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UNIVERSITY OF
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mt
a ny im i i
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THE ANNALS /A\
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MAGAZINE OF NATURAL HISTORY,
INCLUDING
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eee
VOL. XVI.—SEVENTH SERIES.
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ex ceconomid in conservatione, proportione, renovatione, potentia majestatis
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voir qu’elle est le chef-d’ceuvre de la Toute-puissance, et le but auquel se rappor-
tent toutes ses opérations.”—Brucxyer, Théorie du Systeme Animal, Leyden,
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Seglais Ce se s,.% 4» be pylyan powers
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But scatter round ten thousand forms minute
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That drinks the rippling tide: the frozen poles,
Where peril waits the bold adventurer’s tread,
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All, all to us unlock their secret stores
And pay their cheerful tribute.
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Q@H
AG
sA.7
v. /6
CONTENTS OF VOL. XVI.
[SEVENTH SERIES. }
NUMBER XCI.
Page
I, Report on the Schizopods collected by Mr. George Murray,
F.R.S., during the Cruise of the ‘Oceana’ in 1898. By E. W. L.
Horr and W. M. Tarrersart, B.Sc. (Plates I. & Il.)..........
IL. Descriptions of new Coleoptera of the Family Cetoniide from
British New Guinea. By Oniver E. Janson, F.ES.........04.. Il
II, A uew Noctuid from Sierra Leone. By W. J. Hottanp .. 18
IV. A new Genus and Species of Cynipide from South Africa,
representing a new Subfamily. By P. CAMERON............. Hanne 20)
V. Rhynchotal Notes—XXXIII. By W.L. Distant ........ 22
VI. A List of the Freshwater Fishes of Africa. By G. A.
OMIA GMS OCIS a= #5, aries uaed cng dD» ax, g violas nue tladlele daled 60%
VIL. A Revision of the Fishes of the American Cichlid Genus
Cichlosoma and of the Allied Genera, By C, Tarr Reean, B.A... 60
VIII. Revised Nomenclature of the Species described in Bate and
Westwood’s ‘British Sessile-eyed Crustacea.’ By Canon A. M.
TD OTEMIUNi ig nl Ye erred OE OR Bien 7) Bal BJS diel staf Faery ear eee Cn re 78
IX. Kerunia, a Symbiosis of a Hydractinian with a Cephalopod.
By Dr. Francis Baron Nopcsa, Member of the Hungarian
eaoricalmocicty.. (Plate Wi) sais. vaslee on wercests aves oauvee 95
The Nomenclature of Types in Natural History, by Charles
PE MUCHOEL AMA S. Sa UCK MAM.’ - cc wa spelesyeus esas sy cstvns nes 102
NUMBER XCIL.
X. A List of the Batrachians and Reptiles collected by Dr. W. J.
Ansorge in Angola, with Descriptions of new Species. By G. A.
EMORENCHRy E.Ev.o” (elate TV.) 8. alten Gece wlewes etas a 105
XI. On the Internal Parasites of the Tweed Salmon. By James
Rk. Tosu, M.A., D.Sc., Assistant Professor and Lecturer on Natural
History in the University of St. Andrews. (Plate V.) .......... 115
XII. On Hermaphroditism and Vestigial Structures in the Repro-
ductive Organs of Testudo greca. By H. B. Fanruam, B.Se. Lond.,
A.R.C.S., University College, London. (Plate VI.) ........... . 120
126
XIII. Reply to Mr. G. A. Boulenger. By Nits Rostin ......
lV CONTENTS.
XIV. List of the Lizards in the Zoological Museum of Lund, with
Descriptions of new Species. By Nits Rostn, Zool. Inst. Lund.
(Plates VIL—EX.) ci eit oj. oe betapricte ae eon eae ee Cate
XV. Notes on Eastern and Australian Heterocera, with Descriptions
of One new Genus and Thirteen new Species. By Colonel CHaRLzEs
Swinton, M.A. BLS) &¢... cose. goers ene ere eee
XVI. On a new Species of River-Crab from Yunnan. By W. T.
CatMaNn, D.Sc., British Museum (Natural History) .... .........
XVII. On some new Genera and Species of Parasitic Hymenoptera
from Borneo, ‘By BACAMERON -o-5-7my a bs iterate eee
XVIII. On a Second Collection of Mammals obtained by Dr.W. J.
Ansorge in Angola, By OLDFIELD THomas and R. C.WRovuGHrTon.
XIX. Descriptions of Three new Snakes discovered in South
Arabia by Mr. G. W. Bury. By G. A. Bouvenesr, F.R.S.......
XX. Descriptions of new Tailless Batrachians in the Collection
of the British Museum. By G. A. BouLEenGer, F.R.S...........
XXI. Descriptions of new Species of Marine Shells, chiefly from
Ceylon. By G. B.SowERBy PES. (27 se. ee ce py ieee
XXII. Description of a new Shell from the Cape Verd Islands.
By) Gr. yB. SQWERBY, BS. cores, ciate Soetoro eee
XXIII. Descriptions of new Species of Land-Shells from British
New Guinea, and Remarks on Two Species from the Solomon
Islands. By sinGan-A.. Sates: Ooi onc es acl ee eee
XXIV. Notes on the Tabani from the Palearctic Region in the
British Museum Collection. By GErrruDE RicARDO ..........
XXV. Rhynchotal Notes—XXXIV. By W.L. Distanr ....
XXVI. Descriptions and Records of Bees.—I._ By: T., 1; 7As
CocKERELL, University of Colorado” pe cece a ee eee
XXVII. A Revision of the Fishes of the American Cichlid Genus
Cichlosoma and of the Allied Genera. By C. TATE REGAN, B.A. ..
XXVIII. On the Bats of the Rhinolophus philippinensis Group,
with Descriptions of Five new Species. By Knup ANDERSEN ....
New Books :—Preliminary Report on the Geology and Water Re-
sources of Nebraska, west of the One-hundred-and-third
Meridian. By N. H. Darton.—The Land and Sea Mammals
of Middle America and the West Indies. By Davin Giraup
Extiot, F.R.S.E. &e. Vol. IV. Parts I. & 11.—Guide to the
Gallery of Birds in the Department of Zoology of the British
Museum (Natural History)—House, Garden, and Field. By
Page
129
La. Mran, PVRS. oo sy ce ceca 6 oe oe 258—261
Note on the ‘Museum Humfredianum,’ 1779, by C. Davies Sherborn ;
Where are the Types ?, by 8S. S. Buckman
91 ee 262, 264
CONTENTS. ' Vv
NUMBER XCIIL.
Page
XXIX. Rhynchotal Notes—XXXV. By W.L. Disrant...... 265
XXX. On the Bats of the Rhinolophus arcuatus Group, with
“escriptions of Five new Forms. By KNUD ANDERSEN.......... 281
XXXI. On the Bats of the Rhinolophus macrotis Group, with
tescriptions of Two new Forms. By KNup ANDERSEN.......... 289
XXXII. Descriptions and Records of Bees.—II. By T. D. A.
DCKER EEL, University, Of ColoradOn vo. joie os os tra wees ee ays ue 5 202
XXXIII. Descriptions and Records of Bees.—III. By T. D. A.
JOCKBRELE,, Wmiversity Of Colorado cc. csc cic s cee eye eae 301
XXXIV. New Neotropical Chrotopterus, Sciurus, Neacomys,
Joendou, Proechimys, and Marmosu. By OutpFinLtp THomas...... 308
XXXY. On a remarkable new Squirrel from Burma. By Oxp-
. TEI LGAs) eae on peop miedo. Commoon coc clbecoe ucpidrar Coot ar 514
XXXVI. A Revision of the Fishes of the American Cichlid Genus
Nchlosoma and of the Allied Genera. By C. Tarr Reaan, B.A... 316
XXXVII. On new Species of Histertde and Notices of others,
Bp eM ENCES Helis) pi EAGOL NG A ' syeteid ete cicteifels sik swerve 0'oh24 8s 340
XXXVIII. Notes on the Satin Bower-bird (Ptilonorhynchus vio-
taceus). By ARTHUR G. ButuER, Ph.D., F.L.S., F.Z.S., &c....... 350
XXXIX. On Cucumaria Montagu, Fleming. By Canon A. M.
PATepEMNeNNE es CEsLALOT MEL. IM Meretoeea Siehs tate viste -) vats evehela sta afeidy 0 el ed 352
XL. Ophiopsila annulosa (M. Sars), a British Ophiurid. By
PO ena ree eae Vt IN CoM AN Reread Neer ciavehe. -teid! or c/aisre, sfahe ele a yrsksla's. » ¥ «)sbs 360
XLI. A Collection of Fishes made by Dr. H. Gadow in Southern
Mexicans Dy Gr CATH IOMGAN Ae). vhienk: yield Sie ces ose tle 361
XLII. Descriptions of Three new Fishes from Japan, collected by
Mri. Gordon Smith. By C-Tarm Regan, B.A... cece ces 365
New Book :—Memoirs of the Geological Survey of India. Pale-
ontologia Indica. Series XV. Himalayan Fossils. Vol. 1V.
The Fauna of the Spiti Shales. By Dr. Vicror Uutie, Pro-
fessor of Geology in the University of Vienna .............. 565
NUMBER XCIV.
XLII. Descriptions of new Species of Noctuwide in the British
Museum. By Sir Grorexr F. Hampson, Bart., F.Z.8., &. 2.0... 369
XLIV. Some Observations on the Development of an Asterid with
Large Yolky Eges from the Franklin Islands. By E. H. 1enpERson,
M. Se., M.D., McGill University, Montreal. (Plates XII. & XIII.) . 887
XLV. Descriptions and Records of Bees.—IV. By T. D. A.
Secumnmer,, University Of COlOradO: oct. veces cee eect earn nals 392
vi CONTENTS.
Page
XLVI. Some Results of the North-Atlantic Fin-Whale Fishery.
iBy THOMAS SOUTH WEUL, FUZIS! tier cine amcylc: chive te n> iaige sitll 403
XLVII. Onsome Australasian Mammals. By OLpFieLp THomas, 422
XLVII. Notes on the Distribution of some Species of Terrestrial
Tsopoda introduced into Australasia. By Cuarius Curiton,M.A.,
D.Se., F.L.S., Professor of Biology, Canterbury College, New
Fenland. «thas cca awe. sissies Osha COI ete lomantene CRoloer eke ieiekt tee eee 428
XLIX. An undescribed Guereza. By R. LypEKKER.......... 432
L. A Revision of the Fishes of the American Cichlid Genus
Cichlosoma and of the Allied Genera, By C. Tare Ree@an, B.A. .. 433
LI. A New Explanation of the Red Colour in the Hind Wing of
Catocala; Schr. By Ca. SCuAPOSCHNIKOW (21. -5 =. sea dele ele een 445
LIT. On the Mus orthodon of Hensel. By Capt. G. E. H.
BARRETR=ELAMIETONG. «412s ats ore os eles esle hiatonere ere ee kel tele aaa 452
New Books :—Fconomic Resources of the Northern Black Hills. By
J. D. Irvine. With Contributions by 8S. F. Emmons and
T. A. Jacaar, Jr.—lIowa Geological Survey. Vol. XIV.
Annual Report, 1903, with Accompanying Papers. SAMUEL
Carvin, A.M., Ph.D., State Geologist. T. Savace, Assistant
State Geologist. — Maryland Geological Survey. Miocene.
Parts I. & Il.—J. H. Fazer. Souvenirs Entomologiques,
Etudes sur l’Instinct et les Noces des Insectes. (Neuvieme
Series) Were caine ony eres tone ae iar Loeimde wos 453—459
Proceedings of the Geological Society. .....c2:..sevsccssee 460, 461
The Generic Names given by Frisch in 1775, by Oldfield Thomas
and Gerritt S. Miller, Jun.; The Secondary Appendage of the
Upper Antenne as a Character in the Amphipoda, by Alfred
OP Walker’ 0S ita ii dte cra stere cic ete matt neaversteaste saieners 461, 464
NUMBER XCV.
LIII. Descriptions and Records of Bees—V. By T. D. A.
CockKERELE, University of Colorado {52 i.)s. 1): ei eee 465
LIV. Descriptions and Records of Bees—VI. By T. D. A.
CocCKREELL, University of Colotado” (hoi... cens.s0- 0 coon eee 477
LV. Notes on the Forficularia.—IX. On new Species, with
Synonymic Notes. By Matcorm Burr, B.A., F.L.S., F.E.S. .... 486
LVI. On HMipposiderus diadema and its closest Allies. By Kyup
INNDRSION: 9 sph hisiejois soe pisiese'« sea 8s pusle «wenn ele eee ee eee 497
LVI. A Revision of the “Genus” Peneus, with Diagnoses of
some new Species and Varieties. By A. Atcock, M.B., LL.D.,
LDA SR CRY Yk d eer io aL eM Ra ri en Oe todo SO 5b Anbhs oty +c 508
CONTENTS. vil
Page
Museum. By Sir Guorce F. Hampson, Bart., LVL S ESTAS rode Oe 533
LIX. Descriptions of Two new Species of Diurnal Lepidoptera
belonging to the Subfamily Nymphahne. By Hersertr Druce, ae
TGS Cec Sats ook RSIS Ge rong ei0e o pt Se Oia yee sO ea HAS
LX. Description of a new Land-Shell from Christmas Island. =
By Ene@ar A. Smtrn, 1.8.0. 22... cece eee eect eee eens ceees 551
LXI. On remarkable Specimens of Rana esculenta from South- A
western Persia. By G. A. Bourencer, F.R.S. .....+..-..-+++, 552
LXII. Rhynehotal Notes—-XXXVI. By W.L. Distant .... 553
LXIII. Notes on British Copepoda: Change of Names. By
PEG MASE SCONE li) eel eny tere. she sun's asm y wi/eisire as os isle y siete s 567
LXIV. Note on Hevanchus griseus. By C. Tarr Reaan, B.A... 571
LXV. Anew Genus and Two new Species of Bats. By OLDFIELD
THOMAS
New Book :—Coloration in Polistes. By W1LHEI MINE M. EntEMAN. 576
NUMBER XCVI.
LXVI. Descriptions of new Species of Noctwde in the British
Museum. By Sir Grorecs F. Hampson, Bart., F.Z.8., &c. ...... 577
LXVII. On new Species of Histeride and Notices of others. By
(SISLLTN ESPN 8) GUS DEBE oR Se DAR ee a 604
LXVIII. On new and little-known Species of Eastern and Austra-
lian Lepidoptera. By Col. CoarLes Swinuos, M.A., F.L.S., &e... 612
LXIX. Notes on the various Forms of <Arvicanthis pumilio,
SPALEMngs Bye tv Oe WROUGHTON . cc.% tarts se sd. Aeainlpa wet os 629
LXX. On the Habitat of Rana Blanfordii. By G. A. BouLenGEr,
Be EeSelaos ssi. Ae eta bis Heed Ln niece chbaes 1s i et a
640
LXXI. On a small Collection of Fishes from the Kasai River
(Congo); By, G. A. BouLmNGEr, FRiSt fe cousin gots dood hee ciee tb.
LXXII. On a Collection of Fishes from Lake Bangwelo. By
(SUN LELD Ina ahs (G08 oe 1 SS Se , 642
LXXIII. A List of the Species and Subspecies of the Genus
Rhinolophus, with some Notes on their Geographical Distribution.
By Kyup ANDERSEN
LXXIV. On the Oscules of Cinachyra.
By R. Kirxparricx.
(Plate XIV.)
e\ 9) -¢) ein) 'ey'0! 6) @ 8) '0) 0) Rie) «vv! 5) 0) me 3) 0 8
Vill CONTENTS.
Page
LXXV. Rhynchotal Notes —XXXVIL By W.L. Distant .. 668
LXXVI. Note on some British Culicide. By Cuas. 0. WATER- -
ELOU SI, AE aE Sie, fe. dale wks abe eal lo he hare ke arched Paonia ee 674
On the Affinities of Herpetomonas subuiata, and the Phylogeny of
the Trypanosomes, by Louis Léger ...........0.seceersemes 676
TMOCX Pees. tears ONO BAS oso oder auc SOS cara hte tenses Seages vole ONO
PLATES IN VOL. XVI.
oe Schizopods collected by the ‘ Oceana.’
III, Kerunia cornuta, from the Eocene of Egypt.
IV. Batrachians and Reptile from Angola.
V. Internal parasites of the Tweed salmon.
VI. Hermaphroditism &c. in Testudo greeca.
Vit
VII. | Lizards in the Zoological Museum of Lund.
IX.
X. New species of Histeride.
XI, Cucumaria Montagui.
XII.
XIII.
XIV. Cinachyra barbata.
Development of an Asterid with large yolky eggs.
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
(Ct aspbdbasescnondad per litora spargite muscum,
Naiades, et circiim vitreos considite fontes:
Pollice virgineo teneros hic carpite flores:
Floribus et pictum, dive, replete canistrum.
At vos, o Nymphe Craterides, ite sub undas ;
te, recurvato variata corallia trunco
Vellite muscosis e rupibus, et mihi conchas
Ferte, Dee pelagi, et pingui conchylia succo.”
N. Parthenii Giannettasi, Bel. 1.
No. 91. JULY 1905.
I.—Report on the Schizopods collected by Mr. George Murray,
F.R.S., during the Cruise of the ‘Oceana’ in 1898. By
E. W. L. Hour and W. M. Tarrersatt, B.Sc.
[Plates I. & II.]
Tue material dealt with in the following paper was collected
in November 1898 during the cruise of the ‘ Oceana’ in
deep water off the west coast of Ireland some 200 miles west
of Valentia. The area in which the collections were made
lies between lat. 52° 4” 5” N. and 52° 27’ 6’’ N., and long.
12° 27’ W. and 15° 53’ 9” W., the depth of water varying
from 453 to 1835 fathoms. This region is thus partly within
and partly without the British Area as defined by Norman.
The method of collecting was by open serial tow-nets
fished horizontally at known depths and hauled, still fishing,
to the surface. It is obvious therefore that part of the catch
of any net may have been made during its descent or ascent.
We have thus no reliable guide as to the exact depth at
which a species was caught, though a reasonable inference of
the inhabitants of the various zones.of water can be obtained
Ann. & Mag. N, Hist. Ser. 7. Vol. xvi. I
2 Messrs. EK. W. L. Holt and W. M. Tattersall on
by comparisons of the contents of the different nets in each
vertical series.
The classification and terminology used in the present
report are the same as those recently used by us in dealing
with Schizopods from the Atlantic slope *.
The species of Schizopods taken by the Oceana’ number
nine, as follows :—
Division EUCARIDA, Calman.
Order EUPHAUSIACEA, Boas,
Family Euphausiide, Dana.
Subfamily Evruausin», H. & T.
Genus Euphausia, Dana.
E. pellucida, Dana.
Genus Meganyctiphanes, H. & T.
M. norveyica (M. Sars).
Subfamily Nematoscetina, H. & T.
Genus Thysanoessa, Brandt.
ip longicaudata, Kroyer.
T. gregaria ?, G. O. Sars.
Genus Stylocheiron, G. O. Sars.
S, Suhmii, G. O. Sars.
S, abbrevia/um, G. O. Sars.
Division PERACARIDA, Calman.
Order MystpAcEA, Boas.
Family Mysidz, Dana,
Subfamily Lepromystnm, Norman,
Genus Katerythrops, H. & T.
K, Oceane, H. & T.
Family Lophogastride, G. O. Sars.
Genus Gnathophausia, Willemoes-Sulm,
G. drepanephora, HU, & T,
Family Eucopiide, G. O. Sars,
Genus Lucopia, Dana.
E. australis, Dana,
* ‘Report Inland and Sea Fisheries of Iveland for 1902 and 1903,’
pt. ii, App. iv. [1905].
the Schizopods collected by the ‘ Oceana.’ 3
Of these species two were found to be new to science, viz.
Katerythrops Oceane and Gnathophausia drepanephora,
Descriptions and figures of these have already appeared
(H. & T. loc. cit.), but it has been thought advisable to give
brief diagnoses in the present report as well.
Three of the above species were taken by the ‘ Oceana’
for the first time in British waters, viz. :—
Euphausia pellucida.
Thysanoessa gregaria.
Stylocheiron Suhmit.
Stylocheiron abbreviatum and Eucopia australis were ouly
taken in those hauls made outside the British Area, and so
cannot be admitted to the British list on the evidence of the
‘Oceana’ collections, though both species have since been
met with in British waters (H. & T. loc. cit.).
Schizopods occurred in thirty out of a total of forty *
havls with the tow-nets—a very fair proportion. It is
interesting to note that no Sckhizopods occurred in the
surface-hauls at any of the stations, nor in any net fished at
less than 2380 fathoms, while seven of the nine species were
taken only in those nets towed at greater depths than 500
fathoms. Very few hauls were made, however, at less depths
than 500 fathoms, which, in so far as Schizopods are con-
cerned, is to be regretted, since many of the species captured
have been shown to belong essentially to the upper strata by
the collections made in the Bay of Biscay in July 1900 by
Dr. Fowler. It would have been interesting to have seen
whether this holds for the same species in November, in so
far as it may be legitimate to compare July of 1900 with
November of 1899. It is possible that the absence of
Euphausians from the surface and upper waters may be ex-
plained by the consideration that most of the ‘ Oceana’
hauls were made during the daytime, and we have evidence
from Dr. Fowler’s collections that certain Euphausians, such
as Huphausia pellucida, rise to the upper strata by night and
sink by day; or, if this be not admitted as fully proven, it is
at least certain that the more active forms are exceedingly
difficult to catch in the upper strata during daylight, espe-
cially in ordinary fine mesh silk tow-nets, such as were used
by the ‘Oceana.’ We are unable to conelude that the
absence of Schizopods from the nets fished from 230 fathoms
or less furnishes material evidence of the distribution of
* Including some hauls made at St. 1, soundings 89 fath., in which no
Schizopods were taken,
1*
4
N.B.—(a) 1. denotes larva.
(6) Larvz not identified occurred in the following hauls :—4 a, 2 Furcilia ; 4b, 1 Furcilia
(c) The numbers given under the heading “ Fathoms” indicate the depth at which the n
Date.
Noy. 19th,
1898.
| Noy. 20th,
1898.
——— eS
Nov. 21st,
1898.
Nov. 22nd,
1898.
|
|
|
|
Station.
Lat. 52° 4' 5" N.
‘Long. 12° 27’ 0" W.
Lat. 52° 27' 6" N.
Long. 15° 40' 0" W.
Eat. 52° 18) 1 N.
Long. 15° 53' 9" W
Lat. 52° 20' 0” N.
Long. 15° 7' 9” W
Net.
Fathoms.
Table showing the Occurrence 9j
R
1 ierom
ee
3.4 10°0 mm.
14:0 mm
Oro many
2. 4&11 mm.
1. 15 mm
2. 6&9 mm
1. 10mm
2, 8 &13 mm
10. 6-15 mm.
4. 3. 7-10mm
1. 18mm
2. 65&85mm
2. 14 mm.
g) By 10 mm
7 Wllo» 7f tenia
7 6. 7-10mm
wale Simm
Skt Simme
8. 6-16 mm.
D) peel emma:
{ll Brim.
|) Le ealSmme
| 3. 8-15 mm.
1mm:
| 8. 8-15 mm
4. 719 tam
1. 7mm.
| 7. 6-14 mm
9 fil.) bimini:
[gee il 1. 5mm
|
—_—
Messrs. E. W. L. Holt and W. M. Tattersall on
Meganyctiphanes
norvegica.
SS
2.
se eeee
23 & 26 mm. |
Thysanoessa
longicaudata,
ceeeee
seeeee
weeeee
teeeee
6. 6-9 mm. |
7. 7-10 mm.
13. 7-10mm.
seen
Schizopoda in the several Hauls.
4k, 1 Calyptopis; 6d, 1 Calyptopis.
Se. (~
g-8 es
SS 88
sd SS
StS =
SSS RS
Ny B
sopeOe 1. 6mm.
peCeee Ph tay (on ¢( proven
ANOcee i
Soates 2 0.
8. 8mm
tenes
2. 6mm
ee ie
oes 41.
neon 1. 7mm.
1. 8mm.
1. 10mm {oy
ee 2 1.
(Pls G75) mom
|i. 8 mm ite
(Sion
eae ee
3. 7-9 mm eeteae
ree ee
8. 5-8 mm. 7 6-8 mm
1. 65 mm
mone UE.
aah j on a Garam
ee Le
1. 6mm
a ii ;
{ 3. 6-8 mm.
are ay
Dis
{4. 5-7 mm.
AWE
ey:
2. 6&7 mm.
abbreviatum.
Stylocheiron
seen
santas
veeeee
the Schizopods collected by the ‘ Oceana.’
CU
was towed, though not necessarily the depth at which the contained organisms were captured.
| Katerythrops
|
|
Oceane.
seenee
saeeee
drepanephora.
Gnathophausia
teens
Eucopia australis.
saceee
ashe alt © Baipetoce 1. 33mm
gee tS Nee
Per Grin ety Asta MN) “axe
navAne 1. 39 mm aoe
Syrian Le Ota hs | ie Vanni: |
Pe eY BE ge Cesaass 1. 12 mm.
saesée. (ll A meeare | ee
a ee ee |
| |
ep) ACR Seder ede | GL contre |
remedervinriy ae Peceeermn lip ue oaGelae |
|
6 Messrs. E. W. L. Holt and W. M. Tattersall on
these forms in the upper strata. In the work of the ‘ Helga’
the only net which has been found really efficient for their
capture at or near the surface, even at night, has an opening
of about 16 square feet (about eight times that of an ordinary
ring tow-net), with a mesh of mosquito-netting or the largest
size of grit-gauze, which permits of hauling at a speed
which would burst fine bolting-silk.
The details of each haul in which Schizopods occurred,
together with the total number and size of the examples
tuken, are set forth in the table (pp. 4-5).
Euphausia pellucida, Dana.
This species occurred in twenty-five out of thirty hauls
in which Schizopods were taken. None of the specimens
reached the full size of the species. The ‘ Oceana’ collections
first demonstrated the existence of EH. pellucida in British
waters. E. bidentata, Sars, is probably the name most
strictly applicable to the North Atlantic form of E. pellucida.
Meganyctiphanes norvegica (M. Sars).
Only two specimens of this rather common form were
taken by the ‘Oceana.’ ‘They occurred in net 2y fishing
from 650 fathoms to the surface. Station 2 was the station
nearest land, and the absence of this species from the hauls
at the more westerly stations suggests that the latter were
beyond the most seaward limit of its distribution at the time.
It is, however, an active form which might well avoid small
tow-nets.
We have found this species common off the west coast of
Ireland.
Thysanoessa longicaudata (Kroyer).
Synon. 7. tenera, G. O. Sars.
This was by far the most abundant Schizopod taken by
the ‘Oceana’ in point of numbers, though in the number of
hauls in which it occurred it is surpassed by both Euphausia
pellucida and Stylocheiron Suhmii. All the specimens
were more or less badly damaged, and out of a total of 150
not one had a perfect leg of any kind left. It occurred in
twenty out of thirty hauls, fishing from 500-1770 fathoms
to the surface.
the Schizopods collected by the * Oceana.’ 7
Thysanoessa gregaria?, G. O. Sars.
Specimens occurred in nine hauls, in nets fishing between
500-1700 fathoms and the surface. We have noted (/oc. cit.)
that the specimens diverge slightly from Sars’s description
of the type. Probably they will prove referable to Hansen’s
T. parva.
Stylocheiron Suhmii, G. O. Sars.
Synon. See Hansen, Buil. Mus. Ocean. Monaco, no. 30 (1905).
S. Suhmii occurred in twenty-five out of thirty hauls,
in nets fishing between 270-1770 fathoms and the surface.
It was one of the commonest Schizopods taken by
Mr. Murray, who collected it in British waters for the first
time. Several larve of this species in all stages of develop-
ment were also captured during the cruise.
Stylocheiron abbreviatum, G. O. Sars.
Synon. S. chelifer, Chun.
A single male example of this species, measuring 20 mm.,
occurred in net 5 A, fishing from 1410 fathoms to the surface.
This is much larger than Chun’s specimens, which measured
only 14mm., but examples of nearly equal size were captured
by Dr. Fowler in the Bay of Biscay.
Katerythrops Oceana, H. & T.
This is one of the new species taken by the ‘ Oceana.’
Descriptions and figures which have already appeared (H. & T.
loc. cit.) arereproduced, and brief diagnoses of the genus and
species are repeated below.
Kateryturops, H. & T.
Characters of the pleopods in the adult male uncertain ;
pleopods of the female unknown. Other characters as in
Meterythrops, 8.1. Smith, except :—
Antennal scale considerably reduced in length in proportion
to peduncles of antenna and antennule, narrow and feeble, its
outer margin naked, entire, terminating in a small spine, sete
few, confined to the apex and distal third (approximately) of
the inner margin.
Telson possibly without the median sete.
8 Messrs. E. W. L. Holt and W. M. Tattersall on
Katerythrops Oceane, H.& T. (PI. IT.)
Form robust. Carapace much wider than the pleon,
almost entirely covering the thoracic segments; anterior
margin obtusely rounded ; cephalic region inflated and poste-
riorly defined by a well-marked cervical sulcus. Pleon with
the last segment almost as long as the two preceding segments
taken together. Eyes small, remote from each other, sub-
pyriform, the proximal part the broader; visual area
restricted to less than the distal half; cornea not so wide as
the last joint of the antennular peduncle; pigment after
preservation in formalin reddish buff. Peduncle of antennule
at least a fifth longer than the last segment of the pleon,
proportionally stout, its last joint about equal to the two
preceding, beset dorsally between the insertions of the
flagella with a bidentate tubercle, of which the denticles are
nearly in the same dorso-ventral plane. Antennal scale very
short, narrow, and somewhat outwardly curved ; outer margin
entire, naked, terminating in a feeble spme; apex produced
considerably beyond the spine, subacute, setz confined to
the apex and to about the distal third of the imner margin ;
length of scale more than four times (about =) the greatest
width, shghtly less than the combined length of the last two
joints of the antennal peduncle, and but little exceeding the
length of the last joint of the antennular peduncle. An-
tennal peduncle long and proportionally stout, combined length
of the last two joints greater than that of the last joint of
the antennular peduncle.
Exopodites of the thoracic limbs very well developed, with
unusually large flagella. Hndopodites of the first four pairs
moderately long and stout; the tarsus in the third and
fourth pairs consisting of three joints, and succeeded by a
well-developed dactylus; setze not more plumose than in
Parerythrops &c. Pleopods of all five pairs biramous in the
male, the inner ramus bifid. Je/son subtriangular, shorter
than the last segment of the pleon by about two sevenths of
the length of the latter, its sides entire and slightly inflected ;
apex narrowly truncate, armed with two pairs of rather
slender spines, of which the inner are considerably the longer
and stouter ; a median pair of setze possibly present. Outer
uropod the longer, its length including basal articulation
slightly greater than the combined length of the fifth and
sixth segments of the pleon. Length of the type specimen
(an immature male) 6 mm., including antennular peduncles
and uropods.,
the Schizopods collected by the ‘ Oceana,’ 9
A single specimen of this species occurred in each of two
hauls—6 g, fishing from 560 fathoms to the surface, and 4h,
fishing from 1470 fathoms to the surface. It is evidently
one of the few known oceanic or strictly pelagic Mysids.
Gnathophausia drepanephora, H. & T.
This is the second of the new forms discovered by
Mr. Murray. Descriptions and figures of this species have
already appeared. The figure is reproduced and a brief
diagnosis is given below. It belongs to a new section of
the genus Gnathophausia, which may be described as
follows :—
Genus GnatuopHausisA, Willemoes-Suhm.
(Trans. Linn. Soc. Lond. ser. 2, Zool. vol. i.)
Section 4*.
Infero-posterior corners of carapace produced into a spine.
Dorsal keel interrupted anteriorly. Supraorbital spine
small, Antennal scale not jointed at apex. First thoracic
legs with distinctly developed exopodites. Epimeral plates
of last segment not united on the ventral face.
Gnathophausia drepanephora, H.& T. (PI. II.)
Form of body slender. Carapace not very large; dorsal
spine about as long as first segment of pleon ; infero-posterior
corners produced into a spine, bluntly serrulate on ventral
edge, nearly reaching fourth segment of pleon ; upper lateral
keel present; dorsal keel unarmed ; cervical sulcus rather
distinct ; rostrum elongate and slender, as long as the cara-
pace without the infero-posterior spines, distinctly denticulate
on all three edges; supraorbital and antennal spines well
defined, but small. Branchiostegal projections of moderate
proportions, but distinctly poimted. Anterior segments of
pleon without dorsal spines ; epimeral plates produced poste-
riorly into pointed lappets. Eyes very narrow, cornea
scarcely at all expanded, pigment (as preserved in formol)
rather pale brown. Outer flagellum of antennule in male
expanded and flattened at the base, which is beset on the
inner side with a brush-like fringe of fine curling setz.
Antennal scale of moderate size, about four times as long as
broad, tapering distally and very obliquely truncate ; inner
angle produced into a sharp point, outer edge with (about)
three denticulations distally. Telson large and massive,
* Jn sequence to Sars’s Sections 1-3.
10 On the Schizopods collected by the ‘ Oceana.’
with the terminal spines crescent-shaped and denticulate
along the upper face; lateral margins armed for the usual
distance with large spines, separated from each other by
intervals occupied by a few smaller spines. Uropods shorter
than telson ; the proximal joint of outer uropod terminating
externally in a spine about one fourth as long as distal
joint. Colour red. Length 39 mm.
The only known specimen was taken in net 4J/, fished at
1770 fathoms and thence to the surface.
Eucopia australis, Dana.
A single specimen was taken in each of four hauls, in nets
working between 500-1710 fathoms and the surface. One
of the examples measured 33 mm. in length, which is not,
however, quite the full size of the species, since it is known
to reach at least 50 mm.
Previous to the ‘Oceana’ cruise the only record of this
species from near our coasts is one by Calman, who records
its capture by the R. I. A. expedition in 1888 off the west
coast of Ireland in 1020 fathoms. We have since recorded
it from soundings of less than 1000 fathoms.
EXPLANATION OF THE PLATES.
Puare I.
Katerythrops Oceane, H. & T.
Fig. 1. Immature male. Dorsal view.
Fig. 2. Immature male. Lateral view.
Fig. 3. Antenual scale with peduncle.
Fig. 4. Endopodite of the leg of the first pair.
Fig. 5. Pleopod of the first pair, ventral view.
Fig. 6. Telson.
Puate II.
Gnathophausia drepanephora, V1. & T.
Fig. 1. Male. Lateral view.
“g. 2. Base of antennular flagellum of male.
Fig. 3. Antennal scale.
[The Plates are reproduced from the ‘Report on the Sea and Inland
Fisheries of Ireland,’ pt. ii. Appendix iy., 1905.)
Note added in Press——Hansen’s Preliminary Report on
the Schizopoda of the ‘ Princess Alice’ appeared while this
paper was in the press. In correcting proofs we have
adopted his view of the synonymy of the species mentioned.
On new Coleoptera from British New Guinea. 11
Il.— Descriptions of new Coleoptera of the Family Cetoniide
from British New Guinea. By Otiver HK. JANSON,
F.E.S.
THE Coleoptera recently brought home by Mr. A. Pratt and
his son from British New Guinea mainly consisted of a very
fine series of Cetoniide, which I have been fortunate in
obtaining. The collection was made on the ascent of the
Owen Stanley range between Ekeikei (1500 feet) and Mafalu
(8000 feet), chiefly in the vicinity of the village of Babooni,
at an elevation of about 3600 feet. Besides the novelties
now described the collection also contained the following
species :—Ischiopsopha gagatina, Hell.; Lomaptera lina,
Gest., Anne, Hell., satanas, Hell., irtdescens, Hell., fulvi-
cornis, Hell.; Mycterophallus viridula, Kz.; and several
others at present undetermined. Prof. Dr. Heller has kindly
verified the determination of several species for me by a
comparison with his types in the Dresden Museum.
MoROKIA, gen. nov.
Gen. Lomaptere affine. Corpus robustum, ovatum, valde convexum.
Clypeus profunde incisus, lateribus elevatis. Thorax lateribus
regulariter rotundatis, marginatis, Scutellum lobo postico
thoracis haud obtectum, apice acuminatum. LElytra profunde
punctato-sulcata. Pygidium transversum, depressum. Meso-
sterni processus valde productus, oblique directus, apice compressus,
obtusus. Pedes breves, tibiis anticis in utroque sexu bidentatis.
Morokia Meekt, sp. n.
Nigra, nitida, capite punctato, prothorace lateribus crebre punctato ;
elytris in dorso profunde sex-sulcatis, sulcis catenato-punctatis,
apice conjunctim rotundatis.
Long. 35-38 mm. ; lat. 19-21 mm.
36. Abdomine longitudinaliter sulcato, pygidio subter vix impresso.
. Pygidio subter late profunde impresso.
British New Guinea, Morok (A. S. Anthony); Upper
Aroa River (A. S. Meek); Babooni (A. #. Pratt).
Of an ovate form, the body convex and strongly arched
above, the thorax and head being much depressed, deep shiny
black. Head not very closely punctured, slightly convex in
the centre, side margins of the clypeus raised. Thorax
12 Mr. O. E. Janson on new Coleoptera
broadly rounded and margined at the sides, basal angles
rounded and leaving the epimera largely exposed ; basal lobe
Jarge, broadly rounded and slightly emarginate at its apex,
sparsely punctured on the disk and much more coarsely so
towards the sides ; besides this punctuation the whole surface
when examined under a lens is seen to be very densely and
minutely punctured. Elytra narrowed behind and conjointly
rounded at the apex, the apical sutural angles in some
specimens a little rounded, the disk of each with six deep
furrows bearing chain-like rows of coarse semicircular punc-
tures, the interstices very convex and smooth, the sides with
three rows of very coarse punctures merging into a coarse
transverse strigosity towards the apex. Pygidium transverse,
depressed, and coarsely strigose, the underside slightly
impressed in the centre in the male, in the female rather
more obtuse at the apex and with a large deep central
impression on the underside. Beneath coarsely strigose and
punctured at the sides; mesosternal process very large,
obliquely divergent, slightly curved and almost vertical
towards the apex, which is a little compressed and obtuse ;
in some specimens there is a more or less distinct tooth or
keel on the inner edge near the apex. Legs short, femora
strigose and lightly fringed with short black hairs; anterior
tibize with two lateral teeth, which are more or less obsolete
in the male, but large and very acute in the female.
I received a specimen of this remarkable and distinct
insect some years back from Mr. Anthony, but considered it
inadvisable to describe it as new upon a single example.
Messrs. Meek and Pratt have since met with it sparingly.
‘The strongly arched body, ovate form, deeply sulcate elytra,
short legs, &e. will readily distinguish it from Lomaptera.
In one female specimen the pygidium is only slightly
impressed, as in the male.
Lomaptera Pratti, sp. n.
Viridis, nitida, capite subtus, coxis femoribusque anterioribus
rufescentibus et rufo pilosis; prothorace lateribus punctatis ;
elytris seriatim punctatis et transverse strigosis, area basali fere
levi, in apice productis.
Long. 33-36 mm.
¢. Pygidio subter medio subplano, abdomine suleato, tibiis anticis
muticis.
@. Pygidio subter profunde late impresso, tibiis anticis dente parvo
armatis.
from British New Guinea. 13
British New Guinea, Babooni (A. 2. Pratt); Mount
Victoria (A. S. Anthony).
Dark green, in some specimens more or less tinged with
olivaceous ; underside of the head with the palpi and an-
tenn, anterior coxe and femora, intermediate coxe, and in
some specimens the hinder part of the prosternum rufous,
tinged with green. Head finely punctured, and more sparsely
so at the base ; clypeus long, very deeply cleft, and without
any raised margin. Thorax coarsely punctured at the sides,
more finely and very sparsely punctured on the disk, strongly
widened at the base, the basal angles being produced, de-
pressed, and obtuse, the side margins rather strongly raised
on the basal three fourths, the basal lobe long, narrow, and
rounded at its apex. LHlytra with two slightly indicated
discal costze and some rows of coarse punctures, which become
obsolete towards the base and merge into the coarse trans-
verse strigosity with which the apical half is closely covered ;
sides a little dilated at the shoulders, where the marginal
line is deeply impressed, the apex rather strongly produced
towards the suture and slightly emarginate at the sutural
angle. Pygidium transverse, subconical, coarsely strigose,
the underside a little flattened towards the apex in the male,
and in the female with a large central depression, having a
strongly raised semicircular apical border. Underside coarsely
strigose and sparsely pubescent at the sides; mentum, pro-
sternum, and anterior coxee and femora with long rufous hairs ;
mesosternal process long, obliquely divergent, and sharply
keeled on the underside near the apex ; abdomen obliquely
strigose, the fifth segment with very coarse setiferous punc-
tures, and in the male with a strong central furrow ; anterior
tibia simple in the male, and with a slight subapical tooth in
the female.
This fine species is very distinct from any with which I
am acquainted, but from the description of L. Beccari, Gest.,
it would appear to be allied to that species. The colour is
very similar to that of Z. wirens and Latreille’, but rather
more metallic.
Lomaptera lutea, sp. n.
Lutea vel testacea, nitida, prothorace elytrisque opalino-irides-
centibus ; capite postice, prothorace macula magna M-formi,
metasterno in parte mediana, femoribus apice, tibiis anticis et
intermediis, posticis apice tarsisque omnibus nigris; abdomine
14 Mr. O. KE. Janson on new Coleoptera
rufo-testaceo vel piceo; elytris transversim aciculatis, area basali
fere leevi.
Long. 30-32 mm.
3g. Abdomine sulcato, pygidio depressiusculo, subconico, tibiis
anticis simplicibus.
@. Pygidio late producto, apice truncato, tibiis anticis ante apicem
subdentatis.
British New Guinea, Upper Aroa River (A. S. Meek) ;
Babooni (A. £. Pratt).
Very like L. Anne, Hell., but smaller and of a narrower
form, the colour and markings similar, but with a very distinct
opalescent iridescence on the thorax and elytra; the head is
very much more sparsely and finely punctured, especially
towards the apex of the clypeus; the thorax is also less
punctured and narrower at the base (the angles being less
divergent) ; the elytra are a little emarginate at the apex,
with the sutural angle slightly produced; the pygidium of
the male is similar to that of Anne, but in the female it is
less produced than in that species, much broader and almost
truncate at the apex; it is also more coarsely aciculated on
the upperside and more deeply impressed on each side beneath.
The male forceps of the two species are also very different.
The extent of the black markings varies considerably in
the series before me; the margin of the clypeus and the
basal portions of the sides of the thorax and elytra are
generally narrowly edged with black, and in some specimens
this narrow black edging also extends to the apical and basal
margins of the thorax and the basal and sutural margins of
the elytra; there is often a more or less distinct black spot
on each side of the thorax and on the shoulders, and some-
times two small linear marks on the forehead; in several
specimens there are rows of dark punctiform spots on the
elytra. On the underside the black coloration frequently
extends to the prosternum, coxe, abdomen, and other parts,
whilst the tibiee in some are almost entirely black and in
others only partially so.
Mr, Pratt obtained a rather large series of both this
species and Anna, the latter also showing considerable varia-
tion in the extent of the black markings.
Lomaptera semicastanea, Kz.
Besides a considerable series of the New Guinea form of
this species named var. pseudorufa by Dr. Heller, there are
from British New Guinea. 15
also in Mr. Pratt’s collection several others differing so widely
in coloration as to have the appearance of being distinct
species ; but upon careful examination I am unable to find
any other characters to separate them, and must therefore
treat them as varieties; and although I am generally opposed
to the giving of distinctive names to mere varieties, I do so
in this instance, as they are well-marked forms of which no
intermediate variations connecting them occur in the collection.
Var. signata, nov.
Var. pseudorufe simillima, sed clypeo utrinque rufo-notato, pro-
thorace macula magna M-formi nigra. d.
Var. rufipennis, nov.
Nigra, elytris rufis. ¢ Q.
Var. plagiata, nov.
Nigra, elytris macula magna basali rufa. ¢ Q.
Var. unicolor, nov.
Tota aterrima. <9.
Lomaptera rubens, sp. 0.
Nigra, nitida ; prothorace (lateribus antice exceptis) elytrisque rufo-
castaneis, subopalino-iridescentibus ; corpore subtus femoribusque
parce fulvo-pilosis; elytris transverse aciculatis, area ~ basali
remote punctata ; tibiis anticis simplicibus.
Long. 380-32 mm.
3. Pygidio subconico, depressiusculo, strigoso, abdomine late
sulcato.
9. Pygidio obtuse conico producto, subter impresso, lavi, in medio
leviter carinato.
Babooni, British New Guinea (A. ZH. Pratt).
Allied to Z. rufa, Kz.; very similar in coloration, but the
thorax and elytra of a darker or more castaneous red; it is
also larger and of a broader form; the clypeus is more
dilated in front, making the apical lobes less acute ; the thorax
is proportionately broader at the base, rather sparsely punc-
tured, and not strigose at the sides; the elytra are more
strongly sinuate behind the shoulders and not so closely
aciculate ; the pygidium in the male is similar in form, but
16 Mr. O. E. Janson on new Coleoptera
more coarsely and less closely strigose; in the female it is
more broadly rounded at the apex than in this sex of rufa,
and is impressed and smooth on the underside, and with a
slight but distinct narrow, longitudinal, central carina,
becoming obsolete towards the apex : the mesosternal process
is longer and more incurved, and the anterior tibie# of both
sexes are without any indication of lateral teeth.
From var. pseudorufa, Hell., which it is also like in colora-
tion, it may be distinguished by the incurved mesosternal
process, and the different structure of the pygidium and
scarcely emarginate sixth abdominal segment in the female;
the male forceps are also very distinct.
Lomaptera pulchella, sp. n.
Rufo-castanea, subviridescens, nitida; capite basibus et prothorace
disco viridi-sneis ; elytris flavo-virescentibus, sutura late viridi-
marginatis, transversim aciculatis, lateribus subinflexis; pygidio
subconico. o QQ.
Long. 19-21 mm.
Babooni, British New Guinea (A. E. Pratt).
Front of head and the clypeus, sides and front of thorax,
pygidium, underside, antenne, and legs brownish red, with a
greenish reflection ; base of head, the disk and a small spot
on each side of the thorax, and the centre of the body
beneath brassy green; elytra pale yellow, with a greenish
reflection; a broad sutural band, commencing at the base and
gradually narrowed to the apex, dark green; asmall humeral
spot and the anterior half of the extreme outer margin black.
Head very closely punctured on the clypeus, more sparsely
so between the eyes, and smooth at the base. ‘Thorax very
sparsely punctured on the disk, more coarsely and closely
punctured towards the sides, and rugose near the margin ;
basal lobe large, almost impunctate, and slightly notched at
its apex; lateral margins a little reflexed; the basal angles
slightly produced and depressed. Elytra sparsely punc-
tured at the base and in the scutellary region, the rest of
their surface finely transversely aciculated, subcostate at the
sides ; the apex a little rounded, with the sutural angles very
slightly produced and acute. Pygidium transverse, sub-
conical, aciculated both above and below. Underside finely
strigose and with rather sparse grey pubescence ; mesosternal
process long, almost cylindrical, a little curved and obtuse at
the apex ; apical margin of the sixth abdominal segment a
from British New Guinea. 17
little sinuous. Legs strigose and punctured; femora fringed
with long grey pubescence. In the female the pygidium is
more produced and impressed on each side underneath, the
fifth abdominal segment is more strongly punctured and
pubescent, and the anterior tibia have a distinct subapical
tooth which is not present in the male.
A small series of this pretty species was contained in
Mr. Pratt’s collection ; in coloration it would seem to most
nearly resemble L. macrophylla, Gest., a species only known
to me by description ; but the antennal club is of normal size
in both sexes. In some specimens the elytra present rows of
small dark punctiform spots and a greenish mark on the
apical callosities. F
Lomaptera cupriceps, sp. n.
Ainea vel viridi-metallica, iridescens; clypeo igneo-cupreo, apice
rufo-piceo ; elytris omnino transverse aciculatis ; pygidio conico,
supra non carinato; pedibus rufo-piceis, femoribus intermediis
posticisque cupreis. o Q.
Long. 20-21 mm.
Upper Aroa River, British New Guinea (A. S. Meek).
Very like ZL. pygmea, Kz., but with the clypeus more
sparsely punctured, bright coppery red, and piceous towards
the apex ; the thorax more broadly rounded at the sides and
more punctured on the disk ; the elytra more finely aciculated
and with the apical sutural angles in the male acute and a
little more produced, and the mesosternal process rather
smaller and less curved. The pygidium is nearly smooth and
without any longitudinal carina on the upper side; in the
female it is slightly laterally compressed and more pointed at
the apex than in the male. The legs are reddish or piceous,
with the four posterior femora coppery.
In L. pygmea, of which I have a series from Goldie
River, the female has a strong saw-tooth-shaped keel on the
upper side of the pygidium.
Trichaulaw sericea, sp. n.
Nigra, nitida, prothorace sericeo-subopaca ; elytris trisulcatis, sulcis
marginibusque flavo-pilosis. ¢ Q.
Long. 33-35 mm,
Babooni, British New Guinea (A. EZ. Pratt).
Very similar to 7’. Macleayt, Kz., but of a narrower form,
Ann. d& Mag. N. Hist. Ser. 7. Vol. xvi. 2
18 Mr. W. J. Holland on a new
the head less shiny, the clypeus narrower and more parallel-
sided and more closely and shallowly punctured ; the thorax
silky and subopaque and more sparsely punctured, the
basal lobe narrower and more deeply emarginate; the
scutellum distinctly narrower and more acutely produced at
its apex; the hairs in the sulci and at the margins of the
elytra longer and of a yellow colour; the pygidium only
sparsely pilose at the base and apex; the underside is also
more sparsely pubescent, and the centre of the abdomen is
impunctate and entirely without hairs.
This genus has bitherto been known only from Australia,
and it is interesting to find a species so closely allied from
‘the mountains of New Guinea.
III.—A new Noctuid from Sierra Leone.
By W. J. HOLLAND.
THROUGH the kindness of Mr. William Schaus, the Carnegie
Museum in Pittsburgh has received a number of interesting
African moths. One of these, representing a new genus and
species, has been singled out because of its apparent rarity
and rare beauty to bear as its specific designation the name of
the donor of the collection.
I must thank Sir George F. Hampson for saving me the
labour of preparing the following diagnoses of the genus and
species. With that courtesy which marks all his intercourse
with scientific men visiting the British Museum, he, knowing
that my time was taken up with affairs of greater magnitude,
such as the Diplodocus, kindly wrote the description both
of the genus and the species and handed them to me to be
employed in this paper.
NOCTUES.
()UADRIFINZ.
AUCHENISA, gen. nov.
The genus is apparently near Trisula, and is characterized
as follows :—
Proboscis fully developed; palpi obliquely upturned,
rather short, thickly clothed with hair; frons smooth ; eyes
smooth ; antenne of the female bipectinate, with rather long
Noctuid from Sierra Leone. 19
branches, the apical third simple, the basal joint with a large
tuft of hair; head and thorax clothed with hair overlying the
scales; the prothorax and metathorax with crests; tibic
fringed with hair; abdomen with large spreading dorsal crests
of hair on the first three segments. Fore wing with veins 3
and 5 from close to the angle of the cell; 6 from the upper
angle; 9 from 10 anastomosing with 8 to form the areole ;
11 from cell. Hind wing with veins 3-4 from angle of the
cell; 5 from above angle ; 6-7 from angle; 8 anastomosing
with the cell near the base only.
Auchenisa Schaus?, sp. n.
?. Head and thorax pale olive-green; palpi blackish at
the sides ; antennz with the shaft whitish towards the base ;
the branches black, the tufts on the basal joint crimson ;
tegule tipped with crimson; patagia with black spots at
base; prothorax with the crests black; metathorax with
black spot; tibiee with some black hair; tarsi blackish ;
abdomen whitish, dorsally tinged with olive-brown and with
some fuscous in the dorsal crests and on the anal segment ;
there are two lateral series of small black spots. Fore wing
_olive-green, irrorated with white; some white at the base
and two black spots; a subbasal black bar extends from the
subcostal nervure to the submedian fold. There is a broad
antemedial black band, irrorated with white, and with some
green below the cell and above the inner margin. ‘The edges
of this band are irregularly defined by waved white lines,
the band expanding at the imner margin, and with a black
bar beyond the outer white line; reniform green, irrorated
with white, and with the black annulus defined by white,
except above, upright oblong in shape, and with some diffused
Q*
20 Mr. P. Cameron on a new
°
Llack beyond its lower extremity; postmedial line strong,
black, defined by white on the inner side, obliquely waved to
vein 4, then incurved, dentate at veins 3 and 2 and strongly
recurved to the inner margin, at costa expanding into a patch
with some green upon it; there is an ill-defined, irregularly
waved, white subterminal line, with somewhat dentate black
spots beyond it, above and below veins 4 and 2; a terminal
series of black spots defined by white, increasing in size
towards tornus. Hind wing white, the terminal area suffused
with fuscous, narrowing to a point near tornus; two waved
greenish and black subterminal lines from vein 2 to above
tornus; a terminal series of black spots slightly defined by
white. Underside of fore wing fuscous brown, the inner area
white ; both wings with a round black discoidal spot, which
on the fore wing is defined by whitish.
Iixpanse 90 min,
Hab. Sierra Leone (Clements). Type in the Carnegie
Museum, Pittsburgh, U.S.A.
1V.—A new Genus and Species of Cynipide from South
Africa, representing a new Subfamily. By P. Cameron.
PYcNOSTIGMUS, gen. nov.
9. Fore wings with a subcostal nervure ending in a thick,
pyriform, large, horny stigma, more than twice longer than
thick counting from the costa to the hinder part of the wing,
and slightly narrowed aud rounded behind; shortly beyond
the middle an oblique transverse nervure runs from shortly
below the middle to the subcostal nervure. Hind wings with
a subcostal nervure extending from the base to shortly
beyond the middle; its apex is distinctly thickened. An-
tenne 18-jointed, not much thickened towards the apex.
Second abdominal segment very large, the apical ones short,
the fourth shorter than the others. Ovipositor straight,
slightly projecting. Malar space large, longer than the eyes,
which are placed on the top, are apparently bare and parallel.
Clypeus not separated from the face in the middle, its sides
above bordered by a large deep fovea; the apex of the clypeus
(or it may be the labrum) is narrowed, hollowed, transverse,
with the sides rounded. Parapsidal furrows complete, deep.
Scutellum large, rugose, gradually roundly narrowed towards
the apex, where there are two fovee or depressions ; at its
base are two large foveze; the mesonotum is separated from
Genus and Species of Cynipide. ail
it by a transverse furrow. Head and thorax shining, sparsely
punctured and haired.
The legs are moderately long and stout; the hind cox:
are about three times longer than wide; the fore calcari:
curved, slender, simple. The hypopygium is large, but does
not project much; above it is a short stumpy projection—
the apex, apparently, of the penultimate segment. The
abdomen is sessile, the basal segment being very short and
smovth.
The species on which Pycnostigmusine is founded
cannot well be placed in any of the subfam‘lies hitherto
recognized. The alar structure is very peculiar; there is
neither radius nor cubitus, the place of these being taken by
the large horny pyriform stigma, or stigmal branch (if that
is a more appropriate term). This character, with the larger
number of antennal joints, should readily separate it from
the Figitine and Euceeline, with which it is otherwise most
nearly allied. There is no known Hymenopteron with such
a peculiar horny structure on the wings.
Pycnostigmus rostratus, sp. n.
Black, shining, the knees testaceous; the head, antenne,
and legs thickly covered with white pubescence; wings
hyaline, the nervures testaceous, the stigma darker, more
piceous in tint. Face with scattered punctures in the centre,
the sides aciculated; clypeus smooth and shining. — Pro-
notum finely rugose, the apex with pitted punctures ; the
mesonotum with scattered shallow punctures. Scutellum
coarsely, irregularly, rugosely punctured. Metanotum with
two stout keels in the centre, which is irregularly transversely
striated; the apical half on either side of these stoutly
obliquely striated, almost reticulated. Propleure above
coarsely rugosely punctured; the lower part, especially at
the base, striated. Base and lower part of mesopleurze
irregularly punctured-striated, the rest smooth and shining.
Metapleure coarsely aciculated and irregularly striated.
Abdomen smooth and shining. Wings hyaline, iridesceut ;
the apex ciliated. 9.
Length 3 mm.
Cape Town, September (South African Museum).
The antenne are as long as the body, are not much
thickened towards the apex; the pedicle is twice longer
than broad ; the third and fourth joints are about equal i in
length ; the last is conical and twice the length of the
penultimate. Cheeks margiued. The eyes appear to be
slightly hairy ; they are probably rubbed.
22 Mr. W. L. Distant on Homoptera.
V.—Rhynchotal Notes—XXXUI. By W. L. Disrant.
Fam. Cicadide (continued from vol. xv. p. 486).
Subfam. 7rercrvryz (continued).
Division TIBICINARIA.
In this division the abdomen is more or less cylindrical and
attenuated posteriorly; anal appendages usually very pro-
longed and prominent, their lower plate, as a rule, more
longly porrect than their upper one; pronotum generally
more or less distinctly narrowed anteriorly * ; tegmina some-
what tale-like and obscurely wrinkled, their greatest breadth
always more than one third of their length, both tegmina and
wings always prominently sanguineous or reddish ochraceous
at base.
The genera in this division are of a very homogeneous
character and possess a common facies.
Synopsis of Genera.
A. Pronotum about as long as head.
a. Head (including eyes) about as broad as base of
mesonotum,
6. Head (including eyes) as broad or broader than
length of mesonotum (including cruciform eleva-
tion).
ce. Front of head shorter than vertex ............ Tibicina.
aa. Head (including eyes) considerably narrower than
base of mesonotum.
bb. Head (including eyes) equal or almost equal in
breadth to length of mesonotum (including cruci-
form elevation).
ce. Front of head shorter than vertex, its apex more
or less emarginate.
d. Basal cell of tegmina about or almost twice as
oneias broads: .ag. Scie hi ate nae ee Okanagana.
ece. Front of head almost as long as vertex, its apex
not emarginate.
dd. Basal cell of tegmina but little longer than
BEOR YS. Sie ss os oie ah oder ey he ee Ahomana.
B. Pronotum much longer than broad.................. Paharia.
Genus TIBICINA.
Tilicina, Amyot, Ann. Soc. Ent. Fr. 1847, pp. 154 & 350.
Type, 7. hematodes, Scop. (Cicada).
* Except in Tibicina septemdecim, a species which possesses several
aberrant characters.
Mr. W. L. Distant on Homoptera. 23
Tibictna chinensis, sp. n
2. Body and legs black, somewhat thickly greyishly
pilose; lateral margins of face, longitudinal fasciee to femora,
apical areas of tibiz, margins of ‘acetabulze, apices of cox,
abdominal segmental margins beneath, and the same confined
to lateral areas above, testaceous; a central, basal, indented
spot to head, two central longitudinal spots to pronotum, two
discal spots to mesonotum, and the cruciform elevation (ex-
cluding centre) dull ochraceous; tegmina and wings hyaline,
their extreme bases testaceous, venation fuscous, tegmina
with the costal membrane testaceous, the claval area san-
guineous ; wings with the claval area and margins of anal
area fuscous; head about as broad as base of mesonotum ;
lateral pronotal margins distinctly concavely sinuate ; rostrum
reaching the intermediate coxe.
Long., excl. teom., 9 25 mm.; exp. tegm. 62 mm.
Hab. N. China: ‘Ta-chien-lu (Brit. Mus.).
This description is based on two female specimens ; in one
the transverse veins at the bases of second and third apical
areas to tegmina are slightly infuscated, in the second
specimen they are not.
OKANAGANA, gen. nov.
Head (including eyes) considerably narrower than base of
mesonotum and almost equal to its length (including cruci-
form elevation) ; front shorter than vertex, its apex more or
less emarginate, vertex centrally sulcate ; pronotum about as
long as head, its anterior angles in a line with eyes, its poste-
rior angles dilated ; abdomen in male longer, in female about
as long as space between apex of head and base of cruciform
elevation ; tympana completely exposed, tympanal coverings
entirely absent ; ; face more or less centrally sulcate ; rostrum
reaching the intermediate coxee ; opercula small, transverse ;
abdomen beneath with the lateral margins broadly recurved ;
tegmina and wings hyaline; tegmina with the basal cell,
about or almost twice as long as broad, apical areas eight ;
wings with six apical areas.
Type, O. rimosa, Say (Cicada).
AHOMANA, gen. nov.
Head (including eyes) considerably narrower than base of
mesonotum and about equalling its length (including cruciform
elevation), front of head slightly shorter than vertex, its apex
not emarginate ; pronotum about as long as head, its lateral
24 Mr. W. L. Distant on Homoptera.
margins coneavely sinuate ; abdomen in male about as long
as space between apex of head and base of cruciform elevation ;
tympana entirely exposed, tympanal coverings completely
absent ; opercula in male small, broad, transverse ; tegmina
and wings hyaline or semihyaline ; tegmina with the “basal
cell broad, only a little longer than broad, ap‘cal areas eight ;
wings Ww ith six apical areas.
Type, A. neotropicalis, Dist.
Ahomana neotropicalis, sp. n.
Body above black, somewhat thickly greyishly pilose ;
head with a central fascia to front, pronotum with the ante-
rior and posterior margins (narrowly) and the lateral margins
(broadly), posterior margins of meso- and metasterna, ‘and
cruciform elevation (excluding anterior angles), ochraceous ;
body beneath piceous, thickly pilose ; a central longitudinal
fascia to face, rostrum, coxee, and legs ochraceous ; anterior
and intermediate cox pale castaneous, their apices ochra-
ceous ; anterior tibie and tarsi, apices of intermediate tibiz
and the tarsi, piceous ; tegmina and wings semilhyaline, tale-
like, the venation ochraceous ; tegmina with the costal
membrane ochraceous s, the venation on apical area fuscous.
Long., excltegm., ¢ 19, 2 22 mm:; exp. teom aga,
52-55 mm.
flab. Peru: Callao (Brit. Mus.) ; Paraguay (Brit. Mus.).
Ahomana chilensis, sp. n.
3d. Body black; sternum, opercula, and legs ochraceous ;
$
abdomen finely greyishly ‘vilose, sternum longly thickly
pilose; apical areas of intermediate and posterior femora,
posterior tibia (excluding bases and apices), and apical area
of rostrum castaneous, intermediate tibie piceous, their bases
ochraceous (anterior legs mutilated in specimen described) ;
lateral margins of face “ochraceous ; ; head with a short longi-
tudinal, basal, ochraceous fascia; pronotum with the lateral
areas castaneous, its margins, a central longitudinal fascia,
and two central basal spots ochraceous ; mesonotum with two
curved discal linear spots, a spot on each lateral area, and the
cruciform elevation with two central spots and its anterior
angles ochraceous ; tegmina and wings semihyaline, venation
ochraceous or fuscous, extreme bases of both ochraceous ;
tegmina with costal membrane and claval area ochraceous,
the transverse veins at bases of second and third apical areas
slightly infuscate.
Long., excl. tegm., g¢ 22 mm.; exp. tegm. 68 mm.
Hub. Chili (Reed, Brit. Mus.).
Mr. W. L. Distant on Homoptera. 25
PAHARIA, gen. nov.
Head moderately narrow, its breadth (including eyes)
narrower than base of mesonotnm and about equal to its length
(including cruciform elevation), front prominent, as long as
vertex ; pronotum much longer than head, convex, laterally
depressed, obliquely narrowed anteriorly, its posterior angles
strongly lobately produced ; mesonotum strongly deflected on
each side; abdomen (¢) about as long as space between
apex of head and base of cruciform elevation ; tympana com-
pletely exposed, tympanal coverings entirely absent ; abdomen
beneath with the lateral margins strongly recurved and very
prominent; anterior femora robust and strongly spined
beneath ; posterior tibize spined on each side for about half
their length; rostrum reaching the intermediate coxe ;
tegmina and wings tale-like in type, the first broad, with the
apical areas somewhat short and eight in number, the basal
cell about twice as long as broad, narrower at apex than at
base; wings with six apical areas.
Type, P. lacteipennis, Walk. (Cephaloxys).
My knowledge of this genus extends to three species, all
represented by female specimens only.
Division TAPHURARLIA.
Eyes projecting beyond the anterior angles of the pro-
notum ; pronotum subquadrate, not distinctly narrowed
anteriorly, as (with few exceptions) in the Vibicinaria; abdo-
men about as long as space between apex of head and base of
cruciform elevation * ; a more or less distinct posterior meta-
sternal process visible in males between or at the base of the
opercula.
Synopsis of Genera.
A. Wings with six apical areas,
a. Head (including eyes) about as broad as base of
mesonotum,
b. Head with front subprominent, in breadth about
equal to length of lateral margins of vertex ;
space between eyes very much narrower than
anterior margin of pronotum.
ec. Tegmina less than three times longer than
broad,
d. Tegmina with lower veins of radial and
fourth ulnar areas at base widely separated.
* Except in Malagasia and Kanakia,
26 Mr. W. L. Distant on Homoptera.
e. Tegmina with transverse vein at base of
second apical area oblique.
f. Head with vertex longer than front;
CVCSMOMMALN Ane. cone ee een
ee. Teginina with transverse vein | at base of
second apical area vertical.
Sf. ead with front about as long as vertex ;
eyes oblique, their inner margins
ampliated and laminately produced
IWardly?, & ccd Riper so tetent eeeae
dd. Tegmina with lower veins of radial and
fourth ulnar areas at base coubenpae but
not touching. ..... AN eBaNNNie ALIA iy see
ec. Tegmina long “and slender, more than three
times longer ah broad . ahele Sten eenrarete
bb. Head with front prominent, - in breadth con-
siderably narrower than lateral margins of
vertex ; space between eyes only slightly nar-
rower than anterior margin of pronotum . Bene yele
aa. Head (including eyes) broader than base of meso-
notum,
g. Tegmina with transverse vein at base of second
apical area oblique.
h. tront of head about as long as vertex, their
margins continuous.
2. Metasternum in male posteriorly, narrowly,
subtriangularly produced between the
opercula, and occupying basal space be-
tween them.
j. Greatest breadth of tegmina about one
thirdaphem length keg st)a.nii ae acter
hh. Front of head slightly longer ‘than “vertex,
their margins not continuous, but almost at
right angles to each other.
wi. Metasternum in male posteriorly, broadly,
longly, convexly produced between the
opercula, and occupying space between
them.
Jj. Greatest breadth of tegmina more than
one third their length ............
. Metasternum in male not produced between
PALO PENCUlas. Wawa. tahiiceke ee Moke rer-Liay
hhh. Front of head considerably shorter than
vertex, their margins continuous.
wi. Metasternum in male not posteriorly pro-
duced nor occupying space between oper-
cula, which are small, wide apart, and not
reaching basal abdominal segment.
k. Abdomen beneath plobose® ach. memes
kk. Abdomen beneath with the lateral mar-
ginal areas strongly recurved..........
gg. Tegmina with transverse vein at base of second
apical area nearly vertical.
7. Tegmina with the first and second ulnar areas
about as long as, or longer than, remaining
ulnar areas.
m. Head shorter than space betwcen eyes.
Abricta.
Dorachosa,
UVeana.
Parnkalla.
Burbunga.
Trismarcha,
Monomatapa,
Selymbria.
Malagasia.
Kanakia.
Mr. W. L. Distant on Homoptera. 27
n. Front of head a little shorter than vertex,
their margins continuous, the front not
produced, and about, or nearly, twice
broader than LKeNa¥e8 Perens cae h ORS cae Abroma.
nn, Front of head about as long or a little
shorter than vertex, their margins dis-
continuous and almost at right angles to
each other; the front prominent, not
nearly twice as broad as long ........ Lemuriana.
mm. Head about as long asspace between eyes. Taphura.
Wl. First and second ulnar areas to tegmina
shorter than remaining ulnar areas ........ Liyymolpa.
B. Wings with five apical areas.
o. Front of head about twice as broad as either lateral
margin of vertex.
p. Vertex and front of head strongly longitudinally
sulcate; basal margin of face strongly trans-
WETSCLVMITICFASSAEC 5 tte ntelasa so Shae sus legieper sia ore Auta.
oo. Front of head about as wide or little wider than
length of either lateral margin of vertex.
pp. Vertex and front of head not strongly longi-
tudinally suleate; basal margin of face not
strongly merassate..ts<(0% odes aes RRO PAE Panka.
C. Wings with four apical areas ..... POA CERO OS CIO Nelcynda.
Genus ABRICTA.
Tibicen, subgen. Abricta, Stal, Hem. Afr. iv. p. 26 (1866),
Type, A. brunnea, Fabr. (Tettigonia).
Abricta castanea, sp. n.
Body brownish ochraceous, with castancous markings ;
head with the margins of front and the vertical area more or
less castaneous ; pronotum with the anterior and posterior
margins and two central lines widely angulated at base casta-
neous (in some specimens the space included in these lines
is wholly castaneous) ; mesonotum with two short central
obconical and two long sublateral spots, and a large spot in
front of cruciform elevation, castaneous ; posterior margins of
eyes black ; face and space ‘between face and eyes pale. casta-
neous ; sternum and opercula in male ochraceous; tegmina
and wings hyaline, the venation ochraceous or ‘brownish
ochraceous; tegmina with the costal membrane and claval
area ochraceous ; wings with extreme base and base of anal
area ochraceous.
Long., excl. tegm., ¢ 14-15 mm.; exp. tegm. 43 mm.
Hab. North Australia (J. &. Hilsey, Brit. Mus.).
Allied to A. Wellst, Dist.; tegwina unspotted ; opercula
in male longer and much less transverse, Ke,
28 Mr. W. L. Distant on Homoptera.
Abricta Burgesst, sp. n.
Body and legs ochraceous; anal abdominal segment dark
castaneous, its posterior margin ochraceous; head with a
small spot on each lateral margin of front, inner margins of
eyes, the area of the ocelli, four small anterior spots to pro-
notum (two central and two lateral), and the margins of two
central obconical spots to mesonotum, piceous or black; the
inner areas of the black-margined spots, a large oblong spot
on each lateral area, and a triangular basal spot to mesonotum,
brownish ochraceous ; body above and beneath sparingly
greyishly pilose ; two central fascie to face and a central
line to clypeus castaneous ; apices of tibia, tarsi, and rostrum
more or less piceous; tegmina and wings hyaline, the venation
ochraceous; tegmina with the costal membrane ochraceous
and the transverse veins at the bases of the second and third
apical areas slightly infuscate; opercula in male short,
transverse.
Long., exc]. tegm., g 18-19 mm. ; exp. tegm. 60 mm.
Hab. Queensland ; Ripple Creek (Mrs. Jane Burgess,
Brit. Mus.).
In general appearance and abdominal apical markings
resembling Ueana lfuana, Moutr., from which the widely
separated Jower veins of radial and fourth ulnar areas at once
distinguish it.
Genus DORACHOSA.
Dorachosa, Dist. Ann. & Mag. Nat. Hist. (6) x. p. 63 (1892).
Type, D. explicata, Dist.
UEANA, gen. nov.
Head (including eyes) about as wide as base of mesonotum,
the front subprominent, in breadth about equal to the length
of lateral margins of vertex, space between eyes very much
narrower on anterior margin of pronotum, lateral margins
of front and vertex continuous ; pronotum about as long as
head, its lateral posterior angles ampliate ; length of meso-
notum (including cruciform elevation) less than breadth of
head (including eyes) ; abdomen about as long as space
between apex of head and base of cruciform elevation ;
tympana entirely exposed; opercula in male small, trans-
verse; metasternum shortly posteriorly produced in fie male
between the opercula 5 ; anterior femora strongly spined
beneath ; tegmina and wings hyaline; tegmina with eight
apical areas and the bases ee the lower veins of radial anal
hal
Mr. W. L. Distant on Homoptera. 29
fourth ulnar areas contiguous but not touching ; wings with
six apical areas.
Type, U. lifuana, Montr. (Cicada).
PARNKALLA, gen, nov.
Head as long as pronotum, including eyes about as wide
as base of mesonotum, vertex longer than front; pronotum
with the lateral margins nearly straight, the posterior lateral
angles somewhat angularly dilated ; mesonotum longer than
pronotum ; abdomen short, about as long as space between
apex of head and base of cruciform elevation; tympana
eutirely exposed; face globose, transverse striations and
longitudinal sulcation sometimes almost entirely absent or
obsolete ; anterior femora strongly spined beneath; opercula
in male short and transverse ; tegmina long and slender, more
than three times longer than broad, apical areas eight, basal
cell at least twice as long as broad; wings with six apical
areas,
Type, P. Milleri, Dist. (Tibicen).
BuRBUNGA, gen. nov.
Head (including eyes) about as wide or very slightly
wider than base of mesonotum, front of head conically promi-
nent, its breadth at base very considerably narrower than
the length of lateral margins of vertex, in length nearly as
long or a little shorter than vertex, front and vertex together
about as long as pronotum; pronotum with the lateral
margins oblique, the posterior angle widely dilated; abdo-
men short, rcbust, about as long as space between apex of
head and base of cruciform elevation; tympana entirely
exposed, tympanal coverings absent, only denoted by a
rudimentary convexity or ridge; anterior femora shortly or
obsoletely spined beneath ; opercula in male short, basal,
transverse ; posterior tibia strongly spined ; rostrum reaching
or just passing the posterior coxee; face not sulcate; tegmina
and wings hyaline; tegmina somewhat narrow, their greatest
breadth about one tiird their length, apical areas eight,
basal cell longer, but not twice as long as broad; wings with
six apical areas,
Type, B. Gilmoret, Dist. (Tibicen).
Burbunga inornata, sp. n.
. Body brownish ochraceous, sternum and legs ochra-
. . . ? . 5
ceous ; tegmina and wings hyaline, the venation ochraceous ;
50 Mr. W. L. Distant on Homoptera.
tegmina with the transverse veins at the bases of the apical
areas, and small spots at apices of longitudinal veins to apical
areas, obsoletely infuscate ; front of head narrow, prominent,
and conical, space between eyes a little narrower than ante-
rior margin of pronotum ; rostrum passing the posterior coxe ;
opercula short, transverse, their posterior margins moderately
convex, their inner angles well separated.
Long., excl. tegm., ¢ 15 mm.; exp. tegm. 48 mm.
Hab. West Australia: Champion Bay (Brit. Mus.).
Allied to B. Gilmore’, Dist., but with the front of head
more prominent, rostrum a little longer, and much paler in
hue and markings.
Genus TRISMARCHA.
Trismarcha, Karsch, Ent. Nachr. xvii. pp. 348, 349 (1891).
Type, 7’. umbrosa, Karsch.
Trismarcha atrata, sp. 0.
Pronotum castaneous; head, a central longitudinal fascia
(much widened posteriorly) to pronotum, mesonotum, and
abdomen above, piceous ; a marginal spot at anterior angles
of vertex of head, posterior margin of pronotum, spots to
cruciform elevation, and lateral areas of metanotum, ochra-
ceous ; body beneath brownish ochraceous ; face, area between
face and eyes, apex of rostrum, tarsi, lateral margins and a
central longitudinal fascia to abdomen beneath, piceous or
black ; tegmina and wings subhyaline, talc-like, venation
fuscous ; costal membrane to tegmina and basal anal area to
wings fuscous brown.
Long., excl. tegm., ¢ 23 mm.; exp. tegm. 65 mm.
Hab. Congo (Coll. Dist. and Paris Mus.).
Allied to J’. umbrosa, Karsch, but smaller; opercula in
male shorter and narrower, abdomen much more slender,
transverse vein at base of second apical area to tegmina more
vertical, and at fourth area much shorter.
Trismarcha angolensis, sp. 0.
&. Body and legs pale castaneous brown, ochraceously
pilose ; mesonotum with two obscure ochraceously margined
anterior obconical spots; abdomen beneath with the disk
piceous ; tegmina and wings smoky hyaline, the venation
fuscous brown; tegmina with the costal membrane pale
castaneous brown; head (including eyes) a little wider than
anterior margin of pronotum and about as wide as base of
Mr. W. L. Distant on Homoptera. 3
mesonotum ; basal joint of abdomen above centrally, longly,
anteriorly produced; rostrum reaching the posterior cox ;
front of head considerably shorter than vertex, their lateral
margins continuous, front centrally sulcate ; anterior femora
armed with three strong spines beneath ; operecula obliquely
transverse, narrow, inwardly convex, widely separated.
Long., excl. tegm., 3 20 mm.; exp. tegm. 62 mm.
flab. Angola (Dr. Welwitsch, Brit. Mus.).
Allied to 7. eacludens, Walk., but with the basal abdo-
minal segment more anteriorly produced, pronotum more
compressed and narrower, &c.
Genus MONOMATAPA.
Monomatapa, Dist. Ann, & Mag. Nat. Hist. (6) xix. p. 129 (1897).
Type, MZ. insignis, Dist.
Monomatapa socotrana, sp. n.
3. Body above testaceous brown, greyishly pilose; head
black, pronotum with a central hour-glass-shaped fascia (on
each side of which are two short oblique fasciw), the incisures
and lateral margins black ; mesonotum with four obconical
black spots (the two central ones shortest), and two black
spots in front of cruciform elevation ; metanotum and ante-
rior and lateral margins of basal abdominal segment black ;
body beneath pale dull ochraceous; central disk of face, space
between tace and eyes, apex of rostrum, femoral streaks, and
anterior and intermediate tibize and tarsi, piceous or black ;
tegmina and wings semihyaline, tale-like, the venation
fuscous ; opercula not passing basal segment of abdomen ;
rostrum reaching posterior coxe.
Long., excl. tegm., ¢ 16 mm. ; exp. tegm. 50 mm.
LTab. Socotra I. B. Balfour, Brit. Mus.).
Genus SELYMBRIA.
Selymbria, Stal, Ann. Soc. Ent. Fr, (4) i. p. 615 (1861),
Type, S. stigmatica, Germ. (Cicada).
Genus MALAGASIA.
Malagasia, Dist. Trans. Ent. Soc. Lond. 1882, p. 336.
Epora, Stal, Hem. Afr, iv. p. 41 (1861), nom. preeoce.
‘Type, M. inflata, Dist.
32 Mr. W. L. Distant on Homoptera.
Malagasia virescens, sp. n.
3. Body and legs pale virescent ; head with some black
shadings at the area of the ocelli and a short transverse line
at inner margins of eyes black ; tarsi fulvous brown ; tegmina
and wings pale hyaline, the venation virescent or fuscous ;
costal membrane and extreme base of tegmina and extreme
base of wings fulvous brown.
Long., excl. tegm., ¢ 18 mm.; exp. tegm. 60 mm.
Hab. Madagascar (Szkora).
Allied to MZ. Distant’, Karsch, from which it differs by its
totally different colour and markings and by the much greater
length of the second ulnar area.
Genus KANAKIA.
Kanakia, Dist. Ann. & Mag. Nat. Hist. (6) x. p. 62 (1892).
Type, K. typica, Dist.
Genus ABROMA.
Tibicen, snbgen. Abroma, Stal, Hem. Afr. iv. p. 27 (1866).
Type, A. Guerini, Sign. (Cicada).
Abroma Bowringi, sp. n.
?. Head and pronotum pale castaneous brown ; head with
the margins of front and nearly the whole of vertex piceous ;
pronotum with the margins and a central longitudinal fascia
pale ochraceous ; mesonotum ochraceous, with four obconical
piceous spots, the two central ones smallest ; abdomen above
and body beneath and legs dull ochraceous ; face pale casta-
neous, centrally and laterally ochraceous, between face and
eyes piceous ; tegmina and wings hyaline,‘ venation ochra-
cecus or brownish ochraceous, extreme bases of both testa-
ceous; tegmina with tlfe costal membrane and postcostal area
ochraceous ; body sparingly and finely greyishly pilose ;
rostrum reaching the intermediate coxz; head (including
eyes) slightly broader than base of mesonotum ; transverse vein
at base of second apical area to tegmina more or less vertical.
Long., excl. tegm., 9 13 mm.; exp. tegm. 35 mm.
Hab. China: Hong Kong (J. C. Bowring, Brit. Mus.).
LEMURIANA, gen. noy.
Ilead (including eyes) broader than base of mesonotum ;
. . ad .
head with the front prominent, not twice as broad as long,
he 2
3
Mr. W. L. Distant on Homoptera.
©
about as long or a little shorter than vertex, its margins and
those of vertex discontinuous, eyes projecting beyond the
anterior angles of pronotum, which is about as long as head,
its lateral margins sinuate, its posterior angles moderately
ampliate ; mesonotum (including cruciform elevation) nearly
as long as head and pronotum together; abdomen about as
long as space between apex of head and base of cruciform
elevation ; tympana entirely exposed ; opercula in male short,
narrow, transverse ; posterior metasternal process distinct, in
male, produced between the inner angles of the opercula ;
anterior femora strongly spined beneath ; tegmina and wings
hyaline ; tegmina with the two upper ulnar areas as long as
the remaining ulnar areas, apical areas eight; wings with six
apical areas.
Type, LZ. apicalis, Germ. (Cicada).
Lemuriana flavocostata, sp. n.
3g. Body above brownish ochraceous, ochraceously pilose ;
head with centre and apex of front, area of ocelli, and lateral
margins of vertex black ; pronotum with a very large casta-
neous spot occupying the whole of each lateral area and with
a small, central, elongate, black spot ; mesonotum with two
obscure, ochraceously margined, anterior, obconical spots
and two somewhat suffused, discal, black spots; abdominal
segmental margins piceous ; body beneath and legs ochra-
ceous; sternum longly greyishly pilose; face and space
between face and eyes castaneous, central area of face, and an
anterior fascia between face and eyes, black; apex of rostrum
and central segmental ventral spots piceous ; opercula mode-
rately long and broadened posteriorly ; rostrum reaching
posterior cox; tegmina and wings semihyaline, talc-like ;
tegmina with the venation fuscous, the costal membrane
flavescent ; wings with the venation dull ochraceous.
Long., excl. tegm., ¢ 15 mm.; exp. tegm. 48 mim.
Hab. Brit. East Africa: Voi (C. S. Betton, Brit. Mus.).
Lemuriana Sikore, sp. n.
Body castaneous ; head with the anterior angles of vertex,
area of ocelli, and 1 inner and posterior margins of eyes, pro-
notum with two spots on anterior and three transverse spots
on posterior margin, mesonotum with four obconical spots
(those on each lateral area longest, subcbsclete}, a posterior
lateral streak, and two spots in front of cruciform elevation,
black ; abdomen with the segmental margins black, broadest
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 3
34 Mr. W. L. Distant on Homoptera.
at base and apex ; body beneath paler castaneous than above ;
sternum, opercula, and legs ochraceous; a broad, central,
longitudinal fascia to face and a central fascia to abdomen,
black ; tegminaand wings hyaline, venation fuscous ; tegmina
with the costal membrane and claval area olivaceous ; wings
with the inner anal area dark olivaceous ; abdomen above
with a greyish pilose spot on each lateral area of the first and
second segments.
Long., excl. tegm., ¢ 19 mm.; exp. tegm. 58 mm.
Hab. Madagascar (Sikora).
Genus TAPHURA.
Taphura, Stal, Rio Jan. Hem. ii. p. 20 (1858).
Type, 7. misella, Stal (Cicada).
Genus LIGYMOLPA.
Ligymolpa, Karsch, Berl. ent. Zeit. xxxv. pp. 112 & 122 (1890).
Type, L. madegassa, Karsch.
Genus AUTA.
Auta, Dist. Ann. Mus. Civ. Genov. (2*) xvii. p. 381 (1897).
Type, A. insignis, Dist.
PANKA, gen. nov.
Head (including eyes) about as wide or a little wider than
base of mesonotum, its length not equal to breadth between
the inner margins of eyes; pronotum longer than head, its
lateral margins nearly straight, its posterior lateral angles
subdentately produced; abdomen in female a little longer
than space between apex of head and base of cruciform eleva-
tion; anterior femora strongly spined beneath ; tegmina and
wings hyaline; tegmina with eight, wings with five apical
areas.
Type, P. simulata, Dist.
The above generic description is based on two female
specimens ; the venation of the wings is a primary character ;
the characters of the opercula and other abdominal structures
can only be detailed when male specimens are examined.
Panka simulata, sp. n.
Tibicen nubifurca, Dist. (excl. syn.) Mon, Orient. Cicad. p. 131, tab. xiv.
fig, 24a, b (1892).
Hab, Ceylon.
Mr. W. L. Distant on Homoptera. 35
On examination of Walkers type (Cicada nubifurca,
Walk.,= Cicada apicalis, Kirby) I find that the wings have six
apical areas, and it should be included in the genus Abroma,
Stal. The species which I figured (supra) as T. nubifurca,
and which so closely simulates that species, therefore requires
renaming.
Panka africana, sp. n.
3. Body testaceous; front of head, pronotum (excluding
anterior and posterior margins and a central longitudinal
fascia), tympana, apex of abdomen (above and beneath), face,
legs, and opercula, ochraceous ; two anterior obconical spots
to mesonotum, a central longitudinal line to face, and apices
of tibie and tarsi, dark testaceous ; eyes and lateral margins
to vertex of head piceous; tegmina and wings hyaline,
venation mostly fuscous ; tegmina with the costal membrane
ochraceous, claval area purplish; wings with the margins of
the anal areas faintly fuscous.
Long., excl. teom., ¢ 10; exp. tegm. 30 mm.
Hab. West Africa: Fernando Po (c Conradt, Paris Mus.).
Genus NELCYNDA.
Tibicen, subgen. Nelcynda, Stal, difv. Vet.-Ak. Forh. 1870, p. 716.
Type, N. tener, Stal.
Nelcynda madagascariensis, sp. n.
2. Brownish ochraceous; head with the lateral margins
of front, a transverse line between eyes and traversing the
area of the ocelli, inner margins of eyes, and two small basal
spots, black ; pronotum with two central longitudinal spots
on anterior area, two central, elongate, oblique s spots on poste-
rior area, some spots on lateral areas, and the lateral margins,
black ; mesonotum with two small, centr al, anterior, obconical
spots, a sublateral fascia on each side, two spots in front of
and a transverse spot on each side of cruciform elevation,
black ; two central fasciz to face, which are united poste-
riorly, a spot between face and eyes, lateral margins of
clypeus, streaks to femora, and apices of tibie and rostrum,
piceous or black; tegmina and wings hyaline, the venation
fuscous ; costal membrane to tegmina testaceous 5 wings with
four apical areas.
Long., excl. tegm., 2 10 mm.; exp. tegm. 30 mm.
Lab. Madagascar: Fenerive (4. Perrot, Paris Mus.).
Bio
«
36 Mr. G. A. Boulenger on the
VI.—A List of the Freshwater Fishes of Africa.
By G. A. BouLenGer, F.R.S.
So many additions have been made to the African freshwater
fish-fauna during the last few years, over 400 new species
having been described by myself, that a full list, with a brief
indication of the habitats, seems worthy of publication. This
list is intended to serve as a basis for a general account of the
distribution of these fishes which I am now preparing for
the South African Meeting of the British Association.
In order to save space bibliographical references have been
omitted ; the indication of the author for each genus and
species, coupled with the year of publication, should, with the
help of the British Museum Catalogue and the series of
volumes of the ‘ Zoological Record’ (from 1864), suffice to
meet the requirements of the student.
Merely estuarine forms are excluded from tliis list.
Species occurring both in the sea and in fresh water are
marked with an asterisk after the indication of the habitat.
Ord. I. PLAGIOSTOMI.
Fam. 1, CaRCHARIUID™.
1. Carcharias, Cuv. 1817.
1. zambesensis, Peters, 1852. Zambesi.
Fam. 2. Pristip®.
1. Pristis, Latham, 1794.
1. Perroteti, M. & H. 1838. East Coast *.
Ord. II. CROSSOPTERYGII.
Fam. PoLtyprEerip®.
1. Polypterus, Geoffr. 1802.
1. bichir, Geoffr. 1802. Nile, L. Chad.
2. Lapradi, Stdr. 1869. Senegal, Gambia, Niger.
3. congicus, Blgr. 1898. Congo, L. Tanganyika.
4. Endlicheri, Heck. 1849. White Nile, Niger.
5. Delhexi, Blgr. 1899. Congo, L. Chad.
6, ornatipinnis, Blgr. 1902. Congo,
7. Weeksii, Blgr. 1898. Congo, Katanga.
8. senegalus, Cuv. 1829. Nile, L. Rudolf, Senegal, Gambia, Niger.
9. palmas, Ayres, 1850. Liberia, Congo.
10. retropinnis, Vaill. 1899. Congo.
2. Calamichthys, J. A. Smith, 1866.
1. ealabaricus, J. A. Smith, 1866. Niger Delta to Chiloango,
2
—
1p
2
»
Vv.
Freshwater Fishes of Africa. 37
Ord. III. DIPNEUSTI.
Fam. LEPIDOSIRENID®,
1. Protopterus, Ow. 1859.
annectens, Ow. 1839.
. ethiopicus, Heck. 1851.
Dolloi, Bigr. 1900.
Senegal, Gambia, Niger, L. Chad,
Zambesi.
White Nile to Great Lakes, Ugalla
Marshes, E, of L. Tanganyika.
Ogowe, Congo,
Ord. IV. TELEOSTEI.
Subord. MALACOPTERYGII.
Fam. 1.
. Elops, Li. 1766.
1. saurus, L. 1766.
2.
ll
ie
DID ors oo to
co
10.
1
12.
15.
14.
15.
16.
Life
lacerta, C. & V. 1846.
. Megalops, Lacep. 1803.
cyprinoides, Brouss. 1782.
ELoPID2.
E, and W. Coasts, Madagascar *.
W. Coast *.
E. Coast, Madagascar *.
Fam. 2. MormMyrip2&.
Mormyrops, J. Mill. 1848.
deliciosus, Leach, 1810.
anguillordes, L. 1766.
longiceps, Gthr. 1867.
breviceps, Stdr. 1894.
zanclirostris, Gthr. 1867.
engystoma, Blgr. 1898.
parvus, Bier. 1899.
Masuianus, Bler. 1898,
sirenoides, Bler. 1898.
Boulenger, Pellegr. 1900.
curtus, Bley. 1899,
lineolatus, “Blev. 1898,
nigricans, Blyy. 1899.
mucrostoma, Blgr. 1898.
Marie, Schilth. 1891.
attenuatus, Blgr. 1898.
Surcidens, Pellegr. 1900.
2. Petrocephalus, Marcus. 1854.
2.
. bane, Lacep. 1803.
Ansorgit, Blegr. 1902.
Sauvagti, Bley. 1887.
Keatingti, Blgr. 1901.
. Bovei, C. & V. 1846,
esa Sauy. 1885.
simus, Sauy. 1878.
gliroides, Vincig. 1897.
catostoma, Gthr, 1366.
3. Isichthys, Gill, 1863.
1
Henry, Gall, 1863.
4, Marcusenius, Gill, i183.
ip
Mar chit, Sauv. 1878.
Senegal to Congo, Zambesi, Webi
Shebeli, Juba, L. Nyassa.
Lower Nile.
Gold Coast.
St. Paul R. (Liberia).
Gaboon, Ogowe.
Congo.
Nile, L. Chad, Niger.
Lower Niger.
Old Calabar to Congo.
White Nile.
Nile, Senegal, Gambia.
Ogowe, Congo.
Liberia to Cameroon, Ogowe, Congo,
Angola.
Juba.
Rovuma, L. Nyassa.
Liberia to Mayumba.
Ogowe.
38
Mr. G. A. Boulenger on the
4, Marcusenius (con.).
2. sphecodes, Sauy. 1878.
3. brachistius, Gill, 1863.
4. longianalis, Bley. 1901.
5. Kingsleye, Gthr. 1896.
6. Cabre, Blgr. 1900.
7. adspersus, Gthr. 1866,
8. nigripinnis, Blgr. 1899.
9. pulverulentus, Bler. 1899.
10. Lhuystt, Stdr, 1870.
11. Ansorgi, Blgr. 1905.
12. Istdorz, C. & V. 1846.
13. pauciradiatus, Stdr. 1866.
14. Harringtoni, Bler. 1905.
15. Weeksw, Blgr. 1902.
16. plagiostoma, Bley. 1898.
17. tumifrons, Blgr. 1902.
18. Wilverthi, Bley. 1898.
19. discorhynchus, Peters, 1852.
20. Petherict, Blgr. 1898.
21. Budgetti, Blgr. 1904.
22. psittacus, Blgr. 1897.
5. Stomatorhinus, Blgr. 1898.
1. puncticulatus, Blgr. 1899.
2. Walker, Gthr. 1867.
3. humilior, Bley. 1899.
4, Corneti, Blgr. 1899.
5. polylepis, Blgr. 1899.
6. microps, Bley. 1898.
6. Myomyrus, Blgr. 1898.
1. macrodon, Blgr. 1898.
7. Gnathonemus, Gill, 1863.
. niger, Gthr. 1866.
. leopoldianus, Blgr. 1899.
. Bentleyi, Blgr. 1897.
. Fritel, Pellegr. 1904.
Livingstonit, Blgr. 1898.
. Moorv, Gthr. 1867.
. Lambouri, Pellegr. 1904.
. fuscus, Pellegr. 1901.
10. pictus, Marcus. 1864.
ll. Schilthuisia, Blgr. 1899.
12. cyprinoides, L. 1766.
13. senegalensis, Stdr. 1870.
14. Bruyerti, Pellegr. 1904.
15, angolensis, Blgr. 1905.
16. Stanleyanus, Blgr. 1897.
iz. mento, Blgr. 1890.
18. Monteiri, Gthr. 1873.
19. Petersiz, Gthr. 1862.
20. longibarbis, Hilg. 1888.
21 Guill, Blgr. 1904.
92, Ussheri, Gthr. 1867.
23. Abadii, Bley. 1901.
24. Greshoffi, Schilth. 1891.
25, kutuensis, Bley. 1899.
26. tamandua, Gthr. 1864.
CO CONT? OTR Co ND
. macrolepidotus, Peters, 1852.
S. Cameroon to Congo.
Sierra Leone to Congo.
Lower Niger.
Old Calabar, Congo.
Mayumba.
Lagos, Congo.
Congo.
”
Senegal, L. Chad.
Angola.
Nile.
Angola.
White Nile.
Congo.
re Ld
”
Zambesi, L. Nyassa.
White Nile, Blue Nile.
Lower Niger.
Congo,
Congo.
Ogowe.
Congo.
Congo.
Gambia.
Congo.
3?
99
Rovuma.
Zambesi, Katunga.
S. Cameroon to Congo.
Congo.
99
Upper Nile, White Nile.
Congo.
Nile, L. Chad, Niger, Congo.
Senegal, Gambia, L. Chad.
Ivory Coast.
Angola.
Congo.
Liberia, Gaboon.
Congo.
Niger, Calabar, Congo.
L. Victoria.
Lower Niger.
Liberia, Gold Coast.
Upper Niger.
Congo.
”
Niger, Calabar, Congo.
Freshwater
7. Gnathonemus (con.).
Fishes of Africa. 39
Congo.
27. mirus, Blgr. 1898. g
28. elephas, Blgr. 1898. L. Chad, Congo.
29. rhynchophorus, Blzr. 1898. Congo.
30, curvirostris, Blgr. 1898. is
31. zis, Bler. 1902. a
32. numenius, Blgr. 1898.
8. Genyomyrus, Blgr. 1898.
1. Donny?, Blgr. 1898. Congo.
9, Mormyrus, L. 1766.
1. Hasselquisti, C. & V. 1846. Nile.
2. Anchiete, Guim, 1884. Angola.
3. Lacerde, Casteln, 1861. L. Ngami.
4, Guentheri, Blgr. 1898. Niger.
5. macrophthalmus, Gthr. 1866. 5
6. ovis, Blgr. 1898. Congo.
7. caschive, Hasselq. 1757. Nile, L. Chad, Senegal to Congo.
8. rume, C. & V. 1846. Senegal.
9. niloticus, B]. 1801. Nile.
10. longirostris, Peters, 1852. Nile, Congo, Zambesi, L. Mweru.
11. Boze, Pellegr. 1903. Congo.
12, kannume, Forsk. 1775. Nile, L. Victoria, Webi Shebeli.
. caballus, Bley. 1898.
16.
10. Hyperopisus, Gill, 1862.
1. bebe, Lacep. 1803.
2. tenuicauda, Pellegr. 1904,
11. Gymnarchus, Cuv. 1829.
1. niloticus, Cuv. 1829.
Fam. 3.
. Notopterus, Lacep. 1800.
1. afer, Gthr. 1868.
. Xenomystus, Gthr. 1868,
1. nigrt, Gthr. 1868.
. Mildebrandti, Peters, 1882.
. proboscirostris, Blgr. 1898.
tenuirostris, Peters, 1882.
Athi R. (Ukamba).
Congo.
Athi R. (Ukamba).
Nile, L. Chad, Senegal, Gambia, Niger.
L. Chad.
White Nile, L. Chad,
Senegal to Niger.
NororPTERID».
Gambia to Calabar, Congo.
Bahr el Gebel, Liberia to Congo.
Fam, 4. OsTEOGLOSSIDZ.
. Heterotis, Cuv. 1829.
1. niloticus, Cuv. 1829.
Nile, L. Chad, Senegal, Gambia, Niger.
Fam. 5, PANTODONTID&.
. Pantodon, Peters, 1876.
1. Buchholz, Peters, 1876.
Low. Niger to Congo.
Fam. 6. PHRACTOL®MID®.
. Phractolemus, Blgy. 1901.
1. Ansorgu, Blgr. 1901.
Fam. 7
. Clupea, L. 1766.
1. finta, Cuv. 1829.
14 ,
Low. Niger, Congo.
. CLUPEID2.
N. Africa *.
Mr. G. A. Boulenger on the
. Pellonula, Gthr. 1868.
1. vorax, Gthr. 1868. Senegal to Congo.
2. acutirostris, Blgr. 1899. Congo.
. Microthrissa, Blgy. 1902.
1. Royauzxi, Blgr. 1902. Congo.
. Odaxothrissa, Blgr. 1899.
1. losera, Blgr. 1899. Congo.
. Pellona, C. & V. 1847.
1. indica, Swains. 1838. Kingani*.
. Chanos, Lacep. 1803.
1. mossambicus, Peters, 1852. Quisonga (Mozambique).
Fam. 8. SALMONIDZ.
. Salmo, L. 1766.
1. trutta, L. 1766. Atlas of Algeria,
Fam. 9. CRoMERIID.
. Cromeria, Blgr. 1901.
1. nilotica, Blgr. 1901. White Nile.
Subord. I. OSTARIOPHYSIL
Fam. 1. CHARACINIDZ.
. Sarcodaces, Gthr. 1864.
1. odoé, Bl. 1794. Senegal to Congo, L. Chad, L. Ngami.
. Hydrocyon, Cuv. 1817.
1. Forskalit, Cuv. 1817. Nile, Senegal, Niger.
2. lineatus, Blkr, 1863. Blue Nile, White Nile, Liberia, Niger,
Congo, L. Tanganyika and Mweru,
Zambesi, Limpopo, L. Ngami(?). ~
3. vittiger, Bler. 1898, Congo.
4. gohath, Blgr. 1898. 1
5. brevis, Gthr. 1864, Nile, L. Chad, Senegal, Gambia.
. Bryconethiops, Gthr. 1873.
1. microstoma, Gthr. 1873. Cameroon, Congo.
2. Vseuat, Blgr. 1899. Congo.
. Alestes, M. & T. 1846.
1. macrophthalmus, Gthr. 1867. Gaboon to Congo, L. Tanganyika,
L. Mweru.
2. Liebrechtsti, Blgr. 1898. Congo.
0. baremose, Joann, 1835. Nile, L. Rudolf, L. Chad, Senegal to
Niger.
4. dentex, L. 1766, Nile, L. Chad, Senegal to Niger.
5. Stuhlmanni, Pfeff. 1896, Kingani.
6. Tholloni, Pellegr. 1901. Congo.
7. intermedius, Blgr. 1903. Cameroon.
8, nurse, Riipp. 18382. Nile, L. Victoria, L. Rudolf, Senegal to
Cameroon.
9. humilis, Blgr. 1905. Angola.
10. amberi, Peters, 1852. Wami to Zambesi, Angola.
11. lateralis, Blgr. 1900. L. Dilolo, C. Africa.
12. senegalensis, Stdr. 1870. Senegal.
18. Lemairi, Blgr. 1899. L. Mweru.
14. natalensis, Blgr. 1904. Durban.
15. longipinnis, Gthr. 1864. Sierra Leone to Congo,
16. Chapert, Sauv. 1882. Gold Coast.
Freshwater Fishes of Africa. 41
4, Alestes (con.).
17. affinis, Gthr. 1894. Webi Shebeli to Tana.
18. teniurus, Gthr. 1867. Cameroon, Gaboon.
19. Fuchsii, Blgr. 1899. Congo.
20. bimaculatus, Blgr. 1899. x
21. Kingsleye, Gthr. 1896. Ogowe.
22. opisthotenia, Blgr. 1903. Cameroon.
23. brevis, Bler. 1903. Gold Coast, Lagos.
24, macrolepidotus, C. & V. 1849. Nile, L. Chad, Senegal to
Ogowe, L. Tanganyika.
25. grandisquamis, Bley. 1899. Congo.
26. Batesi, Blgr. 1903. Cameroon,
5. Micralestes, Blgr. 1899.
1. acutidens, Ptrs. 1852. Nile, Omo, Niger, Congo, Rovuma,
Zambesi.
2. humilis, Bler. 1899. Congo.
3. holargyreus, Gthr, 1873. ”
4, altus, Blgr. 1899. np
5. Stormst, Blgr. 1902. %
6. interruptus, Bler. 1899. ”
6. Petersius, Hilgend. 1894.
1. conserialis, Hilgend. 1894. Kingani.
2. occidentalis, Gthr. 1899. Gold Coast.
3. caudalis, Bley. 1899. Congo.
4, Leopoldianus, Blgr. 1899. »
5. Hilgendorfi, Blgr. 1899. -
6. modestus, Bley. 1899. 3
7. major, Blgr. 1905. Cameroon,
7. Eugnathichthys, Blgr. 1898.
1. Eetveldii, Blgr. 1898. Congo.
2. macroterolepis, Blgr. 1899. ”
8. Paraphago, Blgy. 1399.
1. rostratus, Bler. 1899. Congo.
9, Mesoborus, Pellegr. 1900.
1. crocodilus, Pellegr. 1900. Congo.
10. Phago, Gthr. 1865
; 1. loricatus, Gthr. 1865. W. Africa.
2. intermedius, Bler. 1899, Congo.
3. Boulengert, Schilth. 1891. a
11. Neoborus, Blgr. 1899.
1. ornatus, Blgr. 1899. Congo.
2. quadrilineatus, Pellegr. 1904. Casamanza.
12. Ichthyoborus, Gthr. 1864,
1. besse, Joann. 1855, Nile, White Nile, L. Chad.
13. Hemistichodus, Pellegr. 1901.
1. Vaillanti, Pellegr. 1901. Ogowe.
. Nannethiops, Gthy. 1871.
1. unitenatus, Gthr. 1871. White Nile, Gold Coast to Congo.
. Neolebias, Stdr. 1894.
~ 1. univittatus, Stdr. 1894, Liberia, Cameroon.
2. trilineatus, Blgr. 1899. Congo,
. Distichodus, M. & 'T. 1845.
1. notospilus, Gthy. 1867. Cameroon to Congo.
2. affims, Gthr. 1873. Congo.
3. altus, Bler, 1899. L. Chad, Congo.
4, nobolt, Bler. 1899. , Congo.
5, hypostomatus, Pellegr. 1900. Ogowe.
42 Mr. G. A. Boulenger on the
16. Distichodus (con.).
6. maculatus, Blgr. 1898. Congo.
7. Petersii, Pfeff. 1896. Kingani R.
8. Antonit, Schilth. 1891. Congo.
9. atroventralis, Blgr. 1898. 5
10. fasciolatus, Blgr. 1898. es
11. mossambicus, Peters, 1852. Zambesi.
12. brevipinnis, Gthr. 1864. White Nile, L. Chad, Senegal, Niger.
13. rostratus, Gthr. 1864. Nile, L. Chad, Senegal, Niger.
14, niloticus, L. 1766. Nile, White Nile, L. Rudolf.
15. engycephalus, Gthr. 1864. Nile, Blue Nile, Niger.
16. seafasciatus, Blgr. 1897. Congo.
17. lusosso, Schilth. 1891. 2
17. Nannocharax, Gthr. 1867.
1. brevis, Blgr. 1902. Congo.
2. fasciatus, Gthr. 1867. Gold Coast, Gaboon, Congo.
3. intermedius, Blgr. 1903. Cameroon.
4. niloticus, Joann. 1835. Nile, White Nile.
5. elongutus, Bley. 1900. Congo.
6. tenia, Blgr. 1902. >
7. dimidiatus, Pellegr. 1904. Casamanza.
18. Xenocharax, Gthr. 1867.
1. spilurus, Gthr. 1867. Cameroon to Congo.
2. crassus, Pellegr. 1900. Congo.
19. Citharidium, Bley. 1902.
1. Ansorgi, Bley. 1902. Low. Niger.
20. Citharinus, Cuv. 1817.
1. citharus, Geoflr. 1809. Nile, L. Chad, Senegal, Gambia, Niger.
2. congicus, Blgr. 1897. Congo.
3. macrolepis, Blgr. 1899. on
4. latus, M. & T. 1846. Nile, Senegal, Niger, Kingani ().
5. gibbosus, Blgr. 1899. Congo, L. Tanganyika.
Fam. 2. CyPpRiNIDZ.
1. Labeo, Cuv. 1817.
1. niloticus, Forsk. 1775. Nile.
2. horie, Heck. 1846. Nile, L. Albert, L. Chad, Senegal,
Gambia, Niger.
3. Steindachnert, Pfeff. 1896. Kingani.
4. altivelis, Peters, 1852. Zambesi, L. Nyassa.
5. Rose, Stdr. 1894. Limpopo R.
6. mesops, Gthr. 1868. L. Nyassa.
7. lineatus, Blgr. 1898. Congo.
8. velifer, Blgr. 1898. *
9. longipinnis, Blgr. 1898. 9
10. coubie, Rupp. 1832. Nile, L. Chad, Senegal, Gambia, Niger.
11. congoro, Peters, 1852, Zambesi.
12. Gregori, Gthr. 1894, Juba to Rovuma.
13. Newmanni, Blgr. 1908. Webi Shebeli,
14. Darlingi, Blgy. 1902. Rhodesia.
15. victorianus, Bley. 1901. L. Victoria.
16, Fuelleborni, Hilg. & Pappenh. 1908. L. Rukwa.
17. cyclorhynchus, Blgr. 1899. Ogowe, Congo,
18. falcipinnis, Blgr. 1908. Congo.
19. Kirkii, Blgr. 1903. ; Royuma.
20. Forskali, Riipp. 1885. Nile.
21. cylindricus, Peters, 1862. Hawash to Zambesi, L. Baringo.
Freshwater Fishes of Africa. 43
1, Labeo (con.).
22. macrostoma, Blezr. 1898. Congo.
23. nasus, Bler. 1899. 73
24. Greenii, Blgr. 1902. ”
25. brachypoma, Gthr. 1868. Lagos, Gold Coast.
26. annectens, Bley. 1903. Cameroon.
27. chariensis, Pellegr. 1904. L. Chad.
28. Lukule, Blgr. 1902. Congo.
29. parvus, Blgr. 1902. ”
30. barbatus, Bigr. 1898. +
31. capensis, A. Smith, 1841. Orange R., Limpopo R.
32. umbratus, A. Smith, 1841. Orange R.
2. Discognathus, Heck. 1845.
1. dembeensis, Riipp. 1837. L. Tsana, Nairobi R. (Kilimandjaro).
2. Johnstoni, Blgr. 1901. L. Victoria.
3. Vinciguerre, Blgr. 1901. Upp. Nile, White Nile.
4, Blanfordii, Blgr. 1901. Eastern watershed of Abyssinia,
Erythreea.
5. Hindi, Bler. 1905. Nyiro R.
6. makiensis, Blgr. 1905. Maki R. (Lake Zwai).
7. quadrimaculatus, Riipp. 1887. L. Tsana, Hawash R.
3. Varicorhinus, Rupp. 1837.
1. beso, Riipp. 1857. L. Tsana, Atbara R., Hawash R.
2. tanganice, Blgr. 1900. L. Tangauyika.
3. maroccanus, Gthr, Morocco.
4. Barbus, Cuv. 1817.
1. callensis, C. & V. 1842. Morocco, Algeria, Tunisia.
2. nasus, Gthr. 1868. Morocco.
3. serra, Peters, 1864. Oliphants R.
4. setivimensis, C. & V. 1842. Morocco, Algeria, Tunisia.
5. Vineiguerre, Pfeff. 1896. Wembere R. (Usambara),
L. Rukwa (?).
6. Welwitschii, Gthr. 1868. Angola.
7. Mattox, Guimar. 1884. 9
8. argenteus, Gthr. 1868. =
9. kurumant, Casteln. 1861. Kuruman R.
10. macropristis, Blgr. 1904. L. Victoria.
11. taitensis, Gthr. 1894. Taita district, inland of Mombasa.
12. paludinosus, Peters, 1852. Hawash to Zambesi, Durban.
13. longicauda, Blgr, 1905, Mozambique.
14, thikensis, Blgr. 1905. Thika (Tana System).
_ 16. Pfeffer, Blgr. 1905°. Unyamwesi district and Kingani.
16. tetraspilus, Pfetf. 1896. Upp. Ituri (Congo System).
17. zanzibaricus, Peters, 1868. Mombasa.
18. Newmayeri, Fisch. 1884. Neuruman (Masai district).
19. carpio, Pfeff. 1896. L. Albert.
20. Percivali, Blgr. 19038. Nairobi (Rufu).
21. lumiensis, Blgr. 1908, Lumi (Rufu).
22. laticeps, Pfeff. 1893. Wami R.
23. Kerstenii, Peters, 1868. | Between Mombasa and Kilimandjaro.
24. salmo, Pfeft. 1896. Pangani.
25. Kesslert, Stdr. 1866. Angola.
26. eutenia, Bler. 1904. =
27. holotenia, Blgr. 1904. Cameroon to Congo.
28. serrifer, Blgr. 1900. L. Tanganyika.
29. Ansorgit, Bley. 1904. Angola.
1 B, altus, Pfeft. 1896, nee Gthr.
44
4, Barbus
Mr. G. A. Boulenger on the
(con.).
30. miolepis, Blgr. 1902. Congo.
31. Guirali, Thomin. 1886. Cameroons, Beuito.
32. pleurogramma, Bler. 1902. L. Tsana.
33. Fritschii, Gthr. 1868. Morocco.
34. Rothschildi, Gthr. 1901. 9
35. Waldoi, Bley. 1902. A
36. atlanticus, Blgr. 1902. %
37. Harterti, Gthr. 1901. es
38. Riggenbachi, Gthr. 1902. ;
39. ksibi, Blgr. 1905. s
40. microterolepis, Blgr. 1902. Maki R. (L. Zwai).
41, macronema, Blgr. 1902, Maki R., Hawash.
. Lueppelli, Blgr. 1902.
3. Holub, Stdr. 1894.
. surkis, Rupp. 1857.
. Harrington, Blgr. 1902.
46. jarsinus, Blgr. 1902,
47.
48.
49,
50,
dl.
52.
55.
54.
55.
56.
57.
58.
59.
60.
61.
62.
63.
64.
65.
66.
67.
68.
69.
70.
(pe
72.
73.
74.
75.
76,
ht
78.
79.
80.
81.
82.
85.
84.
intermedius, Rupp. 1887.
fergussona, Bley. 1901.
eduurdianus, Blgr. 1901.
Breyert, M. Web. 1897.
plagiostomus, Bley. 1902.
plutystomus, Bley. 1902.
gorguart, Rupp. 1837.
bynnt, Forsk. 1775,
kassamensis, Bley, 1902.
gananensis, Vincig. 1895,
brevibarbis, Blgr. 1902.
leptosoma, Bley. 1902.
oreas, Blgr. 1902.
altianalis, Blgr. 1900.
Erlangeri, Blgr. 1908.
marequensis, A. Smith, 1841.
Duchesnii, Bley, 1902.
Radch fit, Bigr. 1903.
mento, Bler. 1902.
Gregort, Blgr. 1902.
hursensis, Bler. 1902.
affinis, Rupp. 1837.
Degen, Bigr. 1902.
nedgia, Riipp. 1837,
labiatus, Bler. 1902,
Reinii, Gthr. 1874,
LTinnelli, Lonnb. 1904.
Bates, Bler. 1903.
tanensis, Gthr. 1894,
Hindi, Blgr. 1902.
oxyrhynchus, Pfett. 1889.
perplevicans, Bley, 1902.
micronema, Bley. 1904.
Compinei, Sauy. 1879.
trimaculatus, Peters, 1852.
nummifer, Blgr. 1904.
Jacksoni, Gthr. 1889,
Pappenheimi, Bier. 1905
Hawash.
Orange R.
L. Tsana.
Hawash.
Jerrer R., near Harrar (Webi Shebeli).
I. Tsana.
L. Albert Edward.
Head-waters of Limpopo.
Hawash.
L. Tsana.
9)
Nile, L. Baringo.
Hawash.
Ganana R. (Jubi).
L. Tsana.
Jerrer R., near Harrar (Webi Shebeli).
L. Kivu and Rusisi.
Head-waters of Webi Shebeli.
“ Tnterior of S. Africa.”
L, Tsana, Mogre (Blue Nile).
L. Victoria.
Hawash R.
Nyuki R. (L. Baringo), Tana R.
Hawash R.
L. Tsana, Hawash, Maki.
L. Tsana.
LE)
Tana.
Morocco.
Cameroon.
”
Tana, Athi R.
” ”
Rufu (Pangani).
Tana.
Cameroon.
Ogowe.
Zambesi, L. Nyassa, Limpopo.
L. Victoria.
Head-waters of Rufu R. (Pangani).
L. Rukwa.
1
* Jacksoni, Hilg. & Pappenh. nec Gthr.
Freshwater Fishes of Africa.
4. Bar bus (con.).
101.
102.
103.
104,
105.
106.
107.
108.
109.
110.
TIDE
INE
nas
114.
115.
116.
We
118.
119.
120.
PAL
1
5. katange, Blgr. 1900.
. platyrhinus, “Bler. 1900.
. capensis,
. Burchelli, A. Smith, 1840.
. Trevelyant, Gthr. 1877.
. Gurney?t, Gthr. 1868.
. Bowkeri, Blgr. 1902.
2. rhodesianus, ‘Bler. 1902.
. uniteniatus, Gthr. 1867.
. lagoensis, Gthr. 1868.
. inermis, Peters, 1852.
}. zambesensis, Peters, 1852.
Of
98.
99.
100.
A. Smith, 1840.
progenys, Blgr. 1903.
golnoides, C. & V. 1842.
ablabes, Blky. 1863.
camptacanthus, Blky. 1868.
teniurus, Blgr. 1903.
Thollonianus, Thomin. 1886.
niyeriensis, Blgr. 1902.
trispilus, Blkr, 1863.
Walkert, Bley. 1904.
Fasciolatus, Gthr. 1868.
guadripunctatus, Pfett. 1896.
Pagenstecheri, Fisch. 1884.
tnnocens, Pfetf. 1896,
lineomaculatus, Bley. 1903.
amphigramma, Bler. 1903.
vulneratus, Casteln. 18617.
viviparus, M. Web. 1897.
perince, Rupp. 1837.
trispilopleura, Blgr. 1902.
humulis, Blgr. 1902.
neglectus, Blgr. 1903.
caudovittatus, Bler. 1902.
congicus, Blgr. 1899.
radiatus, Peters, 1852.
humeralis, Blgr. 1902.
22. Wernert, Blgr. 1905.
1238:
124.
125.
126.
127.
128.
129.
130.
131.
132:
138.
Dogyetti, Blgr. 1904.
pleuropholis, Blgr. 1899.
rapax, Stdr. 1894.
tropidolepis, Blgr. 1900.
afer, Peters, 1864.
anoplus, M. Web. 1897.
anema, Blgr. 1903.
stigmatopygus, Blgr. 1905.
pumilus, Blgr. 1901.
Brazz@, Pellegr. 1902.
je, Blgr. 1903.
5, Rasbora, Blky. 1859.
1. zanzibarensis, Gthr. & Playf. 1866.
6. Phoxinellus, Heck. 18438.
ue
Chargnoni, Vaill. 1904.
45
Lofoi (Katanga).
L. Tanganyika.
Oliphants and Berg R.
Cape Colony.
Berg R., Buffalo R.
Durban.
99
Zambesi.
Angola.
Lag os.
Zambesi.
7
Cameroon.
Cape of Good Hope.
Liberia, Gold Coast, Gaboon.
Fernando Po, Cameroon,
Benito R.
Cameroon.
Benito R.
L. Niger.
Gold Coast, Liberia (¥).
Gold Coast.
Angola.
Rutu R.
Mt. Kilimandjaro.
Wami, L. Rukwa.
Lumi (head-waters of Rufu R.).
Nairobi ( \E
Cape of Good Hope.
Natal.
Nile.
L. Tsana.
Les
Nile.
Congo.
Mozambique.
Congo.
Nile, L. Rudolf, Rovuma.
L. Victoria.
Congo.
KE. Cape Colony, Limpopo.
L. Tanganyika.
Cape C nlony.
Orange R., Klip R., Natal.
Nile.
White Nile.
99
Congo.
S. Cameroon.
Royuma.
Algerian and Tunisian Sahara.
1 multimaculatus, Stdr. 1870.
46 MerG. A. B
7, Alburnus, Weck. 1843.
1. callensis, Guich. 1850.
8. Leptocypris, Blgr. 1900.
1. modestus, “Bler. 1900.
9. Barihus, Ham. Buch. 1822.
: niloticus Joann. 1835,
. Weeksu, Blgr. 1899.
. longirostris, Blgr. 1902.
. microlepis, Gthr. 1864.
. tangamnce, Blgr. 1900.
. Guentheri, Bley. 1896.
. senegalensis, Stdr. 1870,
. Loati, Blgr. 1901.
. Kingsleye, Bigr. 1899.
. Buchholz, Peters, 1876.
. Moor, Bley. 1900.
. ubangensis, Pellegr. 1901.
. Weynsit, Blgr. 1899.
. zambesensis, Peters, 1868.
16. sardella, Gthr. 1868,
10. Chelethiops, Bler. 1899.
1. dibie, Joann, 1835.
2. elongatus, Bley. 1899.
Neobola, Vincig. 1895.
1. Bottegt, Vincig. 1895.
11.
2. argentea, Pellegr. 1904.
. Nemachilus, Hass. 1823.
1. abyssinicus, Blgr. 1902.
. microcephalus, Gthr. 1864.
oulenger on the
Algeria.
Congo.
Nile, Omo, Niger.
Congo.
1G, Nyassa,
L. Tanganyika.
L. Nyassa.
Upp. Shiré.
Senegal.
Nile, Bahr el Gebel, Omo.
Cameroon, Ogowe, Congo.
Ovzowe.
L Tanganyika, L. Rukwa.
Cameroon, Congo.
Congo.
Zambesi.
Shiré, L. Nyassa.
Nile, White Nile, Bahr el Gebel,
Webi Shebeli.
Congo.
Dana and Ganana R. (Juba), Omo,
LL. Rudolf.
L. Victoria.
L. Tsana.
Fam. 3. SILURIDZ.
1. Clarias, Gron. 1781.
1. lazera, C. & V. 1840.
Syria, Nile, L
. Albert Edward, L. Vic-
toria, Senegal, Niger, L. Chad , Congo,
L, Ngami (?).
2. Moorii, Blgr. 1901. L. Albert.
3. mossambicus, Peters, 1852. Mozambique, L. Rukwa.
4, Robecchit, Vincie. 1893. Abyssinia to Zanzibar, lL. Tanganyika.
5. Vinciguerre, Blgr. 1902. Hawash.
6. senegalensis, C. & V. 1840. Senegal, Niger,
7. Budgetti, Blgr. 1900. Gambia.
8. anguillaris, L. 1766. Nile, White Nile.
9. capensis, C. & V. 1849. Cape of Good Hope.
10. gariepinus, Burch. 1822. Zambesi and Katanga to Orange R.
and Natal.
11. tsanensis, Bley. 1902. IL. Tsana.
12. liocephalus, Blgy. 1898. L. Tanganyika.
13. longiceps, Blgr. 1899. Congo.
14. submarginatus, Peters, 1882. Gaboon (?).
15. breviceps, Blgr. 1900. Mayumba.
16. platycephalus, Blgr. 1902. Congo.
17. angolensis, Stdr. 1866. L. Niger to Angola.
. LAberiensis, Stdr. 1894.
Liberia, Sierra Leone, Cameroon,
Benito R.
Ls
oO co -~
“I
9.
10.
19%
Freshwater Fishes of Africa. 47
Clarias (con.).
19. Walkeri, Gthr. 1896.
20. bythipogon, Sauv. 1878.
21. macromystax, Gthr, 1864.
22. Dumerili, Stdr. 1866.
23. Carsonii, Blgr. 1903.
24, leviceps, Gill, 1862.
25. pachynema, Blgr. 1903.
26. Buettikofert, Stdr. 1894.
27. Sale, Hubr. 1881.
28. Theodore, M. Web. 1897.
29. ampleavicauda, Blgr. 1902.
. Allabenchelys, Blgr. 1902.
1. brevior, Bler. 1903.
2. longicauda, Bigr. 1902.
. Clariallabes, Blgr. 1900.
1. melas, Bley. 1887.
. Gymnallabes, Gthr. 1867.
1. typus, Gthr. 1867.
. Channailabes, Gthr. 1873.
1. apus, Gthr. 1873.
. Heterobranchus, Geoffr. 1809.
1. bidorsalis, I. Geoftr. 1827.
2. senegalensis, C. & V. 1840.
3. longifilis, C. & V. 1840.
4. laticeps, Peters, 1852.
5. tsopterus, Blkr. 1863.
. Plotosus, Lacep. 1803.
1. arab, Forsk. 1775.
. Eutropius, M. & T. 1849.
. congolensis, Leach, 1818.
. Grenfelli, Blgr. 1900.
. Adansonii, C. & V. 1839.
. liberiensis, Hubr. 1881,
. altipinnis, Stdr. 1894.
. niloticus, Rupp. 1829.
. Bocagivi, Guimar. 1882.
. Debauwi, Blgr. 1900.
. mentalis, Blgr. 1901.
CO CO NTO) Or B® CO hor
10. depressirostris, Peters, 1852.
11. mandibularis, Gthr. 1867.
12. Mebit, Pfeff. 1896.
13. laticeps, Blgr. 1899.
Schilbe, Cuv. 1817.
1. wranoscopus, Riipp. 1832.
2. Emini, Pfetf. 1895.
3. mystus, L. 1766.
4. senegalensis, C. & V. 1839.
Siluranodon, Blkr. 1858.
1. auritus, Geoffr. 1809.
Physailia, Blgr. 1901.
1. pellucida, Blgr. 1901.
2. somalensis, Vincig. 1897.
3. occidentalis, Pellegr. 1901.
Cameroon, Gabcon, Ogowe.
Calabar to Congo.
Gambia.
Angola.
L. Victoria, N. Nyassaland.
Gold Coast.
Cameroon.
Liberia.
99
Natal.
Congo.
Cameroon.
”
Congo.
Low. Niger, Old Calabar.
Congo, Angola.
Nile.
Nile, L. Chad, Senegal, Nigert
Nile!
Congo, Mozambique.
Gold Coast.
Zanzibar, Madagascar *.
Congo.
C. Chad, Congo.
Senegal, Cameroon.
Liberia, Gaboon, Ogowe.
Liberia to Congo.
Nile, Niger, Old Calabar.
Kwanga R. (Angola).
Congo.
Gold Coast, Cameroon.
Juba, Tana, Zambesi, Katanga,
Limpopo.
Liberia, Gold Coast.
Kingani R.
Congo.
Nile.
Uniamwesi Country.
Nile, L. Chad, Niger to Congo, L. Mweru,
L. Rukwa.
Senegal, Gambia, Niger,
Upp. Zambesi,
Low. Nile.
Upp. Nile.
Ganana (Upp. Juba).
Oowwe.
48 Mr. G. A. Boulenger on the
12. Parailia, Blgy. 1899.
1. congica, Blgr. 1899. Low. Niger, Congo.
2. longifilis, Bler. 1902. Congo.
13. Bagrus, Cuv. 1817.
1. bayad, Forsk. 1775. Nile, L. Chad, Senegal, Niger.
2. urostiyma, Vincig. 1895. Ganana (Upp. Juba).
3. orientalis, Bley. 1962. Pangani.
4, docmae, Forsk. 1775. Nile.
5. ubangensis, Blgr. 1902. Congo.
6. meridionalis, Gthr. 1894. Upp. Shiré.
14, Clarotes, Blky. US5s-
Ml laticeps, Riipp. 1829. Upp. Nile, White Nile, Bahr el Gebel,
40) Chad, Juba, Tana, Niger.
15. Chrysichthys, Blky. 1858.
1. furcatus, Gthr. 1864. Senegal to Congo.
2. Sharpi?, Blgy. 1901. L. Mweru.
3. Cranchit, Leach, 1818. Congo, L. Tanganyika.
4. ornatus, Blgr. 1902. Congo.
5. myriodon, Bler. 1900, L. Tanganyika.
6. brachynema, Bler. 1900. 55
7. punctatus, Blgr. ~ 1899. Congo.
8. Delhexi, Bler. 1859 Bs
9. Wagenaari, Bier. 1899. as
10. auratus, I. Geoffr, 1829. Nile, L. Chad, Niger, Gold Coast.
11. cameronensis, Gthr. 1899, Gambia, Cameroon.
12. Buettikoferi, Stdr. 1894. Liberia, Gold Coast, Niger.
13. Walkert, Gthr. 1899. Gold Coast.
14. coriscanus, Gthr. 1899. Corisco Island.
15. Kingsleye, Gthr. 1899. Ogowe.
16. nigrodigitatus, Lacep. 1803, Senegal, Gambia, Niger.
17. nigrita, Lacep. 1€03. Senegal.
18. acutirostris, Gthr. 1864. Angola.
19. brevibarbis, Bley. 1899. Congo.
20. longibarbis, Bler. 1899. A
16. Gephyroglanis, Blgr. 1899.
: congicus, Bler. 1299. Congo.
2. Sclaterz, Bler. 1901. Vaal R.
3, agoensis, Pellegr. 1900. Ogowe.
4. longipinnis, Bler. 1899. Cameroon to Congo.
17. Lemunema, Sauv. 1884.
1. borbonica, Sauv. 1884. Madagascar, Mauritius.
18. Amphilius, Gthr. 1864.
1. wranoscopus, Pfeff. 1889. Upper ranges of Wami R.
2. grandis, Bley, 1905. Head-waters of Tana System.
3. plutychtr, Gthr. 1864. Head-waters of Rufu R., between
L. Tanganyika and re Nyassa,
Sierra Leone (?).
4, longirostris, Blgr. 1901. S. Cameroon.
5. atesuensis, Bley. 1904. Gold Coast.
6. brevis, Blgr 1902. S. Cameroon, Congo.
7. angustifrons, Blgr. 1902. Congo.
19. Leptoglanis, Blgr. 1902.
1. aenognathus, Blgr. 1902. Congo,
20. Auchenoglanis, Gihr. 1865.
1. biscutatus, 1. Geoffr. 1829. Nile, L. Chad, Senegal.
2. occidentalis, C. & V. 1840. Nile, Senegal to Congo, if Rudolf,
L. Tanganyika.
21,
22,
23,
Ann.
Freshwater Fishes
. Auchenoglanis (con.).
3. punctatus, Blgr. 1902.
4. ubangensis, Blgr. 1902.
5. Ballayi, Sauv. 1873.
6. guttatus, Lonnb. 1895.
Notoglanidium, Gthr. 1902.
1. Walkeri, Gthr. 1902.
Ancharius, Stdr. 1880.
1. fuscus, Stdr. 1880.
Arius, C. & V. 1840.
1. biseutatus, Gthr. 1864.
2. Heudeloti, C. & V. 1840.
3. Parki, Gthr. 1864.
4. Kirkit, Gthr. 1864.
5, faicarius, Richards. 1845.
. Galeichthys, C. & V. 1840.
1. felceps, C. & V. 1840.
. Synodontis, Cuy. 1817.
. caudalis, Blgr. 1899.
. Depauwt, Bley. 1899.
. Tholloni, Blgr. 1901.
. multimaculatus, Bley. 1902.
. granulosus, Bley. 1900.
. schall, Bl. Schn. 1801.
. polyodon, Vaill. 1895.
. omias, Gthr, 1864.
. acanthomias, Blgr. 1899.
10. xiphias, Gthr. 1864.
11. humeratus, C. & V. 1840.
12. angelicus, Schilth. 1891.
13. frontosus, Vaill. 1895.
14. caudovittatus, Blgr. 1901.
15. nigrita, C. & V. 1840.
16. oceliifer, Blgr. 1900.
CO CONT SD Or i OO ND he
of Africa. 49
Congo.
39
Cameroon to Congo.
Cameroon, Congo.
Gold Coast.
Madagascar.
Fernando Po to Congo *.
Senegal *.
Liberia, Lagos, Niger *.
Zambesi *,
Pangani R. *
S. Africa *.
Congo.
”
Ozowe, Congo.
Congo.
L. Tanganyika.
Nile, Senegal, L. Rudolf and Stephanie.
Ogowe.
Niger.
Congo.
Niger.
Upp. Nile.
Congo.
White Nile, Omo R.
White Nile.
White Nile, Senegal, Gambia.
Gambia.
17. zambesensis, Peters, 1864. WebiShebeli to Zambesi, L. Rukwa,
18. multipunctatus, Blgr. 1898.
19. gamhiensis, Gthr. 1864.
20. nebulosus, Peters, 1852.
21. obesus, Blgr. 1898.
22. Greshoffi, Schilth. 1891.
23. eupterus, Blgr. 1901.
24, Afro-Fischeri, Hilgend. 1883.
25, Alberti, Schilth. 1891.
26. serratus, Riipp. 1829.
27. geledensis, Gthr. 1896.
28. filamentosus, Blgr. 1901.
29. labeo, Gthr. 1865.
30. Vaillanti, Blgr. 1897.
31. longirostris, Blgr. 1902.
32. guttatus, Gthr. 1865.
33. Robbianus, J. A. Smith, 1875.
34. Soloni, Blgr. 1899.
35. Smiti, Bler. 1902.
36. ornatipinnis, Blgr. 1899.
37. Fuelleborni, Hilg. & Pappenh. 1905.
38, melanopterus, Blgr. 1902.
L. Nyassa.
L. Tanganyika.
Gambia, Niger.
Zambesi,
Old Calabar, Cameroon, Gaboon.
Congo,
White Nile.
L. Victoria.
Congo.
Nile.
Webi Shebeli.
White Nile.
Niger.
Congo.
”
Lagos, Niger, Gaboon.
Gold Coast, Niger, Calabar.
Congo.
= ”
Congo, Angola.
L. Rukwa.
Low. Niger.
& Mag. N. list. Ser. 7. Vol. xvi. L
a0
25. Synodontis (con.).
39. notatus, Vaill. 1895.
40. nummifer, Blgr. 1899.
41. sorex, Gthr. 1864.
42. pleurops, Bley. 1897.
45. clarias, Li. 1766.
44. decorus, Blgr. 1899.
45. resupinatus, Blgr. 1904.
46. batensoda, Riipp. 1882.
47. membranaceus, Geoffr. 1809.
Microsynodontis, Blgr. 1903.
1. Batesiz, Blgr. 1903.
Chiloglanis, Peters, 1868.
1. Deckenit, Peters, 1868.
2. modjensis, Blgr. 1908.
3. cameronensis, Blgr. 1904.
;
26.
27.
. niloticus, Blgr. 1900.
. brevibarbis, Blgr. 1902.
Batesvi, Blgr. 1904.
Luchilichthys, Blgr. 1900.
1. Guentheri, Schilth. 1891.
2. Royauat, Blgr. 1902.
3. Dybowskit, Vaill. 1892.
9. Atopochilus, Sauv. 1878.
1. Savorgnant, Sauv. 1878.
. Mochocus, Joann. 1835.
1. niloticus, Joann. 1835.
31. Doumea, Sauv. 1878.
1. typica, Sauv. 1878.
. Lhractura, Blgr. 1900.
1. Bovet, Perugia, 1892.
2. Ansorgu, Blgr. 1901.
3. lindica, Blgr. 1902.
4. longicauda, Blgr. 1903.
>
28.
5. scaphirhynchura, Vaill. 1886.
3, Paraphractura, Blgr. 1902.
1. tenwecauda, Blgr. 1902.
. Trachyglanis, Blgr. 1902.
1. minutus, Bley. 1902.
. Belonoglanis, Bley. 1902.
1. tenwis, Bler. 1902.
. Andersonia, Blgr. 1900.
1. leptura, Blgr. 1900.
. Malopterurus, Lacep. 1803.
Mr. G. A. Boulenger on the
m |
Congo.
White Nile.
Congo.
Nile, L. Chad, Senegal, Gambia.
Congo.
Upp. Niger.
White Nile, L. Chad, Senegal,
Gambia, Niger.
White Nile, Senegal, Niger.
Cameroon.
Rufu R. (Pangani).
Modjo R. (Webi Shebeli).
S. Cameroon.
Upp. Nile.
Mathoiya R. (Tana).
S. Cameroon.
Congo.
2)
9
Ogowe.
Upp. Nile, White Nile, L. Rudolf.
S. Cameroon, Ogowe.
Congo.
Low. Niger.
S. Cameroon, Congo.
1
5. Cameroon.
Congo.
Congo.
Congo.
Congo.
Upp. Nile.
1. electricus, Gm. 1789. Nile and Senegal to Congo and Zambesi.
Subord. {1I]. APoDES.
Fam. ANGUILLIDZ.
1. Anguilla, Cuv. 1817.
1. vulgaris, Turt. 1807.
. virescens, Peters, 1852.
. Delalandii, Kaup, 1856.
one) Ee WS)
. Mildebrandti, Sauy. 1891.
. bengalensis, Ham. Buch. 1822.
. amblodon, Gthr. & Playf. 1866.
N. Africa *,
E. Africa, Natal *.
E. Africa *,
Madagascar, S. Africa *,
Seychelles *,
Madagascar *.
Freshwater Fishes of Africa.
Subord. 1V. HaPpLomrt.
Fam. 1. GALAXIID.
1. Galavias, Cuv. 1817.
1. zebratus, Casteln. 186].
2. punctifer, Casteln, 1861.
Cape of Good Hope.
” ”
Fam. 2. KNERIID#.
1, Kneria, Stdr. 1866.
1. angolensis, Stdr. 1866.
2. Spekii, Gthr. 1868.
Angola.
Coast of Zanzibar.
Fam, 3. CyPRINODONTID2.
1. Cyprinodon, Lacep. 1803.
1. calaritanus, Bonelli, 1829.
2. zberus, C. & V. 1846.
3. dispar, Riipp. 1828.
2. Tellia, Gerv. 1855.
1. apoda, Gerv. 1853.
3. Fundulus, Lacep. 1803.
1. divittatus, Lonnb. 1895.
. Loennbergri, Bler. 1903.
. nisorius, Cope, 1877.
. capensis, Garm. 1895.
. orthonotus, Peters, 1844.
. Guenther, Pfeff. 1893.
. melanospilus, Pfeff. 1896.
. tenopygus, Hilg. 1891.
. gularis, Bley. 1901.
10. Sjostedti, Lonnb. 1895.
11. microlepis, Vincig. 1897.
4. Haplochilus, McClell. 1859.
. sptlauchen, A. Dum. 1859.
. Chevahert, Pellegr. 1904.
. macrurus, Blgr. 1904.
. Chaperi, Sauv. 1882.
. Schellert, Bley. 1904.
. senegalensis, Stdr. 1870.
. Marni, Stdr. 1881.
. bifasciatus, Stdy. 1881.
. Loati, Blgr. 1901.
. atripinnis, Pfeff. 1896.
. Johnstont, Gthr. 1893.
. singa, Blgr, 1899.
. Decorsit, Pellegr. 1904.
. Petersii, Sauv. 1882.
. elegans, Blgr. 1899.
. sexfasciatus, Gill, 1865,
. fasciolatus, Gthr. 1866.
. homalonotus, A, Dum, 1861.
6 OO ~1S Or > Co bo
20.
21.
cameronensis, Blgr, 1903.
Antinori, Vincig. 1883.
. nuchimaculatus, Guich. 1866.
Algeria (N. and 8. of Sahara),
Low. Egypt, Somaliland.
Algeria.
Syria and borders of the Red Sea.
Atlas of Algeria.
Cameroon.
S. Cameroon.
Gaboon.
False Bay.
Mozambique.
Zanzibar coast and island.
Zanzibar coast, Seychelles.
L. Victoria.
S. Nigeria.
Cameroon.
Somaliland.
Senegal to Congo.
Congo.
Angola.
Gold Coast.
Low. Egypt.
Senegal.
White Nile.
Upp. Nile, White Nile.
White Nile.
KE. Africa (?).
L. Nyassa.
Congo.
”
Gold Coast.
Congo.
S. Nigeria to Congo.
Sierra Leone.
Madagascar.
”
S. Cameroon, R. Benito,
L. Arsadé (Shoa).
vr
47
Mr. G. A. Boulenger on the
4. Haplochilus (con.).
22. Playfairci, Gthr. 1866. Seychelles,
23. tanganicanus, Blgr. 1898. L. Tanganyika.
5. Procatopus, Bley. 1904.
1, nototenia, Bley. 1904. S. Cameroon,
Subord. V. CATOSTEOMI.
Fam. 1. GASTROSTEIDA,
l. Gastrosteus, L. 1766.
1. aculeatus, L. 1766. Algeria.
Fam. 2, SYNGNATHIDA.
1. Syngnathus, L. 1766.
Ll. algeriensis, Playf. 1871. Algeria.
2. zambesensis, Peters, 1855. Zambesi.
2. Celonotus, Peters, 1855.
1. argulus, Peters, 1855, Johanna (Comoro Is.),
Subord. VI. PERCESOCES,
Fam. 1. ScomMBRESOCID.
1. Hemirhamphus, Cuy. 1817.
1. Commersonit, Cuv. 1817. Licuare R. (Mozambique) *,
Fam. 2. ATHERINIDA,
1. Atherina, Art. 1738.
1. mocho, C. & V. 18365. Algeria, Low. Egypt *.
2. gabonensis, H. W. Fowler, 1904. Gaboon *,
2. Bedotia, Regan, 1903.
1. madagascariensis, Regan, 1903, Madagascar.
Fam. 3. Muaiipm.
1, Mugil, Art. 1738.
1. cephalus, L. 1766. N. and W. Africa *,
2. grandisquamis, C. & V. 1836. Senegal, Gambia *.
3. Constantia, C. & V. 1836. S. Africa *.
4. robustus, Gthr. 1861, Madagascar *,
5. capito, Cuv. 1829. N.W. and S. Africa *,
6. auratus, Risso, 1810. N.W, and 8. Africa, Madagascar *,
7. multilineatus, A. Smith, 1840. S. Africa *.
8. Smithii, Gthr. 1861. S. Africa, Madagascar *.
9. hypselopterus, Gthr. 1861. Niger, Gaboon *,
10. cephalotus, C. & V. 1836. Madagascar *,
11. fuleipinnis, C. & V. 1836. Senegal to Congo *,
2, Agonostomus, Benn. 1830.
a
2.
Telfairit, Benn. 1830. Mozambique, Comoro Is., Madagascar *.
dobuloides, C. & V. 1836. Madagascar *,
Fam. 4. PonyNEMID&.
1. Pentanemus, Art. 1758.
is
quinquarius, L. 1766. W. Africa *,
bo
Freshwater lishes of Africa Oo
. Polynemus, L. 1766.
1. quadrifilis, C. & V. 1829. W. Africa *.
. Galeoides, Gthr, 1860.
1. decadactylus, Bl. 1793, W. Africa *.
Fam, 5. SPHYRENIDA,
. Sphyrena, Art. 1738
1. guachancho, C. & V. 1829. W. Africa *,
Fam. 6, OPHIOCEPHALID®.
. Ophiocephalus, Bl. 1793.
1. obscurus, Gthr. 1861. White Nile, Bahr el Jebel, L. Chad,
Gambia to Congo.
2. insignis, Sauv. 1884, Ogowe, Congo.
3. africanus, Stdr. 1879. Lagos, Niger, “Old. Calabar.
Fam. 7. ANABANTIDZ&.
. Anabas, Cuv. 1817.
1. capensis, C. & V. 1881. Cape of Good Hope.
2. Bainsi, Casteln. 1861.
3. multispinis, Peters, 1846, "Mozambique.
4. nigropannosus, Reichen. 1875. Gaboon to Congo.
5, Pellegrini, Bley. 1902. Congo.
6. congicus, Blgr. 1887. oy
7. maculatus, Vhomin. 1886. S. Cameroon, Gaboon.
8. multifasciatus, Thomin. 1886. S. Cameroon to Congo.
9, fasciolatus, Blgr. 1899. Congo.
10. Petherici, Gthr. 1864. Bahr el Jebel.
ll. Kingsleye, Gthr. 1896. Senegal to Congo.
12. Weeks, Blgr, 1896. LL, Chad, Congo.
13. ocellatus, Pellegr. 1899. Congo.
14, oxyrhynchus, Blgy. 1902. -
Subord. VII. ACANTHOPTERYGII.
Fam. 1. CeENTRARCHID.
_ Kuhlia, Gill, 1861.
1. rupestris, Lacep. 1802. Zanzibar, Madagascar *.
2. tenura, C. & V. 1829. K. Africa, Natal, Seychelles *.
3. caudovittata, Lacep. 1802. Madagascar *.
Fam. 2. NANDID@.
. Polycentropsis, Blgr. 1901.
1. abbreviata, Blgr. 1901. Low. Niger.
Fam. 3. SERRANID ®.
. Morone, Mitchill, 1814.
1. punctata, Bl. 1792. Low. Nile, Senegal *.
. Plesiops, Cuv. 1817.
1. negricans, Rupp. 1828, Johanna (Comoro Is.) *,
Cr
bo
Mr. G. A. Boulenger on the
. Lates, C. & V. 1828.
1. niloticus, Hasselq. 1757. Nile, L. Chad, Senegal, Niger, Congo.
2. microlepis, Bler. 1898. L. Tanganyika.
. Ambassis, C. & V. 1828.
1. Commersonii, C. & V. 1828. Pangani R., Madagascar *.
2. urotenia, Blk. 1852. Seychelles *.
. Apogon, Lacep. 1802.
1. hyalosoma, Blkr. 1851. Seychelles *.
. Therapon, Cuv. 1817.
1. servus, BI. 1790. Mozambique *.
Fam. 4. Scla#NIDZ.
. Otolithus, Cuy. 1829.
1. senegalensis, C. & V. 1833. Senegal to Congo *.
Fam. 5. PristrrPoMATID2.
. Pristipoma, Cuy. 1829.
1. Jubelini, C. & V. 1830. Senegal to Congo *.
. Diagramma, Cuv. 1829.
1. macrolepis, Blgr. 1899. Senegal to Congo *,
Fam. 6. SPARIDZ,
. Chrysophrys, Cuy. 1817.
1. vagus, Peters, 1852. Mozambique *.
Fam. 7. ScoRPIDIDZ.
. Psettus, C. & V. 1831.
1. sebe, C. & V. 1831. Senegal to Congo *.,
2. argenteus, L. 1766. 8. Africa *.
3. falciformis, Lacep. 1802. pee es
Fam. 8. OSPHROMENIDZ.
. Micracanthus, Sauy. 1878.
1. Marchi, Sauv. 1878. Ogowe.
Fam. 9. CICHLIDZ.
. Hemichromis, Peters, 1857.
1. fasciatus, Peters, 1857. Senegal to Congo, L.Chad, Transvaal (?).
2. bimaculatus, Gill, 1862. Sahara, Nile, L. Chad, Senegal to
Congo.
3. angolensis, Stdr. 1865. Angola.
4, Frederici, Casteln. 1861. L. Ngami.
. Paratilapia, Blky. 1868.
1. Polleni, Blk. 1868. Madagascar.
2. Pfeffer’, Blgr. 1898. L. Tanganyika.
3. Moffati, Casteln, 1861. Transvaal.
4, dorsalis, Pellegr. 1902. Congo.
5. cerasogaster, Bilger. 1899. e
6. multicolor, Schoell. 1903. Lower Egypt.
7. longirostris, Hilg. 1888. L. Victoria.
8. Luebberti, Hilg. 1902. S.W. Africa,
9. Demeusii, Blgr. 1899, Congo.
Freshwater Fishes of Africa. 55
2, Paratilapia (con.).
10. moeruensis, Bley. 1899.
11.
12.
13.
14,
15.
16.
WZ:
18.
19.
20.
21,
22,
24,
25.
26.
2%
28.
29)
30.
ol.
32,
33.
afra, Gthr, 1893.
vittata, Bler. 1901.
Carlotte, Bler, 1905.
modesta, Gthr. 1893.
nototenta, Bler. 1902.
dimidiata, Gthr. 1864.
serranus, Pfeff. 1896.
victoriana, Pellegr. 1904,
robusta, Gthr. 1864.
macrocephala, Bley. 1899.
cavifrons, Hilg. 1888.
retrodens, Hile. 1888.
Thumbergi, Casteln. 1861.
Schwebischi, Sauv. 1884.
calliura, Bley. 1901.
intermedia, Gthr. 1864.
longiceps, Gthr. 1864.
ventralis, Bley. 1898.
Dewindti, Bler. 1899.
leptosoma, Bler. 1898.
nigripinnis, Bier, 1901.
stenosoma, Bler. 1901.
Surcifer, Bler. 1898.
3. Nanochromis, Pellegr. 1904.
di.
2,
nudiceps, Bler. 1899,
dimidiatus, Pellegr. 1900.
4. Pelmatochromis, Stdr. 1894,
1. lateralis, "ler. 1898.
: Jentinkt, Stdr, 1894,
. polylepis, Bley. 1900.
. Kingsleye, Bley, 1898.
. Guentheri, Sauy. 1882.
WwW elwitscht, Bley. 1898.
» Pellegrini, Bler. 1902.
: nigrofasciatus, Pellegr. 1900.
. longrrostris, Bley. 1903.
. Boulengeri, Lénnb. 1903.
. subocellatus, Gthr. 1871.
. Ansorgit, Bler. 1901.
. pulcher, Bley. 1901.
. tenatus, Bier. 1901.
- macrops, Blgr. 1898.
. auritus, Bler. 1901.
s multidens, Pelle er, 1900.
; Buettikoferi, Stdr. 1894,
. congicus, Bley, 1897.
5. Gyrtecara, Bigr. 1902.
1
Moorw, Blgy. 1902.
6, Lamprologus, Schilth. 1891.
1. tetracanthus, Blgr. 1899.
D St Coho
; tretocephalus, Blgr. 1899.
. modestus, Bler, 1898.
. tumbanus, Blgr. 1899.
. Mocquardi, Pellegr. 1903.
. congolensis, Schilth, 1891,
L. Mweru.
L. Nyassa.
L. Kivu.
Zambesi.
L. Nyassa and Shivé.
L. Nyassa.
L. Nyassa and Shiré.
L. Victoria.
+P)
L,. Nyassa, Katanga, Zambesi.
L. Mweru.,
L. Victoria.
99
L. Ngami.
Ogowe,
1D Tanganyika.
L. Nyassa, Shiré.
1a Tangan yika.
Congo.
”?
Congo.
Liberia, Gambia.
L. Tanganyika.
Gaboon, Ogowe.
Gold Coast.
Angola.
Low. Niger.
Cameroon to Congo.
Cameroon.
9
Gaboon.
Low. Niger.
”
L Ulaneanesk
1. Tanganyika.
)
C ongo.
Liberia, Congo.
Congo.
L. Nyassa.
L. Tanganyika.
”
>B
Congo.
99
ia
56 Mr. G. A. Boulenger on the
6, Lanpr ologus (con.).
(We)
7. Lemairii, Blgr. 1899.
8. Moorii, Bler. 1898.
9. Hecqui, Blgr. 1899.
10, fasciatus, Bier. 1898.
11. brevis, Blgr. 1899.
12. compressiceps, Bler. 1898.
13. fureifer, Bler. 1898.
14. elongatus, Blor. 1898.
wf . Julidochromis, Bley. “1898,
Ik ornatus, Bler. 1898.
. Telmatochromis, Bler. 1898.
1. temporalis, Blgr. 1898.
2. vittatus, Bler. 1898.
. Tilapia, A. Smith, 1840.
1. Moorti, Bler. 1901.
. Linnellit, Lonnb. 1908.
a
~~
Bloyet?, Sauv. 1883.
Wingati, Bler. 1902.
. Guiarti, Pellegr. 1904.
. Livingston, Gthr. 1893.
. Johnstont, Gthr. 1893.
. microlepis, Bley. 1899.
10. Huntert, Gthr. 1889.
11. negra, Gthr. 1894,
12. shirana, Bley. 1896.
13. mossambica, Ptrs. 1852.
14. nilotica, L. 1766.
CO OIG CUE ood
. Desfontainesi, Lacep. 1802.
L. Tanganyika.
mm =
L. Tanganyika.
L. Tanganyika.
”
L. Tanganyika.
Cameroon.
Algeria, Tunis.
Wami and Kingani R., L. Kivu.
Bahr el Jebel.
L. Victoria,
L. Nyassa and Shiré.
27 99
L. Tanganyika.
Crater lake of Kilimandjaro.
Kibwesi.
Upp. Shiré.
Coast of Zanzibar to Zambesi.
Nile, L. Victoria, L. Albert, L. Tsana, L.
Abeia, L. Kivu, E. Africa to Pangani R.,
Senegal to Niger, L. Chad.
15. natalensis, M. Web. 1897.
16. tanganice, Gthr. 1893.
17. galilea, Art. 1762.
18. flavomarginata, Bley. 1899.
19. Heudelot?, A. Dum, 1859.
20. nigripinnis, A. Dum. 1859,
21. macrocephala, Blky. 1862,
22. Dumerili, Stdr. 1864.
23. lepidura, Blgr, 1899.
24. Boulengeri, Pellegr. 1903.
25, sguamipinnis, Gthr, 1864.
26. Sparrmani, A. Smith, 1840,
27. ovalis, Stdr, 1866.
28. Burtoni, Gthr. 1893.
29. humilis, Stdr. 1866,
30. vorax, Pfeff. 1893.
31. Buettikoferi, Hubr. 1881.
32. Maria, Bley. 1899.
33. dubia, Lonnb. 1904.
34. Cabre, Bley. 1899.
5. Thollon?, Sauv. 1884.
6. crassa, Pellegr. 1903.
7. bilineata, Pellegr. 1900.
8. polyacanthus, Bley. 1899,
Co &8 TE
Coast of Zanzibar, L. Mweru, Congo,
Katanga, Zambesi, Natal.
L. Tanganyika.
Nile, Senegal, Gambia, Niger.
Ogowe, Congo.
Senegal to Congo, L. Chad.
Gaboon.
Gold Coast.
W. Africa (?).
Congo, Angola.
Congo.
L. Nyassa, Shiré.
Angola, Katanga, Zambesi to
Orange R.
Angola, Katanga, Bechuanaland.
ibe Kivu, ii Tanganyika.
Angola.
Zam besi.
Liberia.
Low. Niger.
Cameroon (Crater Lake),
Chiloango.
Ogowe.
Congo.
9
L. Mweru.
9.
Freshwater Fishes of Africa.
Tilapia (con.).
39, Zilli, Gerv. 1848.
40, kotte, Lonnb. 1904. .
41. Horii, Gthr. 1893.
42, Jalle, Bley. 1896.
43. melanopleura, A. Dum. 1859.
44, busumana, Gthr. 1902.
45, Dotlloi, Blgr. 1899.
46, guineensis, Blkx. 1862.
47, pleurotenia, Blgr. 1901.
48. trematocephala, Blgr. 1901.
49, Williamst, Gthr. 1893.
50. calliptera, Gthr. 1895.
51. Kirkit, Gthr. 1893.
52. strigigena, Pfetf. 1893.
53. lethrinus, Gthr. 1893.
54. pectoralis, Pfett. 1895.
55. nuchisquamulata, Hilg. 1888.
56. fasciata, Perugia, 1892.
57. Stormst, Blgr. 1902.
58. philander, M. Web. 1897.
59. acuticeps, Stdr. 1866.
60. Fuelleborni, Hilg. & Pappenh. 1903.
61. Livingstonii, Gthr. 1899.
62. Giardi, Pellegr. 1905.
63. Levaillanti, Casteln. 1861.
64, Andersoni, Casteln. 1861,
65. Dardenni, Bley. 1899.
66. zebra, Bler. 1899.
67. aurata, Blgy. 1897.
68. labiata, Blgr. 1898.
69. rostrata, Blgr. 1899.
70. boops, Blgr. 1901.
fae grandoculis, Bler. 1899.
72. oligacanthus, Blky. 1868.
. Hemitilama, Blgr. 1902.
ik oxyrhynchus, Bler. 1902.
. Stmochromis, Blegr. 1808.
1. diagramma, Gthy. 1898.
. Astatoreochromis, Pellegr. 1904.
1. Alluandi, Pellegr. 1904.
. Tropheus, Blgr. 1898,
ie Moorii, Bler. 1898.
2: annectens, Bler. 1900.
. Docimodus, Bler. 1896,
lis Johnstoni. Bler. 1896.
. Corematodus, Bley. “1896,
ie shiranus, Blegr. 1896.
. Steatocranus, Bler. 1899.
1. gibbiceps, Bler, 1899,
. Petrochromis, Bler, 1898.
1. tanganice, Gthr. 1893.
2. Andersoni, Blgr. 1901,
5. polyodon, Blgr. 1898.
4. nyasse, Bley. 1902,
57
N. Africa to Niger, L. Chad, White Nile,
L. Rudolf.
Cameroon (Crater Lake).
L. Tanganyika.
Zambesi.
Senegal to Congo, L. Chad,
Zambesi.
Gold Coast.
Gaboon, Congo.
Ashantee.
L. Tanganyika.
1b "Ny asSa.
L. Nyassa, Shiré.
White Nile, Wami.
L. Nyassa.
Rufu,
L. Victoria.
Congo.
”
Transvaal, Natal.
Angola, Zambesi.
L. Rukwa.
Zambesi.
”
L. Ngami.
”
L. Tanganyika.
L. Nyassa.
”
L. Tanganyika.
L. Nyassa.
L. Tanganyika,
7
Madagascar.
L. Nyassa.
—
4. Tanganyika,
L. Victoria,
L. Tanganyika.
»
Shiré.
Shiré.
Congo.
L. Tanganyika.
L. Albert.
4. Tanganyika.
L. Nyassa.
—
58 Mr. G. A. Boulenger on the
18. Chilochromis, Blgr. 1902.
1. Duponti, Blgr. 1902. Congo.
19. Asprotilapia, Bler. 1901.
1. leptura, Blgr. 1901. L, Tanganyika.
20. Paretroplus, Blkr. 1868.
1. Dam, Blir. 1868. Madagascar.
2. polyactis, Blky. 1870. 5
21. Eretmodus, Bler. 1898.
1. eyanostictus, Bley. 1898. L. Tanganyika.
22. Spathodus, Blgy. 1900.
1. erythrodon, Blgr. 1900. L. Tanganyika.
23. Perissodus, Bley. 1898.
1. microlepis, Bley. 1898. L. Tanganyika.
24, Plecodus, Blgr. 1898.
1. paradoxrus, Blgr. 1898. L. Tanganyika.
25. Xenochromis, Blgr. 1899.
1. Hecequi, Bley. 1899. L. Tanganyika.
26. Bathybates, Bley. 1898.
1. ferox, Blgr. 1898. L. Tanganyika.
2. fasciatus, Blgr. 1901. ”
27. Trematocara, Blgr. 1899.
1. marginatum, Blgr. 1899. L. Tanganyika.
2, unimaculatum, Bler. 1901. ne
28. Grammatotria, Bley. 1899,
1. Lemairi, Bley. 1899. L. Tanganyika.
29. Ectodus, Blgr. 1898.
1. Descampsii, Blgr. 1898. L. Tanganyika.
2. melanogenys, Blgy. 1898. a
3. longianalis, Bler. 1899. 3
30. Xenotilapia, Blgr. 1899.
1. sima, Blgr. 1899. L. Tanganyika.
2. ornatipinnis, Bley. 1899.
Fam. 10. PLEURONECTID.
1. Citharichthys, Blky. 1862.
1. spilopterus, Gthr. 1862. Niger, Gaboon *.
2. Cynoglossus, Ham. Buch. 1822.
1. senegalensis, Kaup, 1858. Senegal to Ogowe *,
Fam. 11. Gonirp”.
1. Lileotris, Gron. 1763.
1. ophiocephalus, C. & V.1887. Madagascar, Comoro, Seychelles *.
2. senegalensis, Stdr. 1870. Senegal to Ogowe.
3. Lebreton, Stdr. 1870. Senegal, Angola.
4, daganensis, Stdr. 1870. Senegal.
5. Fornasinit, Bianc. 1850. Mozambique.
6. madagascariensts, C. & V. 1887. Madagascar *,
7, fusca, Bl. Schn. 1801. Pangani, Madagascar, Comoro,
Seychelles *.
8. butis, Ham. Buch. 1822. Johanna (Comoro).
9. pectoralis, Regan, 1908, Madagascar.
10, Pisonis, Gimel. 1789. Liberia *.
11. Buettikoferi, Stdr. 1894. Liberia, Gaboon.
12. Monteiri, O’Sh. 1875, Gaboon, Angola.
Freshwater Fishes of Africa. 59
1, Lleotris (con.).
9
(de)
13. africanus, Stdr. 1880, Sierra Leone, Gaboon, Congo *.
14. nanus, Blgy. 1901. Upp. Nile, White Nile.
Gobius, Art. 1788.
1. rhodopterus, C. & V. 1837. Algeria *,
2. giuris, Ham. Buch. 1822. EK. and 8. Africa, Madagascar *.
3. spectabilis, Gthr. 1861. Natal.
4. gymnauchen, M. Web. 1897. Df
5. Dewaalii, M. Web. 1897. 5
6. Gichristi, Blgr. 1898. Cape of Good Hope.
7. paganellus, Li. 1766. Algeria *,
8. Schlegelu, Gthr. 1861. Gold Coast, Niger.
9. eneofuscus, Peters, 1852. Zambesi.
10. guineensis, Peters, 1876. Niger to Ozowe.
1]. macrorhynchus, Blky, 1875. Madagascar.
12. madagascariensis, Blky. 1875. %
13. sambiranoensis, Blkr. 1875. 5
14. hypselosoma, Blkr. 1875. $
15. polyzona, Blky. 1875. as
16. macrorhynchus, Blkr. 1875. ”
17. ocellaris, C. & V. 1837. eed:
Fam. 12. BLENNIIDZ.
. Blennius, Art. 1788.
1. vulgaris, Pollini, 1816. Algeria *.
. Alticus, Blkr. 1875.
1. monochrous, Blkr. 1875. Madagascar *.
. Cristiceps, C. & V. 1886.
1. argentatus, Risso, 1810. Algeria (?) *.
Subord. VIII]. OpistTHomt..,
Fam. MAsTACEMBELIDZ.
. Mastacembelus, Gron, 1781.
1. paucispinis, Blgr. 1899. Congo.
2. frenatus, Blgr. 1901. L. Tanganyika.
3. Moorit, Bler. 1898. c.
4. Marchit, Sauv. 1879. Ozowe.
5. Selatert, Bler. 1903. S. Cameroon.
6. cryptacanthus, Gthr. 1867, Cameroon.
7. congicus, Blgr. 1856. Congo.
8. hberiensis, Blgr. 1898. Liberia.
9. Loennbergit, Blgr. 1898. Niger, Cameroon, L. Chad.
10. Ansorgit, Bley. 1905. Angola.
11. marmoratus, Perugia, 1892, Congo.
12. goro, Blgr. 1902.
138. ellipsifer, Blgr. 1899. L. Tanganyika.
14, tanganice, Gthr. 1898. 93
15. shiranus, Gthr. 1896. L. Nyassa, Shiré.
16. victoria, Blgr. 1904. L. Victoria.
17. teniatus, Blgr. 1901. L. Tanganyika.
18. brachyrhinus, Blgr. 1899. Congo.
19. Greshoffi, Blgr. 1901. =
20. niger, Sauv. 1879, Ogowe.
60 Mr. C. T. Regan on the Dishes of the
1, Mastacembelus (con.).
21. flavomarginatus, Bley. 1898. Cameroon, Gaboon, Ozowe.
22. ngromarginatus, Blgr. 1898. Ashantee.
23. ophidium, Gthr. 1893. L. Tanganyika.
Subord. IX. PLECTOGNATHI.
Fam. TETRODONTID&.
1. Tetrodon, L. 1766.
1. fahaka, Hasselq. 1757. Nile, L. Chad, Senegal, Niger.
2. mbu, Blgr. 1899. Congo.
3. pustulatus, Murr. 1857. Old Calabar, Gaboon.
4, miurus, Bley. 1902. Congo.
VII.—A Revision of the Fishes of the American Cichlid Genus
Cichlosoma and of the Allied Genera. By C. Tarr Regan,
ISgAG
I GRATEFULLY acknowledge the kind assistance given to me
in the prepartion of this paper by Dr. Steindachner, to
whom I am indebted for information as to the number of
gill-rakers in the typical examples of Cichlosoma bifasciatum,
C. lentiginosum, C. altifrons, and C. Sieboldii,and by Dr. Th.
Gill, through whose intervention the British Museum has
acquired typical specimens of Cichlosoma centrarchus, C. bal-
teatum, C. rostratum, and C. basilare. Dr. S. Garman has
very kindly given me information as to the number of gill-
rakers and the length of the dorsal spines in C. pavonaceum.
In the descriptions of the species the number of scales in
a longitudinal series is counted from above the origin of the
lateral line to the base of the caudal fin, in a transverse
series from the base of the first dorsal spine to the lateral
line and thence to the middle of the abdomen. The number
of scales in a transverse series between the lateral line and
the base of the soft dorsal fin does not include the scales
forming the sheath which is usually present at the base of
the fin. The length of the last dorsal spine is measured
from its tip to its actual base, not to the free edge of the
scaly sheath at its base. The caudal peduncle is messured
from the level of the base of the last anal ray to the actual
base of the middle caudal rays, a point which is easily ascer-
tained by bending tlhe fin laterally. I have given a list of
the specimens in the British Museum collection on which
my descriptions are founded, with the total length in milli-
metres of cach,
American Cichlid Genus Cichlosoma. 61
The relations of the genera here dealt with are shown in
the following synopsis :—
Dorsal fin without notch between spinous and soft portions; anal fin
with more than 3 spines; gill-rakers short or of moderate length,
in small or moderate number.
I. Dorsal with XIII-XX 10-16 rays, anal with IV-XII 7-16.
A. Teeth all conical or cylindrical, not compressed.
Posterior (ascending) processes of preemaxillaries shorter
Gear Hepneme sey aio ey tfe, cho) « «9ioay Sea teats «il oped) eh Cichlosoma.
Posterior processes of preemaxillaries as long as the head. Petenza.
B. Teeth mostly conical, but the median teeth of the outer series
compressed, pointed in the young, truncate in the adult.
Herichthys.
C. Teeth all- rather broad and strongly compressed; scales
large, those of the lateral line of the same size as the others.
Teeth entire, with pointed or rounded apices ........ Paraneetroplus.
Teeth entire, truncate, incisor-like.............0+.0. Neetroplus.
Teeth tricuspid, except the median ones of the outer
series, which are entire, truncate, incisor-like .... Herotilapia.
D. Teeth slender, compressed, pointed in the young, rounded in
the adult ; scales rather small, those of the lateral line larger
CHM HE OLHERS erAca ys occas reeds sk cele s Varu.
Il. Dorsal with VIII-XIII 24-31 rays, anal with V-IX 24-82;
teeth conical.
Teeth confined to the anterior part of each jaw ...... Symphysodon.
Teeth extending on to the sides of the jaws.......... Pterophyllum.
CICHLOSOMA.
Cichlasoma, Swains. Nat. Hist. Fish. i. p. 280 (1839); Eigenm. &
Bray, Ann. Ac. N. York, vii. 1894, p. 615; Jord. & Everm. Proc.
U.S. Nat. Mus. xlvii. 1898, p. 1514; Pellegr. Mém. Soe. Zool. France,
xvi. 1903, p. 202 (1904); Meek, Zool. Pub. Columbian Mus. v. 1904,
204
p- 204.
Acara (part.), Heck. Ann. Mus. Wien, ii. 1840, p. 538; Giinth. Cat.
Fish. iv. p. 276 (1862); Steind. Sitzb. Ak. Wien, Ixxi. 1875, p. 65.
Heros, Heck. t.c. p. 862; Gunth. t. c. p. 285; Jord. & Everm. ¢. c.
p. 1526; Pellegr. ¢. c. p. 226.
Hoplarchus, Kaup, Arch. f. Nat. 1860, p. 128.
Theraps, Ginth. ¢. c. p. 284.
Mesonauta, Giinth. t. c. p. 300.
Archocentrus, Gill, Proc. Ac. Philad. 1877, p. 186.
Petenia (part.), Eigenm. & Bray, ¢. c. p. 614; Pellegr. ¢. c. p. 243.
Astronotus (part.), Kigenm, & Bray, ¢. ¢. p. 615.
Astatheros, Pellegr. t. ce. p. 203.
Thorichthys, Meek, t. c. p. 222.
Body deep or elongate, compressed ; scales large or mode-
rate, usually ctenoid. Two lateral lines. Jaws with bands
of small conical teeth, the outer series more or less enlarged
and sometimes forming canines. Mouth small or moderate.
62 Mr. C. T. Regan on the Fishes of the
Premaxillary processes shorter than the head; maxillary
exposed or not. Upper surface of head scaly to the level of
the orbits or beyond; cheeks and opercular bones scaly ;
preeoperculum entire. Gill-rakers rather short and few
(6-15 on the lower part of anterior arch). A single dorsal
fin, with XIV—XIX 7-15 rays; no notch between spinous
and soft portions. Anal with IV—XII 6-14 rays. Pectoral
asymmetrical, with 12-18 rays; ventrals below or a little
behind the base of the pectorals. Caudal rounded, truncate
or emarginate.
Numerous species from Mexico, Central and South
America.
I have examined the skeleton in four species, in which
the number of vertebree is as follows :—
Ci guttulatum 9. ost 14-16 =30,
Ce CeLracanthus @n obsess. lotro oe
C.octofasciatum. ~. . . 184+16=29)
C. Saint (sa) ae oe loo 25,
The ribs are subsessile. The supraoccipital crest is high
and extends forwards on to the frontals, which are hollowed
out anteriorly for the reception of the preemaxillary processes.
In C. guttulatum and C. octofasciatum these are compara-
tively short and the supraoccipital crest extends forward
nearly to the level of the anterior margins of the orbits. In
C. tetracanthus and C. Salvini the premaxillary processes
are longer and the supraoccipital crest terminates above the
middle of the orbits. The parietal crests are well developed
and extend forward to above the middle of the orbits. The
pelvic bones do not diverge anteriorly. In C. tetracanthus
the lower pharyngeals are strongly developed, they are
firmly united by a very sinuous suture, and bear large obtuse
teeth; the fourth vertebra bears a moderately developed
inferior apophysis. In the other three species the suture
between the lower pharyngeals is straight, the teeth are
pointed, whilst the inferior apophysis is feeble or wanting.
The genus Cichlosoma stands in the same relation to Acara
that the African Lamprologus does to Paratilapia, and, as
far as external characters are concerned, it is more difficult
to distinguish between Cichlosoma and Lamprologus than
between Acara and Paratilapia. C. Dovii, for example,
presents considerable similarity to L. congolensis in general
form, structure of the fins, structure of the mouth, dentition,
&e. The most notable difference is that in Lamprologus the
scales on the cheeks and opercular bones are usually
American Cichlid Genus Cichlosoma. 63
deciduous or wanting and the scales on the upper surface of
the head and of the anterior part of the body and on the
lower part of the abdomen are very small, which is not the
case in Cichlosoma. A comparison of the skeletons shows
that the African genus is distinguished by certain features of
specialization, such as the low occipital crest, the absent
parietal crests, the attachment of the ribs near the free ends
of the parapophyses, and the anterior divergence of the
pelvic bones.
A natural arrangement of the numerous species of this
genus is a matter of considerable difficulty. I have tried to
group the species according to their relationships and at the
same time to so define the various sections that they may be
mutually exclusive.
Synopsis of the Species.
I. Teeth of the outer series in the upper jaw usually more or less
distinctly and regularly increasing in size anteriorly (the anterior
pair not strongly enlarged unless “the lips are exceptionally thick).
Teeth of the outer series in the lower jaw either. similar to those
of the upper jaw or with the anterior 2 to 6 on each side enlarged,
subequal, and rather distinct from the smaller lateral teeth.
A. Scales of the lateral line of the same size as those below it on the
side of the body.
1. D. XITV-XVI9-11. A.IV(? V or VI) 8-9. Scales of thoracic
region not much smaller than those on the side of the body.
1. bomaculatum.
2. D. XIV-XVI 10-12. A. VITI-IX 10-12. Body increasing in
depth to below posterior part of dorsal fin.
2. festivum.
3. D. XV—-XIX 7-14. A. VI-XII 6-11. 6 to 8 gill-rakers on
the lower part of the anterior arch. Body ovate. Pectoral
extending to above the origin of anal or beyond. Soft dorsal
scaly at the base. Lips normal.
a. D, XV-XVII 9-11. A. VI-VIII 7-9. 1 or 12 scales be-
tween lateral line and base of anterior part of soft dorsal.
Depth of body 13-21 in the length; fold of the
lower lip not ‘continuous ; ; pectoral extending
toabove origin of anal”. 8 sc uccus sce oss 3. facetum,
Depth of body about 2 in the length ; fold of the
lower lip continuous; pectoral extending a
little beyond origin of arial Olathe HS.) 4, autochthon.
Depth of body 2? in ‘the HOMO CHHT oes: urate a ae ote 5. oblongun.
b. D, XVI-XVIII 11-14, A. VI-VHI 9-11.
21 scales between lateral line and base of anterior
mactot colt, dondal) tu.t ts ss 5, «cits Seven, 6. temporale.
4 scales between lateral }:je and base of anterior
part of soft dorsa} . coryphenoides.
e. D, XVII-XIX 7-10. A. VIII-XII 6-9
D. XVII-XVIIV 8-9, -A. VITI-X 6-8, Snout
much shortey than postorbital part of head.. 8. nigrofasciatwm.
/
,
/
,
64 Mr. C. T. Regan on the Fishes of the
D. XVITI-XIX 9-10. <A. VIII-X 7-8. Snout
not shorter than postorbital part of head.... 9. spilurum.
SV ERE 78. AS I 8 oe os igein 10. spinosissimum.
EP NOVA Xe xO 105 As Xe Oe ae cee pon iene re ll. immaculatum.
4, D. XVI 8-9. A. X-XI 8-9. 15 gill-rakers on the lower part
Of the anterior archin....- cme eel 12. centrurchus,
5. D. XV-XVIII 10-15. A. IV-VII 8-11. Scales of thoracic
region considerably smaller than those on the side of the
body. Either less than 11] gill-rakers on the lower part of
anterior arch ov the depth of body 4 or less than Z its length.
Pectoral not extending to above the anal. Soft dorsal more
or less distinctly scaly at the base.
a. Caudal truncate or rounded.
a. Last dorsal spine 3-3 the length of head; lower jaw
shorter than the upper; depth of body 23-23 in the
length. D. XVII-XVIII 12-15. A. V-VI 8-10.
Length of head 31-83 in the length of the fish... 13. Ezgenmanni.
Length of head 33-4 in the length of the fish .. 14. nebuliferum.
B. Last dorsal spine 2~2 the length of head; jaws equal
anteriorly.
+ Depth of body 13-2 in the length; caudal peduncle 3-$
as long as deep.
D. XVI-XVII-12-14. A. VI-VII 9-10. Last
dorsal spine more than 3 the length of head.
A large dark blotch on the caudal peduncle.. 15. maculicauda.
D. XVII-XVIII 11-13. A. VI-VII 8-9. Last
dorsal spine 4 the length of head. Dark
cross-bars and a dark band from operculum
to ‘base jot caudal. 2A. .c. - cian puis dasa 16. fenestratum.
++ Depth of body 2 in the length; caudal peduncle as long
as deep; last dorsal spine § the length. of head.
1D. VATS. ACME ON ace 17. bifasciatum.
+tt+ Depth of body 21-22 in the length ; caudal peduncle
from 3 to as long as deep; last dorsal spine 2-3 the
length of head.
* 8 to 10 gill-rakers on the lower part of the anterior
arch; anal with 6 or7 spines. 18. guttwlatum.
** 7 or 8 gill-rakers on the lower part of the anterior
arch ; anal with 4 to 6 spines.
t Fold of the lower lip continuous.
19. microphthalmus.
t{ Fold of the lower lip not continuous or sub-
continuous.
§ Caudal peduncle 3 as long as deep.
20. sexfasciatum.
§§ Caudal peduncle from } to as long as deep.
3 scales between lateral line and base of anterior
part of soft dersai; interorbital width 23-8
in the length of head; 2 dark longitudinal
bands or series of spots, one above the upper
lateral line, the other from the middle of side
to fhe base of cad@all oc.) <2. lin sc tee one 21> melanurum.
3 scales between lateral line and base of anterior \
part of soft dorsal; interorbital width 2}-2? \
American Cichlid Genus Cichlosoma. 65
in the length of head ; dark cross-bars and a
dark longitudinal band from operculum to
Rimsencaee ee APRIL PMc erate acon ss eaten vin ats © 5:9 22. Gadovit.
2 or 24 scales between lateral line and base of
anterior part of soft dorsal; a broad dark
band from operculum to middle of side and
thence running upwards to the dorsal fin..,, 23. intermedium.
tttt Depth of body 23-25 in the isagity
eee VLU TS 5 AVL O10 os Sees Sat ne avis 24. Guenthert.
Bee WE ra VE Os tes aie ols Sots ole ass le thea 2 25. pavonaceum.
y- Last dorsal spine 3 the length of head; depth of body
24-23 in the length.
D. XVII11. A. V8. Depth of body 23 in the
Rope ote ens sechie s aeee se onc nb ant 26. Steboldiz.
D. XVI_XVU 12-13. A.V 10. Depth of body
aoe WW HO LONE. 3)5, < 4/5, o:0! a's ia, « AP See 27, Godmanni.
6. Last dorsal spine 3-% the length of head; depth of body
3-33 in the length. D. XV-XVII 12-14. A. IV-V 9-10.
28. wregulare.
6. Caudal emarginate, with rounded lobes.
29. lentiginosum.
6. D, XVIUI-XIX 10-11. A. VII-VII 7-9. 9 or 10 gill-rakers
on the lower part of the anterior arch. Pectoral ex-
tending to above the origin of anal or beyond.
Interorbital width 3 in the length of head; pec-
toral extending to above the origin of anal.. 380. balteatum.,
Interorbital width 31 2% in the length of head ,
pectoral extending a little beyond the origin
Dia aerler, pets SAA“ a-U Nupad ures al aed wie doer sis 31. nicaraguense.
7. D. XIV-XVII 10-14. A. V-VII 8-10. 9 to 13 gill-rakers on
the lower part of the anterior arch. Depth of body ? its
length or more. Pectoral extending beyond origin of anal or
11 to 13 gill-rakers on the lower part of the anterior arch.
Soft dorsal more or less distinctly scaly at the base. Lips
normal.
a. Fourth to sixth dorsal spines longer than the rest.
D; XVI 10-11. A. VI 8-9.
Pectoral extending to above the middle of anal.. 32. Rodertsoni.
Pectoral extending to the posterior end of anal... 33. longimanus.
b. Dorsal spines either subequal from the fifth or sixth or
increasing in length throughout.
a. D. XIV-XV 12-15. A. V8-9. Snout shorter than post-
orbital part of head.
Pectoral extending beyond the origin of anal.... 34. macracanthus.
Pectoral not extending beyond the origin of anal. 35, heterodontus.
8. D. XVI 11. A. V 8-9. Snout longer than postorbital
part of head; pectoral extending nearly to the origin of
Bi ine Caer ideprs econ 36. altifrons.
y. D. XVI-XVII 11-12. A. VII 8-9.
+ -Pectoral extending to above the last anal spine.
37. rostratum.,
~
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. a
66 Mr. C. T. Regan on the Fishes of the
++ Pectoral extending to aboye the third or fourth anal
spine.
Caudal slightly emarginate, with rounded lobes.. 388. margaritiferum.
Caudal rounded ..........00¢ ee porn Seok 39. citrinellum.
8. D. XVII 11-12. A. VII-VIII 8-9. Pectoral extending be-
yond the origin of anal. Lips exceptionally thick.
a: Laps subuormalys oot «2d ne 40. erythreum.
b. Each lip produced as a long fleshy triangular flap.
Depth of body 2} in the length, length of head
TLC HURLY Siu ns ote ies cielo (alec fore ater ein ieee ieee 41. lobochilus.
Depth of body 23 in the length, length of head
ee le ahaa: ies eRe eae el oA. eeleree ete eer 42. labiatum.
9, D. XV-XVII 8-12. A. VI-IX 6-9. Dorsal fin entirely scale-
less. Caudal slightly emarginate, with pointed lobes.
a, Last dorsal spine 3 the length of head; pectoral extending
beyond origin of anal.
Snout not longer than postorbital part of head .. 43. awreum.
Snout longer than postorbital part of head...... 44, affine.
b. Last dorsal spine 2 the length of head or less; pectoral ex-
tending to above originof anal.... 46. cadllolepis.
10, D. XV-XVII 13-14. A. VII-VIII 12-14. Body ovate.
46. severum.
B. Scales of the lateral line larger than those below it on the side of
lite) seri MA aia Hole aie Heid. Rana se 47. psittacum.
Il. Teeth of the outer series in the upper jaw increasing in size ante-
riorly, with the anterior pair more or less distinctly differentiated
as canines. Anterior pair of teeth in the lower jaw, if present,
smaller than the next 1 or 2 pairs, which are more or less strongly
enlarged and canine-like. Lipsnormal. Scales of thoracie region
small.
A. D. XV-XVI 10-12. A. IV 8-10.
Caudal peduncle nearly as long as deep ........ 48. adspersum.
Caudal peduncle 2-3 as long as deep .......... 49, tetracanthus.
B. D. XV-XVI 10-18. A. V (1V-VI) 7-10.
1. Snout nearly equal to postorbital part of head (in the adult) ;
caudal peduncle as long as deep.
Depth of body 22-23 in the length; jaws equal
auberionly er. ei = eee teeters er een eenee 50. estlanum.
Depth of body 23-23 in the length; lower jaw
PKOPERIMO con tere tal a eile gee arse 51. Bartont.
2, Snout shorter than postorbital part of head (in the adult) ;
lower jaw projecting.
Depth of body 2-23 in the length; caudal pe-
duncle 2 as long ‘as deep. <. ce. <1.) demise Sel 52. Beant.
Depth of body 23-2¢ in the length; caudal pe-
duncle as long as deep’... ats: aujerace eine 53. mento.
3. Snout longer than postorbital part of head (in the adult) ;
depth of body 23-3 in the length.
Lower jaw slightly projecting (in the adult) ;
caudal peduncle deeper than long.......... 54. Feste.
Lower jaw a little shorter than the upper (in the
adult) ; caudal peduncle as long as deep.... 55. ornatum.
American Cichlid Genus Cichlosoma. 67
C. D. XVII-XIX 8-10. A. VIII-X 7-8. Przmaxillary processes
extending to above the anterior margin of eye.
56. octofasciatum.
D. D. XV-XVIII 9-12. A, VI-IX 7-10.
1. Preemaxillary processes extending to above anterior 4 of eye.
D. XV-XVII 10-12. A. VI18-8.... 57. urophthalmus.
2. Premaxillary processes extending to above middle of eye or
beyond.
a. Last dorsal spine 3 or nearly 3 the length of head; maxillar
extending to below anterior margin of eye; D. XVI-XVII
9-12.
Snout shorter than postorbital part of head (in the
aha, Uo WEE BUUISE) Teale Sh sear eesee 58. trimaculatum.
Snout as long as postorbital part of head (in the
adult). A. VIDI-IX 7-9 ...., ACs a eee 59. Salvint.
6. Last dorsal spine about 2 the length of head; maxillary ex-
tending to below anterior } of eye; D. XVIII10. A. VII-
VU 8-9; depth of body 23-23 in the length; caudal
peduncle 3-2 as long as deep...... 60. multifasciatum.
c. Last dorsal spine about 3 the length of head.
a. Maxillary extending to below anterior margin or anterior
4 of eye; D. XVII-XVIII 9-11. A. VII-VII 8-9;
depth of body 22-22 in the length.
Caudal peduncle 2 as long asdeep ; depth of prae-
orbital 3-2 the diameter of eye (in specimens
of 120-130 mm. in total length) .......... 61. Friedrichstahli.
Caudal peduncle 3-4 as long as deep; depth of
preeorbital 2 the diameter of eye (in a speci-
men of 115 mm.) or equal to it (in the adult
UD RPE Aranda CY <)'s( o's wie. d uae! gre late ain aiehaiaichel 62. motaguense.
8. Maxillary extending to below middle ofeye. D. XVIII 10.
AE BEEP ios -c seat arent ci sve glans cos ake 63. managuense.
y. Maxillary extending to below anterior 3 of eye.
D. XVIIT 12. A. VI9-10 .... 64. Dovit. :
d, Last dorsal spine 3 or nearly 3 the length of head ; maxillary
extending to below the middle of eye.
D. XV 12-13. A. VI 9-10; scales above lateral
line of the same size as those below it...... 65. spectabile.
D. XV-XVI 10-11. -A. VI 8-9; scales above
lateral line smaller than those belowit .... 66. Kraussiv.
Subgenus CrcHLosoma.
In the more generalized forms the teeth of the outer series
in both jaws rather small, scarcely increasing in size ante-
riorly. In more specialized forms the teeth of the outer
series stronger, regularly and distinctly increasing in size
anteriorly. Sometimes the anterior 3 to 6 teeth on each
side in the lower jaw enlarged, subequal and rather sharply
differentiated from the smaller lateral teeth. In 3 very
specialized species (with exceptionally thick lips) the anterior
pair of teeth im the upper jaw very strong, the anterior 2
pairs in the lower jaw strongly enlarged, subequal.
5x
68 Mr. C. T. Regan on the Fishes of the
Section 1 (Cichlosoma).
Body ovate ; scales of lateral line of the same size as those
above and below it; scales of thoracic region scarcely
smaller than those on the side of the body. Mouth rather
small, moderately protractile; maxillary not exposed; teeth
of the outer series rather small, scarcely increasing in size
anteriorly. Dorsal X1V-XVI 9-11, the soft fin scaly at the
base. Anal IV (? V or VI) 8-9. Caudal rounded.
The single species, from South America, is undoubtedly
closely allied to Acara portalegrensis. On the other hand, it
shows clear affinities with Cichlosoma facetum and its allies.
1. Cichlosoma bimaculatum.
Sciena bimaculata, Linn. Mus. Ad. Fried. i. p. 66 (1754),
Labrus bimaculatus, Linn. Syst. Nat. (ed. x.) p. 285 (1768).
Labrus punctatus (part.), Linn, 7. ¢.
Labrus punctatus, Bloch, Ausl. Fische, vi. p. 20, pl. eexey. (1792) ;
Schneider, Bloch’s Syst. Ichth. p. 251 (1801).
Perca bimaculata, Bloch, t. ec. p. 82, pl. ecex.
Cichla bimaculata, Schneider, o. c. p. 338.
Chromis tenia, Bennett, Proc. Zool. Soc. 1. 18380, p. 112; Storer,
Mem. Ac. Amer, ii. 1846, p. 520.
Acara margarita, Heck. Ann. Mus. Wien, ii. 1840, p. 338.
Acara marginata, Heck. t. c. p. 350.
Acara punctata, Heck. t. e. p. 560,
Acara Gronovii, Heck. t. c. p. 361.
Cychlasoma tenia, Gill, Ann, Lye. N. York, vi. 1858, p. 388,
Heros bimaculatus, Cope, Proc. Ac. Philad. xxiii. 1872, p. 254.
Acara (Heros) bimaculata, Steind. Sitzb. Ak. Wien, Ixxi. 1876, p. 82.
Astronotus (Cichlasoma) bimaculatus, Higenm. & Bray, Ann. Ac. N.
York, vii. 1894, p. 618.
Cichlasoma bimaculatum, Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p. 204 (1904).
Depth of body 1}-2! in the length, length of head 23-3.
Eye much nearer to tip of snout than to extremity of oper-
culum, its diameter 23-4 in the length of head, interorbital
width 2-23. Depth of preorbital equal to the diameter of
eye or less. Maxillary extending to the vertical from ante-
rior margin of eye; premaxillary processes extending nearly
to above anterior margin of eye; jaws equal anteriorly ;
fold of the lower lip not continuous ; cheek with 3 series of
scales; 6 or 7 gill-rakers on the lower part of anterior arch.
Scales 25-27 *, 2 between lateral line and base of anterior
part of soft dorsal. Dorsal XIV-XVI 9-11, commencing
above the opercular cleft, the spincs slightly increasing in
length to the last, which is 2 the length of head. Anal
IV (V-VI1) 8-9. Pectoral as long as the head, extending to
Aimerican Cichlid Geaus Cichlosoma, 69
above the anal spines; ventral often extending to posterior
end of anal. Caudal rounded. Caudal peduncle about ?
as long as deep. Brownish, with 8 dark cross-bars, the third
bearing a blotch below the lateral line; a dark spot below
the posterior part of eye and another on the upper part of
the base of caudal; a longitudinal band from eye to lateral
blotch; each scale of the lower half of the body with a light
spot at its base ; each scale of the posterior half of the body .
with a dark spot; soft dorsal, anal and caudal spotted.
Brazil; Guiana; Trinidad.
1-2. (72 and 83 mm.) Demerara. Dr. Hancock.
3-5, (83-121 mm.) British Guiana. Sir R. Schomburgk,
6-14. (73-141 mm.) Guiana. Berlin Museum.
15. (150 mm.) Trinidad. Zoological Society.
16-17. (93 and 107 mm.) Bahia. Dr. Wucherer.
18-21. (41-56 mm.) Demerara. F. G. Beckford, Esq.
22-56, (40-92 mm.)
37-89, (70-112 mm.) Berbice. F. G. Beckford, Esq.
40-42, (50-105 mm.) Demerara.
43-46, (144-156 mm.) British Guiana.
47. (180 mm.) Trinidad. F. W. Urich, Esq.
48-52. (134-171 mm.) Trinidad. L. Guppy, Esq.
53-64, (72 and 88 mm.) Tabatinga. Mus. Comp. Zool.
All agree in having 4: anal spines, and the numbers 5 and 6
are evidently quite exceptional.
Section 2 (Mesonauta).
Body increasing in depth to below the postcrior part of
the dorsal fin; scales of lateral line of the same size as those
above and below it; scales of thoracic region not very much
smaller than those on the side of the body. Mouth small,
moderately protractile; maxillary very slightly exposed
distally ; teeth of the outer series distinctly increasing in
size anteriorly. Dorsal XIV-XVI 10-12, the soft fin scaly
at the base. Anal VIII-IX 10-12. Caudal rounded.
The single species, from South America, shows relationships
with C. bimaculatum and C. autochthon.
2. Cichlosoma festivum.
Heros festivus, Heck. Ann. Mus. Wien, ii. 1840, p. 376.
Heros insignis, Heck. t. c. p. 378. ; Ne me
Chromys acora, Casteln, Anim. Am, Sud, Poiss. p. 17, pl. ix. fig. 1
(1855). . |
Mesonauta insignis, Giinth. Cat. Fish. iv. p. 800 (1862).
Acara (Heros) festiva, Steind. Sitzb. Ak. Wien, Ixxxi. 1875, p, 93.
Mesonauta festivus, Kigenm. & Bray, Ann, Ac. N. York, vii. 1le04,
p- 619.
Cichlasoma insigne, Pellegr. Mém. Soc. Zool. France, xvi. 1908, p. 221
(1904),
70 Mr. C. T. Regan on the Fishes of the
Depth of body 13-2 in the length, length of head 23-3.
Snout longer than postorbital part of head. Diameter of
eye 22-3} in the length of head, interorbital width 24-24.
Depth of preorbital equal to diameter of eye (adult) or less
(young). Maxillary extending to a little beyond the nostril ;
premaxillary processes not extending to above the eye ; jaws
equal anteriorly ; fold of the lower lip not continuous ; cheek
with 3 series of scales ; 5 or 6 gill-rakers on the lower part
of anterior arch. Sess 27-29 — 34-4 between lateral
line and anterior rays of soft dorsal. Dorsal XIV-XVI
10-12, commencing above or behind axil of pectoral, the
spines increasing in length to the last, which is 3—? the length
of head, the soft fin pomted, much produced in the adult.
Anal VIII-IX 10-12. Pectoral 3-4 the length of head,
extending to above the fifth anal spine; ventral, in the
adult, extending to the caudal. Caudal rounded. Candal
peduncle $ as long as deep. Brownish, with an oblique
blackish stripe from the snout, through the eye, to the tip of
soft dorsal; a dark ocellated spot on the upper 4 of base of
caudal; dorsal and caudal fins spotted.
Guiana; R. Amazon; R. de la Plata.
1. (90 mm.) 3 Zoological Society.
2. (102 mm.) Demerara. F. G. Beckford, Esq.
3. (147 mm.) Rio Negro. Mr. J. C. Antony.
4, (72 mm.) Descalvados, Matto Grosso, Dr. Ternetz.
5-7. (45-56 mm.) Carandasiiho, J Matto Grosso. Dr. A. Borelli.
8. (106 mm.) Tocantins, Brazil. Paris Museum.
Section 3.
Body ovate ; scales of lateral line of the same size as those
above and below it; scales of thoracic region not very much
smaller than those on the sides of the body. Mouth rather
small, moderately protractile; maxillary very slightly ex-
posed distally ; teeth of the outer series moderate, distinctly
enlarged anteriorly. Dorsal XV-XVII 9-11, the soft fin
scaly at the base. Anal VI-VIII 7-9. Pectoral extending
to above the origin of anal or a little beyond. Caudal
rounded.
The three species, from South America, show relationship
to C. bimaculatum in their general form and in the compara-
tively large size of the scales of the thoracic region.
3. Cichlosoma facetum.
Chromis facetus, Jenyns, Voy. ‘ Beagle,’ Fishes, p. 104 (1842).
Heros facetus, Giinth. Cat. Fish, iy. p. 200 (1862) ; Steind. Sitzb. Ak.
Wien, lx. 1869, p. 290, pl. i.
American Cichlid Genus Cichlosoma. 71
Heros Jenynsit, Steind. t. c. p. 292, pl. ii.
Heros acuroides, Hensel, Arch. f. Nat. xxxvi. 1870, p. 54.
Acara faceta, Steind. Sitzb. Ak. Wien, Ixx. 1874, p. 506.
Cichlasoma facetum, Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p. 217 (1904).
Depth of body 13-2! in the length, length of head 23-3.
Eye nearer to tip of snout than to the extremity of oper-
culum, its diameter 3-44 in the length of head, interorbital
width 22-24. Depth of preorbital equal to diameter of eye
(adult), or a little more than } diameter of eye (young).
Maxillary not or scarcely reaching the vertical from anterior
margin of eye; preemaxillary processes extending to above
anterior margin of eye; jaws equal anteriorly ; fold of the
lower lip not continuous ; cheek with 4—5 series of scales,
the lower 1 or 2 rows often deciduous in the adult; 7 or 8
gill-rakers on the lower part of the anterior arch. Scales
26-28 a 14 between lateral line and base of anterior rays
of soft dorsal. Dorsal XV—X VII 9-11, commencing above
the opercular cleft, the spines increasing in length to the
last, which is nearly half the length of head, the soft fin
pointed, when laid back extending to or beyond the middle
of caudal. Anal VI-VIII 7-9. Pectoral ?-+* the length
of head, extending to above the origin of anal; ventral
extending to origin of anal or beyond. Caudal rounded.
Caudal peduncle 2—} as long as deep. Olivaceous, with 6 or
7 dark cross-bars; a dark spot at the base of caudal; fins
dusky.
Rio de la Plata; Rio Grande do Sul.
1. (158 mm.) Buenos Ayres. Mr. White.
2-8,9-11,12-14. (50-122mm.) Rio Grande do Sul. Dr. H. von Ihering.
15-16, (182 &152mm.) Rio Parana. Mr, Salmin,
4. Cichlosoma autochthon.
Heros autochthon, Giinth. Cat. Fish. iv. p. 299 (1862); Kner, Novara
Fische, p. 265 (1869); Pellegr. Mém. Soc. Zool. France, xvi. 1908,
p. 235 (1904).
Acara (Heros) autochthon, Steind. Sitzb, Ak. Wien, Ixx. 1874, p. 502,
pl. i.
