
taitensis, 541.
trigeminus, 547.
virens, 559.
viridis, 560.
viridissimus, 528.
wallacei, 55d.
aanthogaster, 540.
(Cyanotreron) mona-
chus, 550.
ple ite ee ee
Ptilonopus
(Lotreron) iogaster, 559.
(Lamprotreron) —sw-
perbus, 543.
Ptilonorhynchus
melanotis, 99.
Ptilopus
albocinctus, 505, 518,
23, 571, 573.
apicalis, 504, 515, 519,
527, 580, 536.
assimilis, 503, 512, 518,
523, 578, 574, 575.
aurantiicollis, 556.
aurantiifrons, 505, 508,
517, 518, 521, 556.
batilda, 504, 570.
bellus, 506, 508, 517,
522, 563.
bernsteini, 505, 518,
523, 554, 572, 573.
bonapartei, 504, 505,
507, 516, 519, 524,
525.
chaleurus, 505, 515,
519, 529.
chrysogaster, 503, 508,
515, 520, 541.
chrysorrhous, 506, 518,
521, 553.
einetus, 502, 509, 512,
518, 523, 554, 571,
572, 578.
cinctus florensis, 571.
cinctus lettiensis, 506,
572.
clementine, 504, 535,
6.
coralensis, 503, 504, 515,
519, 527, 528, 529,
532.
coronulatus, 99, 505,
511, 517, 518, 520,
546, 547.
corriei, 506, 516, 522,
566.
eyanovirens, 503.
diademata, 502,504,540.
dupetit-thouarsii, 503,
508, 512, 515, 520,
539.
eugenia, 504, 516, 522,
561
ewingit, 503, 504, 507,
518, 519, 526, 532,
533, 534.
fasciatus, 503, 504, 507,
510, 516, 520, 520,
580, 532, 534, 535,
536.
felicie, 504.
INDEX.
Ptilopus
Jischert, 506, 518, 523,
571
flavicollis, 504, 583.
Fformosus, 505, 544, 572,
573.
forsteri, 527.
Furcatus, 503.
geelvinckianus, 506,
511, 517, 522, 560.
geminus, 506, 516, 517,
520, 546, 547.
gestrot, 506, 518, 521,
Date
geversi, 568.
greyi, 504, 507, 516,
519, 523, 524.
gularis, 509, 512, 518,
522, 570, 572, 573.
hugonianus, 568, 569.
humeralis, 505, 517,
521, 548.
humeralis jobiensis, 506,
548, 549.
huitoni, 506, 515, 520,
538.
hyogaster, 502.
insolitus, 505, 508, 512,
513, 516, 521, 549.
togaster, 510, 513, 518,
522, 558, 559.
tonozonus, 99, 547.
tozonus, 99, 505, 517,
521, 547.
Jjambu, 503, 518, 521,
545.
Jobiensis, 506, 508, 517,
521, 548, 549.
Johannis, 506, 516, 522,
564.
layardi, 506, 516, 522,
566, 567.
leclancheri, 504, 518,
522, 568, 569, 573.
lettiensis, 506, 508, 518,
523, 572.
luteovirens, 503, 504,
505, 508, 511, 512,
516, 522, 565, 566,
567.
maculatus, 503.
magnificus, 502, 510,
512, 513, 517, 518,
GBS BB AaT tay
marie, 504, 545.
melanauchen, 552.
melanocephalus, 503,
518, 519, 521, 551,
552, 553, 936, 951.
melanospilus, 506, 518,
521, 552.
1049
Ptilopus
merciert, 504, 508, 515,
520, 536.
miquelt, 563.
monachus, 502, 503,
512, 518, 521, 550.
musschenbroeki, 560,
561.
nanus, 502, 517, 521,
549, 550.
neglectus, 506, 516,
519, 524.
nove-guince, 5d6.
occipitalis, 503, 504,
509, 512, 518, 522,
569, 570.
ochrogaster, 573.
ornatus, 506, 508, 517,
518, 521, 556, 557.
pectoralis, 502, 503,
508, 517, 522, 559.
pelewensis, 505, 516,
519, 531.
perlatus, 502, 509, 512,
513, 517, 518, 522,
557, 558, 573.
perousti, 503, 504, 515,
516, 520, 545.
