INHOUD VAN DEEL 110

BESSELING, A. J. — De watermijten van het Gagelplasje .

CHRYSANTHUS, Fr., O.F.M. Cap. — Spiders from New Guinea IX .

HEERDT, P. F. van & W. BONGERS. — A biocenological investigation of

salt marshes on the south coast of the Isle of Terschelling .

JEEKEL, C. A. W. — A revision of the Papuan Paradoxosomatidae

(Diplopoda, en. in the Museo Civico di Storia Naturale at

Genoa mae med) > ft RES ME

KIRIAKOFF, S. G. — New genera and species of Oriental Notodontidae

(Lepidoptera) . .

LEMPKE, B. J. — Catalogus der Nederlandse ea (Veertiende

Supplement) ee sasa

LIEFTINCK, M. A. — Notes on the genus Austrocnemis Tillyard, 1913

(Odonata, Coenagrionidae)

LUKOSCHUS, F., A. FAIN & M. M. J. BEAUJAN — Beschreibung neuer

Psorergates-Arten (Psorergatidae: Trombidiformes)

OBRAZTSOV, N. S. | — Die Gattungen der palaearktischen Tortricidae. III.

Addenda und Corrigenda. 2. Teil .

OBRAZTSOV, N. S. | — Die Gattungen der palaearktischen Tortricidae. II.

Die Unterfamilie Olethreutinae. 7. Teil. Tribus Eucosmini (Heinr.

1923). Fortsetzung .

PETERS, William L. — New species of Prosopistoma from the Oriental

Region (Prosopistomatoidea: Ephemeroptera)

| \WIEBES, J. T. — Redescription of Sycophaginae from Ceylon and India,

with designation of lectotypes, and a world catalogue of the Otitesellini

(Hymenoptera Chalcicoidea, Torymidae) .

WILLEMSE, Fer — Additional data on some genera and species of Acrididae

(Orthoptera, Acridoidea) from the Indo-Malayan region .

183

107

343

133

381

443

| = 110 AFLEVERING 1 1967

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M. A. LIEFTINCK. — Notes on the genus Austrocnemis Tillyard, 1913 (Odonata,

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NOTES ON THE GENUS AUSTROCNEMIS TILLYARD, 1913

(ODONATA, COENAGRIONIDAE)

M. A. LIEFTINCK

Rijksmuseum van Natuurlijke Historie, Leiden

ABSTRACT

A re-characterization is given of the little known coenagrionid genus Austrocnemis

Tillyard, based on the type species, A. splendida (Martin), from Australia. A second species

occurring in southern Papua, A. maccullochi (Tillyard), thus far considered a member of

Agriocnemis Selys, is also redescribed and transferred to the same genus. The structural

characters of both species are fully illustrated, proving the two insects to be intimately allied.

Austrocnemis is considered a highly specialized member of the ischnurine group of genera,

possessing several characters of its own. Brief notes are given on its peculiar habits and larval

characters, which are unique in the family. The genus may be purely Australo-Papuan in

distribution and restricted to the northern part of the Australian continent and the lowland

lakes of southern New Guinea.

The monobasic genus Azstrocnemis was founded by TILLYARD (1913) for the

reception of Agriocnemis splendida R. Martin (1901), from Victoria, Australia.

Except for the emphasis laid by the author on the remarkable size of the legs for

so small an insect, TILLYARD’s diagnosis of Austrocnemis merely comprises the

following remark: “Characters of Agriocnemis Selys, but with closer venation,

narrower wings, and remarkably long legs.” (the italics are those of TILLYARD).

MARTIN (1901 : 247), besides giving a combined description of the male and

isochromatic female of A. splendida, also referred to a “forme orangée” of the

latter, which he described separately on the next page. A comparative and very

full account of the dichromatism of the female was given some years later by

TILLYARD (1907). This does not, however, explain the author’s reticence in the

generic diagnosis, published six years afterwards, about the existence of a hetero-

chromatic female, with which, as we have seen, TILLYARD was well acquainted.

Later still, the same author gave a well-illustrated description of the semi-

opaque caudal lamellae of the larva of A. splendida in a comprehensive article

on the morphology of the caudal gills of zygopterid larvae (1917 : 98—99 and

626, textfig. 26 and pl. 2 fig. 18—20). Brief characterizations of the adult

dragonfly were published subsequently by MuNz (1919, venation), Ris (1930)

and FRASER (1957, 1960).

Up to this time Austrocnemis splendida (Martin) has remained the only known

member of the genus. However, I have now found that a second species,

Agriocnemis maccullochi Tillyard (1926), described from southern New Guinea,

should be assigned to it as well. The present status of the latter being still un-

known to me at the time of discussing the Coenagrionidae of New Guinea,

A. maccullochi was merely included in a key to the then known Papuan members

2 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 1, 1967

of Agriocnemis and listed as such also in a later paper on the regional fauna

(LIEFTINCK, 1932 : 597; 1949 : 261). Surprisingly, TILLYARD himself did not

recognize this little metallic green insect as Awstrocnemzs in spite of the fact that

a re-examination of the type and one paratype immediately reveal its difference

from Agriocnemis. In point of fact there can be no doubt that A. maccullochi

and splendida are intimately allied species.

In the course of the present study, a number of characters has been noted which

are important in classification but have been overlooked or not fully made use of.

These are laid down in the following augmented descriptions and explained by

illustrations. With regard to Agriocnemis, I have selected New Guinea specimens

of the widely distributed A. pygmaea (Ramb.) for the purpose of comparison

with Austrocnemis (see fig. 6).

Austrocnemis Tillyard, 1913

Stature small and of slender build. Body colour predominantly brilliant metallic

bronze-green. Pale postocular spots wanting; occiput and rear of the head

throughout metallic bronze-black. No light markings on dorsum of pro- and

synthorax.

Head relatively shorter and broader than in Agriocnemis. Inner margin of

compound eye markedly sinuous (S-shaped) in dorsal aspect, the anterior part of

the eye being distinctly broader than the posterior; postocular (epicranial) lobes

accordingly also much wider than usual and more strongly convex posteriorly.

Transverse middorsal area of occiput behind ocellar region small, hind border

perfectly straight. Median lobe of labium with narrowly U-shaped incision, depth

of emargination and total length of lobe in the ratio of 1 : 3.25. Face short.

Labrum and clypeus shaped similarly to Agriocnemis but labrum narrower, not

wider than clypeus. Frons low and rounded without indication of transverse

anterior crest; median sulcus deeply impressed. First two segments of antenna

almost equal in length. Ocelli placed approximately in equilateral triangle.

Pro- and synthorax shaped much as in Agriocnemis; posterior lobe of prothorax

strongly modified in both sexes. Lamina mesostigmalis transverse, anterior ridge

more elevated than posterior, apices rounded and extended laterad and/or hooked

inward.

Legs unusually long and slender; posterior femur reaching to apex of 2nd

abdominal segment. Femoral spines and tibial bristles rather short and fine,

numbering 6—8 on hind femora, 4—7 on hind tibiae (fig. 3 and 5), the terminal

spines of femur not or scarcely longer than width of femur. Claws with short,

triangular inferior tooth much shorter than the strongly curved end-hook.

Shape and venation of wings resembling Agriocnemis but wings less plainly

petiolated, more drawn out and with apices more pointed; nodus situated more

basad, the antenodal and postnodal parts in the ratio of 10 : 17 (in Agriocnemis

pygmaea 10 : 15.8). Arculus at or a little distal to Ax,, its limbs subequal in

length. Ac situated much nearer Ax, than Ax,; length of Ab variable, either

subequal to or a little shorter or longer than Ac. Costal side of fore wing

quadrangle of equal length or a little longer than proximal side and markedly

M. A. LIEFTINCK : Notes on the genus Austrocnemis 3

shorter than distal side; in hind wing its length is about 11%, times (or less) that

of proximal side and almost or fully equals the distal side. Fore wing with 3, hind

wing with 2—3 postquadrangular antenodal cells. M, arises at or little before

Px, in fore wing, at Px, in hind wing; Ma arises 2 (rarely 3) cells further

distad. Medio-anal link variable, normally distinctly broken but occasionally entire

in fore wing. C7, extending over 6—7 marginal cells in fore wing, 5—6 in hind

wing, this vein terminating at level of Px, or slightly beyond in all wings. Post-

nodal cross-veins of first series 7—8 in fore wing, 6—7 in hind wing. Pterostigma

very oblique, braced, subparallel-sided, less deep than the discal cell supporting it;

stigma slightly longer than wide, of equal depth in both pairs of wings. Only 2

poststigmal cells in fore wing, 3 in hinder pair.

Abdomen of normal length, very slender. Posterior border of 10th segment

raised and compressed medially, forming a short dome-shaped protuberance

recalling Ischnura and allied genera.

Penis shaped similarly to Agriocnemzs, with a row of short and fine shaft spines

on either side well before end of first segment and with fewer lateral setae at the

junction of first and second segments; second segment devoid of internal spines

but with well developed external limbus membranosus; third segment at first

rather narrow, then abruptly expanded and recurved, terminating in a pair of

broad divergent flaps whose apices are again tapered and bluntly pointed; lamina

interna conspicuous, slenderly curved and ribbon-like (fig. 4).

Male superior anal appendages subequal in length to segment 10, slightly

longer than inferior pair, armed with a robust hooked inferior process, apical

portion of each with bunch of closely set beard-like hairs (fig. 7 and 11).

Female with apex of 8th sternite not triangularly produced, the posterior border

being straight and entire, but armed with a small median spine or a pair of closely

approximated minute black teeth, which are situated immediately before the sternal

border. Genital valves and anal appendages of normal shape, ventral margin of

valves microscopically crenulate; styli short and thick (fig. 8).

Type species. — Agriocnemis splendida Martin.

Hab. — Australia and New Guinea.

As follows from the above description, Austrocnemis differs from Agriocnemis

in a number of striking characters, the most noteworthy of which are the delicate

bronze-green texture of most parts of the body, the absence of coloured postocular

spots, and the extreme length of the legs. Other peculiarities of structure shared

alike in both sexes are found in the posterior lobe of the prothorax and the

pinched apex of the tenth abdominal tergite. The armature and hairiness of the

male anal appendages and the presence of a rudimentary vulvar spine of the

female are features linking the genus up with Ischnura.

Austrocnemis splendida (Martin, 1901) Fig. 1—4 and 7—8

Material — 4 & (mostly adult, with distinct or slight tendency toward

pruinosity on thorax), 4.Q (adult, isochromatic), 5 9 (id., heterochromatic),

labelled in R. MARTIN's writing: “Agriocnemis n.sp.” or “A. splendida m./

4 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 1, 1967

Fig. 1—4. Austrocnemis splendida (Martin), Alexandra, V.; 1, posterior part of & pro

thorax and lamina mesostigmalis, dorsal view; 2, the same of 92, showing also oute

mesostigmal plate; 3, left hind leg of &, exterior view; 4, lateral view of penis. Fig. 5

Austrocnemis maccullochi (Tillyard), Lake Murray, S. New Guinea, left hind leg of &

paratype, exterior view. Fig. 6. Agriocnemis pygmaea (Ramb.), Merauke, S. New Guinea

left hind leg of 4, exterior view. Fig. 3, 5 and 6 enlarged on the same scale, scale-line =

2.0 mm

M. A. LIEFTINCK : Notes on the genus Austrocnemis 5

Alexandra fév.”, some with suffix “Lagunes”; 3 & (1 incomplete), 1,9 (iso-

chromatic), labelled by the same author “A. splendida m. Queensland”. All in

the Paris Museum. — 1 & 1 2 (ad., isochromatic), Alexandra, Victoria,

XII.1906, ded. R. J. TILLYARD, in Rıs’s collection, Natur-Museum Senckenberg,

Frankfurt. Lectotype & by present designation, “Alexandra, fév. Lagunes”, in

Paris Museum.

Male and female (ad., 1g isochrom.), Alexandra. — Labium pale chrome.

Labrum, mandibles, anteclypeus, genae and frons up to the insertion point of

antennae, bright blue (4) or greenish yellow (2), this colour on either side

extending obliquely upward along margin of compound eye to level of ante-

ocellar depression. Labrum strongly convex at middle, lacking a median impressed

area but with a short black streak or triangle at base; anterior parts fringed with

long bristle-like yellow hairs. Postclypeus dark metallic green, surrounded with

blue (4) or yellow (9), its surface shiny, superficially transversely striate.

Antennae brownish black, the sockets and an apical annule at the first segment

pale. Head otherwise bronze-green, the postocular areas more brilliantly so in

certain lights, with blue and coppery reflections but without forming definite

spots.

Pro- and synthorax with sharply defined yellow green and light blue markings,

as follows: lower part of propleuron and anterior lobe of prothorax narrowly;

apex of mesostigmal lamina; lower two-thirds of mesinfraepisternum; two large

bands running parallel to the sutures and broadly coalescent across second suture,

on metapleurae, the episternal band incomplete above and abruptly narrowed at

level of spiracle, the epimeral band not quite reaching latero-ventral margin;

indistinct confluent yellowish spots on ventral surface. All scuta, scutella and

axillary plates also bright blue in both sexes.

Coxae and trochanters of legs light blue ( 4) or greenish yellow (2), the

anterior half of all coxae dark brown or blackish, a dark spot also on outer face

of trochanters; legs including spines otherwise unicoloured light brown save for

narrow pale rings at extreme base of all femora.

Wing venation pale brown, centre of pterostigma brownish ochreous (4),

or wholly ochreous (2 ). Postnodal cross-veins of first series 7 in fore wing, 6 in

hind wing.

Male abdomen marked with blue, as follows: intersegmental dorsal annule of

1-2; small basal annules, progressively more widely interrupted middorsally and

confluent with narrow yellow stripes along ventral margin at tergites 3—7; a

stripe widening to a definite spot along ventral margin of 8. Apical segments

bright blue; 9 entirely blue except posterior border including the spines finely

black, and the entire dorsal surface of 10, which is bronze-black.