Depth of body about 2 in the length, length of head
about 24. Eye much nearer to tip of snout than to extre-
mity of operculum, its diameter 8-34 in the length of head,
interorbital width about 24. Depth of preorbital 3-3
diameter of eye. Maxillary not or scarcely extending to the
vertical from anterior margin of eye ; preemaxillary processes
extending to above anterior } of eye; lower jaw usually
slightly projecting ; fold of the lower lip continuous ; cheek
¥
72 Mr. C. T. Regan on the Fishes of the
with 3 series of scales ; 7 or 8 gill-rakers on the lower part
of anterior arch. Scales 26-28 2*, 1 between lateral line
and base of anterior part of soft dorsal. Dorsal XV-XVI
(XVII) 9-10, commencing above the pera cleft, the
spines increasing in length to the last, which is $ the length
of head or more; the soft fin pointed, when laid back ex-
tending to the extremity of caudal. Anal VI-VII (VIII)
7-8. Pectoral as long as the head, extending to above the
third or fourth anal spine; ventral extending beyond origin
of anal, sometimes nearly to the caudal. Caudal rounded.
rae ‘peduncle 1-2 as long as deep. Olivaceous, with 6
7 dark cross-bars; a dark spot at the base of the caudal ;
fing dusky.
Eastern Brazil.
1-3. (123-128 mm.) Brazil. Lord Stuart.
4, (113 mm.)
5-6. (82and 123mm.) Theresopolis. ' Dr, E, A. Gotdt.
7. (89 mm.) Porto Real, Prov. Rio M. Hardy du Dréneuf.
Janeiro.
5. Cichlosoma oblongum.
Chromys oblonga, Casteln, Anim, Am. Sud, Poiss. p. 14 (1855),
Heros oblongus, Giinth. Cat. Fish. iv. p. 299 (1862); Pellegr Mém.
Soe. Zool. France, xvi. 1903, p. 286 (1904).
Depth of body 2? in the length, length of head 22. Snout
as long as eye, the ‘diameter of which is 34 in the ‘length of
head, ‘interorbital width 31. Depth of “ preeorbital 2 the
diameter of eye. Maxillary extending nearly to below the
eye ; premaxillary processes extending to above anterior 1 of
eye; Jaws equal anteriorly ; fold of the lower lip subcon-
pee cheek with 14 series of scales, its lower part naked ;
eas rakers on the lower part of anterior arch. Scales
26 =, 15 between lateral line and base of anterior rays of
soft dorsal. Dorsal (XV) XVI 10, commencing above the
opercular cleft, the spines subequal from the fourth to the
twelfth, thence increasing to the last, which is $ the length
of head ; soft fin, when laid back, extending nearly to middle
of caudal. Anal VI9 (VII 7). Pectoral nearly as long as
the head, extending to above the fourth anal spine; ventral
extending a little beyond origin of anal. Caudal slightly
rounded. Caudal peduncle } as long as deep. Brownish,
with about 7 dark cross-bars ; fins blackish.
Southern and Eastern Brazil; Argentina.
J, (101 mm.) Argentina. Captain Vipan.
According to Pellegrin, the type of Chromys oblonga, from
.
American Cichlid Genus Cichlosoma. 73
Tocantins, is in bad condition, but the species is closely allied
to C. autochthon. As the specimen described above agrees
well enough with his description of H. oblongus, I provi-
sionally refer it to that species.
Section 4.
Body ovate; scales of lateral line of the same size as those
above and below it; scales of thoracic region considerably
smaller than those on the sides of the body. Mouth moderate
or rather large (maxillary extending to or nearly to below
the eye, snout longer than postorbital part of head), mode-
rately protractile ; maxillary very slightly exposed distally ;
teeth of the outer series moderate, well developed laterally,
distinctly increasing in size anteriorly. Dorsal XVI-X VIII
11-14, the soft fin scaly at the base. Anal VI-VIII 9-11.
Pectoral extending to above the origin of anal. Caudal
rounded.
The two species, from South America, are modified from
the type represented by C. facetum.
6. Cichlosoma temporale.
Heros temporalis, Ginth. Cat. Fish. iv. p. 287 (1862).
Acara (Heros) crassa, Steind. Sitzb. Ak. Wien, lxxi. 1875, p. 88, pl. v.
Heros Goeldit, Bouleng. Ann. & Mag. Nat. Hist, xx. 1897, p. 298;
Goeldi, Boll. Mus. Para, ii. 1898, pl.
Cichlasoma temporale, Pellegr. Mém. Soc. Zool. France, xvi. 1908,
p. 218 (1904).
Depth of body about 12 (13-2) in the length, length of
head 24-8. Snout longer than postorbital part of head.
Diameter of eye 34-3} in the length of head, interorbital
width 24-23. Depth of preorbital equal to diameter of eye.
Maxillary nearly extending to the vertical from anterior
margin of eye; premaxillary processes not extending to
above the eye ; Jaws equal anteriorly ; fold of the lower lip
not continuous; cheek with 4 or 5 series of scales; 7 or 8
gill-rakers on the lower part of anterior arch. Scales
30-382 —7 23 between lateral line and base of anterior part
of soft dorsal. Dorsal XVI-XVII 11-12, commencing above
the opercular cleft, the spines slightly increasing in length to
the last, which is nearly } the length of head; the soft fin
pointed, when laid back extending beyond the middle of
caudal. Anal VI-VII (VIII) 9 (10). Pectoral 4 the length
of head, extending to above the origin of anal. Ventral
extending to middle of anal or beyond. Caudal rounded.
Caudal peduncle } as long as deep. Brownish, with 3
74 Mr. C. T. Regan on the Fishes of the
blackish spots or blotches, one behind the eye, the second on
the middle of the side, and the third on the upper half of the
base of caudal; a blackish stripe connecting the spots. Fins
dusky.
Brazil ; Guiana.
1. (161 mm.) type of the species, Zoological Society.
2. (159mm.) typeof H. Goeldit, RioCounani,French Dr. E. A. Géldi.
Guiana.
7. Cichlosoma coryphenoides.
Heros coryphenoides, Weck. Ann. Mus. Wien, ii. 1840, p. 873; Ginth.
Cat. Fish. iv. p. 296 (1862).
Heros niger, Heck. t. c. p. 875.
peter coryphenoides, Steind. Sitzb. Ak. Wien, lxxi. 1875,
Grehlasamna coryphenoides, Pellegr. Mém, Soc. Zool. France, xvi. 1903,
p. 219 (1904).
Depth of body 2 in the length, length of head 28. Snout
longer than postorbital part of head. Diameter of eye 33-3?
in the length of head, interorbital width 22-22. Depth of
preorbital 2—} the diameter of eye. Maxillary extending to
the vertical from anterior margin of eye; pramaxillary pro-
cesses not extending to above the eye ; jaws equal anteriorly ;
fold of the lower lip shghtly interrupted ; cheek with 5 series
of scales; 7 or 8 gill-rakers on the lower part of anterior
arch. Scales 31-33 - 4 between lateral line and base of
anterior rays of soft dorsal. Dorsal XVI 12-13 (14), com-
mencing above the opercular cleft, the spines subequal from
the sixth to the thirteenth, thence increasing to the last,
which is more than 4 the length of head ; soft fin, when laid
back, extending beyond middle of caudal. Anal VI (VII)
9-10 (11). Pectoral 2 the length of head, extending to
above origin of anal; ventral extending to middle of anal.
Caudal rounded. Caudal peduncle $ as long as deep.
Brownish, with obscure darker cross-bars ; a blackish blotch
or vertical bar above the lateral line and below the eleventh
to thirteenth dorsal spines ; fins blackish.
Rio Negro.
1-2. (121 and 127 mm.) Rio Negro. Mr. J. C. Antony,
Section 5 (Archocentrus).
Body ovate; scales of lateral line of the same size as those
above and below it; scales of thoracic region considerably
smaller than those on the side of the body. Mouth rather
small, moderately protractile; maxillary not or slightly
American Cichlid Genus Cichlosoma. 75
exposed ; teeth of the outer series distinctly enlarged ante-
riorly. Dorsal XVII-XIX 7-10, the soft fin scaly at the
base. Anal VIII-XII 6-9. Pectoral extending beyond the
origin of anal. Caudal rounded or subtruncate.
The five species, from Guatemala and Nicaragua, are
probably derived from the type represented by C. facetum.
8. Cichlosoma nigrofasciatum.
Heros nigrofasciatus, Ginth. Trans. Zool. Soc. vi. 1869, p. 452, pl. Lxxiv.
fig. 3.
Cichlasoma nigrofasciatum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1525; Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 223
(1904).
Depth of body 2-24 in the length, length of head 23-3.
Snout considerably shorter than postorbital part of head.
Diameter of eye 22-34 in the length of head, interorbital
width 23-3}. Depth of preorbital }—4 the diameter of eye.
Maxillary extending to the vertical from anterior margin of
eye ; premaxillary processes extending to just above anterior
margin of eye ; jaws equal anteriorly ; fold of the lower lip
not continuous ; cheek with 4 or 5 series of scales; 7 or 8
gill-rakers on the lower part of anterior arch. Scales
28-31 545, 2 or 24 between lateral line and base of anterior
part of soft dorsal. Dorsal XVII-XVIII 8-9, commencing
above the opercular cleft, the spines subequal from the sixth
to the fourteenth, thence increasing to the last, which is 2
the length of head ; soft fin, when laid back, extending to
the middle of caudal. Anal VIII-X 6-8. Pectoral a little
shorter than the head, extending to above the third or fourth
anal spine; ventral extending beyond origin of anal,
Caudal rounded. Caudal peduncle 3-3 as long as deep.
Brownish, the body with 7 or 8 black cross-bars which extend
on to the dorsal fin ; a dark bar at the base of caudal.
Guatemala.
1-15, (43-82 mm.) types of the Lake Amatitlan. O. Salvin, Esq.
species.
9. Cichlosoma spilurum.
Heros spilurus, Giinth. Cat. Fish. iv. p. 289 (1862), and Trans. Zool.
Soe. vi. 1869, p. 451, pl. Lxxiii. fig. 1.
Cichlasoma spilurum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898,
p. 1520; Pellegr. Mém. Soc. Zool. France, xvi. 1908, p. 222 (1904).
Depth of body 2 in the length, length of head 3. Snout
as long as or longer than postorbital part of head. Diameter
76 Mr. C. T. Regan on the Fishes of the
of eye 3 in the length of head, interorbital width 22-3.
Depth of przorbital equal to the diameter of eye or less.
Maxillary not extending to below the eye; premaxillary
processes just extending to above anterior margin of eye ; jaws
equal anteriorly ; fold of the lower lip not continuous ; cheek
with 4 or 5 series of scales ; 6 or 7 gill-rakers on the lower
part of anterior arch. Scales 28-30 eis 2 or 24 between
lateral line and base of anterior part of soft dorsal. Dorsal
XVIII-XIX 9-10, commencing above the opercular cleft,
the spines subequal from the sixth to the sixteenth, thence
increasing to the last, which is 3-2 the length of head; soft
fin, when laid back, extending to middle of caudal. Anal
VIII-IX (X 7) 8. Pectoral as long as or longer than the
head, extending to above the third to fifth anal spines ;
ventral extending to middle of anal. Caudal rounded or
subtruncate. Caudal peduncle $-2 as long as deep. Body
with 7 dark cross-bars ; a dark spot or vertical bar on the
base of caudal ; vertical fins dusky, sometimes spotted.
Guatemala.
1-4. (75-90 mm.) types of the Lake Yzabal. O. Salvin, Esq.
species.
5. (92 mm.) Rio Motagua. O. Salvin, Esq.
10. Cichlosoma spinosissimum.
Heros (Cichlasoma) spinosissimus, Vaill. & Pellegr. Bull. Mus. Paris,
1902, p. 87.
Cichlasoma spinosissimum (part.), Pellegr. Mém, Soc. Zool. France, xvi.
1903, p. 224 (1904).
Depth of body 17 1n the length, length of head 3. Diameter
of eye 33 in the length of head. Maxillary not extending to
below the eye; fold of the lower lip not continuous; cheek
with 5 series of scales ; 6 gill-rakers on the lower part of
anterior arch. Scales 28 ;4.. Dorsal XVIII-XIX 7-8,
14-15"
the spines subequal from the fourth, the eighth } the length
of head. Anal XI-XII 7-8. Pectoral # the length of head.
Caudal rounded. Caudal peduncle } as long as deep.
Olivaceous, with small dark spots on the body and soft
vertical fins; a dark longitudinal band from the eye to the
middle of the side ; posterior part of body with 4 or 5 dark
cross-bars ; a dark spot on the base of caudal.
Rio Polochic, Guatemala.
The types (four specimens) measure from 78-100 mm.
American Cichlid Genus Cichlosoma. 77
Very similar to Herotilapia multispinosa, but especially
distinguished by the absolutely conical teeth.
11. Cichlosoma immaculatum.
Cichlasoma spinosissimum, var. immaculatum, Pellegr. Mém. Soe. Zool.
France, xvi. 1903, p. 225 (1904).
Very similar to the preceding species. Dorsal XVIII-
XIX 9-10. Anal IX 9. Scales 29... Cheek with 5or6
13-14"
series of scales. 8 gill-rakers on the lower part of anterior
arch. Uniformly greyish.
Rio Polochie.
Since two specimens agree in these characters, it seems to
me best to regard them as belonging to a distinct species.
12. Cichlosoma centrarchus.
Heros centrarchus, Gill & Bransford, Proc. Ac, Philad. 1877, p. 185.
Cichlasoma centrarchus, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1526; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 224
(1904).
Depth of body nearly 2 in the length, length of head 3.
Snout 2 as long as postorbital part of head. Diameter of
eye 33 in the length of head, interorbital width 24. Depth
of preorbital 3 the diameter of eye. Maxillary not ex-
tending to below the eye; preemaxillary processes extending
to above the anterior margin of eye ; jaws equal anteriorly ;
fold of the lower lip continuous (? or not) ; cheek with 4 or
5 series of scales ; 15 gill-rakers on the lower part of the
anterior arch. Scales 29 = 2 between lateral line and base
of anterior rays of soft dorsal. Dorsal XVI (8) 9, com-
mencing above the opercular cleft, the spines subequal from
the fifth to the fourteenth, the last more than } the length
of head. Anal XI 8(X 9). Pectoral as long as the head,
extending to above the fifth anal spine; ventral extending
to middle of anal. Caudal subtruncate. Caudal peduncle 4
as long as deep. Olivaceous, with 7 or 8 dark brown cross-
bars ; a dark spot at the base of caudal; fins dusky.
Lake of Nicaragua.
1. (140 mm.) one of the types L, Nicaragua. Smithsonian Inst.
of the species.
[To be continued. }
78 Canon A. M. Norman—Revised Nomenclature of
VIII.—Revised Nomenclature of the Species described in Bate
and Westwood’s ‘ British Sessile-eyed Crustacea. By
Canon A. M. Norman, M.A., D.C.L., LL.D., F.R.S.
THE work of Bate and Westwood from the first presented
great difficulties to the student of the Amphipoda and [sopoda
in consequence of the unsatisfactory figures and inadequate
descriptions of many of the species. Moreover, with in-
creasing knowledge of these Crustacea extensive changes in
nomenclature have become necessary. Hence there lies a
heavy stumbling-block in the way of any carcinologist who
commences the study of the species represented in our fauna,
and only an adequate knowledge of all that during recent
years has been written on the subject can at the present time
enable the species described in this work to be correctly
named.
The object of the present paper is to remove, at any rate
partially, this difficulty and bring the nomenclature of Bate
and Westwood into juxtaposition with the names which are
now employed for the several species. I have, of course,
availed myself of all that has been written on the subject by
carcinologists, such as the Rev. T. R. R. Stebbing, Mr. A. O.
Walker, and others in our own country, and by Professor
G. O. Sars and M. Chevreux and others on the Continent, and
I trust that I shall moreover be able to throw fresh light on
the relationship of certain species which have hitherto not
received full elucidation.
There have been very large additions to our knowledge of
the species of these two orders which inhabit our seas during
recent years, but with such additions these notes have nothing
to do.
One new genus is here introduced—Coremapus (kopnpa, a
brush, and zrovs, a foot). Allied to Microdeutopus ; first
gnathopods the largest, complexly subchelate in male and
simply subchelate in female as in that genus, but the second
gnathopods in both sexes narrow, scarcely subchelate,
terminating in a minute nail, the limb forming a brush, the
meros, carpus, and manus being densely setose, especially the
meros, which is much produced over the carpus. ‘Type,
Coremapus (Microdeutopus) versiculatus, Bate.
In the earlier part of this paper the species are taken in the
order in which they are arranged in our authors’ work; the
nomenclature, where necessary, is corrected and a number
prefixed.
In the latter part the species with their corrected names
Crustacea described by Bate and Westwood. 79
are arranged as in the existing leading works upon the
subject, those of Professor G. O. Sars, and the affixed number
is that under which the species occurs in the previously given
list.
If
The Species described in Bate and Westwood, with correction
OO bt
uN
lie
18
of names where such correction is required.
Vout. I.
. Talitrus locusta (Linné), p. 16,= Talitrus locusta (Pallas).
. Orchestia littorea (Montagu), p. 27.
. —— mediterranea (Costa), p. 31.
. ~— Deshayesii, Audouin, p. 36,=Orchestoidea Deshayesii
(Audouin) (Norman, Ann. & Mag. Nat. Hist. ser. 7, vol. v.
p. 189).
. Allorchestes Nilssonii (Rathke), p. 40,= Hyale Nilssoni (Rathke).
imbricatus, Bate, p. 43,= Hyale Lubbockiana (Bate), 3.
. Nicea Lubbockiana, Bate, p. 47,= Hyale Lubbockiana (Bate), 2 .
. Montagua monoculoides (Montagu), p. 54,=Stenothoe monocu-
loides (Montagu).
marina, Bate, p. 58,=Stenothoe marina (Bate).
. —— Alderi, Bate, p. 61,= Metopa Alderi (Bate), 9.
pollexiana, Bate, p. 64,= Metopa norvegica (Lilljeborg)
(Norman, Ann. & Mag. Nat. Hist. ser. 7, vol. vi. p. 41).
. Danaia dubia, Bate, p. 68,= Cressa dubia (Bate).
. Lysianassa Coste, Bate and Westwood (nee Milne-Edwards),
p. 74,=Lysianax septentrionalis, Della Valle.
—— Audouiniana, Bate, p. 79,=Perrierella Audouiniana
(Bate).
atlantica, Bate and Westwood (? Milne-Edwards), p. 82,=
Ambasia Danielssent, Boeck (Norman, Ann. & Mag. Nat.
Hist. ser. 7, vol. v. 1900, p. 144).
longicornis, Bate and Westwood (nec Lucas), p. 85. The
entire figure and the urosome probably taken from Orcho-
mene humilis (Costa), gd, the other dissections from Lysianax
septentrionalis, Della Valle, 6 (see Walker, Ann. & Mag.
Nat. Hist., Feb. 1892; and Norman, ibid. ser. 7, vol. v.
p. 142).
Anonyx longicornis, Bate, p. 91,=Lepidepecreum longicorne
(Bate), 3.
Edwardsi, Bate (nec Kroyer), p. 94,=Orchomene humilis
(A. Costa) (see Norman, Ann. & Mag. Nat. Hist. ser. 7,
vol, v. 1900, p. 202).
80
19.
20.
aL.
22,
23.
Canon A. M. Norman—Revised Nomenclature of
Anonyx obesus, Bate, p. 98,= Acidostoma obesum (Bate).
denticulatus, Bate, p. 101,=Hippomedon denticulatus
(Bate).
Holbolli, Bate and Westwood (nec Kroyer), p. 104,=
Haplonyx cicada (Fabricius).
minutus, Bate and Westwood (nec Kroyer), p. 108. Per-
haps a young male of Orchomene humilis (Norman, Ann. &
Mag. Nat. Hist. ser. 7, vol. v. p. 204).
plautus, Bate and Westwood (nec Kroyer), p. 111, =
Menigrates obtusifrons, Boeck.
longipes, Bate, p. 118,= Tryphosites longipes (Bate), 2.
ampulla, Bate and Westwood (nec Kroyer), p. 116,=
Tryphosites longipes (Bate), 3.
. Callisoma crenata, Bate, p. 120,=Callisoma Hoper, Costa.
. Ampelisca Gaimardi, Bate and Westwood (nec Kroyer), p. 127,
= Ampelisea typica (Bate), 3.
Belliana (Bate), p. 135,= Ampelisca brevicornis (A. Costa).
. Phoxus simplex, Bate, p. 140,==Phowxocephalus simplex (Bate).
Holbotli, Kroyer, p. 143, = Phowocephalus Holbélli (Kroyer).
. —— plumosus, Bate and Westwood (nec Kroyer), p. 146,=
Harpinia neglecta, G. O. Sars.
. Graia imbricata, Bate, p. 152,= Amathilla angulosa (Rathke),
young. I do not, however, consider A. angulosa to be a
distinct species from A. homari (Fabricius).
. Westwoodilla cecula, Bate, p. 155.
hyalina, Bate, p. 158.
Both these Westwoodille, which are the same species, were
so erroneously described that even Bate himself could not
recognize his own genus or species, and the very next species
is called Gdiceros parvimanus. The name to be used for
the species should be therefore, I think, that by which the
species was properly described and by which it could be
recognized, and the species should be called Halimedon
parvimanus (Bate and Westwood).
. Ediceros parvimanus, Bate and Westwood, p. 161, = Halimedon
parvimanus (Bate), =H. Miillerr, Boeck.
. Monoculodes carinatus, Bate, p. 165.
Stimpsoni, Bate, p. 168, = Monoculodes carinatus, Bate.
. Kroyera arenaria, Bate, p. 173,= Pontocrates arenarius (Bate).
altamarina, Bate and Westwood, p. 177,=Pontocrates
altamarinus (B. & W.).
. Amphilochus manudens, Bate, p. 180.
. Darwinia compressa, Bate, p. 184,=Laphystius sturionis,
Kroyer
42
Crustacea described by Bate and Westwood. 81
. Suleator arenarius, Bate, p. 189, = Haustorius arenarius
(Slabber).
43. Urothoe Bairdi, Bate, p. 193,= 3 of U. marinus.
44, -—— marinus, Bate, p. 195, 9.
45. brevicornis, Bate, p. 198, 2.
46. elegans, Bate, p. 200, ¢.
47.
48.
49,
The species of Urothoe require further working out.
Liljeborgia pallida, Bate, p. 203, = Lilljeborgia pallida, Bate.
shetlandica, Bate and Westwood, p. 206,= Cheirocratus
Sundevalli (Rathke), ¢.
Pheedra antiqua, Bate, p. 209. Unknown; might be an
Apherusa, if it were not that it has a secondary appendage
to the antenne.
50. Kinahani, Bate, p. 211,=Lilljeborgia Kinahani (Bate).
51. Isea Montagu, H. Milne-Edwards, p. 215.
52.
53.
54,
55.
56.
Iphimedia obesa, Rathke, p. 219.
eblane, Bate, p. 221.
Otus carinatus, Bate, p. 224, Odius carinatus (Bate).
Pereionotus testudo (Montagu), p. 228.
Acanthonotus Oweniit, Bate, p. 232, = Epimeria cornigera
(Fabricius).
57. Dewvamine spinosa (Montagu), p. 237.
58. tenuicornis, Bate and Westwood (nec Rathke), p. 240,=
Dexamine thea, Boeck.
59. vedlomensis, Bate and Westwood, p. 242,=Taratylus
vedlomensis (B. & W.).
60. Atylus Swammerdamiu (H. Milne-Edwards), p. 246,= Paratylus
Swammerdanit (H. Milne-Edwards).
61, gibbosus, Bate, p. 248, = Tritwta gibbosa (Bate).
62. —— bispinosus, Bate, p. 250,= Apherusa bispinosa (Bate).
68.
69.
. Pherusa bicuspis, Bate (nec Kroyer), p. 253,=Apherusa cirrus
(Bate), ¢ ,=Apherusa borealis, Boeck.
. — fucicola, Leach, p. 255,=Gamimarella brevicaudata,
H. Milne-Edwards, 2 (Walker, Ann. & Mag. Nat. Hist.
ser. 6, vol. vii. p. 418).
. Calliope leviuscula (Kroyer), p. 259,= Callhopius leviusculus
(Kroyer). é,
Ossiani, Bate, p. 261,=Parapleustes latipes (M. Sars).
. — Fingalli, Bate and Westwood, p. 263,= Parapleustes latipes
(M. Sars), adult.
— grandoculis, Bate, p. 265, = Calhiopius leviusculus
(Kroyer), ¢.
Eusirus helvetiv, Bate, p. 267,=Lusirus longipes, Boeck,
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 6
Canon A. M. Norman—Revised Nomenclature of
. Leucothoe articulosa (Montagu), p. 271,=Leucothoe spincarpa
(Abildgaard).
furina (Savigny), p. 274, =? Leucothoe furina (Savigny).
. Gossea microdeutopa, Bate, p. 277. ‘“ Appears to be Apherusa
Jurinii, M.-Edw.” (Walker, Ann. & Mag. Nat. Hist. ser. 6,
vol. xv. 1895, p. 469).
. Aora gracilis, Bate, p. 280.
. Stimpsonia chelifera, Bate, p. 285, = Stimpsonella chelifera (Bate).
. Mierodeutopus gryllotalpa, Bate (nec Costa), p. 289,= Micro-
deutopus damnoniensis, Bate.
76. Websteri, Bate, p. 291,=Lembos Websteri, Bate.
be anomalus, Rathke, 9, p. 293. @ of M. damnoniensis, if
not of M. anomalus.
78. versiculatus, Bate, p. 295, = Coremapus versiculatus
(Bate), 2.
79. Protomedeia hirsutimana, Bate, p. 298,—Leptocheirus pilosus,
Zaddach.
80. Whitei, Bate, p.300,= Cheirocratus Sundevallhi (Rathke), ° .
81. Bathyporeia pilosa, Bate and Westwood (nec Lindstrom), p. 304,
= Bathyporeia Guilliamsona (Bate),=B. norvegica, Sars.
82. —— Robertsoni, Bate, p. 307, 3.
83. pelagica, Bate, p. 309, 3.
84, Niphargus aquilex, Schiddte, p. 315,=Niphargus subterraneus
(Leach).
85 fontanus, Bate, p. 319.
86. Kochianus, Bate, p. 323.
87. Crangonyx subterraneus, Bate, p. 327.
88. Gammarella brevicaudata (H. Milne-Edwards), p. 330, ¢.
89. Normanni, Bate and Westwood, p. 333,= Gammarella
brevicaudata (H. Milne-Edwards), 2.
90. Melita palmata (Montagu), p. 337.
91. obtusata (Montagu), p. 341, ¢.
92. —— proawima, Bate, p. 344,= Melita obtusata (Montagu), ¢.
93. gladiosa, Bate, p. 346.
94. Mera grossimana (Montagu), p. 350.
98.
. Eurystheus erythrophthalmus (Lilljeborg), p.354, = Gammaropsis
maculatus (Johnston), g.
bispinimanus, Bate, p. 357,=Gammaropsis maculatus
(Johnston), 2.
. Amathilla Sabini (Leach), p. 861,=Amathilla homari (Fabri-
cius), var. Amathilla angulosa (Rathke).
Gammarus marinus, Leach, p. 370.
campylops, Leach, p. 375.
Crustacea described by Bate and Westwood. 83
100. Gammarus locusta (Linné), p. 378.
LOL. tenuimanus, Bate, p. 384,=? Mera Batei, Norman (see
Walker, Ann. & Mag. Nat. Hist. ser. 6, vol. xv. 1895, p. 471).
102. Edwardsii, Bate, p. 386 (not as yet recognized by any
other author),
103. pulex (De Geer), p. 388.
104, dubius, Johnston, p. 397, =?
105. maculatus, Johnston, p. 899,=Gammaropsis maculatus
(Johnston).
106. Megamera semiserrata, Bate, p. 401,=Ceradocus semiserratus
(Bate).
107. longimanus (Leach, MS.), p. 403,=Mera othonis (H.
Milne-Edwards), 3.
108. othonis (H. Milne-Edwards), p. 405,= Mera othonis (H.
Milne-Edwards), 9.
109. —— Alderi, Bate, p. 407,= Melita obtusata (Montagu), 9.
brevicaudata, Bate, p. 409, = Elasmopus rapaa, Costa.
111. Hiscladus longicaudatus, Bate and Westwood, p. 412,= Photis
longicaudata (Bate and Westwood).
112. Amphithoe rubricata (Montagu), p. 418.
littorina, Bate, p. 422,=Amphithoe rubricata (Mon-
tagu), 2.
114. —— albomaculata, Bate and Westwood (nec Kroyer), p. 426.
The specimen figured is in my collection. It is Podoceropsis
excavata, Bate, 2. The long sete of the antenne and the
first gnathopods as drawn agree with Podoceropsis, but the
specimen shows nothing of the strong upturned third segment
of metasome or the Amphithoe-like last uropod of Bate and
Westwood’s figure.
gammaroides, Bate, p. 427,= Pleonexes gammarordes,
Bate, 3.
116. Sunamphithoe hamulus, Bate, p. 430,= Pleonexes gammaroides,
Bate, 2.
117. -— conformata, Bate, p. 432,=Swnamphithoe pelagica (H.
Milne-Edwards).
118. Podocerus pulchellus (Leach), p. 4386,= Bruzeliella * falcata
(Montagu), ¢.
* T have examined the types of the genera Podocerus and Jassa in the
British Museum, and fully confirm what Messrs. Stebbing and Walker
have written upon them, that Podocerus variegatus, Leach (as interpreted
by Milne-Edwards’s figure in ‘Régne Animal de Cuvier,’ but not
P. variegatus, B, & W.), is Platophium Darwinii, and that Jassa pelagica,
Leach (but not of B. & W.), is the same as Podocerus capillatus, B. & W.
Bruzelius long ago restricted the use of Jassa to the species last men-
tioned, which has only a rudimentary appendage (scarcely visible with a
high power) to the antennules. This use must be maintained, and not
Parajassa as suggested by Stebbing. I have therefore found it necessary
to institute a genus Bruzeliella, with B. falcata, Montagu, as its type.
6*
115.
84
119;
120.
Canon A. M. Norman—Revised Nomenclature of
Podocerus variegatus, Bate and Westwood (nec Leach), p. 439, =
Bruzeliella pusilla (Sars), Q .
capillatus, Rathke, p. 442,=Jassa pelagica, Leach (nec
Buc Ww):
falcatus (Montagu), p. 445,= Bruzehella falcata (Mon-
tagu), ¢ junior.
pelagicus (Bate and Westwood) (nec Leach), p. 447,=
Bruzeliella falcata (Montagu), 2.
ocius, Bate, p. 450,= Bruzeliella ocia (Bate).
. Cerapus abditus, Templeton, p. 455,=LHrichthonius abditus
Templeton, ¢.
difformis (H. Milne-Edwards), p. 457,= EHrichthonius
difformis, H. Milne-Edwards, ¢.
. Dercothoe punctatus, H. Milne-Edwards, p. 461,= Hrichthonius
difformis, H. Milne-Edwards, 9.
127. Siphoneecetes typicus, Bate and Westwood (nec Kroyer), p. 465,
Siphoneecetes Whiter (Gosse), 3.
128. —— Whitei, Gosse, p. 467,=Siphonacetes Whitei (Gosse), 9,
= 0. Colletti, Boeck.
129. crassicornis, Bate, p. 469,= Cerapus crassicornis (Bate).
130. Nenia tuberulosa, Bate, p.472,= Podoceropsis Sophie, Boeck, 3.
131. rimapalnata, Bate, p. 474,= Podoceropsis excavata (Bate),
oe
132 excavata, Bate, p. 476, = Podoceropsis excavata (Bate), 9 -
133 undata, Bate, p. 477,= Podoceropsis Sophie, Boeck, 9.
134
. Cyrtophium Darwinti, Bate, p. 481,—=Platophium variegatum
(Leach) *.
Cratippus tenuipes, Bate, p. 485,= Colomastix pusilla, Grube.
. Dryope irrorata, Bate, p. 488,= Unciola irrorata (Bate), 2.
crenatipalma, Bate, p. 490,= Unciola irrorata (Bate), 3g.
. Corophium longicorne, Latreille, p. 493,= Corophium volutator
(Pallas), 3.
Bonelli, Bate and Westwood (nec Milne-Edwards),
p- 497,= Corophium affine, Bruzelius, °.
crassicorne, Bruzelius, p. 499, ¢.
. Chelura terebrans, Philippi, p. 503.
Vot. II.
. Listrigonus exulans, Kroyer, p. 5,= Hyperia galba (Montagu),
3 junior.
* I think that, considering the inadequate description of the genus
Podocerus and its erroneous use for nearly one hundred years, the name
ought to be excluded from an altered use.
143.
144.
145.
146.
147.
148.
149.
150,
151.
152.
153.
154.
155.
156.
157.
158.
159.
160.
161.
162,
163.
164.
165.
166.
167.
168.
169.
170.
Wil.
172.
Crustacea described by Bate and Westwood. 85
Listrigonus Kinahant, Bate, p. 8,=Hyperia galba (Montagu),
do mature.
Hyperia galba (Montagu), p. 12, 2.
oblivia, Bate and Westwood (nec Kroyer), p. 16,=
Euthemisto gracilipes (Norman),
Phronima sedentaria (Forskaal), p, 23.
Dulichia porrecta, Bate, p. 31, 3.
falcata, Bate, p. 33, 3.
Proto pedata, Abildgaard, p. 38,= Phtisica marina, Slabber, 9? .
Goodsiri, Bate, p. 42,= Phtisica marina, Slabber, ¢.
Protella phasma (Montagu), p. 45,=Pseudoprotella phasma
(Montagu).
Caprella linearis (Linné), p. 52.
lobata (Miiller), p. 57,=Caprella linearis (Linné), 3.
acutifrons (Latreille), p. 60.
hystrix (Kroyer), Bate and Westwood, p. 63,= Cuprella
septentrionalis, Kroyer, var. nodifera, Mayer.
acanthifera, Leach, p. 65.
— tuberculata, Guerin, p. 68.
equilibra, Say, p. 71.
typica, Kroyer, p. 75,= Periambus typicus (Kroyer).
Cyamus ceti, Martens, p. 85,=Cyamus mysticeti, Liitken.
ovalis, Roussel de Vauzime, p. 91, not British.
gracilis, Roussel de Vauzéme, p. 94, not British.
Thompsoni, Gosse, p. 96,=Platycyamus. Thompsoni
(Gosse).
IsopopDa.
Tanais vittatus (Rathke), p. 125,= Tanais Cavolini, H. Milne-
Edwards.
Dulongit (? Audouin), p. 129.
Leptochelia Edwardsu (Kroyer), p. 134,= Leptochelia Savignit
(Kroyer).
Paratanais forcipatus, Bate and Westwood (nec Lilljeborg),
p. 138,= Puratanais Batei, G. O. Sars.
— rigidus, Bate and Westwood, p. 141,= Leptognathia
rigida (Bate and Westwood).
Apseudes talpa (Montagu), p. 148.
Latrellii, H. Milne-Edwards, p. 153 (vide Norman, Ann,
& Mag. Nat. Hist. ser. 7, vol. iii. p. 327).
Anthura gracilis (Montagu), p. 160, °.
Paranthura Costana, Bate and Westwood, p. 165,= Paranthura
mgropunctata (Lucas), 2.
86
173.
Canon A. M. Norman—Revised Nomenclature of
Anceus mavillaris (Montagu), p. 187,=Gnathia mawvillaris
(Montagu).
Edwardii, Bate, p. 201,= Gnathia ovyurea (Lilljeborg).
Halidaii, Bate and Westwood, p. 203,= Gnathia formica
(Hesse).
. Bopyrus squillarum, Latreille, p. 218.
. Gyge galathee, Bate and Westwood, p. 225,= Gyge branchialis,
Cornalia and Panceri. The specimen which is described
and illustrated at p. 225 was procured by me from Upagebia
stellata, and not, as erroneously recorded by the authors,
from a Galathea.
hippolytes, Kroyer, p. 230,= Bopyroides hippolytes
(Kroyer).
. Phrywxus abdominalis (Kroyer), p. 234.
180. fusticaudatus, Bate and Westwood, p. 238,= Athelges
paguri, Rathke, young.
181. paguri, Rathke, p. 240,= Athelges paguri (Rathke).
182. —— Hyndmanni, Bate and Westwood, p. 243,= Pseudione
Hyndmanni (Bate and Westwood).
183. longibranchiatus, Bate and Westwood, p. 246,= Pleuro-
crypta longibranchiata (Bate and Westwood).
184 galathee, Bate and Westwood (nec Hesse), p. 249,=
Pseudione confusa (Norman).
185. one thoracica (Montagu), p. 255.
199.
200.
. Cryptothiria pygmea (H, Rathke), p. 261,=Liriopsis pygmea
(H. Rathke).
balani (Bate), p. 267,= Cryptothir balani (Bate).
. Adiga bearinata, Leach, p. 278,=4%ga rosacea, Risso. A
doubtful British species (see Norman, Ann. & Mag. Nat. Hist.
ser. 7, vol. xiv. 1904, p. 433).
tridens, Leach, p. 281.
psora (Pennant), p. 283.
monophthalma, Johnston, p. 286.
. Rocinela danmoniensis, Leach, p. 291.
. Cirolana Cranchii, Leach, p. 296.
spinipes, Bate and Westwood, p. 299,= Cirolana borealis,
Lilljeborg.
. Conilera cylindracea (Montagu), p. 304.
. Eurydice pulchra, Leach, p. 310.
. Jera albifrons (Montagu), p.317,=Jera marina (Fabricius).
Nordmanni (Rathke), p. 320.
Munna Kroyer, Goodsir, p. 326, 3.
Whiteana, Bate and Westwood, p. 329, = Munna Kroyeri,
Goodsir, 9.
Crustacea described by Bate and Westwood. 87
. Leptaspidia brevipes, Bate and Westwood, p. 333. I think
that this must be Paramunna bilobata, Sars, and should have
precedence of that name.
. Janira maculosa, Leach, p. 338.
. Asellus aquaticus (Linné), p. 348.
. Limnoria lignorwm (Rathke), p. 351.
. Arcturus longicornis (Sowerby), p. 365,= Astacilla lonyicornis
(Sowerby), 2.
intermedius (H. Goodsir), p. 371, = Astacilla intermedia
(H. Goodsir).
gracilis (H. Goodsir), p. 373,=Astacilla longicornis
(Sowerby), ¢.
. Idotea tricuspidata, Desmarest, p. 379,=Jdotea balthica
(Pallas).
pelagica, Leach, p. 384.
emarginata (Fabricius), p. 386.
. —— linearis (Pennant), p. 388.
parallela, Bate and Westwood, p. 391,=Zenobianu
prismatica (Risso).
acuminata, Leach, p. 394,=Stenosoma acuminatum,
Leach.
appendiculata (Risso), p. 396,=Stenosoma lanciferwin
(Leach).
. Spheroma serratum (Fabricius), p. 405.
. — rugicauda, Leach, p. 408.
. — Hookeri, Leach, p. 410.
. —— curtum, Leach, p. 412,= Cymodoce truncata (Montagu),
o
Prideauxianum, p. 415, = Cymodoce truncata (Mon-
tagu), 2.
. Dynamene rubra, Leach, p. 419, = Nesa bidentata (Adams), 2.
viridis, Leach, p. 421,= Nesa bidentata (Adams), 9.
. —— Montagui, Leach, p. 423,=?? young of Nese bidentata
(Adams), ¢.
. Cymodocea truncata, Leach, p. 426, 3,= Cymodoce truncata,
Leach, 3.
. — emarginata, Leach, p. 428,=C. truncata, Leach, ¢
variety.
. Nesa bidentata (Adams), p. 451, ¢.
. Campecopea hirsuta (Montagu), p. 4384, 3.
Cranchu, Leach, p. 436,= Campecopea hirsuta (Mon-
tagu), 2.
. Ligia oceanica (Linne), p. 444.
88
229.
230.
231.
232.
233.
234.
235.
236.
237.
238.
239.
240.
241.
Canon A. M. Norman— Revised Nomenclature of
Philoscia muscorum (Scopoli), p. 450.
Couchii, Kinahan, p. 452.
Philourgria riparia (Koch), p. 456,= Trichoniscus pusillus,
Brandt.
vwida (Koch), p. 458,= Trichoniscus vividus (Koch).
rosea (Koch), p. 460,= Trichoniscus roseus (Koch).
Platyarthrus Hoffmannseggii, Brandt, p. 464.
Oniscus asellus, Linné, p. 468.
fossor, Koch, p. 471,= Oniscus asellus, Linné.
Porcellio scaber, Latreille, p. 475.
dilatatus, Brandt, p. 478.
putus, Brandt, p. 480.
levis, Latreille, p. 483.
armadilloides, Lereboullet, p. 485,= Cylisticus conveaus
(De Geer).
pruinosus, Brandt, p. 487,=Metoponorthus pruinosus
(Brandt).
cingendus, Kinahan, p. 489,—= Metoponorthus cingendus
(Kinahan),
. Armadillo vulgaris, Latreille, p. 492,= Armadillidium vulgare
(Latreille).
APPENDIX.
. Orchestia brevidigitata, Bate and Westwood, p. 498, = Orchestia
littorea (Montagu), ¢ young.
. Montagua clypeata, Bate and Westwood (nec Kroyer), p. 499,
= Metopa abscisa, Norman.
norvegica, Bate and Westwood (nec Lilljeborg), p. 500,
= Metopa Alderi (Bate), ¢.
. Opis leptochela, Bate and Westwood, p. 501,= Zuonyzx chelatus,
Norman.
quadrumana, Bate and Westwood, p. 503, = Normanion
quadrimanus (Bate and Westwood),
. Ampelisca laevigata, Bate and Westwood (nec Lilljeborg),
p. 904,= Ampelisca tenuicornis, Lilljeborg.
. Haploops tubicola, Lilljeborg, p. 505.
. Monoculodes longimanus, Bate and Westwood, p. 507,= Peri-
oculodes longimanus (Bate and Westwood).
. Kroyera brevicarpa, Bate and Westwood, p. 508,=Synche-
lidium haplocheles (Grube).
. Lepidepecreum carinatum, Bate and Westwood, p. 509,=
Lepidepecreum longicorne (Bate and Westwood), ?.
. Nicippe tumida, Bruzelius, p. 511.
Crustacea described by Bate and Westwood. 89
256. Cheirocratus mantis, Norman, p. 513,= Chirocratus assimilis
(Lilljjeborg), 3.
257. Megamera multidentata, Bate and Westwood, p. 515,= Mera
Batei, Norman.
258. Unciola leucopes, Bate and Westwood (nec Kroyer), p. 517,=
Unciola planipes, Norman.
259. Hyperia tauriformis, Bate and Westwood, p. 519, = Hyperoche
tauriformis (Bate and Westwood).
260 prehensilis, Bate and Westwood, p. 520,=? Hyperia
tauriformis, Bate and Westwood, young (vide Norman, Aun.
& Mag. Nat. Hist. ser. 7, vol. v. 1900, p. 130).
261. cyanece (Sabine), p. 521,=??
262. Themisto crassicornis, Kroyer, p. 522,=LZuthemisto lbellula
(Mandt).
263. Vibilia borealis, Bate and Westwood, p. 524.
264. Tessarops hastata, Norman, p. 530,= Tiron acanthurus, Lillje-
borg.
265. Eriopis elongatus, Bruzelius, p. 530, = Hriopisa elongata
(Bruzelius). ~
266. Mera Loveni, Bruzelius, p. 530.
267. —— Batei, Norman, p. 530.
268. Helleria coalita, Norman, p. 530,= Guernea coalita (Norman).
269. Microprotopus maculatus, Norman, p. 530.
270. Cirolana truncata, Norman, p. 530, = Hurydice truncata
(Norman).
271. Anilocra mediterranea, Leach, p. 530,=Anilocra physodes
(Linné).
1S
Species described by Bate and Westwood in accordance with
present Nomenclature, and arranged after the Monographs
of Professor G. O. Sars *.
I, AMPHIPODA.
Tribe I. Hypuripea.
Hyperia galba (Montagu), 142 and 143 ¢, 144 9.
cyanee, Sabine, 261 ?
Hyperoche tauriformis (B. & W.), 259 (and ? 260),
Juthemisto libellula (Mandt), 262.
gracilipes (Norman), 145,
* The numbers appended to the names are those prefixed to the species
in the preceding list of Bate and Westwood’s species.
90 Canon A. M. Norman—Revised Nomenclature of
Phronima sedentarva (Forskaal), 146.
Vibilia borealis, B. & W., 263.
Tribe Il. GamMaripra.
Talitrus locusta (Pallas), 1.
Orchesteu littorea (Montagu), 2, 245.
mediterranea (Costa), 3.
Orchestoidea Deshayesii (Audouin), 4.
Hyale Nilssoni (Rathke), 5.
Lubbockiana (Bate),6 5,7 2.
Normanion quadrimanus (B. & W.), 249.
Acidostoma obesum (Bate), 19.
Lysianax septentrionalis, Della Valle, 18 and part of 16.
Ambasia Danielsseni, Boeck, 15?
Perrierella Audouiniana (Bate), 14.
Callisoma Hopei, Costa, 26.
Hippomedon denticulatus, Bate, 20.
Orchomene humilis (Costa), 18 and part of 16, and ? 22 junior.
Try phosites longipes (Bate), 24 9,25 ¢.
Haplonyx cicada (Fabricius), 21.
Menigrates obtusifrons, Boeck, 23.
Lepidepecrewm longrcorne (Bate), 17 3, 254 9.
Euonyx chelatus, Norman, 248.
Bathyporera Guilliamsonia (Bate), 81 2.
Robertsoni, Bate, 82 6.
pelagica, Bate, 83 S.
Haustorius arenarius (Slabber), 42.
Urothoe marinus, Bate, 44 9,43 ¢.
brevicornis, Bate, 45 2.
elegans, Bate, 46 S.
Phowocephalus Holbolli (Kroyer), 30.
simplex (Bate), 29.
Harpinia neglecta, G. O. Sars, 31.
Aimpelisca typica (Bate), 27.
—— brevicornis (Costa), 28.
tenuicornis, Lilljeborg, 250.
Huaploops tubicola, Lilljeborg, 251.
Amplhilochus manudens, Bate, 40.
Stenothoe marina (Bate), 9.
monoculoides (Montagu), 8.
Crustacea described by Bate and Westwood.
Metopa Alderi (Bate), 10 9, 247 o.
norvegica (Lilljeborg), 11.
abscisa, Norman, 246.
Cressa dubia (Bate), 12.
Leucothoe spinicarpa (Miller), 70.
——? furina (Savigny), 71.
Monoculodes carinatus, Bate, 36, 37.
Perioculodes longimanus (B. & W.), 252.
Pontocrates arenarius (Bate), 38.
altamarinus (Bate), 39.
Synchelidium haplocheles (Grube), 253.
Halimedon parvimanus (B. & W.), 33, 34, 35.
Parapleustes latipes (M. Sars), 66, 67.
Epimeria cornigera (Fabricius), 56.
Iphimedia obesa, Rathke, 52.
eblane, Bate, 53.
Pereionotus testudo (Montagu), 55.
Odius carinatus (Bate), 54.
Laphystius sturionis, Kroyer, 41.
Tiron acanthurus, Lilljeborg, 264.
Nicippe tumida, Bruzelius, 255.
Eusirus longipes, Boeck, 69.
Apherusa bispinosa (Bate), 62.
cirrus, Bate, 63 3.
Jurinii (H. M.-Edw.) ?, 72.
Calliopius laeviusculus (Kroyer), 65, 68 ¢.
Paratylus Swammerdamii (H. M.-Edw.), 60.
vedlomensis (B. & W.), 59.
Dexamine spinosa (Montagu), 57.
thea, Boeck, 5%.
Triteta gibbosa (Bate), 61 2.
Guernea coalita (Norman), 268.
Amathilla homari (Fabricius), 32 young, 97.
Gammarus marinus, Leach, 98.
campylops, Leach, 99.
locusta (Linné), 100,
—— pulew, De Geer, 103.
— Hdwardsii, Bate ?, 102.
—— “ dubius, Johnston,” 104.
Melita palmata (Montagu), 90.
obtusata (Montagu), 91 and 92 g,109 9.
91
92 Canon A. M. Norman—Zevised Nomenclature of
Melita gladiosa, Bate, 93.
Eriopisa elongata (Bruzelius), 265.
Niphargus subterraneus (Leach), 84.
fontanus, Bate, 85.
Kochianus, Bate, 86.
Crangonyx subterraneus, Bate, 87.
Mera othonis (H. M.-Edw.), 108 9,107 o.
Loveni (Bruzelius), 266.
— Batei, Norman, 101 ?, 257, 267.
grossimana (Montagu), 94.
Ceradocus semiserratus (Bate), 106.
Elasmopus rapax, Costa, 110.
Gammarella brevicaudata, 64 9, 88 5,89 @.
Cheirocratus Sundevalli (Rathke), 48 3g, 80 2.
assimilis (Lilljeborg), 256 3.
Lilljeborgia pallida, Bate, 47 9°.
Kinahani, Bate, 50.
‘“* Phedra antiqua, Bate,” 49.
Iswa Montagui, H. M.-Edw., 51.
Microdeutopus anomalus (Rathke)?, 77 9.
damnoniensis (Bate), 75 3.
Aora gracilis, Bate, 73.
Stimpsonella chelifera (Bate), 74.
Lembos Websterii, Bate, 76.
Coremapus versiculatus (Bate), 78.
Leptocheirus pilosus, Zaddach, 79.
Gammaropsis maculatus (Johnston), 95 3,96 9, 105.
Microprotopus maculatus, Norman, 269.
Photis longicaudata (Bate), 111.
Podoceropsis Sophie, Boeck, 1380 3,183 9.
excavata (Bate), 1381 g§,132 9,114.
Amplhithoe rubricata (Montagu), 112 ¢,113 9.
Pleonexes yammaroides, Bate, 115 ¢,116 @.
Sunamphithoe pelagica (H. M.-Edw.), 117.
Bruzeliella falcata (Montagu), 121 $ junior, 118 g, 122 9°.
pusilla (G. O. Sars), 119 9.
ocia (Bate), 123.
Jassa pelagica, Leach, 120.
Erichthonius abditus (Templeton), 124 3.
difformis (H. M.-Edw.), 125 ¢, 126 9.
Cerapus crassicornis (Bate), 129.
Orustacea described by Bate and Westwood.
Siphonecetes Whitei (Gosse), 127 5,128 2.
Corophium volutator (Pallas), 138 ¢.
affine, Bruzelius, 139 9.
crassicorneé, Bruzelius, 140 3.
Unciola irrorata (Bate), 186 9,137 ¢.
planipes, Norman, 258.
Chelura terebrans, Philippi, 141.
Platophium variegatum (Leach), 134.
Dulichia porrecta, Bate, 147 3.
falcata, Bate, 145 3.
Colomastiv pusilla, Grube, 135.
Tribe III. Capriniipra.
Phtisica marina, Slabber, 149 9,150 6.
Pseudoprotella phasma (Montagu), 151.
Periambus typicus (Kroyer), 159.
Caprella linearis (Linneé), 152 9, 153 ¢.
septentrionalis, Kroyer, var. nodifera, Mayer, 150.
equilibra, Say, 158.
acanthifera, Leach, 156.
——- acutifrons (Latreille), 154.
tuberculata, Guérin, 157.
Cyamus mysticeti, Lutken, 160.
? ovalis, Roussel de Vauzéme, 161.
? gracilis, Roussel de Vauzeme, 162.
Platycyamus Thompsoni (Gosse), 163.
Il. Isopvopa.
Tribe I. CHE LiFERa,
Apseucdes talpa (Montagu), 169.
Latrellii (H. M.-Edw.), 170.
Tanais Cavolini, H. M.-Edw., 164.
Dulonguv (? Audouin), 165.
Leptochelia Savignit (Kroyer), 166.
Paratanais Batei, G. O. Sars, 167.
Leptognathia rigida (B. & W.), 168.
Tribe I]. FLABELLIFERA.
Anthura gracilis (Montagu), 171.
Paranthura nigropunctata (Lucas), 172.
93
94 Crustacea described by Bate and Westwood.
Gnathia maxillaris (Montagu), 173.
oxyurea (Lilljeborg), 174.
formica (Hesse), 175.
Anilocra physodes (Linné), 271.
Aiga tridens, Leach, 189.
psora (Pennant), 190.
monophthalma, Johnston, 191.
—-? rosacea, Risso, 188.
Rocinela danmoniensis, Leach, 192.
Cirolana Cranchii, Leach, 193.
borealis, Lilljeborg, 194.
Conilera cylindracea, Leach, 195.
Eurydice pulchva, Leach, 196.
—— truncata (Norman), 270.
Limnoria lignorum (Rathke), 204.
Spheroma serratum (Fabricius), 215.
rugicauda, Leach, 216,
Hookeri, Leach, 217.
Cymodocea truncata, Leach, 218 and 219 9, 223 and 224 ¢.
Nesa bidentata (Adams), 220 and 221 9, 225 ¢.
Montagu, Leach ?, 222.
Campecopea hirsuta (Montagu), 226 ¢, 227 9.
Tribe III. Varvirpra.
Astacilla longicornis (Sowerby), 205 9, 207 3.
intermedia (Goodsir), 206.
Idotea balthica (Pallas), 208.
pelagica, Leach, 209.
emarginata (Fabricius), 210.
linearis (Pennant), 211.
Zenobiana prismatica (Risso), 212.
Stenosoma acuminatum, Leach, 213.
lanciferum (Leach), 214.
Tribe IV. AseEttora.
Asellus aquaticus (Linné), 203,
Janira maculosa, Leach, 202.
Jera marina (Fabricius), 197.
Nordmanni (Rathke), 198.
Munna Kroyeri, Goodsir, 199 g, 200 ¢.
Leptaspidia brevipes, B. & W., 201.
Dr. F. Baron Nopesa on Kerunia. 95
Tribe V. OnIscorppa.
‘a oceanica (Linné), 228.
honiscus pusillus, Brandt, 231.
— - vividus (Koch), 232.
roseus (Koch), 233.
Oniscus asellus, Linné, 235, 236.
Philoscia muscorum (Scopoli), 229,
Couchii, Kinahan, 230,
Platyarthrus Hoffmannseggvi, Brandt, 234.
Porcellio scaber, Latreille, 237.
dilatatus, Brandt, 238.
prtus, Brandt, 239.
levis, Latreille, 240.
Metoponorihus pruinosus (Brandt), 242.
emngendus (Kinahan), 243.
Cylisticus convexus (De Geer), 241.
Armadillidium vulgare (Latreille), 244.
Tribe VI. Errcartpa.
Bopyrus squillarum, Latreille, 176.
Pseudione Hyndmanni (B. & W.), 182.
confusa (Norman), 184.
Pleurocrypta longibranchiata (B. & W.), 188.
Athelges paguri (Rathke), 181 and 180, the young.
Phry«us abdominalis (Kroyer), 179.
Lone thoracica (Montagu), 185.
Liriopsis pygmea (Rathke), 186.
Cryptothir balani (Bate), 187.
IX. — Kerunia, a Symbiosis of a Hydractinian with a
Cephalopod. By Dr. Francis Baron Nopcsa, Member
of the Hungarian Geological Society.
[Plate IIL]
DurRING arecent visit to Qasr-el-Sagha, in the Fayum district
of Egypt, I succeeded in obtaining a large number of
specimens belonging to the Eocene genus Kerunia uniting
characters which have hitherto not been properly described.
96 Dr. F. Baron Nopesa on Kerunia.
In the first description of Kerunia, given by Professor Mayer-
Eymar, this fossil was regarded as a Cephalopod, but in a
later paper by Dr. Paul Oppenheim it was placed among the
Hydractinians. Remarkable peculiarities in the structure of
this fossil have brought about this discrepancy of views as to
its systematic position in the animal kingdom; but after a
careful study of a fairly complete set of specimens I find
myself in accordance with Mayer-HKymar, as well as with
Oppenheim, and consider Kerunia to be both a Hydractinian
and a Cephalopod.
(A) Kerunia inhabited by a Cephalopod.
Contrary to Oppenheim’s criticism on Mayer-Eymar’s
reconstruction of Kerunia, an examination of several ex-
amples in the British Museum, in the Museum of the
Egyptian Geological Survey at Cairo, in my own collection,
as well as a specimen portrayed in one of Oppenheim’s figures
(Oppenheim, loc. crt. p. 46, fig. 1), all exhibit the accuracy
of Mayer-Eymar’s views, although between the different
specimens there exists a good deal of what may perhaps be
termed individual variation. Kerunia is a bilateral, calca-
reous, mostly recrystallized mass, always showing, however,
a rapidly augmenting, strongly bent, cone-like, median cavity
(Pl. ILI. fig. 6), the outer wall (convex) of which carries
in its median line a row of lofty spines (fig. 9), while from
each side wall of the cavity mentioned one long spine-like
projection is given off (fig. 38). The cone-like cavity, which
evidently contained the soft parts of the animal, shows on its
internal (convex) side a projecting lip, forming the margin of
its opening (fig. 1). The opening itself is sometimes large
and somewhat lobate (fig. 1), although quite as frequently
round and rather constricted (fig. 4). In one case, to be
referred to later on, it is perfectly closed (fig. 10). As shown
in figs. 1-10 of Plate III., Aerunta is a perfectly regularly
built organism, and Oppenheim’s aggressive phrase, “ Ks
bedarf daher eigentlich kaum einer Versicherung, dass auch
mir nichts Aehnliches vorliegt, und dass es die Phantasie des
Autors, nicht wie dieser meint, diejenige der Natur war,
welche geschiftig dieses Fabelwesen geschaffen hat,” is
entirely without foundation,
Looking over my specimens of Kerunia with Mr. R. Bullen
Newton, of the British Museum, to whom all students of
Dibranchiata are greatly indebted for his very useful paper
(written in conjunction with Mr. G. F. Harris) “ A Revision
of the British Kocene Cephalopoda,” he at once indicated to
Dr. F. Baron Nopesa on Kerunia. 97
me the great resemblance that existed between Kerunza and
the Middle Eocene Belosepia found in the Bracklesham beds
of England. Belosepia seems to differ from Kerunia mainly
by the absence of lateral horns and by the fact that the front
median spines of AKerunda are represented in Belosepia by
several rows of irregular tubercles. The last and principal
horn of Kerunia is, however, present in the British Hocene
genus.
Fig. 1. Longitudinal section of Belosepia ({ nat.). Fig. 2. Longitudinal
section of Kerunia (2 nat.).
The unshaded surface indicates the calcareous substance and lines of
growth; the shaded surface (in fig. 2) is the region where the lateral
spines are given off; the black surface shows the chamber containing
the soft parts of the animal. L=ventral lip; S= principal spine in
Kerunia (=rostrum in Belosepia); ch=chamber. (The parts
marked in fig. 2 by dotted lines are not preserved.)
To prove the morphological resemblance of Kerunia and
Belosepia I have thought it best to figure two cross-sections
representing both organisms together (text-figs. 1 & 2).
These exhibit the turned-back ventral lip, separated from the
first spine by a deep transverse cleft, and beyond this first
spine in Belosepia rugosities are visible, while in Merunéa
Ann. & May. N. Hist. Ser. 7. Vol. xvi. 7
98 Dr. F. Baron Nopesa on Kerunia.
some more spines are developed. Internally both fossils are
furnished with the bent cone-like chamber which in Beloseptia
has a much shorter ventral side than characterizes Kerunta ;
therefore in this latter genus the cone appears to be much
more complete and more involute than in Lelosepia. Thus,
though the lateral horns of Keruwnia are absent in Belosepza,
still the similarity is striking. Moreover the bilobate opening
of Kerunza (Pl. III. fig. 1) is fairly safe evidence that the
internal chamber of thisorganism wasinhabited by a Belosepta-
like animal. The growth of Kerunia occurred by thin layers
of calcareous matter superimposed one above the other over the
whole body, so that at the same time the oral opening was
shifted forwards and the different spines augmented in height
and length. No new layers were formed on the inside wall
of the chamber, except close by the mouth, where sometimes
a constriction resulted. That the formation of new layers
was influenced by the animal living in the cone-like chamber
is clearly shown by the fact that in the same degree as the
oral aperture was shifted forward new spines were developed
in the median line, one in front of the other. In some cases,
however, double spines were developed, that 1s, one or two
on each side of the median line (fig. 4).
As the new layers which augmented the size of Kerunia
showed different thicknesses, the first small dorsal spines
slowly got covered up, and it is thus explainable that the
hindermost free spine of the dorsal row (the one nearest to
the ventral lip) is always the strongest, while the more
posterior spines form a breast-like projection. In large or
old specimens the lateral spines show branch-like protu-
berances or outgrowths (fig. 5), and as the number of these
outgrowths augments an irregular body is formed resembling
somewhat the horns of an elk with its pointed ends turned
upwards (fig. 6). On such occasions it also happens that
the basal region of the lateral branches in medium-aged
specimens (fig. 1), which exhibit a certain amount of flat-
tening, becomes perfectly flat and united with the breast-like
projection (fig. 7). Then it is that remarkable asymmetry
occurs, the dorsal spines branch off in quite a posterior
direction high above their base, and frequently again become
irregularly united (fig. 8).
Taking all these facts into consideration, I am unable to
acknowledge Kerunda as a Hydractinian: first, because of its
regular shape; secondly, on account of its external resem-
blance to Belosepia ; and, thirdly, from its regular growth on
the plan of a Belosepia-like organism.
The regular shape of Kerwnda and its external resemblance
Dr. F-. Baron Nopesa on Kerunia. 99
to Belosepia, as well as the continuous development of new
spines near the shifting oral opening, are characters which
prevent us from assuming that it is the result of a Hydrac-
tinian covering some Gastropod or other animal in the
manner quoted by H. J. Carter and others, because in such
cases irregular masses are the only result.
(B) Kerunia a Hydractinian.
Though it was rather easy to prove in the previous remarks
that Kerunia could not be a simple Hydractinian, it is
probably easier to demonstrate that this organism could have
been nothing else. The internal structure of Kerunia
throughout the entire fossilized body, as shown by different
sections, is everywhere exactly the same and without any
sign whatever of a shell-like centre. Agreeing altogether
with Hydrozoan structure and perfectly unlike the dense shell
of Belosepia*, Kerunia is built up of concentric layers, showing
a tubular structure (fig. 11). The tubule of the different
layers are always directed so as to point vertically towards
the external surface of the more internal layer, and through
irregular tubes of similar size they are in communication
with each other. The outermost layer is capable of covering
Balanids and other animals that adhere to its surface, forming
another fact which forcibly reminds us of Hydractinian
affinities.
As already remarked by Oppenheim, there can be distin-
guished on the external surface of Kerunta low spine-like
defences, pits for the single individuals, and small channels
corresponding to the sarcorhiza. The tubercle-like defences
show clearly small convergent riblets, and are mostly accumu-
lated on the bottom or more flattened surface, the pits are
more numerous on the top, while the dense network formed
by the sarcorhiza seems equally spread out over the whole
body ; but none of these elements appear to be present in
the cone-like chamber. Since Oppenheim’s sketch gives but
a poor representation of the delicate surface-sculpture of our
animal, it is considered necessary to introduce a micro-
drawing of a portion of the surface of Kerunia (fig. 12), where
a great resemblance will be observed to Vinasso de Regny’s
illustration of the external structure of Cyclactinia + tncrus-
tans. At the same time it should be observed that Cyclactiniu
incrustans is the single Hydractinian species giving off
Kerunia-like ianieaeione
* Beloptera seems to be less dense in its shell-structure.
+ This genus, according to Oppenheim, belongs to Hydractinia,
100 Dr. F. Baron Nopesa on Kerunia.
A specimen of Kerunza in the British Museum collected by
Dr. C. W. Andrews has the aperture of the conical chamber
not only constricted but perfectly closed up by Hydractinian
structure (fig. 10), proving that the outer layer continued
growing and developing even after the death of the proble-
matical organism that occupied the cavity. In this case,
therefore, the Hydractinian growth along the oral opening
developed irregularly over the surface of the fossil as charac-
terizes a true Hydractinian.
The study of such an example of irregular growth amply
proved by its microscopical structure, both internal and
external, leads to the inference that the animal which formed
these calcareous masses must have been a Hydractinian.
(C) Conclusion: Kerunia a Symbiosis.
There is only one way of merging these two results as set
out in the A and B divisions of this paper, and that is by
assuming that Kerunia resulted from a remarkably close
symbiosis of a Belosepia-like Cephalopod with an encrusting
Hydractinian, in which case symbiosis went so far that the
Hydractinian overtook the labour of building up the protec-
tive shell of the Cephalopod which fixed or controlled to a
certain extent the growth of the Hydractinian.
Only by the death or incapacity of the Cephalopod could the
Hydractinian develop in an irregular, and in consequence
also an asymmetrical, manner.
Perhaps the symbiosis of Hydractinians with Pagurus may
also throw some light on these biological questions.
This explanation of Kerunia is of necessity only a hypo-
thesis, and yet it seems more probable than any other theory
we can advance, since the phenomenon of symbiosis may be
traced among recent Hydractinians, as, for example, H. levi-
spina, which has the tendency to destroy the Gastropod shell
on which it settles. Besides, the symbiosis hypothesis only
can account for the otherwise most remarkable fact that
among the numberless Kerunias I have examined no two
specimens were precisely alike.
The locality where I found the Kerunias here described
and figured was the second low terrace above the temple
ruins of Qasr-el-Sagha. The specimens occur in great quan-
tities in an oyster-bed (Alectryonia Clot-Beyi, Bellardi) and
are always found with their oral openings and the flattened
side downwards, so that the dorsal spines and the ends of the
somewhat twisted lateral horns are turned upwards as weapons
Dr. I’. Baron Nopesa on Kerunia. 101
of defence. Such an occurrence would make it possible that
during life Aerunta existed on a sea-bottom with its head
downwards.
Before finishing this short note I would like to express my
hearty thanks to Mr. K. B. Newton for the great kindness
he has shown by giving me many valuable ‘suggestions as
well as by undertaking ‘ite very tedious work of revising the
manuscript and proof-sheets of this paper.
Literature.
I. E. Epwarps. ‘A Monograph of the Eocene Mollusca of England.
—Part I. Cephalopoda.” Palzeontographical Society, 1849.
H.J. Carter. “ Transformation of an Entire Shell into Chitinous
Structure by the Polype Hydractinia.”. Ann. & Mag. Nat. Hist.
1878, ser. 4, vol. xi. p. 1, pl. 1.
R. B. Newron and G. F. Harris. “A Revision of the British
Eocene Ce Pe Proc. Mal. Soc. London, 1894, vol. i
pelo pla
C. Mayer-Eymar. “ Un ‘singulier Céphalopod de l’Eocéne d’Eeypte
er unia cornuta |.” Eclogze Geologice Helvetiz, 1900, vol. vi.
no. 2, p. 120.
——. “Interessante neue Gastropoden ausdem Untertertiaér Ngyptens.”
Vierteljahr. nat. Ges. Ziirich, 1901, vol. xlvi. p. 30, pl. il.
P.E. Vinassa pt Rea@ny. “Studi sulle Idractinie fossili.” Atti
Real. Accad. Lincei (Roma), Mem. vol. iii. 1901, p. 107, pls. 1.~iii.
P. OPPENHEIM. “ Ueber Kerunia cornuta, Mayer- -Ey mar, aus dem
Kociin Aegyptens.” Centralblatt min. eeol. ‘Paleont. 1902, p. 44
(woodcut figures).
TF. Norcsa. “ Remarques sur Kerunia cornuta.” Bull. Soc. géol.
France, 1905 (in manuscript).
IsXPLANATION OF PLATE III.
Tig. 1. Kerunia, basal view, showing ventral lip and large oral
opening.
Fig. 2. The same, posterior view, showing section of median part of
chamber and fractured surface for main dorsal spine.
Jag. 35, Another specimen, showing breast-like projection and constricted
oral opening.
fig. 4. Another specimen, showing perfectly preserved oral opening
with fractures of numerous symmetrically placed spines.
. Distal end of a lateral spine exhibiting ramification.
An irregular specimen, showing internal chamber and complex
ramification of both lateral branches, dorsal aspect.
The same, giving basal view and showing flattened internal
surface.
Part of a dorsal spine of another specimen exhibiting ramification.
Dorsal aspect of small example, showing dorsal and lateral
spines.
Aw
&
OM N Qr
102 Miscellaneous.
Fig. 10. Front view of same specimen, showing a perfectly closed oral
opening.
Fig. 11. Longitudinal median section of another specimen, illustrating
tubular structure, enlarged.
Fig. 12. Enlarged surface-drawing of the specimen figured in figs. 1
and 2,
The figures are drawn half the natural size of the specimens with the
exception of 11 and 12, which represent magnified structures. Figures 9,
10, and 11 refer to specimens in the British Museum, the remainder being
in the Author's collection.
MISCELLANEOUS.
The Nomenclature of Types in Natural History.
By Cuartes Scuucuerr and 8. 8. Bockman.
Practica work on the arrangement and cataloguing of “ types”
and other museum material has shown us that the present nomen-
clature is not yet sufficient for critically distinguishing all the
different classes of such specimens. Further, some of the terms
which have been proposed for the purpose are already employed in
other ways—for instance, homotype is in use in biology, monotype
is the name of a printing-machine, autotype is the term for a
printing-process. | We wish, therefore, to submit the following
system of nomenclature, and we hope that in making it more com-
plete we have provided a scheme which will render efficient service
in the labelling and registration of types and typical material.
The terms printed in broad-faced letters are the additions or
modifications for which we are at present responsible. A fuller
explanation of all the terms will be found in the ‘ Catalogue of the
Types and Figured Specimens of Invertebrate Fossils in the U.S.
National Museum,’ a work which has been prepared by Charles
Schuchert and is now passing through the press. The present
article gives a synopsis of the terms which it has been found neces-
sary to use in connexion with that and similar work.
We now make another suggestion. After the different terms we
have placed in circles the contractions which we propose should be
used in the actual marking of small specimens, to which it is
impossible or inadvisable to affix the full label. Our plan for such
contractions is this :—For types of the first class two capital letters ;
for those of the second class one capital and one small letter; for
typical specimens two small letters.
In the definitions which follow, the term description indicates
either a description by words or by a picture, or by both combined.
For the sake of accuracy, we suggest that the original description
by words (type-description) be called the protolog, the original
Miscellaneous. 103
description by a picture (type-figure) the protograph. It is
obviously more easy to identify actual types from the latter than
from the former.
Primary Types (Proterotypes).
Material upon which original descriptions of species are based.
Holotype Gn) The only specimen possessed by the nomenclator
at the time; the one specimen definitely selected or indi-
cated by the nomenclator as “the type”; the one specimen
which is the basis for a given or cited protograph.
Syntype (more correct than Cotype) . A specimen of the
original series, when there is no holotype.
Paratype (2) A specimen of the original series, when there is
a holotype.
Lectotype m) A syntype chosen, subsequently to the original
description, to take the place which in other cases a holotype
occupies. (Aexrods, chosen, picked.)
Suretementary Typrs (Apotypes, vice Hypotype in use).
Material upon which supplementary descriptions of species
are based.
Heautotype (vice Autotype, in use) . A specimen identi-
fied with an already described and named species, selected
by the nomenclator himself in illustration of his species,
such specimen not being recognizable as one of the protero-
types. (éavréu, of his own.)
Plesiotype (s.) A specimen idéntified with an already described
and named species, but not selected by the nomenclator
himself.
Neotype (st) A specimen identified with an already described
and named species, selected to be the standard of reference
in eases when the proterotypes are lost, destroyed, or too
104 Miscellaneous.
imperfect for determination, such specimen being from the
same locality and horizon as the holotype or lectotype of
the original species.
Typrcat Specimens (Icotypes). (ét0s, what is like *.) -
Material which has not been used in literature, but serves a
purpose in identification.
Topotype A specimen of a named species from the locality
@)
of the holotype or lectotype—in paleontology from the same
locality and horizon.
Metatype A topotype identified by the nomenclator himself.
@)
Idiotype ( it. ). A specimen identified by the nomenclator
©
himself, but not a topotype. (técos, one’s own.)
Homeeotype (vice Homotype, preoccupied) (ns) A specimen
NY
identified by a specialist after comparison with the holotype
or lectotype. (é6poos, resembling.)
Chirotype A specimen upon which a chironym is based.
©)
(Chironym, a MS. name, Coues, 1884.)
In addition to the above we have the use of the word “ type” in
connexion with genera—a given species is the type of the genus.
The classification of such types is as follows :—
Typrs or GENERA (Genotypes).
Genoholotype. ‘The one species on which a genus is founded ;
of a series of species on which a genus is founded, the one
species stated by the author to be “ the type.”
Genosyntype. One of a series of species upon which a genus is
founded, no one species being the genoholotype.
Genolectotype. ‘The one species subsequently selected out of
genosyntypes to become “the type.”
* eixds (gen, étxdtos), ecko—for evkoro,—to make Icotype for euphony,
Ann.& Mag Nat Hist... 7 Vol XVL PLL.
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EW.L.Holt del.
GM Woodward Lith. Katerythrops Oceanae.
‘ecoydeuevdeap eisneydoyyeun
“yatL paempoom’ WD
“divry rremme NTS 8M ‘TPP HTM 3e80ueedn Oo TSTy
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IT ARES AMEGARAG A RK,
Ann & Mag. Nat. Hist. §.7 Vol. XVI PLA
¢
co plbe GAIG
Bale & Danielsson 1'4imp
P Highley del.et lith .
KERUNIA CORNUTA, Mayer -Eymar from the Eocene of Egypt .
THE ANNALS
MAGAZINE OF NATURAL HISTORY,
[SEVENTH SERIES,]
No. 92. AUGUST 1905.
X.—A List of the Batrachians and Reptiles collected hy
Dr. W. J. Ansorge in Angola, with Descriptions of new
Species. By G. A. BOULENGER, F.R.S.
[Plate IV.]
THE collection made in 1903-1905 by Dr. Ansorge has
considerably added to our knowledge of the Batrachians and
Reptiles of Angola, the study of which has been pursued for
so many years by Professor Barboza du Bocage. Travelling
under somewhat unfavourable conditions for the preservation
of specimens in spirit, Dr. Ansorge has been so fortunate as
to discover as many as four new species. As a contribution
to the knowledge of the exact distribution, a full list of the
species represented in the collection is here given.
Dr. Ansorge has supplied me with the following notes on
some of the localities visited by him :—
Ambaca.—A station of the Loanda to Lucalla railway. Was
visited by Livingstone in 1854, and is mentioned by
him. Is now a very insignificant place, consisting of
some official quarters and half a dozen small shops.
April-May 1903.
Bange Ngola.—The Portuguese were erecting a fort here
when I visited it (end of 1903). It is named, as usual
with African natives, after the important chief who lives
here. In maps the whole of this north-eastern part of
the Loanda province is called the “Jinga country,”
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 8
106 Mr. G. A. Boulenger on
ruled by a number of petty native chiefs apparently
independent of each other.
Bango.—The village of another petty chief in the Jinga
country. End of 1903.
Bihé.—A district in the north-east of the occupied portion of
the Benguella province; roughly only the eastern half
of the Benguella province is only nominally Portuguese.
End of 1904.
Bingondo.—A locality in the northern part of Bihé. The
country is wooded, interspersed with large open glades,
and well populated. October 1904.
Canhoca.—An important station on the Loanda to Lucalla
railway. Nov. 1903—-Feb. 1904.
Duque de Braganga.—Some hundreds of years ago this was
an important Portuguese fort, now it is an insignificant
military station with a dozen shops. It lies on the south
side of the Lucalla River. End of 1903.
Fort Don Carlos.—The most northern fort of the Loanda
province, and in process of erection when I visited it.
It lies at the junction of the Cambo and Cuango rivers.
End of 1903.
Golungo Alto—A rich agricultural district, with a town of
the same name, visited by Livingstone in 1854. Miles
and miles of coffee-plantations are going to waste.
Jan. 1904.
Marimba.—Just conquered by the Portuguese from rebellious
Jingos when I visited it towards the end of 1903. A
fort was being built. Near here is a small lake which I
named ‘‘ Sarmento,’ where I caught the new Cyprino-
dontid fish named Haplochilus macrurus (Blgr. Ann.
& Mag. Nat. Hist. (7) xiv. 1904, p. 19).
Pungo Andongo.—A famous spot for hundreds of years,
known also as the “ pedras nigras” or “ black rocks,”
to which prisoners were transported from Lisbon.
Similar to our “ Botany Bay ” as an exile for prisoners.
It consists of quaint and curious gigantic rocks, hundreds
of feet high, rising suddenly out of the level plain. It
is still important, owing to the sugar-plantations in the
plain. The village of the same name consists of official
quarters, a few shops, and an American Mission. June
and July 1908.
Quilenges.—A district, not far from the ocean, in the south-
western corner of Benguella province. The ruins of an
old fort of the same name still serves as the official
quarters of the officer in charge of the district. It isa
Batrachians and Reptiles from Angola. 107
basin-like depression at the western foot of a lofty chain
of hills. January and February 1905.
Tembo Aluma.—A native village about two hours on foot
from Fort Don Carlos. End of 1903.
BATRACHIA.
ECAUDATA.,
1. Xenopus levis, Daud.
Pungo Andongo.
I have carefully compared the specimens brought home by
Dr. Ansorge with one from Benguella received from Prot.
Barboza du Bocage and with others from various parts of
South Africa, with the result that I cannot regard X. Petersi’,
Bocage, as more thana small race of X. levis.
2. Bufo regularis, Reuss.
Duque de Braganea, Pungo Andongo, Locomi, Canhoca,
Marimba, between Benguella and Bihé.
3. Hemisus marmoratum, Peters.
Semba Acendu.
4. Rana ornatissima, Bocage.
Bingondo in Bihé. Caught in the forest in Brune
In the breeding male a brown callous pad is present on
the inner side of the forearm at the base of the second and
third fingers; similar callosities cover the upper surface of
the three inner fingers. In all other characters this fine frog
agrees with ft. Budgetti as described by me in 1903, and I
must therefore now regard the Gambia specimens as repre-
senting merely a colour-variety of the species previously
described by Barboza du Bocage. R. moeruensis, Blgr., is
also very closely allied to &. ornatissima, but differs in the
inner finger extending beyond the second,
5. Rana mascareniensis, D. & B.
Between Benguella and Bihé.
6. Rana Ansorgit, sp.n. (Pl. IV. fig. 1.)
Vomerine teeth in two short transverse series close to the
inner front edges of the choanee. Head once and a_ half
Sx
108 Mr. G. A. Boulenger on
as long as broad; snout acutely pointed, longer than the
diameter of the orbit, very strongly projecting beyond the
mouth; canthus rostralis feebly marked; loreal region
slightly concave; nostril midway between the eye and the
end of the snout ; interorbital space narrower than the upper
eyelid ; tympanum distinct, three fourths the diameter of the
eye. Fingers moderate, obtusely pointed, first not extending
quite as far as second; toes very long and slender, barely
half webbed ; subarticular tubercles moderate ; a small, oval,
inner metatarsal tubercle. The tibio-tarsal articulation reaches
far beyond the tip of the snout. Foot nearly two thirds the
length of head and body, tibia rather more than two thirds.
Back with eight glandular longitudinal folds, outer strongest ;
a glandular fold from beneath the eye to the shoulder.
Greyish olive above, with black spots; limbs with regular
blackish cross-bars; a black band on the side of the head,
passing through the eye and covering the temple; a vertebral
stripe, the outer glandular fold, and a streak on the upper
lip, from the end of the snout to the shoulder, yellowish
white; lower parts white, edge of lower jaw and side of
throat black.
From snout to vent 40 mm.
A single female specimen from between Benguella and
Bihé.
This new species is intermediate between 2. mascareniensis
and £. stenocephala, Blgr.
7. Rana angolensis, Bocage.
Pungo Andongo.
8. Rana o«yrhynchus, A. Smith.
Duque de Braganga, Cuanza R.
9. Phrynobatrachus natulensis, A. Smith.
Bange Ngola.
10, Arthroleptis xenochirus, sp.n. (Pl. IV. figs. 2, 2a.)
Head moderate, as long as broad; snout obtusely pointed,
projecting beyond the mouth; nostril a little nearer the end
of the snout than the eye; interorbital space as broad as the
upper eyelid; tympanum moderately distinct, two thirds the
diameter of the eye. ‘Third finger extremely long, three
times the length of the second ; toes free ; tips of fingers and
toes very slightly swollen; subarticular tubercles strong; a
small, oval, inner metatarsal tubercle. The tibio-tarsal
Batrachians and Reptiles from Angola. 109
articulation reaches the tympanum. Skin smooth ; sides and
lower belly granulate; a strong fold across the chest.
Greyish brown above, with a vase-shaped dark marking
from between the eyes to the sacral region; some dark spots
on the temple and above the shoulder; rather irregular dark
brown bars across the fore limb and the tibia; lower parts
white. Male with a subgular vocal sac.
From snout to vent 19 mm.
A single male specimen from Marimba.
The longer third finger distinguishes this species from
A, macrodactylus, Blgr.
11. Arthroleptis parvulus, sp.n. (PI. LV. figs. 3-3 0d.)
Head moderate, as long as broad ; snout rounded, projecting
beyond the mouth, shorter than the diameter of the orbit;
nostril a little nearer the end of the snout than the eye; inter-
orbital space a little broader than the upper eyelid ; tympanum
hidden. Fingers moderate, third about once and a half as
long as second ; toes one fourth webbed; tips of fingers and
toes very slightly swollen; subarticular tubercles strong ; a
tarsal and two metatarsal tubercles, small and round. The
tibio-tarsal articulation reaches the eye. Skin smooth or
with small warts on the back. Greyish brown above, with
blackish spots and a triangular blackish marking between the
eyes, with or without a light vertebral line, or with a broad
whitish area, edged with black, covering the top ot the head
and the back; limbs with dark cross-bars ; a white, dark-
edged streak along the back of the thighs ; lower parts white,
with or without brown spots on the breast.
From snout to vent 13 mm.
Four specimens from Bange Ngola.
Closely allied to A. dispar, Peters. Distinguished by the
shorter snout, the more distinct web between the toes, and
the less distinct digital expansions.
12. Rappia marmorata, Rapp.
Tembo Aluma, Bango, Bange Ngola, Locomi, Canhoca,
Golungo Alto, between Benguella and Bihé.
13. Rappia cinnamomeiventris, Bocage.
Canhoca.
This species is, perhaps, not separable from &. picturata,
Peters (ol’vacea, Peters).
110 Mr. G. A. Boulenger on
14. Rappia Bocagit, Std.
Canhoca.
15. Rappia cinctiventris, Cope.
Bange Ngola, Nana Meya.
16. Rappia microps, Gthr.
Fort Don Carlos, Marimba, Bange Ngola.
17. Rappia nasuta, Gthr.
Bange Ngola, Canhoca.
18. Hylambates Anchiete, Bocage.
Between Benguella and Bihé.
REPTILIA.
LACGERTILI A,
1. Hemidactylus mabuia, Mor.
Pungo Andongo.
2. Pachydactylus ocellatus, Cuy.
Cuanza.
3. Agama armata, Peters.
Cuanza, between Benguella and Bihé, between Bihé and
Quilenges.
4. Agama planiceps, Peters.
Pungo Andongo, Ambaca, between Benguella and Bihé,
between Bihé and Quilenges.
5. Agama atricollis, A. Smith.
Duque de Braganca, Bange Ngola, Pungo Andongo,
Cuanza R.
6. Varanus niloticus, L.
Cunga R.
7. Ichnotropis capensis, A. Smith.
Duque de Braganca, Bange Ngola, between Benguella and
Bihé,
Batrachians and Reptiles from Angola. iff
8. Eremias lugubris, A. Smith.
Cuanza R.
9. Gerrhosaurus validus, A. Smith.
Between Bilié and Quilenges.
The single specimen measures 2500 mm. from snout to
vent.
10. Gerrhosaurus nigrolineatus, Hallow.
Dnque de Braganca, Pungo Andongo, between Bihé and
Quilenges.
The frontal shield varies much in form and the frontonasal
is sometimes in contact with the rostral, but the prefrontals
constantly form an extensive suture on the median line.
Femoral pores 14 to 18 on each side.
11. Mabuia quinqueteniata, Licht.
Pungo Andongo.
12. Mabuia Bayonii, Bocage.
Duque de Braganga, Ambaca.
13. Mabuia varia, Peters.
Pungo Andongo, between Benguella and Bihé.
14. Mabuta striata, Peters.
Between Benguella and Bihée.
15. Ablepharus Wahlbergit, A. Smith.
Between Benguella and Bihé.
16. Sepsina angolensis, Bocage.
Pungo Andongo, between Benguella and Bihe.
17. Feylinia Currori, Gray.
Golungo Alto.
RHIPTOGLOSSA.
18. Chameleon gracilis, Hallow.
Duque de Braganga, Pungo Andongo, Canhoca, Marimba.
112 Mr. G. A. Boulenger on
19. Chameleon quilensis, Bocage.
Cuanza R.
20. Chameleon dilepis, Leach.
Between Bihé and Quilenges.
OPHIDIA.
21. Typhlops punctatus, Leach.
Golungo Alto.
22. Glauconia scutifrons, Peters.
Golungo Alto.
23. Tropidonotus olivaceus, Peters.
Pungo Andongo.
24. LHelicops bicolor, Gthr.
Duque de Braganga.
25. Boodon lineatus, D. & B.
Pungo Andongo.
26. Lycophidium capense, A. Smith.
Between Benguella and Bihé.
27. Chlorophis heterolepidotus, Gthr.
Duque de Braganea.
28. Chlorophis irregularis, Leach.
Pungo Andongo.
29. Chlorophis ornatus, Bocage.
Between Benguella and Bile.
30. Dasypeltis scabra, L.
Pungo Andongo.
The specimen (a female), which belongs to the var. pal-
marum, Leach, has 27 scales across the body, 244 ventrals,
and 63 caudals.
31. Leptodira hotamleia, Laur.
Canhoca.
Batrachians and Reptiles from Angola. 113
32. Rhamphiophis acutus, Gthr.
Between Benguella and Bihé.
Upper portion of rostral shield considerably shorter than
its distance from the frontal. Ventrals 168-183, caudals
53-59.
33. Psammophis sibilans, L.
Duque de Braganea, Pungo Andongo.
54. Psammophis Ansorgii, sp.n. (Pl. IV. fig. 4.)
Snout but slightly longer than the eye. Rostral a little
broader than deep, just visible from above; nostril between
two or three shields; internasals two thirds the length of
the prefrontals ; frontal twice as long as broad, longer than
its distance from the end of the snout, as long as the
parietals ; loreal barely once and a half as long as deep ; one
preocular, in contact with the frontal; two postoculars ;
temporals 2+2; seven upper labials, third and fourth
entering the eye; four lower labials in contact with the
anterior chin-shields, which are shorter than the posterior.
Scales in 15 rows. Ventrals 153; anal divided; sub-
caudals 76. Grey above, with a very indistinct streak of
rusty along each side of the back ; no markings on the upper
surface of the head; pre- and postoculars yellowish; a
white black-edged streak on the upper lip, the upper outline
crossing the rostral, the lower half of which is white; lower
parts uniform white.
Total length 700 mm. ; tail 165.
A single male specimen from between Benguella and Bihé.
This very distinct species is most nearly related to the
South-African P. crucifer, Daud., from which it differs in
the larger eye, the shorter snout, and the coloration.
35. Dispholidus typus, A. Smith.
Between Benguella and Bihé.
36. Xenocalamus Mechovit, Peters.
Between Benguella and Bihé.
A single male specimen, measuring 395 mm., tail 30.
Ventrals 217, caudals 25. The two postoculars confluent into
one. Black above, with white spots forming more or less
regular cross-bars ; upper lip, two outer rows of scales, and
lower parts white, each ventral and subcandal shield with
crowded black dots forming cross-bars.
114 On Batrachians and Reptiles from Angola.
37. Miodon collaris, Peters.
Golungo Alto.
A single female specimen, with 233 ventrals and 17 caudals.
88. Aparallactus Bocagit, Blgr.
Between Benguella and Bihé,
A single female specimen, measuring 330 mm., tail 45.
Ventrals 175, caudals 35.
39. Nadia melanoleuca, Hallow.
Pungo Andongo.
40, Nata nigricollis, Reinh.
Golungo Alto.
41. Causus rhombeatus, Licht.
Pungo Andongo, Canhoca, between Benguella and Bihé.
Some of the specimens from between Benguella and Bihé
belong to a colour-variety described by Barboza du Bocage
(Herp. d’ Angola, p. 146), a pair of light lines running along
the back and dividing the large transverse dark dorsal spots.
Specimens of this variety, for which I propose the name
bilineatus, appear to have constantly 17 or 18 rows of scales
only. Ventrals 128-144; caudals 23-30.
42. Bitis Peringueyt, Blgr.
Between Benguella and Bihé.
A single female specimen of this little-known Viper,
measuring 340 mm., tail37. Scales in 27 rows; ventrals 131,
caudals 27. 12 or 14 scales round the eye; two series of
scales between the eye and the upper labials, which are 12 in
number. Coloration as figured by Bocage ( Vipera heraldica),
but the mid-dorsal region and the lateral round spots are
reddish brown.
43, Atheris squamiger, Hallow.
Golungo Alto.
A single female specimen, with 17 scales across the body,
171 ventrals, and 53 caudals.
44, Atractaspis congica, Peters.
Duque de Braganga and Golungo Alto.
Scales 19-21; ventrals 224-235; caudals 18-22.
On the Internal Parasites of the Tweed Salmon. 115
EXPLANATION OF PLATE IV.
Fig. 1. Rana Ansorgit, p. 107, natural size.
Fig. 2. Arthroleptis venochirus, p. 108, natural size.
Fig. 2a. Arthroleptis wenochirus, lower aspect of hand, x 4.
Figs. 3,34, Arthroleptis parvulus, p. 109, natural size.
Fig. 3b. Arthroleptis parvulus, lower aspect of foot, x 4.
Fig. 4. Psammophis Ansorgit, p. 113, upper and side views of head and
anterior part of body, natural size.
XI.—On the Internal Parasites of the Tweed Salmon. By
JAMES R. Tosu, M.A., D.Sc., Assistant Professor and
Lecturer on Natural History in the University of
St. Andrews.
[Plate V.]
SPECIMENS of the parasites mentioned below were. collectod
during the net-fishing season of 1895 at the premises of the
Salmon Fishing Company, Berwick-on-Tweed.
The distinctly marine character of the parasitic guests of
the salmon is an indication of the nature of its food. Fresh-
water parasitic forms are very rare in the salmon, and the
fact that they are practically absent in well-grown fishes
seems to point to the conclusion that salmon do not feed in
the fresh water of a short river like the Tweed except under
extraordinary conditions, when a prolonged stay is imposed
upon them.
The following is a list of Entozoa observed :—
Ascaris capsularia, Rud. Echinorhynchus angustatus, Red.
acuta, Miill. Bothriocephalus intundibuliformis,
obtusocaudata, Zed. Rud.
Distoma varicum, Lud. Tetrarhynchus grossus, Rud.
ocreatum, Rud. macrobothrius, Rud.
Miescheri, Zschokke. Tetrabothrium minimum. Larva.
Echinorhynchus acus, Rud. sp. Larva.
proteus, Westrumb. Tenia sp. Larva.
Ascaris capsularia, Rud. Entoz. t. ii. i. p. 179.
This form occurs very plentitully encapsuled on the pyloric
cxca and mesenteries. ‘lhe average length is about 26 mm.
It is very active when taken out. The usual infection is
trom 20 to 50 in each fish.
Ascaris acuta, Mill. Zool. Dan. vol. iii. p. 53.
This is the parasite that occurs most frequently in the
116 Dr. J. R. Tosh on the
almon, usually in the stomach, seldom below the pylorus.
Many are sexually mature, and if kept in fresh water will
often extrude their eggs, which undergo a direct development.
Ascaris obtusocaudata, Zed., Rud. Entoz. t. i. 1. p. 177.
Only one specimen was obtained from the stomach. It
measures 6 mm. Others may have been overlooked.
Distoma varicum, Rud. Entoz. t. ii. 1. p. 396.
This well-known form is the most constant parasite of the
salmon. It varies in size from 2-3 mm. and occurs as
noticeable yellow specks on the walls of the cesophagus and
stomach, rarely in the intestine. The colour is from the
yellow-brown ova in the uterus. The usual infection is up
to 40 or 50.
Distoma ocreatum, Rud. Entoz. t. ii. i. p. 397.
Large infections of this species were occasionally found
floating in a watery mucus in the stomach above the bend.
It was not observed attached to the stomach-walls like the
preceding species. It occurs more frequently in the sea-trout
than in the salmon, and especially after a diet of herring.
Distoma Mieschert, Zschokke, Verhand. naturforsch. Gesell.
Basel, viii. Theil, 3 Heft.
A few solitary specimens in the gullet adhering to the
mucous membrane. Brown in colour, with stalked ventral
sucker, it varies in length from 9-17 mm. _ The species is
described and figured by Prof. Zschokke from the Rhine
salmon.
Echinorhynchus acus, Rud. Entoz. t. i. i. p. 278.
£. acus was found attached in the lower intestine. The
specimens varied in length from 12-31 mm. Not common.
Echinorhynchus proteus, Westrumb. De Helm. Acanth. p. 37,
no. 66.
E. proteus has been described under many names. It
occurred in the lower intestine with the proboscis buried up
to the neck and with the bulb expanded but out of sight.
Length 6-9 mm. Commoner than £. acus.
Echinorhynchus angustatus, Rud. Hntoz. t. ii. i. p. 266.
E. affinis is given by Rudolphi as a synonym. My
Internal Parasites of the Tweed Salmon. 1g
specimens agreed better with the description of the latter.
Found occasionally in smolts and once in a small salmon.
Length 7-9 mm. A freshwater form.
Bothriocephalus infundibuliformis, Rud. Entoz. t. i. 1. p. 46.
Same as B. proboscideus, Rud. This tapeworm, which
has a wide distribution among marine fishes, is in appearance
the most formidable parasite of the Tweed salmon. It does
not seem to be seriously harmful to its host, and is to be
found in the largest and best-fed fishes, usually in numbers
ranging from 1 to half a dozen. The great infections of
from 100 upwards mentioned by Zschokke as occurring in
the Rhine salmon have not been observed in the case of the
Tweed fishes, but have been noted in the sea-trout of the
same river. In one sea-trout as many as 150 specimens, all
very young, were counted. The head of the tapeworm is
almost invariably attached near the bottom of a pyloric
cecum.
Appended is a table giving the occurrence of this tapeworm
as noted in tlie fishes observed at the Berwick-on-'weed
Salmon Company’s Fish House in 1895 :—
Male Grilse, |Female Grilse.| Male Salmon. Female
Salmon.
7a |S me. | GP RR r= ey Rete a ra
a S| & a a A Az, BS)
=| ise eS fe =| ha = SF
a 8 8 aa a 8 8 4 aa
o 95 Oo 5) 3 o o8 o 25
3 oe 3 BE 5 5B 5 a -
2 20 2 2o 2 Qo 2 = 8
5 ae| 2 a&| g ga| 8 Ea
3 54 =| 54 3 a4 s a+
ZA Zi Z Zi Zi Z A Zi
March....| .. ue ae at 3 1 12 3
70) yl Ue ee Py: i - 15 10 67 92
May os. 5 re ae a 23 8 106 | 34
UNM 8 1 114 3} 18 9 fi) |) BB
uly 2 thie 786 a 20 2 ay. 9 88 94
August ..| 42 5 24 3) 13 5 65 92
September.) 138 G 1 3 4¢ oe ll
October ..| 21 4 3 Ae 34 6 3 4
November.| 12 4 Ay, 20 5 45 | 14
137 ay 69 9 154 53 do2 157
118 Dr. J. R. Tosh on the
Of a total of 892 fishes examined 236, or 26:4 per cent.,
were infected with tapeworm. It would be interesting to
know if the falling off about October were at all constant,
and, if so, whether the rise in November was connected with
a final feast before the dash for the river.
Tetrarhynchus grossus, Rud. Synops. pp. 129 & 448, .pl. i.
fies -o.c0, 105 (Pay)
This form, known to the salmon-fishermen of Berwick-on-
Tweed as a “ flatty,”’ occurs in the abdominal cavity of both
salmon and sea-trout, and is more often dead than alive,
The live form is found almost invariably attached by its
proboscides to the outside of the lower intestine near the
vent. It is cream-coloured and soft, with parallel strize on
the flattened body behind the head. If touched or detached
it contracts actively. When preserved the striated part
becomes rugose and sometimes almost tuberculated. A
preserved specimen sent by the late Dr. Johnston from
Berwick-on-T'weed was described and figured by Dr. Baird
as T’. rugosus *.
If unable to pass to the intestine of another host, e. g.
shark, where it would most likely become sexual, the worm
after some time seems to die, loosens its hold, drifts forward
in the body-cavity, and usually becomes embedded in a soft
organ, such as the liver. There it is pressed out of shape,
becoming often triangular in section or bent on itself. The
head retains most of its former bulk, but the body becomes
stiff and attenuates. Sometimes specimens are found with
the flesh coming away in irregular layers from the body.
The surrounding tissue is often affected by this process of
degeneration. Specimens in this stage were described and
figured by Drummond as 1. solidus T.
In the accompanying Plate the process of degeneration is
illustrated. Fig. 1 is 7. grossus from a preserved specimen ;
fig. 2 has a head and body of TJ. solidus and a tail of
T. grossus, and shows the papilla where the strobila would
have budded.
Von Siebold considers his 7. claviger } to be identical with
T. grossus, and most likely all the species listed by him on
p. 235 (loc. cit.) are synonyms; but he places the ZY. solidus
* Catal. Entoz. Brit. Mus. p. 69, tab. 11. fig. 5.
+ Charlesworth’s Mag. Nat. Hist. vol. ii. (1838).
{ Zeitschr. wiss. Zool. Bd. 11.
Internal Parasites of the Tweed Salmon. we
of Drummond with TJ. megacephalus, which is distinctly
separated from T. grossus by its tapering proboscides. In
the dead form the proboscides are rarely altogether retracted,
and they still resemble those of .7’. grossus.
Tetrarhynchus macrobothrius, Rud. Synops. pp. 131 & 453.
Length 3-8 mm. Specimens occurred free in the body-
cavity, oftenest in the region of the liver and also encysted
on the outer wall of the cesophagus. When liberated from
cysts they were yellow, almost orange-coloured. Those in
the body-cavity were creamy white. '. megabothrius, Rud.,
and T. appendiculatus, Rud., may be the same as the species
in question.
The larva of Tetrabothrium minimum occurred in the mucus
of the pyloric ceca.
Tetrabothrium sp.
A few larval forms were found in fluid in the rectun—
probably more advanced forms of the last species.
Tenia sp.
One larva was observed.
The list of parasites of the Tay salmon given by Prof.
M‘Intosh* corresponds to the above in its thoroughly
marine character.
Gatty Marine Laboratory, St. Andrews,
March 31, 1905.
EXPLANATION OF PLATE V.
Fig. 1. Tetrarhynchus grossus, Rud. Length 30 mm.
Fig. 2. Ditto, Length 38 mm.
Fig. 3. Ditto. Length 29 mm,
Tig. 4. Ditto. Length 24 mm.
Fig. 5. Ditto. Length 30 mm,
All from spirit-specimens.
* Journ, Proc. Linn. Soc. vol. vil. pp. 145-154, 2 woodcuts,
+]
120 Mr. H. B. Fantham on
XII.—On Hermaphroditism and Vestigial Structures in the
Reproductive Organs of Testudo greca. By H. B.
FantHam, B.Sc. Lond., A.R.C.S., University College,
London.
[Plate VI.]
AMONG the various groups of the Vertebrata from time to
time there occur cases of true hermaphroditism, or of the
persistence of some of the genital organs of the opposite sex,
especially in the male. Such occurrences are of interest, and
as apparently few (if, indeed, any) instances have hitherto
been described among the Chelonia, I considered the following
two cases, which recently came under my notice, worthy of
record.
Last summer, while dissecting a specimen of Testudo
greca, an abnormal development of the genitalia was
noticed. Further examination showed that the specimen in
question was hermaphrodite, after comparison with male and
female specimens of this tortoise procured and dissected.
Since that date other adult specimens of Testudo greca have
been examined, about a dozen altogether, of which all but
two possessed the external characters of the male. Among
those with male external features was one showing the
persistence of the Miillerian ducts as slender rudimentary
tubes. ‘To distinguish between these two tortoises, each with
abnormal reproductive organs, the hermaphrodite specimen
first found may be denoted by A, and the second one (a
male with vestigial oviducts) by B. The anatomy and
histology of each may then be dealt with, as far as possible.
I. External Characters and Anatomy.
The external features aiding in the determination of the
sex of atortoise are, according to Gadow [4], as follows :—
The male is slightly smaller in size, has a long tail, and
possesses a concave plastron. ‘The plastron of the female is
flat. Another point of difference between the sexes that I
noticed was in the shape of the posterior end of the plastron,
as viewed ventrally (see text-figures).
Concerning the anatomy of the tortoise, no easily accessible
detailed account in English is known to me. I have used
Bojanus’s figures and explanations of Hmys europea [2].
In Sedgwick’s recently published “ Vertebrata” [10] of
his ‘Student’s Text-book of Zoology, however, g good
general account of the anatomy of the Chelonia jg given,
Hermaphroditism &c. in Testudo greeca. 121
At first it does not appear easy to delineate the relation of
the gonad and the kidney in Testudo. The latter organ
(kidney) is very dorsal and posterior in position and needs
careful dissection, being covered ventrally by a black pig-
mented body. This is the epididymis, lying naturally on
FEMALE
x
QI-
A marked indentation or angle at x in the case of the exoskeleton of the
male. The female is more rounded than the male, especially at Y.
x
the top of the kidney, from the ventral aspect, in a closely
ensheathing and darkly pigmented mesentery (cf. Pl. VI.
fig. 2, ep.). In this mesentery vasa efferentia pass from the
testis, and by it the testes are attached to the kidneys. The
testes (fig. 2, ¢.) are normally brownish-yellow bodies, ovoid
in shape, about 1°6 to 1:8 cm. long, 1:2 to 1:3.cm. broad, and
1 cm. thick. The kidneys (fig. 2, k.) are dark brown,
showing a convoluted ventral surface. There is a slight
orange supra-renal body on the ventral or inner surface of
each kidney, partly covered in the natural position by the
epididymis and testis. The supra-renal body of each side is
apparently the “Ren succenturiatus”’? of Bojanus in his
figures of Emys europea.
Tortoise A.—This specimen in external features possessed
a slightly concave plastron (rather less concave than in the
normal male), had a long tail, and was a little smaller in size
thanthe female specimens compared (though, of course, thismay
be merely a question of age). It had well developed gonads—
the left one of which, at any rate, might be termed an ‘ ovo-
testis’ (see below)—and well developed oviducts (Miillerian
ducts), in addition to well-marked epididymes, vasa deferentia,
and penis, which are, of course, essentially male genitalia.
The epididymes of this specimen were slightly larger than in
some of the normal males examined. The gonads (PI. VI.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 9
122 Mr. Hl. B. Fantham on
fig. 1, g.) attracted attention as soon as seen, being yellowish
brown in colour, rather elongate ovoid in shape, and slightly
larger, darker, and more pointed anteriorly than the testes in
normal male specimens. The gonads were approximately equal
in size and general shape, about 1:9 em. long, 1:1 cm. broad,
and 1 cm. thick. Especially noteworthy was the presence of
a conspicuous yellow egg (fig. 1, 0.) on the ventral surface of
the posterior third of the left gonad (hence use of term
“ ovotestis ” for this gonad). This egg was ‘3 cm. in diameter,
but those found in females of 7. gre@ca were often larger.
The Miillerian ducts were well developed and lay ventral
and to the outer side of each gonad (fig. 1, m.). They were
light yellow in colour and approximately equal in length. and
their external diameter varied from *3 em. anteriorly to
‘6 cm. in the uterine portion. They were attached by a
mesentery posteriorly to the epididymis of either side, and
anteriorly by the broad ligament to the lungs. Hach opened
in front by an ostium, apparently ciliated (fig. 1, 7.), into
the ccelom, considerably in front of the gonads. No eggs,
however, were noticed free in the body-cavity, nor in the
Miillerian ducts, the lumen of each of the latter being con-
tinuous throughout. ‘These oviducts opened into the neck of
the cloaca] bladder, slightly behind and ventral to the ureters
and vasa deferentia, as shown in Pl. VI. fig. 1. The
foldings of these ducts were chiefly confined to their anterior
third.
Tortoise B.—This specimen exhibited all the external
characters of the male, the concavity of its plastron being
well marked. The genitalia (Pl. VI. fig. 2) were of the
normal size, shape, and appearance common to the male
tortoise, each testis being 1°6 cm. long, 1°2 cm. broad, and
1 cm. thick. But in addition there were rudimentary
Miillerian ducts present (fig. 2, m.), opening anteriorly into
the ccelom, that of the left side being slightly longer than that
of the right, but rather narrower in its posterior sixth and
at its entrance to the cloaca (P]. VI. fig. 2). The hinder
portion of the right oviduct being wider (‘3 cm. in external
diameter), its posterior fifth was found to contain functionally
mature spermatozoa, which, however, were not found in the
more constricted hinder part of the left oviduct. Apparently
the basal part of the right rudimentary oviduct functioned as
a sort of vesicula seminalis (¢/. Howes’s observations [6] on a
specimen of Lacerta viridis), the spermatozoa having pro-
bably migrated into it from the vas deferens. Unfortunately
the oviducts of specimen A were not examined for spermatozoa
when fresh, for reasons given below.
ww
Hermaphroditism &e. in Vestudo greca. 12:
II. Histology.
Tortoise A.—Sections of each of the gonads of this speci-
men were prepared and examined. Unfortunately the
material was not fixed immediately after death, the alimentary
canal and vascular system overlying the genitalia ventrally
having been first dissected and examined in detail. On
microscopic examination of the sections the material showed
somewhat poor fixation, as was to be expected under the
circumstances. Delafield’s hematoxylin or iron-hematoxylin
was used for staining, together with eosin or acid fuchsin.
A few transverse sections of a portion across the end of the
anterior third of the right gonad were made, and longitudinal
sections of the whole of the rest of this gonad were cut.
Testicular seminiferous canals and developing spermatozoa
were seen in all the sections, but no developing ova could be
made out with certainty. In longitudinal section the tissue
was seen to be denser in the anterior part of this right
gonad than in the posterior portion.
The left gonad was carefully sectionized longitudinally, as
it showed on its ventral surface an apparent egg. ‘This latter
was, in section, found to be an ovum, the nucleus and yolk
being seen (Pl. VI. fig. 3, n. & y.). Slightly anterior
and dorsal to this ovum, and so within the substance of the
left gonad, another was found developing, after examining a
series of sections. Further, a few groups of bodies resembling
developing “ovarian ova” were seen scattered in separate
groups (follicles) among otherwise testicular tissue, more
Sr a the periphery of the anterior portion of the
gonad (PIP Vi; fie. /4,. ov.).\ The rest “of ‘this organ
consisted of seminiferous canals, connective-tissue stroma, and
a few blood-vessels. I very much regret that circumstances
do not permit of my giving a longer and more detailed account
of the histology of the left gonad, owing to unavoidable delay
in fixation.
The oviducts were lined internally by a glandular epi-
thelium, as is normally the case in the female, the walls being
highly muscular.
Tortoise B.—The right gonad was carefully fixed in corrosive
sublimate and acetic “acid, and sectionized. The left gonad
was teased up. In neither case were any traces of ova found,
only seminiferous canals and developing spermatozoa. The
fact of mature spermatozoa being found in the right rudi-
mentary oviduct directly after death has been already
mentioned. These oviducts, the lumen of each of which was
in some parts very narrow but continuous, were lined by a
glandular epithelium, though the walls were thin.
g#
124 Mr. H. B. Fantham on
Tortoise A, according to the classification of Taruffi (quoted
by Windle [14]), would be a bilateral hermaphrodite.
Tortoise B would perhaps be regarded by him as _ pseudo-
hermaphrodite. However, specimen B was without doubt
functionally a male, while A had a preponderance of male
characters.
IJI. Note on Hermaphroditism in the Vertebrata.
It may at once be said that this is rare, especially in the
Amniota. It occurs in Myeine, which is protandrous. It
occurs casually in some Selachians and in the Sturgeon.
Among Teleostean fishes it constantly occurs in Serranus
and Chrysophrys, while it is occasionally met with in the cod,
herring, mackerel, &c.
Among the Amphibia it is known in Triton (Molge)
teniatus and some frogs. Thereis the remarkable “ Bidder’s
organ” in toads. Among frogs the possession of well-
marked rudiments of Miillerian ducts is fairly common,
while specimens with complete Miillerian ducts or oviducts
are sometimes found together with hermaphrodite gonads
(cf. Marshall [9] and others). Remains of Miillerian ducts
may occur in Lmys europea, Lacerta viridis (cf. Howes [6]),
and L. agilis (cf. Jaquet [8]) among Reptiles, and herma-
phroditism has been recorded in the chaffinch among birds.
A remarkable case of hermaphroditism in the common fowl
was, | believe, lately exhibited at the Royal Society’s Con-
versazione, showing secondary characters of both sexes in
addition to a strictly hermaphrodite gonad.
Professor Howes in 1891 described acase of hermaphroditism
in the codfish [7], and, after discussing the significance of
the phenomenon in the Vertebrates, inclined to the idea that
the ancestral vertebrate was hermaphrodite. Whether this is
likely or not is open to question. The case of Myzine is
usually considered as a secondary acquirement of hermaphro-
ditism, connected with its partially parasitic habit. The
young tadpole of the frog is at one period of its existence
indifferent, which should be carefully distinguished from
hermaphrodite. __
If hermaphroditism be considered the primitive or ancestral
condition of the Vertebrata, then the cases of tortoises A and B
are retrogressive. Stephan [11], in an exhaustive and
valuable paper on this question, inclines to the view that
hermaphroditism in the Vertebrates is not a phenomenon of
retrogression, but a secondary acquirement, substituted for the
dicecious state (Stephan, 1902, page 129).
Hermaphroditism &e. in Vestudo girxca. 125
Again, the female organ (ovary) is perhaps simpler and
more primitive than the male organ (testis), as suggested by
Marshall [9], the ovum being undoubtedly less highly
differentiated than the spermatozoon. A long theoretical
discussion on the question of hermaphroditism in the Verte-
brates would, however, be out of place in this paper.
In the foregoing note only the more commonly occurring
and well known forms of Vertebrates in which hermaphro-
ditism may be found are mentioned. A very full bibliography
and comprehensive treatment of the subject will be found in
Stephan’s paper of 1902. A few other papers, relating
especially to the Reptilia, will be cited in the appended
references,
In the present state of our knowledge only suggestions as
to the significance of hermaphroditism in forms like the
tortoise can be put forward, and the recording of such occur-
rences may be of value in developing that knowledge.
References to Literature.
(1) Acassiz, L. (1857)—Contrib. to Nat. Hist. of U.S.A. vols. i. & ii.
“ Embryology of the Turtle.”
(2) Bosanus, L. H. (1819).—‘ Anatome Testudinis Europe.’ Vilna.
(3) Bronn, H. G., and Horrmann, C. K. (1890).—Klass. u. Ord. des
Thierreichs—Reptilien, vi. 3, “ Schildkroten,” by C. K. Hoffmann.
(4) Gapow, H. (1901)—“ Amphibia and Reptiles” in the Camb.
Nat. Hist. vol. viii. p. 365. is
(5) GeeEenBavr, C, (1901).—Vergl. Anatomie der Wirbelthiere, Bd. ii.
. 533-6.
(6) Bier G. B. (1887).—*On the Vestigial Structures of the
Reproductive Apparatus in the Male of the Green Lizard,’
Journ. of Anat. & Physiol. xxi. pp. 185-9.
(7) Howes, G. B. (1891).—-“ On some Hermaphrodite Genitalia of the
Codtish,” Journ. Linn. Soc., Zool. vol. xxiii. pp. 589-57.
(8) Jaquet, M. (1895).—* Note sur un cas d’hermaphroditisme in-
complet observé chez le Lacerta agilis,’ Bibliogr. Anat. iii.
(Paris) p. 267. .
(9) MarsHarr, A. M. (1884).—“On certain Abnormal Conditions
of the Reproductive Organs in the Frog,” Journ. of Anat. &
Physiol. xviii. pp. 121-45. ‘
(10) SEpewicK, A. (1905).—“ Vertebrata,” vol. ii. of ‘A Student's
Text-book of Zoology,’ pp. 402-15.
(11) SrepHan, P. (1902).—“De Vhermaphrodisme chez les Verié-
brés,’ Annales de la Faculté des Sciences de Marseille, xii.
p- 23-158, with 8 text-figures and | plate.
(12) TARUFFI (1899-1902),—“ Hermaphroditism,’ R. Ace. d. Sci.
dell’ Inst. di Bologna, 8 vols.
(13) WerpERsHErM and Parker (1897).—‘ Comparative Anatomy of
Vertebrates.’
(14) Winniz, B. C. A. (1903).—“ Thirteenth Report on recent
Teratological Literature,” Journ. of Anat, & Physiol. xxxvii.
p. 800.
126 Mr. N. Rosén—Reply to Mr. G. A. Boulenger.
EXPLANATION OF PLATE VI.
6.v. Blood-vessel. ( ov. Ovarian ovum in follicle.
el. Cloaca. | gp. Penis.
ep. Epididymis. |r. Rectum.
f.ep. Follicular epithelium. t. Testis.
Fil. Coelomic aperture in Muller- ts. Seminiferous tubules.
ian duct. | v.ef. Vasa efferentia.
g. Gonad. dS. Aperture of vas deferens
k, Kidney. | contiguous with that of
m. Miullerian duct (oviduct) | ureter.
n. Nucleus. | ©. Aperture of Mullerian duct.
o. Ovum. | y. Yolk.
Fig. 1. Tortoise A, genitalia seen from ventral surface. Cloacal bladder
removed ventrally. Left Miuillerian duct and epididymis
separated and slightly displaced to show kidney dorsal to them.
Penis turned over dorsally to show groove. Nat. size.
Fig. 2. Tortoise B, genitalia, ventral view. Round and broad ligaments
not shown (nor in fig. 1). Cloacal bladder cut away ventrally.
Nat. size.
Fig. 3. Part of a longitudinal section of the left gonad of Tortoise A,
showing an egg on the surface and the relative size of the semi-
niferous tubules of the rest of the gonad. x 12 diameters.
Fig. 4. Part of a longitudinal section of the left gonad of Tortoise A,
showing developing follicle (with ovarian ovum) wedged in
between seminiferous tubules. Slightly diagrammatic. x 100
diameters.
XIII.— Reply to Mr. G. A. Boulenger.
By Nits Rosen.
To the “ Remarks” which Mr. Boulenger in the March
number of these ‘Annals’ has made upon my paper on
snakes belonging to the Museums of Lund and Malmé, and
which at first sight may seem to many to have been totally
annihilating, I beg to give the following reply, which I hope
will fully convince all impartial readers of the groundlessness
of these ‘“‘ Remarks.”
I will begin with Mr. Boulenger’s statement that the snake
I have described as a new genus under the name of Anisodon
Lilheborgi is identical with Psammodynastes pulverulentus,
Boie, and first make a comparative survey of the most
prominent differences between them :—
Mr. N. Rosén—Reply to Mr. G'. A. Boulenger.
Anisodon Lilljeborgi (of which
there are two specimens of
exactly the same shape, one
larger than the other).
(1) Hypapophyses present through-
out the vertebral column, represented
on the posterior dorsal vertebre by a
well-developed crest, projecting below
the condyle.
(2) At least 13 maxillary teeth.
(8) Of the maxillary teeth the
fourth and fifth enlarged.
(4) The last two maxillary teeth
much enlarged and grooved.
(5) Rostral distinctly visible from
ahove.
(6) Frontal not twice as long as
broad.
127
Psammodynastes pulverulentus, Boie
(according to Mr. Boulenger’s
description).
(1) Hypapophyses absent on the
posterior dorsal vertebrae.
(2) 9-11 maxillary teeth.
(3) Of the maxillary teeth the
third or the third and fourth much
enlarged.
(4) The last mucxillary tooth en-
larged and grooved.
(5) Rostral scarcely visible from
above.
(6) Frontal twice to twice anda
half as long as broad.
(7) Temporals 2438
242).
(8) Snout short, profile truncate
or somewhat turned up in the
adult. (Head more distinct from
neck than in 4. Lilijeboryi.)
(9) With or without small
darker and lighter spots above. A
more or less distinct dark streak on
each side of the head, passing
through the eye.
(7) Temporals 2+2. (rarely
(8) Snout pointed.
(9) A dorsal series of rather
large lighter spots, edged with
black. Along the upper labials a
white streak, edged with black.
Assuming that Mr. Boulenger’s description of P. pulveru-
lentus is correct, the difference between these two snakes, as
shown by the above comparison, is so great that it seems
strange that anybody should seriously think of uniting
them. That Mr. Boulenger is, nevertheless, inclined to
do so is unintelligible, all the more so because he has himself
put forth the dentition as well as the hypapophyses as distin-
guishing characteristics of very great value for the classifica-
tion of snakes, the snakes mentioned above differing widely
from each other in these two respects. As to the hypa-
pophyses, I have (¢ Annals,’ Feb. 1905, p. 171) called atten-
tion to the fact that they are not of so much importance as
Mr. Boulenger is inclined to ascribe to them, which particularly
finds expression in his ‘Catalogue of Snakes’ concerning
Colubrine as well as Dipsadomorphinee, since he arranges the
genera of these two subfamilies in two series, ‘‘ according to the
presence or absence of hypapophyses or haemal processes on the
posterior dorsal vertebra” (Cat. Snakes, i. p. 170 segq., iii.
p- 27); and so I have of course not exclusively attached im-
portance to this difference, even if, as is quite natural, | have
128 Mr. N. Rosén—Reply to Mr. G. A. Boulenger.
given it some significance with therest. But to Mr. Boulenger,
who seems still to adhere to it, this difference should be of
such importance that he ought to be very far from thinking
that these two snakes are identical. Upon examination of
the specimens of P. pulverulentus from the Zodlogical Mu-
seum of Copenhagen, which have been kindly placed at my
disposal, I have also found that the difference between this
species and A. Lil/jeborgi, described by me, is really so
great that it is quite out of the question that they are the
same, as Mr. Boulenger maintains, and so I am of opinion
that I have the right still to adhere to its being a new genus,
although under another name (Anisodontes), as I overlooked
the fact that Anisodon has already been applied to a
fossil mammal. However, I shall please Mr. Boulenger with
the information that the difference between them has in some
degree been lessened for Aim by the observations I have
made upon P. pulverulentus. The fact is that I have seen
one specimen with well-developed hypapophyses also on
the posterior dorsal vertebree—this being quite contrary to
Mr. Boulenger’s own statements, so that I expect the
pleasure will not be altogether unmixed to him. Yet it will
be evident from what has been stated above that this has no
real importance so far as this question is concerned, while, on
the other hand, it tends to confirm my opinion that the
hypapophyses have not the great systematic value which
Mr. Boulenger maintains in his Cat. Snakes.
But enough on this subject! Just as strange as this is
Mr. Boulengei’s doubt as to the correctness of my statement
that Chrysopelea ornata sometimes has _ well-developed
hypapophyses also in the posterior region, for one can
hurdly imagine a greater inconsistency than this—first to
maintain the great importance of the hypapophyses as a
systematic characteristic, then, without changing one’s opinion
on this point, to identify one snake having hypapophyses with
another without them, and at last to doubt the statement that
with the same species these apophyses are sometimes present,
sometimes absent! Or have these apophyses a_systema-
tic value for Chrysopelea and other Dipsadomorphine and
Colubrine, but not for my Dipsadomorphine Anisodon ?
Add to this that Mr. Boulenger does not mention the other
species—Helicops modestus, Gthr., H. leopardinus, Schleg.,
and Tretanorhinus intermedius, sp. n. (pp. 170, 171)—which
I have given as a proof of the variation and comparative
unimportance of this feature for classification, and I think that
Mr, Boulenger’s “ Remarks” will appear to be unfounded.
Mr. Boulenger calls upon those who possess specimens of
On the Lizards in the Zoological Museum of Lund. 129
Chrysopelea ornata to examine how the matter stands re -
specting the hypapophyses, and to verify his statement.
An opportunity of complying with his request has kindly
been afforded me as regards specimens belonging to the
Zoological Museum of Copenhagen, and I have thereby found
my statement perfectly verified: there were several specimens,
some with, others without developed hypapophyses in the
posterior region, and I hope that Mr. Boulenger too will
succeed in finding a specimen having these apopliyses.
That Coluber fasciatus should be the same as Drymobius
Boddaertti, var. Rappii, Gthr., seems rather doubtful,
supposing one were to keep to Mr. Boulenger’s own descrip-
tion of it in Cat. Snakes.
Having shown Mr. Boulengei’s criticism in these instances
to be unjustified, | pass over his suppositions, put forward
without argument, as regards the other determinations. [ will
only mention that I have not been “ encouraged &ce.”
XIV.— List of the Lizards in the Zoological Museum of Lund,
with Descriptions of new Species. By Nits Rosin, Zool.
Inst. Lund.
[Plates VIL.-IX.]
Fam. Geckonidz.
1. Gymnodactylus marmoratus, Kuhl.
Java.
2. Gymnodactylus Miliusi’, Bory.
West Australia.
3. Phyllodactylus marmoratus, Gray.
Australia,
4. Ptyodactylus lobatus, Geoftr.
Egy pt.
5. Thecadactylus rapicaudus, Houtt.
West Indies.
6. Hemidactylus frenatus, D. & B.
Java.
7. Hemidactylus Bowringti, Gray.
130 Mr. N. Rosén on the Lizards in the
8. Hemidactylus platyurus, Schn.
Java.
9. Gehyra mutilata, Wiegm.
Java.
10. Gehyra variegata, D. & B.
West Australia.
11. Gehyra australis, Gray.
Australia.
12. Spathoscalabotes mutilatus, Gthr.
Java. Collected (1897) by Dr. Hj. Mdller.
13. Naultinus elegans, Gray.
New Zealand.
14. Gecko verticillatus, Laur.
Java.
15. Gecko monarchus, D. & B.
16. LPtychozoon homalocephalum, Crev.
Java,
17. Tarentola tuberculata, sp. n.
(Pl. VII. fig. 1; woodcut, fig. 1.)
A supraorbital bone. Snout longer than the distance be-
tween the eye and the ear-opening. Jostral scarcely twice
as broad as high. The nostril is pierced between the first
labial and three shields (woodcut, fig. 1). A median cleft in
the upper part of the rostral. ‘T’en upper labials, posterior
very small. Mental not twice as long as it is broad in the
middle. Three chin-shields on each side, in contact with
the lower labials; latter eight or nine. Anterior border of
ear-opening not denticulated. Neck, back, and limbs covered
with large pyramidal tubercles and smaller conical ones: on
the back they are more or less distinctly arranged in twelve
longitudinal rows; the largest tubercles on the back mostly
surrounded by smaller ones. The tail for about two thirds of
its length* with posteriorly directed large spine-like tubercles,
decreasing in size. Greyish brown above, more or less
marbled with darker and lighter.
Algiers (C. Ask, 1895-96).
* In one specimen nearly to the end.
Zoological Museum of Lund. 131
One specimen, with rudiments of claws on the first, second,
and fifth digits.
This new species is very closely allied to 7’. mauritanica,
L. (Boulenger, ‘Catalogue of Lizards,’ i. p. 196). It differs
in the following points :—
(1) The nostril is pierced between the first labial and three
shields ; in 7’. mauritanica, L., and, so far as I know,
in all the other species of Yarentola, it is pierced
between the first labial and two shields.
(2) The median cleft in the upper part of the rostral is
more distinct in this species.
(3) A much greater number of larger tubercles on the
body as well as on the limbs and the tail.
Fig. 1.
Tarentola tuberculata, sp. nu. Upper view of snout,
18. Yarentola annularis, Geoftr.
Egypt.
19. Spherodactylus argus, Gosse.
Fam. Pygopodide.
20. Delma Fraseri, Gray.
Victoria (Australia).
21. Delma lineata, sp.n. (PI. VIII. fig. 1;
woodcut, fig. 2.)
Snout not prominent, a little shorter than the distance
between the orbit and the ear-opening. Tail a little more
than twice as long as head and body. Rudimentary limbs
small, Rostral triangular. Nostril pierced in the lower
portion of the nasal, which forms a suture with its fellow on
the snout. ‘The anterior part of the nasal fused with the
first upper labial. No supranasals. A pair of large plates
between the nasals and the prefrontal; the latter seven-sided,
not so broad as the frontal. Three enlarged scales on the outer
IsZ Mr. N. Roséu on the Lizards in the
side of each parietal. A large loreal. Four small plates in
a line between the orbit and the nasal and some small ones
between the loreal and the orbit. Two supraorbitals. Seven
upper labials; fourth elongate and situated below the orbit,
from which it is separated by a row of small scales. Mental
large, triangular, broader than long. Five lower labials, the
Delma lineata, sp.n. a. Lower view of snout. b. Upper view of
head. c. Side view o head.
first separated from its fellow by a small scale behind the
mental. Sixteen longitudinal rows of scales round the middle
of the body. The enlarged ventral scales decreasing in width
posteriorly, the anterior at least twice as broad as long;
they form 51 pairs. Two large anal scales.
Olive above, lighter beneath, the anterior part of the lower
surfaces yellowish. A broad light (grey ? in spirit!) vertebral
line extending to the end of the tail. The edges of the scales
lighter, on the back forming small longitudinal lines between
the rows of scales.
mm,
Waidthiothead+., ssn cieun ate ase une 5
Mromisnout towvelue +. «cise ces eee fis)
Daal ee fat eR Oe ae 160
Zoological Museum of Lund. 133
Victoria (Australia).
A single specimen.
This species seems to be allied to D. ¢mpar, Fisch.*
(‘ Catalogue of Lizards,’ i. p. 244), from which it differs by
having 16 rows of scales round the middle of the body, the
first lower labial not in contact with its fellow. In D.impar,
Fisch., the prefrontal is a little larger than the frontal,
whereas this lizard has the frontal larger than the preefrontal.
In the coloration there is also some difference. This new
species differs from all previously described species of Delma
by having the anterior part of the nasal fused with the first
upper labial (perhaps an individual character).
Fam. Agamide.
22. Draco volans, L.
Java.
23. Draco fimbriatus, Kuhl.
24. Gonyocephalus Kuhli, Schleg.
Java.
Supraorbitals more or less distinctly keeled. Total length
300 mm.
25. Calotes cristatellus, Kuhl.
Java.
26. Calotes jubatus, D. & B.
Java.
27. Calotes tympanistriga, Gray.
Java.
28. Calotes versicolor, Daud.
29. Calotes ophiomachus, Merr.
30. Calotes aberrans, sp.n. (PI. VIII. fig. 3.)
Upper head-scales rather large, irregular, smooth or feebly
keeled. A rather large tubercle behind the superciliary
edge and a smaller one on its anterior end. A pair of
tubercles on the occiput. Tympanum scale-like. A slight
nuchal crest, composed of a few small spines. No transverse
gular fold. No oblique fold in front of the shoulder.
Dorsal crest indistinct ; a very slightly serrated ridge. 36
* Pseudodelma impar, Fischer, Archiv f. Naturgeschichte, 48 Jahre.
(1882) p. 286.
134 Mr. N. Rosén on the Lizards in the
scales round the middle of the body. Dorsal scales very
large, much larger than the ventrals, pointing backwards and
downwards, smooth. The median series of the dorsal scales
keeled, forming a slight ridge. Ventral scales strongly
keeled, larger on the sides of the belly than on the middle.
The adpressed hind limb reaches the shoulder. Fourth
finger as long as the third or very slightly longer. Tail
compressed, with aslight upper ridge. Caudal scales strongly
keeled.
The specimen is preserved in spirit. The coloration is
indistinct, bluish, marbled white (?}.
mm.
‘otalicngth 2 cone kee ee ee eee 200
Head-is coke SS eee eee ee 19
Tkexoveabfevavounjy 10) Why gausooansboaotoce 81
oredimb:sc ee pace eee ot eee 31
Find dimb ee ee eee eee Eee 3
SPAT GS Be Ay oa eeecta ec kote eee ea EN 119
Java. <A single specimen, collected by Dr. Hj. Moller
(1897).
This species is characterized from all the others by its
short hind limbs. It seems to be most allied to C. tympani-
striga (Gray), from which, however, it is quite distinct.
31. Agama pallida, Reuss.
Egypt.
32. Agama mossambica, Ptrs.
33. Agama stellio, L.
Sinai.
34. Amphibolurus maculatus, Gray.
West Australia. Collected by Dr. N. Holst (1896-97).
Seems to differ a little in coloration.
35. Amphibolurus Holsti*,sp.n. (Pl. IX.)
Head moderately large. Snout as long as the diameter of
the orbit, with subangular canthus rostralis. ‘Tympanum
large. Nostril below the canthus rostralis, much nearer the
eye than the tip of the snout. Upper head-scales subequal,
strongly keeled, smallest on the supraorbital region. Gular
* J take the liberty of naming this lizard after the Swedish geologist
Dr. N. Holst, who has made yery valuable collections of reptiles in
West Australia.
Zoological Museum of Lund. 155
scales much smaller than ventrals, feebly keeled. A dorso-
lateral crest on each side of the neck. A _ well-developed
dorsal crest, beginning on the neck and extending on the
anterior part of the tail, composed of closely-set compressed
spines. Body covered above with very strongly keeled scales,
largest and more strongly keeled on the middle of the back,
not intermixed with enlarged scales. The keels converge
towards the vertebral line. Ventral scales very distinctly
keeled. The hind limb stretched forwards reaches beyond
the orbit. Limbs with keeled scales. About 55 pores, ex-
tending along the whole length of the thighs, scarcely inter-
rupted on the preanal region. Caudal scales equal, keeled.
Brown above, head and body with irregular dark spots
and lines. Four large dark spots on the middle of the ante-
rior part of the back. Tail uniform brown. Lighter beneath.
Lower surface of head with smaller black spots and a large
one in the middle line reaching from snout to the gular sac.
The anterior part of belly and the middle of chest black.
Lower surface of fore limbs black.
mm.
ebeaben et bia ct ater denorcy a) ole i's abs, 0-5 cf ate 350
1 8 [ENG WERE errr a en Dee ae een a 30
1B oe Un ENE RnR COC DROU ACO temeCe 80
Barer Wri hep ts cl eeacev or henesa-oh cic, axek oeehaan ss 42
Utter Mera yo see ey hers see oie a a Se 90
SUUER Ie sen ee eee step ais ay ot errmne ity nica grav er ss « 240
West Australia. A single specimen, collected by Dr. N.
Holst (Dec. 1896).
This species seems to find a systematic position between
A. cristatus, Gray (Cat. Liz. i. p. 383), and A. caudi-
cinctus, Gthr. (Cat. Liz. 1. p. 384). To A. eristatus, Gray, it
is allied by the dorsal crest and the keeled ventral scales, but
these characters are more developed in this new species. In
A. cristatus, Gray, the crest is composed of a few widely
separated spines, the ventral scales are smooth or very feebly
keeled. ‘This species has the crest composed of many spines,
_ «tn contact with each other, the ventral scales distinctly keeled.
In these points it is well separated from A. caudicinctus,
which has a very small nuchal crest, a slight dorsal ridge,
and smooth ventral scales—With A. caudicinctus, Gthr., it
agrees in having the dorsal scales not intermixed with
scattered enlarged scales, asin A. cristatus.
36. Amphibolurus caudicinctus, Gthr.
West Australia. Collected by Dr. N. Holst (1896-97).
Seems to agree with the description of A. caudicinctus,
136 Mr. N. Rosén on the Lizards in the
Gthr. (Cat. Liz. i. p. 384), except the coloration. Lighter
beneath. ‘Tail dark, with narrow indistinct lighter (white ?)
rings. In the ‘ Catalogue of Lizards’ A. caudicinctus, Gthr.,
is described as having regular black rings, narrower than the
interspaces between them.
37. Amphibolurus reticulatus, Gray.
West Australia (VV. Holst, 1896-97).
Twenty-three specimens (all from West Australia) are
preserved in the collections. They vary very much, forming
a continuous series, from such as have no trace of a nuchal
crest to those in which it is distinct and rather well-developed.
In some specimens the ventral scales are feebly but distinetly
keeled. The coloration also is very variable. In other
characters they agree with the description of A. reticulatus,
Gray.—I have not been able to separate them into varieties
or species, and I have therefore referred them all to A. retécu-
latus, Gray.
38. Amphibolurus muricatus, White.
West Australia.
39. Amphibolurus barbatus, Cuv.
Australia.
40. Tympanocryptis cephalus, Gthr.
West Australia (N. Holst).
41. Uromastix spinipes, Daud.
Egypt.
42. Moloch horridus, Gray.
Australia.
Fam. Iguanide.
43. Anolis cristatellus, D. & B.
44, Anolis Leachit, D. & B.
West Indies.
45. Norops auratus, Daud.
46. Polychrus marmoratus, L.
47. Ophryoessa superciliosa, L.
48. Liolemus pictus, D. & B.
Chili.
Zoological Museum of Lund.
49. Amblyrhynchus cristatus, Bell,
Galapagos Islands.
50. Iguana tuberculata, Laur.
D1. Lguana delicatissima, Laur.
52. Sceloporus spinosus, Wiegm.
23. Phrynosoma cornutum, Harl.
N. America,
Fam, Anguide.
54. Ophisaurus apus, Pall.
55. Anguis fragilis, L.
Europe.
Fam. Varanide.
56. Varanus salvator, Laur.
Java,
57. Varanus niloticus, L.
Egypt.
58. Varanus Gouldit, Gray.
59. Varanus caudolineatus, Bler.
W. Australia (Dr. NV. Holst, 1896).
Fam. Teiide.
60. Tupinambis teguixin, L.
S. America.
61. Ameiva surinamensis, Laur.
W. Indies.
62. Ameiva edracantha, Bocourt.
Ecuador.
63. Cnemidophorus lemniscatus, Daud.
Fam. Amphisbenida.
64. Amphisbena fuliginosa, L.
Ecuador.
Ann, & Mag. N. Hist. Ser. 7, Vol. xvi.
10
13 Mr. N. Rosén on the Lizards in the
65. Trogonophis Wiegmanni, Kaup.
Algeria.
Fam. Lacertide.
66. Tachydromus sevlineatus, Daud.
Java.
67. Lacerta ocellata, Daud.
Algeria.
In the ‘Catalogue of Lizards’ (iii. p. 13) a variety of
L. ocellata is described (var. pater), inhabiting Algeria and
Tunis, and characterized by having the dorsal scales more
distinctly keeled and the occipital shield smaller, viz. as broad
as or a little narrower than the frontal. The number of
scales across the middle of the body and the number of
femoral pores seem to be mostly a little larger. Boulenger
says respecting it:—‘‘In some Spanish specimens the occipital
is quite of the same size as in ‘ L. pater, and also the
other characters given as distinctive are by no means constant ;
therefore the Algero-Tunisian form can, in my opinion,
not be specifically separated from the European.”—In the
specimens (from Algeria) in the collections these characters
vary considerably, so that I have not been able to distinguish
the var. pater from the European form.
Spec. a.... 17 femoral pores; occipital broader than frontal.
b. 16-18 ,, os
” ” »
Orage AGsiber a se narrower a
Te SAND ” ” ” ” ”
DEA oS] A 5 broader ss
ienine AS i, ” ” ” ”
68. Lacerta viridis, Laur.
Europe.
69. Lacerta agilis, L.
Europe.
70. Lacerta vivipara, Jacq.
Europe.
71. Lacerta muralis, Laur.
Europe.
72. Psammodromus algirus, L.
Algeria.
Zoological Museum of Lund. 139
73. Acanthodactylus pardalis, Licht.
Algeria.
74, Acanthodactylus boskianus, Daud.
Algeria.
75. Eremias guttulata, Licht.
Algeria.
Fam. Scincidz.
76. Egernia tnornata, sp. n.
(Pl. VII. fig. 2; woodcut, fig. 3.)
Head moderate. Curved groove behind the nostril absent
or very feebly marked. No distinct suture below the nostril.
Fronto-nasal large, larger than the prefrontals, forming
a suture with the rostral, separated from the frontal.
Preefrontals forming a median suture. Frontal about twice
as long as broad, considerably longer than the fronto-parietal.
6 supraoculars, third largest, first and sixth small. 7 upper
Egernia inornata, sp. n. Upper view of head.
labials, fourth, fifth, and sixth below the eye; fourth largest.
3 large temporals. One pair of nuchals. Har-opening with
four obtuse lobules anteriorly. Scales smooth, 42 round the
middle of the body. ‘The adpressed limbs overlap. Fourth
toe long. ‘Tail slightly compressed. Caudal scales
smooth. Upper surface yellowish brown, with irregular
dark brown spots, most numerous on the sides. Yellowish
beneath,
10*
140 Mr. N. Rosén on the Lizards in the
mm
Rotallongth oops essere cero ee 160
Head) nc inces occ ob 15
Wiaidthiofshead see ee eee reli
Rromisnontitowente see eee reer ee 74
Bore dlimb:\...4:5.s¢sleoe eee eee 22
Bind limb: Cites sence eee 32
Tail sae ctiecisenoh sae eeee Ceomene 86
West Australia (Dr. V. Holst, 1897). A single specimen.
Seems to be allied to Z. Whitt, Lacép.
77. Egernia depressa, Gthr.
W. Australia (Dr. N. Holst, 1896).
In one specimen (four spec. in the Museum) some dorsal
scales with five keels and spines.
78. Trachysaurus rugosus, Gray.
Australia.
79. Mabuia Perrotetii, D. & B.
80. Mabuia multifasciata, Kull.
Java.
One specimen with feeb/y keeled subdigital lamellae. Hind
limbs a little longer. In other respects it seems to agree
with MW. multifasciata, Kuhl.
81. Mabuia siamensis, Gthr.
82. Mabuia Sloanit, Daud.
83. Lygosoma (Hinulia) teniolatum, White, var. maculata,
noy. (Pl. VEU dig. 2.)
Rostral and prefrontal narrowly in contact; the pre-
frontals -separated. Hight upper labials, the last three
largest.
Brown above. A narrow vertebral line, edged with white
(in spirit). On each side of the body two longitudinal lines
(white in spirit), between which small spots of the same
colour arranged in a regular line. White beneath.
West Australia.
Two specimens, collected by Dr. N. Holst (1896).
I have not been able to find any structural characters
distinctly separating this lizard from the L. teniolatum,
White, and therefore I believe that it is only a colour-
variety of that species.
Zoological Museum of Lund. 141
34. Lygosoma olivaceum, Gray.
Java.
35. Lygosoma ornatum, Gray.
New Zealand.
$6. Lygosoma Temmincki, D. & B.
Java.
$7. Humeces Schneider’, Daud.
$8. Scincus muscatensis, Murray (?).
89. Chalcides ocellatus, Forsk.
(a) A (Cat. Liz. iii. p. 401).
Algeria, Egypt.
(5) B (Cat. Liz. ii. p. 401).
90. Chalcides tridactylus, Laur.
(a) A (Cat. Liz. iii. p. 404).
Algeria.
(>) B (Cat. Liz. ii. p. 404).
Italy.
(c) Var.nov. Four black longitudinal streaks, the median
pair broken up.
Algeria.
Fam. Chamaleontide.
91. Chameleon vulgaris, Daud.
Algeria, Sinai.
92. Chameleon senegalensis, Daud,
93. Chameleon gallus, Gthr.
Madagascar.
CORRIGENDUM.
Since I finished my descriptions of these lizards, the Ann.
& Mag. Nat. Hist. for December 1904 has been published,
in which Boulenger has described (p. 414) a new lizard
from Western Australia (Amphibolurus Webstert). The
142 Col. C. Swinhoe on
Amphibolurus Holst’, which I have described in this paper
as new, agrees with his species, and therefore the name
“* [Tolstt (sp. n.) ” must be changed to “ Websteri, Blgr.”
EXPLANATION OF THE PLATES.
PuaTeE VIL.
Fig. 1. Tarentola tuberculata, sp. n.
‘ig. 2. Egernia tnornata, sp. n.
Prate VIII.
Fig. 1. Delma lineata, sp. un.
Fig. 2. Lygosoma teniolatum, White, var. maculata, nov.
4g. 3, Calotes aberrans, sp. 1.
PuateE IX,
Amphibolurus Holsti, sp. n.
XV.—Notes on Eastern and Australian Heterocera, with
Descriptions of One new Genus and Thirteen new Species.
By Colonel Cares Swinuog, M.A., F.L.S., &c.
Family Eupterotide.
]. Melanothrix fumosa, nov.
?. Of a uniform dull smoky brown, shafts of the antennze
with an ochreous tinge; palpi blackish above: fore wings
with a broad, white, discal, transverse band, with even sides,
very broad on the costa a little beyond the middle, occupying
at least a fifth of the wing, gradually narrowing hindwards
to the hinder angle, where it is about one eighth of an inch
broad; no other markings. Underside as above, general
tint of colour paler, costal line of fore wings slightly ochreous.
Expanse of wings 2,4, inches.
Brunei, Borneo; one example.
Allied to M. alternans, Pag. Iris, iii. p. 13, from Palawan,
but the white band on the fore wings is much broader and
there are no markings on the hind wings.
Family Drepanulide.
2. Oreta figlina, nov.
¢. Palpi, head, frons, and legs crimson ; antenne, body,
Eastern and Australian Heterocera. 143
and wings above of a uniform brick-red colour: fore wings
with some grey marks on the basal half of the costa, a brown
ence near the apex, and another at one fifth from the APEX 5
a yellow thick line from the apex to the hinder margin
beyond the nnddle, rather concave in shape; a broad shadowy
grey band on the inner side of this line, which is continued
across the centre of the hind wings. Underside more
ochreous and brighter in colour; fore wings and costal
portion of the hind wings with brown striations.
Expanse of wings 14 inch.
Java; one example.
Same shape as O. bicolor, Warren, from Malacca, but
that has no indications of any median band.
Family Arctiide.
3. Diacrisia sumatrana, nov.
¢. Ofauniform dull ochreous brown: fore wings with
faint indications of antemedial, medial, and discal transverse
bands: hind wings with better-defined medial and_post-
medial bands, the latter more or less macular. Underside
slightly paler in colour; a brown mark at the end of each
cell, no other markings; shaft of the antenne pale; legs
without markings ; abdomen with dorsal and lateral indistinct
blackish spots.
Expanse of wings 1,4, inch.
Soekaranda, Sumatra; two examples.
Received under the above name, but I can find no record
of its description; is allied to D. landaka, Moore, from
Java, but that form has crimson sides to the abdomen and
distinct markings, especially on the underside.
Family Lithosiide.
4, Hugoa crassa.
Lyclene crassa, Walker, Journ. Linn. Soe. vi. p. 114 (1862).
Eugoa crassa, mpsn. Phal. ii. p. 547, pl. xxxiv. fig. 28 (1900).
Khasia Hills; two examples.
The type, from Sarawak, is im Mus. Oxon.; the outer
portion of the fore wings beyond the discal band is darker
than usual, but otherwise it appears to me to be identical.
5. Chionema axiologa, nov.
¢. Frons, head, and thorax white ; thorax with lateral
scarlet bands; abdomen grey, with scarlet hairs; antenne
144 Col. C. Swinhoe on
red : fore wings white, bands scarlet, first subbasal and erect,
second antemedial, outwardly curved, banded with black on
the inner side, third postmedial extending from the costa,
where it is nearly bifurcate, towards outer margin below the
middle, then down straight to the hinder margin about one
sixth from the angle, banded with black on the outer side,
fourth band marginal and is continued round the apex and
on the costa till it ends in the arm which nearly forms the
bifurcation of the third band; a large black spot at the upper
end of the cell: hind wings scarlet, with the costal space
broadly white. Underside pale scarlet, the lower portion of
fore and costa of hind wings white, the two lobes on the
fore wings smeared with white; fore legs brown, with white
bands, the other legs and body white.
Expanse of wings 1,5, inch.
Nias; one example.
Resembles C. Plateni, Elwes, from N. Celebes, but that has
two discoidal spots on the fore wings and the subbasal band
of that species is of a different and very abnormal character.
6. Chionema subornata.
Bizone subornata (9), Walker, 11. 550 (1854).
Chionema subornata (2), Hmpsn. Phal. i. p. 821 (1900).
Ceylon; type ? in B. M., Andaman Islands.
The male (hitherto unknown) is very like C. obliquilineata,
Hmpsn., from Sikkim, the type of which is also in the B. M.;
the lobe on the underside distorts the upper surface of the
fore wing and bends the ante- and postmedial bands, and
there are also some indications of the diffused fuscous patch
below the costa towards apex ; it is otherwise marked like
the female. I have received in a collection from Kandy one
iwmale and two females, and | have both sexes from Port Blair,
Andamans, collected by Wimberly.
Family Zygenide.
7. Thyrassia aprepes, nov.
d. Palpi, frons, and collar ochreous; antenne, body, and
wings black ; thorax with an ochreous spot behind; abdomen
with ochreous band: fore wings with an obscure semidiapha-
nous whitish streak from the base, running beneath the
median vein ; a large wedge-shaped hyaline streak at the end
of the cell, four thin streaks beyond it, and three broader
ones below it: hind wings with the basal half hyaline.
Underside: body and legs black.
Eastern and Australian Leterocera. 145
Expanse of wings ,° inch.
Key Island; one example.
Allied to T. Rafflesii, Moore, from Java.
Family Chalcosiide.
8. Soritia bicolor.
Devanica bicolor (2), Moore, Trans. Ent. Soc. 1884, p. 355,
Soritia bicolor, Hmpsn. Moths India, iv. p. 469 (1896).
Soritia viridivena (9), Hmpsn. J. ¢. i. p. 253 (1892)..
Soritia nigribasalis (go), Hmpsn. /. ec.
Cachar, Silhet, Khasia Hills.
The type from Cachar is in the Indian Museum, Calcutta ;
the type of viridivena from Silhet isin the B, M.; Sir George
Hampson has compared the two together, and found them to
be identical. The type of nigribasalis from Sikkim is in
Coll. Elwes—by some printer’s error it was described as a
female. I have received a good many examples of both
sexes of this species from the Khasia Hills,
9. Soritia Bocki, nov.
3 ?. Antenne, head, and abdomen black ; thorax with
the fore half scarlet, the hinder half black: fore wings with
a band at the apex and a large space on the hinder margin
blackish brown, leaving a large ochreous band tinged with
scarlet, which occupies more than half the wing-space, from
the base running along the upper half and bending down to
the outer margin, occupying the whole space below the apical
band : hind wings uniform blackish brown, without markings.
Underside same as above; body and legs black.
Expanse of wings 1, inch.
Hsipaw, Burma. ‘Types in B. M.
Some examples have a stronger suffusion of scarlet on the
band on the fore wings than the types; there are altogether
_three males and two females in the National Collection from
Hsipaw, a male from Hong Kong, and a female from
Shanghai. I have in my own collection a male from Kiang-Si,
which I received as Pidorus Bocki, but I can find no reference ;
it is allied to S. costata, Walker, of which there are many
examples from Northern and Western China; the wings of
that species are much longer, the coloration is deep black
with a bright scarlet narrow band, the head and collar
scarlet, the thorax wholly black.
146 Col. C. Swinhoe on
10. Soritia costimacula.
Soritia costimaculata (3), Auriv. Ent. Tidsk. xv. p. 169 (fig.) (1894) ;
Pag. Semon’s Forschungsreisen, v. pl. xiii. fig. 2 (1895).
Soritia leptalinoides (@ ), Auriv. Lc. (fig.).
Java.
I cannot but believe that these are sexes of the same
species ; the fore wings of the male have the basal streak of
the female, the subcostal discal spot, and a costal spot, indi-
cating the commencement of the medial transverse band, and
on the underside of the hind wings the same peculiar macular
submarginal band; the other differences are no more than is
usual between the sexes in this extraordinary family: the
types of both came from Java, and I have three males and a
female from the same locality.
ll. Pidorus ericydes, nov.
?. Shafts of antenne, frons, head, thorax, and first two
segments of the abdomen bright metallic blue-green; re-
mainder of abdomen dull ochreous; wings white; plumes of
the antennz and basal portions of both wings black, the
latter being well covered with bright metallic blue-green
scales, and on the fore wings is extended along the costa and
joins the large outer black band, which occupies quite a third
of the wing and contains a white subcostal spot near the
apex and two white spots close together near the outer margin
a little above the middle: hind wings with a black outer
marginal band about half the width of the band of the fore
wings, the band narrowing hindwards and terminating in a
point before reaching the anal angle. Underside: body and
legs with metallic blue-green scales: wings as above; both
wings with metallic blue-green scales covering the basal
patch, and similar scales, forming marginal bands.
Expanse of wings 2,°, ‘inches.
Woodlark Island; two examples in B. M., including the
type.
12. Cyclosia papilionaris.
Noctua papilionaris (2), Drury, Ill. Exot. Ent. ii. pl. ii. fig. 4 (1773).
Eterusia ferrea (3), Walker, 11, 451 (1854),
China.
Drury’s type came from China; Walker’s type of ferrea
in the B. M. is from Hong Kong, I have it from Tonkin ;
Drury’s figure is a very good representation of the Chinese
insect, which is very different to the various Indian forms in
Eastern and Australian Fleterocera. 147
both sexes. The allied form C. australinda, Hmpsn., from
S. India, was bred by T. R. Bell and found to be the female
of Pintia latipennis, Hmpsn.
13. Cyclosia venaria.
Phalena venaria, Fabry. Syst. Ent. p. 627 (1775).
Cyclosia papilionaris, Hmpsn. (nec Drury), Moths India, i. p. 269
(1892).
Sikkim, Khasia Hills.
The male is much like ferrea, Walker, but the white spots
are larger and more prominent ; the female has a much more
glossy sheen than papilionaris, Drury ; the white spots beyond
the cell of the fore wings are much broader than in that
species and are spear-shaped ; the black on the hind wings
forms a regular marginal border, instead of running into the
wing above vein 2, right up to the cell, as is the case in the
Chinese form. The type is in the Banksian Cabinet in the
B. M.; it is quite as distinct from papilionaris as are niyres-
cens, Moore, parvula, Butler, australinda, Hmpsn., enodis,
Swinhoe, chartacea, Swinhoe, from each other and from both
papilionaris and venaria.
14. Cyclosia castigata.
Cyclosia castigata (2 ), Walker, xxxi. 115 (1864).
Pintia cyanea (3), Butler, Ann, & Mag. Nat, Hist. (5) xii. p. 160
(1883).
Sumatra, Borneo.
The type from Sumatra is in Mus. Oxon.; Butler’s type
from the same locality is in the B. M.; there are three
females and one male from Borneo in the National Collection
and onein mine. The allied form metachloros, Walker, from
Java, was bred by Horsfield, the sexes showing the same
characteristic differences.
15. Cyclosia sordida.
Pidorus sordidus (2), Walker, Journ. Linn. Soc. vi. p. 98 (1862);
Swinh. Cat. Het. Mus. Oxon. i. p. 67, pl. ii. fig. 4 (1892).
Pintia insularis (3), Swinhoe, /. e. p. 76.
Codane obscurata, Snellen (nee Walker), Tijd. voor Ent. xly. p. 209,
pl. xiv. figs. 4 9,8 ¢§ (1902).
Borneo, Sumatra, Singapore.
The type from Sarawak is in Mus. Oxon., as also is the
type of insularis from Singapore ; there are six males and
two females from the latter locality in the B. M. and one
male from Sumatra: I have males from Singapore, Johore,
148 Col. C. Swinhoe on
Makassar, Celebes, and females from Palembang, Sumatra,
and from Nikko, Japan; the last, collected by Mr. Sanson,
in no way differs from the other Island examples. Piepers
appears to have bred these in Java: I had put them together
in my collection long before I saw Snellen’s paper; by some
curious mistake, though he has figured correctly Walker’s
type of sordida, he has printed a wrong name and a wrong
reference both of the text of Cat. Het. Mus. Oxon. and of
the number of the figure on the plate.
16. Cyclosia distincta.
a rare distincta (2), Guér. Voy. Deless. p. 85, pl. xxiv. fig. 3
ea ee (3), Moore, Cat. Lep. E. I. C. ii. p. 321, pl. viii. a,
fig. 3 (1859).
Eterusia osseata (9), Walker, xxxi. 120 (1864).
Java, Borneo, Sumatra, Mergui.
From all the above localities in the B. M.; in my collec-
tion from Jelebu, Singapore, Sarawak, and Sandakan ; they
are undoubtedly sexes of the same species. Moore’s type
from Java is in the B. M., Walker’s type from Sumatra in
Mus. Oxon. Guérin’s locality Assam is undoubtedly wrong ;
at all events, it has never since been recorded from India
proper. There is an example, however, in Mus. Oxon., taken
by Mouhot in Cambodia, Siam.
17. Isbarta pieridoides.
Epyrgis pieridoides, Herr.-Schaff. Ausser. Schmett. 1, p. 5 (1853),
Java, Borneo, Sumatra, Nias.
The type came from Java. Only females have heretofore
been known; the male is glaucous blue, about half the size of
the female ; it somewhat resembles Jsbarta glauca, Walker,
has the same kind of yellow band on the abdominal margin
of the hind wings below, but otherwise the markings above
and below are identical with those of the other sex except
for some black suffusion on the outer third of the fore wings
above and white submarginal spots: it has heretofore been
mixed up with g/auca in collections ; the female of that form
is not yet known. There is a form of pierzdoides in Sumatra
and another in the island of Nias, in which the females are
very similar, but the males have the thick black vein-
markings on the hind wings very obscure and more white
spots on the fore wings. There are in the B. M. and in my
own collection examples of both sexes of all three forms.
Eastern and Australian Heterocera. 149
18. Isbarta Binghami.
Epyrgis Binghami (9), Butler, Aun, & Mag. Nat. Hist. (5) x. p. 374
(1882),
Isbarta cyanescens (3), Hmpsn. Trans, Ent. Soc, 1895, p. 284.
Tenasserim, Burma.
There can be no doubt these are sexes of the same species ;
the types of both are in the B. M. from Tenasserim, also
three other females from the same locality ; I have a female
in my own collection from the same place.
19. Isbarta curiosa.
7) vi. p. 305 (1900).
Isbarta curiosa, Swinhoe, Ann, & Mag. Nat. Hist. (7)
33, pl. xiv. fig. 3.
Isbarta aspasia, Snellen, Tijd. yoor Ent. 1902, p. 18
Java.
Types (¢ 2) in B. M. Snellen’s types are from the same
locality.
Genus Psapuis.
Walker, ii. p. 433 (1854).
20. Psaphis camadeva.
Gynautocera camadeva (¢), Doubl. Ann. & Mag. Nat, Hist. xix,
p. 75 (1847).
Euschema semiplena (2), Walker, xxxi, 177 (1864).
Penang, Malacca, Borneo.
Doubleday’s type from Penang is in the B. M.; Walker’s
type without locality, from Coll. Norris, is not in the National
Collection, but there is one example from Malacca and
another from Borneo. Rothschild’s species gloriosa, a female
from Sumatra *, must come under this genus; the male,
when found, will certainly resemble camadeva; there is a
female example from Kuching in the B. M.
Family Callidulide.
21. Cleis aureola, nov.
?. Antenne, palpi, frons, and pectus golden yellow;
head, body, and wings above dark chocolate-brown ; fore
wings with a short submarginal band from near the
hinder margin, and hind wings with a broader marginal
band, golden yellow ; the latter is crescent-shaped and does
not touch the margin. Underside: body and legs golden
yellow ; wings same as above, but the golden-yellow band on
* Noy. Zool. vii. p. 275, pl. v. fig. 6.
150 Col. C. Swinhoe on
the fore wings is complete and circles outwardly from the
costa beyond the middle to near the hinder margin, where it
narrows somewhat.
Expanse of wings 14 inch.
Obi Island ; one example.
Nearest to C. posticalis, Guérin, but that has a large
golden-yellow spot near the hinder angle of the fore wings
above and below, and the band on the hind wings is much
broader.
Family Lymantriide.
22. Anthela epicrypha, nov.
2. Pale cinnamon-grey, with a slight pinkish tinge, very
uniform in colour; indications of an antemedial outwardly
curved grey line; a brownish spot at the end of each cell; a
discal sinuous grey line, immediately followed by a crenulate
grey line contaiing dull spots on the veins, both lines bent
inwards on to the costa; on the hind wings these two lines open
out a little towards the costa, and the inner line corresponds
to the outer line of the fore wings. Underside: paler lines as
above, the inner line on both wings thicker and more even ;
an obscure small spot inside each cell in addition to the
spot at the end.
Expanse of wings 3,5 inches.
New South Wales; one example.
I have had this example in my collection for years ; it is
in fine condition, I have carefully gone through the
descriptions of all the described species, examples of which
are not in the B. M., but cannot find any description that fits
it; the nearest is Darala succinia, Lucas (Proc. Linn. Soc.
N.S. W. 1891, p. 290) ; there is nothing like it in the B. M.
Family Hadenide.
23. Polia illoba.
Agrotis illoba, Butler, Ann, & Mag. Nat. Hist. (5) i. p. 162 (1878).
Polta tlloba, Hmpsn, Phal. y. p. 151 (1905).
Khasia Hills.
Not mentioned by Hampson as from India; the type from
Hakodaté, Japan, is in the B. M.; I haveit from Yokohama
and Nikko.
Eastern and Australian Heterocera. 15h
Family Acontiide.
24. Bryophila khasiana.
Bryophila khasiana, Umpsn. Moths India, ii. p. 299 (1894).
Apamea repetita, Swinh. (nec Butler), Ann. & Mag. Nat. Hist. (7) xv.
p. 499 (1905).
Khasia Hills.
Type in Coll. Elwes. Sir George Hampson informs me
that I misidentified this moth. Since writing the above
paper I have received another example from Cherra Punji.
25. Malattha marginalis.
Acontia marginalis, Walker, Journ. Linn. Soe. vil. p. 49 (1864).
Maliattha marginalis, Swinh. Cat. Het. Mus. Oxon. ii. p. 51, pl. i. fig. 9
(1900).
Matattha stolasa, Swinh. Ann. & Mag. Nat. Hist. (7) xv. p. 154 (1905),
Khasia Hills.
The type is in Mus. Oxon. from Sarawak. I overlooked
it when describing stolasa. It is new to the Indian fauna,
Family Sarrothripide.
26. Blenina solomonis, nov.
¢. Palpi brown above, white below and at the sides ;
head, body, and fore wings white, thickly irrorated with dark
grey: fore wings with basal, medial, and submarginal grey
irregular bands, the first outwardly margined by an outwardly
rounded sinuous brown line ; the medial band margined on
both sides in a similar manner, the band much contracted
below the middle and narrow downwards to the hinder
margin; the submarginal band narrow, bifurcate below the
costa, and outwardly margined by a deeply sinuous thick
brown line, the sinuations forming three blunt outward
dentations: hind wings dark brown, without markings.
Underside uniform sordid brown, with some darker shading
in the interior of the fore wings.
Expanse of wings 1,%; inch.
Solomon Islands ; one exainple.
Somewhat similar in the colour and markings on the fore
wings above to B. lichenopa, Meyrick, which I have from
Queensland, but very different on the underside,
152 Col. C. Swinhoe on
Family Gonopteride.
27. Beara nubiferella.
Beara nubiferella, Walker, xxxy. 1704 (1866).
Port Blair, Andamans.
The type from Java is in the B. M. I have it also from
Muok-Lek, Siam, Goping, Perak, Sarawak, and Mackay,
Queensland. It is not mentioned by Hampson.
Genus Erizapa.
Erizada, Walker, xxxii. 506 (1865).
Tinosoma, Umpsn. Moths India, ii. p. 426 (1894).
28. Erizada semifervens.
Gadirtha (?) semifervens, Walker, Journ. Linn. Soc. vii. p. 163 (1864).
Tinosoma semifervens, Swinh. Cat. Het. Mus. Oxon, ii. p. 101 (1900).
Erizada lichenaria, Walker, xxxii. 507.
Tinosoma hyperythrum, Hmpsn. J. e.
Borneo, Java, Siam, Naga Hills.
The type from Sarawak, Borneo, is in Mus. Oxon., the
types of lichenaria from Java in the B. M., and of hypery-
thrum from the Naga Hills in Coll. Elwes; I have it also
from Siam.
Family Quadrifide.
29. Ophiusa rubida.
Ophisma rulida, Walker, Journ. Linn. Soc. vii. p. 179 (1864).
Ophiusa rubida, Swinh. Cat. Het. Mus. Oxon. ii. p. 144 (1900).
Khasia Hills, Jaintia Hills; three examples.
The type from Sarawak, Borneo, is in Mus. Oxon. I have
it also from Java. It is not mentioned by Hampson.
30. Hypetra tepescens.
Athyrma tepescens, Walker, xiv. 1417 (1858).
Port Blair, Andamans.
The type from Penang is in the B. M.; I have it also
from Goping, Perak, Singapore, and Key Island. It is not
mentioned by Hampson.
31. Oromena noduna, nov.
9. Palpi brown above, ochreous grey beneath; head
and thorax brown, speckled with white; abdomen ochreous
Eastern and Australian [leterocera. 153
grey: fore wings with the ground-colour white, irrorated
and thickly marked in parts with blackish brown ; an ante-
medial sinuous brown line, the space inside it with two
brown marks on the costa, one in the middle and a brown
suffusion on the hinder margin which extends to the middle
of the wing; a deep black lunule at the end of the cell;
postmedial, sinuous, double line, with a white centre; the
outer portion of the wing suffused with blackish brown and
containing discal and submarginal white dentated lines ;
cilia brown, with whitish patches; four whitish spots on the
costa near tlie apex: hind aS ochreous grey; a brown
lunule at the end of the cell, indications of a discal sinuous
grey line, and the outer margin broadly brown. eo
ochreous grey ; a brown lunule at the ‘end of each cell ;
grey sinuous discal line and broad brownish- grey borders to
both wings ; legs brown; tarsi with white bands.
expanse of wings 1 ,‘5 inch.
Ceylon.
Allied to O. reliquenda, Walker, from India.
Genus Drtocoma, noy.
¢. Antenne long, two thirds length of costa, with curved
bristles and cilia; palpi upturned, second joint broad,
uniform in width, moderately hairy, reaching vertex of head,
third one third length of second, smooth and blunt ; thorax
with rather long hairs abdomen smooth, extending a little
beyond the hind wings; a sharp frontal tuft; fore tibiz
with long, thick, brush-like hairs; mid and hind tibiz
smooth, the former with one pair, the latter with two pairs
of spurs ; costa of fore wings straight, arched before apex,
which is subfalcate; outer margin slightly rounded ; hinder
margin curved before base: hind wings with the costa arched,
apex acute, outer margin slightly rounded; anal angle
bluntly acute; the wing more or less triangular; neuration
normal.
-Allied to the genus Azazia, Walker.
32. Delocoma marmorea, nov.
gd. Palpi black on the outer and grey on the inner sides ;
body and wings dark lavender-brown; top of head, outer
third of fore wings, and apical third of hind wings lavender-
grey, giving the insect a peculiar marbled appearance : fore
wings with a brown lunule at the end of the cell; ante-
medial, medial, and postmedial highly sinuous dark brown
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. At
154 On Eastern and Australian [leterocera.
lines ; the medial line running through a dark brown fascia
which crosses the middle of both wings, the third line dentate ;
a submarginal sinuous fascia from the costa before apex to
the hinder angle: hind wings with a dentate but not simuous
discal line and a submarginal sinuous line, which does not
enter the white apical space; cilia of both wings whitish.
Underside brown, smeared with whitish scales; a straight
thick brown line, outwardly edged with white, from the costa
of fore wings one fourth from apex to the abdominal margin
of hind wings a little beyond the middle; indications of a
discal band; a thin, brown, marginal band and whitish cilia.
Expanse of wings 2,4, inches.
Sawangan, N. Celebes ; six examples.
In markings unlike anything I have ever seen.
Family Focillide.
33. Zethes subapicalis, nov.
3. Of a uniform ochreous fawn-colour ; first two joints
of palpi brown; thorax blackish brown in front: fore wings
with a small black patch on costa at the base; a black dot
in the middle of the cell, one at the upper and another at
the lower end; a large rounded subapical patch of greyish-
white scales on the outer margin, with some black suffusion
in its interior, and four black dots marked with white close
to the outer margin within the patch; some very minute
dots on the outer margin hindwards ; indistinct sinuous
transverse lines, medial and postmedial: hind wing with a
black dot at lower end of cell, the discoidal vein faintly
marked with black; a medial, indistinct, sinuous line, a
prominent black spot below its commencement on the outer
margin, also some black points along the margin. Underside
brighter and paler, fhe interior of the fore wings and the
whole of the hind wings whitish, the discoidal marks and
rather prominent, discal, sinuous lines pinkish.
Expanse of wings 1,3, inch.
Silchar, Cachar ; one example.
Family Boarmiida.
34. Thinopteryx preétoraria.
Urapteryx pretoraria, Felder, Reise Nov. pl. cxxii. fig. 18 (1873).
N. Celebes ; five pairs.
Type in Coll. Rothschild ; in ‘ Reise Novara’ it is said to
be a male from the East Indies. Dr. Jordan, in answer to
On a new [River-Crab from Yunnan. 155
my enquiries, has kindly informed me that the type is really
a female. It bears two labels: on one there is only the
name of the collector, “ Lorquin”; on the other in red ink
there is the name of the insect, reference to the plate in
‘Reise Novara,’ and the locality “ Molluk ?” ; but he agrees
with me that as Lorquin collected in the Celebes as well as
in other places, the type probably came from that locality.
It is citron-yellow above and below, and my females exactly
resemble the figure; but the male is very differently coloured,
it is ochreous brown above and black below, with yellow
borders, like a large example of 7. nebulosa, Butler, from
India, but the sexes of that form are alike. There is alsoa
citron-yellow form in India, common in the Khasia Hills,
sexes also alike—Z7’. citrina, Warren, Nov. Zool. i. p. 401.
Family Thyridide.
35. Rhodoneura fallax.
Pharambara fallax, Warren, Ann. & Mag. Nat. Hist. (6) xviii. p. 229
(1896).
Khasia Hills.
The type came from New Guinea. I have it also from
the same locality. They are identical.
XVI.—On a new Species of River-Crab from Yunnan, By
W. T. Catman, D.Sc., British. Museum (Natural History).
Tue British Museum has recently received two specimens of
a new species of river-crab collected by Mr. John Graham
from the lake at Yunnan Fu. The species appears to be
most nearly allied to the genus (or subgenus) Parathelphusa,
though differing remarkably from any species hitherto
described from Asia and resembling in some characters the
African group of species forming the subgenus Acantho-
thelphusa of Ortmann. The lake from which the specimens
were taken is situated at an altitude of 6000 feet above sea-
level and drains into the Yang-tze river-system. No species
of Potamonide appear to have been recorded from this part of
Yunnan, although Wood-Mason described three species of
Potamon from Western Yunnan, a district belonging to the
river-system of the [rawady.
Et
156 Dr. W. T. Calman on a .
Parathelphusa spinescens, sp. 0.
Carapace four fifths as long as broad, the greatest breadth
a little in front of the middle, only slightly convex antero-
posteriorly and from side to side. Surface smooth and
polished, with very minute punctuations. The inter-regional
grooves nearly obsolete, the median portion of the cervical
groove not extending beyond the branchio-cardiac grooves on
each side. The distance between the external orbital angles
is a little less than two thirds of the greatest breadth. ‘The
front is rather steeply deflexed, with the margin slightly
everted, extending a little in front of the external orbital
Fig. 1.
Parathelphusa spinescens, §. Nat. size.
angles as seen from above, with a distinct and moderately
deep median notch. Seen from in front it is convex. The
transverse diameter of the orbit is equal to half the width of
the frontal margin. The frontal margin and the upper orbital
margin are granulated. ‘The lower orbital margin is more
distinctly granulated than the upper; its inner angle is not
produced, but there is a prominent, rounded, internal sub-
orbital lobe. The postfrontal crest is nearly obsolete. It is
only represented by the obscurely defined epigastric lobes
which are very slightly rugose in front and are separated by
a shallow mesogastric furrow, showing only in one specimen
a trace of bifurcation.
The antero-laterei margin is convex and is about as long
|
ou
new River-Crab from Yunnan. Lay
as the postero-lateral. The extra-orbital tooth is flattened,
with a slightly convex and minutely serrate outer edge. It
is followed by five or six teeth, which may be unsymmetrical
on the two sides of the carapace. ‘They are slender, spini-
form, nearly straight, widely separate from each other, and
standing nearly at right angles to the margin. The third or
fourth is the longest, and the last is very small or may be
absent on one side.
The lower surface of the carapace is nearly smooth and the
hepatico-branchial groove is ill-defined.
Fig. 2.
Parathelphusa spinescens. a. Abdomen of male. 6, Third maxilliped.
The ischium of the third maxillipeds (fig. 26) has a well-
marked longitudinal groove. The merus is distinctly broader
than long, the antero-external angle rounded and the anterior
margin slightly convex.
Abdomen of male (fig. 2a) with straight sides, converging
but little to the base of telson, which has nearly the form of an
equilateral triangle, the apex rounded and the sides concave.
Merus of chelipeds with a series of granules on its upper
edge, leading to a subterminal spine; lower edge rounded off,
with some obsolescent granules and a subterminal tubercle.
Carpus bearing two stout unequal spines internally. The
right chela of the male is quite smooth, its height equal to
the length of the fingers and two thirds of its length. he
dactylus is slightly arched, so that the fingers gape a little
wien closed, and is very obscurely furrowed. ‘The left chela,
like both chela of the female, has the dactylus nearly
straight and the fingers about three fourths the length of the
palm. ‘The ambulatory legs are moderately long, the merus
without a spine on the upper edge. ‘he dactylus of the
second pair is equal to the propodus, those of the other pairs
are distinctly shorter, and all are strongly spinulose on the
upper surface.
‘The carapace of the male specimen is 29°5 mm. in length
and 37 mm. in breadth ; that of the female is a little smaller.
The colour (in spirit) is yellowish, and the upper surface
158 On a new River-Crab from Yunnan.
of carapace, chelipeds, and legs is abundantly marbled with
orange-red.
This species is distinguished from all the Asiatic represen-
tatives of the genus hitherto described by the possession of
more than four antero-lateral teeth on the carapace and by
having two spines instead of one on the carpus of the
chelipeds. Further, the antero-lateral teeth are much more
slender and spiniform and their number is variable, whereas
Wood-Mason remarks that in this genus these teeth ‘in
point of number and form are as constant for the several
species as are those of the Portunide” *. In all these points
the new form approaches the African group of species referred
to Parathelphusa by A. Milne-Edwards f, but regarded as
forming a subgenus of Potamon, under the name Acantho-
thelphusa, by Ortmann $. But in some, if not all, of the
African species the merus of the chelipeds has a spine on the
anterior and none on the upper edge, and, further, most of
them have a strong and continuous postfrontal crest. Ac-
cording to Ortmann’s arrangement, the chief character sepa-
rating Parathelphusa from Potamon (with its subgenus
Acanthothelphusa) is the less deflexed front. ‘This, however,
is an ill-defined and variable character, and in some un-
doubted species of Parathelphusa (e. g. P. Dayana), as in the
present form, the front is quite as much deflexed as in many
species of Potamon.
Had the present species been found in Africa there can be
little doubt that most carcinologists would have referred it to
Acanthothelphusa. Nevertheless, I do not believe that it 1s
necessary to assume any special relationship between it and
the African species. ‘The differences separating it from some
of thé other Asiatic species of Parathelphusa are compara-
tively small, and the distance is not great even to some of
those species of Potamon in which the antero-lateral margin
is coarsely granulated, as in P. Atkinsonianum and P. denti-
culatum. 'Vhroughout the whole family of the Potamonidze
the generic and specific characters are often very elusive,
phylogenetic conclusions are more than usually hazardous,
and our knowledge of the group appears to be somewhat
inadequate to bear the weight of the geographical speculations
which Ortmann has recently based upon it.
* Wood-Mason, Ann. & Mag. Nat. Hist. (4) xvii. 1876, p. 120.
t¢ A. Milne-Edwards, Bull. Soc. Philom. Paris, x. 1886, p. 148; and
Ann. Sci. Nat. (7) iv. 1887, p. 140.
{ Ortmann, Zool. Jahrb. Abth. f. Syst. x. 1898, p. 300. It is remark-
able that not only in this, but also in a later paper (Proc. Amer. Phil.
Soe. xli. 1902, p. 800), Ortmann appears to have overlooked the second
of Milne-Edwards’s papers quoted above, in which the African species
are figured.
On new Parasiiic Llymenoptera from Borneo. 159
XVII.—On some new Genera and Species of Parasitic
Hymenoptera from Borneo. By P. Cameron.
Evaniide.
Pristaulacus eruthrocephalus, sp. n.
Black ; the head, pro- and mesothorax, antennal scape,
and fore legs red; wings fuscous violaceous, the hinder
pair paler, almost hyaline behind, the stigma dark testaceous,
the recurrent nervure interstitial. ¢ ?.
Length 18-19 mm.; terebra the same.
Kuching, Borneo (R. Shelford, M.A.).
Claws with four teeth, the apical small; the two
middle teeth longer than the basal. Head smooth, shining,
almost bare. Clypeus depressed at the apex in the centre ;
the sides rounded, narrowed to a point; in the centre is a
narrow tooth as viewed from the front, but it is broad
as seen from the sides. The narrowed basal part of the
prothorax is smooth, impunctate, shining; the rest rugosely
punctured; on the sides, at the base of the dilated apical
part, is a stout tooth. Mesonotum rugosely reticulated.
Scutellum with five stout, clearly separated keels, the apex
being also bordered by one. Postscutellum obliquely raised
in the centre ; smooth, the sides stoutly, irregularly striated.
Basal depression of metanotum crenulated ; the rest of it
coarsely transversely reticulated. Pleure reticulated, the
metapleurz more coarsely than the rest. Sternum irregularly
weakly punctured, the central furrow wide, deep, crenulated.
Abdomen smooth and shining: the second transverse cubital
nervure is roundly curved; the first recurrent nervure is
interstitial, the second received shortly behind the base of
the apical third. Legs long, slender; the tibiz sparsely,
the tarsi thickly spinose. The pedicle of antenne is about
three times longer than thick ; the following joint distinctly
shorter than the next, it and the pedicle together being
nearly as long as the third. The hind coxe are finely
transversely striated, weakly at the base, more strongly
towards the apex. Temples wide, rounded.
Braconidae.
Epyia, gen. nov.
Ilind femora stout, but not dilated, the middle irregularly
serrate ; the hind cox about three times longer than thick ;
160 Mr. P. Cameron on new
the basal joint of hind tarsi not longer than the following
two united; calcaria short, claws cleft. Wings with three
cubital cellules—the radial cellule long, the second cubital
cellule behind shorter than the first, narrowed in front,
the transverse cubital nervures converging in front, straight,
oblique; the recurrent nervure is received near the base of
the apical third of the first cubital cellule. Transverse basal
nervure almost interstitial, anal nervure not interstitial.
Middle lobe of mesonotum clearly raised. Metanotum
areolated. Apex of clypeus transverse. Labrum projecting,
rounded. Lower part of frontal depression bordered by a
curved keel; in the centre isa short stout keel, not reaching
to the middle above. Head cubital; temples wide, occiput
margined. Thorax about four times longer than wide.
First abdominal segment ‘broad, longer than the following
two united. Antenne pilose, the first joint of the flagellum
not much shorter than the second. Palpi very long, the
maxillary reaching to the middle coxve, 5-jointed.
The affinities of this genus are with Huscelinus, also from
Borneo. ‘The latter has the hind femora greaily thickened,
the thorax shorter and broader, and the recurrent nervure is
received in the second, not in the first cubital cellule.
Edyia annulicornis, sp. 0.
Black ; a broad ring on the middle of the antenne; the
four front legs except their coxze, basal half (or nearly) of
hind tibize and their tarsi whitish yellow. Wangs hyaline,
the stigma and nervures black. @.
Length 8 mm.
Kuching, August (R. Shelford, M.A.).
Antennz stout, densely pilose, longer than the body.
Head above the antennz smooth, coarsely rugosely punctured,
almost reticulated below. The depressed base of pronctum
is coarsely, irregularly striated, the raised upper part smooth
and shining. Apex of middle lobe of pronotum coarsely,
irregularly reticulated. Scutellum flat, longish, punctured.
Postscutellum with two square aree at the base and two
longer and deeper ones at the apex. ‘There are three rows
of areze on the metanotum, the central longer and narrower
than the others, the central is the longest, the basal the
shortest ; there is a large, irregular, spiracular area, rugose
and irregularly striated. Metapleurz coarsely, closely reti-
culated ; mesopleurze smooth in the centre, strongly
punctured above and below, as is also the case with the
propleure. First abdominal segment irregularly wrinkled ;
Parasitic Hymenoptera from Borneo. 161
there are two keels on the basal half, bordering the ceutre.
The white ring on the antenne is longer than the basal or
apical part. There are five or six small teeth on the
underside of the hind femora in the middle; they are
widely separated and the middle ones are larger and sharper
than the others.
Neotrimorus ? luteus, sp. n.
Luteous, densely covered with long pale hair ; the antenne,
ocelli, and hind tarsi black ; the wings smoky, the basal third
yellowish hyaline, stigma black. Face rugosely punctured,
the rest of the head smooth and shining. Pronotum in front
closely rugosely punctured ; lower part of pleurze bordered
above by a keel, the centre with some stout irregular strive.
Mesothorax smooth, the pleural lower furrow crenulated.
Metanotum finely reticulated; the reticulations round for
the most part ; the metapleuree much more coarsely, irre-
gularly reticulated. The first and the second abdominal
segment to near the apex closely, longitudinally striated, as is
also the base of the third and fourth segments. 2.
Length 8; terebra 4 mm.
Kuching (R. Shelford, M.A.).
In the ‘Records of the Albany Museum,’ i. p. 168, I
described two species of Braconidee under the name of
Trichiobracon, but gave no generic description ; as I was not
sure but that the species might be referred to Acanthobracon,
Szép., non Kriech. There seems to be considerable
structural differences between the species, and it is probable
that they might be split up into two genera. Trimorus,
Kriech., is regarded by Szépligeti* as identical with his
Acanthobracon, but Kriechbaumer says nothing about the
coxee being toothed. The third longitudinal nervure in the
hind wings may be broadly rounded, or geniculated as it is
in T. rujus, Cam. I give a generic description of the
Bornean species meutioned above :—
Radial cellule long, lanceolate. Transverse median
nervure received before the transverse basal; the recurrent
nervure in the first cubital cellule ; accessory nervure inter-
stitial. Hind wings with a third longitudinal nervure
issuing from the lower part of the transverse prebrachial
nervure and reaching close to the apex of the wing; there is
D>?
a large basal cellule. Head cubital, not margined behind.
* Gen. Insect., Braconidie, p. 64.
162 Mr. P. Cameron on new
Eyes oval; the malar space is nearly as long as them.
Mesonotum 38-lobate. Scutellum flat. Median segment
longer than wide; its apex transverse; it is closely reti-
culated. Abdomen sessile; the basal segments closely
striated ; the suturiform articulation deep. Ovipositor long.
Maxillary palpi long. Base of hind coxe with a large
tooth.
The occiput is sharply margined on the lower side. The
mesosternum is bordered by a distinct furrow. Legs short
and stout; their claws simple. Body and legs densely
covered with long hair. Pterostigma lanceolate; the radius
issues from beyond its middle. The second transverse
furrow on the abdomen is distinct. The centre of the
second segment is raised, clearly separated by furrows, which
shape it into a large, transverse, raised, oval area. Sides of
median segment projecting into a blunt tubercle.
Ichneumonide.
JorpPpinNt.
CNOJOPPA, gen. nov.
Scutellum obliquely raised at the base, the apex with a
sharper, steeper slope; the base as long as broad, flat, the
sides and apex bordered by a raised high keel. Areola
widely separated from the base of the segment, 6-angled,
narrowed towards the apex, which is transverse; it is longer
than wide ; the three basal arew are confluent at the base ;
the sides bear longish teeth. Head cubital, about as wide
as long ; the occiput widely, deeply incised; temples as long
as the eyes. Face and clypeus flat; the latter separated at
the sides only, its apex bluntly rounded, very slightly
projecting in the middle. Mandibles long, thick at the base,
largely projecting downwards on the outer side; on the inner
side are two large, sharp-pointed, widely separated teeth.
Malar space large. Petiole long, narrow; postpetiole dilated,
smooth, impunctate; the second and following segment
punctured, the others smooth ; the ventral keel extends to
the apex of the fifth; there are seven segments, the last
being large. Legs long; the hind tarsi twice the length of
their tibiw. Areolet 5-angled; discocubital nervure with
a minute stump of anervure. Antenne longer than the body,
broadly ringed with white, serrate at the apex. Mesonotum
reticulated in the middle. Fore cox small.
Comes near to /meria, Cam.,and Xenojoppa, Cam. (cf. Ann. -
& Mag. Nat. Hist. 1903, xi. p. 174).
Parasitic Hymenoptera from Borneo. 163
Cenojoppa longitarsis, sp. n.
Black; the face, clypeus, orbits (largely below on the
onter), mandibles, palpi, a narrow line on the pronotum, a
broader one on the lower part of the propleure, tegule, a
mark (longer than wide) on the mesonotum, scutellums,
metanotal spines and the part surrounding them, a broad
line on the lower side of the mesopleure, a large square
mark on the apex of metapleurz, the narrowed basal part of
the first abdominal segments and the apices of all broadly,
yellow. Legs yellow; the hind femora fulvous; the hind
coxee (except for a broad mark in the centre above), apex of
hind trochanters below, base and apex of femora and apex of
tibiz, black. Wings hyaline, the stigma and nervures
black) oS:
Length 12 mm.
Kuching, June (R. Shelford, M.A.).
The space separating the mandibular teeth is broad and
rounded. Mesonotum, except on the reticulated central
apical part, closely, shallowly punctured. Scutellum
impunctate. Metanotum smooth at the base, the rest more
or less strongly transversely striated ; the striz on the apex
strong and curved. Propleurz stoutly striated at the base ;
the oblique keel stout ; the meso- and metapleurz for the
greater part striated, the striz on the latter closer, oblique,
and intertwining. Gastrocceeli deep, shining, more or less
irregularly striated; the part between smooth, shining,
irregularly, finely striated.
DrvsEi4, gen. noy.
Scutellum pyramidal, obliquely raised from the base to
the top, which ends in two stout teeth ; the apex has a much
shorter, steeper slope than the base. Metanotum regularly
areolated ; the sides in the middle with longish stout teeth ;
the areola twice longer than wide; the base rounded, the
apex slightly bent inwardly ; there is a longish basal area,
gradually narrowed towards the apex. Spiracles linear,
narrow. ‘Temples long, obliquely narrowed ; occiput deeply
incised. Clypeus transverse at the apex, its sides broadly
rounded. Labrum projecting. Abdominal petiole long,
slender ; the postpetiole gradually dilated towards the apex.
Areolet narrow, five-angled; discocubital nervure not
broken by a stump of a nervure.
The abdomen is sharply narrowed at the base and apex ;
closely punctured; gastroceli shallow, indistinct ; sheaths
164 Mr. P. Cameron on new
of ovipositor as long as the apical three segments united.
Antennz thickened and dilated beyond the middle. Face
and clypeus flat. Hind tarsi much longer than their tibie.
The abdomen is not much longer than the head and thorax
united. Malar space nearly as long as the antennal scape.
The flat face and clypeus in one line, sharply pointed
abdomen with its long projecting ovipositor, stoutly spined
scutellum and metanotum, and violaceous wings should
distinguish this genus readily.
Druseia quadridentata, sp. n.
Black; the face (except for a line in the centre, which
becomes gradually wider from the narrowed top to the apex),
base of labrum, scutellar spines, apex of scutellum below, and
the apices of the three middle segments of the abdomen
narrowly, pale yellow. Wings fuscous violaceous, highly
iridescent, the posterior paler. Basal three segments of
abdomen red. Legs black ; the greater part of the four front
cox, trochanters, tibize and tarsi, and the calcaria whitish
yellow ; the hind coxe, except at the base below, red ; the
rest of the hind legs black, except for a broad pale yellow
line on the base of the tibie behind. 2.
Length 14 mm.
Kuching, March (R. Shelford, M.A.).
Head smooth. Thorax for the greater part finely punc-
tured ; apical slope of metanotum thickly covered with white
hair; the centre transversely, the sides more strongly
obliquely striated; the plenre for the greater part striated.
Petiole smooth ; its base broadly yellow. Transverse median
nervure almost interstitial.
XORIDINI.
Cyanoxorides rufomaculatus, sp. n.
Black, tinged with blue; the thorax, except the mesonotum,
scutellum, and a line down the centre of metanotum red ;
head black, the face, a line on the inner orbits, a broad one
on the lower half of the outer, whitish yellow; the clypeus,
labrum, and malar space dull reddish; the base of first
abdominal segment, a line, dilated at the base in the centre
of the apex, a longish ovate longitudinal mark in the middle
of the second, the apices of the others narrowly and the
ventral surface, white. Four front legs red, the tarsi and
tibiz pale; the middle tibia infuscated behind; hind legs
black, their coxee red; a narrow band on the base of the
Parasitic HTymenoptera from Borneo. 165
tibiz and the tarsi, white. Wings hyaline; the nervures
and stigma black, the latter white at the base. Scape of
antenne and a broad ring on the middle of flagellum
white. @.
Length of body and ovipositor 18 mm.
Kuching (R. Shelford, M.A.).
Face punctured, covered with white pubescence; the
middle near the top projecting ; the black on the front and
vertex distinctly tinged with blue and violaceous tints.
Palpi pale fulvous. Base of mandibles rufous. Base and
sides of mesonotum closely minutely punctured, the rest
reticulated-striated ; the apex in the centre depressed, marked
with four or five longitudinal striz. Scutellum roundly
convex; its apex striated. Postscutellum with the sides
stoutly keeled; its centre with a stout keel. Metanotum
irregularly transversely striated, the base more finely than
the rest; there is one narrow area, coarsely transversely
striated and extending from the base to the apex of the
segment, it is of equal width; there are three lateral aree,
the basal is the larger, the middle is almost square, the
apical triangular. Basal segments of abdomen finely, closely
punctured ; the apex of the first is bounded laterally by a
curved keel, commencing at the sides and reaching close to
the middle at the apex, the oblique furrow on the sides at
the base extending from the inner to the outer and the
furrow nearer the middle extending from the outer to the
inner side are deep; the similar furrows on the third segment
are wider and not so deep, the space between is more raised.
The upper part of the metapleure is bounded by keels ;
except at the base, it is irregularly reticulated; the lower
half is smooth and bounded by a keel below.
Cyanoxorides ceruleus, sp. 0.
Blue, tinged with purple, the antenne broadly ringed with
white; the penultimate abdominal segment for the greater
part clear white ; the four front legs brownish, tinged with
purple; the hinder purple, with the cox reddish fulvous.
Wings hyaline, a cloud along the transverse basal and
median nervures and a larger one in the radial and cubital
cellules. Head smooth; the face keeled in the centre,
sparsely punctured, the furrow separating it from the
clypeus deep. Thorax smooth. Basal central area on
metanctum small, triangular; the two keels running
from it are close together and lead to a longish area, acutely
pointed at base and apex; there are three lateral are on
166 Mr. P. Cameron on new
the metanotum—the basal is the longest, the apical small,
square, its angles denticulate. Basal three segments of
abdomen closely, longitudinally striated. @.
Length 13; terebra 9 mm.
Kuching (R. Shelford, M.A.).
Body and legs covered, but not thickly, with long hairs.
Recurrent nervure broadly rounded. First abdominal seg-
ment nearly as long as the following two united; it is not
keeled and has an oblique furrow on the sides at the apex.
Spiloxorides erythrocephalus, sp. un.
Black ; the head, except the vertex, and the greater part
of the four front legs red ; antennze densely covered with
longish, stiff hair, black, the basal joints reddish. Wings
hyaline, a cloud along the transverse median and the trans-
verse basal nervures (broadest below), and one on the inner
side of the transverse cubital and recurrent nervures, extending
from the hinder part of the radial cellule into the discoidal ;
stigma and costa fuscous, the nervures black. Face strongly,
transversely striated; the front and vertex smooth. The
black of the thorax has a plumbeous hue; smooth; the
propleuree stoutly striated; the apex of mesonotum stoutly
reticulated. Scutellar fovea deep, divided in the middle by
a stout keel. Sides of scutellum stoutly keeled laterally, as
is also the postscutellum. Metanotum regularly areolated,
the petiolar area three times longer than wide, of equal width
throughout; the areola is wider, more irregular, and also
about three times longer than wide; outside these are two
large arez and there are three lateral, the spiracular being
larger, wider, and more irregular than the others; the
posterior median occupies the apex, is strongly, closely,
transversely striated ; the other arez are more irregularly,
sparsely striated. Metapleurs coarsely reticulated. The
pleural furrow is bordered below, especially at the apex.
First abdominal segment irregularly reticulated, more strongly
in the middle than on the sides ; the base in the centre trans-
versely striated ; the middle segments are closely punctured ;
the depressions more or less striated. The apex of the last
segment is white. ¢.
‘Length 10 mm.
Kuching, October (R. Shelford, MA).
CrYPTINA.
LoraDA, gen. nov.
Areolet small, square. Temples not developed, the large
Parasitic Hymenoptera from Borneo. 167
eyes extending to the edge of the occiput, which is transverse.
Eyes large, parallel, not converging. Parapsidal furrows
distinct. Metathorax spined, with one (the basal) trans-
verse keel. Abdominal petiole broadly dilated at the apex.
Metapleural keel obsolete. Thorax three times longer than
wide. First joint of flagellum slightly longer than the
second. Ovipositor short. Hind legs stout, not greatly
lengthened. Claws long. The spiracles on first abdominal
seement are wider from each other than they are from the
apex. Head not wider than the thorax.
The second transverse cubital nervure is faint; the trans-
verse median nervure is received shortly behind the transverse
basal; the transverse median nervure in hind wings is
angularly broken below the middle. Scutellum roundly
convex, the sides not margined. Metathoracic spiracles
small, oval, not much longer than wide.
Of the Oriental genera of Mesostenini the present most
resembles Melcha, which may be known from it by the longer
and thinner abdominal petiole, which is not abruptly dilated
at the apex and whose spiracles are nearer to each other than
to the apex, by there being a complete metapleural keel and
one above it, and by the longer median segment with two
transverse keels. ‘The Oriental genus Skeatia, Cam., has
some affinity with Lotada; but it may be known by the
longer, narrowed, not square areolet, by the long, linear
metathoracic spiracles, by the head being distinctly wider
than the thorax, and by the scutellum being clearly longer
than wide, which is not the case with the present genus, in
which it is, if anything, wider than long.
Lotada maculiceps, sp. n.
Black, clypeus, mandibles, palpi, an interrupted line on
the inner orbits, malar space, a line on the pronotum,
tegule, tubercles, scutellar keels, scutellum, sides of median
segment, apex of basal two abdominal segments broadly, of
the fourth more narrowly, and the whole of the apical two,
yellow. Legs fulvous red; the four front cox and tro-
chanters pale yellow ; the base of hind tibie narrowly, the
apex more broadly, and apex of tarsi, black. Wings hyaline,
the stigma black. Antenne with a broad white band. @.
Length 8 mm.; ovipositor 1 mm.
Kuching (R. Shelford, M.A.).
Face strongly, closely, rugosely punctured ; upper part of
front closely, finely, obliquely striated. Mesonotum minutely
punctured-aciculated, the sides finely closely striated,
168 On new Parasitic Ilymenoptera from Borneo.
Scutellum smooth. Base of metanotum closely punctured,
more or less striated; the rest strongly, closely reticulated ;
spines broad; the apical lateral edges serrate. Propleure
above obliquely rugosely striated, the lower part with stout,
clearly separated keels; the mesopleuree closely rugosely
reticulated- punctured, more or less striated at the apex ; the
metapleurz closely, strongly, obliquely striated-reticulated.
Postpetiole smooth, the middle segments closely punctured.
OPHIONINI.
TRICHIONOTUS, gen. Nov.
Transverse median nervure in hind wings unbroken.
Transverse median nervure in fore wings received shortly
beyond the transverse basal. Scutellum roundly convex ;
its sides keeled. Mesonotum and mesopleure reticu-
lated, the former without parapsidal furrows. Antenne
as long as the body. Basal joint of hind tarsi longer than
the others united. Apex of clypeus rounded. Thorax densely
pilose. Metanotum reticulated, the reticulations hid by the
dense pile. Basal joint of hind trochanters as long as the
second. Discoidal cellule at the apex twice its width at the
base. Eyes large, slightly converging below. Ovipositor
shorter than the apical 1 two segments of the abdomen united.
Mandibular teeth almost equal. Hinder calearia as long as
the second tarsal joint, clearly longer than the breadth of
the apex of the tibiz. Claws pectinated.
Comes close to Trichomma, which may be known from it
by the hairy eyes, strongly converging below, by the basal
joint of the trochanters being more than double the length
of the second, and by the flatter, more oblique, not reti-
culated scutellum, by the more narrowed at the base brachial
cellule, the transverse discoidal nervure being broken farther
up the middle than it is in the present genus, which has
further strongly pectinated claws, while in Trichomma the
spines are very few, if any. The only other genus with
which it could be confounded is Atrometus, but that has the
discoidal cellule sharp-pointed at the base (cf. Schmiede-
knecht, Zeit. f. Hym. u. Dipt. 1902, p. 361).