pictiventris, 507, 515,
516, 519, 530.
ponapensis, 501, 507,
5386.
porphyraceus, 504, 505,
527, 529, 535.
porphyreus, 502, 503,
519, 521, 558, 554,
570.
prasinorrhous, 505, 517,
518, 522, 562.
puella, 512, 516, 517,
573, 574, 575.
pulchellus, 502, 504,
517,521, 5512
pulcherrimus, 503.
purpuratus, 502, 503,
504, 505, 507, 508,
512, 515, 516, 519,
523, 524, 525, 526,
527, 541.
purpureo-leucocephalus,
5038
rarotongensis, 506, 515,
520, 534.
regina, 502, 503, 504,
507, 512, 518, 519,
531, 5382, 542.
rivolit, 503, 516, 522,
561, 562, 563.
roseicapillus, 503, 504,
516, 520, 537.
roseicollis, 503.
1050
Ptilopus
solomonensis, 562.
speciosus, 506, 522, 564,
565.
strophium, 508, 516,
517, 522, 563.
superbus, 121, 126, 502,
503, 504, 511, 512,
516, 517, 518, 520,
542, 548, 544, 545.
swainsoni, 503, 504,
532, 548.
taitensis, 502, 508, 507,
508, 515, 520, 526,
27, 541.
temminckii, 504, 505,
518, 520, 544.
trigeminus, 506, 517,
520, 547.
victor, 506, 508, 511,
516, 522, 566, 568.
virens, 503.
viridis, 506, 518, 522,
560, 561.
viridis geelvinckianus,
506, 560.
viridissimus, 508, 504,
528, 529, 532.
wallacei, 505, 518, 521,
555.
wanthogaster, 502, 504,
507, 518, 520, 540,
541,
swanthorrhous, 506, 508,
518, 521, 553, 951.
zonurus, 506, 518, 522,
558.
Ptilorhis
alberti, 128.
Ptiloris
alberti, 123.
magnificus, 123,
Ptilotis
chrysotis, 124.
filigera, 97, 124.
lewini, 124.
Ptinilopus
marie, 545.
Pudu
humilis, 887, 927.
Puffinuria
garnoti, 739.
Pyecnonotus
barbatus, 129.
nigricans, 356.
Pygarrhicus
albigularis, 433.
1s ornis
Pali. 67, 68.
Pygosceles
teniatus, 653."
INDEX.
’ Pygosceles
wagleri, 653.
Pyranga
bidentata, 55.
erythromelena, 5d.
rubra, 55.
Pyrenestes
capitalbus, 130.
Pyrgisoma
cabanisi, 38, 56.
leucote, 56.
Pyromelana
capensis, 308.
nigriventris, 358.
Pyrrhocentor
melanops, 108, 946.
Pyrrhophzna
riefferi, 88, 70, 71.
Pytelia
melba, 358.
Querquedula
eatoni, 578.
Rallina
euryzonoides, 288, 345.
fasciata, 288.
Rallus
antarcticus, 437.
pectoralis, 120, 127.
Ramphastos
ariel, 988.
tocard, 46.
toco, 933.
Ramphiculus
Jjambu, 554.
occipitalis, 569, 952.
Ramphocelus
passerinit, 5d.
Ramphocenus
semitorquatus, 61.
Ranella
pusilla, 815, 821.
rosea, 815, 816.
Ranghana
punctata, 22, 37.
Rangifer
caribou, 928.
grenlandicus, 928.
tarandus, 887, 927, 928.
Rectes
aruénsis, 96.
dichrous, 96.
dichrous ceramensis, 96.
draschi, 96.
wropygialis, 96.
Remigia
disseverans, 489.
persubtilis, 489.
Retina
rubrivitta, 846, 857.
Rhabdornis
mystacalis, 113, 951.
Rhaphidognatha
hainana, 704, 708.
Rhescipha
elegans, 489.
servia, 489.
Rhinoceros
indicus, 634.
lasiotis, 634, 636.
sondaicus, 634.
sumatrensis, 634, 635,
636, 790.
Rhinolophus
acuminatus, 878.
affinis, 878.
Rhinopalpa4
algina, 645.
amelia, 645.
Rhipidura
assimilis, 87.
griseicauda, 84.
aura, 87.
rufifrons, 84.
setosa, 87, 289, 671.
squamata, 84.