Penis, see generic diagnosis (fig. 4).

Anal appendages (fig. 7). Superior pair thick and blunt, subcylindrical, shorter

than 10th segment, slightly divergent; when viewed from behind each is distinctly

expanded interiorly at extreme base, then diminishes in width, the lower portion

laterally compressed, prolonged ventrad and terminating in a long slightly outbent

vermiform process which curves around the lateral branches of the inferior

appendages; inner surface and apex closely beset with curly pale golden hairs

6 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 1, 1967

Fig. 7—8. Austrocnemis splendida (Martin), Alexandra, V.; 7, terminal segments of &

abdomen, dorsal and right lateral view; 8, the same of ®; 8a, apex of 8th abdominal

sternite, ventral view, showing rudiments of divided vulvar spine

which are increasingly longer towards apex; colour dark with a pale spot at apex

above and with the ventral spur yellow. Basal portion of inferior pair bluish

exteriorly, for the rest yellow, the apices of the outer branches obscured.

Female (isochrom.) — Very similar to male. Prothorax shaped as in fig. 2;

raised lateral lobes of prothoracic hind lobe finely yellow. Anterior rim of lamina

mesostigmalis directed upward, the apical portion of each transverse, bright yellow,

ending in an abruptly incurved cylindrical process (fig. 2). Postnodal cross veins

8 in fore wing, 6—7 in hind wings. Body colour as in male but legs still paler

M. A. LIEFTINCK : Notes on the genus Austrocnemis 7

and side markings of thorax and latero-basal spots of abdominal segments 3—7

enlarged, the latter bright ochreous; intersegmental membranes of 7—9 and

apical spot at sides of 8 light blue; 9 predominantly blue with basal and apical

bronze-black marks slightly varying in extent and either of them indented by blue

in the median line; dorsum of 10 entirely bronze-black, the sides largely light

blue. Apex of segm. 10 pinched, forming a blunt median ridge. Anal appendages

flattened dorsoventrally, tips blunt; colour ochreous, dorsal surface of each brown

in basal half. Genital valves rather short, light ochreous, ventral armature in-

conspicuous, microscopically serrulate; styli gently curved, yellow (fig. 8).

Female (heterochrom.) — All light colours orange-yellow instead of blue

(deep brick-red in the living insect, sec. TILLYARD). Labrum, face except post-

clypeus, as well as entire prothorax, bright orange. Dorsum of synthorax metallic

green with a complete yellow juxtahumeral (mesepisternal) line continued down-

ward along mesinfraepisternal suture; behind this runs a wedge-shaped dark

mesepimeral band, its tapered end pointing upward, which occasionally coalesces

on both ends with the dark colour on dorsum; sides pale except a tiny dark spot

at dorsal end of second suture.

Legs throughout bright orange yellow with blackish spines and bristles.

Abdomen with first two segments largely orange-yellow; 2 with tiny black

basal ring, finely pointing caudad, and a twice broader apical annule which is

prolonged somewhat further at the median line and tapers to a point in the

opposite direction. No sign of pale colour on dorsum of terminal abdominal

segments.

The proportion of the heterochromatic form to the total number of females

that were examined by TILLYARD (1907) was calculated by him as 40 per cent.

All females except one possess a minute, though quite distinct, black spine at

the apex of the eighth abdominal sternite; only in the one specimen figured this

vulvar spine is divided into two, forming two acute denticles (fig. 8a).

Measurements: ¢ abd. + app. 15.0—17.0 mm, hind wing 10.5—11.5 mm;

Q 15.3—16.2 and 11.0—11.5 mm, respectively (TILLYARD, 1907, gives 17—19

mm for the @ abdomen, 11—14 mm for the hind wing).

Hab. — Australia (Victoria and Queensland).

The following interesting observations on the habits of A. splendida are

extracted from TILLYARD’s account of the species. In emphasizing the extra-

ordinary length of the legs, the author writes: “... I should like to point out that

this little insect has a great liking for sitting on the floating leaves of water-

plants. This liking is shared, to some extent, with many other small Agrionidae,

but I remarked, when collecting this species at Alexandra, Victoria, a distinct

difference between the habits of this species and others. Awstroagrion cyane (and,

in other localities, other small species have similar habits), visited the floating

leaves only occasionally, during bright sunshine; and the females also occasionally

oviposited along their edges, but seemed to prefer a less flat surface; as soon as

the wind blew much, or the sun became obscured by cloud, this species made for

the banks, and hid amongst the reeds or grass. But Austrocnemis splendida, using

its long legs to some purpose, would remain sitting on the flat, floating surface,

8 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 1, 1967

facing wind and cloud, with its legs spread widely out, so that it was most difficult

to capture. The best way was to plunge the net into the water under it, and so

secure it at the cost of drenching the net repeated; wavings of the net above it,

failed to frighten it. Towards evening, when the other Agrionidae had retired

into the rushes, these little insects could still be seen, with legs outstretched,

holding on to their favourite floating leaves.” (TILLYARD, 1913 : 456).

Austrocnemis maccullochi (Tillyard, 1926) comb. nov.

Fig. 5 and 9—11

Material. — 1 4 (adult, holotype), South New Guinea, Fly River area, with

five labels, two in A. R. MACCULLOCH's and R. T. TILLYARD's writing: “Western

Reach of Lake Murray, 20.XI.22 (very hard to catch)”; two in TILLYARD’s

writing: “K52345, Agriocnemis maccullochi Till. Holotype ¢” and “Agriocnemis

maccullochi Till. Holotype & R.J.T.'; and one red museum label with: “Austr.

Mus. Collection” in print. — 1 ¢ (adult, paratype), with same locality, date and

identification labels in R. J. TILLyARD’s writing. Holotype in Australian Museum,

Sydney; paratype in Australian National Insect Collection, CSIRO, Canberra.

The following characters, applying partly to either species, may nevertheless

serve to the recognition of A. maccullocht.

Male (holotype). — Whole anterior surface of head including the antennal

sockets and frons laterally, palest blue; postclypeus with transverse patch of

bronze-brown surrounded by a broad area of light colour. On the frons the light

colour extends upward along the eye margin to a level halfway fronto-clypeal

suture and median ocellus. Rear of the head dark save for a fine bluish stripe

bordering the eye margin posteriorly.

Prothorax bronze-green except the anterior lobe entirely blue and a bright

yellow line bordering the propleuron ventrally. Posterior lobe shaped much as in

A. splendida, divided into three parts, the side lobes in the form of narrow

transverse ridges, somewhat more curved and swollen than in that species, the

mid lobe greatly enlarged and strongly hollowed out with raised margins but less

constricted basally than in A. splendida (fig. 10). All dorsal ridges of thorax

cream-coloured, but ante-alar triangles dark.

Light markings at thoracic sides more restricted than in splendida: traces of

cream bordering mesinfraepisternum; a narrow stripe, incomplete above, along

lower two-thirds of mesepimeral (first lateral) suture; and a much wider wedge-

shaped mark, partly bordering the second suture, upon posterior (upper) two-

fifths of metepimeron. Sides for the greater part thinly overlaid with light blue

pruinescence. Lower parts of pleurae fading to brown, the ventral surface being

dark brown.

Coxae brown, touched with yellow below, at trochanters and at bases of femora;

legs otherwise pale brownish with the inner faces more yellowish; spines and

bristles dark brown (fig. 5).

Neuration yellowish brown; arculus at Ax,; nervures Ac and rudiment of Ab

approximately of equal length. Postnodal cross veins 7 in fore wing, 6 in hinder

M. A. LIEFTINCK : Notes on the genus Austrocnemis 9

Fig. 9—11. Austrocnemis maccullochi (Tillyard), Lake Murray, S. New Guinea; 9, general

aspect of & paratype; 10, posterior part of & prothorax and lamina mesostigmalis, dorsal

view (holotype); 11, terminal segments of 4 abdomen, dorsal and right lateral view

(holotype)

pair. Pterostigma grey-brown surrounded by pale yellow. Poststigmal cells 2 in

fore wing, 3 in the hinder pair.

Abdomen bronze-brown, the dark colour becoming gradually more restricted

to dorsum of segments posteriorly; intersegmental ring of segm. 1—2 bright blue,

those of the three terminal segments likewise pale, probably all blue in life.

Segm. 1—3 wholly unmarked and 4—7 with ill-defined pale basal annules only,

the one at 4 finely interrupted middorsally. Segm. 7—10 increasingly more

broadly pale-coloured laterally, but there are no blue dorsal markings on these

segments. i

10 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 1, 1967

Anal appendages brown, the curly pubescence at the distal portion of the

superior pair more golden brown (fig. 11).

The paratype is a slightly more matured specimen, differing from the type only

in its darker colouration, the light blue pruinescence overlaying the prothorax

and parts of the thoracic sides being more strongly marked and concealing most

of the surface.

Measurements: abd. + app. 14.0 mm, hind wing 9.0—9.2 mm.

Female unknown.

Hab. — Lowlands of southern New Guinea.

Superficially, the aspect of A. maccullocht is exactly that of the type species,

splendida. It seems to differ only in details of colouration, slightly smaller size

and, most importantly, in the structure of the prothorax and anal appendages.

These last differences are best understood by comparing the accompanying outline

sketches.

Evidently a very local and easily overlooked insect. Referring to MACCULLOCH's

field note “very hard to catch”, TILLYARD (1926: 161) suggests that “it seems

quite likely that it may have a somewhat similar habit of resting on floating

leaves of aquatic plants, which would make it very difficult to capture”. I may

add that since the time of MACCULLOCH's exploration of the lowlands in southern

Papua, no collecting of Odonata has ever been done in the extensive lake districts

of that country.

RELATIONSHIPS

Considering the evolutionary history of Azstrocnemis, TILLYARD (1913) sug-

gests the unusually long legs of its members to be an archaic character acquired

already by some remote ancestor and preserved by a few present-day descendants

without affecting other parts of the insect. It should be borne in mind, however,

that this one characteristic — possibly to be considered as an adaptation to the

peculiar mode of life described above — is not the only feature by which

Austrocnemis differs from the more advanced ischnurine genera. Whereas the

wing venation is typically that of Agriocnemis, the shape of the head and thoracic

segments, the absence of coloured postocular spots, as well as the scheme of body

colouration in general, are all very unusual, differing markedly from other genera

in the group.1) On the other hand, two features may be mentioned that would

leave little doubt of its connection with the Ischnura-Agriocnemis group of genera,

namely, (1) the existence of two colour forms of the female, (2) the presence

of a rudimentary “vulvar spine” in that sex, and (3) the structure of the male

penile organ. The two last-mentioned characters have not been taken into account

1) Austrocnemis was first placed by TirLYARD & FRASER (1938) in the “Coenagrion

Series’ of their ‘‘Coenagriinae’”’. FRASER (1957), while opposing two categories with

practically identical characters, created two new “‘subfamilies’’, the Ischnurinae and Agriocne-

minae, for a number of interrelated coenagrionid genera, placing Austrocnemis in the last-

mentioned group. Like many other suprageneric taxa proposed by FRASER, these categories

are artificially constructed groups, based on inadequate or poorly defined characters.

M. A. LIEFTINCK : Notes on the genus Austrocnemis 11

earlier but are believed to be of some importance with respect to the classification

of Austrocnemis. On the available evidence I am inclined to regard is as a

specialized offshoot from the main ischnurine stem of the Coenagrionidae. In

fact it stands out as by far the most highly specialized of all known genera of

this assemblage, taking an isolated position amongst them.

A brief note on the larva of Azstrocnemis, commented upon by TILLYARD

(1917) does not seem to be out of place in the present context. As TILLYARD’s

figures well show, the caudal gills are of the denodate form, approaching that

found in the majority of Old World ischnurine genera (including some montane

Ischnura and Oreagrion, see LIEFTINCK, 1959). What is peculiar about these

lamellae is their great length, their semi-opacity, and the wholly uninterrupted

fringe of long hairs all along the margins. Form and structure of the gills do not

correspond with any of the other described genera but most nearly resemble those

of certain (not all) species of Agriocnemis. It would be interesting to obtain

fresh material of the Austrocnemis larva in order to study its mouth-parts, an-

tennae etc., none of these structures having yet been made known.

ACKNOWLEDGEMENTS

I wish to express my best thanks to Dr. J. W. Evans and C. N. SMITHERS, of

the Australian Museum, Sydney, for the loan of the type of Agriocnemis maccul-

lochi Till., and also to Dr. J. A. L. WATSON, of the Division of Entomology,

CSIRO, Canberra, who sent me for comparison one of the topotypes of the same

insect preserved in the National Collection. Through the good offices of Dr. H.

SCHRODER, of the Natur-Museum Senckenberg at Frankfurt, I have been able to

examine some of TILLYARD's specimens of Azstrocnemis splendida (Martin) in

the F. Ris collection. Lastly, I wish to thank Melle S. KELNER-PILLAULT, of the

Muséum National d’Histoire Naturelle, for giving me access to R. MARTIN’s

collection of Odonata to study the type series of the last-mentioned species in the

Paris Museum.

REFERENCES

FRASER, F. C., 1957. A reclassification of the Order Odonata. R. Zool. Soc. N.S. Wales,

Sydney. 133 pp., figs.

, 1960. A handbook of the dragonflies of Australia. R. Zool. Soc. N.S. Wales,

Sydney. 67 pp., figs.

LIEFTINCK, M. A., 1932. The dragonflies (Odonata) of New Guinea and neighbouring

islands. Part I. Descriptions of new genera and species of the families Lestidae

and Agrionidae. Nova Guinea 15 Zool.: 485—602, figs.

——, 1949, The dragonflies (Odonata) of New Guinea and neighbouring islands. Part

VII. Results of the Third Archbold Expedition 1938—1939 and of the Le Roux

Expedition 1939 to Netherlands New Guinea (II. Zygoptera). Nova Guinea, new

ser. 5: 1—271, figs.