Trichionotus reticulatus, sp. n.
Black, the four front legs, hind coxe, apical joint of
trochanters, basal three fourths of hind tibie, and the
calcaria, pale yellow; the mandibles, face (except in the
middle above and below), the inner orbits and the lower
On Mammals from Angolu. 169
fourth of the outer of a bright yellow colour ; the basal half
of the first abdominal segments all round and the apical
below yellow, tinged with fulvous ; the greater part of the
ventral segments, the top and the sides of the middle,
testaceous. ‘Thorax, and especially the median segment,
densely covered with long, pale fulvous pubescence; the
second and following abdominal segments densely covered
with a fulvous pile. Wings hyaline, the stigma testaceous,
the nervures black. ?.
Length 14 mm.; ovipositor 1] mm.
Kuching, March (R. Shelford, M.A.).
Face irregularly rugosely punctured, keeled down the
middle, the lower part with two short, stout keels on either
side of the middle. Clypeus irregularly punctured. Front
with some irregular striz. Centre and sides at the base of
the mesonotum irregularly stoutly reticulated; the sides
raised into a stout keel. Scutellum apparently irregularly
rugose (the sculpture hid by the dense pubescence); the
sides stoutly keeled, and there are two keels on the apical
central half. Median segment coarsely, irregularly rugose.
Upper part of propleure strongly longitudinally striated ;
the mesopleure strongly longitudinally striated all over, the
striz curved and almost forming reticulations in the middle.
Middle tarsi blackish at the apex.
XVIII.—On a Second Collection of Mammals obtained by
Dr. W. J. Ansorge in Angola. By OLDFIELD THOMAS
and Rk. C. WrouGuton.
In the ‘Annals’ for last year * an account was given of the
mammals obtained during 1903 in Northern Angola by
Dr. W. J. Ansorge, and the present paper is based on a
series formed by the same collector during 1904 in the central
part of that country.
Dr. Ansorge was not able to devote so much attention to
mammals as on his previous trip, but the series he obtained
is still a very valuable one and adds considerably to our
knowledge of Angolan mammals. It consists of about 90
specimens belonging to 34 species, of which we have found it
necessary to describe 5 as new, the most notable being the
fine rodent-mole we have named Georychus Ansoryet.
* Ann. & Mag. Nat. Hist. (7) xiii. p. 405 (1904).
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 12
170 =Messrs. O. Thomas and R. C. Wroughton on
1. Galago Monteiri, Bartlett.
g- 15. Ndongo, Benguella.
Very similar in colour to the Nyasan G. Kirkii, Gray.
2. Galago mohoh, Smith.
Benguella.
3. Epomophorus Dobson, Boe.
dé. Kalonga, Bihé. Alt. 6200 feet.
The skull of this interesting species is very different in the
proportions of the palate from that of any other member of
the genus.
4, [hpposiderus caffer, Sund.
9. 58. Caiala.
5. Macroscehdes brachyurus, Boe.
g. 32; 9. 19,33. Caconda, Benguella. Alt. 5700 feet.
6. Macroscelides Schinz’, Noack *.
o. 82,83; 9. 80. Fort Quillenges, Benguella.
7. Crocidura sp.
6. 62.- Caconda.
8. Cynelurus jubatus, Schr.
Benguella.
9. Herpestes Bocaget, sp. n.
=H. gracilis punctulatus, Boc. (nec H. punctulatus, Gray, 1849).
9.17. Caconda, Benguella. Alt. 5700 feet.
This specimen agrees excellently with the description
given by Bocage of his H. g. punctulatus (except that the
head, though dark, is not “presque noire”’), but differs
entirely from H. punctulatus, Gray, the type of which is in
the British Museum. Indeed it much more resembles
HT. ochraceus, Gray. In the true A. punctulatus the speckled
effect produced by the annulation of the individual hairs is
present on the tail both above and below quite to the black
* Macroscelides brachyrhynchus, var. Schinz’, Noack (Zool. Jahrb. 1889,
iy. p. 198).
Mammals from Angola. 171
tip and also on the whole under surface of the body, while in
HZ. ochraceus, as shown by the type, the annulation of the
hairs ceases on the distal half of the tail and is entirely absent
on the belly. The claws, too, are as in ochraceus and in
strong contrast to the much longer, stouter claws of punctu-
latus. In Bocagei the hairs on the cheeks are ringed black
and white, the tips being white; on the crown and nape the
white gives place to brown, while on the back the hairs,
which are 10-15 mm. long, have the basal third or half pale,
then a ring, 3 mm. broad, black, a similar ring pale brown,
and, finally, the tip black; on some of the longer hairs there
is a second black ring with its accompanying brown ring.
This annulation of the hairs is continued on the proximal
third of the upper surface of the tail; the remainder of the
upper surface and the whole under surface of the tail, ex-
cepting the black tip, is chestnut-brown. The under surface
of the body is a reddish buff, the “ roux-orangé uniforme ” of
Bocage’s description ; in ochraceus it is straw-colour or dirty
white.
The skull of Bocaget in size approximates to that of
ochraceus, but in shape it distinctly resembles that of punctu-
latus ; it is narrow and long-snouted, instead of broad and
snub-nosed as in ochraceus, and, moreover, is more delicately
built than in the latter species in all details. The posterior
portion from the parietal suture is unfortunately missing in
the type, but we record the following measurements :—
punctulatus. ochraceus, Bocaget.
Length from parietal suture to mm. mm. mm.
end of nasals............ 38 33 34
Width at the fronto-maxillary
SIGUE SG Gooood aooom oD 12 12 10
Width at the constriction be-
hind the postorbital process. 11 14 12
Length of palate from hen-
SelNOT DT tepsete area crate 38 27 29
Type. 2. B.M. no. 5.5. 9.13. Original number 17.
Taken September 3rd, 1904.
10. Helogale sp.
Young 2. Quillenges.
11. Canis sp. (probably C. adustus, Sund.).
Quite young.
12*
172 Messrs. O. Thomas and R. C. Wroughton on
12. Ictonyx sp.
. Bulu Bulu, Bihé.
The specimen is apparently immature and the skull is
missing.
13. Pecilogale albinucha, Gray.
. 51. Pedreira, Bihé.
+0
14. Funisciurus congicus flavinus, Thos.
. 86, 87,91; 2. 84,92. Quillenges.
. 6. Usolo River, Benguella.
. 1,2. Busolo, Benguella.
. 3. Sand-pits, Benguella.
.4; 9.5. Katenge, Benguella.
. 11. Cabeea de Ladroes, Benguella.
. 10. Eland’s Water, Benguella.
These specimens agree very fairly with the type. The
five specimens from Quillenges, however, show a rather
duller colour and broader lateral black stripe.
Oy Oy OY OY 40 Fy OY
15. Graphiurus Kelleni, Reuvens.
gd. 47; 9.46. Pedreira, Bihé.
We provisionally identify these specimens with @. Kelleni,
described by Reuvens * from Damaraland.
16. Graphiurus angolensis, de Winton.
&. 45. Pedreira, Bihé.
A young individual.
17. Tatera sp.
3. 77,93; 92. 72, 73, 74, 76. Quillenges.
Closely allied to, if not identical with, 7. Lobengula,
de Winton t. Much smaller than 7. valida, Bocage, which
Dr. Ansorge obtained on his previous expedition.
18. Dasymys nudipes, Peters.
@. 14. Kupa, Benguella. Alt. 3000 feet.
19. Arvicanthis pumilio, Sparrm.
gd. 21,24; 2. 16, 27, 63, 64. Caconda.
* ¢ Myoxide,’ 1890, p. 35.
+ Ann. & Mag. Nat. Hist. (7) ii. p. 4 (1898).
Mammals from Angola. 173
Approaching in its dark coloration A. pumilio dilectus,
de Wint., rather than A. p. cinereus, Thos. & Schw.
20. Pelomys campane, Huet.
9.9. Eland’s Water.
?. 12,13. Kupa, Benguella. Alt. 3000 feet.
21. Pelomys frater, Thos.
3d. 52. Pedreira, Bihé,
22. Dendromus Ansorget, sp. n.
dé. 89. Caiala, Bihé.
9. 23. Caconda.
A small species without dorsal stripe. Hair of the back
soft and long (7-8 mm.), shorter and more sparse on the belly.
General colour of upperside of body near ‘ ochraceous buff”
(Ridgway), but the long hair, which is slaty at the base and
fulvous terminally, is easily disarranged, and the animal then
looks darker and duller in colour than it really is. Under
surface only differing from the upper in being rather paler.
Face of the same colour as the body, cheeks rather paler, chin
and throat white. Ears large, only very sparsely covered
with short hairs. Tail long and slender, dark above, pale
below.
Skull short and broad, the orbit very broad anteriorly, so
that the zygomata are parallel in their whole length.
Dimensions of type (measured in the flesh) :—
Head and body 60 mm.; tail 72 ; hind foot 17; ear 10,
Skull: greatest length 19; basilar length 11°5; greatest
breadth 10; nasals, length 7; interorbital breadth 3;
diastema 4°5; length of upper molar series 3:2.
Hab. Caconda, Benguella.
Type. 2. B.M. no. 5. 5. 9. 52. Original number 23.
Collected September 7th, 1904. A second specimen was taken
at Caiala, Bihé, alt. 5600 feet, on the 17th October,
This species most resembles D. messorius, ‘Thos.*, from
the Cameroons. It differs from that species by its smaller
size, longer looser coat, and by the coloration of its under
surface, which is little lighter than the upper, instead of
being nearly white.
23. Dendromus sp.
6. 54. Pedreira, Bihé.
* Ann, & Mag, Nat, Hist. (7) xii. p. 340 (1903).
174 Messrs. O. Thomas and R. C. Wroughton on
This specimen is quite young, but belongs to a form with
black dorsal stripe.
24. Steatomys Bocaget, Thos.
g. 40. Bingondo, Bile.
9. 57. Chiffamba, Bihé. Alt. 4900 feet.
25. Steatomys minutus, sp. n.
3. 81. Fort Quillenges, Benguella.
Smaller than any form of Steatomys as yet described.
Hairs of the back soft, 5-6 mm. long. General colour of
upper surface of the body near “hair-brown” (Ridgway),
paling to drab on the sides ; a sharp line marking the change
to the pure white of the underside. Individual hairs of the
back slate-coloured at their bases, paling to buff and tipped
with black; those of the belly white to their bases. Face
coloured like the back, cheeks and area behind the eyes to the
base of the ears paler. Chin, lips, and “ whisker-patch”
white. Ears moderate, a tuft of fulvous hairs at their base
in front ; a narrow white margin to the conch behind. Fore
legs white, as are also the inner side of the thigh and hind
feet. Tail short, dark above, pale below, especially at its
base.
Dimensions (those of the body taken in the flesh) :—
Head and body 85 mm.; tail 33 ; hind foot 13 ; ear 12.
Skull: greatest length 21:5; nasals 8; interorbital
breadth 4; greatest breadth 11:5; basilar length 17;
diastema 6; palatal foramina 4°5 ; upper molar row 3:4.
Hab. Quillenges, Benguella.
Type. &. B.M. no. 5. 5. 9. 56. Original number 81.
Taken January 11th, 1905.
The nearest ally of this little species seems to be S. pent-
onyx, Scl.*, from the Cape Peninsula.
26. Mus rattus, L.
g. 66; ¢. 65. Caconda, Benguella.
27. Mus. sp. (multimammate).
3. 41. Bingondo, Bihé. Alt. 5100 feet.
g. 48. Pedreira. Bihé. Alt. 5200 feet.
3S. 28,31; 2.29. Caconda, Benguella. Alt. 5700 feet.
* Malacothrix pentonyx, W. Sclat. Ann, 8. Afr. Mus. i. p. 202 (1899) ;
Mamm. 8S. Afr, i. p. 36 (1901).
Mammals from Angola. 17
Or
28. Leggada minutoides, Smith.
$. 55; 9%. 44,49, 50,56. Pedreira, Bihé. Alt. 5200
feet.
¢@. 61. Caconda, Benguella.
9.7. Usolo River, Benguella.
29. Cricetomys Ansorger, Thos.
d. 67; ¢. 20,60. Caconda, Benguella.
30. Georychus Ansorget, sp. n.
3&6. KukemaR., Bihé. Alt.5900 feet. 5th October, 1904.
BME novo). 97.74. . Type.
A very large species of a buffy-yellowish colour, with short
velvety fur.
Size very large, the skull decidedly larger than that of
G. Mechowi, the largest species of the genus hitherto described.
Fur close, short, and crisp; hairs of back about 7 mm. in
length. A few scattered longer bristles present on sides and
rump. General colour above uniform buffy (between buff
and clay-colour of Ridgway), the hairs whitish at their
extreme bases, then dull slaty, then buffy, their extreme tips
inconspicuously darker. Under surface dull greyish white,
with a silvery gloss, not defined at all from the colour of
the sides, the hairs whitish to their roots. Head like body,
but the muzzle darker, with soiled patches on each side of
the mouth, as in G. Mechowi. Upper surface of hands and
feet almost naked, the few hairs whitish. ‘Tail-hairs long,
dull whitish ; extreme tip of tail naked, warty, but whether
this is natural or due to accident we are not able to say.
Skull as compared with that of G. Mechowi decidedly
longer, although in the single specimen the basilar suture is
not closed, while it is in the two adult specimens of G. Mechow?
before us. Nasals broad and heavy. Supraorbital region
rounded, broad, and expanded, the region just behind the
rudimentary postorbital processes markedly inflated; as a
result the greatest interorbital breadth is greater than the
least breadth across the brain-case above the meatus, the
converse being the case in G. Mechow?. Zygomata broadly
expanded, thicker and heavier than in the allied species.
Anteorbital foramina about equally broad and high. Palatal
foramina minute, situated about a quarter of an inch in front
of the anterior cheek-tooth (close to it in G, Mechow?).
Posterior palate short, parallel-sided. Bulle with the part
176 = Messrs. O. Thomas and R. C. Wroughton on
which appears on the occipital surface narrow and high, not
broadly rounded. Lower jaw with its angular region far
broader than in G. Mechowi.
Teeth large and heavy, the incisors very broad.
Dimensions of the type (measured in the flesh) :—
Head and body 260 mm.; tail 24; hind foot, s. u. 40,
c. u. 44,
Skull: greatest length in middle line 58; __ basilar
length 49; zygomatic breadth 43°5; nasals 23°3 x 7:4;
interorbital breadth 19; breadth across postorbital processes
20°6 ; intertemporal constriction 11°3; least breadth above
meatus 19; palate length 37; diastema 22; combined
breadth of two upper incisors 8; height of angular region
of lower jaw, from top of condyle, 27.
Hab. and Type as above.
This fine species may be readily distinguished from
G. Mechowt, Peters, its only near ally, by its larger size,
more inflated interorbital region, and the other details above
recorded. The type of G. Mechowi came from Malanje in
Northern Angola, quite near to Duque de Braganza, whence
Dr. Ansorge obtained in his previous collection the examples
we have used in making the above comparisons.
31. Georychus sp.
6.43; 2. 53. Pedreira, Bihé, 5200 feet.
3-71; ¢. 70. Salamunjamba, Quillenges, Benguella.
9. 34. Bunlhe, Benguella, 6200 feet.
These specimens are smaller than the type of G. Bocogez,
de Wint.*, which is, however, a very old individual.
32. Lepus angolensis, Thos.
gd. 59. Chiyuka, Bihé. Alt. 6000 feet.
33. Lepus Ansorget, sp. n.
a. 37. Caiala, Bihé.
6. 35 (young). Chingwari.
A somewhat larger and more brightly coloured animal than
Lepus angolensis, ‘Thos.
Underfur copious; hair rather harsh, about 20 mm. lonz on
the back. General colour above “ wood-brown ” (Ridgway),
irregularly mottled with dark brown (so dark as to seem black
* Ann. & Mag. Nat. Hist. (6) xx. p. 323 (1897).
Mammals from Angola. Aig
in most lights), the mottling almost absent on the sides.
Individual hairs pale for their basal halves, blackish brown
above with a bright pale buff subterminal ring; underfur
pale slate-colour at its base, merging into a pinkish buff
(near ecru-drab of Ridgway) and ending in a black tip.
Below, the belly and chest are white; the hairs white to
their bases. Face and crown coloured like the back, but the
mottling finer; an obsolescent white frontal patch. Upper
lip and base of whiskers buffy brown. Chin white ; the
hairs with slaty bases. Ears shorter than in angolensis ;
anterior half of their outer surface, for its basal half, coloured
like the head, darker at the tip, at the back of which is a
small black patch; fringe dull buffy, changing to black
about an inch from the tip; posterior edge narrowly edged
with white. Nuchal patch very large, extending along the
median line of the back as far as the points of the ears when
these are laid back. Throat clothed with long hair (27-
30 mm.) forming a cinnamon-coloured ruff (the individual
hairs with short black tips). Fore legs pale brown on the
outer, white on the inner side. Inner side of hind legs white,
a distinct black patch down the centre of the back of the
thigh. ‘Tail black above, white below.
Skull, as compared with that of ZL. angolensis, longer and
narrower, with much more lightly built postorbital wings, of
which the anterior and posterior processes are unusually
slender. Projecting “shoulders” of anterior zygoma-root
much less developed than in the allied species. Bulle
markedly smaller, their antero-posterior diameter 10 mm. as
compared with 12 mm, in the fresh specimen of L. angolensis.
Incisors with the same additional enamel line as described
by Thomas in ZL. angolensis: this line is no doubt present
more often than has been recorded.
Dimensions of the type (the external ones taken by
collector) :—
Head and body 460 mm.; tail 75; hind foot 106; ear 111.
Skull: greatest length 87; basilar length 67; nasals,
diagonal length 388; interorbital breadth 17; greatest
breadth 40; diastema 25; length of upper molar tooth-
row 15.
Hab. Caiala, Bihé. Alt. 6000 feet.
Type. 3. BM. no, 5. 5.9.80. Original number 37.
Collected October 8th, 1904.
This species differs from L. angolensis, Thos., which is also
found in Bihé, by its more rufous coloration, shorter ears,
Jarge nuchal patch, and by the skull-characters mentioned
above. The only other hare described from Angola is
178 Mr. G. A. Boulenger on
L. Sale, Jentink *, from Mossamedes; but from the description
this seems to be a more brightly coloured animal with a much
shorter tail, and a skull smaller in all dimensions except
breadth.
34. Cephalophus Grimmiui, L.
ZG bike:
XIX.— Descriptions of Three new Snakes discovered in South
Arabia by Mr. G. W. Bury. By G. A. BouLenGcer,
F.R.S.
Glauconia Burit.
Head not broader than neck; snout rounded; rostral
broad, more than half the width of the head, extending to
between the eyes; nasal completely divided, in contact with
prefrontal; supraocular present, a little larger than the
frontal; no enlarged parietal; ocular bordering the lip,
between two labials, the first of which is very small and
scarcely deeper than broad. 14 scales round the body.
Diameter of body 52 times in the total length, length of tail
152 times. Caudal spine strong. Pale brown above, yellow
beneath.
Total length 210 mm. ; tail 15.
A single specimen from El Kubar, Upper Hushabi, near
Yemen frontier.
This species is allied to G. Blanfordii, Blgr., from Sind
and Baluchistan, and G. Nursi?, Anders., from Aden. It
differs from both in the much broader rostral and in the
absence of enlarged parietal shields.
Zamenis vartabilis.
Head short; snout truncate, strongly projecting. Rostral
a little broader than deep, the portion visible from above
measuring one third or two fifths its distance from the
frontal; internasals as long as or a little shorter than the
prefrontals ; frontal much broader than the supraocular, once
and one fourth to once and one third as long as broad, longer
than its distance from the end of the snout, shorter than the
parietals ; loreal longer than deep ; one przocular, forming a
* Notes Leyd. Mus. ii. 1880, p. 57.
Three new Snakes from South Arabia, 179
suture with the frontal, with a subocular below it; two
postoculars ; temporals 2+2 or 2+3; eight upper labials,
fourth and fifth entering the eye ; four or five lower labials
in contact with the anterior chin-shields; posterior chin-
shields as long as the anterior and separated from each other
by two series of scales. Scales smooth, in 17 rows. Ventrals
155-169 ; anal divided ; subcaudals 80-90.
Total length 380 mm. ; tail 110.
The specimens at hand, ten in number, exhibit a surprising
amount of colour-variation. They may be arranged under
seven heads :—
A. Olive-grey above, with a dorsal series of large, brown, dark-edged,
transverse spots and a lateral series of similar spots alternating with
them ; two dark bars on the head, the first across the eyes, the
second across the temples and the parietal shields; a third bar
across the nape; lower parts white.
dg. V.157. OC. 88. El Kubar.
@. V.162. C. 84. Schaf Ravine.
. Similar to the preceding in the markings, but reddish brown, with a
bright red vertebral stripe; ventral shields spotted with red on the
sides.
ce. ¢. V.157. C.? Schaf Ravine.
C. Body orange, with a bright red vertebral stripe, the scales and shields
edged with brown; no spots; head olive-grey above, the temples
and nape black.
Q. V.161. C.? El Kubar.
loop | bea
jo ag
. Scales brown or black, with yellowish or pale olive centres ; ventral
and subcaudal shields whitish, edged with brown or blackish; head
olive-grey in front, black behind.
et. ¢ (V. 159, 157; C. 83, 83) & Q (V. 165, 169, 165; C. 80, 89, 82).
El Kubar.
E. Olive-grey above, the scales edged with blackish ; whitish or grey
beneath, the shields edged with dark grey or blackish.
k-l. g. V. 155,161. C.?, 90. El Kubar.
F, Black, scales and shields speckled with white ; head and nape olive-
grey.
m. 9. V.161, C.? El Kubar,
G. Black above, dark grey below; throat and anterior ventral shields
white.
n. 9. V.168. C. 82. El Kubar,
In the low number of its ventral shields Z. variabilis agrees
with Z. brevis, Blgr., from Somaliland, which differs in the
less prominent snout, the smaller rostral shield, the narrower
frontal, the upper labials nine in number, and the scales in
19 rows.
180 Mr. G. A. Boulenger on new
Atractaspis Andersontt,
Snout very short. Portion of rostral seen from above
nearly as long as its distance from the frontal; suture
between the internasals about half as long as that between
the prefrontals ; frontal large, once and one fifth to once and
one fourth as long as broad, much longer than its distance
from the end of the snout, as long as the parietals; one pre-
and one postocular ; temporals scale-like, 2-+3 or 3+4; six
upper labials, third and fourth entering the eye, fourth
largest ; first lower labial forming a suture with its fellow
behind the symphysial; three lower labials in contact with
the chin-shields. Scales in 23 or 25 rows. Ventrals 219-
243 ; anal entire ; subcaudals 28-29, single. Uniform black.
Total length 550 mm. ; tail 45.
Five specimens from E] Kubar.
This species, named in memory of Dr. John Anderson, the
author of several contributions to Arabian herpetology, is
closely allied to A. lewcomelas, Blgr., trom Somaliland, from
which it differs in the broader frontal shield.
In addition to the new snakes here described, Mr. Bury’s
collection contains examples of the following species new to
Arabia:—Boodon lineatus, D. & B., Dasypeltis scabra, L.,
and Nata hate, L.
XX.—Descriptions of new Tarlless Batrachians in the
Collection of the British Museum. By G. A. BOULENGER,
F.R.S.
Arthroleptis Seimundi.
Tongue with a conical median papilla. Head broader than
long ; snout short, broadly rounded, not prominent ; canthus
rostralis distinct ; loreal region slightly concave ; interorbital
space much broader than the upper eyelid ; tympanum very
distinct, two thirds the diameter of the eye. Fingers slender,
with swollen tips, third very long, nearly twice as long as
second, first and second equal ; toes slender, with swollen tips
and arudiment of web; arather large, oval, compressed inner
metatarsal tubercle ; no outer metatarsal tubercle ; no tarsal.
tubercle. The tibio-tarsal articulation reaches the eye. Skin
smooth, granular on the lower belly and on the back of the
thighs. Dark brown above, sides speckled with white; a light
Tailless Batrachians tn the British Museum. 181
line on theeanthus rostralis; loreal and temporal regions black;
a white streak, formed of numerous small spots, from below the
eye to the fore limb, and prolonged on the upper surface of
the latter to a little beyond the elbow ; throat blackish, breast
and limbs dark brown dotted with white, belly and lower
surface of thighs brownish white.
From snout to vent 36 mm.
A single specimen from Fernando Po, collected by Mr. E.
Seimund during the expedition carried out under the auspices
of the Duke of Bedford, the Hon. Walter Rothschild, and
Mrs. Percy Sladen.
This species is intermediate between A. macrodactylus,
Blegr., and A. variabilis, Matsch.
Atelopus tumifrons.
Habit stout, toad-like. Head a little broader than long;
snout very short, not projecting beyond the mouth, above
with a strong rounded swelling extending to between the
eyes ; interorbital space nearly twice as broad as the upper
eyelid. Fingers moderate, with a slight rudiment of web,
first a little shorter than second; toes short, barely half-
webbed ; subarticular tubercles distinct, some double; two
small metatarsal tubercles. The tibio-tarsal articulation
reaches the shoulder. Skin warty, spinulose; the largest
warts, at the angles of the mouth and around tie vent, conical.
Blackish ; sides of snout, tips of digits, lower surface of hands
and feet and of arms, three spots on the chin and throat, and
a large blotch covering the lower belly and the lower side of
the thighs yellow.
From snout to vent 29 mm.
A single female specimen from Pernambuco, presented by
Dr. F. Werner.
Closely related to A. Stelznerit, Weyenb.
OocoRMUS, gen, nov. Cistignathidarum.
Pupil horizontal. Tongue circular, entire, free behind.
Vomerine teeth in a long transverse series behind the choane.
Tympanum hidden under the skin. Fingers and toes free,
the tips not dilated. Outer metatarsals bound together.
Sternum without bony style. Habit stout, Engystoma-like.
Oocorm us m icrops.
Vomerine teeth forming a very open chevron, which is
searcely interrupted in the middle and the branches of which
182 Mr. G. A. Boulenger on new
are slightly curved. Tongue large, covering the whole floor
of the mouth. Head much broader than long; snout very
short, broadly rounded, not projecting beyond the mouth;
eye very small, as large as the tympanum ; interorbital width
more than twice as great as the width of the upper eyelid.
Fingers short, slightly swollen at the end, first and second
equal; subarticular tubercles well developed. ‘Toes short,
perfectly free, not margined, with slightly swollen tips and
strong subarticular tubercles; a large oval inner and asmaller
rounded outer metatarsal tubercle. The tibio-tarsal articula-
tion reaches the tympanum. Skin smooth and shiny; a
strong fold from the eye to the shoulder, forming an angle
behind the tympanum; a feebly marked glandular fold on
each side from the upper eyelid to the sacral region, con-
verging towards its fellow on the scapular region, where it
forms an angle. Brown above, with symmetrical darker
markings, viz. a chevron pointing backwards between the
eyes, an angular band on the outer side of the dorsal glandular
fold, a M-shaped band on the posterior part of the back, an
oblique band on the side, and cross-bars on the limbs; dark
brown beneath.
From snout to vent 34 mm.
Four specimens from the Organ Mountains, Brazil,
presented by Messrs. Wagner Brothers.
Hyla Steinbachi.
Tongue circular, entire or indistinctly nicked, slightly free
behind. Vomerine teeth on a level with the posterior borders
of the very large choane, in two slightly curved oblique
series forming a chevron pointing forwards. Head moderate,
as long as broad; snout rounded, as long as the orbit;
canthus rostralis feeble, slightly curved; loreal region oblique,
concave ; nostril near the tip of the snout ; interorbital region
as broad as the upper eyelid; tympanum distinct, three fifths
the diameter of the eye. Outer fingers with a rudiment of
web ; no distinct rudiment of pollex; toes two-thirds webbed ;
disks barely half the diameter of the eye; subarticular
tubercles well developed ; no tarsal fold. 'Tibio-tarsal articu-
lation reaching a little beyond the tip of the snout. Skin
smooth above, granular on the belly and under the thighs;
heel with a pointed dermal appendage. Pale pinkish grey
above, with scattered darker dots and rather indistinct wavy
longitudinal bands ; a purplish-brown vertebral line, and
another line of the same colour on the canthus rostralis and
Tailless Batrachians tn the British Museum. 183
temple, sometimes continued along the side of the body;
limbs with ill-defined dark cross-bars ; lower parts white.
From snout to vent 35 mm.
Three specimens from the Province Sara, Department
Santa Cruz de la Sierra, Bolivia, collected by Hr. J.
Steinbach.
Most nearly allied to H. boans, Daud., and H. Spegazzintt,
Bler.
Hyla platydactyla.
Tongue subcircular, entire, slightly free behind. Vomerine
teeth just behind the level of the moderately Jarge choana, in
two slightly oblique series converging forwards. Head
moderate, much depressed, a little broader than long; snout
rounded, hardly as long as the orbit; canthus rostralis
distinct ; loreal region very oblique, concave; nostril near
the tip of the snout; interorbital region a little broader than
the upper eyelid ; tympanum rather indistinct, one third the
diameter of the eye. Fingers much flattened, with a short
basal web which extends as a fringe to the disks; toes short,
much flattened, with a deeply notched web extending to the
disks ; latter about half the diameter of the eye; subarticular
tubercles feeble; no tarsal fold. Tibio-tarsal articulation
reaching the eye. Skin smooth above, granular on the belly
and under the thighs. Purplish above, closely dotted with
darker ; upper surface of arms and thighs white, with a
median stripe of closely set dark purple dots; lower parts
white. Male with a feebly developed subgular vocal sac.
From snout to vent 32 mm.
A single male specimen from Merida, Andes of Venezuela,
collected by Sefior Bricefio.
Allied to H, Vilsoniana, Cope (creolica, Werner).
Hyla graminea.
Tongue heart-shaped, free behind. Vomerine teeth in two
widely separated short transverse groups between the very
large choane. Head rather large, much depressed, a little
broader than long; snout rounded, as long as the orbit ;
canthus rostralis indistinct; loreal region very oblique,
slightly concave ; nostril near the end of the snout; inter-
orbital space twice as broad as the upper eyelid; tympanum
very distinct, three fourths the diameter of the eye. Fingers
much flattened, nearly entirely webbed; toes also much
flattened and entirely webbed; disks nearly as large as the
1S4 Mr. G. B. Sowerby on new
tympanum ; subarticular tubercles feeble; an outer tarsal
fold. Tibio-tarsal articulation reaching the tip of the snout.
Skin smooth, finely granulate on the head, coarsely on the
belly and under the thighs; deep wrinkles form an areolation
on each side of the anterior part of the body. Bright green
above, yellowish white beneath; hand and arm, tarsus and
foot, and upper eyelid with a white edge ; arm and thigh
colourless, the latter with a narrow green band in the middle.
From snout to vent 52 mm.
A single specimen from Northern British New Guinea,
altitude 900 feet.
ie nearly allied to H. gracilenta, Peters, and H. chloris,
gr.
XXI.—Deseriptions of new Species of Marine Shells, chiefly
from Ceylon. By G. B. Sowersy, F.L.S.
THE shells here described formed part of the collection of the
late Mr. Hugh Nevill. As very few of them were labelled,
it is impossible to fix their habitats with any certainty, but
the probability is that most, if not all, of them are from
Ceylon.
Fam. Trochide.
Calliostoma planiliratum, sp. n.
Testa conica, elata, angustissime rimata, albida, rufo-fusco et olivaceo
dense strigata et variegata; anfractus 83, primi 2 leves, rotun-
dati, deinde planato declives, liris 7 confertis planulatis cingulati,
cingulis duobus infernis magis elevatis; anfractus ultimus levis-
sime concavus, liris duplicatis; basis planulata, vix convexa,
liris 9, angustis, rotundatis, albo-fusco articulatis, interstitiis
oblique striatis; apertura oblique subtrigona, intus margari-
tacea, haud lirata; columella brevis, rectiuscula, leviter callosa.
Alt. 13, diam. 10 mm.
A high conical shell with an almost flat base. It has very
close flattened spiral ridges, which in the last whorl are some-
times, as in the type, divided by a groove. The colouring
of olivaceous brown, reddish here and there, is mostly dis-
posed in close broad longitudinal streaks, which on the upper
whorls are broken up into square spots, giving the ridges an
articulated appearance.
Although of a not uncommon form, I know of no species
Species of Marine Shells. 185
to which this shell bears a sufficiently close resemblance for
comparison.
In writing the diagnosis of the type specimen I have said
that the lire: of the last whorl are duplicated, or divided by a
groove, but I find this is not a constant character.
Calliostoma Nevilli, sp. n.
Testa conica, solidiuscula, imperforata, albida, fulvo radiata ; spira
elatiuscula, acuta, vix convexa; anfractus 7, planato declives ;
liris 7, confertis conspicue granatis spiraliter cingulati; inter-
stitiis angustis, fulvis; sutura inconspicua; anfractus ultimus
curtus, obtuse angulatus, supra angulum levissime convexus;
basis planato convexa, liris granatis 8 instructa ; columella brevis,
crassa ; apertura oblique subquadrata, intus levis ; labrum crenu-
latum.
Alt. 7?, maj. diam. 7 mm.
A regularly conical shell, the whole surface of which is
covered with closely and beautifully beaded spiral ridges,
The base is not so flat as in the preceding species, and the
spire less elevated.
The shell bears some resemblance to C. scobinatum (Zizi-
phinus scobinatus, Reeve, as of A. Adams), but it is more
broadly conical, and the bead-like pustules are larger and
closer together. Compared with C. fragum, Philippi, it is
also broader, and the beaded ridges are closer together and
more numerous, with very narrow interstices.
Clanculus crassilabrum, sp. n.
Testa turbinata, subovalis, anguste perforata, crassa, albida, maculis
paucis pallide fulvis conspersa; spira elata, leviter convexa;
anfractus 44, primus levis, rotundatus, sequentes convexi, rotun-
dati, dense grano-lirati; liris 8, crassis, pustulis numerosis con-
fertis gemmuliformibus ornatis ; interstitiis angustis, crenulatis;
sutura inepte canaliculata; anfractus ultimus rotundatus, liris
pustulosis circ. 15 instructus ; apertura rotundata; labrum
crassum, crenulatum, intus valde liratum ; columella breviter
expansa, planulata, bidentata, dente postico late rotundato,
antico oblique elongatim producto.
Alt. 7, diam. 64 mm.
A shell of an unusually oval form for the genus, approach-
ing some of the forms of Huchelus, the surface being similar
to that of the much larger Huchelus baccatus, Menke. It has
a much smaller umbilicus than is common in the genus
Clanculus, and is further distinguished by its narrow
obliquely-produced anterior columellar tooth.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 13
156 Mr. G. B. Sowerby on new
Clanculus acutidentatus, sp. 1.
Testa breviter conoidea, profunde umbilicata, rufo-fusca, albo macu-
lata; spira late conica, convexiuscula ; anfractus 5, convexi, liris
spiralibus 5 crenulato-pustulatis cingulati; interstitiis latiuseulis,
oblique striatis; sutura canaliculata; anfractus ultimus brevis,
latiusculus ; basis planato conyexa, liris 9 rugosis instructa;
umbilicus angustus sed profundus, ad marginem crenulatus;
columella obliqua, ad basin acute unidentata; apertura postice
quadrata, antice rotundata, intus lirata; labrum crassiusculum,
crenulatum.
Alt. 5, maj. diam. 6 mm.
Compared with C. ceylanicus, Nevill, this shell is broader
in proportion to its height, and is distinguished from that
species, to which in other respects it is somewhat similar, by
its sharp narrow columellar tooth.
Euchelus cavernosus, sp. n.
Testa globoso-turbinata, crassiuscula, anguste umbilicata, albida;
spira obtusiuscula, convexa; anfractus 5, apicales 2 leves, pel-
lucens, sequentes convexl, tricarinati; carinis rctundatis, nodoso
crenulatis ; interstitiis profundis, liris validis, decussatis ; sutura
canaliculata; anfractus ultimus amplus, rotundatus, carinis 7,
nodulosis, interstitiis profunde foveolatis; apertura subcircularis ;
columella arcuata, levis.
Alt. 73, diam. 74 mm.
A small globular white shell, very conspicuously and _pro-
foundly pitted between the rounded nodulous keels.
The species is similar to #. fuveolatus, A. Adams, but
larger, more globose, and with more nodulous lira.
Omphalius eusculptus, sp. n.
Testa depresse orbicularis, rubro-carnea, fuseo maculata, ad apicem
alba; spira paulo elevata; antractus 43, primi 2 leves, deinde
tricarinati, carinis rotundatis, crenulatis; interstitiis latiusculis,
spiraliter anguste triliratis, eximie oblique decussatim striatis ;
sutura canaliculata; basis convexiuscula, sex-lirata; umbilicus
rotundus, profundus, mediocriter latus; apertura oblique sub-
quadrata, intus plicata, labrum crenulatum ; columella obliqua,
levis.
Alt. 6, maj. diam. 84 mm.
A pretty pink shell with brown spots and a white apex.
The sculpture between the keels is very fine, minute, close-
set, raised, slightly granular strize obliquely traversing three
narrow spiral ridges.
Species of Marine Shells, 187
The shell rather closely resembles T'rochus plebetus, Plil.
(= Clanculus nodiliratus, A. Adams), the principal difference
being that the margin of the umbilicus is smooth, not nodu-
lous or dentate.
Minolia eucoronata, sp. 1.
Testa depresse orbicularis, tenuis, late umbilicata, fulvescens, fusco
strigata ; spira obtusa, vix elevata ; anfractus 31, concavi, bicari-
nati; carinis eximie tuberculosis, sutura indistincta; anfractus
ultimus depressus, granoso triliratus, ad peripheriam carina acuta
crenulata vix nodulosa ; basis convexa, extus levis, area mediana
quadrisuleata et radiatim plicata; umbilicus latus et profundus,
circularis, ad marginem tuberculatus,
Alt. 27, diam. 42 mm.
A little depressed shell, with two beautifully beaded keels
to each whorl, the last whorl exhibiting 3 granular lire
between the keels, the margin of the umbilicus being strongly
nodulous.
I know of no species with which to compare this beautiful
little shell.
Ethalia Nevill, sp. n.
Testa depressa, circularis, alba, pellucida, perforata ; spira convexo-
depressa ; anfractus 5, primus minutissimus, ceeter] leviter con-
vexi, spiraliter minute striati; penultimus puncturatus ; ultimus
striis partim fere obsoletis sculptus, ad peripheriam rotundatus ;
basis convexa, spiraliter leviter striata, striis in regione umbili-
cali magis conspicuis et confertis; umbilicus parvus, circularis,
profundus ; apertura suboblique ovalis, intus levis, peristoma
simplex ; columella brevis, oblique arcuata, leviter incrassata,
Alt. 2, diam. 44 mm.
This species appears to belong to the little group which,
according to Pilsbry, is allied to Te¢nostoma, and erroneously
referred to Hthalia. In the absence of further evidence than
that furnished by the shell, I think it best, however, to place
it in the latter genus. It may belong to Fischer’s subgenus
Pseudorotella, although the umbilicus is open, and not covered
by a callus as in the type of that group (2%. semistriata, Orb.).
Teinostoma pellucidum, sp. n.
Testa minuta, albo-nitens, perdepressa, tenuis, pellucida, imper-
forata; spira convexo-planulata; anfractus 5, leves; sutura
anguste canaliculata; basis planato-convexiuscula, in regione
umbilicali callo lato albo depresso munita; apertura parviuscula,
ovato-subcircularis, yix obliqua; labrum tenue.
Alt. 1, diam. 2} mm.
13*
188 Mr. G. B. Sowerby on new
A little pellucid white shell, with an opaque white callus
covering the umbilical region. Itis very like 7. cancellatum,
Tate, but thinner, and without the sculpture.
Solariella tenuis, sp. n.
Testa conico-orbicularis, depressiuscula, tenuis, late umbilicata,
albida, pallide aurantio strigata; spira obtuse conica, gradata ;
anfractus 4, rotunde-conyexi, spiraliter lirati, liris in anfractu
penultimo 3, in anfractu ultimo numerosis, evanidis; sutura late
impressa ; basis rotunde convexa ; umbilicus perspectivus ; aper-
tura circularis; peristoma continuum, tenue.
Alt. 2, diam. 23 mm.
A delicately coloured, thin, globosely conical shell with a
perspective umbilicus.
I know of no species very similar to this; it might easily
be mistaken for one of the minute Cyclostomide, having some-
thing the look of a Cyclotopsis.
Fam. Littorinide.
Fossarina lineata, sp. n.
Testa oblique ovata, perforata, luteo-albida, lineis virido-fuscis
duplicatis spiraliter cincta, hic illic irr egulariter fusco maculata ;
spira brevis, obtusa; anfractus 4, rotundati, oblique minutissime
striati; anfractus ultimus late obliquus; basis convexa, area
umbilicali alba, oblique late excavata, conspicue filo-striata ;
umbilicus profundus ; apertura subcircularis, ampla, intus viridis,
lineata, levis; labrum tenue; columella arcuata, leviter incras-
sata, alba, postice tenuiter effusa.
Alt. 4, diam. 4 mm,
This shell partakes of the form and general characters
common to the genus Fossarina. It is rounder and less
oblique than /’. Braziert, Angas, and principally characterized
by its greenish-brown spiral lines.
Fam. Fossaride.
Fossarus oblongus, sp. ni.
Testa oblongo-ovata, alba, tenuis, subpellucida, angustissime rimata;
spira brevis, obtusa ; anfractus 33, convexi, supra obtuse angulati,
supra angulum leviter planulati, sutura impressa sejuncti; ultimus
oblongus, spiraliter multiliratus, leviter oblique plicatus ; apertura
elongato-ovata ; labrum tenue; columella planulata, leviter effusa.
Alt, 34, diam. 27 mm.
Species of Marine Shells. 189
An oblong, thin, white, almost transparent shell. It seems
to belong to the same section as F’, reticulatus, S. Wood, but
larger, with a more obtuse spire, and longitudinal plications
much less distinct and more oblique.
Fam. Scutellinide.
Scutellina nobilis, sp. n.
Testa capuliformis, subcircularis, tenuis, lutescens, elatiuscula, antice
valde arcuata, postice concaviuscula, apice acuto, postice inclinato ;
costellis radiantibus numerosis nodulosis, et liris concentricis dis-
tantibus cancellata, ad marginem crenulata.
Long. 28, lat. 26, alt. 17 mm.
This is the largest known species of the genus. Compared
with S. erenudata, Brod., it is much larger, more elevated in
form, and has more distant and much less closely nodulous
ribs. It is further distinguished by the presence of con-
centric rather distant lire crossing the radiating ribs.
Only one specimen found in the Nevill Collection.
Fam. Columbellide.
Columbella perlevis, sp. n.
Testa ovato-fusiformis, aurantia, ad suturam rufo-fusco pauci-
maculata, ad apicem albida; spira elata, acute conica; anfractus 71,
convexi, leves, ultimus supra obtusissime angulatus, infra leviter
contracta, suleata ; apertura oblonga, postice angusta, antice latior ;
labrum leviter incrassatum, rectiusculum, postice angulatum, intus
dentatum ; columella rectiuscula, ad marginem tuberculis ovalis
circ. 7 munita, intus bi-tuberculata; canalis brevis, latiusculus,
vix recurvus.
Long. 13, diam. 63 mm.
A typical Columbella, with a sharply acuminated spire ;
colour bright orange, with a few reddish-brown spots at the
suture and near the base. It differs from C. rustica, Linn.,
in colour, in the relative height of the spire, in the slight but
characteristic angularity of the top of the body-whorl, and in
other respects.
Columbella (Atilia) Macandrewi, sp. n.
Testa acute pyramidata, pallidissime cinerea, longitudinaliter pallide
aurantio strigata, ad suturam et peripheriam albo interruptim
balteata; spira elatissima, acuta; anfractus 9, leves, vix con-
vexi, sutura anguste canaliculata sejuncti; anfractus ultimus
190 Mr. G. B. Sowerby on new
convexus, infra contractus, ad basin valde sulcatus, vix rostratus;
apertura mediocriter lata, intus paucilirata; labrum acutum,
extus incrassatum ; columella rectiusenla ; canalis brevissimus.
Long. 63, diam. 23 mm.
A small very delicately-coloured shell allied to C. eaimia,
Reeve, compared with which species it is smalier, very dif-
ferent in colour and pattern, shorter at the base, and its spire
is more regularly pyramidal.
Fam. Neritide.
Nerita constellata, sp. n.
Testa oblique ovata, solidiuscula, levis, purpurascens, maculis parvis
albidis inzequalibus conspersa; spira brevissima, planulata, fere
immersa; anfractus ultimus rotunde convexus, concentrice ob-
scurissime striatus, subobsolete spiraliter sulcatus; apertura
semicircularis, intus levis; labrum acutum; area columellari
levis, leviter convexa, ad marginem minute irregulariter denti-
culata.
Alt. 63, diam. 9 mm.
This shell bears some resemblance to a large Neritina
fluviatilis, while in form and substance it might almost be a
miniature Nerita Rumphi, Recluz, but the specimens, which
are fairly numerous, are very uniform in size, colour, and
general characters.
The interior of the aperture is quite smooth.
Fam. Scalariide.
Scalaria octogona, sp. n
See 8 p=
Testa pyramidalis, alba, levis; spira elata, acuta; anfractus 8,
rotundati, costis 8 acutis erectis leviter obliquis instructi ; aper-
tura circularis.
Long. 123, diam. 6 mm.
A white Sealaria of simple pyramidal form, with eight
erect sharp scarcely reflexed ribs.
Scalaria interstriata, sp. n.
Testa acuminata, alba, multicostata ; spira acute turrita ; anfractus 9,
convexi, primi 2—3 leves deinde gradatuli, costis elevatis circ. 25,
leviter reflexis, superne acute angulatis instructi, spiraliter striati;
apertura subcircularis ; peristoma mediocriter crassum, postice
angulatum.
Long. 9, diam, 33 mm.
Species of Marine Shells. 191
This species is easily distinguished by the large number
of its prominent slightly reflexed ribs, which are acutely
angularly raised near the suture, the interstices being spirally
striated.
Fam. Buccinide.
Tritonidea (Cantharus) fuscopicta, sp. n.
Testa ovato-acuminata, crassiuscula, alba, maculis parvis fuscis
transversim clongatis picta; spira elata, acuta; anfractus 7,
leviter convexi, spiraliter lirati, liris rotundatis, in anfractu
penultime 4, interstitiis spiraliter striatis; anfractus ultimus
mediocriter amplus, superne leviter inepte canaliculatus, liris
circiter 12 partim leviter crenulatis instructus, ad basin leviter
contractus ; apertura oblongo-ovata, intus lirata ; labrum crassum,
arcuatum, utrinque uni-tuberculatum; columella arcuata, ad
basin plicato-contorta; canalis brevissimus, leviter recurvus.
Long. 14, diam. 7 mm.
I know of no species bearing any near resemblance to this
prettily marked little shell. In proportion to the size of the
body-whorl, its spire is rather unusually long and sharp for
the genus, and the oblong brown markings on the spiral
ridges are very striking.
Fam. Ungulinide.
Dipiodonta Nevill, sp. n.
Testa suborbicularis, tenuis, albida, epidermide tenuis olivacea
induta, leviter inflata, concentrice confertim striata; umbones
obtusi ; margo dorsalis posticus convexo declivis, anticus brevior,
rotundatus ; margo ventralis arcuatus; ligamentum parvum,
immersum. Pagina interna luteo-cinerea; impressiones muscu-
lares indistincti ; cardo normalis.
Diam. antero-post. 16, umbono-marg. 16, crass. 10 mm.
Compared with the typical Diplodonta (D. rotundata),
the umbones are more obtuse and rather more central, the
posterior dorsal margin more sloping; the shell is of a much
thinner substance, and covered with a light olivaceous
epidermis.
Diplodonta (Felania) crebristriata, sp. n.
Testa subtrigono-rotundata, compressiuscula, tenuis, albida, den-
sissime concentrice striata; umbones acutiusculi, elatiuseul,
autice leviter inclinati; margo dorsalis posticus rotundato decliyis,
192 On a new Shell from the Cape Verd Islands,
anticus valde declivis vel subtruncatus ; margo ventralis arcuatus ;
cardo normalis.
Diam. antero-post. 12, umbono-marg. 113, crass. 7 mm.
A thin white shell of simple form and character, more elate
than the typical form of Diplodonta, and with the whole
surface of the valves finely and closely concentrically striated.
The types of all the species described in this paper are in
the British Museum.
XXIL.— Description of a new Shell from the Cape Verd
Islands. By G. B. Sowersy, F.L.S.
Gibbula Murchlandi, sp. n.
Testa perspective umbilicata, late conoidea, rosaceo-albida, maculis
grandibus, minoribus, et minutis, rufo-fuscis ornata; spira sub-
gradata, mediocriter elata, ad apicem acutiuscula; anfractus 6,
ubique spiraliter conferte lirati, primi 3 convexe declives, czeteri
angulati, ad angulum plicato-nodulosi, supra angulum leviter
planulati, infra levissime convexi ; anfractus ultimus latus, curtus,
biseriatim nodosus; basis convexa, oblique striata, spiraliter
valde lirata, liris circ. 10, retrorso complanatis ; umbilicus pro-
fundus, mediocriter latus, spiraliter profunde canaliculatus, funi-
culo elevato crassiusculo reflexo munitus; apertura oblique
subquadrata, intus argentea, levis; labrum acutum; columella
supra arcuata, deinde leviter undulata vel rectiuscula.
Alt. 14, diam. 16 mm.
flab. St. Vincent, Cape Verd Islands.
At first sight this pretty and interesting shell has much
the appearance of a small Gzbbula magus, but it is more elate,
the body-whorl is more nearly rectangular, the base is much
more strongly ridged, and the spiral funicle eutering the
umbilicus much more elevated.
This species has some affinity with Gibbula fanuloides,
Fischer, which belongs to Adams’s subgenus Forskalia, but
it distinctly connects that group with T’rochus magus, Linn.,
which stands as the type of the genus Gzbbula, showing that
the subgenus proposed by H. & A. Adams is not needed.
The specimens were taken in the above-named locality by
Mr. R. Murchland, after whom I have pleasure in naming
the species.
The type is in the British Museum.
Qo
On new Species of Land- Shells. 19
XXIII.—Descriptions of new Species of Land-Shells from
British New Guinea, and Remarks on Two Species from the
Solomon Islands. By Enear A. Situ, 1.8.0.
THE specimens described in this paper were collected by
Mr. A. S. Meek, partly at Choiseul Island, one of the
Solomon Group, in December 1903, and partly during the
first three months of the present year, at Owgarra on
the Anabunga River, “ two days” beyond Mafulu (or
Mafoola), in the Owen Stanley Range, British New Guinea,
at an elevation above 8000 feet.
Papuina Meekiana.
Testa breviter conica, acute carinata, roseo-purpurea, flavo plus
minus obtecta, ad apicem nigrescens; anfractus 5, celeriter
crescentes, tres superiores convexi, leves, cxteri planiusculi,
oblique tenuiter corrugati, ultimus ad peripheriam acute albo-
carinatus, antice prope aperturam subito descendens, infra plani-
usculus, corrugatus, lineisque incrementi radiatim striatus;
apertura intus nigro-purpurea; peristoma saturate nigrum,
nitens, marginibus callo roseo-purpureo tenui junctis, superiore
bisinuato, columellari lato, planato, antice tuberculo parvo
munito.
Diam. maj. 29 mm., min. 23, alt. 19.
Hab. British New Guinea.
This species belongs to the same group as P. naso, Martens,
P. gemina, Fulton, P. Tayloriana, Ad. & Rve., and P. jucunda,
Fulton. With regard to the peristome, it is very like P. naso,
Papuina Meekiana.
but the snout of the body-whorl is more pointed, and the
anterior part of the basal margin of the peristome is quite
different and bears a small tubercle. It is a much smaller
species also. In colour it is very similar to certain forms of
P. Tayloriana, but is subject to some variation. Sometimes
the dark purple ground-colour is almost concealed by a
yellowish coating which shows a disposition to be in some-
what obscure spiral zones, both on the upper and lower
194 Mr. E. A. Smith on
surfaces. In P. naso and P. gemina the last whorl is pecu-
liarly produced beyond the black lip. The present species
has a similar kind of inflation of the keel behind the lip, but
it does not project beyond the peristome. Named in honour
of Mr. Meek, who at considerable personal risk collected in
this dangerous country.
Chloritis (Sulcobasis) globosa, Preston, var. major.
Testa globosa, umbilicata, solidiuscula, saturate fusca, lineis in-
crementi oblique arcuatis striata, sub lente undique microscopice
granulata ; spira breviter convexe conoidea, ad apicem obtusa;
anfractus 5, convexiusculi, ultimus magnus, globosus, antice
prope aperturam breviter descendens; apertura obliqua, late
lunata, intus purpureo-fusca ; peristoma album, mediocriter ex-
pansum et reflexum, marginibus callo tenui junctis, columellari
dilatato, supra umbilicum partim expanso, niveo,
Diam. maj. 40 mm., min. 32, alt. 30. Apertura cum perist.
25 longa, 20 alta.
Hab. British New Guinea.
Like the type of C. globosa, Preston (Proc. Malac. Soc.
vol. v. p. 17, fig.), but larger, the body-whorl being larger in
proportion to the spire and the peristome more widely
expanded. Occurring in a different district from that of the
typical form, which was described from the “ Northern
Coast” of British New Guinea, C. stiruphora, Smith,
another allied form, occurs near Cloudy Bay on the south-
eastern extremity of the island.
Rhyssota bisculptilis.
Testa depresse globosa, mediocriter tenuis, anguste perforata, supra
fuscescens, haud nitida, infra flavescens, in medio pallide fusces-
cens, polita, epidermide tenui, flavo-olivacea, undique induta ;
spira depresse convexa, ad apicem obtusa; anfractus 5, supra
parum conyexi, striis minutis incrementi elevatis confertis pul-
cherrime sculpti, ultimus supra plus minus etiam malleatus, ad
peripheriam obsolete obtuse carinatus, infra striis concentricis
minutis confertis undulatis, et incrementi lineis concinne sculptus ;
apertura obliqua, late lunata, intus fuscescens ; peristoma leviter
incrassatum, simplex, dilute fuscescens, marginibus remotis,
columellari ad insertionem breviter dilatato et reflexo.
Diam. maj. 35 mm., min. 29. Apertura 19 longa, 16 alta.
Hab. British New Guinea.
Nanina campylonota, 'Vapparone Canefri (Ann. Mus. Stor.
nat. Genova, 18838, vol. xix. p. 199, pl. v. fig. 11), appears to
be very similar in form to the present species, but is
new Species of Land- Shells. 195
described as a white shell, and no mention is made of the
beautiful concentric strize on the under surface, nor is
the dullness of the upper side referred to in contrast to the
glossy base.
Hemiplecta Jansoni.
Testa orbicularis, carinata, perforata, supra rufo-fusca, vix nitida,
incrementi lineis striata, striisque minutis costulatis microscopice
sculpta, infra medium versus sensim pallidior, subnitens, striis
incrementi tenuibus sculpta, aliisque concentricis prasertim circa
umbilicum concinne ornata; spira valde depressa, conoidea,
planata; anfractus 5, supra fere plani, supra suturam sub-
depressi, ultimus acute carinatus, haud descendens, supra sulcis
2-3 spiralibus, fere obsoletis, inconspicue impressus; apertura
obliqua, angulatim lunata, purpureo-fusca; peristoma tenue, ad
marginem pallidum, marginibus remotis, columellari leviter in-
crassato, ad insertionem breviter expanso et reflexo.
Diam. maj. 35 mm., min. 28. Apertura 19 longa, 14 alta.
Hab. British New Guinea.
Nanina Cairni, Smith (Ann. & Mag. Nat. Hist. 1887, vol. ix.
p- 417, pl. xv. fig. 5), is closely related to the present species.
The latter has a flatter, more depressed spire, a sharper
peripheral keel, and stronger concentric spiral strize on the
lower surface. Owing to the depression of the spire, the
form of the aperture is different. Named after Mr. Oliver
Janson, through whose agency the Museum was fortunate in
obtaining this and the other interesting species described
in the present paper.
Hemiplecta ougarrana.
Testa parva, anguste perforata, orbicularis, supra subplana, fusca,
infra pallidior, umbilicum versus albida; anfractus 5-52, lente
accrescentes, fere plani, ad suturam filo-carinati, minute oblique
costulato-striati, quasi sericati, ultimus ad peripheriam acute et
compresse carinatus, haud descendens, infra politus, incrementi
lineis tenuibus striatus; apertura obliqua, angulatim lunata,
intus lilacea; peristoma tenue, pallidum, marginibus remotis,
columellari ad insertionem breviter expanso et reflexo.
Diam. maj. 22°5 mm., min. 20), alt. 10°5. Apertura 11-5 longa,
7°5 alta.
Hab. British New Guinea.
Distinguished by its small size, flattened spire, carinate
suture, and very small perforation. Within the aperture
and near the peristome there is a slight shelly thickening
which is peculiar.
196 Miss G, Ricardo on the Tabani from the
H. infelix (Smith *) appears to be the nearest ally of this
species. Its spire, however, is more raised, the peripheral
keel even more acute, the umbilical perforation a trifle
larger, and the form of the aperture rather different.
Papuina lilium, Fulton.
Hab. Choiseul, one of the Solomon Islands.
The exact locality of this species has not hitherto been
recorded. It is remarkable that this very distinct species
should have been considered by Dr. Cox (Proc. Zool. Soc.
1873, p. 567) as a variety of P. wanthochila, Pfr., quoted by
Pilsbry as from Bougainville Island.
The difference in the number of volutions pointed out by
Mr. Fulton is not constant, for both the two specimens in
the Museum have six whorls (not 54). Both are much
finer than the shell figured by him and resemble that
represented in the Proc. Zool. Soc. 1873, pl. xlviil. fig. 7.
Papuina Dampieri (Angas).
Helix (Geotrochus) Dampieri, Angas, Proc. Zool. Soc. 1869, p. 47,
pl. ii. fig. 6.
Papuina Dampieri, Pilsbry, Man. Conch. vol. vii. p. 11.
Hab. Choiseul Island (Meek) ; Louisiade Archipelago
(Angas) ; Choiseul Bay, Bougainville Straits (Guppy) ;
Bougainville Island (Fulton, in Brit. Mus.).
I am inclined to think that the original locality assigned
to this species must be erroneous, for its occurrence at such
a remote place separated by three hundred miles of sea is
most improbable.
XXIV.—Notes on the Tabani from the Palearctic Region in
the British Museum Collection. By GrrrrupE Ricarpo.
Wuite going through the Tabani from the Palearctic Region
in the British Museum Collection I ascertained several
particulars as regards the species described by Walker, and
identified various species in the collection of Diptera from
Algeria by the Rev. A. E. Eaton, and append a few notes
which may be of use. Brauer’s work on the European
Taubani is, of course, of the greatest use, but, as has been
* ¢The Conchologist,’ vol. ii. p. 10); ‘ Journal of Malacology,’ vol. v.
p- 22, pl. ii. figs. 6, 7
Palearctic Region in the British Museum, 197
remarked by others, I cannot follow him satisfactorily in
many of his minute differences, chiefly in the shape of the
palpi and the length of the forehead; in the females they do
not seem marked enough for the identification of species
such as, for example, solstitialis and tropicus.
The following species have been found in Algeria, collected
by the Rev. A. E. Eaton :-—
Atylotus tomentosus, Macq.; Atylotus alexandrinus, Wied. ;
Atylotus rusticus, F.; Tabanus autumnalis, L.; Tabanus
cordiger, Mg.; Tabanus pulchellus, Loew; and one new
species described below.
Atylotus tomentosus, Macq.
One male “near Biskra, along the route of ‘des Zibans’
in the cornfields, on ‘ Ammin Visnoga,’ 18.° v. 93” (Eaton),
94. 114.
One female from Constantine, 28. v. 95 (Haton), 96. 137.
One female ‘from Constantine, on the edge of the pine-
plantations on the top of the hill north of Mansoura,
11. vi. 94” (Haton), 95. 34.
The male agrees with the figure in Explor. d’Algér.
(Lucas) ; the figure in Macqguart’s Dipt. Exot. is different
and apparently depicts a female specimen.
Atylotus alexandrinus, Wied.
One male from Colonne Voirol, Algiers, 5. vi. 93 (Haton),
94. 114.
There is also a male from Tunis (F. Moore), 68. 4.
The third joint of the antenne is wanting; the first
posterior cell of the wings is narrowed at its apex, which
is not mentioned as a characteristic of A. alexandrinus
by any authors ; otherwise these specimens agree with the
description.
Atylotus rusticus, L.
? flaviceps, Zett.; P ruralis, Zett.; Rousselii, Macq.
Two females “ from Constantine, on the edge of the pine-
plantations on the top of the hill north of Mansoura ; visits
© Ferula’” 11. vi. 94” (Eaton), 94, 114.
The Tabanus Rousselii, Macq., a species from Algiers, is,
I consider, a synonym of Atylotus rusticus, F.
These specimens differ from the description of Atylotus
rusticus only by the apex of the third joint of the antenne
being blackish ; the markings on the forehead, which seem
198 Miss G. Ricardo on the Tabani from the
to be variable in this species, appear thus on these speci-
mens :—Frontal callus yellowish brown, oblong, furrowed,
the middle one smaller, oblong, black ; in some females from
Hungary these callosities are both black, and smaller. The
shape of the antenne and palpi agrees with those of
A. rusticus.
Atylotus Eatoni, § 2, sp. 0.
Male (type) ‘from Constantine, Algeria, along the route
de la Corniche, visiting ‘ Ferula scabra, 2. vi. 95”’ (Eaton),
96. 137.
Male from the same place “along the Arab cliffs path
between 9.15 and 10 aM., 22. vi. 947 (Eaton), 94. 114.
Male from the same place “ between the railway and the
route de la Corniche, 18. vi. 95” (Haton), 96. 137.
Male “from Bone, Algeria, by the Bone-Gmelma railway,
23. v. 96” (Eaton), 96. 136.
Female (type) “near the Colonne Voirol, Algiers, among
bushes near the route Madeleine on ‘ Ferula communis,’
18. v. 95” (Eaton), 94. 114.
This species may be distinguished from Afylotus ater,
Rossi, and Atylotus anthracinus, Mg., by the much smaller
tooth of the autenne ; from Aéylotus umbrinus, Mg., by the
absence of any brown cloud or spot on the wings behind the
stigma; from Atylotus alexandrinus, Wied., by the hyaline
wings and the short-haired eyes ; from Atylotus Letourneuzi,
Bigot, by the wholly black tibiz, not “ widely whitish yellow
at base,” as Bigot describes is the case in his species.
Black. Eyes hairy, the upper facets only very slightly
larger; the third joint of the antennz with a small tooth.
Wings hyaline.
Male.—Head not larger than in the female. Eyes with
no band, the facets on the upper part on each side of the
forehead slightly larger; the pubescence is sparse, the
ocelligerous tubercle inconspicuous, yellow-haired, the hind
part of the head with long yellow hairs ; the frontal triangle
is rather protruding, brownish, with silver-grey tomentum.
Antenne black, reddish brown at the base of the third joint ;
the first joint cylindrical, stout, with black hairs; the second
cup-like in shape, with black hairs on each margin; the
third joint with a moderate tooth, the annulate portion as
long as the basal portion. Face blackish, with some grey
tomentum near the eyes, with black hairs intermixed with
some white ones on the inner side; beard blackish brown,
white below; palpi black, with long black hairs. Thorax
Palearctic Region in the British Museum. 199
black, with grey tomentnm and fine short rufous hairs on
the dorsum, some black hairs intermixed, greyer at the sides
and on shoulders, breast- sides blackish brown ;_ halteres
reddish brown. Legs black, the extreme apices ‘of femora
yellowish ; the pubescence is black, long on the cox and
femora. Abdomen black, shining ; pubescence black, the
hairs at the sides black, fulvous on the last two segments, the
underside black. Wings hyaline, narrowly yellow ish brown
on the fore border, the veins reddish brown ; no appendix ;
the first posterior cell not narrowed at its apex.
Length 16, width of head 5 mm.
Female. —Very similar to the male. Frontal stripe
blackish, with grey tomentum; the frontal callus black,
shining, oval, not reaching the ey es, prolonged as a narrow
stripe beyond the middle of the forehead, which is about four
times as long as it is broad; no ocelligerous tubercle; the
frontal triangle is black, shining, with a grey middle line
and grey at the sides, rather protuberant. Hind part of the
head with a black tuft of hairs in the centre and shorter
yellowish hairs beyond. Palpi black, with grey tomentum
and black hairs; the second joint long, rather stouter at
the base, gradually tapering to a point, almost straight.
Abdomen broader than in the male, with plum-coloured
tomentum on the basal segments, most noticeable on the
second one; the fulvous hairs on the last segments are more
numerous ard distributed over the dorsum of the segments ;
and is fainter coloured, nearly white. The hairs on the
thorax are also greyer than i in the male.
Length 15, width of head 5 mm.
Tabanus autumnalis, L.
One female “from Biskra, on the dunes 64 miles from the
town, 18. v. 93” (Eaton), 94. 114.
Tabanus cordiger, Meig.
One female “from Biskra, cn the dunes, caught between
7.15 and 9.30 a.m., 64 miles from the town, 31.2 v. 93”
(Haton), 94, 114.
One female “from Biskra, near Hammam as Salahm,
14. iv. 95” (Eaton), 94. 114.
This species does not appear to have been recorded before
from North Africa,
200 Miss G. Ricardo on the Tabani from the
Tabanus pulchellus, Loew.
One male from Biskra, 28. v. 94 (Haton), 94. 114.
One female from Biskra, 28. v. 94 (Haton), 94. 114.
Two females “from Biskra on the dunes, 7.15—9.30 a.m.,
61 miles from the town, 31. v.? 93 ” (Eaton), 94. 114.
The type came from Asia Minor. ‘These specimens answer
to the description, but the femora are entirely yellow in the
male and female; there are a few black hairs on the palpi of the
female. In the table of the males Brauer places the species
conditionally under the section with long hairs on the back of
the head; this is correct as far as concerns these specimens
from Algeria. Brauer remarks that this species makes the
genus Atylotus no longer tenable, as this species is near
A. rusticus, L., but on account of the naked (or nearly naked)
eyes it must belong to Tabanus in sensu stricto. A note on
the female gives the eyes as light sage-green in life; Brauer
describes them thus, viz. as glass-green.
Tabanus albifacies, Loew.
Two females from Moghara, Egypt, March 1901 (Dr. C. W.
Andrews), 1902. 20, answer to the description given by
Loew; they have a narrow black, not very noticeable, band
below the eyes, not mentioned by Loew.
This species, pulverifer, Wlk., and pulchellus, Loew,
are all near cordiger. These differ by having an appendix
to the wing, and may be distinguished from pulverifer and
pulchellus by their black antenn and darker legs.
The type came from Asia Minor.
The Walker types of Tabanus macrops, nigrifer, and siccus
from Egypt, of Vabanus terminalis from Mount Sinai, and
of Tubanus pallescens and politus from Arabia are not to be
found in the British Museum Collection, and are unknown
to me.
The Tabanus transiens, Walker, from an unnamed locality,
is missing now from the Collection; it was probably a
synonym of T'abanus ater, Rossi, and might in any event be
now expunged from the Catalogue of Tabani.
The specimen named Tabanus connexans, 2, Saunders
Coll., 68. 4, by Walker, from an unknown locality, is nothing
but a specimen of Tabanus bromius, L., and should be added
to the synonyms of that species.
The specimen named Tabanus depressus, 2, by Walker,
from unknown locality, badly preserved, is, I believe, an
Palearctic Region in the British Museum. 201
example of Therioplectes luridus, Fall., and should be added
to the synonyms of Th. luridus.
Tabanus pulverifer, § 2, Walker.
Male (type) from Turkey in Asia (Loftus), 50. 105.
Three females from the same locality (co-types) are nearly
related to Tabanus cordiger, Mg., but belong to a distinct
species ; they differ in the following particulars :—
Antenne wholly red, black at the apex; frontal callus
reddish brown. Coxe and femora reddish yellow, as are the
tibiz, only the anterior tibiz with a brown apex; tarsi
reddish brown, all with white pubescence. The underside
of abdomen has no median black stripe. The species is
much lighter in colour and slighter in form than cordiger ;
the four specimens vary in size from 12-14 mm. In the
female the band under the eyes is narrow and yellowish
brown. In the male the black hairs on the hind part of the
head are not present; the underside of the abdomen is
reddish, greyer at the apex.
Tabanus polygonus, 6 2, Walker.
Male (type) from Bagdad (W. K. Loftus), 50. 105; and
another male from the same locality.
Female (type) from same locality.
The males would come in Brauer’s table near autumnaiis,
spectabilis, and rectus. The female, which is much smaller,
may possibly not belong to the same species and has a slight
resemblance in appearance to Atylotus rusticus, Z.
A redescription may prove of use :—
Reddish-brown species, with grey median triangular spots
on the abdomen. Eyes bare; no long hairs on the hind part
of the head.
Male.—Head large; facets of eyes unequal, sharply
divided, the large facets yellowish, extending over more than
half the eyes, surrounded by a narrow zone of small facets
above and a broad zone on the basal half. Frontal triangle
grey, reddish at apex. Antenne red, with the apex darker;
the third joint with a small tooth, long and slender, the annu-
lated portion not so long as the basal joint ; the ocelligerous
tubercle small, brown. Face whitish, with white hairs ;
beard white ; palpi white, the second joint oval, with white
pubescence. Thorax blackish, with four grey stripes and
greyish pubescence; the shoulders red; the sides reddish,
with black hairs above and whitish ones below ; scutellum
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 14
202 On Vabani from the Palearctic Region.
black, with grey tomentum. Abdomen long and narrow,
reddish, with a row of median grey triangular spots, which
on the second and last segments are surrounded by black
margins; the hind borders of the segments narrowly white,
the pubescence black ; the underside brighter red, with some
irregular black markings. Legs red; the coxee and femora
with longish white hairs ; the tibize with short black pubes-
cence on the sides and some whitish-yellow hairs on their
flat surfaces ; tarsi with black pubescence, pulvilli yellow.
Wings longer than the abdomen, hyaline, the stigma and
veins yellow ; no appendix.
Length 18, width of head 5 mm.
Female.—Kyes not large, the facets equal, with two bands
(in the male I can see no trace of bands). Frontal callus
reddish brown, almost square, a little wider at its base, nearly
touching the eyes ; frontal stripe grey, with black markings
(probably denuded), the middle callus being a black stripe
not joining the frontal callus; tle vertex blackish ; no ocelli-
gerous tubercle. Antenne at base red (the rest wanting).
Frontal triangle grey. Face grey, with white hairs; beard
white ; palpi pale yellow, the second joint long and slender,
at the base stouter, gradually tapering to a point, nearly
straight, with a few black hairs. On the abdomen the grey
triangular spots are replaced by a broad stripe of greyish
tomentum, the black pubescence is longer, the abdomen not
so pointed as in the males. Wings with the first posterior
cell distinctly narrowed at the apex (this is not the case in
the male).
Length 14, width of head 4 mm.
There are several species of Tabani from Japan: a long
series of Tabanus trigonus, Coq., male and female; this is a
very nearly related species to the European Jdovinus and
sudeticus, the brighter yellow colour of the autenne and the
larger tooth of the third joint, with an appendix to the wings,
being the most striking differences. Another species (males
and females) from Korea is very nearly allied to Tabanus
bovinus, Lw., but the tooth of the antenne is larger and
brighter yellow and the wings possess an appendix ; the eyes
of the males having facets all equal in size distinguish it
from Tabanus trigonus, Coq.
Mr. W. L. Distant on Homoptera. 203
XXV.—Rhynchotal Notes—XXXIV. By W. L. Disranr.
Fam. Cicadide (continued from p. 35).
Subfam. 7rercrvrva (continued).
Division PARNISARIA.
This division or large group of genera is allied to that of
Taphuraria, from which it primarily differs by the size and
position of the eyes, which do not project (or, at least, not
distinctly so) beyond the anterior angles of the pronotum ;
the abdomen is usually short, except in a few cases, not
longer than the space between the apex of head and base of
cruciform elevation.
Parnisaria includes a number of genera widely but sparingly
distributed in all the great zoogeographical regions; in the
Palearctic region apparently confined to its extreme eastern
area.
Henicotettix, the type of which I have not seen, probably
comes in this group of genera, but is to be recognized by the
character—“ Tibize anticze ultra insertionem tarsorum in
spinam producte.”
Synopsis of Genera.
1. Tegmina and wings with reticulate venation, giving
the appearance of numerous cellular areas ........ Arcystasia,
2. Tegmina with six, wings with four apical areas .... Derotettix.
3. Tegmina and wings with the venation normal, not
reticulate.
A. Wings with six apical areas.
a. Head (including eyes) more or less narrower than
base of mesonotum.
6. Tegmina about or nearly three times as long
as greatest breadth, costal margin above
radial area neither curved nor gibbous; basal
cell generally about twice as long as broad.
ec. Head with front not prominently projecting,
about as broad as length of vertical margins;
margins of both more or less continuous,
d. Tegmina with the first and second apical
areas longer than eighth.............. Quintilia.
dd. Tegmina with the first and second apical
areas very short, not longer than eighth. Pszlotympana.
cc, Head with front strongly projecting, nar-
rower than length of vertical margins,
which are convex ; margins of both discon-
tinuous, almost at right angles to each other.
e. Tegmina about twice as long as broad ;
abdomen longer than space between apex
14*
204 Mr. W. L. Distant on Homoptera.
of head and base of cruciform elevation ;
opercula in male reaching base of first
abdominal segment ; rostrum reaching
ANECTTNECIAGE COR uals ys oieulely eee Diemenia.
ee. Tegmina about three times as long as
broad; abdomen about as long as space
between apex of head and base of eruci-
form elevation; opercula in male not
reaching base of abdomen; rostrum
passing intermediate cox ............ Koranna,
ece. Head with front strongly projecting,
broader than the length of vertical margins,
which are subangulate Apa ha Oar ee Abagazara,
bb. Teomina little more than twice longer than
ja brentest breadth.
. Costal margin of tegmina above radial area
neither curved nor ‘cibbous Side en eb . Gudanga.
b?, Costal margin of tegmina above radial area
more or less curyed and gibbous.
f. Head with front not prominently produced,
margins of front and vertex continuous.
g. Tegmina with the first and second ulnar
areas narrow, very much longer than
broad ; basal cell broader at base than
apex, not twice as long as broad . Masupha.
ff. Head with front prominently produced,
margins of front and vertex discon-
tinuous.
gg. Tegmina with the first and second ulnar
areas broad, only moderately longer
than broad, placed somewhat obliquely
to each other; basal cell about twice
2s LONE BS DTOAG! 76 .h- sine einer ee . Platypedia.
B. Wings with five apical areas.
h. Ocelli well separated from hase of head.
7. Tegmina with postcostal area narrow, but
distinct.
k. Head with the margins of front and vertex
more or less continuous, front not promi-
mently, projecking To, er ececmei ait Oita Taipinga.
kk. Head with front prominently projecting, the
margins of front and vertex discontinuous
and at right angles to each other........ Adenia.
0. es Tee with posteostal area not distinct . Calyria.
7. Tegmina with postcostal area very broad and
Cele oe sale 3.5.5 silo eee ae ate ieee Parnisa.
hh. Ocelli placed close to base of head
C. Wings with four apical areas.
l. Head with front not prominently produced ; mar-
gins of front and vertex more or less continuous. M, apondera.
Wl. Head with front prominently triangularly pro-
duced; margins of front and vertex discon-
tinuous, somewhat at right angles to each other. Prunasis.
bide gh aianats Callipsaltria.
Mr. W. L. Distant on Homoptera. 205
Genus QUINTILIA.
Tibicen, subgen. Quintilia, Stal, Hem. Afr. iv. p. 28 (1866).
Type, Q. rufiventris, Walk. (Czcada).
Quintilia mogannia, sp. n.
3. Body above black, finely greyishly pilose ; head with
a central line to front, and anterior lateral margins to vertex,
pronotum with the margins, and a central longitudinal fascia
(widened posteriorly, where it contains two small black spots) ,
mesonotum with two central, discal, wedge-shaped fasciz,
the lateral margins, cruciform elevation, and abdomen
with the posterior segmental margins, testaceous; body
and head beneath, sternum, legs, and opercula piceous ;
lateral margins of face and streaks to sternum and femora
testaceous ; intermediate and posterior tibia annulated
apically and basally with pale ochraceous; abdomen beneath
testaceous, with central and marginal piceous spots ; tegmina
and wings hyaline, their basal areas pale fuscous brown,
the venation fuscous; tegmina with the costal membrane
and claval area, and wings with the extreme base, testaceous ;
opercula piceous, with their margins narrowly testaceous ;
rostrum reaching the intermediate coxe; head (including
eyes) a little narrower than base of mesonotum; transverse
vein at base of second apical area to tegmina nearly vertical ;
face moderately centrally sulcate.
?. Body beneath more testaceous than in male, and
abdomen without the piceous spots.
ihoue-, exe). teem., G 15, 9 18 mm. exptegm:, d 9;
40-44 mm.
Hab. China: Hong Kong (J. C. Bowring, Brit. Mus.).
Quintilia maculiventris, sp. n.
Head, pronotum, and mesonotum black; head with a
small central longitudinal spot at base, pronotum with the
margins and a central longitudinal fascia, mesonotum with
two fascie on each lateral area (one marginal, the other and
narrower nearer centre), and the cruciform elevation ochra-
ceous ; abdomen above ochraceous, its base and a central
series of segmental spots black, the margins of the segments
bright testaceous ; body beneath and legs ochraceous; face,
space between face and eyes, spots to acetabule, and central
abdominal segmental spots black, segmental margins paler
testaceous than above; tegmina and wings hyaline ; tegmina
with the veins somewhat broadly margined with fuscous, the
206 Mr. W. L. Distant on Homoptera.
costal membrane ochraceous ; wings pale hyaline, unspotted ;
head (including eyes) a little narrower than base of meso-
notum ; rostrum reaching the intermediate coxe; transverse
vein at base of second apical area to tegmina oblique.
Long., excl. tegm., J 19, ? 16 mm.; exp. tegm., g 39,
9 37 mm.
Hab, Transvaal (Coll. Dist.) ; Natal (Coll. Dist.).
This species has a wonderful resemblance to Melampsalta
leucoptera, Germ.
Genus PSILOTYMPANA.
Psilotympana, Stal, Ann. Soc. Ent. Fr. (4) i. p. 620 (1861).
Type, P. signifera, Germ. (Cicada).
ABAGAZARA, gen. nov.
9. Head considerably longer than pronotum, front longly
and broadly projecting, longer than vertex, and broader than
the length of the lateral margins of vertex, which are a little
discontinuous; ocelli a little nearer to each other than to
eyes; pronotum narrow, transverse, the posterior angles
widely outwardly ampliate ; mesonotum (including cruciform
elevation) shorter than head and pronotum together ; abdomen
about as long as space between apex of head and base of
cruciform elevation; tympana entirely exposed; posterior
tibize spined on each side; anterior femora spined beneath ;
tegmina and wings hyaline, the first nearly three times longer
than greatest breadth, and with eight apical areas; wings
with six apical areas.
Type, A. bicolorata, Dist. (Callipsaltria).
DIEMENIA, gen. nov.
Head with the front strongly convexly projecting, narrower
than length of the vertical margins, which are convex, margins
of both discontinuous, almost at right angles to each other,
ocelli a little nearer to each other than to eyes, head (including
eyes) a little narrower than base of mesonotum; pronotum
about as long as head, lateral margins a little laminate and
slightly convex, posterior angles spatulately produced; meso-
notum (including cruciform elevation) almost as long as
pronotum and head together; abdomen robust, centrally
ridged above, beneath with the disk convex, the marginal
areas recurved, longer than space between apex of head and
base of cruciform elevation; tympana entirely exposed;
opercula in male transverse, about reaching base of first
Mr. W. L. Distant on Homoptera. 207
abdominal segment ; rostrum reaching the intermediate coxee ;
tegmina almost half as broad as long, the postcostal area
broad, basal cell about twice as long as broad, apical areas
eight; wings with six apical areas.
Type, D. coleoptrata, Walk. (Cicada).
KORANNA, gen. nov.
Head with the front strongly projecting, about as broad as
length of the vertical margins, which are convex, margins of
both discontinuous, almost at right angles to each other, head
(including eyes) narrower than base of mesonotum ;_pro-
notum about as long as head, the lateral margins slightly
laminate and convex, the posterior angles strongly spatu-
lately produced ; mesonotum almost as long as pronotui and
head together; abdomen short, robust, as long as space
between apex of head and base of cruciform elevation;
tympana completely exposed ; opercula in male short, trans-
verse, not reaching base of abdomen ; abdomen above convex,
beneath with the disk convex, the lateral areas strongly
recurved ; rostruin passing the intermediate coxe ; tegmina
about three times as long as broad, postcostal area broad,
basal cell broad and about twice as long as broad, apical areas
eight; wings with six apical areas.
Type, K. analis, Dist.
Koranna analis, sp. n.
6. Body above black; head with the vertical margins,
inner margins of eyes, and three small basal spots ochraceous ;
pronotum with the margins, sometimes three central longitu-
dinal spots, and a spot on each lateral area ochraceous ; meso-
notum with two small discal spots (really the apices of two
obsolete obconical spots), lateral margins (narrowly), a spot
at anterior angles, two lateral spots, and base of cruciform
elevation ochraceous ; abdomen with the anal segment and
appendage stramineous, the first with a basal black spot;
body beneath stramineous or pale ochraceous ; face, space
between face and eyes, rostrum (excluding base), lateral areas
of sternum and abdomen piceous and greyishly pilose ;
femora and tibiz pale castaneous, tarsi piceous ; tegmina and
wings semihyaline ; tegmina talc-like, the venation fuscous, a
broad fuscous angulate fascia crossing bases of apical areas,
and small fuscous spots at apices of longitudinal veins to
apical areas ; wings with about basal halves pale ochraceous,
outwardly margined with fuscous, small fuscous spots at
apices of longitudinal veins to apical areas.
208 Mr. W. L. Distant on Homoptera.
Long., exc]. tegm., g 17 mm. ; exp. tegm. 47-50 mm.
Hab. 8. Africa (Dr. Smith and Drége, Brit. Mus.).
GUDANGA, gen. nov.
Head (including eyes) a little narrower than base of meso-
notum, the front broad, depressed, broader than long, almost
continuous with lateral margins of vertex, shorter than
vertex ; pronotum about as long as head, its lateral margins
nearly straight, its posterior angles ampliated ; mesonotum
(including cruciform elevation) a little shorter than head and
pronotum together; abdomen in male longer than space
between apex of head and base of cruciform elevation ;
tympana entirely exposed ; opercula in male short, transverse,
widely separated, not extending beyond basal segment of
abdomen ; rostrum almost reaching posterior coxe ; tegmina
semiopaque, a little more than twice longer than greatest
breadth, costal margin neither arched nor gibbous, basal cell
about twice as long as broad, apical areas eight; wings with
six apical areas (in the typical specimen on which the genus
is founded one wing has six, the other five apical areas) ;
anterior femora not prominently spined beneath.
Type, G. Boulayz, Dist.
Gudanga Boulayi, sp. n.
&. Body black, greyishly pilose; narrow anterior and
posterior margins and a central longitudinal fascia to pro-
notum, cruciform elevation, posterior abdominal segmental
margins both above and beneath, margins of face, and base of
rostrum ochraceous ; opercula greyish; legs piceous; cox
and intermediate and posterior femora beneath more or less
ochraceous; tegmina pale fuscous, mottled with darker
fuscous, the venation paler, the claval area sanguineous at
base, the venation fuscous.
Long., excl. tegm., ¢ 16 mm.; exp. tegm. 36 mm.
Hab. West Australia: Champion Bay (4. du Boulay,
Brit. Mus.).
Genus MASUPHA.
Masupha, Dist. Ann. & Mag. Nat. Hist. (6) ix. p. 817 (1892).
Type, J. ampliata, Dist.
Masupha Dreget, sp. n.
3. Body above black; head with the margins of front
broadly castaneous, a small spot on each vertical margin and
Mr, W. L. Distant on Homoptera. 209
three small spots at base ochraceous; pronotum with the
margins (narrowly),a central discal longitudinal fascia,and the
incisures ochraceous ; mesonotum with two obsolete central
obconical spots, only denoted by their outer margins and
apices, lateral margins (narrowly) and anterior angles and
centre of cruciform elevation ochraceous; abdomen above
black (apex mutilated in type) ; body beneath stramineous;
face piceous ; space between face and eyes, and many irre-
gular spots to sternum, black; legs pale castaneous, apices
to femora ochraceous ; tegmina and wings semihyaline talc-
like, tegmina with the venation brownish ochraceous and
on basal area spotted with fuscous, two angulated fuscous
fascize on apical areas; wings with angulated fuscous spots
on outer area, the outer margin pale hyaline ; costal tegminal
membrane, and extreme bases of both tegmina and wings,
ochraceous.
Long., excl. tegm., ¢ (abdomen mutilated) ; exp. tegm.
49 mm.
Hab, South Africa (Drege, Brit. Mus.).
Genus PLATYPEDIA.
Platypedia, Uhler, Ent. Am. iv. p. 23 (1888).
Type, P. areolata, Uhler (Cicada).
Genus ARCYSTASIA.
Arcystasia, Dist. Proc. Zool. Soc. 1882, p. 133.
Type, A. Goddefroy?, Dist.
Genus DEROTETTIX.
Derotettix, Berg, Anal. Soc. Cient. Argent. xiv. p. 11 (1882).
Type, D. mendosensis, Berg.
Derotettix Wagnert, sp. n.
Head and thorax above olivaceous green; vertex of head,
margins of pronotum, and cruciform elevation pale testaceous ;
area of the ocelli, a central longitudinal fascia to pronotum,
two central, anterior, obconical spots to mesonotum, and dis k
of cruciform elevation black; abdomen pale olivaceous,
thickly greyishly tomentose, segmental margins pale testa -
ceous, and a central longitudinal fascia (broadest at base )
piceous; body beneath and legs ochraceous, greyishly
tomentose ; face olivaceous ; posterior margin of metanotum,
210 Mr. W. L. Distant on Homoptera.
and apices of rostrum and tarsi, piceous ; tegmina and wings
hyaline; tegmina with the costal membrane, postcostal
margin, basal cell, and base and margins of claval area
virescent, venation (excluding that of apical areas) virescent ;
a transverse spot beneath radial area, an irregular transverse
fascia crossing from end of radial area, longitudinal veins to
apical area, and the apical and outer margins pale fuscous;
a basal streak pale sanguineous ; wings with a pale sangui-
neous basal streak, and the outer margins of the apical areas
pale fuscous.
Long., excl. tegm., ¢ 11 mm. ; exp. tegm. 26 mm.
Hab. Chaco dé Santiago del Estero, Bords du Rio Salado
(H. . Wagner, Paris Mus.).
TAIPINGA, gen. nov.
Head (including eyes) a little narrower than base of meso-
notum, front very much shorter than vertex, which is distinctly
longitudinally sulcate at base, margins of front and vertex a
little discontinuous, breadth of front about equal to length of
margins of vertex ; pronotum about equal to, or slightly
longer than, head, the lateral margins slightly ampliate and
convex, the posterior angles ampliate; mesonotum (including
cruciform elevation) about equal in length to head and pro-
notum together ; abdomen longer than space between apex of
head and base of cruciform elevation; tympana_ entirely
exposed ; opercula in male about reaching base of abdomen ;
tegmina and wings hyaline, the first with the postcostal area
moderately broad, the basal cell about twice as long as broad,
and with eight apical areas; wing with five apical areas.
Type, TZ. nana, Walk. (Cicada).
This genus may be divided into two sections by the length
of the upper apical area to the tegmina, which is either longer
than eighth or about the same length. Further generic
division may be necessary when more material can be
examined.
ADENIA, gen. nov.
Head with the front very prominent, broadly conically
produced, very much shorter than vertex, which is sub-
quadrate and with its anterior and lateral margins straight, the
first completely at right angles with front; ocelli slightly
farther apart from each other than from eyes and well separated
from base of head; pronotum about as long as head, its
posterior angles very widely ampliate ; mesonotum (including
Mr. W. L. Distant on Homoptera. 2
cruciform elevation) as long as pronotum and head together,
globosely convex; abdomen somewhat broad, moderately
constricted at base, convex above, flattened beneath, the
second, third, and fourth segments very broad, flat, and tale-
like in appearance, fifth and sixth segments very compressed
and short ; opercula in male very short and transverse, not
reaching base of abdomen; tympana entirely uncovered ;
rostrum short, scarcely reaching intermediate coxee; tegmina
and wings hyaline, the first nearly three times as long as
broad, the postcostal area narrow but distinct, costal membrane
broad and somewhat sinuate, apical areas eight, the upper-
most very short and narrow; wings with five apical areas.
Type, A. Yerburyi, Dist.
Adena Yerburyi, sp. n.
Head and thorax above piceous, thickly greyishly pilose ;
front of head, margins and a central longitudinal fascia to
pronotum, and the margins of two obconical spots to meso-
notum obscurely ochraceous; abdomen above ochraceous,
greyishly pilose, basal area piceous, margins of segments
stramineous, and with a central darker longitudinal fascia ;
head beneath, sternum, and legs ochraceous and _ thickly
greyishly pilose; abdomen beneath with the second, third,
and fourth segments broad and talc-like, their margins and
a central longitudinal fascia pale ochraceous ; tegmina and
wings hyaline, the venation brownish ochraceous ; costal
membrane to tegmina dull ochraceous.
Long., excl. tegm., ¢ 16 mm.; exp. tegm. 40 mm.
flab. Aden (Col. Yerbury, Brit. Mus.).
Genus CALLIPSALTRIA.
Calopsaltria, Stal, Aun. Soc. Ent. Fr. (4) i. p. 620 (1861),
Calhpsatria, Stal, Hem. Afr, iv. pp. 9 & 49 (1866),
Type, C. longula, Stal (Cicada).
Genus CALYRIA.
Calyria, Stal, Rio Jan. Hem. ii. p. 22 (1858).
Type, C. cuna, Walk. (Cicada).
Genus PARNISA.
Parma, Stl, Rio Jan. Hem, ii. p. 21 (1858),
Type, P. proponens, Walk. (Cicada).
212 Mr. W. L. Distant on Homoptera.
MAPONDERA, gen. nov.
Allied to Tatpinga, but differing principally by the venation
of the wings, which possess only four apical areas,
Type, I. pulchella, Stal (Cicada).
In this genus I include the Tibicen (Quintilia) abdominals,
Stal, of which by the kindness of Dr. Aurivillius I have had
the opportunity to examine a co-type which possesses four
apical areas to the wings. Stal (Hem. Afr. iv. p. 37) writes
of this species: ‘Ale areis apicalibus numero varialibus,
tribus; quattuor vel quinque”; but he appears to have been
misled by the striking similarity in colour and markings of
several African species belonging to different genera, ‘These
I hope to figure in my ‘ Insecta ‘Transvaaliensia.’
Genus PRUNASIS.
Prunasis, Stal, Rio Jan. Hem. ii. p. 22 (1858).
Type, P. viridula, Walk. (Cicada).
Stal, in his synopsis of the genera of Cicadide (Hem. Afr.
iv. p. 9, 1866), gives as a distinguishing character of
Prunasis: “ Alis areis apicalibus sex.” This is erroneous, and
he had previously (supra) correctly given in his diagnosis
“ Ale areolis apicalibus quattuor.”
Division CHLOROCYSTARIA.
In this division the abdomen in the males is always more
or less inflated or dilated, with a central longitudinal dorsal
ridge, and longer than the space between the apex of head
and base of cruciform elevation ; the head (including eyes) is
a little narrower than base of mesonotum; tegmina always
longer than body, their greatest width only about a third of
their length.
Synopsis of Genera.
A. Tegmina with eight apical areas.
a, Wings with six apical areas.
6. Opercula in male of ordinary structure.
c. Lateral margins of pronotum obliquely
straight, posterior angles very slightly
amplhate.
d. Head with front not produced and
much broader than long, margins of
front and vertex continuous ...... Stagira.
dd. Head with front much subconically
produced, as long as broad at base,
Mr. W. L. Distant on Homoptera. 213
margins of front and vertex dis-
PTI NMOUS I Salat ot cia stecay sh oy aisha eas s Bavea.
ce, Lateral margins of pronotum a little
convexly sinuate, posterior angles
more or less strongly ampliate.
ddd. Head with front triangularly pro-
duced, as long as broad at base,
margins of front and vertex dis-
continuous.
e. Basal segment of abdomen strongly,
centrally, conically produced. . Musoda.
ee. Basal segment of abdomen not
conically centrally produced . Muda.
dddd, Head with front a little pr oduced,
a little broader than long, margins
of front and vertex discontinuous... Beturia,
bb. Opercula in male convex and inflated,
projecting beyond the lateral margins
of the abdomen.
f. Head with the front short and broad,
much broader than long, margins of
front and vertex continuous ........ Gymnotympana.
Jf. Head with the front narrow, about as
long as broad, margins of front and
vertex discontinuous ..........s00 Kumanga.
aa. Wings with five apical areas ............ Conibosa.
B. Tegmina with thirteen apical areas.
g- Wings with six apical areas .........4.. Chlorocysta.
C. Tegmina with eleven apical areas.
gg. Wings with five apical areas.
h, Apical areas of tegmina and wings of
modorate length: oases. dasee nas se Mardalana.
ggg. Wings with seven apical areas.
hh. Apical areas of tegmina and wings very
POU vartssy geass Seis nainls i slamareaytd Thaumastopsaltria,
DD: Tegmina with the venation of apical two
thirds reticulate, apparently forming nume-
rous cells ; wings with nine apical areas.... Cystopsaltria *.
Genus STAGIRA.
Stagira, Stal, Ann. Soc. Ent. Fr. (4) i. p. 621 (1861).
Type, S. s¢mplex, Germ. (Cicada).
Stagira Darwint, sp. n.
Body pale testaceous ; head beneath, sternum, femora, and
a basal spot to tegmina beneath ochraceous ; tarsi fuscous ;
tegmina and wings hyaline, the venation and costal mem-
brane to tegmina “ochraceous ; lateral areas of the last two
abdominal segments piceous ; rostrum reaching the inter-
mediate cox; space between costa and radial vein r ochraceous;
* I only know this genus from the description and figure (partial) of
Goding and Froggatt,
214 Mr. W. L. Distant on Homoptera.
seventh apical area to tegmina considerably longer than the
eighth ; apical ventral segment centrally carinate, moderately
apically produced.
Long., excl. tegm., g 18 mm.; exp. tegm. 38 mm.
Hab. Mauritius (C. Darwin, Brit. Mus.).
BAVEA, gen. nov.
9. Head (including eyes) narrower than base of meso-
notum, as long as pronotum, the front broadly subconically
produced, about as long as vertex, both centrally longitu-
dinally sulcate, their margins discontinuous, ocelli rather
nearer to eyes than to each other, front ocellus near anterior
margin of vertex, anterior lateral margin of vertex convexly
rounded ; pronotum with the lateral margins obliquely straight,
posterior angles very slightly ampliate ; mesonotum (including
cruciform elevation) much shorter than head and pronotum
together; abdomen (¢) moderately inflated; tegmina and
wings hyaline ; tegmina with eight apical areas and the basal
cell twice as long as broad; wings with six apical areas.
Type, B. concolor, Walk. (Cephaloxys).
Walker gave no habitat to his species which forms the type
of the genus, but the specimen (? ) is labelled South Africa
(Dr. Smith).
Genus Musopa.
Musoda, Karsch, Berl. ent. Zeitschr. xxxv. pp. 112 & 128 (1890).
Type, Jf. favida, Karsch.
The name flavida is based on faded or discoloured speci-
mens; fresh examples are bright grass-green.
Genus B&TURIA.
Beturia, Stal, Hem. Afr. iv. p. 9 (1866).
Type, B. conviva, Stal (Cicada).
Genus Mupa.
Muda, Dist. Ann, Mus. Civ. Genoa, (2*) xvii. p. 384 (1897).
Type, M. concolor, Dist.
Genus GYMNOTYMPANA.
Gymnotympana, Stal, Ann. Soc. Ent. Fr. (4) i. p. 619 (1861),
Type, G. strepitans, Stal (Cicada).
Mr. W. L. Distant on Homoptera. 215
IKUMANGA, gen. nov.
Head with the front narrow, about as long as broad, includ-
ing eyes narrower than the base of mesonotum; pronotum
narrowed anteriorly ; mesonotum (including cruciform eleva-
tion) slightly shorter than head and pronotum together ;
abdomen in male longer than the space between apex of head
and base of cruciform elevation ; tympana entirely exposed ;
opercula completely covering the sonorous cavities and pro-
jecting beyond the lateral margins of the abdomen as seen
from above; rostrum reaching the intermediate coxe ; tegmina
and wings hyaline ; tegmina with eight apical areas, seventh
and eighth about equally long, basal cell about twice as long
as broad; wings with six apical areas.
Type, K. sandaracata, Dist. (Beturia).
CONIBOSA, gen. nov.
Head as long as breadth between eyes, front triangularly
produced, as long as broad at base, its lateral margins and
those of vertex discontinuous, vertex longer than front, ocelli
well separated from basal margin and nearer to each other
than to eyes; pronotum a little longer than head, its lateral
margins straight, its posterior angles very little ampliated ;
mesonotum (including cruciform elevation) scarcely longer
than pronotum ; abdomen somewhat strongly dilated in male,
centrally arched and carinate, longer than space between
apex of head and base of cruciform elevation ; tympana
entirely exposed ; opercula in male type mutilated ; rostrum
reaching the intermediate cox; lower anal appendage to
male long, projecting beyond apex of abdomen; anterior
femora armed with three spines beneath ; tegmina and wings
hyaline ; tegmina with eight, wings with five apical areas ;
basal cell to tegmina not twice as long as broad.
Type, C. oecidentris, Walk. (Cephalorys).
Genus CHLOROCYSTA.
Cystosoma, subgen. Chlorocysta, Westw. Ann, & Mag. Nat. Hist. (2)
vii. p. 208 (1851).
Type, C. vitripennis, Westw. (Cystosoma [Chlorocysta]).
MARDALANA, gen. nov.
Chlorocysta, God. & Frogg. (part.), Proc. Linn. Soc. N.S. Wales, 1904,
p. 658.
Allied to Chlorocysta, Westw., but tegmina with eleven,
and wings with five apical areas; front of head more pro-
duced and less continuous with lateral margins of vertex ;
216 Mr. T. D. A. Cockerell— Descriptions and
ocelli much nearer to each other than to eyes ; lateral margins
of pronotum straighter, posterior angles much less (only
slightly) ampliated ; abdomen in male less inflated.
Type, I. congrua, Walk.
Mardalana congrua.
Cicada congrua, Walk. Journ. Ent. 1. p. 803 (1862).
Chlorocysta macrula, Stal, Tr. Ent. Soc. (3) i. p. 575 (1863); God. &
Frogg. Proc, Linn. Soc. N.S. Wales, 1904, p. 660.
Although I am deferring all synonymical notes for the pub-
lication of my ‘ Catalogue of the Cicadide,’ I think it right to
mention that in reply to Mr. Froggatt’s enquiry I stated that
I could not find Walker’s C. congrua in the Pascoe Collection
(cf. Proc. Linn. Soc. N.S. Wales, 1904, p. 657). It has
since been discovered, and proves to be a female specimen of
Stal’s species, and therefore takes precedence in nomenclature.
Genus THAUMASTOPSALTRIA.
Aecrilla, Stal, Trans. Ent. Soc. Lond. (3) i. p. 575 (1863), nom. preeoce.
Thaumastopsaltria, Kirk. Entomologist, 1900, p. 242, n. nom.
Type, ZT. adipata, Stal (Acrilla).
Genus CYSTOPSALTRIA.
Cystopsaltria, God. & Frogg. Proc. Linn. Soc. N.S. Wales, 1904,
p. 661.
Type, C. immaculata, God. & Frogg.
XXVI.—Descriptions and Records of Bees.—1.
By T. D. A. Cocxrrety, University of Colorado.
Excrpr where the contrary is stated, it is to be understood
that all the Australian species discussed in this series of
papers are in the collection of the British Museum, and all
others in my own collection.
Melissodes kallstremia, sp. n.
3 .—Length about 115 mm.
Black, with the abundant pubescence of the vertex and
thorax above orange-fulvous, without any black; that of
cheeks, face, and pleura whitish ; facial quadrangle not so
broad as long; eyes (dry) silvery grey; clypeus (except
Records cf Bees. PA 7
a black spot on each side) and a large patch on base of
mandibles bright lemon-yellow ; labrum yellowish white.
Antenne long (about 83 mm.) ; third jomt hardly longer
than second ; flagellum entirely very bright fulvous beneath;
fourth joint (first long joint of flagellum) long, much longer
than fifth; joints entirely without longitudinal keels. Meso-
thorax densely punctured; tegule fulvous, with fulvous
hair; hair of legs pale ochraceous (no black), that of hind
tibize and tarsi strongly but briefly plumose; hair on inner
side of tarsi ferruginous ; apices of middle and hind femora,
apices of all the tibiz, and the tarsi entirely, ferruginous ;
hind tibiz and basal joint of tarsi with black specks. Wings
strongly smoky, yellowish basally, nervures and stigma
rather dark ferruginous; second submarginal cell very large,
very nearly as large as the first, receiving the recurrent
nervure near the end; third submarginal cell narrowed
much more than half to marginal. Abdomen rather narrow,
with abundant coarse hair, erect and fulvo-ochraceous on
first segment ; ochraceous on second to fourth, but these
segments have also black hair on the apical middle, and the
fourth has black hair at base; fifth and sixth segments, and
apex, entirely black, with black hair, but pale hair from the
venter showing as a tuft on each side ; apical plate dark red;
hair of yenter pale, with a good deal of black on middle of
fourth and fifth segments, the apical segment dark reddish,
fringed with black. Easily known from M. aurigenia by
the colour of the abdominal pubescence ; from M. grindelie
by the longer antennz, colour of pubescence, &e.
Hab. Mesilla Park, New Mexico, at flowers of Kail-
stremia; July 27 (Cockerell).
It had previously visited an asclepiad, as shown by pollen-
masses adherent to two of the legs.
Melissodes kallstremie, var. phenacoides, nov.
&.—Abdomen broader, with the ochreous pubescence
replaced by white, except that the first segment has the hair
faintly yellowish ; the fifth segment has, in addition to the
black, a few long white hairs; the hair of the vertex and
thorax above is pallid, faintly tinged with ochraceous; the
third antennal joint is a little longer; the nervures are clear
ferruginous ; the claws are larger, those on the hind legs
being very large.
Hab. las Cruces, New Mexico, Aug. 25, 1894 (Cockerell,
2027).
This is the insect which has gone in the New Mexico list
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 15
218 Mr. T. D. A. Cockerell—Descriptions and
as M. communis, Cresson, having been so determined by
Mr. Fox. The total absence of black hair on the thorax,
the very large second submarginal cell, and the entirely
ferruginous tarsi, separate it sufficiently from VM. communis
of the Eastern States. The variety differs from the type
much as M. agilis differs from M. aurigenia.
Xenoglossodes gutierrexi@, sp. n.
?.—Agrees with X. eriocarpi (Ckll.), except in the
following characters :—
Larger (length 10 mm. or rather more) ; flagellum black,
and rather longer; mandibles with no yellow spot or patch ;
upper margin of yellow on clypeus straight ; labrum black ;
hind margin of first abdominal segment much more nar rowly
pallid.
Hab. Fillmore Cajion, Organ Mts., New Mexico, at
flowers of Gutierreziea (the hind Jegs are loaded with the
bright orange pollen), Aug. 29 (C. H. T. Townsend).
Diadasia enavata, Cresson.
At Mesilla, New Mexico, July 18, I took three females at
flowers of Helianthus annuus.
Crocisa Wallacei, sp. n.
$.—Length 9 mm. (but abdomen much retracted in
specimen measured) ; wing 9} mm.
Belongs to the group of C. histrio (scutellum with W-like
margin, abdomen with lateral marks not divided), and has
the blue patches consisting of very bright and shining scales,
in the manner of C. ceruleifrons. Ditters from C. nitidula
by having two blue spots on the scutellum, and the first
abdominal segment with very large quadrangular lateral
blue patches, leaving a black median band and the hind
margin, like a reversed T. The entirely black tarsi, without
any blue scales, and the smaller size separate it from
C. abdominals, Friese (from Java). The abundant blue
spotting on the thorax and other characters separate it
from C. nana, Friese. Apical half of labrum ferruginous ;
clypeus except anterior margin (which is broadly black and
densely punctate) and sides of face up to level of ocelli
covered with very brilliant silvery blue-green scales; middle
of front black and punctured, the punctures of two sizes,
not dense ; antenne black, the flagellum greyish; third
antennal joint a little shorter than fourth; front and vertex
Records of Bees. ZN
with black hair; thorax with blue or greenish-blue patches
as follows: avery large one on upper and a small trans-
verse one on lower part of pleura; a transverse one on each
side of prothorax above, joining one on contiguous part of
mesothorax; an elongate one on anterior middle, a round
one on each side, and an axe-shaped one at each hind corner
of mesothorax; a speck on each axilla; two spots on
scutellum ; interrupted bright blue bands on abdominal
segments 1 to 5, that on 1 as already indicated; venter of
abdomen entirely black, strongly punctate; apex truncate,
faintly subemarginate; tibie largely blue on outer side ;
tegule black, very densely punctured; wings fuliginous,
with a violet lustre.
Hab. Ternate; with numbers 6263 and 92-44.
Named after Dr. A. R. Wallace. In British Museum
Collection. It has a strong suyerficial resemblance to
C. ceruleifrons.
Crocisa lamprosoma, Boisd.
Queensland (/. Saunders) ; three.
Crocisa ceruleifrons, W. F. Kirby.
Friese makes this a synonym of C. quartine, Gribodo, but
Mr. Kirby’s name appears to have priority. It is recorded
from Timor Laut and Celebes; the Australian specimens
before me appear to be certainly the same species, but they
offer varietal or racial characters :—
Var. a.—A little larger; thoracic spots well-defined ; hair
of middle of occiput black, with white on each side ;
apex of male abdomen more narrowly truncate.
Queensland (H. Saunders),2 9,1.
Var. 8. Darwini.—Smaller, not over 9 mm. long; thoracic
spots greenish and less defined, confluent on anterior
part of mesothorax in ¢; occiput with a complete
conspicuous fringe of white hair; apex of male abdomen
more broadly truncate, the truncation concave ; wings
rather paler. The scutellum and tarsi are without
blue. Port Darwin, N. Australia, no. 90-126; one of
each sex. :
Crocisa tincta, sp. n.
? .— Length about 15 mm.
With the abdomen rather long and acuminate; upper
wings dark fuscous, lower hyaline; light spots consisting of
15*
220 Mr. ‘I’. D. A. Cockerell— Descriptions and
hair or elongated scales, which are not shiny; those on head,
thorax, and legs white or practically so, on abdomen pale
blue; margin of scutellum W-like; lateral abdominal patches °
not divided. A species of the C. histrio group, easily known
by its large size, and the absence of light marks on scutellum
or tarsi. Mandibles with a blunt tooth within; eyes straw-
yellow; face, cheeks, and occiput with much white hair;
front strongly and densely punctured; third antennal joint
conspicuously longer than fourth; wae part of pleura
covered with white hair, lower part nude, strongly and
closely punctured, with a small spot of white hair; meso-
thorax with anterior margin except in middle, and extending
on to prothorax, covered with white hair; lateral margins,
and posterior ones except in middle, a median stripe an-
teriorly, and a spot on each side, also marked with hair ;
tibize with large light patches ; tegument of abdomen slightly
purplish on middle of first two segments ; abdominal bands
all very broadly interrupted in middle, that on first segment
forming a large U-like mark on each side, that on fifth
reduced to a pair of large round spots ; venter without light
markings.
Hah. Toowoomba, Australia, no. 93. 189.
T'wo specimens. The locality is in 8.E. Queensland,
Trigona canifrons, Smith.
Adelaide River, Australia, no. 92-4. New to Australia.
The specimen differs from Bingham’s description of cani-
frons in having the hair of the pleura pale cinereous, and
the stigma and nervures fuscons; but a series of 7. canifrons
from Ceylon, received from Mr. Green, shows these charac-
ters, just as in the Australian insect. Judging from the
description, I suspect that 7. biroi, Friese, is a synonym.
Trigona essingtoni, sp. n.
Worker; length 4 mm.
Black (the abdomen brown-black), smooth and shining,
with pale yellow markings as follows: mandibles (except
ferruginous tips), labrum, clypeus (except narrow, black,
anterior and posterior margins, and two large red-brown
spots on disc), wedge-shaped lateral face- marks extending to
level of antenne, ‘tubercles, narrow stripe on each side of
mesothorax, extending to axill, all of scutellum, and apex
of abdomen, the last being thinly pubescent with white hair.
Antenne reddish brown, the flagellum ferruginous beneath ;
Records of Bees. 221
anterior knees with a yellow spot; small joints of tarsi
fulvous; face with a scanty pale pruinose Ole eae
thorax with very scanty pale hair; wings hyaline, nervure
and stigma flavous. Mandibles at apex with a concave
truncation, or minutely bidentate.
Hab. Port Essington, N. Australia, with no. 42. 1.
Two specimens. Easily known from T. australis, Friese,
by the entirely yellow scutellum, pale apex of abdomen,
markings of clypeus, &c.
Nomioides perditellus, sp. n.
? .—Length about 5 mm.
Head and thorax bluish green, shining ;_ pubescence
scanty, white, more abundant on apex and underside of
abdomen ; eyes strongly emarginate; clypeus light yellow
with two broad black bars (sometimes practically “obsolete) ;
mandibles yellowish with ferruginous tips; tubercles and
adjacent upper border of prothorax, and tegule, light
yellow; wings clear, nervures pale testaceous, first recurrent
nervure joining second transverso-cubital ; antenne dark
above and light beneath; femora black, with the knees
yellow ; tibice and tarsi yellow, the middle and hind tibize
largely clouded with black or brown ; abdomen broad,
piceous, with a large transverse yellow mark on each side
of segments 2 to 5, those on 5 nearly meeting to form a
coutinuous band; venter brown.
Hab, Queensland, marked “ Seaf. 1/90,” 438, at 92-16,
two specimens. A third i is marked Queensland, 2200, 75. 39.
Very distinet by the marking of the abdomen ; “es genus
is new to Australia. It is extraordinarily lke some of the
species of the American genus /erdita, agreeing even in
the details of the marking on the clypeus and abdomen ;
but the venation agrees with Halictus.
Thaumatosoma Duboulayi, Smith.
Port Darwin, N. Australia, 92-44; Bandin I., 91-82.
The latter specimen has the fourth and fifth abdominal
segments broadly margined with red.
Celioxys regine, sp. n.
2 .—Length a little over 11 mm.; expanse 195 mm.
Black, even to the tarsi; pubescence on eyes very short
elypeus shining, with extremely large close punctures ;
face covered with white hair ; vertex, mesothorax, and
222 Mr. T. D. A. Cockerell—Deseriptions and
scutellum with extremely large punctures, as close as
possible on scutellum, but well separated on middle of
mesothorax ; lateral teeth of scutellum well-developed,
broad and pointed, not or hardly curved; sides of meta-
thorax, tubercles, and margins of pleura with snow-white
hair; cheeks and occiput also with white hair, but vertex
bare; a small white hair-spot on each side of thorax above
and behind tegulz, and two short white stripes in scutello-
mesothoracie suture ; tegule black; wings strongly in-
fuscated on apical half, the basal part nearly clear; hair on
inner side of hind tarsi fulvous. Abdomen shining, with
strong punctures, quite close at the sides, but widely
separated above; lateral hind margins of the segments with
white hair-bands, which are not continued across the disk,
though on the ventral surface the bands are entire; last
dorsal segment with close minute punctures, strongly keeled
for rather more than two thirds of its length, the keel
abruptly failmg anteriorly, outline of the segment conical,
the apex not turned upwards, and falling far short of the
apex of the last ventral segment; last ventral segment
acuminato-conical in outline, not greatly narrowed or pro-
duced, but conspicuously notched on each side, shghtly con-
cave beneath; penultimate ventral segment with the anterior
part strongly and closely punctured, the posterior part finely
rugose.
Hab. Queensland ; with nos. 320 and 92-16.
Smith long ago remarked that it was singular that
Celiorys should be absent from Australia, while Megachile
is so abundant. It is therefore of special interest to record
two species from that country. W. A. Schulz has recently
described a species (C. Weinlandi) from New Guinea; it is
considerably larger than C. regine, and the last ventral
segment is very slender, hairy at the sides, and not notched.
Celioxys albolineata, sp. n.
2 —Length a little over 9 mm.
Similar to C. regine in appearance, but smaller, and
differmg as follows: middle of mesothorax shinirg, with
sparse small punctures, much smaller than those at the sides ;
scutellum flattened and somewhat concave, shining, and with
very sparse punctures, the disk nearly impunctate, and the
middle hind margin with a deep rounded notch ; on each side of
disk of scutellum, anteriorly, is a rather large oblique patch of
white hair; legs with conspicuous white hair; wings not so
dark ; atdomen with five entire, though narrow, white hair-
Records of Bees. 223
bands ; dorsal punctures sparser; last dorsal segment
narrower, with the keel shorter and less distinct; last ventral
segment not notched at the sides; penultimate ventral segment
with rather sparse, rather elongated punctures all over, or
they may be dense posteriorly.
Hab. Queensland (H. Saunders), with nos. 93-49 and 320.
Another bears no locality-label, but simply the number
1904-27.
Megachile cleomis, var. lippie, Ckll.
Mr. Wilmon Newell kindly sends me a female collected
by Mr. F. Beltran at Monterey, Mexico, where the species
is very destructive to rose-bushes. This extends the known
range about 600 miles S.E. The specimen is peculiar in
having the punctures of the middle of the mesothorax larger
and less dense than normal, and the ventral scopa lacks the
yellowish tinge. Possibly a series would prove that the
Monterey insect should be subspecifically separated.
Bombus gilgitensis, sp. n.
9.—Length about 23 mm.; wing about 16; abdomen
nearly parallel-sided, about 9 mm. (or a little over) wide and
almost 15 long.
Black, with black, dull white, and bright fulvo-ferruginous
hair; hair of head all black, except that on occiput it has a
purplish-brown tint ; hair of legs black, the apices of the tarsal
joints with short deep red spine-like bristles, hair on inner side
of basal joint of hind tarsi a very dark reddish ; thorax in front
with abundant dull white hair, a very broad band of purplish-
black hair between the wings; scutellum with dull white
hair mixed with black; base of abdomen with dull white
hair, covering first segment and basal halt of second, except
at sides, the rest, including apex, black, except the fourth
and fifth segments, which are covered with long fulvo-
ferruginous hair, the base of the fourth being overlapped by
the black of the third; ventral segments with very thin
fringes of long hair, black on the first three segments, fulvo-
ferrugmous on the fourth and fifth. Wings strongly clouded,
with a warm red tint. Head not elongated; malar space
broader than long. Middle of mesothorax with a large nude
shining impunctate area, the region surrounding it strongly
and closely punctured.
Hab. Gilgit, Kashmir; collector unknown tome. (British
Museum Collection.)
224 Descriptions and Records of Bees.
Allied to B. tunicatus, Smith, but the black bands on
thorax and abdomen are broader, and the last abdominal
segment has black hair instead of red. The upper level of
the black of the abdomen, which in twnicatus is straight, in
gilgitensis is strongly concave. The apical segment in gilgit-
ensis is bare above, minutely roughened but not noticeably
punctate, not carinate; at the sides the segment is densely
covered with brown-black tomentum. The wings are not so
dark as those of B. orientalis, Smith.
AXylocopa varipuncta, Patton.
Texas, no. 1977, g. (Colorado Agricultural College
Collection.) New to Texas.
Xylocopa bryorum (Fabr.), subsp. dimidiata (Lep.).
Six 2s, Queensland (E. Saunders) ; Condillac I1., W. Aus-
tralia; Port Darwin. One @, Queensland (EH. Saunders).
The male is a little over 21 mm. long, and the middle tibiz
and tarsi are without dark hair. The female is of the same
size; the yellow pubescence spreads on to the occiput and
the upper part of the pleura, but not at all on to the basal
segment of the abdomen. ‘This insect appears to be inter-
mediate between X. bryorum and X. estuans, as defined by
Bingham ; it is probable that it should be recognized as a
subspecies, In which case the name dimidiata, Lep., is
apparently available.
Lestis erata, Smith.
3 38, Queensland (E. Saunders) ; Sydney. 7 2s, Queens-
land (EH. Saunders) ; N.S. Wales.
Synhalonia lippie (Ckll.).
Synhalonia crenulaticornis, subsp. lippie, Ckll., Ann. & Mag. Nat. Hist.,
July 1904, p. 25.
A female collected by Prof. Townsend at the same time
and place as the original males shows that lippie@ is a distinct
species. Compared with 92 crenulaticornis it is larger (about
12 mm. long), with much longer and ampler wings, which
are not quite so dark, and the pubescence of the thorax above is
white with a very faint yellowish tinge, becoming pronounced
on scutellum, but with no fuscous or black, whereas crenu-
laticornis shows much dark fuscous hair on mesothorax and
scutellum. The abdomen also has more white tomentum,
covering the base of the second and third segments, while
~
On the American Cichlid Genus Cichlosoma. 225
b
the apical band on the second is broadly interrupted in the
middle, and the tomentum is of a sort of bluish white, a
different tint from that of crenulaticornis. The brush at the
end of the basal joint of the hind tarsi, which is black in
crenulaticornis, is reddish brown in lippie.
I have a @S. lippie which I collected at flowers of
Phacelia congesta, at Dripping Spring, Organ Mts., New
Mexico, in August.
XXVII.—A Revision of the Fishes of the American Cichlid
Genus Cichlosoma and of the Allied Genera. By C. Varn
Reean, B.A.
[Continued from p. 77. }
Section 6 (Vheraps).
Body ovate or elongate. Upper profile of snout usually
convex. Scales of the lateral line of the same size as those
above and below it; scales of the thoracic region con-
siderably smaller than those on the side of the body.
Mouth small or moderate (maxillary not extending to below
the eye), moderately protractile (premaxillary processes not
extending to above the eye, in the adult) ; maxillary not
or slightly exposed ; teeth of the outer series moderate,
distinctly enlarged anteriorly. Dorsal XV-XVIII 10-15,
the soft fin more or less distinctly scaly at the base.
Anal IV-VII 8-11. Pectoral not extending to above the
anal. Caudal rounded, truncate, or emarginate with rounded
lobes.
Seventeen species from Mexico and Central America, one
extending into Colombia.
13. Cichlosoma Eigenmanni.
Cichlasoma Eigenmanni, Meek, Zool. Pub. Columbian Mus. iii. 1902,
p: 119, pl. xxx., and vy. 1904, p. 220, pl. xvi.
Depth of body 23-23 in the length, length of head 31-3}.
Snout eqnal to or a little shorter than postorbital part of
head. Diameter of eye 34-41 in the length of head, inter-
orbital width 23-8. Depth of preorbital 1-1} the diameter
of eye. Maxillary not extending to below theeye; lower jaw
shorter than the upper ; fold of the lower lip not continuous ;
cheek with 6 or 7 series of scales; 8 gill-rakers on the lower
226 Mr. C. T. Regan on the Fishes of the
part of anterior arch. Scales 34-35 — 3 or 34 between
lateral line and base of anterior rays of soft dorsal. Dorsal
XVII-XVIII 12-15, commencing above the opercular cleft,
the spines subequal from about ‘the oe to the fifteenth,
thence increasing to the last, which is 4 the length of head ;
soft fin, when laid back, extending to base of caudal.
Anal V-VI 8-10. Pectoral ? the length of head, not
extending so far as the ventral, which nearly reaches the
origin of anal. Caudal truncate. Caudal peduncle about
as long as deep. Olivaceous, with obscure darker cross-bars
and with a blackish longitudinal band from operculum to
base of caudal, where it ends in a dark spot; vertical fins
dusky.
Southern Mexico.
1-4. (112-235 mm.) tio Tonto, La Raya. Dr. H. Gadow.
14. Cichlosoma nebuliferum.
Heros nebulifer, Giinth. Proc. Zool. Soc. 1860, p. 318, and Cat. Fish.
iv. p. 297 (1862).
Heros gibbiceps, Steind. Denkschr, Ak. Wien, xxiii. 1864, p. 66, pl. v.
figs. 1 & 2; Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898,
p. 15 36; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 232 (1904).
Cichlasoma nebuliferum, Jord. & Everm. ¢. e. p. 1524; Pellegr. ¢. ¢.
p. 216; Meek, Zool. Pub. Columbian Mus. v. 1904, p. 220.
Cichlasoma teapae, Everm. & Goldsborough, Bull. US. Fish. Comm.
xxi. 1902, p. 156, fig.; Pellegr. ¢. ¢. p. 213.
Depth of body 23-22 in the length, length of head 33-4.
Snout shorter than postorbital part of head. Diameter of
eye 4—41 in the length of head, interorbital width 3. Depth
of preorbital 1-1} the diameter of eye. Maxillary not nearly
extending to vertical from anterior margin of eye; lower
jaw shorter than the upper; fold of the lower lip not
continuous; cheek with 5 or 6 series of scales; 8 or 9
gill- ao on the lower part of anterior arch. Scales
33-35 ;5°, 34 or 4 between lateral line and base of anterior
part of soft dorsal. Dorsal XVII-XVIII 12-13 (14),
commencing above extremity of operculum, the spines sub-
equal from about the seventh to the fourteenth, which are
3 the length of head, thence increasing to the last, which is
2 the length of head ; the soft fin, when laid back, extending
to base of caudal. Anal VI 9 (10). Pectoral # the length
of head; ventral as long, not quite reaching the origin of
anal. Caudal truncate or subtruncate. Caudal peduncle as
long as or longer than deep. Olivaceous, each scale with a
dark brown intra-marginal line ; a series of dark blotches
American Cichlid Genus Cichlosoma. pA)
on the side, usually more or less confluent to form a longi-
tudinal band ; a dark spot on the base of caudal; vertical fins
dusky, the dorsal with some light spots posteriorly.
Southern Mexico.
1-8. (147-172 mm.) types of the species. Mexico. M. Salle,
15. Cichlosoma maculicauda, sp. n.
Heros parma (part.), Guoth. Cat. Fish. iv. p. 285 (1862), and Trans.
Zool. Soe. vi. 1869, p. 449.
Cichlasoma parma (part.), Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1519.
Cichlasoma fenestratum, var. parma, Pellegr. Mém. Soc. Zool. France,
xvi. 1903, p. 210 (1904).
Depth of body 12-2 in the length, length of head 23-31.
Snout as long as the postorbital part of head (in the aault),
or a little shorter (in the young). Diameter of eye 3-4? in
the length of head, interorbital width 21-23. Depth of
preorbital 2-1} the diameter of eye. Maxillary extending
to between nostril and eye; jaws equal anteriorly; fold of
the lower lip not continuous ; cheek with 4 or 5 series of
scales; 8 or 9 gill-rakers on the lower part of the anterior
areh. ene 32-35 a ae 4 between lateral line aud base of
anterior part of soft dorsal. Dorsal* XVI-XVII 12-14,
commencing above the extremity of operculum, the spines
only slightly i increasing after the sixth, the last 2 or nearly
2 the Ween of head; soft fin, when laid back, extending to
oeete + (young) or beyond the middle (adult) of conde
Anal VI-VII 9-10. Pectoral a little shorter than the head,
extending to above the vent; ventral extending to the
origin of anal or beyond. Caudal subtruncate. Caudal
peduncle 3-? as long as deep. Sides of body and vertical
fins usually with dark spots; a large dark blotch on the
caudal pedunele.
Central America.
1-4, (185-189 mm.) types of the Lake Yzabal. O. Salvin, Esq.
species.
5, (223 mm.) Rio Motagua. O. Salvin, Esq.
6-10. (205-301 mm.) Rio Chagres, O. Salvin, Esq.
11-13. (68-90 mm.) Rio Chagres, O. Salvi in, Esq.
16. Cichlosoma fenestratum.
Chromis fenestrata, Giinth. Proce. Zool. Soe. aut p. 318.
Heros parma (part.), Giinth, Cat. Fish. iv. p. 285 (1862), and Trans.
Zool. Soe. vi. 1869, p. 449.
* Of 13 specimens 10 have XVII 13, 2 have XV II 12, and 1 has
XVI 14 dorsal rays. 8 have VI 10, 4 rae VI 9, and 1 has VII 10
anal rays.
226 Mr. C. T. Regan on the Fishes of the
Heros fenestratus, Giinth. t. ce. p. 286; Steind. Denkschr. Ak. Wien,
xxill. 1864, p. 68, pl. i. fig. 2.
Cichlasoma fenestratum, Jord. & Everm. Bull. US. Nat. Mus. xlvii.
1898, p. 1518.
Cichlasoma parma (part.), Jord. & Everm. t. ec. p. 1519.
Cichlasoma fenestratum (part.), Pellegr. Mém. Soe. Zool. France, xvi.
1903, p. 210 (1904).
Se parma, Meek, Zool. Pub, Columbian Mus, v. 1904, p. 218,
p . XV.
Depth of body 13-2 in the length, length of head 23-81.
Snout shorter than the postorbital part of head. Diameter
of eye 31-42 in the length of head, interorbital width 21-232.
Depth of preorbital 2-14 the diameter of eye. Maxillary
extending to between nostril and eye ; jaws equal anteriorly ;
fold of the lower lip not continuous ; cheek with 5 or 6
series of scales ; 8-10 gill-rakers on the lower part of the
anterior arch. Scales 31-83 2, 3 or 34 between lateral
line and base of anterior part of soft dorsal. Dorsal *
XVII-X VIII 11-13, commencing above the opercular cleft,
the spines only slightly increasing after the sixth, the last
3 the length of head ; soft fin, when laid back, extending to
anterior part (young) or beyond the middle (adult) of
caudal. Anal VI-V1I 8-9. Pectoral a little shorter than
the head, extending to above the vent; ventral extending
nearly to origin of anal or slightly beyond. Caudal sub-
truncate. Caudal peduncle }—} as long as deep. Brownish,
usually with 6 to 8 dark cross-bars aud with a dark longi-
tudinal band from operculum to the basal part of caudal.
Sometimes this band is very strongly marked, and the cross-
bars may then be confined to the upper part of the body, or
may be wanting.
Mexico.
1-2. (98 and 145mm.) types Mexico. M. Salle.
of the species.
8-5. (77-103 mm.) Mexico. M. Sallé.
6-8. (186-228 mm.) types Mexico. Berlin Mus.
of H. parma.
9, (57 mm.) Chimalapa. Dr. A. C. Buller.
11-16. (52-181 mm.) Rio Tonto, La Raya. Dr. H. Gadow.
17. (128 mm.) Motzorongo. Dr. H. Gadow.
17. Cichlosoma bifasciatum.
Heros bifasciatus, Steind. Denkschr. Ak. Wien, xxiii. 1864, pt. ii.
. 60, pl. i.
aan ee bifasciatum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1521; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 217
(1904).
* Of 17 examples 8 have XVII 12, 3 have XVII 11, 2 have XVII 13,
3 have XVIII 12, and 1 has XVIII 18 dorsal rays. 10 have VI 9,
5 have VI 8, one has VII 8, and one VIL 9 anal rays.
American Cichlid Genus Cichlosoma, 229
Depth of body 2 in the length, length of head 34. Snout
as long as the postorbital part of head. Diameter of eye 5
in the length of head. Depth of preorbital 14 the diameter
of eye. Maxillary not extending to below the eye; jaws
equal anteriorly; fold of the lower lip not continuous ;
cheek with 5 or 6 series of scales ; 9 or 10 gill-rakers on the
lower part of the anterior arch. About 34 scales in a longi-
tudinal series. Dorsal XVII 138, commencing above the
opercular cleft, the spimes subequal from the sixth, the last
g the length of head. Anal VI 9. Pectoral not extending
to above the anal. Caudal truncate. Caudal peduncle as
long as deep. Two broad dark longitudinal bands, the
upper from a little behind the origin of the lateral line to
the end of the base of the dorsal fin, the lower from the
pectoral to the base of the caudal.
Mexico.
The type measures 240 mm. in total length.
18. Cichlosoma guttulatum.
Heros guttulatus, Giiuth. Proce. Zool. Soc. 1864, p. 152, and Trans.
Zool. Soc. vi. 1869, p. 466, pl. Ixxviii. fig. 3.
Cichlasoma guttulatum, Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p- 214 (1904).
Depth of body 23-23 in the length, length of head 25-32.
Snout as long or nearly as long as the postorbital part of
head. Diameter of eye 8—42 in the length of head, inter-
orbital width 23-24. Depth of preeorbital 3-14 the diameter
of eye. Maxillary not extending to the vertical from
anterior margin of eye ; jaws equal anteriorly ; fold of the
lower lip not continuous; cheek with 5 series of scales ;
8-10 gill-rakers on the lower part of anterior arch. Scales
31-34 2%,, 33-43 between lateral line and base of anterior
part of soft dorsal. Dorsal XVI-XVIII 12-14, com-
mencing above or in front of the axil of pectoral, the spines
subequal from the fifth or sixth to the fourteenth or fifteenth,
which are 4 the length of head, the last 2 the length of
head ; the soft fin, when laid back, extending to anterior |
vo
of caudal. Anal VI-VII 9-10. Pectoral 3 the length cf
head, not extending as far as the ventral, which nearly or
quite reaches the origin of anal. Caudal subtruncate or
rounded. Caudal peduncle as long or nearly as long as
deep. Brownish, a broad dark band extending from above
pectoral to base of caudal; sides of head spotted; body
often spotted ; sometimes 5 or 6 cross-bars on the upper
part of the body, or a series of blotches along the lateral
line; vertical fins dusky, often with dark spots.
Guatemala ; Southern Mexico.
230 Mr. C. T. Regan on the Fishes of the
1-9. (69-176 mm.) types Lake Amatitlan. O. Salvin, Esq.
of the species.
10. (150 mm.) Nr. Pacific Coast of O. Salvin, Esq.
C. America.
11-19, 20-25. (62-221 mm.) Lake Nacasil. O. Salvin, Esq.
26, (211 mm.) tio de Sarabia. Dr. A. C. Buller.
19. Cichlosoma microphthalmus.
ys
Heros microphthalmus, Giinth, Cat. Fish, iv. p. 295 (1862), and Trans.
Zool. Soc. vi. 1869, p. 464; Jord. & Everm. Bull. U.S. Nat. Mus.
xlvii. 1898, p. 1586; Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p. 282 (1904).
Depth of body 24-23 in the length, length of head 31—
Snout veal as long as postorbital part of head. Dice
of eye 44-5} in the jength of head, interorbital width 22-23.
Depth of prieorbital 14 -1? the diameter of eye. Maxillary
extending to the vertical from anterior mar gin of eye; jaws
equal anteriorly ; fold of the lower lip continuous; cheek
with 5 or 6 series of scales; 7 or 8 gill-rakers on the lower
part of the anterior arch, Scales 32-34 7", 34 or 4: between
lateral line and base of anterior part of soft dorsal. Dorsal
XVIII 11-13, commencing above the axil of pectoral, the
spines subequal from the seventh to the fourteenth, which
are + the length of head, thence increasing to the last, which
is more than. 2 the length of head; the ‘soft fin, when laid
back, extending to anterior part or middle of caudal. Anal
V-VI* 9-11. Pector al # the length of head, not extending
so far as the ventral, which does not reach the origin of anal.
Caudal subtruncate. Caudal peduncle #-¢ as long as deep.
Brownish, usually with 4 or 5 irregular dark cross-bars ; a
dark spot on the base of caudal; vertical fin with series of
dark spots or dark undulating lines.
R. Motagua, Guatemala.
1-3, 4-5. (165-195 mm.) types of the RR. Motagua. _O. Salvin, Esq.
species.
20. Cichlosoma sexfasciatum, sp. n.
Depth of body 2} in the length, length of head 32. Snout
as long as postorbital part of head. Diameter of eye 5} in
the length of head, interorbital width 2}. Depth of pre-
orbital 13 the diameter of eye. Maxillary extending to
between nostril and eye; jaws equal anteriorly; fold of the
lower lip not continuous; cheek with 6 series of scales;
7 or 8 gill-rakers on the lower part of the anterior arch.
* One specimen only has 5 anal spines.
American Cichlid Glenus Cichlosoma. 231
Scales 334, 33 between lateral line and base of anterior
part of soft dorsal. Dorsal XVIII 11, commencing above
the extremity of operculum, the spines slightly increasing in
length to the last, which is a little more than 2 the length of
head ; soft fin, when laid back, extending to anterior } of
caudal. Anal VI 9. Pectoral ? the length of head, not
extending to above the vent; ventrals extending to the
vent. Caudal subtruncate. Caudal peduncle 2 as long as
deep. Olivaceous, with small dark spots; 6 dark cross-bars
on the side, and a dark longitudinal band from operculum to
base of caudal.
Guapote, Mexico.
1, (223 mm.) type of the species. Guapote. Mexico Mus,
21. Cichlosoma melanurum.
Heros melanurus, Giinth. Cat. Fish. iv. p. 288 (1862), and Trans. Zool.
Soe. vi. 1869, p. 450, pl. lxxii. fig. 3.
a ee Steind, Denkschr. Ak. Wien, xxiii. 1864, p. 72,
.1. fig. 8.
Ghuins melanopogon, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1523.
Cichlasoma melanurum, Jord, & Kverm. J. c.; Pellegr. Mém. Soe. Zool.
France, xvi. 1903, p. 212 (1904).
Depth of body 23-22 in the length, length of head 3-31,
Snout as long as postorbital part of head. Diameter of eye
3-5 in the length of head, interorbital width 23-3. Depth
of preorbital 2-1} the diameter of eye. Maxillary not
extending to below the eye; jaws equal antcriorly ; fold of
the lower lip subcontinuous or not continuous; cheek with
5 or 6 series of scales ; 7 or 8 gill-rakers on the lower part
of anterior arch. Scales 31-34 no 3 between lateral line
and base of anterior rays of soft dorsal. Dorsal (XVI)
XVIT-XVIII 11 (12), commencing above the opercular
cleft, the spines subequal from about the sixth to the tenth,
thence sometimes slightly decreasing to the fourteenth or
fifteenth and increasing again to the last, which is 2-1 the
length of head; soft fin, when laid back, extending a little
beyond base of caudal. Anal V-VI 8-9. Pectoral about Pi
the length of head, extending about to above the vent ;
ventral extending to origin of anal. Caudal subtruncate.
Caudal peduncle #—3 as long as deep. Olivaceous, with
2 series of blackish spots which may unite to form con-
tinuous longitudinal bands, the first below the spinous dorsal
and above the lateral line, the second from the middle of
the side to the base of caudal ; vertical fins dusky.
Guatemala ; Southern Mexico.
1-5, (85-260 mm.) types of the species. Lake Peten. O. Salvin, Esq.
bo
Oo
NS
Mr. C. T. Regan on the Fishes of the
22. Cichlosoma Gadovii, sp. n.
? Cichlasoma melanurum (non Giiuth.), Meek, Zool. Pub. Columbian
Mus. y. 1904, p. 219.
Depth of body 24-22 in the length, length of head 3-3}.
Snout a little shorter than postorbital part of head:
Diameter of eve 4-4} in the length of head, interorbital
width 21-22. Depth of preorbital 1-1} the diameter of eye.
Maxillary not extending to below the eye; jaws equal
anteriorly ; fold of the lower lip not continuous; cheek with
5 series of scales ; 2 gill-rakers on the lower part of anterior
arch. Seales 302 =» 8 between lateral line and base of
anterior rays of soft dorsal. Dorsal XVI-XVII 12, com-
mencing above the extremity of operculum, the spines sub-
equal from the sixth or seventh to the thirteenth or four-
teenth, thence increasing to the last, which is a little less
than } the length of head ; the soft fin, when laid back,
extending to anterior $ of caudal. Anal VI 9. Pectoral 2
the length of head, extending nearly to above the vent ;
ventral extending nearly to origin of anal. Caudal sub-
truncate. Caudal peduncle }-+ as long as deep. Olivaceous,
with obscure darker cross-bars “and a dark longitudinal band
from operculum to base of caudal; vertical fins dusky.
Southern Mexico.
1-2. (190 and 241 mm.) types of Motzorongo. Dr. H. Gadow.
the species.
23. Cichlosoma intermedium.
Heros intermedius, Gunth. Cat. Fish. iv. p. 298 (1862), and Trans.
Zool. Soe. vi. 1869, p. 468, pl. xxviii. fig. 1.
Heros angulifer, Giinth. lc. and t.¢. p- 469, pl lixv. sie ae
Acara rectangular, Steind. Denkschr. Ak. Wien, xxiii. ” 1864, py Ot;
pla figs 1.
Guilasoice rectangulare, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1515: Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 205
(1904) ; Meek, Zool. Pub. Columbian Mus. v. 1904, p. 216.
Cichlasoma anguliferum, Jord. & Everm. t. e. P. 1517.
Cichlasoma inter medium, Jord. & Everm. 7. c.; Pellegr. t. c. p. 209.
Depth of body 21-23 in the length, eh of head 3-3}.
Snout as long or nearly as long as postorbital part of head.
Diameter of eye 37-5 in the length of head, interorbital
width 24-24. Depth of preorbital 3-12 the diameter of
eye. Maxillary not extending to the Vertical from anterior
margin of eye; jaws equal anteriorly ; ; fold of the lower lip
not continuous ; cheek with 4-6 series of scales ; 7 or 8 gill-
rakers on the lower part of anterior arch. ee 31-33 2,
American Cichlid Genus Cichlosoma. 233
2 or 24 between lateral line and base of anterior part
of soft dorsal. Dorsal (XVI-XVII) XVIII 10-12 (13),
commencing above the axil of pectoral, the spines subequal
from about the seventh to fourteenth, which are about } the
length of head, thence increasing to the last, which is ? the
length of head or more; the soft fin, when laid back, ex-
tending to anterior } of caudal. Anal (IV) V—VI 8-10 (11).
Pectoral #-3 the length of head, usually not extending so
far as the ventral, which does not quite reach the origin of
anal. Caudal subtruncate. Caudal peduncle a little deeper
than long. Brownish, with a broad dark band running
from the operculum to the middle of the side and then
upwards to the posterior part of the spinous dorsal ; vertical
fins usually with dark stripes or series of spots.
Guatemala.
a. C. intermedium.
Scales forming the dark band each light at the base and
with a blackish vertical stripe near the posterior margin; a
narrow longitudinal band from the angle of the broader
band to the base of caudal, where it forms a vertically
expanded spot.
1-3. (183-155 mm.) types of the Lake Peten. Q. Salvin, Esq.
species.
b. C. anguliferum.
Scales forming the dark band nearly uniformly blackish ;
a dark blotch on the caudal peduncle.
1-2. (96 and 113 mm.) types Rio de Santa Isabel. —O.. Salvin, Esq.
of H. angulifer.
3-5, (134-177 mm.). R. Sta, near Cajabon. J.C. Sarg, Esq.
24. Cichlosoma Guenthevi.
Heros oblongus (non Casteln.), Giinth. Trans. Zool. Soc. vi. 1869, p. 464 ;
Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898, p. 1535.
Cichlasoma Giintheri, Pellegr. Mém. Soc. Zool. France, xv 1903,
p. 215 (1904).
Depth of body 22-23 in the length, length of head 3}-32.
Snout as long or nearly as long as postorbital part of head.
Diameter of eye 32-43 in the length of head, interorbital
width 24-3. Depth of preorbital 1-1} the diameter of eye.
Maxillary not extending to the vertical from anterior margin
of eye; jaws equal anteriorly ; fold of the lower lip con-
tinuous or not; cheek with 5 series of scales; 7 or 8 gill-
rakers on the lower part of anterior arch. Scales 32-33 =
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 16
234 Mr. C. T. Regan on the Fishes of the
34 or 4 between lateral line and base of anterior part of soft
dorsal. Dorsal XVIII 13, commencing above or slightly
before axil of pectoral, the spines subequal from about the
sixth to the fourteenth, which are } the length of head,
thence increasing to the last, which is more than 2 the
length of head; soft fin, when laid back, extending to
anterior part or even to middle of caudal. Anal VI 9 10.
Pectoral 4 the length of head, not extending as far as the
ventral, which nearly or quite reaches the origin of anal.
Caudal subtruncate. Caudal peduncle nearly as long as
deep. Brownish, with 5 or 6 obscure darker cross-hars,
and with a more or less distinct longitudinal band ending in
a dark spot on the base of caudal; vertical fins with series
of dark spots or dark undulating lines.
Rio Motagua, Guatemala.
1-2. (186 and 209 mm.) types of Rio Motagua. O. Salvin, Esq.
the species.
3-4, (108 and 131 mm.) Rio Motagua. O. Salvin, Esq.
25. Cichlosoma pavonaceum.
Heros pavonaceus, Garman, Bull. Mus. Comp. Zool. viii. 1881, p. 93 ;
Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898, p. 1538; Pellegr.
Mém. Soe. Zool. France, xvi. 1903, p. 250 (1904).
Cichlasoma pavonaceum, Meek, Zool. Pub. Columbian Mus. v. 1904,
p- 209.
Depth of body equal to length of head, 23 in the length.
Eye large, its diameter greater than depth of preeorbital.
Maxillary not extending to below anterior margin of eye;
jaws equal anteriorly ; fold of the lower lip not continuous ;
cheek with 5 series of scales ; 7 or 8 gill-rakers on the lower
part of the anterior arch. Scales 82 ( ae Dorsal XVI 12,
the spines subequal from the fifth, the last 2-} the length of
head ; soft fin, when laid back, extending fond base of
caudal. Anal V 8. Pectoral extending to above the vent ;
ventral to or nearly to origin of anal. Caudal rounded.
Brownish, with 10 or 11 obscure dark cross-bars, and with a
series of 4—6 blackish spots, usually ocellated and vertically
expanded, below the dorsal fin in the posterior part of the
body ; a blackish ocellated spot on the upper half of the base
of caudal.
Monclova, Mexico.
The types measure from 30-85 mm. The species is said
to be allied to C. intermedium.
ho
Co
wt
American Cichlid Genus Cichlosoma.
26. Cichlosoma Sieboldii.
Beros Sieboldii, Kner & Steind. Abhandl. Bayern Ak. x. 1864, p. 13,
pl. ii. fig. 2.
Cichlasoma Sieboldti, Jord. & Everm. Bull. 0.8. Nat. Mus. xlvii. 1898,
p. 1516.
Depth of body about 2} in the length, length of head
about 33. Snout as long as postorbital part of head.
Diameter of eye 4—5 in the Jength of head and 13-23 in the
interorbital width. Maxillary not extending to. below the
eye; jaws equal anteriorly; fold of the lower lip not con-
tinuous ; cheek with 5 series of scales; 8 or 9 gill-rakers
on the lower part of the anterior arch. Scales 30-32 =
24 between lateral line and base of anterior part of soft
dorsal. Dorsal XVII 11, the spines increasing in length to
the last, which is about 4+ ‘the length of head ; ‘soft fin, when
laid hack, ee nearly to middle of bard! eal V 8.
Pectoral 2—? the length of head ; ventral extending to the
vent, Caudal subtruncate or slightly rounded. Caudal
peduncle about + as long as deep. Reddish brown; each
scale with a dark spot; 5 or 6 dark cross-bars forming a
series of blotches on the upper lateral line and a second
larger series on the middle of the side; vertical fins dark
greyish with series of blackish spots.
Panama ; Colombia.
The types measure from 125 to 200 mm. in total length.
Cichlosoma Deppii, Heckel, from Mexico, appears to be
closely allied to aos species. D.XVI-XVII 10-11. A. V-
VI8. Se. 30 52; Brownish, with 5 or 6 dark cross-bars
posteriorly, the last forming a spot at the base of caudal.
The synonymy of this species is as follows :—
Heros Deppit, Heck. Ann. Mus. Wien, ii. 1840, p. 382; Ginth. Cat.
Fish. iv. p. 296 (1862).
Heros Montezuma, Heck. t. ec. p. 883; Giinth. . c.
Cichlasoma Deppii, Jord. & Everm, Bull. U.S. Nat. Mus. xlvii. 1898,
p. 1524; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 215 (1904) ;
Meek, Zool. Pub. Columbian Mus. v. 1904, p. 221.
Cichlasoma Montezuma, Jord. & Everm. f. c. p. 1528; Meek, /. ec.
27. Cichlosoma Godmanni.
Heros Godmanni, Giinth. Cat. Fish. iv. p. 296 (1862), and Trans. Zool.
Soe. vi. 1869, p. 466, pi. Ixxiv. fig. 5.
Cichlasoma Godmannt, Tord, & Everm. Bull. U.S. Nat, Mus. xlvii. 1898,
p. 1516; Pellegr. Mem. Soe. Zool. France, xvi. 1903, p- 208 (1904).
Depth of body 22~—2} in the length, length of head 3}.
Snout slightly shorter than postorbital part of head.
Lom
236 Mr. C.'T. Regan on the Fishes of the
Diameter of eye 42 in the length of head, interorbital width
24, Depth of preorbital 14 the diameter of eye. Maxillary
not extending to the vertical from anterior margin of eye;
jaws equal anteriorly or the lower shorter than “the upper ;
fold of the lower lip continuous or not; cheek with 6 or 7
series of scales; 7 or 8 gill-rakers on the lower part of
anterior arch. Scales 32-33 a 3 or 33 between lateral
line and base of anterior part of soft dorsal. Dorsal X VI-—
XVII 12-13, commencing above axil of pectoral, the spines
slightly increasing in length to the last, which is } the length
of head, the soft. fin, w Hen laid backs extending to anterior
i of caudal. Anal V 10. Pector al 3—% the length of head,
not extending so far as the ventral, aie does not reach the
origin of anal. Caudal subtruncate. Caudal peduncle a
little deeper than long. Olivaceous, with a dark longitu-
dinal band along the middle of the side, ending in a blackish
spot on the base of the caudal; a blackish spot above the
origin of the band and below the lateral line; vertical fins
with series of dark spots.
Guatemala.
1-2. (175 and 176 mm.) types of Rio Cahabon. O. Salvin, Esq.
the species.
28. Cichlosoma irregulare.
Theraps trregularis, Giinth. Cat. Fish. iv. p. 284 (1862); Jord. &
Everm. Bull. U.S. Nat. Mus. xlvii. 1898, p. 1540.
Heros tirregularis, Giinth. Trans. Zool. Soe. vi. 1869, p. 467, pl. xxviii.
fig. 2.
Cichlasoma wregulare, Pellegr, Mém. Soc. Zool. France, xvi. 1903,
p. 204 (1904).
Depth of body 3-8 in the length, length of head 31-32.
Snout longer than postorbital part “of head (in the adult).
Diameter of eye 4-5 in the length of head, interorbital
width 3-4. Depth of preeorbital je 1} the diameter of eye.
Maxillary not extending to below the eye; lower jaw shorter
than the upper; fold of the lower lip not continuous; cheek
with 6 or 7 series of scales ; 9-11 gill-rakers on the lower
part of anterior arch. Scales 33-36 a 3 between lateral
line and base of anterior rays of soft dorsal. Dorsal X V-
XVII 12-14, commencing behind the axil of pectoral, the
spines subequal from about the fifth to the thirteenth, thence
usually mcreasing to the last, which is 4-2 the length of
head; soft fin, when laid back, not extending to base of
caudal. Anal*-IV-V 9-10. Pectoral about 2 % the length
* 4 anal spines in the type specimen oft
American Cichlid Genus Cichlosoma. 237
of head, not extending so far as the ventral, which nearly
reaches the origin of anal in the young, but not in the adult.
Caudal subtruncate. Caudal peduncle 13-12 as long as
deep. Brownish, with blackish cross-bars bearing a series
of blotches below the lateral line, which may be more or less
confluent to form a longitudinal band; some light blue spots
on the side of the head. Vertical fins dusky.
Guatemala; Chiapas.
1, (144 mm.) type of — Guatemala. O. Salvin, Esq.
the species.
2. (150 mm.) Guatemala. Q. Salvin, Esq.
3- 5. (112-124 mm.) Rio Geronimo. O. Salvin, Esq.
6. (114 mm.) Lake Yzabal. QO, Salvin, Esq.
7-12. (88-193 mm.) Rio Usamacincta. O. Salvin, Esq.
13-14, (168 and 260 mm.) Lake San Cristobal. F. C. Sarg, Esq.
29. Cichlosoma lentiginosum.
Heros lentiginosus, Steind. Denkschr. Ak. Wien, xxiii. 1864, p. 62,
pl. 1. fig. 1,
Cichlasoma lentiginosum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1624; Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 217
(1904).
Depth of body 23-24 in the length, length of head 3}.
Snout about as long as postorbital part of head. Diameter
of eye nearly 5 in the length of head, interorbital width 24.
Depth of przorbital 14 the diameter of eye. Maxillary not
nearly extending to below the eye ; Jaws eqial anteriorly ;
fold of the lower lip not continuous ; cheek with 6 series of
scales ; 10 gill-rakers on the lower part of the anterior arch.
Silas 34 = Dorsal XVII 13, Oe NG above the
extremity of operculum, the last spine 4 the length of head ;
soft fin, when laid back, extending a little beyond the base
of caudal. Anal VI 9. Pectoral # the length of head,
extending nearly to above the vent; ventral extending
nearly to origin of anal. Caudal slightly emarginate, with
rounded lobes. Caudal peduncle a little longer than deep.
Olivaceous, with 6 or 7 obscure darker cross- ‘bars and with
numerous small dark brown spots; vertical fins with dark
spots.
Mexico.
The typical example measures 215 mm. in total length.
Section 7.
Two species from Lake Nicaragua agree with those of the
preceding section in all essential characters, but have a
238 Mr. C. T: Regan on the Fishes of the
somewhat longer pectoral, which extends to above the origin
of the anal in one, a little beyond in the other. The teeth
of the outer series in the upper jaw are moderate, regularly
increasing in size anteriorly. In the lower jaw ‘the 5 or 6
anterior teeth on each side are distinctly enlarged and differ-
entiated from the smaller lateral teeth. Dorsal XVIII-XIX
10-11. Anal VII-VIII 7-9.
30. Cichlosoma balteatum.
Heros balteatus, Gill & Bransford, Proc. Ac. Philad. 1877, p. 184.
Cichlasoma balteatum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1521; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 220
(1904).
Depth of body 24 in the length, length of head 3+. Snout
a little longer than postorbital part of head. Diameter of
eye 3 in the length of head and equal to the interorbital
width. Depth of przorbital a little less than the diameter
of eye. Maxillary not extending to below the eye; jaws
equal anteriorly ; fold of the lower lip not continuous ;
cheek with 4 or 5 series of scales; 9 or 10 gull- ee on the
lower part of the anterior arch. Scales 34 = 24 between
- lateral line and base of anterior rays of soft dorsal. Dorsal
XVIII (10) 11, the spines subequal from the sixth to the
sixteenth, the last 4 the length of head. Anal VII (7) 8.
Pectoral nearly as long as the head, extending to above the
origin of anal; ventral extending beyond the origin of anal,
Caudal truncate or slightly emarginate. Caudal peduncle
nearly as long as deep. A blackish longitudinal band from
operculum to base of caudal; fins unspotted.
Lake Nicaragua.
1. (180 mm.) one of the types L. Nicaragua. Smithsonian Inst.
of the species.
31. Cichlosoma nicaraguense.
Heros nicaraguensis, Giinth, Proc. Zool. Soc, 1864, p. 153, and Trans.
Zool. Soe. vi. 1869, p. 465, pl. Ixxvii. fig. 1; Jord. & Everm. Bull,
U.S. Nat. Mus. xlvii. 1898, p. 1532.
Cichlasoma nicaraguense, Pellegr. Mém. Soe. Zool. France, xvi. 1903,
p. 221 (1904).
Depth of body 21-2? in the length, length of head 31.
Snout with nearly vertical profile, shorter than postorbital
part of head. Biatover a eye 34-32 in the length of head,
interorbital width 2}- Depth “of preorbital a little
greater than diameter « eye. Maxillary extending nearly
American Cichlid Genus Cichlosoma. 239
to below the anterior margin of eye ; jaws equal anteriorly or
the lower shorter than the upper ; fold of the lower lip not
continuous; cheek with 5 or 6 series of scales; 9 gill-rakers
on the lower part of the anterior arch. Scales 35 7%, 24
between lateral line and base of anterior rays of soft dorsal.
Dorsal X VITI-XIX 10-11, commencing above the opercular
cleft, the spines subequal from the eighth to the fifteenth or
sixteenth, thence increasing to the last, which is a little
more than 4 the length of head; soft fin, when laid back,
extending to anterior + of caudal. Anal VII-VIII 8-9.
Pectoral nearly as long as the head, extending to above the
second or third anal spine; ventral extending beyond origin
of anal. Caudal truncate or slightly emarginate. Caudal
peduncle nearly as long as deep. Upper half of body
brownish, with 5 or 6 dark cross-bars; vertical fins with
dark spots.
Lake Nicaragua.
1. (176 mm.) type of the Lake Nicaragua. Capt. J. M. Dow.
species.
2. (157 mm.) Lake Nicaragua. O. Salvin, Esq.
5. (166 mm.) Lake Nicaragua. Smithsonian Inst.
Section 8 (Astatheros).
Body rather deep (depth 1}—2} in the length). Upper
profile of snout straight, oblique. Scales of the lateral line
of the same size as those above and below it; scales of
thoracic region considerably smaller than those on the sides
of the body. Mouth moderate, moderately protractile ;
maxillary shghtly exposed; teeth of the outer series in both
jaws rather small (16-22 on each side in the upper jaw),
well-developed laterally and only slightly increasing in size
anteriorly. Dorsal XIV-XVI 10-15, the soft fin more or
less distinctly scaly at the base. Anal V—VII 8-9. Caudal
rounded, subtruncate, or slightly emargimate with rounded
lobis.
Six species from Southern Mexico, Central America, and
Colombia, which resemble C. b¢maculatum in their gene-
ralized dentition.
32. Cichlosoma Robertsoni, sp. nu.
Depth of body 2} in the length, length of head 23. Snout
as long as postorbital part of head. Diameter of eye 3}
in the length of head, interorbital width 34. Depth of
preorbital nearly } the diameter of eye. Maxillary not
210 Mr. C. T. Regan on the Fishes of the
extending to below the eye; preemaxillary processes extending
to above anterior margin of eye; jaws equal anteriorly or
the lower very slightly projecting; fold of the lower lip not
continuous ; cheek with 6 series of scales ; iT; gill-rakers on
the lower part of anterior arch. Scales 'a9 5 =, 24 between
lateral line and base of anterior rays of soft dorsal. Dorsal
XVI 11, commencing above the opercular cleft, the spies
subequal from the fourth to the sixth, which are the longest,
thence decreasing to the thirteenth and increasing again to
the last, which is nearly } the length of head; soft fin, when
laid back, extending to anterior } of caudal. Anal VI 9.
Pectoral nearly as long as the head, extending to above
last anal spine; ventral extending beyond origin of anal.
Caudal subtruncate. Caudal peduncle 2 as long as deep.
Olivaceous, with a blackish blotch on the side below the
lateral line and a dark spot on the upper 3 of the base of
caudal; some light blue spots on the cheek ; fins unspotted.
British Honduras.
1. (64 mm.) type of the Stann Creek. Rey. J. Robertson.
Species.
33. Cichlosoma longimanus.
Hleros ee, Giinth. Trans. Zool. Soe. vi. 1869, p. 453, pl. Ixxii.
fig. 2.
Cichlasoma longimanus, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1520; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 212
(1904). .
Depth of body 2} in the length, length of head 23. Snout
as long as postorbital part of head. Diameter of eye 3}
im the length of head and equal to the interorbital width.
Depth of przeorbital equal to diameter of eye. Maxillary
extending nearly to the vertical from anterior margin of eye;
preemaxillary processes extending to above anterior margin
of eye ; luwer jaw very slightly projecting; fold of the lower
lip not continuous ; cheek with 4 series of scales; 12 gill-
rakers on the lower part of anterior arch. Scales 28 2, 23
between lateral line and base of anterior rays of soft dorsal.
Dorsal X V1 10, commencing above the opercular cleft, the
fifth and sixth spines somewhat longer than the succeeding
ones and } the length of head ; the soft fin, when laid back,
extending to the middle of caudal. Anal VI 8. Pectoral
longer than the he ad, extending to the last ray of anal;
ventral extending bey ond origin of anal. Caudal slightly
emarginate, with rounded lobes. Caudal peduncle 2 as long
Americun Cichlid Genus Cichlosoma. 241
as deep. Olivaceous, with a large blackish blotch on the
side, connected to the orbit by an indistinct dark band ;
posterior part of dorsal with oblique stripes or series of
spots.
Lake of Nicaragua.
1, (183 mm.) type of the Lake of Nicaragua. O. Salvin, Esq.
species.
34. Cichlosoma macracanthus. |
Heros macracanthus, Giinth. Proc. Zool. Soc. 1864, p. 153, and Trans.
Zool. Soe. vi. 1869, p. 451.
Cichlasoma macracanthus, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1518; Pellegr. Mém. Soc. Zool. France, xvi. 1908, p. 207
(1904).
Depth of body 13 (adult) to 2 (young) in the length,
length of head 22-3. Snout shorter than postorbital part
of head. Diameter of eye 3-4) in the length of head, inter-
orbital width 24-3. Depth of preorbital 3-1) the diameter
of eye. Maxillary not extending to below the eye; pre-
maxillary processes not extending to above the eye; jaws
equal anteriorly ; fold of the lower lip continuous or not ;
cheek with 4 to 6 series of scales; 11 gill-rakers on the
lower part of anterior arch. Scales 29-31 — 3 or 4
between lateral line and base of anterior rays of soft dorsal.
Dorsal XIV-XV 12-14, commencing above the opercular
cleft, the spines increasing in length to the last, which is
4 (young) to # (adult) the length of head; soft fin, when
laid back, extending to anterior part (young) or posterior
edge (adult) of caudal. Anal V 9-10. Pectoral as long
as the head, extending to above third anal spine; ventral
extending nearly to origin of anal. Caudal subtruncate.
Caudal peduncle ?—2 as long as deep. Olivaceous, with
6 darker cross-bars, the third bearing a blackish blotch
below the lateral line; a blackish spot on the upper part of
the base of caudal ; fins dusky. Adult with nearly uniform
coloration.
Southern Mexico ; Guatemala.
1-7. (69-168 mm.) types of the Huamuchal. O, Salvin, Esq.
species.
8-11. (188-224 mm.) Chiapas. O. Salvin, Esq.
12-18, (53 and 76 mm.) Tequesixtlan. Dr. H. Gadow.
35. Cichlosoma heterodontus.
Heros (Cichlasoma) heterodontus, Vaill. & Pellegr. Bull. Mus. Paris,
1902, p. 86.
Cichlasoma heterodontus, Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p. 225 (1904); Meek, Zool. Pub. Columbian Mus. y. 1904, p. 215.
Cichlasoma Evermanni, Meek. t. ¢. p. 214, fig. 70.
reg On the American Cichlid Genus Cichlosoma.
Depth of body about 2 in the length, length of head
27-24. Snout shorter than postorbital part of head.
Diameter of eye 33-4 in the length of head. Maxillary not
extending to below the eye; jaws equal anteriorly; fold of
the lower lip continuous or not; cheek with 5 series of
scales ; 11] gill-rakers on the lower part of anterior arch.
Scales about 30 *2. Dorsal XIV-XV 12-15, the spines sub-
equal from the fifth, the last 4 the length of head; soft fin,
when laid back, extending beyond the middle of caudal, in
the adult. Anal V 8-9. Pectoral 3-4 the length of head,
extending nearly to above origin of anal; ventral extending
to origin of anal or beyond. Caudal subtruncate. Caudal
peduncle 2—} as long as deep. Olivaceous, with 6 darker
cross-bars ; a blackish spot on the upper part of the base of
caudal ; fins dusky, the vertical ones sometimes with small
dark spots.
Tehuantepec, Southern Mexico.
The specimens described by Meek measure up to 200 mm.
in total length.
As in the closely allied C. macracanthus, and in some
other species of the genus, in adult examples many of the
teeth of the outer series may be worn and end ima flat
circular surface, but this feature is not even of specific,
much less of subgeneric, value.
36. Cichlosoma altifrons.
Heros altifrons, Kner & Steind. Sitzb. Ak. Bayern, 1863, p. 223, and
Abhandl. Ak. Bayern, x. 1866, p. 11, pl. i. fig. 1; Guuth. Trans.
Zool. Soc. vi. 1869, p. 459; Jord. & Everm. Bull. U.S. Nat. Mus.
xlvii. 1898, p. 1538; Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p- 228 (1904).
Depth of body about 24 in the length, length of head
nearly 8. Snout longer than postorbital part of head.
Diameter of eye 4-5 in the length of head and 1} in the
interorbital width. Depth of preorbital greater than
diameter of eye. Maxillary not extending to below the eye ;
jaws equal anteriorly; cheek with 4 or 5 series of scales ;
12 gill-rakers on the lower part of anterior arch. Scales
about 30 = 2} between Jateral line and base of anterior rays
of soft dorsal. Dorsal XVI 11, the spines increasing in
length to the last, which is nearly } the length of head ;
soft fin, when laid back, extending to anterior third of
caudal, Anal V 8-9. Pectoral } the length of head, ex-
tending nearly to above origin of anal; ventral extending a
little beyond origin of anal. Caudal slightly rounded. Caudal
Bats of the Rhinolophus philippinensis Group. 243
peduncle # as long as deep. Olivaceous with 5 or 6 dark
brown cross-bars; light blue spots on the body and soft
vertical fins.
The types from Colombia measure from 115 to 200 mm. in
total length.
37. Cichlosoma rostratum.
Heros rostratus, Gill & Bransford, Proc. Ac. Philad. 1877, p. 481.
Cichlasoma rostratum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1529.
Depth of body 24 in the length, length of head 23.
Snout longer than postorbital part of head. Diameter
of eye 4 in the length of head, interorbital width 32.
Depth of preorbital 1} the diameter of eye. Maxillary
extending to midway between nostril and eye; pre-
maxillary processes not extending to above the eye; jaws
equal anteriorly; fold of the lower lip not continuous ;
cheek with 5 or 6 series of scales; 12 or 13 gill-rakers on
the lower part of the anterior arch. Scales 33 = 33 between
lateral line and base of anterior rays of soft dorsal. Dorsal
XVI 11, commencing above the opercular cleft, the spines
subequal from the sixth to the fourteenth, the last 2 the
length of head. Anal VII 8. Pectoral as long as the head,
extending to above the last anal spine; ventral extending
beyond the origin of anal. Caudal subtruncate. Caudal
peduncle # as long as deep. A blackish blotch below the
fourteenth to the seventeenth scales of the lateral line ; a dark
spot at the base of caudal ; soft dorsal, anal, and caudal with
alternate ight and dark bars or series of spots.
Lake Nicaragua.
1. (186 mm.) one of the types L. Nicaragua, Smithsonian Inst.
of the species.
This species is very closely allied to C. altifrons.
[Te be continued. ]
XXVIII.—On the Bats of the Rhinolophus philippinensis
Group, with Descriptions of Five new Species. By Kynup
ANDERSEN.
Tue conclusions recorded in the present paper are based
on the material in the British Museum, together with some
specimens from the United States National Museum. The
latter were sent for inspection and identification by
Mr. Gerrit 8. Miller, Jr., Washington.
244 Mr. K. Andersen on the Bats
General Characters of the Kh. philippinensis Group.
Diagnosis.—Median anterior nasal swellings large. Palatal
bridge long, on an average more than, and generally con-
siderably more than, 4 the length of the maxillar tooth-row ;
median anterior point opposite the front (or anterior half)
of p*. Posterior connecting-process extremely low and
rounded off, starting from a point considerably below the
summit of the sella. Base of central nose-leaf forming
cup-like or wing-like expansions.
Remarks.—So far as my material goes, the above diagnosis
holds good for all members of the group, the most primitive
(philippinensis, achilles), as well as the highest developed
(trifoliatus, luctus, and all their allies). In these latter the
the skull is rather modified, and some of the modifi-
cations are obviously correlated to the enormous enlar gement
of the nose-leaves and ears. The postnasal depression is
deeply hollowed out, more than in other Rhinolophi. In
front of the median anterior swellings is formed a narrow,
thin brim of bone, the object of which is evidently to increase
the surface of the facial part of the skull (which supports
the large nose-leaves) ; in Rhinolophi with nose-leaves of
normal size there is no such brim of bone, the median
anterior swellings partaking immediately in the formation of
the margin of the nasal openings. The interorbital con-
striction is narrower than in other Rhkinolophi. The cochleze
are, proportionately, slightly larger, the basioccipital, between
them, consequently slightly narrower; but neither are the
cochlez as large nor the basioccipital as narrow (linear)
as in Rh. hipposiderus and its allies.
In most, if not all, species the position of the lower p;
is individually variable (as in other primitive Rhinolophi) :
in the tooth-row, or more or less external to the row. The
upper p* is invariably in the row.
Subdivision of the Group.
The group may conveniently be divided into three
sectious :—the philippinensis section, the sedulus section,
and the ftrifoliatus section. The first of these sections is
represented by Rh. philippinensis and achilles * ; the second
by Rh. sedulus and lanosus; the third by Rh. tr ifoliatus,
solitarius, luctus, perniger, geminus, and Beddomei. They
differ as follows :—
* On Rh. mitratus and Maclaudi, see the “General Remarks” on
954
p. 254.
of the Rhinolophus philippinensis Group. 245
(1) In the philippinensis section the sagittal crest is low
in front, gently sloping towards the postnasal depression
(fig. 1). In the ¢rifoliatus section the crest is high in
front, and abruptly descending towards the postnasal
depression (fig. 2): this causes a very different side view
of the skull.
Fig. 1a. Fig. 2a.
Fig. 1. Skull of Rh. philippinensis, side view. la. The same skull,
upper view. 2. Skull of Rh. trifoliatus, side view. 2a. The same
skull, upper view. The specimens figured are of the same age (teeth
unworn). 2.
In the philippinensis section the teeth are smaller, the
maxillar width narrower, the coronoid process of the
mandible somewhat lower, the temporal fossa narrower, the
zygomatic arches, therefore, very slightly projecting laterally,
making the zygomatic width of the skull but very slightly,
or not at all, larger than the mastoid width (fig. 1a). In the
trifoliatus section the teeth are larger, the maxillar width
broader ; the stronger teeth involve an increase in size of the
temporal muscle, this again a slight heightening of the
coronoid process, a considerable widening out of the temporal
246 Mr. K. Andersen on the Bats
fossa, a projecting of the hinder part of the zygomatic arches,
making the zygomatic width of the skull considerably larger
than the mastoid width (fig. 2 a).
In the philippinensis section the base of the sella forms,
together with the internasal lobes, a cup-like expansion. In
the ¢rifoliatus section the internasal lobes are quite, or
almost quite, normal, the lateral expansions confined to the
sella and wing-like, giving the sella some resemblance to
a Trifolium leaf (hence the name “‘ ¢rifoliatus”’) or a Maltese
cross.
In the philippinensis section the wing-structure * is very
primitive, quite as in other primitive species of the genus
(megaphyllus, borneensis, minor, and many others): the
fourth metacarpal slightly longer than the fifth and the third ;
IV.2+ and V.? but slightly longer than IV." and V.*. In
the trifoliatus section the wing-structure is considerably
modified :—the third metacarpal is much shortened, the foarth
slightly lengthened, the fifth more so, making as a final
result the fi/th metacarpal decidedly the longest of all, and
very much longer than the third (supposing the length of
the forearm, in all species of the group, to be exactly
1000 mm., the fifth metacarpal, in the ¢rzfoliatus section, is
no less than 133 mm. longer than the third, in philippinensis
and achilles 18 mm. only) t. At the same time the first
phalanx of the third finger, in the ¢rifoliatus section, is much
lengthened, chiefly by a removing, in proximal direction, of
the joint between the metacarpal and the first phalanx ; in
other words, the piece by which the third metacarpal has
been shortened has been added to the length of the first
phalanx of the same finger. Thirdly, III.”, IV.’, and V.?,
in the ¢trifoliatus section, are considerably lengthened.
The mechanical reason for this modification of the wing
is, probably, the following :—A lengthening of the distal
phalanges gives a broader wing, consequently an increased
power of flight; the third finger, as being the longest of all
and the nearest to the front margin of the wing (when
expanded), has to sustain the heaviest pressure of the air ;
* The remarks on the wing-structure to be compared, point for point,
with the diagram on p. 247 and the wing-indices on p. 257.
+ For brevity’s sake I call the distal phalanges of the 3rd, 4th, and 5th
fincers III.?, IV.?, and V.?, the proximal phalanges of the same fingers
LY PV.) and V,".
{ An almost exact parallel to this modification of the wing-structure
is seen in Rh. ferrum-equinum compared with the primitive forms of
the group to which that species belongs (f. i. 2h. megaphyllus, borneensis,
&c.).
of the Rhinolophus philippinensis Group. 247
a considerable increase in the length of III.? involves, there-
fore, an increase in the length of the phalanx which supports
III.?, viz. II1.’, and this increase is effected not by a
lengthening of III.‘ in distal direction (which would make
an unproportionately long third finger), but by a lengthening
ao
os
a. Forearm. d3,d4,d5. Third, fourth, and fifth fingers in the philip-
pinensis section. 93,04, 95. The same fingers in the ¢rifoliatus
section, reduced to the same length in proportion to the forearm.
Subdivisions of d and 9, in direction from left to right, indicate the
metacarpal, first and second phalanx.
in proximal direction, 7. e. by a removing backwards of the
joint between the third metacarpal and III.’, which again
makes a shorter third metacarpal.
(2) The sedulus section has the skull and the teeth
essentially as in philippinensis, but the nose-leaves and the
wing:structure as in /rifoliatus.
Only the species of the sedulus and trifoliatus sections
will be described below.
1. Rhinolophus sedulus, sp. n.
Diagnosis.—Cranial characters much as in Rh. philippin-
ensis, but postnasal depression considerably deeper. Nose-
leaves and wing-structure: the ¢rifoliatus type. Forearm
43°5-49°2 mm.
SkullSagittal crest and maxillar width very much as in
philippinensis ; temporal fossa but slightly larger ; postnasal
depression deeply hollowed out as in ¢rifoliatus.
Dentition (two skulls).—p; external; p. and p, almost or
quite in contact ; p? in row, cusp so minute as to be scarcely
observable.
Colour.— ¢ ad., Sarawak; distal epiphyses of metacarpals
ossified ; teeth unworn; skin: General impression a shade
of dark brown, slightly varied with greyish tips to the hairs,
215 Mr. K. Andersen on the Bats
Analysis of colour: the tips of the hairs of the upperside
have a grey iridescence; when keeping the skin so that the
reflections disappear, the true colour proves to be a deep
brown shade of ‘‘ drab”; base of hairs scarcely lighter.
Underside of the same general colour as the upperside, but
without, or almost without, grey reflection. Interfemoral
between the tip of the tail and the calcar, narrowly bordered
with yellow.
Measurements *.—On p. 257.
Type.— ¢ ad. (skin). Sarawak. Collected by Dr. A. R.
Wallace. ‘Tomes Collection (no. 19).
Distribution.—Sarawak (Brit. Mus.) ; Pahang (Un. St.
Nat. Mus.).
Remarks.—Externally this species is exceedingly like a
very small Rh. trifoliatus, but the tibia is shorter. In the
shape of the skull it is very much nearer to Rh. philippin-
ensis and achilles.
2. Rhinolophus lanosus, sp. u.
Diagnosis.—Similar to Rh. sedulus, but very much larger.
Forearm 71°5 mm.
Skull—In the shape of the sagittal crest, the small
temporal fossa, the narrow maxillar width, and the small
teeth, this very large species bears much resemblance to
Rh. philippinensis. But the postnasal depression is deeply
hollowed out as in the sedulus and trifoliatus type (corre-
sponding to the enormously developed nose-leaves).
Dentition (one skull).—p; in row; p, and p, well separated;
p’ in row, cusp extremely small.
Colour.— @ ad.; teeth slightly worn; skin: as in
Rh. sedulus.
Type.— ad. (skin). Kuatun+, N.W. Fokien, China ;
April 4th, 1898. Presented by J.D. La Touche, Esq. Brit.
Mus. no; 98. 11. I. 1.
Remarks.—As proved by the skull, this fine species is an
* Only the following measurements require some explanation :—Zars,
length from base of inner margin to tip. Second phalanx, exclusive of
the cartilaginous terminal rod. Skull, total length, inion to front of
canines. Width of brain-case, above root of zygomata. Mazxillar width,
across antero-external corner of m°. Supraorbital length, from posterior
point of postnasal depression to median anterior point of nasals. Man-
dible, condylus to front of incisors. Upper and lower teeth, exclusive of
incisors.
+ An excellent characteristic of this Chinese village and its sur-
roundings was contributed by Mr. La Touche to the Proc. Zool. Soe. for
1898 (pp. 769-70).
of the Rhinolophus philippinensis Group. 249
offshoot of the sedulus-ty pe, differing in a'l important cranial
characters from the species of the ¢ri/oliatus section. But
externally it is exceedingly like a small Rh. perniger.
3. Rhinolophus trifoliatus, Temm.
Diagnosis.—Cranial and external characters: the ¢rifoliatus-
type (above pp. 245-47). Forearm 47-55 mm.
Skull.—As described and figured above (p. 245).
Dentition (eleven skulls).—p, “vacillating”’ in position :
in row (one skull), or almost in row (one), or half in row
(two), or almost external (one), or quite external (six).
Corresponding to this, p, and p, well separated (five), or
almost in contact (one), or quite in contact (five). p” always
in row. Upper canine and p* well separated. In one
example there is a marked interspace between p* and p* (the
former place of p®, lost in all recent species of the genus).
Colour —Five skins ; ad.; teeth unworn or slightly worn :
Very light-coloured. Upperside, a ‘ wood-brown ” shade
of “ drab,’ somewhat darker on the hinder back; base of
hairs more distinctly light “drab.” Underside ‘ drab,”
somewhat varying in the intensity of the colour, and washed
with “ ecru-drab ” on the throat and breast. Interfemoral,
between the tip of the tail and the calcar, narrowly
bordered with yellow.
Size—The great variation in the size of this species
(forearm 47-55 mm.) is independent of the geographical
habitat and of the sex:—(1) The shape of the skull is
identically the same in al] the individuals examined ; (2) the
size of the skull is much less subject to variation than the
external dimensions (cf. the measurements) ; (3) the smallest
and the largest individuals examined are both from
N. Borneo; (4) the smallest individual in the series (fore-
arm 47 mm., teeth slightly worn) and one of the very
largest (54°3 mm., teeth unworn) are, both of them, males.
All the specimens are full-grown (distal epiphyses of meta-
carpals ankylosed).
Distribution—N. Borneo (Paitan, Kina Balu, Mt. Dulit,
Mt. Mulu, Sarawak); Singapore; Lower Siam (Trong) ;
Tenasserim (Mergui). ? Java (cf. below).
Technical name.—The type specimen of Rh. trifoliatus, as
described by Temminck*, is stated to be from Bantam,
W. Java. I have seen no specimen of this Bat from Java.
It is rather easy indeed to point out some discrepancies
between Temminck’s figure (natural size) and the series
* Temminck, Mon, Mamm. ii. 8° monogr. (1835) pp. 27-28, pl. xxxi.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 17
250 Mr. K. Andersen on the Bats
here under consideration :—The horseshoe is very narrow
(9 mm., as against 10°5-12°4 in the series examined by me);
the tail extremely short (about 21-23 mm., as against
29-36) ; also in the letterpress the tail is stated to be only
“de la longueur du tibia,” whereas in all the individuals I
have seen it is from } to 4 longer than the tibia. On the
other hand, the length of the forearm of the type (“ 1 pouce
10 lignes” = about 50 mm.) and of the metacarpals, pha-
langes, and tibia (on the figure) are quite as in several
Bornean specimens. Also it should be remembered that the
type is a mounted specimen and, according to Jentink *, only
“4-peu-prés adulte”’ ; the differences pointed out above are
of such kind that they may perhaps be due to shrinkage.
At allevents, in the absence of more conclusive evidence, I
think it advisable to retain the name ¢rifoliatus for the
species described above. If not identically the same as
the Bornean and Malaccan bat, the Java form is probably
extremely closely related fT.
4. Rhinolophus solitarius, sp. n.
Diagnosis —Allied to Rh. trifoliatus, but slightly smaller,
with very short tail and tibia. Forearm 465 mm.
Skull.— Smaller than in ¢trifoliatus, but proportionately
broader (compare mazillary, zygomatic, mastoid breadth,
and breadth of brain-case with the measurements of the
trifoliatus skulls, p. 257).
Dentition (one skull).—p; completely wanting, without any
trace of the alveoli (teeth but very slightly worn) ; p, and p,
in contact. This is the only imstance, within the present
group, of a complete obliteration of the middle lower premolar;
the aberration from the general rule ¢ is rather surprising,
and I must leave it open to question if it is, really, a con-
stant (or almost constant) peculiarity of the present species.
* Jentink, “ Cat. syst. Mamm.,’”’ Mus. d’Hist. nat. Pays-Bas, xii. (1888)
p. 160.
+ The following statements in the original description of Rh. trifoliatus
are, evidently, accidental errors only :—‘“ Deux grosses verrues triangu-
laires 4 la machoire supérieure” ; must be “ inférieure.” “ Dans l’adulte
point dincisives supérieures ” ; they must have been overlooked. “ Point
de petite dent anomale eutre la canine et la premiére molaire de la machoire
supérieure ”; it is extremely improbable that p*, which is present in all
species of this group and always situated in the tooth-row, should be
wanting in the type of tr¢foliatus.
{ As ageneral rule, in the genus, the lower p, does not completely dis-
appear until the dentition has arrived at the stage that the upper p? is
external, and even then it is still very often present. In all species of the
present group p* is quite in the tooth row.
of the Rhimolophus philippinensis Group. 251
Also the upper p” is slightly smaller than in ¢rifoliatus, and
consequently the interspace between the canine and p* a
little narrower.
External characters.—In all important respects as Rh. tri-
foliatus, but a trifle smaller than the smallest individuals I
have seen of that species ; tail and tibia extremely short.
Colour (of a spirit-specimen) much as in ¢rifoliatus.
Type.— $ ad. (in alcohol). Tanjong Pamuja, Banka;
June 18th, 1904. Collected by Dr. W. L. Abbott. Un. St.
Nat. Mus. no. 124767.
5. Rhinolophus luctus, Temm.
Diagnosis.—Trifoliatus type. Forearm 65°3-68 mm.
Skull.—As in trifoliatus, but very much larger.
Dentition (five skulls).—p; generally external (four skulls),
sometimes almost in row (one). jp, and py, separated (two),
or almost or quite in contact (three). p? always in row,
cusp rudimentary or altogether wanting. Interspace be-
tween the upper canine and p* rather narrow.
Colour.—(1) 2, rather young, but quite full-grown ; distal
epiphyses of metacarpals ossified; teeth almost unworn;
Selangor, November; skin: Above and below, an exceedingly
dark shade of “ drab,” approaching blackish ; grey iridescence
on the hairs of the upperside.
(2) Aged individuals; teeth slightly worn or well worn;
N. Borneo; four skins: Same style of colour, but decidedly
browner. Details as in Rh. sedulus.
Distribution —N. Borneo (Mt. Dulit, Baram, Lawas) ;
Singapore; Selangor (Semangton). ? Java (cf. below).
Temminck’s Rh. luctus *.—The type specimen + of RA.
luctus is stated to be from Tapos, Java. I have seen no
example of this species from Java, but the original description
of luctus, the measurements (forearm and tail only), and the
figure (natural size) agree very closely ¢ with the species
* Temminck, Mon. Mamm. ii. 8° monogr. (1835) pp. 24-26, pl. xxx.
+ According to Dr. Jentink (Cat. syst. Mamm.,” Mus. d’Hist. nat. Pays-
Bas, xii. (1888) p. 160) there are two type specimens of Rh. luctus in the
Leiden Museum. Technically only one of these examples is the “ type,”
viz. no. “b” (the adult female) ; cf. Temminck (/. c.): “ mesure et de-
scription d'une adulte femelle ” (p. 25); “nos voyageurs n’en trouvérent
quwune femelle” (p. 26). The specimens “a” and “e¢” in Jentink’s
Catalogue are, perhaps, those referred to by Temminck ina later appendix
to the description (op. cit. pp. 30, ¢, d).
{ The measurements given by Temminck are: forearm 63 mm. (“2
pouces 4 lignes”), tail 45 mm. (“1 pounce 8 lignes”). Dr. Jentink has
kindly re-examined the type, and found the length of the forearm as
stated by Temminck,
1 is
De Mr. K. Anlersen on the Bats
here under consideration. If by further examination Java
specimens should prove to differ from the Borneo—Malacca
form, the former will have to stand as RA. luctus, the latter
as Rh. morio, Gray.
Gray’s Rh. morio*.—The type (adult, Singapore ; sent
to Gray in alcohol, subsequently mounted) is in the British
Museum. The light colour, approaching that of RA. tri-
foliatus (and which caused Gray to give it a new name), is
probably due to fading in alcohol; the colour of all the
species of the present group seems to be unusually hable to
fading ; in every other respect Rh. morio is indistinguishable
from Selangor and Borneo specimens.
Alleged occurrence on the Philippine Islands. — Eydoux
and Gervais’s ‘‘ Rh. luctus, varietas rufa,’ from Manila, is
characterized by the authors as follows :—“ Dans la variété
que nous décrivons, quoique la feuille soit parfaitement la
méme [as in Temminck’s Rh /uctus|, le pelage est générale-
ment roussatre” t+. No further desc ription, no measurements,
no figure. By subsequent writers {, noue of whom seem to
have seen the specimen, the name proposed by Eydoux and
Gervais has been referred as a synonym to “ Rh. luctus” 1
very much doubt that this is correct. Neither before 1839
nor since has any bat of the ductus type been recorded from
any of the Philippine Islands. Considering the time at
which Eydoux and Gervais wrote, I should find it very
hazardous to lay much stress on the statement that the nose-
leaves were “ parfaitement ” as in ductus. Of much more
importance is, I think, the fact that in 1839 Rh. philippin-
ensis was not yet described. Perhaps “Rh. luctus, var. rufa”
is nothing but Rh. philippinensis; and, if so, the name
“rufus”? (1839) unfortunately antedates ‘ philippinensis”
(1843). An examination of the type (presumably in the
Paris Museum) would decide the matter.
6. Rhinolophus perniger, Hodgs
Diagnosis.— Allied to RA. luctus, but much larger. Fore-
arm 715-78 mm.
Skull. Quite of the ductus shape, but considerably larger
and with markedly broader nasal swellings.
* J, E. Gray, Ann. & Mag. Nat. Hist. (1) x. (Dec. 1842) p. 257,
+ Eydoux and Gervais, ‘ Voyage autour du monde ...la Favorite,’
vy. pt. 1. (1839) p. 9.
t In his “ Kritische Durchsicht der Ordnung der Flatterthiere” (SB.
Akad. Wien, Bd. lxi. Abth. i., Feb. 1870, p. 194) Dr. Fitzinger, without
having seen the example here under consideration, g gives it a new name—
Aquias FEudouxii—on account of “ die durchaus verschiedene Heimath,
so wie auch der grosse Unterschied in der Farbung.”
cf the Rhinolophus philippinensis Group. 253
Dentition (five skulls).—p, generally external (four skulls),
sometimes almost in row (one) ; pz and p, separated (three,
or almost in contact (two) ; p* in row, cusp rudimentary and
soon worn down. Interspace between the upper canine and
p* generally rather narrow.
External characters.—Considerably larger than Rh. luctus,
with, also proportionately, much longer ears and tibia. It is
in every respect the extreme of the “ ¢rifoliatus type” and
the largest species in the genus. Colour as in RA. luctus.
Distribution.—The Himalayas: Sikkim, Nepal, Masuri.
Technical name.—TVhe types of Hodgson’s Rh. perniyer *
(three adult specimens, Nepal) are in the British Museum.
Remarks.—This species has hitherto wrongly been cou-
founded with Rh. luctus.
7. Rhinolophus geminus, sp. un.
Diagnosis.—Similar to Rh. perniger, but with shorter ears
and much shorter tail. Forearm 73 mm.
Skull. —Much as in perniger, but with markedly narrower
nasal swellings.
Dentition (one skull).—p; almost in row; pz and p, well
separated ; p” in row, cusp extremely small. Interspace
between the upper canine and p* rather wide.
Colour.—The luctus style.
Type.— ? , slightly immature (in alcohol). Kediri, E.Java,
2000-3000 feet, “between March and May, 1878. Presented
by Baron v. Hiigel. Brit. Mus. no. 79. 11. 15. 10.
Remarks.—As might be expected, from the general
character of the mammalian fauna of Java, this species is
much more closely related to the Himalayan form (perniger)
than to Rh. luctus from Borneo and the Malay Peninsula.
8. Rhinolophus Beddomei, sp. n.
Diagnosis.—Allied to Rh. luctus, but rather smaller, with
excessively short metacarpals and tail. Forearm 63°8 mm.
Skull.—Ot the luctus type, but extremely small.
Dentition (one skull).—p; external; p. and p, almost in
contact; p* im row, cusp almost. imperceptible (teeth
unworn) ;
Euternal characters.—This species is almost of the same
size as Kh. luctus, but the metacarpals and tail are much
shorter.
Type.— g ad. (in aleohol). Wynaad, Mysore, S. India.
Collected and presented by Colonel Beddome. Brit. Mus.
NO. Go, Us on L.
* Hodgson, J. A. 8. B. xii. pt. 1, no. 187 (May 1843), p. 414,
254 Mr. K. Andersen on the Bats
General Remarks on the Rh. philippinensis Group *.
In the Philippine Islands and the Austro-Malayan Sub-
region we find preserved the most primitive members of the
whole group.
In all the more important points, cranial and external,
Rh. philippinensis (uuzon) and achilles + (Key Islands) are
either much like or quite on the same level as other primitive
Rhinolophi:—In the general shape of the skull; in the
narrow temporal fossa and low sagittal crest; in the long
palatal bridge; in the dentition—p* less reduced in size,
its cusp well developed, the interspace between the canine
and p* very wide; in the wing-structure—the ratio be-
tween the metacarpals, as well as the proportional length
of the distal phalanges, practically quite as in Rh. megaphyllus
(and its closer allies) or Rh. lepidus (and its modifications) ;
in the number of mental grooves—three, as in all primitive
species of the genus; in the general shape of the ears—very
blunt, emargination of outer margin below the tip very
shallow, the ear therefore not attenuated below the tip. But
the nose-leaves are peculiarly modified { and the ears
unusually large.
This type of bat has found its way to N. India: Rh. mi-
tratus (Chaibassa, on the border of Orissa and Bengal).
The same cup-like expansion of the central nose-leaf, formed
by the base of the sella and the internasal lobes; the same
shape and proportionate size of the ears. But the wing-
structure seems to be more advanced $ and the lateral
mental grooves are obliterated. The skull and dentition are
unknown.
We find the philippinensis-type so far away as Conakry
Island, off the coast of Senegambia: Rh. Maclaudi ||. Iden-
tically the same cup-like (not wing-like) expansion of the
central nose-leaf ; the same shape of the upper (not expanded)
part of the sella—very broad, parallel-margined, the summit
* To be compared with the diagram on p. 247.
+ Thomas, Ann. & Mag. Nat. Hist. (7) v. (1900) p. 145.
¢ It is worth noticing that the connecting-process in Rh. philippinensis
and achilles, although quite of the same style as in the other members of
the group, is slightly less reduced in size, consequently nearer to the
“normal” type.
§ Detailed measurements of the wing of Rh. mitratus are unknown.
According to Blyth (J. A. S. B. xii. (1844) p. 483) the length of the
forearm is 57 mm., the length of the third finger, as a whole, 84°56 mm. ;
this is sufficient to convince me that there must be a considerable
lengthening of III.’.
|| E. de Pousargues, Bull. Mus. d’Hist. nat. 1897, no. 8, pp. 858-381
(Feb, 1898).
of the Rhinolophus philippinensis Group. 255
broadly rounded off, almost truncated ; the same connect ng-
process; the same lancet — peculiarly long, narrow, cuneate ;
the same size and shape of the horseshoe; the enormously
enlarged ears—proportionately of the same size as in achilles
aud essentially of the same shape; the same ratio between
the metacarpals (third metacarpal not shortened). But the
distal phalanges are much lengthened *; the lateral mental
grooves are obliterated (asin Rh. mitratus) ; and the general
size cf the animal is much increased. The dentition of
Rh Maclaudi is more highly developed than in philippinensis
and achilles, and on the same level as in the sedulus and tri-
foliatus section—p* in the tooth-row, very small, the inter-
space between the canine and p* narrow+. The skull is
unknown.—In short: Ri. Maclaudi is an Ethiopian offshoot
of the philippinensis-ty pe, more highly developed than that
species at least in the dentition, the wing-structure, aud the
mental grooves.
The bats of the sedulus section have retained, in all
important respects, the cranial characters of philippinensis
and achilles. But the postnasal depression is deeply hollowed
out (as in the ¢trifoliatus type), and externally, 7. e. in the
nose-leaves (wing-like expansions of the sella), in the wing-
structure, in the mental grooves (one only), and in the more
pomted ears, they are lke ¢rifoliatus and luctus. Two
species only are known—kth, sedulus trom N. Borneo and
the Malay Peninsula, and RA. lanosus from 8. China.
The series of evolution culminates in the ¢rifoliacus section :
also the skull is modified. ‘The six species known are
distributed from N. Borneo to 8. India, from Banka aud
Java to the Himalayas.
* In the trifoliatus section the lengthening of III.* (as pointed out
above) has involved a lengthening of IL.’ in proaimal direction, and
consequently a shortening of the metacarpal. In hk. Maclaudi the
lengthening of III.” has involved a lengthening of ILI.’ in distal direc-
tion; consequently the metacarpal retains its original length. The great
increase in the total length of the third finger resulting herefrom seems
to have been counterbalanced by a lengthening aso of the first phalanx
of the other (fourth and fifth) fingers. ‘These remarks are based on
Pousargues’s measurements of the third and fifth fingers (those of the
fourth tinger are unknown).
+ I am indebted to M. A. Ménégaux, Paris Museum, for the above
details on the dentition of Rh. Maclaud?, as well as for the following
information :—“ M, de Pousargues s’est trompé en disant qu'il y a un
espace entre la canine supérieure et la p? | p* in this paper]; a leur base
la premiére prémolaire [ p*| touche en avant la canine, en arriére la p?
[p*].” By this important correction the dentition of Rh. Maclaudi is
siown to be perfectly in accordance with the usual scheme in the
present group,
256 Bats of the Rhinolophus philippinensis Group.
One of these modifications of the skull, in the ¢rifoliatus
section, is very instructive. When, as is the case in other
Rhinolophi beyond a certain low level of development, the
temporal muscle increases in size, the sagittal crest creeps
more forwards (as in many other Mammalia), reducing the
length of the supraorbital crests, consequently also the
length of the postnasal depression, which is bordered laterally
by the supraorbital crests. In the trifoliatus-type the shape
and size of the nose-leaves seem to require a rather long and
very deep postnasal depression; therefore the temporal
muscle, when growing larger, cannot very well push the
sagittal crest much farther forwards ; stead of that it pushes
the crest upwards, making it unusually high in front. This
accounts for the peculiar shape of the sagittal crest, as
figured on p. 245 (fig. 2).
The subjoined diagram gives a view of the probable
inter-relations of the species :—
perniger.
geminus.
ae
4
luctus.
\| Beddomei.
lanosus. a
solitarius.
ie
Maclaudi.
7 PUBS ee /
|
sedulus.
achilles.
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258 Bibliographical Notices.
BIBLIOGRAPHICAL NOTICES.
Preliminary Report on the Geology and Water Resources of Nebraska,
west of the One-hundred-and-third Meridian. By N. H. Darton.
71 pages, with 43 plates and 23 text-figures. 4to, Government
Printing Office, Washington. 1903.
Tuts is one (No. 17”) of the valuable “ Professional Papers ”
issued by the U.S. Geological Survey, and designed for the supply
of information on the geologica] structure of the country, and the
prospect of obtaining underground water by boring, and surface
water in large quantity by dams and irrigation.
The subject-matters in this memoir are treated in the following
order :—
Topography, pp. 9-14.
Geology, pp. 14-43. The formations taken separately.
1. General Geology of Nebraska, pp. 14-21.
2. Geology of the region west of the 105rd meridian, pp. 21-42.
Underground water, pp. 43-47.
1. General conditions.
2. Water horizons.
3. Deep-seated waters.
Springs, p. 48; Streams, pp. 50-54 ; Irrigation, pp. 54-62.
Climate, p. 62; Timber, p. 62; List of Elevations, pp. 62-°6; Index,
p. 67.
The limits of the district concerned are first of all defined. It
is the north-western part of Nebraska; bounded on the north by
South Dakota, by Wyoming on the west, and by Colorado on the
south. The region is a characteristic part of the Great Plains,
which extend from the Rocky Mountaias eastward. It may be
referred to as having originally sloped gently to the east, and, when
uplifted in early pleistocene times, the rivers excavated valleys
across it, making the chief existing features, portions of the plain
remaining as high tablelands between the valleys.
( Alluvium.
| Sand-hills.