Rhodostethia
rosea, 157, 208, 210.
rossi, 208.
Rhombodera
tectiformis, 580.
Rhyacophilus
glareola, 711.
Rhynchea
australis, 747, 750.
bengalensis, 746, 747.
capensis, 345, 745, '746,
747, 748, 750, 953.
Rhynchobates
djeddensis, 117.
Rhynchocyclus
sulphurescens, 63.
Rhynchonella
nigricans, 413.
psittacea, 400, 418,
414.
rostrum-psittact, 413.
sicula, 403, 418, 414,
416.
Rhynchonycteris
naso, 880.
Rhyparida
costatipennis, 152.
madagascariensis, 152.
nigricollis, 153.
Ricinula
astricta, 807.
Rissa
borealis, 164.
brachyrhynchus, 164.
brevirostris, 164.
Rissa
brunnichii, 158, 164.
gregaria, 164.
kotzebui, 164.
nivea, 164, 177
septentrionalis, 178
tridactyla, 158, 163,
164, 165, 180.
Rissoina
nivea, 867.
Rosema
costalis, 387.
dorsalis, 387.
Sulvipennis, 387.
myops, 387.
zulica, 387.
Rossia
rosea, 208.
Rostellaria
luteostoma, 313.
Rusa
lepida, 902.
peroni, 903.
Ruticilla
mesoleuca, 979.
phenicura, 979.
Salda
aberrans, 462,
471, 477.
Salpinx
crassa, 822, 823, 853.
hobsoni, 695.
klugi, 695, 823.
margarita, 823, 853.
masoni, 823, 853.
minorata, 695, 706.
rhadamanthus, 822,
853.
Saltator
grandis, 38, 56.
magnoides, 56.
Samea
chlorophasma, 493.
Sapphironia
boucardi, 71.
Sarcophanops
steerti, 110.
Sarcops
calvus, 113, 287, 348,
951.
470,
Sarothropyga
rhodopepla, 387.
Satyrus
dorycus, 645.
duponcheli, 645.
verma, 824.
Saucerottia
sophie, 38, 70, 71.
Sauloprocta
melaleuca, 671.
INDEX.
Sauromarptis
gaudichaudi, 94.
tyro, 94.
Sauropatis
chloris, 83, 94, 108,
282, 341, 709, 937.
sancta, 94.
sordida, 94.
Saxicola
rufa, 881.
stapazina, 881, 976.
Sayornis
aquatica, 62.
Scala
friabilis, 866.
(Opalia) australis,
866.
Scalaria
australis, 866.
Friabilis, 866.
Scelida
balyi, 993.
Schismope
atkinsoni, 868.
Schizostoma
behni, 880.
brachyote, 880.
megalotis, 880.
Scissurella
atkinsoni, 868.
Sciurus
adolphei, 663, 664.
e@stuans, 657, 667, 668,
670.
affinis, 660.
albipes, 660.
arizonensis, 659.
aureogaster, 698, 660,
661, 662.
booth, 657, 658, 659,
662, 663, 664.
brunneo-niger, 665,
697, 954.
carolinenis, 657, 658,
659, 669.
chrysosurus, 669.
colliei, 657, 659, 662,
663, 664.
deppei, 668, 669.
dorsalis, 662, 663,
664.
Serruginiventris, 660.
fumigatus, 665, 667.
fisooiiagais, 662,
664.
gerrardi, 657, 658, 665,
666.
gilvigularis, 668.
griseocaudatus,
663, 664.
griseoflavus, 658, 560.
662,
1051
Sciurus
griseogenys, 658, 667,
669.
hoffmanni, 667, 668.
hypopyrrhus, 441, 657,
658, 662, 663, 664,
665, 666.
hyporrhodus, 667.
igniventris, 665, 666.
intermedius, 662, 668,
664.
langsdorffi, 665, 666.
leucogaster, 660.
leucops, 657, 658, 660,
661, 662.
leucotis, 658.
ludovicianus, 660.
melania, 664.
morio, 665, 666, 667.
nebouxit, 441, 664.
nicoyanus, 663.
niger, 661,
nigrescens, 662, 663,
64.
662, 663,
66
pusillus, 658, 668, 670,
954.
pyladei, 663, 664.