—, 1959. On the New Guinea species of Ischnura Charpentier and Oreagrion Ris,

with special reference to the larval forms and notes on the species of adjacent

regions. Nova Guinea, new ser. 10: 213—240, figs.

MARTIN, R., 1901. Les Odonates du continent Australien. Mém. Soc. Zool. France 19:

220—248, fig.

p2 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 1, 1967

Munz, P. A., 1919. A venational study of the suborder Zygoptera (Odonata) with keys for

the identification of genera. Mem. Amer. Ent. Soc. 3: 1—78, figs., 20 pls.

Ris, F., 1930. Drei Notizen über ostasiatische Agrioniden. Arkiv f. Zool. 21A : 1—32, figs.

TILLYARD, R. J., 1907. On dimorphism in the female of Australian Agrionidae. Proc. Linn.

Soc. N.S. Wales 32: 382—390.

—, 1913. On some new and rare Australian Agrionidae (Odonata). Proc. Linn. Soc.

N.S. Wales (1912) 37: 404—479, pls. 44—49.

—, 1917. On the morphology of the caudal gills of the larvae of Zygopterid dragon-

flies. Proc. Linn. Soc. N.S. Wales 42: 13—112, 606—632, pls. 1—6.

—, 1926. On a collection of Papuan dragonflies (Odonata) made by the late Mr. Allan

R. MacCulloch in 1922—3, with descriptions of new species. Rec. Austral. Mus.

15: 157—166, figs.

& F. C. Fraser, 1938. A reclassification of the Order Odonata. 1. Austral. Zool. 9:

125—169, 27 figs.

No. 2. P. J. Brakman, 1966. — Catalogus van Coleoptera uit Nederland en het omlig-

gende gebied (List of the Coleoptera of the Netherlands and adjacent region), 219 pp.,

map. D.FI. 45.— (£4.10—, $ 12.50).

In press: F. Willemse. — Preliminary revision of the genera Stenocatantops Dirsh & Uvarov

and Xenocatantops Dirsh & Uvarov (Orthoptera, Acridiidae, Catantopinae), ca. 200 pp

In preparation: C. A. W. Jeekel. — Nomenclator familiarum et generum Diplopodorum,

about 300 pp.

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_ DEEL 110 AFLEVERING 2 | 1967

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DIE GATTUNGEN DER PALAEARKTISCHEN

TORTRICIDAE

III. ADDENDA UND CORRIGENDA

2. Teil +)

Notes on the Palaearctic Laspeyresiini

VON

NIKOLAUS S. OBRAZTSOV + **)

Editor's note. — The present paper is compiled from notes in English, found

among the papers of the author after his decease. They form a continuation of the

Third Part of the present series which so far has been published in German. How-

ever, it seemed unwise to translate these notes; therefore they are now being

published, except for the usual editorial corrections, in their original state.

Abbreviations used for the sources of studied material

AMNH = American Museum of Natural History, New York, USA.

BMNH = British Museum (Natural History), London, England.

NHMW = Naturhistorisches Museum, Vienna, Austria.

RMNH = Rijksmuseum van Natuurlijke Historie, Leiden, Netherlands.

ZMHB = Zoologisches Museum der Humboldt-Universität, Berlin, East Germany.

ZSBS = Zoologische Sammlung des Bayerischen Staates, Munick, West Germany.

Dichrorampha (Paralipoptycha) plumbana (Scopoli)

Phalaena plumbana Scopoli, 1763, Ent. Carn.: 235, Fig. 592.

For further synonymy cf. Obraztsov, 1958, Tijdschr. Ent. 101: 255.

Dichrorampha (Lipoptycha) resplendana Heinemann, 1863, Schm. Deutschl. Schweiz (2)

1 (1): 236. — Herrich-Schäffer, 1863, Corresp.-Blatt zool.-min. Ver. Regensburg 17 : 124. —

Wocke in Staudinger & Wocke, 1871, Catal. Lep. europ. Faun.: 264, no. 1292. — Obraztsov,

1953, Mitt. Münchn. ent. Ges. 43 : 94. Syn. nov.

Lipoptycha resplendana : Rebel in Staudinger & Rebel, 1901, Catal. Lep. pal. Faun. 2:

128, no. 2303. — Kennel ir» Spuler, 1910, Schm. Eur. 2: 289. — 1921, Pal. Tortr.: 641,

PI. 23 Fig. 54.

Dichrorampha (Paralipoptycha) resplendana : Obraztsov, 1958, Tijdschr. Ent. 101 : 256. —

1961, ibid. 104 : 240. — Hannemann, Die Wickler 7 Dahl, 1961, Tierw. Deutschl. 48 : 84,

Fig. 147, Pl. 11 Fig. 4.

Sericoris rejectana Laharpe, 1858, Nouv. Mém. Soc. Helv. Sci. Nat. 16, Faune Suisse 6:

63. — 1864, Nouv. Mém. Soc. Helv. Sci. Nat. 20, Faune lépid. Suisse, Suppl. 3: 74.

Syn. nov.

Dichrorampha rejectana : Wocke in Staudinger & Wocke, 1871, Cat. Lep. eur. Faun.: 264,

no. 1290. — Rebel iz Staudinger & Rebel, 1901, Cat. Lep. pal. Faun. 2: 128, no. 2301.

*) This study has been carried out with a grant of the National Science Foundation.

**) Formerly Research Associate, the American Museum of Natural History, New York.

14 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Hemimene rejectana: Müller-Rutz in Vorbrodt & Müller-Rutz, 1914, Schm. Schweiz 2.

412. — Kennel, 1921, Pal. Tortr.: 721.

Lipoptycha rejectana: Müller-Rutz, 1922, Mitt. Schweiz. ent. Ges. 13: 232. — 1927,

ibid. 13 : 512. — 1932, ibid. 15: 238. — 1934, ibid. 16: 127, PI. 1 Fig. 19. — Weber,

1945, ibid. 19: 367.

Hemimene alpigenana (nec Heinemann) Müller-Rutz 72 Vorbrodt & Müller-Rutz, 1914,

Schm. Schweiz 2: 412.

Dichrorampha (?Lipoptychodes) rejectana : Obraztsov, 1953, Mitt. Münchn. ent. Ges.

43: 64.

Dichrorampha (Lipoptycha) rejectana: Obraztsov, 1958, Tijdschr. Ent. 101: 252. —

1961, ibid. 104: 240.

Dichrorampha (Paralipoptycha) rejectana : Sauter, 1960, Mitt. Schweiz. ent. Ges. 33 : 273,

Bl. kis, 2.

Dichrorampha (Lipoptycha) saturnana : Heinemann, 1863, Schm. Deutschl. Schweiz (2)

1 (1): 238.

The figures of an adult and the male genitalia of the lectotype of resplendana,

published by HANNEMANN (1961), leave no doubt that this species is a synonym

of plumbana. The color of the wings of plumbana is quite variable, and there is

no need to.reserve the name resplendana for an aberration with reddish gray,

yellowish suffused forewings, and whitish gray hindwings.

At this occasion it may be mentioned that the male genitalia, figured by Han-

NEMANN (1961, Fig. 146) as those of plumbana, belong to Dichrorampha (Pa-

ralipoptycha) aeratana (Pierce & Metcalfe).

SAUTER (1960) selected and examined the lectotype of Serzcoris rejectana, and

found its genitalia being very similar to those of plumbana. The only difference

was in the shape of the cornuti which in rejectana appeared to be irregularly

swollen from one side. Those in the plumbana specimens from Switzerland

SAUTER described as being spindle-shaped, slightly thickened in the middle. He

noticed that in four specimens of rejectana, other than the lectotype, the cornuti

also showed a lateral swelling, although less extreme than in the lectotype. I have

observed in many plumbana specimens from various localities that the shape of

the cornuti is rather variable, and that both shapes described by SAUTER may

sometimes be found in one and the same specimen. Therefore I am satisfied that

rejectana is synonymous with plumbana.

Dichrorampha (Paralipoptycha) discedana (Danilevsky) comb. nov.

Hemimene discedana Danilevsky, 1960, Rev. Ent. URSS 39: 942, Fig. 7.

Dichrorampha (Dichrorampha) montanana (Duponchel)

Ephippiphora montanana Duponchel, 1843, Hist. nat. Lép. France 4: 413, Pl. 83 Fig. 7.

For synonymy and literature cf. Obraztsov, 1958, Tijdschr. Ent. 101: 251, 258.

Hemimene blasiana Kennel, 1919, Mitt. Münch. ent. Ges. 8 [1918]: 89, PI. 3 Fig.

23—24.

Type. Lectotype of blasiana, male (without abdomen), Gustein, Austria |

(ZMHB).

OBRAZTSOV (1958) listed blasiana, as a probable synonym of montanana. The

present examination of the lectotype (the female, mentioned by KENNEL in the

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 15

original description, is missing) has confirmed the above conjecture, as far as this

is possible generally, i.e., without a study of the genitalia. The figure, published

by KENNEL, is decidedly too dark.

Dichrorampha (Dichrorampha) interponana (Danilevsky) comb. nov.

Hemimene interponana Danilevsky, 1960, Rev. Ent. URSS 39 : 939, Fig. 1, 2.

As noted by its author, this species resembles Dichrorampha (D.) simpliciana

(Haworth), by having similar shape and markings of the forewings. However,

judging from the shape of the valva it is related to D. (D.) petiverella (Linné)

and D. (D.) proxima (Danilevsky), and must be placed near these two species.

Dichrorampha (Dichrorampha) ambrosiana (Kennel)

Hemimene ambrosiana Kennel, 1919, Mitt. Münchn. Ent. Ges. 8 [1918]: 90, PI. 3 Fig. 25.

Dichrorampha (Dichrorampha) ambrosiana: Obraztsov, 1953, ibid. 43 : 23.

Dichrorampha (?Dichrorampha) ambrosiana: Obraztsov, 1958, Tijdschr. Ent. 101: 246.

Lectotype: 4, Kentei; lectoparatype: 4, Kentei (ZMHB).

Other specimens examined. 2 ¢ (genitalia of one of them, slide No. 3 — Obr.,

1961), Myski, Kuznetski Alatau, July 20—21, 1956 (M. I. FALKOVITSH)

(ZMHB).

KENNEL removed the terminal segments of the abdomen of the lectotype for

examination of the genitalia, but the genital slide cannot presently be located, as

most of the slides prepared by that author. The only information about the

genitalia of the lectotype is that “this species has other genitalia than Hem. flavi-

dorsana Knaggs and quaestionana Snell. {sic!}, both comparable with it in

markings and colour”. The abdomen of the lectoparatype of ambrosiana is also

missing, and the genitalia can only be studied in the two male specimens available

from Kuznetski Alatau (Myski), which are rather similar to the type specimens

of ambrosiana. These two specimens are slightly smaller than the lectotype and

lectoparatypes of ambrosiana, the forewing being 7.0 mm, instead of 7.5 mm, as

in the types.

The genitalia show relationship to the North American Dichrorampha (Dichro-

rampha) bittana (Busck). Thus, D. ambrosiana has appeared the first known

Palearctic species of the genus, in which the external margin of the basal opening

of the valva has an angulate process directed basad. In D. biffana this process is

much shorter than in ambrosiana, and has a somewhat different shape. Moreover,

the cucullus in bzttana is smaller than in ambrosiana, and not so much expanded

dorsad. The entire male genital structures of ambrosiana are in general more robust

than those of bittana. Externally bettana differs from ambrosiana in having the

dorsal spot of the forewings distinctly larger and obliquely outlined basally.

Dichrorampha (Dichrorampha) larsana (Danilevsky) comb. nov.

Hemimene larsana Danilevsky, 1960, Rev. Ent. URSS 39 : 941, Fig. 4, 5.

16 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Dichrorampha (Dichrorampha) tshetverikovi (Danilevsky) comb. nov.

Hemimene tshetverikovi Danilevsky, 1960, ibid. 39 : 942, Fig. 6.

Dichrorampha (Lipoptycha) alpigenana Heinemann

Dichrorampha (Lipoptycha) alpigenana Heinemann, 1863, Schmett. Deutsch. Schweiz [2]

1 (1): 236. — Wocke in Staudinger & Wocke, 1871, Catal. Lep. eur. Faun.: 264, no. 1293.

Obraztsov, 1958, Tijdschr. Ent. 101: 252. — Hannemann, 1961, Die Wickler 77 Dahl,

Tierw. Deutschl. 48 : 80, Fig. 137, PI. 7 Fig. 10.

Lipoptycha alpigenana : Rebel in Staudinger & Rebel, 1901, Catal. Lep. pal. Faun. 2: 128,

no. 2304. — Mitterberger, 1909, Verz. in Salzburg vork. Microlep.: 172. — Kennel in

Spuler, 1910, Schm. Eur. 2: 289. — 1921, Pal. Tortr.: 642, Pl. 23 Fig. 55. — Osthelder,

1939, Schm. Südbayerns 2: 109.

Dichrorampha (Lipoptychodes) alpigenana: Obraztsov, 1953, Mitt. Miinchn. ent. Ges.

43 : 63, Fig. 26.

Dichrorampha (Lipoptychodes) pfisteri Obraztsov, 1953, ibid. 43 : 65, Fig. 27.

Dichrorampha (Lipoptycha) pfisteri : Obraztsov, 1958, Tijdschr. Ent. 101: 252. — Han-

nemann, 1961, Die Wickler in Dahl, 1961, Tierw. Deutschl. 48 : 82, Fig. 142, PI. 7 Fig. 13.

Hemimene (Lipoptycha) alpigenana : Danilevsky, 1960, Rev. Ent. URSS 39 : 948, Fig. 17.

Hemimene pfisteri : Danilevsky, 1960, ibid. 39 : 948.

DANILEVSKY (1960) examined the genitalia of two males of alpigenana in

WockE's Collection, and one male from Pyhrgas, Upper Austria. He found that

the shape of the cucullus in this species is rather variable, and transitional forms

do exist. For this reason DANILEVSKY concluded that there are grounds for

separating D. pfisteri from D. alpigenana which in his former opinion were

conspecific. For the present this statement seems to be quite satisfactory.