Puuistocens{ Loess.
| Drift.
| Equus-beds.
Ogalalla formation (Pliocene). )
Nuocene j Arikaree formation (Miocene). | = Loup Fork beds.
Gering formation (Miocene).
Eocrne Brule Clay. = White River group. Titanotherium
(Oligocene ?) | Chadron formation. robustum, pl. Xxxv.
( Laramie formation.
| Pierre Clay.
Cretaceous { Niobrara formation. | =Montana group. Inoceramuslabiatus,
| Benton Shales. | Schloth. aud Ostrea congesta Conrad,
| Dakota Sandstone. [pl. xxxvi.
{ Permian limestone.
CARBONIFEROUS + Cottonwood limestone.
Wabaunsee limestone.
Bibliographical Notices. 259
The relative positions, range, and thickness of these formations
are well illustrated by maps and sections: the geography and
geology of the region being shown in plates i. & ii. and ix, to x1,
Xxxvil. & xxxvui.; physical features being richly represented
by numerous figures in the text and in plates.
The Alluvium is chiefly noticeable in the valleys, particularly in
that of the Platte and its tributaries. The ‘‘ Great Sand-hill
district,” in Central Nebaska (of 24,000 square miles), is composed
of wind-borne sand largely derived from the Arikaree formation.
Wide plains of Loess occupy nearly one third of Nebraska, and
consist largely of Boulder-clay. The Ogalalla beds consist of
calcareous grit and pebbles; and they have formed gravel when
disintegrated. The Arikaree formation (about 430 feet) of sand
and concretions, with conglomerate at base, contains also volcanic
ash, and yields abundance of the curious Demondice or * Devil’s
Corkscrews.” The concretions are frequently tubular, and there
are channels in the sand filled with conglomerate. This formation
weathers into the frequent upstanding and isolated pinnacles, stacks,
chimneys, as well as butts and bluffs, forming remarkable and
picturesque features in the region, as shown in many of the plates
and figures. ‘The Gering Sandstone varies considerably in thickness
aud in colour and hardness, and has conglomerate at the base.
The hard pinkish Brule Clay is about 600 feet thick, thinning
eastwards. ‘Then greenish sandy clay (Chadron formation) follows
below, and is said to have yielded remains of 7%tanotherium.
Equivalents of the Laramie formation, extending for some way
under South-western Nebraska, are sandstone and clay. The
Pierre Clay, under the central-western part of the State, is at least
2000 feet thick. The Niobrara Chalky limestones and the Benton
Shales are 450 feet thick to the east and thicker to the west. The
Dakota Sandstone (with a basal Lower Cretaceous sandstone), about
400 feet thick, underlies nearly all Nebraska. “ This formation
carries the great artesian-water supplies, which are so extensively
developed by wells in eastern South Dakota, and on low lands in
Knox, Cedar, and Boyd counties, Nebraska ” (page 9).
Volcanic ash constitutes two widespread beds in the Brule Clay,
some limited and some thick beds in the Gering formation, and a
general admixture as well as several thick bedsin the Arikaree. It
is rare in the Ogalalla, and it occurs in the lower portion of the
Loess in the central and western parts of the State. The ash was
probably blown over from volcanoes in the Rocky Mountains (pages
42 & 43).
Carboniferous limestones, shales, and sandstones underlie all
Nebraska. The Cottonwood Fusulina limestone has been recognized.
In the south-eastern corner of the State the sinkings have proved
about 1200 feet, of which 200 is Permian.
At Lincoln the Lower Limestones (partly Devonian) are at 1635
feet; then Trenton Limestone to 1947. Then follow magnesian
limestones 143 feet, then red sandstone (Cambrian) and red
quartzite (Alongian, pierced for 270 feet), resting on granite.
260 Bibliographical Notices.
A yast amount of valuable information, both special and general
in character, has been collected and published by the State Survey,
as exemplified in this volume ; but it is not yet sufficiently easy to
make a complete and definite judgment on the different districts,
as to the possible underground water-supply, without the further
experience and opinion of experts.
The Land and Sea Mammals of Middle America and the West Indies.
By Davin Giravp Extior, F.R.S.E. &e. Vol. IV. Parts I. & I.
Zoological Series, Field Columbian Museum. Chicago, U.S A.,
1904.
Turse two bulky parts will prove of inestimable value to mam-
malogists, and form a lasting monument to the credit of the author.
The more carefully they are studied, the more one becomes impressed
by the amount of labour and research which must have been
expended on their production.
The present work contains a description of all the mammals
from the northern boundary of Mexico to the Province of Cauea,
S. America, including the islands of the coast, as well as of the
Bahamas and the West Indies.
A feature of these parts, as compared with that already issued
on the mammals north of the area now dealt with, is the addition
of short accounts of the habits of all the more important species,
and many new facts will be found in these statements.
The author will gain the sympathy of a large number of zoolo-
gists in that he has chosen to protest vigorously against the system
now in fashion of multiplying species—a system calculated to do an
immense deal of harm. With some systematists, in fact, it 1s carried
to the very verge of absurdity. Names, the author trenchantly
remarks, ‘“‘are useful for the recognition of specimens possessing
independent distinctive characters, but if an example has none of
these its appellation is of little assistance.” And, again, “ It is, of
course, not to be conceived .... that the mere bestowal of a
name upon a specimen would make it recognisable; and that the
act of naming examples that are separated from their fellows on
account of . ... minute variations cannot be fairly regarded as
an ‘accurate statement of the results of organic evolution.”” As
he points out, ‘‘ Many specimens have been named whose cranial
characters consist altogether in being ‘longer or shorter,’ ‘ broader
or narrower’ than the corresponding parts of some other example,
and it is easily comprehended how slight is the probability that
any specimen can be accurately determined whose characters are
such as those given (the colour of the pelage also being nearly the
same), no topotypes of the forms with which these are compared
by their describer being available, and in many instances no
measurements of the crania having been given.” The lengths to
which some have carried this principle of bestowing names on what
are at most individual variations is appalling. Evidence of this can
be found in plenty in these volumes.
Illustrations have been lavishly distributed, and many, especially
of the crania, are extremely beautiful.
Bibliographical Notices, 261
Guide to the Gallery of Birds in the Department of Zoology of the
British Museum (Natural History). Pp. 228; 24 plates and
7 text-figures. London: Printed by Order of the Trustees of
the British Museum, 1905. Price 2s. 6d.
Tue Bird-Gallery of the British Museum of Natural History is,
without doubt, one of the finest in the world; and this not on
account of any architectural beauties which it possesses, but solely
because of the remarkable character of the specimens selected for
exhibition and the method of their display. Inasmuch as the
arrangement of the Gallery is the work of Mr. W. R. Ogilvie-Grant,
it was but fitting that the preparation of the Guide thereto
should also be entrusted to him. How admirably he has fulfilled his
task will best be appreciated by those who use this book in the
Museum. But the value of this volume is by no means confined to
its use there. On the contrary, it will prove of the greatest service
even to those who have never seen the specimens herein described.
Profusely and beautifully illustrated and crammed with information,
much of which is not to be found elsewhere, it will form a welcome
addition to the library of many a bird-lover throughout the
country. Museum Curators will find it indispensable.
As the visitor works his way down the Gallery, book in hand, he
will find each Order of birds briefly diagnosed, and following this
a description of all the more important species, their habits, and
so on.
The wonderful Nesting-Series of British Birds is described
separately ; and, in addition to this, there is an appendix wherein
will be found a general introduction to the study of birds, which
should prove most valuable to those who are not experts.
The beautiful plates which illustrate this Guide have been
photographed by Mr. R. B. Lodge from specimens actually exhibited
in the Gallery.
House, Garden, and Field. By 1. C. Mratt, F.R.S.
London: Edward Arnold, 1904.
Proressor Mitr has an established reputation asa writer on Natural
History subjects, and this reputation is amply sustained in his
latest work ‘ House, Garden, and Field.’ In the space of some
three hundred pages he has crowded a vast amount of information,
in the form of short essays, on themes of the most varied
description.
Designed more especially for teachers who are compelled, without
special qualification, to give lessons in ‘ Nature Study,’ Prof.
Miall has endeavoured rather to give his readers an insight into
what to observe, and how to observe, than to provide him with
ready-made lessons. As he rightly remarks, the teacher must
himself be an observer if his teaching is to be stimulating. If, as
he says, he“ gets all his knowledge without effort, then the
so-called Nature Study which he dispenses has no more power to
262 Miscellaneous.
excite the love of nature or the spirit of enquiry than a printed
list of the Kings of England with dates.” The introduction, indeed,
to this volume forms an admirable dissertation on the teaching of
this much-abused subject, and should be carefully studied by all
those for whom it is written. The present demand for books on
Nature Study has unfortunately produced an abundant crop of
most pernicious rubbish, which, strange as it may appear, has
been greedily absorbed by those responsible for the education of our
children. Professor Miall’s book is therefore to be welcomed as an
antidote to this poisonous stuff.
We cannot do more here, by way of indicating the nature of these
essays, than particularize one or two. As especially helpful we may
mention that on ‘* A School Course on the Structure and Life of
Insects.” Though short, it is altogether admirable; and his hints
on methods of displaying insect-structures to many people at once
will doubtless be gladly adopted. School Museums; Buttercups—a
Study of Species ; Vacation Rambles; The Summer droop of Boughs ;
The Rock-Barnacle; and The Natural History Excursion, are all
essays of real excellence and full of helpful suggestions.
In short, this is a book which we most cordially reeommend—not
only to teachers, but to dwellers alike in town and country who
have a love of Nature and but little training or practice in the way
to observe.
The illustrations are numerous and good, and the size of the
volume is sufficiently small to be carried conveniently as a guide to
practical work out of doors.
MISCELLANEOUS.
Note on the ‘ Museum Humfredianum, 1779.
By C. Davies SHERporn.
Tue rarity of sale-catalogues of the early collections in which
zoological specimens were preserved is well-known. After diligent
search during the time I was compiling my ‘ Index Animalium,’ one
by one fell into my hands, until at last the ‘Museum Humfredianum’
was the only one to escape me.
Mentioned first by Cobres in his ‘ Delicize Cobresiane,’ 1782,
p- 148; then by Fox in his ‘Synopsis Newcastle Museum,’ 1827,
. 179; it was again listed by Engelmann in ‘ Bibl. Hist. Nat.’
1846, p. 11. These, so far as I know, are the only references to
the Catalogue, though the Museum itself has been referred to many
times (see Murray, ‘ Museums,’ 1904, ii. p. 293 etc.). There is
no doubt that Cobres actually saw the tract, for he gives the
pagination correctly, although he is in error as to the size; but it
is highly probable that Engelmann copied from Cobres. Fox, too,
Miscellaneous. 263
must have handled the book, though he misspells the name,
‘ Humphredianum.’
The reference to Fox and subsequent recovery of the tract I owe
to Dr. David Murray, whose excellent book on ‘ Museums,’ pub-
lished in 1904, I eagerly perused on its appearance as likely to
give me aclue. A letter from Dr. Murray further assisted me, and
I found the Catalogue still preserved in the Hancock Museum,
Newcastle-on-Tyne. It was no doubt the copy marked by George
Allan himself when purchasing from Humphrey’s Museum, as there
are lines in the margin as mentioned by Mr. Fox.
I much regret that I had not found Fox’s reference to this
Catalogue before, as the ‘Mus. Humf.’ has been my “ old man of
the sea” for fifteen years, and | am exceedingly thankful to be able
to shake him off. It is also a somewhat remarkable circumstance
that no one has informed me of the existence of this copy at New-
castle, more especially as I advertised every week for nearly three
years for the loan of a copy in a publication which is, or should be,
in every library or museum of importance.
Considering that I could not see this tract before publishing my
‘Index Animalium,’ it now gives me peculiar satisfaction to
announce to zoologists that the ‘Museum Humfredianum’ has no
zoological value whatsoever. It is merely an auction sale-catalogue
of a general collection of curiosities, without systematic arrange-
ment of the zoological portion, and without generic headings or
bineminal nomenclature of any sort or description.
Indeed, it is now certain that George Humphrey did not employ
definite terms in print until he published his ‘ Museum Calonnianum’
in 1797, and this fact alone will account for the rarity of the ‘ Mus.
Humf, and the comparative abundance of the ‘ Mus. Calonn.’ ‘lhe
former was useless the day after the sale, the latter, being full of
new names, became at once a book of reference.
As the Catalogue has never been properly described, I give the
following as the full text of the title :—
(No. 9, 1779.) | Museum Humrrepianum: | [a line] | A |
CATALOGUE | Of the tare and varuaste | MUSEUM | of |
Mr. Groner Humparey; | Which is presumed to be the most
capital of the kind | ever offered to public Sale in this King-
dom: | Comprehending many Thousand curious Subjects in
NATURAL | HIsToRY, collected with great Judgment and Expence,
from most | Parts of the known World, during a Course of upwards
of thirty | Years: particularly Shells, Corals, and other beautiful
Marine Pro- { ductions; Reptiles, Insects, and other Animals, both
dried and in | Spirits; most of the Species in the Fossil Kingdom ;
among them | many rare and elegant Spars, Crystals (some of them
matchless) | Marbles, Agates, and other Stones ; Ores and Minerals ;
a great | Variety of uncommon Petrefactions of Animals and
Vegetables ; | some elegant Compositions in Shell-work, consisting
of Vases with | Flowers, Temples, &c. the best and most extensive
Collection of the | Cloths, Garments, Ornaments, Weapons of War,
264 Miscellaneous,
Fishing Tackle, and | other singular Inventions of the Natives of
Otahate, Nev Zealand, | and other new discovered Islands in the
South Seas; A Variety of | Mahogany and other Cabinets, adapted
for containing Subjects in | Natural History; a large Range of
elegant Mahogany Cases of the best | Jamaica Wood, nine Feet
high, glazed with large Panes, and might | easily be altered to
Library Cases ; a Mahogany Table, near thirty | Feet long, with
large Show Glasses on the Top, and near 300 Drawers | beneath,
all glazed; a Range of Mahogany Shelves with Cabinets, | con-
taining 128 Drawers, glazed; a perpetual Motion, an artificial
Hail | Storm; sundry Books in Natural History, a Set of six
Columns four- | teen Feet high, and a great Number of other
Articles. | Which will be sold by Avcrron, | By Mr. PATERSON, |
On the Premtses, | No. 70. St. Martin’s Lane, opposite Slaughter’s |
Coffee House, | (By Order of the Asstenrns,) | On Monday, April 5,
1779, and the Twenty-nine | following Days, | To be viewed on
Monday, March the 29th, and to the | Time of Sale. | Catalogues
may be had (Price One Surtune), at | Mr. Paterson's, No. 6 BKing-
Street, Covent Garden; | and at the Museum aforesaid. |
The Catalogue is printed in quarto (not octavo, as invariably
stated), measures 9 x 5? inches, has a Titlepage wich ‘‘ conditions
of sale” on back, and pp. 1-168. The sale occupied the thirty-six
days for which it was catalogued (the titlepage being incorrect on
this point), the last day taking books, of which Humphrey had a
poor lot, the only rarities being seven copies of his own ‘Conchologie,’
of which Allan seems to have purchased one. As this book is stated
in the ‘ Mus. Humf.’ itself to be “‘ Humphrey’s Conchologie,” it con-
firms the opinion expressed in my ‘ Index Animalium,’ 1902, p. xxx,
that Humphrey, and not E. M. DaCosta, was the author of the
book.
The Council of the Hancock Museum has generously agreed to
an exchange with me for this tract, feeling, with me, that so
interesting a curiosity should be in a more accessible position. I
have therefore arranged to deposit the ‘Mus. Humf.’ in the General
Library of the British Museum (Natural History), where it can be
seen by anyone interested.
Where are the Types?
GENTLEMEN,—May I ask if any of your readers can kindly give
me information as to where may be found the types of S. P. Pratt
(Kelloway Ammonites from Christian Malford) and Young and
Bird (Lias fossils in * Geol. of Yorkshire’)? The information is
required for the purposes of the ‘ Paleontologia Universalis,’
8. 5. Buckman,
Westfield, Thame, Oxon.
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MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. 93. SEPTEMBER 1905.
XXIX.—Rhynchotal Notes—XXXV. By W. L. Distant.
Fam. Cicadide (continued from p. 216).
Subfam. 7rercrwrw (concluded).
THIS communication concludes my efforts to revise and
systematize the genera of the family Cicadide. ‘here are a
few genera with which I am still unacquainted— Nosola,
Stal, Nadlistes, Karsch, Plautilla, Stal, Paectira, Karsch,
and Triglena, Fieb.,—but these will, I hope, be located in a
subsequent synonymical catalogue of the family.
Division TETTIGOMYIARIA.
Tegmina broad and short, not or scarcely longer than the
body, sometimes scarcely longer than the abdomen; abdomen
usually inflated, always thickened.
Synopsis of Genera.
A. Bases of upper vein to lower ulnar area and lower
vein to radial area distinctly and well separated at
apex of basal cell.
a, Wings with six apical areas.
6. Tegmina about as long as body.
c. Head as long as breadth between eyes; front
produced and as long as broad ............ Xosopsaltria.
ec. Head considerably shorter than space between
eyes; front not produced, short, transverse,
PARR gue UST oes, <a chala sa ote eye's aei8 6 0s Inyamana.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 18
266 Mr. W. L. Distant on Homoptera.
ece. Head slightly shorter than space between
eyes; front slightly produced, about as long as
LOAM a Wise Mae des Fe em sue dunlehe eee Saticula.
aa. Wings with five, sometimes only four, apical areas.
bb. Tegmina only about two thirds the length of
body SOA AMOI HOMO AG Morr PC Oa On Gazuma.
B. Bases of upper vein to lower ulnar area and lower
vein to radial area with almost a common origin at
mpexicl PSSAL CCl a atem inc rn reir . Tettigomyia.
Genus XOSOPSALTRIA.
Pydna, Stil, Ann. Soc. Ent. Fr. (4) i. p. 621 (1861), nom. preoce.
Xosopsaltria, Kirk. Entomologist, 1904, p. 280, nom. n.
Type, X. lutea, Oliv. (Cicada).
NXosopsaliria fuliginosa, sp. n.
d. Head and pronotum sordidly ochraceous ; head with
marginal and submarginal lines to front, spots at lateral
margins of vertex, and area of the ocelli, pronotum with two
central fasciee with a curved discal fascia on each side, the
incisures and lateral margins, black; mesonotum sordidly
ochraceous, with four large obconical spots (the two central
shortest), a large spot in front of cruciform elevation and the
outer margins of same, black; abdomen sordidly ochraceous,
the posterior segmental margins narrowly testaceous, with
two central longitudinal series of piceous spots ; body beneath
sordidly ochraceous; transverse striz to face, disk of sternum,
legs, and base of metasternum piceous; tibiae (excluding
apices) ochraceous ; tegmina pale shining fuliginous, be-
coming more or less hyaline i in the various areas, the venation
pale fuscous ; ; wings hyaline, the venation fuscous; tegmina
broad, not longer than’the body, their greatest breadth little
less than half their length; abdomen prominently centrally
ridged above ; opercula inwardly oblique, not passing base
of abdomen.
Long., excl. tegm., 21 mm.; exp. tegm. 45 mm.
Hab. Brit. Ef. Africa (C. S. Betton, Brit. Mus.).
INYAMANA, gen. nov.
Head (including eyes) almost as wide as base of meso-
notum, shorter than breadth between eyes, lateral margins of
vertex convex and projecting slightly beyond front, which is
short, transverse, broader than long,. concave, and medially
sulcate, ocelli a little nearer to each other than to eyes; pro-
notum longer than head, the lateral margins a little convex,
nearly straight, the posterior angles “broadly ampliate ;
Mr. W. L. Distant on Homoptera. 267
mesonotum (including cruciform elevation) about as long as
head and pronotum together; abdomen dilated, considerably
longer than space between apex of head and base of cruciform
elevation; tympana entirely exposed; opercula in male
obliquely convex, longer than broad, not passing base of
abdomen; rostrum passing the anterior coxe; tegmina and
wings hyaline; tegmina short, broad, about as long as body ;
basal cell twice as long as broad, postcostal area moderately
broad and cell-like, apical areas eight ; wings with six apical
areas,
Type, Z. hemaris, Dist.
Inyamana hemaris, sp. 0.
Body ochraceous ; head with some small pale fuscous spots
and two short black lines at inner margins of ocelli; pro-
notum with two central longitudinal lines united posteriorly,
on each side of which is a curved discal line, the outer in-
cisures and lateral margins, piceous or black; mesonotum with
four long obconical spots, the two central shortest and placed
close together, and two small spots in front of the cruciform
elevation black ; abdomen with two central broken macular
fascisze and a lateral broken macular fascia on each side black ;
body beneath with the disk of sternum, inner areas of
opeicula, and two central, broken, macular fascize to abdomen
piceous or black ; tegmina and wings hyaline, the venation
mostly fuscous, costal membrane to tegmina ochraceous.
Long., excl. tegm., ¢ ? 14-16 mm.; exp. tegm.34-35 mm.
Hab. Brit. East Africa: Samburu (C. S. Betton, Brit.
Mus.) ; Kondoa (Bloyet, Paris Mus.).
Inyamana ochracea, sp. n.
Body and legs ochraceous, concolorous; tegmina and
wings hyaline, the venation brownish ochraceous; costal
membrane to tegmina ochraceous.
Allied to L. hemaris, but smaller, colour totally different ;
osteostal area to tegmina much broader, lower apical area
much shorter.
Long., excl. tegm., ¢ 9, 11-14mm.; exp. tegm. 25-30 mm.
Hab. Brit. East Africa: Samburu (C. S. Betton, Brit.
Mus.).
Genus SATICULA.
Saticula, Stal, Berl. ent. Zeitschr. x. p. 172 (1866).
Cicada, 5. Gruppe, Hagen, Stett. ent. Zeit. xvii. p. 158 (1856).
Type, S. coriaria, Stal.
18*
268 Mr. W. L. Distant on Homoptera.
GAZUMA, gen. nov.
9. Body elongate, robust ; head (including eyes) slightly
narrower than base of mesonotum, about as long as space
between eyes, front moderately produced, a little broader than
long, margins of front and vertex discontinuous, vertex longi-
tudinally suleate ; pronotum about as long as head, lateral
margins obliquely sinuate, posterior angles moderately
ampliate; mesonotum (including cruciform elevation) about
as long as head and pronotum together; abdomen much
longer than space between apex of head and base of cruciform
elevation ; tympana entirely exposed ; rostrum passing the
anterior coxe; tegmina and wings hyaline ; tegmina short,
broad, only about two thirds the length of body, apical areas
short, eight in number, in some specimens less; wings with
five, in some specimens only four, apical areas.
This description is founded ona series of female specimens.
Type, G. Barrette, Dist.
Gazuma Barrette, sp. n.
9. Body and legs ochraceous, concolorous; tegmina and
wings hyaline, the venation ochraceous ; tegmina with a
slight ochraceous tint, the costal membrane ochraceous ; third
ulnar area very much shorter than fourth; anterior femora
armed with three strong spines beneath.
Long., excl. tegm., 9 10-12 mm.; exp. tegm. 19-20 mm.
Hab. South Africa: Annshaw (Jhiss F. Barrett, Brit.
Mus.).
Gazuma Delalandet, sp. n.
Uniformly brownish ochraceous; tegmina and wings
hyaline, the first a little tinted with very pale brownish, the
venation and costal membrane brownish ochraceous ; wings
with the venation dull ochraceous.
Allied to G. Barrette by venation of tegmina, which are,
however, a little shorter, front of head a little more prominent ;
pronotum deeply, centrally, longitudinally sulcate.
Long., excl. tegm., 9 13 mm.; exp. tegm. 19 mm.
Hab. §. Africa (Delalande, Paris Mus.).
Gazuma pretorie, sp. n.
Closely allied to G. Barrette in general appearance, but
virescent, not ochraceous, and differing structurally from
that species by the length’of the third ulnar area, which is
Mr. W. L. Distant on Homoptera. 269
nearly twice as long as in G. Barrette and almost as long as
the fourth.
Long., excl. tegm., 9 12 mm. ; exp. tegm. 21 mm.
Hab, Transvaal: Pretoria.
Genus TETTIGOMYIA,
Tettigomyia, Amy. & Serv. Hist. Hém. p. 482 (1843).
Type, 7. vespiformis, Amy. & Serv.
Division MELAMPSALTARIA.
This division, though at present containing only three
genera, embraces a large number of species. It is easily
recognized by the venation of the tegmina, in which the
bases of the upper vein to lower ulnar area and the lower
vein to radial area are fused or the first emitted at some
distance from the base of the second. ‘The distribution of
the Melampsaltaria is in the Old. World, though there is a
Nearctic representative ; but it is in Australia where the
greatest number of species is found and where the head-
quarters of the division is clearly indicated.
Synopsis of Genera.
ee taps WIth SIX APICH ALCAS. .)).t on. es oats + «cs yee Melampsalta.
Pavanps with five apical areas... ct. le cant. ieee Pauropsalta.
Cm Wangs with fourapical abeas .oj/.i0% sisa soils else aleve Urabunana.
Genus MELAMPSALTA.
Melampsalta (Melampsalte), Amy. Ann. Soc. Ent. Fr. (2) vy. p. 155
[351] (1847).
Melampsalta, Kolenati, Mél. Ent. vii. p. 27 (1857).
Type, AZ. musiva, Germ. (Cicada).
Considerable difficulty appertains as to the nomenclature
of this genus. It was first named by Amyot (scarcely
described), supra, and takes precedence of the name Cvca-
detta which he used on a succeeding page. Kolenati described
the genus, but had previously also described Cicadctta,
which, if we take Kolenati as the real founder, must have
priority. However, I follow S:al, Karsch, and other writers
in giving the name A/elampsalta precedence, as given by
Amyot. I have failed to separate Cicadetta from JMelam-
psalta: if we compare the type of the first, the Cicada montana,
Scop., with the type of Melampsalta (supra), a difference
appears in the position of emergence of the upper vein of the
lower ulnar area; but much as { endeavoured to thus break
270 Mr. W. L. Distant on Homoptera.
up a congested genus, my efforts failed, and the differences
became evanescent when a long series of species were
examined,
Melampsalta Godingi.
Melampsalta umbrimargo, God. & Froge. Proc. Linn. Soc. N. §,
Wales, 1904, p. 652, t. xviii. fig. 12.
Head black; a central line to front, a spot on each lateral
margin, an elongate central basal spot, a small spot near inner
margin of each eye, and the posterior margin of vertex
ochraceous ; pronotum ochraceous ; two central fascize united
posteriorly and ampliated anteriorly, some irregular diseal
markings, inner posterior margin, and lateral and sublateral
margins black « mesonotum black; the margins of two
central obconical spots, the lateral margins, and cruciform
elevation ochraceous, the last with a black spot on each
anterior angular prolongation ; abdomen black, posterior
segmental margins ochraceous ; sternum very thickly. grey-
ishly pilose; face black, its lateral margins ochraceous ;
opercula in male greyishly yellow, bases piceous ; abdomen
beneath ochraceous, with a central and lateral segmental
series of black spots ; tegmina and wings hyaline, venation
brownish ochraceous ; tegmina with the costal membrane,
apical margin, bases of second, third, fifth, and seventh apical
areas, longitudinal veins to fifth, sixth, and seventh apical
areas, and spots at apices of longitudinal veins to upper four
ulnar areas umber-brown.
Long., excl. tegm., ¢ 184 mm.; exp. tegm. 56 mm.
Hab, Australia: Ardrossan, Yorke Peninsula (Tepper) ;
Gawler, S.A. (Macleay Mus.).
A much smaller species than the Cicada umbrimaryo,
Walk. ; tegmina much shorter and their apices more rounded;
colour and markings also different.
M.lampsalta ethiopica, sp. n.
Body above black, sparingly palely ochraceously pilose ;
ocelli castaneous ; lateral margins of pronotum, lateral
margins of metanotum (narrow ly}, and posterior angles of
cruciform elevation pale ochraceous; body beneath and
opercula pale ochraceous, strong suffusions to femora and
anterior and intermediate tibice, tarsi (more or less), and a
broad, central, longitudinal fascia to abdomen piceous or
black ; tegmina and wings hyaline, the venation fuscous ;
the tegmina with the costal membrane more or less casta-
neous, the upper vein of the lower ulnar area emitted at a
Mr. W. L. Distant on Homoptera. 271
very short distance from base of radial area, the costal
margin moderately arched to apex of radial area, margin of
postcostal area straight; opercula in male somewhat long,
their apices convex and reaching the apex of the second
abdominal segment; rostrum just passing the intermediate
COX.
Long., excl. tegm., ¢ 12 mm.; exp. tegm. 30 mm.
Hab. Brit. East Africa: Zomba (A. Whyte, Brit. Mus.).
Melampsalta Waterhouset, ep. ni.
Body above black, sparingly greyishly pilose; head with
a central fascia (broadened posteriorly) to front, a small spot
on each lateral margin and one at base of vertex, ochraceous ;
pronotum much suffused with ochraceous on the disk of
each lateral area, its extreme posterior margin of the same
colour; disk of cruciform elevation, central margin (inter-
rupted medially) of metanotum, a spot on each side of anal
segment, and anal appendage (excluding base) pale ochra-
ceous ; posterior margins of third, fourth, and fifth abdominal
segments dull ochraceous ; head beneath, sternum, legs, and
rostrum piceous; streaks and apices to anterior and inter-
mediate femora, intermediate tibia, posterior femora and
tibia, opercula in male, and abdomen beneath stramineous ;
posterior femora with some piceous lines; tegmina with a
slight brownish-ochraceous tinge, the venation ochraceous,
fuscous towards apices, the upper vein of lower ulnar area
attached to lower vein of radial area at a short distance from
base; opercula in male short, transverse, not extending
beyond base of abdomen.
Long., excl. tegm., ¢ 13 mm.; exp. tegm. 37 mm.
Hab. Australia: Adelaide (7. Waterhouse, Brit. Mus.).
Melampsaita lactea, sp. n.
Body above black, somewhat strongly ochraceously pilose ;
head with a spot on front and the lateral margins of vertex
and a central basal incised spot to same, margins and a
central linear spot to pronotum, lateral margins of mesonotum,
and the cruciform elevation ochraceous; head beneath,
sternum, and legs piceous or black ; margins and a spot at
base of face, apices and streaks to femora, and annulations
(more or less broad) to tibiae ochraceous ; abdomen beneath
pale ochraceous, lateral marginal and central segmental
series of spots piceous; tegmina talc-like, suffused with pale
fuscous; base and apex of radial area, the last continued to
apex of lower ulnar area, postcostal area, and some pale
272 Mr. W. L. Distant on Homoptera.
suffusions at bases of apical areas lacteous; a black streak in
basal cell, costal membrane, and about basal half of venation
ochraceous, remaining venation brownish ochraceous, costal
edge of postcostal area fuscous, upper vein of lower ulnar
area connected with lower vein of radial area for some distance
from base ; wings hyaline, venation ochraceous, base and
anal area lacteous ; opercula in male black, transverse, poste-
riorly rounded, not passing base of abdomen.
Long., excl. tegm., ¢ 17 mm.; exp. tegm. 39 mm.
Hab. Australia: Melbourne (Brit. Mus.).
Melampsalta issoides,
assotdes, Walk., MS.
Head black, with a linear spot to front and a small rounded
spot at base of vertex ochraceous; pronotum dull testaceous,
the submargins and two central lines dilated posteriorly,
black, the incisures piceous; mesonotum black, with two
very obscure central obconical spots, the margins of which
are dull testaceous, lateral margins, cruciform elevation, and
margins of metanotum, pale testaceous; posterior segmental
margins obscurely testaceous ; body beneath and legs thickly
ochraceously pilose ; tegmina and wings hyaline, the venation
fuscous, a somewhat broad (broadest at apex) oblique fuscous
fascia crossing tegmina at bases of apical areas, more or less
broken posteriorly, the upper vein of lower ulnar area emitted
at only a short distance from base of radial area; opercula in
male small, posteriorly rounded, not extending beyond base
of abdomen,
Long., excl. tegm., ¢ 12 mm.; exp. tegm. 27 mm.
Hab. Australia: New South Wales; Swan River (Brit.
Mus.). West Australia: Perth (7/7. J. Turner, Brit. Mus.).
Genus PAUROPSALTA.
Pauropsalta, God. & Frogg. Proc. Linn. Soe. N. S. Wales, 1904,
p. 615.
Type, P. mneme, Walk. (Cicada),=P. leurensis, God. &
rogg.
Mr, Froggatt has very kindly sent me a number of co-types
of the species he described and identified in conjunction with
Mr. Goding in their monograph of the Australian Cicadide.
I am thus enabled to fix the exact nomenclature of the type
of their genus. All further synonymy will appear in my
synonymic catalogue. It is scarcely to be expected that
Walker’s descriptions allow identifications to be made with
Mr. W. L. Distant on Homoptera. 273
certainty when his types are not available for comparison.
The figure of P. lewrensts given by Goding and Froggatt is
imperfect in the neuration of the tegmina, where the upper
margin of the lower ulnar area is represented by the artist as
emerging close to the apex of the basal cell, whereas it
bifurcates some distance from it.
Pauropsalta stigmatica, sp. n.
@. Head black; a central longitudinal line to front, a
small central spot at base, and a similar spot on each lateral
margin of vertex ochraceous ; pronotum ochraceous, with two
central, longitudinal, piceous lines, angularly united poste-
riorly ‘and ampliately united anteriorly, the incisures and
lateral margins fuscous ; mesonotum black, with two central
pale ochraceous fasciw, which are widened and inwardly
angulated on disk, the lateral margins and cruciform elevation
ochraceous, the last with a central line and apices of the
anterior angles piceous or black ; abdomen black, the poste-
rior segmental margins broadly ochraceous, that on the
posterior segment inedially interrupted ; anal segment testa-
ceous, centrally black; body beneath ochraceous; head
beneath black, lateral margins and an anterior spot to face
ochraceous ; legs more or less suffused with piceous, the
anterior legs most strongly so; tegmina and wings hyaline,
the venation mostly fuscous ; tegmina with the costal’ mem-
brane pale ochraceous, the postcostal area fuscous and
appearing as a dark stigma; wings with five apical areas.
Long., excl. tegm., 9 11 mm.; exp. tegm. 29 mm.
Hab. Adelaide (Ff. Waterhouse, Brit. Mus.).
Pauropsalta Dameli, sp. n.
3. Body above black ; central sulcation at base of vertex
ochraceous ; a spot on each lateral margin of vertex, margins
and a central fascia to pronotum, an irregular discal spot
(mutilated in type by pin) and lateral margins of mesonotum,
cruciform elevation, ‘posterior area of metasternum, abdominal
segmental margins, "and disk of anal segment, testaceous red ; ;
body beneath testaceous ; head beneath (excluding margins
of face) and discal suffusions to sternum piceous; legs,
rostrum (excluding apex), lateral margins of sternum, and
the opercula pale ochraceous ; tezmina and wings hyaline,
the venation ochraceous ; tegmina distinctly arched above
the radial area, with the costal membrane and postcostal area
ochraceous; wings with five apical areas.
Long., excl. teem., 91 mm.; exp. tegm. 22 mm.
Hab. “ Australia” @ idw. Damel, Brit. Mus.).
274 Mr. W. L. Distant on Homoptera.
Genus ?
Cicada complex, Walk. List Hom. i. p. 208 (1850).
The type is unlocalized, without abdomen, and with one
wing mutilated, the remnants preserved being gummed on a
card. The shape and venation of the tegmina resemble those
of Pauropsalta dubia, God. & Frogg., but the one wing
retained possesses only three apical areas. Under such
circumstances it is scarcely possible to generically identify it.
URABUNANA, gen. nov.
Head considerably shorter than pronotum, the front broad
and rounded, its margins discontinuous to, and almost at
right angles with, the lateral margins of vertex, which are
also rounded, vertex much longer than front and with a
central incision at base, ocelli a little nearer to each other than
to eyes, which project beyond the anterior pronotal angles ;
pronotum somewhat quadrangular, its lateral margins straight,
its posterior angles a little ampliated; mesonotum about as
long as pronotum; abdomen about as long or slightly longer
than space between apex of head and base of cruciform
elevation; tympana entirely exposed; opercula in male
obliquely directed inwardly, but not extending beyond the
base of the abdomen; rostrum reaching the intermediate
coxe; anterior femora strongly spined beneath ; tegmina and
wings hyaline; tegmina with eight apical areas, the upper
vein of lower ulnar area emitted at some distance from
base of radial area, where they are fused, postcostal area
moderately broad ; wings with four apical areas.
Type, U, sericervitta, Walk. (Cicada).
Urabunana segmentaria, sp. n.
Body above black, sparingly greyishly pilose; a small
spot at apex and the basal narrow margin of front, a small
spot on each lateral margin, and a central spot at base of
vertex, lateral and posterior margins of pronotum, cruciform
Sleseaon! posterior margin ofl metasternum, and lateral
margins and posterior segmental margins of abdomen ochra-
ceous; body beneath ochraceous; head beneath (excluding
margins of face), clypeus, disk of sternum, basal margins of
opercula, suffusions to coxe and femora, anterior tibie and
tarsi, and base of abdomen black; tegmina and _ wings
hyaline, bases of both narrowly ochraceous; tegmina with
5
the costal membrane, postcostal area, and basal veins pale
Mr. W. L. Distant on Homoptera.
bo
279
ochraceous, remaining venation fuscous ; venation of wings
ochraceous or fuscous ; wings with four apical areas ; anterior
femora with two strong black spines.
Long., excl. tegm., ¢ 10 mm.; exp. tegm. 24 mm.
Hab. Queensland: Peak Downs (Coll. Dist.).
Division HEMIDICTYARIA.
The principal divisional character of the Hemidictyaria is
to be found in the narrow head, which (including eyes) is
about or scarcely more than half the width of mesonotum at
base ; the abdomen (especially in the males) is more or less
globose and usually longitudinally dorsally ridged.
Lacetas, Karsch, from the figure illustrating the description,
apparently belongs to this division, but I have not yet seen
a representative of the genus, and therefore cannot locate it
in the synopsis.
Synopsis of Genera.
A. Venation of tegmina normal, not reticulate.
a. Front of head triangular, deeply and angularly in-
serted in vertex.
6. Head longer than space between eyes.
c. Ocelli placed as near eyes as to each other.
d, Apical areas of tegmina and wings almost as
long as half their | Gitiegn oe Sete 7 pee . Arfaka.
aa. Front of head conical , moderately but not angularly
inserted in vertex.
bb. Head about as long as space between eyes.
ce. Ocelli placed nearer to each other than to eyes.
dd. Apical areas of tegmina and wings shorter
than half their leneth,
e. Lateral margins of pronotum more or less
sinuate.
Jf. Fourth ulnar area of tegmina of normal
BHEDOn cs Wii Ride hebeern «peeercles wigaie's Sapantanga.
Jf. Fourth ulnar area of tegmina with its
anterior margin at base “of third ulnar
area inwardly angulated and emitting a
short rudimentary Ve VENI ee Pete teint . dacatra.
ee. Lateral margins of Prone obliquely
BuPAIO MG, ho Ste oh ee sed v otha Ore eee rots Truana.
eee. Ocelli placed as near to eyes as to each other,
g. Tegmina with the first ulnar area much longer
than third; fourth ulnar area much shorter
than radial area.
h. Norudimentary vein at base of second ulnar
US hetie ag Mees sk MORASS 2 es Dad Be 8 sisetene stds . LPrasia,
gg. Tegmina with the ‘first ulnar area about as
long as third; fourth ulnar area about or
nearly as lone, or a little longer than radial
area,
276 Mr. W. L. Distant on Homoptera.
hh. Rudimentary vein at base of second ulnar
area connecting it more or less imperfectly
with apex of ‘fifth ulnar area, sometimes
almost obsolete .......... Screcea —bemogjite
B. Venation of tegmina more or less reticulate.
zt. Bases of upper vein to lower ulnar area and lower
vein of radial area not amalgamated, the first alone
emitted from end of cell, contiguous to emergence
Ofjseconde:.aeers toes ausgh oe RES tags Siem enoeerom bhoneors Cystosoma.
#. Bases of upper vein to lower ulnar area and lower
vein of radial area amalgamated.
J. Reticulated area of tegmina not more than half
their length and sharply defined by a transverse
LING =, Acopaiee a csestonmeiens Gasper renaserese fale acetone Hemidictya.
Jj. Reticulate area of teemina more than half their
length and not defined by a transverse line .... ovana.
ARFAKA, gen. nov.
Head longer than space between eyes, front long, about as
long as vertex, deeply inserted in vertex and angulate ante-
riorly and posteriorly, breadth (including eyes) little more
than half the breadth of mesonotum at base, eyes large and
oblique, ocelli almost as near to eyes as to each other ; pro-
notum about as long as head, its lateral margins moderately
convex, its posterior angles widely ampliated ; mesonotum
(including cruciform elevation) longer than pronotum ; abdo-
men moderately inflated, attenuated posteriorly, dorsally
medially ridged, longer than space between apex of head
and base of cruciform elevation ; tympana entirely exposed ;
opercula in male extremely small, leaving the cavities
almost entirely exposed; anterior femora only moderately
spined beneath; face much compressed and prominent;
rostrum reaching the intermediate coxe ; tegmina and wings
hyaline; tegmina with eight, wings with six apical areas,
which in both cases are long and occupy at least half their
length.
Type, A. fulva, Walk. (Cephalorys).
SAPANTANGA, @en. nov.
Te)
Head about as long as space between eyes, front long,
about as long as vertex, conical, not angulately immersed in
vertex, breadth (including eyes) about two thirds the width
of mesonotum at base ; pronotum about as long as head, its
lateral margins a little convex, its posterior angles moderately
ampliated ;~ mesonotum (including cruciform elevation) con-
siderably longer than pronotum; abdomen short, robust,
* Characters taken from Westwood’s figure,
Mr. W. L. Distant on Homoptera. 277
attenuated posteriorly, not longer than space between apex
of head and base of cruciform elevation; tympana entirely
exposed ; opercula in male very small, the cavities almost
entirely exposed ; ; rostrum about reaching the anterior coxe ;
face moderately compressed, strongly longitudinally suleate ; :
tegmina and wings hyaline; tegmina with eight, wings with
six apical areas, which in both cases are shorter than half
their length.
Type, S. nutans, Walk. (Cephalowys).
JACATRA, gen. nov.
Head (including eyes) about half the breadth of base of
mesonotum, about as long as broad, front moderately pro-
duced, subtriangular, considerably shorter than vertex, its
lateral ma argins °and those of vertex discontinuous, ocelli
nearer to each other than to eyes, well separated from base of
head; pronotum about as long as head, its posterior angles
somewhat strongly ampliate ; “mesonotum (including cruci-
form elevation) longer than pronotum; abdomen robust,
centrally longitudinally ridged, longer than space between
apex of head and_ base of coe ae elevation; tegmina
hyaline, apical areas eight, first ulnar area longer than ‘third,
fourth ulnar area shorter than radial area, the transverse vein
at base of third ulnar area strongly angulated and emitting a
short rudimentary vein halfway across fourth ulnar area ;
wings with six apical areas.
Type, J. typica, Dist.
Jacatra typica, sp. 0.
Body, legs, tegmina, and wings pale dull ochraceous ;
head slightly tinged with virescent ; pronotum with the
margins and a central longitudinal fascia virescent ; meso-
notum with two anterior central obconical spots, denoted only
by their pale margins, and with a broad virescent fascia on
each lateral area; posterior segmental margins (excluding
basal and apical segments) pale testaceous ; fegmina with a
dull ochraceous tint, the venation and costal membrane a little
darker, the veins ‘also alternately marked with elongate
darker and lighter spots ; wings with a dull ochraceous tint,
the veins darker ; tegmina with the additional curved rudi-
mentary vein very distinct and perfect for halfway across the
apical area of fourth ulnar area.
Long., 3, 20 mm.; exp. tegm. 47 mm.
Habyi Java (Brit. Mus.).
278 Mr. W. L. Distant on Homoptera.
TRUANA, gen. nov.
Head triangular, considerably shorter than the pronotum,
including eyes about half the width of base of mesonotum ;
front a little shorter than vertex, somewhat triangular,
centrally finely sulcate, lateral margins of front and vertex
almost continuous; vertex with two central longitudinal
ridges, exhibiting a deep sulcation between them; front
ocellus placed very near anterior margin, the other two on
each side of ridges, a little nearer to each other than to eyes,
lateral margins and eyes oblique ; pronotum narrowed ante-
riorly, lateral margins oblique, posterior angles a little
subangularly ampliate, centrally, finely, longitudinally sulcate;
mesonotum (including cruciform elevation) a little longer
than pronotum; abdomen short, prominently, centrally,
dorsally ridged; tympana entirely exposed; opercula in
male small, short, not entirely covering the sonorous apertures ;
teemina semiopaque, costal membrane somewhat broad and
concave, postcostal area broad and distinct, apical areas eight ;
wings with six apical areas.
Allied to Lacetas, Karsch, differing principally by the
number of apical veins to the tegmina.
Type, Z. sulcata, Dist.
Iruana sulcata, sp. n.
Somewhat uniformly pale ochraceous ; tegmina semiopaque,
with an ochraceous tint; wings semilyaline, with the venation
ochraceous; head and pronotum more or less centrally longi-
tudinally sulcate, the front finely, the vertex most strongly
sulcate; rostrum reaching the intermediate coxe; other
characters as in generic diagnosis.
Long., excl. tegm., ¢ ¢ 15-16 mm.; exp. tegm. 48—
00 mm.
Hab. Brit. E. Africa: Lagari (C. S. Betton, Brit. Mus.).
Genus PRASIA.
Prasia, Stal, Tr. Ent. Soc. Lond. (3) i. p. 574 (1863).
Drepanopsaltria, Bredd. Hem, Celebes, p. 113 (1902).
Type, P. faticina, Stal.
I am unable to follow our Prussian colleague in separating
his genus Drepanopsaltria from Prasia. (1) He relies on
the presence or absence of the opercula, and gives a figure
representing a species of Prasza without stating what that
species is. Now the type of Prasia—P. faticina, Stal—is in
the British Museum, and is a female specimen, consequently
Mr. W. L. Distant on Homoptera. 279
we cannot refer to the opercula, and I have never seen a male
specimen. Stal, in describing his P. fatiloqua, certainly
writes ‘ operculis nullis,’ but this cannot be accepted as the
type of Prasia, and the true female type has ail the appear-
ance of what Breddin refers to as typical Drepanopsaltria.
(2) To add to the confusion, Breddin gives as the type of
Drepanopsaltria, Lembeja culta, ‘Dist., a species I have never
described ; but it is probable he w as referring to my Prasia
culta (Ann. & Mag. Nat. Hist. (7) i. p. 97, 1898).
Genus LEMBEJA.
Lembeja, Dist. Mon. Orient. Cicad. pp. 103 & 147 (1892), n. nom.
Perissoneura, Dist. Proc. Zool. Soc. Lond. 1885, p. 189, nom. preoce.
Type, L. maculosa, Dist. (Perissoneura).
In reference to this genus, Herr Breddin (Hem. Celebes,
p- 113, 1902) has formulated the dogma ‘Genus Lembejam
Dist. a Prasia Stal naturaliter distingui non posse
existimo.” But this is more of a criticism on the veteran
entomologist Dr. Karsch, of his own Berlin Museum, who has
described two species in that genus.
Genus CYSTOSOMA.
Cystosoma, Westw. Arc. Ent. i. p. 92 (1842),
Type, C. Saundersi, Westw.
Genus HEMIDICTYA.
Hemidictya, Burm. Handb. ii. 1, p. 178 (1839); Amy & Serv. Hist.
Hém. p. 461 (1843),
Type, H. frondosa, Burm.
HOvANA, gen. nov.
Head triangular, about as long as the pronotum, including,
eyes about half the width of mesonotum, front and vertex
subequal in length, eyes outwardly truncate, in a line with
lateral margins of head, base of head frunentes pronotum
with the lateral margins obliquely straight, narrowing to
head, posterior angles prominent, obtusely angulate ; meso-
notum about as long as pronotum ; metanotum triangularly
posteriorly produced ; abdomen (2?) short, robust, shorter
than space between apex of head and posterior angle of
metanotum ; face obliquely compressed; rostrum robust,
reaching intermediate coxe; tegmina opaque, broad, acuminate
at apex, greatest breadth more than half their length, costal
280 Mr. W. L. Distant on Homoptera.
membrane very broad, more than apical half with the venation
reticulate and forming very many cellular areas, upper vein
of lower ulnar area amalgamated at base with lower vein of
radial area; wings hyaline, with eight apical areas.
Type, H. Distant’, Brancsik (Hemidictya).
Division TETTIGARCTARIA.
A division at present only represented by one remarkable
genus, in which the pronotum differs from that in any other
genus of the family by its large size, produced posteriorly
and almost covering the mesonotum.
Genus TETTIGARCTA.
Tettigarcta, White, Eyre’s Tray. in Australia, Append. i. p. 412 (1845).
Type, 7. tomentosa, White.
APPENDIX.
TAP ur oR ARIA,
DULDERANA, gen. nov.
Head broad, eyes projecting beyond the anterior angles of
the pronotum, front broad, considerably shorter than vertex
and centrally longitudinally finely sulcate, margins of front
and vertex moderately discontinuous, ocelli nearer to each
other than to eyes; pronotum about as long as broad, its
lateral margins slightly convex, its posterior angles angu-
lately ampliate ; mesonotum (including cruciform elevation)
about as long as head and pronotum together; metanotum
posteriorly and laterally produced behind the exposed
_tympana; abdomen in male very short, not so long as space
between apex of head and base of cruciform elevation ;
rostrum reaching the posterior coxee; opercula narrow, con-
vexly transverse, obliquely curved inwardly, a_ posterior
metasternal process occupying the space between their inner
angles ; anterior femora with three strong spines beneath ;
tegmina about half as long again as the body, costal margin
suddenly arched near base, third ulnar area very broad and
long, second ulnar area very short, oblique, only about half
the greatest length of first, lower ulnar area very narrow, its
upper margin well separated from lower vein of radial area,
upper vein of penultimate ulnar area emitted at some distance
On the Bats of the Rhinolophus arcuatus Group, 281
from base of radial area, apical areas eight; wings with six
apical areas.
Type, D. truncatella, Walk. (Cicada).
The principal characters of this genus are to be found in
the structure of the tegmina.
Abricta Elseyt, sp. n.
Head black, the front castaneous, the ocelli and a central
spot at base of vertex ochraceous ; pronotum pale castaneous,
with a central longitudinal ochraceous fascia, which is much
ampliated posteriorly and a little so anteriorly, posterior
margin more or less piceous ; mesonotum ochraceous, with
four obconical castaneous spots, of which the two outermost
are much the largest and occupy the whole lateral areas,
cruciform elevation castaneous; abdomen castaneous, greyishly
pilose ; body beneath and legs castaneous, greyishly pilose,
head beneath (excluding face) black; tegmina and wings
hyaline; tegmina with the veins and costal membrane
brownish ochraceous ; wings with the venation ochraceous,
the whole anal area pale g eolden yellow, posteriorly margined
with fuscous; rostrum passing the intermediate coxe ;
opercula in male obliquely directed inwardly, but scarcely
extending beyond base of abdomen.
Long., excl. tegm., g¢ 19 mm.; exp. tegm. 60 mm.
Hab. North Australia: Victoria River (Dr. J. Rk. Elsey,
Gregory Exploration Exped., Brit. Mus.).
Allied to A. ruber, God. & Frogg., but tegmina apically
unspotted, different colour of head and anal area to wings,
shorter ulnar areas to tegmina, &c.
XX X.—On the Bats of the Rhinolophus arcuatus Group, with
Descriptions of Five new Forms. By Kxup ANDERSEN.
General Characters of the Group.
Brief diagnosis —Median anterior nasal swellings large
and abruptly projecting. Sella approximately ovate in shape.
‘Tail much shorter than tibia.
Skull—Chietly characterized by the large, almost semi-
globular, and strongly projecting median nasal swellings.
Palatal bridge rather short, between 4 and 4 the length of
Ann. & Mag. N. Hist. Ser. 7. Vol. ae 19
282 Mr. K. Andersen on the Bats
the maxillar tooth-row ; median anterior point opposite front
of m!, median posterior point level with middle of m’.
Basioccipital not unusually narrowed.
In Rh. arcuatus the premolars and molars are compara-
tively small, the temporal fossa narrow; the zygomatic
breadth of the skull, therefore, practically the same as the
mastoid breadth; the sagittal crest low, very gradually
passing into the supraorbital ridges, Rh. ewryotis shows the
other extreme: the larger teeth have caused a stronger
development of the temporal muscle, a widening out of the
temporal fossa, therefore a markedly larger zygomatic width
of the skull, a heightening of the sagittal crest, especially in
front, making the declivity of this latter towards the post-
nasal depression more abrupt *. These two extremes are
connected together by several intermediate stages.
Teeth —Throughout the whole group the dentition is
rather uniform. All the species have passed the primitive
stage: p; in the tooth-row, this premolar being invariably
external to the row, when not completely wanting. Asa
general rule, p, and p, are in contact, not rarely so strongly
so that their cingula overlap each other; but sometimes
individuals occur in which p, and p, are distinctly separated,
reminiscent of the time when p, was situated in the tooth-
row. ‘The upper p’ is small and always in row.
Nose-leaves.—Chief characters: the shape of the sella and
the connecting-process.
The sella is approximately ovate or ovate-pyriform. The
connecting-process strongly arcuate, almost semicircular in
outline, and starting from the very summit of the sella.
The internasal lobes slightly larger than usual in the genus.
The sella is peculiarly modified in Rh. inops, the connecting-
process in Rh. Creaghi.
There are two well-marked ‘‘ types” of horseshoe in this
group. Inthe more primitive species the median longitudinal
groove separating the two halves of the horseshoe in front is
narrow (linear); in the higher developed forms it is broad,
more or less pentangular in shape. ‘The former condition is
characteristic of the species inhabiting the Philippine Islands
and N.E. Borneo (arcuatus, subrufus, mops, Creaghi), the
latter of the forms distributed over Batchian, Amboina,
Timor Laut, and the Key Islands (euryotis and its local
representatives).
* For similar cranial differences in the Rh. philippinensis group, see
Ann, & Mag. Nat. Hist. for August 1905, p. 245.
of the Rhinolophus arcuatus Group. 283
Mental grooves.—No species passes beyond the primitive
stage: three mental grooves.
Wing-structure (compare the wing-indices below, on p. 287).
—It is a general rule in the genus Rhinolophus that in the
most primitive species the fourth and fifth metacarpals are
equal in length, or, if anything, the fifth a trifle shorter, and
III? less than 14 the length of III.t. All the members of the
Rh. arcuatus group exhibit the slightly more advanced stage
in which the fifth metacarpal is a little /onger than the
fourth, and III.” more than 14 the length of III.".
Tail.—The tail is extremely short, on an average but 3
the length of the lower leg.
Rhinolophus arcuatus exiguus, subsp. n.
Diagnosis.—Similar to the typical Rh. arcuatus (from
Luzon), but with narrower horseshoe and nasal swellings.
Forearm 44°8—46°2 mm.
Skull—General characters as described above (pp. 281-
282). Width of nasal swellings 5-5-2 mm., in typical arcuatus
5°6-5°7 mm.
Dentition (three skulls).—p; external (one skull, teeth
unworn), or quite wanting on one side (one, teeth slightly
worn), or on both sides (one, teeth slightly worn). p, and
ps strongly in contact, their cingula overlapping each other.
External characters.—Al\ essential points as in the typical
form, but horseshoe decidedly narrower (7:7-8°2 mm., as
against 9-10 mm.). Plagiopatagium inserted 1-2°5 mm.
above the ankle. Colour (of spirit-specimens) a shade of
dark brown, as in the dark phase of arcuatus typicus.
Type.— 2 ad. (in alcohol). Zamboanga. Collected by
A. Everett, Esq. Brit. Mus. no. 79. 5. 3. 13.
Range.—Zamboanga. Guimaras.
Rhinolophus subrufus, sp. un.
Diagnosis.—Like Rh. arcuatus, but much larger. Forearm
53°8-57 mm.
Skull—As compared with Rh. arcuatus premolars and
molars comparatively larger; temporal fossa wider, making
the zygomatic width decidedly larger than the mastoid
width ; sagittal crest somewhat higher in front.
Dentition (four skulls).—p; external; p, and p, in contact
or slightly separated.
19*
284 Mr. K. Andersen on the Bats
External characters.—In all essential points like a giant
modification of arcuatus.
Colow.—(1) Red phase. Twoskins, adult; Manila; teeth
unworn. Above and below “ cinnamon-rufous,” a little
lighter on the fore part of the back; base of hairs almost
*“ orange-rufous.”
(2) Dark phase. Oneskin, adult; Mindanao, July; teeth
practically unworn. Upperside a very dark shade of “ Prout’s
brown”; base of hairs and underside “ drab.”
Type.—Ad. (skin). Manila. Presented by J. Gould, Esq.
Brit. Mus. no. 58. 3. 29. 4.
Range.— Luzon (Manila). Mindanao.
Remarks.—This is the species described by Prof. Peters as
** Rh. rufus” *. The name rufus is preoccupied by Eydoux
and Gervais’s “ Rh. luctus, var. rufus” +.
Rhinolophus inops, sp. un.
Diagnosis —Summit of sella forming a triangular pouch.
Forearm 53°8 mm.
Skull. — The arcuatus type, but with slightly wider
temporal fossa ; sagittal crest somewhat higher in front and
a little more abruptly descending towards the postnasal
depression.
Dentition (one skull).—p; external; p, and p, in contact.
External characters—The horseshoe, the connecting-
process, and the lancet quite of the subrufus type, but
summit of sella modified into a small triangular pouch, turning
the opening downwards ; greatest depth of the pouch about
15 mm. Plagiopatagium inserted 1 mm. above the ankle.
Colour (in alcohol) a shade of dark brown.
Type.— g ad. (in alcohol). Mt. Apo, at Jodaya, Min-
danao, 4000 feet; July 8th, 1904. Collected by Dr. E. A,
Mearns (no. 5713). ‘ Bogobo name: Kohs’-set.” U.S.
Nat. Mus. no. 125314.
Remarks.—The shape of the sella, unique in the whole
genus, makes this species readily distinguishable.
* Peters, MB. Akad. Berlin, 1861, p. 710.
+ Conf. Andersen, Ann. & Mag. Nat. Hist., August 1905, p. 252.
of the Rhinolophus arcuatus Group.
285
Measurements * of Kh. arcuatus, subrufus, and inops.
Eh. arcuatus.
| Rh. subrufus. Rh.inops.
typicus exiguu Suess Type
Bed aeceeh de Ate 4skulls, | ~7P*
2 spems., 4 spems.,
2 skulls. 3 skulls.
Min. Max.| Min. Max.| Min. Max.
mm. mm.| mm. mm.| mm mm. | mm.
TTS OME 5 eos, waa sacle s+ fae ise TOG Lie 09-7 | 29 23 | 233
», greatest breadth .......... Se AAs 14 1385 798 |) 183
Nose-leaves, length ............ 15 IgS 14’) 20 | 2188
* breadth of horseshoe.) 9 10 Th 82) | coe 2:8 12:7
MELISS oe Ti shes winyttcre ety hea 458 47:3 | 448 462 | 538 57 53°38
LICL, OREN, 4) tinea deh teen eer ea 17 17°8 | 15 16-2 | 23 2a 16
Mapes ee csc t neo athe OT DO PREG OD 26°2 29:2 24-7
Skull, totallenoth 2)... e.0856: Abe 27. \20 20°8 | 25 24:2
po EO SUGGS 14 98 102] 95 23) |) Is | 1s 11
» zygomatic width ........ 9°8 9°9 | 95 OO 12 12:4 115
», Width of nasal swellings...) 56 5:7 5 oor ee (Al 65
SLUG LN NA A oe 1445 148/134 142/178 186 17-5
BRE RECWIE ees hostels aie el» 8:2 SG) 7:9 8 IOs.) 10:7 10
PPRCMCECOUD 5 «3: talevanciaalo!n do's 4s 88 92) 85 SZ Wi eg EIS). LOO
Rhinolophus euryotis timidus, subsp. nu.
Diagnosis.—Horseshoe narrower (as compared with the
Forearm 55-57°2 mm.
typical form):
Range: Batchian.
10°2-10°7 mm.
This race is characterized externally by the narrower
horseshoe and smaller
ears.
The brain-case is slenderer,
the mandible shorter, and the teeth slightly smaller than
in Rh. e. typicus and prestans. Plagiopatagium inserted
3-45 mm. above the ankle.
Colour.—Ad. skin; teeth unworn. Next to “ Prout’s
brown ” above; base of hairs ‘ drab” ; underside brownish
“drab.” Another skin (ad., teeth unworn) is essentially of
the same colour, but more tinged with “ Mars brown”
above and with “ wood-brown”’ below.
Type.— @ ad. (in alcohol). Batchian. Collected by
Dr. A. R. Wallace. Brit. Mus. no. 60. 1. 10. 5.
* For explanation of measurements, see Ann. & Mag. Nat. Hist,
August 1905, p. 248, footnote.
286 Mr. K. Andersen on the Bats
Rhinolophus euryotis, Temm., typicus.
Diagnosis —Horseshoe broad: about 12 mm.
about 57 mm. Range: Amboina.
The Amboina race of Rh. euryotis differs from the Batchian
form by the broader brain-case, the longer mandible, longer
tooth-rows, broader horseshoe, and larger ears; from the
Key Island race by the smaller and slenderer skull, narrower
nasal swellings, shorter mandible and tooth-rows, and
narrower horseshoe. Plagiopatagium inserted 2-5 mm.
above the ankle.
Forearm
Rhinolophus euryotis prestans, subsp. n.
Diagnosis.—Horseshoe very broad: about 13 mm. Fore-
arm 58mm, Range: Key Islands.
This is the extreme in the size of the horseshoe and the
width of the skull and nasal swellings. Plagiopatagium
inserted very nearly on the ankle.
Type.— $ ad. (in alcohol). Key Islands.
Brit. Mus. no. 99. 12. 4, 4.
Purchased.
Measurements of Rh. euryotis and Subspecies.
Lh. euryotis.
timidus. Sf: prestans.
3 spems., Sf oe a ¢ ad.
3 skulls. : Type.
Min. Max
; mm. mm. | mm. mm.
Bars, lene ths ors ae sie eo temo 20°5 22°8 23
9» greatest breadth.......... 17 19 18
Nose-leav es Mencthe nace «teow ths 17 17°5 19
bs breadth of horseshoe.| 10-2 10-7 118 13
Harearmy. v4. ists bons! eile eis eras 55 O72 568 58
LOIRE, AEP ae ees reer rere? | 18°5 20 215
MEO CE LOD fis cuss cin ni antie sane clecle 6 257 278 28 28
Skull fotallength: - 4. . 2.00 feies 25°1 25°4
ye Mdastoid width 2... ssa Ae Las 12
»» Width of brain-case ...... 104 104 10°8 1
» zygomatic width ........ IVS. 12-24) © 12:2 12°8
oy ig dunxallar width... «im. 2s] 8:9 9 8:7 9°8
width of nasal swellings . 6°8 6°9 68 72
Mandibloget \act® ie .pc le cae | digedin ic 18 18
Uppertceth ee wemcerees a. e | Ov 9°7 10:2 10:2
Lower teeth fis sse oe neces tl 103 10% at 1l
of the Rhinolophus arcuatus Group. 287
Wing-indices.
| |
— eon Rh. Creaghi. Eth. inops. | Rh. euryotis, |
9 ae a 1 spem. lspem. | 6spcms.
Worearm.. .\.... . 1000 1000 1000 | 1000
drd metacarpal. 706 [ee O7 703 702 |
MS SS cet oe 28 286 281 292 289
U0 Ee eee 458 454 481 | 436
4th metacarpal. 728 717 725 | 721
1 ASS Cee 197 211 199 196
LSet este 270 263 | 279 265
5th metacarpal. 742 741 740 | 734.
Wir eal ctsc fe 221 217 205 | 226
Mae dst a3ds O84, i) oN awk? | 240 | 233
General Remarks on the Group.
All the species known of the Rh. arcuatus group are
so far developed as to have the median anterior nasal
swellings more abruptly projecting than usual in the
genus, the middle lower premolar (p;) situated external to
the tooth-row or quite obliterated, the fifth metacarpal
slightly longer than the fourth, and the second phalanx of
the third finger lengthened. All these are secondary
characters. We lack the direct information as to the
affinities of the group, which could have been derived from
species on a more primitive level of development. Never-
theless, there can scarcely be any doubt that ¢he arcuatus
group is more closely related to Rh. philippinensis and
Rh. macrotis than to any other now existing types of the
genus. The strong development of the nose-leaves, the size
of the internasal lobes, the shape of the connecting-process
and the lancet, the broad ears, are characters which seem to
point in that direction.
Rh. arcuatus is unquestionably the most primitive species
of the group, as proved by the slightly less projecting nasal
swellings, the small teeth, the narrow temporal fossa, the
very low sagittal crest, the normal shape of the horseshoe,
the somewhat less pronounced “ovate” shape of the sella.
Of the two local forms here discriminated the southern
(Rh. e. exiguus) is a little higher developed in dentition:
ps is very often lost; the narrow horseshoe is probably a
secondary character.
Rh. subrufus is a modification of the arcuatus type: more
288 On the Bats of the Rhinolophus arcuatus Group.
projecting nasal swellings, larger premolars and molars,
therefore wider temporal fossa and higher sagittal crest;
considerably larger size. There is a certain tendency in the
genus Rhinolophus to a development of a large and a small
representative of quite the same “type”; the difference in
size is so well marked and without any intergradation that
we cannot but recognize them as distinct “ species,”’ although
they sometimes, as in the present case, inhabit the very same
island; thus Rh. subrufus (Luzon) bears very much the
same relation to Rh. arcuatus (Luzon) as Rh. acuminatus
(Java) to Rh. minor (Java), Rh. Rouxi (Himalayas) to
Rh. borneensis, Rh. megaphyllus (Australia) to Rh. simplex
(Lombok).
Rh. Creaghi*, from the extreme north-east of British N.
Borneo (Sandakan), is very closely related to the Philippine
Rh. arcuatus: the same type of skull, if anything a trifle
more advanced in the size of the temporal fossa, but the
difference is extremely small indeed; the same shape and
size of the nasal swellings; the dentition on quite the same
stage as in arcuatus exiguus (p3; rudimentary or wanting) ;
the same shape of the horseshoe, the sella, and the lancet,
but more rounded internasal lobes ; the broad ears charac-
teristic of the group ; the three mental grooves ; the same
wing-structure ; the short tail. But it is unique in the
following points :—The connecting-process is obliterated ;
the median portion of the posterior nose-leaf is peculiarly
inflated and provided with a conical tuft of coarse hairs ;
when pressed backwards the sella leans against this hair-tuft,
which therefore practically replaces the missing connecting-
process.
Rh. inops is in every respect, cranial and external
(apart from the size), very much hke Rh. arcuatus, but
highly peculiar in the shape of the summit of the sella. The
mental grooves do not reach the free border of the lower lip;
it would seem to indicate a beginning obliteration of the
grooves.
The essential cranial character of the group, viz. the
abruptly projecting nasal swellings, culminates in the large
and broad-skulled Rh. euryotis, im which the front part of
the horseshoe is slightly modified. The species reviewed
above are confined to the Philippimes and N.E. Borneo,
Rh. euryotis to the islands between Celebes and New Guinea.
The Rh. arcuatus group, as here defined, has no represen-
tatives in the Ethiopian Region.
* Thomas, Ann. & Mag. Nat. Hist. (6) xviii. (1896) p. 244.
On the Bats of the Rhinolophus macrotis Group. 289
XX XI.—On the Bats of the Rhinolophus macrotis Group, with
Descriptions of Two new Forms. By Knup ANDERSEN.
Rhinolophus hirsutus, sp. n.
Diagnosis —Allied to Rh. macrotis, but with considerably
larger ears and longer tail. Forearm 44°77 mm. Had. Cen-
tral Philippine Islands (Guimaras).
Length of ears 24, greatest breadth of ears about 18 mm. ;
in two adult Rh. macrotis from Masuri the measurements
are 19-20°8 (length) and 15°5-16°5 mm. (breadth). Tail
equal to 14 the length of the lower leg (24 as against 19 mm.) ;
in Rh. macrotis scarcely equal to the lower leg (17 as against
18°8 mm.).
The skull is slightly larger than in Rh. macrotis, but quite
of the same shape. p; half in row; p? in row, with a
distinct, pointed cusp.
Type.— 2 ad. (in alcohol). Guimaras, Central Philippine
Islands. Collected by J. B. Steere, Esq. U.S. Nat. Mus.
no. 105487.
Rhinolophus Pearsoni chinensis, subsp. n.
Diagnosis.—Similar to Rh. Pearsoni from Darjeeling and
Masuri*, but with markedly shorter tibia, slightly smaller
skull, narrower maxillar width, shorter mandible and tooth-
rows. Forearm 52:'7 mm. Hab. Fokien (China).
Length of lower leg 26 mm. (in the typical form 29) ;
maxillar width 9°2 (9:7—-9°8) ; length of mandible 16°8 (17°7-
17°9); maxillar tooth-row 9:5 (9°8-10°2); mandibular
tooth-row, exclusive of incisors, 10°3 (10°8-11°1). Dentition
as in the typical form: p; external; p, and py, almost in
contact ; p? in the tooth-row.
Type.— ad. (skin). Kuatun, Fokien, April 16th, 1898.
Collected and presented by J. De La Touche, Esq. Brit.
Miss no 98s Ia. -2.
* To judge from descriptions and figures of Dobson's Rh. yunanensis,
from Hotha, Yunan (J. A. S. B. xli. pt. 2 (1872), p. 386; and Yunan
Exy. i. (1879) p. 95, pl. iv. fig. 1), and A. Milne-Edwards’s RA. larvatus,
fron. Moupin, Szetchuan (Mamm. Tibet (1872), p. 248, pl. xxxvii. fig. 1,
pl. xxxvii.c. fig. 1), these bats are indistinguishable from the typical
Ith, Pearsont. They are, at all events, different from the Chinese form
described above.
290 Mr. K. Andersen on the Bats
Wing-indices.
| : Lh. ethiops,
| a toe big Rh. Pearsoni. | Rh. HH. ildebrandti,
| 4 spems. ‘| 4 spems. Leh. fumigatus.
25 spems.
IBOTOMEE yee eater ao dsie ee 1000 1000 1000
ord finger, metacarpal . . 703 656 692
5 Ist phalanx. .| 304 333 806
“1 2nd phalanx || 371 452 537
4th finger, metacarpal . . 736 715 726
ra 1st phalanx. .) 215 213 191
,: 2nd phalanx , 245 296 349
5th finger, metacarpal . . | 754 739 742
s Ist phalanx. .| 245 239 238
rs 2nd phalanx . 243 327 | 269
General Remarks on the Rh. macrotis Group.
The skull of RA. macrotis is of the general shape charac-
teristic of all the more primitive species of the genus: very
narrow temporal fossa and low sagittal crest ; nasal swellings
not especially modified, the median anterior swellings but a
trifle larger than usual; basioccipital not more strongly
narrowed than in the majority of species. The palatal bridge
is remarkably long, equal to 4 or # of the maxillar tooth-row.
Dentition primitive: upper canine and p* widely separated ;
p” small (as usual), but with a distinct cusp.
The essential external characters are the long and broad,
almost parallel-margined (tongue-shaped) sella; the long and
dense hairing on the front face of the sella; the very slightly
projecting, rounded connecting-process, starting from a point
below the summit of the sella; the cuneate lancet ; the broad
horseshoe; the three mental grooves; the broad ears; the
quite primitive wing-structure: fifth metacarpal a trifle
shorter than the fourth, III.? (as well as the other distal
phalanges) very short.
Among the now existing types of the genus Rh. macrotis
has no closer relatives than the Rh. philippinensis group.
We find in this group an unusually long palatal bridge ; the
same tendency to enlargement of the nose-leaves ; the shane
of the sella (apart from the lateral expansions, of course)
strongly recalls that of macrotis ; the connecting-process is
broadly rounded and starting from a point below the sunimit
of the sella, as in macrotis ; the lancet long and cuneate ;
the ears very much of the macrotis type ; the mental grooves
of the Rhinolophus macrotis Group. 291
and wing-structure on practically the same stage as in
macrotis. But philippinensis has retained a somewhat more
primitive dentition (p”? unusually well developed), and the
base of the central nose-leaf is peculiarly modified.
Rh. hirsutus is but a Philippine representative of the
macrotis type; the ears are still larger, the tail longer.
Rh. Pearsoni is closely related to Rh. macrotis, but in
several respects considerably more highly developed. The
temporal fossa is decidedly wider ; the sagittal crest higher ;
the palatal bridge shortened. The shape of the nasal
. swellings very much as in macrotis. The only progress in
dentition is a slight reduction of p?; also the cusp is less
distinct ; but the tooth is still completely m row. The
lateral borders of the sella are peculiarly crenulate ; the ears
smaller; the lateral mental grooves quite obliterated. The
third metacarpalis much shortened, the fourth less so, making
the fifth metacarpal decidedly the longest of all; all the
terminal phalanges are much lengthened. On the whole
this wing-structure is rather similar to that of Rh. ferrum-
equinum, a species with which RA. Pearsoni has no closer
affinities.
Rh. ethiops (Angola), Hildebrandti (Mazoe to Uganda),
and fumigatus (Kitui to Abyssinia) are Ethiopian represen-
tatives of Rh. macrotis. There is a most striking resemblance
in the nose-leaves ; the horseshoe, the sella, the connecting-
process, the lancet, differ in nothing but the larger size;
the long and dense hairing on the front face of the sella,
characteristic of macrotis, is retained in the African species ;
the shape of the ears is the same. As in Rh. Pearsoni, the
mental grooves are reduced to one. The wing-structure
differs from that of Rh. macrotis in having the fifth meta-
carpal somewhat longer than the fourth and third (by the
combined effect of a slight shortening of these latter and a
slight lengthening of the former), and the second phalanx of
the third and fourth fingers much lengthened. The modifi-
cation of the skull noticed above when passing from Rh.
macrotis to Rh, Pearson is found again in the Ethiopian
species : wide temporal fossa, more projecting sagittal crest,
shortening of the palatal bridge. In front-of the anterior
nasal swellings is developed a very narrow thin brim of bone,
forming the upper border of the nasal openings (as in the
more differentiated members of the philippinensis group).
The dentition is extremely highly developed: p; and p*
pushed out to the external side of the tooth-row, or com-
pletely lost.
As will be observed from the above, the Ethiopian species
292 Mr. T. D. A. Cockerell—Deseriptions and
here under consideration are on a higher level of development
than the Oriental RA. macrotis : in several important cranial
characters, in the dentition, in the wing-structure. I have
pointed out elsewhere that also the other Ethiopian Rhino-
lophi, without any exception, have their more primitive rela-
tions in the Oriental Region. Thus the general conclusion
arrived at may now be briefly formulated as follows :—A//
the Ethiopian representatives of the genus Rhinolophus are of
Oriental origin.
- The subjoimed diagram gives a view of the probable inter-
relations of the species and groups reviewed above :—
@thiops and allies f.
Pearsoni.
a
7A
arcuatus group fF.
———
=
macrotis and hirsutas.
philippinensis group *.
XX XII.—Descriptions and Records of Bees.—II.
By T. D. A. Cocxzrztt, University of Colorado.
Euryglossa ephippiata, Smith.
Queensland, no. 94. 61, also labelled Ridg. 11.93; 701. 2.
Except for the smaller size, it has a strong superficial
resemblance to Callomelitta picta. The character of the
* Ann. & Mag. Nat. Hist., August 1905, p. 243.
+ Supra, p. 281.
{ Op. cit. (7) xv. (1905) pp. 75-76 (“ Remarks ”).
Records of Bees. 293
relative lengths of the submarginal cells, given by authors to
separate Euryglossa from Prosopis, is hardly distinctive ;
much better is the rounded head of Euryglossa, with the
broad face lacking the characteristic markings of Prosopis.
Euryglossa rubricata, Smith.
Queensland, no. 94. 61, also labelled Ridg. 11.93 ; 859. ¢.
Close to E. ephippiata, but easily separated by the red
antenn (even the scape red), the black axillz, the light
testaceous tegule, the red tibice and tarsi (anterior tibize
with a large black spot), and the mainly dull red abdomen.
Euryglossa subsericea, sp. n.
2? Length between 7 and 8 mm.
Head and thorax shining black ; abdomen very dark blue
with a satiny gloss, hind margins of segments narrowly
black. Mandibles black, with a dark red stain in the middle ;
clypeus with scattered rather feeble punctures; front dullish,
densely striato-punctate; facial foveze linear, as in LH. ephip-
piata; antenne dark, flagellum brownish beneath, the extreme
tip reddened ; mesothorax with distinct but scattered punc-
tures, dense at the sides, the three longitudinal grooves
distinct ; tegule piceous. Wings perfectly hyaline ; stigma
dilute brown, nervures paler ; venation as in FE. ephippiata
(in both the first submarginal cell is very much longer than
second, though the second is very much broader than high).
Enclosure of metathorax smooth and shining; sides of meta-
thorax with scanty whitish hair; legs black, with white
hair; hind spur with large saw-like teeth; scanty hair at
sides of abdomen white; apical fimbria black.
Hab. Queensland, no. 94. 61, also labelled Ridg. 11.93;
702.
Close to EL. depressa, Smith, but smaller, the clypeus less
strongly punctured, the facial foveze not broad. The vertex
has a few whitish hairs (a fringe of pale yellow hairs in
depressa), and the hair on the inner side of the tarsi is
very pale. In E. subsericea, rubricata, and ephippiata the
basal nervure is strongly curved, and falls considerably
short of the transverso-medial.
Euryglossa calliopsiformis, sp. 1.
? .—Length a little over 6 mm.
Black ; head, thorax, and legs marked with bright yellow,
in the manner of a Calliopsis ; face broad, with the clypeus
(except four small black marks, one on each side and two on
294 Mr. T. D. A. Cockerell— Descriptions and
the disk), a broad and low supraclypeal mark, a large suboval
mark on front, and broad band-like lateral face-marks ending
abruptly about the level of the middle of the scape, all bright
chrome-yellow ; the moderately broad and rather short facial
fovez begin where the lateral marks leave off; front granular,
dull ; clypeus shining, sparsely punctured; labrum and
mandibles (except the dark ferruginous tips, which are
bidentate) pale yellowish; posterior orbital margins very
broadly yellow; antennze dark, flagellum pale yellowish-
ferruginous beneath; upper margin of prothorax with a
yellow band (interrupted in middle), continued to tubercles,
which, with a patch below, are yellow; mesothorax dull,
gibbous, with a pair of fine yellowish streaks on the disk and
a yellowish line over each tegula; scutellum with the posterior
margin and a triangular spot on each side yellow; post-
scutellum with a median yellow spot; pleura black, with a
broad transverse yellow band beneath the wings; region
between wings and hind legs pale yellowish; the four
posterior trochanters, the hind femora, a patch on middle
femora behind, the anterior and middle knees, the anterior
and middle tibize in front and a streak behind, all yellow;
anterior and middle tarsi reddish; tegule reddish brown,
with a yellow spot. Wings clear; stigma large, it and the
nervures dilute sepia; second recurrent nervure joining
second submarginal cell a short distance before its end, first
joining the cell slightly beyond the end of its first third ;
basal nervure very strongly bent. Abdomen broad, with
a dull satiny surface, very dark brown, with a whitish
mark on each side of the first segment near the base, a
large lanceolate whitish mark at lateral bases of segments
2 to 4, and a spot in the same region on 5; no hair-bands,
but the apex and venter with a thin pale pilosity ; venter
dark ; hind spur serrate.
Hab. Queensland, no. 94. 61, also labelled Ridg. 11.93 ;
864. One ?.
Allied to E. quadrimaculata, Smith ; the two species form
a group standing apart from typical Huryglossa.
Callomelitta picta, Smith.
Victoria, no. 83. 100; and Franklin, Tasmania, nos. 91-155
and 3468.
The Victoria specimen has a red spot on the middle of the
postscutellum.
Allodape simillima, Smith.
Townsville, Queensland, 5.3.03 (I. P. Dodd); Cossack,
Records of Bees. 295
W.A., no. 91-155; Mor. [=Moreton?] Bay, 1893 (W. W.
Frog oggatt) ; Queensland, no. 319; Queensland (H. Saunders).
Allodape unicolor, Smith.
Queensland, no. 4312; Baudin I., no. 91-82; Montalivet
I., N.W. Australia, no. 90-126.
The last-mentioned specimen differs in having the second
recurrent nervure joining the second submarginal cell almost
at its tip, but it does not appear to be a distinct species.
Anthophora eruginosa, Smith.
Of this wonderful species I have before me a male from
De Freycinet I., N.W. Australia (no. 90-126), and females
from Queensland (H. Saunders, F. P. Dodd), Adelaide River,
and Port Darwin. ‘The following particulars should be added
to Smith’s description :—
? .—Green hair of vertex and above strongly mixed with
black ; pleura with light bluish-green hair ; black on clypeus
more extensive than Smith’s description suggests.
3 .—Clypeus yellow, with the upper lateral margins broadly
black.
Anthophora concinna (Klug).
Queenstown, Cape Colony (ZH. 7. Wells). 2. In British
Museum.
The second s.m. is very broad below. Superficially the
insect looks like a fulvous edition of A. eruginosa.
Lithurgus rubricatus, Smith.
Queensland (EZ. Saunders) ; nos. 93-49 and 324. ?.
Lithurgus atratiformis, sp. n.
?.—Length about 14 mm.
Black, without the distinct hair-bands or red apex of
abdomen seen in L. rubricatus. Head large and round; hair
at sides of face dull white, of region between antenne and
cheeks mixed black and white, of vertex and occiput black,
of anterior middie of clypeus long and reddish, of mandibles
(which are short and obtusely tridentate) reddish ; frontal
tubercle rather prominent, wrinkled and shining, not bifid,
its form much as in L. rubricatus ; vertex as densely punc-
tured as possible; antenne dark, flagellum faintly brownish
beneath ; hair of prothorax and anterior margin of meso-
thorax dull white, of pleura black, of sides of metathorax
296 Mr. T. D. A. Cockerell—Deseriptions and
partly black and partly dull white ; mesothorax anteriorly
very coarsely tuberculate, its posterior third, and the scu-
tellum, dull and densely rugoso-punctate; tegule dark
reddish brown. Wings smoky, nervures black; second re-
current nervure almost meeting second transverso-cubital.
Legs black, with black hair, hind tibiz coarsely tuberculate
on outer side; a small pulvillus present. Abdomen shining
black; the first two segments sericeous and scarcely punc-
tured, the others with minute close punctures ; hair-bands
white, but reduced to hardly visible rudiments, the most
distinct appearing as a fine line on the hind margin of
segment 5; hair of apical segment black, the extreme apex
dark ferruginous ; ventral scopa dark, appearing reddish in
sume lights, a sort of purplish black or dark brown in others ;
venter, underneath the scopa, with three rather narrow clean-
eut white hair-bands.
Hab. N.W. Coast of Australia, no. 69-59.
Allied to the Indian ZL. atratus, Smith, but larger and
more rugose.
Saropoda bombiformis, Smith.
Toowoomba, no. 93.189; 2. Queensland, “ Ridg. 2.92;
691,” and 94.61. ¢@.
The hair of the head and thorax above is strongly mixed
with black in both sexes, but this is not apparent unless the
insect is examined from the side.
Stilpnosoma levigatum, Smith.
Queensland, no. 94. 61, “ Ridgw. 11.93; 480.7 29.
Facial foveze linear, rather short. Thorax with median
and lateral linear grooves. Abdomen varying to dark
purple-blue. The imsect is not lke a Prosopis, but very
closely resembles Euryglossa, from which it appears to be an
offshoot. The venation is like that of Euryglossa, except
that the second recurrent nervure joins the second sub-
marginal cell a considerable distance from its end, whereas
in Euryglossa it joims it at the end.
Parasphecodes talchius, Smith.
Victoria, Australia, no. 85.108. ¢.
The first r. n. meets second t.-c. ; antenne entirely black.
Parasphecodes Froggatti, sp. n.
g —Length about 95 mm.
Head and thorax black, covered with white lanose pubes-
cence; face narrow; lower half of clypeus (except brownish
ltecords of Bees. 297
edge) pale yellow, this sending a pointed projection upwards
in the middle; mandibles dark, reddish in the middle;
antenne broken at fourth joint in type, but apparently all
dark except a small reddish spot at extreme base of flagellum
beneath ; thorax dull, minutely roughened; base of meta-
thorax irregularly ruguloso-plicate, with a sharp rim ;
tegul rufo-fulvous. Wi ings slightly dusky, a little yellowish,
not darkened apically ; nervures and stigma sepia-brown ;
first r.n. meeting second t.-c. Legs black, the anterior and
middle knees, the anterior tibiz broadly in front, and the
anterior and middle tarsi more or less, ferruginous ; abdomen
deep but bright chestnut-red, broad like that of a female
(much broader than in P. talchius) very closely and minutely
punctured, a suffused T-like area on first segment, a suffused
median area on third, and all of fourth and following ones
black.
Hab. Bathurst, N.S.W., Jan. 18, 18938, no. 124
(Froggatt).
Very distinct by its great hairiness and stout abdomen.
Nearest, I think, to P. altichus. The second r. n. and third
t.-c. are strong, not at all weakened as in P. tilachus.
Parasphecodes lichatus, Smith.
Perth, W. Australia ; 93-198. @.
This and the next three species are superficially very
similar; they may be distinguished as follows :—
Abdomen smooth and shining, hind legs clear red... adelaide, Ckll.
Abdomen not so shining, hind legs not red ........ R
1. Vertex and mesothorax t above with much black hair ;
AMA SHIGE VCTYIGRTIC oo ta were ge ape clin avn sty * » lichatus, Sin.
Wertex and mesothorax not 80 5 ...-..,0eccneres 2.
2. Abdomen black beyond third segment............ Loweri, Ckll.
OREUMPSEC TOO TEC <0 ge tanie! a) 4.02 Ss Reteie woh hchaiec alle Tepperi, Ckil,
Parasphecodes adelaide, sp. n.
2? .—Length about 9 mm.
Head and thorax black, with thin whitish pubescence,
dense and ochreous on margin of tubercles ; head large and
broad; clypeus very sparsely punctured, depressed in the
middle, and with a reddish spot; middle of mandibles
reddish ; flagellum dark reddish beneath ; mesothorax
shining, with very minute punctures of different sizes; area
of metathorax slightly irregularly wrinkled, but not con-
spicuously sculptured, its rim rather weak; tegule shining
dark reddish. Wings faintly dusky, not clouded apically ;
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 20
298 Mr. T. D. A. Cockerell—Descriptions and
nervures and stigma reddish; first r.n. joming second s.m.
almost at its end; second s.m. not far from square; second
r. nu. and third t.-c. much weakened. Anterior and middle
legs dark red, hind legs clear red, hair of legs tinged with
yellowish ; hind spur simple ; abdomen convex, shining, dark
chestnut-red strongly suffused with black, from the middle of
the third segment to the apex practically black, except the
coppery hind margins ; surtace of abdomen not punctate.
Hab. Adelaide ; 54-55.
Tn my table of Parasphecodes (Ann. & Mag. Nat. Hist., Sept.
1904, p. 209) this runs to P. tuchilas and P. lichatus, but it
is distinct by the very dark and shining abdomen, wherein,
as also in the weakened outer nervures, it resembles rather
P. tilachus.
Parasphecodes Loweri, sp. 0.
9 —Length about 9} mm.
Robust ; head and thorax black, rather pubescent, the hair
white beneath, yellowish cinereous above, forming a little
dense light patch on the anterior middle of the scutellum ;
head broad, but facial quadrangle longer than broad ; clypeus
shining, with numerous large punctures ; mandibles entirely
black; flagellum faintly reddish beneath; front, vertex,
mesothorax, and scutellum minutely roughened, glistening ;
area of metathorax scarcely defined, with irregular longi-
tudinal wrinkles only on the basal part; upper corners of
posterior truncation rounded ; tegule rather large, dull dark
reddish, the anterior margins narrowly hyaline. Wings with
a rather strong yellowish tinge; nervures and stigma ferru-
ginous ; first r. n. meeting second t.-c., but a little on the
inner side; second s.m. higher than broad, somewhat
narrowed above. Legs dark reddish brown, with abundant
pale pubescence; basal joint of hind tarsi with an apical
fulvous tuft; hind spur simple. Abdomen broad, thinly
pubescent, very minutely and closely punctured; first three
segments chestnut-red, the others black, first segment with a
blackish anchor-shaped mark; hind margins of first three
segments orange. The second r. n. and third t.-c. are
weakened.
Hab. Adelaide, Australia; 52. 124.
Differs from P. tuchilas by the hind margins of abdominal
segments 1 and 2 not darkened, clypeus with numerous
punctures, &c. It approaches P. melbournensis in haying the
enclosure of metathorax diversely sculptured, but differs in
other particulars.
Records of Bees. 299
Parasphecodes Tepperi, sp. ni.
? —Length about 9 mm.
Not so robust as the last species, the abdomen being
noticeably narrower. The followmg differences are also
apparent on comparison :—Head shorter ; clypeus with the
punctures hardly so strong; flagellum pallid beneath except
at base; tegule light rufo-fulvous; scutellum without a
light patch ; area of metathorax rather coarsely longitudi-
nally wrinkled all over; stigma and nervures dark reddish ;
second s.m. nearly square; abdomen rather more shiny,
entirely red, the apex faintly blackish; venter also red ; legs
black.
Hab. Adelaide, Australia; 53.56. Another is marked
simply “ Australia.”
Runs in my table to P. lacthius, Sm., but that has
rufo-testaceous legs and the fourth abdominal segment
fuscous. The names of this and the last species are given
in recognition of the work of two well-known Australian
entomologists.
Sphecodes tasmanie, sp. un.
$ .—Length about 5} mm.
Head and thorax black; head broad, facial quadrangle
broader than long ; face with yellowish pubescence ; mandi-
bles simple, black basally, pale yellowish in the middle,
ferruginous at tips; front and vertex dull, minutely
roughened ; antennz long, entirely black, scape long, joint 3
a trifle longer than 4, 5 about as long as 3 and 4 together,
flagellar joints very distinct; mesothorax and scutellum nude
except for a few pale fulvous hairs (mainly on scutellum),
smooth and shining, with scattered inconspicuous very
minute punctures, only visible with a good lens; sides of
thorax with a very thin griseous or whitish pubescence ; area
of metathorax large, semilunar, dull, with extremely fine
thread-like longitudinal striz ; tegule shining reddish,
marked with darker. Wings clear, strongly iridescent ;
stigma large, it and the nervures rufous ; first sm. much
longer than the other two united; second very narrow, much
higher than broad, narrowed above, receiving the first r.n. at
its middle ; third higher than broad, but larger than second ;
second r.n. and third t.-c. weakened, and lower boundary of
third s.m. almost obsolete. Legs bright ferruginous, the
cox, trochanters, and basal half or more of femora black ;
outer edge of hind tibiz and the last two joints of their tarsi
darkened ; abdomen shining, quite without hair-bands, black
20*
300 Mr. T. D. A. Cockerell—Deseriptions and
basally and apically, with the apical margin of first segment,
the second except a median stain, and the greater part of the
third chestnut-red ; apical plate red, broadly truncate, much
broader than long, suggestive of ‘the genus Proteranea ;
venter red, stained with blackish.
Hab. Hobart, Tasmania, no. 91-155.
Because of the smooth mesothorax, this cannot be the
male of S. antipodes, Smith.
The following three species of Halictus resemble Para-
sphecodes and Sphecodes in having red abdomens :—
Halictus punctatus, Smith, var. exlautus, nov.
@.—Abdomen broad, entirely orange-fulvous (apricot-
colour), except a faint dusky median stain and a small black
spot on extreme side of segments 38 and 4. Mandibles
ferruginous ; facial quadrangle about square ; mesothorax
green, dullish, with large well-separated punctures, median
and lateral grooves distinct ; metathorax black, the enclosure
large but ill-defined, covered with fine raised lines ; ; tubercles
reddish ; pleura black ; stigma very large, honey-colour ; ;
first r. n. joing second s.m. not far from its end ; outer
nervures weak as in Chloralictus ; legs red, hind spur simple.
Length about or not quite 6 mm.
Hab. Australia (no other details known), from F. Smith’s
collection (no. 79. 22).
The typical punctatus has the abdomen dark rufo-piceous
in the middle, red at the sides.
Halictus sphecodopsis, sp. 0.
$ —Very minute; length scarcely 4 mm.
Head and thorax yellowish green, the metathorax a bluer
green; face broad, covered with yellowish hair; broad
anterior margin of clypeus, labrum, and mandibles (except
ferruginous tips) cream-colour; antennze long, lively ferru-
ginous, the flagellum dusky above ; vertex and mesothorax
granular, the latter with a little yellowish pubescence, most
abundant at the anterior corners; area of metathorax
obscurely roughened and wrinkled basally, bounded by a
shining margin; tegule shining fulvous. Wings rather
narrow, clear, strongly iridescent; stigma dark reddish brown,
nervures pale ; first r. n. joiming second s.m. near its end;
outer nervures (as in Chloralictus) much weakened, in fact
almost obsolete. Legs entirely clear orange-ferruginous.
Records of Bees. 301
Abdomen small, fusiform, very shiny, bright ferruginous, the
apical third strongly infuscated ; apex broadly rounded; no
hair-bands; suture between first and second segments a
little depressed.
Hab. Queensland: no. 94.61; “ Ridg. 11.91 ; 710.”
Close to H. sphecodoides, Smith, but the colour of the head
and legs is different, and I do not feel justified in treating it
as the male of that insect, hitherto known only in the female.
Halictus vitripennis, Smith.
A @ from Queensland (Ridg. 11.93; 711) agrees with
Smith’s too brief description and my notes on the type; but
there is just a possibility that actual comparison would reveal
differences. _
XX XIII.—Descriptions and Records of Bees. —II1.
By TT. D. A. Cocxrrett, University of Colorado.
PHENACOLLETES, gen. nov.
Tongue and paraglosse as in Colletes; labial palpi small,
4-jointed, the joimts subequal, but the two middle ones
the shortest ; distance between paraglosse and labial palpi
greater than length of former; blade of maxilla about 23
times as long as wide, rounded at end; maxillary palpi very
small, not half the length of maxilla-blade, -the six joints
stout, the first not easily seen, the second and third cylin-
drical, the fourth and fifth short, the sixth narrow and
pointed ; pubescence short, very finely plumose ; abdomen
free from hair, except a very fine pruinosity, and shaped like
that of a Tachytes or Bembecid ; stigma small and narrow
(about as in Leioproctus as figured by Smith) ; marginal
cell lanceolate, tapering apically ; three submarginal cells—
the first, on cubital nervure, at least as long as the other two
together; the second large, almost square, receiving the first
recurrent nervure at the end of its first third; the third
about twice as long as the second below, but only as long
above, its outer margin regularly curved; the second
recurrent nervure enters it a little before the beginning of
its last fourth, and is practically straight, with only a slight
irregular curvature ; basal nervure falling a very little basad
of transverso-medial.
802 Mr. T. D. A. Cockerell—Z: scriptions and
Phenacolletes mimus, sp. 1.
3 .—Length about 14 mm.; of anterior wing just over 10.
Black, with the large scutellum red varying to black;
head large, circular seen from in front; cheeks broad ;
vertex convex; ocelli well developed, in a low triangle ;
mandibles dark reddish, bidentate, but the inner tooth quite
small ; labrum with a shining elevated C placed with the back
upwards, rather suggestive of a hog’s snout ; face broad, with
the eyes practically parallel (slightly diverging below); the
whole face, up to the ocelli, covered with dense snow-white
pubescence, as also are the cheeks and occiput ; interocellar
region with a tuft of white hair; vertex and mesothorax
exposed, finely granular ; antenne short, entirely dark except
that the apical margin of the scape is reddish, or the scape
may be entirely reddish ; flagellum thick, the middle joints
broader than long, and strictly cylindrical; tubercles more or
less red; pleura, postscutellum, and metathorax with much
white hair; metathorax with a thick white fringe on each
side; its basal enclosure large, shining, and hairless, with a deep
basal pit, and bounded by a strong sulcus ; mesothorax with
three impressed lines in front, and the parapsidal lines fairly
distinct ; tegule yellowish ferruginmous. Wings variably
dusky, inclined to be yellowish; stigma and nervures dark
brown, or the stigma practically black. Legs normal, dark
reddish, with fine white pruinosity, only the anterior femora
having behind a fringe of white hair; hind knee-plates well
developed ; hind tibize rather slender and nearly cylindrical,
the hind legs being quite long ; anterior tibial spurs with a fine
comb, hind spurs simple ; abdomen rather shiny, with very
minute punctures, guite without pubescence except a fine white
pruinosity, especially noticeable at sides ; sutures not depressed ;
seventh dorsal segment with a broad flattened pygidial area;
from the apex projects a small rounded plate. The characters
italicized are likely to be of generic significance, in addition
to those given in the generic diagnosis.
Hab. Turtle Bay, 91-82; three specimens, numbered
1421, 1423, and 1424. Turtle Bay is on the coast of
Australia; I believe off the N.W. coast. The specimens
were collected by Commander J. J. Walker, R.N., on the
‘Penguin’ Expedition.
This remarkable insect caused me much perplexity. The
general appearance and structure of the abdomen resemble
those of some wasp allied to Tachytes ; the legs also are very
wasp-like. On the other hand, the venation is quite that of
a bee, and the mouth-parts are of the Colletid type, which,
Records of Bees. 303
however, is nearly the same as that of Tachytes and its
allies. With a lens, I could not detect any plumosity in the
pubescence, which is very short and like that of a wasp; but
the compound microscope shows that all the hairs are
minutely plumose, and this settles the reference of the genus
to the bees. It is by all means the most wasp-like bee I
know, and I think it affords very strong evidence in favour
of the origin of the Colletid bees from a wasp of the Tachytes-
group. It appears to follow that the bees with pointed
tongues have had a quite different origin, as I have formerly
suggested.
Lithurgus atratiformis, Ckll.
? —A specimen marked 92. 16 and 304 shows that when
quite fresh the second to fifth segments of the abdomen
have extremely narrow but conspicuous white hair-fringes.
Another, with the face narrower than the type, but clearly
the same species, is from Queensland (Gilbert Turner),
labelled also 304.
Bombus terrestris (L.).
New Zealand (H. Saunders); one worker, in British
Museum.
Bombus terrestris, var. audax (Harris).
New Zealand (#. Saunders); one female, in British
Museum.
Bombus hortorum, vay. fidens (Harris).
New Zealand (£. Saunders) ; three females and a worker,
in British Museum. There is also a female labelled
“ Queensland (H. Saunders).”
Bombus hortorum, var.
Similar to var. jfidens, but prothorax, scutellum, and
penultimate abdominal segment more or less distinctly
reddish yellow. I have not found a varietal name for this
form.
New Zealand (HE. Saunders); two females in British
Museum.
Of course, all these Bomdi are the progeny of introduced
individuals. I have credited these and other specimens
to Mr. E. Saunders, following the labels, but I gather that
304 Mr. T. D. A. Cockerell— Descriptions and
Mr. Saunders did not collect them, but only presented
them to the Museum *,
Nomia tenuihirta, Ckll.
Both sexes were taken in Queensland by Gilbert Turner.
The @? is labelled “ Ridg. 4.92” and 869. It differs from
the male by its more robust form, but is in general similar.
Both spurs of the hind tibia are strongly curved at the end.
Nomia Gilberti, sp. n.
Hab. Queensland (Gilbert Turner) ; no. 869, and labelled
« Ridg. 3.92.”
9? .—Length about 85 mm.
Black, with white or yellowish-white pubescence, and
scattered erect black hairs on vertex and occiput ; abdomen
in a bright light appearing very dark reddish, and having
broad but rather thin white hair-bands on the broadly
depressed apical margins of segments 2 to 4, and a patch on
each side of the first segment; fringe of fifth segment dark
grey or blackish ; anterior spur of hind tibia strongly curved,
posterior spur minutely pectinate; legs black, even to the
tarsi, hair on inner side of tarsi black. Mandibles bidentate,
largely dark reddish ; face broad, with coarse yellowish-white
pubescence ; median raised line extending nearly to apex of
clypeus, the latter shining, and curiously longitudinally
ridged in an irregular way; antennz entirely dark; vertex
and mesothorax granulo-punctate. Wings faintly dusky, the
apex conspicuously smoky ; third submarginal cell much
longer than first ; second much higher than broad, but
almost or quite parallel-sided, receiving the first recurrent
nervure near its end.
Superficially, this looks exactly like N. tenuihirta, and
from the fact that the collector gave both the same number
it is probable that he regarded them as the same. They
are, however, clearly distinct, and may be separated as
follows :—
* IT learn from Mr. Chas. O. Waterhouse that all specimens in the
Australian Collection labelled 92. 44, 91.82, 91.155, and 90.126 were
collected by J. J. Walker, on the ‘Penguin’ Expedition ; all labelled
94, 61 and 93.49, from Queensland, were collected by Gilbert Turner ; all
labelled 93.198 by H. W. J. Turner ; all labelled 85. 108 by F. DuBoulay ;
all labelled 85.42 by W. R. Salter; and 1903. 356 by P. F. Dodd.
Records of Bees. 305
N. Gilberti, 9. | N. tenuthirta, 2.
Basal enclosure of metathorax | Basal enclosure of metathorax
nearly or quite as broad (antero- = almost linear at sides; dull,
posteriorly) at sides as the width — granular, without distinct cross-
of the second submarginal cell ;
shining, with irregular but very
distinct cross-ridges.
Tegule piceous, with a whitish Tegule pale testaceous, fuscous
spot on outer margin, basally, and with a whitish
patch posteriorly.
taised facial line hardly invading
ridges,
Raised facial line extending far
down clypeus. clypeus.
Stigma honey-colour, | Stigma dark reddish-piceous.
Wings more hyaline, with the apex | Wings more dusky, the apex uni-
obviously darkened. form with the rest.
In N. Gilberti the triangular area forming the lower
extension of the metathoracic enclosure is perfectly smooth
and shining, and cut off from the ridged area by a transverse
keel ; in N. ¢enuthirta this part is minutely granular like the
rest, and not conspicuously cut off from it.
Nomia semiaurea, sp. n.
? —Length about 10 mm.
Rather broad, the abdomen top-shaped ; black, with some-
times a green tint on middle of first abdominal segment ;
pubescence entirely fulvous or golden, except a very few
scattered dark hairs on mesothorax. Face not very broad ;
inner orbital margins strongly concave above; clypeus
densely covered with elongate punctures (strawberry-
surface) ; face with fulvous tomentum ; supraclypeal area
convex ; longitudinal line very fine, extending from middle
ocellus to beyond middle of clypeus; mandibles bidentate,
more or less stained with dark reddish; flagellum dull dark
brown, delicately pruinose, light at tip; vertex and front
with very minute punctures; mesothorax only sparingly
hairy, the pubescence quite inconspicuous, its surface
minutely roughened but rather shining, with the punctures
large and deep, close at the sides, larger and more separated
in the middle; tubercles densely covered with fulvous
tomentum, as also the postscutellum (which may, however,
be denuded) ; sides of metathorax with long pale fulvous
hair; posterior face very well-defined, shining, with strong
large punctures, and only covered with a very fine tomentum ;
upper part bare and strongly and closely punctured, but the
true enclosure is reduced almost to nothing ; tegule rather
large, orange-fulvous. Wings dusky, especially the apical
part, stigma and nefvures brown; second submarginal cell
306 Mr. T. D. A. Cockerell— Descriptions and
small and narrow, receiving the first recurrent nervure a
little before its end; third with an appendicular apical point.
Tarsiand knees red, and tibi largely so; basal joint of hind
tarsi very broad and flat ; abdomen shining, but strongly and
quite closely punctured, hind margins of segments 1 to 4
broadly depressed and fulvous, those of segments 2 to 4 with
bands of golden-orange appressed hair; fifth segment with a
redder band.
It is not altogether impossible that this may be the 9? of
N. rufocognita, Ckll., but in that species (¢) the middle of
the mesothorax is densely punctured all over, and the first
abdominal segment is much more coarsely and _ closely
punctured. The enclosure of the metathorax in rufocognita,
though a mere line laterally, is divided by little ridges,
which makes it look beaded; this is not at all the case in
semiaured.
Hab. Queensland (Gilbert Turner) ; two, both labelled
Ridg. 4.91 and 617.
From the description, this must be near to Nomia cincta,
Smith, discovered by Wallace on Great Key Island.
Nomia halictella, sp. n.
?.—Length about 8} mm.
Looking very much like a small Halictus calceatus ; the
shining, fulvous-banded abdomen broad in the middle and
narrowing basally, so that in some positions it looks sub-
clavate. Black, with rather light ochreous pubescence ;
head broad, eyes strongly converging below; mandibles
dark reddish in the middle; antennze dark, the flagellum
red at apex; median raised line extending downwards only
to level of antenne; front below ocelli delicately striatulate ;
mesothorax dull, very minutely and closely punctured,
appearing quite hairy when seen from the side, but not so
when seen from above, except at the extreme lateral margins,
and at the scutello-mesothoracie suture, which is full of
ochreous tomentum ; upper border of prothorax, tubercles,
and postscutellum covered with ochreous tomentum ; basal
area of metathorax very narrow (short), with numerous very
fine cross-ridges; posterior face of metathorax dull and
granular, bounded laterally by a ridge only below; tegule
shining rufous, fuscous basally. Wings strongly dusky ;
stigma and nervures light dull brown, stigma large ;
marginal cell very blunt and broadly rounded at apex; first
submarginal cell longer than third; second quite large,
somewhat broader than high, receiving the first recurrent
Records of Bees. 307
nervure near the beginning of its last third. Tarsi dark
reddish ; hair on inner side of basal joint of hind tarsi
(which is rather broad, with the outer apical corner produced
to an acute angle) shining ochreous; second joint of hind
tarsi triangular, the apex produced like that of first joint.
Abdomen shining, with a rather sericeous surface, the
punctures obscure; hind margins of segments rufescent,
those of third and fourth with broad, dense, entire fulvo-
ochraceous hair-bands; on the first and second these bands
only appear laterally, or the band may be nearly complete on
the second; fringe on fifth segment redder; ventral seg-
ments frmged with very long hair.
Hab. Queensland, four specimens (Gilbert Turner). They
all bear the collector’s number 313.
Nomia halictella, var. triangularis, nov.
? .—A little larger (expanse of wings 18 mm.). Stigma
dark reddish; basal enclosure of metathorax much larger in
the middle, being produced behind to form a large triangle,
which is covered with longitudinal ridges, which are not so
fine as those of the type.
This insect is a very puzzling one: the enclosure of meta-
thorax is so different from that of typical halictella, that one
would take it for a distinct species ; but in all other respects
the insect is essentially halictella.
Hab. Queensland (Gilbert Turner), marked 311 Hy.
Halictus urbanus baudinensis, subsp. n.
?.—Length about5 mm. Similar to uwrbanus, with the
same peacock-green mesothorax ; but tibiz and tarsi, and
apical third or fourth of femora, all bright ferruginous ;
hind spur with two long spines ; hind femora somewhat
deformed, appearing as though bent near the middle ; stigma
very dark reddish brown; first r. n. meeting second t.-c. ;
tegule shining fulvous; abdomen beyond the first segment
delicately white-pruinose.
Hab. Baudin I., N.W. Australia (J. J. Walker, 675).
Collected on the ‘ Penguin’ Expedition.
This is very hkely a distinct species, but it is very close to
H. urbanus, and may be treated as a subspecies until more
material has been collected. It has the outer nervures
weakened, as in Chloralictus.
Paracolletes obscurus (Smith).
? .—W. Australia (C. M. Worsfold). The locality is an
308 Mr. O. Thomas on new Neotropical
unexpected one, but the insect runs to obscurus in the table
I made from the types, and agrees sufficiently with Smith’s
description. ‘The abdomen, however, has a narrow white
marginal hair-band on segments 2 to 4. The long black
hair of the head and thorax above is very strongly plumose.
University of Colorado,
Boulder, Colorado, U.S.A.,
May 23, 1905.
XXXIV.—New Neotropical Chrotopterus, Sciurus, Neacomys,
Coendou, Proechimys, and Marmosa. By OLDFIELD
THOMAS.
Chrotopterus auritus guiane, subsp. n.
Ends of wings broadly white for more than an inch at their
tips, both the terminal phalanges of the middle finger and
the membrane itself white, contrasting markedly with the
dark colour of the rest of the membrane; the edge of the
membrane between the fourth and fifth digits also white, and
a slight trace of a whitish edging to the membrane leading to
the foot. Base of thumb almost without hairs. Fur of body
and forearms not extending on to the membranes, which are
practically naked throughout.
~ Other characters as described by Peters, Tomes, and Dobson.
Forearm of type 82 mm.
Hab. Venezuela and Guiana. Type from La Vuelta,
Lower Orinoco, Venezuelan Guiana.
Type. Adult male. B.M. no. 4. 5. 7. 20. Collected 24th
April, 1903, by Mr. 8. M. Klages.
Chrotopterus auritus australis, subsp. n.
White tipping to wings much reduced as compared with
C. a. guiane; the terminal phalanx of the third digit only
whitened, and the membrane only lighter, not white, for about
half an inch at the tip; no whitish line extending along the
hinder edge of the wing-membrane beyond the end of the fourth
digit. Base of thumb with a distinct patch of woolly fur on
the metacarpus. Fur of body extending on to the wing-
membrane behind the elbows below, and on the interfemoral
membrane above behind the thighs and knees.
Forearm of type 83 mm.
Chrotopterus, Sciurus, Neacomys, cc. 309
Hab. Paraguay and Southern Brazil. Type from Concep-
cion, Paraguay; alt. 300 m.
Type. Adult male. B.M. no, 1. 8.11.1. Collected 6th
May, 1900, by Mr. J. Insley. Several specimens also
obtained in Paraguay by Mr. W. Foster.
The differences above described in the amount of white
on the tips of the wings and in the distribution of the fur
seem to be so constant as to indicate a local difference
between the specimens from Guiana and Southern Brazil.
I have seen no examples from Mexico, whence came Peters’s
type, but in his most careful and detailed description he
mentions no white at all on the tips of the wings, nor is any
shown in the plate, and I therefore conclude that the Mexican
Chrotopterus is again different from either of the two races
now described, as was indeed to be expected.
The specimen described and figured by Tomes in 1861*,
and recently acquired with the Tomes collection, proves to be
the southern form, with but little white at the ends of the
wings.
Sciurus Boothie annalium, subsp. n.
General characters as in S. Boothie, but belly grey instead
of white.
Colour of body above as in true Boothie, but less heavily
blackened. Under surface and inner sides of limbs uniformly
dull grizzled greyish (between “ drab-grey”’? and “hair-
brown ’’), not defined laterally from the colour of the sides.
Ears with a narrow black edging ; the small patch behind
their posterior bases dull greyish white. Upper surface of
hands and feet black. Tail-hairs black, annulated along the
middle line below with dull whitish, and tipped with white,
but less broadly than in true Boothie.
Skull as in true Boothie.
Dimensions of the type (measured in the skin) :—
Head and body 250 mm.; tail 250; hind foot, s. u. 51,
c.u. 55 (the second specimen larger: hind foot, s. u. 57,
Gn. GL).
Length of upper tooth-row, exclusive of small premolar, 10.
Hab. Honduras.
Type. Female. B.M. no. 45. 8. 5. 12. Collected by
D. Dyson. Two specimens.
These specimens are those referred to by Gray in his
second description of S. Boothiey, but are so markedly
different, by their greyish bellies, from the type and from a
* P. Z.S. 1861, p. 102, pl. xviii.
+ Ann. & Mag, Nat, Hist. (3) xx. p. 424 (1867).
310 Mr. O. Thomas on new Neotropical
number of adult examples that I have lately had the oppor-
tunity of examining that I think they should receive a special
subspecific name. In his admirable monograph of Central
American Squirrels Mr. Nelson has also referred to these
specimens *,
Neacomys guiane, sp. n.
Very similar to N. spinosus, but conspicuously smaller.
Fur of medium length; hairs and spines of back about
8 mm. long. General colour dark fulvous, heavily lined with
blackish; head and fore-quarters darker; sides brighter
fulvous, especially in front of the hips. Lower flanks greyer,
an ochraceous line edging the white of the belly. Under
surface pure sharply detined white throughout, the hairs white
to their bases. Lips and chin also white. Outer side of
fore limb greyish fulvous, of hind limb clearer fulvous ; inner
side of limbs white. Upper surface of hands and feet dull
white. Tail of medium length, nearly naked, brown above,
whitish below proximally, darkening terminally.
Skull similar to that of N. spinosus, but smaller
throughout.
Dimensions of type (measured in the flesh) :—
Head and body 64 mm.; tail 67; hind foot 18°5.
Skull: greatest length 20; basilar length 15; length of
nasals 7; interorbital breadth 4; breadth of brain-case 10 ;
diastema 5:5; palatal foramina 3; length of upper molar
series 2°5.
Hab. Demerara River, British Guiana. Alt. 120 feet.
Type. Old male. Original number 27. Collected 21st
July, 1905, by Mr. S. B. Warren. One specimen.
The occurrence of Neacomys in Guiana is quite a new
discovery, the nearest locality from which the genus has been
recorded being Bogota. The species may be readily
recognized by its small size, and especially by its small skull
and teeth.
Coendou pruinosus, sp. 0.
A small thickly furred black-and-grey species, with two
sorts of spines as in C, vestitus.
Size about asin C. coudy. Pelage consisting of (1) the
normal spines, about 25 mm. in length, confined to the upper
surface and sides, not occurring on the belly ; (2) long bristle-
spines, attaining 100 mm. in length, at base nearly as thick
as the true spines, but tailing off terminally into a long
* P, Wash, Acad, Sci. i. p. 78 (1899).
Chrotopterus, Sciurus, Neacomys, &c. all
slender point far overtopping the fur; and (3) a thick coat of
woolly hair some 50 mm. in length. General colour of the
animal, as a whole, hoary blackish, the fur (which entirely
hides the spines) uniformly blackish brown, but profusely
lined by the long whitish ends to the bristles. Individually,
on the body, (1) the spines are straw-yellow with blackish
ends ; (2) the bristles are yellowish white basally, have a
broad blackish ring rather beyond their middle, their long
ends being white or yellowish white, contrasting markedly
with the dark fur; (3) the fur is light brown basally,
darkening to blackish brown, and tipped slightly on the dorsal
and broadly on the ventral surface with hoary whitish.
Head with its spines whitish for their basal halves, then
with a broad black ring followed by a white one, the extreme
tip again black; bristles shorter than on the body, their
median dark ring extending downwards nearly to their bases.
Hands and feet blackish, more or less lined with hoary grey.
Tail-base yellowish white above, from the broad whitish ends
to the numerous bristles; below black; the thinly haired
terminal point brown or blackish brown all round.
Skull, as usual, most variable, no two examples resembling
each other, but the dimensions below will give an idea of its
size and shape.
Dimensions of the type (measured by Sr. Bricefio in the
flesh) :—
Head and body 380 mm. ; tail 190; hind foot, s. u. 40,
c. u. 50.
Skull: greatest length 68; basilar length 59; greatest
breadth 40; nasals 20X11:5; interorbital breadth 20;
breadth of brain-case 29; palate length 30; diastema 18 ;
palatal foramina 6 ; length of upper tooth-series 15.
Hab. Merida, Venezuela (Montahas de la Pedregosa).
Alt. 2500 m.
Type. Adult male. B.M. no. 5. 7. 5. 9. Original
number 4. Collected 14th January, 1905, by Sr. Bricefio.
Five specimens.
This well-marked porcupine agrees with C. vestitus, Thos. *,
alone of all described species in the possession of long
spinous bristles in addition to both the normal short spines
and a thick coat of woolly fur. From that animal, however,
it may be readily distinguished by the light colour of the
long ends to these bristles, which in C. vestetus are black.
* Ann, & Mag. Nat, Hist. (7) iv. p. 284 (1899).
312 Mr. O. Thomas on new Neotropical
Proechimys Warrent, sp. 0.
A small species related to P. Cherriei, Thos.
Size quite small, as in P. Cherriec, the hind foot con-
spicuously shorter than in the better-known species of the
P. trinitatis group. Spines thick and strong, about 1°38 mm.
broad by 19 mm. long, very numerous on the back, but
practically absent on the rump and lower flanks. General
colour of the usual dark fulvous, heavily lined with blackish,
the hairs with fulvons subterminal rings, and the spines
greyish white basally, black terminally. Head more greyish
brown. Under surface pure sharply defined snowy white, as
are the hips and inner sides of the limbs. Areas along inner
aspect of forearms and of lower leg thinly haired, almost
naked. Upper sides of hands and feet white. ‘Tail fulvous
like body for its basal inch, then blackish brown above and
white below, the two colours more sharply separated than in
P. Cherrie’; thinly haired, the hairs two or three scales in
length.
Skull very like that of P. Cherriei, agreeing with it in size
and in the smoothness of the brain-case, there being, even in
the oldest specimens, no trace of ridges running across the
parietals. But the palatal foramina are much shorter, widely
open mesially, tapering posteriorly as well as anteriorly,
ending further from the molars, and without any raised
parallel ridges leading back from their edges towards the
front of p*. Pterygoid processes spatulate, about 0°38 mm. broad
at their basal neck, broadening out to 2 mm. at their widest
point ; in P. Cherriet they are almost equally narrow through-
out (U'8 mm. at the neck, 1-0 mm. at broadest point).
Molars rather small, all quadrilaminate as usual.
Dimensions of the type (measured in the flesh) :—
Head and body 228 mm.; tail 185; hind foot (s. u.) 41;
ear 25.
Skull: greatest length 51; basilar length 36°8; greatest
breadth 24°5; nasals 19°5 ; interorbital breadth 11-3; diastema
16°2; palatal foramina 5°1 x 33; length of upper cheek-
tooth series 7.
Hab. Comackka, 80 miles up the Demerara River, British
Guiana. Alt. 50 feet.
Type. Old male. Original number 17. Collected 1st
July, 1905, by Mr. S. B. Warren. Five specimens.
This series of old and young specimens is of interest as
confirming the status of P. Cherriei, which was founded on a
single specimen, whose small size and smooth skull suggested
youth, though the molars were fully worn, From the
Chrotopterus, Sciurus, Neacomys, cc. 313
Orinocan species P. Warreni, though very closely allied, may
be distinguished by its shorter, unrimmed palatal foramina
and broadly spatulate pterygoid processes.
Marmosa cinerea nicarague, subsp. n.
Similar to the Costa Rican MZ. c. Alstonz, Allen *, in size
and general characters. Fur shorter and closer, hairs of back
about 8-9 mm. in length, instead of 11-14, and those of tail-
base only about 7 mm. in length as compared with 12-14.
General colour above greyish brown (nearest to sepia of
Ridgway). Under surface cream-buff, tie hairs of the chest,
middle line of belly, and of inguinal region buffy to their
bases. Hands and feet pale greyish white. Tail very long,
decidedly longer than in A/stonz, dull blackish for about three
fifths of its length, then white, the junction of the two colours
gradual, without marbling ; in Alston? only about one third
of the tail is blackish.
Skull as in the Costa Rica form.
Dimensions of the type (measured in the flesh) :—
Head and body 167 mm.; tail 281; hind foot, s. u. 28°5,
Gu. 30's ear 29:
Skull: basal length 42°7; greatest breadth 25 ; combined
length of three anterior molariform teeth 7°7.
Hab. Bluefields, Nicaragua: sea-level.
Type. Adult male. Original number 5. Collected 9th
January, 1905, by Mr. G. Palmer.
Compared with half a dozen Costa Rican specimens—
practically topotypes—of Dr. Allen’s M/Z. Alston7, this Marmosa
is readily distinguishable by its shorter fur, especially on the
furry tail-base, its longer tail, and the much greater extension
of the black down the latter organ.
Marmosa cinerea demerare, subsp. n.
Size a little smaller than in the other members of the
cinerea group. Fur of medium length ; hairs of back about
10 mm. long. General colour above as usual, near “ broccoli-
brown.” Under surtace broadly washed with ‘‘ cream-buff,”
not sharply defined laterally, the hairs on the chin and down
the centre of the abdomen buffy to their bases. Dark orbital
rings narrow and ill-defined. Muzzle and cheeks pale buffy.
* Caluromys Alstoni, Allen, Bull. Am, Mus. xiii. p. 189 (1900). Iam
still of the opinion that ezmerea and its allies should be placed in Marmosa
rather than in Caluromys. Yor the time being also I prefer to treat both
the Central American forms as subspecies of the Brazilian cinerea.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 21
314 Mr. O. Thomas on a
Hands and feet dull buffy whitish. Tail very long; the
hairs of the basal furry part comparatively short, about as
in M. c. nicarague; naked part brown to within 65 mm.
of the tip, then abruptly white.
Skull as usual, but the molar teeth decidedly smaller than
in the other subspecies.
Dimensions of the type (measured in the flesh) :—
Head and body 166 mm.; tail 250; hind foot (s. u.) 24.
Skull: greatest length 41; basal length 37°3 ; zygomatic
breadth 23°5; combined length of three anterior molariform
teeth 7:1.
Hab. Comackka, 80 miles up Demerara River, British
Guiana.
Type. Adultfemale. Original number 16. Collected 29th
June, 1905, by Mr. 8. B. Warren.
This local representative of the JM. cinerea group differs
from true e¢nerea by the greater extent of the brown part of
its tail and by its smaller teeth. In the former character it
agrees with the Nicaraguan race described above, but by its
small teeth it differs from that as from the other members of
the group. It was quite to be expected that a special race of
M. cinerea should be found in Guiana.
XXXV.—On a remarkable new Squirrel from Burma.
By OLprieLp THOMAS.
Tue British Museum owes to the kindness of Capt. H.
H. Harimegton, of the 90th Punjabis, two specimens, from the
Upper Chindwin River, of a very remarkable squirrel, quite
unlike anything we have seen before. I would propose to
eall it
Sciurus [Haringtont, sp. n.
A pale creamy-buff species with a buffy belly and a whitish
tail.
Size medium. Fur of back rather over 20 mm. in length.
General colour of upper surface “ cream-buff” along the
dorsal area, the buff fading out on the sides, which are dull
whitish. Individually the hairs of the back are whitish grey
basally, with a broad cream-buff subterminal band and a
minute black point. Head creamy whitish, with a slight
buffy suffusion on the crown; the cheeks dull white. Ears
new Squirrel from Burma. 315
whitish, both outside and in. Under surface, from chin to
anus, bright sharply contrasted ochraceous buff (in the type;
the second specimen nearer tawny ochraceous). Lateral line
of demarcation very sharply marked in both specimens, and in
the type emphasized by a blackish line which runs from the
middle of the front of the forearm, across the shoulders, down
the sides and legs to the back of the heel. Fore limbs on
outer side above this line of demarcation creamy whitish,
like the flanks; beyond it, including the hands and the
whole of the inner aspect, ochraceous buffy like the belly, or
slightly paler. Back of upper part of hind leg whitish like
body ; inner side, ankles, and feet buffy like belly, rather
paler on the digits. Tail above and below creamy buff
proximally (the extreme tips of the hairs blackish), lightening
to white terminally.
Skull of the general proportions of that of S. Finlaysont,
erythreus, &c., but there is only one upper premolar in the
single specimen of which the skull has been preserved. This
has the milk-premolar still in place, and the large permanent
premolar just coming up below it, but no trace of the small
p, which usually at this stage is readily discernible.
Dimensions of the type (measured in the flesh) :—
Head and body 250 mm.; tail 265; hind foot (s. u.) 54;
ear 22.
Skull: zygomatic breadth 32; back of postorbital process
to tip of nasals 82°5 ; nasals 16:4 x 7:2; interorbital breadth
18; palate length 23; length of upper tooth-series 10.
Hab. Upper Chindwin River, Burma. Type from
Moungkan; a second specimen from Homalin. About 25°N.,,
95° E.
Type. Immature male. B.M.no. 5. 8.11.1. Collected
14th December, 1904, and eee ‘by Capt. Hii. Ee
Harington.
This very peculiar squirrel is so entirely unlike any known
species that I do not know with what to compare it. Its pale
creamy-buff colour and whitish tail are quite unique, while
no other Oriental squirrel of its size is without the small
upper premolar. This latter character also indicates that
Sciurus Haringtonz is not merely a partially albinistic variation
of some known species, an explanation which its pale colour
might at first sight suggest.
Capt. Harington i is to be congratulated on the discovery of
so striking a new form,
21%
316 Mr. C. T. Regan on the Fishes of the
XXXVI.—A Revision of the Fishes of the American Cichlid
Genus Cichlosoma and of the Allied Genera. By C. Tare
Reean, B.A.
(Continued frem p. 245.]
Section 9.
Two species from Central America agree in most respects
with those of the preceding section, but differ in the some-
what fewer teeth, which are more distinctly enlarged
anteriorly. Dorsal XVI-XVII 31-12. Anal VII 8-9.
38. Cichlosoma margaritiferum.
Heros margaritifer, Giinth. Cat. Fish. iv. p. 287 (1862), and Trans,
Zool. Soe. vi. 1869, p. 450, pl. Ixxi. fig. 2.
Cichlasoma margaritiferum, Jord. & Everm. Bull. U.S. Nat. Mus. xtvii.
1898, p. 1519; Pellegr. Mém. Soc. Zool. France, xvi, 1903, p. 211
(1904).
Depth of body 2} in the length, length of head 3. Snout
slightly longer than postorbital part of head. Diameter of
eye 41 in the length of head, interorbital width 3. Depth
of preorbital 14 the diameter of eye. Maxillary not ex-
tending to the vertical from anterior margin of eye; prie-
maxillary processes not extending to above eye; jaws equal
anteriorly ; fold of the lower lip slightly interrupted ; cheek
with 5 series of scales; 12 gill-rakers on the lower part of
anterior arch. Scales 31 <, 3 between lateral line and base
of anterior rays of soft dorsal. Dorsal XVII 11, commencing
above the opercular cleft, the spines subequal from the sixth,
the last less than } the length of head ; the soft fin, when
laid back, extending to the anterior part of caudal. Anal
VII 9. Pectoral a little shorter than the head, extending
to the third or fourth anal spme; ventral extending to
second anal spine. Caudal slightly emarginate, with rounded
lobes. Caudal peduncle deeper than long. Brownish, with
6 or 7 dark cross-bars and a dark spot at the base of caudal ;
each scale of the side of the body with a white pearl-like
spot; tins, except the pectoral, dusky, the dorsal with some
oblique stripes, the caudal with clear spots.
Guatemala.
1. (164 mm.) type of the species. Guatemala, O. Salvin, Esq.
39. Cichlosoma citrinellum.
Heros citrinellus, Giinth. Proc. Zool. Soc. 1864, p. 153, and Trans. Zool.
Soc. vi. 1869, p. 458, pl. xxi. fig. 1; Jord. & Everm. Bull. U.S, Nat.
American Cichlid Genus Cichlosoma. 317
Mus. xvii. 1898, p. 1534; Pellegr. Mém. Soc. Zool. France, xvi.
1903, p. 237 (1904).
Heros tohochilus (part.), Giiath. Trans. Zool. Soc. vi. 1869, p. 457.
Heros basilaris, Gill & Bransferd, Proc. Ac. Philad. Me ige p. 182
Pellegr. t. ¢. p. 258.
Cichlasoma basilare, Jord. & Tiverm. f. ¢. p. 1532
Depth of body 2-21] in the length, length of head 24-3.
Snout a little shorter than postorbital part of head.
Diameter of eye 37-44 in the length of head, interorbital
width 22-3. Depth of preorbital about equal to the dia-
meter of eye. Maxillary not extending to below the eye;
preemaxillary processes extending to above anterior margin of
eye ; Jaws equal anteriorly ; fold of the lower lip continuous
or subcontinuous; cheek with 4 or 5 series of scales; 9 or
10 gill- es on the lower part of anterior arch. Scales
31-33. om wp 21-3} between lateral line and base of anterior
rays of soft dorsal. Dorsal XVI-XVII 11- 12, commencing
above the opercular cleft, the spines subequal from the sixth
to the fourteenth, thence increasing to the last, which is
2 the length of head ; soft fin, when “laid back, extending to
the middle of caudal or beyond. Anal VII 8-9. Pectoral
as long as the head, extending to above the third or fourth
anal spine ; ventral extending beyond the origin of anal.
Caudal rounded. Caudal peduncle 2 3-4 as long as deep.
Olivaceous or yellowish, the back darker, the dark colour
sometimes extending as cross-bars on the sides of the body ;
sometimes a dark blotch on the middle of the side and a dark
spot at the base of the caudal. Fins pale yellow or wholly
or partly dark, sometimes spotted or barred.
Nicaragua.
1-3. (175-189 mm.) types of — Lake Nicaragua. QO. Salvin, Esq.
the species.
4. (190 mm.) Lake Managua, Capt. J. M. Dow.
5. (180 mm.) one of the Lake Nicaragua. Smithsonian Inst.
types of HZ, basilaris.
6, (188 mm.) Lake Nicaragua, Smithsonian Inst.
Section 10.
Three species from Nicaragua have the general form of
those of the two preceding sections and are evidently closely
allied to C. citrinellum, from which they differ in the excep-
etionally thick lips and in the dentition. Upper jaw with the
anterior pair of teeth strongly enlarged ; lower jaw with the
two anterior pairs strong, subequal. Dorsal XVII 11-12.
Anal VII-VIII 8-9. Caudal rounded or subtruncate.
318 Ma@sl: ee on the Fishes of the
40. Gintosem erythreum.
Heros erythreus, Giinth. Trans. Zool. Soc. vi. 1869, p. 457, a Ixay;
fig. 2; Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898, p-. 16381;
Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 239 (1 904).
Depth of body 23 in the length, length of head 24. Snout
as long as postorbital part of head. Diameter of eye 4) in
the length of head, interorbital width 3. Depth of pre-
orbital equal to diameter of eye. Maxillary not extending
to below the eye ; preemaxillary pr ocesses extending to above
anterior + of eye; jaws equal anteriorly ; lips thick, the
lower with a continuous fold; cheek with 5 series of scales ;
8 gill-rakers on the lower part of anterior arch. Scales 31 ig
3 between lateral line and base of anterior rays of soft
dorsal. Dorsal XVII 12, commencing above the opercular
cleft, the spines subequal from the sixth to the fifteenth,
thence increasing to the last, which is a little more than 4
the length of head. Anal VII 8. Pectoral nearly as long as
the head, extending to above the fourth anal spme; ventral
extending beyond origin of anal. Caudal peduncle 2 as long
as deep. Brownish, obscurely marbled with darker, and
with some dark spots on the posterior part of the body ; fins
pale yellow.
Lake Managua, Nicaragua.
1, (180 mm.) type of the species. Lake Managua. Capt. J. M. Dow.
41. Cichlosoma lobochilus.
Heros lobochilus (part.), Giinth. Trans. Zool. Soe. vi. 1869, p. 457,
pl. Ixxv. fig. 1; Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898,
p- 1530; Pellegr. Méin. Soc. Zool. France, xvi. 1903, p. 240 (1904).
Depth of body 24 in the length, length of head nearly 3
Snout as long as postorbital part of head. Diameter of eye
41 in the length of head, mterorbital width 3. Depth of
preorbital equal to diameter of eye. Maxillary not extend-
ing to below the eye; premaxillary processes extending to
above anterior 4 of eye; jaws equal anteriorly ; upper and
lower lip each forming a large fleshy triangular flap ; cheek
with 5 series of scales; 9 gill-rakers on the lower part of
anterior arch. Scales 31,4, 24 between lateral line and base
of anterior rays of soft dorsal. Dorsal XVII 11, com-
mencing above the opercular cleft, the spines subequal from
the seventh to the fourteenth, thence increasing in length to
the last, which is 4 the length of head; soft fin, when laid
back, extending to middle of caudal. Anal VII 9. Pectoral
nearly as long as the head, extending to above third anal
American Cichlid Genus Cichlosoma. O19
spine ; ventral extending to middle of anal. Caudal rounded.
Caudal peduncle 2 as long as deep. Olivaceous, with 7 dark
cross-bars, the fourth bearing a darker blotch below the
lateral line; a dark spot on the upper 3 of base of caudal ;
vertical fins dusky, the soft dorsal with “obscure darker spots.
Lake Managua, "Nicaragua.
1. (196 mm.) type of the species. Lake Managua. Capt. J. M. Dow.
42. Cichlosoma labiatum.
Heros labiatus, Giinth. Proc. Zool. Soc. 1864, p. 27, pl. iv. fig. 1, and
Trans. Zool. Soc. vi. 1869, p. 456; Jord. & "Everm. Bull. U.S. Nat.
Mus. xlvii. 1898, p. 1530 ; Pellegr, Mém. Soe. Zool. France, xvi.
1903, p. 259 (1904).
Depth of body 24 in the length, length of head 23-23.
Snout as long as or longer than postorbital part of head.
Diameter of eye 44-42 in the length of head, interorbital
width 3-32. Depth of preeorbital equal to or a little more
than diameter of eye. Maxillary not extending to below the
eye ; premaxillary processes extending to above anterior 4
of eye; jaws equal anteriorly ; upper and lower lip each
forming a large fleshy triangular flap ; 4 or 5 series of scales
on the cheek ; 11 or 12 gill-rakers on the lower part of
anterior secha Seales 30 au 3 between lateral lime and
base of anterior rays of soft dorsal. Dorsal XVII 11-12,
commencing above the opercular cleft, the spines subequal
from the seventh to the fourteenth, thence i imcreasing to the
last, which is 1-3 the length of head - isortifiny when laid
back, extending to posterior part of caudal. ‘Anal VII-
VII 8-9. Pectoral + the length of head, extending to
above the fourth or fifth anal spine ; ventral extending
beyond origin or nearly to middle of ell Caudal rounded
or subtruncate. Caudal peduncle 7-4 as long as deep.
Some specimens uniformly pale yellow, others brownish
with obscure darker cross-bars and dark spots on the vertical
fins.
Nicaragua.
1-8. (178-190 mm.) types of Lake Nicaragua. Capt. J. M. Dow.
the species.
4-5. (162 and 178 mm.) Lake Managua. Dr. P. L. Sclater.
Section 11 (Thorichthys).
Body deep or moderately elongate (depth 2-22 in the
length). Upper profile of snout straight, oblique. Scales
of the lateral line of the same size as those above and below
320 Mr. C. T. Regan on the Fishes of the
it; scales of thoracic region considerably smaller than those
on the side of the body. Mouth moderate, moderately
protractile ; maxillary slightly exposed ; teeth of the outer
series small or moderate, in the upper jaw moderately in-
creasing in size anteriorly, in the lower with 4 or 5 anterior
teeth on each side distinctly enlarged. Dorsal XV-X VIL
8-12. Anal VI-IX 6-9. Dorsal and anal fins entirely
scaleless. Caudal slightly emarginate, with pointed lobes.
Three species from Mexico and Central America are allied
to C. macracanthus &c., but differ from all other members
of the genus in the scaleless dorsal and anal fins and in the
form of the caudal.
45. Cichlosoma aureum.
Heros aureus, Gunth. Cat. Fish. iv. p. 292 (1862), and Trans. Zool.
Soc. vi. 1869, p. 455, pl. Ixxiii. fig. 2; Jord. & Everm. Bull. U.S.
Nat. Mus. xlvii. 1898, p. 1533; Pellegr. Mém. Soc. Zool. France,
xvi, 1903, p. 238 (1904).
Heros Helleri, Steind. Denkschr. Ak. Wien, xxiii. 1864, p. 64, pl. iv.
figesk:
Heros maculipinnis, Steind. t. ¢. p. 69, pl. iv. fig. 2; Jord. & Everm.
Bull. U.S. Nat. Mus. xlvii. 1898, p. 1530 ; Pellegr. ¢. ¢. p. 240.
Cichlasoma Helleri, Jord. & Mverm. t. c. p. 1521; Pellegr. t. e. p. 220.
Thorichthys Hellert, Meek, Zool. Pub. Columbian Mus. vy. 1904, p. 225.
Thorichthys Elliott, Meek, 1. ¢. fig. 72.
Depth of body 2-22 in the length, length of head 23-8.
Snout straight, as long as postorbital part of head. Diameter
of eye 22-34 in the length of head, interorbital width 3-33.
Depth of preorbital equal to the diameter of eye or less.
Maxillary not quite extending to below the eye; preemaxillary
processes extending to above anterior 4 of eye; jaws equal
anteriorly ; fold of the lower lip continuous or not; cheek
with 4-6 series of scales ; 9 or 10 gill-rakers on the lower
part of anterior arch. Scales 29-32 a 2 or 3 between
lateral line and base of anterior rays of soft dorsal. Dorsal
XV-XVII 9-12, commencing above the opercular cleft, the
spines subequal from the sixth, the last about 5 the length
of head; soft fin, when laid back, extending to anterior 4 or
middle of caudal. Anal VII-VIII 7-8 (VI-IX 6-9).
Pectoral as long as the head, extending to above middle anal
spines; ventral extending beyond origin of anal. Caudal
slightly emarginate with pointed lobes. Caudal peduncle a
little deeper than long. Olivaceous, with 6 dark cross-bars,
the third bearing a blackish blotch on or below the lateral
line ; suboperculum usually blackish ; head and anterior part
American Cichlid Genus Cichlosoma. 321
of the body with blue spots; vertical fins dusky, often with
clear spots.
Guatemala; Southern Mexico.
1. (108 mm.) type of the Lake Yzabal. O. Salvin, Esq.
species.
2. (95 mim.) R. Motagua, O. Salvin, Esq.
3. (44 mm.) Mexico. M. Sallé.
4, (126 mm.) R. de Sarabia. Dr, A..C. Buller.
5. (92 mm.) San Domingo de Dr. A. €. Buller.
Guzman.
6-9. (83-113 mm.) Motzorongo. Dr. H. Gadow.
10-11. (80 and 109 mm.) R. Tonto. Dr. H. Gadow.
44, Cichlosoma ajjine.
Heros affinis, Gunth, Cat. Fish. iv. p. 292 (1862), and Trans. Zool. Soe.
vi. 1869, p. 455, pl. Ixxix. fig.1; Jord. & Everm. Bull. U.S. Nat. Mus.
xlvii. 1898, p. 1529; Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p- 241 (1904).
Depth of body about 21 in the length, length of head
23-27, Snout longer than postorbital part of head. Dia-
meter of eye 31-32 in the length of head, interorbital width
34-32. Depth of preorbital I-14 the diameter of eye.
Maxillary not extending to the ver tical from anterior margin
OL eve. premaxillary processes not extending to above the
eye; lower jaw projecting ; fold of the lower. hp coutinuous
or not; cheek with 4 or 5 series of scales; 11 or 12 gill-
rakers on the lower part of anterior arch. Scales 27-29 3;
2 between lateral line and base of anterior rays of soft dorsal.
Doisal XVI-XVII 8-9, commencing above the opercular
cleft, the spines subequal from the fifth, the last nearly 1 the
length of head; the soft fin, when laid back, extending to
anterior 1 of caudal. Anal VIII-IX 7-8. Pectoral as long
as the head, extending to first soft ray of anal; ventral
extending bey ond origin of anal. Caudal slightly emarginate,
with pointed lobes. Caudal peduncle a little deeper than long.
Olivaceous, with 6 dark cross-bars, the third bearing a large
blackish blotch on the lateral line ; ; suboperculum usually
blackish ; blue spots on the head and anterior part of body ;
fins, except the pectoral, dusky, the unpaired ones with light
bluish spots enclosed in dark rings.
Guatemala; British Honduras.
1-4, (98-130 mm.) types Lake Peten. O. Salvin, Esq.
of the species.
45. Cichlosoma callolepis.
TTeros callolepis, Regan, Ann. & Mag. Nat. Hist. (7 ) xiii. 1904, p. 258 ;
Pellegr. Mém. Soe. Zool. Fiance, 3 xvi. 1903, p. 238 (1904).
Depth of body about 22 in the length, length of head 3.
322 Mr. C. T. Regan on the Fishes of the
Snout longer than postorbital part of head. Diameter of eye
34 in the length of head and equal to the interorbital width.
Depth of przeorbital equal to diameter of eye. Maxillary not
extending to below the eye ; preemaxillary processes extending
to above anterior margin of eye ; jaws equal anteriorly ; fold
of the lower lip continuous or not ; cheek with 4 or 5 series
of scales; 10 gill- saere on the lower part of anterior arch.
Scales 28-29 1s 2 between lateral line and base of
anterior rays of soft Monee Dorsal XV 9-10, commencing
above or behind axil of pectoral, the spines subequal from the
filth, the last 2 the length of head or less; the soft fin, when
laid back, extending to. anterior 4 of caudal. Anal VI-VII
7-8. Pectoral 3 ihe length of head, extending to above
origin of anal; ventral extending beyond origin of anal.
Caudal slightly emarginate, with pointed lobes. Caudal
peduncle as long as deep. Brownish, with faint traces of
darker cross-bars and with a dark blotch on the lateral line
below the last dorsal spines; small hight blue spots on the
head and at the base of each scale on the body ; fins, except
the pectoral, dusky.
San Domingo de Guzman, Mexico.
1-2. (93 and 100 mm.) San Domingo de Guzman. Dr. A. C. Buller.
types of the species.
Section 12 (Heros).
Body deep. Scales of the lateral line of the same size as
those below it, but rather larger than those above it; scales
of thoracic region considerably smaller than those on the side
of the bedy. Mouth rather small, moderately protractile ;
maxillary slightly exposed ; teeth of the outer series moderate,
distinctly increasing im size anteriorly. Dorsal XV-XVII
13-14, the soft fin scaly at the base. Anal VII-VIII 12-14.
Pectoral extending beyond origin of anal. Caudal sub-
truncate.
The single species, from Brazil and Guiana, is probabiy
derived from the type represented by C. facetum. It leads
to the genera Symphysodon and Pterophyllum.
46. Cichlosoma severum.
Heros severus, Heck. Ann. Mus. Wien, ii. 1840, p. 562
Heros corypheus, Heck. t. c. p. 364.
Heros modestus, Heck. t. ¢. p. 366.
Heros spurius, Heck. t.c. p. 868; Giinth. Cat. Hee iv. p. 293 (1862) ;
Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 256 (1904).
Heros efasciatus, Heck, ¢. c. p. 372; Ginth. ¢. c. p. 294.,
Chromys appendiculata, Casteln, Anim. Am. Sud, Poiss. p- 15, pl. vii.
tig, 3 (1895).
American Cichlid Genus Cichlosoma. 323
Chromys fasciata, Casteln. 0. e. p. 17, pl. ix. fig. 2.
Uaru centrarchoides, Cope, Proc. Ac. Philad. 1872, p. 253, pl.
fig. Dh.
Acara (Heros) spuria, Steind. Sitzb. Ak. Wien, Ixx. 1874, p. 507, pl. iv.
and Ixxi. 1875, p. 83.
Astronotus ( Cichlasoma) severus, Eigenm. & Bray, Ann. Ac. N. York,
vii. 1894, p. 619.
Depth of body 13-14 in the length, length of head 24-3.
Snout longer than postor bital part of head. Diameter of eye
3-33 in the length of head, interorbital width 2 22. Depth
of preorbital #-11 the diameter of eye. Maxillary not
extending to below the eye; praemaexillary processes not
extending to above the eye (in the adult); jaws equal
anteriorly ; fold of the lower lip continuous, somewhat
produced; cheek with 5 or 6 series of scales; 9-11 gill-
rakers on the lower part of anterior arch. Scales 36-42
i» 28-30 in a longitudinal series below the lateral line,
4 or 5 between lateral line and base of anterior rays of soft
dorsal. Dorsal (XV) XVI-X VII 13-14, commencing above
the opercular cleft, the spines subequal from the seventh or
slightly increasing in length to the last, which is }—? the
length of head; soft fin, when laid back, extending to
posterior part of caudal. Anal VII-VIII 12-13 (14).
Pectoral as long as the head, extending to above the fifth
anal spine; ventral extending to middle of anal. Caudal
subtruncate. Caudal peduncle }-? as long as deep. Oliva-
ceous, with obscure darker cross- -bars ; ; a blackish bar from
soft dorsal to posterior part of anal, i sometimes another
on the base of caudal; head and body with spots or vermi-
culations ; vertical fins with dark stripes or series of spots.
Brazil; Guiana.
1-2. (89 and 158 mm.) Rio Cupai.
3. (163 mm.) Guiana. Sir R. Schomburek.
4, (131 mm.) Zoological Society.
5-6. (174 and 192 mm.) Teffé. Prof. A. Agassiz.
7-8. (81 and 111 mm.) Teffé. Dr. J. Bach.
9. (190 mm.) Deffé. Paris Museum.
Section 13 (Hoplarchus).
Body ovate. Scales of the lateral line larger than those
above and below it. Dorsal XV 12-13. Anal V 8-10.
The single species, from South America, appears to be
related to C. severum. It leads to the genus Uaru.
47. Cichlosoma psittacum.
Heros psittacus, Heck. Ann. Wien Mus. ii. 1840, p. 869; Gunth. Cat.
Fish. iv. p. 290 (1862) ; Pellegr. Mém. Soc. Zool. France, xvi. 1908,
p. 242 (1904).
S24 Mr. C. T. Regan on the Fishes of the
Pomotis fasciatus, Schomb. Fish. Guiana, ii, p. 169, pl. xvii. (1843).
Poplanchus pemtacunthus. Kaup, Arch. Nat. 1860, p. 129, pl. vi. fig. 1.
Acara (Heros) psittacus, Steind. Sitzb. Ak. Wien, 1xxi. 1875, p. 87.
Beat of body about 2 in the length, length of head
about 23. Diameter of eye 3-4 in the length ‘of head and
nearly equal to the interorbital width. Depth of preorbital
equal to diameter of eye. Maxillary not extending to below
the eye; fold of the lower lip continuous; cheek with 10-14
series of scales; 7 gill-rakers on the lower part of anterior
arch. Scales 44-48 (probably more if counted above the
lateral line) ;23. Dorsal XV ne the spines increasing
in length to the last, which is } the length of head. Anal
V 8-10. Pectoral nearly as ioe as the head. Caudal
rounded. Caudal peduncle deeper than long. Yellowish or
reddish, with a dark longitudinal band from eye to base of
caudal, where it forms a spot.
R. Amazon ; R. Orinoco.
Heckel’s type measures 130 mm. in total length.
PARAPETENIA, subgen. nov.
Typically with larger, more oblique and more protractile
mouth than in other groups of Cichlosoma and usually with
maxillary more distinctly exposed distally ; but none of these
features are constant. Upper jaw with the anterior pair of
teeth siightly enlarged in the more generalized forms, strongly
enlarged in the more specialized ones. Lower jaw with the
3 anterior teeth on each side more or less canine-like, with
the anterior pair somewhat smaller than the others (corre-
sponding to the enlargement of the anterior pair in the upper
jaw) in the more generalized forms. The anterior pair
wanting, the next pair strong, and the third pair forming
very strong canines in the more specialized forms. Scales
of lateral line of the same size as those below it on the side
of the body; scales of the thoracic region much smaller than
those on the side of the body. Dorsal XV-XIX 8- 13%
posterior part of spinous dorsal and soft fin with a scaly
sheath at the base and with short series of scales on the
interradial membranes. Anal 1V—X 7-10. Caudal rounded
or subtruncate.
48. Cichlosoma adspersum.
Acara adspersa, Giinth. Cat. Fish. iv. p. 282 (1862).
Heros adspersus, Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 227
(1904).
Depth of body 2? in the length, length of head 2+. Snout
American Cichlid Genus Cichlosoma. 325
as long as postorbital part of head. Diameter of eye 51 in
the length of the head, interorbital width 34. Depth of
preorbital 12 the diameter of eye. Maxillary slightly
exposed, extending nearly to below the eye; premaxillary
processes extending to above anterior } of eye; jaws equal
anteriorly, the anterior pair of teeth in the upper jaw the
largest and in the lower jaw somewhat smaller than the next
2 on each side, which are slightly enlarged; fold of the
lower lip continuous; cheek with 7 series of scales ; 2 gill-
rakers on the lower part of anterior arch. Scales 31 2, 34
between lateral line and base of anterior rays of soft derek
Dorsal XV 12, the spines subequal from the sixth to the
twelfth, thence increasing to the last, which is a little more
than } the length of head ; so‘t fin, when laid back, extending
to middle of caudal. Anal 1V 10. Pectoral less than 2 the
length of head, not extending to above the anal ; ventral
extending to origin of anal. Caudal rounded. Caudal
peduncle nearly as long as deep. Olivaceous, with numerous
small blackish spots on head, body, and dorsal and caudal
fins.
Barbadoes.
1, (218 mm.) type of the Barbadoes, Sir R. Schomburgk.
species.
49. Cichlosoma tetracanthus.
Centrarchus tetracanthus, Cuy. & Val. Hist. Nat. Poiss. vii. p- 460
(1881).
Chromis fuscomaculatus, Guichen. Hist. Cuba, Poiss. p. 78, pl. ii. fig. 8
(1853).
Acara fuscomaculata, Giinth. Cat. Fish. iv. p. 282 (1862).
Acara tetracanthus, Steind. Denkschr. Ak. Wi len, xxiii. 1864, p. 60.
Heros tetracanthus, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898,
p- 1539; Eigenm. Bull. U.S, Fish. Comm. 1902, p. 2380, figs. 12
& 13,
Heros fusecmaculatus, Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 227
(1904).
Heros tetracanthus torralbasi, Kigenm. 1. ¢. fig. 11.
Heros tetracanthus griseus, Eigenm. ¢ bc Cups 233, fie. 14,
Hervs tetracanthus latus, Eigenm, t. C. p: iat fic. “15.
Heros tetracanthus cinctus, Eigenm. l. c. fig. x. 16,
Heres nigricans, Eigenm. tc. p- 285, fig. 17.
Depth of body 2-23 in the length, length of head 2
Snout shorter than postorbital part of head. ne of
eye 3-44 in the length of head, interorbital width 31-33
Depth of prorbital 2 —4 the diameter of eye. Maxillary
slightly exposed, extending to below anterior margin of eye ;
premaxillary processes extending to above anterior 1 of eye;
jaws equal anteriorly, or the lower slightly projecting; ‘anterior
326 Mr. C. T. Regan on the Fishes of the
pair of teeth in the upper jaw and 2-3 on each side in the
lower enlarged, forming weak canines ; fold of the lower lip
continuous ; cheek with 7-10 series of scales; 8-10 gill-
neers on the lower part of anterior arch. Scales 28-381
mals a 3 or 4 between lateral line and base of anterior rays of
soft dorsal. Dorsal XV-XVI 10-12, the spines slightly
increasing in length to the last, which is 1—? the length of
head, or sometimes the middle pes subequal ; soft fin,
when laid back, ee to anterior 4 of caudal. Anal*
IV 8-10. Pectoral 2— 3-3 the length of head, usually not
extending to above “ ar ventral not extending to origin
of anal. Caudal subtruncate or rounded. Caudal peduncle
3-3 as long as deep. Olivaceous, with dark spots on head,
body, and vertical fins; sometimes dark cross-bars on the
body and a dark spot below the origin of the lateral line,
another on the middle of the side, and a third on the base of
the caudal.
Cuba.
* (151 mm.) Cuba. Zoological Society.
2. (86 mm.) Puiar del Rio. Protons Eigenmann.
3-4. (80 and 97 mm.) Calabazar. Prof. C Ee Eigenmann.
5-6. (98 and 118mm.) San Antonio. Prof. C. H. Eigenmann.
7-12. (102-154 mm.) Palacios. Prof. C. H. Eigenmann.
13-27. (64-121 mm.) San Cristobal. Prof. C. H. Eigenmann.
50. Cichlosoma istlanum.
? Cichlasoma Steindachneri, Jord. & Snyder, Bull. U.S. Fish. Comm. xix,
1899, p. 143, fig. 20; Jord, & Everm. Bull. U.S. Nat. Mus. xlvii.
1900, Dp 3173; “Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 206
(1904) ; Meek, Zool. ‘Pub. Columbian Mus. v. 1904, p. 211, fi. 67.
Heros istlanus, Jord. & Snyder, t. c. p. 144; Jord. & Everm. t. ¢.
p. 3174; Pellegr. t. c. p. 228.
Cichlasoma istlanum, Meek, é. c. p. 213, fig. 69.
Depth of body 22-22 in the length, length of head 23-24.
Snout as long as postorbital part of head. Diameter of eye
41 in the length of head, interorbital width 33-34. Depth of
preorbital a little less than the diameter of eye. Maxillary
slightly exposed, not extending to below tlie eye; pre-
maxillary processes extending nearly to above middle of
eye; jaws equal anteriorly ; fold of the lower lip sub-
continuous ; upper jaw with the 2 anterior teeth of the outer
series more or less distinctly differentiated as a pair of
canines ; lower jaw with 2 or 3 pairs of rather weak canines ;
* Two specimens from San Cristobal have curiously malformed anal
fins. In each case the posterior part of the fin is w anting ; in one there
are 10 spines and no soft rays, in the other 8 spines and 3 soft rays,
American Cichlid Genus Cichlosoma, 327
cheek with 6 series of scales; 8 gill-rakers on the lower part
of the anterior arch. Scales 30-31 =a 21 between lateral
line and base of anterior rays of soft dorsal. Dorsal XVI 10,
the spines subequal from about the sixth to the thirteenth,
thence increasing to the last, which is 2 the length of head ;
soft fin, when laid back, extending to middle of caudal.
Anal V 8. Pectoral 3 the length of head or more, not
extending to above the anal; ventral extending to origin of
anal or a little beyond. Caudal rounded. Caudal peduncle
as long as deep. Olivaceous, each scale with a small dark
spot at the base; obscure traces of dark cross-bars; a dark
spot below the origin of the lateral line and another on the
middle of the side ; vertical fins with small dark spots.
Rio Ixtla, Mexico.
1-2. (187 and 145 mm.) co-types Rio Ixtla. Dr. D. S. Jordan.
of the species.
With these I have compared several small examples (30-
40 mm.), co-types of C. Steindachneri from the Rio Verde,
which appear to me to present all the characters which might
be expected in young specimens of C.istlanum. The general
form of the body and the number of scales and fin-rays seem
to be the same. The larger eye, shorter snout, smaller
mouth, less produced soft dorsal and anal fins, &c. are
differences due to size. ‘There are several dark cross-bars on
the body and an interrupted longitudinal band bearing a dark
blotch on the middle of the side. The lower lip has the same
structure as in the larger specimens named C. istlanum. I
have retained the name C. istlanum until actual comparison
of adult fish from the two rivers has been made.
51]. Cichlosoma Bartoni.
Acara Bartont, Bean, Proc. U.S. Nat. Mus. 1892, p. 286, pl. xliv. fig. 3.
Cichlasoma Bartoni, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898,
. 1515, fig. 587 ; Peller. Mém. Soe. Zool. France, xvi. 1903, i 206
(1904); Meek, Zool. Pub. Columbian Mus. v. 1904, p. 211, fig. 68.
* Heros (Cichlasoma) labridens, Pellegr. Bull. Mus. Paris, 1903, p. 122.
Cichlasoma labridens, Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 207,
pl. vi. fig. J (1904).
Depth of body 2}—22 in the length, length of bead 23-3.
Snout nearly as long as postorbital part of head. Diameter
of eye 43-53 in the length of head. Depth of przorbital
ereater than diameter of eye. Maxillary not extending to
below the eye; lower jaw projecting ; ; fold of the lower lip
* In placing Heros labridens in the synonymy of C. Bartoni, I follow
Dr. Meek, who has examined a number of specimens.
328 Mr. C. T. Regan on the Fishes of the
not continuous ; anterior teeth forming canines ; cheek with
5 or 6 series of scales; 7 gill- rakers | on the lower part of
anterior arch. Scales about 30 5. Dorsal XV-XVI
10-11, the spines increasing in length to the last, which is
2 the length of head ; soft fin extending a little beyond base
of caudal. Anal V (IV-VI) 8-9. Pectoral 2 the length of
head, not extending to above the anal; ventral extending to
the vent. Caudal rounded. Caudal. peduncle as lone as
deep. Brownish, with about 6 indistinct cross-bars on the
sides ; a dark spot at the base of caudal.
Rio Panuco and its tributaries, Mexico.
The types of the species measure from 90-180 mm. in total
length, those of C. labridens 200-220 mm.
52. Cichlosoma Beani.
Heros Beant, Jord. Proc. U.S. Nat. Mus. 1888, p. 322, and Proe. Cal. Ae.
Sci. 1895, p. 473; Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898,
p. 1538 ; Pelleor. “Mém. Soe. Zool. France, xvi. 1903, p. 250 (1904).
Cichlasoma Beant, Meek, Zool. Pub. Columbian Mus. v. 1904, p. 210.
Depth of body 2-23 in the length, length of head 23-3,
Snout more than 4 the length of head (in the adult), less in
the young. Diameter of eye 33-5 in the length of head,
interorbital width 38-84. Depth of preeorbital a little greater
than diameter of eye (adult), 2 diameter of eye (young).
Maxillary slightly exposed, not extending to below the eye ;
preemaxillary processes extending nearly to above middle of
eye; lower jaw slightly projecting ; fold of the lower lip
continuous ; upper jaw with the 2 anterior teeth of the
outer series more or less distinctly differentiated as a pair of
canines; lower jaw with the 2 or 3 anterior teeth on each
side forming more or less distinct canines; cheek with 6 or
7 series of scales; 8 or 9 gill-rakers on the lower part of
anterior arch. Scales 30-31 = Se, 3 between lateral line and
base of anterior rays of soft docal Dorsal X VI 11-12, the
spines slightly increasing in length to the last, which is about
2 the length of head, or the middle spines sometimes sub-
eae soft fin, ly laid back, extending to anterior } of
caudal or bey ond. Anal V 8-10. Pectoral nearly # ‘the
length of head, not extending to above origin of anal ; ventral
extending to origin of anal. Caudal subtruncate or rounded.
Caudal peduncle about 2 as long as deep. Olivaceous, each
scale with a brown spot ; a dark spot on tlie lateral line below
the posterior dorsal spines and another on the upper part of
the base of caudal; about 8 dark cross-bars in the young ;
vertical fins with small dark spots.
American Cichlid Genus Cichlosoma. 329
Rivers of Sinaloa and Jalisco, Mexico.
1. (67 mm.) co-type of Rio Presidio. Dr. D. 8. Jordan.
the species.
2-4. (104-180 mm.) Rio Presidio. Herr A. Forrer.
53. Cichlosoma mento.
Heros mento, Vaill. & Pellegr. Bull. Mus. Paris, 1902, p. 83; Pellegr.
Mém. Soe. Zool. France, xvi. 1903, p. 229 (1904).
Cichlasoma mento, Meek, Zool. Pub, Columbian Mus. v. 1904, p. 207.
Depth of body 2}—2£ in the length, length of head 24-3.
Snout shorter than postorbital part of head. Diameter of
eye 4-4} in the length of head and 1} in the interorbital
width. Maxillary not extending to below the eye; lower jaw
projecting ; anterior teeth forming canines; fold of the lower
lip continuous ; cheek with 6-8 series of scales; 8 gill-rakers
on the lower part of anterior arch. Scales 29-30 =. Dorsal
XVI 10, the spines only slightly increasing in length after
the fifth, the last } the length of head. Anal V 7-8. Pec-
toral 2 the length of head. Caudal rounded. Caudal
peduncle as long as deep. Violaceous; soft vertical fins
spotted.
Southern Mexico.
The types, from the Rio Negro, measure from 98 to
180 mm.
54. Cichlosoma Feste.
Heros Feste, Bouleng. Boll. Mus. Torin. xiv. 1899, No. 335, p. 6;
Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 250 (1904).
Depth of body 22 in the length, length of head 23. Snout
nearly 4 the length of head. Diameter of eye 52 in the
length of head, interorbital width 3. Depth of preorbital
1} the diameter of eye. Maxillary exposed distally, not
extending to below the eye; premaxillary processes ex-
tending to above anterior 4 of eye; lower jaw slightly
projecting ; fold of the lower hp continuous; upper jaw
with the 2 anterior teeth of the outer series differentiated as
a pair of canines ; lower jaw with the anterior 2 or 3 teeth
on each side forming distinct canines ; cheek with 5 or 6
series of scales, with a naked area between the lowest series
and preoperculum (probably covered in the young with
1 or 2 additional series of scales) ; 9 gill-rakers on the lower
part of anterior arch. Scales 30 = 33 between lateral line
and base of anterior rays of soft dorsal. Dorsal X VI (11) 12,
the spines subequal from the sixth to the fourteenth, thence
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 22
330 Mr. C. T. Regan on the Fishes of the
inereasing to the last, which is $ the length of head; the
soft fin, when laid back, extending nearly to middle of
caudal. Anal V (8) 9. Pectoral # the length of head,
extending to above origin of anal; ventral extending to
origin of anal. Caudal rounded. Caudal peduncle a little
more than 3 as long as deep. Brownish, with 8 blackish
cross-bars ; a blackish ocellated spot on the upper 4 of base
of caudal; vertical fins dusky, the soft dorsal with some
indistinct light spots.
W. Ecuador.
1. (255 mm.) one of the types Rio Guayas, Guayaquil. Dr. H. Festa.
of the species.
55. Cichlosoma ornatum, sp. n.
Depth of body 23 (young)—2% (adult) in the length,
length of head 24-24. Snout nearly } the length of head
(in the adult). Diameter of eye 33-6 in the length of head,
interorbital width 3-34. Depth of preeorbital 3-12 the
diameter of eye. Maxillary slightly exposed, not extending
to below the eye; premaxillary processes extending to above
anterior 1 of eye (adult) or a little beyond (young) ; jaws
nearly equal anteriorly (in the young), or the lower a little
shorter than the upper (in the adult) ; fold of the lower lip
continuous ; upper jaw with the 2 anterior teeth of the outer
series differentiated as a pair of canines; lower jaw with the
2 or 3 anterior teeth on each side forming canines; cheek
with 6 series of scales, the lowest deciduous in the adult;
8 or 9 gill-rakers on the lower part of anterior arch.
Scales 30-32 2, 21-3 between lateral line and base of
anterior rays of soft dorsal. Dorsal XV—XVI 12-13, the
spines subequal from the sixth or seventh to the twelfth to
fourteenth, thence increasing to the last, which is about
4 the length of head (3 in the young, 7 im the adult), the
soft fin, when laid back, extending uearly to middle of
caudal. Anal V 9. Pectoral #-2 the length of head, ex-
tending nearly to above origin of anal; ventral extending to
origin of anal. Caudal rounded. Caudal peduncle as long .
or nearly as long as deep. Olivaceous, with some blackish
cross-bars on the sides of the body; sides of the head with
light blue spots ; each scale with a light blue spot, the spots
of the lower part of the body larger and almost white ;
a blackish spot at the base of pectoral and another on the
upper 4 of the base of caudal; 3 large blackish spots on
the basal part of the dorsal fin, the first at the base of the
9th to 1]th spines (absent in one specimen), the second at
American Cichlid Genus Cichlosoma. 331
the base of the last 2 spines and the first soft ray, the third
on the posterior soft rays; vertical fins with light spots.
N.W. Ecuador.
1-3. (118-300 mm.) types of the Rio Durango. Mr. Rosenberg,
species.
4, (119 mm.) St. Javier. Mr. Rosenberg.
In the adult examples the teeth are obtuse, many ending
in a flat circular surface. On dissection the stomach was
found to be full of broken shells.
These specimens had been referred by Boulenger to
C. Feste, but they differ considerably from that species in
coloration and in the more slender caudal peduncle. More-
over, in C. Feste, when the mouth is closed, the anterior
canines of the upper jaw fit in between the anterior teeth of
the lower jaw ; in C. ornatum, on the contrary, the anterior
teeth of the upper jaw are external to those of the lower
when the mouth is closed, this being especially marked in
the adult fish. In C. Feste the occipital region is more
elevated and convex, and this difference is due neither to
age, sex, nor method of preservation, for the largest specimen
of C. ornatum is a male, and in both cases the occipital crest
can be felt just beneath the skin. Young examples of
C. ornatum (118 to 119 mm. in total length) resemble the
adult of C. Feste in so far as they have a deeper body
and longer dorsal spines than have the adult specimens of
their own species, but the differences between adult examples
of the two species will be evident from the following
measurements :—
C. Feste (3). C. ornatum (S),
mm. mm,
Length to base of caudal .......... 205 245
Wepriot Bod y2 ) sficiscgaiv’ vcsart sstech-s 86 86
MICH PB OLMMCAD Oh 2155 anew a\a's aa 4A sad aia 76 87
Length of caudal peduncle .......... 22 30
Depth of caudal peduncle .......... 50 30
ast dorsal spmMen yes. seins he ss) 5,<eloe 28 25
56. Cichlosoma octofasciatum.
Heros octofusciatus, Regan, Revue Suisse Zool. xi. 1903, p. 417,
pl. xiii. fig. 1 ; Ann. & Mag. Nat. Hist. (7) xiii. 1904, p. 258.
Cichlasoma octofasciatum, Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p- 223 (1904) ; Meek, Zool. Pub. Columbian Mus. y. 1904, p. 218.
Cichlasoma Hedricki, Meek. t. ec. p. 208, fig. 66.
Depth of body 2-2? in the length, length of head 23-3.
Snout shorter than postorbital part of head, Diameter of
eye 3-44 in the length of head, interorbital width about 3,
22%
332 Mr. C. T. Regan on the Fishes of the
Depth of preorbital 3-4 the diameter of eye. Maxillary
scarcely exposed, extending to below the anterior margin of
eye; premaxillary processes extending to above anterior
margin of eye or a little beyond; jaws equal anteriorly or
the lower slightly projecting; fold of the lower lip in-
terrupted or subcontinuous ; upper jaw with the 2 anterior
teeth of the outer series more or less distinctly differentiated
as a pair of canines ; lower jaw with the 3 anterior teeth on
each side forming more or less distinct canines ; cheek with
5 or 6 series of scales; 7 or 8 gill-rakers on the lower part
42-52
of anterior arch. Scales 28-31 33", 33 or 4 between lateral
line and base of anterior rays of soft dorsal. Dorsal X VII-—
XIX 8-10, the spines subequal from about the sixth to the
fourteenth, thence increasing to the last, which is about
2 the length of head ; soft fin, when laid back, extending to
middle of caudal or beyond. Anal VIII-X 7-8. Pectoral
3—4 the length of head, extending to above third anal spine ;
ventral extending well beyond origin of anal. Caudal
rounded. Caudal peduncle $—2 as long as deep. Body with
about 8 dark cross-bars, which become obscure in the adult,
the third bearing a blackish blotch below the lateral line ;
sometimes a longitudinal band from eye to lateral blotch ;
usually some light blue spots on the head and one on each
scale of the side of the body ; a dark spot, often ocellated,
on the upper part of the base of caudal; vertical fins with
series of dark spots.
Southern Mexico; British Honduras (? Jamaica).
1-14, 15-18. (61-150 mm.) British Honduras. Rey. J. Robertson.
19-22, (81-107 mm.) Rio de Sarabia. Dr, A. C. Buller.
57. Cichlosoma urophthalmus.
Heros wrophthalmus, Giinth, Cat. Fish. iv. p. 291 (1862), and Trans.
Zool. Soe, vi. 1869, p. 454, pl. Ixxu. fig. 1; Steind. Denkschr, Ak.
Wien, xxiii. 1864, p. 66, pl. v. fig. 3; Jord. & Everm. Bull. U.S.
Nat. Mus. xlvii. 1898, p. 1557 ; Everm. & Goldsborough, Bull. U.S.
Fish. Comm. xxi. 1902, p. 157, fig.; Pellegr. Mém. Soe. Zool. France,
xvi. 1903, p. 258 (1904).
Heros Troscheli, Steind. Sitzb. Ak. Wien, ly. 1867, p. 524, pl. iv. ;
Jord. & Everm. l. c.; Pellegr. /. ¢.
Depth of body 2-23 in the length, length of head 22-27,
Snout as long as eye, the diameter of which is 31-32 in the
length of head, interorbital width 3-38}. Depth of pre-
orbital 2—% the diameter of eye. Maxillary more or less
exposed distally, extending to below anterior margin of eye ;
premaxillary processes extending to above anterior 1 of
eye or slightly beyond; lower jaw slightly projecting ; fold
9
American Cichlid Genus Cichlosoma. 333
of the lower lip continuous ; upper jaw with the anterior
pair of teeth distinctly differentiated as canines ; lower jaw
with the 2 or 3 anterior teeth on each side forming similar
canines ; cheek with 6 series of scales; 8-10 gill-rakers on
the lower part of anterior arch. Scales 28-31 —- 24 or 3
between lateral line and base of anterior rays of soft dorsal.
Dorsal XV—X VII 10-12, the spines subequal from the sixth,
or else slightly decreasing from the sixth to the fourteenth
and increasing again to the last, which is nearly } the length
of head; the soft fin, when laid back, extending to anterior
4 of caudal or beyond. Anal VI 8-9. Pectoral 4-2 the
length of head, extending to above origin of anal or anterior
anal spines; ventral extending to origin of anal. Caudal
rounded. Caudal peduncle about 2 as long as deep. Oliva-
ceous, with 6 or 7 blackish cross-bars ; a blackish ocellated
spot on the upper half of the base of caudal; vertical fins
dusky.
Guatemala; British Honduras; Yucatan.
1-4, (121-171 mm.) types Lake Peten. O. Salvin, Esq.
of the species.
5. (92 mm.) British Honduras. Rey. J. Robertson.
58. Cichlosoma trimaculatum.
Heros trimaculatus, Ginth. Trans. Zool. Soe. vi. 1869, p. 461, pl. Ixxvi.;
Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898, p. 1529; Pellegr.
Mém. Soe. Zool. Frauce, xvi. 1903, p. 239 (1904).
Depth of body 12 (adult)-24 (young) in the length,
length of head 23-3. Snout shorter than postorbital part
of head. Diameter of eye 34-43 in the length of head,
interorbital width 23-33. Depth of preorbital equal to
(adult) or 5 (young) the diameter of eye. Maxillary exposed
distally, extending to below anterior margin of eye; pre-
maxillary processes extending to above middle of eye or
beyond ; lower jaw slightly projecting ; upper jaw with the
2 anterior teeth of the outer series differentiated as a pair of
canines ; lower jaw with the anterior 2 or 3 teeth on each
side the strongest, forming canines; cheek with 5 or 6 series
of scales ; 8 or 9 gill-rakers on the lower part of anterior arch.
Scales 30-51 = 24 or 3 between lateral line and base of
auterior rays of soft dorsal. Dorsal XVI-X VII 11-12, the
spiues subequal from about the sixth to the fourteenth,
thence increasing to the last, which is 4 or nearly 4 the
length of head; soft fin, when laid back, extending to
middle (young) or posterior end (adult) of caudal. Anal
VI-VII1 8-9. Pectoral 3-7 the length of head, extending
334 Mr. C. 'T. Regan on the Iishes of the
to above origin of anal; ventral extending a little beyond
origin of anal. Caudal rounded. Caudal peduncle nearly
4 as long as deep. Olivaceous, with a blackish spot above
the origin of the lateral line, another on the middle of the
side, and a third on the upper part of the base of caudal ;
vertical fins dusky.
Guatemala ; Southern Mexico.
1. (280 mm.) type of the species. Chiapas. O. Salvin, Esq.
2-4, (68-225 mm.) Huamuchal. O. Salvin, Esq.
59. Cichlosoma Salvini.
Heros Salvini, Giinth, Cat. Fish. iv. p. 294 (1864), and Trans. Zool. Soe.
vi. 1869, p. 460, pl. xxiii. fig. 8; Pellegr. Mém. Soc. Zool. France,
xvi. 1903, p. 241 (1904).
Heros triagramma, Steind. Denkschr. Ak. Wien, xxiii. 1864, p. 70,
pl. ii. fig. 2.
Cichlasoma Salvini, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898,
p. 1528; Meek, Zool. Pub. Columbian Mus. y. 1904, p. 207.
Cichlasoma Moja, pe ce. p. 217, fig. 71.
Depth of body 24-2? in the length, length of head 23-22.
Snout as long as ee era part of head (in the adult).
Diameter of eye 3-4 in the length of head, interorbital
width 8-83. Depth of preeorbital 3-7 the diameter of eye.
Maxillary exposed distally, extending about to below anterior
margin of eye; premaxillary processes extending to above
middle of eye or beyond; lower jaw projecting ; ‘fold of the
lower lip continuous ; upper Jaw with the 2 anterior teeth of
the outer series forming a strong pair of canines; lower
jaw with 2 pairs of strong canines; cheek with 5 series of
scales; 8 or 9 gill-rakers on the lower part of anterior arch.
Scales 28-31 = 2 or 24 between lateral line and base of
anterior rays of soft dorsal. Dorsal * XVI-XVII 9-12, the
spines subequal from the sixth to the fourteenth, thence
increasing in length to the last, which is 4 or nearly 4 the
length be head; the soft fin, hee laid “back, extendme
about to middle of caudal. Anal + VIII-IX 7-9. Pectoral
2 the length of head, extending to above anterior anal spines;
ventral extending beyond origin of anal. Caudal rounded.
Caudal peduncle 4-3 as long as deep. Brownish, with
darker cross-bands which form a series of blackish blotches
along the base of the dorsal and another series below the
lateral line, the latter usually confluent in the adult to form
a longitudinal band from eye to caudal; blue spots on the
* Dorsal with 16 spines in 1 specimen only.
+ Anal with 9 spines in 2 specimens only.
American Cichlid Genus Cichlosoma. 339
cheeks ; sometimes a blackish spot on suboperculum ; vertical
fins often with small dark spots.
Young specimens have the cross-bars more prominent and
the blotches less distinct, except for one on the middle of the
side and another on the base of the caudal.
Guatemala; Southern Mexico; British Honduras.
1-3. (86-115 mm.) types Rio de Santa Isabel. O. Salvin, Esq.
of the species.
4-7, (83-120 mm.) Lake Peten. O. Salvin, Esq.
8-10. (159-155 mm.) Nr. Cajabon. F. C. Sarg, Esq.
11-14. (70-92 mm.) British Honduras. Rey. J. Robertson.
Cichlasoma mojarra, Meck, is founded on a very small
specimen with malformed spinous dorsal. I have carefully
compared the smallest examples in the British Museum
Collection with Dr. Meek’s figure, and I am quite certain as
to the identity of C. mojarra with C. Salvini.
60. Cichlosoma multifasciatum, sp. 1.
Heros Friedrichstahli (part.), Giinth. Trans. Zool. Soc. vi. 1869, p. 459.
Depth of body 24-23 in the length, length of head 23-24.
Snout as long as or shorter than eye, the diameter of which
is 8-32 in the length of head, interorbital width 35-33.
Depth of preorbital 1-1 the diameter of eye. Maxillary
exposed distally, extending to below anterior 4 of eye;
premaxillary processes extending to above posterior } of
eye; lower jaw projecting ; fold of the lower lip continuous ;
upper jaw with a pair of stroug canines situated close
together ; lower jaw with 1-2 pairs of canines, which are
well-se} arated ; cheek with 7 series of scales; 8 gill-rakers
on the lower part of anterior arch. Scales 30-31 SS:
24 between lateral line and base of anterior rays ot soft
dorsal. Dorsal XVIII 10, the spines slightly increasing in
length to the last, which is 2 the length of head or a little
more ; the soft fin, when laid back, extending to anterior 4
of eaudal or beyond. Anal VII-VIII 8-9. Pectoral ? the
length of head, extending to above anterior anal spines;
ventral extending beyond origin of anal. Caudal rounded,
Caudal peduncle 4-3 as long as deep. Olivaceous, with
about 10 blackish cross-bars; a more or less distinct dark
stripe from upper part of eye to operculum and another
from lower part of eye to suboperculum; cheeks and
opercular bones, lower part of body and vertical fins with
dark spots.
Guatemala; British Honduras.
336 Mr. ©. T. Regan on the Fishes of the
1 2. (91 and 134 mm.) types Lake Peten. O. Salvin, Esq.
of the species.
5. (68 mm.) Stann Creek, British Rev. J. Robertson,
Honduras.
61. Cichlosoma Friedrichstahli.
Heros Friedrichstahli, Heck. Ann. Mus, Wien, ii. 1840, p. 881; Ginth.
Cat. Fish. iv. p. 294 (1862).
Heros Friedrichstahti (part.), Giinth. Trans. Zool. Soc. vi. 1869, p. 409.
ae motaguensis (part.), Pellegr. Mém. Soc. Zool, France, xvi. 1903,
984 (1904).
Depth of body 23-22 in the length, length of head 23—
2%. Snout only a little longer than eye, the diameter of
which is nearly 4 in the length of head, and ee to the
interorbital width. Depth of preorbital 4 3-3 diameter of
eye. Maxillary exposed distally, extending to below
anterior margin of eye ; premaxillary processes extending
to above posterior 4 of eye; lower jaw projecting ; fold of
the lower lip continuous; upper jaw with a pair of strong
canines situated close together; lower jaw with 2 pairs of
canines, which are well-separated; cheek with 7 series of
scales ; 6 or 7 gill-rakers, sometimes preceded by 2-3 ae
ments, on the lower part of anterior arch. Scales 30 2;
23 between lateral line and base of anterior rays of soft
dorsal. Dorsal XVIII 9-10, the spines slightly mcereasing
in length to the last, which is a little more than it the length
of head; the soft fin, when laid back, extending to anterior
4 of caudal. Anal VII-VIII 8. Pectoral 2 - the length of
head, extending to above origin of anal ; eae extending
to origin of anal. Caudal rounded. Caudal peduncle 2 as
long as deep. Olivaceous, with 7 or 8 irregular dark cross-
bars forming a series of blotches below the lateral line; a
dark stripe from upper part of eye to operculum, and another
from lower part of eye to suboperculum ; cheeks, opercular
bones, and lower part of body with dark spots; vertical fins
with dark spots.
Lake Peten, Guatemala.
1-2. (121 and 129 mm.) Lake Peten. O. Salvin, Esq.
62. Cichlosoma motaguense.
Heros Boleig HeAES, Ginth. Trans. Zool. Soc. vi. 1869, p. 462, pl. Ixxvii.
fig.2; Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898, p. 1534.
Heros motaguensis (part.), Pellegr. Mém. Soc. Zool. France, xvi. 1903,
p- 234 (1904).
pep of body 22-23 in the length, length of head 23-3.
American Cichlid Genus Cichlosoma. 337
Snout longer than eye, the diameter of which is 4—5 in the
length of head, interorbital width 32-34. Depth of pre-
orbital 3 the diameter of eye (young) or equal to it (adult).
Maxillary more or less exposed distally, extending to below
anterior margin or anterior } of eye; praemaxillary processes
extending to above posterior 4 of eye ; lower jaw projecting ;
fold of the lower lip continuous ; upper jaw with a pair of
strong canines, situated close together ; lower jaw with 1 or
2 pairs of canines, which are well-separated ; cheek with 7
or 8 series of scales; 7 or 8 gill-rakers on the lower part of
anterior arch. Scales 82 ;;%5, 2 or 24 between lateral line
and base of anterior rays of soft dorsal. Dorsal X VII-
XVIII 10-11, the spines slightly increasing in length to
the last, which is } the length of head; the soft fin, when
laid back, extending to anterior } of caudal or beyond.
Anal VII-VIII 8-9. Pectoral rather more than % the
length of head, extending nearly to above origin of anal in
the young, but not in the adult ; ventral extending to origin
of anal or beyond. Caudal rounded. Caudal peduncle
2-4 as long as deep. Olivaceous, with dark cross-bands
forming a series of blotches, which may unite to form a
continuous longitudinal band, below the lateral line; a dark
stripe from upper part of eye to operculum, and another
from lower part of eye to suboperculum ; cheeks, opercular
bones, lower part of body, and vertical fins usually with
dark spots.
Guatemala.
1, 2-4, (115-240 mm.) types Rio Motagua. O. Salvin, Esq.
of the species.
5. (137 mm.) Pacific Slope of Cen- Smithsonian Inst.
tral America.
63. Cichlosoma managuense.
Heros managuensis, Giinth, Trans. Zool. Soc. vi. 1869, p. 463, pl. Ixxvii.
fig. 3; Jord. & Everm. Bull. U.S. Nat. Mus, xlvii. 1898, p. 1533.
Heros motaguensis (part.), Pellegr. Mém, Soc. Zool. France, xvi. 1903,
p. 234 (1904),
Depth of body 23 in the length, length of head 22. Snout
only a little longer than eye, the diameter of which is 44-13
in the length of head, interorbital width 31-33. Depth of
preorbital 2-3} the diameter of eye. Maxillary scarcely
exposed distally, extending to below middle of eye; pre-
maxillary processes extending to above posterior margin of
eye; lower jaw projecting; fold of the lower lip continuous ;
upper jaw with a pair of rather strong canines situated
598 Mr. C. T. Regan on the Fishes of the
close together ; lower with 1 or 2 pairs of canines, which are
well-separated ; cheek with 8 or 9 series of scales; 8 gill-
rakers and 2 rudiments on the lower part of anterior arch.
Seales 33-34 a 24 or 3 between lateral line and base of
anterior part of soft dorsal. Dorsal XVIII 10-11, the
spines slightly increasing in length to the last, which is }
the length of head; tle soft fin, when laid back, extending
to anterior } of caudal. Anal VII 8. Pectoral 2 the length
of head or a little more, extending to above origin of anal;
ventral extending to origin of anal. Caudal rounded.
Caudal peduncle a8 as long as deep. Olivaceous. with 7
irregular dark brown cross-bars forming a series of blotches
below the lateral line; a dark stripe from upper part of eye
to operculum, and another from lower part of eye to sub-
operculum ; vertical fins with dark spots.
Lakes Managua and Nicaragua.
1. (187 mm.) type of the species. Lake Managua, Capt. J. M. Dow.
2. (195 mm.) Lake Nicaragua. Smith-onian Inst.
64. Cichlosoma Dovit.
Heros Dovii, Giinth. Proc. Zool. Soc. 1864, p. 154, and Trans. Zool.
Soe. vi. 1869, p. 461, pl. lxxiii. fig. 4; Jord. & Everm. Bull. U.S.
Nat. Mus. ae 1898, p. 1535 ; Pellegr. Mém. Soc. Zool. France,
xvi. 1903, p. 234 (1904).
Depth of body equal to length of head, 23 in the length.
Snout longer than eye, the diameter of which is 4—44 in the
length of head and oes than the interorbital width.
Depth of preorbital —-? diameter of ae Maxillary exposed
distally, extending to below anterior 4 of eye ; premaxillary
processes extending to above posterior margin of eye; lower
jaw projecting ; fold of the lower lip continuous ; upper
jaw with a pair of strong canines situated close together -
lower jaw with 1 or 2 pairs of canines, which are well
separated ; cheek with 10 series of scales; 8 gill-rakers on
the lower part of anteriorarch. Scales 34-35 5 >: 34 between
lateral line and base of anterior rays of soft dei mall Dorsal
XVIII 12, the spines increasing in length to the last, which
is 4 the length of aoe the soft fin, when laid back,
extending to anterior 4 4 of caudal. Anal VI 9-10. Pectoral
2 the length of head, extending nearly to above origin of
anal ; ventral extending to origin of anal. Caudal rounded.
Caudal peduncle 4-3 as long as deep. Brownish, with
darker cross-bars forming a series of blotches below the
lateral line; a dark stripe from upper part of eye to
American Cichlid Genus Cichlosoma. 339
operculum, and another from lower part of eye to sub-
operculum ; vertical fins obscurely spotted.
Lake Nicaragua.
1-2. (140 and 144 mm.) types —— Lake Nicaragua. O. Salvin, Esq.
of the species.
65. Cichlosoma spectabile.
Petenia spectabilis, Steind. Sitzb. Ak. Wien, Ixxi. 1875, p. 96, pl. iv.
Kigenm. & Bray, Ann. Ac. N. York, vii. 1894, p. 615; Pellegr.
Mém. Soe. Zool. France, xvi. 1903, p. 244 (1904).
Depth of body 12-2 in the length, length of head 23-22.
Snout as long as eye, the diameter of which is 34-4 in the
length of head and equal to the interorbital width, Depth
of preorbital } the diameter of eye. Maxillary exposed,
extending to below the middle of eye; premaxillary pro-
cesses extending about to above posterior margin of eye ;
lower jaw projecting ; anterior canines strong ; cheek with
7 or 8 series of scales. Scales 30 2%. Dorsal XV 12-13,
11-12"
the last spine § the length of head, the soft fin in great part
covered with small scales. Anal VI 9-10. Pectoral nearly
as long as the head. Caudal rounded. Caudal peduncle
about 2 as long as deep. A large dark spot on the middle
of the side,’a smaller one on the upper part of the base of
caudal.
R. Amazon.
The types measure up to about 160 mm. in total length.
This species is closely allied to C. managuense and C. Dovii,
and must certainly be regarded as congeneric with them.
66. Cichlosoma Kraussi.
Petenia Kraussi, Steind. Denkschr. Ak. Wien, xxxix. 1879, p. 28,
pl. ii; Eigenm. & Bray, Ann, Ac. N. York, vii. 1894, p. 615;
Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 244 (1904).
Depth of body 21-21 in the length, length of head 23-22.
Snout a little longer than eye, the diameter of which is
3-44 in the length of head, interorbital width 34-4
Depth of preorbital $ the diameter of eye. Lower jaw
projecting; maxillary exposed, extending to below middle of
eye; preemaxillary processes extending to above the anterior
part of operculum ; anterior canines strong. Cheek with
7 or 8 series of scales; 10 or 11 gill- rakers on the lower
part of anterior arch. Scales above the lateral line smaller
than those below it, about 42 in a longitudinal series above,
about 30 below the lateral line; scales in a transverse
340 Mr. G. Lewis on
bie
series == . Dorsal (XV) XVI (10) 11, the last spine a
little less than $ the length of head, the soft fin scaly at the
base. Anal VI (8) 9. Pectoral nearly as long as the head.
Caudal rounded. Caudal peduncle about 3 as long as deep.
Body with obscure dark cross-bars; a dark spot on the
origin of the lateral line, another on the operculum, a third
on the middle of the side, and a fourth on the upper part of
the base of caudal; vertical fins with dark spots.
Colombia; Venezuela.
1-2. (128 and 188 mm.) Baranquilla, Colombia. Kay Thomson, Eq.
[Te be continued. ]
XXXVII.— On new Species of Histeridie and Notices of others.
By G. Lewis, F.L.S.
[Plate X.]
THE object of this paper—the twenty-fifth of the series—is to
make a few observations on the ‘ Catalogue of the Histeridee ”
published last March, to describe a few species of interest,
and to explain the contents of the Plate now published.
In the ‘ Catalogue’ I did not include the “ catalogue-names”
of Dejean or Marseul. Marseul gave over fifty duplicate
names to species in his Catalogue of 1862, because he con-
sidered that no two species of the same family should bear
similar names; but this rule only obtains generally as
regards genera. The names of Dejean and Marseul are
entered in the Munich Catalogue. Some authors, again,
have given varietal names to specimens, often unique, and
therefore names of individuals, not of races; these names
were treated as synonymic.
I think that Marseul did not always attach sufficient
importance in his arrangement of the genera to the form of
the mesosternum in the Histeridz ; he placed, for instance,
Fachycrerus and Phelister, which have a projecting meso-
sternum, between Platysoma and Omalodes, in which the
mesosternum 1s emarginate; but by associating the two
former genera with Probolosternus and Monoplius, as I have
done, the species seem to occupy a more natural position.
When we find an emarginaticn in the mesosternum of a
species of this family we see an insect with a different mode
of life to those in which the mesosternum is acuminate or
bisinuous, and we can understand the several advantages
new Species of Histeridee. d41
attached to these structures. The mechanical action of the
emarginate mesosternum is that a species, such as those
of Hololepta, Fister, and Paromalus, can move the thorax
easily sideways, as the rounded-off base of the prosternum
works freely to and fro in the mesosternal incision ; and
the mechanical utility of a projecting mesosternum with a
corresponding incision in the keel of the prosternum, such as
that in Trypaneus and Pachycrerus, is that by the close
fitting of the segments the one in the other the column of the
body is strengthened in forcing a way in a direct line through
any obstructed passage they may seek to enter. T'rypanwus
especially finds obstacles in the holes of the Platyp: and other
wood-boring species on which they feed. Marseul, in his
monograph, at first placed the “ Trypanéens” after the
“© Sapriniens,’? but in 1862 he installed them after the
*¢ Fololeptiens,” leaving the genus Teretrius in the “ Abrézns” ;
but I think Teretrius and its allies cannot be separated from
the Trypanez.
I have not overlooked the fact that the genus Hister, as
now constituted with its 365 species, has sometimes a truncate
mesosternum, sometimes a sinuous outline, and sometimes it
is markedly emarginate. Thomson founded the genus
Atholus, of which Hister corvinus, Germ., is the type, but he
associated with it other very different species, such as
HT. bimaculatus, L., and these, I think, cannot be set together.
Hister truncatisternus, Lew., from Central America, 1. obtusi-
sternus, Sch., from Hast Africa, and others from widely
distant places have truncate sterna, but are very dissimilar to
H. corvinus. In the genus Hester the mesosternum varies in
form; in other genera, as in Omalodes, it is similar in all the
species.
Hister orientalis, Payk. (Mon. p. 17, pl. xiii. fig. 7, 1811),
a common species in India and Ceylon, was omitted by an
oversight in my Catalogue.
List of Species, arranged generically.
Hololepta obtusipes, Mars. Eblisia tenuipes, sp. n.
Pygoceelis sp. Macrolister striatisternus, sp. n.
Teretriosoma sexuale, Schaeffer. Hister dauphini, sp. n.
Pleesius hamatus, sp. n. Coryphzeus violaceus, sp. n.
asperimargo, sp. n. Notodoma saturum, Lew.
striatipectus, sp. n. Lissosternus gracilipes, sp. n.
Platysoma rime, sp. n. Saprinus sinaiticus, Crotch.
Cylistosoma epilissum, sp. n. Abrezeus fakir, sp. n.
aie Mr. G. Lewis on
Hololepta obtusipes, Mars.
Marseul only knew the female of this species ; the male
has a carina on the mentum, but it has no thoracic fossettes.
It occurs in Java, Sumatra, and in the Malay Peninsula.
Pya@oca is, Lewis.
The first species of this genus came from Togoland,
Kamerun, and German East Africa, and I have now a species
from Madagascar. A genus extending over so wide an area
is likely to contain very numerous species.
Teretriosoma sexuale, Schaeffer, Journ. N. York Ent. Soc.
x11. p. 203 (1905).
This species differs from 7. conigerum, Lew., in the punc-
tuation of the sternal plates, and it is more robust. The
paper containing the description did not reach me in time
for a reference to it in my Catalogue.
Plesius hamatus, sp. 1.
Oblongo-ovalis, subdepressus, niger, nitidus; fronte stria interrupta ;
pronoto stria marginali antice juncta, laterali valida antice
hamata; elytris striis 1 dorsali integra, 2-5 apicalibus, margine
postice punctulata ; propygidio in medio sparse punctato ; pygidio
dense punctato ; prosterno bistriato; mesosterno antice profunde
marginato.
L. 103-11 mill.
This species is similar to P. levigatus, Mars., but it is
larger and broader; the frontal stria is distinctly interrupted
and widens out on either side of the interruption into a shallow
and rather rugose depression ; the thorax is straighter along
the sides than that of leviégatus and the lateral stria or suleus
reaches as far again round the anterior angle; the elytral
striz, the outer humeral is complete and not widened out like
that of /evigatus, the inner humeral is apical and evanescent
near the middle, the first dorsal is complete and less crenulate,
the other striz are Jess defined but do not differ much from
those of lavigatus, the apices are distinctly but rather
narrowly punctulate ; the propygidium is much less closely
punctured in the middle ; the prosternum has a few punctures
on its lobe and the striw are wider apart than those of devi-
gatus; the mesosternal margination 1s closely similar in both
species,
new Species of Histeride. 343
This is the third species known with a deep marginal stria
or furrow in the mesosternum.
Hab. Tongking (A. Weiss, 1901). ‘Two examples in the
Museum of Paris and one in my own collection.
‘Plesius asperimargo, sp. 0.
Oblongus, subconvexus, niger, nitidus ; fronte transversim punctata,
stria obsoleta; pronoto lateribus conspicue rugoso-punctato ;
elytris striis dorsalibus, 1 integra, 2 striato-punctata; propygidio
utrinque grosse, in medio minor, punctato ; prosterno bistriato,
lobo grosse et profunde punctato.
Ee ii mill,
This species differs from P. Mouhot’, Lew., in being more
oblong and less convex, and the sculpture of the lateral
thoracic margin is close and rugose, resembling that figured
for Platysoma frontale, Payk., in Marseul’s Mon. fig. 13
(1853), and the second dorsal stria and the inner humeral
stria are more punctiform. The thoracic punctuation of
P. Mouhoti is scattered and irregular. The general outline
of P. asperimargo is similar to Placodes caffer, Kr. It is the
eleventh species of Plesius now known,
Hab. Burma.
Plestus striatipectus, sp. n.
P. cossyphi simillimis, sed prosterno bistriato et propygidio
pygidioque multo minus punctatis.
L. 93-103 mill.
This species is extremely similar to P. cossyphus, Mars.,
but the frontal striz are less marked, the lateral thoracic stria
is less deep, the first dorsal stria is represented by punctures
at the base of the elytra, the apical margin of the elytra is
less punctulate, the pygidia are finely and more moderately
punctured, and the prosternum is bistriate.