‘pyrrhonotus, 665.
richardsoni, 662, 664.
rigidus, 662, 6638,
664.
oculatus,
34,
rufo-niger, 657, 658,
666, 667, 669, 954.
socialis, 660, 662.
stramineus, 441, 658,
664, 665.
tephrogaster, 657, 665,
668
tricolor, 665, 667.
variabilis, 657, 658,
665, 666, 668, 954.
variegatoides, 662, 663,
664.
variegatus, 660, 663.
varius, 660, 662.
vulgaris, 662.
yucatanensis, 657, 658,
659.
Scolopax
egocephalus, 288.
cinereus, 711.
incanus, 345, 711.
stenurus, 83.
Scops
brasilianus, 45.
everetti, 936, 942.
lempiji, 942.
leucospilus, 942.
leucotis, 128. .
1052
Scops
morotensis, 78.
wmbratilis, 942.
Scotophilus
temminckit, 876, 879.
Scotosia
catocalaria, 492.
Scyllium
canicula, 116, 117.
Scytalopus
magellanicus, 433.
Selasphorus
flammula, 40, 70.
scintilla, 40, 70.
Selenidera
spectabilis, 47.
Selenophorus
amblyderus, 591.
callistichus, 591.
mitis, 591.
tenuistriatus, 591.
Semele
aphrodite, 859, 864.
aspasia, 860, 864.
hanleyi, 859, 864.
phryne, 860, 864.
Serilophus
rubropygius, 143.
Serpentarius
reptilivorus, 790.
Serpophaga
cinerea, 62, 63.
Setina
discisigna, 35.
irrorella, 35.
nebulosa, 35.
punetilinea, dt,
Setinochroa,
aurantiaca, 35, 37.
infumata, 35, 36.
pallida, 35.
postica, 36.
Setophaga
aurantiaca, 39, 53.
ruticilla, 52.
torguata, 39, 53.
Sicyonia
Jurcata, 310.
ocellata, 310.
Sidyma
abifints, 9:
apicalis, 9, 36.
Sigmodus
graculinus, 357.
Silybura
broughami, 800.
canarica, 155.
dupeni, 801.
guentheri, 801.
levingti, 801.
diura, 155,
INDEX,
| Silybura
macrolepis, 155.
maculata, 154.
madurensis, 802.
melanogaster, 155, 801.
nigra, 154.
nitida, 154.
ocellata, 155.
ochracea, 801.
petersi, 154.
Simareea
basinota, 14, 36.
Simia
satyrus, 789.
Siphonaria
albida, 314, 869.
Siphonia
elastica, 38.
Sistrum
margariticola, 812.
Sisyrophora
pfeiffere, 850, 858.
Sittace
chloroptera, 77.
spixt, 976.
Siurus
aurocapillus, 51.
noveboracensis, 51,
Solenocera
philippii, 302.
siphonocera, 301.
Sospita
lydda, 701.
neophron, 833.
saturata, 701, 707.
Spatula
platalea, 436.
Speiredonia
helicina, 849.
Spermophila
aurita, 58,
corvina, 58.
moreleti, 58.
semicollaris, 58,
Spermospiza
hematina, 130.
Sphecotheres
flaviventris, 124.
Spheniscus
demersus, 653, 654.
humboldti, 116.
magellanicus, 653.
Sphenoproctus
curvipennis, 68.
Spiloglaux
marmoratus, 121.
Spilornis
holospilus, 340, 939.
Spilosoma
menthastri, 382.
obscurum, 382.
Spilosoma
submacula, 882.
Spinachia
vulgaris, 330.
Spinax
niger, 117.
Spirama
helicina, 849, 857.
Spirula
australis, 971, 973.
Spondylus
varius, 967, 968.
Sporophila
othello, 136.
Squatina
angelus, 117.
Steatornis
caripensis, 139,
Stelgidopteryx
Sulvigula, 67.
Sulvipennis, 67.
Stenele
lutescens, 386.
Stenopus
hispidus, 779.
Stenoiis
olivaceus, 589.
Stepnaya
kuritza, 261.
Sterna
anestheta, 273.
bergit, 346, 712, 937,
976.
cerulea, 211, 272.
cantiaca, 193.
cinerea, 212, 278.
media, 193.
philadelphia, 206.
tereticollis, 272.
teretirostris, 211.