The figure of pfisteri, published by HANNEMANN (1961), represents a rather

abnormal specimen with a very dark basal area of the forewings.

Dichrorampha (Lipoptycha) klimeschi Obraztsov

Dichrorampha (Lipoptychodes) klimeschi Obraztsov, 1953, Mitt. Münchn. ent. Ges. 43:

67, Fig. 28.

Dichrorampha (Lipoptycha) klimeschi : Obraztsov, 1958, Tijdschr. Ent. 101 : 252, Fig. 23.

Hemimene klimeschi : Danilevsky, 1960, Rev. Ent. URSS 39: 949.

DANILEVSKY (1960) doubts that &/zmeschi is a species distinct from a/pigenana

and is inclined to treat it as an extreme variety or a local subspecies of the latter

species. I cannot agree with this point of view. In addition to a narrow neck of

the valva and a rather distinctly shaped cucullus, “Hemzimene" klimeschi also

differs from “Hemimene’’ alpigenana externally, having black-coloured veins

before the ocelloid patch of the forewings.

Dichrorampha (Lipoptycha) infuscata (Danilevsky) comb. nov.

Hemimene (Lipoptycha) infuscata Danilevsky, 1960, Rev. ent. URSS 39: 942, Fig. 8, 9.

Dichrorampha (Lipoptycha) tianshanica (Danilevsky) comb. nov.

Hemimene (Lipoptycha) tianshanica Danilevsky, 1960, ibid. 39 : 944, Fig. 10—12.

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae IN)

Dichrorampha (Lipoptycha) alatavica (Danilevsky) comb. nov.

Hemimene (Lipoptycha) alatavica Danilevsky, 1960, bid. 39 : 945, Fig. 13.

Dichrorampha (Lipoptycha) nigrobrunneana (Toll)

Hemimene consortana (nec Stephens) Toll, 1939, Jahresber. Physiogr. Komm. Poln. Ak.

Wiss. 73: 251, 260.

Hemimene nigrobrunneana Toll, 1942, Zeitschr. Wiener ent. Ver. 27 : 169, Pl. 13 Fig. 5, 6,

Pl. 15 Fig. 22, 23. — 1949, Polsk. Pismo Ent. 18 : 160. — 1956, ibid. 24: 37, Pl. 3 Fig.

F4—6; Pl. 7, Fig. 31, 32; PI. 10 Fig. 61. — Danilevsky, 1960, Rev. ent. URSS 39: 947.

Hemimene fusca Danilevsky, 1948, Ent. Obozrenie 30: 78, Fig. 18.

Dichrorampha (Lipoptychodes) fusca: Obraztsov, 1953, Mitt. Münchn. ent. Ges. 43 : 69.

Dichrorampha (Lipoptychodes) nigrobrunneana : Obraztsov, 1953, ibid. 43 : 69.

Dichrorampha (Lipoptycha) fusca: Obraztsov, 1958, Tijdschr. Ent. 101: 252.

Dichrorampha (Lipoptycha) nigrobrunneana : Obraztsov, 1958, ibid. 101: 253.

Dichrorampha (Lipoptycha) alaicana Rebel status nov.

Dichrorampha alaicana Rebel, 1910, Iris 24: 9, Pl. 1 Fig. 6. — Caradja, 1916, ibid.

30073"

Lipoptycha tamerlana Kennel, 1919, Mitt. Münchn. ent. Ges. 8 [1918] : 91, Pl. 3 Fig. 26.

Hemimene alaicana Kennel, 1921, Pal. Tortr.: 721. — Danilevsky, 1960, Rev. Ent.

URSS 39: 947.

Dichrorampha (Dichrorampha) alaicana : Obraztsov, 1953, Mitt. Münchn. ent. Ges. 43 :

60. — 1958, Tijdschr. Ent. 101: 251.

Dichrorampha (Lipoptychodes) tamerlana: Obraztsov, 1953, Mitt. Münchn. ent. Ges.

43°: 70, Fig. 30.

Dichrorampha (Lipoptycha) tamerlana : Obraztsov, 1958, Tijdschr. Ent. 101: 253.

Hemimene (Lipoptycha) tamerlana : Danilevsky, 1960, Rev. Ent. URSS 39: 947.

Dichrorampha (Lipoptycha) eximia (Danilevsky)

Hemimene (Lipoptycha) eximia Danilevsky, 1948, Ent. Obozrenie 30: 76 (external

characters, not genitalia). — 1960, Rev. Ent. URSS 39 : 946, Fig. 15, 16.

Hemimene (Lipoptycha) gracilis (part.): Danilevsky, 1948, Ent. Obozrenie 30: 76, Fig.

17 (genitalia, not external characters; specimen from Daghestan).

Dichrorampha (Lipoptychodes) gracilis : Obraztsov, 1953, Mitt. Münchn. ent. Ges. 43 : 74.

Dichrorampha (Lipoptycha) gracilis (part.): Obraztsov, 1958, Tijdschr. Ent. 101: 253.

Dichrorampha (Lipoptycha) eximia (part.): Obraztsov, 1958, zbid. 101: 254.

Dichrorampha (Lipoptychodes) obraztsovi Amsel, 1959, Bull. Soc. Ent. Egypte 43 : 60,

Pl. 5 Fig. 3. Syn. nov.

AMSEL (1959) compared the male genitalia in the original description of

obraztsovi with those of inconspicua Danilevsky, and found a difference in the

shape of the aedeagus. The new figure of the male genitalia of exzmia, published

recently by DANILEVSKY (1960), shows a great similarity of this species and

obraztsovi, and there is no doubt about their identity. As far as a comparison of

the original descriptions permits one to conclude, externally these two species are

also similar. In all probability they are synonymous. The range of eximia comprises

Crimea, Armenia, Daghestan, and Transilian, and Dzungarian Ala-tau; D.

obraztsovi was described from Haj Omran, Iraq.

Dichrorampha (Lipoptycha) gracilis (Danilevsky)

Hemimene (Lipoptycha) gracilis Danilevsky, 1948, Ent. Obozrenie 30: 76 (external

characters, not genitalia; specimen from Kuldja). — 1960, Rev. Ent. URSS 39 : 945, Fig. 14.

nn.

18 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Hemimene (Lipoptycha) eximia (part.): Danilevsky, 1948, ibid. 30: 77, Fig. 13 (geni-

talia, not external characters).

Dichrorampha (Lipoptychodes) eximia : Obraztsov, 1953, Mitt. Münchn. ent. Ges. 43 : 77.

Dichrorampha (Lipoptycha) gracilis (part.): Obraztsov, 1958, Tijdschr. Ent. 101: 253.

Dichrorampha (Lipoptycha) eximia (part.): Obraztsov, 1958, ibid. 101: 254.

Dichrorampha (Dichrorampha) cinerosana (H.S.)

Grapholitha cinerosana Herrich-Schäffer, 1851, Syst. Bearb. Schm. Eur. 4: 255 [1848,

Tortr., Pl. 41 Fig. 290; non bin.].

Dichrorampha (Dichrorampha) cinerosana: Obraztsov, 1958, Tijdschr. Ent. 101: 247

(synonymy).

Dichrorampha eurychorana Rebel, 1941, Kön. Naturw. Inst. Sofia 14: 2, Fig. 1. —

Thurner, 1941, ibid. 14: 26. Syn. nov.

Dichrorampha (?Dichrorampha) eurychorana : Obraztsov, 1953, Mitt. Münchn. ent. Ges.

53: 29. — 1958, Tijdschr. Ent. 101: 247.

Specimens examined. One female (genitalia on slide No. 633-Obr.), Matka,

Treska Schlucht, Macedonia, May 20—29, 1955 (J. KLIMESCH); one female, the

same data but May 19—29, 1955 (AMNH).

Dr. J. KrimescH (Linz a. d. D.) kindly sent me two female specimens from

Macedonia, which he identified as Dichrorampha eurychorana Rebel. A comparison

with the original description and the photograph of this species, confirmed the

correctness of his identification. REBEL compared his eurychorana with Dichro-

rampha montanana (Duponchel) (= alpestrana Herrich-Schäffer) which it really

somewhat resembles. However, the examination of the genitalia of one of the

received specimens proved beyond any doubt that ewrychorana has nothing in

common with that species, D. montanana, but should be regarded as a synonym

of D. cinerosana.

Dichrorampha (Dichrorampha) chavanneana (Laharpe)

Grapholita chavanneana Laharpe, 1858, Nouv. Mém. Soc. Helv. Sci. Nat. 16, Faune

Suisse 6 : 102. — 1864, ibid. 20, Faune Lép. Suisse, Suppl. 3 : 73.

Dichrorampha (Lipoptycha) alpigenana (part.): Heinemann, 1863, Schm. Deutschl.

Schweiz. [2} 1 (1): 236.

Dichrorampha rhaeticana Frey, 1870, Mitth. Schweiz. Ent. Ges. 3: 247. — 1871, Ent.

Zeitg. Stettin 32 : 104. — 1880, Lep. Schweiz : 331. — Rebel ;» Staudinger & Rebel, 1901,

Cat. Lep. pal. Faun. 2 : 128, no. 2293. Syn. nov.

Dichrorampha chavanneana : Wocke in Staudinger & Wocke, 1871, Cat. Lep. eur. Faun.:

263, no. 1278. — Staudinger, 1871, Berlin. Ent. Zeitschr. 14 [1870]: 284.

Dichrorampha harpeana (part.): Rebel in Staudinger & Rebel, 1901, Cat. Lep. pal. Faun.

2: 128, no. 2294.

Hemimene chavanneana : Kennel in Spuler, 1907, Schm. Eur. 2 : 287. — Müller-Rutz in

Vorbrodt & Müller-Rutz, 1914, Schm. Schweiz 2: 411. — Kennel, 1921, Pal. Tortr.: 632,

Pl. 23 Fig. 38, 39.

Hemimene rhaeticana : Kennel, 1921, op. cit.: 631, Pl. 23 Fig. 37. — Miiller-Rutz, 1927,

Mitt. Schweiz. Ent. Ges. 13 : 232. — 1932, ibid. 15 : 237.

Hemimene bugnionana: Müller-Rutz, 1922, Mitt. Schweiz. Ent. Ges. 13 : 232.

Dichrorampha (Lipoptychodes) bugnionana (part.): Obraztsov, 1953, Mitt. Münchn. ent.

Ges. 43 : 60.

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 19

Dichrorampha (Dichrorampha) rhaeticana : Obraztsov, 1958, Tijdschr. Ent. 101: 251. —

Hannemann, Die Wickler #2 Dahl, 1961, Tierw. Deutschl. 48 : 78, Fig. 132, Pl. 7 Fig. 21.

Dichrorampha (Lipoptycha) bugnionana (part.): Obraztsov, 1958, Tijdschr. Ent. 101:

252. — Hannemann, Die Wickler #2 Dahl, 1961, Tierw. Deutschl. 48: 80.

Types. Type of chavanneana : not seen. Lectotype of rhaeticana, male (genitalia

on slide, No. 5752), “Bernina Alpen von Sils” (BMNH).

Dr. W. SAUTER (Zürich) wrote to me in a letter of May 16, 1962, that he had

retrieved the type of chavanneana. The specimen was in the LAHARPE collection

among the specimens of bugnionana Duponchel, to which also montanana Du-

ponchel was added. An examination of the genitalia of the type specimen of

chavanneana, made by Dr. SAUTER, showed that this species is synonymous with

rhaeticana.

The systematic position of chavanneana and the species itself has for a long time

remained obscure. Most of the authors simply quoted this species from its original

description. HEINEMANN (1863) treated chavanneana as a doubtful synonym of

alpigenana Heinemann. Other authors regarded it to be a distinct species, and

placed it now near agilana Tengström and /igulana Herrich-Schäffer (WOCKE,

1871), then near rhaeticana (KENNEL, 1921), or synonymized it with harpeana

Frey (REBEL, 1901). LAHARPE (1863) refused to recognize chavanneana in his

later publication as distinct, and treated it as the female of bugnionana.

OBRAZTSOV (1953; 1958) followed his view.

The present re-discovery of the type specimen of chavanneana by Dr. SAUTER,

makes this specific name available; this name should replace rhaeticana, used as

yet for the species in question. FREY (1871) established his rhaeticana conditional-

ly, and the original description of this species was very poor. It was given as a

remark to the description of harpeana, and read: “Ich glaube noch einen zweiten,

nahe verwandten Wickler in einigen Exemplaren gleichfalls von den Hochalpen

des Engadin zu besitzen, ein grösseres Thier, mit schmäleren, spitzwinkligeren,

brauneren, ganz verloschen gezeichneten Vorderflügeln. Sollten sich in der Folge

die Artrechte feststellen lassen, so schlage ich den Namen D. Rhaeticana vor.”

In a paper published one year later, FREY (1870) word for word repeated the

above “description”. Only ten years later (1880) he described rhaeficana more

circumstantially, and defined the type lot. He wrote: “Ich fing diese auffallende

hochalpine Art in zwei männlichen Stücken auf der Passhöhe des Bernina. Ein

drittes, wiederum männliches Exemplar erhielt ich aus Wallis durch Anderegg.”

One male specimen from Bernina Alps, originating from FREY's collection, came

through the WALSINGHAM collection to the British Museum (Natural History),

and I had a chance to examine it. This specimen confirms the correctness of the

treatment of rhaeticana in the literature. Slight differences in the shape of the

cucullus of this specimen and of those examined by OBRAZTSOV (1953) and

HANNEMANN (1961) depend on individual variation of this structure. The aede-

agus of rhaeticana has a rather constant shape. The figure of the aedeagus published

by HANNEMANN (1961) is rather simplified, and it lacks the lateral shaft braces,

the most characteristic feature of the aedeagus of rhaeticana.

20 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Laspeyresia generosana (Christoph)

Grapholitha generosana Christoph, 1881, Bull. Soc. Imp. Nat. Moscou 56 : 423. — Rebel,

in Staudinger & Rebel, 1901, Catal. Lep. pal. Faun. 2: 122, no. 2198. — Caradja, 1916,

Iris 30: 69.