The trivial name points to a very important character.
Hab. New Britain (A. Willey, 1895-97). ‘Two examples.
Platysoma rime, sp. n.
Oblongum, depressum, nigrum, nitidum ; fronte punctulata in medio
impressa ; pronoto lateribus punctato; elytris striis 1-3 integris,
4-5 brevissimis; mesosterno marginato; tibiis anticis 4-dentatis.
L. 33-32 mill.
Oblong, depressed, black and shining; the head, surface
finely punctulate, the transverse stria is somewhat obscure
344 Mr. G. Lewis on
but complete; the thorax, the lateral stria is feebly sinuous
and hamate at the basal angle, anteriorly it continues behind
the head, but is obscurely broken behind the eyes, laterally
the thorax is distinctly punctulate ; the elytra, the humeral
strize are wanting, dorsal 1-3 complete, 4—5 apical and very
short and partly panctiform ; the propygidium is impressed
on either side and its punctures are irregular in size and
placement ; the pygidinm is less distinctly impressed on each
side, but the punctuation is more regular, there is no raised
rim; the prosternum, keel is without striz ; the mesosternum
is marginate and widely sinuous; the anterior tibiz are
4-dentate.
This species, with P. r¢marium, Er., and P. novum, Lew.,
belong to an Indian section of the genus in which the thorax
is punctured laterally. P. rime is distinctly less broad than
P. rimarium.
Hab. N.W. India (#. P. Stebbing). Many examples.
Cylistosoma epilissum, sp. n.
Cylindricum, parum elongatum, nigrum, nitidum; fronte excavata,
stria late interrupta ; pronoto stria marginali integra ; elytris striis
brevissimis, suturali evanescenti ; tibiis anticis 4—5-spinosis.
L. 3 mill.
Cylindrical, somewhat elongate, black and shining; the
head, forehead, and clypeus are excavated and the lateral
stria ceases at the edge of the excavation, the surface has a
few fine punctures and a few large ones irregularly mingled
together ; the thorax, marginal stria is complete and some-
what carinate at the base, the surface is punctured like the
head, and there is a fine antescutellar puncture a little distant
from the edge; the scutellum is narrow, elongate, and trian-
gular; the elytra, outer humeral stria is interrupted in the
middle, inner very short, rather deep, and basal, the first
dorsal is basal and nearly dimidiate, 2-3 are represented by
two punctures at the base, the sutural stria is apical, very
faint, and composed of fine points, which cease near the
middle of the dorsum; the pygidia have fine and large
scattered points; the prosternum is bistriate; the meso-
sternum is widely sinuous, not margined anteriorly, but
laterally there is a well-marked stria; the metasternum is
canaliculate ; the tibize are all 4-5-spinose.
There are two species in the Museum of Paris similar to
the above in structure and sculpture ; one of them is coloured
above like a species of Pachycrerus and the other is some-
what metallic.
new Species of Histeride. 345
Hab. Imanombo, Madagascar (Pr. J. Decorse, 1901). In
the Museum of Paris and my own collection. Six examples.
Eblisia tenurpes, sp. n.
Oblonga, parallela, depressa, picea, nitida; fronte impressa, tenuiter
punctulata ; pronoto antice biimpresso; elytris striis 1-4 integris,
5 basi abbreviata, suturali nulla ; pedibus brevibus.
L. 34 mill.
Oblong, parallel, depressed, piceous and shining ; the head
is very finely punctulate, impressed in the middle, the lateral
stria is well marked, but it does not reach the anterior angle
of the head, the frontal stria is much less distinct and abbre-
viated on either side; the thorax is wider than long, the
lateral stria is sinuous, shallow, and there are slight impres-
sions and rugosities along its course, the stria is continued
behind the head, where it is irregularly crenulate ; the elytra,
there are two fine epipleural strie, but no humeral, the dorsal
strie 1-4 are complete, 5 is parallel to the suture, shortened
before the base, where there is a small puncture; the pro-
pygidium is feebly impressed on either side and its punctures
are shallow, not closely set, and some are oval; the pygidium
is more evenly punctured, punctures are circular and on each
side it is somewhat deeply impressed, with the outer rim
raised ; the prosternum has a fine marginal stria round its
base, but the stria does not pass the cox; the mesosternum
‘is emarginate behind the keel and has a fine marginal stria;
the anterior tibize are 5-dentate, the tarsal groove is short
and shallow, and not curved, corresponding in length to the
tarsi, which are all somewhat short ; the legs are rather long
and slender.
I cannot compare this species with any other known. It
is convenient at present to place it in the genus dlisia on
account of its tarsal grooves; but Hdlis¢a and its members
require revision.
flab. Yunnan (Donckier). One example.
Macrolister striatisternus, sp. n.
Oblongo-ovatus, convexiusculus, niger, nitidus; fronte levi, stria
integra antice subrecta; pronoto striis lateralibus integris, stria
interna haud interrupta ; elytris striis 1-3 integris, suturali basi
abbreviata; propygidio pygidioque dense punctatis ; prosterno
inter coxas bistriato; mesosterno emarginato; tibiis anticis
3-dentatis,
L. 11 mill.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi.
bo
Cc
346 Mr. G. Lewis on
Oblong-oval, rather convex, black and shining; the head
is smooth, with the frontal stria complete and nearly straight
anteriorly and somewhat lightly impressed ; the thorax is
slightly ciliate on the lateral edges, the two lateral strie are
parallel to each other and well marked, the inner stria con-
tinues, and is rather fine, behind the head; the elytra, striz,
inner humeral apical and dimidiate, 1-3 complete and strong,
with the edges feebly crenulate, 4 short, apical, and almost
obsolete, sutural fine, shortened apically, and scarcely reaches
the middle; the pygidia are densely punctured ; the prosternum
has two striee between the coxee, which widen out behind, but
do not reach the base, and apparently join base anteriorly ;
the mesosternum is emarginate and the stria is complete and
rather fine, and it does not follow the outline of the emargina-
tion ; the anterior tibiz are strongly 3-dentate.
In outline this species agrees with J. ¢ntrepidus, Lew.,
but the dense punctuation of the pygidia and the prosternal
strie are good distinguishing characters.
Hab. Bihé, Angola.
Hister dauphini, sp. n.
Breviter ovatus, convexiusculus, niger, nitidus; stria frontali
integra antice subrecta; pronoto stria laterali externa brevi,
interna integra; elytris striis validis, 1-3 dorsalibus integris,
humerali interna dimidiata, ceteris nullis ; propygidio pygidioque
parce punctatis; mesosterno emarginato, stria integra; tibiis
anticis 3-dentatis.
L. 8 mill.
Shortly oval, somewhat convex, black and shining ; the
head impunctate and very feebly impressed on its vertex,
siria complete and strong, almost straight in front; the
thorax smooth, outer lateral stria is short and confined to the
anterior angle, inner complete, somewhat deep and a little
sinuous, continued and somewhat crenulate behind the head ;
the elytra, inner humeral stria strong, apical, and dimidiate,
1-3 dorsal similarly strong but complete, 4 is represented by
an apical puncture, all the striz have crenulate edges; the
propygidium and pygidium are moderately punctured; the
prosternum is smooth; the mesosternum is emarginate in
the middle, with a rather fine marginal stria; the anterior
tibie are strongly 3-dentate.
This species is considerably larger than H. sikore, Lew.,
also from Madagasca:.
Hab. Diego Suarez, Madagascar.
new Species of Histeridee. 347
Corypheus violaceus, sp. n.
Oblongo-ovatus, subconvexus, violaceus, nitidus, haud_pilosus ;
pedibus antennisque brunneis; fronte in medio foveolata ; pro-
noto lateribus conspicue rugoso-punctato ad angulos levibus ;
elytris striis humerali interna brevi arcuata, dorsalibus 1-3
integris ; pygidio leviter punctulato; prosterno bistriato.
L. 6 mill.
Oblong-oval, somewhat convex, violet-blue, shining, not
pilose ; legs and antennw obscurely brown, with the thighs
slightly bluish; the head is foveolate on its vertex, with the
frontal stria strong and complete and angulate at the eyes,
surface finely punctulate except in the fovea and in two
anterior impressions, where the points are large; the thorax,
median area very finely punctulate, with a broad lateral band
of “ grater-like ” punctures, the punctures continue narrowly
behind the neck, but the anterior angles are free of points ;
the elytra, surface finely punctulate, striz, inner humeral
well-marked, short and bent, dorsal 1-3 complete and rather
strong, 4 traceable throughout but punctiform on the posterior
half, 5 obsolete, sutural wanting ; the pygidia are very finely
punctulate, with a few larger points in the middle of the base
of the propygidium ; the mesosternum is obscurely marginate ;
the prosternum, anterior lobe markedly sinuous on the ante-
rior edge, bistriate, striz meeting anteriorly.
Corypheus Wallacet, Mars., the type of the genus, is from
. Dorei, N. New Guinea, and it is also recorded by Marseul
from the Fly River; it has five dorsal striz complete and
the sutural stria is entire also, but not well marked at the
base. I have a fourth species of this genus, but the specimen
is not in good condition and the locality is not known to me.
flab, Mount Kina Balu, N. Borneo.
Notodoma saturum, Lew. (Pl. X. fig. 9.)
This species is figured chiefly to show the metasternal
arched stria. WV. bullatum, Mars., has no such stria, and in
N. nigrum, Lew., the only other dark-coloured species
known, the metasternal stria is represented by punctures
only. In Marseul’s description of N. globatum, the type of
the genus, this important character is not noticed, nor is it
given in his figure of the species.
LISSOSTERNUS, gen. nov.
Body ovate, convex ; head retractile; mandibles large and
incurved, acuminate at the tips; antenn, scape dilated,
23%
348 Mr. G. Lewis on
joint 1 globose, 2-7 small and gradually becoming more
transverse, club oval and somewhat elongate ; thorax widest
at the base, suleate laterally; elytra feebly striate; pro-
pygidium rather wide and semicircular in outline ; pygidium
nearly circular ; the prosternum, the keel is relatively wide
and the base has an arched outline to receive the meso-
sternum, which is obtusely projected ; the legs are long, with
the tibiz slightly dilated, tarsal grooves scarcely traceable.
Lissosternus gracilipes, sp.n. (Pl. X. fig. 10.)
Ovatus, convexus, rufo-brunneus; fronte antice sinuata; pronoto
minute tuberculato, stria laterali valida impressa; elytris striis
tenuissimis ; prosterno yix lato, bistriato.
L. 1 mill.
Oval, convex, reddish brown, and shining; the head is
sinuous anteriorly in outline ; the thorax, anterior edge bi-
sinuous, lateral stria well marked, with the outer edge clearly
punctured, on the vertex there are a few minute tubercles,
the four in front of the scutellum are set transversely; the
elytra, the dorsal stria are very faint and shortened, except
the sutural, which is more marked and complete; the pro-
pygidium is semicircular posteriorly, with small tubercles
along the edge and some less defined on its surface; the
prosternum is rather wide, the keel is smooth and polished,
with two lateral strise, which widen out towards the base, the
anterior lobe has shallow punctures ; the mesosternum, along
the anterior edge are a few punctures and on either side a
very fine stria which terminate in a small excavation behind
the acumination, laterally there are two very fine striz and
there is no visible suture to it and the metasternum.
_ The figure of this species is an excellent drawing ; the
insect is probably formicicolous.
Hab. Bahia.
Crotch reported a species of Saprinus from Syria in the
‘Ordnance Survey of Sinai,’ published at Southampton in
1869, in these words :—
“ Saprinus sinatticus, sp. n. The only difference that I
can discover, from description [between it and S. figuratus,
Mars. |, is in the punctuation of the thorax ; in figuratus there
are three separate defined smooth patches, here all three are
united into one, showing only faint indentations to indicate
its possible division.”
At present it is best to consider S. s¢naiticus a nondescript.
The prosternal strie in §. figuratus are so peculiar that I
new Species of UHisteride. 349
think Crotch would have found good specific characters in
the sterna if his species was really distinct ; but there is no
evidence that Crotch examined the undersides of his specimens.
Abreus fakir, sp. n.
Circularis, convexus, niger, subnitidus, supra dense subrugose punc-
tatus, setulis erectis seriatim; antennis pedibusque obscure
rufo-brunneis; fronte punctata haud impressa ; mesosterno
postice crenulato, laterali marginato; metasterno in medio
tenuissime canaliculato.
L, 13-13 mill.
Body nearly circular in outline, convex, black, densely and
somewhat rugosely punctate, with erect sete arranged in lines
on the upper surface ; the head is less coarsely punctured than
the thorax and is not impressed on its vertex ; the thorax,
the basal transverse line is vaguely shown by the disposition
of the punctures; the propygidium is setose, like the elytra,
but the pygidium is without sete and the punctuation is
smaller and clearer; the prosternum is wide and nearly
straight at the base; the mesosternum is crenulate along its
basal edge, the crenulations are about twelve in number, it is
marginate laterally, and the marginal stria continues half
down the metasternum; the metasternum is very feebly
canaliculate longitudinally in the middle ; all the sterna have
simmar large punctures, which are somewhat densely set.
This species may be placed near A. coheres, Lew., but it
is smaller and has a crenulate edge to the base of the meso-
sternum. Paykull’s Indian species extlis has a fine dorsal
punctuation, but it is a species I have not seen.
Hab. Khandesh District, India (T. R. Bell, 1903).
Many examples.
EXPLANATION OF PLATE X.
. Chalcurgus brevipennis, Lew.
. Notodoma saturum, Lew.
rig 10. Lissosternus gr acilipes, sp. n.
Fig. 11. Saprinus inversus, Lew. | Outlines of the sulci in the pygidia of
Fig. 12. aterrimus, Er, the females.
Except for the new species the references will be found in the
Catalogue published last March,
Fig. 1. Hololepta Fee, Lew.
Tig. 2. Lioderma intersectum, Lew.
fig. 3. Hister aino, Lew.
tig. 4. latistrius, Lew.
Lig. 5, Pachycrerus sulcicollis, Lew.
fig. 6. Ditto. Side view of pygidia.
Ing. 7. Pachycrerus princeps, Lew.
8
9
350 Dr. A. G. Butler on
XXX VITI.—WNotes on the Satin Bower-bird (Ptilonorhynchus
violaceus), By ArtHur G. BuTier, Ph.D., F.L.5.,
HZ 5, ee:
In a paper which I published in the ‘ Zoologist’ for July,
1902, p. 257, I called attention to a statement made by
Mr. A. A. C. Le Souéf, Director of the Zoological Gardens
at Melbourne, and stated that the idea that the males of this
species only assumed their full plumage in old age was con-
trary to all our experience of bird-life ; and on pp. 251, 252,
I suggested that the birds studied by Mr. Le Souéf were not
cocks, but old hens which had assumed male plumage owing
to abortion of the functional ovary, a circumstance not
uncommon in birds in captivity, and possibly less so than in
wild birds,
Mr. Le Souéti’s statement is as follows (vide Campbell’s
‘Nests and Eggs of Australian Birds,’ p. 192, footnote) :—
‘“‘ Many years ago I caged a number (at least a dozen) of these
birds at the gardens here, young green birds, caught at
Gembrook, and it was only after the expiration of nearly
etyht years they began to change colour. I think four or five
birds put on the beautiful blue-black plumage, and in a year
or two died off. It is therefore evident that the birds only
come to their full plumage in old age, and that accounts for
the fact that in a flock of say one hundred birds, which we
often used to see at Gembrook, some years ago, there would
be only a very few, not half a dozen, black ones among them.
They die off shortly after the change.”
It is well known that during certain months of the year
many birds travel about in flocks consisting wholly of one
sex, which would explain the occurrence of only six per cent.
hens in male plumage in a flock, without the necessity for
arriving at so improbable a conclusion as that the male of
this species (unlike all others) assumed its attractive plumage
towards the end of its life.
My own experience of Ptilonorhynchus is directly opposed
to Mr. Le Souéi’s view:—On September 18th, 1899, I
purchased a pair of young Satin Bower-birds in the spotted
green plumage, which vaguely resembles the adult hen
plumage (only the latter is not of the same pale-spotted
character). By the end of the following September (1900)
both sexes had attained their full adult plumage—the male
blue-black, the female olive-green, cinnamon, and yellow ;
both sexes with brilliant ultramarine eyes.
The late Mr. Abrahams was of opinion that this species
assumed its full plumage when three years old; but judging
the Satin Bower-bird. 351
by the size of the birds when I bought them and the slight,
though perfectly noticeable, increase of size during the year
in which they emerged from their nestling dress, [ am more
inclined to believe that the adult feathering is perfected by
the end of the second year.
If Mr. Le Souét’s view were correct, my cock bird ought
to have died in 1901 or 1902, yet we are well advanced in
1905, and he shows not the least symptom of failing vigour ;
but, how about the hen?
In the years 1901-4 the cock bird frequently picked up
sticks and straws and pursued the hen, singing vigorously
and going through all the comical contortions and wing-
flappings, side-jumps and dances, peculiar to the species ;
occasionally the hen would chase the cock, with very similar
actions and song: but the aviary in which I kept the pair was
too small for them to make up their minds to serious nest-
building, although they made several attempts to construct a
bower by driving twigs into the earth in two rows converging
at the top.
In 1904 the cock bird began to render the life of the hen
unpleasant by frequently pulling out her flight-feathers.
After pursuing her for a time, if he succeeded in catching
her upon the earth, she would immediately turn upon her back
and oppose him with beak and claws: he would then lie on
his side close to her; and as she turned each would grasp the
other’s claws and pluck viciously at the wings and breast. [
never knew any other bird to fight in this curious fashion.
In the present year the tables were completely turned, the
hen had perfect mastery over the cock, and plucked every
feather out of his wings and tail, so that I had to cage him up
to preserve his life, as the constant drain on his strength was
beginning to make him weak on his legs. In July the hen
began to assume male plumage, which is rapidly developing ;
tlle median and greater coverts are already black, forming a
broad belt across the wings, and the underparts are splashed
with black blotches. It will not surprise me if this bird dies
within the next year or two, but the cock shows no indications
of impending dissolution.
The difference in plumage between nestling and adult birds
is much greater than Gould’s description would lead one to
suppose, the under surface not only being yellower, but
having a distinctly pale-spotted character, owing, perhaps,
to greater width and diffusion of the crescentic markings on
the feathers; moreover, this difference is not confined to the
males, but is, I believe, found in all young birds: I have
examined eight or nine at various times.
352 Canon A. M. Norman on Cucumaria Montagui.
XX XIX.—On Cucumaria Montagu, Fleming.
By Canon A. M. Norman.
[Plate XI.]
In 1808 Montagu described from the coast of Devon a
Holothurian which he named Oucumaria pentactes, var.
In 1828 Fleming, in his ‘ Hist. Brit. Anim., named this
Uolothurian Cucumaria Montagut.
In 1841 Forbes, in his ‘ Hist. Brit. Starfishes, &c.,’ made
Cucumaria Montagui a synonym of Cucumaria pentactes ;
and the left-hand figure of his woodcut possibly represents
the species of Fleming. In 1892 this lead was followed by
Jeffrey Bell in his ‘Cat. British Echinoderms in B.M.’
In 1893 I published a paper “ Cucumaria Montagui (Flem-
ing) and its Synonymy” (Ann. & Mag. Nat. Hist. ser. 6,
vol. xii. p. 469). In that paper I described what I believed
to be this species with great minuteness, especially as regards
the spicula; and inasmuch as I found considerable differences
between the spicula of the largest and the smaller examples,
those differences were pointed out. I presumed that the
smaller specimens were the young state of the larger, though
I purposely abstained from making the direct statement that
they were so, leaving it to future observers to clear up this
matter.
Mr. S. Pace has recently written a paper entitled “ Note
on two Species of Cucumaria from Plymouth, hitherto con-
fused as C. Montagui, Fleming: C. Normand, n. sp., and
C. saxicola, Brady and Robertson” (Journ. Mar. Biol. Assoc.
vol. vii. Dee. 1904, p. 305). I much regret the publication
of that paper, and did what I could to bring Mr. Pace to my
view as regards my specimen A (= C. Montagut) ; and when
he wrote to ask if he might name it after me, my reply was
that of course he could do so if he wished ; but I feared it
would necessitate my committing the act of “ Felo de se.”
That act I am about to be guilty of. I cannot call it “ happy
despatch ” ; for I unwillingly criticize Mr. Pace’s paper as I
am compelled to do.
Mr. Pace writes: “ [Holothuria Montagui, Fleming, affords
a remarkable instance of how much confusion may gather
around a specific name; the species was itself founded on a
misconception, and almost every author who has since made
use of Fleming’s name, or who has attempted to unravel its
synonymy, has but made matters rather more involved.”
Janon A. M. Norman on Cucumaria Montagui. 353
Yet still more involved by the publication of Mr. Pace’s
paper.
Mr. Pace writes: ‘ In 1808 Montagu described and figured
as ‘ Holothuria pentactes, var., a species of Cucumaria which
he had found on the south coast of Devonshire. This species
from Montagu’s excellent description must certainly have
been one of the two forming the subject of this paper; and
which has among other names been known as C. Montagut,
Fleming. Now whatever C. Montagui really may be, it
certainly is not conspecific with Montagu’s supposed variety
of C. pentactes. Of course, it was undoubtedly Fleming’s
intention to honour Montagu by bestowing his name upon
the species discovered by that naturalist, but, unfortunately,
the description of C. Montagut is based. upon specimens of
another species collected in the Firth of Forth. Fleming
makes reference under the name Montagui to Montagu’s
description of the Devonshire Cucumarian, but Montagu’s
actual form is probably the one that is described as Ho/othurta
pentactes in the ‘ History of British Animals.’ Fleming’s
name appears to have dropped into almost complete disuse
until revived by Dr. A. M. Norman. Norman, however,
overlooked the fact that Fleming was not dealing with the
same species as Montagu: and in addition he has failed to
realize that there are two species of Cucumaria of somewhat
similar outward appearance living upon the South Devon
coast. Norman’s Cucumaria Montagui is, in fact, a complex,
and it is mainly as a result of this that: subsequent authors
have experienced so much difficulty in reconciling their ideas
as to the identity of C. Montagui.”
It is inconceivable to me how such a view of Fleming’s
species could have been entertained by Mr. Pace in the
presence of the passages I now proceed to quote from the
descriptions of Montagu and Fleming. From the former
author the passages only are given which bear upon the
description of Fleming; but the passage from Fleming
includes all that he wrote upon the subject, and embraces all
that is important in the description by Montagu.
Montagu, 1808.—Cucumaria pentactes, var. “ Cylindric,
white, covered with a mottled film or epidermis... The
anterior end for an inch or more is of a purplish-brown, and
furnished with eight large and two very small contiguous. . .
tentacula. . . When the animal was alive it was observable
that one of the least arms, or tentacula, was always covering
the mouth, and for that purpose were alternately in motion
... the lips or margin of the aperture white ; the posterior
304 Canon A. M. Norman on Cucumaria Montagu.
end is furnished with a small pentangular opening of a red
colour.”
Fleming, 1828.—JHolothuria Montagu. ‘ Tentacula eight,
with two small contiguous subsidiary ones. Mouth simple.”
(Here reference to Montagu.)
‘The body is cylindric, white, covered with a mottled cuticle ;
towards the head the whole becomes purplish-brown ; margin
of the mouth white; while alive the two small subsidiary
tentacula are alternately in motion, covering the mouth ;
vent pentangular, red. The divisions of the tentacula are
not so numerous as in the preceding species.”
Thus the whole of the description of the species is actually
a quotation from Montagu down to the word “red,” and
then, as though Fleming intended to signify this fact, the
first full stop occurs after “red”; and a capital letter com-
inences the sentence “ The divisions of the tentacula are not
so numerous as in the preceding species.” He gives a locality
cc Hinthseot | dlorths Mir..iNeley ss sbutosut may “be questioned
whether Fleming ever himself saw a specimen which he
would refer to Montagu’s Holothurian, for the sentence
respecting the tentacula might well be written from a com-
parison of Montagu’s figure with the tentacles of the species
which he (Fleming) called Holothuria pentactes. Mr. Pace
writes of ‘ Montagu’s excellent description”; but “ the
description of C, Montagut is based upon specimens of another
species collected in the Firth of Forth.’ It is unnecessary
to write more.—Cucumaria Montagut (Fleming) stet. Cucu-
maria Nermant, Pace, dele.
In my former paper I carefully described the spicula of
Cucumaria Montagut. That description need not be repeated ;
but I have here given illustrations of the spicula, both lower
and upper, of the dermis together with those of the feet and of
the tentacles. Fig. la, 0, and c (Pl. XI.) are normal and
characteristic examples of the spicula of the lower layer.
Specimens B and C,
What are the specimens which in my paper I called B and
C? Are they the young of C. Montagu, or are they a distinct
species as Mr, Pace considers them to be? Pl. XI. fig. 5
represents a specimen, slightly enlarged, which was sent to
me from Plymouth by Mr. Pace himself. Fig. 6 represents
the body-spicula and fig. 7 the spicula of the feet. Of these
figures, fig. 6 a, 6, c represent the spicula with six openings
which distinguish this condition, and are not found, so far as
1 have observed, in full-grown C. Montagu. g, h, 2, k are
«
Canon A. M. Norman on Cucumaria Montagui. 355
irregular forms ; c, d, e spicula with openings as in C. Mon-
togui, cand d showing also the characteristic nodulation.
Mr. Pace suggests that the nodulous spicules had got into
my preparation by accident. This might have been, but was
not the case, as is proved by the fact that whereas the spicule
fig. 6d is one from my original slide of spicula of speci-
men B, fig. 6 e is drawn from a mounting of the spicula of the
specimen fig. 5 which was sent to me by Mr. Pace himself ;
but only very few of these spicula are present in either
specimen.
But while Mr. Pace would reject these nodulous spicula
with four perforations he substitutes for them another; and
says that in specimens similar to my B and C he has found
‘ quite typical ‘ tables,’ which being few in number are easily
overlooked.” They have been overlooked by myself though
diligently sought for, since, if found, they would form an
entirely new feature in the spiculation of Cucumarians, for
there would exist two distinct types of primary body- spicules
in a species, a feature hitherto unknown. May I suggest to
him a source of possible error? When Iwas at Plymouth in
Sept. 1903, two species were brought to me as the two
Cucumarians which besides C. dactea were known at the
Station. On examining one of these and boiling down the
spicule I found the species was no Cucumarian, but Phyllo-
phorus Drummond?, W. Thompson, with its characteristic
“tables.” Perhaps spicules of this species have by some
accident got into Mr. Pace’s mounting; and it is even possible
that he may have employed the very test-tube in which JI had
boiled the £/yllophorus and which may not have been cleaned
with sufficient care *.
The question remains, Are the specimens which I described
as B and Ca mature form, and, if so, are they referable to
Cucumaria saxicola of Brady and Robertson? Let us take
the last part of this question first. I did not neglect to take
C. saxicola into consideration when engaged on my former
paper ; and I wrote to Professor Brady to enquire if he would
allow me to see the type or a mounting of its spicula, but he
no longer had either in his possession. Examining his
description and figures, I came to the conclusion that it was
uot the same as my Band C: Ist, because of the presence
of the stellate-formed upper spicules ; and 2nd, because I saw
no reason to regard it as the same species as my specimens,
* T left a mounting of the spicules of Phyllophorus Drummondi at the
Station together with the specimen itself, but I see that the species is
not inserted in the list of Plymouth Echinoderms published at the end of
last year (Journ. Marine Biol. Assoc. vol. vii. p. 2U6).
356 Canon A. M. Norman on Cucumaria Montagui.
any more than to refer it to the young of C. Planci, Maren-
zeller, to which latter species it appeared to me more likely
to be referable *.
As I told Mr. Pace gn Litt., when young specimens of
C. Montagui, say 14 mm. long, should be found having
spicules agreeing with those of the adult, my view that
B and C are young forms of that species would require to be
reconsidered,
The colours of the two forms are stated by Mr. Pace to
somewhat differ, but not materially. The podia are, he says,
more numerous and in two parallel rows in Montagui, but in
B and C so few as to appear in a single series; this is a
strong point in favour of the latter being immature t+. Again,
he writes: “In sp. 1” (= Montagut) “ the test, being densely
crowded with spicules, is very tough and coriaceous, and its
surface is much wrinkled ; while in sp. 2”? (=my B and C)
“the surface of the body is extremely smooth and delicate,
marked only with transverse strize due to the encircling fibres
of the superficial layer.” Here, again, is a difference such as
is usual and might be expected between mature and immature
examples.
But the main question still remains to be answered: On
what grounds do I presume to suggest that B and C, the
majority of the spicules in which have six perforations, are the
young of a species in which the number of perforations is
almost invariably four, and never six ?
Ist. In Montagut the plate-spicula form a dense regular
layer, over which are developed the campanulate
spicules, but in B and C the spicules are not disposed
in a layer. They are comparatively very few in
number, and are developed at different depths in the
* The spicules of C. Planci are figured by Jeffrey Bell, Cat. Brit.
Echin. B. M. pl. ii. fig. 2; but it is, I think, obvious that there is an
error in the connecting-lines to figures on the plate, and that the three
bottom figures lettered a and also the feet-spicules lettered b on the
middle of the plate, which are joined by connecting-lines with the
spicules of C. Planci, belong to C. Hyndmanni figured on the same plate,
the feet-spicules of which species are peculiarly characteristic. Specimens
in my collection have been examined from Trieste (cotype from Von
Marenzeller), Naples (specimens received many years ago, when the
Holothurians were not well known there, from the Zool. Stat., under the
names C. communis and C. doliolum), Gairloch, Firth of Clyde (Dr. D.
Robertson).
+ It will be seen by my figure 5 (which, slightly magnified, was sent
to me by Mr. Pace, and is the largest specimen known to me having the
spicula of B) that the podia are in double rows, thus showing signs of
more mature growth.
Canon A. M. Norman on Cucumaria Montagui. 357
dermis. This is a difference known to me from the
study of the old and young in other species.
2nd. It is a known fact that spicula found in the young of
certain species (e. g. C. frondosa) entirely disappear
in the adult, or a spicule of another form takes its
place.
3rd. Spicules which are entirely free from nodulation in
some young Cucumarians become highly nodulous or
greatly incrassated in old examples. I will take as
an instance a species I have already referred to,
C. Hyndmanni. The plate-spicules as figured by
Bell, pl. ii. fig. 1a, and also a at bottom of the
plate, show from two to sixteen perforations and are
searcely thickened. I should not have been able to
say positively that these spicula belonged to C. Hynd-
manni had it not been for the peculiarly characteristic
foot-spicules; for the plates of the old specimens are
often all extraordinarily massive, their thickness being
subequal to the breadth; yet other specimens I have,
especially a young one, which have spiculation corre-
sponding with Bell’s figures.
4th. The ventral ambulacra in both A and B bear closely-
set double rows of feet ; dorsal ambulacra with few
irregularly arranged papille or feet. Thus some
authors would place the species in the subgenus
Colochirus.
To sum up my views as regards B and C: I am of opinion
that the figures I give, fig. 6 a, b, f, g, h, 1, k, represent the
primary spicules of the young scattered at depths in the
dermis, which will at a later stage be absorbed; while ce, d,
and e represent some of the earliest spicules developed of
those which will at a later stage constitute the dermal
layer.
I may add that I have studied the spiculation of the Holo-
thuroidea for at least forty years ; that I have mountings of
spicula (often many mountings) of almost all Huropean
shallow-water species and some abyssal forms; and that
while recognizing spiculation as the truest guide to the
determination of species, I know full well that allowance should
be made for variations in respect of individuals and also in
regard to age; and that on these points there remains a
valuable field for study.
After the foregoing was actually printed I happened to
turn to EK. von Marenzeller, ‘ Résultats Camp. scient. Prince
358 Canon A. M. Norman on Cucumaria Montagui.
de Monaco, Contrib. 4 Etude des Holothuries de Atlantique
Nord, 1893, p. 15, and there, under Cucumaria Montagui,
found a reference to a paper of Prof. Jeffrey Bell which had
escaped my memory and to which no reference is made in
that author’s more recent publication ‘Cat. Brit. Echin, in
B.M’ On turning to this paper by Bell, “ On the Spicules of
Cucumaria Hyndmanni and calcigera and two allied Forms,”
Journ. R. Micros. Soc. ser. 2, vol. iii. 1883, I learnt that
types of Montagu’s Cucumarian were in existence in the
British Museum.
I at once went up to the Museum, and with the kind
assistance of Professor Bell examined these interesting speci-
mens, with the following most satisfactory results. The
names and other remarks which I enclose in inverted commas
are in the handwriting of Leach, while the name annexed
has been written on a more recent label.
Bottle 1. “ Holothuria Montagui, 8. Devon, G. Montagu,
Esq., Mus. Leach.” (C. Planez.
This is a large specimen, about six inches, but I
did not measure it. It is my A, with extremely
numerous four-holed spicules in the lower layer, and
with plenty of the characteristic campanulates in the
upper.
Bottle 2. “‘ Holothuria decollata, South Devon, G. Montagu,
Esq., Mus. Leach.” Pentacta Montagut, Gray.
This is a much smaller specimen, agreeing with my
B in spiculation, but smooth four-holed and dumbell
forms most frequent, only a few with six perforations.
Bottle 3. “ Cucumaria communis, South Coast of Devon,
Mus. Leach.” C. Planci.
The spiculation of these three or four small speci-
mens is In entire agreement with my form B.
Thus striking facts confirmatory of my views come out :—
Ist. That at each of three places where adult C. Montagui
have been found—South Devon (Aontagu), Polperro (Norman),
Plymouth (Pace)—there also small specimens have been
found with the spiculation of my B and C.
2nd. That in the British Islands Cucumarians with the
spiculation of B and C have only been found at these three
places.
3rd. That no small examples are known to occur having
the same spiculation as the adult C. Montagu.
I give in the list below some synonymy, but do not refer
to continental authors, leaving them to reconcile differences
now that they will have full illustrations of the spicula of
C. Montagut. In 1886 I sent specimens agreeing with B and
Canon A. M. Norman on Cucumaria Montagui. 359
C to the Vienna Museum as C. Montagui, for I had only one
adult example in my collection, and therefore Marenzeller’s
writings refer to these younger stages, and not to the adult.
Koehler states that he had received a specimen from
Plymouth under the name “ C. pentactes”’ ; this he rightly
referred to C. Montagu?, but he would have been more correct
had he written the young of C. Montagui.
Cucumaria Montagu?, Fleming.
1808. Holothuria pentactes, var., Montagu, Linn. Trans. vol. ix. p. 112,
pl. vii. fig. 4.
1828. Holothuria Montagui, Fleming, Hist. Brit. Anim. p. 483.
1848. Pentacta Montagui, J. K. Gray, List Brit. Animals in B.M. pt. i.
Centroniz, or Radiated Animals, p. 11.
1893. Cucumaria Montagu, Norman, Ann. & Mag, Nat. Iist. ser. 6,
vol. xii. p. 469,
1904. Cucumaria Normani, Pace, Journ. Marine Biol. Assoe. vol. vii.
p. 305.
The Young.
1848. Holothuria decollata, Leach, MS. name only, J. E. Gray, under
Pentacta Montagui as above referred to.
1883. Holothuria decollata=C, Montagut, Bell, Journ. R. Mier. Soc.
ser. 2, vol. iil. p. 3 (separate copy), pl. viii. figs. 4, 4 a.
1893. Cucumaria Montag, specimens B and C, Norman, /. c.
1893. Cucumaria Montagu, Marenzeller, Résultats Camp. scient.
Prince de Monaco, Contrib. 41’ Etude des Holothuries de l’A tlantique
Nord, p. 15, and Ann. & Mag. Nat. Hist. ser. 6, vol. xii. p. 335,
1904. Cucumaria saxicola, Pace (nec Brady and Robertson), /. e.
Of synonyms which I gave in my former paper, Semperia
Drummondii, Hérouard (nec Thompson), has spicules exactly
like those of adult C. Montagui; but C. Lefevre’, Bonnier, is
stated by Koehler not to have the feet arranged as in Colo-
chirus, and therefore to be different from C. Montagui, and
Marenzeller has made it a synonym of C. Koellikeri. I may
add that Marenzeller refers Colochirus Lacazei, Hérouard, as
a synonym to C. Montagut; but I do not think that this can
possibly be the case if fig. 8 of Hérouard correctly represents
one of the upper layer of spicules, and, moreover, there seems
to be considerable difference in other spicules also.
EXPLANATION OF PLATE XI.
» Cucumaria Montagui (Fleming). Spicules of the lower layer of
the adult, a, 6, c being the normal forms.
Ditto. Spicules of sides of feet.
Ditto. Spicules of tentacula.
Ditto. Campanuliform spicules of upper layer of dermis.
Ditto. Immuature “B and C,” slightly enlarged,
Ditto. Dermal spicules: a, b, f, 9,h, 7, k, primary spicules chavac-
teristic of the young; c, d, e, secondary spicules characteristic of
the adult layer.
Fig. 7, Ditto. Spicules of the feet.
I
—
—
hy
a
G2 SUES CORD
360 Canon A. M. Norman on Ophiopsila annulosa.
XL.—Ophiopsila annulosa (M. Sars), a British Ophiurid.
By Canon A. M. Norman.
Ophiopsila annulosa (M. Sars).
1857. Ophianoplus annulosus, M. Sars, Middelhavets Littoral-Fauna,
p. 79, pl. i. figs. 2-7.
1859. Ophiopsila annulosa, Liitken, Addit. ad hist. Ophiuridarum, pt. ii.
. 133.
1869. Ophionoplus annulosus, Brady and Robertson, “ Week’s Dredging
in the West of Ireland,” Ann. & Mag. Nat. Hist. ser. 4, vol. iii.
p- 356, pl. xxii. figs. 1-6.
1904. Ophiopsila aranea, Allen and Pace (viv Forbes), Journ, Marine
Biol. Assoc. vol. vii. pp. 169, 171, and 209,
Although it is thirty-six years ago since Ophiopsila
annulosa was added to our fauna, it remains the most recent
addition to the Ophiuroidea to be met with in our inshore
waters. A few words on its distribution as now known may
not be out of place, as I am able to give two additional
habitats.
1. In 1868, Professor G. 8. Brady and the late Dr. D.
Robertson, when dredging in Birterbuy Bay, Ireland, pro-
cured a single arm of an Ophiurid unknown to them, and
they sent it to mefor determination. This arm (/.¢. pl. xxii.
fig. 1) was figured by them, together with reproductions of
five figures from the plate of M. Sars, and thus the- species
was fully illustrated, and a translation of Sars’s description
given in full.
In 1874, accompanied by my friend Dr. D. Robertson, I
went on a dredging-excursion to the west of Ireland. Round-
stone was one of our quarters; my chief object in going
there was to rediscover the gigantic form of Acera (Acera
bullata, var. Farrani, Norman, Ann. & Mag. Nat. Hist.
ser. 6, vol. vi. p. 68) which Dr. Farran, many years before,
had taken there, and, secondly, to procure Ophiopsila annulosa.
We were not rewarded with the Acera, but procured the
Ophiopsila. Dr. Robertson remembered the place where the
arm had been dredged. It was just at the entrance of
Birterbuy Bay on the west side, and very near the rocks.
A considerable time on two occasions was spent on this spot :
again and again the dredge would catch in the rocks; then,
when we imagined we had passed over a small scrap of sand,
arms and parts of arms would now and then reward us, and
at last a single disk with broken arms came up.
2. Mr. A. G. More was also at Roundstone at the time,
and on seeing the Ophiopsila he said that he had dredged a
On Fishes from Southern Mexico. 361
similar arm in a bay much nearer to Galway. That arm he
subsequently kindly sent me.
3. When at the Biological Station at Plymouth in the
autumn of 1903 I saw a number of fine and perfect specimens
which had been procured and some of these I purchased. The
species had been found to be not uncommon in crevices of the
red sandstone, “ especially in old Pholadidia crypts”’ in 15-
25 fathoms on Mewstone Ledge and Stoke Point Grounds.
It would thus appear to be a lover of hard ground anda
clinger to rocks, which is what we might have been led to
expect from its sturdy build and short and strong arm-spines—
reminding us of species of similar habit, such as the genera
Ophiopholis and Ophiactis. If, therefore, the specimens
dredged by us at Birterbuy were, as we imagined, on sand,
they had probably crept out temporarily from the rocks
close by.
4. I found the Ophiopsila in a fourth locality in the spring
of last year, having dredged a portion of an arm outside
Dartmouth Harbour,
I regard Ophiopsila aranea, Forbes, as a distinct species.
It is of smaller size, and, besides other differences, the arm-
spines are both fewer in number and more slender. The
species inhabits the Adzean and Mediterranean Seas.
XLI.—A Collection of Fishes made by Dr. H. Gadow in
Southern Mexico. By C. Tare Reagan, B.A.
THE collection of fishes made by Dr. H. Gadow, F.R.S., in
Southern Mexico in the summer of 1903 was brought to the
British Museum by him and worked out by me in the autumn
of that year. Publication of the results obtained was held
back in the expectation that another collection would be
received from Dr. Gadow as the result of his visit to Mexico
in 1904. Unfortunately, however, the season proved to be
unfavourable for fish collecting during this second visit.
The Cichlids collected by Dr. Gadow have already been
dealt with in my revision of the genus Cichlosoma. They
include examples of a new species, C. Gadovit, and of two
others, C. Higenmanni, Meek, and C. heterodontus, Pellegr.,
new to the British Museum. Also new to the British
Museum are specimens of Platypecilus Nelsont, Gambusia
fasciata, Heterandria Lutzi, and Pomadasys Temple, species
described by Dr. Meek in his recent work on the fishes of
Mexico.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 24
362 On Fishes from Southern Mexico.
Below I describe a new Cotylopus and make some remarks
about Pseudoxiphophorus bimaculatus and P. pauctradiatus.
Cotylopus punctatus, sp. n.
Depth of body 62 in the length, length of head 53. Snout
14 as long as eye, the diameter of which is 54 in the length
ot head and 11 in the width of the osseous interorbital space.
Lower jaw shorter than the upper; maxillary extending to
below posterior border of eye. Dorsal VI, 1 10; all the rays
of the spinous dorsal, except the first, produced as filaments,
the fourth the longest, when laid back extending beyond the
middle of the second dorsal; rays of the second dorsal in-
creasing in length posteriorly, the last, when laid back, reaching
the caudal. Anal I 11, opposite to the second dorsal; last
anal ray, when laid back, extending 3 of the distance from
its base to the caudal. Pectoral a little longer than the head ;
ventral 2 the length of head. Caudal rounded. 95 scales in
a longitudinal series, 25 between second dorsal and anal fins;
abdomen with a median naked area extending forward from
the vent. Caudal peduncle 12 as long as deep. Head
covered with small dark spots; each scale on the body
bearing a similar spot; very small dark spots on the dorsal
fins and on the base of the pectoral.
A single specimen, 132 mm. in total length, from
‘Tequesixtlan.
‘he outer series of teeth of the lower jaw are completely
concealed in a fleshy pad; but examination of other species
of Cotylopus and Sicydium shows that the extent to which
these teeth are exposed varies considerably and that this
feature must not be regarded as characteristic of the species.
Pseudoxiphophorus pauciradiatus, Regan.
Dr. Meek regards this species as a synonym of Pseudo-
wiphophorus bimaculatus, Heck., but it seems probable that
he has seen specimens of the latter species only.
I have examined a large number of specimens of Pseudo-
atphophorus bimaculatus from Southern Mexico, Guatemala,
and British Honduras. ‘The number of rays in the dorsal
fin is usually 15, but varies from 14 to 17, whilst the head,
snout, and caudal fin appear to be slightly, but constantly,
longer and the anal fin considerably deeper than in P, pauci-
radiatus. Of this latter species I have seen the eight typical
examples from Orizabaand six others collected by Dr. Gadow *,
* Of six specimens of P. pauciradiatus from Orizaba, collected by
Dr. Gadow, one has 13, three have 12, and two have 11 dorsal rays. Of
On new Fishes from Japan. 363
also from Orizaba, which are exactly similar to the types.
The number of rays in the dorsal fin is usually 12, but varie:
from 11 to 13. Each scale on the body has a rather broad
blackish marginal or intramarginal crescent.
In the following table, based only on female specimens,
are given in the first column (A) the number of dorsal rays
and in the others the following measurements in millimetres :—
(B) total length, (C) length to base of caudal, (D) length of
head, (E) distance from tip of snout to origin of dorsal,
(2) longest anal ray.
Ae Bod, C.D, Net.
P. pauciradiatus, two of the types, 13 75 63 15 365 9
from Orizaba. 12 42 36 95 20 55
P. pauciradiatus, two of the speci- (47 72 69 14 365 9
mens from Orizaba, collected E 67 57 135 33. «9
by Dr. Gadow. 7 ny Maley ete
P. bimaculatus, four of the speci- - ie o ibe a e
mens from the Rio Tonto,,;5 79 58 15 30 12
collected by Dr. Gadow. Tas) GOB 13 287° A
P. bimaculatus, two specimens from lazt 2 74 Gl. vi6.. Sint) Ae
San Domingo de Guzman ere ier
Boece i Ao Caller, |? 59 16 30 1
In the typical form of P. bimaculutus the scales of the body
have narrow dark edges, forming a network which usually
fades out below. In five specimens from San Domingo de
Guzman, collected by Dr. A. C. Buller, the scales of the
middle series have very broad dark edges, giving the appear-
ance of a blackish longitudinal band from operculum to base
of caudal. This well-marked variety may be called Pseudo-
viphophorus bimaculatus, var. teniaius.
XLU.—Deseriptions of Three new Fishes from Japan, collected
by Mr. R. Gordon Smith. By C. Tate Recan, B.A.
Since the publication of my report on the fishes collected by
Mr. R. Gordon Smith in the Inland Sea of Japan* the
British Museum has received a further series from him, which
contains examples of three species which appear to be new to
science.
fifteen specimens of P. bimaculatus from the Rio Tonto, also collected by
Dr. Gadow, three have 16, seven have 15, and five have 14 dorsal rays.
* Ann. & Mag. Nat. Hist. (7) xv. 1905, p. 17.
364 On new Fishes from Japan.
Diagramma aporognathus.
Depth of body 31-34 in the length, length of head 33-34.
Snout as long as or ’ slightly longer than eye, the liowaee of
which is 3? in the length of head, interorbital width 34-32.
Jaws equal, or the lower very slightly projecting ; lower jaw
without pores; maxillary extending to below the anterior
margin of eye. 24 gill-rakers on the lower part of the ante-
rior arch. Scales 110-116 25%. Dorsal XIV 17, the fourth
to the seventh spines the longest, } 3-3 the oneal of head ;
rays of the soft fin subequal in length, $ the length of head.
Anal III 8. Pectoral nearly as long as the head. Caudal
apparently emarginate (broken in each specimen). Caudal
peduncle 2} as “long as deep. Greyish, with dark oblique
undulating stripes along the series of scales.
‘T'wo specimens, 260 ‘and 270 mm, in total length, from the
Inland Sea.
Gobius (Ctenogobius) atriceps.
Depth of body 43 in the length, length of head 34. Snout
longer than eye, the diameter of which is 54 in the length of
head and a little less than the interorbital width. Jaws
equal anteriorly ; maxillary not extending to below the
middle of eye. Head naked; nape covered with small
scales. Scales in a longitudinal series 38, in a transverse
series between second dorsal and anal 13. Dorsal VI 12;
rays of the spinous dorsal produced into filaments; rays of
the second dorsal slightly imcieasing in length to the last,
which is 3 the length of head and, when laid back, reaches
the caudal. Analll. Pectoral 4 the length of head; ventral
8 the length of head, extending a little more than 4 of the
distance from its base to the origin of anal. Candal rounded.
Caudal peduncle longer than deep. Olivaceous; head
blackish ; fins dusky, the dorsal and anal light at the base,
the pectoral with a pale transverse bar on its basal part.
A single specimen, 80 mm. in total length, from the
Inland Sea.
Achilognathus longipinnis.
Pharyngeal teeth 5—8S, sickle-shaped, the sides with trans-
verse striz, the edges entire. Depth of body 2-24 in the
length, length of head 4-44. Snout $ the diameter of eye,
which is 24-22 in the length of head and equal to the inter-
orbital width. Lower jaw a little shorter than the upper ;
maxillary hardly reaching the vertical from the anterior
Bibliographical Notice. 365
margin of eye; no barbels. Scales 85-38 = 5L or 6
between lateral line and root of ventral. Dorsal II 14-15,
originating at a point equidistant from tip of snout and base
of caudal; anterior branched rays as long as the head.
Anal II 14-15. Pectoral nearly as long as the head, ex-
tending to the root of the ventral. Ventral 8-rayed, a little
shorter than the pectoral, extending a little beyond the origin
of anal. Caudal deeply forked. Caudal peduncle longer
than deep. Olivaceous above, silvery below; lower parts of
abdomen blackish ; vertical fins dusky, the dorsal and caudal
with some dark spots on the rays, the anal with a blackish
edge; ventral blackish, with the outermost ray white.
Four specimens, 62 to 78 mm. in total length, from the
Yamasabu River, Lake Biwa.
BIBLIOGRAPHICAL NOTICE.
Memoirs of the Geological Survey of India.—Paleontologia Indica.
Series XV. Himalayan Fossils. Vol. lV. The Fauna of the Spite
Shales. By Dr. Vicror Unuie, Professor of Geology in the
University of Vienna. 132 pages, 18 plates, and 10 text-cuts.
Folio. 1904. Geol. Surv. Office, Calcutta; Kegan Paul & Co.,
London ; and Friedliuder, Berlin.
Tue important fossiliferous strata which constitute the basis of this
elaborate and well-illustrated Monograph by Dr. Victor Uhlig, of
Vienna, are met with in limited areas among the much denuded
rocks of the Central Himalaya, especially in the Spiti Valley,
latitude N. 32° 5’, longitude E. 78° 15’, and lat. N. 28° 51’, long.
E. 77° 36’. The beds consist of dark grey and black shales, 300 feet
thick, lying over a limestone and under a sandstone. The former
(part of the Lower Gondwana system) is referred to the Jurassic
epoch, and the latter or Giusmal Sandstone (6U0 feet thick) belongs
to the Upper Gondwana and is referred to the Neocomian. ‘The
relative age of the “ Spiti Fossils” has been the subject of much
controversy, and several eminent palwontologists have assisted the
Geological Survey of India in this investigation with both head and
hand.
In the Introduction Dr. Uhlig makes careful mention of the many
geologists who have advanced our knowledge, special and general,
of the Spiti Fauna through the clouds of doubt and difficulties
encountered in former days; and he gratefully thanks his fellow-
workers and friends in India, Europe, and Britain, by whose help
this comprehensive and really valuable memoir has been perfected
and published. The talented artists and the friendly translator of
his MS. are especially thanked. Indeed everybody who has been
engaged in this good work has to be congratulated on the complete
suecess of their labours,
366 Bibliographical Notice.
The description of the species occupies pages 4 to 132, and in the
Introduction (p. 3) this is apparently referred to as Part I. of the
work, the notes on the natural history of the fossils, their rela-
tionships, stratigraphical position, and discovery being intended for
Part II., but these particulars appear to have been incorporated
with the description of each species.
In his systematic treatment of the species of Ammonites from the
Spiti strata, Dr. Uhlig introduces the Ammonidea (p. 4) by ecare-
fully explaining that, not binding himself to the modern definition
of the genera, he draws up the description of each of the forms with
extreme minuteness, and indicates its ‘‘ apparent position in the
most generally accepted system.” Thus he prefixes his view of the
generic characters at the commencement of the account given of
each group of species,
The classification of the species can be casually seen in the
several tabular lists at pages 77, 82, 83, 89, 93, 95, 96, 98, 99,
101,102, 106, 107, 110, 122,114, 115, 117, U9) 21, tae
126, 127, 129, 1380. The reference-numbers here annexed to the
species, which are arranged according to their natural affinities, will
be of some use to student and reader, especially as there is no
Index issued with this edition of the Memoir.
The following and some other Ammonites (see pages 107-111,
112-118, 119-125, 126-130, 130-132; and the table, infrd, page 368)
from the Spiti beds are described, besides those mentioned at pages
4 to 74 :—
Hoptoceras, Zitie/, p. 18.
AMMONIDEA. : :
: Dieneri, sp. nov., p. 19.
Puytioceras, Suess, p. 4. indicum, sp. noy., p. 21.
plicatum, sp. nov., p. 4. Hecricocrras, Bonarelli, p. 23.
strigile, Blanford, sp., p. 6. Kobelli, Oppel, sp., p. 25.
Lyroceras, Suess, p. 8. latistrigatum, sp. nov., p. 27.
exoticum, Oppel, sp., p. 14. sp. nov. indet., p. 29.
OrrrLia, Waagner, p. 30.
The Oppelie of the Spiti fauna are classified in four groups
according to their lobal structure, as below. It is noted by the
author that ‘‘ the name Oppelia (sensu stricto) should be reserved
for the group of subradiata, as Waagner, in establishing the genus,
had that group of forms above all in his mind.” The generic aspect
is that of a smooth or feebly costate discoidal Ammonite.
I. Group of Oppelia Adolphi, Oppel, p. 31 (Ammonites pictus, Quenstedt ;
Group of the Tenwilobata, Oppel; Streblites, Hyatt), represented by the
under-mentioned forms :—
Page
Oppelia (Streblites) Adolphi, Opp. ...............ssc0esess 42
= 3 Kerafitisas pa OV. mie ceeceeesoreeneeee 44
+ - Griesbachis sp 0 OVela.--c--eaeee ee 47
i * planopictasispanOva .eccsecaseesese: 47
- 5 BPsMOV ING Ob. paso. <.--eewacoet once sete 50
. is MMcOpicta,(spPmnOVawess-heereaasaeees 52
s. Ae inimallayanasisp: MOVse- nsec sesnceeees 51
- Pe substriata, ‘Oppel, sp. .....0.cs0s:0. 54
4 7 Iymamni, Oppel, Spx tenses sens. arene d6
Bibliographical Notice. 367
Page
Oppelia (Streblites) n. sp. indet. aff. Lymani, Oppel ... 56
= f punctatopicta, sp. nOv................ 59
A Bs sphenodoma, sp. nov. ............-.- 58
. Kj platydoma, sp. nov. ............... pO
Ee arr e471 LepLON OMA, Sp) HOVE 8120.9- Ars eo 58
4 i domocrenata, sp. NOV. .............+ 64
4 % PY EULA SPAN OVd ec ececsann<eecsson'- 65
oe a AGUNA ta, SP: OVA ctecvesesccsossyseces 63
. = SP yNOye Wi Oba d- magsied 9-2 haere soe 67
The Adolphi group are closely connected with the European
Tenuilobata by their concordance of the lobal structure, ornamenta-
tion, external shape of the test, development of the carina, and the
form of the body-chamber.
II. Group of Oppelia acucincta (Oppelia, sensu stricto), represented by only
one species: Oppelia acucincta, Strachey, sp. (p. 40).
III. Group of Oppelia nivalis (Neumayria, Bayle), represented by only one
species: Oppelia (Newmayria) nivalis, Stoliczka, sp. (p. 41).
IV. Group of Oppelia (cotraustes) adela, represented by only one species:
Oppelia (Ecotraustes) adela, sp. nov. (p. 41).
In comparing the Spiti Tenwilobata with the European, Dr. Uhlig
finds (pp. 37-38) that the former group had multifarious ramifica-
tions very partially known to us, while in Europe the group is also
manifold, but more closely knit, and there are some analogies and
some divergent characters ; these two geographically distinct series
do not appear to have followed a similar and quite parallel course
of evolution, but were affected “ by partial and provincial tendencies
to mutation, and admitted, even along the same lines of mutation,
of the development of vicarial forms (as, for example, Opp. indupicta
and Frotho).” It is not necessary to ask for a land-barrier to
account for this isolation, for a wide range of ocean may well be
occupied by various and different creatures. It is thought possible,
however, that one at least of the Cutch (Katrol) Ammonites (Opp.
plicodiscus, Waagner) might serve as an intermediate.
Following the genus Oppelia comes the genus <Aspidoceras,
Zittel (pp. 74, 75), represented by Aspidoceras avellanoides, sp. nov.,
founded on a specimen regarded by Stoliczka, in 1865, as Ammonites
liparus, Oppel, and proved on re-examination to be Aspidoceras of
the /nflati group Physoderoceras, Hyatt).
The genus Holostephanus, Neumayr, is recorded next to Aspido-
ceras in order (page 77). In external form it has a thick and costate
shell, tuberculate along the umbilical border. It is divisible into
several groups, one of which has a single representative in the
Spiti Shales (//. Schenkz), while the other groups ‘exhibit an un-
exampled variety of forms.” The alliances of some of these have
been very differently regarded by different writers. Taking Holo-
stzphanus spitiensis (Blanford) as the type of a new subgenus
(Spiticeras, pp. 77 & 82), Dr. Uhlig describes it carefully, and
defines its species in the following order :—
368 Bibliographical Notice.
I. Forms with a broad saddle and slightly pendent, feebly developed,
auxiliary lobes.
(a) Forms of larger growth.
No. Page Page Page
12. Spiticeras Stanleyi (Oppel)......cecsceesseseeee 77 82 107
Lo. PP IMGjSVATI, p> NOV: lercss-cscecncseece 77 82 110
(6) Forms of medium and smaller growth.
1. Spiticeras spitiense (Blanf.) ......ssc..00. Boas STUAd 82 89
3. + subspitiense, Sp. NOV...ececeesseees 77 i) 8!
2, 55 Grote (Oppel) parecescesonesceteee 77 82 92
4, 3 bilobatum, sp. NOV. .........e0.0. 77 82 96
5. i; subbilobatum, sp. nov. ............ 77 2 98
8. 3 binodigrum, sp. NOY. ..+....0+.seeee 77 82 101
ee 5 planamysp, DOWau cesses e-ne-ee ace 77 82 99
9. - CONSELVADS, SP. NOV. ces.sesesee0s-. 11 82 102
10. 3 @autleys(Oppel)ee cer eeeeee hal 82 104
dali: a subcautleyi, sp. NOV. ........-...+« 77 82 106
16. “s Griesbachinisp. WOVe cree cree ee 77 82 114
14. - SELMP LUI) (SU-ACMs)lnansesteceecacsees Maat 82 t12
15. F bulliformie; sp 3moye cs. s-ss-2e- a 2 114
II. Forms with narrow slashed saddles and deeply pendent,
strongly developed, auxiliary lobes.
20. Spiticeras obliquelobatum, sp. nov. ......... Ch 83 122
19. es OppelizapamOvae-scccceshensesersetes 77 83 121
Pele a MG HOTEMAT | NON Boechasconsedasboc 77 83 «124
18. es guttabum (Strach.)........-.--..-+.- (tf 83) eg
17. 3 Bp; HOV. tNdeL, “canteen scersvaseosee Ut” So" AE
III. Isolated and extreme forms.
22. Spiticeras eximium, sp. NOV. ...coe..-ssoeesees 77 83 126
23. x: SpeclOSuM, SP. NOV. «.....2.eseeee- 77 83 127
24, 9 MOVRINGESLs Geos Resse ceasnasesacee sees 7A 83 129
The “important and oft-quoted” Holostephanus (Astieria) Schenki,
Jppel, sp., is refigured and described anew from Oppel’s restoration
of the 'Tobeta specimen, and another fragment is mentioned (pages
130-132). Some observations are made on the standing of the
species among its allies, also on the relationship of some South-
African Ammonites to one of the groups of Holostephanus.
Besides the shape and ornamentation of the shell and the
peculiarities of the septa, the naturalist has to note their successional
features (due to phases of growth), in establishing specific and other
divisions in Ammonites. In estimating the relative value of their
many features and characters, Dr. Uhlig finds breaks and puzzling
contrasts, and it will be quite possible that other Cephalopodists,
following their own views of the relative value of the evidence, may
be unable to accept the classification proposed in this Monograph,
but at all events they will highly appreciate the value of its great
store of illustrated information, so carefully collected and systema-
tically arranged.
Ann. & Mag. Nat. Hist..S.'% Vol_ XVI FLX.
G Lewis.
West,Newman imp.
H. Knight ad nat lith
Histerida.
1, PUXI.
LX\
Ann. & Mag. Nat Hhist..S.7.Vo
Tig aan
een hth.
oa
Tr
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THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. 94. OCTOBER 1905.
X LITT.—Descriptions of new Species of Noctuidiein the British
Museum. By Sir Georce F. Hampson, Bart., F.Z.S., &e.
EurELianZ.
Ingura glauca, sp. nu.
Head, thorax, and abdomen grey, tinged with pale olive-
yellow and irrorated with black, the last with subdorsal and
subventral series of small black spots. Fore wing grey,
largely suffused with pale olive-yellow and slightly irrerated
with black, more thickly on medial area; a black spot at base
of cell with striga from costa above it ; an antemedial black
line angled outwards below costa, then slightly waved and
with traces of an olive line before it; orbicular and reniform
small with olive centres and grey annuli, the former round
or elongate elliptical, sometimes confluent with the latter,
which has its lower extremity produced to a point ; a diffused
waved black medial line; postmedial line double, bent out-
wards below costa, then the inner line excurved in interspaces
and angled inwards on the veins, very oblique below vein 4,
the outer line reduced to black points and a prominent
lunule above inner margin, followed by an oblique olive band
from costa to vein 4, then incurved and reduced to spots ;
some black points on costa towards apex ; an oblique olive
band across apical area from costa to vein 6; a grey and
black apical patch with a pale curved line on its inner edge
and a waved black line on termen; a series of black points
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 25
370 Sir. G. F. Hampson on
just before termen and more prominent spot at submedian
fold; cilia intersected with black near apex. Hind wing pale
suffused with fuscous on the veins and terminal area and
with olive on inner area, more uniformly fuscous in female.
Underside of fore wing with the costal and terminal areas
whitish; hind wing whitish, with traces of medial line, post-
medial series of black points, diffused waved subterminal line,
and waved terminal line.
Hab. Venuzveta, Aroa,1 g type; Br. Guiana, Potaro R.
(Kaye),1 9. Kzxp. 28 mm.
Ingura griseifusa, sp. 0.
Antenne of male bipectinate, the apical part simple.
&. Head and thorax grey, tinged with rufous and irrorated
with a few black scales; abdomen grey irrorated with black.
Fore wing grey-white irrorated with fuscous and slightly
tinged with brown ; a waved black antemedial line ; orbicular
and reniform moderate, rounded, defined by black ; an mdis-
tinct waved medial line; postmedial line double, bent
outwards below costa and incurved below vein 4, with another
indistinct line beyond it; short black streaks above middle
of veins 5, 6; a dark patch from costa before apex; a
crenulate line just before termen. Hind wing fuscous brown,
the interspaces of basal and inner areas paler ; a slight pale
mark above tornus and some black strize on inner margin.
near tornus; the underside paler except terminal area, traces
of waved medial, postmedial, and subterminal lines.
?. Fore wing with the outer edge of reniform indented ;
the line before termen reduced to a series of points.
Hab. Brazit, Rio Janeiro, 1 g type, Organ Mts., Tijuca
(Wagner),1 S$, 1 9; Paracuay, Sapucay (Foster), 1 ¢.
Kap. 32 mm.
Ingura nyctichroma, sp. n.
Antenne of male bipectinate, the apical part simple.
6. Head and thorax dark red-brown irrorated with black ;
tegulee with diffused black medial line; patagia with short
black streaks ; prothorax with pair of white points; pectus
with some white hair; tarsi ringed with white; abdomen
red-brown with dorsal and subdorsal series of black points,
the second and third segments with lateral white lines. Fore
wing red-brown suffused with fuscous; a double minutely
waved antemedial black line ; orbicular absent; reniform a
greyish bar with red-brown line in centre; a waved line from
lower angle of cell to inner margin; postmedial line black,
new Species of Noctuide. 371
slightly bent outwards below costa, angled outwards above
vein 6 and incurved below vein 4, with traces of a waved line
before it; a slight black streak above middle of vein 5; traces
of a pale subterminal line defined on each side by brown
slightly angled outwards at vein 7 and below vein 3 forming
two whitish dentitions ; a whitish patch at apex with short
black streak below it; a series of oblique black strize just
before termen; a fine dark terminal line. Hind wing dark
reddish brown, the interspaces of inner area pale towards
base ; some black striz on inner margin towards tornus; a
fine pale line at base of cilia; the underside grey irrorated
and suffused with brown, a medial line strongly excurved at
middle, traces of a double waved postmedial line with series
of points on the veins and a diffused subterminal line.
Hab. Panama, La Chorrera (Dolby-Tyler), 1 ¢ type.
Exp. 30 mm.
Ingura leucotrigona, sp. n.
9. Head and tegule ochreous tinged with rufous ; thorax
rufous mixed with grey; pectus and legs ochreous tinged
with rufous ; abdomen brown suffused with grey, the anal
tuft and ventral surface whitish tinged with rufous. Fore
wing ochreous tinged with rufous ; the base rufous suffused
with grey ; a triangular white subbasal patch between cell
and vein | before a diffused rufous band from cell to inner
margin with incurved red-brown line on its inner edge; a
red-brown line from below costa beyond middle oblique to
near termen below vein 7, below vein 5 strongly incurved, then
obliquely sinuous to inner margin, the area beyond it dark
brown suffused with grey; a series of black strie before
termen. Hind wing whitish tinged with brown, the terminal
half suffused with dark brown; a diffused grey subterminal
line excurved below costa; a series of dark points just before
termen and striz on termen; the underside yellowish,
irrorated with red-brown ; a brown discoidal lunule, an ill-
defined medial line, an indistinct waved postmedial line with
brown suffusion between it and the diffused grey subterminal
line.
Hab. Borneo, Kuching (Shelford), 1 2? type. Hap.
38 mm.
Ingura poliotis, sp. n.
Antenne of male bipectinate, the apical part simple.
6. Head and thorax rufous mixed with grey and fuscous ;
abdomen rufous tinged with grey and irrorated with black,
slight dorsal black marks on the two medial segments. Fore
De
Me
372 Sir G. F. Hampson on
wing with the basal and costal areas ochreous suffused with
rufous and irrorated with fuscous, the rest of wing grey,
suffused in parts with rufous; a fine strongly incurved ante-
medial line from submedian fold to inner margin, with some
diffused brown beyond it; postmedial line indistinct and very
oblique from costa to below vein 7, then prominent, blackish,
strongly incurved below vein 5 and oblique to inner margin,
with traces of a dentate line beyond it; a grey patch on
apical part of terminal area ; a series of black strie before
termen ; some white points on costa towards apex. Hind
wing fuscous brown, the basal and inner areas rather paler;
a diffused pale subterminal mark towards tornus ; the under-
side grey irrorated with black and suffused with red towards
apex, a discoidal bar, slight sinuous postmedial line, and
diffused subterminal band.
?. Fore wing with the incurved postmedial line less
distinct, the dentate line more prominent; an indistinct
waved subterminal line angled inwards in submedian fold.
flab. Stncarore (Ridley), 1 3,3 9 type. Exp. 28 mm.
Eutelia chrysoterimina, sp. un.
3. Head and thorax deep red-brown mixed with a little
grey, the head and a patch on metathorax redder ; palpi pale
in front ; hind tarsi with black spot at base; abdomen pale
red, the basal half darker with a red patch at base, the crests
and subdorsal series of small spots blackish. Fore wing
deep red-brown suffused with grey on basal half of costal
area, the terminal area below vein 6 broadly golden yellow
suffused with some red, on inner margin extending to before
middle; a subbasal grey striga from costa; an indistinct
highly curved dark antemedial line, below the cell bounding
the dark area; reniform defined by white at sides, very
narrow and constricted at middle; postmedial line almost
obsolete towards costa, below the cell fine double defined
by red-brown on inner side, very oblique and minutely waved,
another line beyond it very oblique from costa to vein 6 near
termen, then inwardly oblique and rather diffused; a
prominent white subterminal bar from costa to vein 6, the
area beyond it deep chocolate with two black lunules defined
by white on termen, the subterminal line then indistinct,
yellow and slightly sinuous; a fine slightly sinuous brown
subterminal line. Hind wing pure white; the costal area
dark brown ; the terminal area below vein 5 fiery red with
sinuous yellow subterminal line; a black spot on inner
margin beyond middle; the underside with the costal and
new Species of Noctuide. 373
terminal areas suffused with fiery red, some black suffusion
in base of cell, an oblique elliptical black discoidal spot
defined by white and with black bar from costa just beyond
it; an indistinct dark postmedial line with blackish spot
below vein 6, an indistinct sinuous yellow subterminal line,
some black points on termen towards apex.
Hab. Fr. Guiana, St. Jean Maroni (Schaus), 1 3g type,
St. Laurent Maroni. Lap. 32-34 mm.
Eutelia caustiplaga, sp. n.
6. Head and thorax bright chocolate-red mixed with a
little grey; palpi with the second joint in front and tuft on
basal joint of antennz white ; pectus, base of legs, and tarsi
white, the tibize with white bases, the tarsi with black spot on
first joint ; abdomen red with deep chocolate-red subdorsal
patches at base, with a little grey before each, the ventral
surface pale grey-brown. Fore wing red with some grey
suffusion in places; the median nervure and veins rising
from it white; subbasal line represented by a grey striga
from costa; antemedial line fine, grey, very oblique from
costa to median nervure, then inwardly oblique, a diffused
oblique grey-white fascia beyond it from costa to lower angle
of cell and a chocolate-red patch on inner area followed by a
triangular white patch with some yellow and a dark striga on
it; reniform narrow, constricted at middle, chocolate-red
with white annulus, a white striga above it from costa, and
a large chocolate-red patch beyond it with a little yellowish
in centre; postmedial line unusually near termen, white,
obliquely curved to vein 6, then inwardly oblique, below vein
3 indistinct, dark, slightly waved, with a white point on its
outer edge at vein 5 and some whitish suffusion below vein 3;
some white points on costa towards apex, which is grey with
a white line across apical area to termen at vein 5; a grey
subterminal line from the white point on postmedial line
angled inwards below vein 4 and below vein 3 incurved,
prominent and white, the area beyond it deep chocolate-red ;
some triangular black points on apical half of termen ; a fine
white line at base of cilia. Hind wing pure white, the
terminal area brown suffused with fiery red; a white
subterminal line from vein 3, angled outwards to near termen
at submedian fold; a fine white line at base of cilia; the
underside with the costal and terminal areas grey; a black
discoidal spot, a small spot at middle of submedian fold ;
traces of two waved postmedial lines with some red on them
at discal and submedian folds, some black points ov termen.
374 Su G. F. Hampson on
Hab. Cussa, Santiago (Schaus), 1 g type, Matanzas.
Exp. 36 mm.
Eutelia pyrastis, sp. n.
Antenne of male bipectinate, the apical part simple.
¢. Head rufous; palpi and basal joint of antennz in front
white ; tegule and prothorax grey, the rest of* thorax
suffused with rufous; abdomen grey slightly tinged with
rufous, rufous dorsal patches on second and third segments and
a point on fifth segment. Fore wing grey suffused in part
with rufous ; the basal area rufous except costal area ; subbasal
line represented by a whitish striga from costa; antemedial
line oblique from costa to median nervure, then obliquely
curved, a yellow patch beyond it on inner area; the veins of
terminal half shghtly streaked with white ; an oblique dark
striga from middle of costa; reniform a narrow lunule
constricted at middle, with dark centre and whitish annulus ;
postmedial line with small white and brown mark on costa,
then indistinct, strongly bent outwards below costa, excurved
and slightly waved to vein 4, then strongiy ineurved and
almost obsolete ; a rufous triangle defined by white from costa
before apex to vein 6, with three small white-defined blackish
marks below it followed by a white line to tornus ; the apical
half of termen grey, the inner half rufous; some black points
on termen. Hind wing with the basal half whitish, the veins
and terminal half brown; some black striz on inner margin
towards tornus; a white bar above tornus; the termen
tinged with red, with some white and three black points at
middle; the underside white slightly irrorated with fuscous,
a small discoidal lunule, indistinct waved medial and post-
medial lines and terminal series of points.
? rather more rufous.
Hab. U.S.A., Florida, Miami (Schaus),1 ¢; Banamas,
Nassau (Sir G. Carter), 1 8,21 2 type. Hapaigiae.
9 386 mm.
Eutelia leucographa, sp. n.
Antenne of male bipectinate, the apical part simple.
Palpi white, the first two jomts and the third joint near
tip brown at sides ; frons white with the frontal tuft tipped
with brown; vertex of head and antenne black; thorax
black with a few white scales and white spot on metathorax ;
pectus white behind ; legs black banded with white ; abdomen
black-brown slighily variegated with white, the ventral
new Species of Noctuidee. 375
surface mostly white. Fore wing red-brown variegated
with white especially on antemedial area and on postmedial
area from vein 3 to inner margin, the white areas with dark
strie; subbasal line represented by a white striga from
costa ; antemedial line unusually near base, strong, black,
slightly angled in cell, then oblique and not quite reaching
inner margin; orbicular an ill-defined white spot with dark
points on it; reniform an elliptical white patch with black
spots in its upper and lower parts; medial line indistinct
except at costa and inner margin, oblique from costa to cell,
then inwardly oblique and obscurely double ; postmedial
line strong, black except between veins 3 and 1], oblique
from costa to vein 6, incurved between veins 3 and 1, defined
on outer side by a fine white line from costa to vein 3;
subterminal line white from costa to vein 3, with a black
band on its inner side expanding towards costa and vein 3,
oblique to vein 6, then outwardly oblique and irregular,
below vein 3 represented by a black spot above tornus ; cilia
chequered brown and whitish. Hind wing white, the
terminal area fuscous, extending to middle at costa and
narrowing to tornus, a slightly waved white subterminal
band from vein 6 to tornus, the area beyond it browner,
two black striz above tornus; a fine crenulate black line;
cilia white at base, blackish at tips; the underside white
irrorated with black, the apical area suffused with brown,
two black discoidal bars, a waved medial line excurved at
middle, double minutely waved postmedial line and traces of
a subterminal line.
?. Fore wing less variegated with white.
Hab. Br. EF. Arrica, El Quaso, Masai (Betton), 3 g,1 2
type; Care Cotony, Steynsberg,2¢. Hap. 36-40 mm.
Eutelia diplographa, sp. n.
Antenne of male bipectinate, the apical part simple.
¢g. Head and thorax grey mixed with brown and fuscous;
palpi with the second joint blackish at sides ; tegule black
at base, with brown medial line and dark tips ; abdomen
grey suffused with fuscous, the ventral surface paler. Fore
wing grey-white suffused with brown especially on basal and
costal areas; subbasal line represented by two striz from
costa; antemedial line double, oblique, the ner line brown,
diffused, the outer strong, black ; orbicular a slight brown
annulus ; reniform defined by fuscous and with some brown
in centre ; medial line strongly excurved round the reniform ;
postmedial line double, the inner line black, the outer brown,
376 Sir G. F. Hampson on
excurved from costa to vein 4, then incurved ; subterminal
lme represented by blackish patches, forming bars from
costa to vein 7 and vein 3 to inner margin, excurved between
those points; a fine waved terminal line; cilia chequered
white and brown. Hind wing white, the terminal area
suffused with brown; the underside white, the costal and
terminal areas irrorated with brown, a discoidal annulus,
crenulate postmedial line with slight streaks on the veins,
traces of subterminal line, and terminal series of points.
2 whiter.
Hab. Stxuim (Pilcher), 13; Perak, Larut Hills (S. S.
Flower), 19 type. Ezp. 40 mm.
Eutelia mesogona, sp. n.
9. Head and thorax ochreous mixed with reddish brown ;
palpi with the second joint brown at sides, the third ochreous,
long and dilated at extremity ; tegule with dark upper edge ;
patagia black at extremity ; pectus whitish; legs whitish,
brown and black ; abdomen whitish tinged with brown and
with small brown subdorsal spots. Fore wing reddish brown
tinged with greenish fuscous ; subbasal line represented by
a slight striga from costa; antemedial line indistinctly
double, slightly waved, oblique from costa to median nervure,
then incurved and with the inner line strong, black; a
medial ochreous band, its outer edge defined by the double
postmedial line, oblique from costa to vein 6, where it is
avutely angled, inwardly oblique below the cell ; orbicular
represented by a small ochreous spot on the dark area, the
reniform slightly defined by brown and rather triangular,
on the ochreous area; traces of a waved medial line; a
somewhat trilobate whitish mark between veins 4 and 2,
with some ochreous below it, before the subterminal line,
which is whitish and very oblique from costa to vein 6, then
ochreous, waved, excurved to vein 3; some white points on
costa towards apex and a whitish apical patch; a terminal
series of small black lunules with ochreous striz beyond
them. Hind wing brown, the basal and inner areas paler ;
some black and whitish striz in submedian fold and on inner
margin towards tornus ; a whitish waved line from vein 3 to
termen near tornus ; ; small black lunules defined by whitish
on termen from apex to submedian fold ; the underside with
the costal half reddish brown, the inner half whitish, a small
black discoidal spot, curved medial line, and three slight
minutely waved lines on terminal area.
Hab. Mavacascar, Betsileo (Cowan), 1 3 type. Eup.
36 mm.
~
new Species of Noctuide. oT
Eutela gerea, sp. n.
Antenne of male strongly serrate and fasciculate, the
apical part simple.
¢. Head, thorax, and abdomen reddish brown suffused
with grey, the last with the dorsal crests small, dark. Fore
wing brownish grey with some dark irroration ; a dark point
at base of cell; antemedial line oblique to subcostal nervure,
hardly traceable except at costa; orbicular represented by a
small spot ; reniform well developed, rounded, rufous defined
by blackish ; medial line oblique from costa to lower angle
of cell, then incurved ; postmedial line indistinctly double,
minutely waved, excurved from below costa to vein 4, then
incurved; a triangular chocolate patch defined by grey on
costa before apex, the subterminal line arising from it and
represented by small rufous dentate marks, excurved at
middle ; a terminal series of small chocolate lunules. Hind
wing rufous, the basal and costal areas paler; the underside
rufous, the inner area whitish, a black discoidal lunule,
curved postmedial and subterminal lines showing through to
upperside.
?. Head, thorax, and fore wing grey-white, the last with
the markings much more distinct, some rufous suffusion
between medial and postmedial lines.
Hab. Mauritius, Curepipe (Tullock), 1 3, 1 2 type.
Exp. 26 mm.
Eutelia flaviluna, sp. n.
Antenne of male serrate and fasciculate.
3. Head, thorax, and abdomen grey tinged with brown
and irrorated with black ; pectus and legs blackish, the
tarsi ringed with white. Fore wing reddish brown suffused
with grey and irrorated with black ; antemedial line double,
the outer line strong, black, oblique from costa to subcostal
nervure where it is angled, then slightly incurved ; orbicular
represented by a slight black streak, the reniform a pro-
minent ochreous lunule defined by black ; medial line rather
indistinct, oblique from costa to median nervure, then
incurved ; postmedial line double, sinuous, oblique from
costa to vein 6, then inwardly oblique; subterminal line
indistinct, ochreous defined by somewhat dentate blackish
marks on inner side, waved, angled inwards to discal fold
and outwards on vein 3; a fine waved black terminal line.
Hind wing fuscous, the inner area slightly paler ; an ill-
defined ochreous line from vein 5 to tornus ; the underside
whitish suffused with brown and irrorated with fuscous, a
378 Sir G. F. Hampson on
small black discoidal spot defined by whitish, a curved medial
line and diffused subterminal band outwardly defined by
whitish on inner half.
Hab. Sincavore (Ridley), 1 g type. Exp. 30 mm.
Eutelia furcicauda, sp. n.
Antenne of male serrate and fasciculate.
3. Head, thorax, and abdomen brick-red tinged with
grey ; palpi in front to near end of second joint, pectus, the
greater part of legs, and ventral surface of abdomen white ;
the lateral anal tufts long. Fore wing brick-red suffused
with grey ; conjoined yellowish-white spots defined by red
and with red centres above and below vein 1 at middle of
inner margin ; a fuscous medial line angled outwards below
costa and ending at the spots on inner margin ; minute
points at angles of cell; a faint postmedial line, excurved
from costa to vein 4, then incurved to near the medial spots ;
costa red towards apex with some yellow points on it; a
slight red streak above vein 7 with two outwardly oblique
yellow strize defined by red from its imner side to vein 5;
subterminal line indistinct red, sinuous, with two obliquely
placed yellow points on it below costa and ending above
tornus; termen strongly angled at vein 3, then oblique,
waved. Hind wing yellowish white, the terminal area brick-
red, brownish at tornus; a yellow point at extremity of
vein 2; termen slightly crenulate, angled at vein 2, then
excised; cilia yellowish and red. Underside of fore wing
with the basal areas of costa and inner margin yellowish, the
disk fuscous irrorated with grey, the terminal area blue-grey,
costa red from middle to near apex, some reddish beyond
cell and a deeper red spot beyond postmedial line; hind
wing white with some black irroration at base of costa and
lower angle of cell, a small red discoidal spot, postmedial
line defined by small red spots before and beyond it, terminal
area suffused with red, some black points on termen.
Hab. Sincavore (Ridley), 3 g type. Lap. 30 mm.
Eutelia cautabasis, sp. n.
Antenne of male minutely serrate and fasciculate.
3. Head and thorax ferruginous red ; patagia with deeper
red patches at shoulders; palpi blackish at base; tarsi
fuscous ringed with white; abdomen pale rufous with some
deeper red on dorsum and some greyish hair at base, the
ventral surface deep rufous. Fore wing with the basal area
«
new Spccies of Noctuide. 379
yellowish with small deep rufous patch below base of costa
and large patch from subcostal nervure to inner margin,
bounded by a red-brown band expanding into a triangular
patch on costa; the rest of wing greenish grey with as slight
rufous tinge ; an indistinct waved rufous medial line excurved
from costa to vein 2 ; postmedial line faint with dark points
on the veins, bent outwards below costa, then slightly curved ;
a small triangular red-brown patch on costa before apex ;
some brown suffusion before termen between veins 4,3; a
terminal series of points; termen angled at vein 3, then
oblique, waved. Hind wing reddish brown, the basal and
inner areas yellowish white; an obliquely curved whitish
medial line ; some slight dark marks on extremity of vein 2
and at tornus ; a fine waved terminal line; cilia rufous; the
underside rufous slightly irrorated with brown, the inner
area whitish, a small discoidal lunule and faint minutely waved
medial and postmedial lines.
Hab. Oup Catasar (Crompton), 1 3 type. Lup.38 mm.
Eutelia holocausta, sp. n.
?. Head, thorax, and base of abdomen fiery rufous ;
tegule with blackish patches at base ; palpi long; abdomen
rufous tinged with grey, with small lateral phocolatemred
spots on third and fourth segments. Tore wing fiery rufous
slightly tinged with grey and irrorated with fuscous; traces
of an antemedial red line, oblique below the cell ; a fiery-red
spot in middle of cell; a faint pale discoidal lunule; traces
of a double postmedial line oblique from costa to vein 6 and
below vein 4; a chocolate-red patch on costa before apex
outwardly defined by an oblique white line with some fuscous
beyond it ; traces of a subterminal line excurved at middle,
then strongly incurved, with minute dentate chocolate marks
before it above and below vein 5, and shght rufous marks
above and below vein 2; termen angled at vein 4, then
oblique. Hind wing fiery rufous, with small chocolate-red
spots on inner margin towards tornus; the underside with
chocolate discoidal spot and minutely waved medial, post-
medial, and subterminal lines, obsolescent on costal half.
Hab, Niceria, Sapele (Sampson), 19 type. Hap. 32 mm.
Eutelia metasarca, sp. 0.
Antenne of male serrate and fasciculate, the apical part
ciliated.
Head, thorax, and abdomen pale red-brown tinged with
380 Sir G. F. Hampson on
grey; palpi black at base ; abdomen with sublateral and two
ventral series of black points on medial segments. Fore
wing grey tinged with pale rufous; a black point in base of
cell; antemedial line indistinct, double, slightly angled out-
wards below costa, then oblique ; a minute annulus in middle
of ecll; reniform D-shaped with brownish centre and white
annulus defined by black ; traces of a medial line; post-
medial line double, oblique from costa to vein 6, then
inwardly oblique, waved ; traces of a pale waved subterminal
line, whiter and with brown shade before it at costa; a ter-
minal series of small black lunules; the termen slightly
angled at vein 4. Hind wing flesh-red, the basal and inner
areas slightly paler ; a terminal series of dark lunules; the
underside whitish irrorated with rufous, a black discoidal
spot on a slightly waved medial line and maculate post-
medial line.
Hab. Nigeria, Sapele (Sampson),1 3,2 2 type. Ezp.
20 mm.
Eutelia melanopis, sp. 1.
Antenne of male serrate and fasciculate, the apical part
simple.
¢. Head, thorax, and abdomen brown suffused with grey;
palpi black at base ; tarsi black with whitish rings; abdomen
with the small dorsal crests black, and paired ventral black
points on basal and three medial segments. Fore wing grey
tinged and irrorated with brown ; antemedial line indistinct,
oblique from costa to submedian fold and with small black
spot on costa beyond it ; a minute annulus in middle of cell;
reniform narrow, black with white annulus and with oblique
black striga above it from costa ; a medial line from cell to
inner margin; postmedial line indistinct, double filled in
with whitish, with black mark on its outer side at costa,
oblique to vein 6 where it is angled, angled at vein 4, then
incurved to below end of cell ; a conical deep chocolate patch
with white points on costa before apex, the subterminal line
arising from its outer side, very indistinct, pale slightly
defined on each side by brown, white at costa, irregularly
waved ; a terminal series of black points ; the termen slightly
angled at vein 4. Hind wing grey-brown; a fine waved
terminal line from apex to submedian fold; some dark
points on inner margin near tornus and on extremity of
vein 2; the underside grey tinged with rufous and irrorated
with fuscous, a black discoidal spot, minutely waved double
postmedial line, and traces of subterminal line with small
black spot at tornus.
Hab. Sierra Leone (Cator),1 3 type. Exp. 24 mm.
new Species of Noctuidee. 3381
Eutelia chlorobasis, sp. n.
@. Head and thorax whitish, mixed with brown; palpi
black at base; abdomen grey, with slight subdorsal series of
fuscous points, small lateral black spots on fourth segment,
and some sublateral spots, the ventral surface suffused with
red and with series of white points. Fore wing pale reddish
brown, slightly irrorated with fuscous; the basal area
suffused with greenish grey ; an oblique blackish antemedial
line, defined by white on inner side except towards costa; a
slight medial line from cell to inner margin ; orbicular and
reniform greenish white defined by fuscous, the former a
minute spot, the latter elliptical, with grey centre; post-
medial line double filled in with grey, oblique from costa to
vein 6, where it is angled, then minutely waved, incurved
below vein 4, some grecnish-grey suffusion beyond it on
costa ; a black patch defined on outer side by a white striga
on costa before apex, the slight grey subterminal line arising
from its outer edge, excurved at middle and incurved below
vein 3, traversing a patch of fuscous suffusion on terminal
area from vein 5 to inner margin; a fine waved black
terminal line. Hind wing brown, suffused with fuscous, with
fine waved dark terminal line. Underside of fore wing with
the terminal area brick-red, with oblique white subapical
striga; hind wing suffused with rufous, a black discoidal
point, fine waved medial and postmedial lines, and terminal
series of lunules.
Hab. Nicerta, Sapele (Sampson), 1 2 type. Ezp.
22 mm.
Eutelia nigridentula, sp. n.
?. Head, thorax, and abdomen red-brown ; palpi whitish,
black at base and on outer side of third joint ; tarsi fuscous,
ringed with white; abdomen with dorsal and subdorsal
series of slight points, the ventral surface whitish, with paired
series of black poimts. Fore wing pale or dark reddish
brown, slightly irrorated with fuscous; a whitish striga in
cell before the double antemedial line, filled in with rufous
and defined by whitish on inner side, angled outwards below
costa, then oblique; orbicular a small black spot; a slight
medial line, excurved below costa and oblique below the cell;
reniform elliptical, chocolate with ochreous annulus defined
by black; postmedial line double filled in with ochreous,
oblique from costa to vein 6, where it is angled, angled
inwards in discal fold and outwards at vein 4, then oblique
waved ; a chocolate patch on costa before apex, defined on
outer side by a whitish lunule, below which is a black dentate
382 Sir G. F. Hampson on
mark, followed by traces of a series of points ; the terminal
area rather darker, with a series of small black lunules on
termen. Hind wing fuscous brown, with fine waved terminal
dark line ; the underside greyish, suffused with rufous and
irrorated with fuscous; a small black discoidal lunule, in-
distinct minutely waved medial and double postmedial lines.
Hab. Sierra Leone (Clements), 1 9 type; Nierrta, Old
Calabar (Crompton), 1 ¢, Sapele (Sampson), 2 9. Exp.
22 mm.
Eutelia albiluna, sp. n.
9. Head and thorax white, slightly mixed with brown ;
abdomen white, tinged with fuscous except at base, where
there are some black scales, the ventral surface white, with
sublateral and two ventral series of black points. Fore wing
whitish, irrorated with fuscous and suffused with fuscous
brown on medial area and beyond the cell; some brownish
suffusion at base of costa; antemedial line indistinctly
double, angled outwards below costa, then oblique ; orbicular
a minute white spot defined by black; an oblique medial
line from cell to inner margin; reniform a prominent white
lunule; postmedial line indistinct, double, oblique from costa
to vein 6, then inwardly oblique, dentate below vein 4; a
black-brown patch below costa towards apex, its outer edge
rather dentate and defined by the whitish subterminal line,
which is definéd by fuscous on inner side, angled inwards in
discal fold, below vein 4 obliquely incurved, strongly defined
by black on inner side and with the area beyond it darker ;
a terminal series of small black lunules. Hind wing fuscous
brown, with some black and white striz on inner margin
near tornus, a black and white subterminal mark on vein 2,
and a fine waved terminal line; the underside whitish,
tinged with brown and irrorated with fuscous, a discoidal
point, slight waved medial and postmedial lines, and traces
of subterminal line.
Hab. Niceria, Sapele (Sampson), 2 9 type. Hap.
22 mm.
Eutelia leucodelta, sp. n.
?. Head, thorax, and abdomen grey-white ; tegule with
slight brown medial line and brownish tips; tibiz with
slight black marks; abdomen with the basal crest tipped
with rufous, the slight dorsal crests rufous, the ventral
surface white, with sublateral and two ventral series of black
points. Fore wing grey-white, the basal and costal areas
suffused with pale olive-brown, slightly irrorated with
new Species of Noctuidae. 383
fuscous ; antemedial line prominently defined by white on
inner side except towards costa, incurved to costa, then
oblique, with a slightly rufous-tinged band beyond it and
black point at imner margin; reniform a C-shaped white
mark incompletely detined by black and with olive centre ;
some rufous suffusion beyond the cell ; postmedial line
double, indistinct, oblique from costa to vein 6, then in-
wardly oblique, dentate below vein 4; an olive patch on
costa before the indistinct, waved, white subterminal line,
which has some slight black marks on it between veins 4
and 2, the area beyond it greyer ; a terminal series of small
black lunules. Hind wing brick-red, the base and termen
whitish; some slight black marks on inner margin towards
tornus; a waved black terminal line. Underside whitish,
irrorated with rufous; fore wing with the disk suffused with
rufous, the white discoidal lunule with black centre, a waved
black postmedial line excurved from costa to vein 4, then
oblique ; hind wing with black discoidal spot, slight waved
medial and double postmedial lines.
Hab. Niceria, Sapele (Sampson), 1 ¢ type. Ezp.
30 mm.
Kutelia ochricostata, sp. un.
?. Head and fore half of thorax rufous, tips of patagia
and hinder part of thorax ochreous white, metathoracic crest
rufous ; palpi blackish at base ; abdomen brown, tinged
with grey and rufous, the dorsal reat dark, eahidoreal series
of slight dark segmental striz. Fore wing red-brown, the
basal and costal areas to near apex ochreous tinged with
rufous ; two oblique black antemedial lines from middle of
cell to inner margin, towards which they separate more
widely ; a slight oblique medial line from cell to inner margin ;
reniform a V-shaped ochreous-white mark filled in with
rufous ; postmedial line indistinct, double, minutely waved,
almost obsolete on costal area, bent outwards below costa,
excurved to vein 4, then incurved ; subterminal line ochreous,
distinct from costa to vein 6, where it is joined by an oblique
ochreous streak from beyond postmedial line, then represented
by slight ochreous and rufous strize incurved below vein 3;
apical | area fuscous grey, with black streaks above veins 6, 7 ;
a terminal series of small dark lunules; cilia with some
ochreous above and below vein 5. Hind wing brown, with
some pale and dark strize on vein 2 and inner margin towards
tornus ; a pale subterminal line towards tornus ; a terminal
series of dark lunules ; the underside suffused with rufous
384 Sir G. F. Hampson on
and irrorated with brown, a dark discoidal lunule, minutely
waved medial and double postmedial line.
Hab. Nigeria, Old Calabar (Crompton), 1 2 type. Ezp.
24 mm.
Eutelia ferridorsata, sp. un.
9. Head blackish, slightly mixed with grey; thorax
ferruginous, the tegule tinged with fuscous; pectus and
legs fuscous and greyish; abdomen pale ferruginous, the
dorsal crests and slight subdorsal segmental striz dark, the
ventral surface suffused with black. Fore wing greenish
grey, suffused with black and rufous and with numerous
dark striz ; a double waved antemedial line; reniform grey,
defined by black, and with some fuscous in centre, narrow,
irregularly elliptical ; postmedial line indistinct, defined by
rufous on outer side, minutely waved, excurved from costa
to vein 4, then incurved; a slight, minutely waved, black
subterminal line, incurved below vein 4, and with the area
beyond it greyish towards tornus ; a terminal series of small
black lunules; cilia rufous and black. Hind wing rufous,
suffused with fuscous, the inner area rather paler; some
dark striz on vein 2 and on inner margin towards tornus ;
a pale mark on termen near tornus ; cilia rufous ; the under-
side rufous irrorated with brown, the inner area whitish, a
discoidal point, medial line excurved below costa, double
minutely waved postmedial line, the inner line indistinct,
traces of a subterminal line, and a terminal series of black
lunules on whitish marks.
Hab. Nigeria, Old Calabar (Sampson), 1 9 type. Exp.
26 mm.
Eutelia cyanolopha, sp. n.
Antenne of male ciliated.
d. Head and thorax rufous, mixed with some whitish ;
tegule with blackish medial line and dark tips ; metathorax
with a pair of blue-black spots; abdomen whitish, dorsally
suffused with rufous, the first segment with a pair of blue-
black spots, the fourth with dorsal white spot. Fore wing
rufous, suffused with greyish ; the cell from near base, the
area beyond it, and the submedian interspace with rubbed
appearance ; subbasal line represented by shght double striz
from costa filled in with grey; antemedial line with two
oblique striz from costa and oblique dark bar to the lobe on
inner margin, with white bar beyond it; reniform an ill-
defined lunule with rufous tuft of scales at upper angle of
new Species of Noctuide. 385
cell; postmedial line faint, oblique, waved, from costa to
vein 6, defined’ by grey on inner side; the terminal half of
inner area with some grey and blackish striz ; some obscure
dark points just before termen and a fine terminal line.
Hind wing semihyaline whitish, the veins and terminal area
suffused with brown; some dark striz on inner margin
towards tornus, and a rufous mark at extremity of vein 2;
cilia whitish; the underside with two black discoidal points,
traces of a curved medial line, and a diffused subterminal
band.
?. Fore wing with the rubbed appearance confined to the
extremity of cell.
Hab. Gouv Coast, Aburi (Johnson), 1 g, 1 2; Br. E.
Arrica, Athi-ya- Mawe (Betton), 1 g type; MasHonaanp,
Umtali (Marshall), 2 8,4 2. Exp. 30-34 mm.
Eutelia albidisca, sp. u.
Antenne of male ciliated.
¢. Head, thorax, and abdomen fuscous, irrorated with
white and a few black scales; metathoracic crest white and
rufous; tarsi ringed with white; abdomen with subdorsal
tufts of white hair at base and white patch on terminal
segment, the crests dark, the ventral surface whitish. Fore
wing greyish fuscous, irrorated with white, the cell and discal
area pure white; a curved black antemedial line bent out-
wards to inner margin; a curved black medial line inter-
rupted in cell; a discoidal lunule and striga beyond upper
angle of cell; postmedial line represented by a black point
on costa, a black striga from below costa to vein 5, and an
indistinctly double sinuous line from cell to inner margin ;
subterminal line shght, white, irregularly dentate, excurved
from below costa to vein 6; a terminal series of small black
lunules. Hind wing white, the terminal area fuscous, broadly
so at costa, narrowing to tornus; the underside with the
costal and terminal areas suffused with grey and irrorated
with brown, a black discoidal lunule, and ill-defined slightly
waved medial and postmedial lines, interrupted at middle
and on inner area.
Hab. Care Cotony, Brak Kloof (Mrs. White), 1 3 type.
Exp. 26 mm.
Eutelia rufula, sp. n.
Antennee of male ciliated.
Head and thorax deep rufous; terminal half of patagia
and mesothorax greyish ochreous; pectus ochreous; legs
Ann. & Mag. N. Hist, Ser. 7, Vol. xvi. 26
386 On new Species of Noctuide.
brown and ochreous, the tarsi blackish with pale rings;
abdomen ochreous, dors ally suffused with rufous and grey
except at base, the slight dorsal crests dark. Fore wing with
the basal half Aaa suffused with violaceous whee ; an
obliquely curved antemedial rufous band; a dark spot in
middle of cell ; an interrupted curved medial line forming
the inner edge of the rufous terminal half; an obscure
discoidal spot defined by blackish points and some rufous ;
some violaceous suffusion before and beyond the postmedial
line, which is double, oblique from costa to vein 6, then
inwardly oblique and sinuous; subterminal line violaceous
erey, defined by obscure dark marks before and beyond it,
slightly angled outwards at vein 7 and excurved at middle ;
a fine black terminal line and whitish line at base of cilia,
which are fuscous. Hind wing fuscous brown, the interspaces
of basal half somewhat paler ; a fine dark terminal line and
pale line at base of cilia; the underside pale, a discoidal
point, curved medial and double postmedial lines.
Hab. Prax (Kiinsiler), 1 9 ; Stincarore (Ridley), 4 3,
1 2 type. Hzp. 22 mm.
Eutelia siccata, sp. n.
?. Head and thorax brown mixed with fuscous ; tegule
with black spots; abdomen grey-brown, irrorated with
fuscous, the shght dorsal crests black, small black subdorsal
spots on first three segments, and some lateral spots. Fore
wing reddish brown, tinged with fuscous, the interspaces
rather paler, giving a ‘streaky appearance ; a diffused blackish »
subbasal mark on inner area; the antemedial area rather
pale; an obscure dark discoidal mark ; the lines obsolete,
except double oblique striz at middle of inner margin, repre-
senting the postmedial line; a small white spot on imner
margin towards tornus ; traces of a subterminal series of
small dark spots; a terminal series of small black lunules.
Hind wing fuscous brown, the basal area paler ; a slight
pale mark on termen near tornus ; the underside with traces
of curved medial and double postmedial lines, the latter with
slight black streaks on median nervures; a terminal series of
small black lunules, the one near tornus with white mark
before it.
Hab. Sterra Leone (Carter), 1 2 type. Hap. 28 mm.
[To be continued. }
.
On the Development of an Asterid. 387
XLIV.— Some Observations on the Development of an
Asterid with Large Yolky Eggs from the Franklin Islands.
By E. H. Henpverson, M.Se., M.D., McGill University,
Montreal *.
[Plates XII. & XIII]
Tue material consisted of a brood of young starfish collected
off the Franklin Islands and belonging to the British
Museum, and the work was carried on in the Zoological
Laboratories of McGill University during the summer
vacations of the years 1902-03-04.
External Appearance.—On examination the specimens
showed evidence of being in the fixing stage, being aggluti-
nated together into masses resembling miniature bunches
of grapes. The five-rayed shape was in all cases evident
and at first sight seemed to be due in part to the pressure of
neighbouring embryos on each other, but on closer exami-
nation was seen to be really due to the assumption of that
typical form of the adult starfish.
When one of these masses was separated with a needle
into its constituent members and examined under a hand-
lens, it became evident that the individual members were
connected together in the region of the preoral lobe (this
arrangement giving them that peculiar bunched appearance),
the preoral lobes being central, while the dorsal portions of the
embryos were outermost.
The individuals themselves were of large size, measuring
almost 2 mm. in diameter ; they were hemispherical in shape,
the ventral surface being flat and showing the radial sym-
metry, the dorsal surface dome-shaped. There were only
two or three members of the collection which did not show
radial symmetry, and these proved to be only masses of
yolk surrounded by a membrane, very likely unfertilized
eggs (Pl. XII. fig. B).
Pl. XII. fig. A shows a group of three, from which others
have been detached, so as to reveal the characteristic union
of the preoral lobes.
Pl. XII. figs. C and D are drawings to represent what were
the earliest stages in the collection. The preoral lobe is still
prominent, showing little or no constriction, while the
hydroceele is just appearing as two longitudinal ridges in each
radius, The five-rayed appearance is seen only from the
* Abstract of a thesis presented for the M.Sc. degree.
388 Dr. E. H. Henderson on the Development
ventral surface, as the arms have not yet grown out and
differentiated themselves from the body, so that the embryo
from above appears spherical. In some instances (fig. D)
there could be seen distinct bosses, or projections, in each
radius along the line of, and just dorsal to, the tube-feet.
These are the arms of the future starfish.
Pl. XII. fig. E (ventral surface) shows the latest stage that
was found, and it existed in the material in great numbers.
The pentamerous arrangement is distinctly seen both from
above and below. ‘The ridges shown in figs. C and D have
developed well-marked tube-feet in fig. E. These can be
counted by a hand-lens and appear as small round white
papille, ten or twelvein each arm. At this stage the preoral
lobe is to a considerable extent atrophied, and is about to
separate off from the young starfish.
I might here state that the adherence of the preoral lobes
to each other is due rather to some agglutinative material,
which forms a coagulum between them, than to any cupping
or suction action, and that they cling to one another thus
till the stalk is excessively atrophied and ready to separate
from the body.
Having described the external appearance, I may now
briefly indicate the results of the study of sections.
Methods adopted—The great difficulty was to get these
large yolky larvee penetrated to the centre with celloidin or
paraffin, so that proper sections might be made. Both the
chloroform-paraffin and the benzol-paraffin methods were
tried, but in each case it was found to be impossible, even
with the greatest care, to get the paraffin to penetrate the
mass of yolk; when the larve were sectioned the centre
would drop out, leaving the outside tissue in the form of
a ring, and so the celloidin method had to be adopted.
Thin, medium, and thick solutions were used, since the more
gradual the change of density in successive solutions the
better is the degree of penetration. It is well to leave
them in each solution at least twenty-four hours, and a longer
time gives better results. Some of my best sections were
made last summer from material that had been lying in a
thin solution of celloidin for over a year.
The sections from a celloidin-paraftin block have a tendency
to curl, a trouble which is remedied in a large measure by
having an exceedingly sharp razor and a Bunsen flame near
by or else bright sunshine playing over the surface of the
microtome.
The sections were cut in varying thickness from 5 » to 10 pu.
of an Asterid with Large Yolky Eggs. 389
In thick sections the yolk is not nearly so apt to be broken
nor is there such a liability in folding, but there is a greater
liability to curl. The sections were then mounted on the slide
in hot water to flatten them. They were stained in hema-
toxylin, then washed in running tap-water till the excess of
stain was removed and until they took on a deep blue colour.
They were then doubly stained in eosin (watery solution).
By such procedure very successful results in staining
were attained and the differentiation between yolk and
tissue shown up clearly and distinctly.
Internal Anatomy.—The type of development conforms
exactly to that shown in Asterina gibbosa as described
by MacBride*. The chief difference is that the embryos
under discussion are much larger in size and have all the
interstices of the body gorged with yolk. The coelomic
cavities, hydrocceles (both right and left), periliemal spaces,
the stone-canal, and pore-canal all have developed in a
similar manner to that which has been described in the
case of Asterina. Pl. XII. fig. F shows the relations of the
celomic spaces. The right posterior ceelom (r.p.c.) is placed
dorsally; the left posterior ccelom (/.p.e.) with its right ventral
horn (.’p.’c.’) extends over to the right on the ventral surface.
Between these two portions of the left ceelom is seen the
remains of the old anterior cceelom, now the axial sinus (az.),
with the stone-canal (sf.c.) on its posterior wall. The develop-
ment of all these struciures is the same as in the case of
Asterina and by their further growth and changes !eads to
similar alterations in the structure of the organism.
The perihzemal spaces are deiived from evaginations of the
celom, four arising from the left posterior aud one from the
anterior celom. PI. XII. fig. F shows one (ph. 1-5) deve-
loped from the left coelom and also one ( ph. 1-2) developed
from the anterior ccelom (az.). These tongue-like projections
of cceelom insinuate themselves between the lobes of the
hydroceele and the ectoderm from either side and finally
meeting beneath the floor of the hydroccele form a septum.
Pl. XIII. fig. G serves to illustrate another point, viz. the
relation of the preoral cavity to the axial sinus. The con-
striction of the preoral lobe or stalk from the body is effected
by opposite sides of the stalk becoming approximated to each
other and folded, so that portions are invaginated into the
interior and destroyed by histolysis. This process causes
the anterior ceelom to become divided into two portions —
* E. W. MacBride, “The Development of Asterina gibbusa,” Q. J. M.S.
1806.
390 Dr. E. H. Henderson on the Development
a stalk-coclom, which is cast off, and a portion remaining
with the body called the axial sinus (a@z.).
The alimentary tract is hard to define in many sections.
No trace of either mouth or anus is found to exist. All that
can be seen of an alimentary system is an extensively folded
and eciled gut-wall lying in the interior yolk. In Pl. XIII.
fig. H I have endeavoured to show its appearance. One is
able to follow its general outline fairly well in some sections,
but the interior within the outer line of the gut-wall reveals
only a disturbed mass of broken-down yolk, portions of gut-
wall, and amcebocytes. The gut-cells are oblong or oval in
shape with irregular ends towards the lumen, and seem to be
engulfing yolk, as particles of yolk can be seen not only
among them but also in their substance. In places where
the lumen can be seen it often appears full of yolk which is
granular, showing that it is being broken up by a process,
perhaps, of digestion and absorption.
The yolk forms by far the largest part of the whole star-
fish, at least nine-tenths of the whole bulk is yolk. Stained
with eosin this becomes easily differentiated from the other
tissues. Itis composed chiefly of large globular or irregularly
shaped masses (PI. XIII. fig. J), which are closely apposed to
one another, leaving few chinks intervening. This yolk not
only fills up the centre but penetrates everywhere ; in fact,
the whole cf the tissues would seem to have been built up
around it, as, indeed, they really are. Itis found among the
ectoderm cells, between the ectodermic wall and the peri-
toneal wall of the coelomic spaces, and among the cells of the
gut-wall, while with the gut it composes the main mass of
the interior. Here it so presses on the coelomic sacs as to
almost obliterate them—the two peritoneal walls being
brought into close apposition, so that the ccelom is not seen
at all as an open space, but only as a layer of peritoneal
tissue in the midst of yolk.
Histology.—The structure of the body-wall differs on the
oral and aboral surfaces. On the dorsum the ectoderm
consists of a single layer of flattened cells, which, as the
ventral surface is approached, gradually merge into long and
narrow cells (PI. XIII. fig. H). On the dorsal surface the
ectoderm les in close relation to the peritoneum, while on
the ventral surface it is separated by a mass of yolk, among
which are found amcebocytes and tissues of the mesenchyme.
On the ventral surface beneath the hydroccele the ectoderm
cells increase in number, and become long and filamentous,
and closely compacted, so that if the section be thick there is
nothing to be seen but a mass of nuclei.
of an Asterid with Large Yolky Eggs. 391
Immediately beneath the bases of the ectoderm cells,
where these abut on the perihzemal spaces, it is possible to
trace evidence of what I conclude to be the nervous system.
This appears as a layer of exceedingly fine fibrils, which
could only be made out by a high power, and which
apparently replaces the mesenchyme intervening between the
ectoderm and the walls of the perihzemal spaces (Pl. XIII.
fig. I).
The cells of the peritoneum vary in shape from flattened
to cubical, and occasionally one sees evidence of fine myo-
epithelial cells, the beginning of muscle-formation. The
ectoderm, too, is often bent into folds, while the peritoneum
remains straight, a fact which may be regarded as evidence
of the contractile power of the latter. This is especially true
of the preoral lobe, the ectoderm of which becomes folded
into pockets, while the peritoneum of the ccelom assumes
a contracted appearauce. These folds are closed off from
the exterior and are then destroyed by amcebocytes.
The cells of the hydroceele vary in appearance from
columnar to flattened or cubical. In the radial water
vascular canal they are flat or cubical, while in the tube-feet
and terminal tentacles they are columnar and increased in
number, so that the nuclei appear several layers deep (Pl. XIII.
fig. 1). Fig. I also shows muscular elements developing
on the aboral side of the walls of the perihzemal spaces
(musc.). These muscles would serve, when in action, to
draw together the ambulacral ossicles of opposite sides and
so close the ambulacral groove in the adult.
Pl. XIII. fig. J shows a curious group of cells projecting
into the axial sinus from its posterior wall. ‘This is the ovoid
gland, and is composed of a mass of primitive germ-cells.
‘These are large and cylindrical in shape, held together by
fibres derived from the mesenchyme. In places the surface of
this group of cells is irregular and folded, and covered with
smaller cells, which are doubtless derived from the peri-
toneum of the axial sinus, which has been pushed in before
it in its growth forward. Further, it is developed from the
wall of the left posterior eccelom, which becomes thickened
and invaginated.
The foregoing are the chief points which could be made
out from the material. I have endeavoured to show, in my
thesis, of which this is an abstract, the close parallelism in
development between this organism and Asterina; while here
I have mentioned them but briefly, and rather dwelt on the
more important differences, and some points of technique,
which I have learned while doing the laboratory work.
392 Mr. T. D. A. Cockerell—Deseriptions and
EXPLANATION OF THE PLATES.
List of Abbreviations employed.
ant.ce., anterior celom; gz., axial sinus; ect., ectoderm; Aist., a
spot on the preoral lobe undergoing histolysis ; hyd., left hydroccele ;
—-V., its five primary lobes; Lp.c., left posterior coelom ; l.'p.'c.’, its right
ventral extension; mes., mesenchyme ; musc., muscular tissue developing
from the wall of the perihemal canal; nerv., nervous fibrils ; per., peri-
toneum ; ph., rudiments of the periheemal spaces; ph. 1-5, that between
lobes I. and V. of the hydroccele ; ph. 1-2, that between lobes I. and II.
of the hydroccele; pr.germ., primitive germ-cells; rp.c., right poste-
rior ccelom ; s¢.c., stone-canal.
Pirate XII,
Fig. A. Three embryos adhering by their preoral lobes.
Fig. B. An unfertilized ege.
Fig. C. The youngest embryo in the collection, showing primary lobes of
the hydroccele.
Fig. D. A slightly older embryo. The rudiments of the arms appear as
bosses above the lobes of the hydroccele.
fig. E. The oldest embryo in the collection. The pentagonal form is
clear, and the rudiments of the tube-feet have appeared.
Fig. F. Transverse section of a comparatively young embryo, showing
relations of ccelomic cavities, the stone-canal as open grooye,
and the perihzmal cavities arising as eyaginations of the
coelom.
Puate XIII.
Fig. G. Portion of another transverse section of an embryo about the
same age as that represented in PI. XII. fig. F, showing the
formation of the axial sinus by constriction from the anterior
ceelom.
Fig. H. Another transverse section, showing gut-wall and its relation to
the yolk.
Fig. 1. Section, more highly magnified, of a small portion of the ventral
body-wall, showing the relations of a lobe of the hydroccele to
the perihemal space and the ectoderm. The beginnings of
nervous and muscular tissue are seen.
Fig. J. Section, highly magnified, of a small portion of the dorsal body-
wall, showing yolk-globules in the dermis and the first rudi-
ment (pr.germ.) of the genital stolon or “ ovoid gland.”
Zoological Laboratory, McGill University.
June 19085.
XLV.—Descriptions and Records of Bees.—1V.
By T. D. A. Cockrre.y, University of Colorado.
Bombus sumatrensis, sp. n.
? Length about 18 mm.
Head elongate, narrow, the greater part of the clypeus,
which is smooth and shining, below the level of the eyes;
Records of Dees. 393
malar space long, smooth and shining; around red tubercle
at extreme base of mandibles; labrum bituberculate, with a
median depression, its lower part with reddish and black
hair ; hair of face and cheeks long and black, that of upper
part of head mixed black and white, the white hairs being
long and mainly on occiput; antennz entirely black, scape
long, strongly curved; first joint of flagellum as long as
third or slightly longer, the second much shorter than either ;
hair of thorax long, dense, yellowish white, with no sign of
any median band or patch. Legs very dark reddish, the hind
tibiz and tarsi quite bright ferruginous; hair of legs black
and orange, the latter mainly on the distal parts; hind
metatarsi very broad. Wings strongly suffused with orange-
brown, nervures ferruginous. Hair of abdomen long, black
on first two segments, with yellowish white toward the sides,
especially on the second ; black on third segment, yellowish
white on fourth and fifth, the colours largely mixed owing to
the overlapping of the long hairs and the presence of more
or jess pale hair even amongst the black; hair of venter
black, except at apex (principally fringe of penultimate
segment), where it is yellowish.
Hab. Sumatra; marked ‘at flower, 2381. X?, 2. 5. 81.”
In collection of British Museum, received in 1892.
This is the first Bombus known from Sumatra, the B. 6elli-
cosus, Sm., supposed to be from ‘‘ Sumatra or India,” bemg
really American according to Bingham. It is closely allied
to B. Mearnsi, Ashmead, from the Philippine Islands. Un-
fortunately B. Mearnsi is known only from a worker which
has been in alcohol; but it seems to be different from the
Sumatra species, and Dr. Ashmead, to whom I showed the
insect now described, expressed the opinion that it was not
the same as his. B. rufipes, Lep., from Java, is also allied,
but is readily distinguished by the black pubescence. Among
the Indian species B. sumatrensis appears to be nearest to
B. funerarius, Sim.
Cladocerapis bipectinatus (Smith).
Lamprocolletes bipectinatus, Smith, Trans. Ent. Soc. Lond. 1856, Proc.
p- dl.
Smith described this remarkable bee as above, and then in
1862, forgetting all about the first description, redescribed
it as L. cladocerus. The species, under the latter name,
became in 1904 the type of my genus Cladocerapis.
39-4 Mr. 1’. D. A. Cockerell—Deseriptions and
ANTHOPHORA, Latr.
The Australian species before me may be separated as
follows :—
Abdomen not, or not obviously, banded;
covered with appressed fulvous hair .... 1.
Abdomen conspicuously banded ........ A
1. Tegument of abdomen black, except broad
hind margins of segments; clypeus yellow,
with two large brown patches.......... scymna, Gribodo, 2.
Tegument of abdomen red ; clypeus with a
large reversed T in y allow rsd adel! rhodoscymna, Ckll., 3.
2. Clypeus light, with a black stripe on each
side of the upper part; males.......:.. 3.
Clypeus with at least large black marks .. 6,
3. Larger, about 13 mm. long ; hair of thorax
above bright orange- -fulvous, with black
intermixed ; abdominal bands bright blue. emendata Gilberti, Cll.
Smaller; abdominal bands greenish or
ochreous or EVM tee Hod GAB pals Roig oo'45 < 4,
4, Hair of thorax above bright orange-fulvous,
with black intermixed ; abdominal bands
emerald-creen shading to blue.......... emendata, Smith.
Hair of thorax above ochreous or light
fulvous, with black intermixed ........ 5.
5. Abdominal bands very light bluish ...... zonata subcerulea (Lep.).
Abdominal bands very light greenish .... zonata subcerulea (Lep ).
6. Sides of clypeus broadly black, leaving a
light pyramid, which is macened on each
side ; abdominal bands dull white ; size
rather small: qMale: cue. tee ae seer Salteri, Ckll,
Clypeus with a lar ge reversed T in yellow,
its stem broadly truncate on upper margin ;
size rather small; males ............-- te
Clypeus with a light reversed T, its stem
tapering above, pointed ; size larger ;
fertales® 9. ccs Seon ee ics cere ere 8.
7. Hair of mesothorax light blue mixed with
Iplack. "a phe eat ee hae etd hares Beery a eee Walkeri, Ckll.
Hair of mesothorax yellow mixed with
LEYS os Sena a DoL Sooo oomaoU De adelaide, Ckll.
8. Hair of mesothorax light ereenish mixed
with black; light hair on outer side of
middle legs strongly suffused with blue;
abdominal bands Loh Wie"... were mies > = ae cingulata (Fabr.).
Hair of mesothorax light blue mixed with
black ; abdominal “bands very bright
@reemish (HUG .uremet ae oo eas i a: Walkeri, Ck.
Hair of mesothorax very bright orange-
fulvous mixed with black; light hair on
outer side of hind legs ‘orange-fulvous ;
larger than the next species .......... emendata Gilberti, Ckll.
Hair of mesothorax paler fulvous, inclining
to greenish and mixed with black ; light
hair on outer side of hind legs white.... zonata (L.).
Records of Bees. 395
Anthophora scymna, Gribodo.
Described from a single ? from Queensland. A @ before
me is from West Australia, “ 68. 6.”
Anthophora rhodoscymna, sp. 0.
3 .—Length about 12}-13 mm.
Similar to A. scymna, but differing thus: tegument of
abdomen entirely bright red, with appressed orange hair,
that on first segment more erect and with some black inter-
mixed; that on hind margins of segments pale golden,
producing faint bands; apex without dark hair; clypeus
with a light orange reversed T, the upwardly-directed stem
tapering to a point; hair of cheeks abundant and snow-
white ; of vertex and occiput pale yellowish and mixed with
black ; of thorax above light yellowish, copiously mixed with
black ; of pleura pale yellowish; tegule apricot-colour ;
wings stained with brown along the veins ; the red legs have
orange pubescence, but the thick fringe on each side of hind
metatarsus and the long inner anterior fringe of hind tibia
are black ; the hind femora have short black hair beneath.
The fourth ventral segment of abdomen has a conspicuous
median tubercle, beset with short ferruginous hair ; the fifth
segment is deeply emarginate and the apical one ends in two
rather short widely separated spines, with a low elevation,
pyramidal in outline, between.
Hab. Queensland (Gilbert Turner); marked “300 Hy.”
Three males.
It is not impossible that this is the undescribed male of
A. scymna, but with the evidence at present available it must
be considered distinct.
The species of the group of A. zonata are very difficult to
classify. Dours, in 1869, recognized only 4. cinctofemorata,
Sichel, with a very strongly punctured abdomen, and
A. zonata (L.), from Australia, the latter being credited with
a number of named varieties. According to the characters
given in descriptions, the various named Australian forms
may be separated thus :—
Size small, usually less than 13 mm......... uP
eyde VEE oat Boo Ghiyr ROIIC OMT Eee Cb Sind) Grane 3.
1. Hair of thorax silvery ; abdominal bands blue,
ULE SOME, SROCMGE sere e Auleis te eye ciorte steerer het | cinctofemorata, Sichel.
Hair of thorax reddish or rufo-fulvous...... 2.
2. Bands of abdomen metallic blue ; light hair of
LEQSOWEHIUGE prow nave Mate € mike teelte ides Wire tae 6 sonata (L.).
396 Mr. T. D. A. Cockerell—Deseriptions and
Bands bluish white; hair of thorax rufo-
LLY OUS, 5, clare.» Meee eet ete RG eee pulchra, Sm.
3. Bands blue or silvery blue; hair of legs on
OWtaide LEPNUSIMONS se fate cls chasers en a vie emendata, Sm.
Bands bluish green or pale blue, not very
metallic ; hair of legs on outside bluish.... edngulata (Fabr.).
A. cinctofemorata is clearly distinct from anything I have
seen. The species before me I classify as follows, recognizing
that some revision will probably be necessary in the light of
fuller information :—
Anthophora zonata (l..).
Three females from Queensland (Gilbert Turner, 298) and
« Australia, 95.64,” agree excellently with zonata. Curiously,
however, the two males accompanying them, collected in
Queensland by Gilbert Turner (297 Hy. and 298), are both
of the variety swbcerulea (Lep.), not hitherto reported from
Australia. The one labelled 298 has pale blue bands, the
other a sort of greenish white.
Anthophora emendata, Smith.
Queensland (Gilbert Turner, 299); 1 2,2 6.
The two males differ greatly in size, but they appear to
belong to the same species, and the collector has given them
the same number. ‘The fifth ventral segment in the maie is
more strongly emarginate than in zonata, so I do not think
emendata can be a mere colour-variety. Smith describes
the female as 6 lines long, and gives good reasons for
supposing that Dours’s measurement of 18 mm. (under the
erroneous name “ cincta, Fabr.”) is a mistake. My female
and one of the males are larger than Smith indicates, being
about 13 mm. long, but the small male agrees with ‘Smith’s
measurement. Smith says the female has the clypeus
reddish yellow, while Dours says it is marked as in zonata;
my ¢ agrees with the latter statement. Smith says that
the ventral surface of the abdomen bas fulvous hair in the
middle, white at the sides; Dours says reddish black in the
middle, white at sides; my insect has it black in the middle,
white at the sides. In view of these discrepancies | will
designate my insect var. Gilberti, taking the female and large
male as typical of the variety.
Anthophora Walkeri, sp. u.
9 .— Length just over 12 mm.
Face-markings &c. agreeing with zonata; flagellum, except
Records of Bees. 397
at base, ferruginous beneath; hair of vertex and thorax
above bluish white very strongly mixed with black, of pleura
bluish white, of cheeks and sides of metathorax white, of
legs as in zonata; abdominal bands strongly metallic, the
first emerald-green, the other three paler, with mother-of-
pearl tints; tegule dark rufous.
3 .—Length about 10 mm.
Face-marks chrome-yellow; clypeus with a large black
mark on each side, shaped like a human foot; hair of hind
tarsi entirely black; abdominal bands emerald-green, the
last two and the third at the sides largely overlaid with
white.
Hab. Baudin Island, Long Reef, Western Australia, 2 ?,
1 ¢, collected by Commander J. J. Walker on the
‘ Penguin’ Expedition.
This exquisite species is appropriately named after its
discoverer, whose collections have made such important
additions to our knowledge of the Australian insect-fauna.
It is nearest to A. cingulata, but distinguished by its smaller
size and strongly metallic bands, and in the male by the
markings of the clypeus. The structure of the male abdomen
beneath is nearly as in zonata. It is to be remarked that
the Australian A. cingulata, as described by Dours, does not
agree with the Indian cingu/ata, described by Bingham. As
the species was originally described from Australia, I take it
that the Doursian description pertains to the true cingulata,
though the length he gives (18 mm.) must be a misprint for
13. Dours says the wings are 10 mm. long, and I find them
so in a specimen 13 mm. long.
Anthophora cingulata (Fabr.).
Clare, South Australia (W. Wesche); two females taken
Nov. 20, 1904.
The hair on the outside of the front and middle legs is
largely bluish green. Froggatt states that this bee is widely
distributed over the southern part of Australia (Tr. Roy.
Soc. Austr., June 1893).
3
Anthophora adelaide, sp. n.
6 .—Length about 11 mm.
Looks like a variety of A. zonata or pulchra, but is distin-
guished as follows: face-marks chrome-yellow; clypeus at
upper sides with large foot-like black patches, with the heel
very long and the toe very short ; hair of head and thorax
above as in zonata, light fulvous strongly mixed with black ;
398 Mr. T. D. A. Cockerell—Descriptions and
abdominal bands light yellowish with a sort of greenish tint ;
fifth ventral segment feebly emarginate, as in zonata, but
with a large, median, subapical, dull black patch where the
sculpture runs in semicircular lines; flagellum ferruginous
beneath ; legs as in zonata.
Hab. Adelaide River (J. J. Walker, 890).
Anthophora Salteri, sp. n.
¢ .—Length about 1] mm.
Looks just like A. adelaide, except that the abdominal
bands are dull white, the foot-like marks on the clypeus have
a short heel and a long thick toe, and the flagellum is only
dull dark brownish beneath. I should hardly venture to
separate it but for the entirely different structure of the fifth
ventral segment of the abdomen, which has a very broad,
deep, rounded emargination, its sides forming nearly a right
angle with the margin on each side, which is furnished with
two or three short but strong spines.
Hab. Paramatta, N.S.W. (W. R. Salter).
Prosopis, Fabr.
All the species in the following table have the abdomen
blue or green :—
First recurrent nervure meeting first trans-
verso-cubital or entering extreme apex of
first submarginal cell; abdomen dark blue.. 1.
Both recurrent nervures received by second
submareimalicells 7. foe cloaks ee onele bie eiee 3.
1. Large, about 11 mm. long; face with three
YELLOW: SEELPOS PSs oranrr om einion\ tee eer mice chiens alcyonea, Erichs., 2.
mall, Jessithan 7mm: longece sans wie ace 2
2. Face all creamy white below level of antennee. adbonitens, Ckll.
Lateral margins of clypeus dark; no light
supraclypeal marks | s82cickh soni ee eee disjuncta, Ckll.
3. Scutellum and postscutellum yellow ........ parallela, Cll.
Scutellum and postscutellum not yellow 4,
4. Tubercles conspicuously marked with yellow
OF Wille tac aatdc eee prea eeclee ee eee 5,
Tubercles not so marked, wholly metallic, or
rarely with a median non-metallic greyish
DANG. 21s. Scio ree Ocean t eee R ee 6.
5. Large; mesothorax black; face-marks and
tubercles very bright yellow.............. alcyonea, Erichs., 3.
Smaller; mesothorax green; face-marks and
mark on tubercles cream-colour .......... varicolor, Smith.
6. Head, thorax, and abdomen bright green .... perviridis, Ck11.
Mbdpmen aeepiplupesye: taka. teen eee if
7. Larger; face with three narrow cream-coloured
AtTiPCSmetee a. dey hese -s Jewkes ps eee reginarum, Ckll.
Smaller; face in male all white below an-
tenne, in female without light markings..,. Turneriana, Ckll.
Records of Bees. 399
Prosopis alcyonea, Erichs.
Albany, W. Australia (J. J. Walker, 4296),1 g ; Franklin,
Tasmania (J. J. Walker, 3469), 1 2.
The difference in venation indicated by the above table is
not specific, I believe, the insects being so similar in all
other respects. The male has the first r. n. nearer the base
of the second s.m. on one side than on the other. Erichson
described only the female; the male has most of the face
bright chrome-yellow, but there is a black band down each
side of the clypeus. The lateral face-marks narrow abruptly
just above the antenne and are then continued very narrowly
nearly to the top of the eyes. There is a yellow mark in
front of the anterior ocellus. The curious shining tubercle
at the base of the metathorax is as in the female. Evidently
P. vidua, Smith, is the male of alcyonea, as Smith himself
surmised; Smith’s type had the venation as in my female.
Prosopis varicolor, Smith.
Townsville, Queensland, 5. 3. 02 (fF. P. Dodd); also
13. 12. O1 (same collector). Both sexes.
This is evidently Smith’s species, but the face-markings
are cream-colour rather than white and the punctures of the
abdomen are quite strong. The male, not hitherto described,
is like the female, but the face-marks are ivory-white, the
lateral marks larger and longer, and the much narrower and
more elongate clypeus has a large white triangle, occupying
the greater part of its disk. The blue and green shades of
the abdomen are very brilliant.
Prosopis albonitens, sp. 0.
6 .—Length a little over 5 mm.
Very closely and minutely punctured; dark deep blue ;
light markings ivory-white, with a canary-yellow suffusion on
anterior and middle tibize. ’ace below antennze all shining
ivory-white, the white ending between the antennz in a
rounded prominence, but along the orbital margins extending
upwards, gradually narrowing to a point, the distance from
which to the top of the eye is about as great as the distance
of the latter from a lateral ocellus; mandibles and scape in
front ivory-white; flagellum brown, pale and yellowish
beneath ; mesothorax convex, very closely and minutely
punctured; hind border of prothorax continued to cover
tubercles, but interrupted in middle, white ; base of meta-
thorax with a very large area, semilunar in form, with the
400 Mr. T. D. A. Cockerell—Descriptions and
ends (sides) shortened, bounded by a sharp ridge, except in
the middle line posteriorly; this area is shining and is
divided into three subequal parts by two longitudinal carine,
the lateral spaces being rather obscurely obliquely ridged.
Legs piceous brown and yellowish white ; femora dark except
the anterior knees ; tibiz light, the anterior ones wholly so,
the middle ones with a dark spot behind, the hind ones with
a very broad, dark, subapical ring; tarsi yellowish white,
with the last two joints of the middle and the last three of
the hind tarsi pale brown; tegule with a white spot in
front; abdomen rather narrow, the hardly noticeable fine
pilosity white as far as the third segment, after that black.
Hab. Queensland (Gilbert Turner, 713, Ridg. 11.91).
Prosopis disjuncta, sp. n.
¢ .—Length about 65 mm.
Strongly punctured, but not so closely as in P. albonitens,
the punctures of the mesothorax in particular being much
larger, more irregular, and less dense; dark blue, the head
aud thorax inclining to steel-blue, the abdomen to deep rich
purple; light markings cream-colour or a very pale primrose ;
wings hyaline, nervures dark. Clypeus light, with a dark
margin, this margin varying in breadth, so that the light
triangle may be very large or rather small; lateral face-
marks broad, bow-shaped, notched above by the antennal
sockets; mandibles and scape wholly dark; flagellum
brown, yellowish ferruginous beneath; middle of front
elevated and rugose, the elevation bounded on each side by
a sharp ridge; at each side of this elevation is a smooth
shining area; hind border of prothorax and large spot on
tubercles light, but the two light areas are not nearly
continuous; basal area of metathorax somewhat dumbbell-
shaped, broader at the sides than in the middle ; tegulze with
asmall light spot. Legs dark, slightly metallic; the anterior
tibia and metatarsus in front, a small spot at base of middle
and hind tibiz, a very small spot at apex of middle tibia,
and short line on its tarsus, yellow; abdomen with the almost
imperceptible pilosity white as far as the fourth segment,
after that black.
Hab. Queensland (Gilbert Turner, 717, Ridg. 11.91),
Two.
Prosopis parallela, sp. n.
2 .—Length nearly 83 mm.
Strongly punctured, the punctures very large and dense
Records of Bees. 401
on mesothorax ; bright Prussian green, the abdomen a bluer
green except near the sutures, the first two segments strongly
suffused with purple above; light markings bright chrome-
yellow, except those of the head, which are pellucid grey,
but one has the upper end of the median facial stripe yellow,
and probably the markings should all be of that colour when
fully mature; wings hyaline, nervures dark, Head elon-
gated, the facial quadrangle nearly twice as long as broad ;
inner orbital margins above gently concave; mandibles
wholly dark, strongly ridged ; light markings of head con-
sisting of a stripe behind the eyes and three nearly parallel
stripes on the face, the middle one going up to the middle
ocellus, the lateral ones failing a short distance before top of
eye; scape long and wholly dark; flagellum ferruginous
beneath ; hind border of prothorax (continuous in middle,
but not reaching tubercles), tubercles, spot on tegule,
triangular spot just below wings, scutellum and_ post-
scutellum (except lateral corners) all yellow; basal area of
metathorax triangular, not bounded by a ridge, but abruptly
defined by the absence of punctures, finely shagreened,
shining green, appearing purplish in some lights; legs
brown-black, a light yellow spot at base of anterior and
middle tibiz ; apex of abdomen with black hairs. ‘The base
of the metathorax is radically different from that of P. dis-
juncta and P. albonitens, but is of quite the same type as
that of P. varicolor, which is closely allied.
Hab. Queensland (Gilbert Turner, 852, Ridg. 11.93, and
272 Hy.). Two.
Prosopis perviridis, sp. 0.
? .—Length about 8 mm.
Strongly punctured, the punctures of the mesothorax well
separated on a shining ground; brilliant green, even
including the legs, except the tarsi, the metallic colour of
which is purplish ; wings hyaline, with dark nervures. Head
broad and not especially long; mandibles and scape dark,
flagellum ferruginous beneath ; median and lateral stripes on
face and stripe behind eyes light chrome-yellow ; the median
stripe is expanded on clypeal margin to look like the head of
a nail in profile; in the supraclypeal area it is represented
by a round mark, and above that it is shaped like a spear-
head, not reaching the ocellus ; the lateral stripes have a
linear extension to the top of the eyes; hind border of
prothorax with two yellow stripes, but tubercles wholly
green; a small light yellow spot just beneath the wings and
a streak on axille; tegule green in front, with no light
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 27
402 Descriptions and Records of Bees.
spot ; scutellum and postscutellum broad and flattened, the
former shining, with widely separated punctures; basal area
of metathorax triangular, large, smooth and shining, bounded
by a groove ; anterior and middle tibiz marked with yellow
at base ; abdomen rather broad, the apical segments strongly
retractile ; the abdomen recalls that of a Ceratina.
Hab. Adelaide River (J. J. Walker, 5759). Collected on
the ‘ Penguin’ Expedition.
Prosopis reginarum, sp. D.
? .—Length nearly 10 mm.
Strongly punctured, the punctures on the mesothorax
close; deep blue, with purple tints, the mesothorax greenish
blue; light markings cream-colour; wings rather dusky,
nervures dark. This has in general the structure and
markings of P. perviridis, but it not only differs in its deep
blue colour, darker wings, and more closely punctured meso-
thorax, but also as follows: flagellum wholly dark; no
light stripe on axille; median facial stripe not reaching far
above antennze, the spear-head mark divided by a dark line.
The apical part of the abdomen has long coarse black hair.
The tubercles are fringed with white pubescence, and have a
minute, hardly visible, light spot.
Hab. Queensland (Gilbert Turner, 270 Hy.).
Prosopis Turneriana, sp. 0.
¢ .—Length 8-9 mm.
Strongly punctured, the punctures very dense on the
mesothorax; head and thorax bluish green, with purple
tints ; abdomen deep shining blue, with purple tints; ante-
rior coxe purple, but their femora olive-green; metallic
tints of middle and hind legs purplish blue; wings dusky at
apex, nervures dark. Head ordinary; face below antenne
white, the white ascending only a short distance above in
the middle line, giving way to a groove which ascends to the
middle ocellus; lateral marks ending very broadly and
bluntly about halfway between the antennz and the top of
the eyes; labrum with a light spot ; mandibles with a small
light stripe; scape creamy white in front, flagellum dull
ferruginous beneath; enclosure of metathorax triangular,
shining, of the type of P. varicolor; no yellow or white
marks on thorax ; tubercles shining reddish purple; tegule
with a light spot in front ; anterior and middle tibize lght
yellow in front ; apex of abdomen with two short widely
separated teeth; last three ventral segments strongly tufted
On the North-Atlantic Fin- Whale Fishery. 403
with black. The hind tibie may have a cream-coloured
stripe on basal half. The face-markings in the specimens
described are more or less discoloured.
? .—Similar, but the face is without light markings and
the area in front of the ocelli is brassy.
Hab. Queensland (Gilbert Turner, 383 Hy., males, and
269 Hy., females). Two of each sex.
The mouth-parts of this species seem not to be those of
true Prosopis; it is probable that the Australian species
assigned to this genus should form the basis of one or two
new genera, but I do not feel able to offer a plan of segre-
gation just at present.
Prosopis Lubbocki, n. n.
Prosopis metallica, Smith, Trans. Ent. Soc. Lond. 1862, p. 59 (Australia,
in coll. of J. Lubbock), g, not 2.
It seems very unlikely that the insects described as the
sexes of P. metallica are conspecific, and it will probably
cause less confusion if the male is separated under the above
name.
Prosopis Frederici, n. n.
Prosopis similis, Smith, Cat. Hym. Brit. Mus. 1853, p. 26 (Australia).
Not of Fabricius, 1793.
Prosopis Cameroni, n. 0.
Prosopis suleifrons, Cameron, Mem. Manchester Soc. 1897, p.51 (New
Zealand). Not of Smith, 1853.
University of Colorado,
Boulder, Colorado, U.S.A.,
June 27, 1905.
XLVI.—Some Results of the North-Atlantic Fin- Whale
Fishery. By THOMAS SOUTHWELL, F.Z.S.
Our knowledge of the specific distinctions, habits, and
distribution of the members of the family of Cetacea known
as the Baleenopteridz, or Fin- Whales, has until quite recently
been lamentably deficient, and even now it cannot be said to
be very complete. This has undoubtedly been owing to the
wide dispersal of these animals over the oceans of the world,
their gigantic size, and to the circumstances under which
if he
404 Mr. T. Southwell—Some Results of the
they until of late occurred having rendered their study a
matter of extreme difficulty ; for although members of the
family have long been more or less familiar objects in all
quarters of the globe, the only opportunities for their scientific
study have been the accidental stranding of individual
examples on the shores of civilized countries ; and these events
have too often occurred in desolate regions, far from the reach
of a competent authority, or even of individuals capable of
recording the features essential to the due recognition of their
species. It thus happened that, mainly owing to hasty
deductions from insufficient materials, species were unduly
multiplied and an almost inextricable state of confusion
arose; this, happily, through the recent labours of British
and Continental cetologists, has at last been reduced to
comparative order.
One great aid to the study of these animals has been the
perfecting by a Norwegian seaman, Captain Svend Foyn,
of Ténsberg, of an apparatus by which these giant whales,
hitherto regarded as too dangerous to be attacked by the
methods employed in the capture of the Polar whale, could be
successfully overcome, and this (although it cannot, in one
respect, be regarded otherwise than with extreme regret, seeing
that it risks the eventual extermination of these interesting
animals) has been the means of providing abundant materials
for a critical study of the various species —facilities which have
been duly utilized by the naturalists of HEurope and America.
The whales when captured are towed to the various shore-
stations, where their carcases are converted into oil and
wanure.
I may here remark that one of the results of the prosecu-
tion of this industry is the revelation of the astonishing
numbers in which these animals occur in summer in the
North Atlantic—a result altogether unlooked for, except
perhaps by the far-seeing Capt. Foyn; and especially has
this been the case with two species, Rudolphi’s Rorqual and
the Blue Whale (B. Sibbald’), which had been regarded as
extreme rarities.
I need not dwell futher on this portion of the subject than
briefly to chronicle the development of the fin-whale fishery ;
the success of Herr Foyn soon led to his example being
extensively followed, and on the expiration of his patent in
1882 many other companies were formed to hunt from the
Finmarken and Murmanski coast, till the Norwegian govern-
ment enacted restrictions which, added to the growing scarcity
of the whales, caused the opening of fresh stations in
Iceland, where the whalers had greater freedom of action,
North-Atlantic Fin- Whale Fishery. AOD
and subsequently of other centres in the Faroe and Shetland
Islands.
On the western side of the Atlantic also, in the year 1898,
a company, under the name of the “Cabot Steam Whaling
Company,” commenced operations, thus becoming the pioneer
of quite a fleet of vessels now engaged in this fishery from
Newfoundland.
It is to a comparison of the results obtained in these three
centres, remote from each other, that I wish now to call
attention. It may here be remarked that just as happened
when the fishery for Balena biscayensis in the temperate
water of the North Atlantic was abandoned in favour of that
of B. mysticetus in the Polar seas, the skill and seamanship
of the old Basque whalers were still requisitioned, so nowin the
extension of this new industry in Hurope and America it is
the Norwegian originators who fill the positions requiring
expertness and experience.
In the follewing notes I purpose to examine, taking each
Species in succession, the results of the operations in the three
great centres of the industry, with a view to ascertaining if
possible whether any racial variation is to be observed in the
members of the same species frequenting the several localities,
as indicated by appreciable differences in their habits,
frequency, or external characters. I shall therefore, avoiding
as much as possible the commercial aspect of the question,
confine my remarks mainly to the comparative abundance of
each species with any departures from the normal type, the
season of their occurrence, proportionate numbers of the
sexes, period of gestation, number of young produced, and
such minor considerations as may help to throw light upon
the identity or otherwise of the races frequenting the localities
under consideration.
As the pioneers of this industry the Norwegians claim our
first attention, and it is desirable to use the statistics of a
somewhat early period in order that they may be as distinct as
ossible from those of the more southern areas, which were
invaded later on. Iam therefore especially fortunate in being
allowed, through the kindness of Mr. Alfred Heneage Cocks,
to use the statistics collected by that gentleman during his
visits to the Finmarken whaling-stations in the years 1883
to 1889 *, on which what follows is mainly based.
The field of operation in the first instance was contiguous
to the coast, the whales, especially the large blue whale
(B. Sibbaldir), penetrating far into the fjords in search of
* ‘ Zovlogist,’ 1884-1890,
406 Mr. T. Southwell—Some Results of the
food; but as the result of persecution they soon discarded
the shore, and the final area in which their pursuit took
place was off the coast of Lapland to the north of 70° N. lat.,
especially between S6:den on the west and Varanger Fjord
on the east, but extending below 70° N. off the Murmanski
coast (Russia) to Swjatoi Nos in about longitude 40° E.
The fin-whales killed, in the order of their frequency, are as
follows :—
BALZNOPTERA MUSCULUS, Linn.
Norwegian Area.
The common rorqual (finhval, sildehval, or rérhval of the
Norwegians). This, the most numerous species, occurs from
March to the early part of September: 473 were killed in the
season of 1885, 646 in 1886, 463 in 1887, and from that
year the numbers began to decline; this decline in the number
of whales, added to the legal restrictions already referred to,
led to the removal of some of the factories to Iceland, the
result of which, so far as known, will be included in the
Shetland fishery.
The common rorqual is so well known a species that no
description of the normal form is needed, but it is subject to
considerable variation both in form and coloration ; the most
important of these is frequently referred to by Mr. Cocks as
well known to the Norwegian whalers under the name of
“ bastarder”’ or bastard whales, and commonly believed to be
the joint offspring of the common rorqual and the blue whale
(B. Sibbaldii). It appears to attain larger dimensions than
the type of the species, and is described as grey rather than
white on the under surface; the anterior baleen-plates are
white, and the remaining portions darker than normal; the
average length of eight females was 68 feet and of five males
70 feet, but 80 feet appears to be no uncommon length.
The “bastards” killed in 1887 averaged nearly 5 feet longer
than those of the normal type.
Another well-recognized variety is known as the “ lodde-
hval,” 7. e. capelan whale. This is the common fin-whale of
the north; it is white under the belly, and disappears at the
end of April. The fin-whale which then comes, Capt.
Ellevsen informed Mr. Cocks, “ has more or less grey colour
among the white, especially on one side [an asymmetrical
disposal of colour common to all the rorquals], its snout is
generally more pointed, and the whale is more slender and
North-Atlantic Fin- Whale Fishery. A)7
longer”; he adds that the whales which eat the capelan or
“‘Jodde” are only in Finmarken waters in the spring, and
“those that eat the ‘kril’ (Calanus finmarchicus), i.e. the
southern form, come later.’ ‘This variety is also known as
the “ langrér” (literally long-reed, as rorqual (rérkval) is the
Finmarken dialect for rérhval or reed-whale: Cocks), and
Capt. Horn, of Yeretiki, informed Mr. Cocks that of sixteen
rorquals captured by him off the Murmanski coast in 1859
all were of this variety, and not one normal fin-whale. ‘The
variation in colour taken by itself would not amount to much,
but in conjunction with several other minor differences in
form, food, and the season of their appearance would seem
to suggest subspecific value.
It is the opinion of experienced Norwegian whalers that
this species pairs in December or January, and that, as a
rule, the foetuses found in April are from 6 to 12 inches long,
whereas in August they are from 6 to 10 feet in length and
at birth 16 to 20 feet; but that there are many exceptions to
this generalization the following examples of actual dates
and measurements of foetuses will show :—
March.—A4 ft.
April.—1 ft. 43 in.-5 ft.
May.—2nd, female in milk ; 7th, 4 ft.; 19th, 4 ft. 2 in.;
20th; 4) tts 2him’, 5) 24st); ft. 2 io. 3 23rd; 1957, in.
June.—2nd, 4 ft. 5 in-3 ft. 8 in.; 10th, 8 ft.; 18th,
2 ft. Lin. ; 20th, 3 ft. ; 25th, 4 ft. 6 in.
July.—1 ft. 10g in. ; Sth, 4 ft. 2 in. and 6 ft.; 7th, 6 ft. 6 in.
andeS sits 1G ing; Sth Gultise9th;6.ft; 6 ins; 12th,
Myite Op inl. 16th; (8 ft.<d ane; .20this 3) ft. Tins;
2Ath, 7 ft.; 28th, 4 ft. 8 in.
August.—4th, 5 ft. 7 in.; 8th, 10 ft. 5 in.
A whale of this species which came on shore at North
Woolwich in November 1899 was delivered after death of
twins; and Mr. Haldane tells me that two foetuses were
taken out of a cow at Faroe last year (1904), also that
Capt. Castberg, of Ronas Voe, met with a similar case
two years ago. ‘hese are the only instances of twins which
have come to my knowledge.
The deductions to be drawn from the above appear to be
that this species in the North-east Atlantic produces a single
young one (on very rare occasions two) at rather irregular
dates in the autumn months, and that the period of gestation
is probably ten months. The destruction of so many gravid
females is a matter of great regret, but seems inevitable
408 Mr. T. Southwell—Some Results of the
under the circumstances. The quantity of oil yielded by one
of these animals varies from 800 to 1400 gallons, but
averages somewhat over 1000 gallons. 65°64 per cent. of
the whales killed in Finmarken, according to Mr. Cocks’s
statistics, were of this species. The average length of a large
number of males was 62 ft. 5 in., and of females 66 ft. 1 in.,
the proportions of the sexes being about equal. The food
consists of herrings, capelan (Mad/otus arcticus), and various
crustaceans (copepods), according to season.
The Shetland Area.
The gradual decrease in the number of the whales and
certain restrictions imposed by the Norwegian government
led Capt. Foyn to establish a station in Iceland about the
year 1883; but here, again, he met with difficulties from the
Danish government, and the enterprise was abandoned before
operations had commenced, but eventually they were success-
fully renewed and spread to the Faroesas well. In 1893 there
were said to be thirty steamers fishing from Iceland and
seven from the Faroes, and in 1902 nearly 2500 whales were
killed by them. In 1903 two stations were opened at Ronas
Voe, a third at Colla Firth in 1904, and subsequently a
fourth at Olna Firth, all on the mainland of Shetland. In
the Hebrides a station was also established at Bunaveneader
in North Harris. It is to the results of the operations in this
more southerly group of stations that we will now refer.
Mr. R. ©. Haldane, of Lochend, Ollaberry, Shetland, has
most kindly allowed me to avail myself of his valuable notes
on the fishery from Shetland, which has been carried on in
his immediate neighbourhood, and as the result of his
enquiries informs me that the Iceland fishery was at first
pursued on the west coast in Ise Fjord, Brede Fjord, &c.,
and gradually spread further west, most whales being taken
along the Greenland ice in July and August; latterly the
east coast has been resorted to. The present areas of the
fishery are, I am informed, as follows :—
Iceland. . . 683°-67° N. lat., 10°-30° W. long.
Faroe . . . 61°-64° - 5°—G° mn
Shetlands ye, G07=62° Wray. 9h OF ae Ls
Before entering upon the comparison of the statistics from
this area of the species under consideration (i. e. B. musculus),
it may be well to mention that a very unlooked-for feature in
the fishery has been the occurrence of a number of sperm-
North-Atlantic Fin- Whale Fishery. 409
whales. In 1903 a sperm-whale 68 feet long was killed by
a Shetland whaler out of a school of five, four of which
escaped, and five or six others were killed by the Iceland
vessels. A considerable school of these valuable animals,
estimated by some at thirty individuals, visited the seas to
the eastward of Iceland, and apparently wintered there, for
five others were killed by the Bunaveneader vessels in the
past season of 1904. But by far the most interesting capture
was a whale killed on 3rd July, 1903, in 61° N. lat. and
4° W. long., with whalebone 7 feet long, evidently an
Atlantic right whale (Balena biscayensis).
Of the first year’s operations at Ronas Voe there are no
statistics further than that 126 whales were killed; these
Mr. Haldane states, with the exception of the sperm-whale
already mentioned, “ one with barnacles ” (probably a hump-
back), and a_ bottlenose, were all the common finner
(B. musculus).
In 1904, 400 finners were killed from Shetland and 37
from Bunaveneader. Of the Shetland finners 236, or 59 per
cent., were males and 164, or 41 per cent., females, showing
a very large majority of males. Of the total catch of 506 of
various species from these islands, 86 (36 per cent.) were
finners. Mr. Haldane gives the average length of 135 males
as 62 ft. 9 in., and of 91 females 66 ft., an excess of 3 ft. 3 in.
over the males. The five longest males were 78, 77, 77, 75,
and 75 feet respectively, and the five longest females 78 (an
exceptionally large animal), 76, 75, 75, and 73 feet; but the
‘ Norrona’ has since killed a male, confidently asserted to be
of this species, 82 feet long.
The average length of the adult female is believed to be
65 or 66 feet, but a foetus 2 feet long was taken from an
example only 50 feet long. In a feetus 16 feet long the
baleen had not begun to show, but in one 22 feet long it was
well developed and the colour of the young one the same as
its mother; at birth the baleen should be quite 2 inches long.
The growth of the calf is very rapid, and suckers 40 feet
long have been seen; the general opinion among the Nor-
wegian whalers is that a calf of this size is not more than a
year old and that the suckers remain with the mother twelve
months. In support of the early sexual maturity of the
female Mr. Haldane mentions the example just referred to,
which, although only 50 feet long, contained a fcetus. The
whalers, who have had great experience, believe the period
of gestation in this species to be eleven months; and a table *
of measurements of eighteen foetuses given by Mr. Haldane,
* Annals of Scot. Nat. Hist., April 1905, p. 70.
410 Mr. T. Southwell—Some Results of the
varying from 6 inches to 16 feet, all obtained in one season
between the months of April and September, indicates that
the dates of birth must be very irregular. In three instances,
mentioned above, twins have been observed in this species.
Two fresh slips of whalebone of B. musculus kindly sent
me by Mr. Haldane were coloured blue-black on the outer
margin, toning down to blue-grey for the first third of their
width, and merging into clear cream-yellow with a few longi-
tudinal stripes of blue-black in the second third, thence to
the inner margin wholly cream-yellow, the hairy fringe the
same colour as the portion from which it took origin.
The slips of bone measured 25 inches from the gum, to
which the bristles added 8 inches, and at the gum they were
12} inches wide. A small anterior slip was 11 inches long
and 4 inches wide, bristles 34 inches long; wholly a clear
pale cream-colour and very transparent.
For about a month from 12th June in 1903 there was an
absence of “kril”’ in the water, and the whales fed largely
on herrings, but in 1904 “kril” (Thysanopoda inermis) was
abundant and they consumed very few herrings; it was
quite evident that the kril was their favourite food, and the
whales which fed on it were in better condition than when of
necessity they were feeding on herrings.
Mr. Haldane informs me that, in addition to the double
row of hairs found under the chin in a finner, he has found
“¢ another double row on the upper jaw, beginning 3 feet from
the snout and running up to just beyond the blow-hole.”
North-west Atlantic.
Passing to the western side of the Atlantic, we have now
to consider the results of the fin-whale fishery from the shores
of Newfoundland,
In the year 1896 the steamship ‘ Cabot,’ of 160 tons, was
fitted out by a company at St. Johns for the purpose of
hunting fin-whales after the manner of the Norwegians.
At first the operations took place in Notre Dame Bay, and
subsequently in Hermitage Bay, but the number of factories
was gradually increased till, according to the Annual Report
of the Newfoundland Department of Fisheries for 1904, there
were fourteen of these establishments in operation, extending
around the island and as far north as Cape Charles on the
Labrador coast ; and five others were expected to commence
operations in the spring of 1905. In the year 1908 there
were killed 858 whales, 345 of which were BL. musculus, and
North-Atlantic Fin-Whale Fishery. 411
in 1904 the number was 1275, of which 690 were the species
under consideration.
I am not able to give the proportion of males to females of
these 1035 “ finners,” but of the total catch of various species
for the two years, numbering 2133 (the sex of one was not
recorded), 1192, or 55°91 per cent., were males, and 940, or
44-09 per cent., were females. It may be well to give here
the numbers of each species (although they will be repeated
as they come separately to be considered). ‘They were as
follows :—
Biumerepee see 2 L035) Rudelphi 5 3 « + 39
Nulphurhobtomsiat +. 5/489.) Sperm. « . ~: 1
Humpbacks*. 4: 52068 | Unknown . ... (.. 4
The preponderance of finners will be noticed here also.
After the middle of July the finners are found in Notre Dame
Bay, where they are most numerous in August; in October
they become scarce and in poor condition, and finally leave
the coast.
In the year 1899 Dr. F. W. True, of the U.S. National
Museum, Washington, paid a first visit to the whaling-station
at Snook’s Arm, Notre Dame Bay, and as the result of this
and subsequent visits there appeared in 1904 a most
exhaustive treatise on the ‘ Whalebone Whales of the
Western North Atlantic,’ wherein, with all the thoroughness
which characterizes the work of that gentleman, he enters
minutely into the history of the species of the whales
frequenting those regions, and compares them with those
inhabiting the Huropean waters. I am thus enabled to avail
myself of Dr. True’s carefully compiled statistics in what
follows.
In 39 examples where the sex was noted 24 were males
and 15 females; the males averaged 58 ft. 7 in. and the
females 62 ft.3 in. Ten of the females killed in the month
of August contained foetuses or were accompanied by young:
of seven foetuses measured by Dr. True, one on the 5th
August measured 6 ft. 5 in., another on 15th August 15 ft.
2 in., and a third on the 27th August 6 ft. 103 in.; it is
evident therefore that, as in the European waters, the date of
pairing must vary considerably. No instance of twin foetuses
is mentioned.
The average length of the longest slip of baleen in six
individuals over 55 feet in length, measured from the gum
and exclusive of the bristles, is given as 214 inches.
The following table shows the average measurements of a
412 Mr. 'T. Southwell—Some Results of the
large number of Norwegian and Shetland finners, which,
although in excess of like averages of the Newfoundland
whales given by Dr. True, is still less than the average of
the Norwegian whales given for comparison in his table ;
this is probably owing to my having rejected the so-called
“ bastard’? whales, the inclusion of which would have
considerably increased the average.
The figures given below for the Newfoundland whales are
from Dr. True’s table and the Report of the Newfoundland
Department of Fisheries, and those for Norway from
Mr. Cocks’s reports.
Balenoptera m usculus.
Length.
Number and proportion of
E SEXES. | Length of
Region. Males. Females. heel Syke
Total. | Males. |Females.| Max. | Med:| Max. | Med.
ft. | ft.in.| ft.in.| ft. in.
Norway ......... 240 118 122 75 | 625° | 77 1°) 66-1 36><12am:
Shetland ...... 400 236 164 78 | 629] 78 O | 66 O | 25x 123 in.
Newfoundland .| 39 24 15 65.757 12 70"S "62" 3 ia,
The “ bastard ” whales are not included in the above: of
these, 16 males averaged 66 ft. 2 in., and 16 females
68 ft. 5 in.; of these several were evidently immature, thus
reducing the average. The longest male measured 72 ft. and
the longest female 80 ft. 6 in.; omitting all those under 65 ft.
as immature, 7 males averaged 69 ft. 5 in., and 13 females
Wut Zain
Mr. Haldane found a feetus in a finner 50 ft. long, but the
ordinary length of a sexually mature female would probably
be over 60 ft.
Of 2893 fin-whales killed in Norway, 189, or 65°64 per
cent., were of this species, and of 2131 killed in Newfoundland
the percentage was 48°56.
The Norwegian foot has been taken as 123 English inches
in the above calculations.
It remains to consider what, if any, is the taxonomic value
North-Atlantic Fin- Whale Fishery. 413
of the variations observed in these animals, and it is con-
venient to do so with regard to the present species, which is
far the best known and with regard to which there is so much
more material on which to base conclusions. Dr. True
minutely examined ten freshly killed specimens in New-
foundland and compared them with Sars’s careful description
of a Norwegian individual, the whole showing conspicuously
the great individual variations to which the species is subject
so far as the intensity and distribution of the body-colours
are concerned, the constant features being a dark left lower
lip, white right lower lip, white anterior right whalebone,
and the marked asymmetry of coloration. Where individuals
differ so materially it would be unsafe to generalize on the
mere shade or disposal of colour; Dr.'True, however, instituted
a very careful fale es of the measurements of the Kast
and West Atlantic fin-whales, and found that the former were
larger in all their proportions than the latter, which is con-
firmed by the foregoing table so far as length is concerned. He
does not from this fact conclude that there exists even sub-
specific differences between those inhabiting the two sides of
the Atlantic ; but I think it clearly demonstrates that the herds
frequenting the two distant areas do not intermingle, their
normal line of migration being south to north in summer,
rarely, if ever, under normal conditions latitudinal, and that
they may be considered racially distinct. Perhaps we may
go even farther than that, and regard the gigantic “ bastard ”’
whale of Mr. Cocks, which has been known to reach the
enormous length of 80 ft. 6 in., as at least a distinct race; and
probably the same may be said of the southern form of
B. musculus, the “ sildehval ” of the Norwegians, which goes
north in summer to occupy the area in the ocean already
vacated by the “ loddehval,’” which has then departed to a
more northerly latitude. That the species may be cosmo-
politan cannot at present be conclusively disproved, but, as
Dr. True very pertinently remarks, “ even should it be
demonstrated that the species of large whalebone whales are
cosmopolitan, it does not follow that the individuals consti-
tuting these several species range throughout the globe, the
probabilities are much against such world-wide movements.”
So long ago as the year 1898* I endeavoured to show with
regard to the Polar right whale that the areas frequented by
separate races or herds of those animals were vastly more
restricted than was generally supposed ; and I think expe-
* “On the Migration of the Right Whale,” Natural Science, June
1808, pp. 897- AL,
414 Mr. T. Southwell—Some Results of the
rience will show the same to be the case with the fin-whales
inhabiting the North Atlantic Ocean.
With regard to the recorded measurements of these animals,
in some instances they are taken in a straight line, in others
along the curves, and till a system is adopted ensuring
uniformity, although useful in their way, they cannot be
regarded as scientifically accurate, and are therefore only
approximately useful for comparison.
BALANOPTERA SIBBALDII.
Norway.
Sibbald’s rorqual, the blaa-hval or blue whale of the
Norwegians and sulphur-bottom of the American whalers, is
the chief prize of the whalemen; it yields an average of
2800 gallons of oil, occasionally even as much as 10,000
gallons. Mr. Cocks states that twenty-five years ago
Varanger Fjord was a favourite resort of this huge animal in
summer, and that it penetrated far into the fjord in search of
the “kril’?? (Thysanopoda tinermis), on which it delighted to
feed; all this is now a thing of the past, constant harrying
has taught the survivors to keep out to sea, and, in addition,
no whales are allowed by law to be captured at a less distance
than 7 kilometres * from the shore.
The time for the appearance of this whale upon the
Finmarken coast is, as a rule, about the middle of June,
exceptional instances have been known in May, and they have
departed by the middle of September—the date of departure
probably being dependent on the falling-off of the food-
supply, and not due, as was formerly supposed, to the
decreasing temperature of the sea. Mr. Robert Gray, when
in the ‘ Eclipse’ whaler on April 27th, met with blue whales
in lat. 70° 14’ N., and speaks of the partiality of this species
for water of very low temperature; its departure, therefore,
from the Finmarken coast cannot be directly attributed to
declining temperature.
Dr. Guldberg describes this species as blue-grey or blue-
black above, and sometimes copper-brown when rolling about
in the sea; it has always a more or less pronounced blue-grey
colour on the back—hence its trivial name. ‘The underparts
are minutely flecked with white and grey, but it varies
considerably in coloration. ‘I'he baleen-plates are black, with
black hairs, about 36 inches long by 24 inches broad at the
gum. Mr. Cocks remarks that “the sex of a blue whale (and
in a less marked degree it is, I believe, true of other species
* This for ten years.
North-Atlantic Fin-Whale Fishery. 415
of Balenoptera and possibly of other whalebone whales) may
be distinguished by the shape of the baleen-plates, which in
a male are long (up to 4 feet including gum) and narrow,
but thick; while in a female they are short and broad, but
thinner.” The baleen is much infested with a species of
copepod (Balenophilus unisetus).
The greatest length given by Mr. Cocks is that of a female
88 ft. 7 in. long, and the sexes occur in about equal numbers.
There is no certain information as to the season of pairing,
but two males are said to have been ‘‘ making overtures” to
a female on August 5th. Capt. Sorensen believes they have
no fixed time for pairing, and foetuses have been found in
Aueust Measmnon lo: ti. 1S, ft. Ii, 9) or 10 ff.,, and
15 ft. 6 in.; Mr. Cocks, however, does not vouch for the
exuctness of these measurements. Calves have been seen
following the mother estimated at 40, 50, or even 60 feet
long. ‘The probable length of the sexually mature female is
about 70 feet. Dr. Guldberg is of opinion that the blue
whale goes with young eighteen or twenty months, and that,
judging mainly from the large size of the young ones which
have been seen following the mother, she produces only
every third year; but perhaps he does not fully allow for the
rapid growth of the young one. ‘Two foetuses, both males
6 and 4 feet long respectively, were found in a female only
60 feet in length which was brought into Eide Fjord, Faroe,
in June 1894, as recorded by Mr. Harvie-Brown in the
Ann. of Scottish Nat. Hist. for April 1905.
No blue whales were taken from Shetland, but, in addition
to that already mentioned as killed from the Faroes in 1894,
forty-two of these whales and five sperm-whales were killed
last season (1904) from the station at Bunaveneader, Harris,
which would seem to indicate that this species follows a more
westerly course on its northward passage than the other
fin-whales.
Newfoundland.
Turning to the consideration of the blue or sulphur-bottom
whale, as it is known by the American whalemen, in the
Western Atlantic, we find, according to Dr. True’s careful
statistics, that, like the common fin-whale, the dimensions of
the Newtoundland examples are on the average considerably
less than those found in the European waters, although
agreeing fairly with the latter in other respects. In colour
there is a close resemblance, subject to the same considerable
individual variations, but all exhibiting more or less of the
characteristic irregular ‘ milk-white spots” below the
pectorals on the fluted sides of the breast described by Sars.
416 Mr. T. Southwell—Some Results of the
The only important difference which Dr. True points out is
in the coloration of the dorsal fin, which in the Newfoundland
specimens ‘is usually more or less white or whitish, except
on the margins, with darker curved lines extending up
vertically from its base”?; but he remarks that this fin is
not fully described in a fresh condition by any European
authority. There seems to be no appreciable difference in the
colour of the baleen. Dr. True arrives at the conclusion
that, notwithstanding its superior dimensions, there is no
sufficient reason for separating the European blue whale from
the American sulphur-bottom.
Balenoptera Sibbaldit.
| Length.
|
|
| Number and proportion of |
' SEES: | Length of
Region. | | Males. Females. | i
f
| |
| Total. | Males. | Females.; Max. | Ay. ax. | Ay. |
|
ene fe he — | ates
| ft.an.,| ft.ims| ft. in) stam:
Norway ......... 104 | 47 | S57 | 8 5/)%5 4 | 887) 77 0) sbx2tie:
| | |
Newfoundland.| 25* | 10 | Leal
| |
727 | 68 8 | 72 2| 68 9 | 23-32 in.
Taking the average length of sexually mature females at
72 feet, Dr. True arrives at an average of 74 ft. 8} in. for
that sex in Newfoundland; subject to the same test the
Norwegian females average 79 ft. 3 in.
Of 2893 fin-whales killed in Norway, 338, or 11°68 per
cent., were of this species; in Newfoundland of 2132, 489, or
22-94 per cent., were B. Sibbaldia.
None of these whales have hitherto been killed in Shetland.
MEGAPTERA LONGIMANA (Rudolphi).
Norway.
The third large species of fin-whale is known to the
British whalemen as the humpback (from its low dorsal fin),
* In another table Dr. True gives the average of a larger number
of these whales from other sources—57 males equalling 70 ft. 8 in. and
35 females 70 ft.; but the above are measurements carefully taken by
himself.
North-Atlantic Fin- Whale Fishery. A417
to the Norwegians as the knélhval, pukkelhval, or troldhval.
Although it was not fully recognized as a Kuropean species
till the year 1829, when Rudolphi described and figured a
specimen east ashore at the mouth of the Elbe, it has been
found fairly plentiful on both sides of the North Atlantic ;
owing, however, to the fact of its sinking when killed and to
its small yield of oil it is not considered a very desirable
capture. It is of moderate length, but bulky in proportion,
and yields from 600 to-800 gallons of oil: Mr. Haldane,
however, heard of a Finmarken specimen, 50 feet long and
16 feet in diameter, which yielded 125 barrels of oil; as the
petroleum barrels used by the Norwegians are of the capacity
of about 40 gallons, this would represent some 5000 gallons!
A marked characteristic of this species is the great length
of its pectoral fins: in a specimen 40 ft. long the pectoral
fin measured 12 ft. in length and 2 ft. 8} in. in its greatest
breadth ; in a 44 ft. example measured by Mr. Cocks the
flipper was 13 ft. 9 in. to the skin of the axilla and 3 ft. 7 in.
in width; the height of the dorsal fin was 9 in. The
pectorals also vary exceedingly in colour, being sometimes
black on the upper surface and white below, or in some
instances entirely white on both surfaces. ‘The body-colours
are equally variable; the throat and nearly all the underside
may be white or entirely black. These variations do not seem
to bear any relation to sex orage. Another peculiarity is the
extent to which they are generally infested by external
parasites—Cyamus in abundance and a cirriped (Diadema
coronula), the latter in turn bearing the curious Conchoderma
auritum. Mr. Cocks also mentions the loud screaming of
the humpback when harpooned, which he compares to the
gruesome sound uttered by a pig when being killed. The food
of this species appears to be rather mixed, consisting of small
fish as well as kril; possibly, as with the common finner,
this may be the result of circumstances. ‘The baleen is
black, with yellowish-brown hairs, and the longest plates
measure about 2 feet.
The humpback comes on the Finmarken coast very early
in the year, having been killed on the 24th of February, and
departs in the middle of September. The date of pairing is
uncertain, as is also the period of gestation; but foetuses have
been found in June (no day mentioned) 12 in. long, again on
June 21st 13 in., July 1st and 15th each 13 in., and on the
28th 174 in. (Norwegian), on August 21st 34} in.—thus
showing a more regular approach to maturity than in most
species.
Capt. Wiborg informed Mr. Cocks that in 1886 he saw a
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 28
418 Mr. T. Southwell—Some Results of the
humpback followed by two calves each 10 Norwegian feet
long, but he does not give the date, and Scammon gives
a beautiful plate of a humpback suckling two calves; the
birth of twins, therefore, in this species seems to be of
oceasional occurrence. [ar more males than females of this
species are killed ; of 37 taken in 1886 28 were males. The
average length of 27 males was 38 ft. 5 in., and of 13 females
42 ft. 3 in. The largest specimen Mr. Cocks records was a
male 53 Norwegian feet (55 ft. 3 in., English) long; he
mentions the girth of a female 52 English feet long as 40 feet,
but this measurement in a dead whale is absolutely valueless,
as distension commences immediately after death. The total
number killed in six years was 817, and the largest number
in a single year 96.
Shetland.
Of the results of the Shetland fishery I have very little
information, and no particulars as to species or measurements
before 1904 ; in that season eight humpbacks were taken by
the Shetland boats and five from Harris in the Hebrides.
The Shetland vessels, Mr. Haldane informs me, fish in about
60° to 62° N. lat. and 0° to 5° W. long.; the Faroe boats
work to the southward and the Shetland vessels as far
as 120 miles north of those islands. Of the Shetland hump-
backs four females measured 30, 39, 40, and 46 feet respec-
tively, three males 47, 49, and 46 feet; the last male had
pectoral fins 13 ft. 10 im. long. The colours varied as already
described. ‘The food of those which Mr. Haldane examined
consisted of kril and a few shrimps; herring also form
part of their diet. The average yield of oil was 1200 to
1600 gallons.
Newfoundland.
In the Newfoundland fishery this species figures largely :
in the season of 1893 287 were killed, and 281 in 1904;
unfortunately no statistics as to sex or dimensions have been
collected at these stations—desiderata which, I hope, will, at
least to some extent, be supplied in the present season,—and
therefore nothing can be added to the investigations made
by Dr. True. That gentleman only had the opportunity of
personally examining three specimens at Snook’s Arm in
August 1899, but of these he gives minute descriptions: he
also gives the dimensions supplied to him of 18 others killed
in the two following years between April 26th and July 6th,
North- Atlantic Fin- Whale Fishery. 419
the bulk (12) being killed in May, 21 in all; of these 10
were males, 4 females, and 7 sex not recorded. By the table
it will be seen that the maximum of the Norwegian specimens
of both sexes considerably exceeded that of those taken in
Newfoundland, but in other respects (colour, dermal tubercles,
dorsal and pectoral fins, and flukes) they present no material
difference. Dr. True is of opinion that the various nominal
species into which the humpbacks of the North Atlantic have
been divided are all referable to one species, and in this he
agrees with Kuropean cetologists; but considering the
extensive range thus assigned to this whale it seems probable
that the individuals inhabiting, say, the West Indies or the
Californian coast are racially distinct from those frequenttng
the Greenland seas.
Megaptera longimana.
Length.
Number and proportion of M
: sexes. | Length of
Region. | Males. | Females, baleen
TspalwnleMiates s Homies Mase | Ary. ||! Mare’ |t Aw.
a Ghded vs F
fim. |) fGen. | tt. 1m.) fits im: |
NOR Way. 1.00025 40 oT 13 M5) ah!) ets) 2 1) fap) (0) | 2) Bs | Seon
|
Shednddiees 4: 7 3 4 |49 0| 474/460] 389
Newfoundiaia:| 14 | 10 4 |46 11|376|466| 416 | 22in.
|
Of 2893 fin-whales killed in Norway, 236, or 8:16 per cent.,
were humpbacks.
BALZNOPTERA BOREALIS (Lesson).
Norway.
This species, known to the Norwegian whalers as the
sejhval (coal-fish whale), until the establishment of the
Finmarken fishery believed to be of excessive rarity, is found
in varying numbers, singly or in schools of perhaps fifty
individuals, every summer in the months of May and June oif
the coast west of the North Cape. Up to the year 1885 it
could only be regarded as a strageler eastward of that cape.
In the year mentioned, however, Mr. Cocks wea asaured that
28"
420 Mr. T. Southwell—Some Results of the
they were commen along the whole coast hunted by the
Norwegian and Russian whalers during nearly the whole
season, and an immense number remained off Sylte Fjord
(Kast Finmarken) for about three weeks in May and June.
They appear, however, to have reached the Russian coast
somewhat later, the first killed by one of the Yeretiki
steamers being on July 10th; the large number of 771 were
killed during that season, these being fairly distributed
among all the vessels of the fleet. In 1886 only 62 were
killed (40 of which fell to one vessel), the whales having
again resorted to their habitat westward of S6réen. In 1887-
they were again abundant, and 245 were killed, 110 of these
by Capt. Bull, chiefly between Loppen Island and Reisen, the
first on May 21st and the last on the 12th of August. The
next year the number fell to 92, but in 1889 it again increased
to about 113—thus showing this species to be a constant
visitor in considerable, but fluctuating, numbers off the
Finmarken coast.
It is not much sought after by the whalers, and owing to
its small size and proportionately small yield of oil is neglected
when other species are to be obtained.
As is the case with all the members of this family, the date
of pairing, period of gestation, and season of calving are not
known with certainty, but the following measurements of
foetuses, with the dates of their occurrence, afford some
indication :—
Date. Length of foetus.
ifs sbay
Jamee2s hes ea ns See LO
July. 3 tee ee a
US op ites een eis 4 ; } two foetuses.
26 8. 5
28 ae 5. 6
ANIC OW eam toes Gare OAR
10 Pac 6 O
10 ext)
15 avis 9 0
A female, 46 ft. 10 in. long, brought into the factory at
Tufjord on the 18th July, 1885, contained twin feetuses of
opposite sexes. Prof. Collett (P. Z. S. 1886, p. 261) mentions
a similar instance of twins ina whale of this species 43 ft. long
captured at the entrance to the Varanger I'jord, which con-
tained two young ones each 6 ft. 7 in. long. So far as lam
North-Atlantic Fin-Whale Fishery. 421
aware, these are the only instances of twins in this species
on record.
In 384 instances where the sex is given, 17 were males and
17 females: the former averaged 45 ft. 4 in. and the latter
46 ft. 7 in. in length ; the longest male was 51 ft. 1 in., and
the longest female 52 ft. 1 in. The measurements are
variously given in Norwegian and English feet; I have
reduced the former to English measurement on the scale of
124 inches to the Norwegian foot.
The yield of oil averages about 200 gallons, rarely as
much as 280 gallons. ‘The food of this species is believed to
consist very largely of “ kril” (Lhysanopoda tnermis), a
small crustacean, and Calanus finmarchicus (a copepod).
Professor Collett visited the factory at Vard6é in 1885, and
contributed an excellent paper on the “‘ external characters ”’
of this species, of which he stated that even then our know-
ledge was very deficient, to the P. Z. 8. 1886, pp. 243-265,
with illustrations.
The baleen is black, with white bristles, and the longest
plate measures 31 inches.
Of 2266 fin-whales killed in Norway in the years 1886,
1887, and 1889 (omitting the abnormal kill of 771 in 1885),
42(), or 18°5 per cent., were Kudolphi’s rorquals.
Shetland.
Of 507 fin-whales killed from Shetland and the Hebrides
in 1904 only 9 were of this species, and I have no particulars
as to sex or measurements further than that two were bulls |
37 and 41 feet long respectively.
Newfoundland.
None of these whales came under the notice of Dr. True
when he visited the Balewna Station at Hermitage Bay, nor
do any appear to have been killed till 1904, when 5 were
obtained trom Balena, 30 from Bay Chaleur, and 3 from
Rose au Rue; there therefore exists no material for comparison
of specimens from the three fishing-centres.
The lesser rorqual (B. rostrata) does not figure in the
returns from any otf the above “ fisheries.”
422 Mr. O. Thomas ox
XLVIJ.—On some Australasian Mammals.
By OLpFIELD THOMAS.
The Nomenclature of the Two Bats of New Zealand.
Tn 1889 *, taking Dobson’s synonymies and identifications of
the two bats found in New Zealand as correct, I amended, on
a purely nomenclatural point, the names he gave to them in
his Catalogue, showing that on his data tuberculatus was to
be used for the Mystacops and morio for the Chalinolobus.
But it has been pointed out to me by my American friends
Messrs. Palmer and Miller that there is an earlier work, most
pertinent to the subject, not quoted by Dobson at all, and
therefore ignored by me, and this upsets the conclusions I
came to. Moreover, Dobson’s identification of the New
Zealand Chalinolobus with one from Australia proves to be
quite incorrect, the former being perfectly distinct from any
occurring on the mainland. This frees the New Zealand bat
from the competition of the earliest name of all, morzo, Gray,
1841 (type locality Tasmania f).
The relations to C. morio of the respective owners of the
names microdon, Tomes, 1859 (South Australia), and signifer,
Dobson (Queensland), may be left for further material to
elucidate, but all three are different from the New Zealand
Chalinolobus, whose skull has a much longer and heavier
muzzle than any of them.
* Ann. & Mag. Nat. Hist. (6) iv. p. 462.
+ In Grey’s ‘ Journals of Expeditions,’ Appendix, ii. p.405. The con-
fusion in Gray’s account of his three species of “ Scotophilus” (S. morio,
Gouldi, and australis) is very great and almost defies elucidation, though
the fault may have been as much the printer’s as the author’s. Among
other things the measurements given as those of S. morio are evidently
those properly belonging to S. Gouldt. As a preliminary step towards
clearing the matter up, I have gone carefully over the specimens which
were before Gray at the time, over his notes in the registers and in his
MS. list of bats, the basis of the published list of 1843. In this way I
have identified no. 37. 4,8. 118 (90. a”) (Dobson’s Chalinolobus tubercu-
latus, e, not f as he stated), from Tasmania, coll. Sir W. J. Hooker, as the
type of S. moro, while no. 41. 1516 (118. a”) e of Dobson’s C. Gouldi
should be taken as the type of the latter species. It was from Launceston,
Tasmania, Gould collection, not, as Dobson says, from Lieut. A. Smith,
whose specimens came in tweuty years later.
Dobson's “ R. Maitland, isq.,” the donor of specimen h, also proves to
be the River Maitland, New South Wales, its locality.
I can identify no individual type of S. australis, which is, however,
evidently a synonyin of moro,
some Australasian Mammals. 423
The point about the double usage of the name tuberculatus
has hitherto been that, as Forster’s description of the Chalino-
fobus was not published till 1844, Gray’s description of the
Mystacops in 1843 (Voy. ‘Sulphur, Mamm. p. 23) had
absolute priority, whether he thought it was Forster’s
tuberculatus or not.
But in the work to which my attention has been drawn,
Dieffenbach’s ‘ New Zealand,’ 1843, Gray, by publishing in
the Appendix (ii. p. 181) five words of description of
Forster’s plate, then unpublished, not only cut him out with
regard to the Chalinolobus, but invalidated his own tubercu-
latus for the Mystacops, which he first used on p. 296 of the
same work, distinctly stating his opinion that it was the same
bat. As a result, tuberculatus being invalidated for the
Mystacops, Hutton’s velutinus will come into force, and the
names and chief synonymy of the two species will be as
follows :—
Chalinolobus tuberculatus, Gray.
Short-eared Bat.
Vespertilio tuberculatus, Gray (ex Forst.), Dieffenb. N. Zealand, ii
p- 181 (1843) ; Forst. Deser. Anim. (ed. Licht.) p. 62 (1844).
Scotophilus tuberculatus, Tomes, P. Z. 8. 1857, p. 135; Hutton, Trans.
N. Z. Inst. iv. p. 185 (1871).
Chalinolobus tuberculatus, Peters, MB. Ak. Berl. 1866, p. 680, and
1867, p. 480; Dobson, Cat. Chir. B. M. p. 248 (1878) (the New
Zealand specimens only).
Chalinolobus morio, Thos, Ann. & Mag. Nat. Hist. (6) iv. p. 462 (1889)
(in part.).
Mystacops velutinus, Hutton.
Long-eared Bat.
Mystacina tuberculata, Gray, Dieffenb. N. Zealand, ii. p. 296 (1843),
nec Vespertilio tuberculatus, id. tom. cit. p. 181; Dobs, Cat. Chir.
B. M. p. 442 (1878). : ¥
Mystacina velutina, Hutton, Tr. N. Z. Inst. iy. p. 185 (1871). i"
Mystacops tuberculatus, Lydekker, in Flow, & Lyd. Mamm. p. 671
(1891).
Dobsonia magna, sp. n.
Similar in all essential respects to D. paliata, Geoff.*, but
very much larger, the skull especially being conspicuously
larger in all dimensions.
* Cephalotes Peronit, Dobs. et auct. al.
424 Mr. O. Thomas on
Colour about as in LD. paliata, the head similarly dark
bistre-brown and the nape drab-brown ; under surface dull
isabella-brown. Back more entirely naked than usual,
absolutely hairless everywhere except just over the base of
the tail.
Skull very similar in its proportions to that of D. paliata,
but very much larger, the difference corresponding with that
found in Nyctymene aello as compared with WV. major, while
Dobson’s Dobsonia minor may similarly rank with the
smallest species, Nyctymene cephalotes, of the allied genus.
These differences may best be seen by the dimensions given
below. ‘Teeth on the whole broader in proportion to their
length than in JD. paliata, but there is a good deal of
variation in this respect in the latter species.
Measurements of the type (those marked with an asterisk
taken by the collector in the flesh) :—
Forearm 145 mm.
Head and body 200* ; tail 26*; ear27*. Third finger:
metacarpus 93, first phalanx 68, second phalanx 103; fifth
finger 188, Lower leg and foot (c. u.) 107.
Skull (measures in brackets those of an adult male skull
from Sumba): greatest length 62 (51); basal length 55:5
(46); muzzle, from tip of nasals to hinder side of postorbital
processes 28°5 (22°5) ; interorbital breadth 10:7 (9°8) ; inter-
temporal breadth 9 (8°3) ; palate length in middle line 29 (23) ;
front of canine to back of m? 26°5 (21:5); length of canine
on hinder edge from cingulum 8°8 (5:2). Lower jaw, condyle
to front of canines 50 (40) ; front of canines to back of m,
28 (22:5).
Hab, Tamata, Northern British New Guinea, near German
frontier. Alt. 100 feet.
Type. Male. Original number 65. Collected 9th Sep-
tember, 1904, by W. Stalker.
Dr. Matschie considers the members of this genus so
variable in size that he unites them all in one species, even
including the little D. minor, Dobs. But none of the dimen-
sions he gives at all approach those of this Papuan example,
and a comparison of the skulls shows such an immense
disproportion between it and those representing true D. paliata,
that it seems necessary to separate it.
No other species has been described from this part of the
Papuan region.
Cr
some Australasian Mammals. 42
Petrogale lateralis Hackett’, subsp. n.
Closely similar to true P. lateralis in external and cranial
characters, including both colour and pattern of markings,
but larger and with larger teeth, especially with larger
anterior premolar or ‘ secator” * (p* of the Catalogue of
Marsupia!s).
Dimensions of the type (measured in skin) :-—
Head and body 660 mm. ; tail 540; hind foot (s. u.) 139;
ear 0d.
Skull: greatest length 107; basal length 93; greatest
breadth 50; front of orbit to back of skull 62; nasals,
length 47, greatest breadth 14°3; interorbital breadth 18:3 ;
diastema 20; palatal foramen 8; large secant premolar 8°3
x41; combined length of three anterior molariform
teeth 18°5.
Hab. Mondrain Island, off the coast of S.E. Western
Australia.
type. AGult male... B.M. no. 9. 42 1,.5.. « Onginal
number 7161. Collected Ist November, 1904, by J. T.
Tunney. Presented by the Western Australian Museum,
Perth. Fourteen specimens examined.
While in colour and other external characters I can find
no difference between the Mondrain-Island rock-wallabies and
the typical specimens of P, lateralis described and figured by
Gould, there is so marked a distinction in the size of the
skulls and teeth, especially of the large secant premolar, that
it is clear that the former cannot be referred to Gould’s
animal,
Among five skulls of P. lateral’s none exceed 95 mm. in
total length, the front of the orbit about 56 mm. from the
back of the skull, and the upper secator 6°8 mm. in length
by 3:1 mm. in breadth, the tooth markedly narrowing
forwards, while in facketti it is a long and comparatively
broad oval, nearly as broad in front as behind. Below, the
* This tooth, the most important for systematic work in the Marsupials,
might conveniently be called the “‘secator,” thus avoiding altogether the
confusing question of its serial homology. It is the “‘P*” of the Cata-
logue of Marsupials, but is now more usually believed to be homologous
with p*, or, more probably, mp%, of Placental mammals, This change of
opinion, even if it were accepted universally—and the question is still
under discussion,—renders the use of a numerical symbol exceedingly
inconvenient for ordinary systematic descriptions ; and it is hoped that a
special name for the tooth will be of as much value to workers on
marsupials as ‘‘carnassial ” has proved to writers on Carnivora,
426 Mr. O. Thomas on
corresponding tooth is also conspicuously broader and
heavier in //acketté than in true lateralis.
I think it probable that P. 1. Hacketti is not a true insular
form, but that it will be found to occur in suitable localities
on the mainland of 8.E. West Australia, diminishing in size
and strength of dentition northwards.
In accordance with a suggestion of Mr. B. H. Woodward’s,
I have named this handsome rock-wallaby in honour of
Dr. Hackett, Chairman of the Western Australian Museum
Conimittee.
Thalacomys * sagitta, sp. 0.
In the Catalogue of Marsupials all the specimens of
Thalacomys—there called Peragale—were referred to a single
species, 7h. lagotis, without distinction between the West-
Australian and South-Australian examples. But the recent
presentation of a fine specimen from South Australia by
Mr. Hillier has drawn my attention to the fact that the form
found in that district is so considerably smaller than the West-
Australian one that the two ought certainly to be separated.
The difference is mainly in the skulls, and may be readily
appreciated by comparing the measurements given below of
the South-Australian animal with those of the typical West-
Australian form published in the Catalogue.
Externally, apart from the corresponding difference in
size, there is little to distinguish the two forms. On the
whole Th. sagitta is a little paler in tone, the black band on
the tail is slightly shorter, being shorter instead of longer
than the white end, and the feet are paler below, the black
only extending about a third of their length underneath the
heel.
Dimensions in the flesh (measured by Mr. Hillier) :—
Head and body 316 mm.; tail 215 ; hind foot 91; ear 79.
Skull: greatest length 85; basal length 76°5; greatest
breadth 38; nasals, length 40, greatest breadth 7°5 ; inter-
temporal breadth 13; palate length 50; basicranial axis 22°5 ;
basifacial axis 54; front of canine to back of last molar 36;
combined length of three anterior molariform teeth 12°5.
Hiab. South Australia. Type from Killalpanima, east of
Lake Eyre.
Type. Adult male. B.M. no. 3. 5. 21.2. Collected 6th
April, 1903, and presented by H. J. Hillier, Esq.
* Cf. Aun. & Mag. Nat. Hist. (7) v. p. 223 (1900).
some Australasian Mammals. 427
Five skulls of this species are in the Museum collection
all of practically the same dimensions, while two from West
Australia are both of the larger size indicated by the
Catalogue measurements.
Th. sagitta is clearly the “common species or § Urgarta,’
Peragale lagotis,” of Prof. Baldwin Spencer’s description * of
a new species of this genus from Central Australia, 7h. mznor,
which he distinguishes, no doubt rightly, by its still smaller
dimensions and its different habits.
Mr. Hillier gives the local name for this animal, as used
by the Diari natives, as ‘* Kapita.”
¢
Phascogale Hillier, sp. n.
Near Ph. cristicauda, but conspicuously paler in colour
throughout.
Size about as in PA. cristicauda. Fur soft and fine; hairs
of back about 9 mm. in length. General colour above a
beautiful soft drab-colour, similar to but decidedly paler than
Ridgway’s ‘ecru-drab” ; a few fine longer hairs black,
contrasting curiously with the general pale colour. Sides
more whitish drab, passing without sharp line of demarcation
into the pure white under surface, on which region, from
chin to anus, the hairs are pure white to their bases. ‘lop of
head like back, but rather more fawn. Cheeks and lips pure
white. Hars thinly clothed with whitish hairs, an incon-
spicuous whitish patch behind their posterior bases. Upper
surface of hands and feet snowy white ; structure of feet
apparently as in PA. cristicauda. ‘Tail slightly incrassated
basally ; crested above for its terminal two inches, the hairs
attaining a length of 16 mm.; nothing that can be called a
crest below, though some of the hairs towards the end
are slightly longer than on the proximal part. In colour
the main part of the tail is rather more rufous than the
general body-colour, though nothing like the corresponding
part in Ph. cristicauda ; under surface indistinctly darker, not
black ; crest glossy black.
Skull unfortunately lost.
Dimensions of the type (measured in the flesh) :—
Head and body 150 mm.; tail 100; hind foot 30; ear 27.
flab, Killalpanima, Lake Eyre East, South Australia.
Type. Male. B.M. no. 5. 8. 28.1. Collected 13th Feb-
ruary, 1905, and presented by H. J. Hillier, Esq.
* P. Roy. Soc. Victoria, ix. p. 6, pl. ii. (1897).
428 Prof. C. Chilton on the Distribution of
This beautiful Phascogale differs so conspicuously from
the Ph. cristicauda, Kufft, as identified by Baldwin Spencer
after comparison with the type, that I have no alternative
but to describe it as distinct, though I have little doubt that
its skull will show it to be closely allied to that animal. I
have much pleasure in naming it after its donor, to whom
the National Museum owes a number of interesting specimens
from the same locality.
XLVIII.—Notes on the Distribution of some Species of
Terrestrial Isopoda introduced into Australasia. By
CHARLES CHILTON, M.A., D.Sc., F.L.8., Professor of
Biology, Canterbury College, New Zealand.
In this short paper I give a few notes on the distribution, so
far as it is at present known, of some species of terrestrial
Isopoda that appear to have been introduced into Australasia,
most probably by the unconscious action of man. I have
already pointed out elsewhere* that the terrestrial Isopoda
are well worthy of study from the point of view of the
geographical distribution of animals, and it is therefore
desirable to place on record any facts illustrating the spread
of different species at the present time. ‘I’wo or three of the
species introduced into Australasia have been described as
new local species, and until these are reduced to the rank
of synonyms lists of the species inhabiting any part of
Australasia are apt to be misleading.
Only references necessary for workers in Australasia have
been given; full references and synonyms can be found in
the works by Dr. Budde-Lund and Professor G. O. Sars.
My collections from Australia are as yet rather meagre,
but Messrs. Helms and Sayce have sent me specimens from
various localities in New South Wales and Victoria, and
Messrs. W. and R. M. Laing from Norfolk Island. To
these gentlemen my thanks are due, and also to Monsieur V.
Bouge, who has taken considerable trouble to send me
specimens from New Caledonia.
* See Trans. Linn. Soc. ser. 2, Zool. viii. p. 100; and ‘New Zealand
Journal of Science,’ 11, (1884) p. 155.
some Species of Terrestrial [sopoda. 429
Porcellio scaber, Latr.
Porcellio scaber, Budde-Lund, Isopoda Terrestria, p. 129.
Porcellio seaber, G. O. Sars, Crustacea of Norway, is ps Lae,
Porcellio graniger, Miers, Cat. New Zealand Crust. p. 99.
Porcellio scaber, Chilton, Trans. Linn. Soe. ser. 2, Zool. viii. p. 139.
Tt will be seen from a reference to the works of Budde-
Lund or Sars quoted above that this species is very widely
distributed and has been described under many names. It is
found ae where in Northern, Western, and Central Europe,
extending far north into Gresnlend and Iceland; less fre-
quently in Southern Hurope; also in North America,
Mexico, &c.; in the islands of St. Paul, St. Croix, in Cape
of Good Hope, and in Kamtschatka. Monsieur Dollfus *
does not give it in his list of terrestrial Isopoda from the
mainland of the North of Africa, but it is known from the
Canary and the Azores Islands. It appears probable, there-
fore, that it belongs originally to the more northern portions
of the continents of Europe, ‘Asia, and perhaps America, and
that it has been unintentionally introduced into the various
islands and other localities in the Southern Hemisphere,
where it is now found.
In Australia it has been recorded by Miers from Melbourne
and Tasmania, and I have had specimens from Sydney in
my collection for many years past, and more recently
Mr, Sayce has sent me a few specimens from “ Forest
Country, Healsville, Victoria”; but it does not appear to be
at all abundant in Australia, for these are the only localities
from which it has been obtained, and it is not represented
from the other places in New South Wales and Victoria
from which I have specimens sent me by Mr. R. Helms and
Mr. Sayce, and it does not appear to occur in Norfolk Island.
In New Zealand, on the other hand, where it has been
long known under the name of Porcellio graniger, Miers, it
is the commonest terrestrial Isopod, and is very abundant
everywhere, more particularly in the neighbourhood of
inhabited places, but also extending sometimes a considerable
distance into the bush; and it is interesting to note that
specimens from New Zealand were already in the collections
of the British Museum in 1847 when White’s ‘ List of the
Crustacea in the British Museum?’ was published. It has not
* Proe, Fourth International Congress of Zoology, p. 250.
+ This statement appears to be in error. The specimens recorded by
White in 1847 were from ‘Van Diemen’s Land.” That from New
Zealand described by Miers was received by the Museum in 1854. {Note
by Dr. W. T. Calman, British Museum (Nat. Hist.).]
430 Prof. C. Chilton on the Distribution of
yet been recorded from the Chathams or any of the outlying
islands of New Zealand.
From the notes given above it appears that Porcellio
scaber thrives best in the colder temperate regions of both
North and South Hemispheres, and becomes less abundant
as the warmer climates are reached.
Dollfus, in his “ Isopodes terrestres du ‘Challenger’ ” *,
when recording this species from the islands of Juan Fernandez
and Tristan d’Acunha, has already drawn attention to the
fact that it is found in several widely remote localities in the
cold temperate regions of the south as well as the north,
although not found in the intermediate warmer regions, and
considers that its abundance excludes the idea of artificial
transport. T think, however, that the additional localities
now given are sufficient to show that it has reached these
places in the south by accidental introduction, as already
explained.
Porcellio levis, Latr.
Porcellio lems, Budde-Lund, Isopoda Terrestria, p. 138.
Porcellio levis, G. O. Sars, Crustacea of Norway, 11. p. 181.
Porcellio obtusify Ons, Haswell, Cat. Australian Crust. p. 280.
Budde-Lund says of this species that it is without doubt
the most widely spread of all the Oniscidae, so that it seems
to be almost cosmopolitan. He does not record it, however,
from Australia, although it had been recorded from Sydney
under the name of Porcellio obtusifrons (a species that
Budde-Lund appears to have overlooked) by Haswell in 1882.
I have had specimens from Sydney for many years com-
pletely agreeing with Haswell’s description, and have no
doubt that they should be referred to P. levis. Besides
these I have specimens sent me by Mr. Sayce labelled
“ From Garden,” presumably from Melbourne, and in 1891
Mr. George Gordon sent ‘me numerous specimens from
Geraldton, in Western Australia, most of them being slightly
larger than those from other localities. It is also to be found
in Norfolk Island and in New Caledonia, where, judging by
the number of specimens sent me by Monsieur V. Bouge, it
appears particularly abundant. Among these were several
albino or partially albino forms similar to those of which
Budde-Lund speaks as being found in Southern Europe.