Stictoplea
grotet, 824, 853.
Stolida
cinerea, 211, 272.
Stonasla
consors, 462, 472.
undulata, 462, 472,
477.
Strepsiceros
imberbis, 441.
kudu, 745.
Strepsilas
interpres, 120, 127,
708, 711.
Strix
lugubris, 940.
sandwichensis, 348.
Struthio
camelus, 790,
Sturnella
ludoviciana, 59.
——
Sturnus
ambiguus, 713.
humei, 712, 713.
indicus, 713.
nitens, 713.
poltaratskyt, 713.
purpurascens, 713.
unicolor, 712.
vulgaris, 712, 713.
Sula
cyanops, 652.
leucogastra, 651.
piscator, 651.
Surendra
quercetorum, 836, 855.
Surniculus
lugubris, 613.
velutinus, 945.
Sylphitreron
perlatus, 558.
Sylvia
blanfordi, 979.
curruca, 979.
melanocephala, 978,979.
rubescens, 979
Symmachia
domitianus, 363.
rhacotis, 363.
Symphedra
dirtea, 629, 699.
pardalis, 699, 706.
Synallaxis
erythrops, 99.
nigrifumosa, 59.
pudica, 59.
Synaptomys
cooperi, 633.
Synchloé
canidia, 701, 707.
limbolaris, 487.
Syngamia
florella, 492.
Syncecus
cervinus, 121, 127.
Syntomis
albifrons, 845, 857, 859. |
atkinsoni, 845, 857.
berinda, 845, 857, 559.
diaphana, 845.
disrupta, 845, 857, 858.
divisa, 845.
fenestrata, 845.
grotet, 844, 857.
libera, 845, 857.
masoni, 845, 857, 858.
melas, 845.
sladeni, 844, 857.
vitreata, 845.
Syrnola
brunnea, 866.
tincta, 866.
|
INDEX.
| Syrrhaptes
heteroclita, 261.
pallasti, 234, 261.
paradoxus, 233, 236.
238, 246, 260, 261.
thibetanus, 234, 236,
238, 260, 263.
Systropha
aureola, 18.
auriflua, 18, 37.
dorsalis, 18.
Tachyeres
cinereus, 437.
Tachyphonus
delattrii, 55.
luetuosus, 55.
Tachys
diploharpinus, 602.
Tzenioptera
obscura, 347.
pyrope, 433.
Tagiades
meetana, 842,
858.
obscurus, 842.
pralaya, 842, 856.
Talegalla
lathami, 127.
Tanagra
cana,
diaconus, 54.
melanoptera, 5D.
palnarum, 5d.
Tantalus
ibis, 625, 627.
loculator, 625, 626, 627,
629.
Tanygnathus
luzonensis,
612, 937.
megalorhynchus, 78.
apes
Alabagella, 870.
Taphozous
melanopogon, 879.
Tapirus
eucogenys, 632.
roulint, 631.
terrestris, 632.
Tarika
nivea, 15.
varanda, 15, 36.
Tatusia
hybrida, 228, 229.
peba, 226, 228, 229.
856,
281, 340,
Ak
| Taxila
drupadi, 832.
fasciata, 882, 855, 858.
Tegulata
basistriga, 22, 36.
protuberans, 25, 36.
tumida, 22.
Proc. Zoou. Soc.—1878, No. LXX.
1053
Telephonus
erythropterus, 357.
Tellina
wroblewskyi, 727, 732.
Temora
velox, 330.
Teracotona
roseata, 382.
submacula, 382.
Terebra
affinis, 805.
brazieri, 866.
exigua, 805.
ustulata, 866.
Terebratella
buckmanni, 405.
frontalis, 406, 407.
marie, 407.
spitzbergensis, 399, 406,
407, 408, 409, 415.
Terebratula
affinis, 404.
ampulla, 403.
aperta, 409.
appressa, 411.
arctica,
benoitiana, 404.
buckmanni, 405.
capsula, 410.
caput-serpentis,399,401,
402, 404, 410, 415.
cardita, 409,
cernica, 404.
cistellula, 410.
cordata, 409.
cranium, 397, 399, 405,
406, 407, 408, 410.
cubensis, 404, 408.
euneata, 410.
davidsoni, 404.
dimidiata, 409.
euthyra, 407, 408.
glabra, 407.
grandis, 403, 404.
irregularis, 405.