?Grapholitha lepidulana Snellen, 1883, Tijdschr. Ent. 26: 221, Pl. 13 Fig. 5—5a.

Laspeyresia generosana : Kennel, 1921, Pal. Tortr.: 666, Pl. 24 Fig. 22. — Obraztsov,

1959, Tijdschr. Ent. 102 : 190, 199, Fig. 42.

Types. Holotype of G. generosana, female (genitalia on slide No. 12-Obr.),

Amur (ZMHB). Lectotype of /epzdulana, female (figured by SNELLEN), Irkutsk,

Siberia; not seen. Lectoallotype of /epzdulana, male (without abdomen), the same

locality, June 12 (Hnr.) (RMNH).

The data on the holotype of generosana were published by OBRAZTSOV (1959).

REBEL (1901) treated /epidulana as a synonym of the former species, and later

authors followed him. I had a chance to study a single male specimen of lepidu-

lana, the lectoallotype, originating from SNELLEN’s collection. Unfortunately, this

specimen lacks the abdomen. The presence of an internal lobe on the dorsum and

the strongly sclerotized veins A, and A, are the striking characters of its hind

wings. Considering the possibility of the presence of SNELLEN’s female /epidulana

in the collection of the Zoological Institute of the Academy of Sciences of the

USSR in Leningrad, it seems sensible to treat that female as lectotype; the figure

accompanying the original description pertains to that specimen. The present,

defective male in that case would become a lectoallotype of G. lepidulana Sn.

Until the genitalia of both /epidulana and generosana are compared, their con-

specificity cannot be considered as proven. The presence of a small, fold-shaped

sac up on the corpus bursae of generosana, rostrad from the cervix bursae, shows

rather decidedly that this is a Laspeyresia species, because similar structures have

been observed only in some species of that genus, and never in the genus Grapho-

litha Treitschke. |

Laspeyresia refrigescens Meyrick

Laspeyresia refrigescens Meyrick, 1924, Exot. Microlep. 3: 68. — Clarke, 1955, Catal.

Meyrick’s Types 1: 269. — 1958, op. cit. 3: 455, Pl. 226 Fig. 3, 3a. — Obraztsov, 1959,

Tijdschr. Ent. 102: 186.

Type. Lectotype, male (genitalia on slide No. 9408, J.F.G.C.), Aswan, Egypt,

1920 (K. J. HaywArp) (BMNH).

In this species, a tuft of hairs in the neck area of the valvae is present, and the

aedeagus has a short dorsal process. In view of these features, it should be placed

near L. arabica Amsel and L. gilviciliana Staudinger.

Laspeyresia interruptana (Herrich-Schäffer)

Tortrix (Grapholitha) interruptana Herrich-Schiffer, 1851, Syst. Bearb. Schm. Eur. 4:

269 (the figure was published uninominally in the same work: 1848, Tortr.: Pl. 29 Fig. 280).

Laspeyresia interruptana : Obraztsov, 1959, Tijdschr. Ent. 102: 191 (literature and

synonymy).

Laspeyresia indivisa Danilevsky, 1963, Rev. Ent. URSS 42: 172, Fig. 10, 11. Syn. nov.

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 21

Recently DANILEVSKY (1963) described the above-mentioned new species

indivisa and wrote as follows: “Obraztsov (1952) erroneously identified the

described species as interruptana H.-Sch. Owing to the kindness of Dr. Hanne-

mann, I had the chance to examine a specimen of interruptana from the collection

of Herrich-Schäffer, preserved in the Zoological Museum of Humboldt University

in Berlin. The specimen has the following labels: 1) “interruptana Khlw.” (in

the handwriting of Herrich-Schäffer?); 2) “Origin”; 3) “Alten”; 4) “Genit.

Unters. No. 1336 Zool. Mus. Berlin.” The genitalia slide-label (balsam) reads:

“No. 1336, Alsen, Coll. Stgr., Typus’. Thus, this specimen must be regarded

as the lectotype of Grapholita interruptana H.-Sch.”

The geographic reference books available to me name two localities in Europe,

known as “Alten”. One of them is in Anhalt (Germany), the other in Northern

Norway. It is therefore difficult to say with certainty which of both might be

implied with the label of the specimens examined by DANILEvsky. The name

“Alsen” written on the slide is probably a misspelling for Alten, although there

is the Island Als in Denmark, known in Germany as Alsen. All of these localities

cannot refer to the original description of interruptana, because Herrich-Schäffer

expressly wrote that this species was known to him only from Bohemia and

Regensburg. In compliance with the above the specimen examined by DANILEVSKY

cannot be a lectotype of znterruptana. Moreover, DANILEVSKY writes that this

species “does not externally differ from duplicana Zett.”. Quite the reverse, the

species figured by Herrich-Schäffer as interruptana has no likeness of duplicana

at all; on the contrary, it matches well with the specimens from Lage (Lippe)

identified by OBRAZTSOV (1952) as interruptana. Until this latter interpretation

of interruptana in the sense of OBRAZTSOV is disproved by the examination of

some authentical material, there is no reason to introduce any new name. Thus,

indivisa Danilevsky should be treated as a synonym of znterruptana Herrich-

Schäffer.

Grapholitha (Grapholitha) orobana Treitschke

Bag

Grapholitha orobana Treitschke, 1830, Schm. Eur. 8: 226.

For further synonymy and literature cf. Obraztsov, 1959, Tijdschr. Ent. 102: 210.

Laspeyresia persicana Osthelder, 1938, Mitt. Münchn. ent. Ges. 28: 28. — Obraztsov,

1959, Tijdschr. Ent. 102: 194. Syn. nov.

Type of persicana. Holotype, male (genitalia on slide No. 1-Obr. 8/29 1958),

Hecarcal-Tal, 28.000— 32.000 ft, Tacht-i-Suleiman, northern section of the Elburz

Mountains, Northern Persia, July 2—7, 1937 (E. PFEIFFER) (ZSBS).

Laspeyresia persicana, described as a distinct species related to dorsana and

orobana, is conspecific with orobana, as is evident from the genitalia. The holotype

of persicana represents an insignificant variety, in which the white dorsal patch is

somewhat modified, but similar specimens are also known from Europe.

Grapholitha (Grapholitha) fimana Snellen

Grapholitha fimana Snellen, 1883, Tijdschr. Ent. 26: 225, Pl. 13 Fig. 7, 7a. — Rebel in

Staudinger & Rebel, 1900, Catal. Lep. pal. Faun. 2: 122, no. 2203.

TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Laspeyresia fimana: Walsingham, 1900, Ann. Mag. Nat. Hist. [7] 6: 432. — Kennel,

1921, Pal. Tortr.: 677, Pl. 24 Fig. 49. — ? Inoue, 1954, Check list Lep. Japan : 92, no. 488.

Grapholitha (Grapholitha) fimana: Obraztsov, 1959, Tijdschr. Ent. 102: 209.

Grapholitha (Grapholitha) timana (misprint, lapsus): Snellen, 1883, Tijdschr. Ent. 26:

183.

Type. Lectotype, male (glued specimen), Island Askold, 1878 (“Eil. Askold

1878 9 D. v. Hed. 134”) (RMNH).

The lectotype is a damaged specimen glued from male and female parts. The

basic portion is of a male, with a right hindwing and a left forewing; the left

hindwing and the abdomen are of a female.

Other specimens examined. One male and one female (genitalia on slides Nos.

6653, 6654), the same data but “1880 (M. JANKOWSK1)” (BMNH).

Male genitalia. Valva with a short, very narrow neck in middle; cucullus ovate.

Aedeagus thick, with one long, strong cornutus, and many fine cornuti, half as

long as the former.

Female genitalia. Lamella antevaginalis narrow, crescentic, with angles directed

caudad; lamella postvaginalis at least four times as broad as lamella antevaginalis,

with a narrow, angulate protuberance directed cephalad. Ostium bursae closer to

posterior edge of seventh sternite. Ductus bursae moderately long, broad, nar-

rowed caudad, with some scobinations cephalad. Cervix bursae as broad as adjacent

portion of ductus bursae.

The authentic records of fimana are only from the Askold Island, but the

species may also be distributed over other localities of south-eastern Siberia. The

specimen from Japan, identified as fimana by WALSINGHAM (1900) has appeared

to be Grapholitha jesonica (Matsumura), discussed in this paper. Until any new

and verified data are present, the records of G. fimana in Hokkaido and Honshu

(INOUE, 1954) cannot be taken for granted.

Grapholitha (Grapholitha) jesonica (Matsumura) comb. nov.

Laspeyresia fimana (part.): Walsingham, 1900, Ann. Mag. Nat. Hist. [7] 6: 432. —

Inoue, 1954, Check list Lep. Japan 1: 92, no. 488.

Laspeyresia jesonica Matsumura, 1931, 6000 Illustr. Ins. Japan-Empire: 1072, Fig. —

Inoue, 1954, Check list Lep. Japan 1 : 92, no. 493. — Obraztsov, 1959, Tijdschr. Ent. 102:

192. — Okano & others, 1959, Iconogr. Ins. japon. color. nat. edita 1: 259, Pl. 174 Fig. 14.

Specimens examined. One female (genitalia on slide No. 6656), Japan, 1886

(PRYER: No. 70125); one female (genitalia on slide No. 6655), Sapporo, Japan,

June 8, 1918 (S. Issıkı) (BMNH).

The type of jesonica remains unknown to me, but after comparing its original

description and the published figure I believe that the above-mentioned two

female specimens in the British Museum belong to this species. One of these is

that misidentified by WALSINGHAM (1900) as fimana Snellen. G. jesonica and

fimana are very similar, but can nevertheless be discriminated even externally.

In G. jesonica the tips of the tegulae are white, and the elongate, oblique spot

on the dorsum of the forewings is clear white. In G. fimana the tegulae are

entirely black, and the mentioned spot is darkly suffused, longer, and less curved

than in jesonica. The costa of the forewings in jesonica is almost concolorous with

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 23

the remaining wing surface, and in the basal half of the forewing there are two

short, white supradorsal streaks. In fzmana, the costa is slightly tinged with white,

and supradorsal streaks are lacking. The ocelloid patch of jesontca is more distinct,

and includes four black, longitudinal streaks; in fzmana there are only two or

three minute, black dots within that patch. In jesonica some rather long, white,

oblique costal streaks of the forewings are sharply defined and arranged in pairs:

there are six of these pairs in the external two-thirds of the costa. In fzmana, the

streaks are very short; only the two pre-apical form a distinct pair; the remaining

streaks rather alternate with black interspaces, not forming pairs.

For the present, only the female genitalia in these two species could be com-

paired; they proved to be quite distinct.

Female genitalia. Lamella antevaginalis elongate-rectangular, with latero-

posterior angles narrowly produced; lamella postvaginalis rotundate-trapezoidal,

with posterior edge deeply concave. Ostium bursae about at middle of seventh

sternite. Ductus bursae short and broad. Cervix bursae bearing light, lateral

scobination, and developed as a somewhat narrowed caudal portion of corpus

bursae.

Grapholitha (Grapholitha) dimidiatana Kennel

Grapholitha dimidiatana Kennel, 1901, Iris 13 [1900]: 300 (cf. Obraztsov, 1959, Tijdschr.

Ent. 102: 211, 214, for synonymy and literature).

Laspeyresia exosticha Meyrick, 1936, Exot. Microlep. 5: 25. — Clarke, 1955, Catal.

Meyrick's Types 1: 138. — Wiltshire, 1939, Trans. Roy. ent. Soc. London 88: 55, PI. 1

Fig. 22. — Obraztsov, 1959, Tijdschr. Ent. 102: 193. Syn. nov.

Types. Lectotype of dimidiatana : male (genitalia on slide No. 23-Obr.), Al-

gezares, Murcia, Spain, 1894 (Kors) (ZMHB). Lectotype of bzpartana Kennel:

female (genitalia on slide No. 22-Obr.), same data (ZMHB). Holotype of intacta

Walsingham: female (genitalia on slide No. 4945), Tetuan, Morocco, April 26,

1902 (WALSINGHAM Coll. No. 87346) (BMNH). Lectotype of exosticha Meyrick:

male, Bhamdoun, Lebanon, April; location unknown.

Specimen examined. One male (genitalia on slide No. 6665), Luize, Lebanon,

April 16, 1934 (E. P. WiLTsHIRE) (BMNH) (identified by E. MEYRICK as

exosticha).

MEYRICK (1936) described exosticha from two specimens (male and female),

taken at Bhamdoun and Luize, Lebanon, in April. WILTSHIRE (1939) confirmed

that only two specimens of this species were taken, and figured a female from

Luize, indicating it as a paratype. Thus, the male from Bhamdoun must be regarded

as the lectotype (MEYRICK did not indicate a holotype), but its present location

is unknown. The male specimen examined for this paper has been identified by

MEYRICK as exosticha; it does not belong to the type lot, although it was probably

taken together with the female paratype figured by WILTSHIRE. This male cor-

responds well with the description and figure of exosticha, and without doubt

belongs to this species. The genitalia and the wing markings are identical with

those of dimidiatana, and this gives the reason to treat exosticha as a synonym of

this species. In the British Museum there is one more female specimen identified

24 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

as exosticha and originating from South Ussuri (genitalia on slide No. 6666). As

it might be supposed in advance, this specimen has nothing in common with

exosticha and, judging from the genitalia, is Pammene orientata Kuznetzov.

Grapholitha (Euspila) callisphena (Meyrick) comb. nov.

Laspeyresia callisphena Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18: 144. —

Clarke, 1955, Catal. Meyrick’s Types 1: 71. — Obraztsov, 1959, Tijdschr. Ent. 102: 192.

Grapholitha (Euspila) critica (Meyrick) comb. nov.

Eucelis critica Meyrick, 1905, Journ. Bombay Nat. Hist. Soc. 16: 587. — Clarke, 1955,

Catal. Meyrick’s Types 1: 103.