It is interesting to note that the species does not appear to
have yet reached New Zealand ; there is no record of its ever
* EXxtr. Bulletin Société d’ Etudes scient. de Paris, xi® Année, 1890,
? ?
some Species of Terrestrial Isopoda. 431
having been found in this Colony, and as I have collected
terrestrial Isopoda for many years, both directly and with the
help of numerous friends, it is hardly likely that it would
have escaped observation if it occurred in any numbers.
Metoponorthus pruinosus (Brandt).
Metoponorthus pruinosus, Budde-Lund, Isopoda Terrestria, p. 169.
Metoponorthus pruinosus, G, O. Sars, Crustacea of Norway, ii. p. 184.
Porcellio zealandicus, Miers, Cat. New Zealand Crustacea, p. 100.
Porcellio zealandicus, Chilton, Trans, Linn, Soe. ser, 2, Zool. viii. p. 141.
This is another species which Budde-Lund says has been
carried by navigation to all parts of the world. It has,
however, not yet been recorded from Australia, and no
specimens have so far reached me from the mainland of that
continent. On the other hand, it appears to be common on
Norfolk Island, whence specimens have been sent to me both
by Mr. W. Laing and Mr. R. M. Laing, and, according to
Monsieur Adrien Dollfus, it also occurs in New Caledonia and
in the Marianna Islands.
A specimen of this species appears to have been collected
in New Zealand before 1847, and was given the name
Porcellio zealandicus in the list published in that year. In
1885 Budde-Lund gave P. zealandicus as a doubtful synonym
of Metoponorthus pruinosus; but as the specimen in the
British Museum was not sufficiently well preserved to render
identification certain, and as other species formerly recorded
had not been met with again in New Zealand, the question
remained doubtful till March 1905, when I received from
Mr. Hutchinson numerous specimens of Metoponorthus prut-
nosus from the shores of a tidal lagoon in Hawke’s Bay, in the
North Island of New Zealand. ‘There is therefore no reason
for doubting that Porcellio zealandicus, Miers, is the same as
Metoponorthus pruinosus and that the species had reached
New Zealand before 1847. It is rather interesting to note
that the species has not spread more widely, for as yet it has
not been met with in any other locality in New Zealand *.
Armadillidium vulgare (Latr.).
Armadillidium vulgare, Budde-Lund, Isopoda Terrestria, p. 66.
Armadilhidium vulgare, G. O. Sars, Crustacea of Norway, ii. p. 189.
Armadillidium vulgare, Chilton, Trans, Linn. Soc. ser. 2, Zool. viii.
p. 142.
* A short paper on the occurrence in New Zealand of this species has
been sent to the Transactions N.Z. Institute, vol. xxxviii.
432 On an undescribed Guereza.
This is another widely distributed species. Budde-Lund
says that it inhabits the whole of Europe and also the neigh-
bouring regions of Asia and Africa, and that it has been
carried to many other places probably by navigation.
Budde-Lund records it also from Melbourne, but so far it
has not been met with from any other part of Australia, and
it is not represented in any of my collections from that
country.
In New Zealand it appears to be abundant in the town of
Nelson; specimens were sent me from there many years ago
by the late Mr. J. C. Gully, and I have recently received
many specimens from Mr, F. G. Gibbs, who tells me that it
appears to be as common in Nelson as Porcellio scaber. I
have also a specimen taken near Mount Egmont and
forwarded to me by the late Mr. S. H. Drew, but up to the
present it has not been met with in any other localities, in
this respect forming a marked contrast to Porcellio scaber,
which has overrun the whole colony.
XLIX.—An undescribed Guereza.
By R. Lypexxer.
Masor Powett-Corron has sent from Qugo, Central Equa-
torial Africa, the skull and skin of an adult Guereza monkey
belonging to the black-and-white group, which appears to
indicate an undescribed form. The specimen closely re-
sembles Colobus palliatus of British Hast Africa (which may
be only a local form of the western C. angolensis), but is
distinguished by the absence of a white superciliary ridge
connecting the two face-tufts and of a white perineal patch,
as well as by the terminal half of the tail being grey, with a
white tip, instead of the whole terminal third being white
and the middle third grey. The new form may be best
regarded as a subspecies of C. palliatus under the name of
‘olobus palliatus Cottoni, and characterized as above. It
adds one more link connecting the wholly black and smooth-
tailed forms of Guereza with the white-tailed C. caudatus,
which has an enormous white mantle and face-tufts and a
white “ flag ” to the tail like that of a setter ; and is therefore
of more interest than the ordinary ‘‘ subspecies.”
—
On the American Otchlid Genus Cichlosoma, &e. +:
L.—A Revision of the Fishes of the American Cichlid Genus
Cichlosoma and of the Allied Genera. By C. Tare
Reean, B.A.
[Concluded from p. 340. |
PETENIA.
Petenia, Giinth. Cat. Fish. iv. p. 391 (1852).
Peteniu (part.), Eigenm. & Bray, Ann. Ac. N. York, vii. 1894, p. 614;
Pellegr. Mém. Soc, Zool. France, xvi. 1903, p. 248 (1904).
Body moderately elongate, compressed ; scales rather large,
ctenoid. Two lateral lines. Jaws with bands of small
conical teeth, the outer series enlarged; canines moderate.
Mouth large ; premaxillaries very protractile, their processes
as long as the head; maxillary considerably exposed ; lower
jaw strongly projecting. Upper surface of the head to the
level of the orbits, cheeks, and opercular bones scaly; pre-
operculum narrow, entire, without distinct lower limb. Gull-
rakers rather short andfew. A single dorsal, with XV-XVI
12-13 rays; anal with V 8-10 rays. Pectoral asymmetrical,
with 14 rays; ventrals below the base of the pectoral.
Caudal rounded.
A single species from Lake Peten, Guatemala, to be
regarded as a specialized form derived from a type similar to
Cichlosoma Dovit.
Petenia splendida.
Petenta splendida, Giinth. Cat. Fish. iv. p. 801 (1862), and Trans. Zool.
Soe. vi. 1869, p. 469, pl. Ixxix. fig. 2; Steind. Denkschr. Ak. Wien,
xxiii. 1864, p. 73, pl. iv. figs. 3&4; Kigenm. & Bray, Ann. Ac.
N. York, vii. 1894, p. 615; Jord. & Everm. Bull. U.S. Nat. Mus.
xlvii. 1898, p. 1513; Everm. & Goldsb. Bull. U.S. Fish. Comm. xxi.
1902, p. 155, fig.; Pellegr. Mém. Soe. Zool. France, xvi. 1903, p. 243
(1904).
Depth of body about 3 in the length, length of head 22-23.
Snout a little shorter than postorbital part of head. Dia-
meter of eye 5-6 in the length of head and ]—14 in the
interorbital width ; depth of przorbital 4—? the diameter of
eye. Lower jaw strongly projecting ; maxillary extending
to below middle of eye or beyond; premaxillary processes
extending nearly to origin of dorsal. Cheek with 7-10
series of scales; 10 or 11 gill-rakers on the lower part of
anterior arch. Scales of the lateral line a little larger than
the others; scales above the lateral line a little smaller
than those below it, about 45 in a longitudinal series above,
Ann. & Mag. N. fist. Ser. 7. Vol, xvi. 29
434 Mr. C. T. Regan on the Fishes of the
40 below the lateral line. Scales in a transverse series
wu: Dorsal XV (XVI) 12-13, the last spine about 2 the
length of head, the soft fin not scaly. Anal V (8) 9-10.
Pectoral nearly 2 the length of head. Caudal rounded.
Caudal peduncle longer than deep. Head, body, and vertical
fins with dark spots; a series of dark blotches along the
middle of the side.
Lake Peten, Guatemala.
1-4, (260-415 mm.) types of Lake Peten. O. Salvin, Esq.
the species.
Hericutuys.
Herichthys, Baird & Girard, Proce. Ac. Philad. 1854, p. 25.
Heros (part.), Gunth. Cat. Fish. iv. p. 290; Jord. & Everm. Bull. U.S.
Nat. Mus. xlvii. 1898, p. 1537; Pellegr. Mém. Soe. Zool. France, xvi.
1903, p. 226 (1904).
Neetroplus (part.), Jord. & Snyder, Bull. U.S. Fish. Comm. xix. 1899,
p. 146; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 245 (1904) ;
Meek, Zool. Pub. Columbian Mus. y. 1904, p. 221.
Cichlasoma (part.), Meek, t. e. p. 205.
Closely allied to Cichlosoma and perhaps most nearly to
C. heterodontus, differing only in that the anterior teeth of
the outer series in each jaw are compressed, pointed in the
young, truncate in the adult.
Three species from Mexico and Central America.
Key to the Species.
I. Anal with 5 (exceptionally 6) spines.
D. XV-XVII 10-12. A. V (VI 7) 8-9. Sec.
27-30 ms Byareshe(atateraiene « Mott orehels het ceakc eisiete 1. cyanoguttatus.
D/XVIAS: SA, V 10 eSeRB 159 Seeds. 85. 2. Bocourti.
II. Anal with 6 or 7 spines.
D. XVI-XVIII 11-12. A. VI-VII 9-10. Se.
BO ge. a5 Agee toe ne eae net 3. Geddesi.
1. Herichthys cyanoguttatus.
Herichthys cyanoguttatus, Baird & Girard, Proc. Ac. Philad. 1854, p. 25,
and Rep. U.S. Mex. Bound. Surv., Zool. p. 30, pl. iv. figs. 9-12 (1859).
Heros cyanoguttatus, Giinth. Cat. Fish. iv. p. 290 (1862); Jord. &
Gilb. Bull. U.S. Nat. Mus. xvi. 1882, p. 608; Jord. & Everm. Bull.
U.S. Nat. Mus. xlvii. 1898, p. 1537; Everm. & Goldsborough, Bull.
US. Fish. Comm. xxi. 1902, p. 157 ; Pellegr. Mém. Soe. Zool. France,
xvi. 1903, p. 231 (1904).
Neetroplus carpintis, Jord. & Snyder, Bull. U.S. Fish. Comm. xix. 1899,
p. 146, fig. 22; Jord. & Everm. t. c. p. 3175; Pellegr. t.c. p. 246;
Meek, Zool. Pub. Columbian Mus, y. 1904, p. 221.
Cichlasoma cyanogutiatum, Meek, t. ec. p. 215,
American Cichlid Genus Cichlosoma, dc. 435
Depth of body 2 in the length, length of head 23-3
Snout shorter than postorbital part of head. Diameter of
eye 3-41 in the length of head, interorbital width 3-34.
Depth of preorbital 4-1) the diameter of eye. Maxillary
slightly exposed, not extending to below the eye; pre-
maxillary processes extending to above anterior 4 of eye or
slightly beyond ; jaws equal anteriorly, or the lower slightly
projecting ; both jaws with the teeth of the outer series
regularly increasing in size anteriorly ; fold of the lower lip
usually more or less distinctly interrupted ; cheek with 4 or
5 series of scales; 7 or 8 aus rakers on the lower part of
anterior arch. Scales 27-30 a ae 2 between lateral line and
base of anterior rays of soft dorsal. Dorsal X V-X VII 10-12,
commencing above the extremity of operculum, the spines
subequal or slightly decreasing from the sixthto thethirteenth,
thence increasing to the last, which is 3 the vote of head ;
the soft fin scaly at the base. Anal V (VI7) 8-9. Pectoral
4 the length of head, extending to above origin of anal ;
ventral extending nearly to origin of anal. ‘Caudal sub-
truncate or rounded. Caudal peduncle 2 as long as deep.
Light blue spots on head, body, and vertical fins ; young
with 7 dark cross- bars, the third bearing a blackish blotch
below the lateral line, ‘and with a dark spot on the base of
caudal.
North-Eastern Mexico and Texas.
1-2. (65 and 118 mm.) Matamoros. Smithsonian Institution.
3-6. (39-62 mm.) co-types Tampico. Dr. D. 8. Jordan.
of N. carpintis.
Heros teporatus, Fowler (Proc. Ac. Philad. 1903, p. 321)
is based on a single specimen of 38 mm. from Tamaulipas.
It appears to differ from H. cyanoguttatus only in the more
slender body (depth 22 in the length).
2. Herichthys Bucourti.
Neetroplus Bocourtt, Vaill. & Pellegr. Bull. Mus. Paris, 1902, p. 85 ;
Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 246 (1904).
Depth of body 2 in the length, length of head 34. Dia-
meter of eye 32 in the length of head. Maxillary not
extending to below the eye; fold of the lower lip slightly
interrupted ; cheek with 6 series of scales; 9 gill-rakers
on the lower part of anterior arch. Scales 314. Dorsal
XVI 13, the spines only slightly increasing in length after
the fifth, the last more than 3 the length of head; soft fin
scaly at the base. Anal V 10. Pectoral a little shorter
20°
436 Mr. C. T. Regan on the Fishes of the
than the head. Caudal slightly rounded. Caudal peduncle
nearly as long as deep. Olivaceous, with obscure darker
cross-bars.
Lake Isabel, Guatemala.
The single known example measures 182 mm. in total
length.
This species is said to be closely allied to the preceding,
differing especially in the larger eye.
3. Herichthys Geddes, sp. n.
Depth of body 21-2} in the length, length of head 23—2%.
Snout as long, or nearly as long, as postorbital part of head.
Diameter of eye 23-34 in the length of head, interorbital
width 31-32. Maxillary not extending to below the eye;
preemaxillary processes extending to above anterior 4 of eye;
jaws equal anteriorly ; both jaws with the teeth of the outer
series regularly increasing in size anteriorly ; fold of the
lower lip not continuous; cheek with 5 or 6 series of scales ;
7 or 8 gill-rakers on the lower part of anterior arch. Scales
30 iy, 2 between lateral line and base of anterior rays of
soft dorsal. Dorsal XVI-X VIII 11-12, commencing above
the opercular cleft, the spines subequal or slightly decreasing
from the fifth or sixth to the fourteenth or fifteenth, the last
2 the length of head; the soft fin scaly at the base. Anal
VI-VII 9-10. Pectoral 3-7 the length of head, extending
to above origin of anal; ventral extending to anterior anal
spines. (? Caudal subtruncate.) Caudal peduncle 3-3 as
long as deep. Brownish, with 7 or 8 cross-bars bearing a
series of blackish blotches below the lateral line; vertical
fins spotted.
Southern Mexico.
1-6. (47-65 mm.) types of Southern Mexico. P. Geddes, Esq.
the species,
PARANEETROPLUS, gen. nov.
Differs from Cichlosoma only in the dentition, all the teeth
being broad, compressed, and pointed or rounded. ‘The
single species, from Mexico, is intermediate between Cichlo-
soma nebuliferum and Neetroplus nematopus.
Paraneetroplus Bulleri, sp. 0.
Depth of body 3 in the length, length of head nearly 4.
Snout a little shorter than postorbital part of head. Diameter
of eye 42 in the length of head, interorbital width 23. Depth
American Cichlid Genus Cichlosoma, te. 437
of preorbital 13 the diameter of eye. Maxillary scarcely
exposed, not nearly extending to below the eye; pre-
maxillary processes not extending to above the eye ; mouth
small, almost transverse, the lower jaw shorter than the
upper; upper jaw with about 20 teeth in the outer series,
the lateral teeth decreasing in size posteriorly ; lower jaw
with a transverse series of about 10 teeth, not continued as a
lateral series; fold of the lower lip not continuous; cheek
with 5 series of scales; 7 gill-rakers ou the lower part of
anterior arch. Scales 34 a3 3 between lateral line and base
of anterior rays of soft dorsal, those of the thoracic region
very small. Dorsal XVIII 12, commencing a little in
advance of the axil of pectoral, the spines subequal from the
fifth to the fifteenth, thence increasing to the last, which
is 3 the length of head; soft fin, when laid back, extending
to base of caudal; posterior part of spinous dorsal and
anterior part of soft fin scaly at the base. Anal VI 9.
Pectoral a little shorter than the head, not nearly extending
to above the anal; ventral extending nearly to origin of
anal. Caudal truncate. Caudal peduncle a little longer
than deep. Olivaceous, with a series of 5 dark blotches on
the side of the body, the last 2 extending upwards as vertical
bars on to the base of the soft dorsal ; a dark spot at the base
of caudal ; vertical fins slightly dusky.
Kio de Sarabia, Mexico.
1, (218 mm.) type of the Rio de Sarabia. Dr. A. C. Buller.
Species.
NEETROPLUS.
Neetroplus, Giinth. Trans, Zool. Soc. vi. 1869, p. 469; Eigenm. &
Bray, Ann. Ac. N. York, vii. 1894, p.618; Jord. & Everm. Bull. US.
Nat. Mus. xlvii. 1898, p. 1541.
Neetroplus (part.), Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 245
(1904).
Differs from Paraneetroplus in that the teeth are not
pointed or rounded, but truncate, incisor-like.
A single species from Nicaragua.
Neetroplus nematopus.
Neetroplus nematopus, Ginth. Trans. Zool. Soc. vi. 1869, p. 470, pl. Ixxiy.
fig. 4; Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1898, p. 1541 ;
Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 245 (1904).
Neetroplus nicaraguensis, Gill & Bransford, Proc. Ac. Philad. 1877,
p. 186,
Depth of body 23 in the length, length of head 34. Snout
as long as postorbital part of head. Diameter of eye 34 in
438 Mr. C. T. Regan on the Fishes of the
the length of head, interorbital width 24. Depth of pre-
orbital a little greater than diameter of eye. Maxillary
extending a little beyond the nostril; preemaxillary processes
not extending to above the eye; jaws nearly equal ante-
riorly ; upper jaw with about 14 teeth in the outer series, the
lateral teeth decreasing in size posteriorly ; lower jaw with a
transverse series of 9 teeth; fold of the lower lip not con-
tinuous; cheek with 5 series of scales; 6 gill-rakers on
the lower part of anterior arch. Scales 33 7, 2} between
lateral line and base of anterior rays of soft dorsal, those of
the thoracic region very small. Dorsal XIX 10, the spines
subequal from the fifth to the sixteenth, the last 2 the length
of head; soft fin, when laid back, extending to middle of
caudal ; dorsal scaly at the base posteriorly. Anal VIII 7.
Pectoral nearly as long as the head, not extending to above
the anal; ventral with the outer ray elongate. Caudal
truncate or even slightly emarginate. Caudal peduncle as
long as deep. Olivaceous, obscurely spotted or marbled
with darker.
Nicaragua.
1. (107 mm.) type of the Lake Managua. Capt. J. M. Dow.
species.
HLerRorivaria.
Herotilapia, Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 247 (1904).
Differs from Cichlosoma only in the dentition; both jaws
with several series of compressed, incisor-like, tricuspid teeth;
the outer series enlarged and with the median teeth simple,
truncate.
The single species is evidently closely related to C. nigro-
fasciatus and its allies.
Herotilapia multispinosa.
Heros multispinosus, Giinth. Trans. Zool. Soc. vi. 1869, p. 453, pl. xxiv.
fig. 2.
Cichlasoma multispinosum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1898, p. 1525.
Herotilapia multispinosa, Pellegr. Mém. Soe. Zool. France, xvi. 1903,
p. 247 (1904).
Depth of body 23 in the length, length of head nearly 3.
Snout a little shorter than eye, the diameter of which is 3}
in the length of head, interorbital width 24. Depth of pre-
orbital nearly 2 the diameter of eye. Maxillary not ex-
tending to below the eye ; preemaxillary processes extending
nearly to above anterior margin of eye; jaws equal
American Oichlid Genus Cichlosoma, &c. 439
anteriorly ; cheek with 3 series of scales; 8 gill-rakers on
the lower part of anterior arch. Scales 28 “=~ Dorsal
XVIII 9, commencing above the opercular cleft, the spines
subequal from the fifth, the last a little more than 3 the
length of head; soft fin scaly at the base, when laid back
extending nearly to middle of caudal. Anal X18. Pectoral
as long as the head, extending to above middle anal spines ;
ventral extending to sixth anal spine. Caudal rounded.
Caudal peduncle 2 as long as deep. Olivaceous, with darker
cross-bars; a dark spot on the middle of the side; a
longitudinal band from eye to lateral spot.
Lake Managua.
1. (87 mm.) type of the species, | Lake Managua. Capt. J. M. Dow.
Uaru.
Uaru, Heck. Ann. Mus. Wien, ii. 1840, p. 830; Giinth. Cat. Fish. iv.
p- 802 (1862); Eigenm. & Bray, Ann. Ac. N. York, vii. 1894,
p- 612; Pellegr. Mém. Soe. Zool. France, xvi. 1908, p. 248 (1904).
Acara (part.), Steind. Sitzb, Ak. Wien, Ixxi. 1876, p. 63.
Closely allied to Cichlosoma, and especially to C. psittacum,
differing in the dentition, the rather slender teeth being
compressed, pointed in the young and with rounded apices
in the adult. Scales rather small, those of the lateral line
larger than the rest.
A single species from the Amazon.
Uaru amphiacanthoides.
Uaru amphiacanthoides, Heck. Ann. Mus. Wien, ii. 1840, p. 381;
Giinth. Cat. Fish. iv. p. 302 (1862); Higenm. & Bray, Ann. Ac,
N. York, vii. 1894, p. 612; Pellegr. Mém. Soc. Zool. France, xvi.
1903, p. 248 (1904).
Uaru obscurum, Ginth. 0. ec.
Acara (Heros) amphiacanthoides, Steind. Sitzb. Ak. Wien, Ixxi. 1875,
. 94.
ones (Heros) imperialis, Steind. Sitzb. Ak. Wien, Ixxx. 1879, p. 161.
Uaru imperialis, Pellegr. t. c. p. 249.
Depth of body 14-1? in the length, length of head 3-3}.
Diameter of eye 34-3* in the length of head, interorbital
width 2-24. Depth of przeorbital 11-1? the diameter of eye.
Jaws equal anteriorly ; maxillary not extending to below
the eye ; fold of the lower lip not continuous ; cheek with
9-12 series of scales; 7 or 8 gill-rakers on the lower part of
anterior arch. Scales in a longitudinal series, 55-60 above
the lateral line, 46-52 below it; scales in a transverse series
1-13" Lateral line 18-21 410-13. Dorsal XV-XVI (13-14)
24-28°
440 Mr. C. T. Regan on the Fishes of the
15-16, the spines subequal from the fifth or sixth, the last
3_2 the length of head, the soft fin scaly at the base. Anal
VIII-IX (X 12-18) 14 (15-16). Pectoral a little longer
than the head. Caudal rounded. Caudal peduncle 4-4 as
long as deep. Brownish, with a broad blackish lateral band,
a blackish spot behind the eye and another on the upper
part of the base of caudal.
R. Amazon.
1. (145 mm.) type of U. obscurum. R. Cupai.
2. (215 mm.) L, Saraca, Prof. A. Agassiz.
3. (198 mm.) Teffé, Paris Museum.
SYMPHYSODON.
Symphysodon, Heck. Ann. Mus. Wien, ii. 1840, p. 882; Giinth. Cat.
Fish, iv. p. 816 (1862) ; Steind. Sitzb. Ak. Wien, lxxi. 1875, p. 106;
Eigenm. & Bray, Ann. Ac. N. York, vii. 1894, p. 623; Pellegr.
Mém, Soc. Zool. France, xvi. 1903, p. 249 (1904).
Closely allied to Cichlosoma, especially to C. severum,
differing in having the teeth confined to the anterior part of
each jaw and in the structure of the fins, the dorsal having
VITI-IX 28-31 rays and the anal VII-IX 26-31. The
body is nearly circular in form, compressed ; the scales are
rather small, those of the lateral line being larger than the
rest.
A single species from the Amazon.
Symphysodon discus.
Symphysodon discus, Heck. Ann. Mus. Wien, ii. 1840, p. 333; Giinth.
Cat. Fish. iv. p. 316 (1862); Steind. Sitzb. Ak. Wien, lxxi. 1875,
p- 106; Pellegr. Mém., Soc. Zool. France, xvi. 19038, p. 250 (1904).
Depth of body 1-1} in the length, length of head 32-33.
Snout as long as postorbital part of head. Diameter of eye
3-34 in the length of head, interorbital width 2-22. Depth
of preorbital 1-11 the diameter of eye. Mouth small,
oblique; maxillary extending to below the nostril; fold of
the lower lip continuous, slightly produced. Cheek with
7 or 8 series of scales; 4 or 5 very short gill-rakers on the
lower part of anterior arch. Scales 65-70 nies 50-55 in a
longitudinal series below the lateral line, 6 between upper
and lower lateral lines. Dorsal VITI-IX 80-31 (VIJ-IX
26-31). Anal VIII 27-380 (VII-IX 26-31). Dorsal and
anal spines graduated, the fins covered with scales in their
basal halves. Pectoral a little longer than the head. Caudal
rounded. Caudal peduncle very short. Body with dark
American Cichlid Genus Cichlosoma, &c. 441
undulating longitudinal stripes and with 9 dark cross-bars,
of which the first, through the eye, the fifth, in the middle of
the length of the body, and the nimth, on the base of the
caudal, are more strongly marked than the others; soft
vertical fins with alternate light and dark spots.
R. Amazon,
1. (110 mm.) R. Cupai.
2. (155 mm.) R. Negro. Mr. J. C. Antony.
3. (170 mm.) Teffé. Dr. J. Bach.
4. (130 mm.) Manaos. Paris Mus.
Pellegrin gives the name equifusciata to a variety without
longitudinal stripes and with all the cross-bars equaliy
distinct.
PTEROPHYLLUM.
Pterophyllum, Heck. Ann. Mus. Wien, ii. 1840, p. 834; Gunth. Cat.
Fish. iv. p. 316 (1862) ; Steind. Sitzb. Ak. Wien, Ixxi. 1875, p. 156;
Eigenm. & Bray, Ann. Ac. N. York, vii. 1894, p. 624; Pellegr.
Mém. Soc. Zool. France, xvi. 1908, p. 251 (1904).
Plataxoides, Casteln. Anim. Am. Sud, Poiss. p. 21, pl. xi. fig. 3 (1855).
Very closely allied to Symphysodon, which it resembles in
the deep nearly circular body and the small scales, but
differing in the dentition, the teeth extending on to the sides
of the jaws. Dorsal with XI-XITI 24-30 rays, anal with
V-VII 24-82.
R. Amazon; R. Orinoco.
Two species.
1. Pterophyllum scalure.
Plataz scalaris, Cuy. & Val. Hist. Nat. Poiss. vii. p. 237 (18381).
Pterophyllum scalare, Heck. Ann. Mus. Wien, 11. 1840, p. 885: Gunth.
Cat. Fish. iv. p. 316 (1862); Steind. Sitzb. Ak. Wien, Ixxi. 1875,
p. 186; Pellegr. Mém. Sve. Zool. France, xvi. 1903, p. 251 (1904).
Plataxoides Dumerili, Casteln. Anim. Am. Sud, Poiss. p. 21, pl. xi.
fig. 3 (1855).
Depth of body a little less than the length, which is 23-3
the length of head. Snout not longer than eye, the diameter
of which is 23-3 in the length of head and equal to the
interorbital width. Depth of preorbital } the diameter of
eye or less. Mouth small, oblique; maxillary extending to
below the nostril; fold of the lower lip continuous ; cheek
with 4 or 5 series of scales; 12-14 gill-rakers on the
lower part of anterior arch. Scales 40-47 =) 83-38 in a
longitudinal series below the lateral line. Lateral line
17-19+9-11. Dorsal XI-XIII 23-27, the spines graduated,
442 Mr. C. T. Regan on the Fishes of the
the last about as long as the head; anterior soft rays con-
siderably produced ; middle part of the fin scaly at the base.
Anal VI 26-29 (V—-VII 24-29). Pectoral as long as the
head ; ventral produced into a long filament. Caudal trun-
cate, the outer rays sometimes produced. Caudal peduncle
3-1 as long as deep. Silvery with 4 well-marked blackish
cross-bars, the first through the eye, the second from spinous
dorsal to origin of anal, the third from soft dorsal to anal,
the fourth on the base of caudal; between these narrower
and less distinct intermediate bars; soft vertical fins some-
times with alternate light and dark stripes.
R. Amazon.
1. (102 mm.) R. Cupai.
2. (81 mm.) Tabatinga. Mus. Comp. Zool.
3. (99 min.) Manaos. Mr. J.C. Antony.
4. (61 mm.) Tonantins. Paris Mus.
5-7. (31-60 mm.) Marajo Id. Dr. E. A. Goldi.
2. Pterophyllum altum.
Pterophyllum altum, Pellegr. Bull. Mus. Paris, 1908, p. 125; Mém.
Soe. Zool. France, xvi. 1903, p. 252, pl. iv. fig. 4 (1904).
Depth of body equal to its length. Cheek with 6-7
series of scales; 11 gill-rakers on the lower part of anterior
arch, Scales about 55 2°, 41-47 in a longitudinal series
below the lateral line. Dorsal XII-XIII 27-30. Anal
(V) VI 28-82. Caudal peduncle % as long as deep. In
other characters similar to the preceding species.
R. Orinoco.
-2, (83 and 85 mm.) co-types of the R. Orinoco. Paris Museum.
species.
APPENDIX.
The British Museum has just received a series of the
Mexican Cichlids collected and described by Dr. 8. E. Meek.
After examination of these it seems useful to make a few
additions and corrections.
Cichlosoma fenestratum (supra, p. 227).
To the synonymy add :—
Cichlasoma melanurum (part.), Meek, Zool. Pub, Columbian Mus. v.
1904, p. 219,
American Cichlid Genus Cichlosoma, cc. 443
After the description of C. melanurum (p. 231) should be
inserted :—
Cichlosoma labridens.
Heros (Cichlasoma) labridens, Pellegr. Bull. Mus. Paris, 1903, p. 122
Cichlusoma labridens, Pellegr. Mém. Soc. Zool. France, xvi. 1903 By
p- 207, pl. vi. fig. 1 (1904).
Cichlasoma Bartoni (non Bean), Meek, Zool. Pub. Columbian Mus, vy.
1904, p. 211.
Cichlosoma Bartoni (part.), Regan, supra, p. 327.
Depth of body 21-2? in the length, length of head 24-31.
Snout a little shorter than postor bital part of head. Diameter
of eye 4-5} in the length of head, interorbital width 3.
Depth of prieorbital 1- 13 the diameter of eye. Maxillary
not extending to below the eye ; jaws equal anteriorly ; fold
of the lower lip subcontinuous or not continuous ; cheek
with 4-6 series of scales ; 6-8 gill-rakers on the lower part
of the anterior arch. Seales 30-32 ap 2) or 3 between
lateral line and base of anterior part of soft dorsal. Dorsal
XV-XVI 10-12, commencing above or in advance of the
axil of pectoral, the spines subequal from the fifth or sixth
to the thirteenth or fourteenth, thence increasing to the
last, which is ? the length of head. Anal V (VI) 8-9.
Pectoral nearly } the length of head, extending about to above
the vent; ventral extending nearly to origin of anal. Caudal
rounded or subtruncate. Caudal peduncle from ? to as long
as deep. Olivaceous, sometimes with 4 or 5 dark cross-bars
on the posterior part of the body ; sometimes the lower part
of the head and abdomen blackish and a large dark blotch
on the caudal peduncle; upper surface and sides of head
often covered with small dark spots ; fins greyish or dusky.
Rio Panuco and its tributaries, Mexico.
The above description is based on six specimens, 80-175 mm.
in length, from the Rio Verde.
This species is very closely allied to C. melanurum.
Cichlosoma Gadovii (p. 232).
The synonymy of this species should read :—
Cichlasoma melanurum (part.), Meek, Zool. Pub. Columbian Mus. vy.
1904, p. 219.
Cichlosoma aureum (p, 320).
Specimens have been received under the names of
Thorichthys Helleri and T. Elliotti. The former have the
444. On the American Cichlid Genus Cichlosoma, ce.
spots on the cheeks blue, often surrounded by a dark ring ;
in the latter the corresponding spots are blackish, many of
them with a blue centre. In all other respects these two
supposed species are absolutely identical.
Cichlosoma istlanum (p. 326).
Several young specimens differ in coloration from examples
of C. Steindachneri of the same size. ‘They have 7 or 8 dark
cross-bars, a dark spot below the origin of the lateral line,
another on the middle of the side, and a third at the base of
the caudal.
After the description of C. istlanum should be inserted :—
Cichlosoma Steindachneri.
Cichlasoma Steindachneri, Jord. & Snyder, Bull. U.S. Fish. Comm. xix.
1899, p. 143, fig. 20; Jord. & Everm. Bull. U.S. Nat. Mus, xlvii.
1900, p. 38173; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 206
(1904) ; Meek, Zool. Pub. Columbian Mus. v. 1904, p. 211, fig. 67.
Cichlosoma istlanum (part.), Regan, supra, p. 326.
Depth of body 24 in the length, length of head 24. Snout
as long as postorbital part of head. Diameter of eye 4 in
the length of head and equal to the imterorbital width.
Depth of przorbital equal to diameter of eye. Maxillary
slightly exposed, extending to between nostril and eye;
premaxillary processes extending to above anterior margin
of eye or slightly beyond; jaws equal anteriorly ; fold of
the lower lip subcontinuous ; upper jaw with the two aute-
rior teeth of the outer series the largest, forming canines ;
lower jaw with 3 pairs of canines, the anterior pair the
smallest ; cheek with 6 series of scales ; 8 serene on the
lower part of the anterior arch. Scales 31 Be 24 between
lateral line and base of anterior rays of soft dorsal. Dorsal
XVI 10, the spines subequal from the sixth to the fourteenth,
thence increasing to the last, which is a little more than }
the length of head ; soft fin, when laid back, extending to
anterior 4 of caudal. Anal V 8. Pectoral 3? the length of
head, extending to above the vent ; ventral eneatine ‘to the
vent. Caudal subtruncate. Caudal peduncle nearly as
long as deep. Olivaceous; sides with about 10 narrow
vertical dark bars ; a series of dark blotches forming an inter-
rupted longitudinal band from operculum to base of caudal ;
sides of head and Jower part of body with small] dark spots ;
On the Red Colour in the Hind Wing of Catocala. 445
spinous dorsal blackish ; soft dorsal, anal, and caudal dusky,
obscurely spotted at the base.
The above description is based on a single specimen,
104 mm. in total length, from the Rio Verde. Smaller
examples (40—55 mm.) ‘have a more slender body (depth 22-3
in the length).
Cichlosoma Bartoni (p. 327).
The synonymy of this species should read :—
Acara Bartoni, Bean, Proc. U.S. Nat. Mus. 1892, p. 286, pl. xliv. fig. 3.
Cichlasoma Barton, Jord. & Everm, Bull. U.S. Nat. Mus. xlvii. 1898,
P. 1515, fig. 587; Pellegr. Mém. Soc. Zool. France, xvi. 1903, p. 206
(1904) ; Meek, Zool. Pub. Columbian Mus. v. 1904, fig. 68.
Cichlosoma Bartoni (part.), Regan, supra, p. 327,
Three young examples of this species, 53-72 mm. in total
length, from the Rio Verde, received as C. Steindachneri,
differ from that species, which they resemble in coloration,
in the larger head (23 in the length), more produced snout,
and larger and more oblique mouth. Scales 32 2*,, 24 be-
tween lateral line and base of anterior part of soft dorsal.
Dorsal XVI 10-11, the last spine + the length of head.
Anal V 8. The coloration is similar to that of the adult
example figured by Bean, but the body is not so deep.
Cichlosoma octofasciatum (p. 331).
A series of specimens received as C. Hedricki includes a
small example in every way similar to the type of the species.
Herichthys cyanoguttatus (p. 434).
Entirely similar examples have been received under the
names Cichlasoma cyanoguttatum and Neetroplus carpintis.
My opinion as to the identity of these species is thus
confirmed.
LI.—A New Haplanation of the Red Colour in the Hind
Wing of Catocala, Schr. By Cu. SCHAPOSCHNIKOW *.
THE question as to how the development of the red colour in
the hind wing of Catoca/a is to be explained has not yet, in
the literature with which I am acquainted, been dealt with
* Translated by E. EK. Austen from the ‘ Biologisches Centralblatt,’
xxiy. Bd. (1904) pp. 514-520,
446 Mr. C. Schaposchnikow on the Red Colour
and answered in the way that I attempt to set forth in its
main outlines in the following pages. My more especial
aim in publishing this little paper, however, is that other
observers may be thereby induced to give utterance to further
remarks and views as to the question under discussion.
All attempts at explaining the significance of the red colora-
tion in the hind wing of Catocala, as recorded in literature,
amount to two modes of regarding it, since the red is put
down either as a “ warning” or as what may be termed an
“attractive” colour. ‘The latter designation implies that, as
interpreted by the authors who hold this view, the signifi-
cance of the red colour consists in attracting the bird which
is pursuing the moth. According to this theory the bird
seizes only the wing, and not the body, of the fugitive insect ;
the portion seized tears away, and thus the victim escapes.
Explanations such as this have been given by Wallace,
Darwin, Poulton, von Bock, Bedar, and others.
The first-mentioned attempt at explaining the red colora-
tion in Catocala as a “ warning” colour is, however, opposed
by important facts, the first of which is that Catocala, on the
approach of an enemy, suddenly departs from its resting-
place when that enemy is still at a considerable distance.
Secondly, it is only during its most rapid flight that the
insect displays the bright colour. These facts can scarcely
be used in support of such an interpretation of the colour as
that it serves to frighten away the enemy, or else to indicate
to it at once that the insect is unfit to eat. Hxperiments,
moreover, lead to the conviction that the pursuing bird seizes
the Catocala in its flight by the brightly coloured wing.
The second attempt at explanation, according to which
the red in Catocala is to be regarded as an “ attractive ”
colour, is chiefly based upon observations upon species of
Agrotis with yellow hind wings, whence it has been trans-
ferred to the species of Catocala with yellow and also to
those with red hind wings. With reference to the observa-
tions upon Agrotis it must not, however, be overlooked that
they were only made in the artificial conditions of an aviary,
Moreover, the external resemblance between Agrotis and
Catocala in no way justifies us in attributing to their similar
coloration the same biological significance in the struggle for
existence, and still less are we entitled to do so seeing that
there is to start with a very considerable difference in the
mode of life of the two genera. Besides this, it is still
doubtful whether the principle in question can even be
applied to all species of Agrotis with yellow hind wings;
there is a series of transitional forms, ranging from the large
in the Hind Wing of Catocala, Schr. 447
Agrotis pronuba, Linn., to the small Agrotés anachoreta,
H.-Sch., and luperinoides, Gn. The hind wings of these
small species of Agrot?s are not less brightly and conspicu-
ously coloured ; and even granting that this feature in the
case of the first-mentioned species (the Large Yellow Under-
wing) is able to divert the attention of a bird so as to cause
it to attempt to seize the insect by its hind wings, this would
still be far too difficult in the case of the two latter species
when on the wing, owing to their very smallness. It would
undoubtedly be easier for the pursuing bird to strike at the
body. But even if we admit the justice of this explanation in
the case of Agrotis pronuba, Linn., it is still impossible to fix
a limit up to which the yellow coloration of the hind wing
may be injurious to the possessor or to decide when its
attractive property may begin to be useful to the insect, for
species of Agrotts occur in all sizes in an ascending series
from the smallest forms. It must also not be overlooked
that, in opposition to the large species which, apart from
accidental contingencies, flies exclusively by night, we find
the above-mentioned small species of Agrotis on the wing by
day, and therefore at the only time when they are in a
position to turn the yellow coloration to account.
It we are unwilling to admit the force of all these state-
ments, there still remains the question as to how we are to
reconcile with the principle of attractive wing-coloration the
bright colour of the dody of various species of Catocala ( pacta,
Linn., neogama, Sm. Abb., magdalena, Streck., Frederic?,
Grote, neonympha, Hbnr., and amica, Hbnr.), of Agrotis
Jimbria, Linn., &e.; for since the wings, which during flight
are in a state of rapid movement, appear less brilliant than
the similarly coloured body, which is more at rest, we are
led, on the basis of the above-mentioned principle, to the
conclusion that this brighter body is calculated to entice
birds to pursuit. Furthermore, how can the black marking
of the hind wing be consistent with the recognition of this
principle? Since, by virtue of well-known physiological
laws, the parts of the wing that lie nearer to the base and
move with less great rapidity appear brighter to us than the
marginal portions of the wing, it follows that the former,
which are adjacent to the body, must, being the brighter,
concentrate the attention of the pursuer upon themselves.
This concentration of the attention is further accentuated and
the capture of the fugitive moth thereby facilitated if the
margins of the wings are darkened, so that the lightness of
the central area is heightened by contrast ; and it is actually
a fact that, in the case of the species of Agrots with yellow
448 Mr. C. Schaposchnikow on the Red Colour
hind wings, we almost always find a more or less broad
black margin. The pursuing bird will accordingly always
endeavour to seize the portion of the wing that lies nearer to
the body, 7. e. the basal half; but if it does so it is doubtful
whether the victim can tear itself free.
Let us now refer to a hypothesis—purely an abstract one,
indeed—which is indispensable if we would explain the bright
coloration as attractive. We actually find in nature a series
of examples showing that in self-defence an animal sacrifices
the portion of its body by which its enemy seizes it; thus,
for instance, the lizard abandons its tail and certain spiders
sacrifice their long legs. In all such cases the part of the
body that is shed is regenerated. In the case of moths an
analogous reproduction naturally cannot take place, and with
them therefore a safeguard of this kind must continue to be
accidental and only possible on one single occasion.
If, however, in spite of all these considerations, we would
still apply the above-mentioned principle to Catocala also, we
shall have to meet the question as to how it is affected by
natural selection.
When motionless, and so long as it remains upon a parti-
cular spot, the moth is unable to make use of the “ attractive ””
coloration ; for this purpose it must be in motion and on the
wing. If, however, during its flight it abandons a portion of
its wings to the pursuing bird, it will scarcely satisfy the
latter by so doing, and can consequently not free itself from
further pursuit. The sacrifice of a part of its wings at most
enables the moth to gain a little time, while, on the contrary,
the lizard, for instance, which sacrifices part of its tail, utilizes
the few moments so gained to disappear into a hole or cleft
in arock. The moth with “attractive” coloration would be
compelled to fly on and to make use of a suitable moment in
order to conceal itself. It follows that in its case there would
have to be developed, firstly, a quick and adroit flight, and,
secondly, the capacity of concealing itself unnoticed. The
struggle for existence was bound first of all to evolve somuch
the more strength and adroitness of flight, since with the
loss of or injury to the hind wing exhibiting the attractive
coloration its attractive 7é/e as such disappears. It follows
that only those individuals would survive that by dint of
concealing themselves unnoticed were able to turn their flight
to the best account and eseape the sacrifice of a portion of
their wings, de. those that made least use of the “ attractive ”
coloration in order to “ attract.” The ultimate object of the
flight would be to settle unobserved by the enemy and to
obtain concealment by means of the mimetic coloration of the
tn the Hind Wing of Catocala, Schr. 449
upper side of the fore wings ; this object is, moreover, alluded
to by von Bock in the passage in which, referring to Catfo-
cala, he states that the moths on being frightened up by him
suddenly disappeared before his eyes, without his having
been able to observe where they had settled. Herein there
lies an inconsistency, however: if the destined function of
the bright colour in the case of Catocala were to turn the
attention of the pursuer to itself, and if the bird then made
use of the chance of seizing its victim by the wing, it would
be impossible for so large a moth as Catocala to conceal itself
unnoticed at so short a distance from its enemy. But what
do we see in reality? ‘The structure of the wings of the red
Catocala, as also the relative size and the shape of its body,
render the insect capable of strong and rapid flight. ‘lhe
coloration of the species in question differs from that of the
other species owing to the hind wings being red on the upper
side and provided with two black transverse bands, while the
grey dark-striped fore wings appear perfectly adapted to the
tree-bark on which the moth usually settles ; the under side
is characterized by a light ground-colour, bordered by two
black bands. By day we generally find our Catocala at rest
on a tree-trunk ; it is excessively shy and flies up if approached
ever so gently. Its flight is not regular, but, on the contrary,
interrupted; it darts from one side to the other, settles
suddenly on a tree, flits up again, and frequently makes one
or two more short flights of this kind before finally settling
down. At times it flies no farther, but merely makes, as it
were, a jump to one side, and settles again upon the same
tree, but on the opposite side and higher up. These irregular
interrupted movements render the insect uncommonly difficult
to capture. On settling it makes a dart with lightning-like
rapidity in the direction of its chosen resting-place, and then
drops down there, having thus again made it extremely
difficult to fix the spot with the eye. The fact that on the
approach of an enemy our moth does not remain quietly
sitting still as other mimetic species are wont to do, but
endeavours to save itself by flying away, certainly points to
the conclusion that in this case flight is of greater importance
for the self-preservation of the insect than the mimetic colora-
tion of the fore wings—that is to say, flight attains the
object of leading the pursuer astray, and then, with the
assistance afforded by the coloration, rendering it possible for
the moth to settle unperceived.
The characteristic flight of our Catocala compels the enemy
to make a special effort with its visual apparatus, in order to
keep its prey within view; in the pursuit the bird fixes its
Ann. & Mig. N. Hist. Ser. 7. Vol. xvi. 30
450 Mr. C. Schaposchnikow on the Red Colour
eyes upon the bright object, as the flying Catocala appears to
it to be. The brighter the effect of its colour, the more diffi-
cult will it be, owing to the contrast of hue, to recognize the
grey patch into which, especially upon grey bark, the moth
converts itself at the moment of settling. Were its colours
wholly grey, the pursuer, fixing its gaze upon this grey
object, would pay most attention to its outlines and not to its
colour; and consequently the object would always remain
visible to the enemy, to whom the spot upon which it had
settled would likewise be apparent. Assuming that its
contrast in colour is absolutely necessary to the moth, it is
evident (1) that moths that have recourse to the above-
described mode of flight, which I would term “ distracting ”
flight, must possess the brightest coloration, and (2) that
those that exhibit brightly coloured hind wings must. also
have a bright-coloured underside to all the wings, in order to
be able to give greater effect to the influence of this coloration
in flight. Since, moreover, the larger the moth the less is
the advantage that it is able to gain from its mimetic colora-
tion, while, on the other hand, it is all the more capable of
saving itself by means of its powers of flight, it follows from
these two hypotheses that the largest species of Catocala
must also show the brightest colours. ‘The red Catocala
chiefly inhabits woods composed of the various deciduous
trees. In perspective a wood appears generally as a green
background (foliage and grass) with some light and dark
streaks (tree-trunks and their shadows) ; consequently as a
contrast of colours we have the red, which serves a comple-
ment of the green. And, again, it is especially in the most
robust representatives of their genus that this red coloration
must display itself. Asa colowr-contrast to the light and dark
stripes in the wood there appear in addition the black- and
white-banded undersides of the fore and hind wings of our
moth, which cause the latter to be easily visible in the light
as well as in the dark interspaces among the trees.
With this appearance of species of Catocala with red hind
wings is also connected their geographical distribution.
Their predominance in the northern half of Europe is espe-
cially characteristic, for among all the species of Catocala
in this region we find only two which show no red. The
conditions presented to them in the area in question are the
following :—the forests are not so dense as in warmer lands,
so that usually sufficient space remains to allow the moth to
make use of its powers of flight ; moreover, a perspective is
produced which, in accordance with the hypothesis expressed
above, enables the moth to bring its corresponding coloration
in the Hind Wing of Catocala, Schr. 451
and marking into play. In these woods the enemies of the
moth, which rests during the day upon the trunks of trees,
are the birds *, both the scansorial species (Picus, Sitta, and
their allies), as also those that pick insects off twigs, or in
other cases search for them upon the ground (Turdus and
others of the same group). At night, when the moth is on
the wing, it is waylaid by bats. From thrush-like birds the
red Catocala is sufficiently protected by the height of its
resting-place ; it scarcely ever settles so near the ground as
to attract the attention of birds of this kind, and, again,
never at such a height as to fall a victim to birds perching
upon twigs. A safe elevation would be at from 1} to 23
metres from the ground, and it is at about this height that,
as a rule, our red Catocala is actually to be seen. More
dangerous to it are the birds that climb and closely search
the tree-trunks (Pécus, Sitta); it is not easy for so large a
moth to escape their notice, but all these birds invariably
make a noise in climbing, and in our Catocala the sense of
hearing appears to be particularly well developed, so that on
the approach of woodpeckers and other climbing birds it is
always possible for it to flit off from the trunk at the right
time. By day it avails itself both of the strength and adroit-
ness of its flight, as also of the corresponding wing-coloration,
in order by these means to be more or less effectually pro-
tected from a bird that may be seeking for it. Though I am
acquainted with a considerable number of cases which point
to bats as being solely responsible for the destruction of the
moth, I am unable to prove a single instance in which a
Catocala has been killed by a bird during the day. Nor is it
difficult to understand why Catocala is chiefly caught by bats,
for bats hunt in the dark and by ear, and against them
Catocala can only make a partial use of its distracting flight,
while its colour is of no use to it at all.
What has been stated above as to the enemies of Catocala
explains at once the precise kind of region in which existence
is possible for a group of moths of fairly considerable size
and powerful musculature, the members of which sit upon
tree-trunks when at rest and possess mimetic fore wings and
brightly coloured hind wings, the coloration of which plays a
distracting part in connexion with a distracting flight. For
the evolution of a group of moths with the above-mentioned
* Lizards and snakes would also come within the category of insect-
enemies which climb without noise ; but the tree-lizards that inhabit the
region in question are too small and not sufficiently numerous, while the
tree-climbing snakes are incapable of ascending trunks as thick as those
usually chosen by the red Catocala for its resting-place. rf
30
452 On the Mus orthodon of Hensel.
characteristics—that is to say, the development of the red
species of Catocala—would be simply impossible, firstly in a
region in which a series of foes exists capable of noiselessly
approaching the moth as it rests upon the tree-trunk, and
secondly where, on being pursued, owing to limited space
and in consequence of the absence of a definite perspective,
the insect cannot terminate its distracting flight and turn its
distracting colour to account. To these circumstances is to
be ascribed the fact that the red Catocala is widely distributed
in the Nearctic and Palearctic Regions, but does not occur
in the Tropics. And it appears to be further deserving of
note that in the Neotropical and Australian Regions, which
have an older character, nothing is known of any represen-
tatives of the genus Catocala at all.
A glance at the recent species of Catocala will serve to
confirm the interpretation here given; almost all the red
species of Catocala are moths of a size which the yellow
species of Catocala in most cases fall far short of. The
marking of the red species of Catocala, consisting of two
black transverse bands, is, almost without exception, absolutely
regular in its occurrence, while the yellow species show
manifold modifications. Another interesting fact is that the
orange-coloured species of Catocala are also of large size,
while their marking, which consists of two black transverse
bands, remains more or less constant; in the case of the
yellow species, on the other hand, it is generally only among
the larger representatives that we find the marking in question
regular in its development. At the same time, however, it is
self-evident that local and historical conditions must have
produced a series of modifications both in size as also in
coloration in the case of the red species of Catocala as well ;
in them nevertheless these moditications are, as has already
been mentioned, much less considerable. As a characteristic
representative of the coloration that we have been investi-
gating, Catocala elocata, Esp., may be taken. This species is
one of the largest of the genus Catocala, and at the same
time one of the commonest of Huropean moths.
LII.—On the Mus orthodon of Hensel.
To the Editors of the ‘Annals and Magazine of
Natural History.’
GrnTLEMEN,—In 1900, while preparing a paper for the
P. Z. S. (pp. 887-428) on Mus sylvaticus and its allies, I
had occasion to examine and discuss the Mus orthodon of
Bibliographical Notices. 453
Hensel. Misled by an authority, who shall be nameless,
and by the fact that an important paper by my friend
Dr. Forsyth Major was published in a foreign tongue with
which I was at that time unacquainted, I erroneously con-
cluded that Dr. Forsyth Major considered Mus orthodon to
be a very near ally of Mus sylvaticus. His real opinion,
published in the Proc. Verb. Soc. Tose. Sci. Nat. 1884, was
very different, as he now informs me, and I believe I am
- right in saying that it coincides with that at which I myself
independently arrived in my paper of 1900.
Although, then, no difference of opinion exists between us
on any question of fact, I fear that those who read my paper
might easily conclude otherwise, and they would certainly
gain a false impression—due to my having followed too
blindly the above-mentioned “ authority ??—of the value of
Dr. Forsyth Major’s work. ‘To remove any such misunder-
standings is the object of this letter, an explanation being
clearly due to Dr. Forsyth Major, against whom I can
formulate no complaint more serious than that so many of
his most luminous papers on Palearctic mammalogy should
have been published in foreign journals often not readily
accessible.
I am, yours truly,
G E. H. Barrert-Hamitton.
BIBLIOGRAPHICAL NOTICES.
Economic Resources of the Northern Black Hills, By J. D. Irvine.
With Contributions by S. F. Emmons and T. A. Jacear, Jr.
Pages 222. 4to. Government Printing Office, Washington,
1904.
Tu1s volume, published as “ No. 26” of the “ Professional Papers :
Series A. Economic Geology, 34; B. Descriptive Geology, 38,”
consists of Part I. General Geology, by T. A. Jaggar, pages 17-41;
and Part II. Mining Geology, by J. D. Irving and 8. F. Emmons ;
214 pages, 19 plates, and 15 text-figures. Lastly, the Index,
8 pages.
This is one of the very useful contributions by the U.S. Geological
Survey to the inhabitants of a definite area of the recognizable
features and structure of the land, systematically considered, as well
as of its products and capabilities.
All notices of the Geology of the district already published are
duly mentioned at page 13.
The Black Hills in South Dakota have a dome-like structure of
454 Bibliographical Notices.
strata in successional order from the Alongkian schistose formation
at the base, up to the Benton (Cretaceous) beds at top. Thus
including: (1) the Cambrian (after the Alongkian) ; (2) the Silurian
or Ordovician ; (3) the Carboniferous ; (4) the Permian and doubtful
Triassic ; (5) the Jurassic; and (6) the Cretaceous. There are some
Tertiary and Quaternary deposits, of limited extent, here and there
on the surface.
The succession, chief characters, physical features, and relative
thicknesses of the several formations are given in the diagram at
page 18, and the table, pages 20 & 21; giving for these
superincumbent formations an aggregate thickness of more than
2000 feet.
These are more or less completely traversed by intrusive igneous
rocks, chiefly porphyritic, in dikes and sills, with laccolites. The
intrusions were probably made in the Eocene period. But the
general uprise of the area was not caused by the forcible addition
of the porphyries and their associated intrusive rocks, but by other
earlier and different crush-movements, intimately connected with
the great movements that elevated the Bighorn Range and
the Rocky Mountains. Many sections well show the extent of
erosion, exposing both the sedimentary and the igneous rocks, and
their local features; also the particular erush-breccias of certain
old limestones. Careful calculations are made as to the formerly
superincumbent formations ; also as to the probable order of events,
and the local results of movements in the various schists and
limestones.
The descriptive Geology of the mining districts (p. 33) treats
concisely of certain sections :—(I.) in the Alongkian Schists near
Homestake and Clover-leaf mines; (II.) in the Lower Palzozoic
Rocks, (1st) on the railroad near Terry Peak, (2nd) near Englewood
in the Eo-Cambrian limestone, and (3rd) on the Bear-Butte Creek
near the town of Galena; (III.) The Tertiary and Quaternary
deposits are not metalliferous, but some of the ‘ancient stream-
gravels may contain placer gold, but they have not been worked to
any extent, as compared with the gravels of the present gulches”
(p. 41).
Part II. Mining Geology begins with the gold-ore of the
Alongkian rocks in the Homestake and Clover-leaf mines (pages 55-
94), treating in full of their history, development, geological features,
and the metbod of working; also of the character and constitution
of the ores and their associated rocks and minerals, and of the
probable order of deposition and metamorphism. ‘Traces of copper
are noticed in the schists at City Creek and Deadwood Hill.
The ore-deposits in the Cambrian rocks of the North Black Hills
(page 98, &c.) are considered both generally and in detail. Of late
years they have given an aggregate output nearly equal to that of the
Homestake Mine. They are the Gold and Silver ores of the Cam-
brian basal conglomerates (pages 104, &c.), the ‘ Refractory Siliceous
Rocks ”’ (page 101, &c.), the Tungsten ores ‘page 162, &c.), and the
Lead and Silver ores (page 169, &c.). The very interesting account
Bibliographical Notices. 455
of the position, characters, and origin of the “ Refractory Siliceous
Ores” is full of information as to discovery, exploitation, extent,
characters, and the value of the several bodies of ore in the Bald
Mountain area (with its subordinate districts), also in the Garden,
Lead, Yellow Creek, and Squaw Creek areas.
The component minerals of these bodies of ore are quartz,
chalcedonic silica, pyrite, arsenpyrite, barite, fluorite, gypsum,
stibnite, and uranium-mica; but they exhibit considerable variations,
due to local alterations and rearrangements of constituents. The
methods of mining and treatment of the ores are referred to at
page 160.
‘The production of these Cambrian Siliceous ores in the North
Black Hills, in 1897, has been reported as amounting to $2.164.287.58
(page 117). The ore-bodies (or “‘shoots”) occur as more or less
banded masses, with a “ channel-like” form, nearly parallel to the
bedding-planes of the strata in which they are found, and are thick
in the centre with vertical extensions, and thin at the periphery.
Their origin is probably due to the passage of mineral waters along
the planes of bedding, and in very numerous cross-fractures, giving
rise also to extensive mineralization of the neighbouring (‘ country”)
rocks. The text-figures at pages 126, 132, and 157, as well as
plates xi. to xvii., are especially illustrative of these “ Refractory
Siliceous ores.” The reason for the name here given to them is
explained at page 111.
Towa Geological Survey.—Vol. XIV. Annual Report, 1903, with
Accompanying Papers. Samurt Catvix, A.M., Ph.D., State
Geologist. T.Savaen, Assistant State Geologist. With 664 pages,
38 plates, and 132 text-figures. Des Moines, Iowa Geological
Survey, 1904.
Tur mineral products of Iowa are generalized at pages 9-26, with
their localities according to Counties, and with the relative value of
the coal, clay, stone, gypsum, lead, zinc, and limestone. Of these
the clays are the most important; and a full account of the clay
industries in the State constitutes the main subject of this useful
volume.
Pages 29-348 treat of the “ Technology of Clay.” This abundant
hydrous silicate of alumina originates in the decomposition of the
felspar in granite and other rocks ; and, being distributed by rain,
river, and sea, is ultimately collected by nature in various modes
and conditions, known to geologists generally, and practically to
agriculturalists, civil engineers, and makers of pottery and other
ceramic ware. For them this chapter proceeds to treat in detail of
the origin and natural history of clay, how it is got, how prepared,
and of its economic uses. The physical properties and consequent
characters of clay are described ; and chemical analyses, both of the
raw andthe burnt material, aregiven. Thus the relative fineness of
grain in the fresh clay (p. 116) is very important in judging the
probable results in proposed manufacture of any kind of brick, tile,
456 Bibliographical Notices.
or pottery, since the various physical conditions directly influence
its fusibility.
Chapter LV. describes processes in the manufacture of clay-wares.
Firstly, the method of getting and preparing the raw material
(pp. 148-184) ; with illustrations of machinery and apparatus. The
making of bricks, tiles, pipes, and pottery follows; with remarks
on glazings (pp. 185-234). The methods of drying, heating, and
burning are described, with notes on fuel and the types of kilns.
Chapter V., “ The Chemistry of Clays” (pp. 322-345), gives the
methods, results, and value of different analytical processes; the
quantitative analyses of samples from seventy-two places in the
State are tabulated, pp. 344 & 345. Chapter VI. (pp. 347-376)
is concerned with the ‘ Selection, Installation, and Care of Plant
Power,” giving clear and useful reports on the engines, boilers, and
associated machinery used in the clay-works—how they should be
chosen, fixed, and utilized to the best advantage. With several
illustrative plates and diagrams.
Chapter VII. (pp. 379-534), “ The Geology of Clay,” deals with
the topographical and geological distribution of clays and shales in
Towa. Plate I. defines the 101 Counties of the State. Plate LI.
shows the outcrop of nine regular geological formations, with a
N.W.-S.E. strike and a general N.E. dip. Plate ILI. indicates
the distribution of seven superficial Drift-deposits and 220 localities
of special clay-pits.
The Map, PI. IL., recognizes the Cretaceous (see also p. 409), the
Carboniferous (Missourian, Des-Moines, Mississippian), Devonian
(p. 325), Silurian, Ordovician, Cambrian, and Alongkian Formations.
At pages 403 &c., after a notice of the Devonian Shales, the Lower
Carboniferous series, including the Kinderhook shales, are described
as being of some value. Inthe Upper Carboniferous (pp. 410-549),
including the Coal-measures, argillaceous beds, especially the fire-
clays, are very valuable and extensive. ‘These fire-clays or under-
clays (pages 411 &e., also p. 325), because of their having lost (by
leaching chiefly) most of their original alkaline constituents, do not
readily fuse, and, being resistant to fire, are well-known as “ fire-
clays.” The characters and uses of these and other clays are carefully
noticed according to localities and Counties. The seventy illustra-
tions (in plates and text-figures) given for the geological sections,
the local works, machinery, and apparatus, are of first-rate quality,
and fit well into their right places, both for artistic and economic
effect. This may also be said of the numerous other illustrations in
the volume, enhancing its value to all engaged in scientific studies
or business pursuits concerned with the handling of clays or with
the trading of its products.
The larger portion of the north-western third of the State of lowa
consists geologically of Cretaceous deposits (p. 509). In the upper
series argillaceous beds predominate, and near the top the marls and
limestones of the Niobrara Chalk appear. In the ‘ Lowa Geological
Survey,’ vol. iii. 185, this formation and its fossils were reported
on; and its Foraminifera were especially noticed in the ‘ Geological
Magazine,’ 1895, pp. 425-428.
Bibliographical Notices, 457
Pleistocene deposits, of glacial origin (Drift, &c.), cover almost all
the surface of the State, with varying depth to as much as 300 feet,
forming most of the landscape features, except where older rocks
give rise to larger hills and valleys. The Drift is a heterogeneous
series of boulder-beds, gravel-trains, and “ more or less sorted over-
wash.” Nearly structureless deposits of Loess (sands, silts, and
clays) occur in the Eastern Counties, and supply abundant material
ready for the clay-works. In most of the Drifts, however, the stones,
boulders, and pebbles give trouble, and, like the many calcareous
concretions, have to be eliminated in preparation for manufacture.
Seven areas of Drift on the map (PI. ILL.) are referred to five out-
spread sheets. Reckoning from above downwards, they are the
Wisconsin and its Moraine, the Iowan, the Loess, the Illinoian and
its Moraine, and the Kansan. At pages 525-554, however, they
are described according to the Counties, and from below upwards, as
follows: the Pre-Kansan or Albertan of Canada, the Kansan,
Illinoian, Iowan, the Loess, and the Red Clay or Gambo.
Chapter VIII.—“ Test of Clay-Products” is the subject of
pages 538-620 : different kinds of test, and different kinds of bricks,
common, hollow, and paving, tested by crushing, breaking, by
absorption, freezing and thawing; followed by a comparison and
summary of tests, with many tables and good diagrams. A
Directory of 294 Iowa clay-workers, arranged in Counties alpha-
betically (pp. 623-642), generally indicates the firm, the locality of
the works, the material used, its geological age, where obtained, the
plan of drying, kind of kilns, and the products. An Index of eight
pages, but not very perfect, completes the volume.
Maryland Geological Survey. Miocene. [Part 1.] Text: pages elv
and 543, with Platesi-ix. [Part II.] Plates x.-exxxy. 8vo.
Baltimore. The Johns-Hopkins Report, 1904.
Tun State of Maryland, occupying a district between the Alle-
ghanies and the Atlantic, and intersected by two great and broad-
mouthed rivers, the Potomac and Chesapeake, has a general slope
from west to east. Its surface is regarded by naturalists as divisible
at the Apalachian region, the Piedmont plateau, and the Coastal
plain, each of them continuing through the adjacent States of
Delaware and Virginia, and thus constituting the ‘‘ Middle Atlantic
Slope,” which comprises in its geology and mineral resources much
that is typical of the entire Atlantic border region (page wviv), A
portion only of the coastal plain in Eastern Maryland is here
described with the Mid-Tertiary or Miocene strata, of which the
surface is mainly composed. These strata and their fossils have
largely attracted the attention of American geologists and others
for two centuries and more according to the Bibliography at
pages wl—laiv, including the earlier sporadic notices and the more
recent systematic and well-nigh exhaustive treatises.
The relation of the Tertiary deposits of the Coastal Plains to the
belt of crystalline rocks in the Piedmont plateau and to the inter-
yening limited series of Secondary strata is referred to (page wut).
458 Bibliographical Notices.
Of the latter the Potomac group is the chief mass, consisting of six
or seven formations. Its lower portion is decidedly of marine
origin and of Cretaceous age, extending northward to New Jersey.
Overlying these are some Kocene formations (Nanjemoy and
Aquia). ‘They are succeeded on the south-east by the Miocene
deposits, formerly known as the Chesapeake group, but now divided
by the State Geological Survey into three well-defined formations—
‘*the Calvert, Chaptank, and St. Mary’s.”
The Chesapeake group consists of variously coloured clays and
sands, sometimes fully charged with shells; especially when
comminuted; these form limestone ledges. The lower part of the
Calvert series is rich in Diatomaceous beds, in places 20-40 feet
thick.
Gravel, sand, and clay, probably of Pliocene age, are noticed in
Maryland. Later deposits of the mid-Atlantic slope, known as the
Columbia group, of Pleistocene age, have been divided by the
Geological Survey into three divisions ; the oldest of these, namely
the Sunderland, consists of gravel, clay, and loam, with ice-borne
boulders on the highest lands on the west shore of Chesapeake Bay.
The Wicomico in formation is similar to the Sunderland and skirts
the high-land capped by the former. The Talbot, somewhat similar
in formation, occupies the low level of the Coastal plain, seldom
rising above 40 feet. It has much peat and boulders and larger
areal extent than other Pleistocenes.
The continuous examination of the fine series of fossiliferous
strata has been well carried out, and its various characters have
been studied both individually and in relation with similar geolo-
gical formations in other parts of the world.
The different areas of local seas during the great period defined
as the Miocene or Mid-Tertiary, their shifting coasts, varied sea-
bottoms and faunal characters, and their successive climatic changes
can be fairly well indicated by geologists careful in work and in
arriving at philosophical conclusions. This aspect of the subject
is elaborated by W. H. Dall and others at pages crxwiv-clv, on the
basis of the comprehensive tabular lists of the Maryland species,
according to local distribution, at pages wciv-caxi, and of the
Mollusca alone in their general geographical and geological distri-
bution (pages cvviv-cavavir).
The systematic paleontology of the Miocene formation of
Maryland is very fully described and illustrated in Part II., pages
1-508, with 132 plates and some figures in the text, Scrupulous
exactitude appears to have been the rule in both descriptions and
drawings, and so abundant and so excellent are the figures that
they may serve the student with materials sufficient (with some
groups of fossils) to make him well acquainted with the facies of
generic and even specific groups. For fossil Vertebrata (Mammalia,
Aves, Reptilia, and Pisces) see plates 10-32; Cirripedia, pls. 33,
34; Ostracoda, pls. 35-38 ; one Cephalopod and many Gasteropods,
pls. 89-63; Amphineura and Scaphopoda, 1 plate; Pelecypoda,
pls. 65-108 ; one Brachiopod and many Bryozoa and one Spirorbis,
Bibliograph ical Notices, 459
pls. 109-118; Echinodermata, pls. 119, 120; Hydrozoa, pl. 121 ;
Anthozoa, pls. 122-129; Radiolaria, pl. 130; Foraminifera,
pls. 131-133; and Diatomace, pls. 134, 135.
The conclusion drawn by W. H. Dall, Part I. page cl, is that the
Miocene of South Europe has a rather more tropical character than
that of Maryland. It is known that a northern fauna invaded the
Miocene area in Europe ; but in the Atlantic region the Chesapeake
formations show that, after an elevation of land and a somewhat
warmer climate, the tropical current was modified and more
temperate conditions prevailed at the end of the Miocene period.
The geological map of the Miocene formations in Maryland
and eight plates (in Part I.) of good photographic views of
local Miocene sections, are useful adjuncts to this interesting
work ; and the same may be said of the full Index, divided into
“general” and “ paleontological,” at pages 509-543 of the same
Part I.
J. H. Fasre. Souvenirs Entomologiques. Etudes sur V Instinet
et les Noces des Insectes. (Neuviéme Série.) 8vo. Paris [1905].
Pp. 374.
Here we have another volume of interesting natural history
romance (‘“ Wahrheit und Dichtung”) from an elegant French
writer, who reminds us somewhat of Alphonse Karr, but is at the
same time a trained and excellent naturalist, whose observations
are always of great interest and value. There is nothing dry or
technical in the writings of Fabre, and they might be read and
appreciated for their delightful style by anyone who can read
French.
The present volume, however, contains less matter than usual
relating to insects in the strict; sense; only the last two of the
twenty-five chapters are devoted to the pretty Coccid, Dorthesia
characias, Latr., which is found on Huphorbia (other species of
the genus are found in Britain on nettle), and to the kermes of the
Ilex, The rest of the volume, exclusive of chapters 13 and 14,
“ Souvenirs mathématiques,” which contain general dissertations,
with which Fabre likes to vary his scientific writings, are devoted
almost entirely to Arachnida: spiders and scorpions, especially the
Lycosa of Narbonne, Epetra fasciata, Clotho Darandi, and the
scorpion of Languedoc (Scorpio occitanus). There are a few illustra-
tions in the text— La promenade & deux,” on p. 305, representing
a pair of scorpions walking hand-in-hand, being perhaps the most
noteworthy. The first volume only of this series has yet been
translated into English; but Fabre is an author whose works lose
so much in translation that it is far better, for those who are able,
to read them in the original, for they are as good literature as good
science,
460 Geological Socrety.
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
February 1st, 1905.—J. E. Marr, Sce.D., F.R.S.,
President, in the Chair.
The following communication was read :—
‘On the Sporangium-like Organs of Glossopteris Browniana
D D5 4 ’
Brongn.’ By HE. A. Newell Arber, M.A., F.L.S., F.G.S.
It has been discovered that some specimens from New South
Wales, on which numerous and well-preserved scale-fronds of
Glossopteris occur, also exhibit impressions of groups of minute
bodies, not unlike the sporangia of certain recent and extinct Ferns
and Cycads. The new organs are elliptical in shape, and measure from
1-2 to 1-5 millimetres along the major axis. They are hollow, sac-
like structures, which open or dehisce longitudinally. Unfortunately,
no trustworthy evidence can be obtained as to their contents. The
fact that they have never been found, except in the closest asso-
ciation with the scale-leaves of Glossopteris, is regarded as an
indication that they may be attributed to that genus; and this
conclusion is supported by the direct evidence of some of the scale-
fronds, which show scars of attachment and fragments of the sac-
like bodies still apparently in continuity. While, in the absence of
recognition of spores, it is impossible to be quite certain that these
sac-like bodies are really sporangia, there is much to be said in
favour of this conclusion. The fact that they were probably borne
on the scale-fronds in sorus-like groups, and that in size, shape,
and mode of dehiscence they are not unlike the sporangia of certain
recent and fossil plants, supports this view. The closest analogy
may probably be found in the microsporangia of Cycads, which are
also exannulate.
An historical sketch is given of the present evidence on the
subject of the fructification of Glossopteris ; and it is shown that no
sporangia have so far been found, and that such evidence as has
been put forward as to the occurrence of sori on the larger fronds
is not altogether satisfactory. It is pointed out that, if the present
conclusion be correct (that the sporangia were borne on the smaller
scale-fronds), Glossopteris cannot be included in any recent family
of the true Ferns. Although the affinities of this genus are still
doubtful, the creation of a new family may not be inadvisable.
Geological Soctety.— Miscellaneous. AGB1
March 8th, 1905.—J. KE. Marr, Sc.D., F.R.S.,
President, in the Chair.
The following communications were read :—
1. ‘Observations on some of the Loxonematid, with Descriptions
of two New Species.’ By Miss Jane Donald.
Shells having more convex whorls, or less sigmoidal lines of
growth than Z. sinuosum, cannot be left within the genus Lowonema.
The two new species described resemble the type in form and in the
sinuosity of the lines of growth; but the whorls are ornamented with
spiral strie, two of which frequently stand out and give the shell a
banded appearance. Both species are of Silurian age: one was
obtained from the Aymestry Limestone near Ledbury, and from
beds of about the same age at Llangadock ; and the other from the
Wenlock Limestone, Dudley.
2. ‘On some Gasteropoda from the Silurian Rocks of Llan-
gadock (Caermarthenshire).’ By Miss Jane Donald.
These fossils occur almost entirely in the state of casts and moulds.
Eleven distinct forms have been made out, referable to seven genera;
but only seven are sufficiently well-preserved for specific determina-
tion. Five of these are new, including one described in the previous
communication; a new genus is described, for the reception of
Euomphalus funatus. The first species described comes nearest to
examples placed in the genus Plethospira, as represented by Pl. Semele,
with which it might be placed in a new subgenus. Three other
species, including two new ones, resemble shells referred to Bemboscia.
The three species are turriculated, and the band is situated on the
angle of the whorls, a little distance above the suture. The specimens
described come from the horizons of the Upper Llandovery, the
Wenlock, the Lower Ludlow (including the Aymestry Limestone),
and the Upper Ludlow.
MISCELLANEOUS.
The Generic Names given by Frisch in 1775.
By Ovprietp THomas and Gerrit 8. Mirier, Jun.
Tur excessively rare ‘Natur-System der vierfiissigen Thiere’*,
published by Frisch in 1775, appears to have remained quite
* ‘Das Natur-System der vierfiissigen Thiere, in Tabellen, darinnen
alle Ordnungen, Geschlechte und Arten, nicht nur mit bestimmenden
Benennungen, sondern beygesetzten unterscheidenden Kennzeichen
angezeigt werden, zum Nutzen der erwachsenen Schuljugend,’ yon J. L.
Frisch. Glogau, 1776.
462 Miscellaneous.
unnoticed, so far as nomenclature is concerned, until 1902, when
Sherborn, in the ‘Index Animalium,’ declared it to be a non-binomial
work. Two years later Palmer, apparently overlooking Sherborn’s
statement, included about a dozen Frisch names in the ‘Index
Generum Mammalium,’ whence a few have recently been introduced
into the nomenclature of mammals. But that this course is quite
unjustifiable seems to us evident after carefully examining a copy of
Frisch’s book which Mr. Sherborn has kindly placed at our disposal.
The work appears to have been intended primarily as a text-book
for pupils in natural history. It deals with mammals and reptiles
only, twenty-three pages being devoted to the former, seven to the
latter. These animals are arranged according to a classification
based on that of Klein, but containing so many new features that
its author did not feel at liberty to connect with it the name of
“the great Pliny of Danzig.” Three categories are recognized—
“ Societas,” ‘* Genus,” and ‘‘ Species,’—and under each category
the names, German and Latin, are arranged column-wise, three
columns to a page. The following are a few examples :—-
Societas. Genus. Species.
Bovina. 1. Bos, der Ochs. 1. Bos vulgaris, gemeiner Ochs.
Ochsenartige.
Ovina. 1. Ovis communis, 1. Ovis vulgaris, gemein Wollen-
Schafartige. gemein Schaf. Schaf.
Tragelaphina. 5, Bubalis, 1. Buselaphus, der Bubalis, der
Gazellenartige. der Stierhirsch. Alten.
8. Tragulus, 1. Guevi, Rehzieglein von Akra.
das Rehzieglein. 2. Guevi Kagor, von Sennegall.
9. Lragus moschifer, Tragus moschifer, Muskusthier.
Muskusbock.
Rhinocerotina. 1. Rhinoceros, Monoceros, et Bi- 1. Rhinoceros unicornis, Nashorn
Nashornartige. cornis, Nashorn mit einem oder mit einem Horn.
2 Hornern.
Felina. 5. Leo, der Liéwe; Leena, die Li- Es sind zwey Rassen yom Lowen.
Katzenartige. winn.
In an unpaged table at the end the whole of the system is
repeated, this time classified into Ordines, Gentes, Societates, and
Genera, the species not being mentioned. The generic names used
in this table are quite commonly different from those in the main
part of the work, and many of them are again double (e. g. Tatu
Armadillus, Araneus mus) or otherwise technically impossible.
At first sight the majority of the names in the second and third
columns of the paged part and in the table at the end have an
attractively Linnean appearance, but on reading the work con-
secutively it becomes evident that those in the third column have
no invariable relation to those in the second, while among the
latter we find such irregularities that all illusion of binomialism
quickly disappears. As names for genera we find such compound
Miscellaneous. 463
terms as “ Ovis communis,” “ Ovis auribus pendentibus,’ “ Tragus
moschifer,” ‘* Mus araneus,” and ‘ Phoca ursus”: while for others
there are alternative names, as ‘“‘ Papio, Pavianus, der Pavian” ;
“‘ Cercopitheca oder Cercopithecus, auch Cebus”; ‘ Phoca minor,
Canis marinus, Seehund” ; ‘* Phoca major, Leo marinus, Seclowe” ;
“ Bos marinus, Vacca marina, der See Ochs oder die See Kuh” ;
“‘ Catus, Felis, der Kater, die Katze”; ‘“‘ Leo, der Lowe, Leena, die
Lowinn.” It seems unnecessary to cite further instances to show
that this system of naming is hopelessly non-Linnean. What
Frisch apparently intended to do was merely to supply his pupils
with Latin equivalents for German vernacular terms; but of the
essential principles of binomial nomenclature he had not the least
idea *.
While we therefore unhesitatingly reject as untenable all of the
names applied to genera + in this work, the circumstances are such
that it seems desirable to publish a complete list of them.
List of Names applied to Genera by Frisch.
MAMMALIA,
1. Equus. Asinus. Bos. Urus. Bubalus. Ovis communis. Ovis auri-
bus pendentibus. Ovis montana. Ovis strepsiceros. Caper. Ibex.
2. Gacella. Adaces. Antilopes. Rupicapra. Bubalis. Cudus. Grimms.
Tragulus. Tragus moschifer. Camelopardalis.
8. Cervus. Dama. Alce. Rangifer. Capreolus. Porcus. Macro-
cephalus. Tagassu. Babirussa.
4. Elephas. Equus Niloticus. ‘Tapirussa. Rhinoceros, Monoceros, &
Bicornis. Camelus, Lama.
5. Tamandua. Quaggelo. Tatu. Erinaceus.
6. Talpa. Mus araneus. Marsupiale. Volucre.
7. Vespertilio. Mus. Rattus.
8. Glis. Sciurus.
9. Cayia. Lepus. Marmota.
10. Castor. Hystrix.
11. Mustela. Putorius. Ichnevmon. Martes.
12. Catus, Felis. Linx. Panthera.
13. Tigris. Leo.
14, Canis. Lupus.
15. Hyena. Vulpes.
16. Coati. Meles. Cibeticum.
17. Lutra. Gulo.
18. Ursus.
19. Tardigradus, Tardipes. Papio, Pavianus.
20, Cercopitheca oder Cercopithecus.
21. Simia.
* Or, if he had any clear notion of it, he deliberately rejected the
system. ‘ Des grossen schwedischen Naturforschers Ritter v. Linné seine
weitliutige und priichtige Systemen, welche er so oft verindert, waren,
nach meiner Einsicht, zu meiner Arbeit unbequem,” he remarks in his
preface (p. ii), and the observation is amply justified.
+ No one bas suggested that the specitic names are valid, and, in fact,
their status is too evident to require any discussion.
46 t Miscellaneous.
22. Phoca minor, Canis marinus. Phoca major, Leo marinus. Phoea ursus,
ursus marinus.
23. Rosmarus. Bos marinus, Vacca marina.
REpPrivia.
24. Testudo digitata. Testudo alata digitis alatis. Orocodilus.
25, Caimanus.
26. Lacerta, Lacertus.
27. Guana, Iguan. Salamandrina. Salamandra. Stellio, Jekko.
28, Cordylus, Uromastix. Scincus. Chalcis, Seps, auch Putria. Chamzleo.
29. Rana.
30. Bufo.
The Secondary Appendage of the Upper Antenne as a Character
in the Amphipoda. By Aurrep O. WALKER.
From the time of Leach (1815) the presence or absence of an
appendage to the upper antenne in the Amphipoda has been used
as a character to separate not only species but genera and families.
As long as the antenne were not examined too closely, and no
notice was taken of the appendage in a rudimentary condition, the
practice worked fairly well, though open to the objection that the
antennse are more often missing in an Amphipod in spirit than any
other part. But when, in consequence of the discovery by Stebbing
of the appendage in certain species of Metopa in the ‘ Challenger ’
Collection (all from one locality), Della Valle, regardless of the fact
that in most species of this genus no appendage has been found,
places two genera so nearly allied as Stenothoé and Metopa in
different families, it is time to enter a protest. Even as a specific
character the appendage is unsatisfactory. Young specimens often
have fewer joints in it than adults, and in Hlasmopus sokotre,
Walker, all the males examined had the appendage, while the
females were without it.
The question has been forced on my notice by the examination
of a species of Iphimedia in the ‘ Discovery’ Collection. A young
specimen, 15 mm. long, has a secondary appendage which is rather
conspicuous, being half as long as the second joint of the flagellum ;
while in another specimen from the same tube, but double the size,
the appendage is reduced to the most rudimentary condition. The
species is in every other respect a typical /phimedia ; yet, according
to G. O. Sars’s definition of the family Iphimedide, the immature
form does not belong to it!
In conclusion, while we can hardly afford to dispense altogether
with the appendage in our diagnoses of species, yet I submit it
should be used with caution and only when it is conspicuous; a
difference in the number of joints should not be considered sufficient
to constitute a distinct species.
Ulecombe Place, Maidstone.
July 15, 1905.
Ant.& Mag. Nat.Hist.5.7. Vol XVI_Pi Xi.
ph.i-5. hyd. ph.-e2 x.
fe
Henderson del. B.&.D.04imp. J.Green lith.
if
Two
Ann .& Mag. Nat. Hist 1S. 7; Vol XVI
——
a
\
BES ey, “5
LOS
#,
ch
‘al tat! ich {
Lib Sos 8
om ety S
{Ba
Henderson del B.& D.14 imp J.Green th.
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”
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. 95. NOVEMBER 1905.
LIII.—Descriptions and Records of Bees.—V.
By T. D. A. Cockrrett, University of Colorado.
Exonevra, Smith.
Abdomen black ; face with a large cream-coloured
Maher ee nes So Gietete et ala ey eer epiiare’se a Tapanw eras Neko ease S botanica, Ckll.
PAU OMNIA Ups eee eve et Waa Pee aisle at ceed Ne if
1. Face entirely dark; size small, length 4mm. .. Froggattit, Friese.
Face with at least a light streak; size larger .. 2.
2. Second abdominal segment with a black patch ;
face with an anchor-shaped cream-coloured
PERE Na a ee recat it racers o Soa hamulata, Ckll.
Second abdominal segment without a black patch ;
face with only a yellow streak.............. bicolor, Smith.
Allied to E. hamulata, but smaller, and with
MUElloWeINNerclasitss iss cakes che toe oe fae tau, Ckll.
Ezxoneura bicolor, Smith.
Franklin and Hobart, Tasmania (J. J. Walker, 3481, 3246,
3248).
Ezoneura Froggattii, Friese.
Moss Bay, 13. 12. 1893 (Froggatt). Two specimens.
Exoneura botanica, sp. n.
? .—Length about 44 mm.
Black, including abdomen ; hind margins of abdominal
segments reddish, especially the third and following ones ;
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 31
466 Mr. T. D. A. Cockerell—Descriptions and
tarsi dull red, as also are the anterior tibie, and an apical
patch on the middle and hind ones; face narrow, the orbits
converging below; an elongated depression at each side of
the antenne; clypeus with a very broad cream-coloured
band, broadest above, with a sudden enlargement something
like the head of a nail; mesothorax smooth and shining ;
tegule margined with reddish. Wings dull hyaline,
iridescent ; stigma and nervures very dark brown. The
tubercles are cream-coloured.
Hab. Botany, New South Wales, 12.10. 1893 (W. W.
Froggatt, 192).
EHxoneura hamulata, sp. n.
? .—Length about 62 mm.
Head and thorax shining black; abdomen chestnut-red
(a rather darker shade than that of dicolor), the first two
segments each with a large black discal patch; legs red, the
coxe, trochanters, aud greater part of the femora black ; hind
tibiee and tarsi with coarse abundant reddish bristles, rufo-
fuscous on the outer side. Head Jarge, face broad, clypeal
cream-coloured band suddenly broadened above, with a sharp
hook on each side. Wings yellowish; stigma and nervures
dark ferruginous, the stigma varying to lighter. The
apical part of the abdomen is roughened, as usual in the
genus.
Hab. Moss Bay, Australia, 13. 12.1893 (W. W. Froggatt).
Two specimens.
Evoneura tau, sp. 0.
Similar to /. hamulata, but smaller (length of anterior
wing 4mm.) ; second submarginal cell much smaller ; clypeal
stripe narrow, with a short cross-bar at the top, which is not
at all hooked ; tubercles yellow. By the light tubercles it
resembles only L. botanica, but that has a black abdomen, dark
stigma, and the clypeal stripe very much broader. /. Frog-
gattii has the comparatively small second submarginal cell,
but that is a much smaller insect.
Hab. Moss Bay, Australia, 13. 12. 1893 (W. W. Froggatt).
It bears the collector’s number 177.
Unfortunately the single example had the abdomen broken
off in transit, and the two basal segments are missing.
Apis mellifera, L.
Perth, W. Australia (H. W. J. Turner) ; Hobart, Tasmania
(J. J. Walker, 3216).
Records of Bees. 467
ProsopPis.
The species discussed below may be separated as follows :—
Abdomen wholly or largely red .............. Le
Abdomen black (in one species reddish black) .. 5.
Abdomen blue; size very small .............. 8.
. Legs, except coxe, entirely bright yellow; scutel-
lum and postscutellum yellow .............. sydneyana, Ckll.
Legs not so; scutellum and postscutellum not
LOI oper she erate tey recs ee SAE afc a oes Shes evehers oles aves 2.
. Mesothorax and scutellum red ...........0.00 lateralis, Smith.
Mesothorax and scutellum black ............-. 3.
. Apical half of abdomen dark fuscous .......... constricta, Ckll.
Apical half of abdomen red like the basal half.. 4
HUprachypeal mark PrOsenh 6.02. ese. acs eens coronata, Ckll.
Supraclypeal mark absent................205- subplebeia, Cll.
. Large and robust, about 12 mm. long ; scutellum
witha lish patel: 222i)... % sie s)eleleysisyy eae 6 percrassa, Ckll.
Much smaller; scutellum whoily black........ 6.
. Yellow of prothorax confined to tubercles, face
yellow nearly up to level of antenne ........ hobartiana, Ckll.
Yellow of prothorax not confined to tubercles .. 7.
Bilarger? clypous blacke! o)).. 0 tiods «a 1a sios0 Bet = amicula, Smith, °.
Smaller; face all yellow below antennz; abdo-
Ret ReU Ish lala ae. aca ciel yala toler ste aa> ois a primulipicta, Ckll.
. Face blue or greenish blue, without light
HET EMSS ion ooh A ebb Dr oneobe nubs solver oer baudinensis, Ckll.
Face with cream-coloured laterai marks ...... albonitens, Ckll., 2.
Prosopis albonitens, Ckll.
The female is like the male, except for the usual sexual
differences. The clypeus is green, and the scape is slender
and dark.
Queensland (Gilbert Turner, Ridg. 11. 93, 712).
Prosopis amicula, Smith.
Queensland (Gilbert Turner, Ridg. 620). One ?, about
6 mm. long.
The basal area of metathorax is irregularly ridged and
bounded by a rim.
Prosopis lateralis, Smith.
Queensland (Gilbert Turner, Ridg. 11. 93, 858).
9? .—The hind tibia has a creamy-white subbasal annulus.
Base of metathorax dull and granular.
Prosopis sydneyana, sp. 0.
g —Length about 6 mm.
Head and thorax black, with yellow markings ; abdomen
ee
468 Nir: SA Cockerell— Descriptions and
with the first two segments red, the others reddish black,
the third red basally at the sides; legs bright yellow.
Face produced, the clypeus prolonged and the malar
space quite large; face below antenne, labrum, and
mandibles bright chrome-yellow; the yellow extends
upwards in the middle line, gradually narrowing, to the
middle ocellus ; laterally, along the orbital margins, it extends
about as far, but ends more bluntly ; ; cheeks entirely black ;
scape yellow, swollen ; flagellum light ferruginous ‘beneath,
darker above ; mesothorax minutely but distinctly and rather
closely punctured; scutellum, postscutellum, collar, tubercles,
tegulee, and a band on each lateral margin of mesothorax bright
yellow ; area of metathorax with regular longitudinal basal
ridges, terminated by a transverse ridge, while at the sides a
curved ridge bounds a larger area. Wings clear hyaline,
iridescent; stigma honey-colour with a narrow fuscous
margin ; nervures fuscous; second s.m. very much broader
than high; first r. n. joining apex of first s.m.; abdomen
punctate.
Hab. Syduey, New South Wales, 1891 (IW. W. Froggatt,
28).
Prosopis constricta, sp. 0.
¢ .— Length about 5 mm.
Head and thorax black, dull, the vertex and meso-
thorax with close minute punctures; abdomen with the
first two segments red, the rest dark fuscous, the fuscous
extending on to the apical part of the second segment.
Clypeus, labrum, and mandibles dull white, the white of
the clypeus notched or constricted just above the middle
by a black spot on each side; no supraclypeal mark ;
lateral marks grey, extending very narrowly some distance
above the antenne; scape rather broad, whitish in
front; flagellum thick, hght ferruginous beneath, greyish
fuscous above, the last joint pointed; sides of collar and
tubercles brownish white ; tegule light brown with a white
spot; area of metathorax semicircular, bounded by a rim,
but not definitely ridged except in the middle, where there
is a strong longitudinal ridge, with a weak one on each
side of it. Wings hyaline and iridescent, nervures and
stigma fuscous; second s.m. comparatively short (narrow),
receiving the first r. n. a short distance from its base, and
the second at its extreme apex. Femora dark brown, the
anterior knees pallid; anterior tibize light amber-colour,
with a fuscous spot behind; middle tibiz light in front and
dark behind, except at extreme apex and base; hind tibize
Records of Bees. 469
dark brown with the base broadly white; tarsi yellowish
white, the hind tarsi fuscous beyond the first jomt ; abdomen
narrow, the punctures so fine as to be hardly visible with a
lens.
Hab. Queensland (Gilbert Turner, Ridg. 10. 91, 708).
Prosopis coronata, sp. 0.
3g .—Similar to P. constricta, but differing thus : clypeus
and lateral face-marks white, suffused with light red ; clypeus
hardly constricted by the black spots, which are minute;
supraclypeal mark present, large, nearly all lght red,
tridentate above, having the appearance of a crown; antenne
entirely clear red, the flagellum scarcely darkened above,
the last joint not so pointed ; collar and tubercles entirely
black ; tegule fuscous, without a white spot; area of meta-
thorax without a median keel, but with strong, oblique,
lateral keels; middle tibize not light in front, except at apex ;
hind tibize without any distinct white base ; abdomen broader,
entirely clear bright red.
Hab. Queensland (Gilbert Turner, Ridg. 11. 91, 707).
Prosopis subplebeia, sp. n.
.—Similar to P. coronata, but differing thus: no supra-
clypeal mark ; clypeus without red, except a little well-defined
patch on each side ; lateral marks grey ; scape with a cream-
coloured patch in front ; collar with a yellowish streak on
each side, and a small light mark on tubercles ; tegule with
a whitish spot ; area of metathorax more regularly sculptured,
the fine lateral keels numerous and regular, and the median
area well ridged ; apex of abdomen somewhat infuscated.
This species is curiously intermediate between coronata and
constricta, but nearest to the former, and apparently distinct.
Hab. Queensland (Gilbert Turner, Ridg. 11. 91, 707).
The collector apparently regarded this and the last as one
species, as he gave them the same number; but the absence
of a supraclypeal mark and decidedly different sculpture of
the metathorax seem to indicate a different species.
Prosopis percrassda, sp. ni.
? .—Length about 12 mm.
Robust, black, strongly punctured; clypeus _ striato-
punctate, with a short groove on each side; no light
face marks, except a dark red spot between the antenne ;
470 Mr. T. D. A. Cockerell—Descriptions and
mesothorax shining between the punctures; disk of
scutellum shining, with the punctures sparse though
large; area of metathorax triangular, with a densely
punctate median area, but otherwise smooth and shining,
posteriorly produced and obtusely angulate; the area is
not bounded by any rim, but sharply by the cessation of
the dense punctures which cover the rest of the metathorax;
a stripe on each side of collar, large spot on tubercles,
triangular spot on the piccous or black tegulz, spot on axille,
large triangular mark on middle of scutellum, and wedge-
shaped mark on middle of postscutellum, all cream-colour,
some of them brighter than others; lateral margins of
metathorax strongly tufted with greyish-white hair. Wings
hyaline, the apex faintly dusky, nervures dark fuscous, stigma
dark ferruginous ; second submarginal cell very broad (long),
about three-quarters length of first, receiving the first r. n.
a long distance from its base, and the second r. n. much
nearer its apex. Legs black, with white pubescence. Abdomen
broad, the apical part thinly but conspicuously hairy ; lateral
hind margins of first segment with dense hair-bands ;
punctures of first segment small, and very close posteriorly,
strongly contrasting with the very large, well-separated
punctures on base of second.
Hab. Queensland (Gilbert Turner, Hy., 273). Presented
to the Museum by Mr. E. Saunders.
Somewhat allied to P. alcyonea.
Prosopis hobartiana, sp. n.
é —Length about 6 mm.
Black, head and thorax dull, the punctures of the vertex and
mesothorax so fine and dense as to produce a minutely granu-
lar effect; head large, broader than thorax; face comparatively
broad, chrome-yellow nearly up to the antenne, including a
supraclypeal mark, the top of which presents a hitle angular
elevation ; laterally the yellow gradually rises, to end in a fine
point on the orbital margin about the level of the antenne ;
labrum and mandibles with much yellow ; scape with a yellow
stripe beneath ; flagellum all dark ; collar dark, but tubercles
yellow with a black dot; area of metathorax formed much
as in percrassc, but the basal part finely granular ; scutellum
sculptured like mesothorax ; tegule piceous. Wings strongly
infuscated, stigma and nervures piceous ; second s.m. much
shorter than first; the two recurrent nervures meeting the
transverso-cubitals. Femora black, the anterior knees pale
reddish; anterior tibize pale reddish with a dusky stripe
Records of Bees. 471
behind ; middle tibize black, with the extreme base yellowish,
and a pale reddish subapical stripe in front ; hind tibiz black,
with about the basal two fifths light yellow ; tarsi with the
first joint yellowish or reddish, the others infuscated ; abdomen
black, rather narrow, moderately shiny.
Hab. Hobart, Tasmania (J. J. Walker, 3249).
Prosopis primulipicta, sp. n.
¢ .—Length about 4) mm.
Slender, black, head and thorax dull, the abdomen slightly
reddish and with a silky lustre; markings bright primrose-
yellow, including labrum, mandibles, face up to level of
antenne, and sending a pointed projection on each side a
little way up orbital margin, collar, and tubercles ; scape with
a yellow stripe beneath; flagellum long, lhght yellowish
fulvous beneath (contrasting with the shining dark funicle),
above dark fuscous; tegule dark. Wings hyaline and
strongly iridescent, the apical field somewhat dusky, and the
upper half of the marginal cell more strongly so; nervures
and stigma piceous ; second s.m. very short, much less than
half length of first, receiving second r.n. at its extreme
apex, but the first r. n. enters the apical part of the first
s.m.; area of the metathorax finely granular, minutely
somewhat plicate basally, defined only by an impressed line ;
femora black, the anterior ones yellow at apex; anterior
tibize yellow, with a large reddish-fuscous spot behind ;
middle tibize black in the middle and yellow at each end,
the yellow connected by a narrow band in front ; hind tibize
black with nearly the basal half yellow; tarsi yellow, the
hind tarsi fuscous with only the basal half of the tirst joint
yellow.
Hab, Queensland (Gilbert Turner, Ridg. 11. 91, 714).
Prosopis baudinensis, sp. n.
? .—Length about 42 mm.
Head, thorax, and abdomen deep blue; sometimes some
green tints on face, which is without light markings ;
antennz dark, the flagellum sometimes more or less ferru-
ginous ; mesothorax shining and strongly punctured ; collar
dark, but tubercles mainly white, with a black dot; base of
metathorax shining, no well-defined enclosure; tegulez
piceous. Wings hyaline and iridescent, nervures and stigma
piceous; second s.m. not far from square, receiving the
recurrent nervures at the extreme base and apex. Legs black ;
middle tibiz with a white spot at base; hind tibie with the
472 Mr. T. D. A. Cockerell—Descriptions and
base broadly white; small joints of anterior tarsi pale
reddish.
Hab. Baudin I., N.W. Australia (J. J. Walker, 673, 674).
Three specimens,
EURYGLOSSA.
The following species, referred to Euryglossa, are probably
not all truly congeneric; but Smith’s species of this genus
also seem to represent more than one generic type, and it will
be best to defer the proposal of segregated genera until more
material is available.
Clypeus with a prominent longitudinal keel;
mesothorax very shiny; abdomen dark red. Froggattiana, Ckll.
Clypeus without a keel; size smaller........ 1.
1. Second and third abdominal segments almost
entirely OFanpe .<\. aes on scree tions cea Bere cygnella, Ckll.
Second and third abdominal segments dark ..
2. Head and thorax very dark green .......... Wr alkeriana, Ckll.
Head: and thorax blacks5 3.2 sinissp ie ee tee
5. Extremely small, about 33 mm. long....... ; a nana, Ckll.
Much larger; hind tarsi (at least) red ...... 4,
4. Mesothorax with sparse feeble punctures .... meglectula, Ckll.
Mesothorax densely punctured.............. 5.
5. Hind tibize strongly infuscated; wings not
Vollowashie. uion oye nee Eien hee cee regine, Ckll.
Tlind tibiz not infuscated; wings yellowish .. adelaide, Ckll.
Euryglossa Froggattiana, sp. u.
? .—Length nearly 9 mm.
Head and thorax shining black ; abdomen broad and
flattened, dark red, clouded with fuscous, especially at the
sides ; head large and broad; clypeus with the form of a
very large equilateral triangle, with a strong and sharp
reddish keel running down its middle, which is elevated ; the
keel continues weakly upwards about ‘halfway to the middle
ocellus, after which it is replaced by a groove; sides of
clypeus and adjacent sides of face with very sparse weak
punctures ; front shining, with numerous exceedingly minute
punctures ; narrow black facial foveze are closely adjacent to
the upper orbital margins, their upper part curving away from
the eye ; scape dark reddish, with scattered hairs ; flagellum
with the apical half thickened, the base reddish, the rest
dark, with the underside dark red; malar space wanting;
mandibles slender, reddish in the middle ; maxillary palpi
6-jointed, ordinary; labial palpi 4-jointed, first joint darkened,
curved, considerably longer than the second, and a little
Records of Bees. 473
longer than 3 and 4 united; mesothorax convex, smooth,
with very few minute punctures ; scutellum broad and
flattened; area of metathorax cup-shaped, smooth and
shining, scarcely defined; margin of tubercles tomentose ;
sides of metathorax with a hairy fringe ;_ tegulz small, light
apricot-colour. Wings dusky hyaline ; stigma dark reddish
brown, nervures lighter; second s.m. hardly half as long as
first, receiving the recurrent nervures not far from its base
and apex; basal nervure falling far short of transverse-
medial. Legs hight red, with thin white hair, the femora
darkened, except at base and apex; hind tibiz tuberculate
on the basal half of the outer edge; pygidial plate extremely
narrow.
Hab. Shoalhaven, New South Wales, 4. 10. 1894 (W. W.
Froggatt, 175).
Euryglossa cygnella, sp. n.
3 .—Length 5 mm.
Slender, black, with the second and third abdominal
segments above orange, except the broad hind margins and
the sides more or less; on tbe ventral side segments 2 to
4 are pale orange ; head and thorax with scanty, but rather
long, dull white hair ; head transversely oval, eyes converging
below ; antenne long, the flagellum stout, and pale yellowish
ferruginous beneath ; mesothorax dull and very densely
punctured ; area of metathorax large, triangular, with a very
narrow basal band enclosed by a raised line, and marked by
numerous little transverse (antero-posterior) ridges ; behind
the basal band the area is covered for a space with very fine
raised lines, and its lateral margins are more or less marked
by little raised lines; tegulz very pale testaceous. Wings
hyaline, stigma large and piceous, nervures very dark
brown ; second submarginal cell about or almost as long as
first, receiving the first r. n. a considerable distance from its
base, and the second very close to its apex. Femora black,
with a little reddish on the knees ; anterior and middle tibize
and tarsi clear ferruginous; hind tibiz ferruginous, with a
dark cloud anteriorly, and conspicuous bristles in little groups
of two or more on the hind edge; _ hind tarsi fuscous ;
abdomen narrow and flattish, the first segment narrow and
long, the black parts with a dull satiny lustre.
Hab. Swan River, Australia, ‘‘ 69. 50.”
Euryglossa Walkeriana, sp. u.
? —Length about 52 mm.
Head and thorax very dark bluish green ; abdomen broad,
ATA Mr, T. D. A. Cockerell—Descriptions and
convex, smooth, and shining black; head wider than thorax,
facial quadrangle nearly square, but perhaps a little broader
than long; mandibles ferruginous in the middle; clypeus
sparsely punctured ; antennz dark, the flagellum red beneath
at apex only; facial foveze distinct, narrowing above,
reaching top of eye; occiput and cheeks with white hair;
tubercles fringed with white hair ; mesothorax convex (almost
gibbous), little hairy, rather shining, finely granular ; area of
metathorax not marked by peculiar sculpture; tegule dark
basally, with broadly hyaline margins. Wings clear hyaline ;
stigma large, very dark brown; nervures pale brown ; second
s.m. less than half length of first, receiving the recurrent
nervures very near its base and apex. Femora black, with
the knees more or less red; anterior tibize and tarsi clear
ferruginous ; middle and hind tibie greatly infuscated ; hind
tarsi dark red.
Hab. Launceston, Tasmania (J. J. Walker, 3855).
Euryglossa pernana, sp. n.
@ .—Length about 3} mm.
Black, the oval abdomen becoming a sort of dull pale
sepia basally, or rather more reddish on middle of first
segment; head ordinary, face flattened or even depressed,
the areas on each side of the clypeus shining silvery with
minute appressed hair; apical half of mandibles bright
ferruginous ; long hairs spring from beneath anterior margin
of clypeus ; facial quadrangle longer than broad; antennz
dark, the flagellum beneath a sort “ot dull yellowish brown ;
head and thorax dull, the punctures too minute to distinguish
with a lens; mesothorax with a complete median groove and
short parapsidal ones; metathorax truncate, the area merely
granular ; tegule brown. Wings dusky hyaline, nervures and
stigma piceous, the stigma rather small; second submarginal
cell higher than broad, the first r. n. meeting first t.-c. Legs
dark, the hind legs with slight reddish and brownish tints.
Hab. Fremantle, W. Australia (J. J. Walker, 1953).
Euryglossa neglectula, sp. n.
9 .—Length about 5} mm.
Shining black, with sparse whitish hair ; abdomen convex
and very broad ; mandibles with a dark red subapical spot ;
flagellum faintly reddish beneath; facial quadrangle a little
broader than long; front shining ; ; ¢clypeus with large
scattered punctures ; facial foveze present but shallow ; meso-
thorax moderately shining, with fine scattered punctures,
Records of Bees, 475
the parapsidal grooves distinct ; area of metathorax triangu-
lar, hardly distinguished, except by its duller surface ;
tubercles frimged with white tomentum; tegule extremely
dark brown. Wings clear hyaline, iridescent ; nervures and
stigma reddish brown, stigma large; second s.m. broader
than high, a little more than half length of first, receiving
the first r. n. a short distance from its base, and the second
almost at its apex; basal nervure strongly bent, only falling
a short distance short of transverse-medial. Legs piceous, the
tarsi ferruginous; abdomen with a very obscure purplish
lustre, sides of its apical part with long hairs.
Hab. Australia (no other particulars known) ; from the
I. Smith collection, accession number 79. 22.
Euryglossa reginé, sp. nv.
3 —Leugth about 6) mm.
Black, head and thorax very densely and minutely punc-
tured ; abdomen granular, with a subsericeous gloss, the hind
margins of the segments narrowly brownish; mandibles
reddish in middle ; flagellum dull ferruginous beneath ; tarsi
ferruginous, as also are the knees, the anterior tibize except a
dark cloud behind, and both ends of the other tibize ; head
broad, but facial quadrangle longer than broad; face with
a good deal of whitish hair; facial foveee linear; area of
metathorax triangular, without any marked peculiarity of
sculpture; tegule very dark brown. Wings dusky hyaline,
nervures and stigma sepia; secoud s.m. much broader than
high, receiving the first r. n. some distance from its base ;
second r.n. meeting second t.-c.; dorsal abdominal segments
overlapping ventral, the overlapping margins broadly whitish
hyaline.
Hab. Queensland (Gilbert Turner, Ridg. 11. 91, 703).
Euryglossa adelaide, sp. n.
? .—Length slightly over 6 mm.
Very close to H#. regine, but differing thus :—Wings
yellowish ; scape, funicle, and flagellum light yellowish
ferruginous beneath, and not more than dark brown above ;
tibize, tarsi, and apices of femora yellowish ferruginous, the
anterior and hind tibize with small and faint dusky clouds ;
tegule yellowish hyaline.
Hab, Adelaide, 8S. Australia ; accession number 59, 52.
It is possible that this is the female of /. regine, but in
view of the differences noted, and the widely different
localities, this seems improbable.
476 Mr. T. D. A. Cockerell—Desertptions and
Euryglossa rejecta, sp. n.
$ .—Length about 65 mm.
Head and thorax black, with long loose pubescence, more
or less fuscous on head and thorax above, otherwise dull
white; head very broad; eyes converging below ; clypeus
with very close, large punctures, the surface between them
shining ; maxillary palpi long and slender ; blade of maxilla
more than twice as long as bread, rounded at end ; mandibles
bidentate, entirely black; antennz black, scape with much
long hair; flagellum rather stout, pruinose, the last joint
with a shining area at the side of the apex; mesothorax
shining, but well punctured ; tegulee very dark brown. Wings
greyish hyaline, iridescent, stigma and nervures piceous ;
second s.m. nearly as long as first, and receiving the recurrent
nervures a moderate distance from its base and apex. Legs
shining black, with thin white hair, spurs creamy white.
Abdomen black, with the hind margins of the segments
broadly ferruginous ; the segments are elevated or gibbous
about the middle, the ferruginous margins thus appearing
to be abruptly depressed ; the apex terminates in a projection,
which might be called spoon-shaped but for its being trun-
cate; on each side of this is an elongate ferruginous Jamina,
and above it a pair of delicate lamelle placed edgeways ;
venter with the basal segment black or almost, the second
and third entirely light yellowish ferruginous, the others
strongly clouded with brown.
Hab. Perth, W. Australia (H. W. J. Turner).
When working on Euryglossa, I put this aside as not
belonging to the genus. I have concluded to leave it there
for the present, however, to be separated when more is known
about the group.
Stilpnosoma semisericea, sp. n.
? .— Length 10 mm.
Head black, large and quadrate ; clypeus very shiny, with
a few scattered punctures; labrum with a very prominent
shining tubercle; mandibles strongly bidentate, slightly
reddish about the middle; mouth-parts practically as in
Euryglossa; blade of maxilla subcircular, broadly truncate,
fringed with rather long hairs; first joint of labial palpi
stout and dark; supraclypeal arca with very distinct, but
small and rather sparse, punctures ; antenne entirely clear
ferruginous, scape long; vertex with a dull silky surface and
very minute punctures; cheeks strongly and closely punc-
tured; thorax black, the anterior part of prothorax, tubercles,
Records of Bees. A477
scutellum, postscutellum, and basal area of metathorax
ferruginous; mesothorax dullish, silky, with minute punctures,
like the vertex ; median and parapsidal incised lines distinct ;
legs red ; hind spur of hind tibia strongly serrate (it is serrate
also in S. levigatum); tegule shining yellowish ferruginous.
Wings yellowish, stigma and nervures ferruginous ; venation
nearly as in S. levigatum, except that the first r.n. meets
the first t.-c., and the upper outer angle of the second s.m.
is an acute angle (an obtuse one in /evigatum) ; the second
r. n. bends inwards, and, as in levigatum, joins the second
s.m. some distance from its end. Abdomen dull, with a sort
of silky surface ; first segment light ferruginous ; second to
fourth very dark brown, nearly black, with the hind margin
broadly ferruginous, the junction of the two colours on the
second and third wavy, with a double curve; fifth segment
ferruginous, black at extreme base; apical segment ferru-
ginous, pointed, with a very narrow pygidial plate ; venter
ferruginous, varied with dark brown.
Hab. Queensland (Gilbert Turner, Ridg. 11. 93, 683), and
another marked ‘‘ Australia” (F. Smith’s collection, 79. 22).
This insect is so unlike S. levigatum in colour and
sculpture, that I thought at first it must represent a distinct
genus; but the other characters are all essentially those of
Stilpnosoma.
Erratum (Ann. & Mag. Nat. Hist., June 1905, p. 528)—For
Melissodes parosete read Melissodes parosele.
University of Colorado,
Boulder, Colorado, U.S.A.,
July 9, 1905,
LIV.—Descriptions and Records of Bees —VI.
By T. D. A. Cocxrrett, University of Colorado.
Paracolletes moretonianus, sp. n.
é.—Length about 11 mm.
Black, with red legs and pale yellow and golden pubescence.
The general appearance of the insect closely simulates
Nomia semiaurea, Ckll., but it is easily known from that
species by the dull, very minutely and densely punctured
first abdominal segment, the second and following segments
478 Mr. T. D. A. Cockerell—Descriptions and
granular instead of strongly punctate, the triangular second
submarginal cell, &c.
Head large, nearly circular seen from in front; inner
orbits parallel; facial quadrangle somewhat longer than
broad; mandibles pale yellowish ferruginous with the
apex black ; face densely covered with yellow hair; scape
red ; flagellum black, its last joint strongly curved inwards,
forming a sort of stout hook; hair of thorax ochreous,
somewhat darker and redder above, that on postscutellum
long; mesothorax and scutellum_ excessively closely
punctured; area of metathorax triangular, smoothish and
shining, its margin minutely beaded, or that effect is pro-
duced by little cross-lines ; tegulz ordinary, light ferruginous.
Wings dusky; nervures and stigma ferruginous, rather dark ;
stigma very small; apex of marginal cell away from costa ;
b. n. meeting t. m. on the distad (apical) side ; second s.m.
triangular, narrowed nearly to a point above, receiving the
first r.n. just before its middle; third s.m. very oblique,
receiving the second r. n. at a moderate (variable) distance
from its end. Legs clear red, with abundant, but mostly
short, very pale yellowish hair ; coxz black, and hind femora
blackened behind ; hind femora somewhat incrassate, with a
posterior keel; hind tibia incrassate; basal joint of hind
tarsi bent. Abdomen black, with the hind margins of the
segments broadly ferruginous ; the whole dorsal surface, but
especially the hind margins (not, however, forming bands),
with a thin beautifully golden tomentum ; apical plate broad
and dark, truncate with rounded edges ; penultimate ventral
segment with a median longitudinal tomentose elevation.
Hab. Moreton Bay, Australia (F. Smith collection, 79. 22).
Another is labelled Queensland (F. Smith collection, 79. 22).
Paracolletes moretonianus is so like a Nomia that I at first
took it for one. In reality, it appears to be closely allied to
P. venustus (Smith), from which it differs by its larger size,
red femora, &c. It should probably be separated generically
from Paracolletes.
Paracolletes bimaculatus (Smith).
g .—Perth, W. Australia; 93-198, collected by H. W. J.
Turner.
The specimen has the tibie largely suffused with black.
The male genitalia of this species are rather striking, the
stipites and sagittz being very large. The former have the
basal half broad, the apical half narrow, like a finger, and
not hairy. The latter, taken together, present a sort of
Records of Bees. 479
broadly-pyriform outline, the inner apical corner of each
being provided with a short spine, as it appears from above,
or lamella, as seen from the side. There is nothing which
can be said to differ essentially from the type of genitalia
found in Colletes. The third submarginal cell is remarkably
large, and the wings are very hairy.
Paracolletes flavomaculatus, sp. n.
¢.—Length about 9 mm.
Slender, shining black ; eyes converging below; facial
quadrangle much longer than broad; face covered with very
hight yellowish hair, dense and with a silky lustre at sides ;
mandibles dark; clypeus with large punctures; antennz
ordinary, black, the flagellum faintly brown beneath ; cheeks
bearded with white hair; vertex and occiput, and also the
shining sparsely punctured mesothorax, with black hair ;
scutellum (except anterior middle) and middle of post-
scutellum covered densely with long, erect, bright, light
orange-fulvous hair, and tubercles covered with hair of the
same colour; metathorax and pleura with sparse dull white
hair, but there is a patch of black just under the wings;
area of metathorax with a slight transverse ridge; tegule
shining piceous. Wingsdusky hyaline; stigma fairly large,
dark reddish; nervures dark brown; b. n. falling a little
short of t. m.; first r. n. joming second s.m. very near its
base; second r. n. joining third s.m. a short distance from
its apex ; third s.m. having at least twice the area of second.
Legs slender, piceous, with rather long thin pale hair.
Abdomen narrow, shining black, punctate, the apical part
with black hair, but no hair-bands ; apical plate strongly
fringed with hair; at the base of the penultimate ventral
segment is a thick suberect fringe of hair, shortest in the
middle.
Hab. Australia (no other particulars known); from the
F. Smith collection ; 79. 22.
By the venation and the black hair on thorax, this is
nearest to P. argentifrons (Smith) ; but it is easily distin-
guished by the patch of fulvous-yellow hair on the scutellum.
Paracolletes cupreus semipurpureus, subsp. n.
? —Length about 94 mm.
Similar to P. cupreus (Sm.), but the caudal fimbria is
brown-black (not pale fulvous), the hair of the abdomen
beneath is white, and the stigma and nervures are amber-
480 Mr DPA, GockersI = Wee ae and
colour. Other characters are: abdomen brilliant crimson-
purple, with some golden tints; clypeus with large well-
separated punctures on a shining ground; mesothorax olive-
green, strongly punctured; second s.m. broad, receiving the
first r. un. about its middle; third s.m. not very much larger
than second, receiving the second r. n. very near its end ;
sides of metathoracic enclosure with numerous raised lines ;
hind spur of hind tibia pectinate with five long spines; all
the tarsi, hind tibie, parts of the other tibiz, and the knees
red.
Hab. Queensland (Gilbert Turner, Ridg. 11. 93, 863).
The typical P. cupreus came from Adelaide; it 1s very
likely that the Queensland insect should be regarded as a
distinct species, but it has all the principal characters of
cupreus, and for the present I treat it as a subspecies. Its
colours remind one of some of the Neotropical species of
Augochlora.
Paracolletes plumosellus, sp. n.
g .—Length 7} mm.
Expanse of wings about 16 mm.; head and thorax bluish
green, the mesothorax with large punctures, the surface
between the punctures minutely sculptured but shining ;
abdomen with a strong purple lustre, the hind margins of
the segments dark reddish; apical plate broadly rounded,
deep red with a hyaline margin; hair of head and thorax
abundant, strongly plumose, dull white with little yellowish
tint, that on the scutellum greyish; antennz rather long,
entirely dark, the third joint with a little protuberance
beneath ; tegule shining rufous. Wings hyaline, stigma and
nervures ferruginous; b.n. meeting t. m.; second s.m. little
narrowed above, receiving the first r.n. well before its
middle ; third s.m. receiving second r. n. very near its end ;
stigma rather large. Legs slender, very dark reddish, with
pale hair.
fab. “ New Holland, 44. 4.”
In all respects this is very similar to P. plumosus (Sm.),
described from a female. Were it not that the venation
ditlers somewhat, I should think plumosellus to be the male
of plumosus ; and as it is, 1 am not wholly without doubt.
Australian students must settle the matter by collecting the
sexes together.
Paracolletes ceruleotinctus, sp. n.
g .—Length about 9 mm.
Slender, the abdomen narrow; head and thorax green,
Records of Bees, 481
with tints of blue, with abundant long pale yellowish hair
(yellowest on face), but the scutellum and’ hind part of
mesothorax (except its hind border) with black hair; facial
quadrangle much longer than broad; mandibles dark,
rufescent apically ; antennz Jong, dark, the third and last
three joints more or less reddish beneath ; vertex and front
strongly punctured, with blue tints, a small shining smooth
space on each side of the ocelli; mesothorax shining olive-
green, with very strong punctures; area of metathorax
triangular, shining, without any transverse ridge or keel;
pleura yellow ish green, very closely and strongly punctured ;
tegule shining piceous. Wings clear, iridescent ; nervures
and stigma very dark red-brown; stigma rather large;
marginal cell long, narrowly obliquely truncate at apex ;
b. n. falling short of t. m.; second s.m. broad, especially
below, receiving the first r. n. well before its middle; third
s.m. receiving the second r.n.ashort distance before its
end. Legs metallic yellowish green ; the anterior tibize
yellowish ferruginous in front; all the claw-joints largely
red ; spurs dark. Abdomen shining ani strongly punctured,
blue-green, with splendid purple tints on the first two
segments ; hair of abdomen pale to a little beyond middle of
third segment, beyond that black or almost, the black hair
at apex quite abundant; apex ending in a broad fan-shaped
plate; last ventral segment shining, raised in the middle,
and having on each side a conspicuous tuft of hair.
Hab. Queenland (Gilbert Turner, “Seaf. 1,91; 4337’),
Two specimens.
A very distinct species, not closely allied to any yet
described ; best known by the narrow, briiliantly coloured
and strongly punctured abdomen.
Paracolletes amabilis (Smith).
Described from “ Australia,” no particular region being
mentioned. A female before me is labelled Queensland,
72,18. The venation differs a little from the type; the b. n.
falling a little short of the t.m., and the second yr. n.
entering the third s.m. a short distance before its end. The
abdomen is olive-green. The insect agrees so well with
amabilis in all its more obvious char acters, that I presume it
is not to be separated, notwithstanding the difference in
venation.
Paracolletes carinatulus, sp. n.
6 .—Length about 8 mm.
Rather slender ;_ head, thorax, and abdomen olive-green;
Ann. & Mag, N. Hist. Ser. 7. Vol. xvi. 32
482 Mr, T. D. A. Cockerell— Descriptions and
anterior and middle femora black, with the extreme apex
red; hind femora, and all the tibize and tarsi, rather dark
chestnut-red ; antenne dark, long, but the flagellum has a
hoary pruinosity, and is reddish beneath at apex; clypeus
olive-green, shining, with sparse punctures; mandibles
bidentate, shghtly reddish towards the apex ; mesothorax
shining and sparsely punctured on middle, but anteriorly
duller with large, closer punctures. Wings hyaline, faintly
dusky at apex ; stigma yellowish ferruginous, nervures dark
fuscous; second s.m., considerably narrowed above, re-
ceiving the first r. n. just before its middle; second r. n.
meeting third t.-c.; b. n. falling a little short of t. m.;
tegule shining dark ferruginous ; pubescence pale, with dark
hairs on vertex and scutellum (not very well seen, as the
insect has been in some liquid) ; area of metathorax with a
strong, sharp, transverse keel; ventral surface of abdomen
noimal.
Hab. Queensland (Gilbert Turner, Ridg. 11. 91, 704).
This is so much like P. amabilis that it might be taken for
its male, but for the sharp keel of the enclosure of meta-
thorax. This character allies it to P. carinatus (Sm.), but
the general coloration suffices to distinguish it from that
species.
Paracolletes viridicinctus, sp. n.
? .—Length a little over 8 mm.
Head and thorax black ; abdomen black with the depressed
lind margins of the segments broadly brassy green; hair
of thorax light grey, but that on the scutellums, mesothorax
(except in front), and vertex (but not occiput) mainly black ;
hair at apex of abdomen thick and black ; venter with three
long greyish-white fringes margining the segments, and a
fourth yellowish. Legs black, the hind tarsivery dark reddish,
with the first jot broadened and flattened; hair of legs mainly
pale, the hind femora above with a large crest of curled
white strongly plumose hair, but the knee capped with
black hair ; hind tibize with light hair except on the upper
surface, where is much black; hind spur pectinate with
many (7 or 8) slender spines; clypeus smooth and bare,
shining, with sparse distinct punctures; antenne black ;
front dull; mesothorax shining, but dull in front, scarcely
punctured ; area of metathorax rather shining but granu-
lar, with a median longitudinal groove, but no transverse
keel; tegule shining black. Wings yellowish, stigma and
nervures red-brown; b.n. meeting t. m.; second s.m. broad,
Records of Bees. ~ 483
receiving the first r. n. at or a little beyond its middle ; third
S.m. receiving second r. n. a short distance from its end.
Hab. Tasmania; 51. 153.
In some ways like P. versicolor (Sm.), but the colour of
the abdomen is different, and there are other differences.
It is also allied to P. chalyheatus (Erichson), and I have
wondered whether it could be a colour-variety of that species ;
but at present there is no reason for assuming this.
Paraco'letes spatulatus, sp. n.
¢ .—Length slightly over 8 mm.
Broad, with rather the shape of a female; head and
thorax black; abdomen dark bluish green, or rather
yellowish green toward the apex, the hind margins of the
segments not reddened. Legs black, the tibiz stained with
red beneath, and the hind tarsi reddened at extreme base ;
hair of legs pale, but on inner side of the tarsi orange-
fulvous. Mandibles strongly bidentate, reddened only at
apex; clypeus shining, with well-separated punctures ;
antenne dark, but flagellum greyish pruinose ; hair of head
and thorax light grey, but on vertex, middle of meso-
thorax, and scutellums there is a good deal of black ;
mesothorax and scutellum shining; area of metathorax
dull, without any distinct transverse keel; tegule shining
black. Wings clear, nervures and stigma dark red-brown ;
b. n. meeting t. m. ; second s.m. broad, receiving first r.n.
about its middle; third s.m. receiving second r. n. near its
end; end of abdomen with some sooty hair, and terminating
with a rounded ferruginous spatulate plate, near the base of
which, on each side, is a little tooth-like projection.
Hab. Blackheath, New South Wales, 1895 (W. W.
Froggatt, 183).
Allied to P. chalybeatus (Erichson).
Paracolletes providellus, sp. n.
$.—Length about 74 mm.
Abdomen not narrow ; head and thorax shining black ;
abdomen shining dark bluish green, with the hind margins
of the segments broadly reddish ; hair of head and thorax
greyish white, a little black or blackish on vertex and
scutellums ; clypeus shining, with well-separated punctures ;
antenne dark; middle of mesothorax scarcely punctured,
very shiny ; area of metathorax dull and granular, with no
sharp transverse keel. Legs black, with light hair, that on
inner side of tarsi yellowish ; tegule dark rufous. Wings
IDF
pac
484 Mr. T. D. A. Cockerell— Descriptions and
hyaline, slightly stained with reddish; stigma dark reddish,
nervures fuscous ; b. n. not quite meeting t. m. ; second s.m.
strongly narrowed above, and receiving the first r. n. about
its middle; third s.m. receiving the second r. n. a short
distance before its end. Hair at apex of abdomen all pale;
venter normal.
Hah. “ Australia” (no more precise locality known),
56. 94.
A second specimen (“ Australia,’ 66. 64) is a little smaller,
with much lighter (reddish amber) stigma and nervures, and
no dark hair on vertex or scutellums ; it is perhaps a distinct
species. By the reddened hind margins of the abdominal
segments, this resembles P. providus (Sm.) and P. frontalis
(Sm.). The dark mandibles &c. readily separate it from
frontalis ; froma providus it appears to be distinguished by
several details, but as that species is known only in the
female, it is perhaps not quite certain that providellus may
not be its male.
Paracolletes obscuripennis, sp. i.
3 —Length about 9 mm.
Black, the abdomen very faintly ochreous, somewhat as in
P. chalybeatus ; pubescence (somewhat spoiled from immer-
sion in a liquid) pale, dense and yellowish (perhaps partly
- stained) on face, but black on vertex and upper part of front,
and largely so on mesothorax and scutellum. Legs black with
pale hair, the tarsi slightly rufescent; tegule dark reddish,
clouded with fuscous. Wings rather light fuliginous
throughout, the stigma and nervures dark; stigma large ;
b. n. not quite meeting t. m., or rather meeting it a little on
one side; second s.m. broad (narrower, but not narrow,
above), receiving first r. n, about its middle; third s.m.
receiving second r, n. near its end. Mandibles dark, biden-
tate; antenne dark, faintly reddened at apex ; maxillary
palpi with the first four joints subequal, and the fifth shortest,
the last three more slender than the first three ; meso-
thorax shining, sparsely punctured ; area of metathorax large,
dullish, humped in the middle, but with no transverse keel ;
abdomen without hair-bands; the hair near the apex is
black ; venter normal.
_ Hab. Tasmania; no other particulars known.
This has the form and appearance of P. viridicinclus, but
it is known from that and from P. chalybeatus by its dark
wings. In P. viridicinctus the metathoracic area is short,
and the posterior lateral margin is a simple grooved line,
Records of Bees. 485
almost straight, but slightly curved inwards; in P. obscuri-
pennis the area is large, and the posterior lateral margin has
a conspicuously beaded appearance and is quite strongly
curved outwards.
Paracolletes colletellus, sp. u.
3d .—Length 8 mm.
Black, very pubescent, the face densely covered with light
yellow hair; vertex and occiput with yellow hair like the
face, but cheeks with abundant white hair, long below,
forming a large beard ; mandibles black basally, rufous
apically, the apical tooth long aud sharp, the inner very
small; flagellum clear red beneath, including the last joint ;
head broad, orbits converging below ; vertex rather dull;
hair of thorax ochreous above, lighter, but not a pure white,
below ; mesothorax shining, but rather closely punctured ;
area of metathorax somewhat shining, ridged transversely ;
tegule shining fulvous. Wings hyaline and iridescent,
stigma and nervures ferruginous ; b. n. falling a little short
of t. m.; second s.m. large and square, except for the fact
that the second t.-c. is bent inwards for its upper half ; first
r.n. joining second s.m. at its extreme base ; second Y. n.
joining third s.m. at its apex. Legs with pale yellowish hair ;
femora dark, reddened at the knees ; tibiz and tarsi clear
ferruginous, but all the tibie with a large black median
patch. Abdomen with the basal half or more of the segments
grey-tomentose, and the apical margins broadly yellowish
hyaline, the junction of the hyaline with the black reddened ;
ventral segments 2 to 5 with the hind margins apparently
deeply excavated or concave in the middle, but the excava-
tions are filled in by hyaline membrane, so that the actual
hind margins of the segments are straight.
Hab. Adelaide River, Australia (J. J. Walker, 5518,
5520) ; two, collected on the § Penguin’ Expedition.
There is some resemblance to P. punctatus (Sm.), known
only in the female, but it does not seem likely that this is its
male. P. colletellus has all the appearance of a small
Colletes, such as C. fodiens, Kirby.
Paracollotes semilautus, sp. 0.
é .—Length about 8 mm.
Black, hairy, the hair in general cinereous or whitish, but
the long hair of the face (abundant but not dense enough to
hide the surface) is mixed with black on the clypeus and
black at the sides, and there is black also on the front and
486 Mr. M. Burr— Notes on the Forficularia.
vertex aud on the scutellum; head large, subquadrate, with
broad cheeks, which are obtusely angled behind; facial
quadrangle much broader than long, the eyes nearly parallel ;
mandibles dark, the extreme apex reddened; clypeus with
many but not very strong punctures ; flagellum dark reddish
beneath; mesothorax hairy, but shining; scutellum very
shiny, sparsely punctate, not obscured by hair; area of
metathorax dullish, not transversely carinate, posterior lateral
margins curved outwards. Legs dark dull red, with hght
hair; tegule shining dark reddish. Wings clear, faintly
dusky at apex, nervures and stigma amber-colour; b. n.
meeting t. m., but not quite squarely; t. m. bent at its
lower end; second s.m. about twice as broad below as
above, and receiving the first r. n. about its middle; third
s.m. receiving second r.n. a short distance from its end.
Abdomen hairy, shining, the hind margins of the segments
rufescent ; the hair at apex more or less fuscous, but not
black.
Hab. “ Australia,” 58. 168.
Easily known by the very broad face, with black hair at
the sides.
LV.—WNotes on the Forficularia—IX. On new WSpccies,
with Synonymic Notes. By MALcoL”m Burr, B.A., F.L.S.,
ORES.
I AM at present engaged upon a general revision of
the Dermaptera (earwigs) of the world, and, through the
generosity of my friends and correspondents, am able to
examine a very large amount of material, including a high
proportion of the types of the described species. Lack of
time, however, will very greatly delay the completion of the
work, and therefore, in order to prevent the confusion likely
to occur through having a large number of novelties named
and described in manuscript, I shall publish the descriptions
of such new species as pass through my hands during the
period of preparation.
Our knowledge of this group of insects has made such
progress in recent years, and so many new forms have been
discovered and described, and many of the old genera con-
tained such heterogeneous groups, that a number of new
genera will be required ; it is not, however, advisable to erect
new genera except in comprehensive works, and so many of
Mr. M. Burr—WNotes on the Forficularia. 487
the new species described in these papers will be provisionally
placed in the existing genera to which they have the strongest
affinities.
Gonolubis Verhoef, sp. n.
Fusco-castaneus, nitidus: antenne ..... (13 segmenta restant) ;
segmento 3 longo, clavyato; 4 quam 3 dimidio breviori, valido,
ovato; 5 quam 3 breviori, ovato; 6 tertium equanti, ovato-
clavato ; ceteris longioribus, ovato-conicis: caput leve, tumidum,
margine postico recto: pronotum subquadratum, margiuibus
omnibus rectis, postice leviter dilatatum, lateribus reflexis ;
planum: abdomen glabrum, lewve, postice valde dilatatum ; seg-
mentum ultimum dorsale quam metanotum duplo latius, magnum,
leeve, latius quam longius, margine postico recto, integro, medio
sutura distincta instructum; lateribus carina _ instructis ;
segmentum penultimum dorsale lateribus carinatis, rugulosis,
postice reflexis, in modum generis <Ancistrogastris; segmentum
ultimum dorsale lateribus oblique truncatis, margine postico recto,
truncato: pygidium minimum, vix perspicuum, transversum :
forcipis bracchia basi valida, triquetra, prope basin margine
superiori dente forti apice obtuso armata ; dehinc attenuata, valde
incurva, bracchio dextro supra sinistrum fortius incurva, apice
decussata. od. Q ignota.
3 °
ong. corporis). aes 1s). >» » LO mm:
AP MMTORGL DISA Pe cats . Pig se
Patria. South Australia (type in Mus. Hope, ¢).
There is a second specimen, agreeing entirely in structure,
but with a thick pale pubescence which is wanting in the
larger specimen, doubtless through being rubbed off; the
length of the body of the smaller one is 7 mm.
This species almost exactly resembles G. Kirbyi, Burr, but
differs in the strong tubercular tooth above the base of the
forceps (unarmed in G. Airbyi) and the presence on the sides
of the antepenultimate dorsal segment of the keel which is
present on the last segment of both species; the sides of the
last two or three seements in this species are more strongly
hooked than in G. Kirby t.
T am pleased to ae it to Dr. Karl Verhoeff, in recog-
nition of his work upon the earwigs ; for though it is not
always possible to agree with his conclusions, the value of his
articles cannot be denied.
Labia tenuipes, sp. n.
Fusco-castanea, valde pubescens: antenne 15-segmentate, nigre ;
segmento 3 clongato, cylindrico, apice vix incrassato ; 4 eylindrico,
488 Mr. M. Burr—WNotes on the Forficularia.
incrassato, quam 3 dimidio breviori; 5 quam 4 sublongiori, apice
incrassato; ceteris elongato-conicis, apicalibus tenuioribus: caput
fusco-castaneum, globosum, suturis haud_ perspicuis, margine
postico recto; totum pilis longis pallidis spissis valde pubescens:
pronotum caput latitudine equans, longins quam latins, rectangn-
lare, margine antico recto, postico rotundato ; lateribus rectis, sat
late reflexis; nigrum: prozona tumida, sutura mediana distincta,
usque ad marginem posticum producta; hac parte tumida tantum
al marginem posticum evanescenti: elytra carinula humerali sub-
obsoleta usque ad marginem posticum elytri percurrenti, punctis
rugulosis parvis seriatim dispositis signata; rugulosa, apice
truncata, lateribus pilis longis pallidis obsita: ale longs, nigree,
rugulose : pedes graciles, longi; valde pubescentes, pilis longis,
pallidis ; femora longa, incrassata ; tibia tenues, femora «quantes,
valde pubescentes, superne deplanatee, sed haud sulcate ; tarsi
gracillimi, uniti tibias longitudine eequantes vel sublongiores ;
segmento primo valde elongato, subtus dense piloso; segmento
secundo minimo, simplici; segmento tertio elongato, quam primum
subbreviori, subtus vix pilosum: abdomen convexum, sat latum,
apicem versus paullo dilatatam, nigrum, minute granulosum,
lateribus pilis longis obsitum; plicis lateralibus parvis sed
distinctis ; segmentum ultimum dorsale magnum, rufescens, latius
quam longius, margine postico subarcuato, medio impresso,
margine ipso reflexo, utrinque tuberculo maguo ruguloso instruc-
tum; segmentum penultimum ventrale amplum, rotundatum,
margine postico medio paullo excavatum : pygidium brevissimum,
haud prominens, crassum, superne incrassatum, margine inferiori
incrassato, subtuberculato: forsipis bracchia basi remota,
elongata, subrecta, basi incrassata et triquetra, pilis brevibus
instructa, basi ipso leviter divergentia, deinde subrecta, elongata,
eracilia, attenuata, medio margine interno dento forti armata.
So. ignota.
O.:
LOD VCOEPOTISH..i-f 2 \- eieaty eCUOATTM,
4s. @LOFCIDIS (2 He: -layiees <i oD
99
Patria. Peru: Chanchamarjo, La Merced, at 3000 metres
(type, onc g, in my collection, taken by W. I’. H. Rosenberg).
This is a very distinct species, resembling no form known
to me: it is characterized by the thick and elongate fourth and
fifth segments of the antenna; by the very hairy body ; by
the granulose elytra and wings ; by the humeral keel of the
elytra, almost obsolete, its position marked by a row of small
tubercles running the entire length of the elytra’s edge; by
the rounded penultimate ventral segment, a little emarginate
posteriorly ; by the form of the pygidium and the long and
slender, slightly curved, almost straight forceps ; especially
by the long and slender feet and the extremely long and very
slender tarsi.
Mr. M. Burr—No¢tes on the Forficularia. 489
Tt will require a new genus for its reception, but its
affinities are with the genus Labia, though it does not
resemble any species known to me.
The unique specimen has the left branch of the forceps
missing, and the right branch appears to have been broken
off at the apex. Otherwise it is in good condition.
Chetospania confusa, sp. n.
Antenne (?) 13-segmentate, fusco-testaceze; caput, pronotum,
elytra, ale atra; ore pallido; pedes testacei, fusco-annulati ;
abdomen rufum; forceps rufo-fuscus. Pronotum margine antico
recto, postico rotundato, metazona angusta: abdomen plicis later-
alibus parvis ; segmentum ultimum dorsale fusco-rufum, magnum,
quadratum, margine postico medio sulculatum, supra insertionem
forcipis tuberculo obtuso utrinque instructo: forcipis bracchia ¢
basi triquetra, typica, subrecta, medio margine interno dentato ;
pygidium breve, latum, margine postico sinuato, angulis posticis
in lobos parvos acutos obliquos productis. 6.
3.
ones COEporis..:....cakse ee oO mom,
Pe VROLCUDIS ners ha erase aera ON” a9
Patiia. Mentawei, Sipora, Sereinu, v.-vi. 1894: taken by
Modigliani, one male (type in coll. mea).
The type of this species has long been in my collection,
labelled ‘Ch. Few, Borm.?” from de Bormans’s own collec-
fon, It Is distinct, however, from Ch, Fee, on account of the
short and broad pygidium of the male, which has the angles
produced into small, sharp tubercles or lobes; otherwise it
resembles Ch. Fee closely, in form and colour. The female
is not yet known.
Mecomera Kervilled, sp. n
Statura majore; colore fusco-rufo: antenn 17-segmentate ; segmento
primo valde incrassato; 3 modice elongato, apice incrassato; 4
quam 3 dimidio breviori, incrassato, ovato ; 5 quam 3 subbreviori,
valido, ovato ; 6 tertium equanti, incrassato, subcylindico ; ceteris
elongatis, cylindricis; brunne: caput leve, deplanatum, rufo-
testaceum, margine postico leviter emarginato: pronotum caput
latitudine equans, deplanatum, testaceum, fusco-variegatum ;
margine antico convexo, in collem producto; margine postico
subrotundato; Jateribus rectis; longius quam Jatius, postice
leviter dilatatum: elytra ampla, deplanata, levia, apice emar-
ginata, fusea: ale longe, leves, fuscee: pedes testaceil, breves,
compressi: tarsorum segmento primo quam tertium distincte
breviorl, secundo parvo, haud Jobato; tarsi dilatati, breves:
490 Mr. M. Burr—Noates on the Forficularia.
abdomen valde depressum, latum, rufum, lateribus infuscatis,
leeve, plicis lateralibus distinctis ; segmentum ultimum dorsale g
magnum, subquadratum, leve, latius quam longius, margine
postico medio impresso, utrinque tuberculato; angulis in tuber-
culum parvum productis; @Q margine postico angustius,
tuberculis obsoletis: pygidium ¢ breve, transversum, margine
postico exciso, angulis in puncta acuta productis; 2 brevissimum,
subglobosum, margine postico leviter emarginato : forcipis bracchia
3 valida, deplanata, lata, rufa, margine interno dente forti
bifido armata, dehinc crenulata, sensim attenuata, apice incurva ;
subcontigua, deplanata, subrecta, margine interno crenulata,
sensim incurva, apice decussata. do 2.
oe 2.
Pons Veorporis’ eee 14-5 mm, 13:5 mm.
Bf foreipis( ey arc ee Oe = a
Patria. Java (1 g,1 2, ex coll. Gadeau de Kerville).
Approaches M. Weisst, Burr, from Tonkin, but larger and
stouter; the coloration is different, and the forceps are shorter
and stouter and more strongly toothed.
I have much pleasure in dedicating this species to my
friend M. Gadeau de Kerville, who, with great generosity, has
placed his entire collection at my disposal.
Ancistrogaster Kervillei, sp. n.
Niger, nitidus, glaber: antenne nigra, 11-segmentate ; segmentis
elongatis, cylindricis; segmentis 3 et + subequantibus, 5
sublongiori: caput tumidum, nigrum, margine postico recto:
pronotum nigrum, lateribus anguste testaceum, caput latitudine
equanti, latius quam longius, margine antico truncato, postico
subconvexo; lateribus rectis ; vix tumidum, deplanatum, sutura
mediana subobsoleta: elytra brevia, latiora quam longiora ;
carina humerali distincta, in medio elytri evanescenti; trapezol-
dalia, margine interno quam externo breviori; margine postico
oblique truncato: ale nulle: pedes typice, longi; tabi superne
integres; tarsorum segmentum primum cetera unita longitudine
zquans: abdomen basi angustum, medio valde dilatatum, apice
angustatum, depressum, nigrum; segmentis 2 et 3 plicis
distinctis; g medio valde dilatatum, segmentis 2-6 lateribus in
tuberculum reflexum acuminatum recurvis ; segmentis 2 penulti-
mis angustioribus, inermibus; segmentum ultimum dorsale ¢
leve, transversum, medio puncto impresso, margine postico
incrassato, tuberculis 2 obtusis paullo elevatis instructum ;
angulis exterioribus posticis acutis: abdomen 2 minus dilatatum,
segmentis lateribus haud reflexis; segmentum ultimum dorsale
2 valdeangustatum,subinerme: pygidium ¢ brevissimum, conicum;
2 brevissimum, conicum: segmentum penultimum ventrale ¢ @
rotundatum; segmentum ultimum ventrale g margine postico medio
Mr. M. Burr—WNotes on the Forficularia. 491
emarginatum, bilobum, lobis rotundatis, fere totum obtectum,
angulis carinulatis; 2 obtectum: forcipis bracchia ¢ depressa, basi
sat incrassata, margine interno prope basin dente parvo armata;
in dimidio primo valde divergentia ; in dimidio apicali repente,
fere angulatim incurva, apice ipso sinuata et mucronata; ad
tertiam partem apicalem margine interno dento obtuso armata;
simplicia, recta, subcontigua, gracilia. ¢ 9.
3. Q.
ous corporis ..5..... 9°d-11*> mm: 10-11 mm.
ee LOL GUIS) 2p" eo ey ccs 3-35 ,, ra
Lat. max. abdominis.... 4°75-5 ,, Sloe
ite ee PL LOLCUDISN. 2 oa. 3 3-3'5
Patria. Venezuela: region of Merida (in coll. Gadeau de
Kerville et mea; typus in coll. mea).
This species I formerly confused with Ancistrogaster
ampennis, Borm., but on a comparison with the original type
of the latter I find it differs in several respects: the forceps
are strongly bowed outwards and then inwards and have but
a small tooth at the base on the inner margin, in A. impennis
there is a powerful tooth at that place and the branches are
gently incurved ; in the new species the extreme tips of the
forceps are strongly sinuate, as the anteapical tooth is obsolete,
but in A. ampennis the two hooks are both strongly developed ;
in A. impennis the elytra have the exterior keel continued
throughout their length, in A. Kervillei it dies out half-
way down the elytra ; in A. ¢mpennis the pronotum is almost
square, in A. Kervillei it is transverse: in the final revision
both these forms will probably require new genera. ,
I dedicate it to my friend M. Gadeau de Kerville, as a
shght token for his generosity in providing me with a large
amount of most interesting material, including three pairs in
good condition, of this new species.
Opisthocosmia Poultoni, sp. n.
Gracilis: antenne ?-segmentate, sat crasse; segmento 3 brevi,
conico ; 4 sublongiori, conico; 5 conico, etiam longiori; nigre
(13 segmenta restant): caput leve, rufum, margine postico recto ;
oculis parvis, nigris: pronotum planum, capite sublatius, margine
antico truncato, postico late rotundato, lateribus rectis; planum,
pallido-flavum, sublatius quam longius, lateribus vix reflexis:
elytra longa, angusta, humeris vix dilatatis, apice truncata,
fusco-castanea: ale longe, pallido-flave : pedes breves, testacei :
abdomen cylindricum, rufum, leve, plicis lateralibus distinctis ;
segmentum ultimum dorsale ¢ quadratum, fuscum, leve, apice
subangustatum, declive, margine postico subsinuato, angulis
oblique truncatis: pygidium brevissimum, transversum, integrum ;
492 Mr. M. Burr—WNotes on the Forficularia.
forcipis bracchia basi remota, gracilia, valde elongata, rotundata ;
margine interno prope basin leviter excavata; ad quartam
partem longitudinis dente parvo armata, dehinc teretia, rectissima,
gracilia, inermia, apice incurva. ¢. @ ignota.
3.
LonGscorponis. 2. osu... -- 12 mm,
so LOLCADIS eye oie tatere So ae
Patria, Batchian (taken by Wallace; type, 1 ¢, in Mus.
Hope, Oxford, “ Stevens, 1860”).
In spite of the age of the type, it is in very good condition,
except that the antenne are somewhat broken.
In form it resembles O. formosa, but differs in colour ; the
broad, flat pronotum is characteristic.
I am very pleased to dedicate it to Professor Poulton, who
has always placed the Hope Collection at my disposal with
the greatest courtesy.
Opisthocosmia formosa, sp. Nn.
Gracilis, elongata: antenne fusce; segmento primo valde
elongato, apice incrassato; 3 valde elongato, pallido, cylin-
drico; 4 eylindrico, quam 3 vix breviori; 5 tertium equanti,
cylindrico; (segmenta 6 restant): caput nigrum, tumidum,
margine postico subemarginato: pronotum capite angustius,
longius quam latius, subovatum, margine antico recto, postico late
rotundato, lateribus subrotundatis; nigrum: prozona tumida;
lateribus reflexis: elytra angusta, carina humerali obsoleta, costa
autem elytri distincte plicata; fusco-castanea, macula humerali
magna rufa ornata: ale longe, nigree, basi macula magna flava
ornate: pedes longi, graciles, testacel, genubus infuscatis :
abdomen gracile, fusco-rufum, leve; segmentum ultimum dorsale
attenuatum, longius quam latius, leve, margine postico sub-
rotundato, subreflexo et subincrassato: pygidium minimum,
globosum : forcipis bracchia basi remota, gracilia, rotundata, valde
elongata, subrecta, margine interno in dimidio basali crenulata ;
ad mediam longitudinem dente parvo subobsoleto armata, dehine
inermia, teretia; usque ad dentem medium subsinuata, ovalem
valde elongatam includentia; dehine subrecta, apice incurva. <6.
© ignota.
Gi
Long. corporis ...... eee so ny
atforcipis “sce Pu 5
Patria, Cameroons (type, one @, ex coll. Gadeau de
Kerville).
This graceful species is the first of the genus recorded from
Africa, if we except the wingless O. Micheli, Burr, from
Abyssinia and OQ. kova, Borm., from Madagascar, both of
Mr. M. Burr—Notes on the Forficularia. 493
which are very different in colour and of much stouter build.
It may be also known by the colour of the elytra and wings,
and the long, slender, almost unarmed forceps.
Anechura sokotrana, sp. n.
Corpus pilis longis pallidis obsitum, fusco-castaneum; antenne
testacee, segmentis gracilioribus; caput ferrugineum, suturis
transversalibus sat distinctis: pronotum quadratum, nigrum,
marginibus rectis, postico rotundato, totum subtumidum, mar-
ginibus reflexis, pallido-testaceis; elytra brevia, nigra, medio
macula magna pallida ornata, margine interno quam externo
multo brevius, margine postico sinuato, angulo interno apieali
rotundato: ale abortive: pedes longi, sat graciles, testacei,
femoribus prope apicem infuscatis: abdomen magnum, rufo-
nigrum, punctulatum, plicis lateralibus sat distinctis; seg-
mentum ultimum dg subquadratum, latius quam _longius,
margine postico superne tuberculis duobus, spiniformibus,
supra forceps spectantibus, armatum; 2 angustum, declive,
inerme: pygidium ¢ quadratum, globosum, lateribus convexis,
margine postico truncato, angulis spinulis armatis; 9 haud
perspicuum. Forcipis bracchia ¢ sat valida, basi ipso paullo
dilatata sed haud deplanata, margine interno basi ipso dente
forti armata, dehine extus ac intus incurva, aream ellipticam
elongatam includentia, inermia; Q recta, inermia, contigua;
forceps rufescens, apice infuscatus.
3. Or.
ORS CORPOLIS: (51s i</< 10's sie vs. « > LG mm, 10 mm.
Jaufp ROECIPIS) © nye spec vos Gres. Sites
Patria. Sokotra (¢ 2 in B.M.).
Anechura Fedtchenkoi, Burr, in Forbes’s Nat. Hist. Sokotra, p. 414
(1903).
The two specimens brought from Sokotra by the Forbes-
Grant Expedition were formerly identified by me as
A. Fedtchenkot (1. ¢.), but, now that I have been able to
examine a good series of that species, a further examination
shows that they are entirely distinct. The forceps are more
strongly bowed, forming an elongated ellipse; the pygidiuin
is different ; the last segment of the abdomen has two tubercles
in the middle, pointing directly backwards and parallel,
whereas in A. Medtchenkoi the tubercles are stronger, situated
at the corners, and point outwards.
Forficula canariensis, sp. n.
F, Lesnei proxime affinis: differt statura majore, robustiori, elytris
brevioribus, apice oblique truncatis, angulis rotundatis, margine
interno quam externo multo brevioribus, corpore minus pubescenti.
Ceteris cum /, Lesnei omnino congruet. @ ignota.
494 Mr. M. Burr—Notes on the Forficularia.
3.
ong. corporis, ....7.'../ Ld mm;
DMT ORCHDIS ons 5 5.0)a: soe” idse | tas
Patria. Teneriffe (¢ in B.M.).
Very closely allied to #. Lesned, agreeing exactly in the
form of the forceps, but somewhat larger and stouter, and the
elytra are trapezoidal instead of square, and almost shdw a
minute scutellum.
Forficula redempta, sp. n.
Statura mediocri: antenne segmento primo testacco: caput rufo-
nigrum, postice subtumidum, suturis obsoletis : pronotum fuscum,
lateribus testaceis, margine antico truncato, postico lateribusque
late rotundatis, sat supra elytra productum ; prozona tumida, a
metazona plana distincte sejuncta: elytra nigra, macula parva
humerali flava ornata, longa, apice truncata: ale long, pallido-
flavee, sutura infuscata: pedes testacei: abdomen nigrum, vix
punctulatum, plicis lateralibus distinctis; segmentum ultimum
dorsale ¢ magnum, subquadratum, paullo latius quam longius,
margine postico medio trituberculato, utrinque ad latera tuberculo
parvo instructum: pygidium haud perspicuum : forcipis bracchia
¢ fusco-rufa, parte dilatata brevi, quartam partem forcipis
attingenti, margine interno crenulato, hac parte deplanata dente
parvo superiori sursum spectanti terminata; dehine attenuata,
gracilia, inermia, arcuatim incurva. ¢. @ ignota.
oun
Long Corporis: jn-.-1,- le saieie, peel Cy
p mALOLOUDIS te toite eae vac Hed adueted as
Forficula smyrnensis, Burr, in Forbes’s Nat. Hist. Sokotra, p. 413,
pl. xxv. fig. 5 (d)} (1903).
Patria. Sokotra: Dahamieh, 8500 feet (g¢, unique, in
B.M.).
Resembles F. smyrnensts, with which it was formerly con-
fused by both M. de Bormans and myseif, but smaller, slighter,
darker : the flattened part of the forceps has a smooth tooth
pointing upwards, and the pronotum is almost circular, instead
of transverse oblong.
Forficula vidua, sp. n.
Haud nitida. Statura majore; colore nigro: antennze (?) 10-seg-
mentate ; segmentis tenuibus, 4 sat elongato, quam 3 vix breviori ;
rufo-nigre: caput nigrum, globosum, suturis vix perspicuis: pro-
notum sublatius quam longius, lateribus parallelis margine postico
rotundato; prozona vix tumida, medio suleulo impresso: elytra
lata, brevia, apice truncata, fusco-castanea, haud nitida: ale
breves, parum prominentes, fusco-castanez : pedes testacei : abdo-
men atrum, yix punctulatum, plicis distinctis ; segmentum ultimum
Mr. M. Burr—WNotes on the Forficularia. AQ
dorsale medio impressum et subtuberculatum: pygidium breve,
conicum, apice fissum: forcipis bracchia sat valida; margine
interno prope basin dilatata et deplanata, hac parte breyi vix
contigua, dente obtuso obsoleto terminata; dehine sensim
incurva, in tertia parte basali denticulo minuto: forceps ipse haud
deplanatus, margine solo interno laminato. <.
ale
Long? corporis’! ..¢ 3'4’..0855 0 12 mm.
Fe LORCIMISN? Oe ht aioe 8 45 ,,
Fatria. Guatemala: Mt. Oltrain (in B.M.).
SYNONYMIC NOTES.
The following descriptions and names appear to me to be
applied to the same species ; I have not been able to verify
all the types, but the quoted localities and the descriptions
leave very little doubt that the names are synonymous.
Anechura eoa, Semenov, Rev. russe d’Ent. 11. p. 100, fig. 2
(1902) = Forficula japonica, Borm. An. Soc. Esp. H. N. ix.
p- 512 (1880). As the name japonica in Forficuls is pre-
occupied by Haan, Rehn (Proc. U.S. Nat. Mus. xxvii. p. 540,
1904) suggests the new specific name athymia. As it falls
into my genus Odontopsalis (Trans. Ent. Soc. London, 1904,
p- 816), which will coincide with true Anechura, Scudder, the
correct name for this insect is Anechura athymia (Relhn).
Nesogaster Fruhstorfert, Verhoeff, Zool. Anzeig. no. 665,
p. 191 (1902) = Labia dolicha, Burr, Ann. & Mag. Nat. Hist.
(6) xx. p. 311 (1897); so that the correct name for this
species is Nesogaster dolichus (Burr).
Sparatta semifulva, Borm. Notes from the Leyden Mus.
vi. p. 183 (1884), and Sphingolabis furcifera, Borm. 1. ce.
p. 194, appear to be respectively the ¢ and the ¢ of thesame
insect. 1 have in my collection an insect which agrees
perfectly with the description of the first, and with an
unpublished drawing of it by de Bormans himself, and another
which agrees equally well with the latter and with an
unpublished coloured drawing of the same: but these two
insects are very obviously the sexes of one species; they
resemble in every particular of locality and coloration, were
taken together, and differ only in the form of the pygidium
and forceps; the “‘semifulva” specimens have all seven
visible abdominal segments (that is, are females), and the
496 Mr. M. Burr—WNotes on the Forficularia.
“ furcifera”’ specimens have always nine (that is, are males) :
it appears, therefore, that de Bormans was mesmerised by the
forceps and pygidium of “ semifulva,” which are unusually
developed for a female, and considered the specimens to be
males. The second tarsal segment, examined by a microscope,
shows practically no dilation; and as furcifera is the type of
Sphingolabis, it effectually removes that genus from the
neighbourhood of Forficula, and conclusively demonstrates
its independence from Apterygida, of which the type,
media, is totally different: unless, therefore, Sphingolabis is
shown to coincide with some other existing genus, it must be
revived for this species, and probably also for S. hawattensis,
Borm., and S. borneensis, Borm. It is wrongly revived by
Verhoeff for S. sansibarica and its allies, as by his very
definition of the new genus he excludes the typical species
of Sphinoglahis ; his genus certainly holds good, but will
require a new name. In the meantime, as the name
“ semifulva”’ occurs a few pages earlier than “ furcifera” in
the same paper, the former name must stand, and therefore,
until the true position of the genus Sphingolabis is satisfac-
torily determined, this insect must be known as Sphingolabis
semifulva, Borm.
Chelisoches vittatus, Burr, Ann. & Mag. Nat. Hist. (7)
xi. p. 274 (1903), is identical with Sphinoglabis variegata,
Kirby, Journ. Linn, Soc., Zool. xxii. p. 326 (1891), as I
have seen the types of both; but it seems that both these
names must fall before the name employed by Fabricius
(Ent. Syst. ii. p. 5, 1793), who described what seems certainly
to be the same insect under the name Forficula flavipennis.
As it is a true Chelisoches, it must be called Chelisoches
flavipennis (Fabr.).
Chelisoches Shelfordi, Burr, Ann. & Mag. Nat. Hist. (7) vi.
p. 96, pl. iv. fig. 4 (1900), and Ch. hercules, Burr, I. ¢.
p. 97, pl. iv. fig. 2, are respectively the 2 and ¢ of the
same species. ‘ihe former name occurs earlier and so must
stand. ‘lhe form of the forceps of Ch. Shelfordi is so
unusual fora @ that I allowed myself to assume it to bea @.
In several large Chelisochide the females look like males
(c.g. Ch. glaucopterus) , and probably Ch. dort, Borm., is the
3 of Ch. superbus, Dohrn.
Royal Societies’ Club,
” St. James’s Street, S.W.
July 1905.
On Hipposiderus diadema and tts closest Allies. 497
LVI.—On Hipposiderus diadema and its closest Allies.
By Knup ANDERSEN.
Hipposiderus diadema, Geoftroy.
Diagnosis (as compared with the three species described
below).—Anteorbital width* of skull between 8°5 and 10mm.;
upper tooth-row (exclusive of BOR 11°3-13°2 mm.
Ears moderate: width 24—27°5 mm.; forearm 75°5-91 mm.
Range.—The Indo- and Austro-Malayan Subregions,
exclusive of Betchian.
Hipposiderus diadema oceanitis, subsp. n.
Diagnosis.—Anteorbital width 8°5 ; upper teeth 11°3-12°1.
Forearm 78°8-79°2.
Details.—Of all the local representatives of the diadema
type, this form has the slenderest skull and the shortest
tooth-rows. But in both of these respects it is almost, or
completely, matched by the smallest individuals of HZ. d.
griseus, from the Philippines, from which it, however, differs
by the somewhat shorter forearm (78°8-79°2 mm., as against
83-88). The three specimens of oceanitis at my disposal
evidently do not show the extremes of individual variation
in size; the external dimensions of this form will, presum-
ably, prove to be very much as in H. d. pullatus from New
Guinea (forearm 75-81 mm.).
The third (external) supplementary leaflet is smaller than
in the other forms of diadema, sometimes so small that it
requires some attention not to be overlooked.
Type.— G ad. (in alcohol). Aola, Guadalecanar, Solomon
Islands. Collected by C. M. Woodford, Esq. Brit. Mus.
no. 88. 1. 5. 23.
Range.—Solomon Islands: Guadalcanar ; Fauro,
Hipposiderus diadema griseus, Meyen.
Diagnosis —Anteorbital width 8-6-9°2; upper teeth 12-
13:2. Forearm 83-88.
Range.—Philippine Islands: Luzon, Catanduanes, Guima-
ras, Leyte, Mindanao (Mt. Apo; Zamboanga).
* The width of the facial portion of the skull, across the front margin
of the orbits, immediately above the upper border of the infraorbital
foramina. —A]l the measurements recorded in the ‘‘ diagnoses” and in the
table, p. 507, are of full-grown individuals, For explanation of some of
the measurements, see Ann, & Mag. Nat. Hist. (7) xvi. (1905) p. 248,
footnote.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. an
498 Mr. K. Andersen on Hipposiderus diadema
Remarks.—8 adult and 5 young specimens *, from all the
above-named islands of the Philippine group, have been
examined. They all agree closely in cranial and external
characters. In the general size of the skull and teeth, as well
as in the anteorbital width, there is scarcely more than an
average difference between this and the foregoing race, but
the forearm is longer. Such at least is the case in all the
specimens just referred to. But one adult example, from
Zamboanga, collected by A. Everett, is quite extraordinarily
small, the forearm measuring only 77°5 mm., the tail 35°8
(in all the others 48-54). The gap between this specimen
and the whole of the other series (the young individuals —
not excepted) is so great that I can hardly believe it to be-
long to the same form. We find in the Philippines, in the
same island, Rhinolophus arcuatus and subrufus, two distinct
species differing in no important respect but size; in Java
Rh. minor and acuminatus, differmg much in the same way ;
in the Solomon Islands H. d. oceanitis and //. dinops,asmall
and a large representative of the same type of bat; it is
therefore not improbable that in the Philippines there occurs
a small and a large form of H. diadema. But to decide tl.is
more ample material is necessary.
Technical name.—The type of “ Rhinolophus griseus”
was from S. Matheo Cave, Luzon+. ‘The figure (half
natural size) gives a tolerably good representation of a
Hf, diadema; the dimensions, as derived from this figure, are
quite as in the Philippine series examined by me. ‘The fur
is stated to be “ ganz aschgrau,” which is certainly not the
case In any specimen or any race of diadema I have seen;
the ears are far too small, the supplementary leaflets not
indicated. Notwithstanding these discrepancies there can,
in my opinion, be no doubt as to the identification of
Meyen’s species.
Hipposiderus diadema pullatus, subsp. nu.
Diagnosis.— Anteorbital width 9°2-9°5; upper teeth 11:8-
12°5. Forearm 75°5-81.
Details—The anteorbital width is slightly larger than in
the foregoing forms, but there is no sharp line of separation,
the maximum of griseus being equal to minimum of pullatus.
* Five of these thirteen specimens were kindly sent me for inspection
by Mr. Gerrit 8. Miller, Washington.
t+ F. J. F. Meyen, “ Beitrige zur Zoologie, gesammelt auf einer Reise
um die Erde,” Nov. Act. Ac. Ces. Leop.-Car. vol. xvi. pt. 2 (1838)
p- 608, pl. xlvi. fig. iv—According to Peters (MB. Ak. Berlin, 1871,
p. 316), the type specimen has been lost.
and its closest Allies. AG9
The general size is as in the Solomon race, markedly smaller
than in griseus.
Type.— § ad. (in alcohol). Haveri, British New Guinea,
700 m., November 1893. Collected by Dr. Lamberto Loria.
Presented by the Genoa Museum. Brit. Mus. no. 97.8.7. 9.
Range.— British New Guinea: Haveri; Chad’s Bay.
Hipposiderus diadema vicarius, subsp. n.
Diagnosis.—Anteorbital width as in H. d. pullatus ; upper
teeth 12°1-13°2. Forearm 80°5-85:2.
Details.—Similar to H. d, pullatus, but on an average
larger. The lateral vertical ridges on the front surface of
the posterior leaf show some tendency to obliteration: in
three out of five spirit-specimens from Sarawak they might
by a hasty examination rather easily be overlooked.
Type.— ad. (in alcohol). Niah Cave, Sarawak. Col-
lected by A. Everett, Esq. Brit. Mus. no. 92. 9. 6. 238.
Range.—N. Borneo (Sarawak); S. Celebes (Kalao) ;
Sumatra.
Remarks.— Six specimens from Borneo, one from Celebes,
and one from Sumatra have been examined. The skull of
the Celebes individual is in all essential points like those
from Borneo (three skulls), only a trifle smaller, with some-
what smaller teeth; the nasal swellings as inflated as in
any Bornean skull, the anteorbital width as great as in the
largest of these latter. The Sumatra skull is slightly more
slender than any from Borneo, the teeth somewhat smaller,
the nasal swellings less inflated. These differences, very
slight as they are, may eventually, when larger series from
these islands are to hand, turn out to be quite individual.
Hipposiderus diadema, Geoffr., typicus.
Diagnosis.—Anteorbital width 9°5-9'8; upper teeth 12°3-
13. Forearm 84°5-91.
Details.—This form is most closely related to vicarius, but
as a rule larger, and with the facial portion of the skull
slightly broader. The size of the teeth is the same as in the
Bornean race. Only skins (six specimens) and skulls (five)
have been examined.
Range.—Java, Timor.
Technical name.—The species was originally described
from Timor*. ‘There is a skin from that island in the
* Geoflroy Saint-Hilaire, “ Sur un genre de Chauve-souris, sous le nom
de Rhinolophes,” Ann. Mus, d’Hist. nat. xx. (1813) pp. 263, 266, pl. vi.
o> o> Me
oo”
500 Mr. K. Andersen on Ilipposiderus diadema
British Museum, closely agreeing with Geoffroy’s figure and
description.
Horsfield’s Rh. nobilis*.—The cotypes + (two skins) of
Horsfield’s “ Rhinolophus nobilis,’ from Java, are in the
British Museum; both of them are young adults, 7. e.
evidently full-grown, but the epiphyses of the metacarpals
not ankylosed and the teeth unworn. The skull of one of
these cotypes, and the skulls of several other examples from
Java, are as inthe Timor bat. The forearm of five Java
skins measures 85-91, of the Timor skin 84°5. It may mean
(if it means anything) that, in a large series, Java specimens
will turn out to be on an average larger than Timor speci-
mens, in which case Horsfield’s name will, of course, have
to stand for the former race {.
Hipposiderus lankadiva, Kelaart.
Diagnosis.—Anteorbital width moderate; 9°2-9°8; teeth
very large: upper row 14-147. Ears moderate: width
about 23-24; size very large: forearm 88-94:°8.
Details.—This species differs from H. diadema (all forms)
in the following particulars :—
* Th. Horsfield, ‘ Zoological Researches in Java and the neighbouring
Islands ’ (London, 1824), no. 7, letterpress (unpaged) ; figs. L, N, O, P,Q
(head and dentition) on black-and-white plate (without number) ; coloured
plate [8] (whole fig.).
+ Nos. A and B in Horsfield’s Cat. Mamm. Mus. East-Ind. Comp.
(London, 1851) p. 55.
{ H. diadema also occurs in the Malay Peninsula. The skull of an
example (skin) in the British Museum, from Gunnong Pulai, Johore
(collected by W. Davison, presented by A. O. Hume), has the anteorbital
width 10 mm., a trifle larger than in the races described above; the size
of the teeth (upper row 13 min.) is quite as in an average form of diadema.,
I should have no hesitation in identifying this bat with Dobson’s
“ H, Masoni,” from Lower Burma, were it not for the reason that Dobson
described this latter as having only one (central) vertical ridge on the front
of the posterior leaf, whereas in the Johore specimen the lateral ridges
are well developed (the other character cf ‘ Masoni” mentioned by
Dobson, viz. a downward projecting process from the symphysis of
the mandible, looks so strange that I suppose it may be an individual
deformity in the only specimen known). The possibility is perhaps not
excluded that Dobson had before him a specimen in which (as I have
pointed out above to be sometimes the case in HZ. d. vicarius) the lateral
ridges were markedly reduced, therefore easily overlooked. The length
of the forearm of the Johore specimen and of the type of “ Masoni” is
almost exactly the same (in the former 86°8, in the latter 85), A safe
identification of the Malacca form would require more material both
from the Malay Peninsula (specimen “c” of ZZ. diadema in Dobson’s
Catalogue (p. 137), from Penang, is a H. armiger) and from the type
locality of “ Mason,”
and tts closest Allies. 501
The skull is very large and heavily built (see measure-
ments on p. 507), but the facial portion (anteorbital width)
comparatively narrow; nasal swellings well developed,
inflated, making the upper surface of the facial portion very
conspicuously convex*; portion between nasal swellings
and sagittal crest convex or flattened, not distinctly concave,
asin H. diadema; hinder part of orbits (temporal fossa)
markedly broader ; sagittal crest higher, more abruptly
descending in front; mandible much longer, the rami
higher. The teeth considerably larger.
The central projection of the posterior leaf is more
prominent than in any other form of the diadema type, the
upper border of the leaf, therefore, trilobate+, in so far
recalling the shape of the posterior leaf in H. armiger. In
H. diadema the central projection is but slightly prominent,
the upper border almost evenly convex, as “a segment of
the circumference of a circle” {. In one specimen of
H. lankadiva there is a minute fourth lateral leaflet, external
to the third ; I have seen a similar individual aberration in
two H. d. vicarius from Borneo.
Range.— Ceylon.
Technical name.—The type locality of H. lankadiva § is
Kandy, Ceylon ||. Two skins from Dr. Kelaart’s collection
are in the British Museum, one of them marked (in Kelaart’s
handwriting ?) “‘ H. lankadiva; Kelaart ; Kandy, 1850,” and
both of them quite as described above. But in his original
description of AH. lankadiva, Kelaart gives as length of the
forearm 3 inches (76°2 mm.), of the third digit 44+ inches
(108 mm.), measurements which, if they are correct (and
there is no reason to doubt them), would indicate a bat much
inferior in size to any “lankadiva” I have seen. There is
* In this respect H. lankadiva is perfectly in accordance with the
other bats described in the present paper; I emphasize this important
point to prevent confusion with H. armiger, a species to which
HI. lankadwva bears a certain superficial resemblance in the general outline
of the posterior leaf, but from which it differs widely in the shape of the
facial portion of the skull. See the ‘‘ General Remarks,” below.
+ The peculiar shape of the posterior leaf in specimens from Ceylon
was pointed out by Blyth in 1863 (Cat. Mamm. Mus. Asiat. Soe. p. 26) ;
it seems to have been overlooked by subsequent writers, who are
unanimeus in putting “ lankadiva” down as a synonym of “ diadema.”
The shape of the leaf is not always easy to ascertain in dried skins ; it is
quite pronounced in two spirit-specimens at my disposal.
{ Dobson, Cat. Chir. Brit. Mus. (1878) p. 137.
§ “ Lanka,” Iam informed on good authority, is a Hindu name of
Ceylon. Is “-diva” the Sanskrit “deva” (cognate with Zeds (Aeds),
ios, deus) ?
\| KE. I. Kelaart, ‘ Prodromus Faunce Ze lanicse * (Colombo, 1852), p. 19.
- 502 Mr. K. Andersen on Hipposiderus diadema
the possibility that two distinct forms of the diadema type, a
small and a large, occur in Ceylon, in which case the name
lankadiva belongs to the former. So long as this remains
uncertain, I prefer to use Kelaart’s name for the form here
under consideration.
Hipposiderus euotis, sp. n.
Diagnosis —Facial portion of skull broad: anteorbital
width 10°3-10°6; teeth moderate: upper row 13°2-1377.
Kars large: width 29-29°8*; size rather moderate: forearm
845-89'2.
Details —The skull of this species bears a certain resem-
blance to that of H/. lankadiva; it is of about the same size,
with very wide temporal fossz (large zygomatic width), and
with approximately the same shape and development of the
sagittal crest; but the facial portion is considerably broader,
the region between the nasal swellings and the sagittal
crest distinctly hollowed out, as in H. diadema, not convex
or flattened as in H. lankadiva. The teeth are markedly
smaller than in this latter.
The shape of the posterior leaf is as in //. diadema.
Type.—Young adult (apparently full-grown), skin. Bat-
chian. Collected by Dr. A. R. Wallace. Tomes Collection
(unregistered).
Hipposiderus dinops, sp. 0.
Diagnosis.—Facial portion of skull excessively broad:
anteorbital width 11; teeth very large: upper row 14°6. Ears
excessively large: width 32:5; size the extreme: forearm
96:5.
Details.—The skull is of the same size as in /7/. lankadiva,
or rather larger, but the facial portion is much broader,
the region behind the nasal swellings distinctly concave ; the
coronoid process of the mandible markedly higher and
broader. The premolars and molars are as in //. lankadiva,
but the upper canines much broader at base.
The lower leg is extraordinarily lengthened: 44 mm., as
against 35°8 in the largest HH. lankadiva. The tail, too, is
markedly longer. The third leaflet external to the horse-
shoe very small. The upper border of the posterior leaf as
in euotis and diadema.
Type.— ¢ ad. (in alcohol). Rubiana +, Solomon Islands.
Collected by C.M. Woodford, Esq. Brit. Mus. no. 88.1.5.22.
* Measurements from two dried skins, therefore probably a little too
small,
+ A minute islet very near the western coast of New Georgia (see
map in C. M. Woodford, ‘A Naturalist among the Head-hunters,’
London, 1890),
and its closest Allies. 503
Colour.
The style of colour is, broadly speaking, the same in all
the forms described above. ‘There is some difference in
details, and there seems to be a certain, though not very
large, amount of individual variation.
This style of colour, in its most pronounced form, might
be described as follows :—
Upperside of the body, anteriorly light-coloured, posteriorly
dark-coloured ; the two colours contrasting, and the line
of demarcation between them sharp; the contrast is simply
due to the fact that, in the hinder part of the upperside,
the dark hair-tips are sufficiently long to completely cover the
light bases of the hairs, in the front part very short or
altogether wanting, exposing the light ground-colour ; the
dark-coloured part of the back corresponds to what I have
called the “ horseshoe-patch” in Rhinolophus. Base of hairs
of upperside rather “ ecru-drab” with a silvery tinge. Two
round patches, whitish or yellowish, on each side of the body,
at the shoulder-region, the one behind the other, rather
close together. A longitudinal stripe, white or yellowish,
bordering each side of the back, along the plagiopatagium.
Underside a shade of “ wood-brown.”
Putting aside some old skins, the colours of which may
not be quite reliable, the chief differences between the species
and subspecies, so far as they can be made out from the
material to hand, seem to be these (all the skins described
are of adult individuals) :—
H. d. vicarius and typicus——Hinder part of back “‘ mars-
brown” washed with “russet.””? Patches and stripes well
marked, sometimes tinged with yellowish.
H. d. griseus——Not essentially different from vicarius ;
the shade of the brown colour of the hinder back varies
somewhat, being sometimes darker, sometimes more tinged
with russet.
FT. d. pullatus (one skin only).—Back very much darker,
approaching “ seal-brown”’?; no contrast between anterior
and posterior part of back. White patches and stripes very
strongly marked *.
Hi. d. oceanitis (one skin).—Essentially as pullatus, but
patches and stripes almost completely obsolete. | Underside
of body considerably darker.
H. lankadiva.—As I], d. vicarius, but hinder part of back
* This skin was obtained on July 10th; teeth somewhat worn; fur
unabraded. Four spirit-specimens obtained in Noyember seem to be of
the same colour.
504 Mr. Kk. Andersen on Hipposiderus diadema
rather dark “ Prout’s brown.” No patches; longitudinal
stripe but very slightly indicated.
H. euotis.—Essentially as H. d. vicarius.
I have seen no skin of //. dinops.
Synopsis of the Species *.
Anteorbital width 85-10; ears moderate : width 23-27°5.
Upper teeth 11:3-13:2; upper border of posterior leaf
forming an are of a circle: Indo-Austro-Malayan
Subregions, exclusive of Batchian ................ diadema.
Upper teeth 14-14°7; upper border of posterior leaf
trilobate: ‘Ceylon * <0. ics .ges ws poem ee ere lankadiva.
Anteorbital width 10°3-11 ; ears large: width 29-32°5.
Upper teeth 13:2-13:7; forearm 84:5-89:2; lower leg
SOBA splsabC MIAN cre Be ses oe ote ts cin aot We ieee euotis.
Upper teeth 14:6; forearm 96; lower leg 44: Solomons. dinops.
General Remarks.
Common characters.—All the bats reviewed above have
been called H. diad:ma. They have the following characters
in common :—
Nasal swellings always distinctly inflated, the upper surface
of the facial portion of the skull never quite flattened, as in
H. armiger and its allies. p* (of course) and p; completely
wanting. The cingula of p, and p, strongly in contact,
rather often a little overlapping each other, very rarely (in
two out of 33 skulls) separated by an extremely narrow
interspace; the tendency to overlapping, and the rarity of
the exceptions to the general rule, may be taken as evidences
that ps has been lost for many generations of ancestors
(compare the pronounced vacillation in this respect in those
highly-developed species of Rhinolophus which have lost ps).
p very small, and, with rare individual exceptions f, situated
in the outer angle formed by the canine and p’, 7. e. on the way
towards complete obliteration. The upper canine and p”, as a
general rule, very distinctly separated ; quite often, however,
the interspace is small or extremely small; very rarely the
cingula are completely in contact {. p. somewhat reduced
in size.
* On H. Masoni and nicobarensis, see the ‘ General Remarks,” below.
+ In one skull, out of 33, p? is almost quite in row; the exception is a
H. d. vicarius from Sumatra.
t The details, from an examination of 33 skulls, are these:—In 16
c and p* are rather widely separated ; in 14 slightly or very slightly ; in
one almost in contact (pwullatus); in one in contact on the one side
(griseus) ; in one in contact on either side (dinops). The variation is
quite individual, the extremes, or approximately the extremes, rather
often occurring in individuals of the same subspecies (griseus, f. 1.).
and its closest Allies. 50d
Third metacarpal longer than fourth, and this latter always
considerably longer than fifth *. Second phalanx of third
digit, with very slight variation, equal to the first phalanx +.
Distal phalanges of fourth and fifth digits much shorter
than the proximal phalanges +. Compare the wing-indices on
p. 507.
Three supplementary leaflets on each side, external to the
horseshoe; sometimes (/ankadiva, d. vicarius) with a slight
trace of a fourth, rarely with some reduction in the size of
the third (dinops, d. oceanitis). Posterior leaf, with very
rare individual exceptions, distinctly broader than the horse-
shoe. Its upper border forming an arc of a circle, rarely
(lankadiva), by a stronger development ‘of the median pro-
jection, distinctly trilobate. Three vertical ridges on the
front face of the posterior leaf; the lateral ridges sometimes
tending towards reduction (d. vicarius), sometimes completely
wanting (Mason, nicobarensis). No frontal pore.
They are all bats of considerable size, the forearm varying
from 66 (nicobarensis) to 96°5 mm. (dinops). ‘The sexes do
not differ appreciably in size, nor is there any other secondary
sexual differences.
Interrelations.—H. diadema is the least modified of the
species: the facial portion of the skull is not excessively
broadened, the teeth not very large, the ears moderate.
We have no means of deciding where this type of bat has
originated, and how it has spread over the large area now
occupied. But it is not unreasonable to suppose that the
Philippine race, with its comparatively slender face, comes
rather near to the ancestral species; and if it is true that
these same islands are also inhabited by a markedly smaller
form, we might with more confidence regard the Philippines
as having, not improbably, been situated within the centre of
dispersal. The equally slender face and rather small teeth
in the extreme eastern form, d. oceanitis from the Solomon
Islands, may, very likely, be due to slight degeneration of
the type, if they are not indications that in this remote and
isolated place some of the primitive characters of the type
have been more faithfully preserved. But the only safe
conclusion to be drawn from our examination of these bats
* Compare the genus Rhinolophus: third metacarpal always the
shortest ; in the more primitive species the fourth, in the more highly-
developed the fifth, the longest. +
} In primitive 2Atnolophi always lengthened, in more differentiated
species much lengthened.
{ In Rhinolophus longer than, or much longer than, the proximal
phalanges,
506 On Hipposiderus diadema and its closest Allies.
is that the Solomon Island and New Guinea forms are most
closely related to each other, likewise the Philippine and
the Celebes-Borneo-Sumatra forms, whereas the Java-Timor
form is a trifle more aberrant from what we may well suppose
to be the “ ancient style.”
In H. d. vicarius we found.a certain tendency to reduction
of the lateral ridges on the front of the posterior leaf. In
H. Masoni, from Lower Burma, they have quite disappeared
(provided Dobson’s description of this bat is correct).
H.. nicobarensis, in which the same is said to be the case,
would seem to be a pigmy form of this modification.
H. lankadiva is most closely related to H. diadema; the
facial width is as in‘an average form of that species, but the
skull is much more heavily built, the teeth markedly larger,
and the shape of the posterior leaf a little aberrant *.
In Batchian (and neighbouring islands?) we find the
diadema type modified into the large-eared and big-faced
H. euotis, but still the teeth have retained their usual size.
This form seems to lead to the very large, big-faced, large-
toothed, large-eared, and exceedingly long-legged H. dinops,
from the Solomon Islands +.
Thus the Solomon Islands, though being the extreme
eastern limit for the group dealt with in this paper, are
inhabited by two modifications of the same type of bat.
But it must be remembered that these two forms (H. d.
oceanitis and H. dinops) have been derived from two different
branches of the fundamental type, the former from the
“ diadema”’ branch, the latter, probably, from the “‘ ewotis ”’
branch, and that they therefore, very hkely, have spread to
these far eastern islands at different times and by different
ways.
* H. diadema has been recorded from the Central Provinces of India
(Dobson, Mon. Asiat. Chir. (1876) p. 200, nos. 293-296) ; I have seen no
specimens from the Indian Peninsula.—Dobson also registers an adult
female, in the collection of the Calcutta Museum, from Darjeeling
(2. ce. no. 292; see also J. Anderson, Cat. Mamm. Ind. Mus. Calcutta,
1881, p. 115). Is that a H. diadema (not a H. armiger) ?
+ The Solomon Islands are indeed peculiarly fertile in the production
of giant species: Mus imperator, Mus rev, Rana Guppy?, to which is now
added Hipposiderus dinops.
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508 Dr. A. Aleock—A Pevision
LVII.—A Revision of the “ Genus” Peneus, with Diagnoses
of some new Species and Varieties. By A. Aucock, M.B.,
ih DBRS:
CONTENTS.
I. Introduction.
If. Definition of the Maniple Penens.
ILI. Diagnoses of the constituent Genera and Tables of the Species of
the several Genera.
IV. Diagnoses of Nine new Forms.
I. InrRopucTION.
The “genus” Peneus (type P. monodon) was established
in the year 1798 by Fabricius for three species from the
“Indian Ocean,” one of which (P. planicornis), since its
antennular flagella are described as compressed, should,
perhaps, be translated to the genus Solenocera of Lucas
(Ann. Soc. Entom. France, 2 sér. vol. viii. 1850, p. 219).
Milne-Edwards (Hist. Nat. Crust. 11. 1837, pp. 411-418)
deals critically with eleven species which he assigns to
Peneus; but two of them have since been transferred to
Solenocera, a genus with which we are not now concerned.
In 1881 in these ‘ Annals’ (vol. viii. ser. 5, p. 169) Spence
Bate published, along with a preliminary notice of the
‘Challenger’ Peneidz, an account of the results of an
examination of Milne-Edwards’s types. In this paper the
“genus” Peneus is left imtact, except that certain forms
with long antennular flagella are referred to as Peneopsis, a
MS. name,of A. Milne-Edwards. Peneopsis was not properly
characterized, nor has its type (P. serraf‘a) ever, as far as I
can ascertain, been described; but if the two species—one
of which bore A. Milne-Edwards’s MS. label ‘ Peneopsis
ocularis ”—described under this name by Faxon (Mem. Mus.
Comp. Zool. Harvard, xviii. 1895, p. 187) may be taken as
typical of the genus, then Peneopsis differs from all Penez
and agrees with all Peneinz except Peneus in having two
arthrobranchiz on the penultimate pair of legs. So that
Peneopsis, like Solenocera, may be left out of consideration
in a review of the phratry or maniple Peneus.
In 1885 S. I. Smith (Proc. U.S. Nat. Mus. vin. p. 170)
proposed to restrict the name Peneus to those species in
which (1) the endopodite of the maxillules is elongate and
segmented, (2) the third maxillipeds have an epipodite, and
(3) the last thoracic somite carries a pleurobranch. To the
of the “Genus” Peneus. 509
species not thus characterized he applied the name Para-
peneus. In this paper the author objected to the action of
Miers and Kingsley in lumping his genus Xiphopeneus (1869)
with Peneus, and emphasized the distinctive characters of
Xiphopeneus.
In 1891 Wood-Mason, in these ‘ Annals’ (ser. 6, vol. viii.
p- 271), pointed out that certain Parapenei differed from the
type species (Parapeneus membranaceus, Risso,=P. longi-
rostris, Lucas) in not possessing the characteristic sutures of
the carapace and in having a filamentous vestige of an
anterior arthrobranch on the penultimate thoracic appendage.
To these forms he gave the name Metapeneus.
Wood- Mason also recognized that Peneus styliferus, Kdw.,
though it possesses the carapacial sutures which are a marked
feature of Parapeneus membranaceus, Risso, is unlike that
species in having curved lamellar exopodites on all the
thoracic legs. He therefore separated P. styliferus and gave
it the MS. name Parapeneopsis, probably because P. styliferus
happens to have the long antennular flagella which Spence
Bate had chosen as the distinctive mark of Peneopsis.
Unfortunately for the aptness of the name Parapeneopsis,
several of the species which must be transferred to this
section or genus have short antennular flagella.
In 1896 de Man (‘ Zoologischer Anzeiger,’ p. 111) published
a description of a new Peneid, which, from the peculiar size
and length of the first pair of chelipeds of the male, he made
the type of a distinct genus, Heteropeneus. 'The observations
of Nobili (Boll. Mus. Torino, 1903, no. 455) seem to show
that the difference between Heteropeneus and Peneus is,
perhaps, rather less than the difference between the latter
genus and Parapeneus, Metapeneus, &c.; so that Heteropeneus
should be taken into the maniple Peneus.
In 1901, in ‘A Catalogue of Indian Deep-sea Crustacea,’
p. 15, I suggested that P. curvirostris, Stimpson (= P. ancho-
ralis, Spence Bate), should be detached from the genus
Parapeneus, where it had been placed by other authors, and
should be made the type of a new subgenus or genus
LTrachypeneus.
In the present paper all the sections—genera or subgenera
—into which the old Fabrician genus has gradually become
split are tabulated and briefly defined, and an attempt. is
made to sift all the species that have been described under
the name Peneus and to distribute them in their proper
sections. It is but an attempt, because to allocate the
species with confidence requires exact information regarding
the grooves aud sutures of the carapace, the presence or
Die) Dr. A. Alcock—A Revision
absence of exopodites, the disposition of epipodites and
branchiz, the form of the endopodite of the maxillule, and
the armature of the telson; and these particulars are often
not to be found in descriptions.
Fortunately, however, [ have not been entirely dependent
on descriptions and figures, for when I was in England in
1897 I was allowed, through the kindness of Professor
Jeffrey Bell, to examine at my leisure the collection of Penet
in the British Museum, which includes the ‘ Challenger’
material determined by Spence Bate, the various species
described and identified by Miers, and Henderson’s Madras
specimens described in the ‘Transactions of the Linnean
Society’ for 1893: all these I went through and tabulated,
specimen by specimen, with an eye to a revision of the
genus. Moreover, in the Indian Museum I have had at my
disposal (1) a collection made about thirty years ago in the
Andamans by that discriminating carcinologist James Wood-
Mason; (2) miscellaneous donations and purchases from
India, China, and Japan; and, chief of all, (3) the many
hundreds of specimens trawled and dredged by the ‘ Investi-
gator’ off all the coasts and islands of British India, from
the Indus Delta to Mergui, during twenty-four years. A
very considerable part of the Indian Museum material had
been sorted and named by Wood-Mason before his death in
1893, and I must add that I have incorporated Wood-
Mason’s rough memoranda and used his MS. names (except
where they have been anticipated) in this paper.
I must also add that the contents of this paper refer
exclusively to the maniple Peneus.
Solenocera, Parasolenocera, Peneopsis, Philonicus, Haliporus,
and Artemisia are excluded, as they all have two arthro-
branchiz on the penultimate thoracic leg, and the first five
have the cervical groove deeply impressed on the tergum of
the carapace.
Funchalia is excluded for the present, as it has long,
sickle-shaped, cross-cutting mandibles.
II. Derinition oF THE Manipie Pevyevs.
Penevs, Fabr.
Peneus Fabricius, Entomol. Syst. Suppl. 1798, p. 408; Latreille, Hist.
Nat. Crust. vi. 1803, p. 246; Leach, Trans. Linn. Soc. xi. 1815,
pp: 336, 347, and Malacost. Podoephth. Brit., text of pl. xlii.; Des-
marest, Consid. Gén. Crust. 1825, p. 224; Milne- Edwards, Hist. Nat.
Crust. 11. 1837, p 411; De Haan, Faun. Japon., Crust. 1849, p. 188;
Dana, U.S. Expl. Exp., Crust. pt. i. 1852, p. 601 ; Bell, Brit. Stalk-
of the “Genus” Peneus. 511
eyed Crust. 1853, p. 317; Heller, Crust. siidl. Europ, 1863, p. 292 ;
Miers, P. Z. S. 1878, p. 298; Boas, Stud. ov. Decapod., Vid. Selsk.
Skr. 6 Rekke, nat. o. math. Afd. 1. 2, 1880, p. 165; Spence Bate,
Ann, & Mag. ‘Nat. Hist. (5) viii. 1881, p. 173, and ‘ Challenger ’
Macrura, 1888, p- 229; Haswell, Cat. Austral. Crust. 1882, p. 198;
S. I. Smith, Proc. U.S. Nat. Mus. viii. 1885, p. 170; Ortmann, in
Bronn’s Thierreich, Malacostraca, pp. 1118-1120; Holmes, Occas.
Papers Calif. Acad. Sci. vii. 1900, p. 217; Kishinouye, Journ.
Fisheries Bureau, Tokyo, viii. no. i (1900) ; Alcock, Cat. Indian
Deep-sea Crust. 1901, p. 13.
Rostrum well developed, laterally compressed. Carapace
with postantennular (antennal) and hepatic spines, sometimes
with a small postocular (orbital) tooth or spine, and some-
times with a spine (branchiostegal) at or near its antero-
inferior angles. The cervical groove is never impressed on the
tergum of the carapace. Abdomen long, with some of its
posterior somites compressed and their terga carinated.
Kyes large. Basal joimt of antennular peduncle hollowed
dorsally to lodge the eye; its outer edge terminates in a
spine, and from the proximal end of its inner edge there
springs a twisted setose scale (antennular scale); the
antennular flagella are cylindrical and tapering and may be
short or long, but are never as long as the body. Antennal
scale large and foliaceous, its outer edge is rigid and
terminates acutely; antennal flagellum very long. The
mandible has a jagged cutting-edge and a broad grinding
crown ; its palp (endopodite) is large and broadly foliaceous,
consisting of two segments, of which the anterior is very
much the larger. The endopodite of the maxillule (first
maxilla) may be long and 2-, 3-, or 4-jointed, or may be
without segmentation and truncated; that of the maxilla
(second) is short.
The endopodite of the first maxillipeds is slender and
5-jointed, that of the second and of the third consists of 7
segments. The exopodite of the second and third maxilli-
peds is very well developed, being curved, compressed,
stiffish, and made up, like the flagella of the antenne, of
numerous small joints. The third maxillipeds are long and
pediform. The first three pairs of legs are chelate, the first
pair usually being the shortest and the third pair usually the
longest. The last. two pairs of legs are monodactylous.
Exopodites are usually present on all or all but the last pair
of thoracic legs, but are sometimes altogether wanting.
No podobranchiz exist on any of “the true legs, and
only one arthrobranch—the posterior one—is present on the
penultimate legs.
The abdominal appendages are of moderate length, the
512 Dr. A. Aleock—A Revision
exopodite being longer than the endopodite. In the first
pair there are no endopodites, but in the male their place is
taken by a pair of more or less rigid, longitudinally pleated,
or convoluted plates, known as the “ petasma” or “andricum,”
which together form a tube or canal. In the second pair
the endopodite carries at its base in the male a fleshy
papilla.
According to Zittel, the first remains of Peneus, so far as
is known at present, appear in the Lithographic Slates of
Bavaria (Jurassic).
The forms included in the maniple Peneuws are found in
greatest abundance off the coasts of the Indo-Pacific, from
the Red Sea and east coast of Africa (as far as 33° 8.) east-
wards to Japan and Australia. Eastwards of this centre a
few species occur in the western meridians of the Pacific, up
to the shores of California and Panama; and westwards of
it three species are found in the Mediterranean, two of
which extend into the N. Atlantic (one of them ranging as
far north as St. George’s Channel), and about ten occur off
the Atlantic coasts of America, from New England, through
the Gulf of Mexico and Caribbean, doubtfully as far south
as the northern end of Patagonia.
Taking the distribution of the several genera of the phratry
or maniple (or subgenera of the genus) in order :—
Peneus (s. ¥.) has the widest range, being found all round
the globe, from the Gulf of Mexico, through the Atlantic
coasts of N. Africa, the Mediterranean, the Red Sea, and
the Indo-Pacific, to California and Panama.
Heteropeneus has been found only in the East Indian
Archipelago (Singapore and Japan).
Parapeneus occurs in the West Indies and off the Atlantic
coast of the U.S. America, in the Mediterranean and its
western approaches, and in Oriental seas from India to Fiji.
Metapeneus: this large genus is almost entirely Indo-
Pacific (Red Sea to Polynesia), two doubtful species being
found in the West Indies.
Parapeneopsis is confined to the Indo-Pacific, ranging
from India to China and Japan.
Xiphopeneus is confined to the Atlantic coasts of sub-
tropical and tropical America.
of the ‘Genus ” Peneus. Sip
Trachypeneus is found, on the one hand, in the West
Indies and neighbouring coasts of America, and, on the
other hand, in Oriental seas from India to Japan.
Atypopeneus is known with certainty only from the Bay of
Bengal, but it probably occurs also in the China Sea.
Key to the Genera of the Maniple Peneus.
I, Rostrum serrated both on its dorsal and on its
ventral edge ; a pleurobranch on the last tho-
racic somite (XIV.); exopodites on all or all
but the last pair of the thoracic legs.
My Hirsiepaie or ehenpedsishort..... 0.0... .53> Peneus (3. ¥.).
2. The first pair of chelipeds of the male are,
typically, stouter and vastly longer than the
second and third pairs ..............+++: Heteropeneus,
II. Rostrum serrated on its dorsal edge only.
1, A pleurobranch on somite XIII. but not on
somite XLV,
i, Exopodites on all or all but the last pair of
bie. CHorncie lees). 11/5. ia neice eapes cas oootorers Metapeneus.
ii. The thoracic legs have no exopodites...... Parapeneus (3. ¥.).
2. No pleurobranchiz on somites XIII. and XLV. ;
all the thoracic legs with exopodites.
i. Epipodites wanting from at least the last
three pairs of thoracic legs.............. Parapeneopsis.
ii, Epipodites absent from only the last two
pairs of legs.
a. Last two pairs of legs of normal form.
A. Antennular flagella short............ Trachypeneus.
B. Antennular flagella much longer than
UHE CATA NACE’ Vira sa Se Sons Coie sas e Atypopeneus.
b. Last two pairs of legs extremely long and
slender (flagelliform) ................ A itphopeneus.
IIL. DraGNosEs OF THE CONSTITUENT GENERA AND TABLES
OF THE SPECIES OF THE SEVERAL GENERA.
1. Penevs, Fabr. (sensu restricto).
Peneus, Fabr., Sidney I. Smith, Proc. U.S. Nat. Mus, viti. 1885, p. 170.
Type, P. caramote, Risso.
Rostrum toothed both dorsally and ventrally. Antero-
inferior angles of carapace not spiniform. Postantennular
sulcus of carapace defined by a dorsal as well as a ventral
ridge.
Antennular flagella short or of moderate length. Endo-
podite of maxillules (first maxille) elongate and distinctly
3-jointed. Exopodites present on all or all but the last pair
of the thoracic legs.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. d4
514 Dr. A. Aleock—A Revision
Epipodites present on all but the last two thoracic ap-
pendages; pleurobranchiz present on the six posterior
thoracic somites,
Andricum symmetrical, simple, pod-shaped ; it consists of
two lobes finely interlocking all along their anterior border,
and capable of loose apposition in more or less of their
posterior border, the opposed faces being concave.