Jelfreysii, 406.
manticula, 404.
michelottiana, 404.
minor, 399, 404.
miocenica, 404.
monstruosa, 412.
orbiculata, 408, 404.
pectiniformis, 409.
plicata, 407.
scobinata, 410.
seminulum, 409, 411.
septata, 399, 405, 406,
407, 408, 412, 415.
septentrionalis, 401.
septigera, 405, 408.
soldaniana, 410.
sphenoidea, 399, 405,
404, 415, 416.
70
1054
Terebratula
subguadrata, 399, 400,
402, 403, 415.
subvitrea, 407.
tenera, 399, 400, 405,
415.
tri ee, 399, 400, 402,
ee 399, 401, 415.
ungula, 409.
urna-antiqua, 409.
vitrea, 399, 403, 404,
406, 407, 410, 415,
416.
Terekia
cinerea, 579, '708, 711.
Terias
esiope, 699, 700, 707.
arcuata, 700, 707
attenuata, 700, 707.
drona, 700.
elathea, 481.
euterpe, 481
formosa, 836, 855.
gnathene, 481.
hainana, 700, 707.
hecabe, 641, 647, 836,
855.
feta, 836, 855.
silhetana, 836, 855.
suava, 836.
subdecorata, 699, 700,
707
wagans, 700.
Terinos
sinha, 828. |
Testudo
greca, 976.
horsfieldi, 975, 976.
pardalis, 790.
Tetracha |
ignea, 587.
Tetragonops i |
Ffrantzii, 39, 47. |
ramphastinus, 952.
Tetrao
alchata, 238, 284, 237,
250.
arenarius, 283, 245,
261
chata, 251 5
Jasciatus, 233, 234,
245
indicus, 233, 254.
namagua, 233, 234,
252.
paradoxus, 233, 234,
237, 260.
senegalus, 233, 243.
subtridactylus, 246.
urogallus, 746. |
|
Tetrodon
nigropunctatus, 133.
INDEX.
Teulisna
plagiata, 24.
tenuisigna, 24, 37.
tetragona, 24.
Thaduka
multicaudata, 836, 855,
858.
Thalassidroma
gracilis, 736.
leucogastra, 735.
melanogastra, 736.
segethi, 736.
tropica, 736,
Thalassina
scorpioides, 778.
Thalasseca
antarctica, 737.
glacialoides, 736.
Thalurania
colombica, 69.
Thamala
miniata, 834, 855,
858.
Thamnobia
lugubris, 803.
Thamnophilus
doliatus, 60.
nevius, 60.
nigricristatus, 60.
punctatus, 60.
simplex, 139.
Thamnotettix
sancte-helene, 462,
476.
Thanaos
indistincta, 694.
obsoleta, 694.
stigmata, 694.
Tharops
eoruscans, 366.
isthmica, 366.
menander, 366,
nitida, 366.
purpurata, 365.
splendida, 366.
Thaumantis
howqua, 827.
louisa, 827, 854.
Thaumatias
cupreiceps, 7}.
Thecidea
mediterranea, 400,
412.
spondylea, 412.
testudinaria, 412.
Thelphusa
depressa, 755.
perlata, 755.
; Theope
aureonitens, 369.
barea, 369.
cenina, 369.
decorata, 369.
Theope
eudocia, 368.
hypoxanthe, 368.
isia, 868
Janus, 369.
Theristicus
melanops, 436.
Thermesia
gemmatalis, 489.
Thestias
sesia, 700, 707.
Thouarsitreron
diademata, 540.
dupetit-thouarsi, 539.
leucocephala, 539.
Thrasaétus
harpyia, 44.
Thriponax
Javensis, 340, 942.
pectoralis, 340, 379.
Thryophilus
castaneus, 5.
modestus, 5}.
pleurostictus, 51,
thoracicus, 51.
Thyca
indica, 839.
pasithoé, 701, 707.
Thyroptera
tricolor, 872, 873.
Thysania
zenobia, 488.
ga
everetti, 611, 612.
Jjavanensis, 612.