Eucosma ludicra Meyrick, 1912, Journ. Bombay Nat. Hist. Soc. 21 : 867. — Clarke, 1955,

Catal. Meyrick’s Types 1: 192.

Eucosma pseudomorpha Meyrick, 1916, Exot. Microlep. 2: 19. — Clarke, 1955, Catal.

Meyrick’s Types 1: 263.

Eucosma trichocrossa Meyrick, 1916, Exot. Microlep. 1: 563. — 1916, op. cit. 2: 19. —

Clarke, 1955, Catal. Meyrick’s Types 1: 315.

Eucosma critica: Clarke, 1958, ibid. 3: 359, Pl. 178 Fig. 1, 1a—b, 2, 2a, 3, 3a, 4, and

4a—c,

Grapholitha (Euspila) conciliata (Meyrick) comb. nov.

Eucosma conciliata Meyrick, 1920, Exot. Microlep. 2: 245. — Clarke, 1955, Catal. Mey-

rick’s Types 1: 97. — 1958, op. cit. 3: 356, Pl. 177 Fig. 1, 1a—b, 2, and 2a.

Laspeyresia heteropa Meyrick, 1920, Exot. Microlep. 2: 351. — Clarke, 1955, Catal.

Meyrick’s Types 1: 158.

Eucosma heteropa: Clarke, 1958, op. cit. 3: 356 (synonym of conciliata).

Grapholitha (Euspila) chelias (Meyrick) comb. nov.

Laspeyresia chelias Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18: 145. — ?1911,

Ann. Transvaal Mus. 2: 229. — ?Janse, 1917, Check list S. Afr. Lep. Heter.: 176. —

Clarke, 1955, Catal. Meyrick’s Types 1: 83. — 1958, loc. cit. 3: 432, Pl. 215 Fig. 4—4c.

Type. Holotype, female (genitalia on slide No. 7378, J.F.G.C.), Maskeliya,

Ceylon, October, 1904 (POLE) (BMNH).

Judging from the structure of the female genitalia (the males of chelias are not

yet known), and the ocelloid patch of the forewings, this species should be

transferred to the subgenus Euspila Hübner of the genus Grapholitha. MEYRICK

(1911) claimed to have chelias from Pretoria, explaining the presence of this

Oriental species in South Africa by a possible artificial introduction. Until the

genitalia of South African specimens are examined, this record remains doubtful.

Grapholitha (Euspila) isacma (Meyrick) comb. nov.

Laspeyresia isacma Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18: 144. — Clarke,

1955, Catal. Meyrick’s Types 1: 176.

Laspeyresia delineana (part.): Meyrick (nec Walker), 1908, Proc. Zool. Soc. London :

721. — 1931, Exot. Microlep. 4: 143. — Clarke, 1958, Catal. Meyrick’s Types 3.: 435,

Pl. 216 Fig. 3—3a.

Grapholitha (Euspila) ?delineana (part.): Obraztsov, 1959, Tijdschr. Ent. 102: 211.

Grapholitha isacma : Bradley, 1961, Bull. Brit. Mus., Ent. 10: 128.

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 25

Type. Lectotype, male (genitalia on slide No. 7384, J.F.G.C.), Khasi Hills,

Assam, September, 1906 (BMNH).

MEYRICK originally described G. zsacma as a distinct species, but later suppres-

sed this name as a synonym of delineana Walker. BRADLEY (1961) compared the

types of these two species, and found them distinct. In G. isacma the sacculus of

the valva is long, with the external angle rotundate-rectangular; in delineana there

is no angle on the end of the sacculus, and the ventral edge of the valva is curved

inwards before the cucullus. This latter is much narrower in #sacma than in

delineana. The “basal opening” of the valva in zsacma is very long. Moreover, the

aedeagus in this species is S-curved, has a rather narrow coecum penis, and a thin,

acute tip; the cornuti are about one-third as long as the aedeagus. In delineana

the aedeagus is straight, somewhat bottle-shaped, widely dilated toward the coecum

penis, and has a blunt, widely opened distal end; the cornuti are half as long as

the aedeagus.

From G. quadristriana Walsingham which was erroneously treated as a synonym

of delineana, G. isacma differs in most of the above-mentioned characters, only

the sacculus in both isacma and quadristriana is angulate externally. But in zsacma

the sacculus is considerably longer, and its external angle is distinctly blunter.

The male genitalia of the type of G. isacma are figured by CLARKE (1958).

Grapholitha (Euspila) quadristriana (Walsingham)

Laspeyresia quadristriana Walsingham, 1900, Ann. Mag. Nat. Hist. [7] 6: 432. —

Matsumura, 1931, 6000 Illustr. Ins. Japan-Empire: 1073, Fig.

Grapholitha quadristriana : Esaki & others, 1932, Iconogr. Ins. Japan: 1465, Fig.

Grapholitha delineana : Issiki & others, 1957, Icones Heteroc. japon. color. nat. 1: 54,

Pl. 8 Fig. 244. — Okano & others, 1959, Iconogr. Ins. japon. color. nat. edita 1: 259,

Ri 74 Kigali.

Grapholitha (Euspila) delineana (part.): Obraztsov, 1959, Tijdschr. Ent. 102: 211.

Types. Lectotype, male (genitalia on slide No. 29), Japan, 1886 (PRYER; WAL-

SINGHAM Coll. No. 70112); lectoallotype: female (genitalia on slide No. 28),

same data (WALSINGHAM Coll. No. 70113).

The genitalia of both type specimens are badly prepared, but they give an idea

about the difference of this species and delineana.

Grapholitha (Euspila) delineana Walker

Grapholitha delineana Walker, 1863, List Specim. Lepid. Ins. 28 : 389.

Grapholitha apicatana Walker, 1863, op. cit. 28: 390. Syn. nov.

Laspeyresia delineana : Caradja, 1932, Bull. Sect. Sci. Acad. Roum. 15 (7—8): 24.

Grapholitha (Euspila) delineana : Obraztsov, 1959, Tijdschr. Ent. 102: 211.

Laspeyresia apicatana : Obraztsov, 1959, ibid. 102: 192.

Types. Lectotype of G. delineana: male (abdomen missing), China; lecto-

paratype: male (genitalia on slide No. 25), China; holotype of apicatana : female

(abdomen missing), China (BMNH).

The holotype of G. apicatana is badly damaged, but its conspecificity with

“Laspeyresia” delineana is quite certain. One male specimen in the British Museum,

26 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Mt. Mlanje, Nyassaland, December 22, 1913 (S. A. NEAVE), identified by MEy-

RICK as delineana, undoubtedly belongs to some other, probably new, species. The

records of delineana from Assam should be referred to G. isacma Meyrick. Those

from New Guinea (DIAKONOFF, 1953, Verh. Ned. Ak. Wet., Nat. [2} 49 (3):

162) need to be verified.

Grapholitha (Euspila) endrosias (Meyrick)

Laspeyresia endrosias Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18 : 145. — Clarke,

1955, Catal. Meyrick’s Types 1: 126. — 1958, op. cit. 3 : 436, Pl. 217 Fig. 3—3a.

Euspila endrosias : Issiki & others, 1957, Icones Heteroc. japon. 1: 54, Pl. 8 Fig. 243.

Grapholitha (Euspila) endrosias : Obraztsov, 1959, Tijdschr. Ent. 102: 211.

Type. Lectotype, male (genitalia on slide No. 7382, J.F.G.C.), Khasi Hills,

Assam, June, 1960 (BMNH).

An examination of the type has confirmed the systematic position of this species.

No specimens from Japan could be studied, but the color photograph published

by Issıkı (1957) fits the type of endrosias well.

Grapholitha (Euspila) rosinana (Kennel) comb. nov.

Laspeyresia rosinana Kennel, 1919, Mitt. Münchn. ent. Ges. 8 [1918]: 88, Pl. 3 Fig. 22. —

Obraztsov, 1959, Tijdschr. Ent. 102 : 195.

Type. Holotype, male (abdomen missing), Sayan Mountains (ZMHB).

The general appearance of the holotype and the markings of the forewings

speak in favour of recognition in rosinana of a species of the above genus and

subgenus. In the catalogue this species might temporarily be placed after G. mun-

dana Christoph which is a Exspzla species with a similar multistrigate dorsal patch

in the forewing.

Pammene blockiana (Herrich-Schäffer)

Textfig. 1—2, Pl. 2 Fig. 1

?Tinea blockiella Rossi, 1794, Mant. Ins. 2: 46. — Werneburg, 1864, Beitr. Schm.kunde

2: 253 (nomen oblitum).

Tortrix (Grapholitha) blockiana Herrich-Schäffer, 1851, Syst. Bearb. Schm. Eur. 4 : 256.

— 1863, Corresp.-Blatt zool.-min. Ver. Regensburg 17: 123.

Tortrix (Semasia) blockiana : Herrich-Schäffer, 1853, Syst. Bearb. Schm. Eur. 6, Syst.

Lepid. Eur.: 45.

Phthoroblastis blockiana : Lederer, 1859, Wien. Ent. Monatschr. 3: 372. — Wocke in

Staudinger & Wocke, 1871, Cat. Lep. eur. Faun.: 260, no. 1206.

Grapholitha (Strophosona) blockiana : Heinemann, 1863, Schm. Deutsch. Schweiz [2] 1

(1): 204.

Pamene blockiana : Rebel in Staudinger & Rebel, 1901, Cat. Lep. pal. Faun. 2: 125, no.

2248. — ?Caradja, 1916, Iris 30: 71.

Pammene blockiana : Kennel, 1921, Pal. Tortr.: 707. — Obraztsov, 1960, Tijdschr. Ent.

103 : 116, 122. — Hannemann, Die Wickler #2 Dahl, 1961, Tierw. Deutschl. 48: 106,

nota, Pl. 22 Fig. 16.

Pammene cupressana Zerny, 1927, Verh. zool.-bot. Ges. Wien 77: (4).

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 247

Fig. 1. Pammene blockiana (Herrich-Schäffer), male, lectotype of P. cupressana Zerny,

genitalia on slide no. V.41

Types. Lectotype of blockiana, female (genitalia on slide no. Mn. 1, made by

V. I. KUZNETZOV), “Tinea Blockiella. Rossi” (ZMHB). Lectotype of cupressana,

male (genitalia on slide, no. V.41), Gargnano, Lake Garda, Italy, July 20, 1913

(M. ZERNY) (NHMW).

Other specimens examined. Two females (allolectotype of cupressana genitalia

on slide no. V.42, and paralectotype), Gargnano, Lake Garda, Italy, July 20, 1913

(M. ZERNY) (NHMW). One female (genitalia on slide no. 13-Obr., 1961),

Treska Gorge, westwards from Skoplje, Macedonia, May 20—30, 1956 (F. Kasy)

(ZMHB).

HERRICH-SCHÄFFER (1851) described blockiana from two specimens in bad

condition, without any record of their origin. One of these specimens is the lecto-

type selected in the present paper; the other specimen is probably missing. The

lectotype consists of a piece of thorax with the head and left wings. The latter are

badly damaged and glued to the thorax. The genitalia slide is also very poor, and

merely includes the bursa copulatrix. The label reads: “Tinea Blockiella. Rossi”.

HERRICH-SCHAFFER apparently believed that no species was published under this

name and decided to publish it as new, changing the ending “-ella” in “-ana”,

in compliance with the tradition of his time. In fact, Rossi (1794) described a

species from Etruria in northern Italy and named it “Tinea blockiella”. This

species remained mysterious to later generations of lepidopterists, but comparing

the original description of bockiella with blockiana is suggestive of the fact that

both these names concern one and the same species. It would nevertheless be un-

wary to restore the priority of the name blockiella on the only basis of such a

suggestion, and I consider it more advisable to treat this name as a “nomen

oblitum”.

Rossi named his blockiella “in honorem Cel. Viri lib. Bar. Block, Dresdensis”.

28 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Fig. 2. Pammene blockiana (Herrich-Schäffer), female, allolectotype of P. cupressana Zerny,

genitalia on slide no. V.42

There is no doubt that also blockiana was dedicated to the same Baron BLOCK in

Dresden. Probably for this reason the later authors associated this name in some

way with Germany which has always been quoted in the literature as the only

locality of blockzana. No authentical records of this species have ever been made

from Germany. On the contrary, this species was found in northern Italy (Lom-

bardy), and re-described as cupressana Zerny. A new record of blockiana in

Macedonia, published for the first time in the present paper, confirms that this

species is of southern origin. The male genitalia of blockiana are very similar to

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 29

those of Pammene crataegophila Amsel from the eastern Mediterranean. The

external angle of the sacculus is more rotundate in blockiana than in the latter

species, and the cucullus is longer. The anal angle of the cucullus is more extended,

the neck of the valva is somewhat broader than in crataegophila, and the setae

in this area are less numerous. The distal portion of the aedeagus is distinctly

broader, and four long, with five minute cornuti, are present. In crataegophila

the cornuti also are of an unequal length, but the long cornuti are distinctly longer

than those in blockiana.

Male genitalia. Valva with an incurved costa and a rather broad neck; sacculus

slightly longer than one-third of entire valva, and ending with a broadly rotundate

projection; a deep, rotundate incision of ventral edge of valva between this

projection and cucullus; latter rather long, directed obliquely upwards, almost

equally broad along its length, with an anal angle distinctly projected and rounded,

and an apical angle broadly rotundate; few rather short setae on sacculus and in

neck area of valva; a well developed corona of moderately long thorns; long setae

on cucullus. Aedeagus rather thick, slightly narrowed distally; four long and five

minute cornuti.

Female genitalia. Ventral plate of seventh abdominal sternite moderate, horizon-

tally crescent. Lamella antevaginalis moderately broad; lamella postvaginalis rather

short. Antrum weakly sclerotized, tubular; signa short, with basal discs rather

broad. Apophyses posteriores much shorter than apophyses anteriores.