The dactylus of the third maxillipeds often shows modifica-
tions of a secondary sexual nature in the adult male.
The branchial formula is as follows :—
Podo- Arthro- Pleuro-
Somite. branchiz. branchive. branchiz. Total.
MEE eae ep. 1 (small) i) We) Mepose
EADIE Petes ees ep.+1 2 2 |! ==.) heplees
LX: cs A ep. 2 1 = ep.td
eer ep. 2 i = ep.43
DI Se ea ep 2 1 = ep.43
7 AS eae ep 2 ] =) iepiniee
XIil. 1 k= 2
KR EVcdisacn bo = 1
Total .... Gep.41 2 6 = Gep.419
In addition to the Indian forms hereafter specified, I have
examined the following species:—P. caramote, P. japonicus,
P. australiensis, P. latisulcatus, P. brasiliensis, P. setifer,
P. stylirostris.
List of the Species of Puneus (sensu restricto).
I. Antennular flagella extremely short ; usually a postocular (as well as
a postantennular) spine and crest on the anterior part of the carapace.
1. Peneus carumote (Risso), Milne-Edwards, Hist. Nat. Crust. ii. p. 413
etsyn. (=P. trisuleatus, Leach, Tr. Linn. Soc. xi. 1815, p. 347.)—
Mediterranean, W. Africa, England (St. George’s Channel).
2. Peneus canaliculatus (Oliv.), Milne-Edwards, op. cit. p. 414. (= P. mar-
ginatus, Randall, Journ. Ac. Nat. Sci. Philad. 1839, p. 146, and
? P. plebejus, Hess, Arch. f. Naturges, xxxi. 1. 1865, p. 168, pl. vii.
fig. 19.)—Indo-Pacific from Red Sea and E. coast of Africa to
Sandwich Is.; also W. coast of Africa.
3. Peneus canaliculatus, var. australiensis, Spence Bate, ‘ Challenger’
Macrura, p. 248, pl. xxxii. fig. 3.
4. Peneus canaliculatus, var. japonicus, Spence Bate, op. cit. p. 245,
pl. xxxi., pl. xxxii. fig. 4, pl. xxxvii. fig. 2.
. Peneus brevirostris, Kingsley, Proc. Ac. Nat. Sci. Philad. (1878) 1879,
p- 98. (According to Miers doubtfully distinct from P. canalicu-
latus.)—W. coast of Nicaragua.
6. Peneus californiensis, Holmes, Occasional Papers Calif. Acad. Sci. vii.
1900, p. 218. (A doubtful species, originally identified by Holmes
with P. canaliculatus.)—California.
Or
=
‘
of the “Genus”? Peneus. 515
. Peneus latisulcatus, Kishinouye, Journ. Fisheries Bureau, Tokyo, viii.
i. 1900, p. 12, pl. ii. fig. 2. (Perhaps a form of P. canaliculatus.)—
Japan.
8. Peneus brasiliensis, Latreille, Milne-Edwards, op. cit. p. 414.—Both
9.
10.
LA
15.
14.
16.
16,
1 Wf
sides of Atlantic in warm latitudes.
II. Antennular flagella not so long as their peduncle; no postocular
spine and crest.
Peneus monodon, Fabr., Milne-Edwards, Hist. Nat. Crust. ii. p. 416.
(=P. esculentus, Haswell, P. L. 8. N.S.W. iv. 1879, p. 38; also
=P. ashiaka, Kishinouye, Journ. Fish. Bur. Tokyo, viii. i. 1900,
p- 14, pl. ii1.)—Indo-Pacifie from E. Africa at least to Japan and
-Australia,
Peneus carinatus, Dana, U.S. Expl. Exp., Crust. pt. i. p. 602, pl. xl.
fiz. 2. (Probably= P. monodon, Fabr.)—Singapore.
Peneus setiferus (L.), Milne-Edwards, op. cit. p. 414. (=P. fluviatilis,
Say, Journ. Acad. Nat. Sci. Philad. 1817, p. 235.)—Atlantic
coasts of America in warm latitudes.
(P. orbignyanus, Latr., is supposed by Milne-Hdwards (op. ett. p. 415)
to be identical with this species.)
Ill. Antennular flagella longer than their peduncle; no postocular
spine and crest.
. Peneus indicus, Milne-Kdwards, Hist. Nat. Crust. it. p. 415.—E. Africa
to the Malay Archipelago and perhaps beyond.
Peneus indicus, var. merguiensis, de Man, Journ, Linn. Soe., Zool. xxii.
1888, p. 287, pl. xviil. fig. 8, pl. xix. fig. 1; and Zool. Jahrb., Abth.
Syst. x. 1888, p. 680.—ALI coasts of India, also Java.
Peneus indicus, var, penicillatus, Wood-Mason, MS.—Coasts of India.
Peneus occidentalis, Streets, Proc. Acad, Nat. Sci. Philad. 1871, p. 243.
(Supposed by Miers to=¢ndicus, Edw.)—Panama.
Peneus stylirostris, Stimpson, Ann. Lyc. Nat. Hist. N. York, x. 1874,
p- 134. (Supposed by Miers to=zndiens, Edw.)—Panama.
Peneus balboe, Faxon, 1893, vide Mem. Mus. Comp. Zool. Harvard,
xviii. 1895, p. 181, pl. xlvii. figs. 1-1 e.—O#f Cocos Island (Panama),
770 fath.
IV. No exopodite on the last pair of thoracic appendages.
Peneus semisulcatus, De Haan, Faun. Japon., Crust. 1850, p. 191,
pl. xlvi. fig. 1.—E. Africa to Australia and Japan.
Peneus tahitensis, Heller, ‘ Novara’ Crust. 1865, pl. xi. fig. 2. (Seems
to be identical with P. semisuleatus, De H.)—Tahiti.
Peneus gracilirostris, Thallwitz, Abh. u. Ber. Zool. Mus. Dresden
(1890-91), 1892, no. 3, p. 3, fig.5. (Probably identical with P. semi-
sulcatus, De H.)—N, Celebes.
Peneus ceruleus, Stebbing, Marine Investig. S. Africa, Crust. pt. iii.
1905, p. 77, pls. xxi., xxi. b’s.—E. coast of 8. Africa, about 33° 8S.
2. HeETEROPENEvS, de Man.
Heteropeneus, de Man, Zool. Anzeiger, 1896, p. 111, and Zool. Jahrb.,
Syst. Abth. x. 1898, p. 684, pl. xxxviii. fig. 75; Nobili, Boll. Mus.
Torino, xviii. 1903, no. 455, p. 4.
Rostrum toothed both dorsally and ventrally. Antero-
inferior angles of carapace not spiniform. Postantennular
34%
D16 Dr. A. Aleock—A Revision
sulcus defined ventrally only, by the buttress of the post-
antennular (antennal) spine.
Antennular flagella short. The first pair of thoracic legs
may, in the adult male, be enormously elongate, especially
as to the propodite ; but in the female, and, as Nobili has
shown, in certain adult males, may be of the ordinary Peneus
form. Exopodites are present on all the thoracic legs.
According to Nobili, epipodites are present on all but the
last two thoracic appendages, and pleurobranchie on the six
posterior thoracic somites.
Andricum symmetrical, simple, much as in Peneus (s. r.).
According to Nobili, the branchial formula is the same as
that of Peneus (s. r.).
Only the following species is known :—
§ §}
Heteropeneus longimanus, de Man, loc. cit.; see also Nobili, loc. cit.—
Jaya Sea; Singapore.
3. Metarenreus, Wood- Mason.
Metapeneus, Wood-Mason, Ann. & Mag. Nat. Hist. (6) vill. 1891,
od
p. 271.
Type, M. affinis, Edw.
Rostrum toothed on its dorsal edge only. Antero-inferior
angles of carapace either rounded or spiniform. — Post-
antennular sulcus defined only ventrally by the buttress of
the postantennular (antennal) spine. No longitudinal or
transverse sutures cn the carapace.
Antennular flagella short or of moderate length. Endo-
podite of maxillules (first maxille) somewhat abbreviated,
two-jointed. Exopodites present on all or all but the last
pair of thoracic legs.
Epipodites absent from the third maxillipeds as well as
from the last two thoracic appendages. No pleurobranch on
the last thoracic somite.
Andricum complicated, symmetrical or asymmetrical: if
symmetrical its distal angles are more or less spout-like ; if
asymmetrical one lobe is either larger or longer than the
other, and both are split up into interleaved convoluted
lobules.
The third maxillipeds never exhibit secondary sexual
characters in the male, but the last pair of thoracic legs
sometimes do.
The branchial formula is : —
of the “Genus” Peneus. 517
Podo- Arthro- Pleuro-
Somite. branchie. branchiz. branchize. Total.
SYA Pelee tee ep. r = ep+r
AA eee ee ep.+1 2 - = ep.t3
| soe enone a 2 1 = 3
os aN cre ep. 2 1 = ep.43
asthe « ep. 2 it = ep.t3
PU eee ep. 2 1 = ep.+5
3.41) RR ae Ee 1 1 = 2
AOR ‘ = 0)
Total .... 5ep.41 ll+r 5 = dep.417+4+r
In addition, a small filamentous vestige of an anterior
arthrobranch is present on the penultimate thoracic somite
in all the species I have examined, which include, besides
the Canadian species, M. Joyneri, M. tenellus, M. Macleayi,
M. philippinensis, M. Richtersi, and M. Batei.
List of the Species of Mutarennvus, Wood-Mason.
I, No marginal subterninal articulating spines on the telson. Last pair
of thoracic leys without exopodite ; their merus, in the adult male, with
a notch and spine or tooth at its proximal end.
1. Metapeneus monoceros, Fabricius, Milne-Edwards, Hist. Nat. Crust.
i. p. 415.—E. Africa to Japan and Australia.
(Metapeneus incisipes, Spence Bate, ‘Challenger’ Macrura, p. 257,
pl. xxiv. fig. 2, seems to be identical with M. monoceros.)
2. Metapeneus affinis, Milne-Edwards, op. cit. p. 416.—Karachi to
Japan.
Ca planicornis, Fabricius, according to Milne-~-Edwards,
closely reseinbles M. affinis, but may possibly, since it has the
antennular flagella compressed, not be a Peneus at all.)
(Metapeneus mutatus, Lanchester, P. Z. 8. 1901, ii. p. 572, pl. xxxiv.
fig. 6, seems to be identical with M. affinis.)
3. Metapeneus Joyneri, Miers, Ann, & Mag. Nat. Hist. (5) v. 1880, p. 458,
pl. xv. figs. 8-10.—Japan.
4, Metapeneus Dobsoni, Miers, P. Z.S. 1878, p. 302, pl. xvii. fig. 2.—
India and Ceylon.
5, Metapeneus brevicornis, Milne-Edwards, op. cit. p. 417. (=. avi-
rostris, Dana, U.S. Expl. Exp., Crust. pt. i. p. 603, pl. xl. fig. 3.)—
Mauritius to Borneo.
(Metapeneus sp., Lanchester, P. Z.S. 1901, ii. p. 571, pl. xxxiv. fig. 7,
does not seem to differ essentially from M. brevicurnis, Edw.)
6. Metapeneus lysianassa, de Man, Journ. Linn. Soc., Zool. xxii. 1888,
p- 290, pl. xix. fig. 1—Orissa to Singapore.
The following species appear to belong to this group, having no movable
spines on the margin of the telson, though the condition of the last pair of
thoracic legs as regards eaopodite §c. is not on record :—
7. Metapeneus tenellus, Spence Bate, ‘Challenger’ Macrura, p. 270,.—
Japan,
8. Metapeneus crucifer, Ortmann, Zool. Jahrb, Syst. Abth. v. 1890,
p. 451, pl. xxxvi. figs. 5 a, b.—Japan.
ols
9.
Dr. A. Aleock—A Revision
Metapeneus Mastersii, Haswell, P. L. S. N.S.W. 1879, p. 42, and
Cat. Austral. Crust. p. 203. (Considered by de Man to be doubt-
fully synonymous with M. monoceros, Fabr.)—Australia.
I. a. Telson without marginal spines ; merus of last pair of thoracic legs
P
10,
il
16.
16.
19,
20.
of male without notch at base.
Metapeneus Deschampsi, Nobili, Boll. Mus. Zool. Torino, xvili. 1908,
no. 452, p. 2, fiz. 1—Pondichery and Mahé. This may possibly
be the non-adult form of M. monoceros, Fabr.
Metapeneus villosus, Guérin, in Voy. ‘ Coquille,’ vol. ii. Zool., Crust.
p. 36, and Icon. Régne Animal, pl. xx. fig. ].—Australia. May
perhaps belong here.
II. Apev of telson with 3 or 4 pairs of lateral marginal spines.
Metapeneus ensis, De Haan, Faun. Japon., Crust. p. 192, pl. xls
fig. 2.—Japan.
(Metapeneus intermedius, Kishinouye, Journ. Fish. Bureau, Tokyo,
viii. 1900, p. 21, is possibly the same as M., ensis (M. monoceros
ensis) of De Haan.)
This species, if my identification be correct, has no exopodite to the
last pair of thoracic legs,
Metapeneus Macleayi, Haswell, P. L. 8. N. S. Wales, iv. 1879, p. 40,
and,Cat. Austral. Crust. p. 201. (No exopodite to last pair of
thoracic legs; related to M. ensis.)—Australia.
. Metapeneus Stebbingi, Nobili, Bull. Mus. d’Hist. Nat. Paris, 1904,
p. 229. (Male with a notch and spine on merus of last pair of
lees; probably related to M. ensis.)—Red Sea, Suez.
. Metapeneus cognatus, Nobili, 1. c. (Belongs to M. ensis group.)—
Djibouti.
. Metapeneus philippinensis, Spence Bate, ‘Challenger’ Macrura,
p- 261, pl. xxxv. figs. 2,38. (All the thoracic legs with exopodites.)
—-Kast Indian Archipelago, 82-150 fathoms. : es
Metapeneus coniger, Wood-Mason, Ann. & Mag. Nat. Hist. (6) vill.
1891, p. 272. (All the thoracic legs with exopodites.)— Off coasts
of India, 68-250 fathoms.
Metapeneus andamanensis, Wood-Mason, t. c. p. 271, Variety of
M. coniger.—Andaman Sea, 100-244 fathoms; and off C. Comorin,
143 fathoms.
7. Metapeneus gracilis, Dana, U.S. Expl. Exp., Crust. pt.i. p. 606, pl. x1.
figs. 7a, b. (Probably belongs to M. philippinensis group.)—Sulu
Sea; Australia.
. Metapeneus Richtersii, Miers, Zool. H.M.S. ‘ Alert,’ p. 564, pl. lui.
fig. A. (Has exopodites on all the thoracic legs, and probably
belongs to the M. philippinensis group.)—Madagascar Seas.
Metapeneus commensalis, Borradaile, P. Z. S. 1898, p. 1001. (Prob-
ably belongs to the M. philippinensis group. )—Rotuma, 8. Pacifie.
Metapeneus stridulans, WWood-Mason, MS. (All the thoracic legs
have exopodites and the antennular flagella are extremely short.)
This species may be the same as M. akayebi, Rathbun, which, as
Miss Rathbun points out, is one of the several species confused by
Spence Bate with M. velutinus, Dana.—Indian Seas, 20-35 fathoms,
. Metapeneus akayeli, Rathbun, Proc. U.S, Nat. Mus. xxvi. 1902, p. 39.
(All the thoracic legs with exopcdites; antennular flagella very
short.)—Japan.
Some of the ‘ Challenger’ specimens identified by Spence Bate with
velutinus probably helong here, according to Miss Rathbun,
of the “Genus” Peneus. 519
22. -Metapeneus mogiensis, Rathbun, 1. c. (As stridu/ans and akayebi
as regards exopodites.)—Japan. ‘Two of Spence Bate’s specimens
of “ M. velutinus” presented to the Indian Museum are this
species.
23. Metapeneus Dalet, Rathbun, t.c. p. 40. (M. akayebi group.)—Japan.
24. ea aca acelivis, Rathbun, t. ep. 41. (MM. akayebt group.) —
apan,
25. Metapeneus consobrinus, Nobili, Bull. Mus. d’Hist. Nat. Paris, 1904,
p- 229. (M. akayebi group.)—Dyjibouti.
26. Bearae Vaillanti, Nobili, l.c. (M. akayebi group.)—Red Sea;
OuezZ.
27. Metapeneus perlarum, Vaillant, Bull. Mus. d’Hist. Nat. Paris, 1905,
p. 158. (M. akayebi group.) —Persian Gulf.
28. Metapeneus lameliatus, De Haan, Faun. Japon., Crust. p. 193, pl. xlvi.
figs. 4,5. (Probably M. akayebi group.)—Japan.
29, Metapeneus Batei, Miers, Zool. H.M.S. ‘ Alert,’ p. 296, pl. xxii. fig. D.
(Probably M. akayebi group.) —Australia.
The following species may possibly come into the M. akayebi alliance :—
30. ? Metapeneus velutinus, Dana, U.S. Expl. Exp., Crust. pt. 1. p. 604,
pl. xl. fig. 4.—Sandwich Islands.
31. ? Metapeneus palmensis, Haswell, P. L. S. N. S. Wales, 1879, p. 48,
and Cat. Austral. Crust. p. 204.—Australian Seas.
32. ? Metapeneus pubescens, Stimpson, Ann. Lyc. Nat. Hist. New York,
x. 1874, p. 133.—St. Thomas, W. Indies.
33. ? Metapeneus Goodei, S. I. Smith, Proc. U.S. Nat. Mus, viii, 1885,
p- 176.—Bermuda; Bay of Panama.
4, Pararensus, S. I. Smith (sensu restricto).
Parapeneus, 8. 1. Smith, Proc, U.S. Nat. Mus. viii, 1885, p. 170.
Type, P. membranaceus, Risso (=P. longirostris, Lucas).
Rostrum toothed dorsally only. Antero-inferior angles of
carapace usually with, sometimes without, a branchiostegal
spine. Postantennular sulcus defined only ventrally, by the
postantennular (antennal) spine.
A longitudinal suture is generally present on either side,
extending from the orbital to the posterior border of the
carapace, and also a vertical suture extending across the
branchiostegite at the level of the second pair of chelipeds.
Antennular flagella of moderate length. Endopodite of
maxillules (first maxilla) abbreviated, unsegmented, the
small terminal segment which is present in Metapeneus not
being differentiated. No exopodites on any of the thoracic
legs.
Epipodites absent from the third maxillipeds, as well as
from the last two thoracic appendages. No pleurobranch on
the last thoracic somite.
Andtricum symmetrical.
520 Dr. A. Aleock—A Revision
The third maxillipeds and last pair of thoracic legs are not
known to show any modifications in the male.
The branchial formula is the same as that of Metapeneus,
but the rudimentary arthrobranch of somite VII. (second
maxillipeds) seems to be absent, and there is no vestigial
filament, representing an anterior arthrobranch, on the
penultimate thoracic somite.
In addition to the Indian species I have examined specimens
of P. membranaceus and P. serratus.
List of the Species of Pararennus, S. I. Smith (sensu restricto).
I. Telson with a single pair of lateral marginal spines, which are fixed ;
carapace with a fine longitudinal fissure, extending on either side from
the orbital to the posterior border ; tip of adult andricum cut up into
spines or hooks and lobules or filaments.
1. Parapeneus membranaceus (Risso), Heller, Crust. siidl. Europa, p. 296,
pl. x. fig. 11. (=P. longirostris, Lucas, Hist. Nat. Anim. Artic. in
Expl. Sci. Algérie, Zool. i. pt. 1, p. 46, pl. iv. fig. 6,=P. Bocager,
Johnson, P. Z. 8. 1863, p. 255, and 1867, p. 900.)—Mediterranean
and its western approaches.
2. Parapeneus politus, S. 1. Smith, Proc. U.S. Nat. Mus. iii. 1881, p. 144.
—wN. Atlantic coast of U.S. America.
. Parapeneus megalops, 8. 1. Smith, Proc. U.S. Nat. Mus. viii. 1885,
p- 172.—W. Indies and neighbourhood, 155-288 fathoms.
. Parapeneus fissurus (Spence Bate), ‘Challenger’ Macrura, p. 263,
pl. xxxvi. fig. 1_—Bay of Bengal to S. Pacific, up to 115 fathoms.
. Parapeneus investigatoris, Alcock & Anderson, Ann. & Mag. Nat.
Hist. (7) 11. 1899, p. 279.—Gulf of Manar, Bay of Bengal, Anda-
man Sea, up to 419 fathoms.
6. Parapeneus americanus, Rathbun, Bull. U.S. Fish. Comm. for 1900
(1901) p. 102. (“ May prove to be a subspecies of P. cnvesti-
gatoris.” )—Porto Rico, 220-225 fathoms.
. Parapeneus longipes, Alcock, infra.—Coasts of India and Ceylon,
74-68 fathoms. Distinguished by the absence of a branchiostegal
spine.
oa —- w
Ly
If. Telson with 2 or 3 pairs of articulating marginal spinelets, in addition
to the fixed pair; carapace without longitudinal sutures; adult
petasma simple, open-pod-shaped.
8. Parapeneus rectacutus (Spence Bate), ‘Challenger’ Macrura, p. 266,
1. xxxvi. fig. 2—Fiji, Philippines, Andamans, Bay of Bengal,
95-418 fathoms.
9, Parapeneus serratus (Spence Bate), op. cit. p. 268, pl. xxxvuii. fig. 1.—
Fiji, 315 fathoms ; Torres Strait, 1400 fathoms,
5. ParareNneopsis, Wood-Mason, MS.
Parapeneopsis, Aleock, Cat. Indian Deep-sea Crust. 1901, p. 14.
Type, P. stylifera, Edw.
Rostrum toothed dorsally only. Antero-inferior angles of
carapace sharp or dentiform. Postantennular sulcus defined
of the Genus” Peneus, 591
ouly ventrally, by the buttress of the postantennular (an-
tennal) spine.
Carapace with longitudinal and transverse sutures as in
most Parapenei, but the longitudinal suture never reaches
the posterior border.
Antennular flagella either long or short. Endopodite of
maxillules (first maxille) short, unsegmented. Petaloid
exopodites are present on all the thoracic legs.
Epipodites absent from the third maxillipeds as well as
from the last three thoracic appendages, sometimes absent
from all the legs. No pleurobranchiz on the last two
thoracic somites.
Andricum symmetrical. The third maxillipeds and fifth
pair of legs are not known to be modified in the male.
The branchial formula is :—
Podo- Arthro- Pleuro-
Somite. branchiz. branchiz. branchie. Total.
Wie es ep. (OT) = ep.
AUN oe ep. +1 2 = ep.+3
Tres ee ar 2 i 3
Eres ats ep. or 0 2 1 = (ep.) +3
». Sree ep. or 0 2 = (ep.) +3
2. | leceaea Ae 2 a 3
2G Ura bic 1 hed 1
2.0 rae we Be oo. = 0
Total.. 4(or2)ep.41 11(+r?) 4 = Sep.t16(+r?)
The vestigial arthrobranch of somite VII. is often absent.
List of the Species of Parareneopsis, Wood- Mason.
I. Epipodites present on the second maxillipeds and first two pairs of legs.
Telson with lateral marginal spinelets.
1. Parapeneopsis stylifera (Kdw.), Milne-Edwards, Hist. Nat. Crust. ii,
p. 418.—Coasts of India.
Parapeneopsis stylifera, var. coromandelica, nov.—East coast of India,
Il. Epipodites present on the second maxillipeds and first two pairs of legs.
Telson with small lateral marginal spinelets only as an occasional
anomaly.
2. Purapeneopsis sculptilis (Heller), ‘ Novara’ Crust. p. 122, pl. xi. fig. 1.
—Seas of India and East Indian Archipelago.
Parapeneopsis sculptilis, var. Hardwickii, Miers, P. Z.S. 1878, p. 300,
pl. xvii. fig. 1.—Coasts of India.
Parapeneopsis sculptilis, var. cultrirostris, nov.—Coromandel coast.
3. Parapeneopsis cornuta (Kishinouye), Journ. Fish. Bur., Tokyo, viii.
1900, no. 1, p. 23.—Japan; India ?
522 Dr. A. Aleock—A Revision
(Parapeneopsis maxillipedo, Al\cock.—Bombay, Madras, Arakan
coast. This species may prove to be identical with P. cornuta,
Kish.)
. Parapeneopsis uncta, Alcock, sp. n.—Ganjam coast.
. Parapeneopsis nana, Alcock, sp. n.—Coromandel coast.
. Parapeneopsts gracillima, Nobili, Boll. Mus. Torino, xviii. no. 447,
1903, p. 4, fig. 1.— Borneo.
ones) a
Ill. Epipodite present on the second mavillipeds only.
. Parapeneopsis acclivirostris, Alcock, sp. n.—Persian Gulf, Coromandel
coast.
8. Parapeneopsis Hungerfordii, Alcock, sp. n.—Hongkong.
6. TrRacuyPENEvs, Alcock.
Trachypeneus, Alcock, Cat. Indian Deep-sea Crust. 1901, p. 15.
Type, 7. curvirostris, Stimpson (= T. anchoralis, Spence
Bate).
Rostrum toothed dorsally only. Antero-inferior angles of
carapace fairly well pronounced. Postantennular sulcus
defined only ventrally.
Carapace with longitudinal and transverse sutures, but
the former is very short, existing only in the orbital region.
Antennular flagella short. Endopodite of maxillules
short, unsegmented. Petaloid exopodites are present on all
the thoracic legs.
Epipodites absent from third maxillipeds, as well as from
the last two thoracic appendages. No pleurobranchiz on
the last two thoracic somites.
Andricum symmetrical. The third maxillipeds and last
thoracic legs are not known to be modified in the male.
The branchial formula is :—
Podo- Arthro- Pleuro-
Somite. branchie. branchie. branchiz. Total.
Ve ee ep. os = ep.
BV anes Sten ep. +1 2 = ep.+3
1D. ne neene = 2 1 = 3
2eaea ade ep 2 iL = ep.+3
DR Seoeton ep 2 1 = ep.+3
2A NGS Se ee ep 2 1 = ep.+3
KGS. BS, 1 +4 = 1
XUV epee ois = 0
Dotal. cr.) ep. 11 4 5ep +16
In addition to the Indian species J have examined specimens
of T. curvirostris (=T. anchoralis) and T. constrictus.
of the “Genus” Peneus. 523
List of the Species of TRacuyrenegus, Alcock.
1. Trachypeneus barbatus (De Haan), Faun. Japon., Crust. p. 192,
pl. xlvi. fig. 8.—Japan. _
(Trachypeneus curvirostris (Stimpson), Proc. Acad. Nat. Sci. Philad,
1860, p.44.—Japan. Probably identical with 7. barbatus =P. affinis
barbatus, De Haan.)
(Trachypeneus granulosus (Haswell), P. L. S. N.S. Wales, 1879,
p. 41, and Cat, Austral. Crust. p. 202.—N.E. Australia. Probably,
as other authors also have thought, identical with 7. curvirostris
= T. barbatus.)
(Trachypeneus anchoralis (Spence Bate), ‘Challenger’ Macrura,
p- 208, pl. xxxv. fig. 1.—Arafura Sea; Japan. Probably, as
other authors have thought, a synonym of 7. granulosus= T. curvi-
rostris &c.)
2. Trachypeneus asper, Alcock, sp. n.—Persian Gulf; Coromandel
coast; Andamans.
3. Trachypeneus constrictus (Stimpson), Ann. Lyc. Nat. Hist. N. Ycrk,
x. 1874, p. 135.—Atlantic coast of U.S. America; West Indies.
Trachypeneus constrictus, var. similis (S. 1. Smith), Proc. U.S. Nat.
Mus. viii. 1885, p. 175.— Atlantic coast of U.S. America; West
Indies,
7. XipHoreneus, S. I. Smith.
Xiphopeneus, 8. I. Smith, Amer. Journ. Sci xlviii. 1869, p. 390; see
also Proc. U.S. Nat. Mus. viii. 1885, p. 1888.
Type, X. Kroyeri, Heller.
Rostrum toothed dorsally only. Antero-inferior angles of
carapace subdentiform. Postantennular sulcus defined only
ventrally.
Carapace with longitudinal and transverse sutures, but the
former is not prolonged to the posterior border.
One of the antennular flagella is very long. Endopodite
of maxillule short, unsegmented. All the thoracic legs have
exopodites.
Epipodites are absent from the third maxillipeds and last
two pairs of thoracic legs. No pleurobranchize on the last
two thoracic somites.
The last two pairs of thoracic legs are of great length, their
three terminal joints forming a long slender flagellum.
The branchial formula is :—
Podo- Arthro- Pleuro-
Somite. branchiz. branchiz. branchie. Total.
VES eats ep. is = ep.+r
NE ree ep.+1 2 = ep. +3
ENC eee sh 2 ] = 3
2. ello ep. 2 1 — ep.+3
po a eae ep. 2 1 = ep.+3
OU rice ss ep. 2 1 = ep.+3
DN a See ae il a = ]
BXCEVEPy hatens's ay, rk ae = 0
Sep.+16+7r
II
Total .... Sep.Fl Ti+y 4
524 Dr. A. Aleock—A Revision
I have examined the specimens in the British Museum.
This genus contains the single species :—
Xiphopeneus Kroyert (Heller), SB. k. Akad. Wiss. Wien, 1862, xlv. B,
Abth. i. p. 425, pL ii. fig. 51. (=X. Harttii, Smith, Amer. Journ,
Sci. xlvili. 1869, p. 890, and Trans. Connect. Acad. ii. 1871, p. 28,
pl. i. fig. 1.)—Brazil; W. Indies.
8. ATYPOPENEUS, gen. nov.
Rostrum toothed dorsally only. Antero-inferior angles of
carapace rectangular. Postantennular sulcus not defined at
all. No longitudinal or transverse carapacial sutures.
Antennular flagella much longer than the carapace. Endo-
podite of maxillules slender. Filamentous exopodites on all
the thoracic legs.
Epipodites absent from third maxillipeds and last two
pairs of thoracic legs. No pleurobranchiz on the last two
thoracic somites.
The andricum is symmetrical.
Type, Peneus compressipes, Henderson, Trans. Linn. Soc.,
Zool. (2) v. 1893, p. 450, pl. xl. figs. 21, 22.
As Henderson surmised, this is a unique form and is
worthy of a separate position.
This genus or section includes, at present, only Henderson’s
Peneus compressipes ; but two of Stimpson’s species from
Hongkong, viz. P. podophthalmus and P. stenodactylus, may
possibly be assigned to it.
9. Sepis INcERTz.
1. Peneus villosus, Guérin, in Voy. ‘Coquille,’ vol. ii. Zool., Crust. p. 36;
and Icon. Regne Anim. pl. xx. fig. 1.—Australia. Probably a
Metapeneus, as the figure shows ventral edge of rostrum smooth,
foliaceous exopodites, and no carapacial sutures.
2. Peneus foliaceus, Risso, Hist. Nat. Eur. mérid. v. 1826, p. 69, pl. ii.
fig. 6.—Mediterranean. Probably a Parapeneus, as the figure
shows no exopodites to the thoracic legs,
3. Peneus tenuis, Dana, U.S. Expl. Exp., Crust. pt. i. p. 605, pl. xl. fig. 6.
—Atlantic coast of Patagonia. Position quite uncertain, except
that it does not belong to Penews (s. r.).
4, Peneus stenodactylus, Stimpson, Proc. Acad. Nat. Sci. Philad. 1860,
p. 43.—Hongkong. Appears to be very closely.related to P. com-
pressipes, Henderson, the type of the section Atypopeneus.
5. Peneus podophthalmus, Stimpson, loc. cit—Hongkong. Also seems to
resemble P. compressipes even more than does P. stenodactylus.
6. Peneus nove-guinee, Haswell, P. L. 8. N.S. Wales, 1879, p. 43; and
Cat. Austral. Crust. p. 203.—New Guinea. Differs from other
Penei in not haying a hepatic spine.
of the Genus”? Peneus. 525
IV. Dracnoses or NinE NEw Forms.
1. Peneus indicus, var. penicillatus, nov.
Peneus penicillatus, Wood-Mason, MS. (name only).
This variety is distinguished by the form of the external
maxillipeds of the male. In these appendages the carpus
and propodite are much shorter and coarser than they are in
indicus and merguiensis, but, on the other hand, the dactylus
is a long tapering joint from 15 to 2} times the length of the
propodite, and the pencil of hairs occupying the groove on
the inner side of the dactylus is of almost corresponding
length.
The rostral crest is not so high as that of indicus, var.
merguiensis, but is higher than that of typical indicus, and
this intermediate form of rostrum also characterizes females
taken in company with males of penicillatus, a fact which
prevents us, for the present, from regarding penicillatus as
merely an allomorphic male of P. indicus.
This form grows to a length of 6 inches. Numerous
specimens have been taken off the Orissa coast and at
Bombay, and a few from Karachi, the Gangetic Delta, and
Mergui (Marine Survey collection).
2. Parapeneus longipes, sp. n.
'P 9
Resembles P. fissurus, Sp. Bate, from which it is distin-
guished by the following characters :—
The rostrum in the female barely reaches the end of the
first joint of the antennular peduncle. There is no trace of
a branchiostegal spine at the antero-inferior angle of the
carapace.
The inner (longer) antennular flagellum is about as long
as its peduncle in the female, and a little longer in the male.
The external maxillipeds reach the tip of the antennal
scale and the last pair of thoracic legs reach a dactylus-
length beyond them.
The andricum is formed on the same plan, but in the
single male obtained it ends in a pair of (median) ragged
petaloid lobes and a pair of (lateral) stiff, curved, horn-like
filaments.
The thelycum consists of a broad longitudinally grooved
plate occupying all the space between the fifth pair of legs,
articulating with a horseshoe-shaped or concave and semi-
circular plate lying between the fourth pair of legs.
The female attains a length of 3} inches.
526 Dr, A. Alcock—A Revision
Thirty-one females and a male have been collected in the
following localities :—Off the Malabar coast (Mangalore) in
21-26 fathoms ; off the Orissa and Ganjam coasts in 20-68
fathoms ; off the Vizagapatam coast in 73-23 fathoms; and
off the south coast of Ceylon in 20 fathoms.
In the single male the rostrum is broken.
This species is distinguished from all those Indian Para-
penet which have longitudinal and transverse carapacial
sutures by the absence of a branchiostegal spine from the
antero-inferior angles of the carapace.
3. Metapeneus stridulans (Wood-Mason, MS.).
Crotalocaris stridulans, Wood-Mason, MS. (name only).
Of this species we possess about 130 specimens dredged
by the ‘ Investigator,’ and one, identified by Spence Bate as
Peneus velutinus, Dana, from the ‘ Challenger ’ collection.
The species may probably prove to be identical with the
Parapeneus akayebi of Miss Rathbun,
It is characterized by the presence in both sexes of a pair
of stridulating-organs, situated one on each side of the
carapace, near the middle of the posterior end of the
branchiostegite, in such a way that the anterior edge of the
first abdominal tergum can play over them. Hach organ
consists of a longitudinal row of vertically disposed mdges,
which vary in number, being usually five, seldom less than
five, and occasionally as many as twelve.
The species is a typical Metapeneus, and belongs to the
same group as MW. philippinensis and coniger.
Integument remarkably thick, hard, and tomentose.
Rostrum nearly straight, uptilted, sometimes reaching to
the end of the antennular peduncle, but often shorter, armed
dorsally with 5-8 teeth, the last of which is small and
isolated (epigastric). No postrostral crest. An indistinct
postocular denticle. Postantennular (antennal) spine very
strong, produced backwards as a strong convexity defining a
broad postantennular sulcus. Hepatic spine small ; cervical
groove present only in its neighbourhood. Branchial region
not defined except by a short crescentie crease below the
hepatic spine. Antero-lateral (antero-inferior) angles of
carapace spiniform., A pair of stridulating-organs as already
defined.
The second abdominal tergum is medially carinated in
less than its posterior half, the third in almost all its extent,
the carina in both cases being sulcate; the fourth to sixth
are all sharply carinated, the carina of the fourth and fifth
of the “Genus” Peueus. 527
being deeply cleft at its after end. The sixth abdominal
somite is nearly twice as long as the fifth, but shorter than
the telson. The telson is about as long as the inner caudal
swimmeret ; it ends very acutely and has near the apex four
pairs of large marginal spines, the last pair being fixed.
Eyes large. Antennular flagella equal, about one third
the length of their peduncle.
The third maxillipeds nearly reach the tip of the antennal
scale; the dactylus is slender and is not much shorter than
the propodite, with which it articulates end on; the basis
bears an antrorse spine. A similar spine is present on the
basis of the first two chelipeds and on the ischium of the
first.
In the female only there 1s a pair of sternal spines between
the second pair of chelipeds.
All the thoracic legs have longish exopodites.
The andricum, which is built in the same way as that of
M. coniger, is asymmetrical, the left lobe being the longer;
the outer lobule of the left lobe ends in a crown of stiffish
filaments.
The thelyecum consists of the following parts :—(1) between
the fifth pair of legs a transverse lamina more or less dis-
tinctly divided into three lobes, the outer of which (abutting
on the fifth legs) are bluntly dentiform ; (2) between the
fourth pair of legs a broad transverse plate, the anterior part
of which shows as a large, smooth, somewhat oval facet ;
(3) in the interval between the fourth and fifth legs a narrow
transverse bar, sinuous and shaped like a very open W.
Large females may attain a length of 3? inches.
The species has been taken in abundarce all along the
east coast of the Peninsula where any rocky patches occur,
from Orissa to Palk Strait, in 20-35 fathoms ; in the Gulf of
Martaban, in 20 fathoms; and at various places in the
Andamans, in 20 fathoms.
4. Parapeneopsis maxillipedo, sp. n. ?
Pan Peneus cornutus, Kishinouye, Journ. Fish. Bureau, Tokyo, viii.
1903, i. p. 23.
Compared with Parapeneopsis stylifera (Edw.) it presents
the following differences :—
The dorsal half of the carapace is tomentose. The rostrum
is recurved at tip, but otherwise is nearly horizontal ; it does
not reach the end of the antennular peduncle in either sex,
and it is armed dorsally with 8-10 teeth (not including the
isolated epigastric tooth), which form a very decided crest,
528 Dew: Milcock =A Revision
The postrostral carina, which is continued right up to the
posterior border of the carapace, is sharp and particularly
prominent.
The antero-inferior angles of the carapace are merely
dentiform; the sinuous subhepatic ridge (defining the
anterior part of the cervical groove) stops far short of the
antero-inferior angle of the carapace.
The longitudinal fissure of the carapace extends only a
short way behind the level of the hepatic spine.
The telson is short and has no marginal spinelets.
The antennular flagella, which are equal, are about two
thirds the length of their peduncle.
All the joints of the third maxillipeds except the dactylus
are abnormally broad, coarse, and tomentose.
The fifth pair of thoracic legs reach only to the middle of
the antennal scale. The basal spines of the chelipeds are
big, and in the female there is one on the third chelipeds as
well as on the first and second.
The andricum has on its outer edges the same basal wing-
like lobule as that of Parapeneopsis sculptilis, Heller, but the
organ ends in a pair of long calipers.
The thelycum is coucave and three-lobed ; the middle lobe
is very large and leaf-like ; the lateral lobes, lying between
the fifth pair of legs, are small and have between them in
the middle line a globous tubercle, behind which is a thick
tuft of long sete.
This species, which attains a length of 44 inches, has been
taken at Bombay, at Madras, and off the Arakan coast. It
may turn out to be Kishinouye’s Parapeneopsis cornuta, a
Japanese species which Nobili has also recorded from
Bombay.
5. Parapeneopsis uncta, sp. n.
Compared with Parapencopsis stylifera (Edw.) this species
exhibits the following points of difference :—
The integument is even thicker and denser, and, though
sparsely punctate, has a polished greasy appearance.
The rostrum has but a faint double curve, and, owing to
the absence of any styliform prolongation, reaches only to the
middle of the second joimt of the antennular peduncle in
both sexes.
The antero-inferior angles of the carapace are sharply
rectangular, not spiniform or dentiform.
As in Parapeneopsis sculptilis, Heller, the postrostral
carina is canaliculate and the sinuous subhepatic ridge
of the “Genus ’’ Peneus. 529
(defining the anterior portion of the cervical groove) does
not reach the antero-inferior angle of the carapace.
Dorsal of the hepatic spine the cervical groove is very
distinct up to the longitudinal suture of the carapace, this
being a quite distinctive feature.
The longitudinal suture of the carapace runs nearly to the
level of the transverse suture.
The sixth abdominal somite is as long as the telson ; the
telson is very short, not reaching the middle of the inner
caudal swimmeret, and is without marginal spinelets.
The antennular flagella are equal and a little shorter than
their peduncle.
The third maxillipeds nearly reach the middle of the
antennal scale and surpass the tips of the fifth thoracic legs.
The spine on the basis of the first chelipeds is very slender
and that on the second chelipeds is not distinguishable.
The andricum, like that of P. scu/ptilis, has on each outer
margin a wing-like basal lobule, beyond which it simply
tapers, to end in four hooks, of which the anterior pair are
small and are concealed by the posterior pair in the flexed
position of the organ.
The thelycum consists of a square plate between the fifth
pair of legs and a semicircular one between the fourth.
Ganjam coast, 10-11 fathoms.
The largest male is 3} inches long.
6. Parapeneopsis nana, sp. n.
This is a small species, the largest female being only
21 inches long, but from the finished form of the andricum
(petasma) I take the representatives of it to be adult.
Compared with Parapeneopsis stylifera (Edw.) it shows
the following differences :—
The rostrum, though in all respects similar, is shorter, so
that its styliform portion does not quite reach the end of the
antennular peduncle. The postrostral carina fades away at
the posterior fourth of the carapace.
The antero-inferior angle of the carapace is sharp-cut, but
not spiniform, and the sinuous subhepatic ridge detining the
anterior part of the cervical groove falls far short of it.
The telson is generally shorter than the sixth abdominal
somite ; its median dorsal groove is short and shallow, and it
has no lateral marginal spinelets,
The antennular flagella, which are equal, are about one
third the length of their peduncle. ‘The fifth pair of legs
reach only to the middle of the antennal scale.
Ann. & Mag. N. Hist. Ser. 7, Vol. xvi. BB,
530 Dr. A. Aleock—A Revision
The andricum is slender and ends in a pair of long,
straight, stiff filaments, which stand out at right angles to
the rest of the organ.
The thelyeum resembles that of Parapeneopsis sculptilis,
Heller, its most conspicuous part being a large leaf-shaped
median plate lying between the fourth pair of thoracic legs.
Numerous specimens have been taken off the Ganjam and
Orissa coasts up to 68 fathoms and at Madras.
7. Parapeneopsis acclivirostris, sp. n.
This small species is quite peculiar among Pene? in having
no epipodites on any of the thoracic legs and no isolated
epigastric tooth.
Compared with Parapencopsis stylifera (Edw.) it also
shows the following points of difference :—
The rostrum in the female, though recurved at tip, is
nearly straight and uptilted; it may reach or may fall short
ot the end of the antennular peduncle; it has seven teeth and
is not produced as a carina behind the gastric region.
The antero-inferior angle of the carapace is sharp-cut but
not spiniform, and the subhepatic ridge, defining the ante-
iror part of the cervical groove, stops far short of it and is
elegantiy ciliated.
'The longitudinal suture of the carapace reaches some way
behind the gastric region.
The sixth abdominal somite is as long as the telson, which
is short and has no marginal spinelets.
The antennular flagella are equal and are not much more
than half the length of their peduncle.
The external maxillipeds and fifth pair of legs reach nearly
to the middle of the antennal scale.
The thelycum consists of a concave semicircular plate
lying between the fourth pair of legs and a squarish plate
occupying the space between the fitth pair of legs.
This species is found in the Persian Gulf, in Palk Strait,
at Madras, and off the Vizagapatam and Ganjam coasts.
All the thirty-four specimens taken are females, and the
largest is only 2} inches long.
8, Parapeneopsis Hungerfordi, sp. n.
This species resembles P. acclivirostris in having only one
epipodite on each side, namely, the one borne by the second
maxillipeds.
of the “‘Genus”’ Peneus. 531
In other respects it resembles P. sculptilis, Heller, except
in the following particulars :—
The antero-inferior angles of the carapace are merely
rectangular, not dentiform, and the antennular flagella of
both sexes are shorter than their peduncle. In these
particulars it resembles P. uncta.
The andricum ends in a pair of large petaloid lobules, the
tips of which are incurved, and its basal lateral lobules have
the free edge deeply notched.
The thelycum is a narrow longitudinal plate, laterally
constricted or notched near the middle and almost cut in
two by a deep longitudinal furrow; it ends posteriorly in a
pair of knob-like facets.
A male and two females from Hongkong, presented by
Surgeon-General R. Hungerford.
In the females the rostrum is like that of many specimens
of P. sculptilis, but in the male it is short, barely reaching
the end of the second joint of the antennular peduncle, and
nearly straight; it is dangerous, however, to settle the
specific form of such a variable structure as the rostrum
from an examination of three specimens.
9. Trachypeneus asper, sp. 0.
Integument very thick and hard, tomentose, finely
scabrous.
Rostrum quite straight, uptilted, strongly soin the female,
not reaching the end of the second joint of the antenuular
peduncle ; dorsally it is armed with nine or ten teeth (not
including the isolated epigastric tooth), which form a crest.
Postrostral carina low, broad, and faint, nearly reaching the
posterior border of the carapace. An orbital spine. A
strong postantennular (antennal) spine, the buttress of
which reaches the hepatic fossa. A fine suture runs along
the anterior part of the floor of the shallow postantennular
sulcus. A similar transverse suture is seen on the branchio-
stegite at the level of the third pair of chelipeds. Hepatic
spme rather small. Antero-inferior angles of carapace
distinctly dentiform. A very indistinct subhepatic groove
(the anterior part of the cervical groove) runs from the base
of the postantennular buttress to the base of the hepatic
spine; dorsal of the hepatic spine the groove cannot be
distinguished.
On the second abdominal tergum there is a median
compressed tubercle. The third to sixth terga are very
30*
532 A Revision of the “Genus” Peneus.
sharply carinated. The fifth abdominal somite is about two
thirds the length of the sixth, the sixth is about as long as
the telson. The telson, which is dorsally grooved, is much
shorter than the inner caudal swimmeret, ends rather
abruptly, and has on either side a very obscure subterminal
marginal spinelet.
The antennular flagella of the male are about three fourths,
those of the female about two thirds the length of their
peduncle ; the lower flagellum is much the coarser. The
third maxillipeds are coarse, except the dactylus, and reach
into the anterior third of the antennal scale. The last pair
of thoracic legs reach at least a dactylus-length beyond the
tip of the antennal scale. A basal spine is present on the
first two pairs of chelipeds. All the thoracic legs have
petaloid exopodites.
The andricum is anchor-shaped.
The thelyecum consists of a transverse bar between the
fifth pair of legs and a coneave semicircular plate between
the fourth.
The female reaches a length of 33 inches. The colours in
life are pink, the abdominal carinz and thoracic appendages
being milk-white.
The species has been taken in the Persian Gulf, off the
Vizagapatam and Ganjam coasts in depths of 20-85 fathoms,
and off the Andamans in 60 fathoms.
It differs from 7. curvirostris, Stimpson (= T. anchoralis,
Spence Bate), of which we have both ‘ Challenger’ specimens
from Japan and other specimens from Hongkong, in the
following particulars :—
The rostrum is quite straight and has more teeth, and the
postrostral carina is much fainter.
The antero-inferior angles of the carapace are sharper aud
the anterior part of the cervical groove is much less distinct.
The antennules are shorter both in their peduncle and in
their flagella, and the fifth pair of legs are louger.
Though the andricum is similar, the thelycum is a good
deal different, specimens of the same size being compared.
On new Species of Noctuide. 533
LVIII.— Descriptions of new Species of Noctuide in the British
Museum. By Sir Georce F, Hampson, Bart., F.Z.S., &e.
[Continued from p. 386.]
STICTOPTERIN #.
Odontodes metamelena, sp. n.
?. Head and thorax reddish brown mixed with grey and
fuscous ; pectus grey ; tarsi black with pale rings; abdomen
fuscous ’ slightly mixed with brown and grey, the ventral
surface grey irrorated with fuscons. Fore wing suffused with
greenish grey ; subbasal line double, the inner line indistinct,
rufous, curved from costa to submedian fold; antemedial line
double, the outer line indistinct, irregularly waved ; orbicular
represented by a grey point defined by black; a sinuous
rufous medial line, oblique from costa to submedian fold,
then inwardly oblique ; reniform well-developed, with rather
irregular brown outline; postmedial line double, dentate,
excurved from costa to vein 4, then incurved; subterminal
line indistinct, pale, irregularly waved, defined by rufous on
inner side, below vein 4 incurved and with broad rufous
band before it extending to postmedial line ; a terminal series
of brown lunules ; a fine pale line at base of cilia. Hind
wing deep black-brown with a slight blue gloss; underside
with the inner area rather paler, traces of a curved medial
line.
Hab. 8.W. New Guinea, Kapaur (Doherty), 1 2 type.
Exp. 50 mm.
Stictoptera crinopis, sp. 0.
9. Head, thorax, and abdomen brown mixed with grey;
frons deep black ; tegulz ochreous at base bounded by a deep
black line, then greenish grey ; metathorax with shght black
erest; pectus whitish ; legs very long, the tarsi fuscous with
pale rings; abdomen with the slight dorsal crests dark.
Fore wing grey suffused with brown and irrorated with black,
the basal, inner, postmedial, and apical areas darker, the
discal area paler ; subbasal line double, oblique, black, from
costa to vein 1; antemedial line double, curved, minutely
waved ; orbicular rather u-shaped ; reniform concave towards
base and with brown central line forming a spot at lower outer
side ; a black streak on vein 2 from cell to the postmedial line,
534 Sir G. F. Hampson on
which is double, minutely dentate, excurved from costa to
vein 4, then incurved, with a rather dentate black mark
beyond it below costa; subterminal line only defined by the
area beyond it being paler except at apex, angled outwards
at vein 7 and excurved at middle; a terminal series of shght
dark lunules. Hind wing with the interspaces of basal half
pale, the veins and terminal half fuscous ; the underside with
dark discoidal lunule showing through to upperside.
Hab. Asuantit, Obuassi (/ergmann), 1 2 type. Hap.
36 mm.
Stictoptera hemithyris, sp. 0.
Head, thorax, and abdomen fuscous brown mixed with some
grey; pectus mostly grey ; abdomen with the dorsal crests
dark, the ventral surface whitish. Fore wing dark reddish
brown slightly tinged with grey, the basal area rather darker,
the discal area paler; a dark elliptical subbasal patch
defined by brown from costa to median nervure ; a slightly
waved antemedial line with striga of raised black scales
beyond it from subcostal nervure to submedian fold; a
medial line, oblique from costa to submedian fold ; reniform
moderate, slightly defined by black ; postmedial line double,
minutely waved, formed of striz of raised black scales, bent
outwards below costa and incurved below vein 3; a similar
single line beyond it defined by ochreous on inner side; a
fine pale subterminal line, dentate at veins 6, 7 and incurved
below vein 3; a terminal series of black striae. Hind wing
with the interspaces of basal half hyaline, the veins and
terminal half fuscous; cilia white at tips; the underside
greyer.
Hab. Punsas, Manpuri, 1 ¢; Bomspay, 1 9; Manras,
Cuddapah (Campbell), 1 2 type. Hap. 24-26 mm.
Stictoptera pheobasis, sp. u.
?. Head and thorax fuscous brown, mixed with some
grey and black ; pectus and femora whitish; tarsi fuscous
ringed with white; abdomen fuscous with slight whitish
segmental lines, a blackish crest on fourth segment, the ventral
surface whitish. Fore wing reddish brown, the basal half
suffused with black, its outer edge obliquely curved;
subbasal line represented by two slight strie from costa;
antemedial line indistinct, waved, defined by whitish on inner
side towards inner margin, some striz of raised black scales
beyond it in and below cell; a medial line bounding the
new Species of Noctuide. 535
dark area, oblique from costa to submedian fold; reniform
incompletely defined by black; postmedial line double, bent
outwards below costa, incurved at discal fold and below vein
4.; subterminal line strongly dentate, forming a dark mark at
costa and incurved below vein 4; a minutely waved dark
line just before termen, on which there is a series of black
strie. Hind wing fuscous, with hyaline streaks in the
interspaces in, below, and beyond cell, and on middle of
inner area; the underside with the inner area whitish.
Hab. Perax, Goping (Kiin-tler), 1 9 type. Hap. 32 mw,
Stictoptera leucostriga, sp. u.
&$. Head and thorax red-brown; pectus whitish; tarsi
fuscous ringed with white; abdomen fuscous brown with
some whitish at base, a blackish crest on fourth segment ; the
ventral surface white. Fore wing red-brown, sometimes
with triangular whitish patch on medial areafrom just above
median nervure to inuer margia; subbasal line represented
by two slight striae from costa; two antemedial indistinct
lines, approximated below cell, then diverging; reniform
with slight ochreous aunulus defined by blackish ; postmedial
line double, strongly bent outwards below costa, then
minutely waved, below vein 4 recurved to lower edge of reni-
form, then excurved again; subt-rmiual line black, its medial
part defined on inner side by a white striga, slightly angled
inwards below costa and incurved below vein 4; some white
points on costa towards apex; a terminal series of black
strie defined by whitish on inner side. Hind wing fuscous,
the interspaces of basal half hyaline; cilia with a fine pale
line at base.
? more fuscous brown; fore wing without the whitish
patch on medial area, the two antemedial lines black on
inner half forming a somewhat conical mark.
Hab. Penance (S. S. Flower), 1 2%; Perax, Goping
(Kiinst/e,) ; Stncavore (Ridley), 1 G,1 9? type. Hap. 24
28 mm.
Stictoptera ferruginea, sp. 0.
¢. Head and thorax rufous; abdomen rufous tinged with
fuscous, the anal tuft ochreous, the ventral surface whitish.
Fore wing rufous ; a faint double subbasal line from costa to
submedian fold, angled inwards below costa; a very indistinct,
double, minutely waved antemedial line with some dark
points on it; a slight discoidal lunule defined by brown; a
faint, double, minutely waved postmedial line, with some dark
536 Sir. G. F. Hampson on
points on its outer edge, oblique aud slightly incurved at
discal fold and in submedian interspace; subterminal line
indistinct ochreous, minutely dentate, defined by slight dark
marks on its inner side below costa and at middle and by a
complete series on outer; a terminal series of dark striz ;
cilia ochreous with dark points at extremity of the veins.
Hind wimg uniform fuscous brown; the underside rather
paler, with traces of curved postmedial line and diffused
subterminal band.
? much darker; fore wing with complete series of dark
spots before subterminal line.
Hab. Borneo, Sarawak (Wallace), 1 9; Puto Laur
(Doherty), 2 3 type. Exp. 80 mm.
Stictoptera anemia, sp. 0D.
9. Palpi with the third joint long and porrect.
Head and thorax whitish tinged with pale brown; frons
white ; abdomen grey-white with slight dark dorsal crests,
the ventral surface white. Fore wing ochreous white; a
very oblique, wedge-shaped, brown mark from costa before
middle to submedian fold, with a slight erect line from its
extremity to inner margin; some slight brownish marks on
terminal half of costa; postmedial line represented by a
series of dark striz from vein 7 to inner margin, excurved
between veins 5 and 3, then incurved and with some greyish
suffusion beyond them; subterminal line whitish, minutely
waved, slightly angled outwards at vein 7 and excurved at
middle, with small dentate black mark before it and streak
beyond it below costa, and slight blackish marks before it
and fuscous suffusion beyond it from vein 5 to inner
margin; a terminal series of small black lunules defined
by white on inner side. Hind wing pale fuscous, the inter-
spaces of basal half hyaline.
Hab. Souomon Is., 1 2 type. Eap. 30 mm.
Stictoptera hypenistis, sp. un.
6. Head and thorax dull brown; abdomen fuscous, the
ventral surface greyish. Fore wing pale reddish brown,
slightly irrorated with fuscous, the terminal area darker ;
traces of a double, irregularly waved antemedial line and of
two irregularly waved medial lines; an indistinct irregular
discoidal mark, constricted at middle; postmedial line oblique,
double, filled in with whitish, the inner line indistinct, the
outer strong and with a series of black points beyond it on
new Species of Noctuide. 537
the veins; a slight dark dentate subterminal line; a slightly
waved black terminal line ; cilia fuscous with a fine pale line
at base. Hind wing fuscous, the interspaces paler and semi-
hyaline, except on terminal area; cilia with a fine pale line at
base ; the underside with traces of curved postmedial line.
Hab. Burma, Hsipaw (De Nicéville), 1 g type. LEzp.
32 mm.
Gyrtona dialeuca, sp. n.
9. Head and thorax red-brown with white stripes on sides
of head and tegule and above patagia; palpi whitish at tips ;
abdomen pale fuscous, whitish at base, the dorsal crests
blackish, the ventral surface white. Fore wing rufous with
whitish fascia from base of costa through the cell to termen
below apex; a white streak on inner margin from near base
to subterminal line, towards which it increases in width; two
rather interrupted medial lines, bent inwards to costa and
terminating at vein 1, where they meet; reniform defined by
slight tufts of raised black scales; postmedial line indistinct,
double, minutely dentate, bent outwards below costa and
oblique below vein 4; subterminal line whitish, excurved at
middie, defined by blackish marks before and beyond it at
costa and middle and a black spot beyond it above tornus ;
a terminal series of blackish strie. Hind wing fuscous
brown, the interspaces of basal half semihyaline.
Hab. Borneo, Sarawak (Wallace), 1 9 type. Lup.
26 mm.
Gyrtona pheozona, sp. n.
Head and thorax reddish brown ; palpi and frons blackish ;
tegule with rather strong black medial line; tarsi blackish
with pale rings; abdomen brown tinged with fuscous, the
ventral surface whitish. Fore wing reddish brown with
slight dark irroration; a subbasal black spot on costa; a
double curved antemedial line followed by a rather obliquely
curved black band with two curved medial lines beyond it ;
medial area faintly tinged with fuscous ; reniform indistinct,
rufous with slight dark outline; antemedial line double,
filled in with ochreous, the outer line deep black, erect,
slightly excurved at middle; terminal area slightly tinged
with fuscous; subterminal line indistinct, greyish, defined
on outer side by fuscous, slightly excurved below costa and
at middle; a terminal series of black points. Hind wing
pale, semihyaline, the veins and terminal area fuscous; the
underside with indistinct curved postmedial line.
Hab. Bomsay, N. Canara, Karwar (Bell), 1 ¢ type;
Manras, Nilgiris, 1 ?. Hzp, 22 mm.
538 Su G. F. Hampson on
Gyrtona glaucobasis, sp. Nn.
9. Head and thorax reddish brown mixed with whitish ;
frons, pectus, and legs whitish; the tarsi fuscous with pale
rings ; abdomen grey tinged with fuscous, the ventral
surface white. Fore wing reddish brown, mostly suffused
with glistening glaucous grey and sparsely irrorated with
black, the antemedial area pale glaucous green; antemedial
line double with strize of raised black scales on its edges,
obliquely curved; reniform very indistinctly defined by
raised black scales on its edges, constricted at middle ; post-
medial line double, defined by striz of raised black scales on
its edges, those on outer side strong, slightly bent outwards
below costa and excurved at middle, with traces of another
minutely waved line beyond it; subterminal line grey defined
by shght blackish marks on outer side, somewhat dentate
and angled outwards at veins 7 and 3,4; a slight glaucous
line before termen ; cilia with a series of minute black points.
Hind wing uniform fuscous; the underside rather paler with
indistinct curved postmedial line.
Hab. Puro Laur (Doherty), 1 9 type. Hap. 22 mm.
Gyrtona chlorograpta, sp. u.
9. Head and thorax grey mixed with fuscous and brown ;
palpi blackish at sides; tegule with medial black line;
abdomen fuscous, the ventral surface whitish. Fore wing
grey suffused with brown and fuscous, leaving the costal half
of wing, except on basal area, paler; an oblique bright green
band from base of costa to inner margin, and a subbasal bar
from costa to cell; an antemedial green band from subcostal
nervure to vein 1, its edges defined by striz of raised black
scales ; a diffused dark spot on middle of costa and traces of
a fine, irregularly waved, medial black line ; a fine minutely
waved postmedial line incurved between vein 4 and sub-
median fold, with another indistinct waved line beyond it ;
a minutely dentate white subterminal line defined by black
on outer side and with blackish patch before it on costa; a
series of small black spots just before termen. Hind wing
uniform fuscous; the underside rather paler.
Hab. Stncarors (Ridley), 1 9 type. Hap. 22 mm.
Gyrtona plumbisparsa, sp. n.
9. Head, thorax, and abdomen dark red-brown; tarsi
with pale rings; abdomen with the slight dorsal crests
new Species of Noctuide. 539
black. Fore wing dark red-brown, suffused with leaden-
silvery scales, especially on inner and terminal areas ; traces
of two curved subbasal lines from costa to vein 1 ; three
indistinct waved antemedial lines with strize of raised black
scales beyond them in and below cell; an indistinct, waved,
brown medial line; two slight, obliquely placed, black dis-
coidal striz ; postmedial line double, smuous, bent outwards
below costa, excurved to vein 4, then retracted to below eud
of cell, and with traces of a waved brown line beyond it;
subterminal line dark defined by leaden-silvery suffusion on
inner side and with dark patch before it on costal area,
excurved at vein 7 and middle; a series of small dark spots
just before termen. Hind wing dark red-brown, the cilia
silvery at tips ; the underside paler.
Hab, Sixuim, 1 ? type. Hap. 24 mm.
Gyrtona polionota, sp. n.
9. Head and thorax pale red-brown suffused with grey ;
abdomen grey, the ventral surface white. Fore wing pale
red-brown, mostly suffused with grey and slightly irrorated
with fuscous; subbasal line represented by slight black
points on the veins; antemedial line indistinct, double,
minutely waved, curved, with some black points on it;
traces of a double medial line; reniform faintly defined by
black points; postmedial line indistinctly double, bent out-
wards below costa and oblique below vein 4, the outer line
somewhat dentate and with some black points on it ; sub-
terminal line represented by a series of black points, angled
outwards at vein 7 and slightly excurved at middle, followed
by an indistinct slightly waved line; a terminal series of
small black lunules. Hind wing fuscous, the interspaces of
basal half pale and thinly scaled.
Hab. Woopvusrk (Meek), 1 2 type. Herp. 80 mm.
Gyrtona polymorpha, sp. n.
6. Head and thorax purplish black-brown with some
rufous on head, tegule, and prothorax ; pectus and hind legs
greyish ; tarsi fuscous with pale rings; abdomen fuscous,
whitish below. Fore wing purplish black-brown with traces
of waved subbasal, antemedial, and medial lines; reniform
defined by black ; postmedial line double, minutely dentate,
bent outwards below costa, excurved to vein 4, then incurved ;
a double minutely waved subterminal line, slightly excurved
at vein 7 and middle; a terminal series of small black
lunules. Hind wing fuscous brown, slightly paler at base ;
the underside paler with indistinct curved postmedial line.
540 Sir G. F. Hampson on
Ab. 1. Fore wing redder brown, the basal area except
towards costa, the medial area except towards costa and
inner margin, and the area between the two subterminal lines
grey-white irrorated with fuscous ; a black patch before the
antemedial line below cell and another beyond it in and
below cell; reniform red-brown largely suffused with black.
? like the typical male, but head, thorax, and fore wing
redder brown.
Ab. 1. Fore wing pale red-brown, the inner half from base
to postmedial line suffused with white.
Ab. 2. Fore wing ochreous brown.
Ab. 3. Fore wing greyish brown, with black point on
antemedial line at vein 1 and patch beyond it in and below
cell.
Ab. 4, Fore wing with the medial and terminal areas dull
reddish brown, the basal area and the area between the two
postmedial lines, except at costa, ochreous brown.
Hab. Mavnritivs, Curepipe (Tullock), 2 g, 4 @ type.
Eup. 30-34 mm.
SARROTHRIPINE.
Hypothripa ruficirra, sp. n.
Head, thorax, and abdomen grey, the two former slightly
mixed with brown. Fore wing grey; a reddish-brown
antemedial shade from costa to submedian fold, with two
lines on it formed of raised black scales; an indistinct
slightly waved medial line; traces of a dark discoidal point ;
postmedial line bent outwards with a downward curve from
costa to vein 6, minutely dentate to vein 4, then strongly
incurved and excurved to inner margin, a rufous shade
beyond it from costa to vein 6; subterminal line in-
distinct, dark, minutely waved, excurved at vein 7 and
middle; a rufous shade at apex; a terminal series of black
points; cilia brownish. Hind wing brownish grey, the veins
and terminal area slightly darker.
Hab. Japan, Yokohama (Pryer), 2d type; Assam, Khasis,
19. xp. 22-24 mm.
Hypothripa miochroa, sp. n.
Palpi with the second joint dilated on inner side at
extremity.
9. Head and thorax greenish grey mostly suffused with
reddish brown; abdomen greyish fuscous. Fore wing
greenish grey, suffused and irrorated with reddish brown,
new Species of Noctuide. 541
especially on basal costal area and disk ; a double minutely
waved subbasal line from costa to submedian fold; a double
minutely waved antemedial line from costa to vein 1; a
whitish discoidal spot with two black points on it; post-
medial line defined by grey on outer side, bent outwards
below costa, then minutely dentate, angled inwards at vein 2;
subterminal line grey defined by fuscous before and beyond
it, slightly excurved at vein 7 and middle; a terminal series
of minute black points. Hind wing uniform fuscous brown ;
the underside paler, except costal area and terminal area to
vein 2.
Hab. Cotomsia, St. Marta (Bouchard), 1 2 type. Exp.
24 mm.
Pardasena ferrigrisea, sp. n.
3d. Head, thorax, and abdomen grey tinged with pale
iron-brown. Fore wing grey suffused with iron-brown; an
indistinct subbasal line from costa to submedian fold, ex-
curved below costa, then incurved ; antemedial line defined
by grey on inner side, minutely waved, oblique from costa
to submedian fold where it is obtusely angled; a slight
discoidal point; postmedial line defined by grey on outer
side, minutely waved, oblique from costa to vein 6, excurved
to vein 4, then incurved ; subterminal line indistinct, diffused,
dark, excurved at vein 7 and middle; a terminal series of
minute points. Hind wing grey suffused with pale brown,
the basal area slightly paler; cilia grey; the underside
paler.
Hab. Gambia, Bathurst (Carter), 1g type. Exp. 18 mm.
Pardasena deleta, sp. n.
6. Head and thorax pale ochreous brown; abdomen
whitish tinged with ochreous. Fore wing pale ochreous
brown; very indistinct sinuous antemedial and medial lines ;
traces of a diffused postmedial line excurved from costa to
vein 4, then oblique; traces of a subterminal series of dark
points ; the termen rather darker. Hind wing semihyaline
white, the terminal area tinged with brown.
Hab. Perv, Callao (J. J. Walker), 1 8 type. Erp.
22 mm.
Giaura leucophea, sp. n.
Head and thorax white and dark brown; palpi with the
second and third joints black except at extremities; antenne
dark; tibiz and tarsi banded with fuscous; abdomen
542 Sir G. F. Hampson on
whitish tinged with fuscous. Fore wing white slightly
irrorated with dark brown; a black point at base of costa ;
a brown patch on costal area with two obliquely placed black
points on it before the oblique, slightly sinuous, black ante-
medial line, with a small black spot beyond it above vein 1 ;
a rather indistinctly double, waved, somewhat oblique medial
line with a blackish patch beyond it on costa and in end
of cell, another patch beyond lower angle of cell before
the rather indistinctly double postmedial line, which is bent
outwards below costa, then sinuous and with a black patch
beyond it on costa; subterminal line angled inwards below
costa and at discal fold, excurved at vein 7, slightly dentate
at veins 4, 3, and with black spot on it at submedian fold ;
a blackish patch at apex and band just before termen down
to vein 5; a terminal series of black points; cilia white
intersected with fuscous. Hind wing pale, wholly suffused
with fuscous.
Hab. Borneo, Kuching, 1 ¢, 1 2 type. Hap. 20 mm.
Giaura nigriscripta, sp. N.
?. Head and thorax whitish grey ; palpi blackish behind ;
pectus and legs white, the fore tibize and tarsi grey in front ;
abdomen grey, the ventral surface white. Fore wing grey
irrorated with fuscous; a slight black streak on base of
median nervure ; a prominent, oblique, black subbasal line
from costa to submedian fold; antemedial line near middle
of wing, fine, black, angled outwards in cell and submedian
fold and inwards on vein 1; claviform represented by two
black streaks extending to the postmedial line; a slight
oblique discoidal shade; postmedial line fine, black, very
oblique from below costa to vein 6 and incurved below vein 4 ;
a subterminal series of slight dark points; a fine terminal
punctiform line. Hind wing white, the termen tinged with
brown from apex to vein 2.
Hab. Br. E. Arrica, Kikuyu (Crawshay), 1 @ type.
Eup. 22 mm.
Giaura strigivenata, sp. n.
?. Head, thorax, and abdomen whitish tinged with
brown ; palpi at sides and fore tibize in front dark brown.
Fore wing whitish irrorated and suffused with reddish brown ;
a diffused dark brown subbasal band from costa to sub-
median fold; a diffused blackish streak above median
nervure and another in terminal part of cell in discal fold;
diffused blackish streaks above vein 1 from before middle
new Species of Noctuide. 543
and on the veins beyond end of cell to the subterminal series
of somewhat dentate small dark marks and more prominent
rounded spot in submedian fold ; a terminal series of slight
points. Hind wing white tinged with brown, the terminal
area suffused with brown.
Hab. New Soutn Waters (Raynor), 1 2 type. Exp.
26 mm.
Giaura lichenigera, sp. un.
Palpi with the second joint broadly rounded with scales in
front.
?. Head and thorax greenish grey more or less mixed
with reddish brown and black ; palpi reddish brown at sides
barred with black ; fore tibize reddish brown in front barred
with black; abdomen grey dorsally tinged with fuscous
brown. Fore wing grey more or less tinged with green and
irrorated with fuscous, sometimes mostly suffused with red-
brown; a diffused black fascia in submedian interspace,
sometimes confined to medial area, ending at postmedial line
or downcurved at extremity and reaching tornus, another
streak above inner margin from near base to postmedial line
sometimes present ; a subbasal line of raised black scales,
oblique from costa to cell where it is angled and ending at
submedian fold; a very oblique sinuous antemedial line re-
tracted to costa; a round tuft of raised scales in end of cell;
postmedial line formed of raised black scales, sinuous, very
oblique from costa to vein 4, below vein 3 retracted to lower
angle of cell, then rather oblique and defined by white on
outer side to inner margin, sometimes with a black patch at
costa; subterminal line whitish with diffused fuscous on
inner side, excurved at vein 7 and at middle, bent inwards
at vein 3, then oblique to near tornus ; a terminal series of
black points sometimes with a fuscous band before them
from below apex to vein 3, Hind wing fuscous brown with
a reddish tinge, the interspaces paler except on terminal
area.
Hab. Sierra Leone (Clements), 1 2; Sixuim, 1 2? ; Sinca-
PORE (Ridley), 3 2 type. Hap. 18-22 mm.
Blenina metanyctea, sp. n.
6. Head and thorax white and green mixed with some
fuscous ; tegula black at outer edge and with black line near
tips; pectus and legs yellowish tinged with brown; fore
tibize and tarsi banded with black; abdomen yellow dorsally
suffused with brown, the crests at base whitish tipped with
544 Sir G. F, Hampson on
black. Fore wing white largely suffused with bright green ;
a diffused, irregular, olive-green and reddish-brown band from
middle of costa to tornus; a black line close to base, angled
in cell and with a black striga beyond it above inner margin ;
a waved, rather inwardly oblique antemedial line from costa
to above inner margin; an indistinct waved medial line;
reniform white irrorated with green, its lower part defined
on inner side by a curved black line and slightly by black
on outer side, its upper part very indistinctly and irregularly
defined by green, with a black striga from costa above it ;
postmedial line indistinct, minutely dentate, strongly in-
curved in submedian interspace ; a minutely dentate sub-
terminal line bent inwards at vein 2, then oblique to near
tornus ; a dark patch at apex and a terminal series of white
spots except towards tornus interrupted by the black termi-
nations of the veins ; cilia yellowish with a series of brown
spots and points at tips to vein 3, below which the termen is
distinctly excised. Hind wing dark brown, the inner
margin yellowish ; cilia yellowish with series of brown
spots to vein 2, whence the termen is excised to tornus ; the
underside yellow, the inner area tinged with brown, an
irregular band from costa to lower angle of cell and a broad
blackish band just before termen from costa to submedian
fold.
Hab. Dutcy New Guinea, Kapaur (Doherty), 1 go type.
Exp. 42 mm.
Blenina brachyptera, sp. n.
9. Head and tegule green; palpi banded with black;
tegule edged with dark brown; thorax reddish brown ;
pectus yellowish; legs yellowish and brown, the fore and
mid tibiz with green patches ; abdomen red-brown, the
ventral surface yellowish except towards extremity. Fore
wing short and broad, the apex and termen rounded, reddish
brown; basal area suffused with green except towards inner
margin ; a double, diffused, black subbasal line from costa to
vein 1, with some whitish on its outer edge; antemedial
line black defined by green on inner side, oblique from costa
to median nervure, then excurved and angled inwards on
vein 1, with slight green streaks on subcostal and median
nervures from it to postmedial line ; a slight discoidal dark
point ; postmedial line black defined by green on outer side,
obliquely excurved and minutely waved from costa to vein 4,
then incurved and with some white beyond the green near
inner margin; a slight, minutely waved, green subterminal
new Species of Noctuidie. 5A5
line, slightly excurved below vein 7, oblique and whitish
from vein 2 to tornus; termen suffused with fuscous.
Hind wing orange-yeilow with terminal black-brown band,
wide at costa, narrowing to vein 2 and expanding towards
tornus, some yellowish points on termen between veins 5
and 3; cilia with a fine yellowish line at base except towards
tornus. Underside of fore wing fuscous, yellow at base.
Hab. New Goines, Humboldt Bay (Doherty), 1 2 type.
Ep. 32 mm.
Blenina triphenopsis, sp. n.
&. Head and thorax greenish grey mixed with black ;
palpi with black points at base of third joint; tips of tegule
and upper edge of patagia sometimes black; tarsi banded
with fuscous ; abdomen orange-yellow tinged with brown
at extremity, the crests at base greenish grey. Fore wing
with the termen slightly oblique towards tornus ; greenish
white irrorated with brown and black; with more or less
prominent black suffusion between medial and postmedial
lines from costa to lower angle of cell and sometimes with
diffused black fascia on inner margin from near base to
subterminal line; traces of a waved line close to base from
costa to submedian fold; an irregularly waved antemedial
line bent outwards below costa, then oblique, ending at vein 1;
medial line minutely waved, oblique from costa to median
nervure ; a black point sometimes present in cell; a slight
white discoidal lunule with slight black streaks from its
lower extremity to the postmedial line, which is minutely
dentate, oblique from costa to vein 7, angled outwards at
vein 4, then incurved, with traces of a minutely dentate line
beyond it from costa to vein 4; a minutely dentate sub-
terminal line, angled outwards at vein 6, excurved at middle
and angled inwards at discal and submedian folds, where
there are blackish patches beyond it; cilia white, brownish
towards apex, with a fine waved black lne through them.
Hind wing orange-yellow with black terminal band, broad
at apex, narrowing to a point before tornus ; cilia orange,
tinged with fuscous at apex and at the angle at vein 2.
Underside orange; fore wing with the terminal half blackish
except on inner area, a black striga from costa above end of
cell with yellow spot beyond it; hind wing with black striga
from costa above end of cell, and terminal band as above.
Hab. Sincarorr (Ridley), 2 g type; New Guinea
(Mathew),1 3. Lxp. 42-45 mm.
Ann, & Mag. N. Hist. Ser. 7. Vol. xvi. 56
546 Sir G. F. Hampson on
Blenina chlorophila, sp. u.
g. Head and thorax bright green slightly irrorated with
white; tegule and patagia with blackish tips ; pectus and
legs yellowish, fore tibie green in front, the tarsi black
ringed with white; abdomen yellow- brown dorsally tinged
with fuscous, the crests green and black, the ventral saris
yellow, blackish at extremity. Fore wing whitish almost
entirely suffused with bright green; the base of inner area
tinged with rufous ; subbasal line oblique, waved, from costa
to submedian fold, ‘with an indistinct green line beyond it
from subcostal nervure to submedian fold; a double medial
line, incurved in cell and submedian interspace, excurved at
median nervure and to inner margin ; a slight black discoidal
bar ; postmedial line double, obliquely excurved and minutely
dentate from costa to vein 4, then single, strongly incurved
in submedian interspace, then oblique to mner margin and
slightly angled outwards on vein 1, a slight line beyond it
from vein 5 to 3, where it is bent inwards, then forming a
green band with blackish spot on it in submedian fold ; sub-
terminal line black, minutely waved, angled outwards at
vein 7, excurved at middle, bent inwards at vein 3 and
oblique to near tornus; a ‘slight black shade at apex and
oblique shade from subterminal line to middle of termen; a
terminal series of black points with white strize between
them; cilia white and green, brownish towards apex, with
series of black points on them. Hind wing black-brown,
the terminal area darker; an orange postmedial band from
vein 6 to 1, incurved below vein 4, obsolescent towards
vein 1, and emitting a slight streak towards base in sub-
median fold; cilia orange, black at apex and tipped with
brown and white above middle. Underside orange; fore
wing with oblique black medial band dilated at middle, the
terminal area black with sinuous inner edge ; ae wing
with black medial line from costa to below vein 2 and black
terminal band narrowing to a point near tornus.
Hab. Stncavore (Ridley), 2 g type. Hxp. 46 mm.
Blenina malachitis, sp. n.
Head and thorax bright sap-green, the latter with slight
dark irroration ; palpi above and antenne black, the palpi
of male with the second joint dilated at base and hollowed
out at sides, the hollow white; pectus and legs yellowish,
fore tibizw green in front, the tarsi brown ringed with white ;
abdomen reddish Brown, the terminal half in male with
new Species of Noctuidie. DAT
lateral fringes of hair curved over dorsum, the ventral surface
whitish, blackish at extremity. lore wing whitish, almost
entirely suffused with emerald-green ; with more or less
blackish suffusion on medial and ‘postmedial area and some-
times some rufous suffusion beyond the cell; an obliquely
curved, slightly waved, black subbasal line from costa to
submedian fold with an indistinct dark bar beyond it from
cell to submedian fold; a more or less distinct double ante-
medial line, angled outwards below costa and cell and at
vein 1; a shght black discoidal bar with whitish point before
it; postmedial line slightly bent outwards below costa,
minutely dentate to vein 3, then strongly incurved and
oblique to inner margin, with another less distinct line
beyond it from costa to vein 3; subterminal line black, green
at middle, very minutely dentate, angled outwards at vein 7,
excurved at middle, bent inwards at vein 2 and oblique to
near tornus 3a blackish patch at apex and below vein 6; the
tips of the veins with slight black streaks with whitish strize
between them; cilia whitish, tinged with green and with
brown towards apex, with series of black points on them.
Hind wing black-brown, darker towards termen; a more
or less developed, curved, yellowish postmedial bar between
veins 5 and 2 emitting a streak towards base below cell; cilia
yellow, blackish at apex and with some dark points at middle.
Underside of fore wing dull yellowish with blackish medial
band, diffused except at costa, terminal area blackish with
diffused inner edge ; hind wing yellowish white with traces of
curved medial line and broad terminal black band from apex
to submedian fold.
Hab. Sincavore (Ridley, Friihstorfer), 2 3, 1 Q type.
Eup. 40 mm.
Blenina miota, sp. un.
3d. Head and thorax pale green mixed with fuscous ;
pectus and legs whitish, the fore tarsi fuscous ringed with
white ; abdomen whitish suffused with fuscous. Fore wing
pale green irrorated with fuscous, the area from middle of
costa and beyond cell suffused with fuscous down to vein 5 ;
an indistinct subbasal black striga from costa; antemedial
line represented by a black striga from costa and point below
median nervure ; an oblique fuscous striga from middle of
costa ; two black discoidal points; postmedial line double,
bent outwards below costa, minutely dentate to vein 3, then
the inner line: obsolete, the outer line erect to inner margin ;
subterminal line formed of small fuscous spots, strongly
36*
548 On new Species of Noctuide.
excurved from vein 5 to 2, where it is bent inwards, then
oblique to tornus; aseries of minute fuscous spots just before
termen, on which there is a series of black strie. Hind wing
brown with a slight reddish tinge ; cilia whitish ; the under-
side whitish, with traces of a curved postmedial line and broad
fuscous terminal band.
Hab. Sinxra Leonz, Pt. Lokko (Penny), 1 ¢ type.
Exp. 28 mm.
Blenina leucotis, sp. n.
2. Head and thorax white, slightly irrorated with fuscous ;
palpi blackish above; fore tarsi banded with black ; abdomen
white. Fore wing glistening white, rather sparsely irrorated
with fuscous ; a minute black streak in base of cell; sub-
basal line angled outwards in cell, then oblique to submedian
fold; antemedial line double, the inner line indistinct,
strongly waved; medial area clouded with black towards
costa and from cell to inner margin and with a diffused black
mark with two points beyond it in submedian interspace ;
postmedial line angled inwards below costa, oblique to vein
6, then indistinct and minutely dentate to vein 3, where it is
bent inwards, then waved to inner margin, a blackish shade
beyond it on costal area followed by two small spots; a
diffused fuscous subterminal line angled inwards in discal
and submedian folds and excurved at middle ; termen clouded
with fuscons, a terminal series of black points. Hind wing
white, the terminal area tinged with fuscous from apex to
vein 2.
Hab. Br. E. Arrica, Uganda Ry., mile 478 (Betton),
lo type, Hay. 300nm.
Blenina brunnescens, sp. n.
¢. Head and thorax white, tinged with pale red-brown;
fore tarsi blackish at extremity; abdomen brownish white.
Fore wing white, suffused with pale red-brown; a slight
blackish streak on base of median nervure; an obliquely
curved antemedial series of four black points from below
costa to median nervure and point on vein 1, an oblique
striga beyond them across base of vein 2; two black dis-
coidal points; a postmedial series of minute streak-like
points, obliquely excurved from costa to vein 4, then oblique ;
traces of a subterminal series of small brown spots; a
terminal series of black points. Hind wing white with a
slight reddish-brown tinge.
Hab. Br. C. Arrica, Nyasa (de Jersey), 1 3 type.
Exp. 22 mm.
On new Species of Diurnal Lepidoptera. 5A9
Barasa tetragramma, sp. 0.
?. Head and thorax white, slightly irrorated with fuscous;
palpi with black mark at side of second joint; fore tibize
blackish on inner side, the tarsi blackish at extremity ;
abdomen white, tinged with grey above, the basal crest
blackish at tip. Fore wing grey-white irrorated with fuscous,
the medial and terminal areas slightly tinged with brown ;
a slight black streak in base of cell; a strong obliquely
curved black subbasal line from costa to submedian fold; a
highly waved medial line with traces of another line before
it ; a slight diffused rufous discoidal mark ; postmedial line
bent outwards below costa, incurved at discal fold and below
vein 4, angled outwards at vein 1; subterminal line strong,
black, not quite reaching costa, on which there is a black
point, met below costa by an oblique shade from apex slightly
excurved at middle ; a terminal series of minute black streaks.
Hind wing white, the apical area tinged with brown down
to vein 2; the underside with the costal area tinged with
brown.
Hab. Fis1, 1 9 type. Hap. 28 mm.
This is the specimen recorded as Argyrothripa nigrostrigata
by Mr. Bethune-Baker (P. Z. S. 1905, 1. p. 93), the type of
which, from New Guinea, has, however, the medial line not
angled inwards below the cell.
[To be continued. |
LIX.—Deseriptions of Two new Species of Diurnal Lepido-
ptera belonging to the Subfamily Nymphaline. By Hersert
Druce, F.L.S. &e.
Perrhanea cubana, sp. n.
Male.—Head, thorax, and abdomen yellowish brown ;
antenne black. Palpi, underside of the thorax, and abdomen
pale brown. Primaries bright red, the costal margin, apex,
and outer margin black, the outer margin spotted with
yellowish brown ; a short black band at the end of the cell,
beyond which a waved rather wide black line crosses the
wing from the costal to the inner margin; two small black
dots close to the apex; the fringe pale reddish yellow:
550 On new Species of Diurnal Lepidoptera.
secondaries, the basal half of the wing bright red, the outer
half deep chrome-yellow ; a submarginal reddish-brown line
extends from the apex to just above the tail, which is black,
irrorated with yellowish-white scales; the fringe almost
white. Underside: primaries, the costal half, apex, and
outer margin greyish black, with the black markings very
similar to those on the upperside; the inner half of the wing
reddish brown, thickly irrorated with black scales: second-
aries blackish from the base to about the middle, beyond
which to the outer margin pale brownish grey, irrorated with
black scales; asmall black spot above the tail.—/emale very
similar to the male, but with much less red on the primaries,
which are chrome-yellow along the outer margin: secondaries
almost entirely chrome-yellow, slightly shaded with red near
the base: on the underside both wings are more yellow than
those of the male.
Expanse, ¢ 34, 2 32 inches.
Ilab, Cuba.
In the collection of Mr. Henley Grose-Smith and Herbert
Druce.
Euphedra Cooksont, sp. 0.
Male.—Head, antenne, thorax, and abdomen black; a
bluish-white dot at the base of the antenne, two in front of
the thorax, one on each side and two at the base of the thorax,
and two on each side of the abdomen nearest the base; the
point of the antennee yellow; palpi, underside of thorax, and
abdomen chrome-yellow ; legs yellowish. Primaries black,
greenish grey along the costal margin ; a white band crosses
the wing near the apex from the costal to the outer margin,
slightly shaded with greenish grey on the inner margin of
the band; the inner margin from the base to near the apex
broadly brownish chrome-yellow, the fringe black : second-
aries brownish chrome-yellow, broadly bordered with black
from the apex to the anal angle; two bluish-grey spots on
the outer margin near the anal angle. Underside: primaries
yellow, shaded with greenish black ; the white band as above ;
three black spots in the cell, the first close to the base and
two close to the end of the cell; the wing is palest on the
inner margin: secondaries chrome-yellow ; a black spot at
the base and a small one in the cell ; a rather wide white
submarginal band extends from the base to the apex, where
it becomes quite narrow ; two indistinct white spots at the
end of the cell and four bluish-white dots extending round
the outer margin from the anal angle—J/emale very similar
On a new Land-Shell from Christmas Island. 551
to the male, with all the white markings on the underside
more distinct.
Expanse, ¢ 31, 2 4 inches.
Hab. N.E. Rhodesia.
LX.—Description of a new Land-Shell from Christmas
Island. By Epoar A. Smita, 1.8.0.
A SMALL collection of non-marine Mollusca, recently sent to
the Natural History Museum for identification by Dr. R.
Hanitsch, of the Raffles Museum, Singapore, contains the
following new species of Jaminia from Christmas Island, in
the Indian Ocean, and also two or three other forms previously
unknown from this remote locality. These will be referred
to in a paper which Dr. Hanitsch proposes tq publish as a
supplement to the ‘ Monograph of Christmas Island’ by
Dr. C. W. Andrews.
Jaminta proscripta.
Testa minuta, oblonga, cylindrica, umbilicata, fusco-cornca, lineis
incrementi tenuibus striata; anfractus 5, perconvexi, sutura
profunda leviter obliqua sejuncti, ultimus antice leviter ascendens,
pone labrum indentatus; labrum pallidum, expansum, subreflexum,
margine externo in medio intus leviter prominens; apertura
quinque-dentata; dentes duo parietales inzequales, columellaris
minutus, palatales duo lamelliformes, subequales, ab margine
labri remoti.
Longit. 2°25 mm., diam. °9.
Hab. Flying Fish Cove.
The type will be preserved in the British Museum.
ay On remarkable Specimens of Rana esculenta.
LXI.—On remarkable Specimens of Rana esculenta from
South-western Persia. By G. A. BOULENGER, F.R.S.
THE collection made by Mr. R. B. Woosnam when accom-
panying Col. Bailward in Persia contains two specimens of
Rana esculenta, male and female, which do not fall under the
definition of any of the races hitherto distinguished, and for
which I think it desirable to propose a new name—var. susana
—the specimens having been captured not far from the ruins
of Susa, at an altitude of about 400 feet.
This variety agrees with var. ridibunda, to which I have
referred the Persian specimens previously examined by me, in
the small size and feeble prominence of the inner metatarsal
tubercle, and with the typical form in the comparatively short
tibiz, which barely meet at the heels when they are laid at
right angles to the body. Besides, the length of the tibia is
less than one half that of head and body, whilst in var. rédi-
bunda it usually measures one half or more. The life-colour
of the malespecimen, obtained in a small stream on March 20th,
is described in Mr. Woosnam’s notes as drab colour, with a
few dark brown spots; head, shoulders, and upper back light
green: the female, caught in the Kerkbah River on March
19th, was of a ereenish-brown colour, with dark brown
blotches and a broad light stripe along the middle of head
and body. Other specimens of /, esculenta, obtained by
Mr. Woosnam at Kermanshah, 6000 feet altitude, are
referable to the var. ridibunda.
I append some measurements of these specimens, in
accordance with the method proposed in the ‘ Proceedings of
the Zoological Society,’ 1891, p. 377 :—
Ly Be 3.5 phos
Var. ridibunda. WKermanshah. ¢.... 67 354 383 9 383
5 5 ais MOD)» 45-~ 40, si meee
. : ier One aot ll aaeee
Wie (IIH. TSUEEA» 6) Guooacooobae 6429 29 ees
7 Fd Be eas en toric 63 29 29 73 3
1, Length (in millimetres) from snout to vent. Length of tibia (in
the flesh). 38. Length of foot measured aa outer metatarsal
tubercle to tip of fourth toe. 4, Length of inner toe. 5. Length
of inner metatarsal tubercle.
Mr. W. L. Distant on Homoptera. 553
LXII.—Rhynchotal Notes —XXXVI. By W. L. Disranr.
Fam. Cicadide.
THERE are a few unique types described by Walker in the
British Museum, which have no locality and are female
specimens. ‘Their generic identification is a matter of doubt
and difficulty, and the following notes have reference to same.
I also append the descriptions of some species from various
habitats.
Purana? hirundo.
Cicada hirundo, Walk. List Hom, i. p. 152 (1850),
This unlocalized type is a female specimen, not “ Mas ” as
stated by Walker, and apparently belongs to the genus
Purana.
Purana ? conifacies.
Cicada conifacies, Walk, List Hom., Suppl. p. 85 (1858).
Dundubia conifacies, Stil, Ofv. Vet.-Ak. Forh. 1862, p. 481,
This species is only represented by a single unlocalized
female specimen, It is apparently a species of Purana.
Diceropyga? nodosa.
Cicada nodosa, Walk. List Hom. i. p. 162 (1850).
A single unlocalized female specimen which I have
doubtfully referred to this genus, the principal characters of
which are found in the male, viz., the opercula and the
structure of the anal abdominal segment.
Diceropyga? acutipennis.
Cicada acutipennis, Walk. Ins. Saund., Hom. p. 20 (1858).
This species is also only represented by a single unlocalized
female specimen, and the remarks appended to the previous
species equally apply to this one.
Terpnosia? stipata.
Cicada stipata, Walk. List Hom. i. p. 155 (1850),
Only represented by a single unlocalized female specimen,
which seems clearly to belong to this genus. Should this
species be proved ultimately to belong to Terpnosta its
specific name must be altered, as it is already used in the
genus.
554 Mr. W. L. Distant on Homoptera.
Subfam. Orcaprv2.
POLYNEURARIA.
Platypleura attenuata, sp. n.
Head and thorax above brownish ochraceous, head with the
anterior margins of front and the lateral margins of vertex
luteous ; base of front, a spot near anterior angle of vertex,
and a transverse fascia between the eyes black; pronotum
with a central longitudinal fascia (not extending through
posterior margin), a longitudinal fasciate spot on each side of
disk, the incisures, and a small marginal spot black ; meso-
notum darker in hue, with four obconical spots, of which the
two central ones are very short, two rounded spots in front of
cruciform elevation, and a spot on each anterior angle of
same, black ; abdomen with the basal segmental areas black,
their apical areas ochraceous ; body beneath piceous, clothed
more or less with very short greyish hairs; a somewhat
macular transverse fascia between eyes at base of face, apices
of anterior and intermediate femora, a longitudinal spot on
each side of anterior femora, and the lateral and posterior
margins of opercula, ochraceous; a spot on each side of
anterior femora near their apices and a longitudinal fascia
beneath them, central longitudinal sulcation to face, and a
suffusion between face and eyes, and the opercula (excluding
margins), black; abdomen beneath piceous, the posterior
segmental margins ochraceous; tegmina and wings hyaline,
the venation fuscous or fuseous brown; tegmina with the
costal membrane. brownish ochraceous, with a spot at about
one third from its base and the whole basal cell piceous ;
claval area brownish, thickly greyishly pilose, and with
some scattered greyish hairs on the basal half of tegmina;
transverse veins at bases of apical areas more or less infus-
cated and submarginal fuscous spots on most of the longitu-
dinal veins to apical areas; wings with the extreme base pale
ochraceous, margins of anal area pale fuscous where the veins
are piceous.
Long., excl. tegm., ¢ 19 mm.; exp. tegm. 66 mm.
Hab. N. Nigeria; Zunguru (A. L. de M. Mynn, Brit.
Mus.).
The lateral margins of the pronotum are angularly
developed, and extend to about half the length of basal cell ;
the whole body above is more or less greyishly pilose, and
the abdomen in male is short and attenuated posteriorly ; the
opercula do not quite mect inwardly, and their lateral and
~
55
Or
Mr, W. L. Distant on Homoptera.
posterior margins are convexly rounded and do not pass the
base of abdomen ; the tegmina are long and narrow, their
greatest breadth being a little less than a third of their length ;
rostrum passing the posterior coxe.
DUNDUBIARIA.
Purana Davidi, sp. n.
Body above brownish ochraceous ; broad lateral margins
to face enclosing a pale spot anteriorly, two transverse
fascize on each lateral area of vertex, and the area of
the ocelli black ; pronotum with the anterior and posterior
margins and a central linear spot to lateral margins, two
central longitudinal fascize (widened and ampliated anteriorly,
united posteriorly), and the incisures black ; mesonotum
with five black longitudinal fascize, the central one continuous,
one on each side shorter and a little curved, the two lateral
fascize much broader, crossing mesonotum, but prominently
broken near base ; a large spot in front of cruciform elevation
and posterior segmental margins black; head _ beneath,
sternum, and legs a little paler than above, abdomen beneath
ochraceous ; face (excluding a basal spot and transverse
striations), space between face and eyes, apices of anterior
and intermediate femora and the upper surfaces of same
(more or less), coxal spots, apex of rostrum, and abdominal
tubercles, piceous or black; tegmina and wings hyaline,
venation testaceous, in parts piceous; tegmina with the
costal membrane testaceous, a distinct stramineous spot at
apex of radial area, and the transverse veins at bases of
second, third, fifth, and seventh apical areas broadly infus-
cated ; blavall areas to both tegmina and wings fuscous ;
opercula i in male transverse, greyish, their extreme margins
piceons, their posterior margins oblique, their posterior
angles just passing base of “second abdominal segment,
outer margins nearly straight, ner angles well separated ;
rostrum reaching apex of basal abdominal segment,
abdominal tubercles very large and prominent; spines to
anterior femora long, piceous, their apices testaceous.
Long., excl. tegm., ¢ 26-33, 9 27; exp. tezgm., ¢ 9,
79-88 mm.
Hab. Kiang-S8i (A. David, 1875, Paris and Brit. Muss.).
Purana pigmentata, sp. n.
Body above brownish ochraceous ; head with some piceous
lateral striz to front; area of the ocelli with a curved fascia
556 Mr. W. L. Distant on Homoptera.
on each side, and inner margins of eyes, black; pronotum
with two central longitudinal fascize (almost united anteriorly
and posteriorly and with a small discal curved fascia on each
side), incisures, narrow lateral and posterior margins, and two
spots on each side of posterior margin, black or piceous ;
mesonotum with a central longitudinal somewhat lanceolate
fascia, on each side of which is a short somewhat oblique
fascia, a longer and broader fascia on each lateral area, and
two rounded spots in front of cruciform elevation, black ;
a broad outer lateral fascia to tympanal coverings, and anal
segment above and beneath, black or piceous, posterior
segmental margins a little paler and with some obscure
lateral and discal fuscous spots; body beneath and legs a
little paler than above; sternum and opercula somewhat
finely greyishly tomentose, sternum with some obscure discal
piceous spots; face with the upper transverse strie (not meeting
centrally) and discal margins of central sulcation piceous,
a black transverse line between face and eyes; abdominal
tubercles pale castaneous ; tegmina and wings hyaline, their
bases narrowly ochraceous, venation ochraceous near base and
afterwards fuscous; tegmina with the costal membrane and
postcostal veins ochraceous, transverse veins at bases of
second and third apical areas slightly infuscated ; opercula
in male oblique, outwardly moderately convex, somewhat
widely separated inwardly, their apices moderately obliquely
truncate and about reaching anterior margin of basal
abdominal segment ; rostrum reaching base of abdomen, its
apex piceous; anterior femora beneath with two strong spines
and a smaller apical one.
Long., excl. tegm., ¢ 17 mm.; exp. tegm. 47 mm.
Hab. Siam, Bangkok; Cambedia; Cochin China, Lakhon.
Allied to P. carmente, Walk., from which it differs by the
concolorous, shorter, and differently shaped opercula in the
male, and also by the broad outer piceous lateral margins of
the tympanal coverings.
Sauda Pratti, sp. n.
3 %. Brownish olivaceous; anterior margin of front
and lateral areas of vertex brownish castaneous, area of
ocelli black ; pronotum with two central fascize ampliated
anteriorly and posteriorly, the incisures, and inner, lateral,
and posterior margins, a central longitudinal line crossing the
posterior margin, and the whole of the anterior lateral angles,
piceous or black; mesonotum with two central obconical
spots, their outer margins continued to anterior angles of
Mr. W. L. Distant on Homoptera. 5d7
cruciform elevation, an obsolete spot on each side of the two
central ones and a similar spot in front of cruciform elevation,
piceous; abdomen above much suffused with piceous ; body
beneath, opercula in male, and legs brownish ochraceous ;
under surfaces of anterior tibie, anterior and intermediate
tarsi, and apex of rostrum piceous; tegmina and wings
hyaline, veins fuscous, a little suffused with pale fuscous
brown along the veins, transverse veins at the bases of
second, third, fifth, and seventh apical areas more distinctly
infuscated ; subclaval areas distinctly fuscous and. inwardly
margined with black ; tegmina with the costal membrane and
basal cell brownish ochraceous ; opercula in male reaching
the penultimate abdominal segment; their apices broad and
subangularly rounded, narrowed towards base, where they
are concavely sinuate on each side, their inner margins
contiguous but not meeting on disk of abdomen, their
surface distinctly granulose, their edges a little recurved and
apices very narrowly cretaceous; rostrum reaching the
posterior coxe.
Llong., excl. tegm., g 48, 2 51; exp. tegm., ¢ 154,
9 168 mm.
Hab. Brit. New Guinea, Ekeikei (Pratt).
Sawda Sharpi, sp. n.
Closely allied to the preceding species (S. Pratti), but
differing by its smaller size, the opercula in male only
reaching the third abdominal segment, and though of same
general shape yet much more narrowed at their bases;
transverse veins at the bases of apical areas to tegmina a little
more infuscated ; pronotum and mesonotum with the piceous
markings obsolete.
Long., excl. tegm., ¢ 47 mm.; exp. tegm,. 132 mm.
Hab. Brit. New Guinea ; Ekeikei (Pratt).
Diceropyga aluana, sp. n.
Head and pronotum ochraceous ; lateral striz to front, a
spot at anterior angles of vertex, area of ocelli, and extreme
basal margin of vertex, piceous or black; anterior margin of
pronotum narrowly black ; mesonotum greenish ochraceous,
sometimes suffused with piceous, with two obconical spots
denoted by their dark outer margins; abdomen in male
brownish ochraceous, palely pilose, with a lateral series of
large black segmental spots on each side and the apex
piceous; in female with the lateral spots smaller, more
elongate, and fasciate, and with a large cretaceous spot at
58 Mr. W. L. Distant on Homoptera.
Or
base of anal segment; body beneath and legs pale ochraceous,
apex of rostrum black ; tegmina and wings hyaline, the
venation ochraceous or ’ Euscous ; tegmina “with’ the costal
membrane greenish ochraceous, a small prominent ochraceous
spot at apex of radial area, a fuscous spot on transverse veins
at bases of second, third, fifth, and seventh apical areas, and
marginal fuscous spots near apices of longitudinal veins to
apical areas; opercula in male extending to about half the
length of abdomen, their apices subacutely rounded, their
outer margins concavely sinuate, their inner margins convex ;
rostrum passing the posterior cox; anterior femora with
two strong spines beneath.
Long., excl. tegm., g 33, 9 24 mm.; exp. tegm., ¢ 2,
76-80 mm.
fab. Solomon Islands; Alu (Brit. Mus.).
Pomponia piceata, sp. n.
Body and legs uniformly piceous brown ; mesonotum with
four darker obconical spots, the two central ones shortest ;
tegmina and wings talc-like, a little opaque, but uncertainly
discerned in a somewhat discoloured spirit-specimen, their
extreme bases piceous ; tegmina with the venation fuscous,
in some parts ochraceous, costal membrane piceous brown,
transverse veins at bases of second, third, fourth, and fifth
apical areas slightly infuscated, and a very obscure marginal
series of pale elongate fuscous spots at apices of longitudinal
veins to apical areas; wings with the venation brownish ;
face globose, the striations robust; rostrum passing the
posterior cox ; opercula in male transverse, their posterior
margins convex, just reaching base of second abdominal
segment, their inner angles a little separated.
Long., excl. tegm., ¢ 42; exp. tegm. 108 mm.
Hab. ‘Tonkin, Cho-Moi (foget, Paris Mus.).
Oncotympana fuscata, sp. n.
Head and pronotum brownish ochraceous; head with curved
striz to front, broad inner area to eyes, and area of ocelli
black ; pronotum with extreme margins, incisures, two broad
central longitudinal fascie (a little ampliated anteriorly, and
much ampliated posteriorly), two spots on disk, and two large
oblique spots on each apical area of posterior margin black ;
mesonotum black, with two subobsolete central obconical
spots, and with the following obscure castaneous maculations,
viz., a very obscure spot in each of the obconical areas, a
spot outside them, a larger spot at their apices, a spot on each
Mr. W. L. Distant on Homoptera. 559
lateral area, and a transverse spot on each side of the lateral
angles to cruciform elevation; cruciform elevation dull
castaneous, its disk and a spot on its anterior angles black ;
abdomen above black, the tympanal coverings dull ochraceous,
the posterior basal segmental margins testaceous; sternum
and opercula greyish ochraceous; legs and abdomen beneath
brownish ochraceous; apices of anterior femora, anterior
tibie (excluding two longitudinal streaks), apical areas of
intermediate tibie, apex of posterior tibiz, anterior tarsi,
intermediate tarsi (excluding base), and obscure lateral spots
to abdomen, black ; spiracular spots to abdomen cretaceous ;
tegmina and wings moderately infuscate, the venation fuscous,
in parts dull ochraceous; tegmina with the costal membrane
testaceous, the transverse veins at the apices of the second,
third, fifth, and seventh apical areas moderately infuscate ;
rostrum passing the posterior coxe ; opercula in male broad,
convex, and overlapping, reaching the base of the second
abdominal segment.
Long., excl. tegm., ¢ 33; exp. tegm. 108 mm.
Hab. North China (Paris Mus.).
Allied to O. maculaticollis, Motsch., from which, apart from
the different coloration, it may be distinguished by the
narrower tegmina and the broader cruciform elevation.
Oncotympana stratoria, sp. n.
Body and legs brownish ochraceous; head with lateral
striz to front, two spots on each lateral margin of vertex,
area of ocelli, and inner margin of eyes black; pronotum
with the lateral margins, incisures, two central longitudinal
fasciz (ampliated anteriorly, and united posteriorly) black ;
mesonotum with four broken and subobsolete obconical spots,
the two central ones smaller and only denoted by their
margins, a small marginal spot before the lateral spots, and
two rounded spots in front of the cruciform elevation black ;
abdomen above with two large spots on basal, and the greater
part of the remaining segments, black; a broad central
fascia to face, apices of anterior and intermediate femora, bases
and apical areas of anterior and intermediate tibia, apex
of rostrum, and lateral abdominal marginal spots, black ;
tegmina and wings hyaline; tegmina with the venation
mostly purplish red on basal half, becoming fuscous towards
apex, a somewhat large fuscous spot on transverse veins to
second, third, fifth, and seventh apical areas, and a marginal
series of minute pale fuscous spots near apices of longitudinal
veins*to apical areas; venation of wings mostly fuscous brown ;
560 Mr. W. L. Distant on Homoptera.
rostrum considerably passing the posterior cox; opercula
centrally overlapping, not quite reaching base of abdomen.
Long., excl. tegm., § 263; exp. tegm. 83 mm.
flab. Yunnan (Paris Mus.).
Allied to O. virescens, Dist., but rostrum larger, opercula
shorter, and colour different.
Subfam. Vrerorvry x.
CARINETARIA.
GUARANISARIA, gen. nov.
Allied to Carineta, but with the tegmina short, not longer
than the body, semiopaque and with an additional vein or
veinlet crossing the third ulnar area. Other structural
characters as in Carineta,
Type, G. disstmilis, Dist.
Guaranisaria dissimilis, sp. 0.
¢. Head and thorax above black, shortly palely pilose ;
a small central spot at base of head, lateral and posterior
margins (very narrow) and the incisures to pronotum, and
Jateral margins to mesonotum ochraceous, the mesonotum
also obsoletely displays the margins of two central obconical
spots; abdomen above ochraceous, the basal segment and
central area of second segment black; exposed tympana
greyish white; body beneath black, the lateral margins of
both sternum and abdomen ochraceous and longly pilose ;
tibia and tarsi stramineous, bases and apices of anterior
and intermediate tibiee, and apices of tarsi piceous or black ;
opercula ochraceous, inwardly, outwardly, and apically
piceous ; tegmina suffused with shining piceous brown, the
costal membrane stramineous, the subclaval area roseate ;
wings hyaline, the venation ochraceous; front robustly
protuberant, about as long as vertex; transverse vein at base
of second tegminal apical area almost vertical; rostrum
reaching intermediate cox; opercula transverse, not meeting
inwardly, their posterior margins rounded and not quite
reaching base of abdomen,
Long., excl. tegm., ¢d 15 mm.; exp. tegm. 34 mm.
Hab. Argentina (Wagner, Brit. Mus.); Paraguay; Sapu-
eay (Coll. Heidemann).
A female specimen from Paraguay sent me for identi-
fication by Mr. Heidemann has the abdomen completely
piceous black above with only marginal lateral ochraceous
streaks, beneath the lateral ochraceous abdominal margins
are traversed by a black median fascia.
Mr. W. L. Distant on Homoptera. 561
TAPHURARIA,
Abroma philippinensis, sp. n.
2. Body pale testaceous; head with a castaneous or
piceous spot at anterior angles of vertex; mesonotum with
four subobsolete obconical spots, of which the two central are
shortest ; sternum somewhat thickly palely pilose; rostrum
with its apex piceous and reaching the intermediate coxe ;
streaks to coxee and femora, and a central fascia to abdomen
beneath, piceous; tegmina and wings hyaline, venation
ochraceous or brownish ochraceous ; costal membrane ochra-
ceous, veins at bases of first and second apical areas and a
subclaval streak pale fuscous; front distinctly finely longi-
tudinally sulecate; first and third apical areas of tegmina
about equal in length and a little longer than second.
Long., excl. tegm., 9 15-17 mm.; exp. tegm. 39-41 mm.
Hab. Philippine Islands, Palaouan (Paris Mus.).
I have not seen a male of this species.
PARNISARIA.
Quintilia Souliz, sp. n.
¢. Body above black, mostly palely pilose ; head with the
apical margin of front, lateral margins of vertex, and a
central spot at base testaceous; pronotum with the margins
and a central fascia which is ampliated posteriorly (where it
contains a black spot) pale testaceous ; mesonotum with the
margins and two central elongate fasciz (each containing a
black spot) testaceous ; abdomen above with the segmental
margins testaceous ; body beneath testaceous ; face (excluding
margins and basal spot), streaks of femora, anterior tibia,
bases and apices of intermediate and posterior tibia, tarsi, and
metasternum black; tegmina and wings hyaline, venation
ochraceous on basal and fuscous on apical areas; tegmina
with the costal membrane ochraceous ; wings with the margins
of the anal area fuscous.
Var. a.— @. Mesonotum with only faint indications of the
pale central elongate fascie.
Var. b.— . Both pronotum and mesonotum without pale
central fasciee.
Long., excl. tegm., ¢ 9 174 mm.; exp. tegm. 39-42 mm.
Hab. Yunnan; Tsé-kou (Mgr. Soulié, Paris Mus.).
Without locality (Brit. Mus.).
Allied to Q. megannia, Dist., but tegmina totally pale
hyaline, wings with only margins of anal area infuscated,
lateral pronotal margins less oblique and more sinuate, &c.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 37
562 Mr. W. L. Distant on Homoptera.
CHLOROCYSTARIA.
AKAMBA, gen. nov.
&@. Head with the front anteriorly and somewhat trian-
gularly produced, centrally distinctly longitudinally impressed,
about as long as vertex, ocelli much nearer to each other than
to eyes, lateral margins of vertex a little upwardly reflexed
and discontinuous with margins of front, including eyes a
little narrower than base of mesonotum ; pronotum consider-
ably shorter than mesonotum, about as long as head, its
lateral margins a little convexly sinuate, its posterior angles
somewhat strongly ampliate; abdomen longer than space
between apex of head and base of cruciform elevation, sub-
globose, with a central dorsal longitudinal ridge, and with a
prominent laterally produced nodosity on each side of basal
segment, the anterior margin of which is a little convex, but not
centrally conically produced ; tympana completely exposed ;
anal segment in male largely produced; rostrum reaching
the intermediate cox ; opercula small, obtusely angulate,
placed at each lateral margin, not extending beyond base of
abdomen; tegmina and wings hyaline, the first with the
second and third ulnar areas shortest, apical areas eight, basal
cell about twice as long as broad; wings with six apical
areas; lees somewhat short and slender, anterior femora with
one moderately strong spine.
Allied to Musoda, Karsch, from which it differs by the
non-conically produced basal abdominal segment and _ its
laterally produced nodosities on each side, &e.
Type, A. ethiopica, Dist.
Akamba cethiopica, sp. n.
¢@. Body and legs ochraceous; ocelli red; pronotum with
the margins, a central longitudinal fascia, on each side of
which is a short curved discal fascia, and the incisures
fuscous brown; mesonotum with two short central obconical
spots, on each side of these a short anterior elongate spot,
followed by three long fascize on each lateral area, fuscous
brown (these spots and markings are sometimes obsolete) ;
posterior abdominal segmental margins red; tegmina and
wings hyaline, the venation ochraceous; tegmina with the
costal membrane and postcostal area ochraceous ; abdomen
above distinctly, finely, palely pilose; lateral pilose protu-
berances to basal abdominal segment with their margins
Mr. W. L. Distant on Homoptera. 565
fuscous ; head with a distinct longitudinal impression on each
side of ocelli.
Long., excl. tegm., ¢ 14 mm.; exp. tegm. 39 mm.
Hab. West Africa; Free Town, Sierra Leone (Z. £.
Austen, Brit. Mus.). B. E. Africa; Samburu (C. S. Betton,
Brit. Mus.).
Gymnotympana olivacea, sp. n.
Body olivaceous ; sternum, opercula, and legs a little paler;
head with the anterior margin of front piceous; anterior
tibiz brownish, striped with fuscous, apices of tarsi fuscous ;
tegmina and wings hyaline, the venation olivaceous; costal
membrane of tegmina olivaceous; rostrum just passing the
anterior tibize, its apex fuscous ; opercula in male with their
apices broadly rounded, inwardly somewhat obliquely sinuate.
Long., excl. tegm., ¢ 2 26-30 mm.; exp. tegm., ¢ 70-78,
2 90 mm.
flab. Brit. Centr. New Guinea; Dinawu, 3600 feet (Pratt).
Genus PLAUTILLA.
Plautilla, St&l, Ofv. Vet.-Ak. Forh. 1865, p. 155; Hem, Afr. iv. p. 2
(1866).
Type, P. stalagmoptera, Stal.
By the kindness of Dr. Handlirsch of the Vienna Museum,
I have been able to examine a cotype, determined by Stal
himself, of P. stalagmoptera, the type of the genus.
¢g. The tympana are entirely uncovered; the opercula in
male large and convex, inflated, and projecting beyond the
lateral margins of the abdomenas in the genus Gymnotympana ;
the tegmina are moderately short and broad, their greatest width
being a little less than half their length and half as broad
again as the wings; tegmina with eight apical areas, bases of
the upper vein to lower ulnar area and lower vein to radial
area fused ; pronotal lateral marginal areas angularly dilated,
thus resembling Zammara, to which Stal allied Plawtilla, but
which belongs to a different subfamily, by the complete
absence of tympanal coverings ; head (including eyes) half the
width of base of mesonotum; abdomen beneath (excluding
apical segment) concave, with a central longitudinal ridge and
the lateral margins recurved.
This genus clearly forms the type of a new division, which
may be called Plautillaria, to be placed between the Melam-
psaltaria and Hemidictyaria. To the first it is allied by the
venation of the tegmina, and to the second by the head
37*
Or
o64 Mr. W. L. Distant on Homoptera.
(including eyes) being only half the width of base of
mesonotum,
I refrain, however, from giving fuller characters, as_there
are apparently other genera to be included and a divisional
diagnosis must not be too narrow or exclusive. Thus the
Cicada humeralis, Walk. (Insect. Saund., Hom. p. 15, 1858),
evidently represents an allied but undescribed genus to
Plautilla, but the unique male type has its abdomen mutilated,
and is therefore not in a condition for generic description.
Fam, Fulgoride.
Subfam. /vreorrm2.
Genus Pyrops.
Pyrops, Spin. Ann. Soc. Ent. Fr. viii. p. 231 (1839); Dist. Tr. Ent.
Soc. 1893, p. 448.
Zanna, Kirk. J. Nat. Hist. Soc. Bomb. xiv. p. 47 (1902).
Pyrops Rendalli, sp. n.
Allied to P. pustulatus, Gerst., from which it differs by
the longer, more robust, and obliquely ascending head ;
colour of tegmina and wings as in Gerstaecker’s species.
I have had in my possession for the last ten years a
mutilated specimen (wanting the abdomen and legs), given
me by Dr. P. Rendall, who captured it at Zomba; in
working over the genus I am now compelled to shortly
describe it as above.
Exp. tegm. 72 mm.
fab. B. C. Africa: Zomba (Dr. P. Rendall, Coll. Dist.).
Length of head (P. pustu/atus, Gerst.) 7-75 mm.; apex
of cephalic process slightly depressed.
Length of head (P. Rendall’, Dist.) 10 mm.; apex of
cephalic process obliquely erect.
Pyrops pulmunculus, sp. n.
Head about twice longer than abdomen, somewhat shining
brown, obscurely mottled with olivaceous, more prominently
so beneath and on apex above, basal area rugose; pro-
notum olivaceous, mottled with brown, rugose ; mesonotum
brownish ochraceous, rugosely punctate ; abdomen brownish
ochraceous, the apical appendage piceous; legs and sternum
much shaded with olivaceous, legs with obscure and irregular
fuscous markings; abdomen beneath greyish, thickly fuscously
punctate; tegmina greyish, with closely reticulate sanguineous
and fuscous veins, not extending to the apical area, which is
Mr. W. L. Distant on Homoptera. 565
a little paler, with more regular transverse veins, costal edge
finely hirsute; wings dull pale ochraceous, the apex very
broadly and the posterior margin moderately pale fuscous ;
head much longer than space between its base and apex of
abdomen.
Long. head, from angle of apex to eyes 17 mm.; eyes to
apex of abdomen 134 mm.; exp. tegm. 44 mm.
Hab, Java: Surabaya (Brit. Mus.).
The principal characters of this species are its very long
cephalic process and the distinctive coloration of the tegmina
and wings.
Pyrops tapirus, sp. n.
Head, pronotum, mesonotum, body beneath, and legs
ochraceous ; head with Jarge coarse black punctures, most
numerous on basal halves of Jateral areas, less numerous on
basal third of upper surface, on apical half of upper surface
two on each outer carination, beneath with about six in sub-
longitudinal series on basal half; body beneath with black
punctate spots, most conspicuously marked on lateral areas
of prosternum, clypeus, and under surface of abdomen ; legs
spotted with black, anterior and intermediate tarsi black ;
abdomen above testaceous, posterior segmental margins
ochraceous, apex piceous, and somewhat longly ochraceously
pilose ; tegmina pale ochraceous, the venation a little darker,
and the whole surface somewhat thickly sprinkled with
small black spots, some a little larger than others; wings
creamy white; the cephalic process is robust and distinctly
slightly recurved downwards at apex, which has its face
marginately but not centrally carinate; the apex of the
rostrum is black and reaches the intermediate coxe; the two
outer longitudinal carinations to the cephalic process both
above and beneath are much waved.
Long. head, from angle of apex to eyes 12 mm.; eyes
to apex of abdomen 19 mm.; exp. tegm. 60 mm.
Hab. Java (Horsfield Coll., Brit. Mus.).
A species to be recognized by the robust cephalic process,
apically recurved, and distinctly and peculiarly maculately
punctured with black.
Pyrops Dalyi, sp. n.
Body and legs pale brownish ochraceous ; cephalic process
finely and thickly blackly punctate on lateral areas, a little
more finely so beneath, and very obsoletely so on discal area
above, the two outer carinz beneath with about six sub-
tubercular reddish-ochraceous spots; clypeus, sternum, and
566 Mr. W. L. Distant on Homoptera.
legs finely blackly punctate, femora with an apical and
anterior and intermediate tibie with two incomplete black
annulations ; abdomen beneath with black spiracular spots,
abdomen above with the anterior segmental margins more or
less piceous; tegmina greyish white, the venation testaceous,
the whole surface more or less sprinkled with minute shining
black spots, some a little larger than others; wings creamy
white, the venation very pale ochraceous; head horizontally
porrect, about as long as space between its base and apex of
abdomen, gradually attenuated anteriorly, its face with its
margins and a central longitudinal line carinate.
Long. head, from angle of apex to eyes 11} mm.; eyes to
apex of abdomen 12 mm.; exp. tegm. 34 mm.
flab. Siam: Lacan (Malcolm Daly, Brit. Mus.).
A small species, to be identified by its long, horizontally
porrect, and anteriorly narrowed cephalic process, with its
bright subtubercular reddish-ochraceous spots beneath.
Genus FULGORA.
Fulgora, Linn. Syst. Nat. ed. xii. i. (2) p. 703 (1767).
Hotinus, Amy. & Serv. Hém. p. 490 (1843).
Pyrops, Kirk. J. Bomb. Nat. Hist. Soc. xiv. p. 47 (1902).
Fulgora siderea, sp. 0.
Body and legs ochraceous ; abdomen above thickly creta-
ceously tomentose ; cephalic process above somewhat thickly
covered with minute white spots, beneath unspotted ; base of
head and a short longitudinal fascia not extending beyond
the latitude of eyes, two spots between eyes, a spot in front,
another behind, and the posterior margins of eyes black ;
pronotum with a black spot near each anterior angle ; meso-
notum with six spots arranged in pairs (the two central
largest) on anterior margin and a large oblique spot on each
lateral margin black; rostrum, anterior and intermediate
tibie, all the tarsi, and transverse fasciz to abdomen beneath
black; tegmina greenish ochraceous, thickly interspersed
with small black maculate markings, four large black spots
on costal membrane, a number of large ochraceous spots
margined with white arranged in somewhat transverse but
much broken series, some of the spots united; wings creta-
ceous white, the upper area (including apex) purplish red,
and containing some irregular black apical spots ; cephalic
process measured from angle of apex to eyes about as long
as from middle of mesonotum to apex of abdomen, nearly its
apical half upwardly recurved, its apex broadly laterally
compressed,
Dr. T. Scott on British Copepoda, 567
Long. ceph. process, from angle of apex to eyes 17 mm. ;
eyes to apex of abdomen 24 mm. ; exp. tegm. 76 mm.
Hab. Borneo (Brit. Mus.).
I have for many years had an unlocalized Specimen in my
own collection.
Fulgora evanescens, sp. n.
Body and legs ochraceous; rostrum, anterior and inter-
mediate tibie, and all the tarsi piceous ; abdomen beneath
with small, scattered, fuscous spots ; cephalic process marked
above and beneath with very small and obscure pale spots ;
tegmina ochraceous, with darker suffusions on apical area
and four or five large darker spots in costal membrane ; there
are also a number of scattered obsolete paler spots with dark
centres, some of which are united ; wings pale hyaline, with
an obscure, pale fuscous, curved, fasciate spot above the apex
of radial area; cephalic process long and slender, only
slightly upwardly recurved, and from its apex to eyes as long
as from eyes to apex of abdomen; rostrum passing the
posterior coxee.
Long. ceph. process, from angle of apex to eyes 19 mm.
eyes to apex of abdomen 19 mm. ; exp. tegm. 70 mm.
Hab. Nias Island.
Allied to #. lauta, Stal.
LXIII.—Notes on British Copepoda: Change of Names.
By Tuomas Scort, LL.D., F.L.S.
In the ‘ Fifteenth Annual Report of the Fishery Board for
Scotland’ (1897), part i. p. 150, I described a species
belonging to the Copepoda under ‘the name of Delavalia
mimica ; but though it exhibited a close affinity with the
genus Delavalia, especially in the structure of the mandibles
and of the fifth pair of thoracic feet, the structure of the
first pair more nearly resembled those of Nitocra or Ametr "ly
the inner branches of that pair being composed of three
instead of two joints. In my remarks on the species [
referred to this difference ; but because the species had such
a general resemblance to Delavalia it was provisionally
ascribed to that genus.
During recent years many specimens of this species, both
from the Scottish and English coasts, have been examined,
568 Dr. T. Scott on British Copepoda.
and they all agree in showing the marked difference in
structure of the first pair of thoracic feet mentioned above, and
I now propose to remove this species to another genus rather
than modify that to which it was provisionally ascribed.
The following is the name and definition of the genus :—
BEATRICELLA *, T. Scott, gen. nov.
Definition of the genus.—Similar to Delavalia,G. 8. Brady,
in general appearance. Antennules eight-jointed. Outer
ramus of the antenne three-jointed. Mandibles as in
Delavalia, the terminal branch of the mandible-palp being
furnished with a long, curved, and moderately stout seta.
Other mouth-appendages as in Delavala.
Both branches of the first pair of thoracic feet three-jointed ;
inner branches longer than the outer ones, first joint longer
than or subequal to the second and third joints. The second
to the fourth pairs as in Delavalia. Fifth pair foliaceous
and somewhat similar to those of the genus mentioned.
Beatricella mimica, T. Scott.
1897. Delavalia mimica, T. Scott, Fifteenth F. B. Report, part iii.
p. 150, pl. i. figs. 1-9.
In this species the first joint of the inner branches of the
first pair of thoracic feet are as long as the entire length of
the three-jointed outer branches, but the second and third
joints are very short, being together scarcely half as long as
the first joint. The fifth pair resemble those of Delavalia
reflexa, G. 8. Brady, but the basal joint bears interiorly a
group of three tolerably short and stout spines and a small
spiniform seta. The fifth pair in the male are smaller than
those of the female ; the basal joint is armed interiorly with
a single moderately large and broad spine of a somewhat
peculiar shape; it is broad and flattened rather than round,
and with the sides parallel except near the distal end, where
they rapidly converge and form a triangular extremity; a
minute seta springs from each of the lateral angles at the
distal end, while the apex is extended to a fine point; the
secondaiy joint is small, subovate in form, and bears three
spines on the oblique distal end of the outer margin, a
moderately long seta at the apex, and a smaller one on the
inner margin.
* Named in compliment to Miss Beatrice Sprague, daughter of
Dr. T. B. Sprague, Edinburgh,—a successful student of the Scottish
freshwater Entomostraca.
Dr. T. Scott on British Copepoda. 569
Remarks.—In Beatricella mimica the first pair of thoracic
feet have a close resemblance to those of certain species of
Dactylopusia and Ameira ; hence the specific name.
Habitat.—Firths of Forth and Clyde, Moray Firth, vicinity
of Plymouth, &c., but not very common *,
Delavalia Normani, T. Scott, nom. nov.
1899. Delavalia Giesbrechti, var., T. Scott, Seventeenth F. B. Report,
part iil. p. 254, pl. xiii. figs. 20-22.
A few specimens of a Delavalia were obtained at Hunterston,
Firth of Clyde, in the autumn of 1898 ; they resembled to
some extent a species previously described by T’.. and A. Scott
under the name of Delavalia Giesbrechti, but differed in one
or two points, and notably in the absence of the peculiar tail-
setee which are so characteristic of D. Giesbrechti; but as it
was considered that the differences observed might, to some
extent at least, be due to difference in haddtat, and as only
one or two specimens had been noticed, they were recorded
simply as “ Delavalia Giesbrechti var.”
Subsequently, however, it was ascertained that the Rev.
Canon A. M. Norman had collected the same form at
Salcombe, South Devon, in 1875, and again in 1903, as well
as at other places on the coasts of Devon and Cornwall.
The examination of these additional specimens showed that
the differences previously observed were constant and not due
to difference in local surroundings ; this form should there-
fore be regarded as a distinct species, and the name I propose
for it is Delavalia Normant.
In this species the two-jointed inner branches are only
slightly longer than the outer, the first joint is robust and
rather more than half the length of the second, while the
second is not only more elongated than the first, but is also
distinctly narrower and tapers slightly towards the distal
extremity ; the apical seta, which is fully longer than the
second joint, is moderately stout and spiniform, and there are
three setze on the inner margin of the second and one on the
first joint.
In the fifth pair in the female the basal joint is short and
furnished interiorly with four tolerably long plumose sete
along its lower edge; they are arranged in pairs, one pair
being near the inner angle, the other close behind the
* A second species—Delavalia (Beatricella) emula, T.Scott,—in which
the three joints of the inner branches of the first pair of thoracic feet are
nearly of equal length, is described in the Eleventh Report of the Fishery
Board for Scotland, pt. iii. p, 204, pl. iv. figs, 36-47 (1893),
570 Dr. T. Scott on British Copepoda.
secondary joint, so that there is a comparatively wide space
between the two pairs. The secondary joint, which is
lamelliform and with nearly parallel sides, is about one and a
half times longer than broad, the width slightly increasing
toward the distal end; it is provided with five sete, one on
the outer margin and four round the distal extremity ; the
marginal seta, which springs from a notch on the outer edge,
is stout and spiniform ; the outermost and innermost of the
four apical setee are of moderate length, but the other two
are short.
The fureal joints, which are rather shorter than the last
abdominal segment, are comparatively widely apart and also
somewhat divergent.
THOMPSONULA, T. Scott, gen. nov.
My friend the late Mr. I. C. Thompson, of Liverpool,
whose interesting discoveries have added so much to our
knowledge of the Copepod fauna of the British Islands,
described in 1899, under the name of Jonesiella hyene, a
somewhat curious form that had recently been observed in
some collections from Port Erin Bay, Isle of Man. This
form was also subsequently obtained in the Firth of Forth
and at some other places around our shores, and its distribu-
tion as now known is fairly extensive.
Although this form was included in the genus Jonesiedla,
G. &. Brady, it differs from the typical members of that
genus in one or two important particulars, and especially in
the structure of the first pair of thoracic feet. In the generic
definition given by Professor Brady the inner branches of the
first pair are described as “ two-jointed, and bearing long
terminal sete, but no claws”; in Jonesiella hyene, on the
other hand, the inner branches are distinctly three-jointed.
In my remarks on this species in the ‘Kleventh Annual
Report of the Fishery Board for Scotland,’ part iil. p. 203
(1893), it is stated that ‘‘ these differences may render it
necessary to modify the generic description or to remove
Jonesiella hyene to another genus” *, I now propose to
adopt the latter of these suggestions, and have named this
new genus after my late friend.
Definition of the genus Thompsonula—Somewhat similar
* It is interesting to note that Mr. I. C. Thompson also realized the
difficulty referred to here, but was inclined “ to slightly modify an existing
genus rather than make a new one.”
Mr. C. T. Regan on Hexanchus griseus. 571
to Jonesiella in general appearance. Antennules (anterior
antenne) short and composed of six joints. Antenne (poste-
rior antenne) with the outer ramus well developed, three-
jointed, middle joint short. Mandibles distinctly two-branched.
First pair of thoracic feet with both branches three-jointed,
the inner being more elongated than the outer branches ;
other thoracic feet somewhat similar in structure to those of
Jonesiella.
Thompsonula hyene (I. C. Thompson).
1889. Jonesiella hyene, I, C. Thompson, Proc. Biol. Soc. Liverpool,
vol, iii. p. 193, pl. ix. figs. 1-10.
1893. Jonesiella hyene, T, Scott, Eleventh F. B. Report, pt. iii. p. 202,
pl. ili. figs. 1-6.
In this species the basal joint of the short six-jointed
antennules is very stout and comparatively large; it extends
forward, while the remaining joints curve outwards, so that
the last three are nearly at right angles to the basal joint.
The rostrum, which extends forward between the basal joints
of the antennules, reaches to beyond the extremity of these
joints. The furcal joints are extremely short, and as the
last abdominal segment is tolerably broad, the posterior has
an abruptly truncated appearance.
Further particulars concerning this interesting species will
be found in the papers referred to above.
LXIV.—WNote on Hexanchus griseus.
By C. Tate Reean, B.A.
Tue British Museum has recently received, from Captain
Alan Owston, a shark of the genus Hewvanchus, captured in
Japanese seas.
The specimen measures 2100 mm. in total length, and for
comparison with it I had a small example (600 mm.) of
Hexanchus corinus, Jordan and Gilbert, from the Pacific
Coast of the United States, as well as specimens of the
Atlantic H. griseus.
The Japanese fish closely resembles the one named
H. corinus in most respects, but differs notably in the dentition.
In the lower jaw the median tooth has about 4 distinct cusps
on each side and 3 minute ones on the upper edge, whilst the
lateral teeth have from 8 to 10 cusps each. In the upper jaw
572 Mr. O. Thomas on a new Genus and
the median pair of teeth are slender and unicuspid, the next
pair have a minute basal cusp, which becomes well-developed
and pointed on the next 3 or 4 on each side. In the smaller
fish (A. cor’nus) the lower median tooth has 2 or 3 cusps on
each side and none on the upper edge, whilst each lateral
tooth has only 6 cusps ; the second pair of teeth in the upper
jaw are unicuspid, and the next 3 have only a very small and
obtuse basal cusp.
In the larger shark the fins are relatively less developed
and consequently they are more widely separated from each
other. The anal begins below the last } of the dorsal and
the length of its base is less than its distance from the caudal,
which fin is ? the total length of the fish, whereas in the
smaller example the origin of the anal is only a little behind
the middle of the dorsal, the length of its base is equal to its
distance from the caudal, which is more than 4 the length of
the fish.
Examination of a series of examples from the Mediter-
ranean and the Atlantic shows conclusively that these
differences are due to the size of the fish and are not specific.
The British Museum has large specimens agreeing in denti-
tion and the position of the fins with the Japanese example,
and smaller ones which resemble the Californian fish in these
respects. Moreover, after a careful comparison, I am unable
to find any reason for referring the Pacific and Atlantic
specimens to different species, and I am forced to conclude
that there is but one living species of Hewxanchus, viz.
H. griseus, L., which has a wide distribution.
LXV.—A new Genus and Two new Species of Bats.
By OLpFIELD T'HOMAS.
Homops, gen. nov.
In 1900 * Dr. Scharff described a bat from Benin under the
name of Mormopterus Whitleyi and was good enough to
transfer the typical specimen to the British Museum.
Later on Dr. W. J. Ansorge obtained on the Lower Niger
two examples of the same bat, and in examining these I find
that a mistake has inadvertently been made in the dental
formula given, and that, instead of being a Mormopterus, this
* Ann, & Mag. Nat. Hist. (7) vi. p. 569.
g Pp
Two new Species of Bats. 573
bat belongs to quite a different group, hitherto unknown in
the Old World.
For it proves to have the characteristic incisors of Molossus,
the upper pair being large, nearly filling up the space between
the canines, and touching each other in the middle line. ‘The
lower ones are only two in number, not four as_originally
stated, and while appearing from the front to be of the normal
shape—¢. e., narrow, broadening upward, each with a deep
central notch dividing it into two cusps, of which the inner is
the higher—they are curiously deep antero-posteriorly, deeper
than broad, their hinder part running back between the
canines.
The premaxillary region of the palate is very completely
ossified, in great contrast to all the members of the Nyeti-
nomus group, the enly remnant of the premaxillary notch
being two minute foramina on each side of the middle line,
between and opposite the centre of the canines.
The skull is smooth and rounded, not broadened and
flattened across the muzzle as it is in M/yopterus, nor specially
crested as in Molossus. ‘The base of the skull has a narrow
median ridge between two well-defined sphenoid pits.
Under these circumstances it is clear that this bat cannot
be referred to Nyctinomus or Mormopterus, but is more nearly
allied to Molossus, Myopterus, and their allies. The peculiar
shape of its lower incisors, however, and the detailed structure
of its skull prevent its being referred to any one of the
existing American genera, and I would therefore propose to
erect for its reception a special genus, which might bear the
name of Homops.
It is also to be noted that, owing to the bad condition of
his specimens, Dr. Scharff described the underside of Homops
Whitleyt as “of a light reddish-yellow colour”; but
Dr. Ansorge’s fresh examples show that this is pure white,
as are also the wing-membranes above and below, the digits,
and the underside of the interfemoral ; the upper surface of
the forearms, tibiz, and interfemoral are pale whitish
brown.
Vespertilio Matschiet, sp. n.
A small sandy-coloured species, in general external appear-
ance very like Scotophilus Schlieffeni.
Size small. ur of medium length, hairs of back about
4 mm. long; strictly confined to the body, not extending on
to the limbs or membranes, except slightly at the base of the
574. Mr. O. Thomas on a new Genus and
tail. General colour of fur uniform sandy buff above, rather
paler below, strongly contrasting with the blackish ears and
wings; the hairs unicolor throughout. Ears rather short ;
inner margin strongly convex at base, almost straight above
to the rounded tip; outer margin flattened mesially, convex
below and ending in a long low antitragal lobe. Tragus
short, its inner margin slightly concave, tip rounded, outer
margin convex with a well-marked rounded basal lobule.
Wings to the metacarpus. Extreme tip of tail projecting.
Skull slender, with a small brain-case, but widely expanded
zygomata. Muzzle remarkably flattened, the space between
the orbits quite flat or even concave, the strong supraorbital
ridges rising up higher than its central line; as a result the
upper profile-line of the skull is decidedly concave in its
anterior half, only becoming gradually convex over the brain-
case. Nasal notch sharply angular behind.
Inner upper incisor conical, unicuspid in the type, but a
supplementary cusp may have been worn off; outer incisor
similar in shape, about one fourth the size of the inner.
Lower incisors crowded, overlapping, tricuspid, the outer
notch on each deeper than the inner one.
Dimensions of the type (measured on the skin) :—
Forearm 85 mm.
Head and body (c.) 41; tail (c.) 32; third finger, meta-
carpus 35, Ist phalanx 11, 2nd phalanx 13; tibia 14.
Skull: greateat length 12°3 ; basal length in middle line
10; front of canine to back of m’ 4°5; front of lower canine
to back of mz 4°7.
Hab. Jimel, near Aden. Alt. 850 m.
Type. Male. B.M. no. 99. 11. 6. 19. Collected 16th
August, 1899, by W. Dodson. One specimen.
This little bat has so remarkable a resemblance to Scoto-
philus Schliefent that it was included among the specimens
of that species recorded in my paper on the Dodson mammals
from Aden*. Now, however, that its skull has been looked
at it proves to be a Vespertilio, in which genus its peculiar
colour distinguishes it from all described species except
V. Flowert t, from which its smooth forearms and unusually
flattened muzzle readily separate it.
I have named this interesting species in honour of my
friend Dr. Paul Matschie, of the Berlin Museum, whose notes }
* P. Z. S. 1900, p. 99.
+ Glauconycteris Floweri, de Wint. Ann. & Mag, Nat. Hist. (7) vii.
p. 45 (1901).
} SB. Ges. nat. Fr. Berl. 1893, p. 26.
Two new Species of Bats. 579
on Scotophilus Schlieffent and other mammals from Aden
formed the first contribution to the mammalogy of that
region.
Flesperoptenus Tomesi, sp. n.
A large species allied to H. Tickelli, but wholly dark brown,
like a Noctule.
Size about as in H. Tickelli, the skull rather larger but the
wing-bones shorter. Fur close and fine, the hairs of the
back about 4-5 mm. in length, but those of the nape much
longer, attaining 8-10 mm. Base of interfemoral slightly
haired above and below, rest of the membranes naked.
General colour above uniform dark brown (“ burnt umber,”
but darker), the hairs darkest at their bases, rather lighter
mesially, and with dark tips, but the difference is scarcely
perceptible. Under surface similarly uniform brown (about
matching Ridgway’s burnt umber) throughout. Wing-
membranes everywhere dark brown.
Ears short, rounded above, with a smali rounded basal
lobule at the base of their inner margin ; outer margin ending
in a long low antitragal lobule. Tragus very broad, its
inner margin faintly concave, tip pointed; outer margin
very convex, strongly bowed opposite the base of the inner
margin ; outer basal lobule rounded ; a narrow band of fine
hair passing across the front face of the tragus at its broadest
point. Wings to the end of the metatarsus. Postcalcareal
lobule narrow. Penis long, with a very long bone, similarly
Y-shaped to that of H. Tickell’, but twice as long (10 mm.
in length’in the type) and much thicker; whether the glands
and prepuce are similar to those of H. Tickelli cannot be
determined on the dried type.
Skull rather larger than that of ZZ. Tickell’, the sagittal
crest rather lower, but the posterior “ helmet ” more projected
backward.
Teeth larger and heavier than those of H/. Tickelli. I
large in section, its postero-external side almost touching the
canine outside 2*, which is also very large, placed quite
behind 7*, opposite the middle of the canine, and is separated
from the inner corner of the premolar by a distance only equal
to its own diameter. Premolar much extended transversely.
Below, the incisors are all heavier than in H. Tickelli, 73 being
patticularly large.
Dimensions of the type (measured in skin) :—
Forearm 51 mm.
Head and body 74; tail (vertebre in place) 44; ear 14;
ys Bibliographical Notice.
tracus, inner edge 5, breadth 3; third finger, metacarpus 45,
Ist phalanx 22, 2nd phalanx 22; fifth finger 58; lower leg
and foot (c. u.) 32.
Skull: greatest length 20°5; zygomatic breadth 15°5;
occiput to nasal notch 17; front of canine to back of m* 8:3.
Hab. Malacca.
Type. Adult male. Original number 190 A. From the
collection of the late Mr. R. F. Tomes.
This fine bat, which is distinguishable at the first glance
from its only near ally, H. Tickedli, by its uniform dar
colour, was recognized by Mr. ''omes as a new species, and
marked by him with a name which I would have adopted
did it go euphoniously with the long generic name Llespero-
ptenus. Bat as it does not I am glad to commemorate
Mr. Tomes’s great interest in bats, and the work he did on
them, by naming this species in his honour.
BIBLIOGRAPHICAL NOTICE.
Coloration in Polistes. By Witurtmine M. Enreman. Published
by the Carnegie Institution of Washington, November 1904.
Sm. 4to. Pp. 88, col. pls. vi., and 27 figures in the text.
Poxistes is a handsome and widely-distributed genus of wasps
allied to Vespa; it is, however, unrepresented in Britain, and only
one species (P. gallicus, L.) is found in Europe, though this has
been met with as far north as Lapland. After a general account of
the genus the present essay records the results of a careful investi-
gation into individual variation in the colour-pattern of Polistes ;
ontogenesis of the colour-pattern ; physical and chemical nature of
the pigment; geographical distribution of the types of colour-
marking; laws governing colour-differentiation ; and considerations
with respect to various theories of evolution. Two of the plates
are devoted to colour-patterns, two to figures of various species of
the genus, and two to maps showing distribution. The parallel
drawn between Eurasian and American variation in the genus is
specially interesting.
We also meet with occasional references to colour-patterns in
other Vespide, as well as in butterflies &c. The authoress’s con-
clusions are finally summed up under fourteen heads, too long to
quote here, and we must confine ourselves to calling attention to a
work which should not be overlooked by biologists (whether specially
hymenopterists or not) who are interested in coloration, geographical
distribution, and the other factors bearing on evolution which are
here discussed.
THE ANNALS
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. 96. DECEMBER 1905.
LXVI.—Descriptions of new Species of Noctuide in the
British Museum. By Sir Grorce F. Hampson, Bart.,
PAS: Se:
{Concluded from p, 549. |
Genus EL#0GNATHA, Nov.
Type H. argyritis.
Proboscis absent ; palpi slender, the second joint reaching
vertex of head and with tuft of hair on inner side at
extremity, the third long and somewhat dilated at extremity ;
frons smooth, rounded; eyes large; antennx of female
laminate; head and thorax clothed chiefly with scales, the
pro- and metathorax with spreading crests; fore tibix
fringed with long hair on outer side; abdomen with dorsal
crest at base and slight subdorsal tufts of hair. Fore wing
elongate, the inner margin strongly arched, the apex rounded
and the termen obliquely curved; vein 2 from middle of
cell, curved at base; 3 and 5 from close to angle of cell; 6
from upper angle or from areole, which is rather long and
formed by 9 from 10 anastomosing with 7, 8; 11 from cell.
Hind wing with veins 3 and 5 stalked, 4 absent; 6, 7 stalked ;
8 anastomosing with the cell to near extremity.
Eleognatha argyritis, sp. n.
9. Head and thorax silvery white ; palpi with the second
joint dark brown at sides; tips of tegule and patagia dark
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 38
578 Sir G. F. Hampson on
brown; fore tibia and tarsi banded with fuscous; abdomen
white, dorsally tinged with brown. Fore wing silvery white ;
some black points at base; silvery-blue streaks in base of
submedian fold and above base of inner margin, black at
extremities ; a waved black subbasal line from costa to just
above inner margin ; a large rufous patch on middle of costa,
defined on inner side by the antemedial line, which is angled
outwards on subcostal nervure and ends at submedian fold ;
medial black striz from cell and vein 1 ; a quadrate black spot
in middle of cell and obliquely curved discoidal bar, with
two slight oblique marks beyond it ; postmedial line irregu-
larly waved, oblique from costa to vein 6, then inwardly
oblique to vein 2, retracted to cell, then excurved and angled
inwards ih submedian fold, a brown patch beyond it from
costa to vein 6, constricted at middle and with oblique bar
from its lower edge; subterminal line rather slight and
punctiform, angled outwards at vein 7 and inwards below 7
and 3, some brown beyond it at apex and an oblique streak
from it to termen above tornus; a terminal series of black
points. Hind wing pale fuscous.
Hab. Panama, Cana Mines (Tylecote), 1 9 type.
Hep. 30 mm.
Eleognatha pheostrota, sp. n.
@. Head and thorax clothed with silvery-white, reddish-
brown, and black scales; palpi with the second joint blackish
at sides; pectus and legs grey, the fore and mid tarsi
fuscous above; abdomen grey, dorsally suffused with reddish
brown. Fore wing grey, irrorated with black and suffused
with dark brown except on terminal area; a very indistinct
antemedial series of dark spots; a black pomt in middle of
cell and another on discocellulars ; postmedial line bent
outwards below costa, then somewhat dentate, incurved
between veins 5 and 8, then retracted to the cell, bent
outwards between veins 2 and ], then retracted to before
middle of wing and bent outwards again to inner margin ;
subterminal line fuscous defined by white on inner side,
minutely waved, angled outwards at veins 7, 6, retracted at
vein 3 and ending at tornus ; a terminal series of prominent
black spots. Hind wing dark reddish brown ; cilia whitish
at tips.
Habe Panama, Cana Mines (Tylecote), 1 2 type. Exp.
36 mm.
new Species of Noctuide. 579
Iscadia diopis, sp. un.
Iscadia aperta, Druce, Biol. Centr.-Am., Het. ii. p. 474 (part.).
2. Head and thorax clothed with fuscous, red-brown, and
grey scales; palpi with the second joint white in front, black
at sides except at tips; pectus and legs grey-brown; abdo-
men fuscous brown, the ventral surface whitish. Fore wing
grey irrorated with black and suffused in parts with dark
red-brown ; some rufous suffusion at base followed hy dark
red-brown except towards costa ; antemedial line indistinctly
double, waved, strongly excurved below cell, followed by
some dark red-brown suffusion ; orbicular and reniform round
brownish ocelli defined by fuscous and with spots of raised
black scales in centre ; an indistinct irregular medial line
angled outwards below costa; postmedial line double, the
outer line less distinct, irregularly waved, strongly bent
outwards below costa, angled inwards at discal fold and bent
inwards at vein 2 to below end of cell, some dark brown
suffusion beyond it at discal fold and on inner area; sub-
terminal line grey, defined on inner side by slight dentate
dark marks and on outer by dark brown suffusion, minutely
dentate, angled outwards at vein 7 and excurved at middle ;
a series of small black marks just before termen connected
by short streaks with a terminal series of small lunules.
Hind wing fuscous brown ; cilia greyish at tips.
Hab. Costa Rica, Sta. Clara Valley, 1200 feet (Zurcher),
1 ¢ type. Hap. 50 mm.
Tscadia poliopepla, sp. n.
?. Head and thorax grey, mixed with dark brown ; palpi
with the second joint white in front, black at sides except at
tips ; vertex of head blackish; tegulze with black medial
line; abdomen grey, the dorsal crests blackish at tips.
Fore wing grey irrorated with black and suffused with brown
except on basal and terminal areas, the basal area whiter; an
obliquely curved dark antemedial line ; a diffused dark line
forming the inner edge of the brownish area; a black point
in end of cell and two black discoidal points on a faint
whitish bar; postmedial line obliquely excurved from costa
to vein 4, then strongly incurved and sinuous, angled inwards
just below submedian fold, followed by another minutely
dentate line bent inwards below vein 3; subterminal line
white, very irregular, oblique at costa, angled inwards below
vein 7 and at discal fold, interrupted and bent inwards to
the second postmedial line and vein 2, where it forms a
38*
| 580 Sur G. I. Hampson on
whitish mark irrorated with black, then oblique and sinuous
to tornus, some short black streaks beyond it towards apex ;
a series of prominent black points just before termen
forming two striez above tornus. Hind wing white, the
veins and marginal areas slightly tinged with brown; the
underside with brown patch on costa before apex.
Hab. Braz, Rio Janeiro (EL. Wilson), 1 2 type. Hap.
44 mm.
Plotheia nephelozona, sp. un.
9. Head and thorax grey, tinged with pale red-brown ;
abdomen whitish. Fore wing grey, tinged with pale red-
brown and slightly irrorated ‘with black; a broad diffused
irregular medial fuscous band expanding on costa towards
apex and on inner margin towards base, with a very small
elongate tuft of raised scales on it in middle of cell; two
minute black discoidal points ; faint traces of a subterminal
line; a very slight brown shade on apical part of terminal
area; a terminal series of minute black points. Hind wing
semihyaline white; the termen slightly tinged with brown
towards apex and with some slight points at middle; the
underside with the costal area slightly tinged with brown.
Hab. Br. KE. Arrica, Kikuyu, Nairobi plains (Crawshay),
1 9 type. Exp. 30 mm.
Plotheia canofusa, sp. n.
?. Head, thorax, and abdomen grey-white slightly mixed
with brown ; fore tarsi tinged with fuscous. Fore wing pale
brownish ochreous suffused with grey-white on basal and
costal areas and slightly irrorated with fuscous; antemedial
line indistinctly double, oblique, waved, angled inwards in
submedian fold and on vein 1 and retracted to inner margin,
towards which there is some fuscous on its inner side; an
oblique pale fuscous shade from costa beyond middle to
antemedial line below the cell; two minute black discoidal
spots ; postmedial line double, bent outwards below costa,
then minutely waved, excurved to vein 4, then incurved to
near antemedial line, some grey points beyond it on costa ;
subterminal line faint, dark, oblique from costa to vein 7
and with black spot on it above tornus; a terminal series of
slight black striz. Hind wing semihyaline white, the costal
and terminal areas tinged with brown; a fine dark terminal
line; cilia white, tinged with brown at apex.
Hab. Care Corony, Aunshaw (Miss F. Barrett), 1 2 type.
Exp, 28 mm.
new Species of Noctuide. 581
Plotheia ferrofusa, sp. u.
d. Head and thorax pale grey, the sides of palpi, head,
and thorax more or less deeply tinged with ferruginous ;
fore legs tinged with ferruginous in front; abdomen pale
grey slightly tinged with brown on dorsum. Fore wing pale
grey, the basal inner area, medial area except at inner
margin, postmedial area at middle, and terminal area except
at apex and tornus suffused with bright ferruginous ; a small
fuscous spot below costa near base and subbasal spots below
costa and in cell; a double slightly waved antemedial line,
strongly excurved from costa to submedian fold, then angled
inwards on vein 1; a small tuft of whitish scales in middle
of cell and small black discoidal tuft ; postmedial line double,
arising near antemedial line, very strongly bent outwards
below costa, oblique to veins 4, 3, where it is minutely den-
tate, then bent inwards to vein 2 and erect to inner margin ;
subterminal line whitish defined by dark brown on inner
side, minutely waved, excurved below costa and at middle ;
a fine blackish terminal line slightly defined by grey on inner
side. Hind wing semihyaline white, the termen narrowly
reddish brown.
Hab. Manas, Belgaum (Watson), 1 ¢; Cuyton, Colombo
(Green), 1 g type. Hap. 20 mm.
Plotheia leucogonia, sp. n.
?. Head and thorax grey slightly tinged with red-brown ;
patagia with brown line near tips ; abdomen grey, the crests
dark at tips. Fore wing with the basal and inner areas grey
slightly tinged with red-brown, the rest of wing ochreous
suffused with rufous, the grey area defined by a black-brown
line from costa to submedian fold, in which it forms a streak
to postmedial line, emitting a slight striga from its lower
edge before middle; a slight oblique red-brown discoidal
bar ; postmedial line indistinctly double, very oblique from
costa to vein 6, then excurved, slightly angled inwards in
discal fold, at vein 2 retracted to near cell, slightly angled
outwards at submedian fold and excurved to inner margin ;
subterminal line represented by some diffused rufous, ex-
curved at vein 7 and middle; a small triangular white mark
from termen to postmedial line at vein 2; a tuft of dark
scales above tornus ; a terminal series of black striz; cilia
rufous. Hind wing white, the terminal area slightly tinged
with brown; the underside with the costal area slightly
tinged with red-brown. aes
582 Sir G. F. Hampson on
Hab. Br. EB. Arrica, Teita (Jackson), 1 2? type. Hap.
20 mm.
Subsp. 1. Head, thorax, and abdomen browner ; fore wing
with the triangular white mark rather larger; hind wing
wholly suffused with dark brown.
Hab. Sterna Leone (Clements), 1 ¢.
Subsp. 2. Fore wing without the triangular white mark ;
hind wing whitish suffused with pale brown.
Hab. Gamsia, Bathurst (Sir G. Carter), 1 9. Eap.
16 mm.
Plotheia gerea, sp. 0.
d. Head, thorax, and abdomen grey slightly tinged with
fuscous ; tegule with fine oblique dark streaks, patagia with
black npper edge and fine oblique streak across tips. Fore
wing grey slightly suffused with fuscous brown and finely
irrorated with black; obliquely placed subbasal fuscous
points in and below cell; an indistinct double antemedial
line, slightly angled inwards on median nervure and oblique
below submedian fold; a small black discoidal bar; post-
medial line rather indistinct, double, oblique from costa to
vein 3, where it is bent inwards to near angle of cell, then
oblique and slightly waved to inner margin; an indistinct
diffused brown subterminal line, excurved below costa and
at middle; a terminal series of slight dark striez. Hind
wing whitish tinged with brown especially on terminal
area.
Hab. Qurenstand, Townsville (Dodd), 2 3 type. Hap.
24 mm.
Plotheia diplographa, sp. u.
Headand thorax grey slightly tinged with fuscous; abdomen
grey-white. Fore wing grey-white suffused with pale brown ;
traces of a double subbasal line, bent outwards from costa to
subcostal nervure, then slightly sinuous ; a prominent double
black antemedial line almost at middle of wing, somewhat
oblique towards costa, incurved from middle of cell to sub-
median fold, emitting a slight black streak in submedian
fold towards postmedial line; the disk sometimes suffused
with rufous, a small brown discoidal spot; postmedial line
indistinctly double, sinuous, oblique from costa to vein 6 and
bent inwards below vein 4; an indistinct, minutely dentate,
whitish subterminal line defined on inner side by slight
dentate black marks, angled outwards at vein 7 and excurved
at middle; a fine black terminal line. Hind wing white, the
terminal area shghtly tinged with brown.
new Species of Noctuide. 583
Hab. Punsas, Kangra (Hocking), 1 ¢ ; Manras, Nilgiris
(Hampson), 1 3 type. Hap. 26 mm.
Nanaguna brunnea, sp. n.
g. Head and thorax pale reddish brown more or less
tinged with fuscous; abdomen pale reddish brown. Fore
wing pale reddish brown slightly irrorated with black, the
inner area to postmedial line and sometimes the inner half
of terminal area whitish; the costal area near base and at
middle, the cell and area just below it sometimes suffused
with black ; subbasal line represented by an inwardly oblique
striga from costa and a very oblique streak from cell to
antemedial line at vein 1, obscured in the black suffused
specimens ; antemedial line almost medial, strongly angled
outwards on subcostal nervure and in submedian fold and
inwards in cell and on vein 1; traces of a discoidal annulus ;
postmedial line with series of black points on it and defined
by whitish on outer side, very oblique from costa to vein 6,
then inwardly oblique and incurved below vein 2, where there
is some brownish or fuscous beyond it; some slight fuscous
and whitish points on costa towards apex; traces of a waved
grey subterminal line with two obliquely placed fuscous
points beyond it at vein 2 and sometimes a point on it in
submedian fold; a terminal series of black points; cilia
ochreous with series of brownish or fuscous spots. Hind
wing whitish uniformly suffused with pale reddish brown;
the underside whitish, the costal area tinged with brown.
Hab. Assam, Khasis, 1 g type; Puno Laur (Doherty),
1g. Exp. 26 mm.
Nanaguna albisecta, sp. n.
@. Head and thorax whitish, slightly tinged with brown ;
tegule at tips, slight marks on patagia, and a patch on meso-
thorax black; fore tibiz: and tarsi banded with fuscous.
Fore wing pale olive-grey, slightly irrorated with black ; some
rufous suffusion below the cell near base and below middle
of terminal area; a diffused white fascia from costa near
base through the cell and thence to apex, defined below by a
black streak ending in a black spot at lower angle of cell; a
medial black line, oblique and punctiform from costa to the
white fascia, waved below the cell; a rather punctiform black
postmedial line defined by whitish on outer side, very oblique
from costa to vein 6, then inwardly oblique, some blackish
points beyond it on costa and slight streaks from vein 6 to
submedian fold; a small wedge-shaped white spot on middle
584 ; Sir G. F. Hampson on
of termen ; a terminal series of black points ; cilia pale with
some dark spots towards apex and a black patch above toruus.
Hind wing whitish tinged with brown, especially on the
veins and terminal area.
Hab. Assam, Khasis; Cryton, Puttalam (Pole), 1 ? type ;
SamBawa (Doherty),1 9. Eap. 20 mm.
Dilophothripa olivia, sp. n.
6. Head and thorax whitish, mixed with pale olive ;
palpi at sides and antenne blackish; tegule at tips and
outer edge of patagia brown; abdomen whitish. Fore wing
pale olive; a black patch on costa near base and some fiery-
red marks on inner area before the indistinct, oblique, waved
antemedial line; two minute black points in middle of cell
and two on discocellulars, and two more prominent obliquely
placed points below middle of cell; postmedial line fuscous
defined by white on inner side, very minutely waved, very
oblique from costa to vein 6, then inwardly oblique, a fuscous
patch beyond it on costa with white streak below it curving
up to apex ; some white points on costa towards apex; a
fuscous mark beyond the postmedial line in submedian fold ;
subterminal line white with fuscous marks on its inner side,
excurved below costa and at middle; a te:minal series of
black points. Hind wing white, slightly tinged with brown.
Hab. Borneo, Paulo Laut (Doherty). Eap. 18 mm.
Sarrothripa mesoplaga, sp. 0.
?. Head and thorax grey-white, mixed with fuscous ;
fore tarsi fuscous with white rigs; abdomen greyish fuscous,
ventrally whitish. Fore wing rather silvery white ; a fuscous
subbasal band defined by waved black lines from costa to
submedian fold; antemedial line oblique from costa to
median nervure and angled outwards on subcostal and
median uervures, then incurved; a double waved medial
line ; a fuscous patch on costa from the inner medial to
beyond the postmedial line extending to median nervure; a
white discoidal elliptical spot defined by black ; a double
waved postmcdial line, bent outwards below costa, angled
outw:rds at vein 6, and incurved below vein 4; a dentate
subterminal line, bent inwards between veins 6 and 4 ; some
fuscous suffusion on inner and terminal are.is except at apex ;
a terminal punctiform line. Hind wing pale fuscous.
Hab. Sixearore (Ridley), 1 3 type. Lap. 22 mm.
new Species of Noctuidee. 58d
ACONTIANE.
Paraxestis mesogonia, sp. Nn.
3. Head and thorax ochreous, tinged with rufous ; pectus
and legs whitish; abdomen ochreous, the ventral surface
white. Fore wing ochreous, the basal and costal areas to
;ostmedial line irrorated with brown, the medial area (except
towards costa) and the terminal area suffused with rufous; a
brown antemedial line obtusely angled in cell, then oblique
and slightly sinuous; a brown postmedial line angled at vein
6, then oblique; traces of a subterminal line towards costa
and an ill-defined spot on inner margin ; cilia rufous, mixed
with brown. Hind wing semihyaline yellowish white with a
slight rufous tinge towards termen.
?. Fore wing with the medial area suffused with dark
brown, the terminal area tinged with brown ; hind wing
more ochreous, the termen brownish.
Hab. Axsyssinia (Gerrard), 2 3,1 9 type. Eap., 3 24,
9 26 mm.
Parawxestis glaucescens, sp. n.
?. Head and thorax olive-ochreous, with a slight rufous
tinge; palpi, tibie, and tarsi rufous; abdomen ochreous,
tinged with rufous. Fore wing olive-ochreous, suffused with
rufous and with a silky gloss ; traces of slightly curved ante-
and postmedial rufous lines; termen and cilia deeper rufous.
Hind wing ochreous, tinged with rufous, especially on
terminal area.
Hab, Oxuv Catasar (Crompton), 1 2 type. Hap. 22 mm.
Arcyophura piperita, sp. nu.
?. Head ochreous ; thorax ochreous suffused with brown ;
pectus, legs, and abdomen whitish. Fore wing ochreous,
thickly irrorated with brown, the terminal half tinged with
rufous, and the terminal area very minutely striated with
brown ; an indistinct antemedial line slightly excurved below
costa, then oblique ; an indistinct postmedial line excurved
from below costa to vein 5, then slightly incurved, faint traces
of a subterminal line. Hind wing semihyaline white, the
termen tinged with ochreous.
Hab. Nigeria, Borgu, Yelwa Lake (Migeod),1 ¢ type.
Exp. 22 mm.
586 Sir G. F. Hampson on
Erizada rufa, sp. 0.
¢. Head and thorax red-brown; pectus and legs whitish,
fore tibie rufous, the tarsi banded with rufous; abdomen
whitish, dorsally tinged with red, except at base. Fore wing
rufous, the basal costal area brown ; a brown patch on costa
from middle to near apex, with a very oblique diffused rufous
band from it to inner margin; cilia deep rufous at base,
whitish at tips. Hind wing pale red, the basal and costal
areas slightly paler. Underside of fore wing suffused with
fiery red except at marginal areas.
Hab. Suita (Wallace),1 8 type. Exp. 22 mm.
Genus PoGonozADA, nov.
Proboscis fully developed ; palpi porrect, extending about
the length of head, the second joint thickly scaled above;
antenne of male almost simple; mid and hind tibiz thickly
fringed with hair above, the latter with the spurs long and
with tufts of long hair between them, the first joint of hind
tarsi with very large fringes of hairabove. Fore wing broad,
the costa and inner margin arched, the apex and termen
rounded; vein 2 from well before angle of cell; 3 and 4
from angle ; 5 from just above angle; 6, 7,8, 9, 10, 11 from
cell. Hind wing short and contorted, the tornus lobed, the
inner and terminal areas clothed with rough hair; the
underside with large tufts of rough scales on tornal area ;
the lower part of cell short ; veins 3 and 5 from near angle ;
6, 7 stalked and bent downwards, separating widely after
origin; 8 approximated to cell to middle, then curved
upwards.
Pogonozada distorta, sp. n.
gd. Head, thorax, and abdomen rufous, the thorax
dorsally tinged with fuscous. Fore wing rufous, slightly irro-
rated with darker brown, the inner area tinged with fuscous ;
some slight oblique brown strize from basal half of costa ;
four obliquely curved faint brown lines from terminal half of
costa towards tornus; a slight black streak on vein 2 towards
tornus, with strize from its extremities to inner margin ; cilia
tinged with fuscous. Hind wing rufous, suffused in parts
with brown.
Hab. C. Cuina, Ichang (Mrs. Pratt), 1 ¢ type.
Eep. 18 mm.
new Species of Noctuide. 587
Tortriciforma chloroplaga, sp. n.
?. Head and thorax grey-white ; palpi, pectus, legs, and
abdomen ochreous, tinged with rufous. Fore wing ochreous
white, the base and terminal area more grey-white, the costa
faintly tinged with rufous ; a small fiery-red streak on base
of inner margin; an irregularly elliptical dark sap-green
subbasal patch from costa to median nervure, its lower part
suffused with some reddish brown on subcostal nervure; a
black discoidal point; a faint diffused brownish medial line
from vein 2 to inner margin; an oblique sap-green streak
from costa just beyond middle to vein 7, with traces of a
sinuous postmedial line arising from it, some rufous on costa
beyond it with some grey below it before the subterminal
line, which is represented by a series of small obscure
blackish marks strongly angled outwards at vein 7 and
middle; a whitish subapical spot. Hind wing yellow, the
inner area tinged with fiery red; a red discoidal point ; a
diffused red subterminal line and narrow terminal band.
Underside of fore wing tinged with fiery red; hind wing with
some red irroration on costal area, a red discoidal lunule,
sinuous red postmedial and subterminal lines from costa to
vein 5, some grey on termen.
Hab. Prrax, Larut Hills (S. S. Flower), 1 9 type. Ezp.
24 mm,
Metaptya lucida, sp. n.
6. Head, thorax, and abdomen brownish grey; palpi
white in front; tarsi with fuscous bands. Fore wing
ochreous brown, suffused with glistening grey and irrorated
with a few black scales; a fine, obliquely curved, whitish
antemedial line; a slight black point at upper angle of cell;
a fine whitish postmedial line excurved from costa to vein 4,
then incurved; some slight black streaks on costa towards
apex; subterminal line represeuted by some black scales, a
small spot below costa and rather bifid spot between veins 5
and 3; cilia blackish at the angle at vein 4. Hind wing
ochreous brown suffused with glistening grey; aslight black
and white striga on inner margin above tornus. Underside
fuscous brown, the inner area of fore wing and hind wing
(except terminal area) whitish, both wings with rather diffused,
slightly sinuous postmedial line and dark point at upper
angle of cell.
Hab. Batt, Low Country (Doherty), 1 g type. Hap.
26 mm.
588 Sir G. F. Hampson on
Urbona microcyma, sp. n.
dg. Head and thorax grey-white largely suffused with
rufous or dark brown ; palpi with the second joint blackish
above; tarsi banded with fuscous ; abdomen brownish white.
Fore wing whitish, tinged with rufous and more or less
thickly irrorated with fuscous ; a black point in middle of
cell and two discoidal points or smal] spots; an oblique
minutely dentate brown line from costa near apex to middle
of inner margin, sometimes slightly defined by white on outer
side. Hind wing whitish tinged with brown; the underside
whiter, sometimes with traces cf postmedial line on costal
half. .
?. Head, thorax, and fore wing much darker red-brown;
abdomen fuscous; fore wing with the discoidal points
conjoined into a lunule.
Hab. Nicerta, Akasa (Lugard), 1 ¢ type; Sapele
(Sampson), 5 g,3 2 ; Old Calabar (Crompton),1 3. Exp.
28-32 mm.
Westermannia cuprea, sp. n.
¢. Head, thorax, and abdomen white, slightly tinged with
brown ; palpi blackish at sides. Fore wing golden cupreous,
the inner area white, slightly tinged with brown from base
of costa to inncr margii near tornus, which is greyish; the
costal area, the cell before antemedial line, and the area
beyond the cell to postmedial line down to vein 2 suffused
with blue-grey ; the veins slightly streaked with white, more
prominently on postmedial area from costa to vein 6; an
oblique loop-shaped subbasal mark defined by a fine white
line from subcostal nervure to the inner white area; ante-
medial line pure white, inwardly oblique and incurved from
below costa to the white area; orbicular and reniform finely
defined by white, the former somewhat elongate elliptical,
the latter with its lower part defined by grey, with white
point on its inner lower edge and small elliptical spot on its
outer ; postmedial line white, oblique from costa to vein 7,
where it is acutely angled outwards, then incurved to vein 3,
where it is less acutely angled, and again incurved to inner
margin ; asubterminal series of small black spots, with fine,
oblique, slightly waved, white line on their outer edge from
costa to vein 7; the white streaks on veins 7,6 meeting at
termen in an acute angle in the interspace. Hind wing
pale brown.
Hab. Asuanti, Obuassi (Bergmann), 1 3 type. Eap.
32 mm.
new Species of Noctuide. 589
Westermannia leucogonia, sp. ni.
2. Head, thorax, and abdomen pale rufous. Fore wing
pale rufous, with slight dark irroration ; an oblique white
subbasal line from costa to submedian fold, with a small
round rufous spot defined by a fine white annulus beyond it
in cell; the area to antemedial line suffused with grey-white
from costa to submedian fold; antemedial line white, ex-
tremely oblique from costa to median nervure near lower
angle of cell, then incurved to vein 1, where it meets the
postmedial line and terminates; a pear-shaped rufous
discoidal spot defined by a fine white annulus, some grey
suffusion beyond upper and lower angles of cell, the veins
from lower angle streaked with white ; postmedial line white,
very oblique from costa to vein 7, oblique below vein 3 to
vein 1, where it terminates, curving round to join the ante-
medial line; a triangular white patch beyond it from costa to
vein 7, then slightly defined by grey on outer side and below
vein 3 by white ; a somewhat irregular subterminal series of
slight blackish spots; a fine pale line at base of cilia, Hind
wing pale rufous, the costal area paler; the underside
whitish, with slight brown disco:dal spot.
Hab. Oxtv CaraBar (Crompton), 1 3 type. Exp. 30 mm.
Westermannia agrapha, sp. un.
@. Head, thorax, and abdomen pale rufous; palpi in
front, pectus, legs, aud ventral surface of abdomen whitish.
Fore wing pale rufous ; a fine white streak from base of costa
along vein | to tornus, emitting a slight streak along median
nervure to the slight very oblique white antemedial line, the
angle between these lines tinged with grey; a rather trian-
gular cupreous discoidal spot defined by a fine whitish
annulus ; postmedial line indistinct, fine, whitish, oblique
from costa to vein 6, and below vein 4 bent inwards to
below end of cell, then erect to inner margin ; terminal area
suffused with golden cupreous ; a white subterminal point
on costa. Hind wing pale rufous, the underside whitish.
Hab. Sierra Leone (Foxcroft), 1 9 type. Hap. 26 mm.
Westermannia argyroplaga, sp. u.
6. Head and tegule white, palpi above and antenne
rufous, the tegulze with some rufous scales at edges ; thorax
white suffused with rufous; pectus and legs pale rufous;
abdomen white tinged with rufous. Fore wing pale rufous,
with some darker rufous irroration ; a large oblique quadrate
590 Sir G, F. Hampson on
silvery-white patch extending on costa from base to near
middle and down to vein 1 and below end of cell, where it is
confluent with a broad, curved, silvery-white band from
costa beyond middle, extending down to submedian fold, and
emitting a short spur inwards below costa, its outer edge
slightly indented at discal fold; traces of a dark rufous
subterminal line, incurved at submedian fold; a fine dark
terminal line. Hind wing white. Underside of fore wing
whitish, suffused with fuscous except inuer and terminal
areas,
Hab. Ucanpna, Marengi (Christy), 1 g type. Hap.
28 mm.
Westermannia trigonica, sp. 0.
g. Head, thorax, and abdomen pale brown ; palpi greyish
in front, blackish at sides; pectus and legs greyish; abdo-
men greyish at base, the ventral surface greyish, with dark
patch at middle. Fore wing with the basal area white tinged
with brown; a large triangular golden-bronze patch ex-
tending on costa from near base to beyond middle, its apex
at middle of inner margin, defined by white lines, the area
beyond it white shading into golden bronze on terminal
area; an indistinct rather maculate subterminal line un-
usually remote from termen, angled outwards at vein 7 and
inwards in discal and submedian folds and excurved at
middle. Hind wing pale golden bronze. Underside of fore
wing pale golden bronze, with the inner area whitish.
?. Fore wing with the basal and postmedial areas much
browner.
Hab. Niceria, Old Calabar (Crompton), 1 3 type;
Akassa (Lugard), 1 3g; Sapele (Sampson), 1 9. Exp.
28 mm.
Westermannia areogramma, sp. 0.
9. Head, thorax, and abdomen whitish suffused with
rufous. Fore wing pale rufous irrorated with darker rufous ;
an indistinct double oblique line from costa before middle to
inner margin beyond middle; an indistinct, rather large,
oblique, elliptical discoidal spot defined by a slight pale
annulus, with some slight dark suffusion beyond its lower
extremity ; an indistinct double postmedial line excurved
from costa to vein 3, then bent inwards to below end of cell,
and almost meeting the antemedial line at inner margin; a
slight pale line at base of cilia. Hind wing white, faintly
new Species of Noctuide. 591
tinged with rufous. Underside of fore wing whitish slightly
tinged with rufous.
Hab. Br. C. Arrica, Chiromo (de Jersey), 1 @ type.
Exp. 26 mm.
Westermannia nocturna, sp. n.
3. Head, thorax, and abdomen pale greyish brown.
Fore wing with the inner. margin highly arched, the termen
obliquely curved ; pale reddish brown suffused with greyish
and irrorated with a few dark scales; an oblique brown
striga from middle of costa; a slight curved discoidal bar ;
an oblique brown streak from costa beyond middle to vein 6,
with a greyer patch beyond it; traces of a subterminal line
excurved below costa and at middle. Hind wing uniform,
rather pale red-brown; the underside greyer, tinged with
fuscous from lower angle of cell to termen.
Hab. Nicerta, Old Calabar (Crompton),1 g type. Exp.
38 mm.
Westermannia pheopepla, sp. n.
?. Head, thorax, and abdomen pale grey-brown ; pectus
and ventral surface of abdomen whitish. Fore wing pale
greyish fuscous brown, the costal area (except on medial area)
paler, with an ochreous tinge and slight dark irroration ;
antemedial line indistinct, dark, very slightly incurved ; an
obscure dark discoidal bar ; postmedial line indistinct, dark,
diffused, oblique from costa to vein 6, then inwardly oblique,
a faint dark spot beyond it at vein 4. Hind wing fuscous
brown, with a faint reddish gloss. Underside pale grey ;
fore wing suffused with fuscous except marginal areas.
Hab. Nicerta, Sapele (Sampson), 1 2 type. Lap.
28 mm.
Westermannia stalactitis, sp. n.
9. Head and thorax greyish brown; palpi whitish in
front ; abdomen fuscous brown. Fore wing pale reddish
brown with a cupreous tinge, the basal area glossed with
grey ; a somewhat oblique slightly sinuous antemedial line,
with a minute dark wedge-shaped mark from costa before it ;
a wedge-shaped chocolate patch from middle of costa to
vein 1, its edges defined by whitish lines and waved, its
point bent outwards and acute; three white discoidal points,
the lowest rather larger ; a sinuous postmedial line, oblique
from costa to vein 6, excurved at median nervules, then
incurved ; a somewhat semicircular chocolate patch on.costa
592 Sir G. F. Hampson on
towards apex, with two pale points on costa, its edge defined
by a pale line, slightly indented on inner side and angled
outwards on outer; a subterminal series of minute black
and white spots, the spot below costa slightly larger; cilia
with a fine pale line at base. Hind wing fuscons brown with
a reddish gloss ; the underside whitish, suffused and irrorated
with brown down to median nervure and vein 2; an indistinct
curved postmedial line.
Hab. Nicertia, Old Calabar (Crompton), 1 2 type. Exp.
28 mm.
Westermannia geminastra, sp. 0.
?. Head and thorax pale reddish brown ; palpi whitish
in front; abdomen fuscous brown. Fore wing reddish
brown with a cupreous gloss; an oblique, slightly sinuous,
brown antemedial line, defined by whitish on inner side ;
two pure white discoidal spots, the lower rather larger and
placed on a diffused paler rufous patch; postmedial line
brown, defined by whitish on outer side, excurved from costa
to vein 4, then incurved ; a fine pale line at base of cilia.
Hind wing pale with a reddish gloss; a fine pale line at base
of cilia; the underside whitish tinged with reddish brown.
Hab. Niceria, Old Calabar (Crompton), 1 ¢ type. Eap.
24mm,
Westermannia digramma, sp. v\.
g. Head and thorax pale red-brown ; abdomen greyish
brown, the dorsal crest and anal tuft dark at tips, the genital
tufts whitish. Fore wing pale rufous; an oblique, straight,
dark brown antemedial line ; two black discoidal points with
a few white scales on them ; postmedial line shghtly excurved
from costa to vein 6, then oblique; aslight pale line at base
of cilia. Hind wing brown with a silky gloss ; the underside
whitish slightly irrorated with brown.
Hab. Niceria, Old Calabar (Crompton), 1 3 type. Exp.
28 mm.
Westermannia selenitis, sp. 1.
@. Head and thorax grey mixed with brown and fuscous ;
palpi blackish above; abdomen whitish, dorsally tinged with
fuscous. Fore wing pale rufous sparsely irrorated with
black ; traces of an oblique slightly sinuous antemedial line ;
a small blackish spot in middle of cell and crescentic discoidal -
spot; postmedial line arising from costa towards apex,
fuscous, defined by whitish on outer side, slightly exeurved
new Species of Noctuidee. 593
from costa to vein 4, then slightly incurved ; a fine pale line
at base of cilia. Hind wing W hitish, uniformly tinged with
pale brown; the underside rather whiter.
Hab. Nicerta, Sapele (Sampson), 1 Sg type. Hxp.30 mm.
Aiteta costiplaga, sp. n.
@. Head and thorax rufous; palpi with the extremity
of second and third joints white : frons with white patch
below; vertex of head and thorax with whitish patches ;
abdomen pale rufous. Fore wing pale brownish rufous
slightly tinged with grey ; antemedial line indistinct, out-
wardly oblique, arising from a slight olive triangular ‘patch
on costa ; postmedial line indistinct, inwardly oblique, arising
from apex of a triangular olive patch on costa; traces of a
sinuous punctiform subterminal line ; cilia grey at base, with
brown line through them and w hitish tips. Hind wing and
underside of both wings orange.
Hab. Nigeria, Onitsba, 1 2 type. LHzp.30 mm.
Aiteta meterythra, sp. n.
$. Head and thorax deep rufous; fore tibia and tarsi
and mid and hind tibiz streaked with rufous, the fringe of
very long hair on hind tibiz tinged with fuscous ; abdomen
orange-red, the base and extremity of dorsum tinged with
brown. Fore wing deep rufous, slightly suffused with grey
except on medial area and with slight dark irroration ;
antemedial line defined by grey on inner side, outwardly
oblique, almost straight; a slight blackish point in cell
towards extremity and blackish discoidal bar; postmedial
line defined by greyish on outer side, with some deeper
rufous suffusion on each side of it, slightly oblique from costa
to vein 6 and incurved below vein 4; subterminal line fine,
blackish, slightly waved, angled outwards at vein 7, excurved
at middle, and ending on termen above tornus. Hind wing
and underside of both wings fiery red.
?. Thorax and abdomen sometimes dorsally tinged with
grey ; fore wing with the ground-colour deep rufous or more
or less entirely suffused with grey.
Hab. Nicerta, Sapele (Sampson), 1 g, 2 2 type;
Old Calabar (Crompton),3 ¢%. Exp. g 30, 2 26-28 mm.
Carea acypera, sp. n.
3. Fore wing with the apex acute, the termen nearly
straight.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 39
594 Sir G. F. Hampson on
Head, thorax, and abdomen pale grey-brown ; palpi with
the third joint blackish ; pectus, legs, and ventral surface of
abdomen whitish. Fore wing pale grey-brown, sparsely
irrorated and striated with raised black scales; a slightly
more prominent black discoidal point; cilia white at base,
brown at tips except towards tornus. Hind wing greyish
brown with blackish discoidal point; cilia whitish; the
underside brownish white, irrorated and striated with raised
black scales, the black discoidal spot prominent.
Hab. Bau (Doherty), 1 g type. Hap. 30 mm.
Carea tumida, sp. 0.
Fore wing with the costa strongly arched and forming a
lobe towards apex in male, slightly in female, the termen
evenly curved.
Head and thorax deep rufous; palpi blackish; frons
whitish at base, with blackish bar above and between an-
tenne ; tegule with black line near base ; fore legs blackish
in front ; pectus whitish ; abdomen rufous, dorsally tinged
with fuscous. Fore wing bright rufous, suffused with silky
grey on inner area, as a wedge-shaped patch from basal half
of costa to near tornus and from postmedial line to near
termen, subbasal line represented by an oblique black striga
from costa; an indistinct, oblique, slightly curved ante-
medial line; postmedial line double, black, from costa
beyond middle to tornus; an indistinct subterminal line,
somewhat oblique and maculate from costa to vein 6 and
obscurely angled inwards at vein 5. Hund wing fiery red,
the inner area fuscous brown. Underside fiery red; fore
wing with the apical area suffused with grey; hind wing
tinged with grey.
Hab. Punane (S. S. Flower), 1 9; Sincapore (Ridley),
1 ¢; Borneo, Sadong (Shelford), 1 g type, 1 9. Kap.
43 mm.
Carea diplogramma, sp. 0.
Fore wing with the apex somewhat produced and acute,
the termen excurved at middle.
?. Head and tegule dark red-brown ; frons greyish at
base; thorax rufous; pectus and legs pale; abdomen
dorsally fuscous, ventrally rufous. Fore wing rufous, the
costal and imner areas slightly tinged with brown; an
indistinct oblique antemedial line, slightly angled outwards
at median nervure ; a slight discoidal striga; a double lime
from costa near apex to tornus. Hind wing fiery red, the
new Species of Noctuidee. 595
inner area fuscous brown. Underside pale fiery red; fore
wing with the costal area greyish ; hind wing with the inner
area whitish, traces of a dark subterminal line except
towards tornus.
Hab. Mauacca, Johore (Ridley), 1 9 type. Hxp.40 mm.
Carea holophea, sp. u.
Fore wing with the apex somewhat produced and acute,
the termen excurved at middle.
2. Head and tegule black-brown; palpi rufous, greyish
in front; frons at base and basal joint of antennz in front
whitish; thorax rufous more or less tinged with grey;
pectus and legs greyish, the fore legs dark in front ; abdomen
fuscous, ventrally rufous. Fore wing rufous more or less
suffused with grey ; an indistinct dark subbasal line from
costa to submedian fold; a pale green subbasal patch on
inner margin ; antemedial line fine, black, obliquely excurved
from pulncactal nervure to inner margin; a large triangular
patch of blackish suffusion on costa before the double oblique
black postmedial line, and extending to below lower angle of
cell; a slight fuscous tinge on termen below apex. Hind
wing fuscous brown with a slight rufous tinge ; termen and
cilia rufous. Underside fiery red, the costal area tinged with
grey towards apex; hind wing with the inner half whitish.
Hab. Matacca, Johore (Ridley), 1 9; Perak Hills
(Ridley), 1 9 type. Ezp. 34 mm.
Carea bwwiata, sp. n.
dg. Head and thorax fiery red mixed with some yellow;
fore tibize and tarsi tinged with fuscous ; pectus and mid and
hind legs whitish ; abdomen grey tinged with rufous, the
anal tuft blackish. Fore wing with the apex slightly
produced, the termen nearly evenly curved, fiery red
sparsely irrorated with black; the base of inner area
yellowish, the apical area tinged with grey; an oblique
straight dark line from costa ‘before middle to middle of
inner margin ; an oblique line from costa beyond middle to
tornus, slightly excurved below costa; the inner margin
dark except towards base.. Hind wing pale fiery red; the
underside whitish, with the costal and terminal areas irrorated
with brown.
?. Fore wing with the ground-colour nearly uniform
fiery red; the lines slightly more diffused, the postmedial
line hardly excurved below costa; a dark discoidal point,
39*
596 Sir G. F. Hampson on
Hab. Penane (S. S. Flower), 1 g type; Perax, Larut
Hills (S. S. Flower),1 9. Eap., g 32, 2? 40 mm.
Carea endophea, sp. n.
g. Head and thorax deep chocolate-red ; fore tibize and
tarsi pure white in front ; mid and hind tibiz mostly white,
the tarsi ringed with white; abdomen fuscous, the anal tuft
white, the ventral surface fiery red. Fore wing deep
chocolate-red ; the costal edge pure white, the medial and
terminal areas with a more or less prominent greyish gloss ;
two indistinct rather diffused antemedial lines, very slightly
excurved below costa and incurved in submedian interspace ;
a black discoidal point surrounded by a grey annulus; an
indistinct somewhat diffused postmedial line, slightly angled
outwards below costa and at middle, with a still less distinct
diffused red line beyond it, slightly excurved from costa to
middle, then incurved ; an indistinct, dark, somewhat dentate
subterminal line with series of white points on its outer edge,
slightly excurved at vein 7 and middle. Hind wing fiery
red, the inner area to cell and vein 2 fuscous brown.
Underside of both wings fiery red, the apices tinged with
chocolate, the inner area of hind wing greyish.
Hab. Travancorr, Pirméd (Imray), 4 @ type. Lap.
44 mm.
Carea pallida, sp. un.
9. Head and thorax pale red-brown; abdomen pale
brown ; pectus and ventral surface of abdomen whitish.
Fore wing pale red-brown, finely and thickly irrorated with
dark brown; the costal edge pure white; an indistinct
oblique antemedial line, very slightly excurved below costa
and incurved in submedian interspace; a black discoidal
point ; postmedial line rather indistinct, slightly excurved
below costa and at middle, angled inwards at discal fold and
incurved below vein 4; traces of a subterminal line excurved
at vein 7 and middle, cilia whitish. Hind wing white,
faintly tinged with red and the inner area with fuscous; the
underside white faintly tinged with red.
Hab. Burma, N. Shan States (Bingham),1 2 type. Eap.
38 mm.
Carea leucocraspis, sp. 0.
gd. Head and thorax bright rufous; fore tibiee and tarsi
white in front; mid tibize white at base; abdomen pale
new Species of Noctuide. 597
fuscous brown. Fore wing bright rufous, the costal edge
pure white; a very indistinct antemedial line oblique from
costa to submedian fold; a very faint postmedial line
excurved below costa and at middle; traces of a sinuous
subterminal line. Hind wing fiery red, the inner area pale
fuscous brown ; the underside fiery red, with the basal and
inner areas whitish.
Hab. Borneo, Sandakan (Pryer), 1 3 type. Exp.
42 mm.
Carea rubrifusa, sp. 0.
6. Head and thorax bright rufous; pectus, legs, and
abdomen whitish brown. Fore wing whitish brown tinged
with rufous, the basal area (except at base of inner margin)
suffused with bright rufous to the antemedial line, which is
oblique, angled outwards below costa and at median nervure
and inwards in cell and submedian fold ; a minute discoidal
point; postmedial line rufous, obliquely incurved from
costa to vein 6, where it is acutely angled outwards, then
incurved, some rufous suffusion beyond it; an indistinct,
somewhat diffused, subterminal line, incurved at discal fold ;
some rufous suffusion on termen from apex to vein2. Hind
wing whitish suffused with pale red. Underside whitish
tinged with red.
Hab. Borneo, Sandakan (Pryer), 1 3 type. Exp.
40 mm.
Carea auritincta, sp. 1.
d. Palpi with the third jomt tufted with hair on inner
side ; hind tibiz not dilated, without groove and tuft of
hair; fore wing short and broad, the termen erect, curved.
Head and thorax rufous with a silky gloss ; palpi in front,
pectus, and legs white; fore and mid tibiz partly rufous,
the tarsi rufous ringed with white ; abdomen pale rufous,
the ventral surface white. Fore wing rufous with a greenish-
golden gloss, especially towards costa; the costa with ante-
medial and medial black and white points, towards apex with
dark brown streaks interrupted by three white points. Hind
wing fiery orange-red; cilia white at tips. Underside of
fore wing orange-red, the imner area and costal area to
beyond middle suffused with white; hind wing white tinged
with red.
Hab. Nigeria, Old Calabar (Crompton), 1 3 type. Exp.
26 mm.
598 Sir G. F. Hampson on
Carea gamma, sp. 1.
3. Head, tegule, and patagia greyish brown, the tegul
with deep chocolate medial patch; thorax leaden grey ;
pectus pale ; coxe fiery red, the tufts of hair on hind tibie
fuscous ; abdomen greyish on dorsum, reddish at sides and
below, the crests at base black, the extremity blackish.
Fore wing greyish brown to an oblique line from middle of
costa to inner margin before middle, the basal costal area
suffused with reddish brown; the terminal half black-brown ;
a black point defined by a slight pale annulus in end of cell ;
a slight silvery gamma-shaped mark on discocellulars with
black point between its arms, its tail extending obliquely
beyond lower angle of cell; an indistinct black postmedial
line oblique from costa to vein 4, then bent inwards to
vein 2 and outwards towards tornus, a diffused blackish
fascia from it to termen above vein 5. Hind wing yellow
suffused with fiery red, especially towards apex. Underside
of fore wing fiery red.
Hab, Nicertia, Sapele (Sampson), 1 ¢ type. Hap. 32mm.
Carea metaleuca, sp. n.
?. Head and thorax bright rufous; palpi at base, pectus,
legs, and abdomen white. Fore wing bright rufous with
very slight dark irroration on costal, inner, and terminal
areas ; subbasal line represented by a black striga from costa ;
antemedial line fuscous, defined by white on inner side,
slightly bent inward to costa, then oblique ; a black discoidal
point ; postmedial line fuscous, defined by white on outer
side, oblique and slightly excurved below costa and at middle ;
two white points on costa towards apex ; a subterminal series
of slight dark spots, somewhat excurved at vein 7 and middle.
Hind wing white tinged with yellow, the inner area slightly
with brown. Underside white.
Hab. Br. C. Arrica, Chiromo (de Jersey), 1 2 type.
Hap. 24 mm. '
Carea mesocausta, sp. n.
dg. Head, thorax, and abdomen grey; pectus and base
of legs white; tarsi red-brown ringed with white ; abdomen
with a little fiery red on terminal segment, the anal tuft
white at tip, the ventral surface whitish tinged with red
towards extremity. Fore wing grey sparsely irrorated with
black, the medial area strongly suffused with red-brown,
with some large black-pencilled scales below the cell; an
new Species of Noctuide. 599
oblique antemedial line, excurved in cell and incurved in
submedian interspace ; an oblique black discoidal bar, defined
by grey on outer side; an oblique postmedial line ending at
tornus, excurved below costa and at middle; a subterminal
punctiform fuscous line ending on termen above tornus,
angled outwards at vein 7 and excurved at middle; a terminal
series of black points; cilia white intersected by pale
brownish. Hind wing fuscous brown; the underside
whitish suffused with fuscous.
Hab. Borneo, Sandakan (Pryer), 1 & type. zap.
28 mm.
Carea albipicta, sp. n.
@. Head and thorax grey-white mixed with purplish red ;
palpi rufous ; pectus and legs whitish, tarsi rufous ringed
with white; abdomen white. Fore wing white, the costal,
inner, and terminal areas suffused with purple-grey and
irrorated with rufous; some rufous suffusion at base of
costa; the medial and postmedial areas (except towards costa
and inner margin) striated with bright ferruginous ; an ill-
defined waved antemedial line, diffused towards costa and
somewhat inwardly oblique; a ferruginous discoidal bar; a
somewhat oblique, sinuous, white postmedial band, incurved
in submedian interspace ; a dark brown subterminal line,
excurved at vein 7 and middle; a dark brown terminal line
expanding into small spots in the interspaces ; cilia ferru-
ginous. Hind wing semihyaline white. Underside of fore
wing white, the costal area suffused with grey and red-brown.
Hab. ist, | 2 type. Hzp. 32 mm.
Maurilia albirivula, sp. n.
¢. Head and thorax pale rufous; second joint of palpi
and tegule slightly tipped with white; pectus and legs
whitish ; abdomen fuscous brown, the ventral surface
whitish. Fore wing pale rufous, the medial area dark
reddish brown, extending below vein 6 to just’ beyond post-
medial line and below vein 4 to termen; a small patch of
brown suffusion in cell before the antemedial line, which is
white defined by brown on outer side, oblique, waved, more
strongly incurved in submedian interspace ; postmedial line
white defined by brown on each side, slightly waved, oblique
from costa to vein 4; some whitish points on costa towards
apex ; a subterminal series of small obscure fuscous spots
except towards costa, excurved below vein 4 to near termen;
some brown suffusion on terminal area below apex, with
600 Sir G. F. Hampson on
slightly darker streaks on the veins. Hind wing fuscous
brown, the cilia ochreous white. Underside of fore wing
fuscous brown, the inner area and hind wing whitish.
Hab. Nigeria, Old Calabar (Crompton), 1 2 type. Exp.
30 mm.
Maurilia heterochroa, sp. n.
Head, thorax, and fore wing varying from bright rufous
to greyish rufous or greyish fuscous; palpi irrorated with
white ; frons with white patch; pectus and base of legs
white ; tibiz and tarsi white mixed with rufous; abdomen
fuscous, the anal tuft rufous, the ventral surface white tinged
with rufous towards extremity. Fore wing with double,
waved, very oblique antemedial line; black points in end of
cell and on discocellulars; a double, somewhat waved post-
medial line, oblique from costa to vein 6, slightly angled
inwards in discal fold, then excurved and incurved in sub-
median interspace ; a slight somewhat sinuous fuscous shade
from apex to antemedial line below the cell; a subterminal
punctiform black line slightly excurved at vein 7 and in-
curyed at discal fold; the termen rather darker from just
below apex to vein 2. Hind wing fuscous brown to deep
fuscous; cilia greyish at tips. Underside of fore wing
fuscous black, the costal and terminal areas red, with some
whitish on costa towards apex, inner area whitish; hind
wing red, the basal and inner areas grey-white.
Ab. 1.—Fore wing with the inner area from near base to
subterminal line and extending to upper angle of cell suttused
with dark red-brown.
Ab. 2.—Fore wing with this area deep ferruginous red,
with a large, darker ferruginous, somewhat triangular patch
on it beyond the cell.
Ab. 3.—Fore wing greyish fuscous.
Ab. 4.—Fore wing greyish fuscous, with a similar dark
fuscous patch on inner area.
Ab. 5.—Fore wing greyish fuscous, with diffused black
wedge-shaped patch beyond the cell.
Hab. Nicerta, Old Calabar (Rutherford, Crompton), 4 @,
2 2 type; Sapele (Sampson), 2 $. Ep. 36-40 mm.
Maurilia phea, sp. n.
g. Olive-fuscous; head, thorax, and fore wing sometimes
tinged with dull rufous; pectus and base of legs and
abdomen whitish. Fore wing with two dark, oblique, waved
ante- and postmedial lines, the latter somewhat excurved
new Species of Noctuide. 60L
below costa and at middle and ending near tornus ; a slight
oblique dark discoidal bar; a subterminal series of small
fuscous spots, somewhat oblique from costa to vein 6, slightly
incurved at discal fold and excurved at middle. Hind wing
and underside uniform fuscous brown.
Hab. Nicerta, Old Calabar (Miss Kingsley, Sampson,
Crompton), 4 3 type. Hzxp, 32-38 mm.
Tyana acypera, sp. n.
Tyana pustulifera, Leech, Trans, Ent. Soc. 1899, p. 203 (nee WIk.).
3d. Hind wing with a large glandular swelling at tornus
clothed with flocculent hair on underside, the inner margin
fringed with long hair; fore wing with the apex extremely
produced and acute, the termen almost straight.