Timetes
corita, 270.
phiale, 270.
Tinamus
Frantzii, 41.
Tinea
pulchella, 298.
Tinnunculus
cinnamominus, 435.
dominicensis, 485.
isabellinus, 435.
leucophrys, 435.
sparverioides, 435.
sparverius, 434.
Tinoleus
eburneigutta, 851,
858.
Tityra
personata, 65.
Tivela
undulosa, 870.
Tmolus
columella, 481.
pan, 481.
Tockus
melanoleucus, 308.
Todirhamphus
sanctus, 121.
Todirostrum
cinereum, 62.
Todus
viridis, 101.
Tolypeutes
conurus, 222, 223, 224,
225, 228.
muriet, 223, 224, 225.
tricinctus, 222, 228,
224, 225, 226, 227,
228.
Torinia
perspectiviunculus, 816, |
821
planulata, 816.
Tornatina
fusiformis, 869.
Totanus
glareola, 80.
ariseopygius, 127.
incanus, 121, 127, 345, |
301, 711.
Trabala
vishnu, 848, 857.
Tragelaphus
angasi, 884.
decula, 884.
spekit, 884.
sylvaticus, 884.
Tragulus
kanchil, 882.
stanleyanus, 682.
Trepsichrois
midamus, 823, 8538.
thoosa, 638.
Treroleema
leclancheri, 504, 568,
569.
Treron
delalandit, 359.
nasica, 623.
Trichoglossus
cyanogrammus,
579.
massene, 93.
multicolor, 126.
nigrigularis, 93.
subplacens, 671.
swainsonii, 126.
Trichostoma
abboti, 616.
rufifrons, 611, 616.
Trigla
pini, 218.
Trigonia
margaritacea, 871.
Tringa
acuminata, 289, 290,
albescens, 708, 712.
fasciata, 2383, 254.
fuscicollis, 4388,
interpres, 711.
platyrhyncha, 712.
93,
|
INDEX.
Tringoides
hypoleucus, 83, 90, 100,
289, 345, 624, 711,
937.
macularius, 44.
Trinophylum
cribratum, 720,
Triphoris
scitulus, 867.
sculptus, 817.
Tripura
prasend, 8.
Triton
laciniatuim, 815.
(Epidromus) bednalli,
864.
(Simpulum) strange,
816, 821.
Trivia
australis, 867.
Trochilus
colubris, 70.
Trochita
calyptreformis, 867.
Trochus
calyptreformis, 867.
(Clanculus) microdon,
818, 821.
(Forskahlia) pulcherri-
mus, 818, 821.
(Monilea) calyeulus,
818.
——) masoni, 818.
(Polydonta) maculatus,
730.
(Thalotia) yokohamen-
sis, 727, 732.
Troglodytes
hypaedon, 51.
intermedius, 51.
Trogon
massena, 48.
puella, 48.
Tropidorhynchus
buceroides, 124.
plumigenys, 88, 289.
Trygon
dupetit-thouarsi, 539.
| Trygonorhina
Ffasciata, 117.
Tudicula
arnigera, 610.
inermis, 610.
spinosa, 610.
(Murex) spzrillus, 611.
Turacena
crassirostris, 673.
| Turbinella
Sfastigium, 812.
incarnatus, 813.
| Turbonilla
acicularis, 867.
angast, 867.
|
|
1055
Turbonilla
fusca, 867.
hofmani, 867.
nitida, 867.
Turdus
armillaris, 79.
chrysolaus, 429.
falklandicus, 431.
grayt, 38, 50.
icterorhynchus, 129.
leucauchen, 50.
mandarinus, 578.
nigrescens, 40, 50.
obscurus, 429.
obsoletus, 40, 50.
pelios, 129.
plebeius, 50.
varius, 429.
Turnix
maculosa, 578.
melanota, 127.
Turritella
infraconstricta, 817,
821,
Turtur
dussumier?, 288, 344,
711, 937, 952.
Typhis
angasi, 864.
Tyranniscus
parvus, 63.
Tyrannus
meiancholicus, 64.
Urania
sloanus, 490.
Urapteryx
podahriata, 850, 858.
Uromys
rufescens, 274.
Urospatha
marti, 39, 49.
Urospiza
poliocephala, 91, 92.