Leguminivora glycinivorella (Matsumura)

Pee nice 2

Grapholitha glycinivorella Matsumura, 1900, Ent. Nachr. 26: 197. — 1901, Illustr.

Zeitschr. Ent. 6: 23. — Esaki & others, 1932, Iconogr. Ins. Jap., Ist ed.: 1465, Fig. —

Uchida, 1949, Illustr. pocket book Jap. ins.: 302, Fig. — Esaki & others, 1950, Iconogr.

Ins. Japan, 2nd ed.: 479, Fig. — Inoue, 1954, Check list Lep. Jap.: 93. — Mutuura,

1953, Publ. Ent. Agr. Naniwa Univ. 1: 12, Pl. 4 Fig. 1, Pl. 5 Fig. 4. — Issiki & others,

1957, Icon. Heteroc. japon. color. nat. 1: 56, Pl. 8 Fig. 248.

Laspeyresia parastrepta Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18: 147. —

Janse, 1917, Check list S. Afr. Lep. Heter.: 176. — Meyrick, 1936, Iris 50 : 157. — Clarke,

1955, Catal. Meyrick's Types 1: 235. — 1958, op. cit. 3: 448, Pl. 223 Fig. 2—2a. —

Obraztsov, 1959, Tijdschr. Ent. 102: 194. Syn. nov.

Laspeyresia anticipans Meyrick, 1927, Exot. Microlep. 3: 342. — Clarke, 1955, op. cit.

1: 48. — 1958, op. cit. 3: 432, Pl. 215 Fig. 1—1c. Syn. nov.

Laspeyresia glycinivorella: Kuwayama, 1928, Journ. Coll. Agr. Sapporo 19: 261, Fig. —

Matsumura, 1931, 6000 Illustr. Ins. Japan-Empire : 1072, Fig. — Kurentsov, 1950, Soobshtch.

Dalnevost. Fil. Akad. Nauk : 32.

Laspeyresia zygogramma Meyrick in Caradja & Meyrick, 1935, Mater. Faun. chin. Prov.:

63. — Clarke, 1955, op. cit. 1: 332. — Obraztsov, 1959, Tijdschr. Ent. 102 : 195. Syn. nov.

?Enarmonia parastrepta : Ghesquiére, 1940, Ann. Mus. Congo Belge (C) (3) 7 (1): 106.

Leguminivora glycinivorella : Obraztsov, 1960, Tijdschr. Ent. 103 : 131, Fig. 81—85.

Types. I did not see the type of g/ycinivorella. Lectotype of parastrepta, male

(genitalia on slide, no. 7379, J.F.G.C.), Khasi Hills, Assam, September, 1906

(BMNH). Holotype of anticipans, female (genitalia on slide, no. 7404, J.F.G.C.),

Pollachi, Coimbatore district, S. India, April 30, 1923 (“bred in August from

30 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

larva webbing flowers of mango”, MENON) (BMNH). Holotype of zygogramma,

Lungtan, China, September 27 (CARADJA Collection; not seen).

Other specimens examined. One male (genitalia on slide, no. 2, 2/4 1958-

Obr.), Taikyu, Korea; one female (genitalia on slide, no. 1, 5/3 1958-Obr.),

Kotoni, Sapporo, Japan; both of the above specimens in the United States National

Museum. One male (genitalia on slide, no. 6662), Nanking, China, September 25,

1933 (HONE); identified as zygogramma (det. E. MEYRICK) (BMNH).

The identity of L. g/ycinivorella Matsumura has been established with the data

from literature. The types of the synonyms were also studied.

This species is known from Japan, Korea, China, southeastern Siberia, Assam,

and S. India. The records from Congo and S. Africa (‘‘parastrepta’: JANSE, 1917;

GHESQUIERE, 1940) are rather doubtful.

Leguminivora ptychora (Meyrick) comb. nov.

Laspeyresia ptychora Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18: 147. — 1911,

ibid. 22 : 288. — 1911, Proc. Linn. Soc. N. S. Wales 36 : 288. — 1916, Exot. Microlep. 1:

565. — ?Janse, 1917, Check list S. Afr. Lep. Heter.: 176. — Fletcher, 1920, Mem. Dept.

Agr. India, Ent. 6: 63. — Clarke, 1955, Catal. Meyrick’s Types 1: 264. — 1958, op. cit.

3: 452, Pl. 225 Fig. 1—1a.

?Enarmonia piychora : Ghesquière, 1940, Ann. Mus. Congo Belge (C) (3) 7 (1): 106.

Enarmonia pseudonectis (part.): Diakonoff, 1949, Bijdr. Dierk. 28: 138.

Lectotype. Male (genitalia on slide, No. 7377, J.F.G.C.), Dibidi, North Coorg,

August 10, 1906 (NEWCOME) (BMNH).

Differs from glycinivorella in having a shorter valva, with the cucullus less

distinctly separated, and a somewhat longer aedeagus. This is a distinct species

and not a synonym of L. pseudonectis Meyrick; the latter is synonymous with

L. tricentra Meyrick (CLARKE, 1958). The records of L. ptychora from S. Africa

and Congo (JANSE, 1917; GHESQUIÈRE, 1940) need to be confirmed.

Leguminivora tricentra (Meyrick) comb. nov.

Laspeyresia tricentra Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 17: 734. — ? 1908,

Proc. Zool. Soc. Lond. 1908 : 721. — 1929, Ann. Soc. Ent. France 98 : 721. — 1935, Iris

50 : 157. — Clarke, 1955, Catal. Meyrick’s Types 1: 315. — 1958, op. cit. 3 : 460, Pl. 229

Fig. 1—3a.

Laspeyresia crocopa Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18: 146. — ?Janse,

1917, Check list S. Afr. Lepid. Heter.: 176. — Clarke, 1955, Catal. Meyrick's Types 1: 104.

Laspeyresia pseudonectis Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 18: 146. —

Fletcher, 1920, Mem. Dept. Agr. India, Ent. 6 : 66, PI. 15. — Clarke, 1955, Catal. Meyrick's

Types! 1 :7263.

Enarmonia pseudonectis (part): Diakonoff, 1949, Bijdr. Dierk. 28: 138. — Teotia &

Pathax, 1957, Ann. Zool., Agra 2: 65—25, 11 Fig.

Types. Lectotype of tricentra, male (genitalia on slide no. 7415, J.F.G.C.),

Maskeliya, Ceylon, April, 1906 (J. PoLE). Holotype of crocopa, male (genitalia

on slide no. 7376, J.F.G.C.), Galboda, Ceylon, November, 1901 (J. POLE).

Holotype of pseudonectis, male (without abdomen), Surat, Bombay, October 27,

1904 (H. MAXWELL-LEFROY). (All in BMNH).

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 31

Other specimen examined: one male (genitalia on slide no. 7383, J.F.G.C.),

Pusa, Bihar (BMNH).

I am satisfied to be able to confirm the above synonymy, established by CLARKE

(1958). This species is distinct from the remaining known species of Legumini-

vora in having a long, narrow, serrate process, originating from the anellus above

the basis of the coecum penis. The presence of this process would hardly justify

a subgeneric separation of L. fricentra, until the female genitalia of this species

are studied. The material of /rzcentra in the British Museum is limited to specimens

from India and Ceylon. The record from China (MEYRICK, 1935) could not be

verified, but seems to be probable. That from S. Africa (JANSE, 1917) is rather

doubtful.

Genus Articolla Meyrick

Articolla Meyrick, 1907, Journ. Bombay Nat. Hist. Soc. 17 : 976. — 1909, ibid. 19 : 590.

— Fletcher, 1929, Mem. Dept. Agr. India, Ent. 11: 25. — Turner, 1946, Trans. R. Soc. S.

Austral. 70 : 218. — Meyrick, 1911, Proc. Linn. Soc. N. S. Wales 36 : 285. — Diakonoff,

1952, Zool. Meded. 31: 169. — 1953, Verh. Ned. Ak. Wet., Nat. [2] 49 (3): 88. —

Clarke, 1958, Catal. Meyrick’s Types 3: 303.

Type-species: Articolla cyclidias Meyrick, 1907.

There is no doubt about the close relationship of the present genus and Mafsu-

muraeses Issiki. The distinction is formed by vein Rs in the forewing of Articolla

being approximated to R,, R, and R, shortly stalked, the upper internal vein

of discal cell being absent, and the vein Cu, originating more distad, closer to the

end of the discal cell. In the hind wings of Articolla the veins R and M,, and

Ms and Cu,, are stalked, and the vein Cu, is slightly more approximated to the

end of discal cell, while in Matsumuraeses the veins Rand M,, and M; and Cu,,

are distinctly separated, although they are closely approximated to each other at the

origin, especially the latter pair of the mentioned veins. The male genitalia are

very similar in both genera, and only the presence of a thorn-like spine on the

cucullus of the valva separates Articolla from Matsumuraeses. The female genitalia

of Articolla cyclidias are not yet studied. Those of A. prospera show some sclero-

tization of the antrum and the cervix bursae. The fact of prospera and cyclidias

being congeneric cannot be hold as proven, until the female genitalia of the latter

species (or the male genitalia of the former) are examined 1). The systematic

position of two more species (myriolychna Turner and scioessa Turner), described

as members of the genus Articolla, is merely provisional, and needs verification.

Genus Mastumuraeses Issiki

Semasia (not Stephens): Matsumura, 1900, Ent. Nachr. 26 : 197. — 1901, Illustr. Zeitschr.

Ent. 6: 23. — Kennel, 1900, Iris 13: 14J. — 1916, Pal. Tortr.: 513. — Yasuda, 1956,

Publ. Ent. Agr. Univ. Osaka 2: 15, 16.

Eucelis (part.): Walsingham, 1900, Ann. Mag. Nat. Hist. [7} 6: 407. — Yasuda, 1956,

Publ. Ent. Agr. Univ. Osaka 2: 16, 17.

Eucosma (part.): Meyrick, 1908, Journ. Bombay Nat. Hist. Soc. 18: 613. — 1916, Exot.

1) The species prospera Meyrick has recently been separated in a new genus, Artiphanes

Diakonoff, 1966, Zool. Meded. 85: 65. — Ed.

32 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Microlep. 2: 17. — Fletcher, 1920, Mem. Dept. Agr. India, Ent. 6: 49. — Meyrick, 1922,

Exot. Microlep. 2: 515. — 1935, in Caradja & Meyrick, Mater. Faun. chin. Prov.: 55. —

Diakonoff, 1949, Bijdr. Dierk. 28 : 136. — Viette, 1952, Bull. Soc. Ent. France 57: 151. —

Clarke, 1955, Catal. Meyrick’s Types 1: 199, 204, 319, — 1958, op. cit. 3 : 372. — Yasuda,

1956, Publ. Ent. Agr. Univ. Osaka 2: 16, 17.

Argyroploce (part.): Janse, 1917, Check list S. Afr. Lepid. Heter.: 175. — Meyrick, 1927,

Exot. Microlep. 3 : 340. — Caradja, 1939, Iris 53: 12. — Clarke, 1955, Catal. Meyrick’s

Types 1: 162.

Thiodia (not Hübner): Matsumura, 1931, 6000 Illustr. Ins. Japan-Empire: 1076. —

Obraztsov, 1952, Ent. News 63 : 148.

Endothenia (not Stephens): Meyrick, 1936, Exot. Microlep. 5: 24.

Lathronympha (not Meyrick): Issiki, 1950, Iconogr. Ins. Japan, 2nd ed.: 480. — Inoue,

1954, Check list Lep. Jap.: 95. — Mutuura, 1953, Publ. Ent. Agr. Naniwa Univ. 1: 1, 2,

12. — Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 15217:

Laspeyresia (part.): Inoue, 1954, Check list Lep. Jape ot

Matsumuraeses : Issiki & others, 1957, Icones Heteroc. japon. color. nat. 1: 57, —

Obraztsov, 1960, Tijdschr. Ent. 103 : 134.

Olethreutes (part.): Clarke, 1958, Catal. Meyrick’s Types 3: 519, 559.

Allohermenias (nec Diakonoff): Razowski, 1960, Polsk. Pismo Ent. 30 : 385.

Type-species, Semasia phaseoli Matsumura, 1900.

The author is satisfied to be able to publish a few additional data on the genus

Matsumuraeses and to establish new synonymies. In the shape of the valvae this

genus might remind one of some species treated in the recent literature as

belonging to the genera Hermenias Meyrick, 1911 and Allohermenias Diako-

noff, 1953. These two genera are, however, typical members of the tribe Eucos-

mini, and structurally have not much in common with Matsumuraeses. They have

the vein M, of the hindwing distinctly approximated at origin to the common

stalk of veins M; and Cu,. The terminal segment of the labial palpus in these two

genera is distinctly longer than in Matsumuraeses, the vein R, of the forewing far

remote from Ry, the veins Ry and R, are stalked, and the males have a costal fold

in the forewing. These characters distinguish Hermenias and Allohermenias from

Matsumuraeses. It may be the right place to remark that most of the species

treated recently as belonging to the genus Hermenias (Diakonoff, 1953) differ

from the type-species of this genus, H. epidola Meyrick, in having distinctly

broader forewings and the hindwings without a dorsal fold including a long hair

pencil 1).

Matsumuraeses phaseoli (Matsumura)

Semasia phaseoli Matsumura, 1900, Ent. Nachr. 26: 197 (July, 1900). — 1901, Illustr.

Zeitschr. Ent. 6: 23. — Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 16.

Semasia elutana Kennel, 1900, Iris 13 : 147, Fig. 23 (August, 1900). — 1916, Pal. Tortr.:

513, PI. 19 Fig. 86.

Eucelis falcana Walsingham, 1900, Ann. Mag. Nat. Hist. [7] 6: 407 (October, 1900). —

Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 16417:

Eucelis ochreocervina Walsingham, 1900, Ann. Mag. Nat. Hist. [7] 6: 407 (October,

1900). — Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 17.

Eucosma trophiodes Meyrick, 1908, Journ. Bombay Nat. Hist. Soc. 18: 613. — Clarke,

1955, Catal. Meyrick’s Types 1: 319.