Head and thorax bright green; palpi red, white below; frons
yellow, with crimson line above; antenne red ringed with
white ; pectus and legs white, the fore tibie and tarsi red ;
abdomen dorsally brown, ventrally whitish. Fore wing bright
green; the costal edge brown irrorated with grey, the extreme
tip of apex crimson; brown points at angles of cell; cilia
yellow. Hind wing pure white, the hair on inner margin
slightly tinged with fuscous.
?. Fore wing with the apex less produced, the termen
rounded ; vertex of head crimson and yellow; tegule and
base of patagia yellow, with some crimson on shoulders;
abdomen white, with minute crimson tuft at base. Fore
wing with the ground-colour whitish, almost completely
covered with green irroration, the costa and inner margin
white ; a small crimson annulus with yellow centre at lower
angle of cell, sometimes filled in with crimson.
Hab. Cunrr. Cutna, Omei-shan, 1 ¢ type; Pu-tsu-fong,
2 9.. Ezp. 38 mm.
Ariola xanthobasis, sp. u.
3. Head and thorax pale yellow; palpi, tegule (except at
middle), and metathorax tinged with rufous ; pectus whitish,
legs mostly dark brown; abdomen fuscous brown, ochreous
at base, the claspers large, white at sides, the ventral surface
white. Fore wing with the basal area pale yellow, bounded
by a curved, rather diffused, brilliant silver line from costa
before middle to middle of inner margin, the costa tinged
with rufous; the rest of wing uniform very deep chocolate-
brown. Hind wing uniform fuscous brown; the underside
pale, the costal and terminal areas broadly suffused with
602 Sir G. F. Hampson on
fuscous, leaving a diffused whitish patch on costal area before
apex.
9. Pectus and legs more ochreous ; hind wing with a
shght rufous tinge.
Hab. Niceria, Akassa (Lugard), 1 3 type; Sapele
(Sampson),1 2. Hap. 24 mm.
Sinna floralis, sp. n.
g. Head and thorax bright yellow; third joint of palpi
and antenne blackish, brilliant crimson stripes on back of
head and tegule, on dorsum of thorax, and obliquely from
base of fore wing across patagia to metathorax ; pectus, legs,
and abdomen white, the last with broad black-brown band
on ventral surface of penultimate segment, the extremity
blackish. Fore wing whitish suffused with yellow, the apical
area from costa beyond middle to above tornus deep golden
yellow ; three quadrate black spots on basal half of costa
with brijliant crimson bands arising from them, oblique and
not reaching inner margin, but bifurcating and uniting to
form reticulations in and below cell, enclosing three irregular
patches of the ground-colour and a point at lower angle of
cell; an oblique black fascia on medial part of vein 3; a
black spot at apex and a curved series of three spots from
costa before apex to termen above middle; a black bar on
termen at tornus. Hind wing nearly pure white. Under-
side white ; fore wing with oblique blackish bar from middle
of costa expanding into an elliptical discoidal patch, the spots
on terminal area as above.
Hab. Penane (S. 8S. Flower), 1 3 type. Hap. 40 mm.
Sinna atrifusa, sp. 0.
¢. Head and thorax white; antennz brown; tegule and
shoulders with small fuscous spots with some crimson scales
on them; abdomen white, dorsally suffused with fuscous
except at base, leaving whitish segmental lines and dorsal
streak. Fore wing black-brown; an oblique white fascia
from base of costa along inner margin to before middle ; the
costal edge white, expanding into a slight patch just before
middle; an almost obsolete oblique crimson fascia from
below costa near base, and thence above vein 1] to near
termen joined by traces of medial, postmedial, and subterminal
crimson bands; an apical yeilowish-white patch bounded
on inner side by an obliquely curved black band from just
below costa to near termen; the termen narrowly yellowish
new Species of Noctuide. 603
white to submedian fold. Hind wing pale ochreous yellow,
the costal area whiter. Underside pale ochreous yellow, the
fore wing tinged with fuscous except apical area.
Hab. Java, Arjuno, 3000 feet (Doherty), 1 g type. Exp.
40 mm.
Titulcia meterythra, sp. 0.
?. Head and thorax dull yellow, irrorated with fuscous,
the vertex of head, collar, and patagia marked with white ;
abdomen rufous. Fore wing with the basal area, costa, and
termen greenish yellow irrorated with fuscous ; two subbasal
white points; an irregular antemedial white band; a broad
postmedial fuscous-brown band with sinuous edges, narrowing
to costa and followed by a white band not reaching costa.
Hind wing rufous, the costal area white,
Hab. Borneo, Pulo Laut (Doherty), 1 ¢ type. Ezp.
20 mm.
Siglophora hematica, sp. n.
3. Head, thorax, and abdomen yellow suffused with red,
the base of abdomen yellow with dorsal red crest. Fore
wing yellow suffused with red, especially on terminal half,
leaving yellow ante- and postmedial semicircular patches on
costa and small medial and apical spots; traces of an
obliquely incurved autemedial line and of waved medial and
postmedial lines; a slight brownish discoidal spot; termen
narrowly greyish fuscous from apex to vein 3. Hind wing
with the basal half yellow, the apical half red. Underside
with the red areas deeper fiery red.
Hab. Borneo, Pulo Laut (Doherty), 1 3 type. Ezp.
22 mm.
; :
Chandica semiflava, sp. n.
¢. Head whitish mostly suffused with crimson; palpi in
front, base of frons, and basal joint of antennez in front white,
antenne reddish; tegule white edged with rufous and fuscous ;
patagia rufous and fuscous ; thorax whitish ; pectus, legs,
and abdomen white, fore tibiz crimson in front, abdomen
dorsally brownish tinged with crimson except at base. Fore
wing red-brown to beyond middle, shading into blackish, its
outer edge oblique from costa to vein 6; terminal area pale
bright yellow, with some crimson suffusion on edge of dark
area, from apex to angle of dark area, and as a very ill-defined
line from this angle to termen above tornus; a terminal
series of minute crimson points. Hind wing with the basal
604 Mr. G. Lewis on
half whitish tinged with fuscous, the terminal area pale
yellow with a faint crimson tinge on edge of basal area and
on termen at apex and middle.
Hab. Borneo, 1 2 type. Hap. 20 mm.
Cacyparis cyclops, sp. 0.
¢@. Head and thorax olive-brown mostly suffused with
brilliant silver; palpi and legs yellowish; abdomen reddish
brown, the extremity and ventral surface yellow. Fore wing
grey thickly irrorated and striated with fuscous, the costal
area to middle and at apex and the middle of inner margin
olive-brown ; brilliant silver irroration from base of costa
along inner margin to middle, where there is a boss of black
scales suffused with silver, with a silver point beyond it; an
oblique silver line from costa before middle to median
nervure ; a silver striga from middle of costa and boss of
silver scales in upper angle of cell; a round blue-black sub-
apical ocellus defined by a yellow annulus, and with minute
yellow streak in centre, some silver before the ocellus; the
termen pale yellow-brown, with a maculate silver line on
termen and a small fuscous patchabove tornus. Hind wing
pale brown suffused with fuscous, the costal area ochreous to
near apex. Underside ochreous; fore wing with broad
subterminal band of fuscous suffusion not reaching costa or
inner margin.
?. Abdomen and hind wing ochreous yellow, the latter
with some fuscous suffusion on basal inner area and a large
subapical patch.
Hab. Sou.omons, Guadalcanar I. (Meek), 1 3,1 2 type;
Isabel I. (Meek), 1 8, 1 9; Rendover I. (Meek), 1 ¢@.
Exp. 40 mm.
LXVII.— On new Species of Histeridee and Notices of others.
By G. Lewis, F.L.S.
THE last paper on the Histeride was published in the
September number of this Magazine; the present one is
the twenty-sixth of the series.
List of Species.
Pygoccelis strigosifrons. Omalodes bullatus.
Teretrius distinctus, Lew. tuberculatus,
Platysoma leyidorsum. Hister impiger.
rosselense. Pachycreerus pullus, Gerst.
rufopygum, Paratropus chelonitis.
—— pygidiale. testudo, Gerst.
dorsale. Plegaderus rumaniz.
Omalodes striatipectus, Lew., 1900. Saprinus ceeruleatus.
new Species of Histeride. 605
Pygocelis strigosifrons, sp. n.
Cylindricus, niger, nitidus; fronte dense strigosa; rostro apice
truncato, leviter sinuato ; pronoto ante medium partim strigoso ;
prosterno marginato; pygidio valide punctato; tibiis anticis
4-dentatis.
L, 33 mill.
Cylindrical, black and shining; the head, the anterior part
is densely strigose with a small fovea on its vertex, behind
which there is a small smooth space; the thorax is con-
spicuously sinuous at the sides with the lateral striz well-
marked, surface rather closely and coarsely punctured,
punctures shallow and crescent-shaped anteriorly, others
before the middle but not close to the sides are strigose ; the
elytral punctures are less close and smaller; the pygidium
is coarsely and densely subocellately punctured and in the
female but slightly concave on its disk ; the prosternum is as
long again as broad, marginate on its four sides and slightly
sinuous at the coxee, where the margin is reddish, the anterior
and posterior edges are both sinuous, the surface has some-
what shallow and elongate points; the mesosternum, the
anterior part projects somewhat in the middle and the marginal
stria 1s interrupted behind the prominence, the lateral margin
is reddish near the cox, and on the surface the punctures are
relatively fewer and less close (yet evenly set) than those of
the prosternum and are more conspicuously cucullate; the
metasternum is similarly punctured ; the anterior tibiz are
4-dentate.
The prosternum of this species is shorter than that of
P. africanus, Lew., and the species is narrower and not so
large as P. usambicus, Kolbe.
Hab. Antongil Bay, Madagascar (Mocquerys).
Teretrius distinctus, Lew., 1902.
This species, a native of Cape Colony, has been bred by
Mr. C. O. Waterhouse in the British Museum from Acacia-
bark infested with Xylopertha and Sinoxylon.
Platysoma levidorsum, sp. n.
Oblongo-ovatum, convexiusculum, nigrum, nitidum ; fronte plana,
stria integra ; pronoto stria laterali ante basin sinuata; elytris
striis 1 integra, 2 basi abbreviatis, 3 basi obliqua, tenuiter
impressa ; propygidio impunctato, utrinque leviter foveolato ;
606 Mr. G. Lewis on
pygidio laterali conspicue impresso, vix dense punctato, margine
externo densissime punctulato; prosterno haud striato; meso-
sterno sinuato et marginato; tibiis anticis 4-dentatis.
L. 53 mill.
This species closely resembles P. sesquistriatum, Mars., but
differs by the frontal stria not being broken, the thoracic stria
being sinuous before the base, the second dorsal stria is not
interrupted near the middle, and the outer margin of the pygi-
dium is densely punctulate. The propygidium is without a
“transverse train of punctures.”
Fab. Dutch New Guinea.
Platysoma rosselense, sp. 1.
Ovato-parallelum, subdepressum, nigrum, nitidum ; fronte concava,
stria transversa integra; pronoto stria laterali valida, antice
late interrupta, utrinque subsinuata ; elytris striis 1-3 integris,
sinuatis, 4 brevi apicali; propygidio basi irregulariter punctato,
utrinque foveolato; pygidio basi conspicue bifoveolato, sparse
punctato; prosterno inter coxas sinuato; mesosterno sinuato
marginatoque ; tibiis anticis 4-dentatis.
L. 33 mill.
The elytral striation of this species and its facies generally
correspond with P. Paugam?, Le Guillou: the head is, how-
ever, more transverse, the lateral thoracic stria is not parallel
to the edge but sinuous, the propygidium is wider and partly
smooth; the pygidium is bifoveolate at the base, with rather
fine punctures, instead of coarse closely set points; and the
prosternal keel is sinuous between the coxe.
flab. Rossel Island, Louisiade Archipelago.
Platysoma rufopygum, sp. 0.
Ovale, subdepressum, nigro-piceum, nitidum ; fronte concava, stria
integra; antennis, mandibulis, femoribus pygidioque rufo-
brunneis; elytris striis 1-4 integris, 5 basi abbreviata, suturali
ante medium terminata.
L, 23 mill.
Oval, rather depressed, piceous, with the antennze, mandibles,
and legs, especially the thighs, reddish brown, the anterior
angles and lateral margins of the thorax with the pygidia
and abdominal segments are also reddish brown ; the head
anteriorly concave, stria complete, surface visibly punctured ;
the thorax, marginal stria complete, strongest behind the head,
lateral margins broadly, but not densely, punctured; the
elytra, striz 1-4 complete, 5 well shortened before the base,
new Species of Uisteride. 607
sutural is apical and does not reach the middle; the pygidia
are rather coarsely, evenly, and rather closely punctured; the
propygidium has two rather shallow foveee which are apical
rather than basal; the prosternum is smooth and without
striz ; the mesosternum is sinuous and very clearly marginate ;
the anterior tibiz are 4-dentate.
In form this species agrees closely with P. debile, Mars.,
but the dorsal strie and the coloration are very different. In
outline it is also similar to P. depressum, Lec.
Hab. Palembang, Sumatra.
Platysoma pygidiale, sp. n.
Ovatum, convexiusculum, nigro-piceum, nitidum; fronte plana,
stria integra late sinuata; pronoto impunctato, stria integra
margine laterali parallela; elytris striis 1-3 integris, 4 apicali,
5 dimidiata; propygidio transyersim grosse punctato; pygidio
anterius punctato, sed apice in totum levi; prosterno angustato,
haud striato; mesosterno sinuato, marginato; tibiis anticis
4-5-dentatis.
L, 24-31 mill.
This species frequently has a short discal sutural stria,
which is, however, not constant in length. P. pygidiale is
similar to P. elingue, Lew., from Ceylon in its general facies,
while the sculpture of the pygidium agrees closely with that
of P. Confucii, Mars. he epistoma is wide in both elingue
and pygidiale and the fifth dorsal stria turns slightly away
from the suture anteriorly, especially in elingue, in which
species the stria is longer.
Hab. Sumatra (H, Rolle). I have many examples, but no
special locality.
Platysoma dorsale, sp. n.
Ovatum, convexiusculum, nigrum, nitidum ; fronte impressa, stria
recta; pronoto stria integra, parum forti, margine laterali
parallela ; elytris striis validis, impunctatis, 1-3 integris, 4 dimi-
diata, 5 ultra medium abbreviata, antice sinuata, suturali brevi
media ; propygidio sat dense punctato ; pygidio dense punctato,
margine parte exteriore late levi; prosterno inter coxas bistriato ;
mesosterno emarginato, stria integra, in medio retrorsum acumi-
nata; tibiis anticis 4- 5-dentatis.
L. 32 mill.
This species may be recognised from the known species
which resemble it by its short discal sutural stria, by the
fifth dorsal stria being anteriorly sinnous, by the prosternal
strie being confined to the region of the posterior lobe and
being joined before and behind, and by the mesosternal stria
608 Mr. G. Lewis on
being angulate in the middle. It may be grouped with
P. integrum and silvestre, Sch., and thugnaum and elingue,
Lew., and others, which are oval and more or less convex,
with the lateral thoracic stria parallel to the margin.
Hab. Anamalai Mountains, 8. India (Andrewes, No. 692).
Omalodes striatipectus, Lew., 1900 = O. amazonius, Mars.,
1861.
Omalodes bullatus, sp. n.
Oblongo-ovatus, parum conyexus, niger, nitidus; fronte leviter
punctulata, canaliculata, stria retrorsum acuminata; pronoto
levi, stria marginali integra; elytris striis punctatis, 1-2 in-
tegris, 3 basi dimidiata; propygidio tuberculato, sat dense sub-
tiliter punctulato ; prosterno bistriato ; mesosterno stria marginali
late interrupta; tibiis anticis 4-dentatis.
L. 53 mill.
This species is more oblong than O. tuberculipygus, Sch.,
and the mesosternal marginal stria is not complete. The
prosternal striw are as long as the keél and diverge from each
other equally before and behind. The tubercles on the pro-
pygidium also are more prominent than the other three
species known with tubercles. 0. tuberculipygus, Sch., is the
only species at present known with a tuberculate propygidium
and a complete mesosternal stria, but this latter character was
not noticed by Schmidt.
Hab, Jatahy, Province of Goyaz, Brazil.
Omalodes tuberculatus, sp. n.
Ovatus, parum conyexus, niger, nitidus; fronte leviter punctata,
eanaliculata, stria retrorsum acuminata ; pronoto levi, stria mar-
ginali integra; elytris striis punctatis, 1-3 integris, suturali
postice punctis indicatis; propygidio tuberculato; prosterno
bistriato, sed striis brevibus ; tibiis anticis valde 4-dentatis.
L. 53 mill.
This species differs from O. bullatus in being more oval, in
having three dorsal strize complete, and the prosternal striz
are short and occupy nearly one half (the anterior part) of
the keel only. In this last respect they are similar to
O. tuberosus, Lew., which is, however, a larger species, with
a more circular outline, being described as “ breviter ovatus.”
Flab. French Guiana.
new Species of Histeridee. 609
Fister impiger, sp. n.
Breviter ovalis, parum convexus, niger, nitidus ; stria frontali semi-
circulari ; pronoto stria laterali unica integra post oculos sinuata ;
elytris striis subcrenulatis, 1-3 integris, 4 basi vix abbreviata
rudimento aucta, 5-6 ultra medium abbreviatis; propygidio
pygidioque dense punctatis ; prosterno inter coxas bistriato.
L. 33 mill.
HH. impiger is shorter and much more convex than HZ. navus,
Mars., and néponicus, Lew. It is most similar to the last,
but differs from it in having the thorax less wide, and the
lateral stria is incurved at the base and there is a shallow ante-
scutellar fovea near but not on the edge; the elytral fifth and
sixth strize are longer and the punctuation of the pygidia is
closer and rather finer. ‘The prosternal striae are short and
intercoxal, like those of navus; in niponicus they extend
along the sides nearly to the end of the keel.
Hab. Yunnan (Donckier),
Pachycrerus pullus, Gerstiicker.
Platysoma pullum, Gerst. Archiv fiir Naturg. xxxiii. p. 31 (1867) ; Lew,
Ann. & Mag. Nat. Hist. ser. 7, vol. iv. p. 11 (1899).
I copy here Gerstiicker’s description for the sake of
reference :—
“ Platysoma pullum, Gerstiicker.
“Oblongum, cylindricum, nigrum, nitidum, sneo-micans; pro-
thorace estriato, subtiliter punctulato ; elytrorum stria suturali
utrinque, secunda pone basin abbreviata, sequentibus subintegris.
** Long. 2 mill.
“, minimis hucusque cognitis speciebus. Caput con-
fertim subtiliter punctulatum, fronte utrinque sulcata, sulcis
clypeum versus convergentibus. Prothorax transversus,
lateribus parallelus, angulis anticis late rotundatis, posticis
rectis, basi in medio angulata, supra ubique punctis sub-
tilibus, basin versus majoribus obsitus. Coleoptera prothorace
dimidio longiora, parallela, apicem versus crebre punctata:
singula 6-striata, striis evidenter punctatis, suturali basi
apiceque, 2* haud procul pone basin, 6* antrorsum dimidiatim
abbreviata, 5° omnino integra, 3* et 4* utrinque parum abbre-
viatis. Propygidium fortius, pygidium subtilius crebre
punctata, nitida wneo-micantia: pedes sanguinei.
“Specimen unicum ad Kisuani, Masailand (mens Oct.
1862), captam.”
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 40
610 Mr. G. Lewis on
Paratropus chelonitis, sp. n.
Breviter ovalis, nigro-piceus, supra impunctatus, vix nitidus ; elytris
striis dorsalibus 1-4 integris, 5 versus, suturali ultra medium
abbreviatis ; propygidio basi tenuissime punctato.
L. 4 mill.
In outline this species is much less oval than P. ovddes,
Mars., from Senegal, of which there are three examples in
the Museum of Paris, but otherwise the excellent figures in
Marseul’s monograph well represent the sculpture both above
and below. I have seen specimens of P. chelonitis labelled
P. testudo, Gerst., but the elytral strie differ from Gerstiicker’s
species in the first four being complete. Gerstiicker, in
counting the dorsal strie, called the sutural the first ; his
third and fifth strie correspond, therefore, with the fourth
and second of Marseul and other writers on the Histeride.
Marseul (Ann. Soe. ent. France, Bull. p. lxvi, 1871) recorded
this species, of which there is a single specimen in the
Museum of Paris, as referable to P. testudo, but the subjoined
diagnosis of Gerstiicker’s will show he was mistaken. The
Pelorurus also noticed by Marseul (/. c.) from Abyssinia was
described by Schmidt in 1890 as P. formosus; there are
specimens also of it in the Paris Museum. Pelorurus
glaucopterus, Mars., which I have lately received from Beira,
is much larger than P. formosus and has a red pygidium.
Hab. Abyssinia (Raffray). In my own collection and in
the Museum of Paris.
Paratropus tesiudo, Gerst. Beitr. Ins. Zanz. p. 87 (1866).
Breviter ovalis, nigro-piceus, supra levis, nitidus; antennis pedi-
busque rufo-brunneis; clytris stria suturali basin versus,
secunda arcuata ante medium abbreviatis, tertio quintoque
integris.
L. 4 mill.
Hab. Kisuani, KE. Africa.
Plegaderus rumanie, sp. n.
P. discist simillimus, sed angustior; prosterno lobo impunctato et
margine antice utrinque sinuato ; tibiis anticis abrupte dilatatis.
L. 1} mill.
Above, this species is extremely similar to P, discisus, Er.,
but it is narrower in form generally, the lateral margin of
the thorax is less wide and the punctuation of its surface is
finer and much more sparse, especially behind the transverse
sulcus, and the inner elytral oblique stria is well marked and
sulciform ; in P. discisus the stria is obsolete. The elytral
new Species of Histeride. 611
surface also in P. rumanie is less closely punctured, and
some of the punctures near the scutellum are much larger
than those of P. discisus. Beneath, the differences are
greater. The anterior prosternal lobe is smooth and convex
(similar to that of nétidus, Horn) and slightly sinuous on
either side of the anterior margin, and the median canalicu-
lation of the metasternum is wide and deep throughout its
course and markedly smooth. The anterior tibiz also are
dilated abruptly, not rounded off like those of P. discisus.
The anterior lobe of the prosternum of P. discisus is very
clearly and closely punctured, but this character is not noticed
by Marseul in his monograph of 1856, p. 272.
Hab. Comana Vlasca, Rumania (A. ZL. Montandon).
Saprinus ceruleatus, sp. n.
Circularis, convexiusculus, ceeruleus, antennis pedibusque rufo-
brunneis, supra undique dense strigosus vel punctatus, striis
dorsalibus (subhumerali et suturali exceptis) obsoletis; pygidio
dense punctato.
L. 3 mill.
Almost circular in outline, rather convex, blue, and some-
what shining; the head is densely and rugosely punctured, with
a lateral stria over the eyes only, there is a slight impression
on the vertex; the thorax, marginal stria complete, surface
rugosely punctate behind the head, markedly and densely
strigose on the other parts; the elytra, strie, outer humeral
fine and complete, continuing along the apex and joined to
the sutural, which is obliterated by the sculpture close to the
scutellum, there are several indications of striae on the
humeral area, the oblique humeral being the most distinct,
the surface is densely and rugosely punctate and is without
the marked strigosity of the thorax; the pygidia are densely
punctate ; the prosternum, the keel is widely sinuous laterally,
with microscopic strigosity and irregular punctures on its
surface, the base widens out and is hatchet-shaped, the
strie do not follow the outline at the base, but they are
similarly joined before and behind in the form of an arch;
the mesosternum is nearly straight anteriorly, but the marginal
stria is widely sinuous, the surface has shallow punctures,
not closely set, on the metasternum the points are smaller ;
the antenne are reddish brown, with the club and scape
darker.
I do not know of any species similar to this.
Hab. Plateau de PAndroy, Région d’Ambovombe, Mada-
gascar.
40*
612 Col. C. Swinhoe on
LXVIII.—On new and little-known Species of Eastern and
Australian Lepidoptera. By Colonel CuarLes SWINHOE,
M.A., F.LS., &e.
Family Hesperiidae.
Casyapa dissimilis, nov.
¢. Antenne ochreous with brown bands; palpi, frons,
and head dark ochreous ; thorax, abdomen, and wings of a
uniform ochreous brown, entirely covered with dark ochreous
scales, without any markings ; cilia pale ochreous. Under-
side dark ochreous brown, without the ochreous suffusion
and also without any markings; fore legs with long dark
ochreous hairs.
@. Antenne brown, a long white space before the hooked
tip ; palpi dark ochreous; head, body, and wings of a
uniform dark olive-brown: fore wings with a broad, nearly
uniform white band, from the costa beyond the middle
to the outer margin above the hinder angle, where it is
joined to a curved white spot; cilia of both wings brown.
Underside pale, but more blackish: fore wings with the
band as above, the hinder margin and base suffused with
grey; hind wings with about the outer half coloured like
the fore wings, the abdominal half suffused with grey: body
grey, legs dark ochreous.
Expanse of wings, ¢ 3, 9 34%, inches.
German N. Guinea ; one pair.
Allied to C. nevifera, Mab., from Batjan, but can easily
be separated by the absence of the strong black mark at the
end of the cell of the fore wings in the male, which is so
prominent a character in Mabille’s species. There is a pair
from N. Guinea in the B.M. unnamed.
Tagiades curiosa, nov.
g. Antenne black, white on the underside of the club ;
palpi, head, thorax, and fore wings dark olive blackish brown
with an olive tint ; frons white, containing a brown band;
palpi white beneath: fore wings with an obscure discal
band, of three obscure duplex spots, rather near the outer
margin, rising from near the hinder margin more than
halfway up the wing; spots white, all small and linear,
with the exception of the outer three, which are round—one
at the end of the cell, two above it, one below it, all in an
oblique line, one above but outside the last, two above this
Eastern and Australian Lepidoptera. 613
one, near the costa, and the outer three subapical, near
together but almost at right angles to each other: hind
wings with the upper half of the same colour as the fore
wings but paler, lower half pure white; marginal border
black, composed of large spots joined together; an obscure
spot opposite the apex and a prominent one below it, both
rather close to the margin ; cilia brown, white towards anal
angle of hind wings: abdomen with the basal half brown,
the rest white. Underside black: fore wings with the spots
as above, some white suffusion at the end of the cell, and a
broad submarginal band of broad, duplex, obscure white
spots, which curves inwards below the apex, the duplex
spot near the hinder margin the largest and most pro-
minently white; hind wings pure white, with a broad
dark black costal border, the : spots as above : body and legs
white.
Expanse of wings 2,7, inches.
German N. Guinea; one example.
Allied to 7. clericus, Butler, which has a black abdomen
and has no smeared whitish mar kings on the outer portions of
the fore wings below. There is an “example from Aru Isl. in
the B.M. unnamed.
Genus IMA, nov.
Antenne slender, two thirds the length of the costa,
club long, tip curved and of moderate length ; palpi up-
turned, rising as high as the vertex, densely covered with
thick hair, third joint short, smooth, and with a blunt point ;
thorax and abdomen moderately stout, the abdomen reaching
a little lower than the hind wings ; legs hairy, tarsi smooth
and without spurs: fore wings wie the costa arched at the
base, outer margin oblique, apex consequently rather pro-
duced, hinder margin straight, slightly longer than the outer
margin, hinder angle rounded : hind wings with the costa
and apex rounded, outer margin waved below the middle,
anal angle blunt, abdominal margin waved; vein 5 from
above the middle of cell.
Lima jovina, nov.
3 9. Of a uniform dark bright olive-brown colour; palpi
below white, with brown hairs ; fore wings with a broad
ochreous band from the costa of fore wings ‘at the middle to
near the hinder margin before the hinder angle, the band is
fairly uniform in breadth but has irregular margins on both
614 Col. C. Swinhoe on
its sides and is excavated on its inner side on the submedian
vein; cilia of fore wings brown, of hind wings white.
Expanse of wings, ¢ 2,3,, 2 2 4; inches.
East Celebes ; one pair.
I received these three years ago from a dealer on the
Continent under the name of Hidari joviana, and there are
two males in the Crowley Collection in the B.M. as H. Irvint,
but I can find no reference for either name; I have gone
through a great deal of literature, and can only come to the
conclusion that the species has never been described and
published. Except on the waved outer margin of the hind
wings it has the shape of Erionota thrax, but this and the
third joint of the palpi separate it from the genus Hrionota ;
the neuration is similar, but so it is in many genera of this
family.
Telesto uniformis, nov.
g. Palpi beneath ochreous grey with brown hairs ; frons
with an ochreous-grey ring, brown in the centre; antennz
blackish brown with ochreous-grey dots on the underside
and with the underside of the club suffused with the same
colour; head, body, and wings of a uniform dark blackish
brown, without any markings, except for the sexual and
linear brand, which runs in a more or less waved line from
near the origin of vein 4 ot the fore wings to the hinder
margin beyond the middle and is grey in colour; underside
paler, without any markings.
Expanse of wings 1,4, inch.
Ké Island ; two examples.
Telesto saxuda.
Hesperilla saxula, Mab. Ann. Soc. Ent. Belg. 1891, p. Ixxxi, 9.
¢. Of a uniform olive-brown colour; palpi white below,
black above, last joint black: fore wings with the sexual
linear brand pale, nearly white, outwardly lined with black,
a white dot outside its upper end, two subapical white dots,
rather far apart from each other; no other markings; cilia
a little paler than the wings, with a tinge of ochreous.
Underside paler than above, the hinder marginal space of
fore wings pale, the three white dots as above; fore legs and
body with white hairs.
Expanse of wings 1,3, inch.
Mackay, Queensland ; three examples.
Mabille’s female type came from Cooktown, Queensland,
Eastern and Australian Lepidoptera. 615
and his description fits my examples very well, considering
the usual sexual differences in this family.
In his Monograph of the Hesperiide in Wytsman’s
‘Genera Insectorum,’ fase. 174, p. 182 (1904), by some
extraordinary error, Mabille puts his species under Godman
and Salvin’s genus Halotis, with Costa Rica as its locality,
In the ‘ Biologia,’ Insecta, Lep.-Rhop. ii. p. 505, pl. xev.
ff. 42, 43, 44, g (1900), a Hesperiid from Costa Rica is
described and figured as the type of the genus Halotis; but
neither the description nor the figures represent the Queens-
land insect. One of the ‘ Biologia’ examples, it is said,
is labelled as having been compared by Salvin with the type
of Hesperia saxula, Mab., a description of which could not
be found; this must refer to some Hesperiid from Costa
Rica so named by Mabille, which never was described and
published ; it can have no reference to the Cooktown insect.
Mimas melie.
Mimas melie, de Nicé. Journ. Bomb. N. H. Soe. ix. p. 394, pl. Q.
f. 55, Q (1895).
6. Both wings and cilia dark brown with an ochreous
tinge: fore wings with obscure ochreous streaks from the
base in the interspaces, the two lowest extending to the
discal dull ochreous oblique band, which is composed of
three spots, the lowest the largest: hind wings with the
inner portion of the wing smeared with dull ochreous,
extending on the abdominal margin from the base to one
third from the anal angle, narrowing inwards to a large
square ochreous spot in the upper disk. Underside rich
purplish brown, markings pale ochreous yellow, a large spot
at the end of the cell, a large space commeucing with the
uppermost spot of the discal band, widening hindwards and
extending on the hinder margin from the middle of the
wing to the hinder angle, a duplex subapical spot in a
line with the inner margin of the band, and four lnear
subcostal spots close together, a little on the inner side
of the wing: hind wings with two large spots—one below
the origin of vein 2, and the other towards the outer margin
in the interspace above vein 4: palpi, head, and body with
ochreous hairs; legs dull ochreous grey.
Expanse of wings 14% -1,%) inch.
Humboldt Bay, N. Guinea; two examples.
The female, as described and figured by de Nicéville, has no
band on the fore wings, but the peculiarly marked underside
is identical ; vein 5 of the hind wings is also absent, and the
616 Col. C. Swinhoe on
neuration similar. Both are from the same locality; I
therefore cannot help being convinced that they are sexes of
the same species. One example has on the hind wings below
an extra dot near each spot.
Padraona tranquilla, nov.
3 ¢. Palpi brown with ochreous hairs, white beneath and
at the tips; head, body, and wings red-brown with a dull
olive tinge; head and thorax with ochreous hairs: fore
wings with a thin, oblique, dull ochreous discal band com-
posed of six spots close together and three similarly coloured
incurved subcostal dots near apex: hind wings with a short
discal band of five spots close together; cilia of both wings
ochreous. Underside: fore wings nearly all black, a sub-
costal ochreous streak from base to beyond the middle, some
ochreous suffusion on the apical space, the discal band and
dots as above: hind wings ochreous, the band as above but
with a black outline; both wings with the outer marginal
line black, cilia ochreous: face and pectus white ; thorax
grey; legs grey with white stripes ; abdomen white.
Expanse of wings 1,4) inch.
Milne Bay, N. Guinea; one male. Type female in B.M.
This is not a typical Padraona, there being a sex-mark
consisting of a line of raised scales on the fore wing of the
male, bordering the inner sides of the first four spots forming
the discal band. When more examples are procured, it will
come into a new genus between Padraona, Moore, and
Oxybydastes, Heron.
Padraona tabla, nov.
¢. Dark brown, tinged with ochreous ; markings orange-
ochraceous : fore wings with a small spot at the upper end
of cell, two obscure streaks from the base in the two lowest
interspaces touching the discal band, which is linear, out-
wardly oblique, and consists of three spots; four small
subapical spots in pairs, almost at right angles to each
other ; these spots are obsolete in one example: hind wings
with a discal band, broader than the band of the fore wings,
consisting of five conjoined spots ; cilia of both wings orange-
ochreous ; palpi, frons, and head with ochreous hairs. Under-
side bright orange-brown, the whole surface suffused with
orange-colour ; spots as above, with the addition of a spot
at the end of the cell of the hind wings.
Expanse of wings 14 inch.
Eastern and Australian Lepidoptera. 617
Humboldt Bay, N. Guinea (Doherty) ; two examples.
Resembles P. concinna, Elwes, from the Nilgiri Hills, of
which I have many examples.
Padraona batchiana, nov.
3d. Dark brown, suffused with ochreous; palpi and frons
with bright orange-ochreous hairs; wings with dark orange-
ochreous markings: fore wings with some obscure longi-
tudinal streaks near the base and an obscure streak along
the hinder margin, on the inner side of the discal band,
which is outwardly oblique and composed of three spots, the
lowest lunular, the middle spot with its inner edge a little
more inwards than the others, and two small linear spots
close together below the costa, almost in a line with the
band ; hind wings with an upper discal short band of rather
larger spots divided by the veins into four spots; cilia of
both wings orange-ochreous. Underside bright orange-
brown; spots on the fore wings same as above, the lowest
spot of the band with two suffused white spots in a line
towards the base, some black suffusion above these spots, and
some more at the hinder angle of the wing: hind wings
without markings ; some indistinct, suffused ochreous stripes
in the interspaces on the outer portions of both wings.
Expanse of wings 1,’) inch.
Batjan (Doherty) ; one example.
Padraona silativa, nov.
3g. Ochreous brown; palpi at the sides and frons orange-
ochreous, some hairs of the same colour on the head and on
the sides, basal end of thorax, and anal tuft; wings marked
with orange-ochreous: fore wings with a thin costal streak
on basal half, a broad basal streak in the cell, divided by
a black line, its upper part running nearly to the end of the
cell; two basal streaks on the two lowest interspaces, reaching
the discal band, which is oblique and consists of five large
spots, the uppermost ones the smallest, then two dots joining
it to the three linear subapical spots, which are close
together: hind wings with a wedge-shaped spot in the
cell, with its point inwards, a discal band divided into five
spots and a dot by the veins, some long ochreous hairs
forming two obscure stripes near the abdominal margin ;
cilia of the fore wings brown, ochreous towards hinder
angle ; cilia of the hind wings entirely ochreous. Underside
paler; markings as above, except that the wedge-shaped spot
is replaced by a small yellow spot, and there are some obscure
618 Col. C. Swinhoe on
streaks in the interspaces on the outer portions of the wings ;
pectus ochreous; legs striped with ochreous ; tarsi entirely
ochreous.
Expanse of wings 1,’ inch.
Humboldt Bay, New Guinea ; one example.
Resembles P. sperthias, Felder; has a very obscure
sinuous sex-mark of andraconia scales on the inner side
of the discal band of the fore wings.
Ismene lysima, nov.
@. Of a uniform dark pink-brown with an ochreous tinge,
clubs of the antenne ochreous beneath, as also is the under-
side of the palpi; both wings with some dark orange suffusion
in the basal portions, formed by obscure streaks in the
interspaces, no other markings. On the underside the wings
are slightly paler, a broad dull orange-red streak along the
costa of fore wings and some obscure streaks of the same
colour in the interspaces on the outer portions of the wing:
hind wings with a broad space on the costa at the base
smeared with the same colour; no other markings: legs
streaked with ochreous.
Expanse of wings 1,9) inch.
Ké Island ; one example.
Hasora palinda, nov.
g. Ofa uniform dark pink-brown ; underside of palpi
ochreous, some blackish suffusion at the anal angle of hind
wings; wings without any markings above. On the underside
the fore wings have a pale streak at the end of the cell and a
pale transverse submarginal streak on the upper half; the
hinder marginal area whitish: hind wings with a pale discal
thin band, quite straight from the costa near apex, ter-
minating in a short white mark where it joins the large
black rounded space on the anal angle ; a whitish spot on the
abdominal margin, just outside the black mark: legs with
long ochreous hairs.
Expaunse of wings 2,45 inches.
Soekaboeni, Java; one example.
Like H. Ribbei, Plotz, but that form has the hind wings
glossed with blue below.
Family Aganaide.
Asota latiradia, nov.
9. Palpi with first two joints ochreous, brown stripes at
Eastern and Australian Lepidoptera. 619
the sides above, the upper third white beneath, last joint
black, white beneath ; frons, head, and thorax dull ochreous,
tinged with greenish grey ; thorax with six small black spots
in front ; abdomen bright orange-ochreous, with short black
dorsal bands on the segments: fore wings purplish brown,
basal patch coloured like the thorax, containing small black
spots, three subcostal, one below near the base, and three in
a row on the outer margin of the patch, all the spots circled
with white, the white on the outer spots angled outwards ; a
very broad white medial streak, its upper part occupying
half the cell, its lower part gradually produced almost to a
point along vein 2 about one sixth from the outer margin,
the outer edge irregular; vein 1 and the hinder margin
white for about half the distance from the base: hind wings
white, with a black marginal border, broad at the apex, evenly
and gradually attenuated to the anal angle, which it does not
quite reach.
Expanse of wings 2,3, inches.
Babber Island, west of Timor-Laut; one example.
There is nothing like it in my collection, but Dr. Jordan
tells me it somewhat resembles streaked specimens of
A, timorana, Roths. Nov. Zool. iv. p. 349.
Asota symmorpha, nov.
6 2. First two joints of the palpi, frons, head, thorax,
and abdomen bright orange-ochreous ; the palpi with a streak
above and the last joint black ; two large black spots on the
collar and two behind them; thorax with a broad black
longitudinal stripe ; abdomen with broad black segmental
bands above and below, disconnected at the sides and at the
middle below: fore wings purplish brown with the basal
patch very small, containing two large black spots ; a white
narrow stripe along the median vein, with a round club at
the end, extending a little beyond the end of the cell, and
in the male not reaching the base: hind wings white, a
small blackish spot at the end of the cell, and broad mar-
ginal black borders, slightly broadest at the apex, running
up the abdominal margin.
Expanse of wings 2;£, inches.
Saparua Island ; one pair.
Allied to A. doryca, Boisd., which I have from Ké, N. Ire-
land, N. Guinea, and Fergusson Islands, but differs in its
bright yellow body and has no streaks on its veins and no
points extending from the club; the bands on the abdomen
of the female are less pronounced than those in the male.
620 Col. C. Swinhoe on
Family Arctiide.
Diacrisia aurapsa, nov.
9. Antenne ochreous; palpi, head, thorax, and fore
wings dark brown with a pinkish-ochreous tinge; abdomen
dull crimson above, with short segmental dorsal bands,
leaving a broad crimson band on each side. Body below and
legs dark brown without markings: fore wings with very
indistinct traces of markings, a brown spot at upper end of
cell, and two lines or thin bands across the disk, arising
rather close together on the hinder margin beyond the
middle, separating upwards, the outer line apparently reaching
the apex: hind wings paler brown without markings.
Expanse of wings 1,8, inch.
Saparua Island ; one example.
Allied to D. landaka, Moore, from Java.
Family Drepanulida.
Tridrepana adelpha, nov.
¢@. Of a uniform pale ochreous colour ; shafts of the
antennze, costal line of fore wings, and cilia of both wings
dark pinkish brown; a pale purplish spot in the cell of fore
wings and another at the end; both wings with traces of
antemedial and postmedial recurved lunular lines; a large
pale purplish patch in the centre of each wing, each con-
taining three brown spots: fore wings with a narrow brown
mark below the apex, limited by a brown submarginal line
which runs near the margin from the middle of the wing to
the apex. Underside much paler, the patches on the wings
showing through ; no other markings.
Expanse of wings 1,2, inch.
Khasia Hills; many examples.
Allied to T. sadana, Moore, from Sikhim, and generally
mixed up with that species, but no one with a series of each
before him could possibly put them together.
Family Acontiide.
Erastria crotopha, nov.
d @. Palpi, head, and thorax olive-brown, a white spot
on the frons, some white dots on the thorax; abdomen and
the tufts dark brown, some whitish at the sides, the first two
segments and anal tuft also whitish: fore wings olive-
ochreous, irrorated with olive-brown, bands and markings
Eastern and Australian Lepidoptera. 621
dark olive-brown, a sinuous subbasal line, inside of which is
a patch on the costa and a small spot, also on the costa
between it and the medial band, which is contracted above
the middle and is slightly outwardly curved ; a discal crenu-
late thin band, a large subapical costal patch, and two
smaller costal patches between it and the medial band; a
paler patch at the apex, some markings below it and a black
line on the margin of disconnected lunules, inwardly edged
with whitish; cilia with a brown line and pale spots: hind
wings pale brown without markings. Underside pale brown ;
a cell-spot on the hind wings and a brown, outwardly curved,
discal line across both wings.
Expanse of wings, ¢ 1,4, 2 14%, inch.
Khasia Hills; five males and one female.
Allied to E.aurata, Moore, from Sikhim ; the female is paler
than the male. There are two examples from the Khasia
Hills in the B.M., Drawer 181.
Eublemma atimeta, nov.
3 ¢. Frons and head whitish ; body and wings of a uni-
form dull whitish ochreous: fore wings with the costal line
dull pink, two trausverse oblique lines, rather indistinct, the
first from near the hinder margin before the middle to near
the costa beyond the middle, the other a little beyond,
parallel, extending to the apex, and apparently duplex; the
outer margin shaded with pinkish ochreous, with cilia of the
same colour, and a row of duplex points near the outer margin :
hind wings without markings : abdomen below whitish towards
the extremity.
Expanse of wings ,/, inch.
Padang, Sumatra ; one pair.
Tarache procrita, nov.
3d. Frons and vertex of head white; collar broadly dark
chestnut-brown ; thorax brown witha white band in front:
fore wings with the ground-colour white, the bands and
markings dark chestnut-brown ; a patch on the costa before
the middle, complete in some examples, broken in the type ;
a broad medial band, more or less dislocated and much con-
stricted at the end of the cell, widening on the hinder margin
with a white spot within it, absent in some specimens; a
dislocated marginal band, connected above the middle with
the medial band; a white spot at the apex ; the white portions
of the wing more or less irrorated and marked with brown
622 Col. C. Swinhoe on
atoms: hind wings and abdomen dull brown. Underside
with the fore wings dull brown; a white dot on the costa
before the apex and a white spot at the apex: hind wings
dull whitish suffused with pale brown, a brown dot at the
end of the cell ; medial and discal brownish thin bands.
Expanse of wings ,8, inch.
Khasia Hills; several examples.
Somewhat resembles 7. noloides, Butler, from Japan.
Rivula magniplaga, nov.
&. Palpi with black stripes above and below and ochreous
brown at the outer sides, ochreous inside, whitish at the tips ;
frons and vertex greyish brown, collar pale; thorax and fore
wings brownish ochreous: fore wings with a broad, even,
outwardly oblique, medial black band, which only reaches
the middle of the end of the cell and is limited on its outer
side by a white line which is abruptly angled inwards on to
the costa ; from the angle of this line there is a brown patch
which widens on to the outer margin below the apex, and
from its middle to the hinder margin there is a thin, some-
what sinuous, brown band; on the margin within the patch
is a white line and the cilia of both wings are brown: hind
wings ochreous grey, without markings. Underside much
paler than above, with a large brownish triangular patch
occupying the greater part of the basal half of the fore wing ;
tarsi with white spots.
Expanse of wings 1 inch.
Khasia Hills ; two examples.
Family Quadrifide.
Genus Masca, Walker.
Masca, Walker, xvi. 8 (1858).
Phagytra, Walker, xxxiy. 1508 (1865).
Masca abactalis.
Masca abactalis, Walker, xvi. 9.
Khasia Hills ; one pair.
The type from Singapore is in Mus. Oxon. ; it is not men-
tioned by Hampson in his ¢ Moths of India.’
In Cat. Het. Mus. Oxon. i. p. 165 (1900) I put Phagytra
leucogastralis, Walker, xxxiv. 1508, type from Java in B.M.,
= Metria platypoda, Felder, Reise Nov., Lep. pl. 120. f£. 44
(1873), type Amboina, in Coll. Rothschild, as a synonym to
the above ; but in this I think I was wrong: leucogastralis
Eastern and Australian Lepidoptera. 623
is a darker insect with the hind wings more heavily bordered.
My Khasia Hills examples are identical with Walker’s
Singapore type; I have abductalis also from Singapore, Obi
Island, and Waigiu Island, and /euwcogastralis from Gilolo,
Fergusson Island, and Amboina.
Family Focillide.
Egnasia rudmuna, nov.
3S. Palpi, antenne, head, thorax, and fore wing ochreous
brown: fore wings with ante- and postmedial outwardly
curved whitish lines, inwardly lined with brown, the former
with an outward brown spot on the costa, the latter with a
costal spot on each side; both lines rather acutely bent in-
wards below the costa ; the first lime with a white spot just
outside it in the cell ; the second line with a much larger
and distorted white spot inside it at the end of the cell; a
submarginal brightly sinuous white line, curved round a
large blackish subapical spot, and some black lunules on the
inner side of the line; some marginal black lunular points
outwardly marked with white; marginal line ochreous; cilia
ochreous, with a pale brown interline or band: hind wings
pale ochreous grey, brownish on the outer portions; indi-
cations of a waved pale submarginal thin band ; two black
white-pointed spots, one above the other, near the abdominal
margin above the anal angle; marginal line black, lunular ;
cilia as in the forewings. Underside ochreous grey: fore wings
with indications of the upperside markings: hind wings with
three transverse, highly sinuous, brown lines with some very
prominent black spots on them, including a lunular mark at
the end of the cell; antennz bipectinate, with stiff bristles
and cilia: fore wings narrow, not angulated, outer margin
oblique ; hind wings normal.
Expanse of wings 1,2, inch.
Kina Balu ; three examples.
Rhesala tenuilinea, nov.
$. Ofa uniform grey-brown: fore wings with five white
costal points on the apical third; both wings with five thin,
sinuous, and outwardly curved brown lines, subbasal, ante-
medial, medial, discal, and submarginal, at equal distances
apart, the third running through an 8-shaped orbicular mark,
the last composed of dots: hind wings with three similar
lines, corresponding to the last three of the fore wings ; both
wings with a brown marginal crenulated line; a similar line
624 Col. ©. Swinhoe on
in the cilia, which have ochreous spots at the tips. Underside
paler ; costa of fore wings and basal portions of both wings
with some whitish suffusion; lines as above, reniform and
orbicular white and prominent ; tarsi banded with white.
Expanse of wings 1 inch.
Padang, W. Sumatra; one example.
Near R. commoda, Walker, and R. nigriceps, Hmpsn.
Family Deltoidide.
Bocana subalbida, nov.
d. Antenne white, tinged with pink ; palpi pure white be-
neath ; palpi above, head, body, and wings pale pinkish brown,
almost a rosy tint : fore wings with a white dot near base of
cell, a large ear-shaped white mark at the end; antemedial,
postm edial , and discal, transverse, slightly sinuous brown lines,
terminating in white dots on the costa, the first crossing the
white dot, the second outwardly curved: hind wings with the
costal space broadly white; an indistinct cell-mark and
three transverse brown lines, medial, discal, and submarginal ;
marginal line of both wings brown; abdomen with a pure
white anal tuft. Underside: fore wings with the upper half
pale pinkish brown, the lower part white, unmarked ; hind
wings pure white on the basal portions, the outer pale pinkish-
brown lines as above, a prominent black cell- spot ; body aud
legs pure white.
Expanse of wings 1,7, inch.
Batjan ; one example. There is one from Amboina and
another from Fergusson Island in the B.M. unnamed,
Deltoid drawer 36.
Oxvenanus magniplaga, nov.
3. Antenne, palpi, head, body, and wings pale brown,
tinged with pinkish and ochreous : fore wings with sinuous,
ante- and postmedial transverse grey lines, the former inwardly
and the latter outwardly oblique, being rather close together
on the hinder margin of the wing; the former ends in a black
spot on the costa with a black dot below it inside the cell; a ver
large ear-shaped black patch at the end of the cell, extending
up to the costa, edged outwardly with whitish, with a sinuous
grey line running Aoean from it to the hinder margin ; a broad
pale space across ‘the disk of the wing, which is continued across
the middle of the hind wings, through which is a sinuous grey
line; the hind wings otherwise unmarked; marginal line of
Eastern and Australian Lepidoptera. 625
both wings ochreous, with black lunular points. Underside
much paler, with corresponding transverse lines and a black
spot at the end of each cell.
Expanse of wings 14 inch.
Khasia Hills ; one example.
One example from Sikkim in the B.M., Deltoid drawer 37,
with Mosopia megaspila, Walker, Hampson, Moths India, iii.
p- 50; but it is quite distinct from Walker’s Penang type.
Genus Lienicrpa, nov.
6. Antenne with minute cilia; palpi as long as the head
and thorax, second joint porrect, with long brush-like hair
above, a small brush standing out at the end below, third joint
nearly as long as the second, abruptly erect, short thick hair on
the underside, a thick brush of long hair on the upperside ;
thorax and abdomen smooth, the latter not extending beyond
the wings ; legs moderately hairy ; hind tibiz with two pairs
of spurs: hind wing with vei 5 from near lower end of cell :
fore wing long, rather narrow, apex acute, angled at the
middle of the outer margin ; costa rounded at the base: hind
wings with the apex somewhat produced, the outer margin
nearly straight, rounded at the anal angle.
Type, L. echana, nov.
Lignicida echana, nov.
3. Antenne and palpi red-brown; body and wings of a
uniform ochreous brown: fore wings with four rather large
black spots on the costa, at nearly equal distances apart, being
terminations of very indistinct, highly sinuous, transverse
brown lines, the third the most distinct and continued across
the middle of the hind wing; a faint indication of a pale
discal band across both wings between the last two lines,
marginal lunular black points ; cilia pale, containing a thick
blackish line. Underside paler and more ochreous, the last
two transverse lines prominent, and a brown dot at the end of
each cell; body and legs without markings.
Expanse of wings 12 inch.
Palawan (Waterstradt); one example.
There is an example from Bali in B.M. unnamed, Deltoid
drawer 14.
Family Hypenide.
Britha biguttata.
Britha biguttata, Walker, xxiv, 1147 (1865); Hmpsn. Moths India,
iii. p. 94 (1895).
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. Al
626 Col. C. Swinhoe on
Herminia (2) incertalis, Walker, xxxiv. 1518.
Hypena colabalis, Felder, Reise Nov., Lep. pl. cxx. f. 29 (1878).
Type, Moreton Bay, in B.M.
Type (incertalis), Java, in B.M.
Type (colabalis), Amboina, in coll. Rothschild. I have it
from Queensland, Pulo Laut, Sarawak, and Ceylon. Felder’s
name and reference have been heretofore overlooked.
Family Epiplemide.
Epiplema pecilaria, nov.
g. Palpi black, a white band at the base of third joint ;
frons white ; head and collar chocolate-pink, the latter with
a white line in it; thorax and a broad band on hinder
margin of fore wings pure white; the remamder of the fore
wings with the ground-colour white but thickly irrorated
with chocolate-pink, especially on the outer portions of the
wing ; a white spot in the cell, divided by a black mark,
another at the end; a pink indistinct band from the middle
of the costa, angled outwards, and is lost in the brownish
suffusion in the disk ; this is margined outwardly by a white
band, which is indistinctly traceable in an attenuated and
reeurved form to the hinder angle, where there are three
black spots in a line on its outer edge; outside this band on
the apical portion of the wing there is some dark brown
suffusion ; marginal line and cilia pale pink with some brown
spots: hind wings with the costal third pure white, its outer
edge irregular and very sinuous, the remainder of the wing
brown-pink; a short white band, lined with black, at the
middle of the abdominal margin; some blackish transverse
short lines, a white dot in the disk; abdomen pinkish grey.
Underside: fore wings suffused with dark brown, a white
band on the hinder margin; hind wings white, slightly
suffused with pinkish grey on the lower portions; body and
legs white.
Expanse of wings 141, inch.
Padang, W. Sumatra; one example.
Fore wings with the outer margin rounded ; hind wings
with two tails. Superficially hke a very large example of
E. desistaria, Walker.
Family Boarmiide.
Tristrophis veneris.
Urapterya veneris, Butler, Ann. & Mag. N. H. (5) i. p. 892 (1878).
Eastern and Australian Lepidoptera. 627
Tristrophis veneris, Butler, Journ. Linn. Soc., Zool. xvii. p. 200, pl. ix.
fig. 84 (1883) ; Swinhoe, Cat. Het. Mus, Oxon. ii. p. 230 (1900).
Jaintia Hills; one example.
The type from Yokohama is in the B.M. I have it also
from the same place and from Asama Yama. It is not
recorded by Hampson.
Pogonopygia nigralbata.
Pogonopygia nigralbata, Warren, Noy. Zool. i. p. 681 (1894) ; Hmpsn.
Moths India, iii. p. 619 (1895).
Jaintia Hills; one example.
I have it also from Japan. There is no specific difference
between the Japanese and Indian example, and Sir George
Hampson was therefore wrong in doubting the Tring locali-
ties. Warren quotes Japan, Central China (many examples),
and Khasia Hills (two examples) as his localities, but he
does not state the locality of the specimen on which he bases
his type.
Leptesthes circumflexaria.
Ennomos circumflexaria, Kollar, Hiigel’s Kasch. iv. p. 485 (1842).
Lagyra megaspila, Moore, P. Z. 8. 1867, p. 616.
Hyposidra (Leptesthes) megaspila, Hmpsn. Moths India, iii. p. 215
(1895).
Kollar’s type came from Massuri; Moore’s type from
Bengal is in the B.M. I have it from the Khasia Hills.
L. circumflexaria is not mentioned by Hampson.
Medasina pallidimargo, nov.
@. Frons and head white; body and wings grey, thickly
irrotated with dark brown: fore wings with an outwardly
eurved, antemedial, black band, bluntly angled below the
costa, and again below the cell, where it is bent inwards and
then straight to the hinder margin, one fourth from base,
and includes a black spot inside the cell; a straight black
band from the middle of the costa to a black spot at the end
of the cell; two short, black, costal bands between this and
the apex, at equal distances apart, the last being the termi-
nation of an irregular band which limits the white outer
marginal space, this limitation is bent inwards on vein 3 and
then is continued in a somewhat sinuous form to the hinder
margin near the hinder angle ; in some examples there is a
discal lunular black line from the penultimate costal band
which is continued across the hind wing a little beyond the
41*
628 On Eastern and Australian Lepidoptera.
middle, and on this wing there is a black spot at the end of
the cell, and a submarginal irregular and indistinct band,
limiting the white marginal space, which is more covered with
irrorations than it is on the fore wings ; ciliaof both wings
brown with white spots: the underside is paler and whiter,
markings mostly visible, the white marginal space much
clearer.
Expanse of wings 245 inches.
Kashmir; nine examples, all females.
Allied to nothing I know of.
Abraxas languidata.
Abraxas languidata, Walker, xxiv. 1122 (1862).
Calabraxas languidata, Cotes & Swinhoe, Cat. Moths India, ii. no, 8726
(1888) ; Hmpsn. Moths India, iii. p. 518 (1895).
Khasia Hills ; a fine series.
I have it also from Japan ; there is no specific difference.
Walker’s type is recorded from Nepal (Hearsay), but Hampson
states it is from Japan and omits it from the Indian record.
Hearsay never collected in Japan.
Family Sterrhide.
Emmiltis elyra, nov.
g. Of a uniform dull yellow colour; frons black with a
yellow band: wings with a black spot at the end of each
cell; a pale grey, straight, thin band from near apex of fore
wings to the abdominal margin of hind wings before the
middle ; a crenulated grey line from near apex of fore wing
to the hinder margin near the angle, then across the disk of
the hind wings; on the fore wings inside this line is a row of
black spots, some of which on the lower portion of the wing
are large and prominent ; on the hind wing there is a sub-
marginal similar line, rather close to the discal line, with
four rather large black spots inside it. Underside paler,
cell-spots on all the wings; on the fore wings the two
bands are visible and some brown suffusion at the base.
Expanse of wings 1,45 inch.
Kashmir ; one example.
The hind wings are angled at the middle.
Anthometra unipuncta, nov.
¢@. Of a uniform ochreous-white colour; top of head
black ; costal space of fore wings slightly irrorated with
ochreous grey, and a very slight tinge of that colour on all
On the various Forms of Arvicanthis pumilio. 629
the margins of both wings; a black spot at the end of each
cell; no other markings.
Expanse of wings ,§, inch.
Kashmir; one example.
Family Geometride.
Microloxia vestigiata, nov.
3. Palpi above orange-red, whitish beneath ; antenne
orange, shafts and base white ; frons orange-red, with a pale
band ; top of head and abdomen ochreous white ;_ thorax and
wings of a uniform pale grass-green ; costal line of fore
wings pale ochreous white; cilia white, no markings above or
below : body below white; legs tinged with ochreous.
Expanse of wings 1,4, inch.
Kashmir ; three examples.
LXIX.— Notes on the various Forms of Arvicanthis pumilio,
Sparrm. By R. C. WRouGHtTon.
Tue Natural History Museum possesses a very fine series of
specimens of this common South-African mouse, amounting
to over 130 individuals from more than thirty localities.
The majority of these and the best prepared have been
obtained by Mr. C. H. B. Grant in connexion with the
exploration of the mammal fauna of South Africa now being
carried on by Mr. C. D. Rudd, by whom they have been
presented to the Museum. Those from Deelfontein were
obtained by the same collector working for Col. Sloggett,
and other smaller series are due to Mr. F. C. Selous, Mr. J.
ffolliott Darling, Sir H. H. Johnston, Mr. R. B. Woosnam,
Dr. W. J. Ansorge, Mr. F. J. Jackson, and others.
A detailed examination of this collection has satisfied me
that the general pattern of the coloration in this species (or
group of species) is, without exception, absolutely constant
in all the forms, which are spread over Africa south of the
Equator.
The coloration, on the other hand, is almost as variable as
the pattern is constant. From one or two localities there are
series which are fairly evenly coloured, but in the majority
of cases there are almost as many variations of colour as
there are individuals in a series.
I have found, however, that the size of the bulle furnishes
630 Mr. R. C. Wroughton on the
a reliable character on which to base a differentiation of
forms, which agrees very fairly with their geographical
distribution, and have thus been able to distinguish four
groups with bulle whose antero-posterior lengths are
respectively 6°5, 6, 5:5, and 5 mm.
Group I. covers the country, roughly speaking, south of
25° 8. lat. and west of 25° E. long.
Group II. runs along the coast from the Cape to Zululand,
but a form of this group crops up in Angola. The area
directly north of Group I. is unrepresented in the collection
except by a single specimen from Matopo, but this specimen
falls also into Group II.
Group. III. occupies the whole of the country north of
25° 8. lat. and east of 25° E. long, except in two small
isolated patches.
Group IV., so far, is represented by specimens from two
isolated areas, viz. :—
1. Basutoland, and a small area adjoining, along the crest
of the Drakensberg.
2. The Mlangi Plateau, Nyasaland.
The specimens from both these localities are labelled
* 6000’ Alt.,” and possibly even this altitude is a compara-
tively lower level surrounded by still greater heights; it
certainly is so in the case of the Basutoland specimens. In
any case, however, this group is evidently a high-level form.
Each of these groups contains, as will be seen, more than
one form, and I have found it very difficult to decide as to
the systematic status which should be given to these forms.
I think that each of these four groups should probably rank
as a species ; but if pumilio is once broken up consistency
would require that other species besides these should be formed
(e. g. pumilio and vittatus, both taken at Cape Town, &c.).
I have come to the conclusion that, in view of the absolute
identity of pattern, the variability of coloration, and the
difficulty of deciding the inter-relationship of the different
forms, the simpler and safer way is to call them all subspecies
of the original species pumilio.
The following is a key to the various forms I have been
able to differentiate :—
A, Antero-posterior length of bullz 6°5 mm.
a. Hind foot 27 mm. or more......... wee. bechuane, Thos.
6. Hind foot less than 27 mm.
a, Length of tooth-row 5 mm. —
a’. Hind foot 25 mm. or more.......... cinereus, Thos.& Schw.
6. Hind foot at most 25 mm.:......... mertdionalis, subsp. n.
b', Length of tooth-row 466 mm. ...,.... grique, subsp. n.
various Forms of Arvicanthis pumilio, 631
B. Antero-posterior length of bullae 6 mm.
a. Length of tooth-row 4°8 mm.
a‘, Basilar length of skull 24 mm......... pumilio, Sparrm.
6‘. Basilar length of skull 21 mm. ........ (?) vittatus, Wagner.
6. Length of tooth-row less than 4°8 mm.
a’, Hind foot 23 mm.; basilar length of
skull 23 mm.; length of tooth-row
ERAHAEB Se tha Peak «sheets? a cede. 050 angole, subsp. n.
b', Hind foot 22 mm.; basilar length of
skull 21°5 mm.; length of tooth-row
PENTA (oie aol We te aco Slave 21> a's aile 4 2a intermedius, subsp. un.
ce’. Hind foot 21 mm.; basilar length of
skull 20 mm.; length of tooth-row
Ee PEBINIE Mo tsira) es or rey Ai ayata olay ale # o/s Seek oe (Matopo specimen.)
C. Antero-posterior length of bullae 5-5 mm.
a. Hind foot 22 mm.; basilar length of skull
24 mm.; length of tooth-row 45 mm. ., Chake, subsp. n.
6. Hind foot 20 mm.; basilar length of skull
23°5 mm.; length of tooth-row 45mm... diminutus, Thos.
e. Hind foot 20 mm.; basilar length of skull
22 mm.; length of tooth-row 4.4mm. ., dilectus, de Wint.
D. Antero-posterior length of bulle 5 mm.
a. Brighter coloured. Basutoland.......... Moshesh, subsp. un.
6. Sombre-coloured. Nyasa’ ........0050. nyas@, subsp. u.
Arvicanthis pumilio bechuane.
Isomys pumilio bechuane, Thomas, P. Z. S. 1892, p. 551.
57.1.28.10. ‘ Bechuanaland.” (Type.)
81.3.8.8. 2. Great Namaqualand.
The type specimen is a very old one, and the skull is not
available, but the enormous hind foot is alone sufficient to
differentiate this form from any other.
Arvicanthis pumilio cinereus.
Arvicanthis pumilio cinereus, Thos. & Schw. P. Z.S. 1903, vol. ii. p. 336.
4,2.3.68-74. Klipfontein, Namaqualand. Alt, 3104’.
Rudd Collection.
98.9.3.5-7. Garies, Namaqualand. (Noskulls.) Dr. R.
Broom.
In the original description the length of the hind foot is
given, from the collector’s label, as 28 mm., but this measure-
ment is not, I think, reliable ; measured on the specimen, the
majority of the series show a hind foot of 26 mm. or slightly
over. Judging from the other dimensions recorded by the
collector, the tail in this form is slightly donger than the head
and body.
The following are the normal dimensions of an adult
individual :—
Head and body 120 mm. ; tail 125; hind foot 26.
632 Mr. R. C. Wroughton on the
Skull: basilar length 24; length of tooth-row 5; ant.-
post. length of bulle 6:5.
The following are measurements taken on a few selected
specimens :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
AOS 60. Gs N.Old ® ewer ok 26 24:5 5 65
AO At Ase MOGs satecel> ae 26 24 5 65
ASS 10. ii) A RODE ie Geer 26 23 5 G5
OSAON DAD: His aeons cee 27 ty y P
Arvicanthis pumilio meridionalis, subsp. n.
95.9.3.11-12. Rondebosch, Cape Town. F. C. Selous.
3. 7.2.32, 84. Tokai Retreat, Cape Town. Alt. 600’.
Rudd Exploration.
2.9.1.63. Deelfontein. Col. Sloggett.
The normal dimensions of this form are :—
Head and body 125 mm.; tail 120; hind foot 25.
Skull: basilar length 24; length of tooth-row 5; bulla 6°5.
This form differs but little in dimensions from einereus
except that the hind foot is slightly but constantly shorter.
Whereas cinereus, however, is one of the few forms which
are constant in their coloration, in which there is no trace of
rufous or fulvous, meridionalis, on the other hand, has no
two individuals alike, though all are markedly tinged with
fulvous.
The following are dimensions taken from selected specimens
of meridionalis :—
Basilar
Hind length of Length of
foot. skull, tooth-row. Bulle.
95 OS eliaic. MOld wie uen 248 5 65
95.9.8.12. g. Adult.. 25 23-5 5 65
(Type) 3.7.2.84. 9. V.old.. 25 24 5 65
2.9,1.63. g. Adult .. 24 23 5 65
Arvicanthis pumilio grique, subsp. n.
4,10.1.61, 4.4.8. 15-21. Kuruman, Bechuanaland.
Alt. 4000’.
* These age classes throughout this paper are :— Young: basilar suture
not closed. Y. adult: all teeth unworn. Adult: posterior molar worn,
Old all teeth slightly worn. V. old: teeth worn flat,
various Forms of Arvicanthis pumilio. 633
4.1.6.4-7. Abraham’s Kraal, Modder River. Alt.
4500’.
297.1.13.1. Kimberley, O.R.C.
This form is exceptionally constant in its coloration, but
the modifying tinge is rufous rather than the ordinary fulvous.
The normal dimensions may be taken to be as follows :—
Head and body 115 mm. ; tail 110; hind foot 24.
Skull: basilar length 22; length of tooth-row 4°6;
bullee 6°5.
The following are actual measurements of some selected
specimens :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
det, G50 D.:Oldvises © 24 22 46 65
(Type) 4.4.8.15. 3. Viold.. 24 22 46 65
4,4,8.20. 9. Y.adult, 23 20:5 46 65
Gein tig, Old...) 24 23 46 65
The Kimberley specimen has an abnormal length of skull,
but as by its other measurements and geographically it
belongs here, I have ranked it as grique.
Arvicanthis pumilio.
Mus pumilio, Sparrman, Vet.-Akad. Handl. 1784, p. 339.
Arvicola pumilio, Desmarest, Mamm. 1822, p. 285.
Mus pumilio major, Brants, Muiz, 1827, p. 103.
Mus lineatus, Cuy. Mamm. 1829, pl. clxi.
Mus Donovani, Donovan, Nat. Repos. 1834, vol. iil. pl. Lxxiii.
Mus septemvittatus, Schinz, Synops. Mamm., ii. 1845, p. 155.
Mus striatus, Sundeval, Gify. Vet.-Ak. For. Stockh. 1846, p. 88.
The Mus pumilio major of Brants is said to come from
the country interior to Cape Town and it is stated that
it does not occur in Cape Town itself, while the habitat of
Mus Donovant is given as “ the same part of Africa” as that
from which Sparrman’s Mus pumilio came. There is
nothing in either description by which any particular form
can be recognized, and both descriptions give a length of
head and body of 52 inches (say 145 mm.). No specimen in
the Museum collection reaches even 130 mm. Under these
circumstances there is nothing for it but to rank both names
as synonyms of “ pumilio.” As Sparrman erected pumilio
on a quite immature specimen, I am compelled to select as
the representative of typical pumilio the specimen which
nearest coincides with it in habitat. Sparrman states that
634 Mr. R. C. Wroughton on the
he obtained his specimens “in the Sitsicamma Forest hard
by Slangen River,’ and his map shows that the “ Sitsi-
camma” district extended along the shore of Algoa Bay as
far as Cape Padrone, just short of which “Slang River”
entered the sea. Inthe Museum collection are some specimens
received from the Grahamstown Museum and_ labelled
“ Uitenhage ” and “Fish River”; these, therefore, being
practically topotypes, may be accepted as typical pumuilio.
This being so, I recognize the following specimens in the
collection as being typical pumilo :—
97.11.5. 26-30. Uitenhage, C.C.
5.5.7.67-75. Knysna, C.C. Alt. 1400’.
3.6.2.14, 15. Port St. John, Pondoland.
4.12.8.81. Unmfolosi, Zululand. Alt. 230!. ;
3.7.2. 23, 25, 26, 28. Newlands, Cape Town. Alt. 700’.
3.7.2. 29-31. Tokai Retreat, Cape Town. Alt. 50-500".
The normal dimensions of A. pumilio may be taken to be
as follows :—
Head and body 120 mm.; tail 116; hind foot 24.
Skull: basilar length 24 ; length of tooth-row 4°8 ; bulle 6.
The series from Knysna are quite constant in colour
amongst themselves and show almost no tinge of either
fulvous or rufous ; they are probably a forest form, and it is
noticeable that they come from a markedly higher elevation
than the others. ‘The remainder show the usual individual
variation in the amount of fulvous colouring. he following
are measurements of some selected specimens :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
O7, Mii. 6.980 .4 Visoles se 24 ? 4:8 6
5.5.7.70. G. Yoadult..., 24 23 48 6
BTS Hae Veold ae: 24 24-3 4:8 6
S)Gh2c1b.W OLN Violdl: aoe. 24 24 48 6
A519) 8 SlciOM POld yey! set 24 24-3 4:8 6
51712. 998 4G a Adult hi. 24 24 48 6
BET ose lee ait MOldie, eer 24 23:5 48 6
Unfortunately the Grahamstown Museum skulls are much
broken and in no case is the “ basilar length of skull ” avail-
able ; but the one whose dimensions are given above has a
‘“‘oreatest length” of 30 mm., which is about the same as
that of all the others in the table.
various Forms of Arvicanthis pumilio. 635
The following specimens—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
Site Ween, Meadult; a... 5.0',5. 24 20°5 4:8 6
Onlg oe SA adult: cere sees 24 20°5 4:8 6
Oe eran NOUNS srr. sclera: «ea 24 19 4:8 6
Diieovods AGUIE',, Jc pe cing sae 24 21°6 48 6
—among those taken at Cape Town seem to point to a distinct
form having a far shorter head than pumilio. Unfortunately
all the specimens are comparatively young. I have given
the antero-posterior length of bulla in these specimens as 6,
but it is, 1 believe, slightly less, though greater than 5:5 ;
this measurement is a difficult one to take uniformly, and I
have not dared to record differences of less than 4 mm.
Should the receipt of more specimens, especially old ones,
show later that this is a good form, the name of vittatus,
Wagner (Arch. Naturg. vill. 1842, p. 11), is, perhaps, avail-
able for it, the name having been based on specimens from
the Cape ‘* Promontory ”’; till then I fear vdttatus must rank
as a synonym of pumilio.
Arvicanthis pumilio intermedius, subsp. n.
ZeOa ll. 62565, oe
3.1.4. 52-55. Deelfontein.
SG eco pars) on
This is a form which combines some characters from each
of its neighbours, grigue on the north and pumilio on the
east. From the former it differs by its smaller hind foot and
bulle and from the latter by its smaller hind foot and tooth-
row. It is most variable in its coloration, no two specimens
being exactly alike, and, judging from collector’s measure-
ments, the tail is slightly longer than the head and body.
The normal dimensions may be taken as :—
Head and body 105 mm.; tail 108; hind foot 22.
Skull: basilar length 21:5; length of tooth-row 4:6;
bullee 6.
The measured dimensions of some selected specimens are
as follows :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
2.9.1.69. 9. Old.... 22 ? 4-6 6
2.9.1.70. 9. Y.adult. 22 20° 4-6 6
2.9.1.71. 9. Adult.. 22 21 4-6 6
(Type) 3.1.4.53. 3. V.old.. 22 215 46 6
i7.reb, 2. Olde ae 22 215 4-6 6
636 Mr. R. C. Wroughton on the
Arvicanthis pumilio angola, subsp. n.
5.5.9.41-46. Caconda, Angola. Alt. 4700'.
This form, though belonging to the group with medium
bullz, is completely separated from the other members of the
group noticed above. It is rather smaller all round than
pumilio and has a fairly constant though dull coloration,
resembling that of dilectus. The tail is apparently markedly
shorter than the head and body, much more so than in any
other form.
The normal dimensions may be put as :—
Head and body 118 mm.; tail 95; hind foot 23.
Skull: basilar length 23 ; tooth-row 4°7; bull 6.
The skulls are unfortunately much broken and the actual
basilar length cannot be measured in any one of them; but
the greatest length and zygomatic breadth are about 28 and
13°5 mm. against 30 and 14 in pumilio. I have therefore
taken 23 mm. as the probable “basilar length of skull.”
The following measurements are from selected specimens :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
5.5.9.48. 2. Adult .. 23 2 4-7 6
(Type) 5.5.9. 45 Old a 28 ? 4-7 6
There is a single specimen from Matopo with dimensions
as follows :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
AO ASSiis Dey pAGult ie sks s 21 19°5 45 6
and the label gives—head and body 89 mm. ; tail 96.
I do not venture to give a specific name to this form
represented by only a single specimen (and that a compara-
tively young individual), but have thought the existence of
this specimen worthy of record as seeming to indicate that
the country west of dilectus and north of Group II. will be
found, when examples from it are available, to be peopled
by a form with medium-sized bulla, linking up this specimen
with angole.
Arvicanthis pumilio Chaka, subsp. n.
5.1.68-74. Sibudeni, Zululand. Alt. 3500’.
12:
4,
4.12.5.27-29. Estcourt, Natal. Alt. 4500’.
various Forms of Arvicanthis pumilio. 637
4,6.6.14,15. Notinsila, Pondoland.
4.9.1.62, 65-67. Wakkerstroom, Transvaal. Alt. 6000’.
4.9.1.61. Zuurbron, Transvaal. Alt. 4600’.
This form has slightly larger teeth than the other forms
with small bulle. It appears to range between the
Drakensberg and the coast at somewhat high elevations,
meeting the form of Basutoland and the Eastern Transvaal
at Wakkerstroom. The Pondoland and Natal specimens are
dull-coloured as compared with the more northern individuals,
but there is much variation in colour.
The normal dimensions may be taken as :—
Head and body 115 mm.; tail 105; hind foot 22.
Skull: basilar length 23 ; tooth-row 4°6 ; bulle 5:5.
The following are some measurements of selected speci-
mens :—
Basilar
Hind lengthof Length of
foot. skull. tooth-row. Bulle.
4.5.1.65. 2. Y.adult.. 22 22 46 55
AES C. uivald, 340422 24 46 55
(Type) 4.5.1.72. d. Old...... 22 23 46 55
4.66.14. g.. V.old. .. 22 235 46 55
4,9.1.62. g. V.old .. 22 24 46 55
4.12,5.27. g. Adult.. 22 22°5 4-6 55
I have named this form after Chaka, the famous Zulu
chief.
Arvicanthis pumilio dilectus,
Arvicanthis pumilio dilectus, de Wint. P. Z. S. 1896, p. 803.
95.11.3.25-27. Mazoe, Mashonaland. Alt. 4000’.
1115, 1117, 1118, 1122, 1123, 1135, 1140, 1150. Zout-
pansberg, Transvaal. Alt. 4400’. Rudd Exploration.
98.4.4.22. Krugersdorp, Transvaal. Alt. 4900’.
The normal dimensions of this form may be taken as :—
Head and body 105 mm.; tail 95; hind foot 20.
Skull: basilar length 22; tooth-row 4°4; bulle 55.
Mr. de Winton based his subsp. dzlectus on the specimens
from Mazoe and spoke of them as dull-coloured ; the specimens
from the N.W. Transvaal are brighter and curiously resemble
A, p. grique in coloration, though they are darker than that
form.
The following are some measurements from selected
specimens :—
638 Mr. Rh. C. Wroughton on the
Basilar
Hind length of Length of
foot. skull, tooth-row. Bulle.
(Type) 95.11.8.25. g. Old... 20 22 44 55
95.11.3.26. 3. V.old. 20 22:3 44 Bb
95.11. 3.27. g. Y.adult. 20 21 4-4 55
IIT! SV Told oy 20 225 4-4 Bb
1135;.3, Adult. 40, 20 21:5 4-4 55
T110 Or Old ae eeee 20 21:5 4-4 55
98, 4.4.22. g. V.old. 20 225 44 55
Arvicanthis pumilio diminutus.
Isomys pumilio diminutus, Thos. P. Z. S. 1892, p. 551.
93. 2.3.39, 40. Mianzini, Masailand.
99. 8.4.85-88. Ravine Station, B.E.A.
The normal dimensions are :—
Head and body 94 mm. ; tail 62; hind foot 20.
Skull: basilar length 23°5; tooth-row 4°5; bulle 5:5.
The coloration, as shown by the half-dozen specimens, is
very constant, showing a bright golden tinge throughout the
dorsal ground-colour and bright, almost white, interspaces
between the dorsal stripes.
Though not markedly larger than délectus in any one
dimension, it is distinctly though slightly larger all round.
The following are some actual measurements :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
93. 2.3. 39. ? Oldia yaar ete stoke 20 23 4:5 55
QO NB A.150: Gye Older tettoletels 20 23°5 45 od
99,8. 4,865.9. 7Y.adulf ~... 20 20 45 55
Arvicanthis pumilio Moshesh, sp. n.
4.12.5.48-51. Maseru, Basutoland. Alt. 5000’.
4.9.1.59, 60. Zuurbron, Transvaal. Alt. 4600’.
4.9.1 ib G3) 68. Wakkerstroom, Transvaal. Alt. 6000’.
98.3.23.7. Potchefstruom, Transvaal.
97. 820.1. " Krugersdorp, Transvaal. Alt. 4900’.
4.5.1.69, 74. Sibudeni, Zululand. Alt. 3500’.
This is a form with very small bullz, which seems to have
its habitat in the country along the crest of the Drakensberg.
At Krugersdorp it appears to meet dilectus, the form of the
N.W. Transvaal &c., and at Wakkerstroom and Sibudeni
the below Berg form (Chaka). Whether it is or is not
found immediately northwards, however, there is unfortu-
nately no evidence to show; a very similar form has been
various Forms of Arvicanthis pumilio. 639
found on the high plateau south of Lake Nyasa (see below).
It is fairly constant in coloration, showing but a slight fulvous
tinge over the ground-colour of the back; the dorsal stripes
are black and the interspaces differ in no way from the
general dorsal coloration.
The normal measurements may be taken as :—
Head and body 110 mm.; tail 90; hind foot 21.
Skull: basilar length 22; tooth-row 4°5 ; bull 5.
The following are some actual measurements :-—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
4.9.1.59. 9. Y.adult; 21 20°5 4:5 5
Ayr bat. TAD te Vesold ss” Ot 22 4:5 5)
(Type) 4.12.5.48. g. Adult.. 21 21°5 4:5 5
A000. , 9 4 Oldies ol 21:5 4:5 5
OBia.zaule Go. sVsold, . 21 21:5 45 5
97.8. 5.7 Gre NGroldin | ek 22°5 4:5 5
I have named this species after Moshesh, the founder of
the present ruling dynasty of Basutoland.
Arvicanthis pumilio nyase, subsp. n.
92.8.1.41-44. Mlanje Plateau, Nyasa. Alt.6000’. Sir
H. H. Johnston.
So far as the measurements go, on which I have chiefly
relied, I can find no difference between this form and Moshesh.
However, I regard the 600 or 700 miles which separate
their habitats as a strong argument for separating the forms.
nyasce is more sombre-coloured, ¢. e. less fulvous tinged, has
a stouter broader skull, and a slight difference in the shape of
the posterior molar, which is pentagonal or subcircular in
horizontal section, while in Moshesh this tooth is laterally
compressed in an oblique direction across the jaw, sloping
backwards and slightly outwards. The skulls available are
much broken, but I quote the following measurements :—
Basilar
Hind length of Length of
foot. skull. tooth-row. Bulle.
ie Older, ee 21 ? 4:5 P
PS tOld isa a8: 21 22 4°5 5
640 Mr. G. A. Boulenger on
LXX.—On the Habitat of Rana Blanfordu.
By G. A. BouLencer, F.R.S.
In the ‘Catalogue of Tailless Batrachians in the British
Museum,’ published in 1882, I described and figured a new
frog under the name of Rana Blanfordii, from two specimens
presented by the late Dr. Blanford in 1880 along with a
collection of reptiles and batrachians, partly from Darjeeling,
partly from Muscat, partly from other localities. The
specimens were in a bottle with a Bufo Andersonw, Blgr.,
a toad which has since been rediscovered at Muscat; and
Dr. Blanford himself informed me that all three specimens
probably came from Muscat, although he was not abso-
lutely certain of the locality. A query was therefore
inserted after the indication of the habitat, which follows
the description. I am now convinced that the two frogs
came from Darjeeling, having received a third specimen of
the same species from Capt. F. Wall, C.M.Z.S., who
obtained it in the Himalayas, at Mussoorie, at an altitude
of 7000 to 8000 feet. As I pointed out a short time ago in
these ‘Annals’ (xv. 1905, p. 378), 2. Blanfordii is closely
allied to R. Pleskec, Gthr., from Tibet, but it has longer hind
limbs and the toes are dilated into small disks at the extre-
mity. R. vicina, Stol. (1872), of which a description and
figure have been given by Mr. W. L. Sclater in 1892 (P. Z.8.
p. 842, pl. xxiv. fig. 1), is still more closely related to
R. Blanfordit, and I should have felt inclined to unite the
two except for the statement that the toes of A. vicina are
‘‘ webbed to the extreme tips.”
LX XI.—On a small Collection of Fishes from the Kasai River
(Congo). By G. A. BOULENGER, F.R.S.
Tn concluding my Address to the Zoological Section of the
British Association at Cape Town in August last, I men-
tioned the fishes of the southern tributaries of the Congo
River as among the most important desiderata in our know-
ledge of the freshwater fish-fauna of Africa. I was therefore
delighted to hear, on my return home, that Dr. J. L. Todd
and the regretted Dr. J. E. Dutton had collected some fishes
for the Congo State Museum, among which was a series
from Lusambo, on the Kasai River. These specimens are
Fishes from the Kasai River (Congo). 641
now in my hands for study, and I here offer a list of the
species represented in the Kasai collection :—
1. Polypterus ornatipinnis, Blgr.
This species was described in 1902 from a single specimen
from Monsembe, presented to the British Museum by the
Rev. J. H. Weeks. The specimen from Lusambo is a little
larger than the type, measuring 46 centimetres. 10 dorsal
spines; 63 scales along the body, 24 between the occiput and
the first dorsal spine, 42 round the middle of the body.
2. Mormyrops attenuatus, Bler.
ro)
A single specimen, measuring 185 mm. D. 41; A. 64.
Lat. 1. 90. Depth of body 8 times in total length, length of
head 54 times.
3. Gnathonemus elephas, Bler.
4. Hydrocyon lineatus, Bikr.
5. Alestes Liebrechtsii, Bler.
6. Alestes Fuchsii, Blgr.
7. Distichodus Antoni, Schilth.
8. Distichodus fusciolatus, Bler.
9. Distichodus sexfasciatus, Ble.
10. Clarias lazera, C. & V.
11. Chrysichthys Duttont, sp. n.
Depth of body 44 times in total length, length of head 3?
times. Head once and a half as long as broad, with granular
rugosities above; eye one fourth length of head, nearly
equalling interorbital width; snout broadly rounded, ex-
tending a little beyond lower jaw; vomero-pterygoid teeth
forming a long but very narrow band ; nasal barbel 3 diameter
of eye; maxillary barbel a little longer than head, extending
nearly to extremity of pectoral spine. Dorsal I 6; spine
strong, serrated behind; first soft ray as long as head ;
adipose fin as long as deep, its base a little more than its
distance from the rayed dorsal. Anal IV 7. Pectoral spine
3 length of head, very strongly serrated behind. Caudal fin
deeply notched, with rounded lobes. Caudal peduncle nearly
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 42
642 Mr. G. A. Boulenger on
once and a half as long as deep. Brown above, white
beneath.
Total length 130 mm. A single specimen.
Most nearly related to C. punctatus, Blgr., but maxillary
barbels and soft rays of dorsal fin much longer.
12. Auchenoglanis occidentalis, C. & V.
13. Synodontis angelicus, Schilth.
14. Synodontis Greshoffi, Schilth.
15. Paratilapia Toddi, sp. n.
Depth of body equal to length of head, 22 times in total
length. Eye 2 length of snout, 4 times in length of head,
exceeding a little interorbital width ; maxillary extending to
below anterior border of eye; teeth small, in 3 or 4 rows; 6
series of scales on the cheek. Gill-rakers short, the larger
T-shaped, 13 on lower part of anterior arch. Dorsal XVI 10;
spines subequal in length, 4 length of head; longest soft rays
not quite half length of head. Anal III 7. Pectoral ob-
scurely pointed, 2 length of head. Ventral reaching origin
of anal. Caudal rounded. Caudal peduncle a little deeper
than long. Scales ctenoid, 30 ae lateral lines o. Dark
olive-brown above, whitish beneath; soft dorsal and caudal
fins barred with blackish; a few large ocellar dark spots on
the anal fin.
Total length 150 mm. A single specimen.
Most nearly related to P. macrocephala, Blgr., but scales
larger, especially on the back.
16. Pelmatochromis lateralis, Bler.
LXXII.—On a Collection of Fishes from Lake Bangwelo.
By G. A. BoULENGER, F.R.S.
Ir is a very great satisfaction to me to be able to fill up a
second desideratum in our knowledge of the ichthyology of
the Congo basin by offering a list of fishes from Lake
Bangwelo. No sooner had I reported on the Kasai fishes
enumerated in the preceding paper than I received from
Mr. Frank H. Melland a series of fishes collected by him,
with the cooperation of the Rev. Father Foulon, in Lake
Fishes from Lake Banguwelo. 643
Bangwelo, the fish-fauna of which was totally unknown.
The native names have been supplied by Father Foulon.
The specimens have been presented to the British Museum
by Mr. Melland.
1. Gnathonemus macrolepidotus, Peters.
Native name: JZutesa.
2. Gnathonemus Monteir’, Gthr.
Native name: Lukesu.
3. LHydrocyon lineatus, Blkr.
Native name: Mehene or Manda. Specimens have been
caught averaging up to 51 lbs. 11 oz., but the fish is said to
run a great deal beyond this weight.
4, Alestes macrophthalmus, Gthr.
Native name: Lumanse.
5. Alestes imberi, Peters.
Native name: Lusaku.
6. Alestes grandisquamis, Bler.
Native name: Mutula.
7. Distichodus maculatus, Blgr.
Native name: Chikamalunski.
8. Labeo altivelis, Peters.
Native name: Pumbu.
9. Barbus banqwelensis, sp. n.
Depth of body equal to length of head, 4 times in total
length. Snout rounded, slightly projecting beyond the lower
jaw, equal to diameter of eye and to interorbital width, 32
times in length of head; barbels two pairs, very small,
barely 4 diameter of eye. Dorsal III 7, equally distant
between end of snout and root of caudal, originating above
first ray of ventral ; last simple ray flexible, not enlarged, a
little shorter than head. Anal III 5, longest ray not quite 3
length of head. Pectoral 3 length of head. Caudal peduncle
twice as long as deep. Scales 28 a, 2 between lateral line
42*
644 Mr. G. A. Boulenger on
and root of ventral, 12 round caudal peduncle. Silvery,
brownish on the ae posterior border of scales blackish; a
black streak along the side of head, passing through the eye ;
fins greyish.
Total length 90 mm.
A single specimen. Native name: Busonga.
Allied to B. congicus, Blgr. Distinguished by the shorter
barbels and the more numerous scales in the lateral line.
10. Clarias Mellandi, sp. n.
Depth of body 54 times in total length, length of head 34
times. Head once ‘and 2 as long as broad, feebly granulate
above ; occipital process ang ular ; frontal fontanelle 1 Jength
of head; occipital fontanelle very small ; diameter of eye 3s
times in length of snout, 54 times in interorbital ici
Vomerine teeth granular, forming a large semielliptic patch,
squarely tr uncate behind, its longitudinal diameter 3 times
that of the band of premaxillary teeth. Nasal barbel 2
length of head, maxillary barbel 2 length of head. Gill-
rakers long, about 35 on first arch. Dorsal 65, its distance
from the oeehna process + length of head, its distance from
the caudal fin 4+ length of head. Anal 55, very narrowly
separated from ane caudal. Pectoral spine "serrated on the
outer border, # the length of the fin. Ventrals slightly
nearer end ot snout than caudal. Olive-brown above,
marbled with darker, whitish beneath ; a dark streak on each
side of the throat.
Total length 345 mm.
A single specimen. Native name: Muta or Mulenga.
Said to reach a weight of over 6 Ibs.
A very distinct species, unique in the form of the patch of
vomerine teeth.
11. Clarias Fouloni, sp. n.
Depth of body 6} times in total length, length of head 5
times. Head once ama 4 as long as “broad, smooth above ;
occipital process angular ; frontal foufanelie 4 length a
head, twice as long as the occipital fontanelle ;- diameter of
eye 3 times in length of snout, 5 times in interorbital ae
Vomerine teeth conical, forming a crescentic band, which,
the middle, is a little broader than the band of premasillary
teeth. Nasal barbel 3 length of head, maxillary barbel +
Jength of head. Gill- ieee about 20 on first arch. Dorsal
83, its distance from the occipital process 4 length of head ;
sin 68; both dorsal and anal embracing the root of She
Fishes from Lake Bangwelo. 645
eaudal. Pectoral spine not serrated on the outer border, 2
the length of the fin. Ventrals twice as distant from the
extremity of the caudal as from the end of the snout.
Blackish brown, belly paler brown.
Total length 200 mm.
A single specimen. Native name: Mulenfi.
Closely allied to C. Theodore, M. Weber, and C. ampleai-
cauda, Bley. Distinguished from both by the longer head.
12. Schilbe mystus, L.
Native name: Lupata.
13. Chrysichthys mabust, sp. n.
Depth of body 43 times in total length, length of head 3}
times. Head once and a half as lone as broad, with fechle
granular rugosities above ; diameter of eye 6 times in length
of head, twice and 3 in interorbital width; snout broadly
rounded, extending a little beyond lower jaw; vomero-
pterygoid teeth forming a long but narrow band, narrowly
interrupted in the middle; nasal barbel nearly as long as eye ;
maxillary barbel half length of head, not reaching base of
pectoral spine. Dorsal 163 spine strong, serrated behind ;
second soft ray longest, a little shorter than head ; adipose
fin a little longer than deep, its base ? its distance from the
rayed dorsal. Anal IV 7. Pectoral spine half length of
head, very strongly serrated behind. Caudal fin deeply
notched, with pointed lobes. Caudal peduncle once and 2
long as deep. Olive-brown above, white beneath.
Total length 250 mm.
A single specimen. Native name: Alubuszt.
‘his species is characterized by its short maxillary barbels,
combined with long dorsal rays and pointed caudal lobes.
14. Auchenoglanis occidentalis, C. & V.
Native name: Bowa.
15. Synodontis nigromaculatus, sp. n.
Depth of body equal to length of head, 4 times in total
length. Head a little longer than broad; snout rounded ;
diameter of eye 6 times in length of head, 25 times in length
of snout and in interorbital width ; maxillary barbel without
distinct fringe, reaching middle of pectoral spine; mandibular
barbels with long branches. About 30 slender mandibular
teeth. Humeral process sharply pointed, without keel.
646 Mr. G. A. Boulenger on
Dorsal I 7; spine as long as head, smooth in front, serrated
behind. Adipose fin 4 times as long as deep, twice and $ as
long as its distance from the rayed dorsal. Anal IV 7.
Pectoral spine as long as head, strongly serrated on both
sides. Caudal deeply forked. Skin without villosities.
Greyish above, white beneath, covered all over with round
black spots, some of which, on the posterior part of the body,
are confluent.
Total length 210 mm.
A single specimen. Native name: Chirima.
Closely allied to S. zambesensis, Peters. Distinguished by
the strong serrature on the anterior border of the pectoral
spine.
16. Anabas multispinis, Peters.
Native name: Lukomo.
17. Paratilapia robusta, Gthr.
Native names: Nsuku, Polwe, Mubantasa.
18. Paratilapia Mellandi, sp. n.
Depth of body 2} times in total length, length of head 3
times. Hye 2 length of snout, 4 times in length of head,
equal to interorbital width; maxillary not extending to below
anterior border of eye; teeth small, in 3 or 4 rows; 4 series
of scales on the cheek. Gill-rakers short, 13 on lower part
of anterior arch. Dorsal XV 13, spines equal in length from
the 7th, which measures 2 length of head ; longest soft ray 2
length of head. Anal JIT 10. Pectoral acutely pointed,
slightly shorter than head. Ventral reaching origin of anal.
Caudal rounded. Caudal peduncle as long as deep. Scales
eycloid, 37 2; lateral lines =. Brownish; dorsal fin edged
with yellowish ; soft dorsal and candal with numerous round
blackish spots; anal with round white spots; ventral
blackish.
Total length 150 mm.
A single specimen. Native name: Jd/dilia.
Closely related to P. moeruensis, Blgr., but soft dorsal rays
more numerous and pectoral fin longer.
19, Pelmatochromis lateralis, Blgr.
Native name: Nsangula.
20. Tilapia natalensis, M. Weber.
Native names: MMutuba (young) and Kamba (full-grown).
Fishes from Lake Bangwclo. 647
21. Tilapia melanopleura, A. Dum. (lata, Gthr.,
Rendalli, Blgr.).
Native name: Lupondo.
22. Tilapia Sparrmant, A. Smith.
Native name: Chikundu.
23. Tilapia Foulont, sp. n.
Depth of body 21 times in total length, length of head 34
times. Hye 2 length of snout, 4 times in length of head, 2
interorbital width ; mouth small, maxillary not “extending to
below anterior border of eye ; teeth small, in 4 rows; ‘two
series of scales on the cheek. Glill-rakers short, 10 on lower
part of anterior arch. Dorsal XV ob spines increasing in
length to the last, which measures 2 length of head; longest
soft ray # length. of head. Anal Til 10. Pectoral acutely
pointed, as long as head. Ventral reaching origin of anal.
Caudal rounded, subtruncate. Caudal , peduncle a little
deeper than long. Scales cycloid, 29 2; lateral lines 3.
Brownish ; a black opercular spot; soft dorsal with rather
indistinct round blackish spots.
Total length 140 mm.
A single specimen. Native name: Mituku.
Distinguished from 7. Sparrmant by the smaller mouth
and the longer pectoral fin.
24. Mastacembelus signatus, sp. n.
Depth of body 104 times in total length, length of head 7
times. Vent equally distant from end of snout and base of
caudal. Length of head 22 times in its distance from vent,
4 times its distance from first dorsal spine ; snout 3 times as
long as eye, ending in a trifid appendage which is twice as
long as eye; mouth extending to below nostril ; a strong
preeorbital spine; three strong preopercular spines. Vertical
fins united with the rounded caudal. Dorsal XXIX 85.
Anal III 85. Pectoral about 4 length of head. About 20
scales between origin of soft dorsal and lateral line. Brownish
above, yellow beneath ; a series of 14 large X-shaped
blackish markings on each side of the body.
Total length 275 mm.
A sing!c specimen.
Closely allied to AL. congicus, Blgr.
648 Mr. K. Andersen on the Species and
LXXIIT.—A List of the Species and Subspecies of the Genus
Rhinolophus, with some Notes on their Geographical
Distribution. By KNuD ANDERSEN. .
THE present paper gives a brief summary of the systematic,
phylogenetic, and zoogeographical conclusions at which I
have arrived by a study of the bats of the genus Rhinolophus
preserved in the British Museum and the United States
National Museum. For the details that have served as a
basis for the conclusions I must refer to my former papers on
this subject *.
A Systematic List of the Species and Subspecies.
A. The Rhinolophus simplex Group.
Rh. simplex, K. And.—Lombok fF.
. Rh. megaphylius, J. EH. Gray.—Eastern Australia; Louisiade Archi-
pelago.
2 a. Ph. ~megaphyllus f. typica.—Eastern Australia (Queensland,
N. 8. Wales).
2b, Rh. megaphyllus monachus, K. And.—Louisiade Archipelago
(St. Aignan’s Isl.).
? Rh. keyensis, Ptrs.—“ Key Islands.
. Rh. truncatus, Ptvs—Batchian.
Rh. nanus, K. And.—Goram Island.
. Rh. celebensis, K, And.—Celebes (Makassar, Menado).
oe
”
Or co
* Andersen and Matschie, “ Ueber einige geographische Formen der
Untergattung Furyalus,” SB. Ges. naturf. Fr. Berlin, 1904,
no. 5, pp. 71-83.
Andersen, ‘‘ Five new Rhinolophi from Africa,” Ann. & Mag. Nat.
Hist. (7) xiv., Nov. 1904, pp. 3878-888.
Td., “ On von Heuglin’s, Riippell’s, and Sundevall’s Types of African
Rhinolophi,” t. c., Dec. 1904, pp. 451-458.
Id., “‘ Further Descriptions of new Rhinolophi from Africa,” op. ett.
(7) xv., Jan. 1905, pp. 70-76.
Id., “On the Bats of the Rhinolophus philippinensis Group, with
Descriptions of Five new Species,” op. cit. (7) xvi., Aug. 1905,
p. 243-257.
Id., “ On the Bats of the RAinolophus arcuatus Group, with Descrip-
tions of Five new Forms,” ¢. ¢., Sept. 1905, pp. 281-288.
Id., “ On the Bats of the Rhinolophus macrotis Group, with Descrip-
tions of Two new Forms,” ¢. ¢., Sept. 1905, pp. 289-292.
Id., “On some Bats of the Genus Rhinolophus, with Remarks on
their Mutual Affinities, and Descriptions of Twenty-six new
Forms,” Proc. Zool. Soc. 1905, ii. (Oct.) pp. 75-145, pls. iii, iv.
+ When not otherwise stated, the record of the geographical distri-
bution of the species and subspecies is based exclusively on examples
examined by myself (a few localities quoted from literature are printed
between inverted commas).
Subspecies of the Genus Rhinolophus. 649
. Rh, borneensis, Ptrs—N. Borneo; S. Natunas; Karimata Archi-
pelago.
6a. Rh. borneensis £. typiea.—N. Borneo; Labuan; Banguey.
6b. Rh. borneensis spadix*, Miller—S. Natunas (Sirhassen) ; Kari-
mata Archipelago (Karimata, Pulo Sarutu).
. Lh. virgo, K. And.-—Luzon.
Rh, malayanus, Bonhote.—Malay Peninsula (Jalor); ? Siam (Laos
Mts.).
Rh. nereis, K. And.—Anambas Archipelago (Pulo Siantan).
Rh. simulator, K. And.—Mashonaland (Mazoe).
. Rh, Denti, Thos.—Bechuanaland (Kuruman) ; Wakkerstroom (Zuur-
bron).
. Rh. stheno, K. And.—Malay Peninsula (Selangor, Penang).
. Rh, Rouxi, Temm.—From 8. China, through the Himalayas, to the
Indian Peuinsula and Ceylon.
13a. Rh. Rouxi sinicus, K. And.—Lower Yangtse Valley.
136. Rh. Rouxi f. typica.—Himalayas (Darjeeling, Nepal, Masuri) ;
S. India (Nilghiri, Kanara); Ceylon.
. Eh. capensis, Lchtst.—S. Cape Colony,
. Rh. Thomasi, K&. And.—Burmah (Karin Hills).
. Rh. affinis, Horsf.—From the N.W, Himalayas to 8. China; through
Indo-China, the Malay Peninsula, and N. Natunas, to Sumatra,
Java, and Lombok.
16a. Rh. affinis himalayanus, K. And.—Himalayas (Masuri, Nepal,
Darjeeling) ; S. China (Nanking).
16. Rh. affinis tener, K. And.—Pegu.
16c. Rh. affinis macrurus, K. And.—Burmah (Karin Hills).
16d. Rh. affinis superans, K, And.—Lower Siam ; Malay Peninsula ;
Sumatra.
16e. Lh. affinis nesites, K. And.—N. Natunas (Bunguran Isl.).
16 f. Eh. affinis f. typica.—Java.
16g. Eh. affinis princeps, K. And.—Lombok.
. Rh, andamanensis +, Dobson.—S. Andamans.
. Rh. clivosus, Cretzschm.—* Arabia (Mohila)”; Berbera.
. Rh. Darlingi, &%. And.—Mashonaland (Mazoe); Angola.
. Rh. acrotis, Heugl—From Erythrea to Lower Egypt.
20a, Rh. acrotis f. typiea.—Erythrea.
20 6. Rh. acrotis Andersoni t, Thos.—Eastern Egyptian Desert.
20¢. Rh. acrotis brachygnathus, K. And.—Lower Egypt.
21. Rh, ferrum-equinum, Schreb.—F rom 8. China and Japan, through
the Himalayas, the Mediterranean Subregion (exclusive of
Egypt) and Central Europe, to 8. England.
21a. Kh. ferrum-equnum nippon, Temm.—S, China (Shanghai) ;
Pt. Hamilton ; Japan.
21b. Rh. ferrum-equinum tragatus, Hodgs.—Darjeeling ; Nepal.
2le. Rh. ferrum-equinum regulus, K. And.—Almora; Masuri.
21d. Rh. ferrum-equnum proximus, K, And.—Gilgit.
2le. Rh. ferrum-equinum f. typica—From Transcaspia and the
Euphrates Valley, through Southern and Central Europe,
exclusive of the Spanish Peninsula.
21 f. Rh. ferrum-equinum obscurus, Cabrera.—Spanish Peninsula
(with Balearics) ; Algeria,
* Doubtfully distinct from the typical form of Rh. borneensis.
+ Perhaps a local form of RA. affinis.
{ Doubtfully distinct from the typical 2h. acrotis.
650 Mr. K. Andersen on the Species and
22. Rh. augur, K. And.—Orange River tract ; Natal ; Lower Zambesi.
22a. Rh. augur f. typica—Orange River tract (Transvaal, Orange
River Colony, Bechuanaland, Namaqualand). i
22 b. Rh. augur zuluensis, K. And.—Zululand ; Natal; Pondoland ;
K. Williamstown.
92 ¢.- Rh. ae zambesiensis, K. And.—Lower Zambesi tract (Mazoe,
\yasa).
23, Rh. Deckent, Ptrs.—Ukambani tract; Zanzibar coast.
B. The Rhinolophus lepidus Group.
24. Rh. lepidus, Blyth —S. India (Wynaad) ; Ganges Valley.
25. Rh. monticola, K. And.—Masuri.
26, Rh. refulyens, K. And.—Malay Peninsula (Perak, Selangor),
27, Rh. acuminatus, Ptrs.—Jaya; Lombok.
27 a. Rh. acuminatus f. typica.—Java.
27 b. Rh. acuminatus audax, K. And.—Lombok.
28. Rh. sumatranus, K. And.—Sumatra.
29. Rh. calypso, K. And.—Engano.
30. Rh. minor, Horsf—Java; ? Siam; ? Darjeeling.
31. Rh. minutus, Miller, nec Montagu.u—Anambas Archipelago.
32. Rh. cornutus, Temm.—Japan.
32a. Rh. cornutus pumilus, K. And.—Loo-choo Islands (Okinawa) ;
? S. China (Foochow).
32b. Rh. cornutus f. typica—Japan proper.
33. Rh. gracilis, K. And.—Malabar coast.
34. Rh. subbadius, Blyth.—‘‘ Nepal”; “ Assam (Garo Hills).”
35. Rh. monoceros, K. And.—Formosa.
36. Rh. empusa, K. And.—Nyasa.
37. Rh. Andreinii*®, Senna.—“ Erythrea.”
38. Rh. Blasii, Ptrs.—Mediterranean Subregion.
39. Rh. Landeri, Martin.—Fernando Po; Gaboon.
40, Rh. lobatus, Ptrs.—Zambesi tract (Shupanga, Shire, Nyasa) ; Ukam-
bani tract.
41. Rh. Dobsoni, Thos.—Kordofan.
42, Rh. euryale, J. H. Blasius—Mediterranean Subregion.
42a. Rh. euryale judaicus, K. And. & Mtsch.—Euphrates Valley ;
Palestine; Lower Egypt.
42b. Rh. euryale Mehelyi, Mtsch.—Dobrudsha ; N. Bulgaria.
42c, Rh. euryale f. typica—Dalmatia; Po Valley; Liguria.
42d. Rh. euryale tuscanus, K. And. & Mtsch.—Tuscany (Pisa) ;
Latium (Roma).
42 e. Rh. euryale carpetanus, Cabrera.—Guadiana Valley.
42 f. Rh. euryale Cabrere, K. And. & Mtsch.—Tajo Valley (Madrid,
Cintra).
429. Rh. euryale atlanticus, K. And. & Mtsch.—France ; Galizia.
42h. Rh. euryale barbarus, K. And. & Mtsch.—Morocco to Tunisia
(coast form).
42%. Rh. euryale meridionalis, K. And. & Mtsch.—A]geria (probably
mountain form).
* Stated to differ from Rh. Blasii in the shape of the sella (Angelo
Senna, “Contributo alla conoscenza dei Chirotteri Eritrei,” Archivio
Zoologico, ii. pt. 3, pp. 256-260, pl. xvi. fig. 1, pl. xviii. figs. 7-16 ;
Sept. 50, 1905).
43,
44,
45,
4G.
47.
48.
49,
Subspecies of the Genus Rhinolophus. G51
©. The Rhinolophus midas Group.
Rh. midas, K. And.—Persian Gulf (Jask).
Rh. hipposiderus, Bechst.—From Gilgit to Ireland, from the Baltic
to Sennar.
44a. Rh. hipposiderus minimus, Heug].—Erythrea and Sennar; the
Mediterranean Subregion.
446, Rh. hipposiderus f. typica—F rom the extreme N.W. Himalaya
(Gilgit), through N.W. Persia (Urmi) and Armenia (Van),
over the whole of Central Europe.
44¢, Rh. hipposiderus minutus, Montagu.—England; Wales; Ireland.
? Rh. phasma, Cabrera.— Central Spain (Tajo Valley).”
D. The Rhinolophus philippinensis Group.
Rh. philippinensis, Waterh.—Philippines.
Rh. achilles, Thos.—Key Islands.
Rh, mitratus, Blyth.— N. India (Chaibassa).”
Rh. Maclaudi, Pous.—‘‘ Conakry Island” (off Senegambia).
Rh. sedulus, K. And.—N. Borneo ; Malay Peninsula (Pahang).
50. Rh. lanosus, K. And.—N.W. Fokien.
51. Rh. trifoliatus, Temm.—Java; Sumatra; N. Borneo; Malay Penin-
52.
53.
54.
55.
56.
57.
58.
59.
60.
61.
62.
63.
64.
65.
66.
67.
68.
sula; Lower Siam; Tenasserim.
Rh. solitarius, K. And.—Banka.
Rh. Beddomei, K. And.—S. India (Wynaad).
Rh. luctus, Temm.—Java; N. Borneo; Malay Peninsula,
Rh. geminus, K. And.—Java.
Lh. perniger, Hodgs.—Himalayas (Sikkim, Nepal, Masuri).
KE. The Rhinolophus macrotis Group.
Rh. macrotis, Hodgs.—Masuri ; Nepal.
Rh. hirsutus, K. And.—Philippines.
Rh. ethiops, Ptrs.— Angola.
Rh. Hildebrandti, Ptrs.—Zambesi tract (Mazoe, Nyasa) ; Ukambani
tract (Taita, Machakos, Kenya).
Rh, eloquens, K. And.—Uganda.
Rh. fumigatus, Riipp.— British East Africa ; Abyssinia ; Somaliland ;
Erythrea.
62a. Rh. fumigatus exsul, K. And.—British East Africa.
62 6. Rh. fumigatus f. typica—Abyssinia (Shoa, Adowa) ; Somali-
land (Pozzi Dass, Jifa Medir); Erythrea.
Rh, Pearsoni, Horsf—Himalayas, eastwards to Fokien.
63a. Rh. Pearsoni f. typica—Himalayas (Masuri, Darjeeling) ;
“Yunan”; “ Szetchuen.”
63 b. Rh. Pearsoni chinensis, K. And.—Fokien.
F. The Rhinolophus arcuatus Group,
Eh. arcuatus, Ptrs.—Philippines.
64a. RA. arcuatus f. typica— Luzon.
64 b. Rh. arcuatus exiguus, K. And.—Zamboanga; Guimaras,
Rh, subrufus, K. And.—Philippines.
Rh. inops, K. And,—Mindanao,
Rh. Creaghi, Thos.—N. Borneo.
Rh. celophyllus, Ptrs—Malay Peninsula (Kedah) ; “ Lower Burmah
(Moulmein) ” ; Upper Burmah (Tsigine).
652 Mr. K. Andersen on the Species and
69. Rh. ewyotis, Temm.—Batchian ; Amboina; Key Islands.
69a. Rh. euryotis timidus, K. And.—Batchian.
69 b. Rh. euryotis f. typica.—Amboina.
69 ¢, Lh. euryotis prestans, K. And.—Key Islands.
G. Incert@ sedis.
(70) Rh. angolensis *, Seabra.—“ Angola (Hanha).”
(71) Rh. alcyonet, Temm.—* Gold Coast.”
A Geographical Review of the Species, with some Notes on
their probable Interrelations.
Bats, as being possessed of a greater facility of locomotion
than other mammals, are commonly supposed to be deceptive
guides for the zoogeographer. It may well be that this
is in part, perhaps chiefly, due to the fact that very often
distinct, and sometimes widely distinct, species have been
covered by one technical nameft. If we draw the lines
of separation between the species (and their local modifica-
tions) somewhat more closely in accordance with the lines
drawn by nature, we shall, no doubt, find that in most
instances bats are as good and reliable zoogeographical
guides as other small but non-flymg mammals. Such
at least is the case with the bats of the genus here under
consideration. There is a great similarity between the
Rhinolophus fauna of N. Borneo and that of the Malay
Peninsula (see below), but hardly greater than between the
mammalian faunas of these countries in general. In the
Philippines, on the other hand, we find a remarkable
assemblage of very primitive Lhinolophi, most of them
o . .
essentially different from those of the opposite continent,
* The “lobo central do appendice nasal” is described by Seabra as
“bifurcado como no Rh. Blasii” (Jorn. Sci. Math. Phys. Nat. Lisboa,
(2) v. Dec. 1898, p. 250). If this means that the connecting-process is
high and pointed and the sella deltoid (triangular, with pointed summit),
Rh. angolensis is certainly a distinct species and of much interest as a
West-African representative of the empusa type, which as yet, within
the Ethiopian Region, is known from Nyasaland and Erythrea only.
+ There is not in the original description of the only known specimen
of this bat (Le'den Museum) one’single word of any value for identifying
the species or determining its affinities. It is as thoroughly unknown as
if it had never been recorded.
{ E.g.: Rh. “ ferrum-equinum,” made up of Rh. ferrum-equinum, augur,
acrotis, and fumigatus, and therefore distributed over the whole of the
Ethiopian and the whole temperate part of the Palearctic Region;
Rh. “affinis” as a collective name for Rh. borneensis, stheno, Rouxi, and
affinis; Rh.“ minor” for Rh. lepidus, monticola, refulgens, minor, cornutus,
gracilis, and subbadius; Ke.
Subspecies of the Genus Rhinolophus. 653
only one species (A7rsutus), itself a primitive form, being a
genuine Himalayan type, though as a species quite distinct ;
this, again, is perfectly in accordance with the general
character of the Philippine fauna. The immigration of
Rhinolophine types from south into the Philippines, and the
radiation from these islands southwards into the Austro-Indo-
Malayan Archipelago, have by no means been greater than
of other mammals—rodents, f. 1. A very narrow tract of
water can form an apparently insurmountable barrier for the
spreading of a Rhinolophus (Ith. ferrum-equinum in England,
not in Ireland), as it has formed for the voles. The Rh/no-
lophus fauna of Lower Egypt* is markedly different from
that of Palestine; not even the direct land-connexion has
caused a more extensive interchange of species than in the
case of non-flying mammals. All this—and a series of
similar instances could be adduced—tends to show that for
the spreading of the RAinolophi their power of flight has been
a factor of very little importance ; their present distribution,
like that of non-flying mammals, has been determined by the
history of the type to which the species belongs and the
geological history of the continent or island in question.
AUSTRALIA :—Lh, megaphyllus typicus.—The only Austra-
lian species is most closely related to Rh. simplex, from
Lombok.
LovuistapE ARCHIPELAGO :—LZh. megaphyllus monachus.—
The Louisiade form seems to be a not quite perfectly differ-
entiated offshoot of the Australian species.
New GuIneA.—As yet no species is known from New
Guinea, although the genus is represented both east (Louis-
iade Archipelago), south (Australia, Key Islands), and west
(Moluccas) of the island.
Kry Isuanps :—“ Lh. keyensis” ; Ith. achilles; Rh. eury-
otis prestans.— Rh, achilles is a peculiar modification of the
philippinensis type. Lh. euryotis prestans has its nearest,
scarcely more than subspecifically distinct, allies in Amboina
and Batchian. ‘ 2h. keyensis,’ a still very imperfectly
known form, is probably closely related to RA. stmplex and
megaphyllus. Vhe Rhinolophus fauna of the Key Islands,
therefore, points partly north-westwards, to the Moluccas and
the Philippines, partly westwards.
* Of the four Palestine species (2A. ferrum-equinum, Blasi, euryale
J i ? ’ X
Jjudaicus, hipposiderus minimus), one only (euryale judaicus), so far as I
know, has spread from the Asiatic side of the Mediterranean to Lower
Egypt. The only other species recorded from Lower Egypt (2A. acrotis
oy Sie eee 5, 5 eye
is unknown in Syria and Palestine.
654 Mr. K. Andersen on the Species and
GorAM :—Lh. nanus.—A representative of the common
Austro-Malayan simplex type.
AMBOINA:—Lh. euryotis typicus.—This form has its
closest, only subspecifically distinct, allies to the north
(Batchian) and to the south-east (Key Islands) ; but the
euryotis type belongs to a group of the genus (the arcuatus
group) which now has its most primitive representatives in
the Philippines.
BaTcHIAN :—Rh. truncatus; Rh. euryotis timidus.—Rh.
truncatus is a well-marked species of the widely distributed
simplex type. Eh. euryotis points, as already stated, in the
last instance northwards, to the Philippines.
Lombok :—Lh. simplex, Rh. affinis princeps; Rh. acumina-
tus audax.—h. simplex seems to be the most primitive
member of the section which I have proposed to call the
Lh. simplex group ; it has very close relatives in (probably)
the: whole of the Austro-Malayan and Indo-Malayan sub-
regions. Lh. affinis princeps is the extreme south-eastern
outpost of a species now distributed from the Himalayas
through Indo-China, Sumatra, and Java; the Lombok form
seems to be more closely related to the Malacca-Sumatra race
(Rh. a. superans) than to the Java race (Lh. a. typicus).
Rh. acuminatus audaz is a local form of a Java species.
-THE Auvstro-MALAYAN SUBREGION.—Out of 69 species
known, only 8 are found in this subregion (9, if Rh. keyensis
is regarded as a species). Of these 8 species, two (Rh. affinis
princeps, Rh. acuminatus audax) are south-eastern outposts
of Indo-Malayan or Indo-Chinese species. Of the remaining
6 no less than 4 (Lh. simplex, megaphyllus, truncatus, nanus)
are representatives of the simplex type, which also numbers
several very primitive species in the Indo-Malayan Archi-
pelago. The last two species (2th. achilles and euryotis) can
be traced back to the Philippines.
CEeLEBES :—Z?h. celebensis, a representative of the simplewx
type, in certain cranial characters rather intermediate between
the Austro-Malayan and the genuine Indo-Malayan species
of the s¢mplex group.
PHILIPPINES :—Lth. virgo; Rh.philippinensis; Rh.arcuatus,
Rh. subrufus, Rh. inops; Rh. hirsutus—The Philippine
Rhinolophus tauna 1s remarkable for its richness in primitive,
even extremely primitive, types, and the total absence of
highly differentiated forms. 2h. virgo is closely related to
Rh. borneensis, both of them species on a low level of develop-
ment. Lh. philippinensis is the most primitive representative
known of the philippinensis group; so far as concerns the
Subspecies of the Genus Rhinolophus. 655
dentition, it has apparently remained ona slightly lower level
than any other species of the genus. 2h. arcuatus and sub-
rufus are the most primitive members of the arcwatus group ;
Eh. inops a representative of the same group, chiefly charac-
terized by its peculiarly modified sella. RA. hirsutus is a
very primitive species of the macrotis group, closely related
to the Himalayan 2h. macrotis—-Rh. philippinensis and
Rh. arcuatus cannot be brought into close genetic connexion
with any other known bat; in the absence of paleontological
evidence to the contrary, we may therefore regard them as
autochthonous Philippine types—7. e. as the least modified
survivors of types which have originated in the Philippines,
or, more likely, in a tract of land of which the Philippines
are the relicts. We can still trace their radiation out from
that centre: the phzlippinensis type has spread both south-
wards, to the Key Islands (Lh. achilles), and westwards,
through India (22. mitratus) as far as the Ethiopian Region
(Rh. Maclaudi), while a third offshoot has given rise to
the slightly more aberrant Indo-Malayan sedulus-trifoliatus
branch ; the arcuatus type has spread southwards and become
differentiated into the comparatively rather highly developed
Austro-Malayan 2h. euryotis. The presence of the simplex
type (th. virgo) in the Philippines is evidence of an immi-
gration ¢néo the islands from the south; the close relationship
between the Himalayan Rh. macrotis and the Philippine
Lh. hirsutus points to a former connexion with the continent.
N. Bornzo :—RaA, borneensis (typicus); Rh. sedulus, Rh.
trifoliatus, Rh. luctus ; Rh. Creaghi.—Rh. borneensis is a bat
of the s¢mplea type, slightly more advanced than RA, cele-
bensis. Lith. sedulus, trifoliatus, and luctus are members of
the philippinensis group; the former species in its cranial
characters rather primitive, in its essential external characters
close to trifoliatus ; Lh. trifoliatus and luctus are more highly
developed species of the group. Rh. Creaghi is a peculiar
modification of the arcuatus type.—The fauna points partly
(Rh. borneensis) eastwards, to Celebes and the Austro-
Malayan islands, partly and most decidedly north-eastwards,
to the Philippines (all the other species). It is very closely
connected with the Lhinolophus fauna of the Malay Peninsula,
no less than three species (sedulus, trifoliatus, luctus) being
common to both countries.
S. NATUNAS AND KARIMATA ARCHIPELAGO :—RA. borne-
ensis spadia, extremely closely related to (or identical with)
the Bornean form of the species.
Maay PeninsuuA, Lower Sram, SoutuH TENASSERIM :—
th. malayanus, Rh, stheno, Kh, affinis superans ; Rh. reful-
656 Mr. K. Andersen on the Species and
gens; Rh. sedulus, Rh. trifoliatus, Rh. luctus; Rh. celo-
phyllus.— The first three species belong to the simplex
group: Lh. malayanus is very closely related to Rh. borneen-
sis; Ith, stheno a more thorough modification of the borneensis
type; Rh. affints superans is but a local race of a Himalayan
species. Lh, refulgens, a bat of the lepidus group, has its
closest relative in the Himalayas (RA. monticola). Rh.
sedulus, trifoliatus, and luctus, all of the philippinensis group,
are common to Borneo and the Malay Peninsula. 2h. eelo.
phyllus is a highly peculiar species of the arcuwatus group,
probably rather closely related to the Bornean 2h. Creaghi.—
Of the eight species here under consideration, six (22h. malay-
anus, stheno, sedulus, trifoliatus, luctus, calophyllus) bear
evidence of the very close faunistic connexion between Borneo
and the Malay Peninsula; the remaining two (affinis, reful-
gens) are but) slightly modified immigrants from the north.
Sourn Anpamans :—“ Rh. andamanensis. ”_Although as
yet very imperfectly known, this bat is undoubtedly closely
related to Lh. affinis superans from the Malay Peninsula.
SUMATRA :—RA, affinis superans; Lh. sumatranus; Rh.
trifoliatus.—Rh. affinis superans and Lh. trifoliatus are
common to Sumatra and the Malay Peninsula. 2h. suma-
tranus belongs to a small section of the lepzdus group, closely
connected with Rh. refulgens from the Malay Peninsula.
ENGANO :—Lh. calypso.—It is worth noticing that the
only Lhinolophus as yet known from Engano is closely related
to, but specifically distinct from, 2A. sumatranus.
‘BANKA:—Rh. solitarius, a local representative of the
philippinensis type, closely allied to, but specifically distinet
from, Lh, trifoliatus from the Malay Peninsula and Sumatra.
JAVA:—Tih. affints typicus; Rh. minor, Rh. acuminatus
typicus; Rh. trifoliatus, Rh. luctus, kh. geminus.—The Java
form of RA. affints seems to be closer related to the Hima-
layan race than to Rh. a. superans from Sumatra and the
Malay Peninsula. 2h. minor is either identically the same
species as found in Siam and Darjeeling or a very closely
allied form. A. acuminatus has no closer relative than
Rh. sumatranus. Lh. trifoliatus and luctus are common to
Java, Borneo, and Malacca. Rh. geminus, a bat of the
luctus type, is very closely related to the Himalayan Rh. per-
niger.—As a summary: of six species, three (2A. afjnis,
minor, geminus) point to a closer faunistic affinity between
Java and the Indo-Chinese and Himalayan tracts than
between Java and the geographically nearer Sumatra,
Malacca, and Borneo; the remaining three are common
Indo- Malay: an types.
Subspecies of the Genus Rhinolophus. 657
N. Natunas:—RA. affinis nesites, an apparently well-
differentiated form, most closely related to Rh. a. superans
from Malacca.
ANAMBAS ISLANDS :—Rh. nerets; Rh. “ minutus.”’—The
two species point to a connexion both with Borneo and with
the continent, the former being an offshoot of the borneensis
type, the latter of the minor type.
Tue Inpo-MALAYAN Suprecion.—Of 69 species known,
26 (38 per cent.) are found in this subregion, and no less
than 24* are, as species, apparently autochthonous; of the
remaining two, one (Rh. afinis) is certainly, the other (Rh.
minor) probably, Indo-Chinese.—To form a clearer idea of
the affinities and probable origin of this fauna it is best,
however, to consider the primary groups of species repre-
sented within the subregion ; we then arrive at the conclusion
that all the species of the simplex group (seven in number ;
see footnote) probably, in the very last instance, are
descendants of Austro-Malayan types; that the five species
of the lepidus group and the only species of the macrotis
group can be ultimately traced back to some part of what we
now call Indo-China; whereas the eleven species of the
philippinensis and arcuatus groups may very likely have
originated from purely autochthonous types. If this be so,
we have as a total result 15 species which (at least as
“‘ types ”) can be traced back to places outside the subregion
as against 1] apparently purely autochthonous.
TENASSERIM TRACT (including Karennee) :—Rh. Thomas?,
Rh. affinis macrurus; Rh. celophyllus—Rh. Thomast is a
very peculiar modification of the Chinese and Himalayan
Rouwxi type ; Rh. affints macrurus a local representative of a
Himalayan species. Rh. cwelophyllus has come from the south
(Malay Peninsula).
PEGU TRACT :—Rh. affinis tener, very closely related to the
Himalayan form of Rh. affinis.
ASSAM TRACT :—Rah. subbadius, also known from Nepal.
SourH CHINA AND Formosa:—Rh. Roux sinicus, Rh.
affinis himalayanus, Rh. ferrum-equinum nippon ; Rh. cornutus
pumilus, Rh. monoceros; Rh, lanosus; Rh. Pearsoni chinensis.
* Seven species of the s*mplex group: Rh. eelebensis, borneensis, virgo,
malayanus, nereis, stheno, andamanensis. Five of the lepidus group :
Rh, refulgens, acuminatus, sumatranus, calypso, “ minutus.” Six of the
philippinensis group: Rh, philippinensis, sedulus, trifoliatus, solitarius,
luctus, geminus. Five of the arcuatus group: Rh. arcuatus, subrufus,
tnops, Creaghi, celophyllus. One of the macrotis group: Rh. hirsutus.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 43
658 Mr. K. Andersen on the Species and
—Three of these species (Rh. affinis, ferrum-equinum, Pear-
soni) are most probably of Himalayan or, at least, Indo-
Chinese origin; Rh. cornutus has no closer relative than the
Himalayan Rh. minor ; Rh. monoceros, known from Formosa
only, is a modification of the Himalayan Rh. subbadius.
Thus, five out of the seven species point westwards ; with
the two remaining, Rh. Rouat and Rh. lanosus, the case is
different—the former species, though also found throughout
the Himalayas, is most closely related to RA. borneensis, the
latter to the Bornean Rh. sedulus.
Souty Korea, Loo-cuoo ISLANDS, AND JAPAN PROPER :—
Rh. ferrum-equinum nippon; Rh. cornutus.—Both species are
undoubtedly immigrants from China.
HimaLayas:—Rh, Rouaxi typicus, Rh. affin’s himalayanus,
Rh. ferrum-cquinum tragatus and regulus; Rh. monticola,
Rh. minor (2), Rh. subbadius; Rh. perniger; Rh. macrotis,
Rh. Pearsoni typicus.—Four of these species (RA. affinis,
ferrum-equinum, macrotis, Pearsoni) may very likely be of
Himalayan origin; the two former have spread far beyond
this tract. Rh. monticola, minor (?), and subbadius may also,
as species, be of Himalayan origin, but they have slightly
more primitive allies in the Indian Peninsula. Rh. Rowxtis,
as already stated, probably an immigrant from east, derived
from the dorneensis type. Rh. perniger is most closely related
to Rh. geminus from Java.
THE HIMALAYAN AND INDO-CHINESE SUBREGION (in-
cluding Korea and Japan).—Of 69 species known, 14 (7. e.
20 per cent.) occur in this subregion, but one of them (RA,
celophyllus) is probably a direct immigrant from south, The
four forms of the simplex group (Rouwi, Thomasi, affnis,
ferrum-equinum) have, most probably, as species originated
within the area; when traced back to their remotest origin,
they are descendants of a more eastern type. ‘The same is
the case with the representatives of the philippinensis group
(lanosus, perniger). The five species of the leprdus group
(monticola, minor, cornutus, subbadius, monoceros) seem to
have a slightly more primitive relative in the Indian Penin-
sula. Rh. macrotis is the only Indo-Chinese species which I
fail to trace back to any other known type of the genus*;
it may be the very primitive survivor of a genuine (autoch-
thonous) Himalayan type; in any case, its origin evidently
* It is highly probable that the macrotis type originated from an
ancient philippinensis-like bat which had not acquired the peculiar
specialization of the nose-leaves characteristic of all the now-existing
representatives of the phzlippinensis group (see my paper on the Rh, ma-
crotis group, loc. cit. pp. 290-292).
Subspecies of the Genus Rhinolophus. 659
dates back to a period when the distribution of land and
water in this part of the world was essentially different from
what it is nowadays, for we find representatives of the macrotis
type in the now thoroughly isolated Philippine Islands (RA,
hirsutus) and in a vast part of the Hthiopian Region (RA.
ethiops, Hildebrandtt, eloquens, fumigatus). Rh. Pearsoni is
a comparatively highly developed Himalayan and 8. Chinese
modification of the macrotis type.
Gances VALLEY: — Rh. lepidus; Rh. mitratus. — The
former is a very primitive (perhaps the most primitive)
member of the lepidus group; the latter a representative of
the philippinensis group, much more closely related to the
Indo-Austro-Malayan Rh. philippinensis and achilles (and
the Ethiopian Rh. Maclaudi) than to the geographically
nearer Himalayan form of the same group.
Soutn Inp1a :—Rh. Rouaxi typicus; Rh. lepidus; Rh.
Beddomei.—Rh. Rouxt is no doubt an immigrant from the
Himalayas, where identically the same race occurs. RA.
lepidus is also found in the Ganges tract. Rh. Beddomer is
closely allied to Rh. luctus from Borneo and the Malay
Peninsula.
CEYLON :—Rh. Rouaxt typicus, common to Ceylon and
S. India. (A bat of the philippinensis type occurs in Ceylon,
presumably Rh. Beddomei; I have seen a very young indi-
vidual only.)
Marazpar Coast:—Rh. gracilis, a bat of the probably
Himalayan mznor type.
THe INDIAN AND CEYLONESE SuBREGIONS. — Only five
species occur, one of them (Roux?) Indo-Chinese. Rh. gracilis
points northwards ; Rh. mitratus and Beddomet to the Indo-
Malayan countries. One species (RA. lepidus) may represent
a purely autochthonous type.
SoMALILAND, EryTHREA, ABYSSINIA, AND BAHR-EL-
ABIAD TRACT:—Ra. clivosus, Rh. acrotistypicus; Rh. Andreinii,
Rh. Dobsoni; Rh. hipposiderusminimus; Rh. fumigatus typicus.
—The first two species are modifications of the Himalayan
affinis type. Rh. Andreinii (very closely related to 2h. Blasit)
and Rh. Dobsoni (very close to Rh. lobatus) point back
to the Himalayan Rh. subbadius. Lh. hipposiderus has no
nearer known ally than the Persian Rh. midas, and the
particular race (minimus) here under consideration is the
same as now distributed over the Mediterranean countries.
Rh. fumigatus is a very highly developed species of the
Himalayan macrotis type.
4.3*
660 Mr. K. Andersen on the Species and
UGANDA :—Rh. eloquens, a bat of the macrotis type, in
certain characters rather intermediate between Rh. Hilde-
brandti and Rh. fumigatus.
UKAMBANI TRACT AND ZANZIBAR Coast :—Rh. Deckent;
Rh. lobatus; Rh. Hildebrandti, Rh. fumigatus exsul.—Rh.
Deckeni is an Ethiopian representative of the Oriental ferrum-
equinum type. Rh. lobatus belongs to a small group of
Ethiopian species (Landeri-lobatus- Dobson?) which have their
more primitive counterpart in the Himalayan Rh. subbadius.
Rh. Hildebrandti and fumigatus can be traced back ultimately
to a bat like Rh. macrotis.
ZAMBESI TRACT :—Rh, simulator, Rh. Darlingt, Rh. augur
cambesiensis; Rh. lobatus, Rh. empusa; Rh. Hildebrandti.—
Rh. simulator is a bat of the borneensis type; Rh. Darling?
of the Himalayan affinis type; Rh. augur of the Oriental
jferrum-quinum type. Rh. empusa is an Ethiopian repre-
sentative of the Rh. Blasit stage, which, however, again
leads back to the Oriental ménor-subbadius stage. ‘The two
remaining species (lobatus, Hildebrandti) are common to this
and the toregoing tract.
Limpopo TRACT :—Rh. augur typicus ; on the species, see
Zambesi tract, above.
ZULULAND, NATAL, EASTERN CAPE CoLony :—RA. augur
zuluensis.—This small, but zoogeographically rather well-
marked, district is inhabited by a special race of the wide-
spread Kthiopian RA. augur.
S.W. Cape Cotony :—RaA. capensis, an Ethiopian repre-
sentative of the Oriental Rh. Rouaxt type.
ORANGE River TRACT:—RhA. Denti, Rh. augur typicus.—
Rh. Denti, closely related to Rh. s¢mulator from the Gambesi
tract, is a bat of the borneensis type. On the affinities of
Rh. augur, see the Zambesi tract above.
BENGUELA AND LOANDA:—Rh. Darlingi; Rh. angolensis ;
Rh. ethiops.—Rh. ethiops is a highly developed representa-
tive of tle Himalayan macrotis type. Rh. Darlingi is
common to this district and the Zambesi tract. Rh. ango-
lensis is unknown to me (but see footnote above on p. 652).
Lower Guinea :—Rh. Landeri, closely related to the
Eastern Ethiopian RA. lobatus and Dobsoni, all of them bats
of the Oriental subbadius type.
Gop Coast :—Rh. alcyone; unknown to me.
GAMBIA TRACT :—h, Maclaudi, a bat of the Indo- Malayan
philippinensis type.
Tue ETHIOPIAN REGION:—19 out of 69 known * species
* Leaving the imperfectly known 2h. angolensis and the practically
quite unknown RA. aleyone out of consideration.
Subspecies of the Genus Rhinolophus. 661
have as yet been recorded from the Ethiopian Region. To
sum up the probable affinities of these species: Rh. Denti
and simulator represent the borneensis type; Rh. capensis the
Rousxi type; Rh. clivosus, Darlingi, and acrotis the affints
type; Rh. augur and Deckent the ferrum-equinum type ; Rh.
empusa and Andreini? one branch, Rh. Landeri, lobatus, and
Dobsoni another branch, of the m/nor-subbadius type; Rh.
hipposiderus the midas type; Rh. Maclaudi the philippinensis
type; Rh. ethiops, Hildebrandti, eloquens, and fumigatus the
macrotis type.—Thus, the distribution of the primary groups of
the genus within the Ethiopian Region is, broadly speaking, as
follows :—the simplex group (8 species) from the Cape Colony
to Lower Egypt (beyond the limits of the Region), and on
the western side of the Continent as far north as Angola; the
macrotis group (4 species) from Abyssinia to the Lower
Zambesi, across the Continent to Angola; the lepidus group
(4 species) in a broad tract across the Continent from about
15° N. to 20° S.; the mzdas group (1 species) confined to
the extreme north-eastern corner ; the philippinensis group
(1 species) to the north-western corner (probably of wider
distribution).—It is a matter of some zoogeographical
importance that all the Ethiopian species of the genus Rhino-
lophus, without exception, also have representatives in the
Oriental Region; but still more important is the fact that all
the Ethiopian species have more primitive representatives in
S. Asia or the Indo-Malayan Archipelago. In view of this,
and bearing in mind that in the absence of all paleontological
evidence we have to base our conclusions exclusively on what
we know about the now-existing forms, we are justified in
supposing that all the Ethiopian Rhinolophi are, in the last
instance, derived from Oriental forms. The passage from
the Oriental to the Ethiopian Regions must have been
considerably easier in past times than now,
Eastern EayptiAn Desert:—Rh. acrotis Andersoni.
The species is Ethiopian.
Lower Eqaypt:—Rh. acrotis brachygnathus ; Rh. euryale
judaicus.— Rh. acrotis is undoubtedly an immigrant from the
Ethiopian Region. Rh. euryale has come from the Asiatic
side of the Mediterranean; examples from Lower Egypt are
indistinguishable from the Palestine-Kuphrates race; the
species does not seem to have spread south of Lower Hgypt.
Tue MEDITERRANEAN SUBREGION (exclusive of Lower
Egypt) :— Rh. clivosus, Rh. ferrum-equinum (proximus,
typicus, and obscurus) ; Rh. midas, Rh. hipposiderus minimus ;
Rh, Blasti, Rh. euryale—Rah., clivosus is known only from
662 Mr. R. Kirkpatrick on
the border districts of the Ethiopian and Palearctic Regions
(Red Sea coasts), Rh. midas from the shore of the Persian
Gulf. These, as well as the four truly “ Mediterranean ””
species, are undoubtedly of Oriental origin. Worth noticing
is the close faunistic connexion between the Spanish Penin-
sula and N.W. Africa (Algeria) : the same race (ohscurus) of
Jerrum-equinum.
CenTRAL Evurope:—Rh. ferrum-equinum typicus; Rh.
hipposiderus typicus.—The Central European Rh. hipposiderus
is slightly different from the Mediterranean form.
Britisu Istanps :—Rh. ferrum-equinum; Rh. hipposiderus
minutus.—Both of the Central European species have reached
the British Islands. Rh. hippostderus, as being the more
hardy of the two species, as having spread over the whole of
England and to several places in Ireland, and as having
become to a certain slight degree different from the conti-
nental form, was probably the earliest comer. The range of
Rh. ferrum-equinum is restricted to the southern part of
England.
THE WHOLE AREA OF THE GEeNuUS.—AI] the now-existing
species can be referred to six “types.” All the types can be
traced back to some part or other of the Oriental Region.
From there they have spread eastwards as far as Hastern
Australia and Japan, south-westwards over the whole of the
Ethiopian Region, westwards to Southern and Central
Europe.
LXXIV.—On the Oscules of Cinachyra.
By R. Kirkpatrick.
[Plate XIV.]
WHILE engaged in the investigation of specimens of
Cinachyra barbata, Sollas, obtained by the ¢ Discovery ’ from
the Antarctic, I was led to examine examples of that species
obtained by the ‘Challenger’ from Kerguelen and described
by Sollas in his Report on the Tetractinellida.
Specimens of this species are spheroidal or ovoidal in
shape and with a root-tuft ; the surface bristles with a pile-
like coat of spicules, which are mostly protrizenes. Arranged
round the sides of the sponge are flask-shaped recesses with
oval or circular orifice and with the margins guarded by a
the Oscules of Cinachyra. 663
fringe of needles rising above the general surface; on the
upper part of the surface are smaller orifices, likewise
surrounded by a spicular fringe.
Sollas (9, pp. xxxv, 27) regarded the flask-shaped recesses
as being either vestibules or cloacas, though he failed to find
any structural differences in them, and for convenience of
description he called them all oscules. The smaller orifices
and pits near the summit were considered to be young
undeveloped vestibules and cloacas of the same kind as those
on the rest of the surface.
The good state of preservation of numerous specimens from
the Antarctic has enabled me to clearly distinguish two kinds
of depressions—vestibular and cloacal; and later I was able
to note the same distinction in the ‘ Challenger’ specimens.
Cinachyra barbata in its adult condition (PI. XIV. fig. 1)
shows three zones :—(1) a basal root-tuft zone; (2) a broad
“ equatorial ” zone of poral vestibules ; and (3) a superior or
“polar” zone of oscules (sensu stricto). In good specimens
these zones are clearly obvious to the naked eye, for the
surface pile of spicules is higher and looser in the poral than
in the oscular zone and slopes downwards ; the pile in the
oscular area is shorter and denser and points vertically
upwards, presenting, in fact, a stubble-like appearance.
The orifices of the poral vestibules are usually wide open,
though even large ones may be closed; but in almost every
instance the oscules are tightly shut, the marginal fringes
being closed over them in the form of conical stacks.
The shape of many of the specimens from Kerguelen is
oval and the poral zone and oscules are arranged obliquely
to the long axis. Often, too, at first sight no trace of oscular
fringes or oscules is apparent ; but by rubbing off the surface
pile at the superior end of the body, so as to leave a tonsure-
like patch, the oscules become visible as minute dark pin-
points on the surface (Pl. XIV. fig. 3).
Prof. Sollas has given such a full account of the poral
vestibules that there is no need to say anything further
about them here. The oscules are few in number in com-
parison with the vestibules ; in one specimen there are seven
of the former arranged in an open spiral on the summit and
over forty of the latter.
Usually the oscules appear as low monticules each with a
shallow crater-like cup, but occasionally they may be flush
with the surface. On making a section so as to cut vertically
through an oscule, usually, owing to the extreme state of
contraction, no passage can be seen, but simply a break in
the continuity of the white cortex, the fleshy tissues of the
664 Mr. R. Kirkpatrick on
oscular channel being pale reddish. On dissection and
magnification, however, asimple canal with deep longitudinal
folds is easily made out; below the cortex the passage leads
into a main exhalant canal coming from the centre and into
large lateral canals (Pl. XIV. fig. 2).
As regards the homologies of the poral vestibules and the
shallow, crater-like, oscular depressions, these both seem to
be ectodermal invaginations, and the vertical intracortical
preoscular tube to be the terminal exhalant canal (or gastral
cavity ?).
Lendenfeld (7, p. 26), who records eight species of Cina-
chyra, gives as the definition of the genus :— Ausstrémungs-
6ffhnungen klein, zu Gruppen vereint am grunde kahler,
schalen-, kelch-, oder sackférmiger Hinsenkungen der
Oberfliiche.” Obviously this definition must be emended,
since in C. barbata, the type of the genus, the oscules are
simple and separate and open each into a single intracortical
channel, which latter receives the main exhalant canals. If
for ‘ Ausstrémungséffnungen” the word ‘ Poren” were
substituted, the definition might stand.
On referring to the descriptions and figures of other species
of Cinachyra it seems to me possible that misinterpretations
concerning the pores and oscules may have been made in
some of them also. In Cinachyra Schulzet, Keller, from the
Ried Sea, Keller (3, p. 387, pl. xix. figs. 41, 42) figures a
sponge with numerous depressions all apparently alike in
general character: fig. 42 depicts an enlarged view of a
section showing the surface of one of these depressions ;
here we see pores with inhalant canals radiating out from
them into the body of the sponge ; at the base of the depres-
sion are slightly larger orifices opening into wider canals.
Keller assumes that the larger orifices are the oscular openings
of exhalant canals. ‘The latter, except that they are a little
wider, look very like the inhalant ones, and, & priori, it seems
unlikely that excrementitious orifices should be situated in
the floor of a deep pit surrounded by inhalant orifices, natural
selection tending to keep the inhalant and exhalant systems
each out of the way of the other. A comparison with
C. barbata would lead one to suggest that the depressions in
C. Schulzed are poral vestibules, and that extremely con-
tracted oscules situated elsewhere (probably near the summit)
have escaped notice.
Cinachyra eurystoma, Keller (3, p. 338, pl. xix. figs. 46,
48), from the Red Sea, looks like a young immature form ;
young specimens of C. barbata have several relatively large
the Oscules of Cinachyra. 665
vestibules, with, perhaps, only one minute oscule. The
figure of a section of a depression (3, fig. 48) shows small
openings leading each by a narrow tube to moniliform canals ;
possibly here also the vestibules are purely poral, and a
careful search might reveal a much contracted oscule. Simi-
larly the circular zone of depressions shown in the figure of
C. trochiformis, Keller (3, p. 340, pl. xix. figs. 44, 45), from
the Red Sea, is possibly vestibular, and an examination of
the conical summit might show the existence of oscules in the
form of minute contracted points.
Cinachyra amboinensis (Kieschnick), from Amboina (4,
p. 556), is globular or egg-shaped and has a zone of poral
depressions and a single osculum at the summit.
Cinachyra hirsuta (Dendy) (2, p. 75), from the Gulf of
Manaar, is spheroidal and firmly attached by a broad base to
a piece of rock. Here there is a zone of depressions and one
large osculum situated to one side of the summit. Dendy
believed that some of the depressions were pore-areas, others
oscular areas; after an examination of the specimen I
have come to the conclusion that all the depressions are
poral vestibules. ‘The variation in shape and depth, some
being shallow and hemispherical, others deep and tubular,
appears to me to result from differences in age and in the
degree of contraction.
Cinachyra Voeltzkowt, Lendenfeld (6, p. 101, pl. ix. fig. 39),
from Zanzibar, appears, at first sight, to present in the
arrangement of its pores and oscules a complete exception to
what is found in other species of Cinachyra. he three speci-
mens are spherical. ‘Two of the examples are only 8 mm. in
diameter and have only one depression each ; a larger one,
2 cm. in diameter, has fifteen depressions. Lendenfeld
observes (6, p. 101) :—“ An der oberfliche finden sich
allenthalben Poren, doch sind in den Weingeistexemplaren
nur jene an den konvexen, pelztragenden Teilen offen, die,
wie es scheint kleineren und zahlreicheren, in den kahlen
Vertiefungen dagegen, geschlossen. Von den ersteren ziehen
offene Kaniile herab, welche die rinde durchsetzen.”
Lendenfeld regards the “ Poren” on the general surface
as inhalant and those in the depressions as exhalant, and
describes one of the depressions themselves as “ glattwandiger
praeoscularraum” (/. c, p. 129). He observes (i. c. p. 103)
that very possibly pores are present on the surface of C. bar-
666 Mr. R. Kirkpatrick on
bata outside the depressions and that they have been overlooked
by Sollas. I have carefully searched for these pores and
have not been able to detect any; indeed, it would be
surprising if pore-canals could penetrate the dense palisade of
cortical oxeas. Since the depressions are smooth-walled and
the pores in their walls are smaller and more numerous than
those on the general surface, I think that probably the
depressions are poral vestibules similar to those of C. barbata,
that some of the ‘‘Poren”? on the surface are oscules,
possibly much contracted, and that this kind will be found
occupying a more or less definite area.
An eighth species of Cinachyra mentioned in Lendenfeld’s
list, viz. C. robusta (Carter), from Mergui (1, p. 79), remains
to be noticed. The Natural History Museum possesses one
half of the type specimen. Judging from its appearance, I
should take it to be a macerated specimen of Tetil/la which
has become much worn down with age and rough usage from
strong currents, so that the whole of the cortex has become
denuded, leaving large open spaces and caverns between the
radiating fibres of the skeleton. Carter (7. ¢. p. 79) writes :—
“‘ The spicules of the interior, which project so abundantly as
to produce a hispid condition of the surface, are so matted
together by the mud in which the sponge has grown on the
subjacent rock that, in taking off this crust, the ‘forks’ and
‘anchors,’ together with the projecting ends of the ‘ body-
spicules,’ all come away with it?’ Sollas’s designation
(9, p. 48), viz. Tetilla robusta, of Carter’s Tethya cranium,
var. robusta, seems to me correct.
Apparently only one of the seven species at present retained
in the genus Cinachyra has a cortex with a dense palisade of
oxeas, though several of the others appear to possess a thick
fibrous cortex.
Very young specimens of Cinachyra are conical and
have only one large poral vestibule, situated inferiorly and at
one side, the one oscule being at or near the summit. This
asymmetrical arrangement calls to mind the sponge Spongo-
cardium Gilchristi, Kirkp. (5, p. 224), from South Africa.
I now think that Spongocardium must be regarded as a
synonym of Fangophilina, O. Schmidt (8, p. 73, pl. x. fig. 3),
in spite of the fact that FW, submersa has a well-developed
root-tuft, and #. Gulchristt appears not to possess this
appendage, though a root-tuft may have been torn off in
dredging.
the Oscules of Cinachyra. 667
F, Gilchristi has two deep depressions, poral vestibular
and cloacal, the former having smooth walls perforated by
minute pores and the latter having orifices of oscules in its
base.
Judging from his description and figure, Schmidt, I believe,
mistook the nature of these depressions, describing the poral
as cloacal and vice versé ; for the smooth-walled deep depres-
sion of £, Gilchristi is undoubtedly a poral vestibule and not
acloaca. Without a knowledge of the development it would
be difficult to say whether the openings in the floor of the
cloacal depression in the latter species are to be regarded as
an assemblage of oscules or as orifices of exhalant canals
opening into a cloaca terminating in one oscule; the first
hypothesis seems the more probable one.
To sum up: the depressions on the surface of the seven
species of Cinachyra are probably in every instance poral
vestibules, the oscules being separate and distinct.
Taterature.
(1) Canter, H. J. “ Report on the.... Sponges from.... Mergui
Archipelago....” Journ. Linn. Soc., Zool. 1887, vol. xxi.
(2) Denny, A. “ Report on a Second Collection of Sponges from the
Gulf of Manaar.” Ann. & Mag. Nat. Hist. 1889, (6) iii.
(3) Ketter, C. ‘Die Spongienfauna des rothen Meeres.—II.”
Zeitschr. f. wiss. Zool. 1891, Bd. lii.
(4) Krescunicx, O. “ Kieselschwiimme von Amboina. Zool.
Forschungsreisen Semon.” Denkschr, Gesellsch. Jena, 1900,
Bd. v., Text.
(5) Kirkpatrick, R. “ Descriptions of South African Sponges —I.”
‘Marine Investigations in South Africa,’ 1902.
(6) LenpDENFELD, R. von. “Spongien von Sansibar.” Abhand.
Senckenberg. Gesellsch. 1899, Bd. xxi.
(7) ‘Das Tierreich.’ Lief. 19. 1903. Porifera Tetraxonia.
(8) Scumipt, O. ‘Die Spongien des Meerbusen yon Mexico.’ 1879.
(g) Sottas, W. J. ‘ Report on the Tetractinellida collected by
H.MLS. ‘Challenger.’ ” Zoology, vol. xxv. 1888.
EXPLANATION OF PLATE XIV,
Fig. 1, Cinachyra barbata, Sollas, from the Antarctic, natural size.
a, zone of poral vestibules; }, zone or area of oscules.
Fig. 2. Vertical section through an oscule, x 5. a, oscule; 8, cortex ;
c, main excurrent canal; d, foraminiferan shell.
Fig. 3. Cinachyra barbata from Kerguelen (‘Challenger’), The surface
pile of spicules has been rubbed off in the oscular region to
show the oscules, which are here contracted to pin-points.
a, oscules,
668 Mr. W. L. Distant on Homoptera.
LXXV.—Rhynchotal Notes —XX XVII.
By W. L. Disranrt.
Fam. Cicadide.
POLYNEURARIA.
Platypleura lineatella, sp. n.
@. Body and legs very pale tawny brown; head with a
very obscure piceous fascia between the eyes ; pronotum with
the central anterior edge of the posterior margin and the
Jateral dilated areas pale ochraceous; mesonotvm with sub-
obsolete traces of ill-defined obconical spots ; abdomen above
thickly palely pilose, with a central segmental longitudinal
series of black spots, the segmental margins and the anal
segment also black; body beneath and legs concolorous ;
tegmina and wings subhyaline, talc-like, the venation greenish
ochraceous, their extreme bases and the basal cell of tegmina
ochraceous.
Long., excl. tegm., 22 mm. ; exp. tegm. 72 mm.
Hab. Sunda Islands ; Salayer (Lveritt).
I am compelled to found this species on a female specimen,
in order that it may find a place in my catalogue. The
lateral margins of the pronotum are broadly angularly
ampliate, and the rostrum extends beyond the middle of the
abdomen ; the head (including eyes) is about as wide as the
mesonotum ; the apical segment of the abdomen beneath has
a small fuscous spot on each side.
Platypleura lindiana, sp. n.
Body and legs dull dark obscure ochraceous ; head with a
basal fascia to front, a transverse line at each anterior angle
of vertex, a linear transverse fascia between eyes, and inner
margins of eyes black ;.pronotum with a transverse fascia
near anterior margin and a central longitudinal fascia black ;
mesonotum with four anterior obconical spots, of which the
two central ones are shortest, a central lanceolate spot, and a
small spot near each anterior angle of the cruciform elevation
black ; abdomen above with the bases of the segments broadly
black ; head beneath with a broad black transverse fascia
between the eyes; tegmina pale subhyaline, nearly basal half
pale tawny, radial area containing two large pale spots, be-
neath which are other two pale spots in the fourth ulnar area ;
the remaining subhyaline area with small subobsolete darker
spots, the transverse veins at the bases of the first, second,
Mr. W. L. Distant on Homoptera. 669
third, fourth, fifth, and seventh apical areas infuscated, and a
double series of small submarginal fuscous spots placed one
on each side of the longitudinal veins to apical areas ; wings
very pale ochraceous, “about apical third hyaline, the outer
margin of the ochraceous area slightly tinged with fuscous ;
opercula short, transverse, their anterior angles well separated;
rostrum passing posterior coxee, its apex piceous; lateral
margins of pronotum ampliate and broadly angulated ; head
(including eyes) about as wide as mesonotum.
Long., excl. tegm., ¢ 163 mm.; exp. tegm. 56 mm.
Hab, B. East Africa; Lindi.
Belonging to the group of species represented by P, Afzeli,
Stal.
Platypleura mira.
Platypleura mira, Dist. Ann. & Mag. Nat. Hist. (7) xiv. p. 333 (1904).
Platypleura laotiana, MS., Paris Mus.
Since describing the male (supra) I have seen the other
sex, also contained in the Paris Museum.
2, Differs from @ inthe tint of the tegmina, the ground-
colour of which is greyish brown and not brownish ochraceous :
the two oblique piceous fascize are also deeper and brighter i in
hue, and the outer one distinct and not fused with the dark
markings on the apical area.
Platypleura Murchisoni, sp. n.
3S. Head, thorax, body beneath, and legs pale tawny ;
abdomen black, the tympana pale tawny ; body somewhat
thickly, finely, greyishly pilose; mesonotum with four
anterior obconical spots, denoted by their black margins, the
two central ones much the shorter, and with two small, rounded,
black spots in front of the er uciform elevation ; tegmina very
pale tawny, obscurely mottled with paler or grey vish suffusions ;
the radial area greyish with two very pale fuscous spots, the
innermost preceded by a similar spot on costal membrane,
the outer halves of the first to third ulnar areas and the
whole of the apical areas greyish, semiopaque, with very
obscure pale tawny macular mottlings, principally on the
longitudinal veins, and a double series of small outer sub-
marginal fuscous spots placed one on each side of the
longitudinal veins to apical areas; wings pale ochraceous,
the apical area and the subposterior margin pale tawny
brown, apex of anal area and posterior and apical margins
greyish, semiopaque ; head (including eyes) about as wide as
670 Mr. W. L. Distant on Homoptera.
mesonotum, lateral margins of the pronotum ampliate and
convex; opercula short, transverse, their inner margins
slightly overlapping, their posterior margins broadly convex ;
rostrum reaching basal segment of abdomen.
Long., excl. tegm., ¢ 22 mm.; exp. tegm. 65 mm.
Hab. Transvaal ; Murchison Range (C. R. Jones, Brit.
Mus.).
Allied to P. stridula, Linn.
Platypleura Ridleyana, sp. n.
?. Body and legs pale castaneous brown; posterior
abdominal segmental margins somewhat broadly olivaceous
green; tegmina hyaline, the costal membrane ochraceous,
the venation (excluding that of basal cell) testaceous red,
about basal half more or less opaque, a large quadrangular
black spot before middle of radial area and a smaller fuscous
spot near its extremity, an oblong spot at bases of first and
second ulnar areas, two similar spots at base of third ulnar
area, a spot at base of the long sixth and of the seventh and
eighth apical areas, two spots in fourth ulnar area, the inner-
most of which is very large, and a spot at apex of fifth ulnar
area, dark fuscous or piceous ; a fuscous fascia crossing bases
of first to fifth apical areas connected with an irregular apical
fascia of the same colour, a double series of fuscous marginal
spots on the longitudinal veins to apical areas; wings piceous,
with the venation testaceous-red, the base broadly opaque
creamy yellow, the apical half of anal area and more than
posterior margin pale hyaline; head (including eyes) about as
wide as mesonotum, lateral margins of the pronotum broadly
ampliate and obtusely angulate; rostrum about reaching the
middle of abdomen.
Long., excl. tegm., 9 173 mm. ; exp. tegm. 52-56 mm.
Hab. Straits of Malacca; Ding Ding Islands (H. N.
Ridley, Brit. Mus.). North Borneo (Banguey, Paris Mus.).
Platypleura Harmandi, sp. n.
Head ochraceous; a large spot at base of front, a transverse
fascia between eyes and the inner margin of eyes black ;
pronotum brownish ochraceous, slightly obscurely piceous on
anterior and posterior margins, two small obsolete piceous
spots before middle of posterior margin ; mesonotum ochra-
ceous, with four anterior obconical spots, of which the two
central are shortest, a central discal lanceolate spot, a discal
spot to cruciform elevation, and a small spot in front of each
Mr. W. L. Distant on Homoptera. 671
anterior angle to same, black ; abdomen and tympana piceous,
thickly finely ochraceously tomentose on the segmental
margins ; head beneath, sternum, and opercula thickly finely
tawny pilose ; legs tawny brown; abdomen beneath brown,
its lateral margins cretaceously tomentose; face with the
central sulcation and transverse strize black and between face
and eyes a fascia of the same colour; tegmina very pale
tawny brown, becoming paler and greyish on the upper half
of radial area, in the upper ulnar area, and particularly so in
the apical areas; wings orange- yellow, about apical half
dark castaneous, containing two pairs of orange-yellow spots
placed near anterior and posterior margins. Head (including
eyes) about as wide as mesonotum ; lateral margins of pro-
notum ampliate and broadly angulate; opercula short,
rounded, not quite meeting centrally and not passing base of
abdomen ; rostrum reaching the posterior cox, its apex
black.
Long., excl. tegm., g 16 mm.; exp. tegm. 47 mm.
Hab. Cochin China; Lakhon (Har mand, Paris Mus.).
Allied to P. ar minops, Noualh.
Kongota Muri, sp. n.
?. Head, pronotum, and mesonotum ochraceous, the latter
inclining to clivaceous; front of head with a large discal
spot anteriorly connected with two narrow, lateral, marginal
fascie, black ; vertex with a short transverse fascia on ‘each
side of base of front and a discal transverse fascia passing
through the area of the ocelli and connected with the eyes at
lateral margins black ; pronotum with a central lanceolate
fascia not extending behind the middle, a short curved fascia
on each side of disk, all the fissures, two small rounded spots
at centre of posterior margin, and the posterior lateral
margins black ; mesonotum with a narrow, central, longitu-
dinal fascia, two short curved fascia at anterior margin, a
transverse series of four spots in front of cruciform elevation,
a spot on anterior angles of same, and a transverse spot on
each side of it, black ; abdomen above black, a large basal
spot and the segmental margins pale castaneous ; body
beneath and legs ochraceous, inclining to virescent; waved
fascize between the eyes crossing anterior portion ‘of face,
margin of face before clypeus, apical annulations to femora,
basal annulations and apices to tibie, and apices of tarsi
black ; tegmina brownish ochraceous, a little paler on costal
membrane and apical area, with piceous spots and suffusions,
a double subapical marginal series of piceous spots being very
672 Mr. W. L. Distant on Homoptera.
distinct, and between the outermost are a series of larger
obscure ochraceous spots, in the radial area a piceous spot at
base, and three central, longitudinal, piceous fascie; wings
orange-yellow, the five upper radial areas with castaneous
ray-like fascia, and a narrow submarginal fascia of the same
colour.
Long., excl. tegm., 23 mm.; exp. tegm. 73 mm.
Hab. Natal; Durban (fF. Murr, Brit. Mus.).
This is the second described species in the genus, and
is not to be confused with K. punctigera, Walk.; the type
specimen being a female the structural characters of the
opercula cannot be described.
Fam. Fulgoridz.
Subfam. /urgorrv 2.
Genus PHRICTUS.
Phrictus, Spin. Ann. Soc. Ent. Fr. viii. p. 216 (1839).
Phrictus auromaculatus, sp. n.
Body above dull brownish ochraceous; head above with
two small spots on disk of vertex, the lateral angular areas
of same, and the lateral marginal areas black; pronotum
(excluding disk) suffused with black or piceous; mesonotum
with three large black spots; abdomen above (excluding
segmental and lateral margins and a broken, longitudinal,
central fascia) black ; body beneath, rostrum, and legs black
or piceous ; cephalic process beneath ochraceous ; coxal and
sternal spots and a lateral abdominal series of spiracular spots
somewhat silvery white; tegmina with about basal two thirds
brownish ochraceous, with the thickly reticulate venation
piceous, that on the costal membrane virescent, divided by
transverse black spots, the whole speckled with small distinet
yellow spots, apical area brownish ochraceous, with a few
scattered, irregular, piceous spots; wings with about basal
half bright yellow, with some scattered small white spots and
two oblique black maculate fascia ; anal area fuscous brown,
with the fine reticulate venation greyish; the apical area
black, speckled with small bluish spots; cephalic process
prominently angulate on each side in front of eyes, and then
narrowed before apex, which is broadly widened and some-
what upwardly raised, the two central longitudinal ridges
Mr. W. L. Distant on Homoptera. 673
are crenulate, the anterior apical transverse ridge angularly
sinuate, and centrally united with the carinate anterior margin
by a longitudinal ridge, beneath including face having “its
margins entire, centrally ridged on its dilated anterior area,
where the margins are also carinate, its surface finely rugulose ;
posterior tibia with five long spines; rostrum reaching
middle of abdomen.
Long., excl. tegm., 27-30 mm.; exp. tegm. 65-67 mm.
Hab. Bolivia (I. Steinbach, Brit. Mus.).
The cephalic process is shorter than in any of the previously
described species and the coloration is very distinct.
Genus EPpisctus.
Episcius, Spin. Ann, Soc. Ent, Fr, viii. p. 249 (1839).
Episcius bolivianus, sp. n.
Head above, pronotum, and mesonotum testaceous brown $
abdomen above castaneous, its apex tomentosely cretaceous :
body beneath and legs brownish ochraceous, the femora and
abdomen more or less suffused with piceous ; tegmina pale
umber-brown, a little more than basal half, only reaching
costal margin at extremity, dark fuscous nora. margins w ith
scattered minute fuscous strize and a few also on apical area ;
wings with about basal two thirds purplish red, here and
there slightly suffused with pale fuscous, outwardly broadly
and irregularly margined with fuscous, the posterior margin
of anal area greyish, apical area hyaline; vertex longer than
broad, moderately concave, rugulose; pronotum finely rugu-
lose, centrally tricarinate, the area bounded by the carinations
quadrangular ; mesonotum very finely rugulose, tricarinate,
the central carinations very much waved and almost meeting
anteriorly ; rostrum slightly passing the middle of abdomen ;
legs marginally longly pilose, posterior tibiz with seven
spines.
Long., excl. tegm., 23 mm.; exp. tegm. 58 mm.
Ilab. Bolivia (L. Steinbach, Brit. Mus.).
Apart from the different coloration, this species may be
distinguished from £. platyrhina, Germ., by the quadrangular
carinate area of the pronotum; in Germar’s species it is
anteriorly convexly rounded.
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. Ad
(or)
~i
He
Mr. C. O. Waterhouse on
LX XVI.— Note on some British Culicidee.
By Cuas. O. WaTERHOUSE, F.E.S.
TAvinG had occasion to collect and make observations on the
larvee of Culicide, I was fortunate enough to rear froma larva
found in the New Forest a species which appears to be new
and of which I subjoin a description kindly drawn up for me
by Mr. F. V. Theobald. I also reared the male of Culee
diversus, 'Vheobald, which was not previously known. My
attention was first directed to the larva of C. Waterhouset by
the length of time it remained under water without coming
to the surface for air, ‘This larva, which was of a very pale
green, almost always rested in deep water on its back or in
an oblique position. The longest time I ever actually watched
it was half an hour; but on one occasion I was working close
by the aquarium for two hours, and looked at it at frequent
intervals. Having carefully noted its position, I feel certain
that it never moved during the whole of that time. This is
in great contrast with the larva of C. cantans, which rarely
remains down in the water for more than half a minute.
The following are Mr. Theobald’s descriptions :
Culex Waterhouse, sp. n.
Thorax black, clothed with pale golden to creamy scales.
Abdomen deep brown, with pale basal bands. Palpi of male
brown, with yellowish mottling in places, two basal white
bands to the two apical segments, and two yellow bands
towards the base. Legs deep brown, femora and tibiz
mottled with yellow; metatarsi and first two tarsals of fore
and mid legs with narrow basal white bands; in the hind
legs the white bands are broad. Apical segment of male
claspers with spines; median processes broadly expanded
apically. Fore and mid ungues unequal, hind ungues equal,
all uniserrated.
3. Head deep brown, clothed with rather large, creamy,
curved scales, long, narrow, ochreous, upright, forked scales
in front, with rather broader and shorter ones behind, and flat,
creamy, lateral scales. Palpi with the two apical segments
nearly the same length, deep brown, with an irregular, white,
scaled basal area to each, and with blackish hair-tufts; the
long antepenultimate segments with two broad, creamy, scaled
bands, the two apical segments with black hair-tufts and
also black hairs on the apex of the antepenultimate. An-
tenne with very deep brown plume-hairs. Proboscis deep
brown, unbanded.
Thorax black, with rather large, curved, pale golden scales,
some British Culicide. 675
somewhat paler over the roots of the wings; scutellum testa-
céous, deep brown along the border of the mid-lobe, and with
a deep brown patch on each side between the mid- and side-
lobes, clothed with rather large, narrow, curved scales of
similar hue to those of the mesothorax; posterior border-
bristles dense, pale golden; pleurze deep brown, with pale
creamy flat scales ; metanotum brown. Abdomen blackish,
with basal creamy bands and pale brown to dull golden hairs ;
basal segment with many pale scales.
Legs with the femora and tibix mottled brown and yellow,
the fore and mid metatarsi with narrow basal white bands,
also the first two tarsals; in the hind legs the bands are
broader and are present on all the segments ; fore and mid
ungues large, unequal, both uniserrated, the hind equal and
uniserrated.,
Wings with rather broad lateral vein-scales, especially on
the branches of the second long vein; many scales on the
first long vein somewhat Tentorhynchus-like, First sub-
marginal cell considerably longer and narrower than the
second posterior cell, their bases nearly level; stem of the
first submarginal cell nearly as long as the cell; stem of the
second posterior longer than the cell; mid cross-vein longer
than either the supernumerary or posterior cross-veins, the
latter about its own length distant from the mid cross-vein.
Male genitalia with claspers long and broad, spiny towards
the apex, the apical segment short and thick ; one side of the
apex of the basal lobes armed with short hook-like spines; a
large pineapple-shaped densely tuberculate process from the
base of each side; the two median processes long, composed
of two segments, the apical one broadly knife-shaped and
curved on its inner edge.
Length 55 mm.
Habitat. New Forest, Brockenhurst (C. O. Waterhouse).
Observations. Described from a single male easily told
from all other dark-banded-legged European species by the
very marked male genitalia. ‘I'he male genitalia of the type
are preserved in balsam. The squamose characters of the
wing exclude it from Cu/ea; but until female material is
examined I prefer to leave it provisionally in that hetero-
genous genus.
Culex diversus, Vheobald.
Mon. Culicid. p. 73 (1901) (Q only described).
3. Palpi deep violet-black, with traces of yellow scales at
the base of the two apical segments and on the antepenulti-
mate segment, the penultimate and apex of the antepenultimate
676 Miscellaneous.
with dense dusky hair-tufts, the apical one with scanty hairs,
the two apical segments and apex of the antepenultimate
slightly swollen, the last two of nearly equal length, apical
seement blunt; the hair-tuft on apex of antepenultimate
seoment very dense. Proboscis deep brown, with dull
yellow scales at the base. Antenne deep brown, with deep
brown plume-hatrs.
Thorax as in the female. Abdomen blackish, with basal
pale bands, moderately hairy, hairs pale. Genitalia densely
hairy, hairs golden; apical joint of claspers curved, with
the terminal segment long and bent at the tip, a few short
spines below its junction with the larger part of the clasper,
basal lobe densely hairy in places, especially on the inner side ;
between the basal lobes a long dense mass of golden hairs,
two long curved processes with short spines on their inner
lower edge, and a curious group of flattened curved spines
forming a prominent object between them and the basal lobes.
Legs deep blackish brown, except the femora, which are
pale beneath. Fore and mid ungues unequal, the larger
biserrated, the smaller uniserrated; hind ungues large,
simple, uniserrated.
Wings with the first submarginal cell longer and narrower
than the second posterior cell, its base nearer the apex of the
wing, its stem a little longer than the cell; stem of the
second posterior cell also longer than the cell ;_ posterior cross-
vein about its own length distant from the mid cross-vein.
Length 5:5 to 6 mm.
Hatitat. New Forest, Brockenhurst.
Observations. Bred by Mr. Waterhouse. The male
genitalia are very marked.
MISCELLANEOUS.
On the Affinities of Herpetomonas subulata *, and the Phylogeny of
the Trypanosomes. By Louis Lécer.
Tur structure of Herpetomonas subulata points to its extremely
close affinity with certain species of Crithidia previously described
by me. To Crithidia minuta of Tabanus tergestinus, Egg., espe-
cially, the resemblance of the young gregarine and monad forms is
such that it is impossible to distinguish these two Flagellata in
these stages.
* [This parasite is described by M. Léger in the number of the Comptes
Rend. Hebd. des Séances de la Soc. de Biologie (Dec. 50, 1904, pp. 613-
615) from which the present extract is taken. He states (loc. eit. p. 613)
that he met with it in the alimentary canal of Tabanus glaucopis, Mg.
(a species of horse-fly which occurs in Great Britain), in the south of
France. According to M. Léger.it is not common, and he found it only
four times in sixty specimens of 7Jabanus and Haematopota coliected in
autumn upon cattle and horses,—TRANSL. |
Ann. & Mag. Nat. Hist. §.Z Vol. XVI. PIX
P.Hiahley, de et lith
Cinaohnyra Nharoata.
Misecllaneous. 677
This peculiarity has led me to take up afresh the study of
C. minuta in the monad form, which I had not met with in its
fully developed state at the time of my first observations. Asa
result I was able to convince myself that in this form C. minuta
exhibits all the characters of Herpetomonas, and should conse-
quently be assigned to this latter genus. H. minuta is distinguished
from H. subulata only by being shorter and not tapering posteriorly.
The genus Crithidia must therefore be restricted at the present
time to two species—C. fasciculata of Anopheles and C. campanu-
lata of Chironomus—which are characterized by the bulky, piriform,
or campanulate form of their body.
There is little further need to insist on the weakness of this
systematic arrangement, which, although necessary, 1s undoubtedly
destined to have to submit to serious attacks in proportion as we
become better acquainted with the development of these Flagellata.
A certain number of these crithidian or herpetomonad forms found
in the biting insects are probably stages of Hemoflagellates met
with in Vertebrates. This opinion, which I advanced as early as
1902, has been justified by the admirable researches of Schaudinn *,
which showed that Trypanosoma noctue multiplies in Culex in a
erithidian form. In the case of the Crithidia of Anopheles the fact
is scarcely open to question, and as regards the herpetomonad
forms found in species of Tabanus and Hematopota, by reason of
their hosts’ mode of feeding, aud of the number, form, and minute
size of the parasites, it is also very possible that we have to deal
with stages of Hemoflagellates +; but this is not certain, for we
know that species of Herpetomonas are found in insects that do not
bite (Musca, Sarcophaga, Pollenia, and Fucellia, according to recent
observations of my own, &c.), in which their entire life-cycle takes
place (cf. Herpetomonas musce domestice, according to Prowazek).
The establishment of this fact, taken in conjunction with the
close relationship between the three genera, Herpetomonas, Crithidia,
and Trypanosoma, which has recently been well brought to light by
Laveran and Mesnil {, and further strengthened as it is by the
striking similarity of their young stages and the morphology of
Herpetomonas subulata, should lead us to search among the species
of Herpetomonas for the ancestors of the Trypanosomes with a
morphologically anterior flagellum (of the type of Prypanosoma noctuce
according to Schaudinn) §.
While at first performing their entire life-cycle in non-biting
insects these herpetomonad forms underwent progressive modifica-
tion in those among the latter that became hematophagous. Here
a much richer nutritive medium constituted by the blood, which
* Schaudinn, ‘ Arbeiten a. d. kaiserlichen Gesundheitsamte,’ Bd, xx,
(1904).
+ From this point of view one cannot refrain from noticing the great
resemblance between the little fixed forms of Herpetomonas subulata and
Piroplasma.,
{ Laveran and Mesnil, ‘Trypanosomes et Trypanosomiases’ (Paris,
1904),
§ Schaudinn, oc. cat.
678 Miscellaneous.
was absorbed and then digested, provoked an enormous multiplica-
tion of the parasite and at the same time prepared it to live in a
new medium, the circulating blood of Vertebrates, into which it
was bound finally to make its way, by reason of its minute size
and its host’s mode of feeding. 'Thus was brought into existence
the type of Trypanosome with anterior flagellum by progressive
development of an undulating membrane, which, with Senn * and
Laveran and Mesnilt, I regard as a character of the adaptive
order, in relation to the consistency of the medium in which the
parasites live.
The Trypanosomes of the blood, therefore, represent merely a
partial and secondary adaptation of a parasite primitively intes-
tinal or enteroccelomic of Invertebrates, and this explains why they
have to return into the latter in order to perform their sexual
reproduction.
The same considerations are, moreover, applicable to the Plasmo-
dium of malaria, in which schizogony alone takes place in the
blood of man, the sexual reproduction requiring the return of the
parasite into the mosquito.
As regards the Trypanosomes with a morphologically posterior
flagellum (of which, according to Schaudinn{, Trypanosoma
Ziemanni is the type), the lineage of these appears to be philo-
genetically very different from that of the former. We know asa
matter of fact from Schaudinn’s observations that they attach
themselves by the pole opposite to the flagellum ; now, since in the
case of Herpetomonas and Crithidia the fixative rostrum is homolo-
gous with an anterior flagellum, it is reasonable to suppose that
these Trypanosomes are derived from forms primitively provided
with two oppositely directed flagella (but not bipolar)—such as the
species of Z'rypanoplasma, for example.
As the result of a morphological study of the latter genus § I
had been led to consider Trypanosomes in general as species of
Trypanoplasma which had lost their anterior flagellum. This mode
of regarding them is, however, too exclusive, as I have shown
above, but I think that it may still apply to the Trypanosomes with
a posterior flagellum. It follows then that we have in the Trypano-
somes two very distinct types, which have sprung from different
stems, to which there correspond, moreover, according to the
researches of Schaudinn ||, two different types of development of
the ookinete.
One of the most important points in connexion with the morpho-
logy and the classification of the Trypanosomes would therefore be
to determine their pole of fixation, which, as regards the majority of
species, is at present unknown.—Comptes Rendus Hebdomadaires
des Séances de la Société de Biologie, tome lvii. pp. 615-617.
* Senn, ‘ Archiv f. Protist.’ Bd. L. Heft 2, 1902, p. 353.
+ Laveran and Mesnil, loc. cit,
¢ Schaudinn, Joe. cit.
§ Léger, Comptes Rendus Acad. d. Se., March 28 and April 4, 1904.
|| Schaudinn, loc. eit.
679
INDEX to VOL. XVI.
ABAGAZARA, characters of the new
genus, 206.
Abreeus, new species of, 349.
Abricta, new species of, 27, 281.
Abroma, new species of, 32, 561.
Achilognathus, new species of, 364.
Adenia, characters of the new genus,
210.
Abhomana, characters of the new
genus, 25,
Aiteta, new species of, 593.
Akamba, characters of the
genus, 562.
Alcock, Dr. A., revision of the
genus Peneus, 508.
Amphipoda, on the secondary ap-
pendage of the upper antenne as a
character in the, 464.
Ancistrogaster, new species of, 490.
Anechura, new species of, 493.
Andersen, K., on the bats of the
Rhinolophus philippinensis group,
243; on the bats of the Rhino-
lophus arcuatus group, 281; on
the bats of the Rhinolophus
macrotis group, 289; on Hippo-
siderus diadema, 497; on the
species and subspecies of Rhino-
lophus, 648.
Anthela, new species of, 150,
Anthometra, new species of, 628,
Anthophora, new species of, 395.
ceruginosa, note on, 295.
Arber, I. A. N., on the sporangium-
like organs of Glossopteris Browni-
ana, 460.
Arcyophora, new species of, 585.
Arfaka, characters of the new genus,
276.
Aviola, new species of, 601.
Armadillidium vulgare, on the dis-
tribution of, 431,
Arthroleptis, new species of, 108,
180.
Aryicanthis pumilio, on the various
forms of, 629,
Asota, new species of, 618.
Asterid, on the development of an,
with large yolky eggs, 387.
new
Atelopus, new species of, 181.
Atractaspis, new species of, 180.
Atylotus, new species of, 198.
Atyopeneus, characters of the new
genus, 524.
Auchenisa, characters of the new
genus, 18.
Baleenopteride,
species of, 403,
Barasa, new species of, 549,
Barbus, new species of, 642.
Barrett-Hamilton, Capt. G. E. H.,
on the Mus orthodon of Hensel,
452.
Bate and Westwood’s ‘ British
Sessile-eyed Crustacea,’ revised
nomenclature of the species de-
scribed in, 78.
Batrachians, new, 107, 180.
Bavea, characters of the new genus,
214,
Beatricella, characters of the new
genus, 568,
Blenina, new species of, 151, 543.
Bocana, new species of, 624.
Bombus, new species of, 223, 392.
Books, new :—Darton’s Preliminary
Report on the Geology and Water
Resources of Nebraska, 258 ;
Elliot's Land and Sea Mammals
of Middle America and the West
Indies, 260 ; Guide to the Gallery
of Birds in the British Museum,
261; Miall’s House, Garden, and
Field, 261; Uhlig’s Fauna of the
Spiti Shales, 365; Irving’s
Economie Resources of — the
Northern Black Hills, 453 ; Iowa
Geological Survey, vol. xiv., 455;
Maryland Geological Survey,
Miocene, 457; Fabre’s Souvenifs
Entomologiques, 459; Enteman’s
Coloration in Polistes, 576.
Bothriocephalus infundibuliformis,
note on, 117.
Boulenger, G. A., on the freshwater
fishes of Africa, 36 ; on batrachians
and reptiles from Angola, 105; on
new snakes from 8. Arabia, 178;
observations on
680
on new tailless batrachians, 180;
on specimens of Rana esculenta
from Persia, 552; on the habitat of
Rana Blanfordii, 640; on fishes
from the Kasai River, 640; on
fishes from Lake Bangwelo, 642.
Bower-bird, note on the satin,
350,
‘British Sessile-eyed Crustacea,’
revised nomenclature of the species
described in, 78.
Buckman, 8. 8., on the nomenclature
of types in natural history, 102 ;
on the types of S. P. Pratt and
Young and Bird, 264,
Burbunga, characters of the new
genus, 29,
Burr, M., on new Forficularia, 486.
Butler, Dr. A. G., on the satin bower-
bird, 350.
Cacyparis, new species of, 604.
Czenojoppa, characters of the new
genus, 162.
Calliostoma, new species of, 184.
Calman, Dr. W. T., on a new river-
crab from Yunnan, 155.
Calotes, new species of, 133.
Cameron, P., on a new genus and
species of Cynipidee, 20; on new
parasitic hymenoptera from Borneo,
159.
Carea, new species of, 598.
Casyapa, new species of, 612.
Catocala, on the red colour in the
hind wing of, 445.
Cheetospania, new species of, 489.
Chandica, new species of, 605.
Chilton, Prof. C., on the distribution
of some terrestrial Isopoda intro-
duced into Australasia, 428.
Chionama, new species of, 145.
Chloritis, new species of, 194.
Chlorocystaria, characters of the new
division, 212.
Chrotopterus,
308.
Chrysichthys, new species of, 641,
645.
Cicadide, revision of the, 22, 203,
265, 553, 668.
Cichlosoma, revision of the genus,
60, 225, 316, 433 ; new species of,
227, 330.
Cinachyra, on the oscules of, 662.
Clanculus, new species of, 185.
Clarias, new species of, 644,
Cleis, new species of, 149.
new subspecies of,
INDEX.
Cockerell, T. D. A., descriptions and
records of bees, 216, 292, 301, 392,
465, 477.
Coelioxys, new species of, 221.
Coendou, new species of, 310.
Coleoptera, new, 11, 340, G04.
Colobus, new subspecies of, 452.
Columbella, new species of, 189.
Conibosa, characters of the new
genus, 215.
Copepoda, notes on British, 567.
Coryphieus, new species of, 347,
Cotylopus, new species of, 562.
Crocisa, new species of, 218.
Crustacea, new, 8, 155, 525, 569; on
the nomeuclature of the British
sessile-eyed, 78.
Cucumaria Montagui, note on, 552.
Culex, new species of, 674.
Culicid, on some British, 674.
Cyanoxorides, new species of, 164.
Cylistosoma, new species of, 544.
Delavalia, new species of, 569.
Delma, new species of, 131.
Delocoma, characters of the new
genus, 153.
Dendromus, new species of, 175.
Dermaptera, new, 486.
Derotettix, new species of, 209.
Diacrisia, new species of, 145, 620.
Diagramma, new species of, 364.
Diceropyga, new species of, 557.
Diemenia, characters of the new
genus, 206,
Dilophothripa, new species of, 584.
Diplodonta, new species of, 191.
Diptera, new, 198, 674.
Distant, W. L., Rhynchotal notes,
22, 203, 265, 553, 668.
Dobsonia, new species of, 425.
Donald, Miss J., observations on some
of the Loxonematide, 461; on
some Gasteropoda from Llanga-
dock, 461.
Druce, H., on new Nymphaline,
549.
Druseia, characters of the new genus,
165.
Dulderana, characters of the new
genus, 280.
Kblisia, new species of, 345,
Hdyia, characters of the new genus,
159,
Eeernia, new species of, 139.
Kgnasia, new species of, 625,
Eleognatha, characters of the new
genus, 577,
END EX:
Emmiltis, new species of, 628,
EKomops, characters of the new genus,
572.
Kpiplema, new species of, 626,
Episcius, new species of, 673.
Hrastria, uew species of, 620,
Erizada, new species of, 586.
Ethalia, new species of, 187.
Kublemma, new species of, 621,
Kuchelus, new species of, 186.
Kupheedra, new species of, 550,
Euryglossa, new species of, 292, 472.
Kutelia, new species of, 372.
Hxoneura, new species of, 465.
Fantham, H. B., on hermaphroditism
and vestigial structures in the
reproductive organs of Testudo
greca, 120.
Fin-whale fishery, on some results of
the North-Atlantic, 403.
Fishes, on the treshwater, of Africa,
36; new, 227, 530, 361, 363, 643 ;
from the Kasai River, on, 640,
Forficula, new species of, 493,
Forficularia, notes on, 486,
Fossarina, new species of, 188.
Fossarus, new species of, 188.
Frisch’s generic names, on, 461.
Fulgora, new species of, 566,
Gazuma, characters of the new genus,
268.
Geological Society, proceedings of
the, 460.
Georychus, new species of, 175.
Giaura, new species of, 541.
Gibbula, new species of, 192.
Glauconia, new species of, 178.
Gnathophausia, new species of, 9.
Gobius, new species of, 564.
Gonolabis, new species of, 487.
Guaranisaria, characters of the new
genus, 560,
Gudanga, characters of the new
genus, 208.
Guereza, on a new, 452.
Gymnotympana, new species of, 563.
Gyrtona, new species of, 537.
Halictus, new species of, 300, 307.
TLampson, Sir G. F., on new species
of Noctuide, 869, 533, 577.
Hasora, new species of, 618.
Hemidictyaria, characters of the new
division, 275.
Hemiplecta, new species of, 195,
Henderson, Dr. E. H., on the de-
velopment of an asterid with
- large yolky eggs, 387.
681
Herichthys, new species of, 436.
Herpestes, new species of, 170,
Herpetomonas subulata, on the affini-
ties of, 676.
Hesperoptenus, new species of, 575.
Hexanchus griseus, note on, 571.
Hipposiderus diadema and its allies,
remarks on, 497.
Tlister, new species of, 346, 609.
Histeridze, new, 340, 604.
Holland, W. J., on a new noctuid
from Sierra Leone, 18.
Holt, EK. W. L., on the schizopods
collected by the ‘Oceana,’ 1.
Homoptera, new, 23, 205, 265, 553,
668.
Hovana, characters of the new genus,
279.
Hyla, new species of, 182.
Hymenoptera, new, 20, 159, 216,
292, 301, 392, 465, 477.
Hy pothripa, new species of, 540.
Ilma, characters of the new genus,
613, ;
Ingura, new species of, 369,
Inyamana, characters of the new
genus, 266.
Iruana, characters of the new genus,
278.
Iscadia, new species of, 579.
I.mene, new species of, 618.
[sopoda, on the distribution of some
terrestrial, 428.
Jacatra, characters of the new genus,
Hills
Jamiiia, new species of, 551.
Janson, O. I., on new Cetontida
from British New Guinea, 11.
Katerythrops, characters of the new
genus, 7.
Kerunia, on the Eocene genus, 93.
Kirkpatrick, R., on the oscules of:
Cinachyra, 662.
Kongota, new species of, 671.
Koranna, characters of the new
genus, 207.
Kumanga, characters of the new
genus, 215,
Labia, new species of, 487.
Léger, L., on the affinities of Herpeto-
monas subulata, and the phylogeny
of the Trypanosomes, 676.
Lemuriana, characters of the new
genus, 32.
Lepidoptera, new, 18, 142, 369, 533,
549, 577, 612.
Lepus, new species of, 176,
Ann. & Mag. N. Hist. Ser. 7. Vol. xvi. 45
682
Lewis, G., on new species of His-
teridze, 340, 604.
Lignicida, characters of the new
genus, 625,
Lissosternus, characters of the new
genus, 347.
Lithurgus, new species of, 295.
Loiada, characters of the new genus,
166.
Lomaptera, new species of, 12.
Lydekker, R., on an undescribed
Guereza, 432.
Lygosoma tzeniolatum, new variety
of, 140.
Macrolister, new species of, 345.
Malagasia, new species of, 32.
Mammals, new, 169, 243, 281, 289,
308, 314, 423, 432, 497, 572, 632.
Mapondera, characters of the new
genus, 212.
Mardalana, characters of the new
genus, 215.
Marmosa, new subspecies of, 313.
Mastacembelus, new species of, 647.
Masupha, new species of, 208.
Maurilia, new species of, 599.
Mecomera, new species of, 489.
Medasina, new species of, 627.
Megaptera longimana, observations
on, 416.
Melampsalta, new species of, 270.
Melampsaltaria, characters of the
new division, 269.
Melanothrix, new species of, 142.
Melissodes, new species of, 216.
Metapeneus, new species of, 526.
Metaptya, new species of, 587.
Metoponorthus pruinosus, on the
distribution of, 431.
Microloxia, new species of, 629.
Miller, G. S., jun., on the generic
names of Frisch, 461.
Minolia, new species of, 187.
Mollusea, new, 184, 192, 198, 551.
Monomatapa, new species of, 31.
Morokia, characters of the new
genus, 11.
Mus orthodon, note on, 452.
‘Museum Humfredianum,’ note on
the, 262.
Nanaguna, new species of, 583.
Neacomys, new species of, 310,
Neleynda, new species of, 35.
Neotrimorus, new species of, 161.
Nerita, new species of, 190.
Noctuidee, new, 369, 533, 577.
Nomia, new species of, 304.
INDEX.
Nomioides, new species of, 221.
Nopesa, Dr. F. B., on Kerunia, a
symbiosis of a hydractinian with a
cephalopod, 93.
Norman, Canon A. M., revised
nomenclature of the species de-
scribed in Bate and Westwood’s
‘ British Sessile-eyed Crustacea,’
78; on Cucumaria Montagui, 352;
on Ophiopsila anuulosa, 360.
Notodoma saturum, note en, 347.
Odontodes, new species of, 553.
Okanagana, characters of the new
genus, 23,
Omalodes, new species of, 608.
Omphalius, new species of, 186,
Oncotympana, new species of, 558.
Oocormus, characters of the new
genus, 181.
Ophiopsila annulosa, note on, 360.
Opisthocosmia, new species of, 491.
Oreta, new species of, 142.
Oromena, new species of, 152.
Oxzenanus, new species of, 624.
Padraona, new species of, 616.
Paharia, characters of the new genus,
25.
Panka, characters of the new genus,
34.
Papuina, new species of, 193.
Paracolletes, new species of, 477.
Paraneetroplus, characters of the
new genus, 436,
Parapeneopsis, new species of, 527.
Parapeneus, new species of, 525,
Parapetenia, characters of the new
subgenus, 324.
Parasites of the Tweed salmon, on
the, 115.
Parasphecodes, new species of, 296.
Parathelphusa, new species of, 156.
Paratilapia, new species of, 642, 646,
Paratropus, new species of, 610.
Paraxestis, new species of, 585.
Pardasena, new species of, 541.
Parnisaria, characters of the new
division, 203.
Parnkalla, characters of the new
genus, 29.
Pauropsalta, new species of, 273.
Peneus, revision of the genus, 508.
Perrhanza, new species of, 549.
Petrogale, new subspecies of, 425.
Phascogale, new species of, 427.
Phenacolletes, characters of the new
genus, 301.
Phrictus, new species of, 672.
_Yy{
INDEX. 683
Pidorus, new species of, 146.
Plesius, new species of, 342.
Platypleura, new species of, 554, 668.
Platysoma, new species of, 343, 605.
Plegaderus, new species of, 610.
Plotheia, new species of, 580.
Pogonozada, characters of the new
genus, 580.
Pomponia, new species of, 558.
Porcellio, on the distribution of
species of, 429.
Pristaulacus, new species of, 159.
Proechimys, new species of, 312.
Prosopis, new species of, 3899, 467.
Psammophis, new species of, 113.
Pseudoxiphophorus _ pauciradiatus,
observations on, 5362.
Ptilonorhynchus violaceus, note on,
350.
Purana, new species of, 555.
Pycnostigmus, characters of the new
genus, 20.
Pygoccelis, new species of, 605.
Pyrops, new species of, 564.
Quintilia, new species of, 205, 561.
Rana, new species of, 107.
— Blanfordii, on the habitat of,
640.
esculenta, on specimens of, from
Persia, 552.
Regan, C. Tate, revision of the
genus Cichlosoma &c., 60, 225,
316, 433 ; on fishes from Southern
Mexico, 361; on new fishes from
Japan, 363 ; on Hexanchus griseus,
571.
Reptiles, new, 113, 180, 178.
Rhesala, new species of, 625.
Rhinolophus, new species of, 247,
283, 289; list of the species and
subspecies of, 648.
Rhyssota, new species of, 194.
Ricardo, Miss G., on the Palearctic
Tabani, 196.
Rivula, new species of, 622.
Rosén, N., on the snakes in the mu-
seums of Lund and Malmé, 126;
on the lizards in the museum of
Lund, 12%).
Salmon, on the internal parasites of
the Tweed, 115.
Sapantanga, characters of the new
genus, 276.
Saprinus, new species of, 611.
Sarrothripa, new species of, 584,
Sawda, new species of, 556.
Scalaria, new species of, 190,
Schaposchnikow, Ch., on the red
colour in the hind wing of Cate-
cala, 445,
Schizopods collected during the
cruise of the ‘ Oceana,’ on the, 1.
Schuchert, C., on the nomenclature
of types in natural history, 102.
Sciurus, new species of, 509, 514.
Scott, Dr. T., notes cn British
Copepoda, 567.
Scutellina, new species of, 189.
Sherborn, C. D., note on the ‘ Mu-
seum Humfredianum,’ 262.
Siglophora, new species of, 603.
Sinna, new species of, 602.
Smith, E. A., on new Jand-shells
from British New Guinea &c.,
193; on a new land-shell from
Christmas Island, 551.
Solariella, new species of, 188.
Soritia, new species of, 145.
Southwell, T., on some results of the
North-Atlantic fin-whale fishery,
403.
Sowerby, G. B., on new marine
shells, 184; on a new shell from
the Cape Verd Islands, 192.
Spheeodes, new species of, 299.
Spiloxorides, new species of, 166.
Stagira, new species of, 213.
Steatomys, new species of, 174.
Stictoptera, new species of, 533.
Stilpnosoma, new species of, 476.
Swinhoe, Col. C., on Eastern and
Australian Heterocera, 142, 612.
Synodontis, new species of, 645.
Tabani, revision of the Palearctic,
196.
Tagiades, new species of, 612.
Taipinga, characters of the new genus,
2
Taphuraria, characters of the new
division, 25.
Tarache, new species of, 621.
Tarentola, new species of, 130.
Tattersall, W. M., on the schizopods
collected by the ‘ Oceana,’ 1.
Teinostoma, new species of, 187.
Telesto, new species of, 614.
Testudo greeca, on hermaphroditism
and vestigial structures in the
reproductive organs of, 120,
Tetrarhynchus grossus, note on, 118.
Tettigarctaria, characters of the new
division, 280.
Tettigomyieria, characters of the new
division, 266.
684
Thalacomys, new species of, 426.
Thomas, O., on mammals from An-
gola, 169; on new neotropical
Chrotopterus, Sciurus, &e., 308 ;
on a new squirrel from Burma,
314; on some Australasian mam-
mals, 422; on the generic names
of Frisch, 461; on a new genus
“and two new species of bats, 572.
Thomsonula, characters of the new
genus, 570. ‘
Thyrassia, new species of, 144.
Tibicina, new species of, 23.
Tilapia, new species of, 647.
Titulcia, new species of, 603.
Tortriciforma, new species of, 587.
Tosh, Dr. J. R., on the internal para-
sites of the Tweed salmon, 115.
Trachypeneus, new species of, 531,
Trichaulax, new species of, 17.
Trichiobracon, characters of the
genus, 161.
Trichionotus, characters of the new
genus, 168.
Tridrepana, new species of, 620.
Trigona, new species of, 220.
Prismarcha, new species of, 30.
INDEX.
Tritonidea, new species of, 191.
Trypanosomes, on the phylogeny of
the, 676.
Tyana, new species of, 601.
Types, on the nomenclature of, 102.
Ueana, characters of the new genus,
28.
Urabunana, characters of the new
genus, 274,
Urbona, new species of, 588.
Vespertilio, new species of, 573.
Walker, A. O., on the secondary
appendage of the upper antennze
as a character in the Amphipoda,
464.
Waterhouse, C. O., on some British
Culicidee, 674.
Westermannia, new species of, 588.
Whales, observations on species of,
403. ‘
Wroughton, R. C., on mammals from
Angola, 169; on the various forms
of Arvicanthis pumilio, 629.
Xenoglossodes, new species of, 218.
Xosopsaltria, new species of, 266.
Zamenis, new species of, 178.
Zethes, new species of, 154.
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