Urospizias
iogaster, 80.
poliocephalus, 91.
spilothorax, 92.
Ursus
americanus, 878.
arctos, 378.
Feroz, 378.
isabellinus, 378.
malayanus, 378.
maritimus, 378.
ornatus, 378.
piscator, 378.
syriacus, 378.
tibetanus, 378.
Urubitinga
ghiesbreghti, 44.
Utethesia
pulchella, 704, 708.
Vamuna
- bipars, 10, 36.
maculata, 10, 36.
remelanda, 10.
Vanellus
cayennensis, 437, 625,
627, 628.
occidentalis, 437.
Vanessa
amelia, 645.
Vates
ashmolianus, 584.
Venus
levigata, 870.
undulosa, 870.
variabilis, 870.
Vermetus
gigas, 966, 967.
Vernonia
wollastoniana, 462, 470,
477.
Vespertilio
adversus, 876.
muricola, 879.
murinus, 872.
temminckit, 876.
Vesperugo
abramus, 879.
nanus, 879.
Vidua :
principalis, 130.
Vireosylvia
flavifrons, 53.
flavoviridis, 53.
josephe, 53.
olivacea, 53.
philadelphica, 53.
Volatinia
jacarina, 57.
Volvocivora
cerulescens, 283, 284.
mindanensis, 936, 947.
morio, 948.
Volvula
rostrata, 869.
Vulpes
cana, 892.
Vultur
atratus, 435.
aura, 435.
papa, 435.
Waldheimia
cranium, 399, 407.
davidsoniana, 407.
Sa
Printed by TayLor and Fr
INDEX.
Waldheinia
floridana, 407, 408.
kergquelensis, 411.
peloritana, 408.
raphaélis, 407.
Xantholema
hemacephala, 944.
rosea, 282,
Xanthotenia
busiris, 827, 854.
Xanthotis
chrysotis, 97.
Jiligera, 97.
Xema
atricilla, 194.
bonapartii, 207.
brunneicephala, 197,
198.
caniceps, 200.
capistrata, 200.
cirrhocephala, 196, 197,
201, 202, 2038.
collaris, 209, 210,
crassirostris, 193.
Franklini, 195.
Surcata, 157, 210.
gelastes, 192.
genet, 192.
glaucodes, 203.
ichthyaetus, 198.
Jjamesont, 185.
kittlitzii, 205.
lambruschinii, 192.
leucophthalma, 193.
melanocephala, 199.
minuta, 206.
pheocephala, 204.
pileata, 200.
pipixcan, 195.
ridibunda, 200.
sabinti, 141, 158, 209, |
211.
Xenelaphus
anomalocera, 924.
chilensis, 924.
huamel, 924.
leucotis, 924.
Xenophora
solarioides, 818.
Xenurus
unicinctus, 228, 229.
Xiphopeneus
harttii, 305.
Xiphorhynchus
pusillus, 60.
Xylophis
indicus, 576.
Ml } =U
ancis, Redayi
Hs,
| Xystosomus
belti, 601.
olivaceus, 601.
Yphthima
baldus, 825, 853.
methora, 825, 853.
pandocus, 696, 706.
zodiaca, 696, 706.
Ypsipetes
anomala, 491.
Yungipicus
aurantiiventris, 943.
Ffuscoalbidus, 943.
validirostris, 943.
Zadadra
distorta, 25, 37.
Zamenis
cliffordi, 977, 978.
elegantissimus, 977.
Zebronia
plutusalis, 849, 857.
Zemeros
albipunctata, 701.
confucius, 701, 707.
Alegyas, 701, 832, 855.
Zenaidura
carolinensis, 43.
Zeocephus
cinnamomeus, 616.
eyanescens, 616.
Zethera
diademoides, 824, 853,
858.
Zeuxidia
amethystus, 826.
horsfieldi, 639.
masont, 826, 854.
Ziziphinus
jucundus, 798.
Zonotrichia
canicapilla, 432.
pileata, 38, 57, 482.
vuleani, 40, 57.
Zophoéssa
mutrheadi, 696, 706.
Zosterops
austeni, 286.
chloris, 84.
everetti, 110, 950.
intermedia, 85.
luteus, 121, 125.
nigrorum, 280, 286.
Zygena
malleus, 116.
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