!) These species have recently been separated in a new genus, Neohermenias Diakonoff.

1966, Zool. Meded. 85: 73. — Ed.

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 33

Argyroploce trophiodes : ?Janse, 1917, Check list S. Afr. Lep. Heter.: 175. — Caradja,

1939, Iris 53: 12.

Eucosma metacritica Meyrick, 1922, Exot. Microlep. 2: 515. — 1935, in Caradja & Mey-

rick, Mater. Fauna chin. Prov.: 55. — Viette, 1952, Bull. Soc. Ent. France 57: 151. —

Clarke, 1955, Catal. Meyrick’s Types 1: 204. Syn. nov.

Thiodia azukivora Matsumura, 1931, 6000 Illustr. Ins. Japan-Empire: 1076, Fig. —

Obraztsov, 1952, Ent. News 63: 148.

Eucosma falcana : Diakonoff, 1949, Bijdr. Dierk. 28 : 136.

Lathronympha phaseoli : Issiki & others, 1950, Iconogr. Ins. Japan, 2nd ed.: 480, Fig. —

Mutuura, 1953, Publ. Ent. Agr. Naniwa Univ. 1: 1, 2, 12, PI. 1 Fig. 1; Pl. 3 Fig. 5; Pl. 4

Fig. 2; PI. 5 Fig. 5. — Inoue, 1954, Check list Lep. Jap.: 95, no. 509. — Yasuda, 1956,

Publ. Ent. Agr. Univ. Osaka 2: 15, 17, Fig. 1—4. — Kuroko, 1957, Enum. Ins. Montis

Hikosan, Lep. 1: 10.

Lathronympha phaeoli {sic}: Mutuura, 1953, Publ. Ent. Agr. Naniwa Univ. 1: 1.

Eucosma melaneura (lapsus): Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 16, 17.

Laspeyresia falcana : Inoue, 1954, Check list Lep. Jap.: 91, no. 481.

Laspeyresia ochreocervina : Inoue, 1954, Check list Lep. Jap.: 91, no. 482.

Semasia azukivora : Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 15, 16.

Eucelis ochreoservina (misprint): Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 16.

Matsumuraeses phaseoli : Issiki & others, 1957, Icon. Heteroc. japon. color. nat. 1: 57,

PI. 8 Fig. 257, 258. — Okano, 1959, Icon. Ins. japon. color. nat. 1: 260, PI. 174 Fig. 20. —

Obraztsov, 1960, Tijdschr. Ent. 103 : 134, Fig. 86—91.

Olethreutes trophiodes : Clarke, 1958, Catal. Meyrick’s Types 3 : 559, Pl. 278 Fig. 1—1a.

Allohermenias metacritica : Razowski, 1960, Polsk. Pismo Ent. 30: 385, Fig. 10.

Matsumuraeses (Semasia) elutana : Kuznetsov, 1962, Trudy Zool. Inst. Akad. Nauk SSSR

30 : 346, Fig. 7.

Types. Types of phaseoli Matsumura and az4kivora Matsumura, not studied.

Lectotype of elutana, female (genitalia on slide figured by KUZNETZOV, 1962),

Amur, in the Zoological Institute of the Academy of Sciences in Leningrad. Holo-

type of falcana Walsingham, female (genitalia on slide, No. 5701), Japan

(PRYER, 1886; No. 70560). Holotype of ochreocervina Walsingham, male

(genitalia on slide, No. 5699), Goorais Valley, 7000 feet, Kashmir, August 1887

(LEECH; No. 60137). Lectotype of trophiodes Meyrick, male (genitalia on slide,

No. 7173, G.F.G.C.), Hakgala, Ceylon, April 1907 (E. E. GREEN). The types of

falcana, ochreocervina and trophiodes are in the British Museum (Natural History).

Lectotype of metacritica Meyrick, male (genitalia on slide, No. 3756; figured by

RAZOWSKI, 1960), Zikawy, China, in the Muséum National d’Histoire Naturelle,

Paris.

Other specimens examined. One male (genitalia on slide made by C. HEINRICH,

October 16, 1923), Japan; one male (genitalia on slide, made by N. OBRAZTSOV,

April 16, 1957), Matsukuri, Japan; both of the above specimens are in the United

States National Museum. One female (genitalia on slide, No. 6668), “aK 5.7”

(BMNH).

The identity of phaseoli and azukivora has been established from the literature

data. In one of his recent papers KUZNETZOV (1962) established the synonymy

of phaseoli and elutana, giving preference to the latter name. In fact, four of the

seven known synonyms of phaseoli were published in the same year, and the name

phaseoli Matsumura is the oldest. As yet, phaseoli is the only Matsumuraeses

species, the female of which is known. Therefore I give a description of the female

genitalia of this species.

34 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Female genitalia. Papillae anales rather small, narrowed rostrally and moderately

dilated caudally; apophyses posteriores shorter than apophyses anteriores. Ostium

bursae located at postsegmental margin of seventh abdominal sternite and preceded

by a well developed ventral plate dilated laterally and deeply excavated caudally;

antrum short, infundibuliform, slightly sclerotized; lamella postvaginalis elongate,

chiefly membranous, dilated and sclerotized caudo-laterally. Ductus bursae narrow,

rather long; cervix bursae very short; corpus bursae ovate, membranous; two signa

shaped as strong, curved thorns.

Matsumuraeses melanaula (Meyrick) comb. nov.

Eucosma melanaula Meyrick, 1916, Exot. Microlep. 2: 17. — Fletcher, 1920, Mem. Dept.

Agr. India, Ent. 6: 49, PI. 1—2. — Clarke, 1955, Catal. Meyrick’s Types 1: 199. — 1958,

op. cit. 3: 372, PI. 185 Fig. 1—la.

Eucosma falcana (part.): Diakonoff, 1949, Bijdr. Dierk. 28 : 136.

Eucosma melaneura (lapsus): Yasuda, 1956, Publ. Ent. Agr. Univ. Osaka 2: 16, 17.

Matsumuraeses (Semasia) elutana (part.): Kuznetzov, 1962, Trudy Zool. Int. Akad. Nauk

SSSR 30 : 346.

Type. Lectotype, male (genitalia on slide No. 7045, J.F.G.C.), Pusa, Bengal,

January 3, 1911 (T. B. FLETCHER); in the British Museum.

This distinct species was erroneously synonymized with falcana Walsingham.

The latter is conspecific with Matsumuraeses phaseoli (Matsumura), and differs

from melanaula in having the cucullus of the valva spatulate, with the external

margin straight. In melanaula the cucullus is rather ovate, rounded externally. On

the photograph of the male genitalia of melanaula, published by CLARKE (1958),

the socii are hardly visible. In fact, they only are a little smaller and narrower

than in phaseoli.

Matsumuraeses hoplista (Meyrick) comb. nov.

Argyroploce hoplista Meyrick, 1927, Exot. Microlep. 3: 340. — Clarke, 1955, Catal.

Meyrick's Types 1: 162.

Endothenia hoplista : Meyrick, 1936, Exot. Microlep. 5 : 24.

Olethreutes hoplista : Clarke, 1958, Catal. Meyrick’s Types 3: 519, Pl. 258 Fig. 4—4a.

Type. Lectotype, male (genitalia on slide No. 7307, J.F.G.C.), Sinabaeng,

Sumatra, 3300 feet, bred July, 1925 (FULMEK) (BMNH).

This species differs from the other of the same genus in having a short uncus

with lateral tufts of long hairs. In falcana a similar structure is slightly indicated

by a little tubercle, located at the posterior margin of the tegumen and covered by

the socii. The socii of hoplista are well developed and obliquely directed towards

the sides, as if they were moved apart by the uncus. On the slide the uncus is

hardly dicernible, but turning of the slide and examination of the genitalia from

the ventral surface of the tegumen make the uncus well visible.

Matsumuraeses monstruosana Kuznetzov

Semasia elutana (part.): Kennel, 1900, Iris 13 : 147, Fig. 22. — 1916, Pal. Tortr.: 513,

PI. 19 Fig. 85.

Matsumuraeses monstruosana Kuznetzov, 1962, Trudy Zool. Inst. Akad. Nauk SSSR 30:

346, Fig. 8.

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae 35

This species is known to me only from the original description and the

published figures of the genitalia. It differs much from the remaining known

species of Matsumuraeses in having the cucullus of the valva with the apex turned

upwards, and the socii represented by two separate groups of setae located on a

common plate. The ostium bursae differs from that in phaseoli, and rather reminds

one of that in some Laspeyresia species. The lamella postvaginalis is dilated

caudally, and is somewhat differently shaped than in phaseoli. In view of the

above it is doubtful that monstruosana belongs to Matsumuraeses, and quite pos-

sible that this species requires description of a new genus.

Parapammene dicastica (Meyrick) comb. nov.

Pammene dicastica Meyrick, 1922, Exot. Microlep. 2: 526. — Clarke, 1955, Catal. Mey-

fick’s Types 1: 115. — 1958, op. cit. 3: 563, PI. 280 Fig. 1—1b.

Holotype. Male (genitalia on slide No. 7353, J.F.G.C.), Murree, Punjab, 7,500

feet, June 1918 (Durr) (BMNH).

Parapammene homotorna (Meyrick)

Pammene homotorna Meyrick, 1912, Journ. Bombay Nat. Hist. Soc. 21: 874. — Clarke,

1955, Catal. Meyrick’s Types 1: 162. — 1958, op. cit. 3 : 563, Pl. 280 Fig. 4, 4a.

Parapammene homotorna : Kuznetzov, 1962, Trudy Zool. Inst. Akad. Nauk SSSR 30 : 349.

Lectotype. Male (genitalia on slide No. 7360, J.F.G.C.), Khasi Hills, Assam,

June 1906 (BMNH).

Parapammene isocampta (Meyrick) comb. nov.

Pammene isocampta Meyrick, 1914, Exot. Microlep. 1: 196. — Clarke, 1955, Catal.

Meyrick’s Types 1: 177. — 1958, op. cit. 3 : 564, Pl. 281 Fig. 1, la.

Holotype. Male (genitalia on slide No. 7347, J.F.G.C.), Paradeniya, Ceylon,

1913 (A. RUTHERFORD) (BMNH).

Parapammene leucodora (Meyrick) comb. nov.

Pammena {sic} leucodora Meyrick, 1928, Exot. Microlep. 3 : 447.

Pammene leucodora: Clarke, 1955, Catal. Meyrick’s Types 1: 185. — 1958, op. cit. 3:

564, Pl. 281 Fig. 2—2b.

Lectotype. Male (genitalia on slide No. 7354, J.F.G.C.), Los Bafios, Philip-

pines, 1927 (BAKER) (BMNH).

Parapammene marmaranthes (Meyrick) comb. nov.

Pammene marmaranthes Meyrick, 1933, Exot. Microlep. 4: 422. — Clarke, 1955, Catal.

Meyrick’s Types 1: 197. — 1958, op. cit. 3: 564, Pl. 281 Fig. 3, 3a.

Lectotype. Female (genitalia on slide No. 7346, J.F.G.C.), Java, January, 1932

(L. G. E. KALSHOVEN) (BMNH). |

36 TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2, 1967

Parapammene oxystaura (Meyrick) comb. nov.

Pammene oxystaura Meyrick in Caradja & Meyrick, 1935, Mater. Faun. chin. Prov.: 62.

— Clarke, 1955, Catal. Meyrick's Types 1: 231. — 1958, op. cit. 3: 564, PI. 281 Fig.

4—4b. — Obraztsov, 1960, Tijdschr. Ent. 103 : 121.

Lectotype. Male (genitalia on slide no. 7345, J.F.G.C.), Tienmushan, China,

5.300 feet, September 1932 (CARADJA). Lectoparatype. Female (genitalia on slide

no. 6667) (BMNH).

Selania extinctana (Chrétien) comb. nov.

Grapholitha extinctana Chrétien, 1915, Ann. Soc. Ent. France 84: 306.

Laspeyresia extinctana: Obraztsov, 1959, Tijdschr. Ent. 102: 186. — Razowski, 1961,

Acta Zool. Cracoviensia 5: 674, PI. 88 Fig. 9.

Type. Holotype, male (genitalia on slide No. 3794), Gafsa, Tunisia, January 2,

1909 (Museum National d’Histoire Naturelle, Paris).

CHRETIEN (1915) described this species as being related to Laspeyresia cap-

paridana (Zeller). The male genitalia, recently figured by RAZOWSKI (1961),

distinctly show that extinctana is a Selania species. Judging from the above-

mentioned description and figure of the male genitalia, extinctana has two big

almost equally long cornuti, and about ten much smaller ones. The same number

of cornuti has been observed in Selania vana (Kennel), but the long cornuti of

this species are not of the same length. Moreover, the cucullus of vana bears two

small thorns, in addition to the anal thorn; this character, unfortunately, does not

show in the photograph of the genitalia, published by OBRAZTSOV (1960, Pl. 13

Ere 3).

TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL. 2 1967 PLAAT 1

Pl. 1. Laspeyresia persicana Osthelder [= Grapholitha (Grapholitha) orobana Treitschke],

4, holotype, above, total aspect; below, genitalia. (Author phot.)

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae

ca e nt tn ann nn sue la a eee en nn m nm nn m nn nn un

PLAAT 2

2 1967

TIJDSCHRIFT VOOR ENTOMOLOGIE, DEEL 110, AFL

Pl. 2, Fig. 1. Pammene blockiana (Herrich-Schäffer), ©, “Treska-Schlucht, W. v. Skopje,

Macedonia, 20—30.V.1956” (F. Kasy) (ZMHB). Fig. 2. Laspeyresia zygogramma Meyrick,

slide no. 6662, Nanking, China, 25.IX.1933 (Höne) (BMNH).

(Author phot.)

male genitalia

N. S. OBRAZTSOV : Die Gattungen der palaearktischen Tortricidae

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In preparation: C. A. W. Jeekel. — Nomenclator familiarum et generum Diplopodorum,

about 300 pp.

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